id	sid	tid	token	lemma	pos
alces-263	1	1	39	39	NUM
alces-263	1	2	revisiting	revisit	VERB
alces-263	1	3	the	the	DET
alces-263	1	4	recruitment	recruitment	NOUN
alces-263	1	5	-	-	PUNCT
alces-263	1	6	mortality	mortality	NOUN
alces-263	1	7	equation	equation	NOUN
alces-263	1	8	to	to	PART
alces-263	1	9	assess	assess	VERB
alces-263	1	10	moose	moose	NOUN
alces-263	1	11	growth	growth	NOUN
alces-263	1	12	rates	rate	NOUN
alces-263	1	13	ian	ian	PROPN
alces-263	1	14	w.	w.	PROPN
alces-263	1	15	hatter	hatter	PROPN
alces-263	1	16	nature	nature	PROPN
alces-263	1	17	wise	wise	ADJ
alces-263	1	18	consulting	consulting	NOUN
alces-263	1	19	,	,	PUNCT
alces-263	1	20	308	308	NUM
alces-263	1	21	uganda	uganda	PROPN
alces-263	1	22	avenue	avenue	PROPN
alces-263	1	23	,	,	PUNCT
alces-263	1	24	victoria	victoria	PROPN
alces-263	1	25	,	,	PUNCT
alces-263	1	26	british	british	PROPN
alces-263	1	27	columbia	columbia	PROPN
alces-263	1	28	v9a	v9a	VERB
alces-263	1	29	5x7	5x7	NUM
alces-263	1	30	,	,	PUNCT
alces-263	1	31	canada	canada	PROPN
alces-263	1	32	abstract	abstract	NOUN
alces-263	1	33	:	:	PUNCT
alces-263	1	34	hatter	hatter	NOUN
alces-263	1	35	and	and	CCONJ
alces-263	1	36	bergerud	bergerud	NOUN
alces-263	1	37	(	(	PUNCT
alces-263	1	38	1991	1991	NUM
alces-263	1	39	)	)	PUNCT
alces-263	1	40	developed	develop	VERB
alces-263	1	41	a	a	DET
alces-263	1	42	recruitment	recruitment	NOUN
alces-263	1	43	-	-	PUNCT
alces-263	1	44	mortality	mortality	NOUN
alces-263	1	45	(	(	PUNCT
alces-263	1	46	r	r	NOUN
alces-263	1	47	-	-	PUNCT
alces-263	1	48	m	m	NOUN
alces-263	1	49	)	)	PUNCT
alces-263	1	50	equation	equation	NOUN
alces-263	1	51	to	to	PART
alces-263	1	52	estimate	estimate	VERB
alces-263	1	53	the	the	DET
alces-263	1	54	annual	annual	ADJ
alces-263	1	55	finite	finite	ADJ
alces-263	1	56	rate	rate	NOUN
alces-263	1	57	of	of	ADP
alces-263	1	58	change	change	NOUN
alces-263	1	59	(	(	PUNCT
alces-263	1	60	λ	λ	NOUN
alces-263	1	61	)	)	PUNCT
alces-263	1	62	in	in	ADP
alces-263	1	63	a	a	DET
alces-263	1	64	moose	moose	NOUN
alces-263	1	65	(	(	PUNCT
alces-263	1	66	alces	alce	NOUN
alces-263	1	67	alces	alce	NOUN
alces-263	1	68	)	)	PUNCT
alces-263	1	69	population	population	NOUN
alces-263	1	70	from	from	ADP
alces-263	1	71	a	a	DET
alces-263	1	72	single	single	ADJ
alces-263	1	73	estimate	estimate	NOUN
alces-263	1	74	of	of	ADP
alces-263	1	75	calf	calf	NOUN
alces-263	1	76	recruitment	recruitment	NOUN
alces-263	1	77	and	and	CCONJ
alces-263	1	78	adult	adult	NOUN
alces-263	1	79	mortality	mortality	NOUN
alces-263	1	80	.	.	PUNCT
alces-263	2	1	i	i	PRON
alces-263	2	2	present	present	VERB
alces-263	2	3	and	and	CCONJ
alces-263	2	4	assess	assess	VERB
alces-263	2	5	an	an	DET
alces-263	2	6	alternative	alternative	ADJ
alces-263	2	7	formulation	formulation	NOUN
alces-263	2	8	of	of	ADP
alces-263	2	9	the	the	DET
alces-263	2	10	r	r	PROPN
alces-263	2	11	-	-	PUNCT
alces-263	2	12	m	m	NOUN
alces-263	2	13	equation	equation	NOUN
alces-263	2	14	and	and	CCONJ
alces-263	2	15	compare	compare	VERB
alces-263	2	16	it	it	PRON
alces-263	2	17	with	with	ADP
alces-263	2	18	the	the	DET
alces-263	2	19	original	original	NOUN
alces-263	2	20	.	.	PUNCT
alces-263	3	1	a	a	DET
alces-263	3	2	modification	modification	NOUN
alces-263	3	3	to	to	ADP
alces-263	3	4	the	the	DET
alces-263	3	5	r	r	NOUN
alces-263	3	6	-	-	PUNCT
alces-263	3	7	m	m	NOUN
alces-263	3	8	equations	equation	NOUN
alces-263	3	9	is	be	AUX
alces-263	3	10	provided	provide	VERB
alces-263	3	11	to	to	PART
alces-263	3	12	accommodate	accommodate	VERB
alces-263	3	13	early	early	ADV
alces-263	3	14	to	to	ADP
alces-263	3	15	mid	mid	ADJ
alces-263	3	16	-	-	NOUN
alces-263	3	17	winter	winter	NOUN
alces-263	3	18	composition	composition	NOUN
alces-263	3	19	surveys	survey	NOUN
alces-263	3	20	where	where	SCONJ
alces-263	3	21	recruitment	recruitment	NOUN
alces-263	3	22	is	be	AUX
alces-263	3	23	measured	measure	VERB
alces-263	3	24	when	when	SCONJ
alces-263	3	25	calves	calf	NOUN
alces-263	3	26	are	be	AUX
alces-263	3	27	less	less	ADJ
alces-263	3	28	than	than	ADP
alces-263	3	29	1	1	NUM
alces-263	3	30	year	year	NOUN
alces-263	3	31	-	-	PUNCT
alces-263	3	32	of	of	ADP
alces-263	3	33	-	-	PUNCT
alces-263	3	34	age	age	NOUN
alces-263	3	35	.	.	PUNCT
alces-263	4	1	an	an	DET
alces-263	4	2	example	example	NOUN
alces-263	4	3	with	with	ADP
alces-263	4	4	the	the	DET
alces-263	4	5	modified	modified	ADJ
alces-263	4	6	r	r	NOUN
alces-263	4	7	-	-	PUNCT
alces-263	4	8	m	m	NOUN
alces-263	4	9	equation	equation	NOUN
alces-263	4	10	illustrates	illustrate	VERB
alces-263	4	11	estimation	estimation	NOUN
alces-263	4	12	of	of	ADP
alces-263	4	13	λ	λ	PROPN
alces-263	4	14	for	for	ADP
alces-263	4	15	the	the	DET
alces-263	4	16	female	female	ADJ
alces-263	4	17	component	component	NOUN
alces-263	4	18	of	of	ADP
alces-263	4	19	two	two	NUM
alces-263	4	20	moose	moose	NOUN
alces-263	4	21	populations	population	NOUN
alces-263	4	22	under	under	ADP
alces-263	4	23	recent	recent	ADJ
alces-263	4	24	study	study	NOUN
alces-263	4	25	in	in	ADP
alces-263	4	26	british	british	PROPN
alces-263	4	27	columbia	columbia	PROPN
alces-263	4	28	,	,	PUNCT
alces-263	4	29	canada	canada	PROPN
alces-263	4	30	.	.	PUNCT
alces-263	5	1	due	due	ADP
alces-263	5	2	to	to	ADP
alces-263	5	3	potential	potential	ADJ
alces-263	5	4	biases	bias	NOUN
alces-263	5	5	with	with	ADP
alces-263	5	6	estimating	estimate	VERB
alces-263	5	7	recruitment	recruitment	NOUN
alces-263	5	8	and	and	CCONJ
alces-263	5	9	mortality	mortality	NOUN
alces-263	5	10	rates	rate	NOUN
alces-263	5	11	,	,	PUNCT
alces-263	5	12	the	the	DET
alces-263	5	13	calculation	calculation	NOUN
alces-263	5	14	of	of	ADP
alces-263	5	15	λ	λ	PROPN
alces-263	5	16	with	with	ADP
alces-263	5	17	the	the	DET
alces-263	5	18	r	r	PROPN
alces-263	5	19	-	-	PUNCT
alces-263	5	20	m	m	NOUN
alces-263	5	21	equation	equation	NOUN
alces-263	5	22	should	should	AUX
alces-263	5	23	be	be	AUX
alces-263	5	24	verified	verify	VERB
alces-263	5	25	with	with	ADP
alces-263	5	26	periodic	periodic	ADJ
alces-263	5	27	density	density	NOUN
alces-263	5	28	surveys	survey	NOUN
alces-263	5	29	whenever	whenever	SCONJ
alces-263	5	30	possible	possible	ADJ
alces-263	5	31	.	.	PUNCT
alces-263	6	1	the	the	DET
alces-263	6	2	r	r	NOUN
alces-263	6	3	-	-	PUNCT
alces-263	6	4	m	m	NOUN
alces-263	6	5	equation	equation	NOUN
alces-263	6	6	is	be	AUX
alces-263	6	7	most	most	ADV
alces-263	6	8	useful	useful	ADJ
alces-263	6	9	for	for	ADP
alces-263	6	10	estimating	estimate	VERB
alces-263	6	11	λ	λ	PROPN
alces-263	6	12	when	when	SCONJ
alces-263	6	13	moose	moose	NOUN
alces-263	6	14	density	density	NOUN
alces-263	6	15	surveys	survey	NOUN
alces-263	6	16	are	be	AUX
alces-263	6	17	not	not	PART
alces-263	6	18	feasible	feasible	ADJ
alces-263	6	19	or	or	CCONJ
alces-263	6	20	an	an	DET
alces-263	6	21	estimate	estimate	NOUN
alces-263	6	22	of	of	ADP
alces-263	6	23	the	the	DET
alces-263	6	24	adult	adult	NOUN
alces-263	6	25	survival	survival	NOUN
alces-263	6	26	rate	rate	NOUN
alces-263	6	27	is	be	AUX
alces-263	6	28	available	available	ADJ
alces-263	6	29	.	.	PUNCT
alces-263	7	1	alces	alce	NOUN
alces-263	7	2	vol	vol	NOUN
alces-263	7	3	.	.	PUNCT
alces-263	8	1	56:39–47	56:39–47	NUM
alces-263	8	2	(	(	PUNCT
alces-263	8	3	2020	2020	NUM
alces-263	8	4	)	)	PUNCT
alces-263	8	5	key	key	ADJ
alces-263	8	6	words	word	NOUN
alces-263	8	7	:	:	PUNCT
alces-263	8	8	alces	alce	NOUN
alces-263	8	9	alces	alce	NOUN
alces-263	8	10	,	,	PUNCT
alces-263	8	11	finite	finite	ADJ
alces-263	8	12	rate	rate	NOUN
alces-263	8	13	of	of	ADP
alces-263	8	14	change	change	NOUN
alces-263	8	15	,	,	PUNCT
alces-263	8	16	mortality	mortality	NOUN
alces-263	8	17	rate	rate	NOUN
alces-263	8	18	,	,	PUNCT
alces-263	8	19	population	population	NOUN
alces-263	8	20	growth	growth	NOUN
alces-263	8	21	rate	rate	NOUN
alces-263	8	22	,	,	PUNCT
alces-263	8	23	recruitment	recruitment	NOUN
alces-263	8	24	mortality	mortality	NOUN
alces-263	8	25	equation	equation	NOUN
alces-263	8	26	,	,	PUNCT
alces-263	8	27	recruitment	recruitment	NOUN
alces-263	8	28	rate	rate	NOUN
alces-263	8	29	,	,	PUNCT
alces-263	8	30	survival	survival	NOUN
alces-263	8	31	rate	rate	NOUN
alces-263	8	32	hatter	hatter	NOUN
alces-263	8	33	and	and	CCONJ
alces-263	8	34	bergerud	bergerud	NOUN
alces-263	8	35	(	(	PUNCT
alces-263	8	36	1991	1991	NUM
alces-263	8	37	)	)	PUNCT
alces-263	8	38	developed	develop	VERB
alces-263	8	39	a	a	DET
alces-263	8	40	recruitment	recruitment	NOUN
alces-263	8	41	-	-	PUNCT
alces-263	8	42	mortality	mortality	NOUN
alces-263	8	43	(	(	PUNCT
alces-263	8	44	r	r	NOUN
alces-263	8	45	-	-	PUNCT
alces-263	8	46	m	m	NOUN
alces-263	8	47	)	)	PUNCT
alces-263	8	48	equation	equation	NOUN
alces-263	8	49	to	to	PART
alces-263	8	50	estimate	estimate	VERB
alces-263	8	51	the	the	DET
alces-263	8	52	finite	finite	ADJ
alces-263	8	53	rate	rate	NOUN
alces-263	8	54	of	of	ADP
alces-263	8	55	change	change	NOUN
alces-263	8	56	or	or	CCONJ
alces-263	8	57	growth	growth	NOUN
alces-263	8	58	rate	rate	NOUN
alces-263	8	59	(	(	PUNCT
alces-263	8	60	λ	λ	NOUN
alces-263	8	61	)	)	PUNCT
alces-263	8	62	of	of	ADP
alces-263	8	63	an	an	DET
alces-263	8	64	ungulate	ungulate	ADJ
alces-263	8	65	population	population	NOUN
alces-263	8	66	from	from	ADP
alces-263	8	67	estimates	estimate	NOUN
alces-263	8	68	of	of	ADP
alces-263	8	69	juvenile	juvenile	ADJ
alces-263	8	70	recruitment	recruitment	NOUN
alces-263	8	71	(	(	PUNCT
alces-263	8	72	r	r	NOUN
alces-263	8	73	)	)	PUNCT
alces-263	8	74	and	and	CCONJ
alces-263	8	75	adult	adult	NOUN
alces-263	8	76	mortality	mortality	NOUN
alces-263	8	77	(	(	PUNCT
alces-263	8	78	m	m	NOUN
alces-263	8	79	)	)	PUNCT
alces-263	8	80	rates	rate	NOUN
alces-263	8	81	where	where	SCONJ
alces-263	8	82	λ	λ	X
alces-263	8	83	=	=	SYM
alces-263	8	84	(	(	PUNCT
alces-263	8	85	1−m)/(1−r	1−m)/(1−r	NUM
alces-263	8	86	)	)	PUNCT
alces-263	8	87	.	.	PUNCT
alces-263	9	1	this	this	DET
alces-263	9	2	equation	equation	NOUN
alces-263	9	3	has	have	AUX
alces-263	9	4	been	be	AUX
alces-263	9	5	used	use	VERB
alces-263	9	6	commonly	commonly	ADV
alces-263	9	7	with	with	ADP
alces-263	9	8	populations	population	NOUN
alces-263	9	9	of	of	ADP
alces-263	9	10	moose	moose	NOUN
alces-263	9	11	(	(	PUNCT
alces-263	9	12	alces	alce	NOUN
alces-263	9	13	alces	alce	NOUN
alces-263	9	14	)	)	PUNCT
alces-263	9	15	(	(	PUNCT
alces-263	9	16	e.g.	e.g.	ADV
alces-263	9	17	,	,	PUNCT
alces-263	9	18	gasaway	gasaway	NOUN
alces-263	9	19	et	et	PROPN
alces-263	9	20	al	al	PROPN
alces-263	9	21	.	.	PROPN
alces-263	9	22	1992	1992	NUM
alces-263	9	23	,	,	PUNCT
alces-263	9	24	boertje	boertje	PROPN
alces-263	9	25	et	et	PROPN
alces-263	9	26	al	al	PROPN
alces-263	9	27	.	.	PROPN
alces-263	9	28	1996	1996	NUM
alces-263	9	29	,	,	PUNCT
alces-263	9	30	kunkel	kunkel	PROPN
alces-263	9	31	and	and	CCONJ
alces-263	9	32	pletscher	pletscher	NOUN
alces-263	9	33	1999	1999	NUM
alces-263	9	34	,	,	PUNCT
alces-263	9	35	hayes	hayes	PROPN
alces-263	9	36	et	et	PROPN
alces-263	9	37	al	al	PROPN
alces-263	9	38	.	.	PROPN
alces-263	9	39	2000	2000	NUM
alces-263	9	40	,	,	PUNCT
alces-263	9	41	kuzyk	kuzyk	INTJ
alces-263	9	42	et	et	PROPN
alces-263	9	43	al	al	PROPN
alces-263	9	44	.	.	PROPN
alces-263	9	45	2019b	2019b	NOUN
alces-263	9	46	,	,	PUNCT
alces-263	9	47	severud	severud	NOUN
alces-263	9	48	et	et	PROPN
alces-263	9	49	al	al	PROPN
alces-263	9	50	.	.	PROPN
alces-263	9	51	2019	2019	NUM
alces-263	9	52	)	)	PUNCT
alces-263	9	53	,	,	PUNCT
alces-263	9	54	caribou	caribou	NOUN
alces-263	9	55	(	(	PUNCT
alces-263	9	56	rangifer	rangifer	VERB
alces-263	9	57	tarandus	tarandus	NOUN
alces-263	9	58	)	)	PUNCT
alces-263	9	59	(	(	PUNCT
alces-263	9	60	e.g.	e.g.	ADV
alces-263	9	61	,	,	PUNCT
alces-263	9	62	bergerud	bergerud	NOUN
alces-263	9	63	and	and	CCONJ
alces-263	9	64	elliott	elliott	PROPN
alces-263	9	65	1986	1986	NUM
alces-263	9	66	,	,	PUNCT
alces-263	9	67	seip	seip	NOUN
alces-263	9	68	and	and	CCONJ
alces-263	9	69	cichowski	cichowski	PROPN
alces-263	9	70	1996	1996	NUM
alces-263	9	71	,	,	PUNCT
alces-263	9	72	mcloughlin	mcloughlin	PROPN
alces-263	9	73	et	et	PROPN
alces-263	9	74	al	al	PROPN
alces-263	9	75	.	.	PROPN
alces-263	9	76	2003	2003	NUM
alces-263	9	77	,	,	PUNCT
alces-263	9	78	hebblewhite	hebblewhite	NOUN
alces-263	9	79	et	et	PROPN
alces-263	9	80	al	al	PROPN
alces-263	9	81	.	.	PROPN
alces-263	9	82	2007	2007	NUM
alces-263	9	83	,	,	PUNCT
alces-263	9	84	sorensen	sorensen	PROPN
alces-263	9	85	et	et	PROPN
alces-263	9	86	al	al	PROPN
alces-263	9	87	.	.	PROPN
alces-263	9	88	2008	2008	NUM
alces-263	9	89	,	,	PUNCT
alces-263	9	90	latham	latham	PROPN
alces-263	9	91	et	et	PROPN
alces-263	9	92	al	al	PROPN
alces-263	9	93	.	.	PROPN
alces-263	9	94	2011	2011	NUM
alces-263	9	95	,	,	PUNCT
alces-263	9	96	decesare	decesare	VERB
alces-263	9	97	et	et	PROPN
alces-263	9	98	al	al	PROPN
alces-263	9	99	.	.	PROPN
alces-263	9	100	2012	2012	NUM
alces-263	9	101	,	,	PUNCT
alces-263	9	102	hervieux	hervieux	PROPN
alces-263	9	103	et	et	PROPN
alces-263	9	104	al	al	PROPN
alces-263	9	105	.	.	PROPN
alces-263	9	106	2013	2013	NUM
alces-263	9	107	,	,	PUNCT
alces-263	9	108	2014	2014	NUM
alces-263	9	109	,	,	PUNCT
alces-263	9	110	serrouya	serrouya	PROPN
alces-263	9	111	et	et	PROPN
alces-263	9	112	al	al	PROPN
alces-263	9	113	.	.	PROPN
alces-263	9	114	2017	2017	NUM
alces-263	9	115	)	)	PUNCT
alces-263	9	116	,	,	PUNCT
alces-263	9	117	elk	elk	PROPN
alces-263	9	118	(	(	PUNCT
alces-263	9	119	cervus	cervus	X
alces-263	9	120	elaphus	elaphu	NOUN
alces-263	9	121	)	)	PUNCT
alces-263	9	122	(	(	PUNCT
alces-263	9	123	e.g.	e.g.	ADV
alces-263	9	124	,	,	PUNCT
alces-263	9	125	kunkel	kunkel	PROPN
alces-263	9	126	and	and	CCONJ
alces-263	9	127	pletscher	pletscher	NOUN
alces-263	9	128	1999	1999	NUM
alces-263	9	129	,	,	PUNCT
alces-263	9	130	devore	devore	PROPN
alces-263	9	131	et	et	PROPN
alces-263	9	132	al	al	PROPN
alces-263	9	133	.	.	PROPN
alces-263	9	134	2018	2018	NUM
alces-263	9	135	)	)	PUNCT
alces-263	9	136	,	,	PUNCT
alces-263	9	137	black	black	NOUN
alces-263	9	138	-	-	PUNCT
alces-263	9	139	tailed	tail	VERB
alces-263	9	140	deer	deer	NOUN
alces-263	9	141	(	(	PUNCT
alces-263	9	142	odocoileus	odocoileus	PROPN
alces-263	9	143	hemionus	hemionus	PROPN
alces-263	9	144	columbianus	columbianus	PROPN
alces-263	9	145	)	)	PUNCT
alces-263	9	146	(	(	PUNCT
alces-263	9	147	e.g.	e.g.	ADV
alces-263	9	148	,	,	PUNCT
alces-263	9	149	hatter	hatter	NOUN
alces-263	9	150	and	and	CCONJ
alces-263	9	151	janz	janz	PROPN
alces-263	9	152	1994	1994	NUM
alces-263	9	153	)	)	PUNCT
alces-263	9	154	,	,	PUNCT
alces-263	9	155	and	and	CCONJ
alces-263	9	156	white	white	ADJ
alces-263	9	157	-	-	PUNCT
alces-263	9	158	tailed	tail	VERB
alces-263	9	159	deer	deer	NOUN
alces-263	9	160	(	(	PUNCT
alces-263	9	161	odocoileus	odocoileus	ADJ
alces-263	9	162	virginianus	virginianus	PROPN
alces-263	9	163	)	)	PUNCT
alces-263	9	164	(	(	PUNCT
alces-263	9	165	e.g.	e.g.	ADV
alces-263	9	166	,	,	PUNCT
alces-263	9	167	kunkel	kunkel	PROPN
alces-263	9	168	and	and	CCONJ
alces-263	9	169	pletscher	pletscher	NOUN
alces-263	9	170	1999	1999	NUM
alces-263	9	171	,	,	PUNCT
alces-263	9	172	patterson	patterson	PROPN
alces-263	9	173	et	et	PROPN
alces-263	9	174	al	al	PROPN
alces-263	9	175	.	.	PROPN
alces-263	9	176	2002	2002	NUM
alces-263	9	177	)	)	PUNCT
alces-263	9	178	.	.	PUNCT
alces-263	10	1	refinements	refinement	NOUN
alces-263	10	2	have	have	AUX
alces-263	10	3	been	be	AUX
alces-263	10	4	made	make	VERB
alces-263	10	5	by	by	ADP
alces-263	10	6	several	several	ADJ
alces-263	10	7	authors	author	NOUN
alces-263	10	8	to	to	PART
alces-263	10	9	improve	improve	VERB
alces-263	10	10	its	its	PRON
alces-263	10	11	utility	utility	NOUN
alces-263	10	12	as	as	ADP
alces-263	10	13	an	an	DET
alces-263	10	14	ungulate	ungulate	ADJ
alces-263	10	15	population	population	NOUN
alces-263	10	16	assessment	assessment	NOUN
alces-263	10	17	tool	tool	NOUN
alces-263	10	18	(	(	PUNCT
alces-263	10	19	decesare	decesare	NOUN
alces-263	10	20	et	et	PROPN
alces-263	10	21	al	al	PROPN
alces-263	10	22	.	.	PROPN
alces-263	10	23	2011	2011	NUM
alces-263	10	24	,	,	PUNCT
alces-263	10	25	hervieux	hervieux	PROPN
alces-263	10	26	et	et	PROPN
alces-263	10	27	al	al	PROPN
alces-263	10	28	.	.	PROPN
alces-263	10	29	2013	2013	NUM
alces-263	10	30	)	)	PUNCT
alces-263	10	31	.	.	PUNCT
alces-263	11	1	the	the	DET
alces-263	11	2	primary	primary	ADJ
alces-263	11	3	utility	utility	NOUN
alces-263	11	4	of	of	ADP
alces-263	11	5	the	the	DET
alces-263	11	6	r	r	PROPN
alces-263	11	7	-	-	PUNCT
alces-263	11	8	m	m	NOUN
alces-263	11	9	equation	equation	NOUN
alces-263	11	10	is	be	AUX
alces-263	11	11	that	that	SCONJ
alces-263	11	12	it	it	PRON
alces-263	11	13	calculates	calculate	VERB
alces-263	11	14	λ	λ	PROPN
alces-263	11	15	from	from	ADP
alces-263	11	16	a	a	DET
alces-263	11	17	single	single	ADJ
alces-263	11	18	estimate	estimate	NOUN
alces-263	11	19	of	of	ADP
alces-263	11	20	recruitment	recruitment	NOUN
alces-263	11	21	provided	provide	VERB
alces-263	11	22	that	that	SCONJ
alces-263	11	23	the	the	DET
alces-263	11	24	adult	adult	NOUN
alces-263	11	25	mortality	mortality	NOUN
alces-263	11	26	rate	rate	NOUN
alces-263	11	27	is	be	AUX
alces-263	11	28	known	know	VERB
alces-263	11	29	.	.	PUNCT
alces-263	12	1	it	it	PRON
alces-263	12	2	is	be	AUX
alces-263	12	3	particularly	particularly	ADV
alces-263	12	4	appropriate	appropriate	ADJ
alces-263	12	5	when	when	SCONJ
alces-263	12	6	there	there	PRON
alces-263	12	7	are	be	VERB
alces-263	12	8	few	few	ADJ
alces-263	12	9	other	other	ADJ
alces-263	12	10	cost	cost	NOUN
alces-263	12	11	-	-	PUNCT
alces-263	12	12	effective	effective	ADJ
alces-263	12	13	alternatives	alternative	NOUN
alces-263	12	14	.	.	PUNCT
alces-263	13	1	one	one	NUM
alces-263	13	2	drawback	drawback	NOUN
alces-263	13	3	of	of	ADP
alces-263	13	4	using	use	VERB
alces-263	13	5	the	the	DET
alces-263	13	6	r	r	NOUN
alces-263	13	7	-	-	PUNCT
alces-263	13	8	m	m	NOUN
alces-263	13	9	equation	equation	NOUN
alces-263	13	10	with	with	ADP
alces-263	13	11	moose	moose	NOUN
alces-263	13	12	is	be	AUX
alces-263	13	13	that	that	SCONJ
alces-263	13	14	recruitment	recruitment	NOUN
alces-263	13	15	is	be	AUX
alces-263	13	16	commonly	commonly	ADV
alces-263	13	17	measured	measure	VERB
alces-263	13	18	from	from	ADP
alces-263	13	19	composition	composition	NOUN
alces-263	13	20	surveys	survey	NOUN
alces-263	13	21	conducted	conduct	VERB
alces-263	13	22	during	during	ADP
alces-263	13	23	early	early	ADJ
alces-263	13	24	or	or	CCONJ
alces-263	13	25	mid	mid	NOUN
alces-263	13	26	-	-	NOUN
alces-263	13	27	winter	winter	NOUN
alces-263	13	28	before	before	SCONJ
alces-263	13	29	calves	calf	NOUN
alces-263	13	30	are	be	AUX
alces-263	13	31	recruited	recruit	VERB
alces-263	13	32	into	into	ADP
alces-263	13	33	the	the	DET
alces-263	13	34	adult	adult	NOUN
alces-263	13	35	population	population	NOUN
alces-263	13	36	at	at	ADP
alces-263	13	37	1	1	NUM
alces-263	13	38	year	year	NOUN
alces-263	13	39	-	-	PUNCT
alces-263	13	40	of	of	ADP
alces-263	13	41	-	-	PUNCT
alces-263	13	42	age	age	NOUN
alces-263	13	43	(	(	PUNCT
alces-263	13	44	hatter	hatter	NOUN
alces-263	13	45	and	and	CCONJ
alces-263	13	46	bergerud	bergerud	NOUN
alces-263	13	47	1991	1991	NUM
alces-263	13	48	)	)	PUNCT
alces-263	13	49	.	.	PUNCT
alces-263	14	1	this	this	PRON
alces-263	14	2	is	be	AUX
alces-263	14	3	because	because	SCONJ
alces-263	14	4	aerial	aerial	ADJ
alces-263	14	5	spring	spring	NOUN
alces-263	14	6	surveys	survey	NOUN
alces-263	14	7	are	be	AUX
alces-263	14	8	generally	generally	ADV
alces-263	14	9	impractical	impractical	ADJ
alces-263	14	10	due	due	ADJ
alces-263	14	11	to	to	ADP
alces-263	14	12	lack	lack	NOUN
alces-263	14	13	of	of	ADP
alces-263	14	14	snow	snow	NOUN
alces-263	14	15	-	-	PUNCT
alces-263	14	16	cover	cover	NOUN
alces-263	14	17	,	,	PUNCT
alces-263	14	18	poor	poor	ADJ
alces-263	14	19	visibility	visibility	NOUN
alces-263	14	20	,	,	PUNCT
alces-263	14	21	revisiting	revisit	VERB
alces-263	14	22	the	the	DET
alces-263	14	23	r	r	NOUN
alces-263	14	24	-	-	PUNCT
alces-263	14	25	m	m	NOUN
alces-263	14	26	equation	equation	NOUN
alces-263	14	27	.	.	PUNCT
alces-263	15	1	–	–	PUNCT
alces-263	15	2	ian	ian	PROPN
alces-263	15	3	w.	w.	PROPN
alces-263	15	4	hatter	hatter	PROPN
alces-263	15	5	alces	alces	PROPN
alces-263	15	6	vol	vol	NOUN
alces-263	15	7	.	.	PROPN
alces-263	16	1	56	56	NUM
alces-263	16	2	,	,	PUNCT
alces-263	16	3	2020	2020	NUM
alces-263	16	4	40	40	NUM
alces-263	16	5	and	and	CCONJ
alces-263	16	6	dispersal	dispersal	NOUN
alces-263	16	7	of	of	ADP
alces-263	16	8	moose	moose	NOUN
alces-263	16	9	from	from	ADP
alces-263	16	10	winter	winter	NOUN
alces-263	16	11	ranges	range	NOUN
alces-263	16	12	(	(	PUNCT
alces-263	16	13	gasaway	gasaway	NOUN
alces-263	16	14	et	et	PROPN
alces-263	16	15	al	al	PROPN
alces-263	16	16	.	.	PROPN
alces-263	17	1	1986	1986	NUM
alces-263	17	2	)	)	PUNCT
alces-263	17	3	.	.	PUNCT
alces-263	18	1	the	the	DET
alces-263	18	2	purposes	purpose	NOUN
alces-263	18	3	of	of	ADP
alces-263	18	4	this	this	DET
alces-263	18	5	paper	paper	NOUN
alces-263	18	6	were	be	AUX
alces-263	18	7	to	to	PART
alces-263	18	8	review	review	VERB
alces-263	18	9	the	the	DET
alces-263	18	10	original	original	ADJ
alces-263	18	11	r	r	NOUN
alces-263	18	12	-	-	PUNCT
alces-263	18	13	m	m	NOUN
alces-263	18	14	equation	equation	NOUN
alces-263	18	15	(	(	PUNCT
alces-263	18	16	hatter	hatter	NOUN
alces-263	18	17	and	and	CCONJ
alces-263	18	18	bergerud	bergerud	NOUN
alces-263	18	19	1991	1991	NUM
alces-263	18	20	)	)	PUNCT
alces-263	18	21	,	,	PUNCT
alces-263	18	22	compare	compare	VERB
alces-263	18	23	it	it	PRON
alces-263	18	24	with	with	ADP
alces-263	18	25	an	an	DET
alces-263	18	26	alternative	alternative	ADJ
alces-263	18	27	formulation	formulation	NOUN
alces-263	18	28	,	,	PUNCT
alces-263	18	29	identify	identify	VERB
alces-263	18	30	a	a	DET
alces-263	18	31	modification	modification	NOUN
alces-263	18	32	that	that	PRON
alces-263	18	33	accommodates	accommodate	VERB
alces-263	18	34	moose	moose	NOUN
alces-263	18	35	composition	composition	NOUN
alces-263	18	36	surveys	survey	NOUN
alces-263	18	37	before	before	SCONJ
alces-263	18	38	calves	calf	NOUN
alces-263	18	39	are	be	AUX
alces-263	18	40	1	1	NUM
alces-263	18	41	year	year	NOUN
alces-263	18	42	-	-	PUNCT
alces-263	18	43	of	of	ADP
alces-263	18	44	-	-	PUNCT
alces-263	18	45	age	age	NOUN
alces-263	18	46	,	,	PUNCT
alces-263	18	47	and	and	CCONJ
alces-263	18	48	provide	provide	VERB
alces-263	18	49	an	an	DET
alces-263	18	50	example	example	NOUN
alces-263	18	51	of	of	ADP
alces-263	18	52	its	its	PRON
alces-263	18	53	use	use	NOUN
alces-263	18	54	.	.	PUNCT
alces-263	19	1	the	the	DET
alces-263	19	2	r	r	NOUN
alces-263	19	3	-	-	PUNCT
alces-263	19	4	m	m	NOUN
alces-263	19	5	equation	equation	NOUN
alces-263	19	6	the	the	DET
alces-263	19	7	r	r	NOUN
alces-263	19	8	-	-	PUNCT
alces-263	19	9	m	m	NOUN
alces-263	19	10	equation	equation	NOUN
alces-263	19	11	enables	enable	VERB
alces-263	19	12	the	the	DET
alces-263	19	13	calculation	calculation	NOUN
alces-263	19	14	of	of	ADP
alces-263	19	15	λ	λ	PROPN
alces-263	19	16	from	from	ADP
alces-263	19	17	annual	annual	ADJ
alces-263	19	18	recruitment	recruitment	NOUN
alces-263	19	19	and	and	CCONJ
alces-263	19	20	adult	adult	NOUN
alces-263	19	21	mortality	mortality	NOUN
alces-263	19	22	rates	rate	NOUN
alces-263	19	23	.	.	PUNCT
alces-263	20	1	this	this	DET
alces-263	20	2	equation	equation	NOUN
alces-263	20	3	expressed	express	VERB
alces-263	20	4	as	as	ADP
alces-263	20	5	a	a	DET
alces-263	20	6	difference	difference	NOUN
alces-263	20	7	equation	equation	NOUN
alces-263	20	8	(	(	PUNCT
alces-263	20	9	serrouya	serrouya	PROPN
alces-263	20	10	et	et	PROPN
alces-263	20	11	al	al	PROPN
alces-263	20	12	.	.	PROPN
alces-263	20	13	2017	2017	NUM
alces-263	20	14	)	)	PUNCT
alces-263	21	1	is	be	AUX
alces-263	21	2	:	:	PUNCT
alces-263	21	3	nt+1	nt+1	X
alces-263	21	4	=	=	SYM
alces-263	22	1	nt	not	PART
alces-263	22	2	+	+	CCONJ
alces-263	22	3	rnt+1	rnt+1	NOUN
alces-263	22	4	‒	‒	X
alces-263	22	5	mnt	mnt	NOUN
alces-263	22	6	(	(	PUNCT
alces-263	22	7	1	1	NUM
alces-263	22	8	)	)	PUNCT
alces-263	22	9	where	where	SCONJ
alces-263	22	10	r	r	NOUN
alces-263	22	11	is	be	AUX
alces-263	22	12	the	the	DET
alces-263	22	13	proportion	proportion	NOUN
alces-263	22	14	of	of	ADP
alces-263	22	15	juveniles	juvenile	NOUN
alces-263	22	16	at	at	ADP
alces-263	22	17	the	the	DET
alces-263	22	18	end	end	NOUN
alces-263	22	19	of	of	ADP
alces-263	22	20	their	their	PRON
alces-263	22	21	first	first	ADJ
alces-263	22	22	year	year	NOUN
alces-263	22	23	of	of	ADP
alces-263	22	24	life	life	NOUN
alces-263	22	25	(	(	PUNCT
alces-263	22	26	i.e.	i.e.	X
alces-263	22	27	,	,	PUNCT
alces-263	22	28	year	year	NOUN
alces-263	22	29	t1	t1	NOUN
alces-263	22	30	)	)	PUNCT
alces-263	22	31	and	and	CCONJ
alces-263	22	32	m	m	PROPN
alces-263	22	33	is	be	AUX
alces-263	22	34	the	the	DET
alces-263	22	35	adult	adult	NOUN
alces-263	22	36	(	(	PUNCT
alces-263	22	37	1	1	NUM
alces-263	22	38	+	+	NUM
alces-263	22	39	year	year	NOUN
alces-263	22	40	-	-	PUNCT
alces-263	22	41	old	old	ADJ
alces-263	22	42	males	male	NOUN
alces-263	22	43	and	and	CCONJ
alces-263	22	44	females	female	NOUN
alces-263	22	45	)	)	PUNCT
alces-263	22	46	mortality	mortality	NOUN
alces-263	22	47	rate	rate	NOUN
alces-263	22	48	during	during	ADP
alces-263	22	49	the	the	DET
alces-263	22	50	year	year	NOUN
alces-263	22	51	(	(	PUNCT
alces-263	22	52	i.e.	i.e.	X
alces-263	22	53	,	,	PUNCT
alces-263	22	54	from	from	ADP
alces-263	22	55	year	year	NOUN
alces-263	22	56	t0	t0	PROPN
alces-263	22	57	to	to	ADP
alces-263	22	58	t1	t1	PROPN
alces-263	22	59	)	)	PUNCT
alces-263	22	60	.	.	PUNCT
alces-263	23	1	rearranging	rearrange	VERB
alces-263	23	2	eq.1	eq.1	PROPN
alces-263	23	3	and	and	CCONJ
alces-263	23	4	solving	solve	VERB
alces-263	23	5	for	for	ADP
alces-263	23	6	λ	λ	PROPN
alces-263	23	7	=	=	SYM
alces-263	23	8	nt+1	nt+1	PROPN
alces-263	23	9	/	/	SYM
alces-263	23	10	nt	not	PART
alces-263	23	11	yields	yield	NOUN
alces-263	23	12	:	:	PUNCT
alces-263	23	13	λ	λ	X
alces-263	23	14	=	=	SYM
alces-263	23	15	(	(	PUNCT
alces-263	23	16	1	1	NUM
alces-263	23	17	‒	‒	NUM
alces-263	23	18	m)/(1	m)/(1	NOUN
alces-263	23	19	‒	‒	ADP
alces-263	23	20	r	r	NOUN
alces-263	23	21	)	)	PUNCT
alces-263	23	22	.	.	PUNCT
alces-263	24	1	(	(	PUNCT
alces-263	24	2	2	2	X
alces-263	24	3	)	)	PUNCT
alces-263	24	4	as	as	ADP
alces-263	24	5	1	1	NUM
alces-263	24	6	–	–	PUNCT
alces-263	24	7	m	m	VERB
alces-263	24	8	is	be	AUX
alces-263	24	9	the	the	DET
alces-263	24	10	adult	adult	NOUN
alces-263	24	11	survival	survival	NOUN
alces-263	24	12	rate	rate	NOUN
alces-263	24	13	,	,	PUNCT
alces-263	24	14	eq	eq	NOUN
alces-263	24	15	.	.	PROPN
alces-263	24	16	2	2	NUM
alces-263	24	17	also	also	ADV
alces-263	24	18	equals	equal	VERB
alces-263	24	19	:	:	PUNCT
alces-263	24	20	λ	λ	X
alces-263	24	21	=	=	PUNCT
alces-263	24	22	s/(1	s/(1	PROPN
alces-263	24	23	‒	‒	ADP
alces-263	24	24	r	r	NOUN
alces-263	24	25	)	)	PUNCT
alces-263	24	26	(	(	PUNCT
alces-263	24	27	3	3	X
alces-263	24	28	)	)	PUNCT
alces-263	24	29	where	where	SCONJ
alces-263	24	30	s	s	NOUN
alces-263	24	31	is	be	AUX
alces-263	24	32	the	the	DET
alces-263	24	33	annual	annual	ADJ
alces-263	24	34	probability	probability	NOUN
alces-263	24	35	of	of	ADP
alces-263	24	36	adult	adult	NOUN
alces-263	24	37	survival	survival	NOUN
alces-263	24	38	.	.	PUNCT
alces-263	25	1	here	here	ADV
alces-263	25	2	,	,	PUNCT
alces-263	25	3	i	i	PRON
alces-263	25	4	refer	refer	VERB
alces-263	25	5	to	to	ADP
alces-263	25	6	eq	eq	NOUN
alces-263	25	7	.	.	PROPN
alces-263	25	8	3	3	NUM
alces-263	25	9	as	as	ADP
alces-263	25	10	the	the	DET
alces-263	25	11	type	type	NOUN
alces-263	25	12	1	1	NUM
alces-263	25	13	r	r	NOUN
alces-263	25	14	-	-	PUNCT
alces-263	25	15	m	m	NOUN
alces-263	25	16	equation	equation	NOUN
alces-263	25	17	.	.	PUNCT
alces-263	26	1	if	if	SCONJ
alces-263	26	2	hunting	hunting	NOUN
alces-263	26	3	occurs	occur	VERB
alces-263	26	4	,	,	PUNCT
alces-263	26	5	then	then	ADV
alces-263	26	6	s	s	VERB
alces-263	26	7	must	must	AUX
alces-263	26	8	account	account	VERB
alces-263	26	9	for	for	ADP
alces-263	26	10	both	both	DET
alces-263	26	11	non	non	ADJ
alces-263	26	12	-	-	ADJ
alces-263	26	13	hunting	hunting	ADJ
alces-263	26	14	(	(	PUNCT
alces-263	26	15	mn	mn	NOUN
alces-263	26	16	)	)	PUNCT
alces-263	26	17	and	and	CCONJ
alces-263	26	18	hunting	hunt	VERB
alces-263	26	19	(	(	PUNCT
alces-263	26	20	mh	mh	NOUN
alces-263	26	21	)	)	PUNCT
alces-263	26	22	mortality	mortality	NOUN
alces-263	26	23	rates	rate	NOUN
alces-263	26	24	:	:	PUNCT
alces-263	26	25	s	s	AUX
alces-263	26	26	=	=	SYM
alces-263	26	27	(	(	PUNCT
alces-263	26	28	1‒	1‒	NUM
alces-263	26	29	mh	mh	PROPN
alces-263	26	30	)	)	PUNCT
alces-263	26	31	×	×	PROPN
alces-263	26	32	(	(	PUNCT
alces-263	26	33	1‒	1‒	NUM
alces-263	26	34	mn	mn	NOUN
alces-263	26	35	)	)	PUNCT
alces-263	26	36	(	(	PUNCT
alces-263	26	37	4	4	X
alces-263	26	38	)	)	PUNCT
alces-263	26	39	alternative	alternative	ADJ
alces-263	26	40	formulations	formulation	NOUN
alces-263	26	41	of	of	ADP
alces-263	26	42	the	the	DET
alces-263	26	43	r	r	PROPN
alces-263	26	44	-	-	PUNCT
alces-263	26	45	m	m	NOUN
alces-263	26	46	equation	equation	NOUN
alces-263	26	47	where	where	SCONJ
alces-263	26	48	r	r	NOUN
alces-263	26	49	is	be	AUX
alces-263	26	50	the	the	DET
alces-263	26	51	juvenile	juvenile	ADJ
alces-263	26	52	:	:	PUNCT
alces-263	26	53	adult	adult	NOUN
alces-263	26	54	ratio	ratio	NOUN
alces-263	26	55	and	and	CCONJ
alces-263	26	56	adults	adult	NOUN
alces-263	26	57	refer	refer	VERB
alces-263	26	58	to	to	ADP
alces-263	26	59	both	both	DET
alces-263	26	60	sexes	sex	NOUN
alces-263	26	61	combined	combine	VERB
alces-263	26	62	(	(	PUNCT
alces-263	26	63	guthery	guthery	NOUN
alces-263	26	64	and	and	CCONJ
alces-263	26	65	shaw	shaw	PROPN
alces-263	26	66	2013	2013	NUM
alces-263	26	67	,	,	PUNCT
alces-263	26	68	devore	devore	PROPN
alces-263	26	69	et	et	PROPN
alces-263	26	70	al	al	PROPN
alces-263	26	71	.	.	PROPN
alces-263	26	72	2018	2018	NUM
alces-263	26	73	)	)	PUNCT
alces-263	26	74	are	be	AUX
alces-263	26	75	:	:	PUNCT
alces-263	26	76	nt+1	nt+1	PROPN
alces-263	26	77	=	=	SYM
alces-263	26	78	nts(1+r	nts(1+r	PROPN
alces-263	26	79	)	)	PUNCT
alces-263	26	80	(	(	PUNCT
alces-263	26	81	5	5	NUM
alces-263	26	82	)	)	PUNCT
alces-263	26	83	and	and	CCONJ
alces-263	26	84	λ	λ	X
alces-263	26	85	=	=	SYM
alces-263	26	86	s(1+r	s(1+r	PROPN
alces-263	26	87	)	)	PUNCT
alces-263	26	88	(	(	PUNCT
alces-263	26	89	6	6	X
alces-263	26	90	)	)	PUNCT
alces-263	26	91	i	i	PRON
alces-263	26	92	refer	refer	VERB
alces-263	26	93	to	to	ADP
alces-263	26	94	eq	eq	NOUN
alces-263	26	95	.	.	PROPN
alces-263	26	96	6	6	NUM
alces-263	26	97	as	as	ADP
alces-263	26	98	the	the	DET
alces-263	26	99	type	type	NOUN
alces-263	26	100	2	2	NUM
alces-263	26	101	r	r	NOUN
alces-263	26	102	-	-	PUNCT
alces-263	26	103	m	m	NOUN
alces-263	26	104	equation	equation	NOUN
alces-263	26	105	.	.	PUNCT
alces-263	27	1	the	the	DET
alces-263	27	2	r	r	PROPN
alces-263	27	3	-	-	PUNCT
alces-263	27	4	m	m	NOUN
alces-263	27	5	equation	equation	NOUN
alces-263	27	6	has	have	AUX
alces-263	27	7	also	also	ADV
alces-263	27	8	been	be	AUX
alces-263	27	9	used	use	VERB
alces-263	27	10	to	to	PART
alces-263	27	11	estimate	estimate	VERB
alces-263	27	12	λ	λ	PROPN
alces-263	27	13	for	for	ADP
alces-263	27	14	the	the	DET
alces-263	27	15	female	female	ADJ
alces-263	27	16	segment	segment	NOUN
alces-263	27	17	of	of	ADP
alces-263	27	18	the	the	DET
alces-263	27	19	population	population	NOUN
alces-263	27	20	as	as	SCONJ
alces-263	27	21	growth	growth	NOUN
alces-263	27	22	rates	rate	NOUN
alces-263	27	23	are	be	AUX
alces-263	27	24	largely	largely	ADV
alces-263	27	25	determined	determine	VERB
alces-263	27	26	by	by	ADP
alces-263	27	27	females	female	NOUN
alces-263	27	28	(	(	PUNCT
alces-263	27	29	caughley	caughley	NOUN
alces-263	27	30	1977	1977	NUM
alces-263	27	31	)	)	PUNCT
alces-263	27	32	and	and	CCONJ
alces-263	27	33	because	because	SCONJ
alces-263	27	34	many	many	ADJ
alces-263	27	35	studies	study	NOUN
alces-263	27	36	focus	focus	VERB
alces-263	27	37	on	on	ADP
alces-263	27	38	adult	adult	NOUN
alces-263	27	39	female	female	ADJ
alces-263	27	40	mortality	mortality	NOUN
alces-263	27	41	rates	rate	NOUN
alces-263	27	42	(	(	PUNCT
alces-263	27	43	hervieux	hervieux	PROPN
alces-263	27	44	et	et	PROPN
alces-263	27	45	al	al	PROPN
alces-263	27	46	.	.	PROPN
alces-263	27	47	2013	2013	NUM
alces-263	27	48	,	,	PUNCT
alces-263	27	49	2014	2014	NUM
alces-263	27	50	,	,	PUNCT
alces-263	27	51	kuzyk	kuzyk	INTJ
alces-263	27	52	et	et	PROPN
alces-263	27	53	al	al	PROPN
alces-263	27	54	.	.	PROPN
alces-263	27	55	2019a	2019a	NUM
alces-263	27	56	,	,	PUNCT
alces-263	27	57	2019b	2019b	NUM
alces-263	27	58	)	)	PUNCT
alces-263	27	59	.	.	PUNCT
alces-263	28	1	equations	equation	NOUN
alces-263	28	2	7–12	7–12	PROPN
alces-263	28	3	apply	apply	VERB
alces-263	28	4	specifically	specifically	ADV
alces-263	28	5	to	to	ADP
alces-263	28	6	females	female	NOUN
alces-263	28	7	,	,	PUNCT
alces-263	28	8	although	although	SCONJ
alces-263	28	9	they	they	PRON
alces-263	28	10	are	be	AUX
alces-263	28	11	easily	easily	ADV
alces-263	28	12	modified	modify	VERB
alces-263	28	13	for	for	ADP
alces-263	28	14	males	male	NOUN
alces-263	28	15	or	or	CCONJ
alces-263	28	16	both	both	DET
alces-263	28	17	sexes	sex	NOUN
alces-263	28	18	combined	combine	VERB
alces-263	28	19	(	(	PUNCT
alces-263	28	20	hatter	hatter	NOUN
alces-263	28	21	and	and	CCONJ
alces-263	28	22	bergerud	bergerud	NOUN
alces-263	28	23	1991	1991	NUM
alces-263	28	24	)	)	PUNCT
alces-263	28	25	.	.	PUNCT
alces-263	29	1	recruitment	recruitment	NOUN
alces-263	29	2	in	in	ADP
alces-263	29	3	the	the	DET
alces-263	29	4	type	type	NOUN
alces-263	29	5	1	1	NUM
alces-263	29	6	model	model	NOUN
alces-263	29	7	is	be	AUX
alces-263	29	8	estimated	estimate	VERB
alces-263	29	9	as	as	ADP
alces-263	29	10	:	:	PUNCT
alces-263	29	11	=	=	PUNCT
alces-263	29	12	×	×	NOUN
alces-263	29	13	×	×	NOUN
alces-263	29	14	+	+	CCONJ
alces-263	29	15	r	r	NOUN
alces-263	29	16	j	j	PROPN
alces-263	30	1	j	j	PROPN
alces-263	30	2	c	c	X
alces-263	30	3	/f	/f	PROPN
alces-263	31	1	c	c	NOUN
alces-263	31	2	/f	/f	NOUN
alces-263	31	3	1	1	NUM
alces-263	31	4	(	(	PUNCT
alces-263	31	5	7	7	NUM
alces-263	31	6	)	)	PUNCT
alces-263	31	7	and	and	CCONJ
alces-263	31	8	in	in	ADP
alces-263	31	9	the	the	DET
alces-263	31	10	type	type	NOUN
alces-263	31	11	2	2	NUM
alces-263	31	12	model	model	NOUN
alces-263	31	13	as	as	ADP
alces-263	31	14	r	r	NOUN
alces-263	31	15	=	=	SYM
alces-263	31	16	c	c	PROPN
alces-263	31	17	×	×	PROPN
alces-263	31	18	j	j	PROPN
alces-263	31	19	/	/	SYM
alces-263	31	20	f	f	PROPN
alces-263	31	21	(	(	PUNCT
alces-263	31	22	8)	8)	NUM
alces-263	31	23	where	where	SCONJ
alces-263	31	24	c	c	PROPN
alces-263	31	25	is	be	AUX
alces-263	31	26	the	the	DET
alces-263	31	27	proportion	proportion	NOUN
alces-263	31	28	of	of	ADP
alces-263	31	29	recruited	recruit	VERB
alces-263	31	30	juveniles	juvenile	NOUN
alces-263	31	31	that	that	PRON
alces-263	31	32	are	be	AUX
alces-263	31	33	female	female	ADJ
alces-263	31	34	and	and	CCONJ
alces-263	31	35	j	j	PROPN
alces-263	31	36	/	/	SYM
alces-263	31	37	f	f	PROPN
alces-263	31	38	is	be	AUX
alces-263	31	39	the	the	DET
alces-263	31	40	ratio	ratio	NOUN
alces-263	31	41	of	of	ADP
alces-263	31	42	juveniles/1	juveniles/1	PROPN
alces-263	31	43	+	+	NOUN
alces-263	31	44	year	year	NOUN
alces-263	31	45	-	-	PUNCT
alces-263	31	46	old	old	ADJ
alces-263	31	47	females	female	NOUN
alces-263	31	48	.	.	PUNCT
alces-263	32	1	calf	calf	NOUN
alces-263	32	2	:	:	PUNCT
alces-263	32	3	cow	cow	NOUN
alces-263	32	4	ratios	ratio	NOUN
alces-263	32	5	(	(	PUNCT
alces-263	32	6	j	j	NOUN
alces-263	32	7	/	/	SYM
alces-263	32	8	f	f	PROPN
alces-263	32	9	)	)	PUNCT
alces-263	32	10	for	for	ADP
alces-263	32	11	moose	moose	NOUN
alces-263	32	12	are	be	AUX
alces-263	32	13	usually	usually	ADV
alces-263	32	14	measured	measure	VERB
alces-263	32	15	in	in	ADP
alces-263	32	16	early	early	ADJ
alces-263	32	17	or	or	CCONJ
alces-263	32	18	mid	mid	NOUN
alces-263	32	19	-	-	NOUN
alces-263	32	20	winter	winter	NOUN
alces-263	32	21	when	when	SCONJ
alces-263	32	22	aerial	aerial	ADJ
alces-263	32	23	survey	survey	NOUN
alces-263	32	24	conditions	condition	NOUN
alces-263	32	25	are	be	AUX
alces-263	32	26	optimal	optimal	ADJ
alces-263	32	27	for	for	ADP
alces-263	32	28	determining	determine	VERB
alces-263	32	29	herd	herd	ADJ
alces-263	32	30	composition	composition	NOUN
alces-263	32	31	and	and	CCONJ
alces-263	32	32	abundance	abundance	NOUN
alces-263	32	33	(	(	PUNCT
alces-263	32	34	gasaway	gasaway	NOUN
alces-263	32	35	et	et	PROPN
alces-263	32	36	al	al	PROPN
alces-263	32	37	.	.	PROPN
alces-263	32	38	1986	1986	NUM
alces-263	32	39	)	)	PUNCT
alces-263	32	40	.	.	PUNCT
alces-263	33	1	however	however	ADV
alces-263	33	2	,	,	PUNCT
alces-263	33	3	these	these	DET
alces-263	33	4	surveys	survey	NOUN
alces-263	33	5	do	do	AUX
alces-263	33	6	not	not	PART
alces-263	33	7	provide	provide	VERB
alces-263	33	8	an	an	DET
alces-263	33	9	accurate	accurate	ADJ
alces-263	33	10	measure	measure	NOUN
alces-263	33	11	of	of	ADP
alces-263	33	12	recruitment	recruitment	NOUN
alces-263	33	13	at	at	ADP
alces-263	33	14	1	1	NUM
alces-263	33	15	yearof	yearof	NOUN
alces-263	33	16	-	-	PUNCT
alces-263	33	17	age	age	NOUN
alces-263	33	18	since	since	SCONJ
alces-263	33	19	calves	calf	NOUN
alces-263	33	20	die	die	VERB
alces-263	33	21	at	at	ADP
alces-263	33	22	a	a	DET
alces-263	33	23	higher	high	ADJ
alces-263	33	24	rate	rate	NOUN
alces-263	33	25	than	than	ADP
alces-263	33	26	cows	cow	NOUN
alces-263	33	27	during	during	ADP
alces-263	33	28	winter	winter	NOUN
alces-263	33	29	(	(	PUNCT
alces-263	33	30	ballard	ballard	PROPN
alces-263	33	31	et	et	PROPN
alces-263	33	32	al	al	PROPN
alces-263	33	33	.	.	PROPN
alces-263	33	34	1991	1991	NUM
alces-263	33	35	,	,	PUNCT
alces-263	33	36	kuzyk	kuzyk	INTJ
alces-263	33	37	et	et	PROPN
alces-263	33	38	al	al	PROPN
alces-263	33	39	.	.	PROPN
alces-263	33	40	2019b	2019b	NUM
alces-263	33	41	)	)	PUNCT
alces-263	33	42	.	.	PUNCT
alces-263	34	1	failing	fail	VERB
alces-263	34	2	to	to	PART
alces-263	34	3	account	account	VERB
alces-263	34	4	for	for	ADP
alces-263	34	5	the	the	DET
alces-263	34	6	differential	differential	ADJ
alces-263	34	7	winter	winter	NOUN
alces-263	34	8	mortality	mortality	NOUN
alces-263	34	9	between	between	ADP
alces-263	34	10	calves	calf	NOUN
alces-263	34	11	and	and	CCONJ
alces-263	34	12	cows	cow	NOUN
alces-263	34	13	results	result	NOUN
alces-263	34	14	in	in	ADP
alces-263	34	15	a	a	DET
alces-263	34	16	biased	biased	ADJ
alces-263	34	17	estimate	estimate	NOUN
alces-263	34	18	of	of	ADP
alces-263	34	19	r	r	NOUN
alces-263	34	20	,	,	PUNCT
alces-263	34	21	which	which	PRON
alces-263	34	22	by	by	ADP
alces-263	34	23	definition	definition	NOUN
alces-263	34	24	is	be	AUX
alces-263	34	25	measured	measure	VERB
alces-263	34	26	when	when	SCONJ
alces-263	34	27	calves	calf	NOUN
alces-263	34	28	are	be	AUX
alces-263	34	29	1	1	NUM
alces-263	34	30	-	-	PUNCT
alces-263	34	31	year	year	NOUN
alces-263	34	32	-	-	PUNCT
alces-263	34	33	ofage	ofage	NOUN
alces-263	34	34	.	.	PUNCT
alces-263	35	1	calf	calf	NOUN
alces-263	35	2	:	:	PUNCT
alces-263	35	3	cow	cow	NOUN
alces-263	35	4	ratios	ratio	NOUN
alces-263	35	5	from	from	ADP
alces-263	35	6	these	these	DET
alces-263	35	7	surveys	survey	NOUN
alces-263	35	8	alces	alce	NOUN
alces-263	35	9	vol	vol	NOUN
alces-263	35	10	.	.	PROPN
alces-263	35	11	56	56	NUM
alces-263	35	12	,	,	PUNCT
alces-263	35	13	2020	2020	NUM
alces-263	35	14	revisiting	revisit	VERB
alces-263	35	15	the	the	DET
alces-263	35	16	recruitment	recruitment	NOUN
alces-263	35	17	-	-	PUNCT
alces-263	35	18	mortality	mortality	NOUN
alces-263	35	19	.	.	PUNCT
alces-263	36	1	–	–	PUNCT
alces-263	36	2	ian	ian	PROPN
alces-263	36	3	w.	w.	PROPN
alces-263	36	4	hatter	hatter	PROPN
alces-263	36	5	41	41	NUM
alces-263	36	6	must	must	AUX
alces-263	36	7	be	be	AUX
alces-263	36	8	adjusted	adjust	VERB
alces-263	36	9	to	to	PART
alces-263	36	10	account	account	VERB
alces-263	36	11	for	for	ADP
alces-263	36	12	this	this	DET
alces-263	36	13	differential	differential	ADJ
alces-263	36	14	mortality	mortality	NOUN
alces-263	36	15	.	.	PUNCT
alces-263	37	1	the	the	DET
alces-263	37	2	adjustment	adjustment	NOUN
alces-263	37	3	of	of	ADP
alces-263	37	4	r	r	NOUN
alces-263	37	5	(	(	PUNCT
alces-263	37	6	r	r	NOUN
alces-263	37	7	'	'	PUNCT
alces-263	37	8	)	)	PUNCT
alces-263	37	9	for	for	ADP
alces-263	37	10	the	the	DET
alces-263	37	11	type	type	NOUN
alces-263	37	12	1	1	NUM
alces-263	37	13	model	model	NOUN
alces-263	37	14	is	be	AUX
alces-263	37	15	:	:	PUNCT
alces-263	37	16	=	=	PUNCT
alces-263	37	17	×	×	NOUN
alces-263	37	18	×	×	NOUN
alces-263	37	19	×	×	NOUN
alces-263	37	20	×	×	NOUN
alces-263	37	21	+	+	CCONJ
alces-263	37	22	r	r	NOUN
alces-263	37	23	'	'	PUNCT
alces-263	37	24	c	c	NOUN
alces-263	38	1	j	j	PROPN
alces-263	39	1	f	f	INTJ
alces-263	39	2	sf	sf	INTJ
alces-263	39	3	c	c	PROPN
alces-263	39	4	j	j	PROPN
alces-263	39	5	f	f	X
alces-263	39	6	sj	sj	INTJ
alces-263	39	7	sj	sj	PROPN
alces-263	39	8	/	/	PUNCT
alces-263	39	9	(	(	PUNCT
alces-263	39	10	/	/	SYM
alces-263	39	11	)	)	PUNCT
alces-263	39	12	w	w	PROPN
alces-263	39	13	w	w	PROPN
alces-263	39	14	w	w	PROPN
alces-263	39	15	(	(	PUNCT
alces-263	39	16	9	9	NUM
alces-263	39	17	)	)	PUNCT
alces-263	39	18	and	and	CCONJ
alces-263	39	19	the	the	DET
alces-263	39	20	adjustment	adjustment	NOUN
alces-263	39	21	for	for	ADP
alces-263	39	22	the	the	DET
alces-263	39	23	type	type	NOUN
alces-263	39	24	2	2	NUM
alces-263	39	25	model	model	NOUN
alces-263	39	26	is	be	AUX
alces-263	39	27	:	:	PUNCT
alces-263	39	28	=	=	PUNCT
alces-263	39	29	×	×	NOUN
alces-263	39	30	×r	×r	NOUN
alces-263	39	31	'	'	PART
alces-263	39	32	c	c	NOUN
alces-263	40	1	j	j	PROPN
alces-263	40	2	f	f	X
alces-263	40	3	sj	sj	PROPN
alces-263	40	4	sf/	sf/	PROPN
alces-263	40	5	/w	/w	PUNCT
alces-263	41	1	w	w	PROPN
alces-263	41	2	(	(	PUNCT
alces-263	41	3	10	10	NUM
alces-263	41	4	)	)	PUNCT
alces-263	41	5	where	where	SCONJ
alces-263	41	6	sjw	sjw	PROPN
alces-263	41	7	is	be	AUX
alces-263	41	8	the	the	DET
alces-263	41	9	winter	winter	NOUN
alces-263	41	10	calf	calf	NOUN
alces-263	41	11	survival	survival	NOUN
alces-263	41	12	rate	rate	NOUN
alces-263	41	13	and	and	CCONJ
alces-263	41	14	sfw	sfw	PROPN
alces-263	41	15	is	be	AUX
alces-263	41	16	the	the	DET
alces-263	41	17	winter	winter	NOUN
alces-263	41	18	cow	cow	NOUN
alces-263	41	19	survival	survival	NOUN
alces-263	41	20	rate	rate	NOUN
alces-263	41	21	.	.	PUNCT
alces-263	42	1	these	these	DET
alces-263	42	2	survival	survival	NOUN
alces-263	42	3	rates	rate	NOUN
alces-263	42	4	must	must	AUX
alces-263	42	5	be	be	AUX
alces-263	42	6	measured	measure	VERB
alces-263	42	7	from	from	ADP
alces-263	42	8	the	the	DET
alces-263	42	9	end	end	NOUN
alces-263	42	10	of	of	ADP
alces-263	42	11	the	the	DET
alces-263	42	12	winter	winter	NOUN
alces-263	42	13	survey	survey	NOUN
alces-263	42	14	to	to	PART
alces-263	42	15	just	just	ADV
alces-263	42	16	before	before	SCONJ
alces-263	42	17	calves	calf	NOUN
alces-263	42	18	become	become	VERB
alces-263	42	19	1	1	NUM
alces-263	42	20	year	year	NOUN
alces-263	42	21	-	-	PUNCT
alces-263	42	22	of	of	ADP
alces-263	42	23	-	-	PUNCT
alces-263	42	24	age	age	NOUN
alces-263	42	25	,	,	PUNCT
alces-263	42	26	and	and	CCONJ
alces-263	42	27	is	be	AUX
alces-263	42	28	typically	typically	ADV
alces-263	42	29	accomplished	accomplish	VERB
alces-263	42	30	through	through	ADP
alces-263	42	31	telemetry	telemetry	NOUN
alces-263	42	32	studies	study	NOUN
alces-263	42	33	of	of	ADP
alces-263	42	34	radio	radio	NOUN
alces-263	42	35	-	-	PUNCT
alces-263	42	36	collared	collar	VERB
alces-263	42	37	animals	animal	NOUN
alces-263	42	38	(	(	PUNCT
alces-263	42	39	pollock	pollock	NOUN
alces-263	42	40	et	et	PROPN
alces-263	42	41	al	al	PROPN
alces-263	42	42	.	.	PROPN
alces-263	42	43	1989	1989	NUM
alces-263	42	44	)	)	PUNCT
alces-263	42	45	.	.	PUNCT
alces-263	43	1	the	the	DET
alces-263	43	2	finite	finite	ADJ
alces-263	43	3	rate	rate	NOUN
alces-263	43	4	of	of	ADP
alces-263	43	5	change	change	NOUN
alces-263	43	6	for	for	ADP
alces-263	43	7	the	the	DET
alces-263	43	8	type	type	NOUN
alces-263	43	9	1	1	NUM
alces-263	43	10	model	model	NOUN
alces-263	43	11	is	be	AUX
alces-263	43	12	then	then	ADV
alces-263	43	13	estimated	estimate	VERB
alces-263	43	14	as	as	ADP
alces-263	43	15	:	:	PUNCT
alces-263	43	16	λ	λ	X
alces-263	43	17	=	=	PRON
alces-263	43	18	sf/(1	sf/(1	PUNCT
alces-263	43	19	‒	‒	ADP
alces-263	43	20	r	r	NOUN
alces-263	43	21	'	'	PUNCT
alces-263	43	22	)	)	PUNCT
alces-263	43	23	(	(	PUNCT
alces-263	43	24	11	11	NUM
alces-263	43	25	)	)	PUNCT
alces-263	43	26	and	and	CCONJ
alces-263	43	27	for	for	ADP
alces-263	43	28	the	the	DET
alces-263	43	29	type	type	NOUN
alces-263	43	30	2	2	NUM
alces-263	43	31	model	model	NOUN
alces-263	43	32	as	as	ADP
alces-263	43	33	:	:	PUNCT
alces-263	43	34	λ	λ	X
alces-263	43	35	=	=	PUNCT
alces-263	43	36	sf(1	sf(1	PROPN
alces-263	43	37	+	+	CCONJ
alces-263	43	38	r	r	NOUN
alces-263	43	39	'	'	PUNCT
alces-263	43	40	)	)	PUNCT
alces-263	43	41	(	(	PUNCT
alces-263	43	42	12	12	NUM
alces-263	43	43	)	)	PUNCT
alces-263	43	44	where	where	SCONJ
alces-263	43	45	sf	sf	PROPN
alces-263	43	46	is	be	AUX
alces-263	43	47	the	the	DET
alces-263	43	48	annual	annual	ADJ
alces-263	43	49	cow	cow	NOUN
alces-263	43	50	survival	survival	NOUN
alces-263	43	51	rate	rate	NOUN
alces-263	43	52	.	.	PUNCT
alces-263	44	1	figure	figure	NOUN
alces-263	44	2	1	1	NUM
alces-263	44	3	illustrates	illustrate	VERB
alces-263	44	4	estimates	estimate	NOUN
alces-263	44	5	of	of	ADP
alces-263	44	6	change	change	NOUN
alces-263	44	7	in	in	ADP
alces-263	44	8	λ	λ	PROPN
alces-263	44	9	when	when	SCONJ
alces-263	44	10	there	there	PRON
alces-263	44	11	is	be	VERB
alces-263	44	12	differential	differential	ADJ
alces-263	44	13	overwinter	overwinter	ADJ
alces-263	44	14	mortality	mortality	NOUN
alces-263	44	15	between	between	ADP
alces-263	44	16	cows	cow	NOUN
alces-263	44	17	and	and	CCONJ
alces-263	44	18	calves	calf	NOUN
alces-263	44	19	(	(	PUNCT
alces-263	44	20	sjw	sjw	PROPN
alces-263	44	21	/sfw	/sfw	PROPN
alces-263	44	22	)	)	PUNCT
alces-263	44	23	.	.	PUNCT
alces-263	45	1	for	for	ADP
alces-263	45	2	example	example	NOUN
alces-263	45	3	,	,	PUNCT
alces-263	45	4	λ	λ	PRON
alces-263	45	5	ranged	range	VERB
alces-263	45	6	from	from	ADP
alces-263	45	7	0.98	0.98	NUM
alces-263	45	8	(	(	PUNCT
alces-263	45	9	sjw	sjw	NOUN
alces-263	45	10	/sfw	/sfw	PROPN
alces-263	46	1	=	=	PUNCT
alces-263	46	2	0.50	0.50	NUM
alces-263	46	3	)	)	PUNCT
alces-263	46	4	to	to	ADP
alces-263	46	5	1.06	1.06	NUM
alces-263	46	6	(	(	PUNCT
alces-263	46	7	sjw	sjw	PROPN
alces-263	46	8	/sfw	/sfw	PROPN
alces-263	46	9	=	=	PUNCT
alces-263	46	10	1.00	1.00	NUM
alces-263	46	11	)	)	PUNCT
alces-263	46	12	when	when	SCONJ
alces-263	46	13	assuming	assume	VERB
alces-263	46	14	j	j	PROPN
alces-263	46	15	/	/	SYM
alces-263	46	16	f	f	PROPN
alces-263	46	17	=	=	SYM
alces-263	46	18	0.35	0.35	NUM
alces-263	46	19	,	,	PUNCT
alces-263	46	20	c	c	NOUN
alces-263	46	21	=	=	SYM
alces-263	46	22	0.5	0.5	NUM
alces-263	46	23	,	,	PUNCT
alces-263	46	24	and	and	CCONJ
alces-263	46	25	sf	sf	NOUN
alces-263	46	26	=	=	NOUN
alces-263	46	27	0.90	0.90	NUM
alces-263	46	28	.	.	PUNCT
alces-263	47	1	the	the	DET
alces-263	47	2	above	above	ADJ
alces-263	47	3	equations	equation	NOUN
alces-263	47	4	do	do	AUX
alces-263	47	5	not	not	PART
alces-263	47	6	account	account	VERB
alces-263	47	7	for	for	ADP
alces-263	47	8	uncertainty	uncertainty	NOUN
alces-263	47	9	in	in	ADP
alces-263	47	10	the	the	DET
alces-263	47	11	parameters	parameter	NOUN
alces-263	47	12	used	use	VERB
alces-263	47	13	to	to	PART
alces-263	47	14	estimate	estimate	VERB
alces-263	47	15	λ	λ	PROPN
alces-263	47	16	.	.	PUNCT
alces-263	48	1	in	in	ADP
alces-263	48	2	order	order	NOUN
alces-263	48	3	to	to	PART
alces-263	48	4	determine	determine	VERB
alces-263	48	5	the	the	DET
alces-263	48	6	95	95	NUM
alces-263	48	7	%	%	NOUN
alces-263	48	8	ci	ci	NOUN
alces-263	48	9	for	for	ADP
alces-263	48	10	λ	λ	PROPN
alces-263	48	11	in	in	ADP
alces-263	48	12	eq	eq	ADP
alces-263	48	13	.	.	PROPN
alces-263	48	14	11	11	NUM
alces-263	48	15	or	or	CCONJ
alces-263	48	16	12	12	NUM
alces-263	48	17	,	,	PUNCT
alces-263	48	18	the	the	DET
alces-263	48	19	se	se	PROPN
alces-263	48	20	must	must	AUX
alces-263	48	21	be	be	AUX
alces-263	48	22	measured	measure	VERB
alces-263	48	23	for	for	ADP
alces-263	48	24	j	j	PROPN
alces-263	48	25	/	/	SYM
alces-263	48	26	f	f	PROPN
alces-263	48	27	,	,	PUNCT
alces-263	48	28	sf	sf	PROPN
alces-263	48	29	,	,	PUNCT
alces-263	48	30	sfw	sfw	PROPN
alces-263	48	31	,	,	PUNCT
alces-263	48	32	sjw	sjw	PROPN
alces-263	48	33	,	,	PUNCT
alces-263	48	34	and	and	CCONJ
alces-263	48	35	c.	c.	PROPN
alces-263	48	36	a	a	DET
alces-263	48	37	number	number	NOUN
alces-263	48	38	of	of	ADP
alces-263	48	39	researchers	researcher	NOUN
alces-263	48	40	including	include	VERB
alces-263	48	41	caughley	caughley	NOUN
alces-263	48	42	(	(	PUNCT
alces-263	48	43	1977	1977	NUM
alces-263	48	44	)	)	PUNCT
alces-263	48	45	,	,	PUNCT
alces-263	48	46	gasaway	gasaway	NOUN
alces-263	48	47	et	et	PROPN
alces-263	48	48	al	al	PROPN
alces-263	48	49	.	.	PROPN
alces-263	49	1	(	(	PUNCT
alces-263	49	2	1986	1986	NUM
alces-263	49	3	)	)	PUNCT
alces-263	49	4	,	,	PUNCT
alces-263	49	5	skalski	skalski	NOUN
alces-263	49	6	et	et	PROPN
alces-263	49	7	al	al	PROPN
alces-263	49	8	.	.	PUNCT
alces-263	50	1	(	(	PUNCT
alces-263	50	2	2005	2005	NUM
alces-263	50	3	)	)	PUNCT
alces-263	50	4	,	,	PUNCT
alces-263	50	5	and	and	CCONJ
alces-263	50	6	pollock	pollock	NOUN
alces-263	50	7	et	et	PROPN
alces-263	50	8	al	al	PROPN
alces-263	50	9	.	.	PROPN
alces-263	51	1	(	(	PUNCT
alces-263	51	2	1989	1989	NUM
alces-263	51	3	)	)	PUNCT
alces-263	51	4	provide	provide	VERB
alces-263	51	5	methods	method	NOUN
alces-263	51	6	and	and	CCONJ
alces-263	51	7	examples	example	NOUN
alces-263	51	8	for	for	ADP
alces-263	51	9	making	make	VERB
alces-263	51	10	these	these	DET
alces-263	51	11	calculations	calculation	NOUN
alces-263	51	12	.	.	PUNCT
alces-263	52	1	fig	fig	NOUN
alces-263	52	2	.	.	PUNCT
alces-263	53	1	1	1	X
alces-263	53	2	.	.	X
alces-263	53	3	contour	contour	NOUN
alces-263	53	4	plot	plot	NOUN
alces-263	53	5	depicting	depict	VERB
alces-263	53	6	the	the	DET
alces-263	53	7	range	range	NOUN
alces-263	53	8	of	of	ADP
alces-263	53	9	population	population	NOUN
alces-263	53	10	growth	growth	NOUN
alces-263	53	11	rates	rate	NOUN
alces-263	53	12	(	(	PUNCT
alces-263	53	13	λ	λ	X
alces-263	53	14	)	)	PUNCT
alces-263	53	15	based	base	VERB
alces-263	53	16	on	on	ADP
alces-263	53	17	plausible	plausible	ADJ
alces-263	53	18	ranges	range	NOUN
alces-263	53	19	in	in	ADP
alces-263	53	20	annual	annual	ADJ
alces-263	53	21	cow	cow	NOUN
alces-263	53	22	survival	survival	NOUN
alces-263	53	23	(	(	PUNCT
alces-263	53	24	sf	sf	PROPN
alces-263	53	25	)	)	PUNCT
alces-263	53	26	and	and	CCONJ
alces-263	53	27	differential	differential	ADJ
alces-263	53	28	winter	winter	NOUN
alces-263	53	29	mortality	mortality	NOUN
alces-263	53	30	between	between	ADP
alces-263	53	31	calves	calf	NOUN
alces-263	53	32	and	and	CCONJ
alces-263	53	33	cows	cow	NOUN
alces-263	53	34	(	(	PUNCT
alces-263	53	35	sjw	sjw	PROPN
alces-263	53	36	/	/	SYM
alces-263	53	37	sfw	sfw	PROPN
alces-263	53	38	)	)	PUNCT
alces-263	53	39	.	.	PUNCT
alces-263	54	1	the	the	DET
alces-263	54	2	midwinter	midwinter	PROPN
alces-263	54	3	calf	calf	NOUN
alces-263	54	4	:	:	PUNCT
alces-263	54	5	cow	cow	NOUN
alces-263	54	6	ratio	ratio	NOUN
alces-263	54	7	(	(	PUNCT
alces-263	54	8	j	j	PROPN
alces-263	54	9	/	/	SYM
alces-263	54	10	f	f	PROPN
alces-263	54	11	)	)	PUNCT
alces-263	54	12	was	be	AUX
alces-263	54	13	0.35	0.35	NUM
alces-263	54	14	,	,	PUNCT
alces-263	54	15	and	and	CCONJ
alces-263	54	16	the	the	DET
alces-263	54	17	calf	calf	NOUN
alces-263	54	18	sex	sex	NOUN
alces-263	54	19	ratio	ratio	NOUN
alces-263	54	20	was	be	AUX
alces-263	54	21	50:50	50:50	NUM
alces-263	54	22	.	.	PUNCT
alces-263	55	1	revisiting	revisit	VERB
alces-263	55	2	the	the	DET
alces-263	55	3	r	r	NOUN
alces-263	55	4	-	-	PUNCT
alces-263	55	5	m	m	NOUN
alces-263	55	6	equation	equation	NOUN
alces-263	55	7	.	.	PUNCT
alces-263	56	1	–	–	PUNCT
alces-263	56	2	ian	ian	PROPN
alces-263	56	3	w.	w.	PROPN
alces-263	56	4	hatter	hatter	PROPN
alces-263	56	5	alces	alces	PROPN
alces-263	56	6	vol	vol	NOUN
alces-263	56	7	.	.	PROPN
alces-263	56	8	56	56	NUM
alces-263	56	9	,	,	PUNCT
alces-263	56	10	2020	2020	NUM
alces-263	56	11	42	42	NUM
alces-263	56	12	following	follow	VERB
alces-263	56	13	latham	latham	PROPN
alces-263	56	14	et	et	PROPN
alces-263	56	15	al	al	PROPN
alces-263	56	16	.	.	PROPN
alces-263	57	1	(	(	PUNCT
alces-263	57	2	2011	2011	NUM
alces-263	57	3	)	)	PUNCT
alces-263	57	4	and	and	CCONJ
alces-263	57	5	hervieux	hervieux	PROPN
alces-263	57	6	et	et	PROPN
alces-263	57	7	al	al	PROPN
alces-263	57	8	.	.	PROPN
alces-263	58	1	(	(	PUNCT
alces-263	58	2	2013	2013	NUM
alces-263	58	3	)	)	PUNCT
alces-263	58	4	,	,	PUNCT
alces-263	58	5	one	one	NUM
alces-263	58	6	method	method	NOUN
alces-263	58	7	of	of	ADP
alces-263	58	8	estimating	estimate	VERB
alces-263	58	9	the	the	DET
alces-263	58	10	95	95	NUM
alces-263	58	11	%	%	NOUN
alces-263	58	12	ci	ci	NOUN
alces-263	58	13	for	for	ADP
alces-263	58	14	λ	λ	PROPN
alces-263	58	15	is	be	AUX
alces-263	58	16	to	to	PART
alces-263	58	17	randomly	randomly	VERB
alces-263	58	18	draw	draw	VERB
alces-263	58	19	from	from	ADP
alces-263	58	20	each	each	DET
alces-263	58	21	year	year	NOUN
alces-263	58	22	’s	’s	PART
alces-263	58	23	annual	annual	ADJ
alces-263	58	24	survival	survival	NOUN
alces-263	58	25	and	and	CCONJ
alces-263	58	26	recruitment	recruitment	NOUN
alces-263	58	27	distributions	distribution	NOUN
alces-263	58	28	(	(	PUNCT
alces-263	58	29	i.e.	i.e.	X
alces-263	58	30	,	,	PUNCT
alces-263	58	31	x̅	x̅	PROPN
alces-263	58	32	and	and	CCONJ
alces-263	58	33	se	se	X
alces-263	58	34	)	)	PUNCT
alces-263	58	35	a	a	DET
alces-263	58	36	large	large	ADJ
alces-263	58	37	number	number	NOUN
alces-263	58	38	of	of	ADP
alces-263	58	39	times	time	NOUN
alces-263	58	40	(	(	PUNCT
alces-263	58	41	e.g.	e.g.	ADV
alces-263	58	42	,	,	PUNCT
alces-263	58	43	10,000	10,000	NUM
alces-263	58	44	)	)	PUNCT
alces-263	58	45	using	use	VERB
alces-263	58	46	monte	monte	PROPN
alces-263	58	47	carlo	carlo	PROPN
alces-263	58	48	simulation	simulation	PROPN
alces-263	58	49	.	.	PUNCT
alces-263	59	1	survival	survival	NOUN
alces-263	59	2	rates	rate	NOUN
alces-263	59	3	should	should	AUX
alces-263	59	4	be	be	AUX
alces-263	59	5	drawn	draw	VERB
alces-263	59	6	from	from	ADP
alces-263	59	7	a	a	DET
alces-263	59	8	beta	beta	ADJ
alces-263	59	9	distribution	distribution	NOUN
alces-263	59	10	(	(	PUNCT
alces-263	59	11	values	value	NOUN
alces-263	59	12	range	range	VERB
alces-263	59	13	from	from	ADP
alces-263	59	14	0–1	0–1	NOUN
alces-263	59	15	)	)	PUNCT
alces-263	59	16	and	and	CCONJ
alces-263	59	17	calf	calf	NOUN
alces-263	59	18	:	:	PUNCT
alces-263	59	19	cow	cow	PROPN
alces-263	59	20	(	(	PUNCT
alces-263	59	21	j	j	PROPN
alces-263	59	22	/	/	SYM
alces-263	59	23	f	f	PROPN
alces-263	59	24	)	)	PUNCT
alces-263	59	25	ratios	ratio	NOUN
alces-263	59	26	from	from	ADP
alces-263	59	27	a	a	DET
alces-263	59	28	lognormal	lognormal	ADJ
alces-263	59	29	distribution	distribution	NOUN
alces-263	59	30	(	(	PUNCT
alces-263	59	31	values	value	NOUN
alces-263	59	32	>	>	X
alces-263	59	33	0	0	NUM
alces-263	59	34	)	)	PUNCT
alces-263	59	35	.	.	PUNCT
alces-263	60	1	for	for	ADP
alces-263	60	2	a	a	DET
alces-263	60	3	quick	quick	ADJ
alces-263	60	4	and	and	CCONJ
alces-263	60	5	simple	simple	ADJ
alces-263	60	6	comparison	comparison	NOUN
alces-263	60	7	,	,	PUNCT
alces-263	60	8	the	the	DET
alces-263	60	9	normal	normal	ADJ
alces-263	60	10	approximation	approximation	NOUN
alces-263	60	11	may	may	AUX
alces-263	60	12	be	be	AUX
alces-263	60	13	used	use	VERB
alces-263	60	14	to	to	PART
alces-263	60	15	determine	determine	VERB
alces-263	60	16	if	if	SCONJ
alces-263	60	17	two	two	NUM
alces-263	60	18	estimates	estimate	NOUN
alces-263	60	19	(	(	PUNCT
alces-263	60	20	est	est	X
alces-263	60	21	)	)	PUNCT
alces-263	60	22	(	(	PUNCT
alces-263	60	23	e.g.	e.g.	ADV
alces-263	60	24	,	,	PUNCT
alces-263	60	25	j	j	PROPN
alces-263	60	26	/	/	SYM
alces-263	60	27	f	f	PROPN
alces-263	60	28	,	,	PUNCT
alces-263	60	29	sf	sf	PROPN
alces-263	60	30	,	,	PUNCT
alces-263	60	31	sfw	sfw	PROPN
alces-263	60	32	,	,	PUNCT
alces-263	60	33	sjw	sjw	PROPN
alces-263	60	34	,	,	PUNCT
alces-263	60	35	or	or	CCONJ
alces-263	60	36	λ	λ	NOUN
alces-263	60	37	)	)	PUNCT
alces-263	60	38	are	be	AUX
alces-263	60	39	significantly	significantly	ADV
alces-263	60	40	different	different	ADJ
alces-263	60	41	using	use	VERB
alces-263	60	42	=	=	PUNCT
alces-263	60	43	−	−	PROPN
alces-263	61	1	+	+	CCONJ
alces-263	61	2	>	>	X
alces-263	61	3	z	z	PROPN
alces-263	61	4	est	est	X
alces-263	61	5	est	est	X
alces-263	61	6	var	var	PROPN
alces-263	61	7	est	est	X
alces-263	61	8	var	var	PROPN
alces-263	61	9	est	est	X
alces-263	61	10	(	(	PUNCT
alces-263	61	11	)	)	PUNCT
alces-263	61	12	(	(	PUNCT
alces-263	61	13	(	(	PUNCT
alces-263	61	14	)	)	PUNCT
alces-263	61	15	(	(	PUNCT
alces-263	61	16	)	)	PUNCT
alces-263	61	17	)	)	PUNCT
alces-263	61	18	1.961	1.961	NUM
alces-263	61	19	2	2	NUM
alces-263	61	20	1	1	NUM
alces-263	61	21	2	2	NUM
alces-263	61	22	where	where	SCONJ
alces-263	61	23	var	var	NOUN
alces-263	61	24	=	=	SYM
alces-263	61	25	se2	se2	PROPN
alces-263	61	26	(	(	PUNCT
alces-263	61	27	sinclair	sinclair	PROPN
alces-263	61	28	et	et	PROPN
alces-263	61	29	al	al	PROPN
alces-263	61	30	.	.	PROPN
alces-263	61	31	2006	2006	NUM
alces-263	61	32	)	)	PUNCT
alces-263	61	33	.	.	PUNCT
alces-263	62	1	if	if	SCONJ
alces-263	62	2	estimates	estimate	NOUN
alces-263	62	3	are	be	AUX
alces-263	62	4	not	not	PART
alces-263	62	5	significantly	significantly	ADV
alces-263	62	6	different	different	ADJ
alces-263	62	7	,	,	PUNCT
alces-263	62	8	they	they	PRON
alces-263	62	9	may	may	AUX
alces-263	62	10	be	be	AUX
alces-263	62	11	merged	merge	VERB
alces-263	62	12	to	to	PART
alces-263	62	13	produce	produce	VERB
alces-263	62	14	a	a	DET
alces-263	62	15	more	more	ADV
alces-263	62	16	precise	precise	ADJ
alces-263	62	17	estimate	estimate	NOUN
alces-263	62	18	of	of	ADP
alces-263	62	19	λ	λ	NOUN
alces-263	62	20	using	use	VERB
alces-263	62	21	the	the	DET
alces-263	62	22	procedure	procedure	NOUN
alces-263	62	23	outlined	outline	VERB
alces-263	62	24	by	by	ADP
alces-263	62	25	sinclair	sinclair	PROPN
alces-263	62	26	et	et	PROPN
alces-263	62	27	al	al	PROPN
alces-263	62	28	.	.	PROPN
alces-263	63	1	(	(	PUNCT
alces-263	63	2	2006:234	2006:234	NUM
alces-263	63	3	)	)	PUNCT
alces-263	63	4	.	.	PUNCT
alces-263	64	1	an	an	DET
alces-263	64	2	example	example	NOUN
alces-263	64	3	factors	factor	NOUN
alces-263	64	4	affecting	affect	VERB
alces-263	64	5	moose	moose	NOUN
alces-263	64	6	population	population	NOUN
alces-263	64	7	declines	decline	VERB
alces-263	64	8	in	in	ADP
alces-263	64	9	british	british	PROPN
alces-263	64	10	columbia	columbia	PROPN
alces-263	64	11	,	,	PUNCT
alces-263	64	12	canada	canada	PROPN
alces-263	64	13	are	be	AUX
alces-263	64	14	currently	currently	ADV
alces-263	64	15	being	be	AUX
alces-263	64	16	investigated	investigate	VERB
alces-263	64	17	with	with	ADP
alces-263	64	18	population	population	NOUN
alces-263	64	19	dynamics	dynamic	NOUN
alces-263	64	20	studied	study	VERB
alces-263	64	21	intensively	intensively	ADV
alces-263	64	22	(	(	PUNCT
alces-263	64	23	kuzyk	kuzyk	INTJ
alces-263	64	24	et	et	PROPN
alces-263	64	25	al	al	PROPN
alces-263	64	26	.	.	PROPN
alces-263	64	27	2019a	2019a	NUM
alces-263	64	28	,	,	PUNCT
alces-263	64	29	2019b	2019b	NUM
alces-263	64	30	)	)	PUNCT
alces-263	64	31	.	.	PUNCT
alces-263	65	1	estimates	estimate	NOUN
alces-263	65	2	of	of	ADP
alces-263	65	3	j	j	PROPN
alces-263	65	4	/	/	SYM
alces-263	65	5	f	f	PROPN
alces-263	65	6	,	,	PUNCT
alces-263	65	7	sf	sf	PROPN
alces-263	65	8	,	,	PUNCT
alces-263	65	9	sfw	sfw	NOUN
alces-263	65	10	,	,	PUNCT
alces-263	65	11	and	and	CCONJ
alces-263	65	12	sjw	sjw	PROPN
alces-263	65	13	were	be	AUX
alces-263	65	14	available	available	ADJ
alces-263	65	15	for	for	ADP
alces-263	65	16	3	3	NUM
alces-263	65	17	consecutive	consecutive	ADJ
alces-263	65	18	years	year	NOUN
alces-263	65	19	in	in	ADP
alces-263	65	20	the	the	DET
alces-263	65	21	bonaparte	bonaparte	PROPN
alces-263	65	22	study	study	PROPN
alces-263	65	23	area	area	NOUN
alces-263	65	24	(	(	PUNCT
alces-263	65	25	2016–17	2016–17	NOUN
alces-263	65	26	to	to	ADP
alces-263	65	27	2018–19	2018–19	NUM
alces-263	65	28	)	)	PUNCT
alces-263	65	29	and	and	CCONJ
alces-263	65	30	2	2	NUM
alces-263	65	31	consecutive	consecutive	ADJ
alces-263	65	32	years	year	NOUN
alces-263	65	33	in	in	ADP
alces-263	65	34	the	the	DET
alces-263	65	35	prince	prince	PROPN
alces-263	65	36	george	george	PROPN
alces-263	65	37	south	south	PROPN
alces-263	65	38	study	study	PROPN
alces-263	65	39	area	area	NOUN
alces-263	65	40	(	(	PUNCT
alces-263	65	41	2017–18	2017–18	NUM
alces-263	65	42	to	to	ADP
alces-263	65	43	2018–19	2018–19	NUM
alces-263	65	44	)	)	PUNCT
alces-263	65	45	(	(	PUNCT
alces-263	65	46	table	table	NOUN
alces-263	65	47	1	1	NUM
alces-263	65	48	)	)	PUNCT
alces-263	65	49	,	,	PUNCT
alces-263	65	50	and	and	CCONJ
alces-263	65	51	the	the	DET
alces-263	65	52	proportion	proportion	NOUN
alces-263	65	53	of	of	ADP
alces-263	65	54	8	8	NUM
alces-263	65	55	-	-	PUNCT
alces-263	65	56	month	month	NOUN
alces-263	65	57	-	-	PUNCT
alces-263	65	58	old	old	ADJ
alces-263	65	59	female	female	ADJ
alces-263	65	60	calves	calf	NOUN
alces-263	65	61	(	(	PUNCT
alces-263	65	62	c	c	NOUN
alces-263	65	63	)	)	PUNCT
alces-263	65	64	was	be	AUX
alces-263	65	65	documented	document	VERB
alces-263	65	66	from	from	ADP
alces-263	65	67	the	the	DET
alces-263	65	68	sex	sex	NOUN
alces-263	65	69	ratio	ratio	NOUN
alces-263	65	70	of	of	ADP
alces-263	65	71	radio	radio	NOUN
alces-263	65	72	-	-	PUNCT
alces-263	65	73	collared	collar	VERB
alces-263	65	74	calves	calf	NOUN
alces-263	65	75	from	from	ADP
alces-263	65	76	2016–17	2016–17	NUM
alces-263	65	77	to	to	ADP
alces-263	65	78	2019–20	2019–20	NUM
alces-263	65	79	.	.	PUNCT
alces-263	66	1	calf	calf	NOUN
alces-263	66	2	sex	sex	NOUN
alces-263	66	3	ratios	ratio	NOUN
alces-263	66	4	were	be	AUX
alces-263	66	5	not	not	PART
alces-263	66	6	significantly	significantly	ADV
alces-263	66	7	different	different	ADJ
alces-263	66	8	from	from	ADP
alces-263	66	9	50:50	50:50	NUM
alces-263	66	10	in	in	ADP
alces-263	66	11	either	either	CCONJ
alces-263	66	12	study	study	NOUN
alces-263	66	13	area	area	NOUN
alces-263	66	14	(	(	PUNCT
alces-263	66	15	bonaparte	bonaparte	NOUN
alces-263	66	16	:	:	PUNCT
alces-263	66	17	χ2	χ2	PROPN
alces-263	66	18	=	=	PUNCT
alces-263	66	19	0.45	0.45	NUM
alces-263	66	20	,	,	PUNCT
alces-263	66	21	p	p	X
alces-263	66	22	=	=	NOUN
alces-263	66	23	0.50	0.50	NUM
alces-263	66	24	,	,	PUNCT
alces-263	66	25	n	n	PROPN
alces-263	66	26	=	=	SYM
alces-263	66	27	80	80	NUM
alces-263	66	28	;	;	PUNCT
alces-263	66	29	prince	prince	PROPN
alces-263	66	30	george	george	PROPN
alces-263	66	31	south	south	PROPN
alces-263	66	32	:	:	PUNCT
alces-263	67	1	χ2	χ2	PROPN
alces-263	67	2	=	=	PROPN
alces-263	67	3	0.24	0.24	NUM
alces-263	67	4	,	,	PUNCT
alces-263	67	5	p	p	X
alces-263	67	6	=	=	NOUN
alces-263	67	7	0.62	0.62	NUM
alces-263	67	8	,	,	PUNCT
alces-263	67	9	n	n	NOUN
alces-263	67	10	=	=	NUM
alces-263	67	11	66	66	NUM
alces-263	67	12	)	)	PUNCT
alces-263	67	13	,	,	PUNCT
alces-263	67	14	so	so	SCONJ
alces-263	67	15	c	c	PROPN
alces-263	67	16	was	be	AUX
alces-263	67	17	set	set	VERB
alces-263	67	18	equal	equal	ADJ
alces-263	67	19	to	to	ADP
alces-263	67	20	0.5	0.5	NUM
alces-263	67	21	.	.	PUNCT
alces-263	68	1	differential	differential	ADJ
alces-263	68	2	winter	winter	NOUN
alces-263	68	3	survival	survival	NOUN
alces-263	68	4	(	(	PUNCT
alces-263	68	5	sjw	sjw	PROPN
alces-263	68	6	/	/	SYM
alces-263	68	7	sfw	sfw	PROPN
alces-263	68	8	)	)	PUNCT
alces-263	68	9	varied	varied	ADJ
alces-263	68	10	from	from	ADP
alces-263	68	11	0.47–0.89	0.47–0.89	NUM
alces-263	68	12	in	in	ADP
alces-263	68	13	the	the	DET
alces-263	68	14	bonaparte	bonaparte	PROPN
alces-263	68	15	study	study	NOUN
alces-263	68	16	area	area	NOUN
alces-263	68	17	and	and	CCONJ
alces-263	68	18	from	from	ADP
alces-263	68	19	0.78–0.87	0.78–0.87	PROPN
alces-263	68	20	in	in	ADP
alces-263	68	21	the	the	DET
alces-263	68	22	prince	prince	PROPN
alces-263	68	23	george	george	PROPN
alces-263	68	24	south	south	PROPN
alces-263	68	25	area	area	NOUN
alces-263	68	26	.	.	PUNCT
alces-263	69	1	growth	growth	NOUN
alces-263	69	2	rates	rate	NOUN
alces-263	69	3	based	base	VERB
alces-263	69	4	on	on	ADP
alces-263	69	5	unadjusted	unadjusted	ADJ
alces-263	69	6	recruitment	recruitment	NOUN
alces-263	69	7	rates	rate	NOUN
alces-263	69	8	from	from	ADP
alces-263	69	9	the	the	DET
alces-263	69	10	hatter	hatter	NOUN
alces-263	69	11	and	and	CCONJ
alces-263	69	12	table	table	NOUN
alces-263	69	13	1	1	NUM
alces-263	69	14	.	.	NOUN
alces-263	69	15	moose	moose	NOUN
alces-263	69	16	population	population	NOUN
alces-263	69	17	parameters	parameter	NOUN
alces-263	69	18	including	include	VERB
alces-263	69	19	calf	calf	NOUN
alces-263	69	20	:	:	PUNCT
alces-263	69	21	cow	cow	NOUN
alces-263	69	22	ratio	ratio	NOUN
alces-263	69	23	(	(	PUNCT
alces-263	69	24	j	j	PROPN
alces-263	69	25	/	/	SYM
alces-263	69	26	f	f	PROPN
alces-263	69	27	)	)	PUNCT
alces-263	69	28	,	,	PUNCT
alces-263	69	29	annual	annual	ADJ
alces-263	69	30	cow	cow	NOUN
alces-263	69	31	survival	survival	NOUN
alces-263	69	32	rate	rate	PROPN
alces-263	69	33	(	(	PUNCT
alces-263	69	34	sf	sf	PROPN
alces-263	69	35	)	)	PUNCT
alces-263	69	36	,	,	PUNCT
alces-263	69	37	winter	winter	NOUN
alces-263	69	38	cow	cow	NOUN
alces-263	69	39	survival	survival	NOUN
alces-263	69	40	rate	rate	NOUN
alces-263	69	41	(	(	PUNCT
alces-263	69	42	sfw	sfw	PROPN
alces-263	69	43	)	)	PUNCT
alces-263	69	44	,	,	PUNCT
alces-263	69	45	and	and	CCONJ
alces-263	69	46	winter	winter	NOUN
alces-263	69	47	calf	calf	NOUN
alces-263	69	48	survival	survival	NOUN
alces-263	69	49	rate	rate	NOUN
alces-263	69	50	(	(	PUNCT
alces-263	69	51	sjw	sjw	PROPN
alces-263	69	52	)	)	PUNCT
alces-263	69	53	within	within	ADP
alces-263	69	54	the	the	DET
alces-263	69	55	bonaparte	bonaparte	NOUN
alces-263	69	56	and	and	CCONJ
alces-263	69	57	prince	prince	PROPN
alces-263	69	58	george	george	PROPN
alces-263	69	59	south	south	PROPN
alces-263	69	60	study	study	PROPN
alces-263	69	61	areas	area	NOUN
alces-263	69	62	of	of	ADP
alces-263	69	63	british	british	PROPN
alces-263	69	64	columbia	columbia	PROPN
alces-263	69	65	,	,	PUNCT
alces-263	69	66	canada	canada	PROPN
alces-263	69	67	;	;	PUNCT
alces-263	69	68	data	datum	NOUN
alces-263	69	69	is	be	AUX
alces-263	69	70	from	from	ADP
alces-263	69	71	kuzyk	kuzyk	INTJ
alces-263	69	72	et	et	PROPN
alces-263	69	73	al	al	PROPN
alces-263	69	74	.	.	PROPN
alces-263	70	1	(	(	PUNCT
alces-263	70	2	2019	2019	NUM
alces-263	70	3	)	)	PUNCT
alces-263	70	4	.	.	PUNCT
alces-263	71	1	merged	merge	VERB
alces-263	71	2	refers	refer	NOUN
alces-263	71	3	to	to	ADP
alces-263	71	4	combined	combine	VERB
alces-263	71	5	estimates	estimate	NOUN
alces-263	71	6	for	for	ADP
alces-263	71	7	2017–18	2017–18	NUM
alces-263	71	8	and	and	CCONJ
alces-263	71	9	2018–19	2018–19	NOUN
alces-263	71	10	.	.	PUNCT
alces-263	72	1	a.	a.	PROPN
alces-263	72	2	bonaparte	bonaparte	PROPN
alces-263	72	3	study	study	PROPN
alces-263	72	4	area	area	PROPN
alces-263	72	5	year	year	NOUN
alces-263	72	6	calf	calf	NOUN
alces-263	72	7	:	:	PUNCT
alces-263	72	8	cow	cow	NOUN
alces-263	72	9	ratio	ratio	NOUN
alces-263	72	10	annual	annual	ADJ
alces-263	72	11	cow	cow	NOUN
alces-263	72	12	survival	survival	PROPN
alces-263	72	13	winter	winter	PROPN
alces-263	72	14	cow	cow	PROPN
alces-263	72	15	survival	survival	PROPN
alces-263	72	16	winter	winter	PROPN
alces-263	72	17	calf	calf	PROPN
alces-263	72	18	survival	survival	PROPN
alces-263	72	19	j	j	PROPN
alces-263	72	20	/	/	SYM
alces-263	72	21	f	f	PROPN
alces-263	72	22	se	se	PROPN
alces-263	72	23	n	n	PROPN
alces-263	72	24	sf	sf	PROPN
alces-263	72	25	se	se	X
alces-263	72	26	n	n	X
alces-263	72	27	sfw	sfw	PROPN
alces-263	72	28	se	se	PROPN
alces-263	72	29	n	n	X
alces-263	72	30	sjw	sjw	PROPN
alces-263	72	31	se	se	PROPN
alces-263	72	32	n	n	PROPN
alces-263	72	33	2016–17	2016–17	NOUN
alces-263	72	34	0.13	0.13	NUM
alces-263	72	35	0.028	0.028	NUM
alces-263	72	36	208	208	NUM
alces-263	72	37	0.91	0.91	NUM
alces-263	72	38	0.057	0.057	NUM
alces-263	72	39	79	79	NUM
alces-263	72	40	0.96	0.96	NUM
alces-263	72	41	0.028	0.028	NUM
alces-263	72	42	52	52	NUM
alces-263	72	43	0.45	0.45	NUM
alces-263	72	44	0.051	0.051	NUM
alces-263	72	45	20	20	NUM
alces-263	72	46	2017–18	2017–18	NUM
alces-263	72	47	0.32	0.32	NUM
alces-263	72	48	0.047	0.047	NUM
alces-263	72	49	256	256	NUM
alces-263	72	50	0.98	0.98	NUM
alces-263	72	51	0.022	0.022	NUM
alces-263	72	52	53	53	NUM
alces-263	72	53	0.96	0.96	NUM
alces-263	72	54	0.022	0.022	NUM
alces-263	72	55	52	52	NUM
alces-263	72	56	0.85	0.85	NUM
alces-263	72	57	0.069	0.069	NUM
alces-263	72	58	20	20	NUM
alces-263	72	59	2018–19	2018–19	NOUN
alces-263	72	60	0.28	0.28	NUM
alces-263	72	61	0.056	0.056	NUM
alces-263	72	62	148	148	NUM
alces-263	72	63	0.95	0.95	NUM
alces-263	72	64	0.026	0.026	NUM
alces-263	72	65	70	70	NUM
alces-263	72	66	0.95	0.95	NUM
alces-263	72	67	0.028	0.028	NUM
alces-263	72	68	63	63	NUM
alces-263	72	69	0.80	0.80	NUM
alces-263	72	70	0.073	0.073	NUM
alces-263	72	71	20	20	NUM
alces-263	72	72	merged	merge	VERB
alces-263	72	73	0.30	0.30	NUM
alces-263	72	74	0.036	0.036	NUM
alces-263	72	75	0.97	0.97	NUM
alces-263	72	76	0.017	0.017	NUM
alces-263	72	77	0.96	0.96	NUM
alces-263	72	78	0.017	0.017	NUM
alces-263	72	79	0.83	0.83	NUM
alces-263	72	80	0.050	0.050	NUM
alces-263	72	81	b.	b.	PROPN
alces-263	72	82	prince	prince	PROPN
alces-263	72	83	george	george	PROPN
alces-263	72	84	south	south	PROPN
alces-263	72	85	study	study	PROPN
alces-263	72	86	area	area	NOUN
alces-263	72	87	year	year	NOUN
alces-263	72	88	calf	calf	NOUN
alces-263	72	89	:	:	PUNCT
alces-263	72	90	cow	cow	NOUN
alces-263	72	91	ratio	ratio	NOUN
alces-263	72	92	annual	annual	ADJ
alces-263	72	93	cow	cow	NOUN
alces-263	72	94	survival	survival	PROPN
alces-263	72	95	winter	winter	PROPN
alces-263	72	96	cow	cow	PROPN
alces-263	72	97	survival	survival	PROPN
alces-263	72	98	winter	winter	PROPN
alces-263	72	99	calf	calf	PROPN
alces-263	72	100	survival	survival	PROPN
alces-263	72	101	j	j	PROPN
alces-263	72	102	/	/	SYM
alces-263	72	103	f	f	PROPN
alces-263	72	104	se	se	PROPN
alces-263	72	105	n	n	PROPN
alces-263	72	106	sf	sf	PROPN
alces-263	72	107	se	se	X
alces-263	72	108	n	n	X
alces-263	72	109	sfw	sfw	PROPN
alces-263	72	110	se	se	PROPN
alces-263	72	111	n	n	X
alces-263	72	112	sjw	sjw	PROPN
alces-263	72	113	se	se	PROPN
alces-263	72	114	n	n	X
alces-263	72	115	2017–18	2017–18	NUM
alces-263	72	116	0.34	0.34	NUM
alces-263	72	117	0.040	0.040	NUM
alces-263	72	118	375	375	NUM
alces-263	72	119	0.79	0.79	NUM
alces-263	72	120	0.049	0.049	NUM
alces-263	72	121	54	54	NUM
alces-263	72	122	0.90	0.90	NUM
alces-263	72	123	0.041	0.041	NUM
alces-263	72	124	40	40	NUM
alces-263	72	125	0.70	0.70	NUM
alces-263	72	126	0.071	0.071	NUM
alces-263	72	127	20	20	NUM
alces-263	72	128	2018–19	2018–19	NUM
alces-263	72	129	0.31	0.31	NUM
alces-263	72	130	0.056	0.056	NUM
alces-263	72	131	168	168	NUM
alces-263	72	132	0.79	0.79	NUM
alces-263	72	133	0.052	0.052	NUM
alces-263	72	134	55	55	NUM
alces-263	72	135	0.85	0.85	NUM
alces-263	72	136	0.052	0.052	NUM
alces-263	72	137	47	47	NUM
alces-263	72	138	0.74	0.74	NUM
alces-263	72	139	0.076	0.076	NUM
alces-263	72	140	19	19	NUM
alces-263	72	141	merged	merge	VERB
alces-263	72	142	0.33	0.33	NUM
alces-263	72	143	0.033	0.033	NUM
alces-263	72	144	0.79	0.79	NUM
alces-263	72	145	0.036	0.036	NUM
alces-263	72	146	0.88	0.88	NUM
alces-263	72	147	0.032	0.032	NUM
alces-263	72	148	0.72	0.72	NUM
alces-263	72	149	0.052	0.052	NUM
alces-263	72	150	alces	alce	NOUN
alces-263	72	151	vol	vol	NOUN
alces-263	72	152	.	.	PROPN
alces-263	72	153	56	56	NUM
alces-263	72	154	,	,	PUNCT
alces-263	72	155	2020	2020	NUM
alces-263	72	156	revisiting	revisit	VERB
alces-263	72	157	the	the	DET
alces-263	72	158	recruitment	recruitment	NOUN
alces-263	72	159	-	-	PUNCT
alces-263	72	160	mortality	mortality	NOUN
alces-263	72	161	.	.	PUNCT
alces-263	73	1	–	–	PUNCT
alces-263	73	2	ian	ian	PROPN
alces-263	73	3	w.	w.	PROPN
alces-263	73	4	hatter	hatter	PROPN
alces-263	73	5	43	43	NUM
alces-263	73	6	bergerud	bergerud	NOUN
alces-263	73	7	(	(	PUNCT
alces-263	73	8	1991	1991	NUM
alces-263	73	9	)	)	PUNCT
alces-263	73	10	r	r	X
alces-263	73	11	-	-	PUNCT
alces-263	73	12	m	m	NOUN
alces-263	73	13	equation	equation	NOUN
alces-263	73	14	were	be	AUX
alces-263	73	15	higher	high	ADJ
alces-263	73	16	than	than	ADP
alces-263	73	17	those	those	PRON
alces-263	73	18	based	base	VERB
alces-263	73	19	on	on	ADP
alces-263	73	20	adjusted	adjust	VERB
alces-263	73	21	recruitment	recruitment	NOUN
alces-263	73	22	rates	rate	NOUN
alces-263	73	23	(	(	PUNCT
alces-263	73	24	table	table	NOUN
alces-263	73	25	2	2	NUM
alces-263	73	26	)	)	PUNCT
alces-263	73	27	.	.	PUNCT
alces-263	74	1	estimates	estimate	NOUN
alces-263	74	2	of	of	ADP
alces-263	74	3	λ	λ	PROPN
alces-263	74	4	based	base	VERB
alces-263	74	5	on	on	ADP
alces-263	74	6	adjusted	adjust	VERB
alces-263	74	7	recruitment	recruitment	NOUN
alces-263	74	8	rates	rate	NOUN
alces-263	74	9	were	be	AUX
alces-263	74	10	identical	identical	ADJ
alces-263	74	11	between	between	ADP
alces-263	74	12	the	the	DET
alces-263	74	13	type	type	NOUN
alces-263	74	14	1	1	NUM
alces-263	74	15	and	and	CCONJ
alces-263	74	16	type	type	NOUN
alces-263	74	17	2	2	NUM
alces-263	74	18	r	r	NOUN
alces-263	74	19	-	-	PUNCT
alces-263	74	20	m	m	NOUN
alces-263	74	21	equations	equation	NOUN
alces-263	74	22	.	.	PUNCT
alces-263	75	1	the	the	DET
alces-263	75	2	95	95	NUM
alces-263	75	3	%	%	NOUN
alces-263	75	4	ci	ci	NOUN
alces-263	75	5	for	for	ADP
alces-263	75	6	λ	λ	PROPN
alces-263	75	7	,	,	PUNCT
alces-263	75	8	based	base	VERB
alces-263	75	9	on	on	ADP
alces-263	75	10	adjusted	adjust	VERB
alces-263	75	11	recruitment	recruitment	NOUN
alces-263	75	12	rates	rate	NOUN
alces-263	75	13	,	,	PUNCT
alces-263	75	14	ranged	range	VERB
alces-263	75	15	from	from	ADP
alces-263	75	16	<	<	X
alces-263	75	17	1.0	1.0	NUM
alces-263	75	18	to	to	PART
alces-263	75	19	>	>	X
alces-263	75	20	1.0	1.0	NUM
alces-263	75	21	for	for	ADP
alces-263	75	22	each	each	DET
alces-263	75	23	year	year	NOUN
alces-263	75	24	in	in	ADP
alces-263	75	25	both	both	DET
alces-263	75	26	populations	population	NOUN
alces-263	75	27	,	,	PUNCT
alces-263	75	28	except	except	SCONJ
alces-263	75	29	for	for	ADP
alces-263	75	30	2017–18	2017–18	NUM
alces-263	75	31	in	in	ADP
alces-263	75	32	the	the	DET
alces-263	75	33	bonaparte	bonaparte	PROPN
alces-263	75	34	study	study	PROPN
alces-263	75	35	area	area	NOUN
alces-263	75	36	.	.	PUNCT
alces-263	76	1	in	in	ADP
alces-263	76	2	this	this	DET
alces-263	76	3	area	area	NOUN
alces-263	76	4	estimates	estimate	NOUN
alces-263	76	5	of	of	ADP
alces-263	76	6	λ	λ	NOUN
alces-263	76	7	were	be	AUX
alces-263	76	8	lower	low	ADJ
alces-263	76	9	in	in	ADP
alces-263	76	10	2016–17	2016–17	NOUN
alces-263	76	11	than	than	ADP
alces-263	76	12	in	in	ADP
alces-263	76	13	2017–18	2017–18	NUM
alces-263	76	14	(	(	PUNCT
alces-263	76	15	z	z	NOUN
alces-263	76	16	=	=	SYM
alces-263	76	17	2.64	2.64	NUM
alces-263	76	18	,	,	PUNCT
alces-263	76	19	p	p	NOUN
alces-263	76	20	=	=	NOUN
alces-263	76	21	0.008	0.008	NUM
alces-263	76	22	)	)	PUNCT
alces-263	76	23	and	and	CCONJ
alces-263	76	24	also	also	ADV
alces-263	76	25	between	between	ADP
alces-263	76	26	2016–17	2016–17	NOUN
alces-263	76	27	and	and	CCONJ
alces-263	76	28	2018–19	2018–19	NUM
alces-263	76	29	(	(	PUNCT
alces-263	76	30	z	z	NOUN
alces-263	76	31	=	=	NUM
alces-263	76	32	1.82	1.82	NUM
alces-263	76	33	,	,	PUNCT
alces-263	76	34	p	p	X
alces-263	76	35	=	=	NOUN
alces-263	76	36	0.068	0.068	NUM
alces-263	76	37	)	)	PUNCT
alces-263	76	38	.	.	PUNCT
alces-263	77	1	growth	growth	NOUN
alces-263	77	2	rates	rate	NOUN
alces-263	77	3	were	be	AUX
alces-263	77	4	not	not	PART
alces-263	77	5	different	different	ADJ
alces-263	77	6	between	between	ADP
alces-263	77	7	2017–18	2017–18	NUM
alces-263	77	8	and	and	CCONJ
alces-263	77	9	2018–19	2018–19	NOUN
alces-263	77	10	(	(	PUNCT
alces-263	77	11	z	z	NOUN
alces-263	77	12	=	=	SYM
alces-263	77	13	1.00	1.00	NUM
alces-263	77	14	,	,	PUNCT
alces-263	77	15	p	p	X
alces-263	77	16	=	=	NOUN
alces-263	77	17	0.32	0.32	NUM
alces-263	77	18	)	)	PUNCT
alces-263	77	19	.	.	PUNCT
alces-263	78	1	merging	merge	VERB
alces-263	78	2	these	these	DET
alces-263	78	3	years	year	NOUN
alces-263	78	4	produced	produce	VERB
alces-263	78	5	a	a	DET
alces-263	78	6	more	more	ADV
alces-263	78	7	precise	precise	ADJ
alces-263	78	8	estimate	estimate	NOUN
alces-263	78	9	of	of	ADP
alces-263	78	10	λ	λ	PROPN
alces-263	78	11	(	(	PUNCT
alces-263	78	12	table	table	NOUN
alces-263	78	13	2	2	NUM
alces-263	78	14	)	)	PUNCT
alces-263	78	15	.	.	PUNCT
alces-263	79	1	growth	growth	NOUN
alces-263	79	2	rates	rate	NOUN
alces-263	79	3	were	be	AUX
alces-263	79	4	similar	similar	ADJ
alces-263	79	5	in	in	ADP
alces-263	79	6	the	the	DET
alces-263	79	7	prince	prince	PROPN
alces-263	79	8	george	george	PROPN
alces-263	79	9	south	south	PROPN
alces-263	79	10	area	area	PROPN
alces-263	79	11	between	between	ADP
alces-263	79	12	2017	2017	NUM
alces-263	79	13	–	–	SYM
alces-263	79	14	2018	2018	NUM
alces-263	79	15	and	and	CCONJ
alces-263	79	16	2018–19	2018–19	NUM
alces-263	79	17	(	(	PUNCT
alces-263	79	18	z	z	NOUN
alces-263	79	19	=	=	SYM
alces-263	79	20	0.034	0.034	NUM
alces-263	79	21	,	,	PUNCT
alces-263	79	22	p	p	X
alces-263	79	23	=	=	NOUN
alces-263	79	24	0.97	0.97	NUM
alces-263	79	25	)	)	PUNCT
alces-263	79	26	;	;	PUNCT
alces-263	79	27	merging	merge	VERB
alces-263	79	28	these	these	DET
alces-263	79	29	years	year	NOUN
alces-263	79	30	also	also	ADV
alces-263	79	31	produced	produce	VERB
alces-263	79	32	a	a	DET
alces-263	79	33	more	more	ADV
alces-263	79	34	precise	precise	ADJ
alces-263	79	35	estimate	estimate	NOUN
alces-263	79	36	of	of	ADP
alces-263	79	37	λ	λ	PROPN
alces-263	79	38	.	.	PUNCT
alces-263	79	39	estimates	estimate	NOUN
alces-263	79	40	of	of	ADP
alces-263	79	41	growth	growth	NOUN
alces-263	79	42	rates	rate	NOUN
alces-263	79	43	from	from	ADP
alces-263	79	44	density	density	NOUN
alces-263	79	45	surveys	survey	NOUN
alces-263	79	46	in	in	ADP
alces-263	79	47	the	the	DET
alces-263	79	48	bonaparte	bonaparte	PROPN
alces-263	79	49	study	study	PROPN
alces-263	79	50	area	area	NOUN
alces-263	79	51	during	during	ADP
alces-263	79	52	2013–18	2013–18	NUM
alces-263	79	53	(	(	PUNCT
alces-263	79	54	λ	λ	X
alces-263	79	55	=	=	NOUN
alces-263	79	56	0.97	0.97	NUM
alces-263	79	57	)	)	PUNCT
alces-263	79	58	were	be	AUX
alces-263	79	59	within	within	ADP
alces-263	79	60	the	the	DET
alces-263	79	61	range	range	NOUN
alces-263	79	62	calculated	calculate	VERB
alces-263	79	63	from	from	ADP
alces-263	79	64	recruitment	recruitment	NOUN
alces-263	79	65	and	and	CCONJ
alces-263	79	66	mortality	mortality	NOUN
alces-263	79	67	(	(	PUNCT
alces-263	79	68	λ	λ	X
alces-263	79	69	=	=	SYM
alces-263	79	70	0.93–1.12	0.93–1.12	NOUN
alces-263	79	71	)	)	PUNCT
alces-263	79	72	,	,	PUNCT
alces-263	79	73	and	and	CCONJ
alces-263	79	74	estimates	estimate	NOUN
alces-263	79	75	of	of	ADP
alces-263	79	76	population	population	NOUN
alces-263	79	77	growth	growth	NOUN
alces-263	79	78	from	from	ADP
alces-263	79	79	density	density	NOUN
alces-263	79	80	surveys	survey	NOUN
alces-263	79	81	in	in	ADP
alces-263	79	82	the	the	DET
alces-263	79	83	prince	prince	PROPN
alces-263	79	84	george	george	PROPN
alces-263	79	85	south	south	PROPN
alces-263	79	86	study	study	PROPN
alces-263	79	87	area	area	NOUN
alces-263	79	88	during	during	ADP
alces-263	79	89	2012–17	2012–17	NUM
alces-263	79	90	(	(	PUNCT
alces-263	79	91	λ	λ	X
alces-263	79	92	=	=	NOUN
alces-263	79	93	0.91	0.91	NUM
alces-263	79	94	)	)	PUNCT
alces-263	79	95	were	be	AUX
alces-263	79	96	comparable	comparable	ADJ
alces-263	79	97	to	to	ADP
alces-263	79	98	the	the	DET
alces-263	79	99	r	r	PROPN
alces-263	79	100	-	-	PUNCT
alces-263	79	101	m	m	NOUN
alces-263	79	102	equation	equation	NOUN
alces-263	79	103	(	(	PUNCT
alces-263	79	104	λ	λ	X
alces-263	79	105	=	=	NOUN
alces-263	79	106	0.90	0.90	NUM
alces-263	79	107	)	)	PUNCT
alces-263	79	108	.	.	PUNCT
alces-263	80	1	discussion	discussion	NOUN
alces-263	80	2	the	the	DET
alces-263	80	3	original	original	ADJ
alces-263	80	4	formulation	formulation	NOUN
alces-263	80	5	of	of	ADP
alces-263	80	6	the	the	DET
alces-263	80	7	r	r	PROPN
alces-263	80	8	-	-	PUNCT
alces-263	80	9	m	m	NOUN
alces-263	80	10	equation	equation	NOUN
alces-263	80	11	developed	develop	VERB
alces-263	80	12	by	by	ADP
alces-263	80	13	hatter	hatter	NOUN
alces-263	80	14	and	and	CCONJ
alces-263	80	15	bergerud	bergerud	NOUN
alces-263	80	16	(	(	PUNCT
alces-263	80	17	1991	1991	NUM
alces-263	80	18	)	)	PUNCT
alces-263	80	19	requires	require	VERB
alces-263	80	20	an	an	DET
alces-263	80	21	adjustment	adjustment	NOUN
alces-263	80	22	to	to	ADP
alces-263	80	23	the	the	DET
alces-263	80	24	recruitment	recruitment	NOUN
alces-263	80	25	rate	rate	NOUN
alces-263	80	26	when	when	SCONJ
alces-263	80	27	calf	calf	NOUN
alces-263	80	28	:	:	PUNCT
alces-263	80	29	cow	cow	NOUN
alces-263	80	30	ratios	ratio	NOUN
alces-263	80	31	are	be	AUX
alces-263	80	32	measured	measure	VERB
alces-263	80	33	in	in	ADP
alces-263	80	34	early	early	ADJ
alces-263	80	35	-	-	PUNCT
alces-263	80	36	to	to	ADP
alces-263	80	37	-	-	PUNCT
alces-263	80	38	mid	mid	NOUN
alces-263	80	39	winter	winter	NOUN
alces-263	80	40	.	.	PUNCT
alces-263	81	1	this	this	DET
alces-263	81	2	adjustment	adjustment	NOUN
alces-263	81	3	accounts	account	VERB
alces-263	81	4	for	for	ADP
alces-263	81	5	differential	differential	ADJ
alces-263	81	6	overwinter	overwinter	ADJ
alces-263	81	7	survival	survival	NOUN
alces-263	81	8	of	of	ADP
alces-263	81	9	calves	calf	NOUN
alces-263	81	10	and	and	CCONJ
alces-263	81	11	adults	adult	NOUN
alces-263	81	12	which	which	PRON
alces-263	81	13	may	may	AUX
alces-263	81	14	be	be	AUX
alces-263	81	15	made	make	VERB
alces-263	81	16	using	use	VERB
alces-263	81	17	either	either	CCONJ
alces-263	81	18	the	the	DET
alces-263	81	19	type	type	NOUN
alces-263	81	20	1	1	NUM
alces-263	81	21	or	or	CCONJ
alces-263	81	22	type	type	NOUN
alces-263	81	23	2	2	NUM
alces-263	81	24	r	r	NOUN
alces-263	81	25	-	-	PUNCT
alces-263	81	26	m	m	NOUN
alces-263	81	27	equations	equation	NOUN
alces-263	81	28	.	.	PUNCT
alces-263	82	1	failure	failure	NOUN
alces-263	82	2	to	to	PART
alces-263	82	3	account	account	VERB
alces-263	82	4	for	for	ADP
alces-263	82	5	differential	differential	ADJ
alces-263	82	6	survival	survival	NOUN
alces-263	82	7	results	result	NOUN
alces-263	82	8	in	in	ADP
alces-263	82	9	an	an	DET
alces-263	82	10	overestimate	overestimate	NOUN
alces-263	82	11	of	of	ADP
alces-263	82	12	λ	λ	NOUN
alces-263	82	13	.	.	PUNCT
alces-263	83	1	users	user	NOUN
alces-263	83	2	of	of	ADP
alces-263	83	3	the	the	DET
alces-263	83	4	r	r	PROPN
alces-263	83	5	-	-	PUNCT
alces-263	83	6	m	m	NOUN
alces-263	83	7	equations	equation	NOUN
alces-263	83	8	may	may	AUX
alces-263	83	9	prefer	prefer	VERB
alces-263	83	10	the	the	DET
alces-263	83	11	type	type	NOUN
alces-263	83	12	2	2	NUM
alces-263	83	13	model	model	NOUN
alces-263	83	14	since	since	SCONJ
alces-263	83	15	most	most	ADJ
alces-263	83	16	moose	moose	NOUN
alces-263	83	17	biologists	biologist	NOUN
alces-263	83	18	define	define	VERB
alces-263	83	19	calf	calf	NOUN
alces-263	83	20	recruitment	recruitment	NOUN
alces-263	83	21	as	as	ADP
alces-263	83	22	the	the	DET
alces-263	83	23	calf	calf	NOUN
alces-263	83	24	:	:	PUNCT
alces-263	83	25	cow	cow	NOUN
alces-263	83	26	ratio	ratio	NOUN
alces-263	83	27	rather	rather	ADV
alces-263	83	28	than	than	ADP
alces-263	83	29	%	%	NOUN
alces-263	83	30	calves	calf	NOUN
alces-263	83	31	and	and	CCONJ
alces-263	83	32	because	because	SCONJ
alces-263	83	33	calculations	calculation	NOUN
alces-263	83	34	are	be	AUX
alces-263	83	35	simpler	simple	ADJ
alces-263	83	36	.	.	PUNCT
alces-263	84	1	table	table	NOUN
alces-263	84	2	2	2	NUM
alces-263	84	3	.	.	NOUN
alces-263	84	4	moose	moose	NOUN
alces-263	84	5	recruitment	recruitment	NOUN
alces-263	84	6	(	(	PUNCT
alces-263	84	7	r	r	NOUN
alces-263	84	8	)	)	PUNCT
alces-263	84	9	,	,	PUNCT
alces-263	84	10	adjusted	adjust	VERB
alces-263	84	11	moose	moose	NOUN
alces-263	84	12	recruitment	recruitment	NOUN
alces-263	84	13	(	(	PUNCT
alces-263	84	14	r	r	NOUN
alces-263	84	15	’	'	PUNCT
alces-263	84	16	)	)	PUNCT
alces-263	84	17	,	,	PUNCT
alces-263	84	18	and	and	CCONJ
alces-263	84	19	population	population	NOUN
alces-263	84	20	growth	growth	NOUN
alces-263	84	21	rates	rate	NOUN
alces-263	84	22	(	(	PUNCT
alces-263	84	23	λ	λ	X
alces-263	84	24	)	)	PUNCT
alces-263	84	25	with	with	ADP
alces-263	84	26	monte	monte	PROPN
alces-263	84	27	carlo	carlo	PROPN
alces-263	84	28	simulated	simulate	VERB
alces-263	84	29	confidence	confidence	NOUN
alces-263	84	30	intervals	interval	NOUN
alces-263	84	31	(	(	PUNCT
alces-263	84	32	95	95	NUM
alces-263	84	33	%	%	NOUN
alces-263	84	34	ci	ci	NOUN
alces-263	84	35	)	)	PUNCT
alces-263	84	36	for	for	ADP
alces-263	84	37	the	the	DET
alces-263	84	38	type	type	NOUN
alces-263	84	39	1	1	NUM
alces-263	84	40	and	and	CCONJ
alces-263	84	41	type	type	NOUN
alces-263	84	42	2	2	NUM
alces-263	84	43	r	r	NOUN
alces-263	84	44	-	-	PUNCT
alces-263	84	45	m	m	NOUN
alces-263	84	46	equations	equation	NOUN
alces-263	84	47	within	within	ADP
alces-263	84	48	the	the	DET
alces-263	84	49	bonaparte	bonaparte	NOUN
alces-263	84	50	and	and	CCONJ
alces-263	84	51	prince	prince	PROPN
alces-263	84	52	george	george	PROPN
alces-263	84	53	south	south	PROPN
alces-263	84	54	study	study	PROPN
alces-263	84	55	areas	area	NOUN
alces-263	84	56	of	of	ADP
alces-263	84	57	british	british	PROPN
alces-263	84	58	columbia	columbia	PROPN
alces-263	84	59	,	,	PUNCT
alces-263	84	60	canada	canada	PROPN
alces-263	84	61	.	.	PUNCT
alces-263	84	62	merged	merge	VERB
alces-263	84	63	refers	refer	NOUN
alces-263	84	64	to	to	ADP
alces-263	84	65	combined	combine	VERB
alces-263	84	66	estimates	estimate	NOUN
alces-263	84	67	for	for	ADP
alces-263	84	68	2017–18	2017–18	NUM
alces-263	84	69	and	and	CCONJ
alces-263	84	70	2018–19	2018–19	NOUN
alces-263	84	71	.	.	PUNCT
alces-263	85	1	a.	a.	PROPN
alces-263	85	2	bonaparte	bonaparte	PROPN
alces-263	85	3	study	study	PROPN
alces-263	85	4	area	area	NOUN
alces-263	85	5	year	year	NOUN
alces-263	85	6	r	r	X
alces-263	85	7	-	-	PUNCT
alces-263	85	8	m	m	NOUN
alces-263	85	9	equation1	equation1	NOUN
alces-263	85	10	type	type	NOUN
alces-263	85	11	1	1	NUM
alces-263	85	12	r	r	NOUN
alces-263	85	13	-	-	PUNCT
alces-263	85	14	m	m	NOUN
alces-263	85	15	equation	equation	NOUN
alces-263	85	16	type	type	NOUN
alces-263	85	17	2	2	NUM
alces-263	85	18	r	r	NOUN
alces-263	85	19	-	-	PUNCT
alces-263	85	20	m	m	NOUN
alces-263	85	21	equation	equation	NOUN
alces-263	85	22	type	type	NOUN
alces-263	85	23	1	1	NUM
alces-263	85	24	&	&	CCONJ
alces-263	85	25	2	2	NUM
alces-263	85	26	r	r	NOUN
alces-263	85	27	(	(	PUNCT
alces-263	85	28	eq	eq	NOUN
alces-263	85	29	.	.	NOUN
alces-263	85	30	7	7	X
alces-263	85	31	)	)	PUNCT
alces-263	85	32	λ	λ	NOUN
alces-263	85	33	(	(	PUNCT
alces-263	85	34	eq	eq	NOUN
alces-263	85	35	.	.	NOUN
alces-263	85	36	3	3	X
alces-263	85	37	)	)	PUNCT
alces-263	85	38	r	r	NOUN
alces-263	85	39	’	'	PUNCT
alces-263	85	40	(	(	PUNCT
alces-263	85	41	eq	eq	NOUN
alces-263	85	42	.	.	NOUN
alces-263	85	43	9	9	X
alces-263	85	44	)	)	PUNCT
alces-263	85	45	λ	λ	NOUN
alces-263	85	46	(	(	PUNCT
alces-263	85	47	eq	eq	NOUN
alces-263	85	48	.	.	NOUN
alces-263	85	49	11	11	NUM
alces-263	85	50	)	)	PUNCT
alces-263	85	51	r	r	NOUN
alces-263	85	52	’	'	PUNCT
alces-263	85	53	(	(	PUNCT
alces-263	85	54	eq	eq	NOUN
alces-263	85	55	.	.	NOUN
alces-263	85	56	10	10	NUM
alces-263	85	57	)	)	PUNCT
alces-263	85	58	λ	λ	NOUN
alces-263	85	59	(	(	PUNCT
alces-263	85	60	eq	eq	NOUN
alces-263	85	61	.	.	PROPN
alces-263	85	62	12	12	NUM
alces-263	85	63	)	)	PUNCT
alces-263	85	64	95	95	NUM
alces-263	85	65	%	%	NOUN
alces-263	85	66	ci	ci	NOUN
alces-263	85	67	2016–17	2016–17	NOUN
alces-263	85	68	0.061	0.061	NUM
alces-263	85	69	0.97	0.97	NUM
alces-263	85	70	0.030	0.030	NUM
alces-263	85	71	0.93	0.93	NUM
alces-263	85	72	0.030	0.030	NUM
alces-263	85	73	0.93	0.93	NUM
alces-263	85	74	0.79–1.02	0.79–1.02	NUM
alces-263	85	75	2017–18	2017–18	NUM
alces-263	85	76	0.138	0.138	NUM
alces-263	85	77	1.13	1.13	NUM
alces-263	85	78	0.124	0.124	NUM
alces-263	85	79	1.12	1.12	NUM
alces-263	85	80	0.142	0.142	NUM
alces-263	85	81	1.12	1.12	NUM
alces-263	85	82	1.04–1.18	1.04–1.18	NUM
alces-263	85	83	2018–19	2018–19	NOUN
alces-263	85	84	0.123	0.123	NUM
alces-263	85	85	1.08	1.08	NUM
alces-263	85	86	0.105	0.105	NUM
alces-263	85	87	1.06	1.06	NUM
alces-263	85	88	0.118	0.118	NUM
alces-263	85	89	1.06	1.06	NUM
alces-263	85	90	0.98–1.14	0.98–1.14	NUM
alces-263	85	91	merged	merge	VERB
alces-263	85	92	0.132	0.132	NUM
alces-263	85	93	1.11	1.11	NUM
alces-263	85	94	0.116	0.116	NUM
alces-263	85	95	1.09	1.09	NUM
alces-263	85	96	0.131	0.131	NUM
alces-263	85	97	1.09	1.09	NUM
alces-263	85	98	1.04–1.14	1.04–1.14	NUM
alces-263	85	99	1estimated	1estimate	VERB
alces-263	85	100	using	use	VERB
alces-263	85	101	the	the	DET
alces-263	85	102	original	original	ADJ
alces-263	85	103	r	r	NOUN
alces-263	85	104	-	-	PUNCT
alces-263	85	105	m	m	NOUN
alces-263	85	106	equation	equation	NOUN
alces-263	85	107	from	from	ADP
alces-263	85	108	hatter	hatter	NOUN
alces-263	85	109	and	and	CCONJ
alces-263	85	110	bergerud	bergerud	NOUN
alces-263	85	111	(	(	PUNCT
alces-263	85	112	1991	1991	NUM
alces-263	85	113	)	)	PUNCT
alces-263	85	114	.	.	PUNCT
alces-263	86	1	b.	b.	PROPN
alces-263	86	2	prince	prince	PROPN
alces-263	86	3	george	george	PROPN
alces-263	86	4	south	south	PROPN
alces-263	86	5	study	study	PROPN
alces-263	86	6	area	area	NOUN
alces-263	86	7	year	year	NOUN
alces-263	86	8	r	r	X
alces-263	86	9	-	-	PUNCT
alces-263	86	10	m	m	NOUN
alces-263	86	11	equation1	equation1	NOUN
alces-263	86	12	type	type	NOUN
alces-263	86	13	1	1	NUM
alces-263	86	14	r	r	NOUN
alces-263	86	15	-	-	PUNCT
alces-263	86	16	m	m	NOUN
alces-263	86	17	equation	equation	NOUN
alces-263	86	18	type	type	NOUN
alces-263	86	19	2	2	NUM
alces-263	86	20	r	r	NOUN
alces-263	86	21	-	-	PUNCT
alces-263	86	22	m	m	NOUN
alces-263	86	23	equation	equation	NOUN
alces-263	86	24	type	type	NOUN
alces-263	86	25	1	1	NUM
alces-263	86	26	&	&	CCONJ
alces-263	86	27	2	2	NUM
alces-263	86	28	r	r	NOUN
alces-263	86	29	(	(	PUNCT
alces-263	86	30	eq	eq	NOUN
alces-263	86	31	.	.	NOUN
alces-263	86	32	7	7	X
alces-263	86	33	)	)	PUNCT
alces-263	86	34	λ	λ	NOUN
alces-263	86	35	(	(	PUNCT
alces-263	86	36	eq	eq	NOUN
alces-263	86	37	.	.	NOUN
alces-263	86	38	3	3	X
alces-263	86	39	)	)	PUNCT
alces-263	86	40	r	r	NOUN
alces-263	86	41	’	'	PUNCT
alces-263	86	42	(	(	PUNCT
alces-263	86	43	eq	eq	NOUN
alces-263	86	44	.	.	NOUN
alces-263	86	45	9	9	X
alces-263	86	46	)	)	PUNCT
alces-263	86	47	λ	λ	NOUN
alces-263	86	48	(	(	PUNCT
alces-263	86	49	eq	eq	NOUN
alces-263	86	50	.	.	NOUN
alces-263	86	51	11	11	NUM
alces-263	86	52	)	)	PUNCT
alces-263	86	53	r	r	NOUN
alces-263	86	54	’	'	PUNCT
alces-263	86	55	(	(	PUNCT
alces-263	86	56	eq	eq	NOUN
alces-263	86	57	.	.	NOUN
alces-263	86	58	10	10	NUM
alces-263	86	59	)	)	PUNCT
alces-263	86	60	λ	λ	NOUN
alces-263	86	61	(	(	PUNCT
alces-263	86	62	eq	eq	NOUN
alces-263	86	63	.	.	PROPN
alces-263	86	64	12	12	NUM
alces-263	86	65	)	)	PUNCT
alces-263	86	66	95	95	NUM
alces-263	86	67	%	%	NOUN
alces-263	86	68	ci	ci	NOUN
alces-263	86	69	2017–18	2017–18	NUM
alces-263	86	70	0.145	0.145	NUM
alces-263	86	71	0.93	0.93	NUM
alces-263	86	72	0.117	0.117	NUM
alces-263	86	73	0.90	0.90	NUM
alces-263	86	74	0.133	0.133	NUM
alces-263	86	75	0.90	0.90	NUM
alces-263	86	76	0.77–1.01	0.77–1.01	NUM
alces-263	86	77	2018–19	2018–19	NOUN
alces-263	86	78	0.134	0.134	NUM
alces-263	86	79	0.91	0.91	NUM
alces-263	86	80	0.119	0.119	NUM
alces-263	86	81	0.90	0.90	NUM
alces-263	86	82	0.135	0.135	NUM
alces-263	86	83	0.90	0.90	NUM
alces-263	86	84	0.76–1.02	0.76–1.02	NUM
alces-263	86	85	merged	merge	VERB
alces-263	86	86	0.142	0.142	NUM
alces-263	86	87	0.92	0.92	NUM
alces-263	86	88	0.119	0.119	NUM
alces-263	86	89	0.90	0.90	NUM
alces-263	86	90	0.135	0.135	NUM
alces-263	86	91	0.90	0.90	NUM
alces-263	86	92	0.81–0.98	0.81–0.98	NUM
alces-263	86	93	1	1	NUM
alces-263	86	94	estimated	estimate	VERB
alces-263	86	95	using	use	VERB
alces-263	86	96	the	the	DET
alces-263	86	97	original	original	ADJ
alces-263	86	98	r	r	NOUN
alces-263	86	99	-	-	PUNCT
alces-263	86	100	m	m	NOUN
alces-263	86	101	equation	equation	NOUN
alces-263	86	102	from	from	ADP
alces-263	86	103	hatter	hatter	NOUN
alces-263	86	104	and	and	CCONJ
alces-263	86	105	bergerud	bergerud	NOUN
alces-263	86	106	(	(	PUNCT
alces-263	86	107	1991	1991	NUM
alces-263	86	108	)	)	PUNCT
alces-263	86	109	.	.	PUNCT
alces-263	87	1	revisiting	revisit	VERB
alces-263	87	2	the	the	DET
alces-263	87	3	r	r	NOUN
alces-263	87	4	-	-	PUNCT
alces-263	87	5	m	m	NOUN
alces-263	87	6	equation	equation	NOUN
alces-263	87	7	.	.	PUNCT
alces-263	88	1	–	–	PUNCT
alces-263	88	2	ian	ian	PROPN
alces-263	88	3	w.	w.	PROPN
alces-263	88	4	hatter	hatter	PROPN
alces-263	88	5	alces	alces	PROPN
alces-263	88	6	vol	vol	NOUN
alces-263	88	7	.	.	PROPN
alces-263	89	1	56	56	NUM
alces-263	89	2	,	,	PUNCT
alces-263	89	3	2020	2020	NUM
alces-263	89	4	44	44	NUM
alces-263	89	5	there	there	PRON
alces-263	89	6	are	be	VERB
alces-263	89	7	4	4	NUM
alces-263	89	8	primary	primary	ADJ
alces-263	89	9	methods	method	NOUN
alces-263	89	10	currently	currently	ADV
alces-263	89	11	used	use	VERB
alces-263	89	12	to	to	PART
alces-263	89	13	estimate	estimate	VERB
alces-263	89	14	λ	λ	PROPN
alces-263	89	15	for	for	ADP
alces-263	89	16	moose	moose	NOUN
alces-263	89	17	populations	population	NOUN
alces-263	89	18	.	.	PUNCT
alces-263	90	1	the	the	DET
alces-263	90	2	growth	growth	NOUN
alces-263	90	3	rate	rate	NOUN
alces-263	90	4	may	may	AUX
alces-263	90	5	be	be	AUX
alces-263	90	6	measured	measure	VERB
alces-263	90	7	from	from	ADP
alces-263	90	8	2	2	NUM
alces-263	90	9	or	or	CCONJ
alces-263	90	10	more	more	ADJ
alces-263	90	11	population	population	NOUN
alces-263	90	12	surveys	survey	NOUN
alces-263	90	13	over	over	ADP
alces-263	90	14	time	time	NOUN
alces-263	90	15	(	(	PUNCT
alces-263	90	16	van	van	PROPN
alces-263	90	17	ballenberghe	ballenberghe	NOUN
alces-263	90	18	1983	1983	NUM
alces-263	90	19	)	)	PUNCT
alces-263	90	20	,	,	PUNCT
alces-263	90	21	from	from	ADP
alces-263	90	22	calf	calf	NOUN
alces-263	90	23	recruitment	recruitment	NOUN
alces-263	90	24	and	and	CCONJ
alces-263	90	25	adult	adult	NOUN
alces-263	90	26	mortality	mortality	NOUN
alces-263	90	27	rates	rate	NOUN
alces-263	90	28	(	(	PUNCT
alces-263	90	29	hatter	hatter	NOUN
alces-263	90	30	and	and	CCONJ
alces-263	90	31	bergerud	bergerud	NOUN
alces-263	90	32	1991	1991	NUM
alces-263	90	33	)	)	PUNCT
alces-263	90	34	,	,	PUNCT
alces-263	90	35	from	from	ADP
alces-263	90	36	survival	survival	NOUN
alces-263	90	37	and	and	CCONJ
alces-263	90	38	fecundity	fecundity	NOUN
alces-263	90	39	schedules	schedule	NOUN
alces-263	90	40	(	(	PUNCT
alces-263	90	41	van	van	PROPN
alces-263	90	42	ballenberghe	ballenberghe	NOUN
alces-263	90	43	1983	1983	NUM
alces-263	90	44	)	)	PUNCT
alces-263	90	45	,	,	PUNCT
alces-263	90	46	or	or	CCONJ
alces-263	90	47	from	from	ADP
alces-263	90	48	fitting	fitting	ADJ
alces-263	90	49	population	population	NOUN
alces-263	90	50	models	model	NOUN
alces-263	90	51	to	to	ADP
alces-263	90	52	multiple	multiple	ADJ
alces-263	90	53	sources	source	NOUN
alces-263	90	54	of	of	ADP
alces-263	90	55	observed	observed	ADJ
alces-263	90	56	data	datum	NOUN
alces-263	90	57	(	(	PUNCT
alces-263	90	58	kuzyk	kuzyk	NOUN
alces-263	90	59	et	et	PROPN
alces-263	90	60	al	al	PROPN
alces-263	90	61	.	.	PROPN
alces-263	90	62	2018	2018	NUM
alces-263	90	63	)	)	PUNCT
alces-263	90	64	.	.	PUNCT
alces-263	91	1	the	the	DET
alces-263	91	2	primary	primary	ADJ
alces-263	91	3	advantage	advantage	NOUN
alces-263	91	4	of	of	ADP
alces-263	91	5	the	the	DET
alces-263	91	6	r	r	PROPN
alces-263	91	7	-	-	PUNCT
alces-263	91	8	m	m	NOUN
alces-263	91	9	model	model	NOUN
alces-263	91	10	is	be	AUX
alces-263	91	11	that	that	SCONJ
alces-263	91	12	it	it	PRON
alces-263	91	13	provides	provide	VERB
alces-263	91	14	an	an	DET
alces-263	91	15	estimate	estimate	NOUN
alces-263	91	16	of	of	ADP
alces-263	91	17	λ	λ	PROPN
alces-263	91	18	from	from	ADP
alces-263	91	19	a	a	DET
alces-263	91	20	single	single	ADJ
alces-263	91	21	,	,	PUNCT
alces-263	91	22	late	late	ADJ
alces-263	91	23	winter	winter	NOUN
alces-263	91	24	herd	herd	NOUN
alces-263	91	25	composition	composition	NOUN
alces-263	91	26	survey	survey	NOUN
alces-263	91	27	when	when	SCONJ
alces-263	91	28	the	the	DET
alces-263	91	29	annual	annual	ADJ
alces-263	91	30	adult	adult	NOUN
alces-263	91	31	mortality	mortality	NOUN
alces-263	91	32	rate	rate	NOUN
alces-263	91	33	is	be	AUX
alces-263	91	34	known	know	VERB
alces-263	91	35	.	.	PUNCT
alces-263	92	1	it	it	PRON
alces-263	92	2	may	may	AUX
alces-263	92	3	also	also	ADV
alces-263	92	4	be	be	AUX
alces-263	92	5	preferred	prefer	VERB
alces-263	92	6	where	where	SCONJ
alces-263	92	7	moose	moose	NOUN
alces-263	92	8	density	density	NOUN
alces-263	92	9	surveys	survey	NOUN
alces-263	92	10	are	be	AUX
alces-263	92	11	not	not	PART
alces-263	92	12	feasible	feasible	ADJ
alces-263	92	13	,	,	PUNCT
alces-263	92	14	such	such	ADJ
alces-263	92	15	as	as	ADP
alces-263	92	16	in	in	ADP
alces-263	92	17	densely	densely	ADV
alces-263	92	18	forested	forest	VERB
alces-263	92	19	areas	area	NOUN
alces-263	92	20	or	or	CCONJ
alces-263	92	21	where	where	SCONJ
alces-263	92	22	moose	moose	NOUN
alces-263	92	23	occur	occur	VERB
alces-263	92	24	at	at	ADP
alces-263	92	25	very	very	ADV
alces-263	92	26	low	low	ADJ
alces-263	92	27	density	density	NOUN
alces-263	92	28	(	(	PUNCT
alces-263	92	29	severud	severud	NOUN
alces-263	92	30	et	et	PROPN
alces-263	92	31	al	al	PROPN
alces-263	92	32	.	.	PROPN
alces-263	92	33	2019	2019	NUM
alces-263	92	34	)	)	PUNCT
alces-263	92	35	.	.	PUNCT
alces-263	93	1	guthery	guthery	NOUN
alces-263	93	2	and	and	CCONJ
alces-263	93	3	shaw	shaw	PROPN
alces-263	93	4	(	(	PUNCT
alces-263	93	5	2013	2013	NUM
alces-263	93	6	)	)	PUNCT
alces-263	93	7	note	note	VERB
alces-263	93	8	that	that	SCONJ
alces-263	93	9	the	the	DET
alces-263	93	10	r	r	PROPN
alces-263	93	11	-	-	PUNCT
alces-263	93	12	m	m	NOUN
alces-263	93	13	equation	equation	NOUN
alces-263	93	14	is	be	AUX
alces-263	93	15	a	a	DET
alces-263	93	16	tautology	tautology	NOUN
alces-263	93	17	and	and	CCONJ
alces-263	93	18	thus	thus	ADV
alces-263	93	19	inevitably	inevitably	ADV
alces-263	93	20	true	true	ADJ
alces-263	93	21	and	and	CCONJ
alces-263	93	22	does	do	AUX
alces-263	93	23	not	not	PART
alces-263	93	24	require	require	VERB
alces-263	93	25	empirical	empirical	ADJ
alces-263	93	26	verification	verification	NOUN
alces-263	93	27	.	.	PUNCT
alces-263	94	1	however	however	ADV
alces-263	94	2	,	,	PUNCT
alces-263	94	3	survey	survey	VERB
alces-263	94	4	timing	timing	NOUN
alces-263	94	5	and	and	CCONJ
alces-263	94	6	survey	survey	NOUN
alces-263	94	7	bias	bias	NOUN
alces-263	94	8	can	can	AUX
alces-263	94	9	affect	affect	VERB
alces-263	94	10	the	the	DET
alces-263	94	11	accuracy	accuracy	NOUN
alces-263	94	12	of	of	ADP
alces-263	94	13	λ	λ	PROPN
alces-263	94	14	calculations	calculation	NOUN
alces-263	94	15	.	.	PUNCT
alces-263	95	1	serrouya	serrouya	PROPN
alces-263	95	2	et	et	PROPN
alces-263	95	3	al	al	PROPN
alces-263	95	4	.	.	PROPN
alces-263	96	1	(	(	PUNCT
alces-263	96	2	2016	2016	NUM
alces-263	96	3	)	)	PUNCT
alces-263	96	4	compared	compare	VERB
alces-263	96	5	population	population	NOUN
alces-263	96	6	growth	growth	NOUN
alces-263	96	7	rates	rate	NOUN
alces-263	96	8	for	for	ADP
alces-263	96	9	caribou	caribou	NOUN
alces-263	96	10	between	between	ADP
alces-263	96	11	abundance	abundance	NOUN
alces-263	96	12	surveys	survey	NOUN
alces-263	96	13	and	and	CCONJ
alces-263	96	14	the	the	DET
alces-263	96	15	r	r	PROPN
alces-263	96	16	-	-	PUNCT
alces-263	96	17	m	m	NOUN
alces-263	96	18	equation	equation	NOUN
alces-263	97	1	and	and	CCONJ
alces-263	98	1	found	find	VERB
alces-263	98	2	that	that	SCONJ
alces-263	98	3	the	the	DET
alces-263	98	4	r	r	PROPN
alces-263	98	5	-	-	PUNCT
alces-263	98	6	m	m	NOUN
alces-263	98	7	equation	equation	NOUN
alces-263	98	8	overestimated	overestimate	VERB
alces-263	98	9	λ	λ	INTJ
alces-263	98	10	compared	compare	VERB
alces-263	98	11	to	to	ADP
alces-263	98	12	survey	survey	NOUN
alces-263	98	13	-	-	PUNCT
alces-263	98	14	based	base	VERB
alces-263	98	15	λ	λ	NOUN
alces-263	98	16	.	.	PUNCT
alces-263	99	1	they	they	PRON
alces-263	99	2	proposed	propose	VERB
alces-263	99	3	3	3	NUM
alces-263	99	4	possible	possible	ADJ
alces-263	99	5	reasons	reason	NOUN
alces-263	99	6	including	include	VERB
alces-263	99	7	:	:	PUNCT
alces-263	99	8	1	1	X
alces-263	99	9	)	)	PUNCT
alces-263	99	10	measurement	measurement	NOUN
alces-263	99	11	of	of	ADP
alces-263	99	12	recruitment	recruitment	NOUN
alces-263	99	13	at	at	ADP
alces-263	99	14	10	10	NUM
alces-263	99	15	-	-	PUNCT
alces-263	99	16	months	month	NOUN
alces-263	99	17	of	of	ADP
alces-263	99	18	age	age	NOUN
alces-263	99	19	rather	rather	ADV
alces-263	99	20	than	than	ADP
alces-263	99	21	as	as	ADP
alces-263	99	22	1	1	NUM
alces-263	99	23	year	year	NOUN
alces-263	99	24	-	-	PUNCT
alces-263	99	25	old	old	ADJ
alces-263	99	26	,	,	PUNCT
alces-263	99	27	2	2	NUM
alces-263	99	28	)	)	PUNCT
alces-263	99	29	adult	adult	NOUN
alces-263	99	30	survival	survival	NOUN
alces-263	99	31	estimates	estimate	VERB
alces-263	99	32	that	that	PRON
alces-263	99	33	are	be	AUX
alces-263	99	34	biased	bias	VERB
alces-263	99	35	towards	towards	ADP
alces-263	99	36	more	more	ADV
alces-263	99	37	mature	mature	ADJ
alces-263	99	38	animals	animal	NOUN
alces-263	99	39	,	,	PUNCT
alces-263	99	40	and	and	CCONJ
alces-263	99	41	3	3	X
alces-263	99	42	)	)	PUNCT
alces-263	99	43	errors	error	NOUN
alces-263	99	44	in	in	ADP
alces-263	99	45	herd	herd	NOUN
alces-263	99	46	composition	composition	NOUN
alces-263	99	47	surveys	survey	NOUN
alces-263	99	48	(	(	PUNCT
alces-263	99	49	e.g.	e.g.	ADV
alces-263	99	50	,	,	PUNCT
alces-263	99	51	sightability	sightability	NOUN
alces-263	99	52	differences	difference	NOUN
alces-263	99	53	between	between	ADP
alces-263	99	54	barren	barren	ADJ
alces-263	99	55	females	female	NOUN
alces-263	99	56	and	and	CCONJ
alces-263	99	57	those	those	PRON
alces-263	99	58	with	with	ADP
alces-263	99	59	offspring	offspring	NOUN
alces-263	99	60	)	)	PUNCT
alces-263	99	61	.	.	PUNCT
alces-263	100	1	they	they	PRON
alces-263	100	2	found	find	VERB
alces-263	100	3	that	that	SCONJ
alces-263	100	4	the	the	DET
alces-263	100	5	r	r	PROPN
alces-263	100	6	-	-	PUNCT
alces-263	100	7	m	m	NOUN
alces-263	100	8	equation	equation	NOUN
alces-263	100	9	explained	explain	VERB
alces-263	100	10	60	60	NUM
alces-263	100	11	%	%	NOUN
alces-263	100	12	of	of	ADP
alces-263	100	13	the	the	DET
alces-263	100	14	variation	variation	NOUN
alces-263	100	15	in	in	ADP
alces-263	100	16	survey	survey	NOUN
alces-263	100	17	-	-	PUNCT
alces-263	100	18	based	base	VERB
alces-263	100	19	λ	λ	PROPN
alces-263	100	20	and	and	CCONJ
alces-263	100	21	was	be	AUX
alces-263	100	22	a	a	DET
alces-263	100	23	better	well	ADJ
alces-263	100	24	predictor	predictor	NOUN
alces-263	100	25	of	of	ADP
alces-263	100	26	this	this	DET
alces-263	100	27	parameter	parameter	NOUN
alces-263	100	28	compared	compare	VERB
alces-263	100	29	to	to	ADP
alces-263	100	30	other	other	ADJ
alces-263	100	31	approaches	approach	NOUN
alces-263	100	32	.	.	PUNCT
alces-263	101	1	users	user	NOUN
alces-263	101	2	of	of	ADP
alces-263	101	3	the	the	DET
alces-263	101	4	r	r	PROPN
alces-263	101	5	-	-	PUNCT
alces-263	101	6	m	m	NOUN
alces-263	101	7	equation	equation	NOUN
alces-263	101	8	should	should	AUX
alces-263	101	9	ensure	ensure	VERB
alces-263	101	10	that	that	SCONJ
alces-263	101	11	herd	herd	NOUN
alces-263	101	12	composition	composition	NOUN
alces-263	101	13	surveys	survey	NOUN
alces-263	101	14	are	be	AUX
alces-263	101	15	representative	representative	ADJ
alces-263	101	16	of	of	ADP
alces-263	101	17	the	the	DET
alces-263	101	18	population	population	NOUN
alces-263	101	19	and	and	CCONJ
alces-263	101	20	measure	measure	VERB
alces-263	101	21	recruitment	recruitment	NOUN
alces-263	101	22	just	just	ADV
alces-263	101	23	before	before	SCONJ
alces-263	101	24	calves	calf	NOUN
alces-263	101	25	become	become	VERB
alces-263	101	26	1	1	NUM
alces-263	101	27	year	year	NOUN
alces-263	101	28	-	-	PUNCT
alces-263	101	29	of	of	ADP
alces-263	101	30	-	-	PUNCT
alces-263	101	31	age	age	NOUN
alces-263	101	32	.	.	PUNCT
alces-263	102	1	alternatively	alternatively	ADV
alces-263	102	2	,	,	PUNCT
alces-263	102	3	the	the	DET
alces-263	102	4	modified	modified	ADJ
alces-263	102	5	r	r	NOUN
alces-263	102	6	-	-	PUNCT
alces-263	102	7	m	m	NOUN
alces-263	102	8	equation	equation	NOUN
alces-263	102	9	may	may	AUX
alces-263	102	10	be	be	AUX
alces-263	102	11	used	use	VERB
alces-263	102	12	to	to	PART
alces-263	102	13	account	account	VERB
alces-263	102	14	for	for	ADP
alces-263	102	15	differential	differential	ADJ
alces-263	102	16	winter	winter	NOUN
alces-263	102	17	mortality	mortality	NOUN
alces-263	102	18	between	between	ADP
alces-263	102	19	calves	calf	NOUN
alces-263	102	20	and	and	CCONJ
alces-263	102	21	adults	adult	NOUN
alces-263	102	22	when	when	SCONJ
alces-263	102	23	survival	survival	NOUN
alces-263	102	24	estimates	estimate	NOUN
alces-263	102	25	are	be	AUX
alces-263	102	26	available	available	ADJ
alces-263	102	27	.	.	PUNCT
alces-263	103	1	in	in	ADP
alces-263	103	2	addition	addition	NOUN
alces-263	103	3	,	,	PUNCT
alces-263	103	4	users	user	NOUN
alces-263	103	5	need	need	VERB
alces-263	103	6	to	to	PART
alces-263	103	7	be	be	AUX
alces-263	103	8	careful	careful	ADJ
alces-263	103	9	to	to	PART
alces-263	103	10	ensure	ensure	VERB
alces-263	103	11	that	that	SCONJ
alces-263	103	12	estimates	estimate	NOUN
alces-263	103	13	of	of	ADP
alces-263	103	14	adult	adult	NOUN
alces-263	103	15	survival	survival	NOUN
alces-263	103	16	from	from	ADP
alces-263	103	17	radio	radio	NOUN
alces-263	103	18	-	-	PUNCT
alces-263	103	19	collared	collar	VERB
alces-263	103	20	animals	animal	NOUN
alces-263	103	21	include	include	VERB
alces-263	103	22	a	a	DET
alces-263	103	23	representative	representative	ADJ
alces-263	103	24	sample	sample	NOUN
alces-263	103	25	of	of	ADP
alces-263	103	26	their	their	PRON
alces-263	103	27	standing	stand	VERB
alces-263	103	28	age	age	NOUN
alces-263	103	29	distribution	distribution	NOUN
alces-263	103	30	.	.	PUNCT
alces-263	104	1	this	this	PRON
alces-263	104	2	may	may	AUX
alces-263	104	3	require	require	VERB
alces-263	104	4	estimating	estimate	VERB
alces-263	104	5	survival	survival	NOUN
alces-263	104	6	rates	rate	NOUN
alces-263	104	7	over	over	ADP
alces-263	104	8	multiple	multiple	ADJ
alces-263	104	9	years	year	NOUN
alces-263	104	10	.	.	PUNCT
alces-263	105	1	in	in	ADP
alces-263	105	2	alberta	alberta	PROPN
alces-263	105	3	,	,	PUNCT
alces-263	105	4	the	the	DET
alces-263	105	5	population	population	NOUN
alces-263	105	6	trend	trend	NOUN
alces-263	105	7	for	for	ADP
alces-263	105	8	boreal	boreal	ADJ
alces-263	105	9	caribou	caribou	NOUN
alces-263	105	10	was	be	AUX
alces-263	105	11	monitored	monitor	VERB
alces-263	105	12	using	use	VERB
alces-263	105	13	the	the	DET
alces-263	105	14	r	r	NOUN
alces-263	105	15	-	-	PUNCT
alces-263	105	16	m	m	NOUN
alces-263	105	17	equation	equation	NOUN
alces-263	105	18	based	base	VERB
alces-263	105	19	on	on	ADP
alces-263	105	20	annual	annual	ADJ
alces-263	105	21	surveys	survey	NOUN
alces-263	105	22	of	of	ADP
alces-263	105	23	recruitment	recruitment	NOUN
alces-263	105	24	and	and	CCONJ
alces-263	105	25	an	an	DET
alces-263	105	26	ongoing	ongoing	ADJ
alces-263	105	27	,	,	PUNCT
alces-263	105	28	intensive	intensive	ADJ
alces-263	105	29	ratio	ratio	NOUN
alces-263	105	30	telemetry	telemetry	NOUN
alces-263	105	31	program	program	NOUN
alces-263	105	32	with	with	ADP
alces-263	105	33	adult	adult	NOUN
alces-263	105	34	females	female	NOUN
alces-263	105	35	(	(	PUNCT
alces-263	105	36	hervieux	hervieux	PROPN
alces-263	105	37	et	et	PROPN
alces-263	105	38	al	al	PROPN
alces-263	105	39	.	.	PROPN
alces-263	105	40	2013	2013	NUM
alces-263	105	41	)	)	PUNCT
alces-263	105	42	.	.	PUNCT
alces-263	106	1	annual	annual	ADJ
alces-263	106	2	changes	change	NOUN
alces-263	106	3	in	in	ADP
alces-263	106	4	λ	λ	PROPN
alces-263	106	5	were	be	AUX
alces-263	106	6	used	use	VERB
alces-263	106	7	to	to	PART
alces-263	106	8	calculate	calculate	VERB
alces-263	106	9	realized	realize	VERB
alces-263	106	10	population	population	NOUN
alces-263	106	11	change	change	NOUN
alces-263	106	12	,	,	PUNCT
alces-263	106	13	which	which	PRON
alces-263	106	14	were	be	AUX
alces-263	106	15	the	the	DET
alces-263	106	16	successive	successive	ADJ
alces-263	106	17	product	product	NOUN
alces-263	106	18	of	of	ADP
alces-263	106	19	λ	λ	PROPN
alces-263	106	20	calculated	calculate	VERB
alces-263	106	21	from	from	ADP
alces-263	106	22	the	the	DET
alces-263	106	23	first	first	ADJ
alces-263	106	24	year	year	NOUN
alces-263	106	25	of	of	ADP
alces-263	106	26	monitoring	monitor	VERB
alces-263	106	27	up	up	ADP
alces-263	106	28	to	to	ADP
alces-263	106	29	and	and	CCONJ
alces-263	106	30	including	include	VERB
alces-263	106	31	the	the	DET
alces-263	106	32	most	most	ADV
alces-263	106	33	recent	recent	ADJ
alces-263	106	34	year	year	NOUN
alces-263	106	35	’s	’s	PART
alces-263	106	36	λ	λ	NOUN
alces-263	106	37	calculation	calculation	NOUN
alces-263	106	38	.	.	PUNCT
alces-263	107	1	because	because	SCONJ
alces-263	107	2	this	this	DET
alces-263	107	3	approach	approach	NOUN
alces-263	107	4	may	may	AUX
alces-263	107	5	compound	compound	VERB
alces-263	107	6	errors	error	NOUN
alces-263	107	7	in	in	ADP
alces-263	107	8	relative	relative	ADJ
alces-263	107	9	abundance	abundance	NOUN
alces-263	107	10	over	over	ADP
alces-263	107	11	time	time	NOUN
alces-263	107	12	(	(	PUNCT
alces-263	107	13	serrouya	serrouya	PROPN
alces-263	107	14	et	et	PROPN
alces-263	107	15	al	al	PROPN
alces-263	107	16	.	.	PROPN
alces-263	107	17	2017	2017	NUM
alces-263	107	18	)	)	PUNCT
alces-263	107	19	,	,	PUNCT
alces-263	107	20	periodic	periodic	ADJ
alces-263	107	21	surveys	survey	NOUN
alces-263	107	22	of	of	ADP
alces-263	107	23	absolute	absolute	ADJ
alces-263	107	24	abundance	abundance	NOUN
alces-263	107	25	should	should	AUX
alces-263	107	26	be	be	AUX
alces-263	107	27	performed	perform	VERB
alces-263	107	28	to	to	PART
alces-263	107	29	help	help	VERB
alces-263	107	30	validate	validate	VERB
alces-263	107	31	long	long	ADJ
alces-263	107	32	-	-	PUNCT
alces-263	107	33	term	term	NOUN
alces-263	107	34	population	population	NOUN
alces-263	107	35	trends	trend	NOUN
alces-263	107	36	based	base	VERB
alces-263	107	37	on	on	ADP
alces-263	107	38	the	the	DET
alces-263	107	39	r	r	PROPN
alces-263	107	40	-	-	PUNCT
alces-263	107	41	m	m	NOUN
alces-263	107	42	equation	equation	NOUN
alces-263	107	43	.	.	PUNCT
alces-263	108	1	the	the	DET
alces-263	108	2	r	r	PROPN
alces-263	108	3	-	-	PUNCT
alces-263	108	4	m	m	NOUN
alces-263	108	5	equation	equation	NOUN
alces-263	108	6	may	may	AUX
alces-263	108	7	provide	provide	VERB
alces-263	108	8	relatively	relatively	ADV
alces-263	108	9	large	large	ADJ
alces-263	108	10	confidence	confidence	NOUN
alces-263	108	11	intervals	interval	NOUN
alces-263	108	12	for	for	ADP
alces-263	108	13	λ	λ	PROPN
alces-263	108	14	.	.	PUNCT
alces-263	109	1	in	in	ADP
alces-263	109	2	order	order	NOUN
alces-263	109	3	to	to	PART
alces-263	109	4	improve	improve	VERB
alces-263	109	5	the	the	DET
alces-263	109	6	accuracy	accuracy	NOUN
alces-263	109	7	and	and	CCONJ
alces-263	109	8	precision	precision	NOUN
alces-263	109	9	of	of	ADP
alces-263	109	10	λ	λ	PROPN
alces-263	109	11	,	,	PUNCT
alces-263	109	12	it	it	PRON
alces-263	109	13	is	be	AUX
alces-263	109	14	important	important	ADJ
alces-263	109	15	to	to	PART
alces-263	109	16	ensure	ensure	VERB
alces-263	109	17	that	that	SCONJ
alces-263	109	18	samples	sample	NOUN
alces-263	109	19	of	of	ADP
alces-263	109	20	recruitment	recruitment	NOUN
alces-263	109	21	and	and	CCONJ
alces-263	109	22	mortality	mortality	NOUN
alces-263	109	23	rates	rate	NOUN
alces-263	109	24	are	be	AUX
alces-263	109	25	representative	representative	ADJ
alces-263	109	26	and	and	CCONJ
alces-263	109	27	that	that	SCONJ
alces-263	109	28	sample	sample	NOUN
alces-263	109	29	sizes	size	NOUN
alces-263	109	30	are	be	AUX
alces-263	109	31	sufficiently	sufficiently	ADV
alces-263	109	32	large	large	ADJ
alces-263	109	33	.	.	PUNCT
alces-263	110	1	decesare	decesare	NOUN
alces-263	110	2	et	et	PROPN
alces-263	110	3	al	al	PROPN
alces-263	110	4	.	.	PROPN
alces-263	111	1	(	(	PUNCT
alces-263	111	2	2012	2012	NUM
alces-263	111	3	)	)	PUNCT
alces-263	111	4	employed	employ	VERB
alces-263	111	5	elasticity	elasticity	NOUN
alces-263	111	6	and	and	CCONJ
alces-263	111	7	life	life	NOUN
alces-263	111	8	-	-	PUNCT
alces-263	111	9	stage	stage	NOUN
alces-263	111	10	simulation	simulation	NOUN
alces-263	111	11	analysis	analysis	NOUN
alces-263	111	12	for	for	ADP
alces-263	111	13	a	a	DET
alces-263	111	14	woodland	woodland	NOUN
alces-263	111	15	caribou	caribou	NOUN
alces-263	111	16	population	population	NOUN
alces-263	111	17	in	in	ADP
alces-263	111	18	alberta	alberta	PROPN
alces-263	111	19	and	and	CCONJ
alces-263	111	20	found	find	VERB
alces-263	111	21	that	that	SCONJ
alces-263	111	22	adult	adult	NOUN
alces-263	111	23	female	female	ADJ
alces-263	111	24	survival	survival	NOUN
alces-263	111	25	and	and	CCONJ
alces-263	111	26	recruitment	recruitment	NOUN
alces-263	111	27	rates	rate	NOUN
alces-263	111	28	were	be	AUX
alces-263	111	29	nearly	nearly	ADV
alces-263	111	30	equivalent	equivalent	ADJ
alces-263	111	31	drivers	driver	NOUN
alces-263	111	32	of	of	ADP
alces-263	111	33	population	population	NOUN
alces-263	111	34	growth	growth	NOUN
alces-263	111	35	.	.	PUNCT
alces-263	112	1	this	this	PRON
alces-263	112	2	suggests	suggest	VERB
alces-263	112	3	that	that	SCONJ
alces-263	112	4	increased	increase	VERB
alces-263	112	5	sampling	sample	VERB
alces-263	112	6	to	to	PART
alces-263	112	7	improve	improve	VERB
alces-263	112	8	precision	precision	NOUN
alces-263	112	9	of	of	ADP
alces-263	112	10	λ	λ	PROPN
alces-263	112	11	should	should	AUX
alces-263	112	12	be	be	AUX
alces-263	112	13	directly	directly	ADV
alces-263	112	14	towards	towards	ADP
alces-263	112	15	both	both	DET
alces-263	112	16	population	population	NOUN
alces-263	112	17	parameters	parameter	NOUN
alces-263	112	18	.	.	PUNCT
alces-263	113	1	alternatively	alternatively	ADV
alces-263	113	2	,	,	PUNCT
alces-263	113	3	annual	annual	ADJ
alces-263	113	4	estimates	estimate	NOUN
alces-263	113	5	of	of	ADP
alces-263	113	6	j	j	PROPN
alces-263	113	7	/	/	SYM
alces-263	113	8	f	f	PROPN
alces-263	113	9	,	,	PUNCT
alces-263	113	10	sf	sf	PROPN
alces-263	113	11	,	,	PUNCT
alces-263	113	12	sfw	sfw	NOUN
alces-263	113	13	,	,	PUNCT
alces-263	113	14	and	and	CCONJ
alces-263	113	15	sjw	sjw	PROPN
alces-263	113	16	may	may	AUX
alces-263	113	17	be	be	AUX
alces-263	113	18	merged	merge	VERB
alces-263	113	19	over	over	ADP
alces-263	113	20	multiple	multiple	ADJ
alces-263	113	21	years	year	NOUN
alces-263	113	22	to	to	PART
alces-263	113	23	improve	improve	VERB
alces-263	113	24	precision	precision	NOUN
alces-263	113	25	,	,	PUNCT
alces-263	113	26	providing	provide	VERB
alces-263	113	27	these	these	DET
alces-263	113	28	parameters	parameter	NOUN
alces-263	113	29	are	be	AUX
alces-263	113	30	not	not	PART
alces-263	113	31	significantly	significantly	ADV
alces-263	113	32	different	different	ADJ
alces-263	113	33	between	between	ADP
alces-263	113	34	years	year	NOUN
alces-263	113	35	.	.	PUNCT
alces-263	114	1	biologists	biologist	NOUN
alces-263	114	2	have	have	AUX
alces-263	114	3	been	be	AUX
alces-263	114	4	tasked	task	VERB
alces-263	114	5	with	with	ADP
alces-263	114	6	ensuring	ensure	VERB
alces-263	114	7	moose	moose	NOUN
alces-263	114	8	objectives	objective	NOUN
alces-263	114	9	for	for	ADP
alces-263	114	10	both	both	DET
alces-263	114	11	conservation	conservation	NOUN
alces-263	114	12	alces	alce	NOUN
alces-263	114	13	vol	vol	NOUN
alces-263	114	14	.	.	PROPN
alces-263	115	1	56	56	NUM
alces-263	115	2	,	,	PUNCT
alces-263	115	3	2020	2020	NUM
alces-263	115	4	revisiting	revisit	VERB
alces-263	115	5	the	the	DET
alces-263	115	6	recruitment	recruitment	NOUN
alces-263	115	7	-	-	PUNCT
alces-263	115	8	mortality	mortality	NOUN
alces-263	115	9	.	.	PUNCT
alces-263	116	1	–	–	PUNCT
alces-263	116	2	ian	ian	PROPN
alces-263	116	3	w.	w.	PROPN
alces-263	116	4	hatter	hatter	PROPN
alces-263	116	5	45	45	NUM
alces-263	116	6	and	and	CCONJ
alces-263	116	7	sustainable	sustainable	ADJ
alces-263	116	8	use	use	NOUN
alces-263	116	9	are	be	AUX
alces-263	116	10	met	meet	VERB
alces-263	116	11	.	.	PUNCT
alces-263	117	1	these	these	DET
alces-263	117	2	challenges	challenge	NOUN
alces-263	117	3	have	have	AUX
alces-263	117	4	recently	recently	ADV
alces-263	117	5	intensified	intensify	VERB
alces-263	117	6	with	with	ADP
alces-263	117	7	broad	broad	ADJ
alces-263	117	8	scale	scale	NOUN
alces-263	117	9	moose	moose	NOUN
alces-263	117	10	declines	decline	NOUN
alces-263	117	11	occurring	occur	VERB
alces-263	117	12	in	in	ADP
alces-263	117	13	parts	part	NOUN
alces-263	117	14	of	of	ADP
alces-263	117	15	north	north	PROPN
alces-263	117	16	america	america	PROPN
alces-263	117	17	(	(	PUNCT
alces-263	117	18	timmermann	timmermann	VERB
alces-263	117	19	and	and	CCONJ
alces-263	117	20	rodgers	rodger	VERB
alces-263	117	21	2017	2017	NUM
alces-263	117	22	,	,	PUNCT
alces-263	117	23	kuzyk	kuzyk	INTJ
alces-263	117	24	et	et	PROPN
alces-263	117	25	al	al	PROPN
alces-263	117	26	.	.	PROPN
alces-263	117	27	2018	2018	NUM
alces-263	117	28	)	)	PUNCT
alces-263	117	29	.	.	PUNCT
alces-263	118	1	the	the	DET
alces-263	118	2	r	r	NOUN
alces-263	118	3	-	-	PUNCT
alces-263	118	4	m	m	NOUN
alces-263	118	5	equation	equation	NOUN
alces-263	118	6	provides	provide	VERB
alces-263	118	7	a	a	DET
alces-263	118	8	simple	simple	ADJ
alces-263	118	9	and	and	CCONJ
alces-263	118	10	relatively	relatively	ADV
alces-263	118	11	inexpensive	inexpensive	ADJ
alces-263	118	12	method	method	NOUN
alces-263	118	13	to	to	PART
alces-263	118	14	rapidly	rapidly	ADV
alces-263	118	15	assess	assess	VERB
alces-263	118	16	moose	moose	NOUN
alces-263	118	17	population	population	NOUN
alces-263	118	18	trends	trend	NOUN
alces-263	118	19	when	when	SCONJ
alces-263	118	20	adult	adult	NOUN
alces-263	118	21	survival	survival	NOUN
alces-263	118	22	rates	rate	NOUN
alces-263	118	23	are	be	AUX
alces-263	118	24	available	available	ADJ
alces-263	118	25	.	.	PUNCT
alces-263	119	1	however	however	ADV
alces-263	119	2	,	,	PUNCT
alces-263	119	3	numerous	numerous	ADJ
alces-263	119	4	biases	bias	NOUN
alces-263	119	5	may	may	AUX
alces-263	119	6	exist	exist	VERB
alces-263	119	7	in	in	ADP
alces-263	119	8	estimating	estimate	VERB
alces-263	119	9	recruitment	recruitment	NOUN
alces-263	119	10	and	and	CCONJ
alces-263	119	11	mortality	mortality	NOUN
alces-263	119	12	rates	rate	NOUN
alces-263	119	13	that	that	PRON
alces-263	119	14	can	can	AUX
alces-263	119	15	substantially	substantially	ADV
alces-263	119	16	affect	affect	VERB
alces-263	119	17	estimates	estimate	NOUN
alces-263	119	18	of	of	ADP
alces-263	119	19	λ	λ	PROPN
alces-263	119	20	(	(	PUNCT
alces-263	119	21	serrouya	serrouya	PROPN
alces-263	119	22	et	et	PROPN
alces-263	119	23	al	al	PROPN
alces-263	119	24	.	.	PROPN
alces-263	119	25	2017	2017	NUM
alces-263	119	26	,	,	PUNCT
alces-263	119	27	severud	severud	NOUN
alces-263	119	28	et	et	PROPN
alces-263	119	29	al	al	PROPN
alces-263	119	30	.	.	PROPN
alces-263	119	31	2019	2019	NUM
alces-263	119	32	)	)	PUNCT
alces-263	119	33	.	.	PUNCT
alces-263	120	1	thus	thus	ADV
alces-263	120	2	,	,	PUNCT
alces-263	120	3	it	it	PRON
alces-263	120	4	is	be	AUX
alces-263	120	5	important	important	ADJ
alces-263	120	6	to	to	PART
alces-263	120	7	assess	assess	VERB
alces-263	120	8	the	the	DET
alces-263	120	9	accuracy	accuracy	NOUN
alces-263	120	10	and	and	CCONJ
alces-263	120	11	precision	precision	NOUN
alces-263	120	12	of	of	ADP
alces-263	120	13	both	both	DET
alces-263	120	14	parameters	parameter	NOUN
alces-263	120	15	when	when	SCONJ
alces-263	120	16	calculating	calculate	VERB
alces-263	120	17	λ	λ	NOUN
alces-263	120	18	using	use	VERB
alces-263	120	19	the	the	DET
alces-263	120	20	r	r	NOUN
alces-263	120	21	-	-	PUNCT
alces-263	120	22	m	m	NOUN
alces-263	120	23	equation	equation	NOUN
alces-263	120	24	and	and	CCONJ
alces-263	120	25	to	to	PART
alces-263	120	26	ensure	ensure	VERB
alces-263	120	27	that	that	SCONJ
alces-263	120	28	over	over	ADP
alces-263	120	29	-	-	PUNCT
alces-263	120	30	winter	winter	NOUN
alces-263	120	31	differential	differential	NOUN
alces-263	120	32	mortality	mortality	NOUN
alces-263	120	33	between	between	ADP
alces-263	120	34	cows	cow	NOUN
alces-263	120	35	and	and	CCONJ
alces-263	120	36	calves	calf	NOUN
alces-263	120	37	is	be	AUX
alces-263	120	38	incorporated	incorporate	VERB
alces-263	120	39	.	.	PUNCT
alces-263	121	1	if	if	SCONJ
alces-263	121	2	the	the	DET
alces-263	121	3	r	r	NOUN
alces-263	121	4	-	-	PUNCT
alces-263	121	5	m	m	NOUN
alces-263	121	6	equation	equation	NOUN
alces-263	121	7	is	be	AUX
alces-263	121	8	used	use	VERB
alces-263	121	9	to	to	PART
alces-263	121	10	monitor	monitor	VERB
alces-263	121	11	trends	trend	NOUN
alces-263	121	12	over	over	ADP
alces-263	121	13	multiple	multiple	ADJ
alces-263	121	14	years	year	NOUN
alces-263	121	15	,	,	PUNCT
alces-263	121	16	then	then	ADV
alces-263	121	17	estimates	estimate	NOUN
alces-263	121	18	of	of	ADP
alces-263	121	19	λ	λ	PROPN
alces-263	121	20	should	should	AUX
alces-263	121	21	be	be	AUX
alces-263	121	22	validated	validate	VERB
alces-263	121	23	periodically	periodically	ADV
alces-263	121	24	with	with	ADP
alces-263	121	25	surveys	survey	NOUN
alces-263	121	26	of	of	ADP
alces-263	121	27	absolute	absolute	ADJ
alces-263	121	28	abundance	abundance	NOUN
alces-263	121	29	.	.	PUNCT
alces-263	122	1	acknowledgements	acknowledgement	NOUN
alces-263	122	2	g.	g.	PROPN
alces-263	122	3	kuzyk	kuzyk	PROPN
alces-263	122	4	,	,	PUNCT
alces-263	122	5	c.	c.	PROPN
alces-263	122	6	proctor	proctor	PROPN
alces-263	122	7	,	,	PUNCT
alces-263	122	8	and	and	CCONJ
alces-263	122	9	s.	s.	PROPN
alces-263	122	10	marshall	marshall	PROPN
alces-263	122	11	provided	provide	VERB
alces-263	122	12	access	access	NOUN
alces-263	122	13	to	to	ADP
alces-263	122	14	the	the	DET
alces-263	122	15	british	british	PROPN
alces-263	122	16	columbia	columbia	PROPN
alces-263	122	17	moose	moose	PROPN
alces-263	122	18	data	datum	NOUN
alces-263	122	19	sets	set	NOUN
alces-263	122	20	used	use	VERB
alces-263	122	21	in	in	ADP
alces-263	122	22	the	the	DET
alces-263	122	23	example	example	NOUN
alces-263	122	24	.	.	PUNCT
alces-263	123	1	m.	m.	NOUN
alces-263	123	2	scheideman	scheideman	PROPN
alces-263	123	3	kindly	kindly	ADV
alces-263	123	4	produced	produce	VERB
alces-263	123	5	the	the	DET
alces-263	123	6	estimates	estimate	NOUN
alces-263	123	7	of	of	ADP
alces-263	123	8	cow	cow	PROPN
alces-263	123	9	winter	winter	NOUN
alces-263	123	10	survival	survival	NOUN
alces-263	123	11	rates	rate	NOUN
alces-263	123	12	and	and	CCONJ
alces-263	123	13	provided	provide	VERB
alces-263	123	14	the	the	DET
alces-263	123	15	winter	winter	NOUN
alces-263	123	16	calf	calf	NOUN
alces-263	123	17	sex	sex	NOUN
alces-263	123	18	ratios	ratio	NOUN
alces-263	123	19	.	.	PUNCT
alces-263	124	1	i	i	PRON
alces-263	124	2	appreciate	appreciate	VERB
alces-263	124	3	the	the	DET
alces-263	124	4	constructive	constructive	ADJ
alces-263	124	5	reviews	review	NOUN
alces-263	124	6	from	from	ADP
alces-263	124	7	m.	m.	NOUN
alces-263	124	8	carstensen	carstensen	NOUN
alces-263	124	9	and	and	CCONJ
alces-263	124	10	2	2	NUM
alces-263	124	11	anonymous	anonymous	ADJ
alces-263	124	12	reviewers	reviewer	NOUN
alces-263	124	13	which	which	PRON
alces-263	124	14	greatly	greatly	ADV
alces-263	124	15	improved	improve	VERB
alces-263	124	16	an	an	DET
alces-263	124	17	earlier	early	ADJ
alces-263	124	18	draft	draft	NOUN
alces-263	124	19	of	of	ADP
alces-263	124	20	this	this	DET
alces-263	124	21	manuscript	manuscript	NOUN
alces-263	124	22	.	.	PUNCT
alces-263	125	1	this	this	DET
alces-263	125	2	study	study	NOUN
alces-263	125	3	is	be	AUX
alces-263	125	4	dedicated	dedicate	VERB
alces-263	125	5	to	to	ADP
alces-263	125	6	the	the	DET
alces-263	125	7	late	late	PROPN
alces-263	125	8	w.	w.	PROPN
alces-263	125	9	a.	a.	NOUN
alces-263	125	10	bergerud	bergerud	PROPN
alces-263	125	11	who	who	PRON
alces-263	125	12	developed	develop	VERB
alces-263	125	13	the	the	DET
alces-263	125	14	proof	proof	NOUN
alces-263	125	15	for	for	ADP
alces-263	125	16	the	the	DET
alces-263	125	17	original	original	ADJ
alces-263	125	18	r	r	NOUN
alces-263	125	19	-	-	PUNCT
alces-263	125	20	m	m	NOUN
alces-263	125	21	equation	equation	NOUN
alces-263	125	22	.	.	PUNCT
alces-263	126	1	references	reference	NOUN
alces-263	126	2	ballard	ballard	PROPN
alces-263	126	3	,	,	PUNCT
alces-263	126	4	w.	w.	PROPN
alces-263	126	5	b.	b.	PROPN
alces-263	126	6	,	,	PUNCT
alces-263	126	7	j.	j.	PROPN
alces-263	126	8	s.	s.	PROPN
alces-263	126	9	whitman	whitman	PROPN
alces-263	126	10	,	,	PUNCT
alces-263	126	11	and	and	CCONJ
alces-263	126	12	d.	d.	PROPN
alces-263	126	13	j.	j.	PROPN
alces-263	126	14	reed	reed	PROPN
alces-263	126	15	.	.	PUNCT
alces-263	127	1	1991	1991	NUM
alces-263	127	2	.	.	PUNCT
alces-263	128	1	population	population	NOUN
alces-263	128	2	dynamics	dynamic	NOUN
alces-263	128	3	of	of	ADP
alces-263	128	4	moose	moose	NOUN
alces-263	128	5	in	in	ADP
alces-263	128	6	south	south	ADJ
alces-263	128	7	-	-	PUNCT
alces-263	128	8	central	central	ADJ
alces-263	128	9	alaska	alaska	NOUN
alces-263	128	10	.	.	PUNCT
alces-263	129	1	wildlife	wildlife	NOUN
alces-263	129	2	monographs	monograph	NOUN
alces-263	129	3	114	114	NUM
alces-263	129	4	.	.	PUNCT
alces-263	130	1	bergerud	bergerud	NOUN
alces-263	130	2	,	,	PUNCT
alces-263	130	3	a.	a.	NOUN
alces-263	130	4	t.	t.	PROPN
alces-263	130	5	,	,	PUNCT
alces-263	130	6	and	and	CCONJ
alces-263	130	7	j.	j.	PROPN
alces-263	130	8	p.	p.	PROPN
alces-263	130	9	elliott	elliott	PROPN
alces-263	130	10	.	.	PROPN
alces-263	131	1	1986	1986	NUM
alces-263	131	2	.	.	PUNCT
alces-263	132	1	dynamics	dynamic	NOUN
alces-263	132	2	of	of	ADP
alces-263	132	3	caribou	caribou	NOUN
alces-263	132	4	and	and	CCONJ
alces-263	132	5	wolves	wolf	NOUN
alces-263	132	6	in	in	ADP
alces-263	132	7	northern	northern	ADJ
alces-263	132	8	british	british	PROPN
alces-263	132	9	columbia	columbia	PROPN
alces-263	132	10	.	.	PUNCT
alces-263	133	1	canadian	canadian	ADJ
alces-263	133	2	journal	journal	PROPN
alces-263	133	3	of	of	ADP
alces-263	133	4	zoology	zoology	NOUN
alces-263	133	5	64	64	NUM
alces-263	133	6	:	:	PUNCT
alces-263	133	7	1515–1529	1515–1529	NUM
alces-263	133	8	.	.	PUNCT
alces-263	134	1	doi	doi	NOUN
alces-263	134	2	:	:	PUNCT
alces-263	134	3	10.1139	10.1139	NUM
alces-263	134	4	/	/	SYM
alces-263	134	5	z86	z86	PROPN
alces-263	134	6	-	-	PUNCT
alces-263	134	7	226	226	NUM
alces-263	134	8	boertje	boertje	NOUN
alces-263	134	9	,	,	PUNCT
alces-263	134	10	r.	r.	PROPN
alces-263	134	11	d.	d.	PROPN
alces-263	134	12	,	,	PUNCT
alces-263	134	13	p.	p.	PROPN
alces-263	134	14	valkenburg	valkenburg	PROPN
alces-263	134	15	,	,	PUNCT
alces-263	134	16	and	and	CCONJ
alces-263	134	17	m.	m.	PROPN
alces-263	134	18	e.	e.	PROPN
alces-263	134	19	mcnay	mcnay	PROPN
alces-263	134	20	1996	1996	NUM
alces-263	134	21	.	.	PUNCT
alces-263	135	1	increases	increase	NOUN
alces-263	135	2	in	in	ADP
alces-263	135	3	moose	moose	NOUN
alces-263	135	4	,	,	PUNCT
alces-263	135	5	caribou	caribou	NOUN
alces-263	135	6	and	and	CCONJ
alces-263	135	7	wolves	wolf	NOUN
alces-263	135	8	following	follow	VERB
alces-263	135	9	wolf	wolf	PROPN
alces-263	135	10	control	control	NOUN
alces-263	135	11	in	in	ADP
alces-263	135	12	alaska	alaska	PROPN
alces-263	135	13	.	.	PUNCT
alces-263	136	1	journal	journal	PROPN
alces-263	136	2	of	of	ADP
alces-263	136	3	wildlife	wildlife	NOUN
alces-263	136	4	management	management	NOUN
alces-263	136	5	80	80	NUM
alces-263	136	6	:	:	PUNCT
alces-263	136	7	474–479	474–479	NUM
alces-263	136	8	.	.	PUNCT
alces-263	137	1	doi	doi	NOUN
alces-263	137	2	:	:	PUNCT
alces-263	137	3	10.2307/3802065	10.2307/3802065	NUM
alces-263	137	4	caughley	caughley	NOUN
alces-263	137	5	,	,	PUNCT
alces-263	137	6	g.	g.	PROPN
alces-263	137	7	1977	1977	NUM
alces-263	137	8	.	.	PUNCT
alces-263	138	1	analysis	analysis	NOUN
alces-263	138	2	of	of	ADP
alces-263	138	3	vertebrate	vertebrate	ADJ
alces-263	138	4	populations	population	NOUN
alces-263	138	5	.	.	PUNCT
alces-263	139	1	john	john	PROPN
alces-263	139	2	wiley	wiley	PROPN
alces-263	139	3	&	&	CCONJ
alces-263	139	4	sons	sons	PROPN
alces-263	139	5	,	,	PUNCT
alces-263	139	6	london	london	PROPN
alces-263	139	7	,	,	PUNCT
alces-263	139	8	united	united	PROPN
alces-263	139	9	kingdom	kingdom	PROPN
alces-263	139	10	.	.	PUNCT
alces-263	140	1	decesare	decesare	PROPN
alces-263	140	2	,	,	PUNCT
alces-263	140	3	n.	n.	PROPN
alces-263	140	4	j.	j.	PROPN
alces-263	140	5	,	,	PUNCT
alces-263	140	6	m.	m.	NOUN
alces-263	140	7	hebblewhite	hebblewhite	PROPN
alces-263	140	8	,	,	PUNCT
alces-263	140	9	m.	m.	PROPN
alces-263	140	10	bradley	bradley	PROPN
alces-263	140	11	,	,	PUNCT
alces-263	140	12	k.	k.	PROPN
alces-263	140	13	smith	smith	PROPN
alces-263	140	14	,	,	PUNCT
alces-263	140	15	d.	d.	PROPN
alces-263	140	16	hervieux	hervieux	PROPN
alces-263	140	17	,	,	PUNCT
alces-263	140	18	and	and	CCONJ
alces-263	140	19	l.	l.	PROPN
alces-263	140	20	neufeld	neufeld	PROPN
alces-263	140	21	.	.	PUNCT
alces-263	141	1	2012	2012	NUM
alces-263	141	2	.	.	PUNCT
alces-263	142	1	estimating	estimate	VERB
alces-263	142	2	ungulate	ungulate	ADJ
alces-263	142	3	recruitment	recruitment	NOUN
alces-263	142	4	and	and	CCONJ
alces-263	142	5	growth	growth	NOUN
alces-263	142	6	rates	rate	NOUN
alces-263	142	7	using	use	VERB
alces-263	142	8	age	age	NOUN
alces-263	142	9	ratios	ratio	NOUN
alces-263	142	10	.	.	PUNCT
alces-263	143	1	journal	journal	NOUN
alces-263	143	2	of	of	ADP
alces-263	143	3	wildlife	wildlife	NOUN
alces-263	143	4	management	management	NOUN
alces-263	143	5	76	76	NUM
alces-263	143	6	:	:	PUNCT
alces-263	143	7	144–153	144–153	NUM
alces-263	143	8	.	.	PUNCT
alces-263	144	1	doi	doi	NOUN
alces-263	144	2	:	:	PUNCT
alces-263	144	3	10.1002	10.1002	NUM
alces-263	144	4	/	/	SYM
alces-263	144	5	jwmg.244	jwmg.244	PROPN
alces-263	144	6	devore	devore	NOUN
alces-263	144	7	,	,	PUNCT
alces-263	144	8	r.	r.	PROPN
alces-263	144	9	m.	m.	PROPN
alces-263	144	10	,	,	PUNCT
alces-263	144	11	m.	m.	PROPN
alces-263	144	12	j.	j.	PROPN
alces-263	144	13	butler	butler	PROPN
alces-263	144	14	,	,	PUNCT
alces-263	144	15	m.	m.	PROPN
alces-263	144	16	c.	c.	PROPN
alces-263	144	17	wallace	wallace	PROPN
alces-263	144	18	,	,	PUNCT
alces-263	144	19	and	and	CCONJ
alces-263	144	20	s.	s.	PROPN
alces-263	144	21	g.	g.	PROPN
alces-263	144	22	liley	liley	PROPN
alces-263	144	23	.	.	PUNCT
alces-263	145	1	2018	2018	NUM
alces-263	145	2	.	.	PUNCT
alces-263	146	1	population	population	NOUN
alces-263	146	2	dynamics	dynamic	NOUN
alces-263	146	3	model	model	NOUN
alces-263	146	4	to	to	PART
alces-263	146	5	inform	inform	VERB
alces-263	146	6	harvest	harvest	NOUN
alces-263	146	7	management	management	NOUN
alces-263	146	8	of	of	ADP
alces-263	146	9	a	a	DET
alces-263	146	10	small	small	ADJ
alces-263	146	11	elk	elk	NOUN
alces-263	146	12	herd	herd	NOUN
alces-263	146	13	in	in	ADP
alces-263	146	14	central	central	ADJ
alces-263	146	15	new	new	PROPN
alces-263	146	16	mexico	mexico	PROPN
alces-263	146	17	.	.	PUNCT
alces-263	147	1	journal	journal	PROPN
alces-263	147	2	of	of	ADP
alces-263	147	3	fish	fish	NOUN
alces-263	147	4	and	and	CCONJ
alces-263	147	5	wildlife	wildlife	NOUN
alces-263	147	6	management	management	NOUN
alces-263	147	7	9	9	NUM
alces-263	147	8	:	:	PUNCT
alces-263	147	9	531–544	531–544	NUM
alces-263	147	10	.	.	PUNCT
alces-263	147	11	doi	doi	NOUN
alces-263	147	12	:	:	PUNCT
alces-263	147	13	10.3996/012018	10.3996/012018	NUM
alces-263	147	14	-	-	PUNCT
alces-263	147	15	jfwm-008	jfwm-008	NOUN
alces-263	147	16	gasaway	gasaway	NOUN
alces-263	147	17	,	,	PUNCT
alces-263	147	18	w.	w.	PROPN
alces-263	147	19	c	c	PROPN
alces-263	147	20	,	,	PUNCT
alces-263	147	21	r.	r.	PROPN
alces-263	147	22	d.	d.	PROPN
alces-263	147	23	boertje	boertje	PROPN
alces-263	147	24	,	,	PUNCT
alces-263	147	25	d.	d.	PROPN
alces-263	147	26	v.	v.	PROPN
alces-263	147	27	grangaard	grangaard	PROPN
alces-263	147	28	,	,	PUNCT
alces-263	147	29	d.	d.	PROPN
alces-263	147	30	g.	g.	PROPN
alces-263	147	31	kelleyhouse	kelleyhouse	PROPN
alces-263	147	32	,	,	PUNCT
alces-263	147	33	r.	r.	PROPN
alces-263	147	34	o.	o.	PROPN
alces-263	147	35	stephenson	stephenson	PROPN
alces-263	147	36	,	,	PUNCT
alces-263	147	37	and	and	CCONJ
alces-263	147	38	d.	d.	PROPN
alces-263	147	39	g.	g.	PROPN
alces-263	147	40	larson	larson	PROPN
alces-263	147	41	.	.	PROPN
alces-263	147	42	1992	1992	NUM
alces-263	147	43	.	.	PUNCT
alces-263	148	1	the	the	DET
alces-263	148	2	role	role	NOUN
alces-263	148	3	of	of	ADP
alces-263	148	4	predation	predation	NOUN
alces-263	148	5	in	in	ADP
alces-263	148	6	limiting	limit	VERB
alces-263	148	7	moose	moose	NOUN
alces-263	148	8	at	at	ADP
alces-263	148	9	low	low	ADJ
alces-263	148	10	densities	density	NOUN
alces-263	148	11	in	in	ADP
alces-263	148	12	alaska	alaska	PROPN
alces-263	148	13	and	and	CCONJ
alces-263	148	14	yukon	yukon	PROPN
alces-263	148	15	and	and	CCONJ
alces-263	148	16	implications	implication	NOUN
alces-263	148	17	for	for	ADP
alces-263	148	18	conservation	conservation	NOUN
alces-263	148	19	.	.	PUNCT
alces-263	149	1	wildlife	wildlife	NOUN
alces-263	149	2	monographs	monograph	NOUN
alces-263	149	3	120	120	NUM
alces-263	149	4	:	:	PUNCT
alces-263	149	5	1–59	1–59	NOUN
alces-263	149	6	.	.	PUNCT
alces-263	150	1	_	_	PUNCT
alces-263	151	1	_	_	PUNCT
alces-263	152	1	_	_	PUNCT
alces-263	153	1	_	_	PUNCT
alces-263	154	1	_	_	PUNCT
alces-263	154	2	,	,	PUNCT
alces-263	154	3	s.	s.	PROPN
alces-263	154	4	d.	d.	PROPN
alces-263	154	5	dubois	dubois	PROPN
alces-263	154	6	,	,	PUNCT
alces-263	154	7	d.	d.	PROPN
alces-263	154	8	j.	j.	PROPN
alces-263	154	9	reed	reed	PROPN
alces-263	154	10	,	,	PUNCT
alces-263	154	11	and	and	CCONJ
alces-263	154	12	s.	s.	PROPN
alces-263	154	13	j.	j.	PROPN
alces-263	154	14	harbo	harbo	PROPN
alces-263	154	15	.	.	PUNCT
alces-263	155	1	1986	1986	NUM
alces-263	155	2	.	.	PUNCT
alces-263	156	1	estimating	estimate	VERB
alces-263	156	2	moose	moose	NOUN
alces-263	156	3	population	population	NOUN
alces-263	156	4	parameters	parameter	NOUN
alces-263	156	5	from	from	ADP
alces-263	156	6	aerial	aerial	ADJ
alces-263	156	7	surveys	survey	NOUN
alces-263	156	8	.	.	PUNCT
alces-263	157	1	biological	biological	ADJ
alces-263	157	2	papers	paper	NOUN
alces-263	157	3	of	of	ADP
alces-263	157	4	the	the	DET
alces-263	157	5	university	university	PROPN
alces-263	157	6	of	of	ADP
alces-263	157	7	alaska	alaska	PROPN
alces-263	157	8	.	.	PUNCT
alces-263	158	1	number	number	NOUN
alces-263	158	2	22	22	NUM
alces-263	158	3	.	.	PUNCT
alces-263	159	1	institute	institute	PROPN
alces-263	159	2	of	of	ADP
alces-263	159	3	arctic	arctic	PROPN
alces-263	159	4	biology	biology	PROPN
alces-263	159	5	,	,	PUNCT
alces-263	159	6	fairbanks	fairbanks	PROPN
alces-263	159	7	,	,	PUNCT
alces-263	159	8	alaska	alaska	PROPN
alces-263	159	9	,	,	PUNCT
alces-263	159	10	usa	usa	PROPN
alces-263	159	11	.	.	PROPN
alces-263	159	12	guthery	guthery	PROPN
alces-263	159	13	,	,	PUNCT
alces-263	159	14	f.	f.	PROPN
alces-263	159	15	s.	s.	PROPN
alces-263	159	16	,	,	PUNCT
alces-263	159	17	and	and	CCONJ
alces-263	159	18	j.	j.	PROPN
alces-263	159	19	h.	h.	PROPN
alces-263	159	20	shaw	shaw	PROPN
alces-263	159	21	.	.	PROPN
alces-263	160	1	2013	2013	NUM
alces-263	160	2	.	.	PUNCT
alces-263	161	1	density	density	NOUN
alces-263	161	2	dependence	dependence	NOUN
alces-263	161	3	:	:	PUNCT
alces-263	161	4	applications	application	NOUN
alces-263	161	5	in	in	ADP
alces-263	161	6	wildlife	wildlife	NOUN
alces-263	161	7	management	management	NOUN
alces-263	161	8	.	.	PUNCT
alces-263	162	1	journal	journal	NOUN
alces-263	162	2	of	of	ADP
alces-263	162	3	wildlife	wildlife	NOUN
alces-263	162	4	management	management	NOUN
alces-263	162	5	77	77	NUM
alces-263	162	6	:	:	PUNCT
alces-263	162	7	33–38	33–38	NUM
alces-263	162	8	.	.	PUNCT
alces-263	163	1	doi	doi	NOUN
alces-263	163	2	:	:	PUNCT
alces-263	163	3	10.1002	10.1002	NUM
alces-263	163	4	/	/	SYM
alces-263	163	5	jwmg.450	jwmg.450	NOUN
alces-263	163	6	hatter	hatter	NOUN
alces-263	163	7	,	,	PUNCT
alces-263	163	8	i.	i.	PROPN
alces-263	163	9	w.	w.	PROPN
alces-263	163	10	,	,	PUNCT
alces-263	163	11	and	and	CCONJ
alces-263	163	12	w.	w.	PROPN
alces-263	163	13	a.	a.	NOUN
alces-263	163	14	bergerud	bergerud	PROPN
alces-263	163	15	.	.	PUNCT
alces-263	164	1	1991	1991	NUM
alces-263	164	2	.	.	PUNCT
alces-263	165	1	moose	moose	NOUN
alces-263	165	2	recruitment	recruitment	NOUN
alces-263	165	3	,	,	PUNCT
alces-263	165	4	adult	adult	NOUN
alces-263	165	5	mortality	mortality	NOUN
alces-263	165	6	and	and	CCONJ
alces-263	165	7	rate	rate	NOUN
alces-263	165	8	of	of	ADP
alces-263	165	9	change	change	NOUN
alces-263	165	10	.	.	PUNCT
alces-263	166	1	alces	alce	NOUN
alces-263	166	2	27	27	NUM
alces-263	166	3	:	:	PUNCT
alces-263	166	4	65–73	65–73	NUM
alces-263	166	5	.	.	PUNCT
alces-263	167	1	_	_	PUNCT
alces-263	168	1	_	_	PUNCT
alces-263	169	1	_	_	PUNCT
alces-263	170	1	_	_	PUNCT
alces-263	170	2	_	_	PUNCT
alces-263	170	3	,	,	PUNCT
alces-263	170	4	and	and	CCONJ
alces-263	170	5	d.	d.	PROPN
alces-263	170	6	w.	w.	PROPN
alces-263	170	7	janz	janz	PROPN
alces-263	170	8	.	.	PUNCT
alces-263	171	1	1994	1994	NUM
alces-263	171	2	.	.	PUNCT
alces-263	172	1	apparent	apparent	ADJ
alces-263	172	2	demographic	demographic	ADJ
alces-263	172	3	changes	change	NOUN
alces-263	172	4	in	in	ADP
alces-263	172	5	black	black	ADJ
alces-263	172	6	-	-	PUNCT
alces-263	172	7	tailed	tail	VERB
alces-263	172	8	deer	deer	NOUN
alces-263	172	9	associated	associate	VERB
alces-263	172	10	with	with	ADP
alces-263	172	11	wolf	wolf	PROPN
alces-263	172	12	control	control	NOUN
alces-263	172	13	on	on	ADP
alces-263	172	14	northern	northern	PROPN
alces-263	172	15	vancouver	vancouver	PROPN
alces-263	172	16	island	island	PROPN
alces-263	172	17	.	.	PUNCT
alces-263	173	1	canadian	canadian	PROPN
alces-263	173	2	revisiting	revisit	VERB
alces-263	173	3	the	the	DET
alces-263	173	4	r	r	PROPN
alces-263	173	5	-	-	PUNCT
alces-263	173	6	m	m	NOUN
alces-263	173	7	equation	equation	NOUN
alces-263	173	8	.	.	PUNCT
alces-263	174	1	–	–	PUNCT
alces-263	174	2	ian	ian	PROPN
alces-263	174	3	w.	w.	PROPN
alces-263	174	4	hatter	hatter	PROPN
alces-263	174	5	alces	alces	PROPN
alces-263	174	6	vol	vol	NOUN
alces-263	174	7	.	.	PROPN
alces-263	175	1	56	56	NUM
alces-263	175	2	,	,	PUNCT
alces-263	175	3	2020	2020	NUM
alces-263	175	4	46	46	NUM
alces-263	175	5	journal	journal	NOUN
alces-263	175	6	of	of	ADP
alces-263	175	7	zoology	zoology	NOUN
alces-263	175	8	72	72	NUM
alces-263	175	9	:	:	PUNCT
alces-263	175	10	878–884	878–884	NUM
alces-263	175	11	.	.	PUNCT
alces-263	176	1	doi	doi	NOUN
alces-263	176	2	:	:	PUNCT
alces-263	176	3	10.1139	10.1139	NUM
alces-263	176	4	/	/	SYM
alces-263	176	5	z94	z94	NUM
alces-263	176	6	-	-	PUNCT
alces-263	176	7	119	119	NUM
alces-263	176	8	hayes	haye	NOUN
alces-263	176	9	,	,	PUNCT
alces-263	176	10	r.	r.	PROPN
alces-263	176	11	d.	d.	PROPN
alces-263	176	12	,	,	PUNCT
alces-263	176	13	a.	a.	PROPN
alces-263	176	14	m.	m.	PROPN
alces-263	176	15	baer	baer	PROPN
alces-263	176	16	,	,	PUNCT
alces-263	176	17	u.	u.	PROPN
alces-263	176	18	wotschikowsky	wotschikowsky	PROPN
alces-263	176	19	,	,	PUNCT
alces-263	176	20	and	and	CCONJ
alces-263	176	21	a.	a.	PROPN
alces-263	176	22	s.	s.	PROPN
alces-263	176	23	harestad	harestad	PROPN
alces-263	176	24	.	.	PUNCT
alces-263	177	1	2000	2000	NUM
alces-263	177	2	.	.	PUNCT
alces-263	178	1	kill	kill	VERB
alces-263	178	2	rate	rate	NOUN
alces-263	178	3	by	by	ADP
alces-263	178	4	wolves	wolf	NOUN
alces-263	178	5	on	on	ADP
alces-263	178	6	moose	moose	NOUN
alces-263	178	7	in	in	ADP
alces-263	178	8	the	the	DET
alces-263	178	9	yukon	yukon	PROPN
alces-263	178	10	.	.	PUNCT
alces-263	179	1	canadian	canadian	PROPN
alces-263	179	2	journal	journal	PROPN
alces-263	179	3	of	of	ADP
alces-263	179	4	zoology	zoology	NOUN
alces-263	179	5	78	78	NUM
alces-263	179	6	:	:	PUNCT
alces-263	179	7	49–59	49–59	NOUN
alces-263	179	8	.	.	PUNCT
alces-263	180	1	doi	doi	NOUN
alces-263	180	2	:	:	PUNCT
alces-263	180	3	10.1139	10.1139	NUM
alces-263	180	4	/	/	SYM
alces-263	180	5	z99	z99	PROPN
alces-263	180	6	-	-	PUNCT
alces-263	180	7	187	187	NUM
alces-263	180	8	hebblewhite	hebblewhite	NOUN
alces-263	180	9	,	,	PUNCT
alces-263	180	10	m.	m.	NOUN
alces-263	180	11	,	,	PUNCT
alces-263	180	12	j.	j.	PROPN
alces-263	180	13	whittington	whittington	PROPN
alces-263	180	14	,	,	PUNCT
alces-263	180	15	m.	m.	PROPN
alces-263	180	16	bradley	bradley	PROPN
alces-263	180	17	,	,	PUNCT
alces-263	180	18	g.	g.	PROPN
alces-263	180	19	skinner	skinner	PROPN
alces-263	180	20	,	,	PUNCT
alces-263	180	21	a.	a.	NOUN
alces-263	180	22	dibb	dibb	PROPN
alces-263	180	23	,	,	PUNCT
alces-263	180	24	and	and	CCONJ
alces-263	180	25	c.	c.	PROPN
alces-263	180	26	a.	a.	PROPN
alces-263	180	27	white	white	PROPN
alces-263	180	28	.	.	PUNCT
alces-263	181	1	2007	2007	NUM
alces-263	181	2	.	.	PUNCT
alces-263	182	1	conditions	condition	NOUN
alces-263	182	2	for	for	ADP
alces-263	182	3	caribou	caribou	NOUN
alces-263	182	4	persistence	persistence	NOUN
alces-263	182	5	in	in	ADP
alces-263	182	6	the	the	DET
alces-263	182	7	wolf	wolf	PROPN
alces-263	182	8	-	-	PUNCT
alces-263	182	9	elk	elk	NOUN
alces-263	182	10	caribou	caribou	NOUN
alces-263	182	11	systems	system	NOUN
alces-263	182	12	of	of	ADP
alces-263	182	13	the	the	DET
alces-263	182	14	canadian	canadian	ADJ
alces-263	182	15	rockies	rockie	NOUN
alces-263	182	16	.	.	PUNCT
alces-263	183	1	rangifer	rangifer	VERB
alces-263	183	2	17(special	17(special	NUM
alces-263	183	3	issue	issue	NOUN
alces-263	183	4	):	):	PUNCT
alces-263	183	5	79–91	79–91	NUM
alces-263	183	6	.	.	PUNCT
alces-263	184	1	doi	doi	NOUN
alces-263	184	2	:	:	PUNCT
alces-263	184	3	10.7557/2.27.4.322	10.7557/2.27.4.322	NUM
alces-263	184	4	hervieux	hervieux	PROPN
alces-263	184	5	,	,	PUNCT
alces-263	184	6	d.	d.	PROPN
alces-263	184	7	,	,	PUNCT
alces-263	184	8	m.	m.	NOUN
alces-263	184	9	hebblewhite	hebblewhite	PROPN
alces-263	184	10	,	,	PUNCT
alces-263	184	11	n.	n.	PROPN
alces-263	184	12	j.	j.	PROPN
alces-263	184	13	decesare	decesare	PROPN
alces-263	184	14	,	,	PUNCT
alces-263	184	15	m.	m.	PROPN
alces-263	184	16	russell	russell	PROPN
alces-263	184	17	,	,	PUNCT
alces-263	184	18	k.	k.	PROPN
alces-263	184	19	smith	smith	PROPN
alces-263	184	20	,	,	PUNCT
alces-263	184	21	s.	s.	PROPN
alces-263	184	22	robertson	robertson	PROPN
alces-263	184	23	,	,	PUNCT
alces-263	184	24	and	and	CCONJ
alces-263	184	25	s.	s.	PROPN
alces-263	184	26	boutin	boutin	PROPN
alces-263	184	27	.	.	PUNCT
alces-263	185	1	2013	2013	NUM
alces-263	185	2	.	.	PUNCT
alces-263	186	1	widespread	widespread	ADJ
alces-263	186	2	declines	decline	NOUN
alces-263	186	3	in	in	ADP
alces-263	186	4	woodland	woodland	PROPN
alces-263	186	5	caribou	caribou	PROPN
alces-263	186	6	(	(	PUNCT
alces-263	186	7	rangifer	rangifer	VERB
alces-263	186	8	tarandus	tarandus	PROPN
alces-263	186	9	caribou	caribou	PROPN
alces-263	186	10	)	)	PUNCT
alces-263	186	11	continue	continue	VERB
alces-263	186	12	in	in	ADP
alces-263	186	13	alberta	alberta	PROPN
alces-263	186	14	.	.	PUNCT
alces-263	187	1	canadian	canadian	PROPN
alces-263	187	2	journal	journal	PROPN
alces-263	187	3	of	of	ADP
alces-263	187	4	zoology	zoology	NOUN
alces-263	187	5	91	91	NUM
alces-263	187	6	:	:	PUNCT
alces-263	187	7	872–882	872–882	NUM
alces-263	187	8	.	.	PUNCT
alces-263	188	1	doi	doi	NOUN
alces-263	188	2	:	:	PUNCT
alces-263	188	3	10.1139/	10.1139/	NUM
alces-263	188	4	cjz-2013	cjz-2013	NOUN
alces-263	188	5	-	-	PUNCT
alces-263	188	6	0123	0123	NUM
alces-263	188	7	_	_	PUNCT
alces-263	189	1	_	_	PUNCT
alces-263	190	1	_	_	PUNCT
alces-263	191	1	_	_	PUNCT
alces-263	192	1	_	_	PUNCT
alces-263	192	2	,	,	PUNCT
alces-263	193	1	_	_	PUNCT
alces-263	194	1	_	_	PUNCT
alces-263	195	1	_	_	PUNCT
alces-263	196	1	_	_	PUNCT
alces-263	197	1	_	_	PUNCT
alces-263	197	2	,	,	PUNCT
alces-263	197	3	d.	d.	PROPN
alces-263	197	4	stepnisky	stepnisky	PROPN
alces-263	197	5	,	,	PUNCT
alces-263	197	6	m.	m.	NOUN
alces-263	197	7	bacon	bacon	NOUN
alces-263	197	8	,	,	PUNCT
alces-263	197	9	and	and	CCONJ
alces-263	197	10	s.	s.	PROPN
alces-263	197	11	boutin	boutin	PROPN
alces-263	197	12	.	.	PUNCT
alces-263	198	1	2014	2014	NUM
alces-263	198	2	.	.	PUNCT
alces-263	199	1	managing	manage	VERB
alces-263	199	2	wolves	wolf	NOUN
alces-263	199	3	(	(	PUNCT
alces-263	199	4	canis	canis	NOUN
alces-263	199	5	lupus	lupus	NOUN
alces-263	199	6	)	)	PUNCT
alces-263	199	7	to	to	PART
alces-263	199	8	recover	recover	VERB
alces-263	199	9	threatened	threaten	VERB
alces-263	199	10	woodland	woodland	NOUN
alces-263	199	11	caribou	caribou	PROPN
alces-263	199	12	(	(	PUNCT
alces-263	199	13	rangifer	rangifer	VERB
alces-263	199	14	tarandus	tarandus	PROPN
alces-263	199	15	caribou	caribou	PROPN
alces-263	199	16	)	)	PUNCT
alces-263	199	17	in	in	ADP
alces-263	199	18	alberta	alberta	PROPN
alces-263	199	19	.	.	PUNCT
alces-263	200	1	canadian	canadian	PROPN
alces-263	200	2	journal	journal	PROPN
alces-263	200	3	of	of	ADP
alces-263	200	4	zoology	zoology	NOUN
alces-263	200	5	92	92	NUM
alces-263	200	6	:	:	PUNCT
alces-263	200	7	1029–1037	1029–1037	NUM
alces-263	200	8	.	.	PUNCT
alces-263	201	1	doi	doi	NOUN
alces-263	201	2	:	:	PUNCT
alces-263	201	3	10.1139	10.1139	NUM
alces-263	201	4	/	/	SYM
alces-263	201	5	cjz-2014	cjz-2014	NOUN
alces-263	201	6	-	-	PUNCT
alces-263	201	7	0142	0142	NUM
alces-263	201	8	kunkel	kunkel	PROPN
alces-263	201	9	,	,	PUNCT
alces-263	201	10	k.	k.	PROPN
alces-263	201	11	,	,	PUNCT
alces-263	201	12	and	and	CCONJ
alces-263	201	13	d.	d.	PROPN
alces-263	201	14	h.	h.	PROPN
alces-263	201	15	pletscher	pletscher	PROPN
alces-263	201	16	.	.	PUNCT
alces-263	202	1	1999	1999	NUM
alces-263	202	2	.	.	PUNCT
alces-263	203	1	species	species	NOUN
alces-263	203	2	-	-	PUNCT
alces-263	203	3	specific	specific	ADJ
alces-263	203	4	population	population	NOUN
alces-263	203	5	dynamics	dynamic	NOUN
alces-263	203	6	of	of	ADP
alces-263	203	7	cervids	cervid	NOUN
alces-263	203	8	in	in	ADP
alces-263	203	9	a	a	DET
alces-263	203	10	multi	multi	ADJ
alces-263	203	11	-	-	ADJ
alces-263	203	12	predator	predator	ADJ
alces-263	203	13	ecosystem	ecosystem	NOUN
alces-263	203	14	.	.	PUNCT
alces-263	204	1	journal	journal	PROPN
alces-263	204	2	of	of	ADP
alces-263	204	3	wildlife	wildlife	NOUN
alces-263	204	4	management	management	NOUN
alces-263	204	5	63	63	NUM
alces-263	204	6	:	:	PUNCT
alces-263	204	7	1082–1093	1082–1093	NUM
alces-263	204	8	.	.	PUNCT
alces-263	205	1	doi	doi	NOUN
alces-263	205	2	:	:	PUNCT
alces-263	205	3	10.2307/3802827	10.2307/3802827	NUM
alces-263	205	4	kuzyk	kuzyk	INTJ
alces-263	205	5	,	,	PUNCT
alces-263	205	6	g	g	PROPN
alces-263	205	7	,	,	PUNCT
alces-263	205	8	i.	i.	PROPN
alces-263	205	9	hatter	hatter	PROPN
alces-263	205	10	,	,	PUNCT
alces-263	205	11	s.	s.	PROPN
alces-263	205	12	marshall	marshall	PROPN
alces-263	205	13	,	,	PUNCT
alces-263	205	14	c.	c.	PROPN
alces-263	205	15	procter	procter	PROPN
alces-263	205	16	,	,	PUNCT
alces-263	205	17	b.	b.	PROPN
alces-263	205	18	cadsand	cadsand	PROPN
alces-263	205	19	,	,	PUNCT
alces-263	205	20	d.	d.	PROPN
alces-263	205	21	lirette	lirette	PROPN
alces-263	205	22	,	,	PUNCT
alces-263	205	23	h.	h.	PROPN
alces-263	205	24	schindler	schindler	PROPN
alces-263	205	25	,	,	PUNCT
alces-263	205	26	m.	m.	PROPN
alces-263	205	27	bridger	bridger	PROPN
alces-263	205	28	,	,	PUNCT
alces-263	205	29	p.	p.	PROPN
alces-263	205	30	stent	stent	NOUN
alces-263	205	31	,	,	PUNCT
alces-263	205	32	a.	a.	NOUN
alces-263	205	33	walker	walker	PROPN
alces-263	205	34	,	,	PUNCT
alces-263	205	35	and	and	CCONJ
alces-263	205	36	m.	m.	NOUN
alces-263	205	37	klaczek	klaczek	PROPN
alces-263	205	38	.	.	PROPN
alces-263	206	1	2018	2018	NUM
alces-263	206	2	.	.	PUNCT
alces-263	207	1	moose	moose	NOUN
alces-263	207	2	population	population	NOUN
alces-263	207	3	dynamics	dynamic	NOUN
alces-263	207	4	during	during	ADP
alces-263	207	5	20	20	NUM
alces-263	207	6	years	year	NOUN
alces-263	207	7	of	of	ADP
alces-263	207	8	declining	decline	VERB
alces-263	207	9	harvest	harvest	NOUN
alces-263	207	10	in	in	ADP
alces-263	207	11	british	british	PROPN
alces-263	207	12	columbia	columbia	PROPN
alces-263	207	13	.	.	PUNCT
alces-263	208	1	alces	alce	NOUN
alces-263	208	2	54	54	NUM
alces-263	208	3	:	:	PUNCT
alces-263	208	4	101–119	101–119	NUM
alces-263	208	5	.	.	PUNCT
alces-263	209	1	_	_	PUNCT
alces-263	210	1	_	_	PUNCT
alces-263	211	1	_	_	PUNCT
alces-263	212	1	_	_	PUNCT
alces-263	213	1	_	_	PUNCT
alces-263	213	2	,	,	PUNCT
alces-263	213	3	c.	c.	PROPN
alces-263	213	4	procter	procter	PROPN
alces-263	213	5	,	,	PUNCT
alces-263	213	6	s.	s.	PROPN
alces-263	213	7	marshall	marshall	PROPN
alces-263	213	8	,	,	PUNCT
alces-263	213	9	and	and	CCONJ
alces-263	213	10	d.	d.	PROPN
alces-263	213	11	hodder	hodder	PROPN
alces-263	213	12	.	.	PUNCT
alces-263	214	1	2019a	2019a	NUM
alces-263	214	2	.	.	PUNCT
alces-263	215	1	factors	factor	NOUN
alces-263	215	2	affecting	affect	VERB
alces-263	215	3	moose	moose	NOUN
alces-263	215	4	population	population	NOUN
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alces-263	215	6	in	in	ADP
alces-263	215	7	british	british	PROPN
alces-263	215	8	columbia	columbia	PROPN
alces-263	215	9	:	:	PUNCT
alces-263	215	10	updated	update	VERB
alces-263	215	11	research	research	NOUN
alces-263	215	12	design	design	NOUN
alces-263	215	13	.	.	PUNCT
alces-263	216	1	wildlife	wildlife	NOUN
alces-263	216	2	bulletin	bulletin	NOUN
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alces-263	216	4	.	.	PUNCT
alces-263	216	5	b-128	b-128	PROPN
alces-263	216	6	.	.	PUNCT
alces-263	217	1	british	british	PROPN
alces-263	217	2	columbia	columbia	PROPN
alces-263	217	3	ministry	ministry	PROPN
alces-263	217	4	of	of	ADP
alces-263	217	5	forests	forest	NOUN
alces-263	217	6	,	,	PUNCT
alces-263	217	7	lands	land	NOUN
alces-263	217	8	and	and	CCONJ
alces-263	217	9	natural	natural	ADJ
alces-263	217	10	research	research	NOUN
alces-263	217	11	operations	operation	NOUN
alces-263	217	12	and	and	CCONJ
alces-263	217	13	rural	rural	ADJ
alces-263	217	14	development	development	NOUN
alces-263	217	15	,	,	PUNCT
alces-263	217	16	victoria	victoria	PROPN
alces-263	217	17	,	,	PUNCT
alces-263	217	18	british	british	PROPN
alces-263	217	19	columbia	columbia	PROPN
alces-263	217	20	,	,	PUNCT
alces-263	217	21	canada	canada	PROPN
alces-263	217	22	.	.	PUNCT
alces-263	218	1	_	_	PUNCT
alces-263	219	1	_	_	PUNCT
alces-263	220	1	_	_	PUNCT
alces-263	221	1	_	_	PUNCT
alces-263	222	1	_	_	PUNCT
alces-263	222	2	,	,	PUNCT
alces-263	223	1	_	_	PUNCT
alces-263	224	1	_	_	PUNCT
alces-263	225	1	_	_	PUNCT
alces-263	226	1	_	_	PUNCT
alces-263	227	1	_	_	PUNCT
alces-263	227	2	,	,	PUNCT
alces-263	228	1	_	_	PUNCT
alces-263	229	1	_	_	PUNCT
alces-263	230	1	_	_	PUNCT
alces-263	231	1	_	_	PUNCT
alces-263	232	1	_	_	PUNCT
alces-263	232	2	,	,	PUNCT
alces-263	232	3	h.	h.	PROPN
alces-263	232	4	schindler	schindler	PROPN
alces-263	232	5	,	,	PUNCT
alces-263	232	6	h.	h.	PROPN
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alces-263	232	8	,	,	PUNCT
alces-263	232	9	m.	m.	NOUN
alces-263	232	10	scheideman	scheideman	NOUN
alces-263	232	11	,	,	PUNCT
alces-263	232	12	and	and	CCONJ
alces-263	232	13	d.	d.	PROPN
alces-263	232	14	hodder	hodder	PROPN
alces-263	232	15	.	.	PUNCT
alces-263	233	1	2019b	2019b	NOUN
alces-263	233	2	.	.	PUNCT
alces-263	234	1	determining	determine	VERB
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alces-263	234	3	affecting	affect	VERB
alces-263	234	4	moose	moose	NOUN
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alces-263	234	6	change	change	NOUN
alces-263	234	7	in	in	ADP
alces-263	234	8	british	british	PROPN
alces-263	234	9	columbia	columbia	PROPN
alces-263	234	10	:	:	PUNCT
alces-263	234	11	testing	test	VERB
alces-263	234	12	the	the	DET
alces-263	234	13	landscape	landscape	NOUN
alces-263	234	14	change	change	NOUN
alces-263	234	15	hypothesis	hypothesis	NOUN
alces-263	234	16	.	.	PUNCT
alces-263	235	1	wildlife	wildlife	NOUN
alces-263	235	2	working	working	NOUN
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alces-263	235	4	.	.	PUNCT
alces-263	236	1	no	no	INTJ
alces-263	236	2	.	.	PUNCT
alces-263	237	1	wr-127	wr-127	NOUN
alces-263	237	2	.	.	PUNCT
alces-263	238	1	progress	progress	PROPN
alces-263	238	2	report	report	NOUN
alces-263	238	3	:	:	PUNCT
alces-263	238	4	february	february	PROPN
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alces-263	238	6	–	–	PUNCT
alces-263	238	7	april	april	PROPN
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alces-263	238	9	.	.	PUNCT
alces-263	239	1	british	british	PROPN
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alces-263	239	4	of	of	ADP
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alces-263	239	6	,	,	PUNCT
alces-263	239	7	lands	land	NOUN
alces-263	239	8	and	and	CCONJ
alces-263	239	9	natural	natural	ADJ
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alces-263	239	12	and	and	CCONJ
alces-263	239	13	rural	rural	ADJ
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alces-263	239	15	,	,	PUNCT
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alces-263	239	18	british	british	PROPN
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alces-263	239	20	,	,	PUNCT
alces-263	239	21	canada	canada	PROPN
alces-263	239	22	.	.	PUNCT
alces-263	240	1	latham	latham	PROPN
alces-263	240	2	,	,	PUNCT
alces-263	240	3	a.	a.	PROPN
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alces-263	240	5	,	,	PUNCT
alces-263	240	6	m.	m.	PROPN
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alces-263	240	9	,	,	PUNCT
alces-263	240	10	n.	n.	PROPN
alces-263	240	11	a.	a.	PROPN
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alces-263	240	13	and	and	CCONJ
alces-263	240	14	s.	s.	PROPN
alces-263	240	15	boutin	boutin	PROPN
alces-263	240	16	.	.	PUNCT
alces-263	241	1	2011	2011	NUM
alces-263	241	2	.	.	PUNCT
alces-263	242	1	invading	invade	VERB
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alces-263	242	3	-	-	PUNCT
alces-263	242	4	tailed	tail	VERB
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alces-263	242	7	wolf	wolf	PROPN
alces-263	242	8	-	-	PUNCT
alces-263	242	9	caribou	caribou	NOUN
alces-263	242	10	dynamics	dynamic	NOUN
alces-263	242	11	in	in	ADP
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alces-263	242	13	alberta	alberta	PROPN
alces-263	242	14	.	.	PUNCT
alces-263	243	1	journal	journal	PROPN
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alces-263	243	4	management	management	NOUN
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alces-263	243	6	:	:	PUNCT
alces-263	244	1	204–212	204–212	NUM
alces-263	244	2	.	.	PUNCT
alces-263	244	3	doi	doi	NOUN
alces-263	244	4	:	:	PUNCT
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alces-263	244	9	,	,	PUNCT
alces-263	244	10	p.	p.	PROPN
alces-263	244	11	d.	d.	PROPN
alces-263	244	12	,	,	PUNCT
alces-263	244	13	e.	e.	PROPN
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alces-263	244	15	,	,	PUNCT
alces-263	244	16	b.	b.	PROPN
alces-263	244	17	wynes	wynes	PROPN
alces-263	244	18	,	,	PUNCT
alces-263	244	19	and	and	CCONJ
alces-263	244	20	s.	s.	PROPN
alces-263	244	21	boutin	boutin	PROPN
alces-263	244	22	.	.	PUNCT
alces-263	245	1	2003	2003	NUM
alces-263	245	2	.	.	PUNCT
alces-263	246	1	declines	decline	NOUN
alces-263	246	2	in	in	ADP
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alces-263	246	4	of	of	ADP
alces-263	246	5	woodland	woodland	PROPN
alces-263	246	6	caribou	caribou	PROPN
alces-263	246	7	.	.	PUNCT
alces-263	247	1	journal	journal	PROPN
alces-263	247	2	of	of	ADP
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alces-263	247	4	management	management	NOUN
alces-263	247	5	67	67	NUM
alces-263	247	6	:	:	PUNCT
alces-263	247	7	755–761	755–761	NUM
alces-263	247	8	.	.	PUNCT
alces-263	248	1	doi	doi	NOUN
alces-263	248	2	:	:	PUNCT
alces-263	248	3	10.2307/3802682	10.2307/3802682	PROPN
alces-263	248	4	patterson	patterson	PROPN
alces-263	248	5	,	,	PUNCT
alces-263	248	6	b.	b.	PROPN
alces-263	248	7	r.	r.	PROPN
alces-263	248	8	,	,	PUNCT
alces-263	248	9	b.	b.	PROPN
alces-263	248	10	a.	a.	PROPN
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alces-263	248	12	,	,	PUNCT
alces-263	248	13	b.	b.	PROPN
alces-263	248	14	a.	a.	PROPN
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alces-263	248	16	,	,	PUNCT
alces-263	248	17	d.	d.	PROPN
alces-263	248	18	g.	g.	PROPN
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alces-263	248	20	,	,	PUNCT
alces-263	248	21	and	and	CCONJ
alces-263	248	22	l.	l.	PROPN
alces-263	248	23	k.	k.	PROPN
alces-263	248	24	benjamin	benjamin	PROPN
alces-263	248	25	.	.	PROPN
alces-263	248	26	2002	2002	NUM
alces-263	248	27	.	.	PUNCT
alces-263	249	1	proximate	proximate	NOUN
alces-263	249	2	factors	factor	NOUN
alces-263	249	3	limiting	limit	VERB
alces-263	249	4	population	population	NOUN
alces-263	249	5	growth	growth	NOUN
alces-263	249	6	of	of	ADP
alces-263	249	7	white	white	ADJ
alces-263	249	8	-	-	PUNCT
alces-263	249	9	tailed	tail	VERB
alces-263	249	10	deer	deer	NOUN
alces-263	249	11	in	in	ADP
alces-263	249	12	nova	nova	PROPN
alces-263	249	13	scotia	scotia	PROPN
alces-263	249	14	.	.	PUNCT
alces-263	250	1	journal	journal	PROPN
alces-263	250	2	of	of	ADP
alces-263	250	3	wildlife	wildlife	NOUN
alces-263	250	4	management	management	NOUN
alces-263	250	5	66	66	NUM
alces-263	250	6	:	:	PUNCT
alces-263	250	7	511–521	511–521	NUM
alces-263	250	8	.	.	PUNCT
alces-263	251	1	doi	doi	NOUN
alces-263	251	2	:	:	PUNCT
alces-263	251	3	10.2307/	10.2307/	NUM
alces-263	251	4	3803184	3803184	NUM
alces-263	251	5	pollock	pollock	NOUN
alces-263	251	6	,	,	PUNCT
alces-263	251	7	k.	k.	PROPN
alces-263	251	8	h.	h.	PROPN
alces-263	251	9	,	,	PUNCT
alces-263	251	10	s.	s.	PROPN
alces-263	251	11	r.	r.	PROPN
alces-263	251	12	winterstein	winterstein	PROPN
alces-263	251	13	,	,	PUNCT
alces-263	251	14	c.	c.	PROPN
alces-263	251	15	m.	m.	NOUN
alces-263	251	16	bunck	bunck	PROPN
alces-263	251	17	,	,	PUNCT
alces-263	251	18	and	and	CCONJ
alces-263	251	19	p.	p.	PROPN
alces-263	251	20	d.	d.	PROPN
alces-263	251	21	curtis	curtis	PROPN
alces-263	251	22	.	.	PROPN
alces-263	251	23	1989	1989	NUM
alces-263	251	24	.	.	PUNCT
alces-263	252	1	survival	survival	NOUN
alces-263	252	2	analysis	analysis	NOUN
alces-263	252	3	in	in	ADP
alces-263	252	4	telemetry	telemetry	NOUN
alces-263	252	5	studies	study	NOUN
alces-263	252	6	:	:	PUNCT
alces-263	252	7	the	the	DET
alces-263	252	8	staggered	staggered	ADJ
alces-263	252	9	entry	entry	NOUN
alces-263	252	10	design	design	NOUN
alces-263	252	11	.	.	PUNCT
alces-263	253	1	journal	journal	NOUN
alces-263	253	2	of	of	ADP
alces-263	253	3	wildlife	wildlife	NOUN
alces-263	253	4	management	management	NOUN
alces-263	253	5	53	53	NUM
alces-263	254	1	:	:	PUNCT
alces-263	254	2	7–15	7–15	VERB
alces-263	254	3	.	.	PUNCT
alces-263	255	1	doi	doi	NOUN
alces-263	255	2	:	:	PUNCT
alces-263	255	3	10.2307/	10.2307/	NUM
alces-263	255	4	3801296	3801296	NUM
alces-263	255	5	seip	seip	NOUN
alces-263	255	6	,	,	PUNCT
alces-263	255	7	d.	d.	PROPN
alces-263	255	8	r.	r.	PROPN
alces-263	255	9	,	,	PUNCT
alces-263	255	10	and	and	CCONJ
alces-263	255	11	d.	d.	PROPN
alces-263	255	12	b.	b.	PROPN
alces-263	255	13	cichowski	cichowski	PROPN
alces-263	255	14	.	.	PUNCT
alces-263	256	1	1996	1996	NUM
alces-263	256	2	.	.	PUNCT
alces-263	257	1	population	population	NOUN
alces-263	257	2	ecology	ecology	NOUN
alces-263	257	3	of	of	ADP
alces-263	257	4	caribou	caribou	PROPN
alces-263	257	5	in	in	ADP
alces-263	257	6	british	british	PROPN
alces-263	257	7	columbia	columbia	PROPN
alces-263	257	8	.	.	PUNCT
alces-263	258	1	rangifer	rangifer	VERB
alces-263	258	2	9	9	NUM
alces-263	258	3	:	:	PUNCT
alces-263	258	4	73–80	73–80	NUM
alces-263	258	5	.	.	PUNCT
alces-263	259	1	doi	doi	NOUN
alces-263	259	2	:	:	PUNCT
alces-263	259	3	10.7557/2.16.4.1223	10.7557/2.16.4.1223	PROPN
alces-263	259	4	serrouya	serrouya	PROPN
alces-263	259	5	,	,	PUNCT
alces-263	259	6	r.	r.	PROPN
alces-263	259	7	,	,	PUNCT
alces-263	259	8	s.	s.	PROPN
alces-263	259	9	gilbert	gilbert	PROPN
alces-263	259	10	,	,	PUNCT
alces-263	259	11	r.	r.	PROPN
alces-263	259	12	s.	s.	PROPN
alces-263	259	13	mcnay	mcnay	PROPN
alces-263	259	14	,	,	PUNCT
alces-263	259	15	b.	b.	PROPN
alces-263	259	16	n.	n.	PROPN
alces-263	259	17	mclellan	mclellan	PROPN
alces-263	259	18	,	,	PUNCT
alces-263	259	19	d.	d.	PROPN
alces-263	259	20	c.	c.	PROPN
alces-263	259	21	heard	hear	VERB
alces-263	259	22	,	,	PUNCT
alces-263	259	23	d.	d.	PROPN
alces-263	259	24	r.	r.	PROPN
alces-263	259	25	seip	seip	PROPN
alces-263	259	26	,	,	PUNCT
alces-263	259	27	and	and	CCONJ
alces-263	259	28	s.	s.	PROPN
alces-263	259	29	boutin	boutin	PROPN
alces-263	259	30	.	.	PUNCT
alces-263	260	1	2017	2017	NUM
alces-263	260	2	.	.	PUNCT
alces-263	261	1	comparing	compare	VERB
alces-263	261	2	population	population	NOUN
alces-263	261	3	growth	growth	NOUN
alces-263	261	4	rates	rate	NOUN
alces-263	261	5	between	between	ADP
alces-263	261	6	census	census	NOUN
alces-263	261	7	and	and	CCONJ
alces-263	261	8	recruitment	recruitment	NOUN
alces-263	261	9	-	-	PUNCT
alces-263	261	10	mortality	mortality	NOUN
alces-263	261	11	models	model	NOUN
alces-263	261	12	.	.	PUNCT
alces-263	262	1	journal	journal	NOUN
alces-263	262	2	of	of	ADP
alces-263	262	3	wildlife	wildlife	NOUN
alces-263	262	4	management	management	NOUN
alces-263	262	5	81	81	NUM
alces-263	262	6	:	:	PUNCT
alces-263	262	7	297–305	297–305	NUM
alces-263	262	8	.	.	PUNCT
alces-263	263	1	doi	doi	NOUN
alces-263	263	2	:	:	PUNCT
alces-263	263	3	10.1002	10.1002	NUM
alces-263	263	4	/	/	SYM
alces-263	263	5	jwmg.21185	jwmg.21185	PROPN
alces-263	263	6	alces	alce	NOUN
alces-263	263	7	vol	vol	NOUN
alces-263	263	8	.	.	PROPN
alces-263	264	1	56	56	NUM
alces-263	264	2	,	,	PUNCT
alces-263	264	3	2020	2020	NUM
alces-263	264	4	revisiting	revisit	VERB
alces-263	264	5	the	the	DET
alces-263	264	6	recruitment	recruitment	NOUN
alces-263	264	7	-	-	PUNCT
alces-263	264	8	mortality	mortality	NOUN
alces-263	264	9	.	.	PUNCT
alces-263	265	1	–	–	PUNCT
alces-263	265	2	ian	ian	PROPN
alces-263	265	3	w.	w.	PROPN
alces-263	265	4	hatter	hatter	PROPN
alces-263	265	5	47	47	NUM
alces-263	265	6	severud	severud	NOUN
alces-263	265	7	,	,	PUNCT
alces-263	265	8	w.	w.	PROPN
alces-263	265	9	j.	j.	PROPN
alces-263	265	10	,	,	PUNCT
alces-263	265	11	g.	g.	PROPN
alces-263	265	12	d.	d.	PROPN
alces-263	265	13	delgiudice	delgiudice	PROPN
alces-263	265	14	,	,	PUNCT
alces-263	265	15	and	and	CCONJ
alces-263	265	16	j.	j.	PROPN
alces-263	265	17	k.	k.	PROPN
alces-263	265	18	bump	bump	PROPN
alces-263	265	19	.	.	PUNCT
alces-263	266	1	2019	2019	NUM
alces-263	266	2	.	.	PUNCT
alces-263	267	1	comparing	compare	VERB
alces-263	267	2	survey	survey	NOUN
alces-263	267	3	and	and	CCONJ
alces-263	267	4	multiple	multiple	ADJ
alces-263	267	5	recruitment	recruitment	NOUN
alces-263	267	6	-	-	PUNCT
alces-263	267	7	mortality	mortality	NOUN
alces-263	267	8	models	model	NOUN
alces-263	267	9	to	to	PART
alces-263	267	10	assess	assess	VERB
alces-263	267	11	growth	growth	NOUN
alces-263	267	12	rates	rate	NOUN
alces-263	267	13	and	and	CCONJ
alces-263	267	14	population	population	NOUN
alces-263	267	15	projections	projection	NOUN
alces-263	267	16	.	.	PUNCT
alces-263	268	1	ecology	ecology	NOUN
alces-263	268	2	and	and	CCONJ
alces-263	268	3	evolution	evolution	NOUN
alces-263	268	4	9	9	NUM
alces-263	268	5	:	:	SYM
alces-263	268	6	12613–12622	12613–12622	NUM
alces-263	268	7	.	.	PUNCT
alces-263	269	1	doi	doi	NOUN
alces-263	269	2	:	:	PUNCT
alces-263	269	3	10.1002	10.1002	NUM
alces-263	269	4	/	/	SYM
alces-263	269	5	ece3.5725	ece3.5725	ADJ
alces-263	269	6	sinclair	sinclair	NOUN
alces-263	269	7	,	,	PUNCT
alces-263	269	8	a.	a.	PROPN
alces-263	269	9	r.	r.	PROPN
alces-263	269	10	e.	e.	PROPN
alces-263	269	11	,	,	PUNCT
alces-263	269	12	j.	j.	PROPN
alces-263	269	13	m.	m.	PROPN
alces-263	269	14	fryxell	fryxell	PROPN
alces-263	269	15	,	,	PUNCT
alces-263	269	16	and	and	CCONJ
alces-263	269	17	g.	g.	PROPN
alces-263	269	18	caughley	caughley	PROPN
alces-263	269	19	.	.	PUNCT
alces-263	270	1	2006	2006	NUM
alces-263	270	2	.	.	PUNCT
alces-263	271	1	wildlife	wildlife	NOUN
alces-263	271	2	ecology	ecology	NOUN
alces-263	271	3	,	,	PUNCT
alces-263	271	4	conservation	conservation	NOUN
alces-263	271	5	,	,	PUNCT
alces-263	271	6	and	and	CCONJ
alces-263	271	7	management	management	NOUN
alces-263	271	8	.	.	PUNCT
alces-263	272	1	second	second	ADJ
alces-263	272	2	edition	edition	PROPN
alces-263	272	3	.	.	PUNCT
alces-263	273	1	blackwell	blackwell	PROPN
alces-263	273	2	publishing	publishing	PROPN
alces-263	273	3	,	,	PUNCT
alces-263	273	4	hoboken	hoboken	PROPN
alces-263	273	5	,	,	PUNCT
alces-263	273	6	new	new	PROPN
alces-263	273	7	jersey	jersey	PROPN
alces-263	273	8	,	,	PUNCT
alces-263	273	9	usa	usa	PROPN
alces-263	273	10	.	.	PROPN
alces-263	273	11	skalski	skalski	PROPN
alces-263	273	12	,	,	PUNCT
alces-263	273	13	j.	j.	PROPN
alces-263	273	14	r.	r.	PROPN
alces-263	273	15	,	,	PUNCT
alces-263	273	16	k.	k.	PROPN
alces-263	273	17	e.	e.	PROPN
alces-263	273	18	ryding	ryding	PROPN
alces-263	273	19	,	,	PUNCT
alces-263	273	20	and	and	CCONJ
alces-263	273	21	j.	j.	PROPN
alces-263	273	22	j.	j.	PROPN
alces-263	273	23	millspaugh	millspaugh	PROPN
alces-263	273	24	.	.	PUNCT
alces-263	274	1	2005	2005	NUM
alces-263	274	2	.	.	PUNCT
alces-263	275	1	wildlife	wildlife	NOUN
alces-263	275	2	demography	demography	NOUN
alces-263	275	3	:	:	PUNCT
alces-263	275	4	analysis	analysis	NOUN
alces-263	275	5	of	of	ADP
alces-263	275	6	sex	sex	NOUN
alces-263	275	7	,	,	PUNCT
alces-263	275	8	age	age	NOUN
alces-263	275	9	,	,	PUNCT
alces-263	275	10	and	and	CCONJ
alces-263	275	11	count	count	NOUN
alces-263	275	12	data	datum	NOUN
alces-263	275	13	.	.	PUNCT
alces-263	276	1	elsevier	elsevi	ADJ
alces-263	276	2	academic	academic	PROPN
alces-263	276	3	press	press	PROPN
alces-263	276	4	,	,	PUNCT
alces-263	276	5	new	new	PROPN
alces-263	276	6	york	york	PROPN
alces-263	276	7	,	,	PUNCT
alces-263	276	8	new	new	PROPN
alces-263	276	9	york	york	PROPN
alces-263	276	10	,	,	PUNCT
alces-263	276	11	usa	usa	PROPN
alces-263	276	12	.	.	PROPN
alces-263	276	13	sorensen	sorensen	PROPN
alces-263	276	14	,	,	PUNCT
alces-263	276	15	t.	t.	PROPN
alces-263	276	16	,	,	PUNCT
alces-263	276	17	p.	p.	PROPN
alces-263	276	18	d.	d.	PROPN
alces-263	276	19	mcloughlin	mcloughlin	PROPN
alces-263	276	20	,	,	PUNCT
alces-263	276	21	d.	d.	PROPN
alces-263	276	22	hervieux	hervieux	PROPN
alces-263	276	23	,	,	PUNCT
alces-263	276	24	e.	e.	PROPN
alces-263	276	25	dzus	dzus	PROPN
alces-263	276	26	,	,	PUNCT
alces-263	276	27	j.	j.	PROPN
alces-263	276	28	nolan	nolan	PROPN
alces-263	276	29	,	,	PUNCT
alces-263	276	30	b.	b.	PROPN
alces-263	276	31	wynes	wynes	PROPN
alces-263	276	32	,	,	PUNCT
alces-263	276	33	and	and	CCONJ
alces-263	276	34	s.	s.	PROPN
alces-263	276	35	boutin	boutin	PROPN
alces-263	276	36	.	.	PUNCT
alces-263	277	1	2008	2008	NUM
alces-263	277	2	.	.	PUNCT
alces-263	278	1	determining	determine	VERB
alces-263	278	2	sustainable	sustainable	ADJ
alces-263	278	3	levels	level	NOUN
alces-263	278	4	of	of	ADP
alces-263	278	5	cumulative	cumulative	ADJ
alces-263	278	6	effects	effect	NOUN
alces-263	278	7	for	for	ADP
alces-263	278	8	boreal	boreal	ADJ
alces-263	278	9	caribou	caribou	NOUN
alces-263	278	10	.	.	PUNCT
alces-263	279	1	journal	journal	PROPN
alces-263	279	2	of	of	ADP
alces-263	279	3	wildlife	wildlife	NOUN
alces-263	279	4	management	management	NOUN
alces-263	279	5	72	72	NUM
alces-263	279	6	:	:	PUNCT
alces-263	280	1	900–905	900–905	NUM
alces-263	280	2	.	.	PUNCT
alces-263	281	1	doi	doi	NOUN
alces-263	281	2	:	:	PUNCT
alces-263	281	3	10.2193/2007	10.2193/2007	NUM
alces-263	281	4	-	-	SYM
alces-263	281	5	079	079	NUM
alces-263	281	6	timmermann	timmermann	PROPN
alces-263	281	7	,	,	PUNCT
alces-263	281	8	h.	h.	PROPN
alces-263	281	9	r.	r.	PROPN
alces-263	281	10	,	,	PUNCT
alces-263	281	11	and	and	CCONJ
alces-263	281	12	a.	a.	PROPN
alces-263	281	13	r.	r.	PROPN
alces-263	281	14	rodgers	rodgers	PROPN
alces-263	281	15	.	.	PROPN
alces-263	282	1	2017	2017	NUM
alces-263	282	2	.	.	PUNCT
alces-263	283	1	the	the	DET
alces-263	283	2	status	status	NOUN
alces-263	283	3	and	and	CCONJ
alces-263	283	4	management	management	NOUN
alces-263	283	5	of	of	ADP
alces-263	283	6	moose	moose	NOUN
alces-263	283	7	in	in	ADP
alces-263	283	8	north	north	PROPN
alces-263	283	9	america	america	PROPN
alces-263	283	10	–	–	PUNCT
alces-263	283	11	circa	circa	NOUN
alces-263	283	12	2015	2015	NUM
alces-263	283	13	.	.	PUNCT
alces-263	284	1	alces	alce	NOUN
alces-263	284	2	53	53	NUM
alces-263	284	3	:	:	PUNCT
alces-263	284	4	1–22	1–22	NOUN
alces-263	284	5	.	.	PUNCT
alces-263	285	1	van	van	PROPN
alces-263	285	2	ballenberghe	ballenberghe	NOUN
alces-263	285	3	,	,	PUNCT
alces-263	285	4	v.	v.	ADP
alces-263	285	5	1983	1983	NUM
alces-263	285	6	.	.	PUNCT
alces-263	286	1	rate	rate	NOUN
alces-263	286	2	of	of	ADP
alces-263	286	3	increase	increase	NOUN
alces-263	286	4	in	in	ADP
alces-263	286	5	moose	moose	NOUN
alces-263	286	6	populations	population	NOUN
alces-263	286	7	.	.	PUNCT
alces-263	287	1	alces	alce	NOUN
alces-263	287	2	19	19	NUM
alces-263	287	3	:	:	PUNCT
alces-263	287	4	98–117	98–117	NUM
alces-263	287	5	.	.	PUNCT