id	sid	tid	token	lemma	pos
alces-23	1	1	alces	alce	NOUN
alces-23	1	2	vol	vol	NOUN
alces-23	1	3	.	.	PROPN
alces-23	2	1	45	45	NUM
alces-23	2	2	,	,	PUNCT
alces-23	2	3	2009	2009	NUM
alces-23	2	4	hundertmark	hundertmark	ADJ
alces-23	2	5	genetic	genetic	ADJ
alces-23	2	6	diversity	diversity	NOUN
alces-23	2	7	in	in	ADP
alces-23	2	8	introduced	introduce	VERB
alces-23	2	9	populations	population	NOUN
alces-23	2	10	137	137	NUM
alces-23	2	11	reduced	reduce	VERB
alces-23	2	12	genetic	genetic	ADJ
alces-23	2	13	diversity	diversity	NOUN
alces-23	2	14	in	in	ADP
alces-23	2	15	two	two	NUM
alces-23	2	16	introduced	introduce	VERB
alces-23	2	17	and	and	CCONJ
alces-23	2	18	isolated	isolated	ADJ
alces-23	2	19	moose	moose	NOUN
alces-23	2	20	populations	population	NOUN
alces-23	2	21	in	in	ADP
alces-23	2	22	alaska	alaska	PROPN
alces-23	2	23	kris	kris	PROPN
alces-23	2	24	j.	j.	PROPN
alces-23	2	25	hundertmark	hundertmark	PROPN
alces-23	2	26	institute	institute	PROPN
alces-23	2	27	of	of	ADP
alces-23	2	28	arctic	arctic	PROPN
alces-23	2	29	biology	biology	NOUN
alces-23	2	30	and	and	CCONJ
alces-23	2	31	department	department	NOUN
alces-23	2	32	of	of	ADP
alces-23	2	33	biology	biology	NOUN
alces-23	2	34	and	and	CCONJ
alces-23	2	35	wildlife	wildlife	NOUN
alces-23	2	36	,	,	PUNCT
alces-23	2	37	university	university	NOUN
alces-23	2	38	of	of	ADP
alces-23	2	39	alaska	alaska	PROPN
alces-23	2	40	fairbanks	fairbanks	PROPN
alces-23	2	41	,	,	PUNCT
alces-23	2	42	po	po	PROPN
alces-23	2	43	box	box	PROPN
alces-23	2	44	757000	757000	NUM
alces-23	2	45	,	,	PUNCT
alces-23	2	46	fairbanks	fairbank	NOUN
alces-23	2	47	,	,	PUNCT
alces-23	2	48	alaska	alaska	PROPN
alces-23	2	49	99775	99775	NUM
alces-23	2	50	,	,	PUNCT
alces-23	2	51	usa	usa	PROPN
alces-23	2	52	.	.	PROPN
alces-23	2	53	kris.hundertmark@alaska.edu	kris.hundertmark@alaska.edu	PROPN
alces-23	2	54	abstract	abstract	NOUN
alces-23	2	55	:	:	PUNCT
alces-23	3	1	i	i	PRON
alces-23	3	2	examined	examine	VERB
alces-23	3	3	indices	index	NOUN
alces-23	3	4	of	of	ADP
alces-23	3	5	genetic	genetic	ADJ
alces-23	3	6	diversity	diversity	NOUN
alces-23	3	7	in	in	ADP
alces-23	3	8	2	2	NUM
alces-23	3	9	isolated	isolated	ADJ
alces-23	3	10	moose	moose	NOUN
alces-23	3	11	(	(	PUNCT
alces-23	3	12	alces	alce	NOUN
alces-23	3	13	alces	alce	NOUN
alces-23	3	14	)	)	PUNCT
alces-23	3	15	populations	population	NOUN
alces-23	3	16	in	in	ADP
alces-23	3	17	alaska	alaska	PROPN
alces-23	3	18	that	that	PRON
alces-23	3	19	were	be	AUX
alces-23	3	20	founded	found	VERB
alces-23	3	21	by	by	ADP
alces-23	3	22	low	low	ADJ
alces-23	3	23	numbers	number	NOUN
alces-23	3	24	of	of	ADP
alces-23	3	25	individuals	individual	NOUN
alces-23	3	26	to	to	PART
alces-23	3	27	determine	determine	VERB
alces-23	3	28	effects	effect	NOUN
alces-23	3	29	of	of	ADP
alces-23	3	30	founding	founding	NOUN
alces-23	3	31	and	and	CCONJ
alces-23	3	32	infer	infer	VERB
alces-23	3	33	whether	whether	SCONJ
alces-23	3	34	subsequent	subsequent	ADJ
alces-23	3	35	gene	gene	NOUN
alces-23	3	36	flow	flow	NOUN
alces-23	3	37	has	have	AUX
alces-23	3	38	occurred	occur	VERB
alces-23	3	39	with	with	ADP
alces-23	3	40	surrounding	surround	VERB
alces-23	3	41	moose	moose	NOUN
alces-23	3	42	populations	population	NOUN
alces-23	3	43	.	.	PUNCT
alces-23	4	1	kalgin	kalgin	PROPN
alces-23	4	2	island	island	PROPN
alces-23	4	3	is	be	AUX
alces-23	4	4	a	a	DET
alces-23	4	5	small	small	ADJ
alces-23	4	6	,	,	PUNCT
alces-23	4	7	predator	predator	NOUN
alces-23	4	8	-	-	PUNCT
alces-23	4	9	free	free	ADJ
alces-23	4	10	island	island	NOUN
alces-23	4	11	in	in	ADP
alces-23	4	12	cook	cook	NOUN
alces-23	4	13	inlet	inlet	NOUN
alces-23	4	14	that	that	PRON
alces-23	4	15	was	be	AUX
alces-23	4	16	founded	found	VERB
alces-23	4	17	by	by	ADP
alces-23	4	18	6	6	NUM
alces-23	4	19	moose	moose	NOUN
alces-23	4	20	(	(	PUNCT
alces-23	4	21	3	3	NUM
alces-23	4	22	females	female	NOUN
alces-23	4	23	)	)	PUNCT
alces-23	4	24	in	in	ADP
alces-23	4	25	the	the	DET
alces-23	4	26	late	late	ADJ
alces-23	4	27	1950s	1950s	NUM
alces-23	4	28	;	;	PUNCT
alces-23	4	29	its	its	PRON
alces-23	4	30	population	population	NOUN
alces-23	4	31	has	have	AUX
alces-23	4	32	since	since	SCONJ
alces-23	4	33	undergone	undergo	VERB
alces-23	4	34	dramatic	dramatic	ADJ
alces-23	4	35	fluctuations	fluctuation	NOUN
alces-23	4	36	.	.	PUNCT
alces-23	5	1	berners	berner	NOUN
alces-23	5	2	bay	bay	PROPN
alces-23	5	3	is	be	AUX
alces-23	5	4	an	an	DET
alces-23	5	5	isolated	isolated	ADJ
alces-23	5	6	population	population	NOUN
alces-23	5	7	along	along	ADP
alces-23	5	8	the	the	DET
alces-23	5	9	coast	coast	NOUN
alces-23	5	10	of	of	ADP
alces-23	5	11	southeastern	southeastern	ADJ
alces-23	5	12	alaska	alaska	PROPN
alces-23	5	13	that	that	PRON
alces-23	5	14	was	be	AUX
alces-23	5	15	founded	found	VERB
alces-23	5	16	by	by	ADP
alces-23	5	17	21	21	NUM
alces-23	5	18	calves	calf	NOUN
alces-23	5	19	introduced	introduce	VERB
alces-23	5	20	in	in	ADP
alces-23	5	21	1958	1958	NUM
alces-23	5	22	-	-	SYM
alces-23	5	23	1960	1960	NUM
alces-23	5	24	.	.	PUNCT
alces-23	6	1	genetic	genetic	ADJ
alces-23	6	2	attributes	attribute	NOUN
alces-23	6	3	of	of	ADP
alces-23	6	4	those	those	DET
alces-23	6	5	populations	population	NOUN
alces-23	6	6	were	be	AUX
alces-23	6	7	compared	compare	VERB
alces-23	6	8	to	to	ADP
alces-23	6	9	a	a	DET
alces-23	6	10	population	population	NOUN
alces-23	6	11	in	in	ADP
alces-23	6	12	yukon	yukon	PROPN
alces-23	6	13	flats	flat	NOUN
alces-23	6	14	in	in	ADP
alces-23	6	15	central	central	ADJ
alces-23	6	16	alaska	alaska	NOUN
alces-23	6	17	that	that	PRON
alces-23	6	18	served	serve	VERB
alces-23	6	19	as	as	ADP
alces-23	6	20	an	an	DET
alces-23	6	21	outbred	outbred	ADJ
alces-23	6	22	control	control	NOUN
alces-23	6	23	.	.	PUNCT
alces-23	7	1	indices	index	NOUN
alces-23	7	2	from	from	ADP
alces-23	7	3	11	11	NUM
alces-23	7	4	microsatellite	microsatellite	NOUN
alces-23	7	5	markers	marker	NOUN
alces-23	7	6	indicated	indicate	VERB
alces-23	7	7	substantial	substantial	ADJ
alces-23	7	8	effects	effect	NOUN
alces-23	7	9	of	of	ADP
alces-23	7	10	founding	founding	NOUN
alces-23	7	11	and	and	CCONJ
alces-23	7	12	subsequent	subsequent	ADJ
alces-23	7	13	isolation	isolation	NOUN
alces-23	7	14	.	.	PUNCT
alces-23	8	1	heterozygosity	heterozygosity	NOUN
alces-23	8	2	and	and	CCONJ
alces-23	8	3	allelic	allelic	ADJ
alces-23	8	4	diversity	diversity	NOUN
alces-23	8	5	,	,	PUNCT
alces-23	8	6	both	both	PRON
alces-23	8	7	of	of	ADP
alces-23	8	8	which	which	PRON
alces-23	8	9	are	be	AUX
alces-23	8	10	reduced	reduce	VERB
alces-23	8	11	by	by	ADP
alces-23	8	12	genetic	genetic	ADJ
alces-23	8	13	bottlenecks	bottleneck	NOUN
alces-23	8	14	,	,	PUNCT
alces-23	8	15	were	be	AUX
alces-23	8	16	significantly	significantly	ADV
alces-23	8	17	lower	low	ADJ
alces-23	8	18	in	in	ADP
alces-23	8	19	the	the	DET
alces-23	8	20	introduced	introduce	VERB
alces-23	8	21	populations	population	NOUN
alces-23	8	22	than	than	ADP
alces-23	8	23	the	the	DET
alces-23	8	24	yukon	yukon	PROPN
alces-23	8	25	flats	flat	VERB
alces-23	8	26	population	population	NOUN
alces-23	8	27	.	.	PUNCT
alces-23	9	1	kalgin	kalgin	PROPN
alces-23	9	2	island	island	NOUN
alces-23	9	3	diversity	diversity	NOUN
alces-23	9	4	was	be	AUX
alces-23	9	5	significantly	significantly	ADV
alces-23	9	6	lower	low	ADJ
alces-23	9	7	than	than	ADP
alces-23	9	8	that	that	PRON
alces-23	9	9	for	for	ADP
alces-23	9	10	berners	berner	NOUN
alces-23	9	11	bay	bay	NOUN
alces-23	9	12	,	,	PUNCT
alces-23	9	13	and	and	CCONJ
alces-23	9	14	was	be	AUX
alces-23	9	15	likely	likely	ADJ
alces-23	9	16	due	due	ADP
alces-23	9	17	to	to	ADP
alces-23	9	18	the	the	DET
alces-23	9	19	smaller	small	ADJ
alces-23	9	20	founding	founding	NOUN
alces-23	9	21	size	size	NOUN
alces-23	9	22	and	and	CCONJ
alces-23	9	23	subsequent	subsequent	ADJ
alces-23	9	24	population	population	NOUN
alces-23	9	25	fluctuations	fluctuation	NOUN
alces-23	9	26	.	.	PUNCT
alces-23	10	1	neither	neither	CCONJ
alces-23	10	2	introduced	introduce	VERB
alces-23	10	3	population	population	NOUN
alces-23	10	4	exhibited	exhibit	VERB
alces-23	10	5	evidence	evidence	NOUN
alces-23	10	6	of	of	ADP
alces-23	10	7	gene	gene	NOUN
alces-23	10	8	flow	flow	NOUN
alces-23	10	9	from	from	ADP
alces-23	10	10	surrounding	surround	VERB
alces-23	10	11	populations	population	NOUN
alces-23	10	12	.	.	PUNCT
alces-23	11	1	managers	manager	NOUN
alces-23	11	2	should	should	AUX
alces-23	11	3	consider	consider	VERB
alces-23	11	4	the	the	DET
alces-23	11	5	isolation	isolation	NOUN
alces-23	11	6	of	of	ADP
alces-23	11	7	those	those	DET
alces-23	11	8	populations	population	NOUN
alces-23	11	9	when	when	SCONJ
alces-23	11	10	assessing	assess	VERB
alces-23	11	11	risks	risk	NOUN
alces-23	11	12	to	to	ADP
alces-23	11	13	population	population	NOUN
alces-23	11	14	viability	viability	NOUN
alces-23	11	15	and	and	CCONJ
alces-23	11	16	crafting	craft	VERB
alces-23	11	17	management	management	NOUN
alces-23	11	18	strategies	strategy	NOUN
alces-23	11	19	.	.	PUNCT
alces-23	12	1	alces	alce	NOUN
alces-23	12	2	vol	vol	NOUN
alces-23	12	3	.	.	PUNCT
alces-23	13	1	45	45	NUM
alces-23	13	2	:	:	SYM
alces-23	13	3	137	137	NUM
alces-23	13	4	-	-	SYM
alces-23	13	5	142	142	NUM
alces-23	13	6	(	(	PUNCT
alces-23	13	7	2009	2009	NUM
alces-23	13	8	)	)	PUNCT
alces-23	13	9	key	key	ADJ
alces-23	13	10	words	word	NOUN
alces-23	13	11	:	:	PUNCT
alces-23	13	12	alaska	alaska	PROPN
alces-23	13	13	,	,	PUNCT
alces-23	13	14	alces	alce	NOUN
alces-23	13	15	alces	alce	NOUN
alces-23	13	16	,	,	PUNCT
alces-23	13	17	bottleneck	bottleneck	NOUN
alces-23	13	18	,	,	PUNCT
alces-23	13	19	gene	gene	NOUN
alces-23	13	20	flow	flow	NOUN
alces-23	13	21	,	,	PUNCT
alces-23	13	22	insular	insular	ADJ
alces-23	13	23	,	,	PUNCT
alces-23	13	24	introduced	introduce	VERB
alces-23	13	25	population	population	NOUN
alces-23	13	26	,	,	PUNCT
alces-23	13	27	moose	moose	NOUN
alces-23	13	28	.	.	PUNCT
alces-23	14	1	one	one	NUM
alces-23	14	2	of	of	ADP
alces-23	14	3	the	the	DET
alces-23	14	4	primary	primary	ADJ
alces-23	14	5	concerns	concern	NOUN
alces-23	14	6	of	of	ADP
alces-23	14	7	conservation	conservation	NOUN
alces-23	14	8	biology	biology	NOUN
alces-23	14	9	is	be	AUX
alces-23	14	10	the	the	DET
alces-23	14	11	loss	loss	NOUN
alces-23	14	12	of	of	ADP
alces-23	14	13	genetic	genetic	ADJ
alces-23	14	14	diversity	diversity	NOUN
alces-23	14	15	through	through	ADP
alces-23	14	16	genetic	genetic	ADJ
alces-23	14	17	drift	drift	NOUN
alces-23	14	18	in	in	ADP
alces-23	14	19	small	small	ADJ
alces-23	14	20	populations	population	NOUN
alces-23	14	21	.	.	PUNCT
alces-23	15	1	in	in	ADP
alces-23	15	2	cases	case	NOUN
alces-23	15	3	where	where	SCONJ
alces-23	15	4	populations	population	NOUN
alces-23	15	5	are	be	AUX
alces-23	15	6	isolated	isolate	VERB
alces-23	15	7	,	,	PUNCT
alces-23	15	8	thus	thus	ADV
alces-23	15	9	preventing	prevent	VERB
alces-23	15	10	immigration	immigration	NOUN
alces-23	15	11	from	from	ADP
alces-23	15	12	neighboring	neighboring	NOUN
alces-23	15	13	populations	population	NOUN
alces-23	15	14	,	,	PUNCT
alces-23	15	15	genetic	genetic	ADJ
alces-23	15	16	drift	drift	NOUN
alces-23	15	17	occurs	occur	VERB
alces-23	15	18	at	at	ADP
alces-23	15	19	a	a	DET
alces-23	15	20	maximum	maximum	ADJ
alces-23	15	21	rate	rate	NOUN
alces-23	15	22	depending	depend	VERB
alces-23	15	23	on	on	ADP
alces-23	15	24	population	population	NOUN
alces-23	15	25	size	size	NOUN
alces-23	15	26	.	.	PUNCT
alces-23	16	1	loss	loss	NOUN
alces-23	16	2	of	of	ADP
alces-23	16	3	diversity	diversity	NOUN
alces-23	16	4	from	from	ADP
alces-23	16	5	drift	drift	NOUN
alces-23	16	6	is	be	AUX
alces-23	16	7	compounded	compound	VERB
alces-23	16	8	in	in	ADP
alces-23	16	9	populations	population	NOUN
alces-23	16	10	that	that	PRON
alces-23	16	11	are	be	AUX
alces-23	16	12	founded	found	VERB
alces-23	16	13	by	by	ADP
alces-23	16	14	low	low	ADJ
alces-23	16	15	numbers	number	NOUN
alces-23	16	16	of	of	ADP
alces-23	16	17	individuals	individual	NOUN
alces-23	16	18	due	due	ADJ
alces-23	16	19	to	to	ADP
alces-23	16	20	demographic	demographic	ADJ
alces-23	16	21	and	and	CCONJ
alces-23	16	22	genetic	genetic	ADJ
alces-23	16	23	bottlenecks	bottleneck	NOUN
alces-23	16	24	.	.	PUNCT
alces-23	17	1	loss	loss	NOUN
alces-23	17	2	of	of	ADP
alces-23	17	3	genetic	genetic	ADJ
alces-23	17	4	diversity	diversity	NOUN
alces-23	17	5	has	have	AUX
alces-23	17	6	been	be	AUX
alces-23	17	7	related	relate	VERB
alces-23	17	8	to	to	ADP
alces-23	17	9	loss	loss	NOUN
alces-23	17	10	of	of	ADP
alces-23	17	11	fitness	fitness	NOUN
alces-23	17	12	(	(	PUNCT
alces-23	17	13	reed	reed	NOUN
alces-23	17	14	and	and	CCONJ
alces-23	17	15	frankham	frankham	PROPN
alces-23	17	16	2003	2003	NUM
alces-23	17	17	)	)	PUNCT
alces-23	17	18	.	.	PUNCT
alces-23	18	1	in	in	ADP
alces-23	18	2	ungulates	ungulate	NOUN
alces-23	18	3	,	,	PUNCT
alces-23	18	4	studies	study	NOUN
alces-23	18	5	have	have	AUX
alces-23	18	6	found	find	VERB
alces-23	18	7	correlations	correlation	NOUN
alces-23	18	8	between	between	ADP
alces-23	18	9	indices	index	NOUN
alces-23	18	10	of	of	ADP
alces-23	18	11	diversity	diversity	NOUN
alces-23	18	12	and	and	CCONJ
alces-23	18	13	reduction	reduction	NOUN
alces-23	18	14	in	in	ADP
alces-23	18	15	juvenile	juvenile	ADJ
alces-23	18	16	survival	survival	NOUN
alces-23	18	17	(	(	PUNCT
alces-23	18	18	coulson	coulson	PROPN
alces-23	18	19	et	et	PROPN
alces-23	18	20	al	al	PROPN
alces-23	18	21	.	.	PROPN
alces-23	18	22	1999	1999	NUM
alces-23	18	23	,	,	PUNCT
alces-23	18	24	mainguy	mainguy	NOUN
alces-23	18	25	et	et	PROPN
alces-23	18	26	al	al	PROPN
alces-23	18	27	.	.	PROPN
alces-23	18	28	2009	2009	NUM
alces-23	18	29	,	,	PUNCT
alces-23	18	30	silva	silva	PROPN
alces-23	18	31	et	et	PROPN
alces-23	18	32	al	al	PROPN
alces-23	18	33	.	.	PROPN
alces-23	18	34	2009	2009	NUM
alces-23	18	35	)	)	PUNCT
alces-23	18	36	,	,	PUNCT
alces-23	18	37	variation	variation	NOUN
alces-23	18	38	in	in	ADP
alces-23	18	39	horn	horn	NOUN
alces-23	18	40	/	/	SYM
alces-23	18	41	antler	antler	NOUN
alces-23	18	42	growth	growth	NOUN
alces-23	18	43	(	(	PUNCT
alces-23	18	44	scribner	scribner	PROPN
alces-23	18	45	and	and	CCONJ
alces-23	18	46	smith	smith	PROPN
alces-23	18	47	1990	1990	NUM
alces-23	18	48	,	,	PUNCT
alces-23	18	49	von	von	PROPN
alces-23	18	50	hardenberg	hardenberg	PROPN
alces-23	18	51	et	et	PROPN
alces-23	18	52	al	al	PROPN
alces-23	18	53	.	.	PROPN
alces-23	18	54	2007	2007	NUM
alces-23	18	55	)	)	PUNCT
alces-23	18	56	,	,	PUNCT
alces-23	18	57	and	and	CCONJ
alces-23	18	58	parasite	parasite	ADJ
alces-23	18	59	resistance	resistance	NOUN
alces-23	18	60	(	(	PUNCT
alces-23	18	61	coltman	coltman	NOUN
alces-23	18	62	et	et	PROPN
alces-23	18	63	al	al	PROPN
alces-23	18	64	.	.	PROPN
alces-23	18	65	1999	1999	NUM
alces-23	18	66	)	)	PUNCT
alces-23	18	67	.	.	PUNCT
alces-23	19	1	thus	thus	ADV
alces-23	19	2	,	,	PUNCT
alces-23	19	3	genetic	genetic	ADJ
alces-23	19	4	diversity	diversity	NOUN
alces-23	19	5	of	of	ADP
alces-23	19	6	populations	population	NOUN
alces-23	19	7	should	should	AUX
alces-23	19	8	be	be	AUX
alces-23	19	9	a	a	DET
alces-23	19	10	primary	primary	ADJ
alces-23	19	11	concern	concern	NOUN
alces-23	19	12	in	in	ADP
alces-23	19	13	ungulate	ungulate	ADJ
alces-23	19	14	management	management	NOUN
alces-23	19	15	,	,	PUNCT
alces-23	19	16	particularly	particularly	ADV
alces-23	19	17	with	with	ADP
alces-23	19	18	small	small	ADJ
alces-23	19	19	and	and	CCONJ
alces-23	19	20	isolated	isolated	ADJ
alces-23	19	21	populations	population	NOUN
alces-23	19	22	.	.	PUNCT
alces-23	20	1	valuable	valuable	ADJ
alces-23	20	2	insight	insight	NOUN
alces-23	20	3	may	may	AUX
alces-23	20	4	be	be	AUX
alces-23	20	5	gained	gain	VERB
alces-23	20	6	from	from	ADP
alces-23	20	7	studying	study	VERB
alces-23	20	8	wild	wild	ADJ
alces-23	20	9	populations	population	NOUN
alces-23	20	10	with	with	ADP
alces-23	20	11	known	know	VERB
alces-23	20	12	demographic	demographic	ADJ
alces-23	20	13	histories	history	NOUN
alces-23	20	14	to	to	PART
alces-23	20	15	determine	determine	VERB
alces-23	20	16	effects	effect	NOUN
alces-23	20	17	of	of	ADP
alces-23	20	18	population	population	NOUN
alces-23	20	19	size	size	NOUN
alces-23	20	20	on	on	ADP
alces-23	20	21	genetic	genetic	ADJ
alces-23	20	22	diversity	diversity	NOUN
alces-23	20	23	.	.	PUNCT
alces-23	21	1	introduced	introduce	VERB
alces-23	21	2	populations	population	NOUN
alces-23	21	3	often	often	ADV
alces-23	21	4	act	act	VERB
alces-23	21	5	as	as	ADP
alces-23	21	6	natural	natural	ADJ
alces-23	21	7	experiments	experiment	NOUN
alces-23	21	8	in	in	ADP
alces-23	21	9	that	that	DET
alces-23	21	10	regard	regard	NOUN
alces-23	21	11	,	,	PUNCT
alces-23	21	12	particularly	particularly	ADV
alces-23	21	13	when	when	SCONJ
alces-23	21	14	founding	found	VERB
alces-23	21	15	population	population	NOUN
alces-23	21	16	size	size	NOUN
alces-23	21	17	is	be	AUX
alces-23	21	18	known	know	VERB
alces-23	21	19	,	,	PUNCT
alces-23	21	20	as	as	ADV
alces-23	21	21	well	well	ADV
alces-23	21	22	as	as	ADP
alces-23	21	23	demographic	demographic	ADJ
alces-23	21	24	trends	trend	NOUN
alces-23	21	25	since	since	SCONJ
alces-23	21	26	founding	found	VERB
alces-23	21	27	.	.	PUNCT
alces-23	22	1	i	i	PRON
alces-23	22	2	compared	compare	VERB
alces-23	22	3	2	2	NUM
alces-23	22	4	small	small	ADJ
alces-23	22	5	,	,	PUNCT
alces-23	22	6	isolated	isolate	VERB
alces-23	22	7	,	,	PUNCT
alces-23	22	8	introduced	introduce	VERB
alces-23	22	9	populations	population	NOUN
alces-23	22	10	of	of	ADP
alces-23	22	11	moose	moose	NOUN
alces-23	22	12	(	(	PUNCT
alces-23	22	13	alces	alce	NOUN
alces-23	22	14	alces	alce	NOUN
alces-23	22	15	)	)	PUNCT
alces-23	22	16	in	in	ADP
alces-23	22	17	alaska	alaska	PROPN
alces-23	22	18	to	to	PART
alces-23	22	19	determine	determine	VERB
alces-23	22	20	the	the	DET
alces-23	22	21	effect	effect	NOUN
alces-23	22	22	of	of	ADP
alces-23	22	23	their	their	PRON
alces-23	22	24	respective	respective	ADJ
alces-23	22	25	demographic	demographic	ADJ
alces-23	22	26	histories	history	NOUN
alces-23	22	27	on	on	ADP
alces-23	22	28	indices	index	NOUN
alces-23	22	29	of	of	ADP
alces-23	22	30	genetic	genetic	ADJ
alces-23	22	31	diversity	diversity	NOUN
alces-23	22	32	,	,	PUNCT
alces-23	22	33	and	and	CCONJ
alces-23	22	34	to	to	PART
alces-23	22	35	infer	infer	VERB
alces-23	22	36	the	the	DET
alces-23	22	37	degree	degree	NOUN
alces-23	22	38	to	to	PART
alces-23	22	39	which	which	PRON
alces-23	22	40	gene	gene	NOUN
alces-23	22	41	flow	flow	NOUN
alces-23	22	42	has	have	AUX
alces-23	22	43	affected	affect	VERB
alces-23	22	44	diversity	diversity	NOUN
alces-23	22	45	.	.	PUNCT
alces-23	23	1	i	i	PRON
alces-23	23	2	also	also	ADV
alces-23	23	3	compared	compare	VERB
alces-23	23	4	those	those	DET
alces-23	23	5	populations	population	NOUN
alces-23	23	6	to	to	ADP
alces-23	23	7	an	an	DET
alces-23	23	8	outbred	outbred	ADJ
alces-23	23	9	moose	moose	NOUN
alces-23	23	10	population	population	NOUN
alces-23	23	11	to	to	PART
alces-23	23	12	demonstrate	demonstrate	VERB
alces-23	23	13	the	the	DET
alces-23	23	14	extent	extent	NOUN
alces-23	23	15	to	to	PART
alces-23	23	16	which	which	PRON
alces-23	23	17	the	the	DET
alces-23	23	18	introduced	introduce	VERB
alces-23	23	19	populations	population	NOUN
alces-23	23	20	have	have	AUX
alces-23	23	21	lost	lose	VERB
alces-23	23	22	diversity	diversity	NOUN
alces-23	23	23	.	.	PUNCT
alces-23	24	1	genetic	genetic	ADJ
alces-23	24	2	diversity	diversity	NOUN
alces-23	24	3	in	in	ADP
alces-23	24	4	introduced	introduce	VERB
alces-23	24	5	populations	population	NOUN
alces-23	24	6	hundertmark	hundertmark	NOUN
alces-23	24	7	alces	alce	NOUN
alces-23	24	8	vol	vol	NOUN
alces-23	24	9	.	.	PROPN
alces-23	25	1	45	45	NUM
alces-23	25	2	,	,	PUNCT
alces-23	25	3	2009	2009	NUM
alces-23	25	4	138	138	NUM
alces-23	25	5	study	study	NOUN
alces-23	25	6	area	area	NOUN
alces-23	25	7	moose	moose	NOUN
alces-23	25	8	populations	population	NOUN
alces-23	25	9	in	in	ADP
alces-23	25	10	kalgin	kalgin	NOUN
alces-23	25	11	island	island	NOUN
alces-23	25	12	and	and	CCONJ
alces-23	25	13	berners	berners	PROPN
alces-23	25	14	bay	bay	PROPN
alces-23	25	15	,	,	PUNCT
alces-23	25	16	alaska	alaska	PROPN
alces-23	25	17	have	have	VERB
alces-23	25	18	similar	similar	ADJ
alces-23	25	19	histories	history	NOUN
alces-23	25	20	.	.	PUNCT
alces-23	26	1	they	they	PRON
alces-23	26	2	both	both	PRON
alces-23	26	3	were	be	AUX
alces-23	26	4	established	establish	VERB
alces-23	26	5	through	through	ADP
alces-23	26	6	introduction	introduction	NOUN
alces-23	26	7	of	of	ADP
alces-23	26	8	individuals	individual	NOUN
alces-23	26	9	from	from	ADP
alces-23	26	10	southcentral	southcentral	ADJ
alces-23	26	11	alaska	alaska	PROPN
alces-23	26	12	in	in	ADP
alces-23	26	13	the	the	DET
alces-23	26	14	late	late	ADJ
alces-23	26	15	1950s	1950s	NUM
alces-23	26	16	.	.	PUNCT
alces-23	27	1	the	the	DET
alces-23	27	2	population	population	NOUN
alces-23	27	3	on	on	ADP
alces-23	27	4	kalgin	kalgin	PROPN
alces-23	27	5	island	island	PROPN
alces-23	27	6	(	(	PUNCT
alces-23	27	7	60º	60º	PROPN
alces-23	27	8	27’n	27’n	PROPN
alces-23	27	9	,	,	PUNCT
alces-23	27	10	152º	152º	PROPN
alces-23	27	11	00’w	00’w	NUM
alces-23	27	12	)	)	PUNCT
alces-23	27	13	was	be	AUX
alces-23	27	14	founded	found	VERB
alces-23	27	15	by	by	ADP
alces-23	27	16	6	6	NUM
alces-23	27	17	moose	moose	NOUN
alces-23	27	18	(	(	PUNCT
alces-23	27	19	3	3	NUM
alces-23	27	20	females	female	NOUN
alces-23	27	21	)	)	PUNCT
alces-23	27	22	transported	transport	VERB
alces-23	27	23	to	to	ADP
alces-23	27	24	the	the	DET
alces-23	27	25	island	island	NOUN
alces-23	27	26	in	in	ADP
alces-23	27	27	1957	1957	NUM
alces-23	27	28	-	-	SYM
alces-23	27	29	1959	1959	NUM
alces-23	27	30	(	(	PUNCT
alces-23	27	31	burris	burris	PROPN
alces-23	27	32	and	and	CCONJ
alces-23	27	33	mcknight	mcknight	NOUN
alces-23	27	34	1973	1973	NUM
alces-23	27	35	)	)	PUNCT
alces-23	27	36	.	.	PUNCT
alces-23	28	1	the	the	DET
alces-23	28	2	population	population	NOUN
alces-23	28	3	in	in	ADP
alces-23	28	4	berners	berner	NOUN
alces-23	28	5	bay	bay	PROPN
alces-23	28	6	(	(	PUNCT
alces-23	28	7	58º	58º	PROPN
alces-23	28	8	45’n	45’n	PROPN
alces-23	28	9	,	,	PUNCT
alces-23	28	10	134º	134º	PROPN
alces-23	28	11	50’w	50’w	PROPN
alces-23	28	12	)	)	PUNCT
alces-23	28	13	was	be	AUX
alces-23	28	14	founded	found	VERB
alces-23	28	15	by	by	ADP
alces-23	28	16	15	15	NUM
alces-23	28	17	calves	calf	NOUN
alces-23	28	18	in	in	ADP
alces-23	28	19	1958	1958	NUM
alces-23	28	20	and	and	CCONJ
alces-23	28	21	6	6	NUM
alces-23	28	22	additional	additional	ADJ
alces-23	28	23	calves	calf	NOUN
alces-23	28	24	in	in	ADP
alces-23	28	25	1960	1960	NUM
alces-23	28	26	(	(	PUNCT
alces-23	28	27	burris	burris	PROPN
alces-23	28	28	and	and	CCONJ
alces-23	28	29	mcknight	mcknight	NOUN
alces-23	28	30	1973	1973	NUM
alces-23	28	31	)	)	PUNCT
alces-23	28	32	.	.	PUNCT
alces-23	29	1	moreover	moreover	ADV
alces-23	29	2	,	,	PUNCT
alces-23	29	3	both	both	DET
alces-23	29	4	populations	population	NOUN
alces-23	29	5	are	be	AUX
alces-23	29	6	seemingly	seemingly	ADV
alces-23	29	7	isolated	isolate	VERB
alces-23	29	8	from	from	ADP
alces-23	29	9	neighboring	neighboring	NOUN
alces-23	29	10	moose	moose	NOUN
alces-23	29	11	populations	population	NOUN
alces-23	29	12	.	.	PUNCT
alces-23	30	1	kalgin	kalgin	PROPN
alces-23	30	2	island	island	NOUN
alces-23	30	3	is	be	AUX
alces-23	30	4	located	locate	VERB
alces-23	30	5	in	in	ADP
alces-23	30	6	cook	cook	NOUN
alces-23	30	7	inlet	inlet	NOUN
alces-23	30	8	which	which	PRON
alces-23	30	9	is	be	AUX
alces-23	30	10	characterized	characterize	VERB
alces-23	30	11	by	by	ADP
alces-23	30	12	strong	strong	ADJ
alces-23	30	13	tidal	tidal	ADJ
alces-23	30	14	currents	current	NOUN
alces-23	30	15	that	that	PRON
alces-23	30	16	have	have	AUX
alces-23	30	17	kept	keep	VERB
alces-23	30	18	large	large	ADJ
alces-23	30	19	mammals	mammal	NOUN
alces-23	30	20	,	,	PUNCT
alces-23	30	21	including	include	VERB
alces-23	30	22	predators	predator	NOUN
alces-23	30	23	,	,	PUNCT
alces-23	30	24	from	from	ADP
alces-23	30	25	colonizing	colonize	VERB
alces-23	30	26	the	the	DET
alces-23	30	27	island	island	NOUN
alces-23	30	28	.	.	PUNCT
alces-23	31	1	nonetheless	nonetheless	ADV
alces-23	31	2	,	,	PUNCT
alces-23	31	3	the	the	DET
alces-23	31	4	short	short	ADJ
alces-23	31	5	distance	distance	NOUN
alces-23	31	6	from	from	ADP
alces-23	31	7	mainland	mainland	NOUN
alces-23	31	8	to	to	ADP
alces-23	31	9	island	island	NOUN
alces-23	31	10	(	(	PUNCT
alces-23	31	11	<	<	X
alces-23	31	12	10	10	NUM
alces-23	31	13	km	km	NOUN
alces-23	31	14	)	)	PUNCT
alces-23	31	15	has	have	AUX
alces-23	31	16	fueled	fuel	VERB
alces-23	31	17	speculation	speculation	NOUN
alces-23	31	18	that	that	SCONJ
alces-23	31	19	gene	gene	NOUN
alces-23	31	20	flow	flow	NOUN
alces-23	31	21	is	be	AUX
alces-23	31	22	possible	possible	ADJ
alces-23	31	23	.	.	PUNCT
alces-23	32	1	berners	berner	NOUN
alces-23	32	2	bay	bay	PROPN
alces-23	32	3	is	be	AUX
alces-23	32	4	separated	separate	VERB
alces-23	32	5	from	from	ADP
alces-23	32	6	neighboring	neighboring	NOUN
alces-23	32	7	moose	moose	NOUN
alces-23	32	8	populations	population	NOUN
alces-23	32	9	by	by	ADP
alces-23	32	10	rugged	rugged	ADJ
alces-23	32	11	coastline	coastline	NOUN
alces-23	32	12	characterized	characterize	VERB
alces-23	32	13	by	by	ADP
alces-23	32	14	mature	mature	ADJ
alces-23	32	15	spruce	spruce	NOUN
alces-23	32	16	-	-	PUNCT
alces-23	32	17	hemlock	hemlock	NOUN
alces-23	32	18	(	(	PUNCT
alces-23	32	19	picea	picea	NOUN
alces-23	32	20	sitchensistsuga	sitchensistsuga	NOUN
alces-23	32	21	heterophylla	heterophylla	PROPN
alces-23	32	22	)	)	PUNCT
alces-23	32	23	forest	forest	NOUN
alces-23	32	24	that	that	PRON
alces-23	32	25	is	be	AUX
alces-23	32	26	avoided	avoid	VERB
alces-23	32	27	by	by	ADP
alces-23	32	28	moose	moose	NOUN
alces-23	32	29	(	(	PUNCT
alces-23	32	30	hundertmark	hundertmark	NOUN
alces-23	32	31	et	et	PROPN
alces-23	32	32	al	al	PROPN
alces-23	32	33	.	.	PROPN
alces-23	32	34	1990	1990	NUM
alces-23	32	35	)	)	PUNCT
alces-23	32	36	.	.	PUNCT
alces-23	33	1	predators	predator	NOUN
alces-23	33	2	of	of	ADP
alces-23	33	3	moose	moose	NOUN
alces-23	33	4	occurring	occur	VERB
alces-23	33	5	in	in	ADP
alces-23	33	6	berners	berner	NOUN
alces-23	33	7	bay	bay	PROPN
alces-23	33	8	are	be	AUX
alces-23	33	9	wolves	wolf	NOUN
alces-23	33	10	(	(	PUNCT
alces-23	33	11	canis	canis	NOUN
alces-23	33	12	lupus	lupus	NOUN
alces-23	33	13	)	)	PUNCT
alces-23	33	14	,	,	PUNCT
alces-23	33	15	brown	brown	ADJ
alces-23	33	16	bears	bear	NOUN
alces-23	33	17	(	(	PUNCT
alces-23	33	18	ursus	ursus	PROPN
alces-23	33	19	arctos	arctos	PROPN
alces-23	33	20	)	)	PUNCT
alces-23	33	21	,	,	PUNCT
alces-23	33	22	and	and	CCONJ
alces-23	33	23	black	black	ADJ
alces-23	33	24	bears	bear	NOUN
alces-23	33	25	(	(	PUNCT
alces-23	33	26	u.	u.	NOUN
alces-23	33	27	americanus	americanus	PROPN
alces-23	33	28	)	)	PUNCT
alces-23	33	29	.	.	PUNCT
alces-23	34	1	the	the	DET
alces-23	34	2	moose	moose	NOUN
alces-23	34	3	population	population	NOUN
alces-23	34	4	on	on	ADP
alces-23	34	5	kalgin	kalgin	PROPN
alces-23	34	6	island	island	NOUN
alces-23	34	7	has	have	AUX
alces-23	34	8	undergone	undergo	VERB
alces-23	34	9	dramatic	dramatic	ADJ
alces-23	34	10	fluctuations	fluctuation	NOUN
alces-23	34	11	in	in	ADP
alces-23	34	12	population	population	NOUN
alces-23	34	13	size	size	NOUN
alces-23	34	14	due	due	ADP
alces-23	34	15	to	to	ADP
alces-23	34	16	density	density	NOUN
alces-23	34	17	-	-	PUNCT
alces-23	34	18	dependent	dependent	ADJ
alces-23	34	19	effects	effect	NOUN
alces-23	34	20	of	of	ADP
alces-23	34	21	habitat	habitat	NOUN
alces-23	34	22	and	and	CCONJ
alces-23	34	23	periods	period	NOUN
alces-23	34	24	of	of	ADP
alces-23	34	25	intense	intense	ADJ
alces-23	34	26	harvest	harvest	NOUN
alces-23	34	27	,	,	PUNCT
alces-23	34	28	increasing	increase	VERB
alces-23	34	29	in	in	ADP
alces-23	34	30	size	size	NOUN
alces-23	34	31	to	to	ADP
alces-23	34	32	an	an	DET
alces-23	34	33	estimated	estimate	VERB
alces-23	34	34	212	212	NUM
alces-23	34	35	individuals	individual	NOUN
alces-23	34	36	in	in	ADP
alces-23	34	37	1982	1982	NUM
alces-23	34	38	,	,	PUNCT
alces-23	34	39	declining	decline	VERB
alces-23	34	40	to	to	ADP
alces-23	34	41	8	8	NUM
alces-23	34	42	in	in	ADP
alces-23	34	43	1986	1986	NUM
alces-23	34	44	,	,	PUNCT
alces-23	34	45	and	and	CCONJ
alces-23	34	46	increasing	increase	VERB
alces-23	34	47	since	since	SCONJ
alces-23	34	48	then	then	ADV
alces-23	34	49	(	(	PUNCT
alces-23	34	50	bowyer	bowyer	PROPN
alces-23	34	51	et	et	PROPN
alces-23	34	52	al	al	PROPN
alces-23	34	53	.	.	PROPN
alces-23	34	54	1999	1999	NUM
alces-23	34	55	)	)	PUNCT
alces-23	34	56	.	.	PUNCT
alces-23	35	1	the	the	DET
alces-23	35	2	berners	berners	PROPN
alces-23	35	3	bay	bay	PROPN
alces-23	35	4	population	population	NOUN
alces-23	35	5	is	be	AUX
alces-23	35	6	stable	stable	ADJ
alces-23	35	7	and	and	CCONJ
alces-23	35	8	thought	think	VERB
alces-23	35	9	to	to	PART
alces-23	35	10	be	be	AUX
alces-23	35	11	close	close	ADJ
alces-23	35	12	to	to	ADP
alces-23	35	13	carrying	carry	VERB
alces-23	35	14	capacity	capacity	NOUN
alces-23	35	15	at	at	ADP
alces-23	35	16	120	120	NUM
alces-23	35	17	-	-	SYM
alces-23	35	18	150	150	NUM
alces-23	35	19	individuals	individual	NOUN
alces-23	35	20	(	(	PUNCT
alces-23	35	21	barten	barten	NOUN
alces-23	35	22	2008	2008	NUM
alces-23	35	23	)	)	PUNCT
alces-23	35	24	.	.	PUNCT
alces-23	36	1	both	both	DET
alces-23	36	2	populations	population	NOUN
alces-23	36	3	support	support	VERB
alces-23	36	4	limited	limited	ADJ
alces-23	36	5	harvest	harvest	NOUN
alces-23	36	6	.	.	PUNCT
alces-23	37	1	the	the	DET
alces-23	37	2	moose	moose	NOUN
alces-23	37	3	population	population	NOUN
alces-23	37	4	in	in	ADP
alces-23	37	5	yukon	yukon	PROPN
alces-23	37	6	flats	flat	NOUN
alces-23	37	7	(	(	PUNCT
alces-23	37	8	66º	66º	NUM
alces-23	37	9	10	10	NUM
alces-23	37	10	’	'	PUNCT
alces-23	37	11	n	n	CCONJ
alces-23	37	12	,	,	PUNCT
alces-23	37	13	149º	149º	NUM
alces-23	37	14	00	00	NUM
alces-23	37	15	’	'	PUNCT
alces-23	37	16	w	w	NOUN
alces-23	37	17	)	)	PUNCT
alces-23	37	18	occurs	occur	VERB
alces-23	37	19	in	in	ADP
alces-23	37	20	lowland	lowland	PROPN
alces-23	37	21	boreal	boreal	PROPN
alces-23	37	22	forest	forest	NOUN
alces-23	37	23	along	along	ADP
alces-23	37	24	the	the	DET
alces-23	37	25	yukon	yukon	PROPN
alces-23	37	26	river	river	PROPN
alces-23	37	27	in	in	ADP
alces-23	37	28	central	central	ADJ
alces-23	37	29	alaska	alaska	PROPN
alces-23	37	30	.	.	PUNCT
alces-23	38	1	although	although	SCONJ
alces-23	38	2	the	the	DET
alces-23	38	3	population	population	NOUN
alces-23	38	4	exists	exist	VERB
alces-23	38	5	in	in	ADP
alces-23	38	6	a	a	DET
alces-23	38	7	large	large	ADJ
alces-23	38	8	contiguous	contiguous	ADJ
alces-23	38	9	area	area	NOUN
alces-23	38	10	of	of	ADP
alces-23	38	11	suitable	suitable	ADJ
alces-23	38	12	moose	moose	NOUN
alces-23	38	13	habitat	habitat	NOUN
alces-23	38	14	,	,	PUNCT
alces-23	38	15	it	it	PRON
alces-23	38	16	occurs	occur	VERB
alces-23	38	17	at	at	ADP
alces-23	38	18	extremely	extremely	ADV
alces-23	38	19	low	low	ADJ
alces-23	38	20	density	density	NOUN
alces-23	38	21	(	(	PUNCT
alces-23	38	22	caikoski	caikoski	NOUN
alces-23	38	23	2008	2008	NUM
alces-23	38	24	)	)	PUNCT
alces-23	38	25	presumably	presumably	ADV
alces-23	38	26	due	due	ADP
alces-23	38	27	to	to	ADP
alces-23	38	28	predation	predation	NOUN
alces-23	38	29	(	(	PUNCT
alces-23	38	30	bertram	bertram	PROPN
alces-23	38	31	and	and	CCONJ
alces-23	38	32	vivion	vivion	NOUN
alces-23	38	33	2002	2002	NUM
alces-23	38	34	)	)	PUNCT
alces-23	38	35	and	and	CCONJ
alces-23	38	36	poaching	poach	VERB
alces-23	38	37	of	of	ADP
alces-23	38	38	female	female	ADJ
alces-23	38	39	moose	moose	NOUN
alces-23	38	40	(	(	PUNCT
alces-23	38	41	caikoski	caikoski	NOUN
alces-23	38	42	2008	2008	NUM
alces-23	38	43	)	)	PUNCT
alces-23	38	44	.	.	PUNCT
alces-23	39	1	nonetheless	nonetheless	ADV
alces-23	39	2	,	,	PUNCT
alces-23	39	3	yukon	yukon	PROPN
alces-23	39	4	flats	flat	NOUN
alces-23	39	5	is	be	AUX
alces-23	39	6	an	an	DET
alces-23	39	7	open	open	ADJ
alces-23	39	8	population	population	NOUN
alces-23	39	9	as	as	SCONJ
alces-23	39	10	opposed	oppose	VERB
alces-23	39	11	to	to	ADP
alces-23	39	12	the	the	DET
alces-23	39	13	presumably	presumably	ADV
alces-23	39	14	closed	close	VERB
alces-23	39	15	nature	nature	NOUN
alces-23	39	16	of	of	ADP
alces-23	39	17	the	the	DET
alces-23	39	18	berners	berner	NOUN
alces-23	39	19	bay	bay	NOUN
alces-23	39	20	and	and	CCONJ
alces-23	39	21	kalgin	kalgin	VERB
alces-23	39	22	island	island	NOUN
alces-23	39	23	populations	population	NOUN
alces-23	39	24	.	.	PUNCT
alces-23	40	1	although	although	SCONJ
alces-23	40	2	the	the	DET
alces-23	40	3	yukon	yukon	PROPN
alces-23	40	4	flats	flat	VERB
alces-23	40	5	population	population	NOUN
alces-23	40	6	was	be	AUX
alces-23	40	7	not	not	PART
alces-23	40	8	the	the	DET
alces-23	40	9	source	source	NOUN
alces-23	40	10	of	of	ADP
alces-23	40	11	founders	founder	NOUN
alces-23	40	12	for	for	ADP
alces-23	40	13	either	either	CCONJ
alces-23	40	14	berners	berner	NOUN
alces-23	40	15	bay	bay	NOUN
alces-23	40	16	or	or	CCONJ
alces-23	40	17	kalgin	kalgin	PROPN
alces-23	40	18	island	island	NOUN
alces-23	40	19	,	,	PUNCT
alces-23	40	20	it	it	PRON
alces-23	40	21	serves	serve	VERB
alces-23	40	22	as	as	ADP
alces-23	40	23	a	a	DET
alces-23	40	24	good	good	ADJ
alces-23	40	25	example	example	NOUN
alces-23	40	26	of	of	ADP
alces-23	40	27	an	an	DET
alces-23	40	28	outbred	outbred	ADJ
alces-23	40	29	alaskan	alaskan	ADJ
alces-23	40	30	moose	moose	NOUN
alces-23	40	31	population	population	NOUN
alces-23	40	32	and	and	CCONJ
alces-23	40	33	should	should	AUX
alces-23	40	34	serve	serve	VERB
alces-23	40	35	as	as	ADP
alces-23	40	36	a	a	DET
alces-23	40	37	suitable	suitable	ADJ
alces-23	40	38	control	control	NOUN
alces-23	40	39	population	population	NOUN
alces-23	40	40	in	in	ADP
alces-23	40	41	lieu	lieu	NOUN
alces-23	40	42	of	of	ADP
alces-23	40	43	samples	sample	NOUN
alces-23	40	44	from	from	ADP
alces-23	40	45	south	south	ADJ
alces-23	40	46	-	-	PUNCT
alces-23	40	47	central	central	ADJ
alces-23	40	48	alaska	alaska	PROPN
alces-23	40	49	.	.	PUNCT
alces-23	41	1	indeed	indeed	ADV
alces-23	41	2	,	,	PUNCT
alces-23	41	3	schmidt	schmidt	PROPN
alces-23	41	4	et	et	PROPN
alces-23	41	5	al	al	PROPN
alces-23	41	6	.	.	PROPN
alces-23	42	1	(	(	PUNCT
alces-23	42	2	2009	2009	NUM
alces-23	42	3	)	)	PUNCT
alces-23	42	4	found	find	VERB
alces-23	42	5	little	little	ADJ
alces-23	42	6	difference	difference	NOUN
alces-23	42	7	in	in	ADP
alces-23	42	8	levels	level	NOUN
alces-23	42	9	of	of	ADP
alces-23	42	10	diversity	diversity	NOUN
alces-23	42	11	among	among	ADP
alces-23	42	12	6	6	NUM
alces-23	42	13	moose	moose	NOUN
alces-23	42	14	populations	population	NOUN
alces-23	42	15	distributed	distribute	VERB
alces-23	42	16	widely	widely	ADV
alces-23	42	17	within	within	ADP
alces-23	42	18	alaska	alaska	NOUN
alces-23	42	19	,	,	PUNCT
alces-23	42	20	demonstrating	demonstrate	VERB
alces-23	42	21	that	that	DET
alces-23	42	22	location	location	NOUN
alces-23	42	23	of	of	ADP
alces-23	42	24	the	the	DET
alces-23	42	25	population	population	NOUN
alces-23	42	26	is	be	AUX
alces-23	42	27	less	less	ADV
alces-23	42	28	important	important	ADJ
alces-23	42	29	than	than	ADP
alces-23	42	30	demographic	demographic	ADJ
alces-23	42	31	history	history	NOUN
alces-23	42	32	.	.	PUNCT
alces-23	43	1	methods	method	NOUN
alces-23	43	2	samples	sample	NOUN
alces-23	43	3	for	for	ADP
alces-23	43	4	genetic	genetic	ADJ
alces-23	43	5	analysis	analysis	NOUN
alces-23	43	6	were	be	AUX
alces-23	43	7	acquired	acquire	VERB
alces-23	43	8	either	either	CCONJ
alces-23	43	9	as	as	ADP
alces-23	43	10	tissue	tissue	NOUN
alces-23	43	11	from	from	ADP
alces-23	43	12	hunters	hunter	NOUN
alces-23	43	13	(	(	PUNCT
alces-23	43	14	kalgin	kalgin	NOUN
alces-23	43	15	island	island	NOUN
alces-23	43	16	and	and	CCONJ
alces-23	43	17	berners	berners	PROPN
alces-23	43	18	bay	bay	PROPN
alces-23	43	19	)	)	PUNCT
alces-23	43	20	or	or	CCONJ
alces-23	43	21	as	as	ADP
alces-23	43	22	blood	blood	NOUN
alces-23	43	23	samples	sample	NOUN
alces-23	43	24	from	from	ADP
alces-23	43	25	captured	capture	VERB
alces-23	43	26	animals	animal	NOUN
alces-23	43	27	(	(	PUNCT
alces-23	43	28	yukon	yukon	PROPN
alces-23	43	29	flats	flat	NOUN
alces-23	43	30	)	)	PUNCT
alces-23	43	31	.	.	PUNCT
alces-23	44	1	samples	sample	NOUN
alces-23	44	2	for	for	ADP
alces-23	44	3	each	each	DET
alces-23	44	4	population	population	NOUN
alces-23	44	5	were	be	AUX
alces-23	44	6	collected	collect	VERB
alces-23	44	7	within	within	ADP
alces-23	44	8	a	a	DET
alces-23	44	9	single	single	ADJ
alces-23	44	10	year	year	NOUN
alces-23	44	11	.	.	PUNCT
alces-23	45	1	dna	dna	PROPN
alces-23	45	2	extraction	extraction	NOUN
alces-23	45	3	and	and	CCONJ
alces-23	45	4	genotyping	genotyping	NOUN
alces-23	45	5	were	be	AUX
alces-23	45	6	conducted	conduct	VERB
alces-23	45	7	under	under	ADP
alces-23	45	8	contract	contract	NOUN
alces-23	45	9	in	in	ADP
alces-23	45	10	one	one	NUM
alces-23	45	11	of	of	ADP
alces-23	45	12	two	two	NUM
alces-23	45	13	laboratories	laboratory	NOUN
alces-23	45	14	:	:	PUNCT
alces-23	45	15	kalgin	kalgin	NOUN
alces-23	45	16	island	island	NOUN
alces-23	45	17	and	and	CCONJ
alces-23	45	18	berners	berners	PROPN
alces-23	45	19	bay	bay	NOUN
alces-23	45	20	samples	sample	NOUN
alces-23	45	21	were	be	AUX
alces-23	45	22	analyzed	analyze	VERB
alces-23	45	23	at	at	ADP
alces-23	45	24	wildlife	wildlife	NOUN
alces-23	45	25	genetics	genetic	NOUN
alces-23	45	26	international	international	ADJ
alces-23	45	27	(	(	PUNCT
alces-23	45	28	nelson	nelson	PROPN
alces-23	45	29	,	,	PUNCT
alces-23	45	30	british	british	PROPN
alces-23	45	31	columbia	columbia	PROPN
alces-23	45	32	,	,	PUNCT
alces-23	45	33	canada	canada	PROPN
alces-23	45	34	)	)	PUNCT
alces-23	45	35	,	,	PUNCT
alces-23	45	36	whereas	whereas	SCONJ
alces-23	45	37	yukon	yukon	PROPN
alces-23	45	38	flats	flat	NOUN
alces-23	45	39	moose	moose	NOUN
alces-23	45	40	were	be	AUX
alces-23	45	41	analyzed	analyze	VERB
alces-23	45	42	at	at	ADP
alces-23	45	43	the	the	DET
alces-23	45	44	department	department	NOUN
alces-23	45	45	of	of	ADP
alces-23	45	46	biological	biological	ADJ
alces-23	45	47	sciences	science	NOUN
alces-23	45	48	,	,	PUNCT
alces-23	45	49	university	university	NOUN
alces-23	45	50	of	of	ADP
alces-23	45	51	alberta	alberta	PROPN
alces-23	45	52	(	(	PUNCT
alces-23	45	53	calgary	calgary	PROPN
alces-23	45	54	,	,	PUNCT
alces-23	45	55	alberta	alberta	PROPN
alces-23	45	56	,	,	PUNCT
alces-23	45	57	canada	canada	PROPN
alces-23	45	58	)	)	PUNCT
alces-23	45	59	.	.	PUNCT
alces-23	46	1	two	two	NUM
alces-23	46	2	samples	sample	NOUN
alces-23	46	3	from	from	ADP
alces-23	46	4	yukon	yukon	PROPN
alces-23	46	5	flats	flat	NOUN
alces-23	46	6	were	be	AUX
alces-23	46	7	also	also	ADV
alces-23	46	8	analyzed	analyze	VERB
alces-23	46	9	by	by	ADP
alces-23	46	10	wildlife	wildlife	NOUN
alces-23	46	11	genetics	genetic	NOUN
alces-23	46	12	international	international	ADJ
alces-23	46	13	to	to	PART
alces-23	46	14	ensure	ensure	VERB
alces-23	46	15	consistency	consistency	NOUN
alces-23	46	16	in	in	ADP
alces-23	46	17	allele	allele	NOUN
alces-23	46	18	calling	call	VERB
alces-23	46	19	and	and	CCONJ
alces-23	46	20	warrant	warrant	VERB
alces-23	46	21	comparison	comparison	NOUN
alces-23	46	22	of	of	ADP
alces-23	46	23	genotypes	genotype	NOUN
alces-23	46	24	from	from	ADP
alces-23	46	25	the	the	DET
alces-23	46	26	two	two	NUM
alces-23	46	27	labs	lab	NOUN
alces-23	46	28	.	.	PUNCT
alces-23	47	1	i	i	PRON
alces-23	47	2	analyzed	analyze	VERB
alces-23	47	3	19	19	NUM
alces-23	47	4	samples	sample	NOUN
alces-23	47	5	from	from	ADP
alces-23	47	6	kalgin	kalgin	PROPN
alces-23	47	7	island	island	NOUN
alces-23	47	8	,	,	PUNCT
alces-23	47	9	8	8	NUM
alces-23	47	10	from	from	ADP
alces-23	47	11	berners	berner	NOUN
alces-23	47	12	bay	bay	NOUN
alces-23	47	13	,	,	PUNCT
alces-23	47	14	and	and	CCONJ
alces-23	47	15	28	28	NUM
alces-23	47	16	from	from	ADP
alces-23	47	17	yukon	yukon	PROPN
alces-23	47	18	flats	flat	NOUN
alces-23	47	19	.	.	PUNCT
alces-23	48	1	loci	loci	PROPN
alces-23	48	2	bl42	bl42	PROPN
alces-23	48	3	,	,	PUNCT
alces-23	48	4	bm4513	bm4513	NOUN
alces-23	48	5	,	,	PUNCT
alces-23	48	6	bm888	bm888	PROPN
alces-23	48	7	,	,	PUNCT
alces-23	48	8	bm1222	bm1222	PROPN
alces-23	48	9	,	,	PUNCT
alces-23	48	10	bm203	bm203	PROPN
alces-23	48	11	,	,	PUNCT
alces-23	48	12	bm848	bm848	PROPN
alces-23	48	13	(	(	PUNCT
alces-23	48	14	bishop	bishop	PROPN
alces-23	48	15	et	et	PROPN
alces-23	48	16	al	al	PROPN
alces-23	48	17	.	.	PROPN
alces-23	48	18	1994	1994	NUM
alces-23	48	19	)	)	PUNCT
alces-23	48	20	,	,	PUNCT
alces-23	48	21	fcb193	fcb193	PROPN
alces-23	48	22	(	(	PUNCT
alces-23	48	23	buchanan	buchanan	PROPN
alces-23	48	24	and	and	CCONJ
alces-23	48	25	crawford	crawford	PROPN
alces-23	48	26	1993	1993	NUM
alces-23	48	27	)	)	PUNCT
alces-23	48	28	,	,	PUNCT
alces-23	48	29	rt5	rt5	NOUN
alces-23	48	30	,	,	PUNCT
alces-23	48	31	rt9	rt9	NOUN
alces-23	48	32	,	,	PUNCT
alces-23	48	33	rt24	rt24	PROPN
alces-23	48	34	,	,	PUNCT
alces-23	48	35	and	and	CCONJ
alces-23	48	36	rt30	rt30	PROPN
alces-23	48	37	(	(	PUNCT
alces-23	48	38	wilson	wilson	PROPN
alces-23	48	39	et	et	PROPN
alces-23	48	40	al	al	PROPN
alces-23	48	41	.	.	PROPN
alces-23	48	42	1997	1997	NUM
alces-23	48	43	)	)	PUNCT
alces-23	48	44	were	be	AUX
alces-23	48	45	used	use	VERB
alces-23	48	46	to	to	PART
alces-23	48	47	characterize	characterize	VERB
alces-23	48	48	genetic	genetic	ADJ
alces-23	48	49	diversity	diversity	NOUN
alces-23	48	50	.	.	PUNCT
alces-23	49	1	populations	population	NOUN
alces-23	49	2	were	be	AUX
alces-23	49	3	tested	test	VERB
alces-23	49	4	to	to	PART
alces-23	49	5	ensure	ensure	VERB
alces-23	49	6	compliance	compliance	NOUN
alces-23	49	7	with	with	ADP
alces-23	49	8	hardyweinberg	hardyweinberg	PROPN
alces-23	49	9	equilibrium	equilibrium	NOUN
alces-23	49	10	using	use	VERB
alces-23	49	11	an	an	DET
alces-23	49	12	exact	exact	ADJ
alces-23	49	13	chialces	chialce	NOUN
alces-23	49	14	vol	vol	NOUN
alces-23	49	15	.	.	PROPN
alces-23	50	1	45	45	NUM
alces-23	50	2	,	,	PUNCT
alces-23	50	3	2009	2009	NUM
alces-23	50	4	hundertmark	hundertmark	ADJ
alces-23	50	5	genetic	genetic	ADJ
alces-23	50	6	diversity	diversity	NOUN
alces-23	50	7	in	in	ADP
alces-23	50	8	introduced	introduce	VERB
alces-23	50	9	populations	population	NOUN
alces-23	50	10	139	139	NUM
alces-23	50	11	square	square	ADJ
alces-23	50	12	test	test	NOUN
alces-23	50	13	implemented	implement	VERB
alces-23	50	14	in	in	ADP
alces-23	50	15	software	software	NOUN
alces-23	50	16	genepop	genepop	NOUN
alces-23	50	17	(	(	PUNCT
alces-23	50	18	raymond	raymond	NOUN
alces-23	50	19	and	and	CCONJ
alces-23	50	20	rousset	rousset	VERB
alces-23	50	21	1995	1995	NUM
alces-23	50	22	)	)	PUNCT
alces-23	50	23	.	.	PUNCT
alces-23	51	1	diversity	diversity	NOUN
alces-23	51	2	was	be	AUX
alces-23	51	3	expressed	express	VERB
alces-23	51	4	as	as	ADP
alces-23	51	5	allelic	allelic	ADJ
alces-23	51	6	richness	richness	NOUN
alces-23	51	7	(	(	PUNCT
alces-23	51	8	number	number	NOUN
alces-23	51	9	of	of	ADP
alces-23	51	10	alleles	allele	NOUN
alces-23	51	11	per	per	ADP
alces-23	51	12	locus	locus	NOUN
alces-23	51	13	,	,	PUNCT
alces-23	51	14	a	a	PRON
alces-23	51	15	)	)	PUNCT
alces-23	51	16	,	,	PUNCT
alces-23	51	17	observed	observe	VERB
alces-23	51	18	heterozygosity	heterozygosity	NOUN
alces-23	51	19	(	(	PUNCT
alces-23	51	20	ho	ho	INTJ
alces-23	51	21	)	)	PUNCT
alces-23	51	22	and	and	CCONJ
alces-23	51	23	expected	expect	VERB
alces-23	51	24	heterozygosity	heterozygosity	NOUN
alces-23	51	25	(	(	PUNCT
alces-23	51	26	he	he	PRON
alces-23	51	27	)	)	PUNCT
alces-23	51	28	based	base	VERB
alces-23	51	29	on	on	ADP
alces-23	51	30	allele	allele	NOUN
alces-23	51	31	frequencies	frequency	NOUN
alces-23	51	32	assuming	assume	VERB
alces-23	51	33	hardy	hardy	ADJ
alces-23	51	34	-	-	PUNCT
alces-23	51	35	weinberg	weinberg	PROPN
alces-23	51	36	equilibrium	equilibrium	NOUN
alces-23	51	37	.	.	PUNCT
alces-23	52	1	because	because	SCONJ
alces-23	52	2	estimates	estimate	NOUN
alces-23	52	3	of	of	ADP
alces-23	52	4	allelic	allelic	ADJ
alces-23	52	5	richness	richness	NOUN
alces-23	52	6	are	be	AUX
alces-23	52	7	related	relate	VERB
alces-23	52	8	to	to	ADP
alces-23	52	9	sample	sample	NOUN
alces-23	52	10	size	size	NOUN
alces-23	52	11	,	,	PUNCT
alces-23	52	12	we	we	PRON
alces-23	52	13	standardized	standardize	VERB
alces-23	52	14	our	our	PRON
alces-23	52	15	estimates	estimate	NOUN
alces-23	52	16	by	by	ADP
alces-23	52	17	using	use	VERB
alces-23	52	18	rarefaction	rarefaction	NOUN
alces-23	52	19	(	(	PUNCT
alces-23	52	20	kalinowski	kalinowski	NOUN
alces-23	52	21	2005	2005	NUM
alces-23	52	22	)	)	PUNCT
alces-23	52	23	to	to	PART
alces-23	52	24	express	express	VERB
alces-23	52	25	the	the	DET
alces-23	52	26	expected	expect	VERB
alces-23	52	27	number	number	NOUN
alces-23	52	28	of	of	ADP
alces-23	52	29	alleles	allele	NOUN
alces-23	52	30	per	per	ADP
alces-23	52	31	locus	locus	NOUN
alces-23	52	32	based	base	VERB
alces-23	52	33	on	on	ADP
alces-23	52	34	a	a	DET
alces-23	52	35	sample	sample	NOUN
alces-23	52	36	of	of	ADP
alces-23	52	37	8	8	NUM
alces-23	52	38	individuals	individual	NOUN
alces-23	52	39	from	from	ADP
alces-23	52	40	each	each	DET
alces-23	52	41	population	population	NOUN
alces-23	52	42	(	(	PUNCT
alces-23	52	43	the	the	DET
alces-23	52	44	smallest	small	ADJ
alces-23	52	45	sample	sample	NOUN
alces-23	52	46	size	size	NOUN
alces-23	52	47	of	of	ADP
alces-23	52	48	our	our	PRON
alces-23	52	49	3	3	NUM
alces-23	52	50	populations	population	NOUN
alces-23	52	51	)	)	PUNCT
alces-23	52	52	.	.	PUNCT
alces-23	53	1	estimates	estimate	NOUN
alces-23	53	2	of	of	ADP
alces-23	53	3	allelic	allelic	ADJ
alces-23	53	4	richness	richness	NOUN
alces-23	53	5	were	be	AUX
alces-23	53	6	compared	compare	VERB
alces-23	53	7	between	between	ADP
alces-23	53	8	population	population	NOUN
alces-23	53	9	pairs	pair	NOUN
alces-23	53	10	using	use	VERB
alces-23	53	11	a	a	DET
alces-23	53	12	sign	sign	NOUN
alces-23	53	13	test	test	NOUN
alces-23	53	14	.	.	PUNCT
alces-23	54	1	estimates	estimate	NOUN
alces-23	54	2	of	of	ADP
alces-23	54	3	inbreeding	inbreede	VERB
alces-23	54	4	(	(	PUNCT
alces-23	54	5	fis	fis	PROPN
alces-23	54	6	)	)	PUNCT
alces-23	54	7	were	be	AUX
alces-23	54	8	calculated	calculate	VERB
alces-23	54	9	by	by	ADP
alces-23	54	10	genepop	genepop	NOUN
alces-23	54	11	.	.	PUNCT
alces-23	55	1	population	population	NOUN
alces-23	55	2	differentiation	differentiation	NOUN
alces-23	55	3	was	be	AUX
alces-23	55	4	assessed	assess	VERB
alces-23	55	5	via	via	ADP
alces-23	55	6	nei	nei	PROPN
alces-23	55	7	’s	’s	PART
alces-23	55	8	unbiased	unbiased	ADJ
alces-23	55	9	genetic	genetic	ADJ
alces-23	55	10	distance	distance	NOUN
alces-23	55	11	(	(	PUNCT
alces-23	55	12	nei	nei	NOUN
alces-23	55	13	1978	1978	NUM
alces-23	55	14	)	)	PUNCT
alces-23	55	15	,	,	PUNCT
alces-23	55	16	pairwise	pairwise	NOUN
alces-23	55	17	fst	fst	NOUN
alces-23	55	18	,	,	PUNCT
alces-23	55	19	and	and	CCONJ
alces-23	55	20	comparison	comparison	NOUN
alces-23	55	21	of	of	ADP
alces-23	55	22	allele	allele	NOUN
alces-23	55	23	frequencies	frequency	NOUN
alces-23	55	24	of	of	ADP
alces-23	55	25	populations	population	NOUN
alces-23	55	26	.	.	PUNCT
alces-23	56	1	significance	significance	NOUN
alces-23	56	2	of	of	ADP
alces-23	56	3	differences	difference	NOUN
alces-23	56	4	based	base	VERB
alces-23	56	5	on	on	ADP
alces-23	56	6	pairwise	pairwise	NOUN
alces-23	56	7	fst	fst	NOUN
alces-23	56	8	estimates	estimate	NOUN
alces-23	56	9	was	be	AUX
alces-23	56	10	estimated	estimate	VERB
alces-23	56	11	from	from	ADP
alces-23	56	12	a	a	DET
alces-23	56	13	permutation	permutation	NOUN
alces-23	56	14	test	test	NOUN
alces-23	56	15	conducted	conduct	VERB
alces-23	56	16	in	in	ADP
alces-23	56	17	software	software	NOUN
alces-23	56	18	fstat	fstat	NOUN
alces-23	56	19	(	(	PUNCT
alces-23	56	20	goudet	goudet	NOUN
alces-23	56	21	1995	1995	NUM
alces-23	56	22	)	)	PUNCT
alces-23	56	23	.	.	PUNCT
alces-23	57	1	significance	significance	NOUN
alces-23	57	2	of	of	ADP
alces-23	57	3	pairwise	pairwise	NOUN
alces-23	57	4	comparisons	comparison	NOUN
alces-23	57	5	of	of	ADP
alces-23	57	6	allele	allele	NOUN
alces-23	57	7	frequencies	frequency	NOUN
alces-23	57	8	was	be	AUX
alces-23	57	9	conducted	conduct	VERB
alces-23	57	10	as	as	ADP
alces-23	57	11	a	a	DET
alces-23	57	12	chi	chi	ADJ
alces-23	57	13	-	-	PUNCT
alces-23	57	14	square	square	ADJ
alces-23	57	15	test	test	NOUN
alces-23	57	16	for	for	ADP
alces-23	57	17	each	each	DET
alces-23	57	18	locus	locus	NOUN
alces-23	57	19	and	and	CCONJ
alces-23	57	20	significance	significance	NOUN
alces-23	57	21	values	value	NOUN
alces-23	57	22	were	be	AUX
alces-23	57	23	combined	combine	VERB
alces-23	57	24	via	via	ADP
alces-23	57	25	fisher	fisher	PROPN
alces-23	57	26	’s	’s	PART
alces-23	57	27	method	method	NOUN
alces-23	57	28	(	(	PUNCT
alces-23	57	29	fisher	fisher	PROPN
alces-23	57	30	1948	1948	NUM
alces-23	57	31	)	)	PUNCT
alces-23	57	32	by	by	ADP
alces-23	57	33	genepop	genepop	NOUN
alces-23	57	34	to	to	PART
alces-23	57	35	compute	compute	VERB
alces-23	57	36	a	a	DET
alces-23	57	37	population	population	NOUN
alces-23	57	38	-	-	PUNCT
alces-23	57	39	wide	wide	ADJ
alces-23	57	40	significance	significance	NOUN
alces-23	57	41	level	level	NOUN
alces-23	57	42	.	.	PUNCT
alces-23	58	1	results	result	NOUN
alces-23	58	2	and	and	CCONJ
alces-23	58	3	discussion	discussion	NOUN
alces-23	58	4	all	all	DET
alces-23	58	5	populations	population	NOUN
alces-23	58	6	were	be	AUX
alces-23	58	7	in	in	ADP
alces-23	58	8	hardy	hardy	ADJ
alces-23	58	9	-	-	PUNCT
alces-23	58	10	weinberg	weinberg	NOUN
alces-23	58	11	equilibrium	equilibrium	NOUN
alces-23	58	12	(	(	PUNCT
alces-23	58	13	kalgin	kalgin	NOUN
alces-23	58	14	island	island	NOUN
alces-23	58	15	:	:	PUNCT
alces-23	58	16	χ2	χ2	PROPN
alces-23	58	17	=	=	SYM
alces-23	58	18	15.4	15.4	NUM
alces-23	58	19	,	,	PUNCT
alces-23	58	20	p	p	X
alces-23	58	21	=	=	NOUN
alces-23	58	22	0.75	0.75	NUM
alces-23	58	23	;	;	PUNCT
alces-23	58	24	berners	berners	PROPN
alces-23	58	25	bay	bay	NOUN
alces-23	58	26	:	:	PUNCT
alces-23	59	1	χ2	χ2	PROPN
alces-23	59	2	=	=	SYM
alces-23	59	3	24.5	24.5	NUM
alces-23	59	4	,	,	PUNCT
alces-23	59	5	p	p	X
alces-23	59	6	=	=	NOUN
alces-23	59	7	0.14	0.14	NUM
alces-23	59	8	;	;	PUNCT
alces-23	59	9	yukon	yukon	PROPN
alces-23	59	10	flats	flat	NOUN
alces-23	59	11	:	:	PUNCT
alces-23	59	12	χ2	χ2	PROPN
alces-23	59	13	=	=	SYM
alces-23	59	14	18.8	18.8	NUM
alces-23	59	15	,	,	PUNCT
alces-23	59	16	p	p	X
alces-23	59	17	=	=	NOUN
alces-23	59	18	0.66	0.66	NUM
alces-23	59	19	)	)	PUNCT
alces-23	59	20	.	.	PUNCT
alces-23	60	1	all	all	DET
alces-23	60	2	loci	locus	NOUN
alces-23	60	3	were	be	AUX
alces-23	60	4	polymorphic	polymorphic	ADJ
alces-23	60	5	in	in	ADP
alces-23	60	6	yukon	yukon	PROPN
alces-23	60	7	flats	flat	NOUN
alces-23	60	8	whereas	whereas	SCONJ
alces-23	60	9	one	one	NUM
alces-23	60	10	locus	locus	NOUN
alces-23	60	11	(	(	PUNCT
alces-23	60	12	rt9	rt9	NOUN
alces-23	60	13	)	)	PUNCT
alces-23	60	14	was	be	AUX
alces-23	60	15	monomorphic	monomorphic	ADJ
alces-23	60	16	in	in	ADP
alces-23	60	17	both	both	DET
alces-23	60	18	kalgin	kalgin	NOUN
alces-23	60	19	island	island	NOUN
alces-23	60	20	and	and	CCONJ
alces-23	60	21	berners	berner	NOUN
alces-23	60	22	bay	bay	NOUN
alces-23	60	23	populations	population	NOUN
alces-23	60	24	.	.	PUNCT
alces-23	61	1	all	all	DET
alces-23	61	2	populations	population	NOUN
alces-23	61	3	differed	differ	VERB
alces-23	61	4	from	from	ADP
alces-23	61	5	each	each	DET
alces-23	61	6	other	other	ADJ
alces-23	61	7	in	in	ADP
alces-23	61	8	allelic	allelic	ADJ
alces-23	61	9	richness	richness	NOUN
alces-23	61	10	(	(	PUNCT
alces-23	61	11	p	p	X
alces-23	61	12	<	<	X
alces-23	61	13	0.001	0.001	NUM
alces-23	61	14	;	;	PUNCT
alces-23	61	15	table	table	NOUN
alces-23	61	16	1	1	NUM
alces-23	61	17	)	)	PUNCT
alces-23	61	18	.	.	PUNCT
alces-23	62	1	the	the	DET
alces-23	62	2	3	3	NUM
alces-23	62	3	populations	population	NOUN
alces-23	62	4	shared	share	VERB
alces-23	62	5	at	at	ADP
alces-23	62	6	least	least	ADV
alces-23	62	7	one	one	NUM
alces-23	62	8	allele	allele	NOUN
alces-23	62	9	at	at	ADP
alces-23	62	10	each	each	DET
alces-23	62	11	locus	locus	NOUN
alces-23	62	12	,	,	PUNCT
alces-23	62	13	but	but	CCONJ
alces-23	62	14	allele	allele	NOUN
alces-23	62	15	frequencies	frequency	NOUN
alces-23	62	16	differed	differ	VERB
alces-23	62	17	in	in	ADP
alces-23	62	18	pairwise	pairwise	NOUN
alces-23	62	19	comparisons	comparison	NOUN
alces-23	62	20	(	(	PUNCT
alces-23	62	21	p	p	X
alces-23	62	22	<	<	X
alces-23	62	23	0.0001	0.0001	NUM
alces-23	62	24	)	)	PUNCT
alces-23	62	25	.	.	PUNCT
alces-23	63	1	the	the	DET
alces-23	63	2	extent	extent	NOUN
alces-23	63	3	of	of	ADP
alces-23	63	4	the	the	DET
alces-23	63	5	reduction	reduction	NOUN
alces-23	63	6	in	in	ADP
alces-23	63	7	diversity	diversity	NOUN
alces-23	63	8	undergone	undergo	VERB
alces-23	63	9	by	by	ADP
alces-23	63	10	kalgin	kalgin	NOUN
alces-23	63	11	island	island	PROPN
alces-23	63	12	and	and	CCONJ
alces-23	63	13	berners	berners	PROPN
alces-23	63	14	bay	bay	NOUN
alces-23	63	15	populations	population	NOUN
alces-23	63	16	is	be	AUX
alces-23	63	17	illustrated	illustrate	VERB
alces-23	63	18	by	by	ADP
alces-23	63	19	the	the	DET
alces-23	63	20	occurrence	occurrence	NOUN
alces-23	63	21	of	of	ADP
alces-23	63	22	private	private	ADJ
alces-23	63	23	alleles	allele	NOUN
alces-23	63	24	(	(	PUNCT
alces-23	63	25	those	those	PRON
alces-23	63	26	occurring	occur	VERB
alces-23	63	27	in	in	ADP
alces-23	63	28	only	only	ADV
alces-23	63	29	one	one	NUM
alces-23	63	30	population	population	NOUN
alces-23	63	31	;	;	PUNCT
alces-23	63	32	table	table	NOUN
alces-23	63	33	1	1	NUM
alces-23	63	34	)	)	PUNCT
alces-23	63	35	,	,	PUNCT
alces-23	63	36	wherein	wherein	SCONJ
alces-23	63	37	yukon	yukon	PROPN
alces-23	63	38	flats	flat	NOUN
alces-23	63	39	had	have	VERB
alces-23	63	40	17	17	NUM
alces-23	63	41	alleles	allele	NOUN
alces-23	63	42	not	not	PART
alces-23	63	43	found	find	VERB
alces-23	63	44	in	in	ADP
alces-23	63	45	the	the	DET
alces-23	63	46	other	other	ADJ
alces-23	63	47	populations	population	NOUN
alces-23	63	48	.	.	PUNCT
alces-23	64	1	kalgin	kalgin	PROPN
alces-23	64	2	island	island	PROPN
alces-23	64	3	exhibited	exhibit	VERB
alces-23	64	4	the	the	DET
alces-23	64	5	least	least	ADJ
alces-23	64	6	diversity	diversity	NOUN
alces-23	64	7	,	,	PUNCT
alces-23	64	8	as	as	SCONJ
alces-23	64	9	measured	measure	VERB
alces-23	64	10	by	by	ADP
alces-23	64	11	allelic	allelic	ADJ
alces-23	64	12	richness	richness	NOUN
alces-23	64	13	and	and	CCONJ
alces-23	64	14	heterozygosity	heterozygosity	NOUN
alces-23	64	15	,	,	PUNCT
alces-23	64	16	followed	follow	VERB
alces-23	64	17	by	by	ADP
alces-23	64	18	berners	berner	NOUN
alces-23	64	19	bay	bay	PROPN
alces-23	64	20	and	and	CCONJ
alces-23	64	21	yukon	yukon	PROPN
alces-23	64	22	flats	flat	NOUN
alces-23	64	23	(	(	PUNCT
alces-23	64	24	table	table	NOUN
alces-23	64	25	1	1	NUM
alces-23	64	26	)	)	PUNCT
alces-23	64	27	.	.	PUNCT
alces-23	65	1	inbreeding	inbreede	VERB
alces-23	65	2	coefficients	coefficient	NOUN
alces-23	65	3	for	for	ADP
alces-23	65	4	all	all	DET
alces-23	65	5	populations	population	NOUN
alces-23	65	6	were	be	AUX
alces-23	65	7	very	very	ADV
alces-23	65	8	close	close	ADJ
alces-23	65	9	to	to	ADP
alces-23	65	10	zero	zero	NUM
alces-23	65	11	,	,	PUNCT
alces-23	65	12	indicating	indicate	VERB
alces-23	65	13	no	no	DET
alces-23	65	14	evidence	evidence	NOUN
alces-23	65	15	of	of	ADP
alces-23	65	16	inbreeding	inbreede	VERB
alces-23	65	17	.	.	PUNCT
alces-23	66	1	all	all	DET
alces-23	66	2	populations	population	NOUN
alces-23	66	3	differed	differ	VERB
alces-23	66	4	from	from	ADP
alces-23	66	5	each	each	DET
alces-23	66	6	other	other	ADJ
alces-23	66	7	in	in	ADP
alces-23	66	8	pairwise	pairwise	NOUN
alces-23	66	9	comparisons	comparison	NOUN
alces-23	66	10	of	of	ADP
alces-23	66	11	genetic	genetic	ADJ
alces-23	66	12	distance	distance	NOUN
alces-23	66	13	and	and	CCONJ
alces-23	66	14	fst	fst	NOUN
alces-23	66	15	(	(	PUNCT
alces-23	66	16	table	table	NOUN
alces-23	66	17	2	2	NUM
alces-23	66	18	)	)	PUNCT
alces-23	66	19	;	;	PUNCT
alces-23	66	20	the	the	DET
alces-23	66	21	largest	large	ADJ
alces-23	66	22	differences	difference	NOUN
alces-23	66	23	were	be	AUX
alces-23	66	24	between	between	ADP
alces-23	66	25	the	the	DET
alces-23	66	26	2	2	NUM
alces-23	66	27	introduced	introduce	VERB
alces-23	66	28	populations	population	NOUN
alces-23	66	29	.	.	PUNCT
alces-23	67	1	it	it	PRON
alces-23	67	2	is	be	AUX
alces-23	67	3	striking	strike	VERB
alces-23	67	4	that	that	SCONJ
alces-23	67	5	kalgin	kalgin	VERB
alces-23	67	6	island	island	NOUN
alces-23	67	7	and	and	CCONJ
alces-23	67	8	berners	berners	PROPN
alces-23	67	9	bay	bay	NOUN
alces-23	67	10	populations	population	NOUN
alces-23	67	11	differed	differ	VERB
alces-23	67	12	to	to	ADP
alces-23	67	13	such	such	DET
alces-23	67	14	a	a	DET
alces-23	67	15	degree	degree	NOUN
alces-23	67	16	considering	consider	VERB
alces-23	67	17	that	that	SCONJ
alces-23	67	18	their	their	PRON
alces-23	67	19	founding	founding	NOUN
alces-23	67	20	individuals	individual	NOUN
alces-23	67	21	came	come	VERB
alces-23	67	22	from	from	ADP
alces-23	67	23	the	the	DET
alces-23	67	24	same	same	ADJ
alces-23	67	25	general	general	ADJ
alces-23	67	26	area	area	NOUN
alces-23	67	27	.	.	PUNCT
alces-23	68	1	sample	sample	NOUN
alces-23	68	2	size	size	NOUN
alces-23	68	3	of	of	ADP
alces-23	68	4	the	the	DET
alces-23	68	5	berners	berner	NOUN
alces-23	68	6	bay	bay	PROPN
alces-23	68	7	population	population	NOUN
alces-23	68	8	was	be	AUX
alces-23	68	9	less	less	ADJ
alces-23	68	10	than	than	ADP
alces-23	68	11	ideal	ideal	ADJ
alces-23	68	12	but	but	CCONJ
alces-23	68	13	the	the	DET
alces-23	68	14	samples	sample	NOUN
alces-23	68	15	were	be	AUX
alces-23	68	16	obtained	obtain	VERB
alces-23	68	17	opportunistically	opportunistically	ADV
alces-23	68	18	and	and	CCONJ
alces-23	68	19	there	there	PRON
alces-23	68	20	was	be	VERB
alces-23	68	21	a	a	DET
alces-23	68	22	low	low	ADJ
alces-23	68	23	probability	probability	NOUN
alces-23	68	24	of	of	ADP
alces-23	68	25	obtaining	obtain	VERB
alces-23	68	26	additional	additional	ADJ
alces-23	68	27	samples	sample	NOUN
alces-23	68	28	from	from	ADP
alces-23	68	29	the	the	DET
alces-23	68	30	few	few	ADJ
alces-23	68	31	hunters	hunter	NOUN
alces-23	68	32	that	that	PRON
alces-23	68	33	harvest	harvest	VERB
alces-23	68	34	moose	moose	NOUN
alces-23	68	35	there	there	ADV
alces-23	68	36	.	.	PUNCT
alces-23	69	1	nonetheless	nonetheless	ADV
alces-23	69	2	,	,	PUNCT
alces-23	69	3	estimates	estimate	NOUN
alces-23	69	4	of	of	ADP
alces-23	69	5	observed	observed	ADJ
alces-23	69	6	and	and	CCONJ
alces-23	69	7	expected	expect	VERB
alces-23	69	8	heterozygosity	heterozygosity	NOUN
alces-23	69	9	,	,	PUNCT
alces-23	69	10	as	as	ADV
alces-23	69	11	well	well	ADV
alces-23	69	12	as	as	ADP
alces-23	69	13	fis	fis	PROPN
alces-23	69	14	and	and	CCONJ
alces-23	69	15	fst	fst	NOUN
alces-23	69	16	that	that	PRON
alces-23	69	17	are	be	AUX
alces-23	69	18	based	base	VERB
alces-23	69	19	on	on	ADP
alces-23	69	20	heterozygosity	heterozygosity	NOUN
alces-23	69	21	,	,	PUNCT
alces-23	69	22	are	be	AUX
alces-23	69	23	not	not	PART
alces-23	69	24	related	relate	VERB
alces-23	69	25	to	to	ADP
alces-23	69	26	sample	sample	NOUN
alces-23	69	27	size	size	NOUN
alces-23	69	28	and	and	CCONJ
alces-23	69	29	would	would	AUX
alces-23	69	30	not	not	PART
alces-23	69	31	be	be	AUX
alces-23	69	32	expected	expect	VERB
alces-23	69	33	to	to	PART
alces-23	69	34	change	change	VERB
alces-23	69	35	predictably	predictably	ADV
alces-23	69	36	with	with	ADP
alces-23	69	37	an	an	DET
alces-23	69	38	increase	increase	NOUN
alces-23	69	39	population	population	NOUN
alces-23	69	40	n	n	CCONJ
alces-23	69	41	a	a	DET
alces-23	69	42	a8	a8	PROPN
alces-23	69	43	private	private	ADJ
alces-23	69	44	alleles	allele	NOUN
alces-23	70	1	ho	ho	INTJ
alces-23	71	1	he	he	PRON
alces-23	71	2	fis	fis	PROPN
alces-23	71	3	kalgin	kalgin	PROPN
alces-23	71	4	island	island	NOUN
alces-23	71	5	19	19	NUM
alces-23	71	6	2.9	2.9	NUM
alces-23	71	7	2.7	2.7	NUM
alces-23	71	8	1	1	NUM
alces-23	71	9	0.47	0.47	NUM
alces-23	71	10	0.45	0.45	NUM
alces-23	71	11	-0.01	-0.01	NUM
alces-23	71	12	berners	berner	NOUN
alces-23	71	13	bay	bay	NOUN
alces-23	71	14	8	8	NUM
alces-23	71	15	3.1	3.1	NUM
alces-23	71	16	3.1	3.1	NUM
alces-23	71	17	2	2	NUM
alces-23	71	18	0.53	0.53	NUM
alces-23	71	19	0.49	0.49	NUM
alces-23	71	20	-0.03	-0.03	NUM
alces-23	71	21	yukon	yukon	PROPN
alces-23	71	22	flats	flat	VERB
alces-23	71	23	28	28	NUM
alces-23	71	24	5.5	5.5	NUM
alces-23	71	25	4.2	4.2	NUM
alces-23	71	26	17	17	NUM
alces-23	71	27	0.67	0.67	NUM
alces-23	71	28	0.64	0.64	NUM
alces-23	71	29	-0.02	-0.02	NUM
alces-23	71	30	table	table	NOUN
alces-23	71	31	1	1	NUM
alces-23	71	32	.	.	PUNCT
alces-23	72	1	indices	index	NOUN
alces-23	72	2	of	of	ADP
alces-23	72	3	genetic	genetic	ADJ
alces-23	72	4	diversity	diversity	NOUN
alces-23	72	5	for	for	ADP
alces-23	72	6	11	11	NUM
alces-23	72	7	microsatellite	microsatellite	NOUN
alces-23	72	8	loci	locus	NOUN
alces-23	72	9	measured	measure	VERB
alces-23	72	10	in	in	ADP
alces-23	72	11	3	3	NUM
alces-23	72	12	alaskan	alaskan	ADJ
alces-23	72	13	moose	moose	NOUN
alces-23	72	14	populations	population	NOUN
alces-23	72	15	.	.	PUNCT
alces-23	73	1	kalgin	kalgin	PROPN
alces-23	73	2	island	island	PROPN
alces-23	73	3	and	and	CCONJ
alces-23	73	4	berners	berners	PROPN
alces-23	73	5	bay	bay	NOUN
alces-23	73	6	populations	population	NOUN
alces-23	73	7	show	show	VERB
alces-23	73	8	limited	limited	ADJ
alces-23	73	9	diversity	diversity	NOUN
alces-23	73	10	due	due	ADP
alces-23	73	11	to	to	ADP
alces-23	73	12	founder	founder	NOUN
alces-23	73	13	events	event	NOUN
alces-23	73	14	and	and	CCONJ
alces-23	73	15	lack	lack	NOUN
alces-23	73	16	of	of	ADP
alces-23	73	17	gene	gene	NOUN
alces-23	73	18	flow	flow	NOUN
alces-23	73	19	with	with	ADP
alces-23	73	20	neighboring	neighboring	NOUN
alces-23	73	21	populations	population	NOUN
alces-23	73	22	.	.	PUNCT
alces-23	74	1	parameters	parameter	NOUN
alces-23	74	2	are	be	AUX
alces-23	74	3	:	:	PUNCT
alces-23	74	4	n	n	CCONJ
alces-23	74	5	=	=	SYM
alces-23	74	6	sample	sample	NOUN
alces-23	74	7	size	size	NOUN
alces-23	74	8	,	,	PUNCT
alces-23	74	9	a	a	DET
alces-23	74	10	=	=	PUNCT
alces-23	74	11	allelic	allelic	ADJ
alces-23	74	12	richness	richness	NOUN
alces-23	74	13	(	(	PUNCT
alces-23	74	14	alleles	allele	NOUN
alces-23	74	15	/	/	SYM
alces-23	74	16	locus	locus	NOUN
alces-23	74	17	)	)	PUNCT
alces-23	74	18	,	,	PUNCT
alces-23	74	19	a8	a8	PROPN
alces-23	74	20	=	=	PUNCT
alces-23	74	21	estimate	estimate	NOUN
alces-23	74	22	of	of	ADP
alces-23	74	23	a	a	DET
alces-23	74	24	standardized	standardize	VERB
alces-23	74	25	to	to	ADP
alces-23	74	26	a	a	DET
alces-23	74	27	sample	sample	NOUN
alces-23	74	28	size	size	NOUN
alces-23	74	29	of	of	ADP
alces-23	74	30	8	8	NUM
alces-23	74	31	,	,	PUNCT
alces-23	74	32	ho	ho	PROPN
alces-23	74	33	=	=	NUM
alces-23	74	34	observed	observed	ADJ
alces-23	74	35	heterozygosity	heterozygosity	NOUN
alces-23	74	36	,	,	PUNCT
alces-23	74	37	he	he	PRON
alces-23	74	38	=	=	AUX
alces-23	74	39	expected	expect	VERB
alces-23	74	40	heterozygosity	heterozygosity	NOUN
alces-23	74	41	,	,	PUNCT
alces-23	74	42	and	and	CCONJ
alces-23	74	43	fis	fis	PROPN
alces-23	74	44	=	=	SYM
alces-23	74	45	inbreeding	inbreede	VERB
alces-23	74	46	coefficient	coefficient	NOUN
alces-23	74	47	.	.	PUNCT
alces-23	75	1	genetic	genetic	ADJ
alces-23	75	2	diversity	diversity	NOUN
alces-23	75	3	in	in	ADP
alces-23	75	4	introduced	introduce	VERB
alces-23	75	5	populations	population	NOUN
alces-23	75	6	hundertmark	hundertmark	NOUN
alces-23	75	7	alces	alce	NOUN
alces-23	75	8	vol	vol	NOUN
alces-23	75	9	.	.	PROPN
alces-23	76	1	45	45	NUM
alces-23	76	2	,	,	PUNCT
alces-23	76	3	2009	2009	NUM
alces-23	76	4	140	140	NUM
alces-23	76	5	in	in	ADP
alces-23	76	6	sample	sample	NOUN
alces-23	76	7	size	size	NOUN
alces-23	76	8	.	.	PUNCT
alces-23	77	1	of	of	ADP
alces-23	77	2	the	the	DET
alces-23	77	3	indices	index	NOUN
alces-23	77	4	of	of	ADP
alces-23	77	5	diversity	diversity	NOUN
alces-23	77	6	that	that	PRON
alces-23	77	7	i	i	PRON
alces-23	77	8	examined	examine	VERB
alces-23	77	9	,	,	PUNCT
alces-23	77	10	only	only	ADV
alces-23	77	11	allelic	allelic	ADJ
alces-23	77	12	richness	richness	NOUN
alces-23	77	13	is	be	AUX
alces-23	77	14	affected	affect	VERB
alces-23	77	15	by	by	ADP
alces-23	77	16	sample	sample	NOUN
alces-23	77	17	size	size	NOUN
alces-23	77	18	(	(	PUNCT
alces-23	77	19	kalinowski	kalinowski	NOUN
alces-23	77	20	2005	2005	NUM
alces-23	77	21	,	,	PUNCT
alces-23	77	22	pruett	pruett	NOUN
alces-23	77	23	and	and	CCONJ
alces-23	77	24	winker	winker	NOUN
alces-23	77	25	2008	2008	NUM
alces-23	77	26	)	)	PUNCT
alces-23	77	27	,	,	PUNCT
alces-23	77	28	which	which	PRON
alces-23	77	29	is	be	AUX
alces-23	77	30	why	why	SCONJ
alces-23	77	31	i	i	PRON
alces-23	77	32	used	use	VERB
alces-23	77	33	a	a	DET
alces-23	77	34	rarefaction	rarefaction	NOUN
alces-23	77	35	method	method	NOUN
alces-23	77	36	so	so	SCONJ
alces-23	77	37	that	that	SCONJ
alces-23	77	38	estimates	estimate	NOUN
alces-23	77	39	of	of	ADP
alces-23	77	40	richness	richness	NOUN
alces-23	77	41	among	among	ADP
alces-23	77	42	populations	population	NOUN
alces-23	77	43	could	could	AUX
alces-23	77	44	be	be	AUX
alces-23	77	45	compared	compare	VERB
alces-23	77	46	.	.	PUNCT
alces-23	78	1	results	result	NOUN
alces-23	78	2	from	from	ADP
alces-23	78	3	a	a	DET
alces-23	78	4	simulation	simulation	NOUN
alces-23	78	5	study	study	NOUN
alces-23	78	6	indicated	indicate	VERB
alces-23	78	7	that	that	SCONJ
alces-23	78	8	mean	mean	ADJ
alces-23	78	9	estimates	estimate	NOUN
alces-23	78	10	of	of	ADP
alces-23	78	11	ho	ho	PROPN
alces-23	78	12	and	and	CCONJ
alces-23	78	13	he	he	PRON
alces-23	78	14	did	do	AUX
alces-23	78	15	not	not	PART
alces-23	78	16	change	change	VERB
alces-23	78	17	significantly	significantly	ADV
alces-23	78	18	for	for	ADP
alces-23	78	19	sample	sample	NOUN
alces-23	78	20	sizes	size	NOUN
alces-23	78	21	ranging	range	VERB
alces-23	78	22	from	from	ADP
alces-23	78	23	5	5	NUM
alces-23	78	24	-	-	SYM
alces-23	78	25	100	100	NUM
alces-23	78	26	individuals	individual	NOUN
alces-23	78	27	,	,	PUNCT
alces-23	78	28	and	and	CCONJ
alces-23	78	29	that	that	SCONJ
alces-23	78	30	estimates	estimate	NOUN
alces-23	78	31	were	be	AUX
alces-23	78	32	more	more	ADV
alces-23	78	33	consistent	consistent	ADJ
alces-23	78	34	over	over	ADP
alces-23	78	35	a	a	DET
alces-23	78	36	range	range	NOUN
alces-23	78	37	of	of	ADP
alces-23	78	38	sample	sample	NOUN
alces-23	78	39	sizes	size	NOUN
alces-23	78	40	for	for	ADP
alces-23	78	41	populations	population	NOUN
alces-23	78	42	with	with	ADP
alces-23	78	43	low	low	ADJ
alces-23	78	44	genetic	genetic	ADJ
alces-23	78	45	diversity	diversity	NOUN
alces-23	78	46	as	as	SCONJ
alces-23	78	47	compared	compare	VERB
alces-23	78	48	with	with	ADP
alces-23	78	49	high	high	ADJ
alces-23	78	50	diversity	diversity	NOUN
alces-23	78	51	(	(	PUNCT
alces-23	78	52	pruett	pruett	NOUN
alces-23	78	53	and	and	CCONJ
alces-23	78	54	winker	winker	NOUN
alces-23	78	55	2008	2008	NUM
alces-23	78	56	)	)	PUNCT
alces-23	78	57	.	.	PUNCT
alces-23	79	1	clearly	clearly	ADV
alces-23	79	2	,	,	PUNCT
alces-23	79	3	moose	moose	NOUN
alces-23	79	4	populations	population	NOUN
alces-23	79	5	in	in	ADP
alces-23	79	6	kalgin	kalgin	NOUN
alces-23	79	7	island	island	NOUN
alces-23	79	8	and	and	CCONJ
alces-23	79	9	berners	berners	PROPN
alces-23	79	10	bay	bay	PROPN
alces-23	79	11	show	show	NOUN
alces-23	79	12	severely	severely	ADV
alces-23	79	13	reduced	reduce	VERB
alces-23	79	14	diversity	diversity	NOUN
alces-23	79	15	relative	relative	ADJ
alces-23	79	16	to	to	ADP
alces-23	79	17	yukon	yukon	PROPN
alces-23	79	18	flats	flat	NOUN
alces-23	79	19	,	,	PUNCT
alces-23	79	20	a	a	DET
alces-23	79	21	result	result	NOUN
alces-23	79	22	that	that	SCONJ
alces-23	79	23	arguably	arguably	ADV
alces-23	79	24	stems	stem	VERB
alces-23	79	25	from	from	ADP
alces-23	79	26	genetic	genetic	ADJ
alces-23	79	27	bottlenecks	bottleneck	NOUN
alces-23	79	28	associated	associate	VERB
alces-23	79	29	with	with	ADP
alces-23	79	30	introduction	introduction	NOUN
alces-23	79	31	.	.	PUNCT
alces-23	80	1	moreover	moreover	ADV
alces-23	80	2	,	,	PUNCT
alces-23	80	3	kalgin	kalgin	PROPN
alces-23	80	4	island	island	NOUN
alces-23	80	5	moose	moose	NOUN
alces-23	80	6	were	be	AUX
alces-23	80	7	significantly	significantly	ADV
alces-23	80	8	less	less	ADV
alces-23	80	9	diverse	diverse	ADJ
alces-23	80	10	that	that	SCONJ
alces-23	80	11	berners	berner	NOUN
alces-23	80	12	bay	bay	PROPN
alces-23	80	13	moose	moose	NOUN
alces-23	80	14	;	;	PUNCT
alces-23	80	15	this	this	DET
alces-23	80	16	difference	difference	NOUN
alces-23	80	17	is	be	AUX
alces-23	80	18	likely	likely	ADJ
alces-23	80	19	a	a	DET
alces-23	80	20	function	function	NOUN
alces-23	80	21	of	of	ADP
alces-23	80	22	the	the	DET
alces-23	80	23	smaller	small	ADJ
alces-23	80	24	founding	founding	NOUN
alces-23	80	25	size	size	NOUN
alces-23	80	26	of	of	ADP
alces-23	80	27	the	the	DET
alces-23	80	28	kalgin	kalgin	PROPN
alces-23	80	29	island	island	NOUN
alces-23	80	30	population	population	NOUN
alces-23	80	31	combined	combine	VERB
alces-23	80	32	with	with	ADP
alces-23	80	33	its	its	PRON
alces-23	80	34	marked	mark	VERB
alces-23	80	35	fluctuations	fluctuation	NOUN
alces-23	80	36	.	.	PUNCT
alces-23	81	1	genetic	genetic	ADJ
alces-23	81	2	differentiation	differentiation	NOUN
alces-23	81	3	among	among	ADP
alces-23	81	4	the	the	DET
alces-23	81	5	3	3	NUM
alces-23	81	6	populations	population	NOUN
alces-23	81	7	was	be	AUX
alces-23	81	8	much	much	ADV
alces-23	81	9	greater	great	ADJ
alces-23	81	10	than	than	ADP
alces-23	81	11	that	that	PRON
alces-23	81	12	reported	report	VERB
alces-23	81	13	among	among	ADP
alces-23	81	14	6	6	NUM
alces-23	81	15	moose	moose	NOUN
alces-23	81	16	populations	population	NOUN
alces-23	81	17	from	from	ADP
alces-23	81	18	alaska	alaska	PROPN
alces-23	81	19	(	(	PUNCT
alces-23	81	20	fst	fst	NOUN
alces-23	81	21	range	range	NOUN
alces-23	81	22	=	=	NOUN
alces-23	81	23	0.014	0.014	NUM
alces-23	81	24	-	-	PUNCT
alces-23	81	25	0.109	0.109	NUM
alces-23	81	26	;	;	PUNCT
alces-23	81	27	schmidt	schmidt	PROPN
alces-23	81	28	et	et	PROPN
alces-23	81	29	al	al	PROPN
alces-23	81	30	.	.	PROPN
alces-23	81	31	2009	2009	NUM
alces-23	81	32	)	)	PUNCT
alces-23	81	33	.	.	PUNCT
alces-23	82	1	that	that	DET
alces-23	82	2	study	study	NOUN
alces-23	82	3	found	find	VERB
alces-23	82	4	a	a	DET
alces-23	82	5	remarkable	remarkable	ADJ
alces-23	82	6	lack	lack	NOUN
alces-23	82	7	of	of	ADP
alces-23	82	8	differentiation	differentiation	NOUN
alces-23	82	9	among	among	ADP
alces-23	82	10	moose	moose	NOUN
alces-23	82	11	across	across	ADP
alces-23	82	12	a	a	DET
alces-23	82	13	large	large	ADJ
alces-23	82	14	geographic	geographic	ADJ
alces-23	82	15	scale	scale	NOUN
alces-23	82	16	,	,	PUNCT
alces-23	82	17	and	and	CCONJ
alces-23	82	18	was	be	AUX
alces-23	82	19	contrary	contrary	ADJ
alces-23	82	20	to	to	ADP
alces-23	82	21	other	other	ADJ
alces-23	82	22	studies	study	NOUN
alces-23	82	23	(	(	PUNCT
alces-23	82	24	broders	broder	NOUN
alces-23	82	25	et	et	PROPN
alces-23	82	26	al	al	PROPN
alces-23	82	27	.	.	PROPN
alces-23	82	28	1999	1999	NUM
alces-23	82	29	,	,	PUNCT
alces-23	82	30	wilson	wilson	PROPN
alces-23	82	31	et	et	PROPN
alces-23	82	32	al	al	PROPN
alces-23	82	33	.	.	PROPN
alces-23	82	34	2003	2003	NUM
alces-23	82	35	)	)	PUNCT
alces-23	82	36	.	.	PUNCT
alces-23	83	1	the	the	DET
alces-23	83	2	relative	relative	ADJ
alces-23	83	3	difference	difference	NOUN
alces-23	83	4	among	among	ADP
alces-23	83	5	the	the	DET
alces-23	83	6	3	3	NUM
alces-23	83	7	study	study	NOUN
alces-23	83	8	populations	population	NOUN
alces-23	83	9	would	would	AUX
alces-23	83	10	be	be	AUX
alces-23	83	11	unexpected	unexpected	ADJ
alces-23	83	12	if	if	SCONJ
alces-23	83	13	the	the	DET
alces-23	83	14	kalgin	kalgin	NOUN
alces-23	83	15	island	island	NOUN
alces-23	83	16	and	and	CCONJ
alces-23	83	17	berners	berners	PROPN
alces-23	83	18	bay	bay	NOUN
alces-23	83	19	populations	population	NOUN
alces-23	83	20	were	be	AUX
alces-23	83	21	open	open	ADJ
alces-23	83	22	and	and	CCONJ
alces-23	83	23	exchanged	exchange	VERB
alces-23	83	24	individuals	individual	NOUN
alces-23	83	25	with	with	ADP
alces-23	83	26	neighboring	neighboring	NOUN
alces-23	83	27	populations	population	NOUN
alces-23	83	28	.	.	PUNCT
alces-23	84	1	thus	thus	ADV
alces-23	84	2	,	,	PUNCT
alces-23	84	3	these	these	DET
alces-23	84	4	differences	difference	NOUN
alces-23	84	5	presumably	presumably	ADV
alces-23	84	6	indicate	indicate	VERB
alces-23	84	7	the	the	DET
alces-23	84	8	strong	strong	ADJ
alces-23	84	9	effect	effect	NOUN
alces-23	84	10	of	of	ADP
alces-23	84	11	founding	found	VERB
alces-23	84	12	combined	combine	VERB
alces-23	84	13	with	with	ADP
alces-23	84	14	genetic	genetic	ADJ
alces-23	84	15	drift	drift	NOUN
alces-23	84	16	associated	associate	VERB
alces-23	84	17	with	with	ADP
alces-23	84	18	isolation	isolation	NOUN
alces-23	84	19	from	from	ADP
alces-23	84	20	neighboring	neighboring	NOUN
alces-23	84	21	populations	population	NOUN
alces-23	84	22	.	.	PUNCT
alces-23	85	1	reduction	reduction	NOUN
alces-23	85	2	in	in	ADP
alces-23	85	3	heterozygosity	heterozygosity	NOUN
alces-23	85	4	associated	associate	VERB
alces-23	85	5	with	with	ADP
alces-23	85	6	founder	founder	NOUN
alces-23	85	7	effect	effect	NOUN
alces-23	85	8	can	can	AUX
alces-23	85	9	be	be	AUX
alces-23	85	10	calculated	calculate	VERB
alces-23	85	11	as	as	ADP
alces-23	85	12	:	:	PUNCT
alces-23	85	13	he	he	PRON
alces-23	85	14	=	=	PUNCT
alces-23	85	15	ho(1	ho(1	NOUN
alces-23	85	16	–	–	PUNCT
alces-23	85	17	1/2n	1/2n	NUM
alces-23	85	18	)	)	PUNCT
alces-23	85	19	.	.	PUNCT
alces-23	86	1	where	where	SCONJ
alces-23	86	2	:	:	PUNCT
alces-23	86	3	ho	ho	PROPN
alces-23	86	4	is	be	AUX
alces-23	86	5	the	the	DET
alces-23	86	6	heterozygosity	heterozygosity	NOUN
alces-23	86	7	of	of	ADP
alces-23	86	8	the	the	DET
alces-23	86	9	source	source	NOUN
alces-23	86	10	population	population	NOUN
alces-23	86	11	,	,	PUNCT
alces-23	86	12	he	he	PRON
alces-23	86	13	is	be	AUX
alces-23	86	14	the	the	DET
alces-23	86	15	expected	expect	VERB
alces-23	86	16	heterozygosity	heterozygosity	NOUN
alces-23	86	17	of	of	ADP
alces-23	86	18	the	the	DET
alces-23	86	19	founded	found	VERB
alces-23	86	20	population	population	NOUN
alces-23	86	21	at	at	ADP
alces-23	86	22	the	the	DET
alces-23	86	23	time	time	NOUN
alces-23	86	24	of	of	ADP
alces-23	86	25	founding	founding	NOUN
alces-23	86	26	,	,	PUNCT
alces-23	86	27	and	and	CCONJ
alces-23	86	28	n	n	PRON
alces-23	86	29	is	be	AUX
alces-23	86	30	the	the	DET
alces-23	86	31	number	number	NOUN
alces-23	86	32	of	of	ADP
alces-23	86	33	individuals	individual	NOUN
alces-23	86	34	introduced	introduce	VERB
alces-23	86	35	.	.	PUNCT
alces-23	87	1	using	use	VERB
alces-23	87	2	yukon	yukon	PROPN
alces-23	87	3	flats	flat	NOUN
alces-23	87	4	as	as	ADP
alces-23	87	5	a	a	DET
alces-23	87	6	proxy	proxy	NOUN
alces-23	87	7	for	for	ADP
alces-23	87	8	the	the	DET
alces-23	87	9	source	source	NOUN
alces-23	87	10	population	population	NOUN
alces-23	87	11	(	(	PUNCT
alces-23	87	12	ho	ho	PROPN
alces-23	87	13	=	=	SYM
alces-23	87	14	0.67	0.67	NUM
alces-23	87	15	;	;	PUNCT
alces-23	87	16	table	table	NOUN
alces-23	87	17	1	1	NUM
alces-23	87	18	)	)	PUNCT
alces-23	87	19	,	,	PUNCT
alces-23	87	20	kalgin	kalgin	VERB
alces-23	87	21	island	island	NOUN
alces-23	87	22	with	with	ADP
alces-23	87	23	a	a	DET
alces-23	87	24	founding	found	VERB
alces-23	87	25	size	size	NOUN
alces-23	87	26	of	of	ADP
alces-23	87	27	6	6	NUM
alces-23	87	28	would	would	AUX
alces-23	87	29	have	have	VERB
alces-23	87	30	he	he	PRON
alces-23	87	31	=	=	NUM
alces-23	87	32	0.61	0.61	NUM
alces-23	87	33	which	which	PRON
alces-23	87	34	is	be	AUX
alces-23	87	35	30	30	NUM
alces-23	87	36	%	%	NOUN
alces-23	87	37	greater	great	ADJ
alces-23	87	38	than	than	ADP
alces-23	87	39	ho	ho	PROPN
alces-23	87	40	for	for	ADP
alces-23	87	41	kalgin	kalgin	NOUN
alces-23	87	42	(	(	PUNCT
alces-23	87	43	0.47	0.47	NUM
alces-23	87	44	;	;	PUNCT
alces-23	87	45	table	table	NOUN
alces-23	87	46	1	1	NUM
alces-23	87	47	)	)	PUNCT
alces-23	87	48	.	.	PUNCT
alces-23	88	1	similarly	similarly	ADV
alces-23	88	2	,	,	PUNCT
alces-23	88	3	he	he	PRON
alces-23	88	4	for	for	ADP
alces-23	88	5	the	the	DET
alces-23	88	6	initial	initial	ADJ
alces-23	88	7	population	population	NOUN
alces-23	88	8	in	in	ADP
alces-23	88	9	berners	berner	NOUN
alces-23	88	10	bay	bay	PROPN
alces-23	88	11	would	would	AUX
alces-23	88	12	be	be	AUX
alces-23	88	13	expected	expect	VERB
alces-23	88	14	to	to	PART
alces-23	88	15	be	be	AUX
alces-23	88	16	0.66	0.66	NUM
alces-23	88	17	or	or	CCONJ
alces-23	88	18	24	24	NUM
alces-23	88	19	%	%	NOUN
alces-23	88	20	higher	high	ADJ
alces-23	88	21	than	than	ADP
alces-23	88	22	observed	observed	ADJ
alces-23	88	23	(	(	PUNCT
alces-23	88	24	0.53	0.53	NUM
alces-23	88	25	;	;	PUNCT
alces-23	88	26	table	table	NOUN
alces-23	88	27	1	1	NUM
alces-23	88	28	)	)	PUNCT
alces-23	88	29	.	.	PUNCT
alces-23	89	1	the	the	DET
alces-23	89	2	differences	difference	NOUN
alces-23	89	3	between	between	ADP
alces-23	89	4	the	the	DET
alces-23	89	5	current	current	ADJ
alces-23	89	6	heterozygosity	heterozygosity	NOUN
alces-23	89	7	in	in	ADP
alces-23	89	8	the	the	DET
alces-23	89	9	introduced	introduce	VERB
alces-23	89	10	populations	population	NOUN
alces-23	89	11	and	and	CCONJ
alces-23	89	12	the	the	DET
alces-23	89	13	expected	expect	VERB
alces-23	89	14	heterozygosity	heterozygosity	NOUN
alces-23	89	15	based	base	VERB
alces-23	89	16	on	on	ADP
alces-23	89	17	number	number	NOUN
alces-23	89	18	of	of	ADP
alces-23	89	19	founders	founder	NOUN
alces-23	89	20	can	can	AUX
alces-23	89	21	be	be	AUX
alces-23	89	22	explained	explain	VERB
alces-23	89	23	by	by	ADP
alces-23	89	24	genetic	genetic	ADJ
alces-23	89	25	drift	drift	NOUN
alces-23	89	26	occurring	occur	VERB
alces-23	89	27	in	in	ADP
alces-23	89	28	the	the	DET
alces-23	89	29	interim	interim	NOUN
alces-23	89	30	.	.	PUNCT
alces-23	90	1	moreover	moreover	ADV
alces-23	90	2	,	,	PUNCT
alces-23	90	3	the	the	DET
alces-23	90	4	severity	severity	NOUN
alces-23	90	5	of	of	ADP
alces-23	90	6	a	a	DET
alces-23	90	7	genetic	genetic	ADJ
alces-23	90	8	bottleneck	bottleneck	NOUN
alces-23	90	9	is	be	AUX
alces-23	90	10	directly	directly	ADV
alces-23	90	11	related	relate	VERB
alces-23	90	12	to	to	ADP
alces-23	90	13	the	the	DET
alces-23	90	14	duration	duration	NOUN
alces-23	90	15	of	of	ADP
alces-23	90	16	the	the	DET
alces-23	90	17	bottleneck	bottleneck	NOUN
alces-23	90	18	(	(	PUNCT
alces-23	90	19	nei	nei	PROPN
alces-23	90	20	et	et	PROPN
alces-23	90	21	al	al	PROPN
alces-23	90	22	.	.	PROPN
alces-23	90	23	1975	1975	NUM
alces-23	90	24	)	)	PUNCT
alces-23	90	25	,	,	PUNCT
alces-23	90	26	suggesting	suggest	VERB
alces-23	90	27	that	that	SCONJ
alces-23	90	28	both	both	DET
alces-23	90	29	populations	population	NOUN
alces-23	90	30	grew	grow	VERB
alces-23	90	31	slowly	slowly	ADV
alces-23	90	32	after	after	ADP
alces-23	90	33	founding	founding	NOUN
alces-23	90	34	,	,	PUNCT
alces-23	90	35	thus	thus	ADV
alces-23	90	36	extending	extend	VERB
alces-23	90	37	the	the	DET
alces-23	90	38	length	length	NOUN
alces-23	90	39	of	of	ADP
alces-23	90	40	the	the	DET
alces-23	90	41	bottleneck	bottleneck	NOUN
alces-23	90	42	;	;	PUNCT
alces-23	90	43	the	the	DET
alces-23	90	44	kalgin	kalgin	PROPN
alces-23	90	45	island	island	NOUN
alces-23	90	46	population	population	NOUN
alces-23	90	47	was	be	AUX
alces-23	90	48	likely	likely	ADV
alces-23	90	49	affected	affect	VERB
alces-23	90	50	by	by	ADP
alces-23	90	51	a	a	DET
alces-23	90	52	second	second	ADJ
alces-23	90	53	bottleneck	bottleneck	NOUN
alces-23	90	54	when	when	SCONJ
alces-23	90	55	the	the	DET
alces-23	90	56	population	population	NOUN
alces-23	90	57	declined	decline	VERB
alces-23	90	58	abruptly	abruptly	ADV
alces-23	90	59	to	to	ADP
alces-23	90	60	the	the	DET
alces-23	90	61	founding	found	VERB
alces-23	90	62	size	size	NOUN
alces-23	90	63	(	(	PUNCT
alces-23	90	64	8	8	NUM
alces-23	90	65	individuals	individual	NOUN
alces-23	90	66	)	)	PUNCT
alces-23	90	67	in	in	ADP
alces-23	90	68	the	the	DET
alces-23	90	69	1980s	1980s	NUM
alces-23	90	70	.	.	PUNCT
alces-23	91	1	interpopulation	interpopulation	NOUN
alces-23	91	2	distances	distance	NOUN
alces-23	91	3	reported	report	VERB
alces-23	91	4	for	for	ADP
alces-23	91	5	moose	moose	NOUN
alces-23	91	6	in	in	ADP
alces-23	91	7	canada	canada	PROPN
alces-23	91	8	(	(	PUNCT
alces-23	91	9	broders	broder	NOUN
alces-23	91	10	et	et	PROPN
alces-23	91	11	al	al	PROPN
alces-23	91	12	.	.	PROPN
alces-23	91	13	1999	1999	NUM
alces-23	91	14	)	)	PUNCT
alces-23	91	15	were	be	AUX
alces-23	91	16	much	much	ADV
alces-23	91	17	lower	low	ADJ
alces-23	91	18	than	than	ADP
alces-23	91	19	those	those	PRON
alces-23	91	20	reported	report	VERB
alces-23	91	21	here	here	ADV
alces-23	91	22	;	;	PUNCT
alces-23	91	23	distances	distance	NOUN
alces-23	91	24	were	be	AUX
alces-23	91	25	0.013	0.013	NUM
alces-23	91	26	-	-	PUNCT
alces-23	91	27	0.298	0.298	NUM
alces-23	91	28	,	,	PUNCT
alces-23	91	29	however	however	ADV
alces-23	91	30	the	the	DET
alces-23	91	31	latter	latter	ADJ
alces-23	91	32	value	value	NOUN
alces-23	91	33	was	be	AUX
alces-23	91	34	essentially	essentially	ADV
alces-23	91	35	a	a	DET
alces-23	91	36	comparison	comparison	NOUN
alces-23	91	37	of	of	ADP
alces-23	91	38	2	2	NUM
alces-23	91	39	different	different	ADJ
alces-23	91	40	subspecies	subspecie	NOUN
alces-23	91	41	representing	represent	VERB
alces-23	91	42	moose	moose	NOUN
alces-23	91	43	from	from	ADP
alces-23	91	44	cape	cape	PROPN
alces-23	91	45	breton	breton	PROPN
alces-23	91	46	island	island	PROPN
alces-23	91	47	,	,	PUNCT
alces-23	91	48	nova	nova	PROPN
alces-23	91	49	scotia	scotia	PROPN
alces-23	91	50	(	(	PUNCT
alces-23	91	51	introduced	introduce	VERB
alces-23	91	52	from	from	ADP
alces-23	91	53	alberta	alberta	PROPN
alces-23	91	54	)	)	PUNCT
alces-23	91	55	and	and	CCONJ
alces-23	91	56	moose	moose	VERB
alces-23	91	57	from	from	ADP
alces-23	91	58	the	the	DET
alces-23	91	59	avalon	avalon	PROPN
alces-23	91	60	peninsula	peninsula	PROPN
alces-23	91	61	,	,	PUNCT
alces-23	91	62	newfoundland	newfoundland	PROPN
alces-23	91	63	(	(	PUNCT
alces-23	91	64	introduced	introduce	VERB
alces-23	91	65	from	from	ADP
alces-23	91	66	nova	nova	PROPN
alces-23	91	67	scotia	scotia	PROPN
alces-23	91	68	kalgin	kalgin	PROPN
alces-23	91	69	island	island	PROPN
alces-23	91	70	berners	berners	PROPN
alces-23	91	71	bay	bay	PROPN
alces-23	91	72	yukon	yukon	PROPN
alces-23	91	73	flats	flat	NOUN
alces-23	91	74	kalgin	kalgin	VERB
alces-23	91	75	island	island	NOUN
alces-23	91	76	0.518	0.518	NUM
alces-23	91	77	0.365	0.365	NUM
alces-23	91	78	berners	berner	NOUN
alces-23	91	79	bay	bay	PROPN
alces-23	91	80	0.301	0.301	NUM
alces-23	91	81	0.31	0.31	NUM
alces-23	91	82	yukon	yukon	PROPN
alces-23	91	83	flats	flat	VERB
alces-23	91	84	0.197	0.197	NUM
alces-23	91	85	0.151	0.151	NUM
alces-23	91	86	table	table	NOUN
alces-23	91	87	2	2	NUM
alces-23	91	88	.	.	PUNCT
alces-23	91	89	indices	index	NOUN
alces-23	91	90	of	of	ADP
alces-23	91	91	population	population	NOUN
alces-23	91	92	differentiation	differentiation	NOUN
alces-23	91	93	based	base	VERB
alces-23	91	94	on	on	ADP
alces-23	91	95	11	11	NUM
alces-23	91	96	microsatellite	microsatellite	NOUN
alces-23	91	97	markers	marker	NOUN
alces-23	91	98	measured	measure	VERB
alces-23	91	99	in	in	ADP
alces-23	91	100	3	3	NUM
alces-23	91	101	alaskan	alaskan	ADJ
alces-23	91	102	moose	moose	NOUN
alces-23	91	103	populations	population	NOUN
alces-23	91	104	.	.	PUNCT
alces-23	92	1	nei	nei	PROPN
alces-23	92	2	’s	’s	PART
alces-23	92	3	(	(	PUNCT
alces-23	92	4	1978	1978	NUM
alces-23	92	5	)	)	PUNCT
alces-23	92	6	unbiased	unbiased	ADJ
alces-23	92	7	genetic	genetic	ADJ
alces-23	92	8	distance	distance	NOUN
alces-23	92	9	is	be	AUX
alces-23	92	10	above	above	ADP
alces-23	92	11	the	the	DET
alces-23	92	12	diagonal	diagonal	ADJ
alces-23	92	13	and	and	CCONJ
alces-23	92	14	fst	fst	NOUN
alces-23	92	15	is	be	AUX
alces-23	92	16	below	below	ADP
alces-23	92	17	the	the	DET
alces-23	92	18	diagonal	diagonal	NOUN
alces-23	92	19	.	.	PUNCT
alces-23	93	1	based	base	VERB
alces-23	93	2	on	on	ADP
alces-23	93	3	fst	fst	NOUN
alces-23	93	4	estimates	estimate	NOUN
alces-23	93	5	,	,	PUNCT
alces-23	93	6	all	all	DET
alces-23	93	7	populations	population	NOUN
alces-23	93	8	differ	differ	VERB
alces-23	93	9	significantly	significantly	ADV
alces-23	93	10	(	(	PUNCT
alces-23	93	11	p	p	X
alces-23	93	12	<	<	X
alces-23	93	13	0.001	0.001	NUM
alces-23	93	14	)	)	PUNCT
alces-23	93	15	.	.	PUNCT
alces-23	94	1	alces	alce	NOUN
alces-23	94	2	vol	vol	NOUN
alces-23	94	3	.	.	PROPN
alces-23	95	1	45	45	NUM
alces-23	95	2	,	,	PUNCT
alces-23	95	3	2009	2009	NUM
alces-23	95	4	hundertmark	hundertmark	ADJ
alces-23	95	5	genetic	genetic	ADJ
alces-23	95	6	diversity	diversity	NOUN
alces-23	95	7	in	in	ADP
alces-23	95	8	introduced	introduce	VERB
alces-23	95	9	populations	population	NOUN
alces-23	95	10	141	141	NUM
alces-23	95	11	and	and	CCONJ
alces-23	95	12	new	new	PROPN
alces-23	95	13	brunswick	brunswick	PROPN
alces-23	95	14	)	)	PUNCT
alces-23	95	15	.	.	PUNCT
alces-23	96	1	distances	distance	NOUN
alces-23	96	2	reported	report	VERB
alces-23	96	3	by	by	ADP
alces-23	96	4	wilson	wilson	PROPN
alces-23	96	5	et	et	PROPN
alces-23	96	6	al	al	PROPN
alces-23	96	7	.	.	PROPN
alces-23	97	1	(	(	PUNCT
alces-23	97	2	2003	2003	NUM
alces-23	97	3	)	)	PUNCT
alces-23	97	4	were	be	AUX
alces-23	97	5	comparable	comparable	ADJ
alces-23	97	6	to	to	ADP
alces-23	97	7	those	those	PRON
alces-23	97	8	reported	report	VERB
alces-23	97	9	here	here	ADV
alces-23	97	10	;	;	PUNCT
alces-23	97	11	however	however	ADV
alces-23	97	12	,	,	PUNCT
alces-23	97	13	the	the	DET
alces-23	97	14	largest	large	ADJ
alces-23	97	15	fst	fst	NOUN
alces-23	97	16	value	value	NOUN
alces-23	97	17	(	(	PUNCT
alces-23	97	18	0.3013	0.3013	NUM
alces-23	97	19	)	)	PUNCT
alces-23	97	20	occurred	occur	VERB
alces-23	97	21	between	between	ADP
alces-23	97	22	populations	population	NOUN
alces-23	97	23	in	in	ADP
alces-23	97	24	newfoundland	newfoundland	NOUN
alces-23	97	25	and	and	CCONJ
alces-23	97	26	riding	ride	VERB
alces-23	97	27	mountain	mountain	NOUN
alces-23	97	28	national	national	ADJ
alces-23	97	29	park	park	PROPN
alces-23	97	30	,	,	PUNCT
alces-23	97	31	manitoba	manitoba	PROPN
alces-23	97	32	and	and	CCONJ
alces-23	97	33	was	be	AUX
alces-23	97	34	a	a	DET
alces-23	97	35	comparison	comparison	NOUN
alces-23	97	36	across	across	ADP
alces-23	97	37	a	a	DET
alces-23	97	38	subspecies	subspecie	NOUN
alces-23	97	39	boundary	boundary	ADJ
alces-23	97	40	.	.	PUNCT
alces-23	98	1	thus	thus	ADV
alces-23	98	2	,	,	PUNCT
alces-23	98	3	the	the	DET
alces-23	98	4	level	level	NOUN
alces-23	98	5	of	of	ADP
alces-23	98	6	differentiation	differentiation	NOUN
alces-23	98	7	observed	observe	VERB
alces-23	98	8	between	between	ADP
alces-23	98	9	the	the	DET
alces-23	98	10	2	2	NUM
alces-23	98	11	introduced	introduce	VERB
alces-23	98	12	alaska	alaska	NOUN
alces-23	98	13	populations	population	NOUN
alces-23	98	14	was	be	AUX
alces-23	98	15	equal	equal	ADJ
alces-23	98	16	to	to	ADP
alces-23	98	17	or	or	CCONJ
alces-23	98	18	greater	great	ADJ
alces-23	98	19	than	than	ADP
alces-23	98	20	that	that	PRON
alces-23	98	21	observed	observe	VERB
alces-23	98	22	between	between	ADP
alces-23	98	23	subspecies	subspecie	NOUN
alces-23	98	24	elsewhere	elsewhere	ADV
alces-23	98	25	on	on	ADP
alces-23	98	26	the	the	DET
alces-23	98	27	continent	continent	NOUN
alces-23	98	28	.	.	PUNCT
alces-23	99	1	comparisons	comparison	NOUN
alces-23	99	2	of	of	ADP
alces-23	99	3	levels	level	NOUN
alces-23	99	4	of	of	ADP
alces-23	99	5	diversity	diversity	NOUN
alces-23	99	6	between	between	ADP
alces-23	99	7	studies	study	NOUN
alces-23	99	8	employing	employ	VERB
alces-23	99	9	different	different	ADJ
alces-23	99	10	sets	set	NOUN
alces-23	99	11	of	of	ADP
alces-23	99	12	molecular	molecular	ADJ
alces-23	99	13	markers	marker	NOUN
alces-23	99	14	require	require	VERB
alces-23	99	15	caution	caution	NOUN
alces-23	99	16	because	because	SCONJ
alces-23	99	17	of	of	ADP
alces-23	99	18	the	the	DET
alces-23	99	19	different	different	ADJ
alces-23	99	20	levels	level	NOUN
alces-23	99	21	of	of	ADP
alces-23	99	22	variation	variation	NOUN
alces-23	99	23	inherent	inherent	ADJ
alces-23	99	24	in	in	ADP
alces-23	99	25	different	different	ADJ
alces-23	99	26	loci	locus	NOUN
alces-23	99	27	.	.	PUNCT
alces-23	100	1	nonetheless	nonetheless	ADV
alces-23	100	2	,	,	PUNCT
alces-23	100	3	if	if	SCONJ
alces-23	100	4	loci	locus	NOUN
alces-23	100	5	are	be	AUX
alces-23	100	6	truly	truly	ADV
alces-23	100	7	neutral	neutral	ADJ
alces-23	100	8	and	and	CCONJ
alces-23	100	9	markers	marker	NOUN
alces-23	100	10	conform	conform	VERB
alces-23	100	11	to	to	ADP
alces-23	100	12	the	the	DET
alces-23	100	13	same	same	ADJ
alces-23	100	14	model	model	NOUN
alces-23	100	15	of	of	ADP
alces-23	100	16	mutation	mutation	NOUN
alces-23	100	17	,	,	PUNCT
alces-23	100	18	estimates	estimate	NOUN
alces-23	100	19	of	of	ADP
alces-23	100	20	population	population	NOUN
alces-23	100	21	differentiation	differentiation	NOUN
alces-23	100	22	should	should	AUX
alces-23	100	23	be	be	AUX
alces-23	100	24	broadly	broadly	ADV
alces-23	100	25	comparable	comparable	ADJ
alces-23	100	26	between	between	ADP
alces-23	100	27	studies	study	NOUN
alces-23	100	28	.	.	PUNCT
alces-23	101	1	i	i	PRON
alces-23	101	2	have	have	AUX
alces-23	101	3	shown	show	VERB
alces-23	101	4	that	that	SCONJ
alces-23	101	5	2	2	NUM
alces-23	101	6	small	small	ADJ
alces-23	101	7	,	,	PUNCT
alces-23	101	8	introduced	introduce	VERB
alces-23	101	9	moose	moose	NOUN
alces-23	101	10	populations	population	NOUN
alces-23	101	11	in	in	ADP
alces-23	101	12	alaska	alaska	PROPN
alces-23	101	13	underwent	undergo	VERB
alces-23	101	14	extreme	extreme	ADJ
alces-23	101	15	reductions	reduction	NOUN
alces-23	101	16	in	in	ADP
alces-23	101	17	genetic	genetic	ADJ
alces-23	101	18	diversity	diversity	NOUN
alces-23	101	19	associated	associate	VERB
alces-23	101	20	with	with	ADP
alces-23	101	21	founding	found	VERB
alces-23	101	22	and	and	CCONJ
alces-23	101	23	subsequent	subsequent	ADJ
alces-23	101	24	genetic	genetic	ADJ
alces-23	101	25	drift	drift	NOUN
alces-23	101	26	.	.	PUNCT
alces-23	102	1	it	it	PRON
alces-23	102	2	is	be	AUX
alces-23	102	3	highly	highly	ADV
alces-23	102	4	unlikely	unlikely	ADJ
alces-23	102	5	that	that	SCONJ
alces-23	102	6	either	either	CCONJ
alces-23	102	7	population	population	NOUN
alces-23	102	8	experiences	experience	VERB
alces-23	102	9	gene	gene	NOUN
alces-23	102	10	flow	flow	NOUN
alces-23	102	11	with	with	ADP
alces-23	102	12	neighboring	neighboring	NOUN
alces-23	102	13	populations	population	NOUN
alces-23	102	14	;	;	PUNCT
alces-23	102	15	otherwise	otherwise	ADV
alces-23	102	16	,	,	PUNCT
alces-23	102	17	the	the	DET
alces-23	102	18	level	level	NOUN
alces-23	102	19	of	of	ADP
alces-23	102	20	diversity	diversity	NOUN
alces-23	102	21	in	in	ADP
alces-23	102	22	the	the	DET
alces-23	102	23	introduced	introduce	VERB
alces-23	102	24	populations	population	NOUN
alces-23	102	25	would	would	AUX
alces-23	102	26	be	be	AUX
alces-23	102	27	greater	great	ADJ
alces-23	102	28	and	and	CCONJ
alces-23	102	29	more	more	ADV
alces-23	102	30	similar	similar	ADJ
alces-23	102	31	with	with	ADP
alces-23	102	32	that	that	PRON
alces-23	102	33	of	of	ADP
alces-23	102	34	yukon	yukon	PROPN
alces-23	102	35	flats	flat	NOUN
alces-23	102	36	.	.	PUNCT
alces-23	103	1	managers	manager	NOUN
alces-23	103	2	of	of	ADP
alces-23	103	3	the	the	DET
alces-23	103	4	kalgin	kalgin	NOUN
alces-23	103	5	island	island	NOUN
alces-23	103	6	and	and	CCONJ
alces-23	103	7	berners	berners	PROPN
alces-23	103	8	bay	bay	NOUN
alces-23	103	9	populations	population	NOUN
alces-23	103	10	should	should	AUX
alces-23	103	11	consider	consider	VERB
alces-23	103	12	the	the	DET
alces-23	103	13	degree	degree	NOUN
alces-23	103	14	of	of	ADP
alces-23	103	15	isolation	isolation	NOUN
alces-23	103	16	and	and	CCONJ
alces-23	103	17	paucity	paucity	NOUN
alces-23	103	18	of	of	ADP
alces-23	103	19	genetic	genetic	ADJ
alces-23	103	20	variation	variation	NOUN
alces-23	103	21	in	in	ADP
alces-23	103	22	those	those	DET
alces-23	103	23	populations	population	NOUN
alces-23	103	24	when	when	SCONJ
alces-23	103	25	assessing	assess	VERB
alces-23	103	26	risks	risk	NOUN
alces-23	103	27	to	to	ADP
alces-23	103	28	population	population	NOUN
alces-23	103	29	viability	viability	NOUN
alces-23	103	30	and	and	CCONJ
alces-23	103	31	crafting	craft	VERB
alces-23	103	32	management	management	NOUN
alces-23	103	33	strategies	strategy	NOUN
alces-23	103	34	.	.	PUNCT
alces-23	104	1	as	as	ADP
alces-23	104	2	an	an	DET
alces-23	104	3	example	example	NOUN
alces-23	104	4	,	,	PUNCT
alces-23	104	5	recovery	recovery	NOUN
alces-23	104	6	of	of	ADP
alces-23	104	7	diversity	diversity	NOUN
alces-23	104	8	can	can	AUX
alces-23	104	9	probably	probably	ADV
alces-23	104	10	be	be	AUX
alces-23	104	11	accomplished	accomplish	VERB
alces-23	104	12	only	only	ADV
alces-23	104	13	with	with	ADP
alces-23	104	14	introduction	introduction	NOUN
alces-23	104	15	of	of	ADP
alces-23	104	16	additional	additional	ADJ
alces-23	104	17	individuals	individual	NOUN
alces-23	104	18	rather	rather	ADV
alces-23	104	19	than	than	ADP
alces-23	104	20	relying	rely	VERB
alces-23	104	21	on	on	ADP
alces-23	104	22	immigration	immigration	NOUN
alces-23	104	23	from	from	ADP
alces-23	104	24	surrounding	surround	VERB
alces-23	104	25	populations	population	NOUN
alces-23	104	26	.	.	PUNCT
alces-23	105	1	acknowledgements	acknowledgement	NOUN
alces-23	105	2	funding	funding	NOUN
alces-23	105	3	was	be	AUX
alces-23	105	4	provided	provide	VERB
alces-23	105	5	by	by	ADP
alces-23	105	6	a	a	DET
alces-23	105	7	grant	grant	NOUN
alces-23	105	8	from	from	ADP
alces-23	105	9	federal	federal	ADJ
alces-23	105	10	aid	aid	NOUN
alces-23	105	11	in	in	ADP
alces-23	105	12	wildlife	wildlife	NOUN
alces-23	105	13	restoration	restoration	NOUN
alces-23	105	14	to	to	ADP
alces-23	105	15	the	the	DET
alces-23	105	16	alaska	alaska	PROPN
alces-23	105	17	department	department	PROPN
alces-23	105	18	of	of	ADP
alces-23	105	19	fish	fish	NOUN
alces-23	105	20	and	and	CCONJ
alces-23	105	21	game	game	NOUN
alces-23	105	22	.	.	PUNCT
alces-23	106	1	i	i	PRON
alces-23	106	2	thank	thank	VERB
alces-23	106	3	n.	n.	NOUN
alces-23	106	4	barten	barten	PROPN
alces-23	106	5	and	and	CCONJ
alces-23	106	6	k.	k.	PROPN
alces-23	106	7	white	white	PROPN
alces-23	106	8	for	for	ADP
alces-23	106	9	providing	provide	VERB
alces-23	106	10	samples	sample	NOUN
alces-23	106	11	of	of	ADP
alces-23	106	12	berners	berner	NOUN
alces-23	106	13	bay	bay	PROPN
alces-23	106	14	moose	moose	NOUN
alces-23	106	15	.	.	PUNCT
alces-23	107	1	references	reference	NOUN
alces-23	107	2	barten	barten	VERB
alces-23	107	3	,	,	PUNCT
alces-23	107	4	n.	n.	PROPN
alces-23	107	5	l.	l.	PROPN
alces-23	107	6	2008	2008	NUM
alces-23	107	7	.	.	PUNCT
alces-23	108	1	unit	unit	NOUN
alces-23	108	2	1c	1c	NUM
alces-23	108	3	moose	moose	NOUN
alces-23	108	4	.	.	PUNCT
alces-23	109	1	pages	page	NOUN
alces-23	109	2	27	27	NUM
alces-23	109	3	-	-	SYM
alces-23	109	4	52	52	NUM
alces-23	109	5	in	in	ADP
alces-23	109	6	p.	p.	PROPN
alces-23	109	7	harper	harper	PROPN
alces-23	109	8	,	,	PUNCT
alces-23	109	9	editor	editor	NOUN
alces-23	109	10	.	.	PUNCT
alces-23	110	1	moose	moose	PROPN
alces-23	110	2	management	management	PROPN
alces-23	110	3	report	report	NOUN
alces-23	110	4	of	of	ADP
alces-23	110	5	survey	survey	NOUN
alces-23	110	6	-	-	PUNCT
alces-23	110	7	inventory	inventory	NOUN
alces-23	110	8	activities	activity	NOUN
alces-23	110	9	1	1	NUM
alces-23	110	10	july	july	PROPN
alces-23	110	11	2005	2005	NUM
alces-23	110	12	-	-	SYM
alces-23	110	13	30	30	NUM
alces-23	110	14	june	june	PROPN
alces-23	110	15	2007	2007	NUM
alces-23	110	16	.	.	PUNCT
alces-23	111	1	alaska	alaska	PROPN
alces-23	111	2	department	department	PROPN
alces-23	111	3	of	of	ADP
alces-23	111	4	fish	fish	NOUN
alces-23	111	5	and	and	CCONJ
alces-23	111	6	game	game	NOUN
alces-23	111	7	,	,	PUNCT
alces-23	111	8	juneau	juneau	PROPN
alces-23	111	9	,	,	PUNCT
alces-23	111	10	alaska	alaska	PROPN
alces-23	111	11	,	,	PUNCT
alces-23	111	12	usa	usa	PROPN
alces-23	111	13	.	.	PROPN
alces-23	111	14	bertram	bertram	PROPN
alces-23	111	15	,	,	PUNCT
alces-23	111	16	m.	m.	NOUN
alces-23	111	17	,	,	PUNCT
alces-23	111	18	and	and	CCONJ
alces-23	111	19	m.	m.	NOUN
alces-23	111	20	vivion	vivion	NOUN
alces-23	111	21	.	.	PUNCT
alces-23	112	1	2002	2002	NUM
alces-23	112	2	.	.	PUNCT
alces-23	113	1	moose	moose	NOUN
alces-23	113	2	mortality	mortality	NOUN
alces-23	113	3	in	in	ADP
alces-23	113	4	eastern	eastern	ADJ
alces-23	113	5	interior	interior	PROPN
alces-23	113	6	alaska	alaska	PROPN
alces-23	113	7	.	.	PUNCT
alces-23	114	1	journal	journal	PROPN
alces-23	114	2	of	of	ADP
alces-23	114	3	wildlife	wildlife	NOUN
alces-23	114	4	management	management	NOUN
alces-23	114	5	66	66	NUM
alces-23	114	6	:	:	PUNCT
alces-23	114	7	747	747	NUM
alces-23	114	8	-	-	SYM
alces-23	114	9	756	756	NUM
alces-23	114	10	.	.	PUNCT
alces-23	115	1	bishop	bishop	PROPN
alces-23	115	2	m.	m.	PROPN
alces-23	115	3	d.	d.	PROPN
alces-23	115	4	,	,	PUNCT
alces-23	115	5	s.	s.	PROPN
alces-23	115	6	m.	m.	PROPN
alces-23	115	7	kappes	kappes	PROPN
alces-23	115	8	,	,	PUNCT
alces-23	115	9	j.	j.	PROPN
alces-23	115	10	w.	w.	PROPN
alces-23	115	11	keele	keele	PROPN
alces-23	115	12	,	,	PUNCT
alces-23	115	13	r.	r.	PROPN
alces-23	115	14	t.	t.	PROPN
alces-23	115	15	stone	stone	PROPN
alces-23	115	16	,	,	PUNCT
alces-23	115	17	s.	s.	PROPN
alces-23	115	18	l.	l.	PROPN
alces-23	115	19	f.	f.	PROPN
alces-23	115	20	sunden	sunden	PROPN
alces-23	115	21	,	,	PUNCT
alces-23	115	22	g.	g.	PROPN
alces-23	115	23	a.	a.	PROPN
alces-23	115	24	hawkins	hawkins	PROPN
alces-23	115	25	,	,	PUNCT
alces-23	115	26	s.	s.	PROPN
alces-23	115	27	solinas	solinas	PROPN
alces-23	115	28	-	-	PUNCT
alces-23	115	29	toledo	toledo	PROPN
alces-23	115	30	,	,	PUNCT
alces-23	115	31	r.	r.	PROPN
alces-23	115	32	fries	fries	PROPN
alces-23	115	33	,	,	PUNCT
alces-23	115	34	m.	m.	PROPN
alces-23	115	35	d.	d.	PROPN
alces-23	115	36	gross	gross	PROPN
alces-23	115	37	,	,	PUNCT
alces-23	115	38	j.	j.	PROPN
alces-23	115	39	yoo	yoo	PROPN
alces-23	115	40	,	,	PUNCT
alces-23	115	41	and	and	CCONJ
alces-23	115	42	c.	c.	PROPN
alces-23	115	43	w.	w.	PROPN
alces-23	115	44	beattie	beattie	PROPN
alces-23	115	45	.	.	PUNCT
alces-23	116	1	1994	1994	NUM
alces-23	116	2	.	.	PUNCT
alces-23	117	1	a	a	DET
alces-23	117	2	genetic	genetic	ADJ
alces-23	117	3	linkage	linkage	NOUN
alces-23	117	4	map	map	NOUN
alces-23	117	5	for	for	ADP
alces-23	117	6	cattle	cattle	NOUN
alces-23	117	7	.	.	PUNCT
alces-23	118	1	genetics	genetic	NOUN
alces-23	118	2	136	136	NUM
alces-23	118	3	:	:	PUNCT
alces-23	118	4	619	619	NUM
alces-23	118	5	-	-	SYM
alces-23	118	6	639	639	NUM
alces-23	118	7	.	.	PUNCT
alces-23	119	1	bowyer	bowyer	PROPN
alces-23	119	2	,	,	PUNCT
alces-23	119	3	r.	r.	PROPN
alces-23	119	4	t.	t.	PROPN
alces-23	119	5	,	,	PUNCT
alces-23	119	6	m.	m.	PROPN
alces-23	119	7	c.	c.	PROPN
alces-23	119	8	nicholson	nicholson	PROPN
alces-23	119	9	,	,	PUNCT
alces-23	119	10	e.	e.	PROPN
alces-23	119	11	m.	m.	PROPN
alces-23	119	12	molvar	molvar	PROPN
alces-23	119	13	,	,	PUNCT
alces-23	119	14	and	and	CCONJ
alces-23	119	15	j.	j.	PROPN
alces-23	119	16	b.	b.	PROPN
alces-23	119	17	faro	faro	PROPN
alces-23	119	18	.	.	PUNCT
alces-23	120	1	1999	1999	NUM
alces-23	120	2	.	.	PUNCT
alces-23	121	1	moose	moose	PROPN
alces-23	121	2	on	on	ADP
alces-23	121	3	kalgin	kalgin	PROPN
alces-23	121	4	island	island	NOUN
alces-23	121	5	:	:	PUNCT
alces-23	121	6	are	be	AUX
alces-23	121	7	density	density	NOUN
alces-23	121	8	-	-	PUNCT
alces-23	121	9	dependent	dependent	ADJ
alces-23	121	10	processes	process	NOUN
alces-23	121	11	related	relate	VERB
alces-23	121	12	to	to	PART
alces-23	121	13	harvest	harvest	VERB
alces-23	121	14	?	?	PUNCT
alces-23	122	1	alces	alce	NOUN
alces-23	122	2	35	35	NUM
alces-23	122	3	:	:	PUNCT
alces-23	122	4	73	73	NUM
alces-23	122	5	-	-	SYM
alces-23	122	6	89	89	NUM
alces-23	122	7	.	.	PUNCT
alces-23	123	1	broders	broder	NOUN
alces-23	123	2	,	,	PUNCT
alces-23	123	3	h.	h.	PROPN
alces-23	123	4	g.	g.	PROPN
alces-23	123	5	,	,	PUNCT
alces-23	123	6	s.	s.	PROPN
alces-23	123	7	p.	p.	PROPN
alces-23	123	8	mahoney	mahoney	PROPN
alces-23	123	9	,	,	PUNCT
alces-23	123	10	w.	w.	PROPN
alces-23	123	11	a.	a.	PROPN
alces-23	123	12	montevecchi	montevecchi	PROPN
alces-23	123	13	,	,	PUNCT
alces-23	123	14	and	and	CCONJ
alces-23	123	15	w.	w.	PROPN
alces-23	123	16	s.	s.	PROPN
alces-23	123	17	davidson	davidson	PROPN
alces-23	123	18	.	.	PUNCT
alces-23	123	19	1999	1999	NUM
alces-23	123	20	.	.	PUNCT
alces-23	124	1	population	population	NOUN
alces-23	124	2	genetic	genetic	ADJ
alces-23	124	3	structure	structure	NOUN
alces-23	124	4	and	and	CCONJ
alces-23	124	5	the	the	DET
alces-23	124	6	effect	effect	NOUN
alces-23	124	7	of	of	ADP
alces-23	124	8	founder	founder	NOUN
alces-23	124	9	events	event	NOUN
alces-23	124	10	on	on	ADP
alces-23	124	11	the	the	DET
alces-23	124	12	genetic	genetic	ADJ
alces-23	124	13	variability	variability	NOUN
alces-23	124	14	of	of	ADP
alces-23	124	15	moose	moose	NOUN
alces-23	124	16	,	,	PUNCT
alces-23	124	17	alces	alce	NOUN
alces-23	124	18	alces	alce	NOUN
alces-23	124	19	,	,	PUNCT
alces-23	124	20	in	in	ADP
alces-23	124	21	canada	canada	PROPN
alces-23	124	22	.	.	PUNCT
alces-23	125	1	molecular	molecular	ADJ
alces-23	125	2	ecology	ecology	NOUN
alces-23	125	3	8	8	NUM
alces-23	125	4	:	:	PUNCT
alces-23	125	5	1309	1309	NUM
alces-23	125	6	-	-	SYM
alces-23	125	7	1315	1315	NUM
alces-23	125	8	.	.	PUNCT
alces-23	126	1	buchanan	buchanan	PROPN
alces-23	126	2	f.	f.	PROPN
alces-23	126	3	c.	c.	PROPN
alces-23	126	4	,	,	PUNCT
alces-23	126	5	and	and	CCONJ
alces-23	126	6	a.	a.	NOUN
alces-23	126	7	m.	m.	PROPN
alces-23	126	8	crawford	crawford	PROPN
alces-23	126	9	.	.	PUNCT
alces-23	126	10	1993	1993	NUM
alces-23	126	11	.	.	PUNCT
alces-23	127	1	ovine	ovine	ADJ
alces-23	127	2	microsatellites	microsatellite	NOUN
alces-23	127	3	at	at	ADP
alces-23	127	4	the	the	DET
alces-23	127	5	oarfcb11	oarfcb11	NOUN
alces-23	127	6	,	,	PUNCT
alces-23	127	7	oarfcb128	oarfcb128	PROPN
alces-23	127	8	,	,	PUNCT
alces-23	127	9	oarfcb193	oarfcb193	PROPN
alces-23	127	10	,	,	PUNCT
alces-23	127	11	oarfcb266	oarfcb266	PROPN
alces-23	127	12	and	and	CCONJ
alces-23	127	13	oarfcb304	oarfcb304	NUM
alces-23	127	14	loci	locus	NOUN
alces-23	127	15	.	.	PUNCT
alces-23	128	1	animal	animal	NOUN
alces-23	128	2	genetics	genetic	NOUN
alces-23	128	3	24	24	NUM
alces-23	128	4	:	:	SYM
alces-23	128	5	145	145	NUM
alces-23	128	6	.	.	PUNCT
alces-23	129	1	burris	burris	PROPN
alces-23	129	2	,	,	PUNCT
alces-23	129	3	o.	o.	PROPN
alces-23	129	4	e.	e.	PROPN
alces-23	129	5	,	,	PUNCT
alces-23	129	6	and	and	CCONJ
alces-23	129	7	d.	d.	PROPN
alces-23	129	8	e.	e.	PROPN
alces-23	129	9	mcknight	mcknight	PROPN
alces-23	129	10	.	.	PUNCT
alces-23	130	1	1973	1973	NUM
alces-23	130	2	.	.	PUNCT
alces-23	131	1	game	game	NOUN
alces-23	131	2	transplants	transplant	NOUN
alces-23	131	3	in	in	ADP
alces-23	131	4	alaska	alaska	PROPN
alces-23	131	5	.	.	PUNCT
alces-23	132	1	game	game	PROPN
alces-23	132	2	technical	technical	ADJ
alces-23	132	3	bulletin	bulletin	PROPN
alces-23	132	4	4	4	NUM
alces-23	132	5	.	.	PUNCT
alces-23	133	1	alaska	alaska	PROPN
alces-23	133	2	department	department	PROPN
alces-23	133	3	of	of	ADP
alces-23	133	4	fish	fish	NOUN
alces-23	133	5	and	and	CCONJ
alces-23	133	6	game	game	NOUN
alces-23	133	7	,	,	PUNCT
alces-23	133	8	juneau	juneau	PROPN
alces-23	133	9	,	,	PUNCT
alces-23	133	10	alaska	alaska	PROPN
alces-23	133	11	,	,	PUNCT
alces-23	133	12	usa	usa	PROPN
alces-23	133	13	.	.	PROPN
alces-23	133	14	caikoski	caikoski	PROPN
alces-23	133	15	,	,	PUNCT
alces-23	133	16	j.	j.	PROPN
alces-23	133	17	r.	r.	PROPN
alces-23	133	18	2008	2008	NUM
alces-23	133	19	.	.	PUNCT
alces-23	134	1	units	unit	NOUN
alces-23	134	2	25a	25a	NOUN
alces-23	134	3	,	,	PUNCT
alces-23	134	4	25b	25b	NOUN
alces-23	134	5	,	,	PUNCT
alces-23	134	6	and	and	CCONJ
alces-23	134	7	25d	25d	NUM
alces-23	134	8	moose	moose	NOUN
alces-23	134	9	.	.	PUNCT
alces-23	135	1	pages	page	NOUN
alces-23	135	2	617	617	NUM
alces-23	135	3	-	-	SYM
alces-23	135	4	647	647	NUM
alces-23	135	5	in	in	ADP
alces-23	135	6	p.	p.	PROPN
alces-23	135	7	harper	harper	PROPN
alces-23	135	8	,	,	PUNCT
alces-23	135	9	editor	editor	NOUN
alces-23	135	10	.	.	PUNCT
alces-23	136	1	moose	moose	PROPN
alces-23	136	2	management	management	PROPN
alces-23	136	3	report	report	NOUN
alces-23	136	4	of	of	ADP
alces-23	136	5	survey	survey	NOUN
alces-23	136	6	-	-	PUNCT
alces-23	136	7	inventory	inventory	NOUN
alces-23	136	8	activities	activity	NOUN
alces-23	136	9	1	1	NUM
alces-23	136	10	july	july	PROPN
alces-23	136	11	2005	2005	NUM
alces-23	136	12	-	-	SYM
alces-23	136	13	30	30	NUM
alces-23	136	14	june	june	PROPN
alces-23	136	15	2007	2007	NUM
alces-23	136	16	.	.	PUNCT
alces-23	137	1	alaska	alaska	PROPN
alces-23	137	2	department	department	PROPN
alces-23	137	3	of	of	ADP
alces-23	137	4	fish	fish	NOUN
alces-23	137	5	and	and	CCONJ
alces-23	137	6	game	game	NOUN
alces-23	137	7	,	,	PUNCT
alces-23	137	8	juneau	juneau	PROPN
alces-23	137	9	,	,	PUNCT
alces-23	137	10	alaska	alaska	PROPN
alces-23	137	11	,	,	PUNCT
alces-23	137	12	usa	usa	PROPN
alces-23	137	13	.	.	PROPN
alces-23	137	14	coltman	coltman	PROPN
alces-23	137	15	,	,	PUNCT
alces-23	137	16	d.	d.	PROPN
alces-23	137	17	w.	w.	PROPN
alces-23	137	18	,	,	PUNCT
alces-23	137	19	j.	j.	PROPN
alces-23	137	20	g.	g.	PROPN
alces-23	137	21	pilkington	pilkington	PROPN
alces-23	137	22	,	,	PUNCT
alces-23	137	23	j.	j.	PROPN
alces-23	137	24	a.	a.	PROPN
alces-23	137	25	smith	smith	PROPN
alces-23	137	26	,	,	PUNCT
alces-23	137	27	and	and	CCONJ
alces-23	137	28	j.	j.	PROPN
alces-23	137	29	m.	m.	PROPN
alces-23	137	30	pemberton	pemberton	PROPN
alces-23	137	31	.	.	PUNCT
alces-23	138	1	1999	1999	NUM
alces-23	138	2	.	.	PUNCT
alces-23	139	1	parasite	parasite	NOUN
alces-23	139	2	-	-	PUNCT
alces-23	139	3	mediated	mediate	VERB
alces-23	139	4	selection	selection	NOUN
alces-23	139	5	against	against	ADP
alces-23	139	6	inbred	inbreed	VERB
alces-23	139	7	soay	soay	PROPN
alces-23	139	8	sheep	sheep	NOUN
alces-23	139	9	in	in	ADP
alces-23	139	10	a	a	DET
alces-23	139	11	free	free	ADJ
alces-23	139	12	-	-	PUNCT
alces-23	139	13	living	live	VERB
alces-23	139	14	,	,	PUNCT
alces-23	139	15	island	island	NOUN
alces-23	139	16	population	population	NOUN
alces-23	139	17	.	.	PUNCT
alces-23	140	1	evolugenetic	evolugenetic	ADJ
alces-23	140	2	diversity	diversity	NOUN
alces-23	140	3	in	in	ADP
alces-23	140	4	introduced	introduce	VERB
alces-23	140	5	populations	population	NOUN
alces-23	140	6	hundertmark	hundertmark	NOUN
alces-23	140	7	alces	alce	NOUN
alces-23	140	8	vol	vol	NOUN
alces-23	140	9	.	.	PROPN
alces-23	141	1	45	45	NUM
alces-23	141	2	,	,	PUNCT
alces-23	141	3	2009	2009	NUM
alces-23	141	4	142	142	NUM
alces-23	141	5	tion	tion	NOUN
alces-23	141	6	53	53	NUM
alces-23	141	7	:	:	SYM
alces-23	141	8	1259	1259	NUM
alces-23	141	9	-	-	SYM
alces-23	141	10	1267	1267	NUM
alces-23	141	11	.	.	PUNCT
alces-23	142	1	coulson	coulson	PROPN
alces-23	142	2	,	,	PUNCT
alces-23	142	3	t.	t.	PROPN
alces-23	142	4	,	,	PUNCT
alces-23	142	5	s.	s.	PROPN
alces-23	142	6	albon	albon	PROPN
alces-23	142	7	,	,	PUNCT
alces-23	142	8	j.	j.	PROPN
alces-23	142	9	slate	slate	PROPN
alces-23	142	10	,	,	PUNCT
alces-23	142	11	and	and	CCONJ
alces-23	142	12	j.	j.	PROPN
alces-23	142	13	pemberton	pemberton	PROPN
alces-23	142	14	.	.	PUNCT
alces-23	143	1	1999	1999	NUM
alces-23	143	2	.	.	PUNCT
alces-23	144	1	microsatellite	microsatellite	ADJ
alces-23	144	2	loci	locus	NOUN
alces-23	144	3	reveal	reveal	VERB
alces-23	144	4	sex	sex	NOUN
alces-23	144	5	-	-	PUNCT
alces-23	144	6	dependent	dependent	ADJ
alces-23	144	7	responses	response	NOUN
alces-23	144	8	to	to	ADP
alces-23	144	9	inbreeding	inbreede	VERB
alces-23	144	10	and	and	CCONJ
alces-23	144	11	outbreeding	outbreede	VERB
alces-23	144	12	in	in	ADP
alces-23	144	13	red	red	ADJ
alces-23	144	14	deer	deer	NOUN
alces-23	144	15	calves	calf	NOUN
alces-23	144	16	.	.	PUNCT
alces-23	145	1	evolution	evolution	NOUN
alces-23	145	2	53	53	NUM
alces-23	145	3	:	:	SYM
alces-23	145	4	1951	1951	NUM
alces-23	145	5	-	-	SYM
alces-23	145	6	1960	1960	NUM
alces-23	145	7	.	.	PUNCT
alces-23	146	1	fisher	fisher	PROPN
alces-23	146	2	,	,	PUNCT
alces-23	146	3	r.	r.	PROPN
alces-23	146	4	a.	a.	PROPN
alces-23	146	5	1948	1948	NUM
alces-23	146	6	.	.	PUNCT
alces-23	147	1	combining	combine	VERB
alces-23	147	2	independent	independent	ADJ
alces-23	147	3	tests	test	NOUN
alces-23	147	4	of	of	ADP
alces-23	147	5	significance	significance	NOUN
alces-23	147	6	.	.	PUNCT
alces-23	148	1	american	american	PROPN
alces-23	148	2	statistician	statistician	PROPN
alces-23	148	3	2	2	NUM
alces-23	148	4	:	:	SYM
alces-23	148	5	30	30	NUM
alces-23	148	6	.	.	PUNCT
alces-23	148	7	goudet	goudet	NOUN
alces-23	148	8	,	,	PUNCT
alces-23	148	9	j.	j.	PROPN
alces-23	148	10	1995	1995	NUM
alces-23	148	11	.	.	PUNCT
alces-23	149	1	fstat	fstat	PROPN
alces-23	149	2	(	(	PUNCT
alces-23	149	3	version	version	NOUN
alces-23	149	4	1.2	1.2	NUM
alces-23	149	5	):	):	PUNCT
alces-23	149	6	a	a	DET
alces-23	149	7	computer	computer	NOUN
alces-23	149	8	program	program	NOUN
alces-23	149	9	to	to	PART
alces-23	149	10	calculate	calculate	VERB
alces-23	149	11	f	f	PROPN
alces-23	149	12	-	-	PUNCT
alces-23	149	13	statisitcs	statisitcs	PROPN
alces-23	149	14	.	.	PUNCT
alces-23	150	1	journal	journal	PROPN
alces-23	150	2	of	of	ADP
alces-23	150	3	heredity	heredity	NOUN
alces-23	150	4	86	86	NUM
alces-23	150	5	:	:	PUNCT
alces-23	150	6	485	485	NUM
alces-23	150	7	-	-	SYM
alces-23	150	8	486	486	NUM
alces-23	150	9	.	.	PUNCT
alces-23	151	1	hundertmark	hundertmark	PROPN
alces-23	151	2	,	,	PUNCT
alces-23	151	3	k.	k.	PROPN
alces-23	151	4	j.	j.	PROPN
alces-23	151	5	,	,	PUNCT
alces-23	151	6	w.	w.	PROPN
alces-23	151	7	l.	l.	PROPN
alces-23	151	8	eberhardt	eberhardt	PROPN
alces-23	151	9	,	,	PUNCT
alces-23	151	10	and	and	CCONJ
alces-23	151	11	r.	r.	PROPN
alces-23	151	12	e.	e.	PROPN
alces-23	151	13	ball	ball	PROPN
alces-23	151	14	.	.	PUNCT
alces-23	152	1	1990	1990	NUM
alces-23	152	2	.	.	PUNCT
alces-23	153	1	winter	winter	NOUN
alces-23	153	2	habitat	habitat	NOUN
alces-23	153	3	use	use	NOUN
alces-23	153	4	by	by	ADP
alces-23	153	5	moose	moose	NOUN
alces-23	153	6	in	in	ADP
alces-23	153	7	southeastern	southeastern	ADJ
alces-23	153	8	alaska	alaska	PROPN
alces-23	153	9	:	:	PUNCT
alces-23	153	10	implications	implication	NOUN
alces-23	153	11	for	for	ADP
alces-23	153	12	forest	forest	NOUN
alces-23	153	13	management	management	NOUN
alces-23	153	14	.	.	PUNCT
alces-23	154	1	alces	alce	NOUN
alces-23	154	2	26	26	NUM
alces-23	154	3	:	:	PUNCT
alces-23	154	4	108	108	NUM
alces-23	154	5	-	-	SYM
alces-23	154	6	114	114	NUM
alces-23	154	7	.	.	PUNCT
alces-23	155	1	kalinowski	kalinowski	PROPN
alces-23	155	2	,	,	PUNCT
alces-23	155	3	s.	s.	PROPN
alces-23	155	4	t.	t.	PROPN
alces-23	155	5	2005	2005	NUM
alces-23	155	6	.	.	PUNCT
alces-23	156	1	hp	hp	VERB
alces-23	156	2	-	-	PUNCT
alces-23	156	3	rare	rare	ADJ
alces-23	156	4	1.0	1.0	NUM
alces-23	156	5	:	:	PUNCT
alces-23	156	6	a	a	DET
alces-23	156	7	computer	computer	NOUN
alces-23	156	8	program	program	NOUN
alces-23	156	9	for	for	ADP
alces-23	156	10	performing	perform	VERB
alces-23	156	11	rarefaction	rarefaction	NOUN
alces-23	156	12	on	on	ADP
alces-23	156	13	measures	measure	NOUN
alces-23	156	14	of	of	ADP
alces-23	156	15	allelic	allelic	ADJ
alces-23	156	16	richness	richness	NOUN
alces-23	156	17	.	.	PUNCT
alces-23	157	1	molecular	molecular	ADJ
alces-23	157	2	ecology	ecology	NOUN
alces-23	157	3	notes	note	VERB
alces-23	157	4	5	5	NUM
alces-23	157	5	:	:	SYM
alces-23	157	6	187	187	NUM
alces-23	157	7	-	-	SYM
alces-23	157	8	189	189	NUM
alces-23	157	9	.	.	PUNCT
alces-23	158	1	mainguy	mainguy	NOUN
alces-23	158	2	,	,	PUNCT
alces-23	158	3	j.	j.	PROPN
alces-23	158	4	,	,	PUNCT
alces-23	158	5	s.	s.	PROPN
alces-23	158	6	d.	d.	PROPN
alces-23	158	7	côté	côté	PROPN
alces-23	158	8	,	,	PUNCT
alces-23	158	9	and	and	CCONJ
alces-23	158	10	d.	d.	PROPN
alces-23	158	11	w.	w.	PROPN
alces-23	158	12	coltman	coltman	PROPN
alces-23	158	13	.	.	PUNCT
alces-23	159	1	2009	2009	NUM
alces-23	159	2	.	.	PUNCT
alces-23	160	1	multilocus	multilocus	NOUN
alces-23	160	2	heterozygosity	heterozygosity	NOUN
alces-23	160	3	,	,	PUNCT
alces-23	160	4	parental	parental	ADJ
alces-23	160	5	relatedness	relatedness	NOUN
alces-23	160	6	and	and	CCONJ
alces-23	160	7	individual	individual	ADJ
alces-23	160	8	fitness	fitness	NOUN
alces-23	160	9	components	component	NOUN
alces-23	160	10	in	in	ADP
alces-23	160	11	a	a	DET
alces-23	160	12	wild	wild	ADJ
alces-23	160	13	mountain	mountain	NOUN
alces-23	160	14	goat	goat	NOUN
alces-23	160	15	,	,	PUNCT
alces-23	160	16	oreamnus	oreamnus	ADJ
alces-23	160	17	americanus	americanus	NOUN
alces-23	160	18	population	population	NOUN
alces-23	160	19	.	.	PUNCT
alces-23	161	1	molecular	molecular	ADJ
alces-23	161	2	ecology	ecology	NOUN
alces-23	161	3	18	18	NUM
alces-23	161	4	:	:	SYM
alces-23	161	5	2297	2297	NUM
alces-23	161	6	-	-	SYM
alces-23	161	7	2306	2306	NUM
alces-23	161	8	.	.	PUNCT
alces-23	162	1	nei	nei	PROPN
alces-23	162	2	,	,	PUNCT
alces-23	162	3	m.	m.	NOUN
alces-23	162	4	1978	1978	NUM
alces-23	162	5	.	.	PUNCT
alces-23	163	1	estimation	estimation	NOUN
alces-23	163	2	of	of	ADP
alces-23	163	3	heterozygosity	heterozygosity	NOUN
alces-23	163	4	and	and	CCONJ
alces-23	163	5	genetic	genetic	ADJ
alces-23	163	6	distance	distance	NOUN
alces-23	163	7	from	from	ADP
alces-23	163	8	a	a	DET
alces-23	163	9	small	small	ADJ
alces-23	163	10	number	number	NOUN
alces-23	163	11	of	of	ADP
alces-23	163	12	individuals	individual	NOUN
alces-23	163	13	.	.	PUNCT
alces-23	164	1	genetics	genetic	NOUN
alces-23	164	2	89	89	NUM
alces-23	164	3	:	:	PUNCT
alces-23	164	4	538	538	NUM
alces-23	164	5	-	-	SYM
alces-23	164	6	590	590	NUM
alces-23	164	7	.	.	PUNCT
alces-23	165	1	_	_	PUNCT
alces-23	166	1	_	_	PUNCT
alces-23	167	1	_	_	PUNCT
alces-23	168	1	_	_	PUNCT
alces-23	169	1	_	_	PUNCT
alces-23	169	2	,	,	PUNCT
alces-23	169	3	t.	t.	PROPN
alces-23	169	4	maruyama	maruyama	NOUN
alces-23	169	5	,	,	PUNCT
alces-23	169	6	and	and	CCONJ
alces-23	169	7	r.	r.	PROPN
alces-23	169	8	chakraborty	chakraborty	PROPN
alces-23	169	9	.	.	PUNCT
alces-23	170	1	1975	1975	NUM
alces-23	170	2	.	.	PUNCT
alces-23	171	1	the	the	DET
alces-23	171	2	bottleneck	bottleneck	NOUN
alces-23	171	3	effect	effect	NOUN
alces-23	171	4	and	and	CCONJ
alces-23	171	5	genetic	genetic	ADJ
alces-23	171	6	variability	variability	NOUN
alces-23	171	7	in	in	ADP
alces-23	171	8	populations	population	NOUN
alces-23	171	9	.	.	PUNCT
alces-23	172	1	evolution	evolution	NOUN
alces-23	172	2	29	29	NUM
alces-23	172	3	:	:	SYM
alces-23	172	4	1	1	NUM
alces-23	172	5	-	-	SYM
alces-23	172	6	10	10	NUM
alces-23	172	7	.	.	PUNCT
alces-23	173	1	pruett	pruett	PROPN
alces-23	173	2	,	,	PUNCT
alces-23	173	3	c.	c.	PROPN
alces-23	173	4	l.	l.	PROPN
alces-23	173	5	,	,	PUNCT
alces-23	173	6	and	and	CCONJ
alces-23	173	7	k.	k.	PROPN
alces-23	173	8	winker	winker	PROPN
alces-23	173	9	.	.	PUNCT
alces-23	173	10	2008	2008	NUM
alces-23	173	11	.	.	PUNCT
alces-23	174	1	the	the	DET
alces-23	174	2	effects	effect	NOUN
alces-23	174	3	of	of	ADP
alces-23	174	4	sample	sample	NOUN
alces-23	174	5	size	size	NOUN
alces-23	174	6	on	on	ADP
alces-23	174	7	population	population	NOUN
alces-23	174	8	genetic	genetic	ADJ
alces-23	174	9	diversity	diversity	NOUN
alces-23	174	10	estimates	estimate	NOUN
alces-23	174	11	in	in	ADP
alces-23	174	12	song	song	NOUN
alces-23	174	13	sparrows	sparrows	PROPN
alces-23	174	14	melospiza	melospiza	PROPN
alces-23	174	15	melodia	melodia	PROPN
alces-23	174	16	.	.	PROPN
alces-23	174	17	journal	journal	PROPN
alces-23	174	18	of	of	ADP
alces-23	174	19	avian	avian	ADJ
alces-23	174	20	biology	biology	NOUN
alces-23	174	21	39	39	NUM
alces-23	174	22	:	:	PUNCT
alces-23	174	23	252	252	NUM
alces-23	174	24	-	-	SYM
alces-23	174	25	256	256	NUM
alces-23	174	26	.	.	PUNCT
alces-23	175	1	raymond	raymond	PROPN
alces-23	175	2	m.	m.	PROPN
alces-23	175	3	,	,	PUNCT
alces-23	175	4	and	and	CCONJ
alces-23	175	5	f.	f.	PROPN
alces-23	175	6	rousset	rousset	PROPN
alces-23	175	7	.	.	PUNCT
alces-23	176	1	1995	1995	NUM
alces-23	176	2	.	.	PUNCT
alces-23	177	1	genepop	genepop	NOUN
alces-23	177	2	(	(	PUNCT
alces-23	177	3	version	version	NOUN
alces-23	177	4	1.2	1.2	NUM
alces-23	177	5	):	):	PUNCT
alces-23	177	6	population	population	NOUN
alces-23	177	7	genetics	genetic	NOUN
alces-23	177	8	software	software	NOUN
alces-23	177	9	for	for	ADP
alces-23	177	10	exact	exact	ADJ
alces-23	177	11	tests	test	NOUN
alces-23	177	12	and	and	CCONJ
alces-23	177	13	ecumenicism	ecumenicism	NOUN
alces-23	177	14	.	.	PUNCT
alces-23	178	1	journal	journal	NOUN
alces-23	178	2	of	of	ADP
alces-23	178	3	heredity	heredity	NOUN
alces-23	178	4	86	86	NUM
alces-23	178	5	:	:	SYM
alces-23	178	6	248	248	NUM
alces-23	178	7	-	-	SYM
alces-23	178	8	249	249	NUM
alces-23	178	9	.	.	PUNCT
alces-23	179	1	reed	reed	PROPN
alces-23	179	2	,	,	PUNCT
alces-23	179	3	d.	d.	PROPN
alces-23	179	4	h.	h.	PROPN
alces-23	179	5	,	,	PUNCT
alces-23	179	6	and	and	CCONJ
alces-23	179	7	r.	r.	PROPN
alces-23	179	8	frankham	frankham	PROPN
alces-23	179	9	.	.	PUNCT
alces-23	180	1	2003	2003	NUM
alces-23	180	2	.	.	PUNCT
alces-23	181	1	correlation	correlation	NOUN
alces-23	181	2	between	between	ADP
alces-23	181	3	fitness	fitness	NOUN
alces-23	181	4	and	and	CCONJ
alces-23	181	5	genetic	genetic	ADJ
alces-23	181	6	diversity	diversity	NOUN
alces-23	181	7	.	.	PUNCT
alces-23	182	1	conservation	conservation	NOUN
alces-23	182	2	biology	biology	NOUN
alces-23	182	3	17	17	NUM
alces-23	182	4	:	:	PUNCT
alces-23	182	5	230	230	NUM
alces-23	182	6	-	-	SYM
alces-23	182	7	237	237	NUM
alces-23	182	8	.	.	PUNCT
alces-23	183	1	schmidt	schmidt	PROPN
alces-23	183	2	,	,	PUNCT
alces-23	183	3	j.	j.	PROPN
alces-23	183	4	i.	i.	PROPN
alces-23	183	5	,	,	PUNCT
alces-23	183	6	k.	k.	PROPN
alces-23	183	7	j.	j.	PROPN
alces-23	183	8	hundertmark	hundertmark	PROPN
alces-23	183	9	,	,	PUNCT
alces-23	183	10	r.	r.	PROPN
alces-23	183	11	t.	t.	PROPN
alces-23	183	12	bowyer	bowyer	PROPN
alces-23	183	13	,	,	PUNCT
alces-23	183	14	and	and	CCONJ
alces-23	183	15	k.	k.	PROPN
alces-23	183	16	g.	g.	PROPN
alces-23	183	17	mccracken	mccracken	PROPN
alces-23	183	18	.	.	PUNCT
alces-23	183	19	2009	2009	NUM
alces-23	183	20	.	.	PUNCT
alces-23	184	1	population	population	NOUN
alces-23	184	2	structure	structure	NOUN
alces-23	184	3	and	and	CCONJ
alces-23	184	4	genetic	genetic	ADJ
alces-23	184	5	diversity	diversity	NOUN
alces-23	184	6	of	of	ADP
alces-23	184	7	moose	moose	NOUN
alces-23	184	8	in	in	ADP
alces-23	184	9	alaska	alaska	PROPN
alces-23	184	10	.	.	PUNCT
alces-23	185	1	journal	journal	PROPN
alces-23	185	2	of	of	ADP
alces-23	185	3	heredity	heredity	NOUN
alces-23	185	4	100	100	NUM
alces-23	185	5	:	:	SYM
alces-23	185	6	170	170	NUM
alces-23	185	7	-	-	SYM
alces-23	185	8	180	180	NUM
alces-23	185	9	.	.	PUNCT
alces-23	186	1	scribner	scribner	PROPN
alces-23	186	2	,	,	PUNCT
alces-23	186	3	k.	k.	PROPN
alces-23	186	4	t.	t.	PROPN
alces-23	186	5	,	,	PUNCT
alces-23	186	6	and	and	CCONJ
alces-23	186	7	m.	m.	PROPN
alces-23	186	8	h.	h.	PROPN
alces-23	186	9	smith	smith	PROPN
alces-23	186	10	.	.	PUNCT
alces-23	186	11	1990	1990	NUM
alces-23	186	12	.	.	PUNCT
alces-23	187	1	genetic	genetic	ADJ
alces-23	187	2	variability	variability	NOUN
alces-23	187	3	and	and	CCONJ
alces-23	187	4	antler	antler	PROPN
alces-23	187	5	development	development	NOUN
alces-23	187	6	.	.	PUNCT
alces-23	188	1	pages	page	NOUN
alces-23	188	2	460	460	NUM
alces-23	188	3	-	-	SYM
alces-23	188	4	473	473	NUM
alces-23	188	5	in	in	ADP
alces-23	188	6	g.	g.	PROPN
alces-23	188	7	a.	a.	NOUN
alces-23	188	8	bubenik	bubenik	PROPN
alces-23	188	9	and	and	CCONJ
alces-23	188	10	a.	a.	PROPN
alces-23	188	11	b.	b.	PROPN
alces-23	188	12	bubenik	bubenik	PROPN
alces-23	188	13	,	,	PUNCT
alces-23	188	14	editors	editor	NOUN
alces-23	188	15	.	.	PUNCT
alces-23	189	1	horns	horn	NOUN
alces-23	189	2	,	,	PUNCT
alces-23	189	3	pronghorns	pronghorn	NOUN
alces-23	189	4	and	and	CCONJ
alces-23	189	5	antlers	antler	NOUN
alces-23	189	6	.	.	PUNCT
alces-23	190	1	springer	springer	NOUN
alces-23	190	2	-	-	PUNCT
alces-23	190	3	verlag	verlag	PROPN
alces-23	190	4	,	,	PUNCT
alces-23	190	5	new	new	PROPN
alces-23	190	6	york	york	PROPN
alces-23	190	7	,	,	PUNCT
alces-23	190	8	usa	usa	PROPN
alces-23	190	9	.	.	PROPN
alces-23	190	10	silva	silva	PROPN
alces-23	190	11	,	,	PUNCT
alces-23	190	12	a.	a.	PROPN
alces-23	190	13	d.	d.	PROPN
alces-23	190	14	,	,	PUNCT
alces-23	190	15	j.-m	j.-m	PROPN
alces-23	190	16	.	.	PUNCT
alces-23	191	1	gaillard	gaillard	PROPN
alces-23	191	2	,	,	PUNCT
alces-23	191	3	n.	n.	PROPN
alces-23	191	4	g.	g.	PROPN
alces-23	191	5	yoccoz	yoccoz	PROPN
alces-23	191	6	,	,	PUNCT
alces-23	191	7	a.	a.	PROPN
alces-23	191	8	j.	j.	PROPN
alces-23	191	9	m.	m.	PROPN
alces-23	191	10	hewison	hewison	PROPN
alces-23	191	11	,	,	PUNCT
alces-23	191	12	m.	m.	NOUN
alces-23	191	13	galan	galan	PROPN
alces-23	191	14	,	,	PUNCT
alces-23	191	15	t.	t.	PROPN
alces-23	191	16	coulson	coulson	PROPN
alces-23	191	17	,	,	PUNCT
alces-23	191	18	d.	d.	PROPN
alces-23	191	19	allaine	allaine	PROPN
alces-23	191	20	,	,	PUNCT
alces-23	191	21	l.	l.	PROPN
alces-23	191	22	vial	vial	PROPN
alces-23	191	23	,	,	PUNCT
alces-23	191	24	d.	d.	PROPN
alces-23	191	25	delorme	delorme	PROPN
alces-23	191	26	,	,	PUNCT
alces-23	191	27	g.	g.	PROPN
alces-23	191	28	van	van	PROPN
alces-23	191	29	laere	laere	ADV
alces-23	191	30	,	,	PUNCT
alces-23	191	31	f.	f.	PROPN
alces-23	191	32	klein	klein	PROPN
alces-23	191	33	,	,	PUNCT
alces-23	191	34	and	and	CCONJ
alces-23	191	35	g.	g.	PROPN
alces-23	191	36	luikart	luikart	PROPN
alces-23	191	37	.	.	PUNCT
alces-23	192	1	2009	2009	NUM
alces-23	192	2	.	.	PUNCT
alces-23	193	1	heterozygosity	heterozygosity	NOUN
alces-23	193	2	-	-	PUNCT
alces-23	193	3	fitness	fitness	NOUN
alces-23	193	4	correlations	correlation	NOUN
alces-23	193	5	revealed	reveal	VERB
alces-23	193	6	by	by	ADP
alces-23	193	7	neutral	neutral	ADJ
alces-23	193	8	and	and	CCONJ
alces-23	193	9	candidate	candidate	NOUN
alces-23	193	10	gene	gene	NOUN
alces-23	193	11	markers	marker	NOUN
alces-23	193	12	in	in	ADP
alces-23	193	13	roe	roe	PROPN
alces-23	193	14	deer	deer	NOUN
alces-23	193	15	from	from	ADP
alces-23	193	16	a	a	DET
alces-23	193	17	long	long	ADJ
alces-23	193	18	-	-	PUNCT
alces-23	193	19	term	term	NOUN
alces-23	193	20	study	study	NOUN
alces-23	193	21	.	.	PUNCT
alces-23	194	1	evolution	evolution	NOUN
alces-23	194	2	63	63	NUM
alces-23	194	3	:	:	PUNCT
alces-23	194	4	403	403	NUM
alces-23	194	5	-	-	SYM
alces-23	194	6	417	417	NUM
alces-23	194	7	.	.	PUNCT
alces-23	195	1	von	von	PROPN
alces-23	195	2	hardenberg	hardenberg	PROPN
alces-23	195	3	,	,	PUNCT
alces-23	195	4	a.	a.	PROPN
alces-23	195	5	,	,	PUNCT
alces-23	195	6	b.	b.	PROPN
alces-23	195	7	bassano	bassano	PROPN
alces-23	195	8	,	,	PUNCT
alces-23	195	9	m.	m.	NOUN
alces-23	195	10	festabianchet	festabianchet	NOUN
alces-23	195	11	,	,	PUNCT
alces-23	195	12	g.	g.	PROPN
alces-23	195	13	luikart	luikart	PROPN
alces-23	195	14	,	,	PUNCT
alces-23	195	15	p.	p.	PROPN
alces-23	195	16	lanfranchi	lanfranchi	PROPN
alces-23	195	17	,	,	PUNCT
alces-23	195	18	and	and	CCONJ
alces-23	195	19	d.	d.	PROPN
alces-23	195	20	coltman	coltman	PROPN
alces-23	195	21	.	.	PUNCT
alces-23	196	1	2007	2007	NUM
alces-23	196	2	.	.	PUNCT
alces-23	197	1	age	age	NOUN
alces-23	197	2	-	-	PUNCT
alces-23	197	3	dependent	dependent	ADJ
alces-23	197	4	genetic	genetic	ADJ
alces-23	197	5	effects	effect	NOUN
alces-23	197	6	on	on	ADP
alces-23	197	7	a	a	DET
alces-23	197	8	secondary	secondary	ADJ
alces-23	197	9	sexual	sexual	ADJ
alces-23	197	10	trait	trait	NOUN
alces-23	197	11	in	in	ADP
alces-23	197	12	male	male	ADJ
alces-23	197	13	alpine	alpine	NOUN
alces-23	197	14	ibex	ibex	PROPN
alces-23	197	15	,	,	PUNCT
alces-23	197	16	capra	capra	PROPN
alces-23	197	17	ibex	ibex	PROPN
alces-23	197	18	.	.	PUNCT
alces-23	198	1	molecular	molecular	ADJ
alces-23	198	2	ecology	ecology	NOUN
alces-23	198	3	16	16	NUM
alces-23	198	4	:	:	SYM
alces-23	198	5	1969	1969	NUM
alces-23	198	6	-	-	SYM
alces-23	198	7	1980	1980	NUM
alces-23	198	8	.	.	PUNCT
alces-23	199	1	wilson	wilson	PROPN
alces-23	199	2	,	,	PUNCT
alces-23	199	3	g.	g.	PROPN
alces-23	199	4	a.	a.	PROPN
alces-23	199	5	,	,	PUNCT
alces-23	199	6	c.	c.	PROPN
alces-23	199	7	strobeck	strobeck	PROPN
alces-23	199	8	,	,	PUNCT
alces-23	199	9	l.	l.	PROPN
alces-23	199	10	wu	wu	PROPN
alces-23	199	11	,	,	PUNCT
alces-23	199	12	and	and	CCONJ
alces-23	199	13	j.	j.	PROPN
alces-23	199	14	w.	w.	PROPN
alces-23	199	15	coffin	coffin	PROPN
alces-23	199	16	.	.	PUNCT
alces-23	200	1	1997	1997	NUM
alces-23	200	2	.	.	PUNCT
alces-23	201	1	characterization	characterization	NOUN
alces-23	201	2	of	of	ADP
alces-23	201	3	microsatellite	microsatellite	ADJ
alces-23	201	4	loci	locus	NOUN
alces-23	201	5	in	in	ADP
alces-23	201	6	caribou	caribou	PROPN
alces-23	201	7	(	(	PUNCT
alces-23	201	8	rangifer	rangifer	VERB
alces-23	201	9	tarandus	tarandus	NOUN
alces-23	201	10	)	)	PUNCT
alces-23	201	11	,	,	PUNCT
alces-23	201	12	and	and	CCONJ
alces-23	201	13	their	their	PRON
alces-23	201	14	use	use	NOUN
alces-23	201	15	in	in	ADP
alces-23	201	16	other	other	ADJ
alces-23	201	17	artiodactyls	artiodactyls	NOUN
alces-23	201	18	.	.	PUNCT
alces-23	202	1	molecular	molecular	ADJ
alces-23	202	2	ecology	ecology	NOUN
alces-23	202	3	6	6	NUM
alces-23	202	4	:	:	SYM
alces-23	202	5	697	697	NUM
alces-23	202	6	-	-	SYM
alces-23	202	7	699	699	NUM
alces-23	202	8	.	.	PUNCT
alces-23	203	1	wilson	wilson	PROPN
alces-23	203	2	,	,	PUNCT
alces-23	203	3	p.	p.	PROPN
alces-23	203	4	j.	j.	PROPN
alces-23	203	5	,	,	PUNCT
alces-23	203	6	s.	s.	PROPN
alces-23	203	7	grewal	grewal	PROPN
alces-23	203	8	,	,	PUNCT
alces-23	203	9	a.	a.	NOUN
alces-23	203	10	rodgers	rodgers	PROPN
alces-23	203	11	,	,	PUNCT
alces-23	203	12	r.	r.	PROPN
alces-23	203	13	rempel	rempel	PROPN
alces-23	203	14	,	,	PUNCT
alces-23	203	15	j.	j.	PROPN
alces-23	203	16	saquet	saquet	PROPN
alces-23	203	17	,	,	PUNCT
alces-23	203	18	h.	h.	PROPN
alces-23	203	19	hristienko	hristienko	PROPN
alces-23	203	20	,	,	PUNCT
alces-23	203	21	f.	f.	PROPN
alces-23	203	22	burrows	burrows	PROPN
alces-23	203	23	,	,	PUNCT
alces-23	203	24	r.	r.	PROPN
alces-23	203	25	peterson	peterson	PROPN
alces-23	203	26	,	,	PUNCT
alces-23	203	27	and	and	CCONJ
alces-23	203	28	b.	b.	PROPN
alces-23	203	29	n.	n.	PROPN
alces-23	203	30	white	white	PROPN
alces-23	203	31	.	.	PUNCT
alces-23	204	1	2003	2003	NUM
alces-23	204	2	.	.	PUNCT
alces-23	205	1	genetic	genetic	ADJ
alces-23	205	2	variation	variation	NOUN
alces-23	205	3	and	and	CCONJ
alces-23	205	4	population	population	NOUN
alces-23	205	5	structure	structure	NOUN
alces-23	205	6	of	of	ADP
alces-23	205	7	moose	moose	NOUN
alces-23	205	8	(	(	PUNCT
alces-23	205	9	alces	alce	NOUN
alces-23	205	10	alces	alce	NOUN
alces-23	205	11	)	)	PUNCT
alces-23	205	12	at	at	ADP
alces-23	205	13	neutral	neutral	ADJ
alces-23	205	14	and	and	CCONJ
alces-23	205	15	functional	functional	ADJ
alces-23	205	16	dna	dna	PROPN
alces-23	205	17	loci	loci	PROPN
alces-23	205	18	.	.	PUNCT
alces-23	206	1	canadian	canadian	ADJ
alces-23	206	2	journal	journal	PROPN
alces-23	206	3	of	of	ADP
alces-23	206	4	zoology	zoology	NOUN
alces-23	206	5	81	81	NUM
alces-23	206	6	:	:	PUNCT
alces-23	206	7	670	670	NUM
alces-23	206	8	-	-	SYM
alces-23	206	9	683	683	NUM
alces-23	206	10	.	.	PUNCT