ALCES 50, 2014 A Journal Devoted to the Biology and Management of Moose CHIEF EDITOR PETER J. PEKINS University of New Hampshire SUBMISSIONS EDITOR ROY V. REA UNIVERSITY OF NORTHERN BRITISH COLUMBIA BUSINESS EDITOR ARTHUR R. RODGERS Ontario Ministry of Natural Resources ASSOCIATE EDITORS EDWARD M. ADDISON Ecolink Science VINCE F. J. CRICHTON Manitoba Conservation MURRAY W. LANKESTER Lakehead University (retired) BRIAN E. MCLAREN Lakehead University RON MOEN University of Minnesota GERALD W. REDMOND Maritime College of Forest Technology KRISTINE M. RINES New Hampshire Fish and Game Printed at Lakehead University Thunder Bay, Ontario, Canada (called Proceedings of the North American Moose Conference from 1972 through 1980) ISSN 0835-5851