id	sid	tid	token	lemma	pos
alces-297	1	1	113	113	NUM
alces-297	1	2	estimating	estimate	VERB
alces-297	1	3	sustained	sustained	ADJ
alces-297	1	4	yields	yield	NOUN
alces-297	1	5	for	for	ADP
alces-297	1	6	moose	moose	NOUN
alces-297	1	7	in	in	ADP
alces-297	1	8	central	central	ADJ
alces-297	1	9	british	british	PROPN
alces-297	1	10	columbia	columbia	PROPN
alces-297	1	11	using	use	VERB
alces-297	1	12	a	a	DET
alces-297	1	13	predator	predator	NOUN
alces-297	1	14	-	-	PUNCT
alces-297	1	15	prey	prey	NOUN
alces-297	1	16	model	model	NOUN
alces-297	1	17	ian	ian	PROPN
alces-297	1	18	w.	w.	PROPN
alces-297	1	19	hatter	hatter	PROPN
alces-297	1	20	nature	nature	PROPN
alces-297	1	21	wise	wise	ADJ
alces-297	1	22	consulting	consulting	NOUN
alces-297	1	23	,	,	PUNCT
alces-297	1	24	49	49	NUM
alces-297	1	25	-	-	SYM
alces-297	1	26	640	640	NUM
alces-297	1	27	upper	upper	ADJ
alces-297	1	28	lakeview	lakeview	NOUN
alces-297	1	29	road	road	NOUN
alces-297	1	30	,	,	PUNCT
alces-297	1	31	invermere	invermere	PROPN
alces-297	1	32	,	,	PUNCT
alces-297	1	33	british	british	PROPN
alces-297	1	34	columbia	columbia	PROPN
alces-297	1	35	v0a	v0a	PROPN
alces-297	1	36	1k3	1k3	NUM
alces-297	1	37	,	,	PUNCT
alces-297	1	38	canada	canada	PROPN
alces-297	1	39	abstract	abstract	NOUN
alces-297	1	40	:	:	PUNCT
alces-297	1	41	one	one	NUM
alces-297	1	42	of	of	ADP
alces-297	1	43	the	the	DET
alces-297	1	44	fundamental	fundamental	ADJ
alces-297	1	45	principles	principle	NOUN
alces-297	1	46	of	of	ADP
alces-297	1	47	wildlife	wildlife	NOUN
alces-297	1	48	harvesting	harvesting	NOUN
alces-297	1	49	is	be	AUX
alces-297	1	50	that	that	SCONJ
alces-297	1	51	it	it	PRON
alces-297	1	52	must	must	AUX
alces-297	1	53	result	result	VERB
alces-297	1	54	in	in	ADP
alces-297	1	55	a	a	DET
alces-297	1	56	sustained	sustained	ADJ
alces-297	1	57	yield	yield	NOUN
alces-297	1	58	(	(	PUNCT
alces-297	1	59	sy	sy	NOUN
alces-297	1	60	)	)	PUNCT
alces-297	1	61	,	,	PUNCT
alces-297	1	62	a	a	DET
alces-297	1	63	harvest	harvest	NOUN
alces-297	1	64	that	that	PRON
alces-297	1	65	can	can	AUX
alces-297	1	66	be	be	AUX
alces-297	1	67	taken	take	VERB
alces-297	1	68	year	year	NOUN
alces-297	1	69	after	after	ADP
alces-297	1	70	year	year	NOUN
alces-297	1	71	without	without	ADP
alces-297	1	72	jeopardizing	jeopardize	VERB
alces-297	1	73	future	future	ADJ
alces-297	1	74	harvests	harvest	NOUN
alces-297	1	75	.	.	PUNCT
alces-297	2	1	predator	predator	NOUN
alces-297	2	2	-	-	PUNCT
alces-297	2	3	prey	prey	NOUN
alces-297	2	4	models	model	NOUN
alces-297	2	5	are	be	AUX
alces-297	2	6	rarely	rarely	ADV
alces-297	2	7	incorporated	incorporate	VERB
alces-297	2	8	into	into	ADP
alces-297	2	9	estimates	estimate	NOUN
alces-297	2	10	of	of	ADP
alces-297	2	11	sys	sys	NOUN
alces-297	2	12	for	for	ADP
alces-297	2	13	moose	moose	NOUN
alces-297	2	14	,	,	PUNCT
alces-297	2	15	despite	despite	SCONJ
alces-297	2	16	predation	predation	NOUN
alces-297	2	17	of	of	ADP
alces-297	2	18	moose	moose	NOUN
alces-297	2	19	by	by	ADP
alces-297	2	20	wolf	wolf	PROPN
alces-297	2	21	(	(	PUNCT
alces-297	2	22	canis	canis	NOUN
alces-297	2	23	lupus	lupus	NOUN
alces-297	2	24	)	)	PUNCT
alces-297	2	25	,	,	PUNCT
alces-297	2	26	grizzly	grizzly	NOUN
alces-297	2	27	bear	bear	PROPN
alces-297	2	28	(	(	PUNCT
alces-297	2	29	ursus	ursus	PROPN
alces-297	2	30	arctos	arctos	PROPN
alces-297	2	31	)	)	PUNCT
alces-297	2	32	,	,	PUNCT
alces-297	2	33	and	and	CCONJ
alces-297	2	34	black	black	ADJ
alces-297	2	35	bear	bear	NOUN
alces-297	2	36	(	(	PUNCT
alces-297	2	37	u.	u.	NOUN
alces-297	2	38	americanus	americanus	PROPN
alces-297	2	39	)	)	PUNCT
alces-297	2	40	throughout	throughout	ADP
alces-297	2	41	much	much	ADJ
alces-297	2	42	of	of	ADP
alces-297	2	43	western	western	ADJ
alces-297	2	44	north	north	PROPN
alces-297	2	45	america	america	PROPN
alces-297	2	46	.	.	PUNCT
alces-297	3	1	a	a	DET
alces-297	3	2	simple	simple	ADJ
alces-297	3	3	predator	predator	NOUN
alces-297	3	4	-	-	PUNCT
alces-297	3	5	prey	prey	NOUN
alces-297	3	6	model	model	NOUN
alces-297	3	7	was	be	AUX
alces-297	3	8	parameterized	parameterized	ADJ
alces-297	3	9	from	from	ADP
alces-297	3	10	a	a	DET
alces-297	3	11	stable	stable	ADJ
alces-297	3	12	moose	moose	NOUN
alces-297	3	13	-	-	PUNCT
alces-297	3	14	wolf	wolf	NOUN
alces-297	3	15	-	-	PUNCT
alces-297	3	16	bear	bear	NOUN
alces-297	3	17	system	system	NOUN
alces-297	3	18	in	in	ADP
alces-297	3	19	central	central	ADJ
alces-297	3	20	british	british	PROPN
alces-297	3	21	columbia	columbia	PROPN
alces-297	3	22	during	during	ADP
alces-297	3	23	1987–1998	1987–1998	NUM
alces-297	3	24	.	.	PUNCT
alces-297	4	1	modelled	model	VERB
alces-297	4	2	moose	moose	NOUN
alces-297	4	3	,	,	PUNCT
alces-297	4	4	wolf	wolf	NOUN
alces-297	4	5	,	,	PUNCT
alces-297	4	6	and	and	CCONJ
alces-297	4	7	harvest	harvest	NOUN
alces-297	4	8	parameters	parameter	NOUN
alces-297	4	9	compared	compare	VERB
alces-297	4	10	favourably	favourably	ADV
alces-297	4	11	with	with	ADP
alces-297	4	12	observed	observed	ADJ
alces-297	4	13	parameters	parameter	NOUN
alces-297	4	14	when	when	SCONJ
alces-297	4	15	the	the	DET
alces-297	4	16	annual	annual	ADJ
alces-297	4	17	rate	rate	NOUN
alces-297	4	18	of	of	ADP
alces-297	4	19	wolf	wolf	PROPN
alces-297	4	20	removal	removal	NOUN
alces-297	4	21	(	(	PUNCT
alces-297	4	22	human	human	NOUN
alces-297	4	23	-	-	PUNCT
alces-297	4	24	caused	cause	VERB
alces-297	4	25	wolf	wolf	PROPN
alces-297	4	26	mortality	mortality	NOUN
alces-297	4	27	)	)	PUNCT
alces-297	4	28	was	be	AUX
alces-297	4	29	31	31	NUM
alces-297	4	30	%	%	NOUN
alces-297	4	31	.	.	PUNCT
alces-297	5	1	sy	sy	PROPN
alces-297	5	2	curves	curve	NOUN
alces-297	5	3	were	be	AUX
alces-297	5	4	modelled	model	VERB
alces-297	5	5	by	by	ADP
alces-297	5	6	incrementally	incrementally	ADV
alces-297	5	7	increasing	increase	VERB
alces-297	5	8	wolf	wolf	PROPN
alces-297	5	9	removal	removal	NOUN
alces-297	5	10	rates	rate	NOUN
alces-297	5	11	from	from	ADP
alces-297	5	12	0	0	NUM
alces-297	5	13	to	to	PART
alces-297	5	14	40	40	NUM
alces-297	5	15	%	%	NOUN
alces-297	5	16	while	while	SCONJ
alces-297	5	17	maintaining	maintain	VERB
alces-297	5	18	selective	selective	ADJ
alces-297	5	19	moose	moose	NOUN
alces-297	5	20	harvests	harvest	NOUN
alces-297	5	21	of	of	ADP
alces-297	5	22	16	16	NUM
alces-297	5	23	%	%	NOUN
alces-297	5	24	bulls	bull	NOUN
alces-297	5	25	,	,	PUNCT
alces-297	5	26	2	2	NUM
alces-297	5	27	%	%	NOUN
alces-297	5	28	cows	cow	NOUN
alces-297	5	29	and	and	CCONJ
alces-297	5	30	9	9	NUM
alces-297	5	31	%	%	NOUN
alces-297	5	32	calves	calf	NOUN
alces-297	5	33	.	.	PUNCT
alces-297	6	1	sys	sys	PROPN
alces-297	6	2	displayed	display	VERB
alces-297	6	3	an	an	DET
alces-297	6	4	s	s	NOUN
alces-297	6	5	-	-	PUNCT
alces-297	6	6	shape	shape	NOUN
alces-297	6	7	curve	curve	NOUN
alces-297	6	8	with	with	ADP
alces-297	6	9	wolf	wolf	PROPN
alces-297	6	10	removal	removal	NOUN
alces-297	6	11	rates	rate	NOUN
alces-297	6	12	,	,	PUNCT
alces-297	6	13	a	a	DET
alces-297	6	14	hook	hook	NOUN
alces-297	6	15	-	-	PUNCT
alces-297	6	16	shape	shape	NOUN
alces-297	6	17	curve	curve	NOUN
alces-297	6	18	with	with	ADP
alces-297	6	19	wolf	wolf	PROPN
alces-297	6	20	densities	density	NOUN
alces-297	6	21	,	,	PUNCT
alces-297	6	22	and	and	CCONJ
alces-297	6	23	were	be	AUX
alces-297	6	24	linearly	linearly	ADV
alces-297	6	25	related	relate	VERB
alces-297	6	26	to	to	ADP
alces-297	6	27	moose	moose	NOUN
alces-297	6	28	density	density	NOUN
alces-297	6	29	.	.	PUNCT
alces-297	7	1	optimal	optimal	ADJ
alces-297	7	2	harvests	harvest	NOUN
alces-297	7	3	included	include	VERB
alces-297	7	4	a	a	DET
alces-297	7	5	moderate	moderate	ADJ
alces-297	7	6	harvest	harvest	NOUN
alces-297	7	7	of	of	ADP
alces-297	7	8	bulls	bull	NOUN
alces-297	7	9	(	(	PUNCT
alces-297	7	10	16–21	16–21	NUM
alces-297	7	11	%	%	NOUN
alces-297	7	12	)	)	PUNCT
alces-297	7	13	,	,	PUNCT
alces-297	7	14	a	a	DET
alces-297	7	15	nil	nil	NOUN
alces-297	7	16	-	-	PUNCT
alces-297	7	17	to	to	ADP
alces-297	7	18	-	-	PUNCT
alces-297	7	19	very	very	ADV
alces-297	7	20	low	low	ADJ
alces-297	7	21	harvest	harvest	NOUN
alces-297	7	22	of	of	ADP
alces-297	7	23	cows	cow	NOUN
alces-297	7	24	(	(	PUNCT
alces-297	7	25	0–0.2	0–0.2	NOUN
alces-297	7	26	%	%	NOUN
alces-297	7	27	)	)	PUNCT
alces-297	7	28	,	,	PUNCT
alces-297	7	29	and	and	CCONJ
alces-297	7	30	moderate	moderate	ADJ
alces-297	7	31	-	-	PUNCT
alces-297	7	32	to	to	ADP
alces-297	7	33	-	-	PUNCT
alces-297	7	34	high	high	ADJ
alces-297	7	35	harvests	harvest	NOUN
alces-297	7	36	of	of	ADP
alces-297	7	37	calves	calf	NOUN
alces-297	7	38	(	(	PUNCT
alces-297	7	39	15–43	15–43	NUM
alces-297	7	40	%	%	NOUN
alces-297	7	41	)	)	PUNCT
alces-297	7	42	when	when	SCONJ
alces-297	7	43	wolf	wolf	PROPN
alces-297	7	44	removal	removal	NOUN
alces-297	7	45	rates	rate	NOUN
alces-297	7	46	were	be	AUX
alces-297	7	47	≥	≥	NUM
alces-297	7	48	20	20	NUM
alces-297	7	49	%	%	NOUN
alces-297	7	50	.	.	PUNCT
alces-297	8	1	higher	high	ADJ
alces-297	8	2	cow	cow	NOUN
alces-297	8	3	harvest	harvest	NOUN
alces-297	8	4	rates	rate	NOUN
alces-297	8	5	(	(	PUNCT
alces-297	8	6	2	2	NUM
alces-297	8	7	%	%	NOUN
alces-297	8	8	)	)	PUNCT
alces-297	8	9	could	could	AUX
alces-297	8	10	be	be	AUX
alces-297	8	11	accommodated	accommodate	VERB
alces-297	8	12	without	without	ADP
alces-297	8	13	substantially	substantially	ADV
alces-297	8	14	lowering	lower	VERB
alces-297	8	15	sys	sys	NOUN
alces-297	8	16	if	if	SCONJ
alces-297	8	17	calf	calf	NOUN
alces-297	8	18	harvest	harvest	NOUN
alces-297	8	19	rates	rate	NOUN
alces-297	8	20	were	be	AUX
alces-297	8	21	reduced	reduce	VERB
alces-297	8	22	.	.	PUNCT
alces-297	9	1	optimal	optimal	ADJ
alces-297	9	2	harvest	harvest	NOUN
alces-297	9	3	rates	rate	NOUN
alces-297	9	4	did	do	AUX
alces-297	9	5	not	not	PART
alces-297	9	6	improve	improve	VERB
alces-297	9	7	yields	yield	NOUN
alces-297	9	8	over	over	ADP
alces-297	9	9	bull	bull	NOUN
alces-297	9	10	-	-	PUNCT
alces-297	9	11	only	only	ADV
alces-297	9	12	hunting	hunting	NOUN
alces-297	9	13	when	when	SCONJ
alces-297	9	14	wolf	wolf	PROPN
alces-297	9	15	removal	removal	NOUN
alces-297	9	16	rates	rate	NOUN
alces-297	9	17	were	be	AUX
alces-297	9	18	0–10	0–10	NOUN
alces-297	9	19	%	%	NOUN
alces-297	9	20	and	and	CCONJ
alces-297	9	21	management	management	NOUN
alces-297	9	22	constraints	constraint	NOUN
alces-297	9	23	were	be	AUX
alces-297	9	24	placed	place	VERB
alces-297	9	25	on	on	ADP
alces-297	9	26	adult	adult	NOUN
alces-297	9	27	sex	sex	NOUN
alces-297	9	28	ratios	ratio	NOUN
alces-297	9	29	.	.	PUNCT
alces-297	10	1	this	this	DET
alces-297	10	2	study	study	NOUN
alces-297	10	3	supports	support	VERB
alces-297	10	4	previous	previous	ADJ
alces-297	10	5	findings	finding	NOUN
alces-297	10	6	that	that	SCONJ
alces-297	10	7	the	the	DET
alces-297	10	8	optimal	optimal	ADJ
alces-297	10	9	harvest	harvest	NOUN
alces-297	10	10	strategy	strategy	NOUN
alces-297	10	11	for	for	ADP
alces-297	10	12	moose	moose	NOUN
alces-297	10	13	should	should	AUX
alces-297	10	14	primarily	primarily	ADV
alces-297	10	15	target	target	VERB
alces-297	10	16	bulls	bull	NOUN
alces-297	10	17	and	and	CCONJ
alces-297	10	18	calves	calf	NOUN
alces-297	10	19	,	,	PUNCT
alces-297	10	20	whereas	whereas	SCONJ
alces-297	10	21	cows	cow	NOUN
alces-297	10	22	should	should	AUX
alces-297	10	23	be	be	AUX
alces-297	10	24	harvested	harvest	VERB
alces-297	10	25	minimally	minimally	ADV
alces-297	10	26	.	.	PUNCT
alces-297	11	1	however	however	ADV
alces-297	11	2	,	,	PUNCT
alces-297	11	3	for	for	ADP
alces-297	11	4	low	low	ADJ
alces-297	11	5	-	-	PUNCT
alces-297	11	6	density	density	NOUN
alces-297	11	7	,	,	PUNCT
alces-297	11	8	predator	predator	NOUN
alces-297	11	9	limited	limited	ADJ
alces-297	11	10	moose	moose	NOUN
alces-297	11	11	populations	population	NOUN
alces-297	11	12	,	,	PUNCT
alces-297	11	13	bull	bull	NOUN
alces-297	11	14	-	-	PUNCT
alces-297	11	15	only	only	ADV
alces-297	11	16	harvests	harvest	NOUN
alces-297	11	17	may	may	AUX
alces-297	11	18	provide	provide	VERB
alces-297	11	19	equivalent	equivalent	ADJ
alces-297	11	20	yields	yield	NOUN
alces-297	11	21	while	while	SCONJ
alces-297	11	22	maintaining	maintain	VERB
alces-297	11	23	higher	high	ADJ
alces-297	11	24	moose	moose	NOUN
alces-297	11	25	and	and	CCONJ
alces-297	11	26	wolf	wolf	NOUN
alces-297	11	27	densities	density	NOUN
alces-297	11	28	.	.	PUNCT
alces-297	12	1	alces	alce	NOUN
alces-297	12	2	vol	vol	NOUN
alces-297	12	3	.	.	PUNCT
alces-297	13	1	57	57	NUM
alces-297	13	2	:	:	PUNCT
alces-297	13	3	113–129	113–129	NUM
alces-297	13	4	(	(	PUNCT
alces-297	13	5	2021	2021	NUM
alces-297	13	6	)	)	PUNCT
alces-297	13	7	key	key	ADJ
alces-297	13	8	words	word	NOUN
alces-297	13	9	:	:	PUNCT
alces-297	13	10	alces	alce	NOUN
alces-297	13	11	alces	alce	NOUN
alces-297	13	12	,	,	PUNCT
alces-297	13	13	bears	bear	NOUN
alces-297	13	14	,	,	PUNCT
alces-297	13	15	harvest	harvest	NOUN
alces-297	13	16	rate	rate	NOUN
alces-297	13	17	,	,	PUNCT
alces-297	13	18	moose	moose	NOUN
alces-297	13	19	,	,	PUNCT
alces-297	13	20	optimal	optimal	ADJ
alces-297	13	21	harvests	harvest	NOUN
alces-297	13	22	,	,	PUNCT
alces-297	13	23	predation	predation	NOUN
alces-297	13	24	rate	rate	NOUN
alces-297	13	25	,	,	PUNCT
alces-297	13	26	predator	predator	NOUN
alces-297	13	27	-	-	PUNCT
alces-297	13	28	prey	prey	NOUN
alces-297	13	29	model	model	NOUN
alces-297	13	30	,	,	PUNCT
alces-297	13	31	sustained	sustained	ADJ
alces-297	13	32	yield	yield	NOUN
alces-297	13	33	,	,	PUNCT
alces-297	13	34	wolves	wolf	NOUN
alces-297	13	35	,	,	PUNCT
alces-297	13	36	yield	yield	NOUN
alces-297	13	37	-	-	PUNCT
alces-297	13	38	density	density	NOUN
alces-297	13	39	curve	curve	NOUN
alces-297	13	40	one	one	NUM
alces-297	13	41	of	of	ADP
alces-297	13	42	the	the	DET
alces-297	13	43	fundamental	fundamental	ADJ
alces-297	13	44	principles	principle	NOUN
alces-297	13	45	of	of	ADP
alces-297	13	46	wildlife	wildlife	NOUN
alces-297	13	47	harvesting	harvesting	NOUN
alces-297	13	48	is	be	AUX
alces-297	13	49	that	that	SCONJ
alces-297	13	50	it	it	PRON
alces-297	13	51	must	must	AUX
alces-297	13	52	result	result	VERB
alces-297	13	53	in	in	ADP
alces-297	13	54	a	a	DET
alces-297	13	55	sustained	sustained	ADJ
alces-297	13	56	yield	yield	NOUN
alces-297	13	57	(	(	PUNCT
alces-297	13	58	sy	sy	NOUN
alces-297	13	59	)	)	PUNCT
alces-297	13	60	,	,	PUNCT
alces-297	13	61	a	a	DET
alces-297	13	62	harvest	harvest	NOUN
alces-297	13	63	that	that	PRON
alces-297	13	64	can	can	AUX
alces-297	13	65	be	be	AUX
alces-297	13	66	taken	take	VERB
alces-297	13	67	year	year	NOUN
alces-297	13	68	after	after	ADP
alces-297	13	69	year	year	NOUN
alces-297	13	70	without	without	ADP
alces-297	13	71	jeopardizing	jeopardize	VERB
alces-297	13	72	future	future	ADJ
alces-297	13	73	harvests	harvest	NOUN
alces-297	13	74	(	(	PUNCT
alces-297	13	75	sinclair	sinclair	PROPN
alces-297	13	76	et	et	PROPN
alces-297	13	77	al	al	PROPN
alces-297	13	78	.	.	PROPN
alces-297	13	79	2006	2006	NUM
alces-297	13	80	)	)	PUNCT
alces-297	13	81	.	.	PUNCT
alces-297	14	1	caughley	caughley	NOUN
alces-297	14	2	(	(	PUNCT
alces-297	14	3	1976	1976	NUM
alces-297	14	4	)	)	PUNCT
alces-297	14	5	proposed	propose	VERB
alces-297	14	6	a	a	DET
alces-297	14	7	general	general	ADJ
alces-297	14	8	model	model	NOUN
alces-297	14	9	for	for	ADP
alces-297	14	10	harvesting	harvesting	NOUN
alces-297	14	11	ungulates	ungulate	NOUN
alces-297	14	12	that	that	PRON
alces-297	14	13	produces	produce	VERB
alces-297	14	14	a	a	DET
alces-297	14	15	yield	yield	NOUN
alces-297	14	16	-	-	PUNCT
alces-297	14	17	density	density	NOUN
alces-297	14	18	curve	curve	NOUN
alces-297	14	19	with	with	ADP
alces-297	14	20	a	a	DET
alces-297	14	21	bell	bell	NOUN
alces-297	14	22	-	-	PUNCT
alces-297	14	23	shape	shape	NOUN
alces-297	14	24	,	,	PUNCT
alces-297	14	25	slightly	slightly	ADV
alces-297	14	26	skewed	skewed	ADJ
alces-297	14	27	to	to	ADP
alces-297	14	28	the	the	DET
alces-297	14	29	right	right	NOUN
alces-297	14	30	,	,	PUNCT
alces-297	14	31	and	and	CCONJ
alces-297	14	32	with	with	ADP
alces-297	14	33	the	the	DET
alces-297	14	34	maximum	maximum	ADJ
alces-297	14	35	sustained	sustained	ADJ
alces-297	14	36	yield	yield	NOUN
alces-297	14	37	(	(	PUNCT
alces-297	14	38	msy	msy	PROPN
alces-297	14	39	)	)	PUNCT
alces-297	14	40	occurring	occur	VERB
alces-297	14	41	at	at	ADP
alces-297	14	42	~	~	PUNCT
alces-297	14	43	70	70	NUM
alces-297	14	44	%	%	NOUN
alces-297	14	45	of	of	ADP
alces-297	14	46	carrying	carry	VERB
alces-297	14	47	capacity	capacity	NOUN
alces-297	14	48	(	(	PUNCT
alces-297	14	49	k	k	NOUN
alces-297	14	50	)	)	PUNCT
alces-297	14	51	.	.	PUNCT
alces-297	15	1	crête	crête	PROPN
alces-297	15	2	(	(	PUNCT
alces-297	15	3	1987	1987	NUM
alces-297	15	4	)	)	PUNCT
alces-297	15	5	expanded	expand	VERB
alces-297	15	6	on	on	ADP
alces-297	15	7	caughley	caughley	PROPN
alces-297	15	8	’s	’s	PART
alces-297	15	9	model	model	NOUN
alces-297	15	10	and	and	CCONJ
alces-297	15	11	proposed	propose	VERB
alces-297	15	12	two	two	NUM
alces-297	15	13	yield	yield	NOUN
alces-297	15	14	-	-	PUNCT
alces-297	15	15	density	density	NOUN
alces-297	15	16	curves	curve	NOUN
alces-297	15	17	for	for	ADP
alces-297	15	18	moose	moose	NOUN
alces-297	15	19	(	(	PUNCT
alces-297	15	20	alces	alce	NOUN
alces-297	15	21	alces	alce	NOUN
alces-297	15	22	)	)	PUNCT
alces-297	15	23	,	,	PUNCT
alces-297	15	24	one	one	NUM
alces-297	15	25	with	with	ADP
alces-297	15	26	wolves	wolf	NOUN
alces-297	15	27	(	(	PUNCT
alces-297	15	28	canis	canis	NOUN
alces-297	15	29	lupus	lupus	NOUN
alces-297	15	30	)	)	PUNCT
alces-297	15	31	and	and	CCONJ
alces-297	15	32	bears	bear	NOUN
alces-297	15	33	(	(	PUNCT
alces-297	15	34	ursus	ursus	PROPN
alces-297	15	35	spp	spp	PROPN
alces-297	15	36	.	.	PUNCT
alces-297	15	37	)	)	PUNCT
alces-297	16	1	present	present	ADJ
alces-297	16	2	(	(	PUNCT
alces-297	16	3	predator	predator	NOUN
alces-297	16	4	limited	limited	ADJ
alces-297	16	5	k	k	PROPN
alces-297	17	1	=	=	SYM
alces-297	17	2	400	400	NUM
alces-297	17	3	moose/1000	moose/1000	NOUN
alces-297	17	4	km2	km2	PROPN
alces-297	17	5	)	)	PUNCT
alces-297	17	6	,	,	PUNCT
alces-297	17	7	and	and	CCONJ
alces-297	17	8	the	the	DET
alces-297	17	9	other	other	ADJ
alces-297	17	10	without	without	ADP
alces-297	17	11	predators	predator	NOUN
alces-297	17	12	(	(	PUNCT
alces-297	17	13	food	food	NOUN
alces-297	17	14	-	-	PUNCT
alces-297	17	15	limited	limit	VERB
alces-297	17	16	k	k	NOUN
alces-297	17	17	=	=	SYM
alces-297	17	18	1000–2000	1000–2000	NUM
alces-297	17	19	moose/1000	moose/1000	NOUN
alces-297	17	20	km2	km2	NOUN
alces-297	17	21	)	)	PUNCT
alces-297	17	22	.	.	PUNCT
alces-297	18	1	the	the	DET
alces-297	18	2	predator	predator	NOUN
alces-297	18	3	-	-	PUNCT
alces-297	18	4	limited	limit	VERB
alces-297	18	5	curve	curve	NOUN
alces-297	18	6	implied	imply	VERB
alces-297	18	7	that	that	SCONJ
alces-297	18	8	lowering	lower	VERB
alces-297	18	9	moose	moose	NOUN
alces-297	18	10	density	density	NOUN
alces-297	18	11	to	to	ADP
alces-297	18	12	200–300	200–300	NUM
alces-297	18	13	moose/1000	moose/1000	NOUN
alces-297	18	14	km2	km2	X
alces-297	18	15	through	through	ADP
alces-297	18	16	hunting	hunting	NOUN
alces-297	18	17	will	will	AUX
alces-297	18	18	increase	increase	VERB
alces-297	18	19	sys	sys	NOUN
alces-297	18	20	about	about	ADV
alces-297	18	21	4	4	NUM
alces-297	18	22	-	-	NOUN
alces-297	18	23	fold	fold	ADJ
alces-297	18	24	.	.	PUNCT
alces-297	19	1	that	that	PRON
alces-297	19	2	is	is	ADV
alces-297	19	3	,	,	PUNCT
alces-297	19	4	by	by	ADP
alces-297	19	5	intensively	intensively	ADV
alces-297	19	6	harvesting	harvesting	NOUN
alces-297	19	7	moose	moose	NOUN
alces-297	19	8	,	,	PUNCT
alces-297	19	9	wolf	wolf	NOUN
alces-297	19	10	numbers	number	NOUN
alces-297	19	11	and	and	CCONJ
alces-297	19	12	predation	predation	NOUN
alces-297	19	13	rates	rate	NOUN
alces-297	19	14	should	should	AUX
alces-297	19	15	decline	decline	VERB
alces-297	19	16	,	,	PUNCT
alces-297	19	17	and	and	CCONJ
alces-297	19	18	moose	moose	NOUN
alces-297	19	19	growth	growth	NOUN
alces-297	19	20	rates	rate	NOUN
alces-297	19	21	modelling	model	VERB
alces-297	19	22	sustained	sustain	VERB
alces-297	19	23	yields	yield	NOUN
alces-297	19	24	for	for	ADP
alces-297	19	25	moose	moose	NOUN
alces-297	19	26	–	–	PUNCT
alces-297	19	27	hatter	hatter	NOUN
alces-297	19	28	.	.	PUNCT
alces-297	20	1	alces	alce	NOUN
alces-297	20	2	vol	vol	NOUN
alces-297	20	3	.	.	PROPN
alces-297	20	4	57	57	NUM
alces-297	20	5	,	,	PUNCT
alces-297	20	6	2021	2021	NUM
alces-297	20	7	114	114	NUM
alces-297	20	8	and	and	CCONJ
alces-297	20	9	harvest	harvest	NOUN
alces-297	20	10	should	should	AUX
alces-297	20	11	increase	increase	VERB
alces-297	20	12	.	.	PUNCT
alces-297	21	1	gasaway	gasaway	ADV
alces-297	21	2	et	et	PROPN
alces-297	21	3	al	al	PROPN
alces-297	21	4	.	.	PROPN
alces-297	22	1	(	(	PUNCT
alces-297	22	2	1992	1992	NUM
alces-297	22	3	)	)	PUNCT
alces-297	22	4	rejected	reject	VERB
alces-297	22	5	the	the	DET
alces-297	22	6	concept	concept	NOUN
alces-297	22	7	of	of	ADP
alces-297	22	8	a	a	DET
alces-297	22	9	bell	bell	NOUN
alces-297	22	10	-	-	PUNCT
alces-297	22	11	shaped	shape	VERB
alces-297	22	12	yield	yield	NOUN
alces-297	22	13	-	-	PUNCT
alces-297	22	14	density	density	NOUN
alces-297	22	15	curve	curve	NOUN
alces-297	22	16	for	for	ADP
alces-297	22	17	moose	moose	NOUN
alces-297	22	18	in	in	ADP
alces-297	22	19	the	the	DET
alces-297	22	20	presence	presence	NOUN
alces-297	22	21	of	of	ADP
alces-297	22	22	wolves	wolf	NOUN
alces-297	22	23	and	and	CCONJ
alces-297	22	24	bears	bear	NOUN
alces-297	22	25	in	in	ADP
alces-297	22	26	alaska	alaska	PROPN
alces-297	22	27	and	and	CCONJ
alces-297	22	28	yukon	yukon	PROPN
alces-297	22	29	,	,	PUNCT
alces-297	22	30	and	and	CCONJ
alces-297	22	31	presented	present	VERB
alces-297	22	32	an	an	DET
alces-297	22	33	approximate	approximate	ADJ
alces-297	22	34	sy	sy	NOUN
alces-297	22	35	curve	curve	NOUN
alces-297	22	36	for	for	ADP
alces-297	22	37	moose	moose	NOUN
alces-297	22	38	where	where	SCONJ
alces-297	22	39	sustainable	sustainable	ADJ
alces-297	22	40	harvests	harvest	NOUN
alces-297	22	41	increased	increase	VERB
alces-297	22	42	more	more	ADV
alces-297	22	43	gradually	gradually	ADV
alces-297	22	44	with	with	ADP
alces-297	22	45	moose	moose	NOUN
alces-297	22	46	density	density	NOUN
alces-297	22	47	.	.	PUNCT
alces-297	23	1	their	their	PRON
alces-297	23	2	empirical	empirical	ADJ
alces-297	23	3	data	datum	NOUN
alces-297	23	4	suggested	suggest	VERB
alces-297	23	5	that	that	SCONJ
alces-297	23	6	wolves	wolf	NOUN
alces-297	23	7	and	and	CCONJ
alces-297	23	8	bears	bear	NOUN
alces-297	23	9	remained	remain	VERB
alces-297	23	10	effective	effective	ADJ
alces-297	23	11	predators	predator	NOUN
alces-297	23	12	on	on	ADP
alces-297	23	13	calves	calf	NOUN
alces-297	23	14	at	at	ADP
alces-297	23	15	very	very	ADV
alces-297	23	16	low	low	ADJ
alces-297	23	17	densities	density	NOUN
alces-297	23	18	,	,	PUNCT
alces-297	23	19	and	and	CCONJ
alces-297	23	20	that	that	DET
alces-297	23	21	sys	sys	NOUN
alces-297	23	22	were	be	AUX
alces-297	23	23	lower	low	ADJ
alces-297	23	24	than	than	SCONJ
alces-297	23	25	predicted	predict	VERB
alces-297	23	26	by	by	ADP
alces-297	23	27	crête	crête	PROPN
alces-297	23	28	’s	’s	PART
alces-297	23	29	(	(	PUNCT
alces-297	23	30	1987	1987	NUM
alces-297	23	31	)	)	PUNCT
alces-297	23	32	predator	predator	NOUN
alces-297	23	33	-	-	PUNCT
alces-297	23	34	limited	limited	ADJ
alces-297	23	35	,	,	PUNCT
alces-297	23	36	yield	yield	NOUN
alces-297	23	37	-	-	PUNCT
alces-297	23	38	density	density	NOUN
alces-297	23	39	curve	curve	NOUN
alces-297	23	40	.	.	PUNCT
alces-297	24	1	gasaway	gasaway	NOUN
alces-297	24	2	et	et	PROPN
alces-297	24	3	al	al	PROPN
alces-297	24	4	.	.	PROPN
alces-297	25	1	(	(	PUNCT
alces-297	25	2	1992	1992	NUM
alces-297	25	3	)	)	PUNCT
alces-297	25	4	concluded	conclude	VERB
alces-297	25	5	that	that	SCONJ
alces-297	25	6	predation	predation	NOUN
alces-297	25	7	by	by	ADP
alces-297	25	8	lightly	lightly	ADV
alces-297	25	9	harvested	harvest	VERB
alces-297	25	10	wolf	wolf	NOUN
alces-297	25	11	,	,	PUNCT
alces-297	25	12	grizzly	grizzly	NOUN
alces-297	25	13	bear	bear	NOUN
alces-297	25	14	(	(	PUNCT
alces-297	25	15	u.	u.	PROPN
alces-297	25	16	arctos	arctos	PROPN
alces-297	25	17	)	)	PUNCT
alces-297	25	18	,	,	PUNCT
alces-297	25	19	and	and	CCONJ
alces-297	25	20	black	black	ADJ
alces-297	25	21	bear	bear	NOUN
alces-297	25	22	(	(	PUNCT
alces-297	25	23	u.	u.	NOUN
alces-297	25	24	americanus	americanus	PROPN
alces-297	25	25	)	)	PUNCT
alces-297	25	26	populations	population	NOUN
alces-297	25	27	in	in	ADP
alces-297	25	28	alaska	alaska	PROPN
alces-297	25	29	and	and	CCONJ
alces-297	25	30	yukon	yukon	PROPN
alces-297	25	31	lowered	lower	VERB
alces-297	25	32	and	and	CCONJ
alces-297	25	33	maintained	maintain	VERB
alces-297	25	34	moose	moose	NOUN
alces-297	25	35	populations	population	NOUN
alces-297	25	36	within	within	ADP
alces-297	25	37	a	a	DET
alces-297	25	38	low	low	ADJ
alces-297	25	39	density	density	NOUN
alces-297	25	40	dynamic	dynamic	ADJ
alces-297	25	41	equilibrium	equilibrium	NOUN
alces-297	25	42	(	(	PUNCT
alces-297	25	43	ldde	ldde	ADJ
alces-297	25	44	,	,	PUNCT
alces-297	25	45	≤	≤	NUM
alces-297	25	46	417	417	NUM
alces-297	25	47	moose/1000	moose/1000	NOUN
alces-297	25	48	km2	km2	PROPN
alces-297	25	49	)	)	PUNCT
alces-297	25	50	,	,	PUNCT
alces-297	25	51	and	and	CCONJ
alces-297	25	52	that	that	SCONJ
alces-297	25	53	intensive	intensive	ADJ
alces-297	25	54	harvesting	harvesting	NOUN
alces-297	25	55	of	of	ADP
alces-297	25	56	predators	predator	NOUN
alces-297	25	57	,	,	PUNCT
alces-297	25	58	rather	rather	ADV
alces-297	25	59	than	than	ADP
alces-297	25	60	moose	moose	NOUN
alces-297	25	61	,	,	PUNCT
alces-297	25	62	was	be	AUX
alces-297	25	63	required	require	VERB
alces-297	25	64	to	to	PART
alces-297	25	65	elevate	elevate	VERB
alces-297	25	66	sys	sys	PROPN
alces-297	25	67	.	.	PROPN
alces-297	25	68	hatter	hatter	NOUN
alces-297	25	69	(	(	PUNCT
alces-297	25	70	1999	1999	NUM
alces-297	25	71	)	)	PUNCT
alces-297	25	72	developed	develop	VERB
alces-297	25	73	a	a	DET
alces-297	25	74	preliminary	preliminary	ADJ
alces-297	25	75	yield	yield	NOUN
alces-297	25	76	-	-	PUNCT
alces-297	25	77	density	density	NOUN
alces-297	25	78	curve	curve	NOUN
alces-297	25	79	for	for	ADP
alces-297	25	80	moose	moose	NOUN
alces-297	25	81	in	in	ADP
alces-297	25	82	north	north	NOUN
alces-297	25	83	-	-	PUNCT
alces-297	25	84	central	central	ADJ
alces-297	25	85	british	british	PROPN
alces-297	25	86	columbia	columbia	PROPN
alces-297	25	87	(	(	PUNCT
alces-297	25	88	bc	bc	PROPN
alces-297	25	89	)	)	PUNCT
alces-297	25	90	that	that	PRON
alces-297	25	91	was	be	AUX
alces-297	25	92	similar	similar	ADJ
alces-297	25	93	in	in	ADP
alces-297	25	94	shape	shape	NOUN
alces-297	25	95	to	to	ADP
alces-297	25	96	the	the	DET
alces-297	25	97	yield	yield	NOUN
alces-297	25	98	-	-	PUNCT
alces-297	25	99	density	density	NOUN
alces-297	25	100	curve	curve	NOUN
alces-297	25	101	for	for	ADP
alces-297	25	102	alaska	alaska	PROPN
alces-297	25	103	and	and	CCONJ
alces-297	25	104	yukon	yukon	PROPN
alces-297	25	105	(	(	PUNCT
alces-297	25	106	gasaway	gasaway	NOUN
alces-297	25	107	et	et	PROPN
alces-297	25	108	al	al	PROPN
alces-297	25	109	.	.	PROPN
alces-297	25	110	1992	1992	NUM
alces-297	25	111	)	)	PUNCT
alces-297	25	112	,	,	PUNCT
alces-297	25	113	suggesting	suggest	VERB
alces-297	25	114	that	that	SCONJ
alces-297	25	115	some	some	DET
alces-297	25	116	moose	moose	NOUN
alces-297	25	117	populations	population	NOUN
alces-297	25	118	in	in	ADP
alces-297	25	119	bc	bc	PROPN
alces-297	25	120	may	may	AUX
alces-297	25	121	also	also	ADV
alces-297	25	122	exist	exist	VERB
alces-297	25	123	within	within	ADP
alces-297	25	124	a	a	DET
alces-297	25	125	ldde	ldde	NOUN
alces-297	25	126	.	.	PUNCT
alces-297	26	1	although	although	SCONJ
alces-297	26	2	moose	moose	NOUN
alces-297	26	3	are	be	AUX
alces-297	26	4	limited	limit	VERB
alces-297	26	5	by	by	ADP
alces-297	26	6	wolf	wolf	NOUN
alces-297	26	7	and	and	CCONJ
alces-297	26	8	bear	bear	VERB
alces-297	26	9	predation	predation	NOUN
alces-297	26	10	throughout	throughout	ADP
alces-297	26	11	much	much	ADJ
alces-297	26	12	of	of	ADP
alces-297	26	13	north	north	PROPN
alces-297	26	14	america	america	PROPN
alces-297	26	15	(	(	PUNCT
alces-297	26	16	bergerud	bergerud	PROPN
alces-297	26	17	et	et	PROPN
alces-297	26	18	al	al	PROPN
alces-297	26	19	.	.	PROPN
alces-297	26	20	1983	1983	NUM
alces-297	26	21	,	,	PUNCT
alces-297	26	22	gasaway	gasaway	NOUN
alces-297	26	23	et	et	PROPN
alces-297	26	24	al	al	PROPN
alces-297	26	25	.	.	PROPN
alces-297	26	26	1983	1983	NUM
alces-297	26	27	,	,	PUNCT
alces-297	26	28	1992	1992	NUM
alces-297	26	29	,	,	PUNCT
alces-297	26	30	crête	crête	NOUN
alces-297	26	31	1987	1987	NUM
alces-297	26	32	,	,	PUNCT
alces-297	26	33	messier	messy	ADJ
alces-297	26	34	1994	1994	NUM
alces-297	26	35	,	,	PUNCT
alces-297	26	36	van	van	PROPN
alces-297	26	37	ballenberghe	ballenberghe	NOUN
alces-297	26	38	and	and	CCONJ
alces-297	26	39	ballard	ballard	PROPN
alces-297	26	40	1994	1994	NUM
alces-297	26	41	)	)	PUNCT
alces-297	26	42	,	,	PUNCT
alces-297	26	43	predator	predator	NOUN
alces-297	26	44	-	-	PUNCT
alces-297	26	45	prey	prey	NOUN
alces-297	26	46	models	model	NOUN
alces-297	26	47	are	be	AUX
alces-297	26	48	rarely	rarely	ADV
alces-297	26	49	used	use	VERB
alces-297	26	50	in	in	ADP
alces-297	26	51	determining	determine	VERB
alces-297	26	52	sys	sys	NOUN
alces-297	26	53	for	for	ADP
alces-297	26	54	moose	moose	NOUN
alces-297	26	55	.	.	PUNCT
alces-297	27	1	van	van	PROPN
alces-297	27	2	ballenberghe	ballenberghe	NOUN
alces-297	27	3	and	and	CCONJ
alces-297	27	4	dart	dart	NOUN
alces-297	27	5	(	(	PUNCT
alces-297	27	6	1982	1982	NUM
alces-297	27	7	)	)	PUNCT
alces-297	27	8	used	use	VERB
alces-297	27	9	a	a	DET
alces-297	27	10	simple	simple	ADJ
alces-297	27	11	conceptual	conceptual	ADJ
alces-297	27	12	model	model	NOUN
alces-297	27	13	to	to	PART
alces-297	27	14	examine	examine	VERB
alces-297	27	15	harvest	harvest	NOUN
alces-297	27	16	yields	yield	NOUN
alces-297	27	17	subject	subject	ADJ
alces-297	27	18	to	to	ADP
alces-297	27	19	wolf	wolf	NOUN
alces-297	27	20	and	and	CCONJ
alces-297	27	21	bear	bear	VERB
alces-297	27	22	predation	predation	NOUN
alces-297	27	23	and	and	CCONJ
alces-297	27	24	found	find	VERB
alces-297	27	25	that	that	SCONJ
alces-297	27	26	bull	bull	NOUN
alces-297	27	27	-	-	PUNCT
alces-297	27	28	only	only	ADJ
alces-297	27	29	hunts	hunt	NOUN
alces-297	27	30	provided	provide	VERB
alces-297	27	31	an	an	DET
alces-297	27	32	equivalent	equivalent	ADJ
alces-297	27	33	numerical	numerical	ADJ
alces-297	27	34	yield	yield	NOUN
alces-297	27	35	to	to	ADP
alces-297	27	36	either	either	ADV
alces-297	27	37	-	-	PUNCT
alces-297	27	38	sex	sex	NOUN
alces-297	27	39	hunts	hunt	NOUN
alces-297	27	40	,	,	PUNCT
alces-297	27	41	but	but	CCONJ
alces-297	27	42	had	have	VERB
alces-297	27	43	a	a	DET
alces-297	27	44	much	much	ADV
alces-297	27	45	higher	high	ADJ
alces-297	27	46	margin	margin	NOUN
alces-297	27	47	of	of	ADP
alces-297	27	48	safety	safety	NOUN
alces-297	27	49	for	for	ADP
alces-297	27	50	management	management	NOUN
alces-297	27	51	errors	error	NOUN
alces-297	27	52	.	.	PUNCT
alces-297	28	1	more	more	ADV
alces-297	28	2	recently	recently	ADV
alces-297	28	3	,	,	PUNCT
alces-297	28	4	attention	attention	NOUN
alces-297	28	5	has	have	AUX
alces-297	28	6	been	be	AUX
alces-297	28	7	given	give	VERB
alces-297	28	8	to	to	ADP
alces-297	28	9	optimizing	optimize	VERB
alces-297	28	10	sys	sys	PROPN
alces-297	28	11	,	,	PUNCT
alces-297	28	12	or	or	CCONJ
alces-297	28	13	obtaining	obtain	VERB
alces-297	28	14	the	the	DET
alces-297	28	15	maximum	maximum	ADJ
alces-297	28	16	yield	yield	NOUN
alces-297	28	17	by	by	ADP
alces-297	28	18	selectively	selectively	ADV
alces-297	28	19	harvesting	harvesting	NOUN
alces-297	28	20	bulls	bull	NOUN
alces-297	28	21	,	,	PUNCT
alces-297	28	22	cows	cow	NOUN
alces-297	28	23	,	,	PUNCT
alces-297	28	24	and	and	CCONJ
alces-297	28	25	calves	calf	NOUN
alces-297	28	26	at	at	ADP
alces-297	28	27	different	different	ADJ
alces-297	28	28	rates	rate	NOUN
alces-297	28	29	.	.	PUNCT
alces-297	29	1	sæther	sæther	PROPN
alces-297	29	2	et	et	PROPN
alces-297	29	3	al	al	PROPN
alces-297	29	4	.	.	PROPN
alces-297	30	1	(	(	PUNCT
alces-297	30	2	2001	2001	NUM
alces-297	30	3	)	)	PUNCT
alces-297	30	4	concluded	conclude	VERB
alces-297	30	5	that	that	SCONJ
alces-297	30	6	the	the	DET
alces-297	30	7	optimal	optimal	ADJ
alces-297	30	8	harvest	harvest	NOUN
alces-297	30	9	strategy	strategy	NOUN
alces-297	30	10	in	in	ADP
alces-297	30	11	northern	northern	ADJ
alces-297	30	12	norway	norway	NOUN
alces-297	30	13	with	with	ADP
alces-297	30	14	few	few	ADJ
alces-297	30	15	large	large	ADJ
alces-297	30	16	predators	predator	NOUN
alces-297	30	17	,	,	PUNCT
alces-297	30	18	but	but	CCONJ
alces-297	30	19	within	within	ADP
alces-297	30	20	a	a	DET
alces-297	30	21	fluctuating	fluctuate	VERB
alces-297	30	22	environment	environment	NOUN
alces-297	30	23	,	,	PUNCT
alces-297	30	24	involved	involve	VERB
alces-297	30	25	a	a	DET
alces-297	30	26	high	high	ADJ
alces-297	30	27	harvest	harvest	NOUN
alces-297	30	28	of	of	ADP
alces-297	30	29	calves	calf	NOUN
alces-297	30	30	and	and	CCONJ
alces-297	30	31	bulls	bull	NOUN
alces-297	30	32	,	,	PUNCT
alces-297	30	33	and	and	CCONJ
alces-297	30	34	that	that	SCONJ
alces-297	30	35	cows	cow	NOUN
alces-297	30	36	should	should	AUX
alces-297	30	37	hardly	hardly	ADV
alces-297	30	38	be	be	AUX
alces-297	30	39	harvested	harvest	VERB
alces-297	30	40	.	.	PUNCT
alces-297	31	1	nilsen	nilsen	PROPN
alces-297	31	2	et	et	PROPN
alces-297	31	3	al	al	PROPN
alces-297	31	4	.	.	PROPN
alces-297	32	1	(	(	PUNCT
alces-297	32	2	2005	2005	NUM
alces-297	32	3	)	)	PUNCT
alces-297	32	4	considered	consider	VERB
alces-297	32	5	moose	moose	NOUN
alces-297	32	6	populations	population	NOUN
alces-297	32	7	in	in	ADP
alces-297	32	8	south	south	ADJ
alces-297	32	9	-	-	PUNCT
alces-297	32	10	eastern	eastern	ADJ
alces-297	32	11	norway	norway	NOUN
alces-297	32	12	where	where	SCONJ
alces-297	32	13	due	due	ADJ
alces-297	32	14	to	to	ADP
alces-297	32	15	strict	strict	ADJ
alces-297	32	16	management	management	NOUN
alces-297	32	17	control	control	NOUN
alces-297	32	18	there	there	PRON
alces-297	32	19	was	be	VERB
alces-297	32	20	no	no	DET
alces-297	32	21	numerical	numerical	ADJ
alces-297	32	22	response	response	NOUN
alces-297	32	23	by	by	ADP
alces-297	32	24	wolves	wolf	NOUN
alces-297	32	25	,	,	PUNCT
alces-297	32	26	and	and	CCONJ
alces-297	32	27	found	find	VERB
alces-297	32	28	that	that	SCONJ
alces-297	32	29	in	in	ADP
alces-297	32	30	the	the	DET
alces-297	32	31	presence	presence	NOUN
alces-297	32	32	or	or	CCONJ
alces-297	32	33	absence	absence	NOUN
alces-297	32	34	of	of	ADP
alces-297	32	35	predation	predation	NOUN
alces-297	32	36	,	,	PUNCT
alces-297	32	37	a	a	DET
alces-297	32	38	high	high	ADJ
alces-297	32	39	proportion	proportion	NOUN
alces-297	32	40	of	of	ADP
alces-297	32	41	calves	calf	NOUN
alces-297	32	42	in	in	ADP
alces-297	32	43	the	the	DET
alces-297	32	44	harvest	harvest	NOUN
alces-297	32	45	gave	give	VERB
alces-297	32	46	the	the	DET
alces-297	32	47	highest	high	ADJ
alces-297	32	48	sys	sys	PROPN
alces-297	32	49	.	.	PUNCT
alces-297	33	1	xu	xu	PROPN
alces-297	33	2	and	and	CCONJ
alces-297	33	3	boyce	boyce	PROPN
alces-297	33	4	(	(	PUNCT
alces-297	33	5	2010	2010	NUM
alces-297	33	6	)	)	PUNCT
alces-297	33	7	considered	consider	VERB
alces-297	33	8	moose	moose	NOUN
alces-297	33	9	populations	population	NOUN
alces-297	33	10	in	in	ADP
alces-297	33	11	central	central	ADJ
alces-297	33	12	alberta	alberta	PROPN
alces-297	33	13	subject	subject	PROPN
alces-297	33	14	to	to	ADP
alces-297	33	15	predation	predation	NOUN
alces-297	33	16	and	and	CCONJ
alces-297	33	17	stochastic	stochastic	ADJ
alces-297	33	18	weather	weather	NOUN
alces-297	33	19	events	event	NOUN
alces-297	33	20	,	,	PUNCT
alces-297	33	21	and	and	CCONJ
alces-297	33	22	also	also	ADV
alces-297	33	23	concluded	conclude	VERB
alces-297	33	24	that	that	SCONJ
alces-297	33	25	when	when	SCONJ
alces-297	33	26	optimizing	optimize	VERB
alces-297	33	27	total	total	ADJ
alces-297	33	28	yield	yield	NOUN
alces-297	33	29	,	,	PUNCT
alces-297	33	30	bulls	bull	NOUN
alces-297	33	31	and	and	CCONJ
alces-297	33	32	calves	calf	NOUN
alces-297	33	33	should	should	AUX
alces-297	33	34	be	be	AUX
alces-297	33	35	subject	subject	ADJ
alces-297	33	36	to	to	ADP
alces-297	33	37	intense	intense	ADJ
alces-297	33	38	harvest	harvest	NOUN
alces-297	33	39	,	,	PUNCT
alces-297	33	40	with	with	ADP
alces-297	33	41	a	a	DET
alces-297	33	42	low	low	ADJ
alces-297	33	43	harvest	harvest	NOUN
alces-297	33	44	of	of	ADP
alces-297	33	45	females	female	NOUN
alces-297	33	46	.	.	PUNCT
alces-297	34	1	while	while	SCONJ
alces-297	34	2	the	the	DET
alces-297	34	3	effect	effect	NOUN
alces-297	34	4	of	of	ADP
alces-297	34	5	predators	predator	NOUN
alces-297	34	6	on	on	ADP
alces-297	34	7	sys	sys	NOUN
alces-297	34	8	was	be	AUX
alces-297	34	9	considered	consider	VERB
alces-297	34	10	by	by	ADP
alces-297	34	11	nilsen	nilsen	PROPN
alces-297	34	12	et	et	PROPN
alces-297	34	13	al	al	PROPN
alces-297	34	14	.	.	PROPN
alces-297	35	1	(	(	PUNCT
alces-297	35	2	2005	2005	NUM
alces-297	35	3	)	)	PUNCT
alces-297	35	4	and	and	CCONJ
alces-297	35	5	xu	xu	PROPN
alces-297	35	6	and	and	CCONJ
alces-297	35	7	boyce	boyce	PROPN
alces-297	35	8	(	(	PUNCT
alces-297	35	9	2010	2010	NUM
alces-297	35	10	)	)	PUNCT
alces-297	35	11	,	,	PUNCT
alces-297	35	12	neither	neither	DET
alces-297	35	13	study	study	NOUN
alces-297	35	14	modelled	model	VERB
alces-297	35	15	predator	predator	NOUN
alces-297	35	16	-	-	PUNCT
alces-297	35	17	prey	prey	NOUN
alces-297	35	18	dynamics	dynamic	NOUN
alces-297	35	19	in	in	ADP
alces-297	35	20	detail	detail	NOUN
alces-297	35	21	.	.	PUNCT
alces-297	36	1	the	the	DET
alces-297	36	2	purpose	purpose	NOUN
alces-297	36	3	of	of	ADP
alces-297	36	4	this	this	DET
alces-297	36	5	study	study	NOUN
alces-297	36	6	was	be	AUX
alces-297	36	7	to	to	PART
alces-297	36	8	develop	develop	VERB
alces-297	36	9	a	a	DET
alces-297	36	10	model	model	NOUN
alces-297	36	11	that	that	PRON
alces-297	36	12	explicitly	explicitly	ADV
alces-297	36	13	considered	consider	VERB
alces-297	36	14	predator	predator	NOUN
alces-297	36	15	-	-	PUNCT
alces-297	36	16	prey	prey	NOUN
alces-297	36	17	interactions	interaction	NOUN
alces-297	36	18	over	over	ADP
alces-297	36	19	a	a	DET
alces-297	36	20	range	range	NOUN
alces-297	36	21	of	of	ADP
alces-297	36	22	fall	fall	NOUN
alces-297	36	23	wolf	wolf	NOUN
alces-297	36	24	densities	density	NOUN
alces-297	36	25	,	,	PUNCT
alces-297	36	26	and	and	CCONJ
alces-297	36	27	to	to	PART
alces-297	36	28	use	use	VERB
alces-297	36	29	the	the	DET
alces-297	36	30	model	model	NOUN
alces-297	36	31	to	to	PART
alces-297	36	32	investigate	investigate	VERB
alces-297	36	33	how	how	SCONJ
alces-297	36	34	predation	predation	NOUN
alces-297	36	35	affects	affect	VERB
alces-297	36	36	sys	sys	NOUN
alces-297	36	37	and	and	CCONJ
alces-297	36	38	optimal	optimal	ADJ
alces-297	36	39	harvesting	harvesting	NOUN
alces-297	36	40	of	of	ADP
alces-297	36	41	moose	moose	NOUN
alces-297	36	42	in	in	ADP
alces-297	36	43	central	central	ADJ
alces-297	36	44	bc	bc	PROPN
alces-297	36	45	.	.	PUNCT
alces-297	37	1	i	i	PRON
alces-297	37	2	considered	consider	VERB
alces-297	37	3	the	the	DET
alces-297	37	4	optimal	optimal	ADJ
alces-297	37	5	harvest	harvest	NOUN
alces-297	37	6	to	to	PART
alces-297	37	7	be	be	AUX
alces-297	37	8	the	the	DET
alces-297	37	9	highest	high	ADJ
alces-297	37	10	sustained	sustained	ADJ
alces-297	37	11	yield	yield	NOUN
alces-297	37	12	of	of	ADP
alces-297	37	13	moose	moose	NOUN
alces-297	37	14	from	from	ADP
alces-297	37	15	all	all	DET
alces-297	37	16	sex	sex	NOUN
alces-297	37	17	/	/	SYM
alces-297	37	18	age	age	NOUN
alces-297	37	19	classes	class	NOUN
alces-297	37	20	,	,	PUNCT
alces-297	37	21	as	as	SCONJ
alces-297	37	22	opposed	oppose	VERB
alces-297	37	23	to	to	ADP
alces-297	37	24	the	the	DET
alces-297	37	25	greatest	great	ADJ
alces-297	37	26	carcass	carcass	NOUN
alces-297	37	27	weight	weight	NOUN
alces-297	37	28	or	or	CCONJ
alces-297	37	29	maximizing	maximize	VERB
alces-297	37	30	hunting	hunt	VERB
alces-297	37	31	opportunity	opportunity	NOUN
alces-297	37	32	.	.	PUNCT
alces-297	38	1	study	study	NOUN
alces-297	38	2	area	area	NOUN
alces-297	38	3	the	the	DET
alces-297	38	4	19,000	19,000	NUM
alces-297	38	5	km2	km2	PROPN
alces-297	38	6	study	study	NOUN
alces-297	38	7	area	area	NOUN
alces-297	38	8	,	,	PUNCT
alces-297	38	9	hereafter	hereafter	ADV
alces-297	38	10	referred	refer	VERB
alces-297	38	11	to	to	ADP
alces-297	38	12	as	as	SCONJ
alces-297	38	13	the	the	DET
alces-297	38	14	prince	prince	PROPN
alces-297	38	15	george	george	PROPN
alces-297	38	16	study	study	PROPN
alces-297	38	17	area	area	NOUN
alces-297	38	18	(	(	PUNCT
alces-297	38	19	pgsa	pgsa	NOUN
alces-297	38	20	)	)	PUNCT
alces-297	38	21	,	,	PUNCT
alces-297	38	22	was	be	AUX
alces-297	38	23	located	locate	VERB
alces-297	38	24	around	around	ADP
alces-297	38	25	prince	prince	PROPN
alces-297	38	26	george	george	PROPN
alces-297	38	27	(	(	PUNCT
alces-297	38	28	53	53	NUM
alces-297	38	29	°	°	NUM
alces-297	38	30	54’n	54’n	NUM
alces-297	38	31	×	×	NOUN
alces-297	38	32	122	122	NUM
alces-297	38	33	°	°	NOUN
alces-297	38	34	41’w	41’w	NUM
alces-297	38	35	)	)	PUNCT
alces-297	38	36	in	in	ADP
alces-297	38	37	central	central	ADJ
alces-297	38	38	bc	bc	PROPN
alces-297	38	39	(	(	PUNCT
alces-297	38	40	fig	fig	NOUN
alces-297	38	41	.	.	PUNCT
alces-297	38	42	1	1	NUM
alces-297	38	43	)	)	PUNCT
alces-297	38	44	.	.	PUNCT
alces-297	39	1	the	the	DET
alces-297	39	2	terrain	terrain	NOUN
alces-297	39	3	is	be	AUX
alces-297	39	4	flat	flat	ADJ
alces-297	39	5	to	to	PART
alces-297	39	6	rolling	roll	VERB
alces-297	39	7	and	and	CCONJ
alces-297	39	8	forests	forest	NOUN
alces-297	39	9	are	be	AUX
alces-297	39	10	mainly	mainly	ADV
alces-297	39	11	hybrid	hybrid	ADJ
alces-297	39	12	white	white	ADJ
alces-297	39	13	-	-	PUNCT
alces-297	39	14	engelmann	engelmann	NOUN
alces-297	39	15	spruce	spruce	NOUN
alces-297	39	16	(	(	PUNCT
alces-297	39	17	picea	picea	NOUN
alces-297	39	18	glauca	glauca	NOUN
alces-297	39	19	x	x	PART
alces-297	39	20	engelmannii	engelmannii	PROPN
alces-297	39	21	)	)	PUNCT
alces-297	39	22	and	and	CCONJ
alces-297	39	23	subalpine	subalpine	PROPN
alces-297	39	24	fir	fir	PROPN
alces-297	39	25	(	(	PUNCT
alces-297	39	26	abies	abie	NOUN
alces-297	39	27	lasiocarpa	lasiocarpa	VERB
alces-297	39	28	)	)	PUNCT
alces-297	39	29	with	with	ADP
alces-297	39	30	extensive	extensive	ADJ
alces-297	39	31	successional	successional	ADJ
alces-297	39	32	stands	stand	NOUN
alces-297	39	33	of	of	ADP
alces-297	39	34	lodgepole	lodgepole	ADJ
alces-297	39	35	pine	pine	PROPN
alces-297	39	36	(	(	PUNCT
alces-297	39	37	pinus	pinus	NOUN
alces-297	39	38	contorta	contorta	PROPN
alces-297	39	39	)	)	PUNCT
alces-297	39	40	.	.	PUNCT
alces-297	40	1	forestry	forestry	NOUN
alces-297	40	2	is	be	AUX
alces-297	40	3	a	a	DET
alces-297	40	4	prominent	prominent	ADJ
alces-297	40	5	industrial	industrial	ADJ
alces-297	40	6	activity	activity	NOUN
alces-297	40	7	and	and	CCONJ
alces-297	40	8	cutblocks	cutblock	NOUN
alces-297	40	9	are	be	AUX
alces-297	40	10	common	common	ADJ
alces-297	40	11	throughout	throughout	ADP
alces-297	40	12	the	the	DET
alces-297	40	13	area	area	NOUN
alces-297	40	14	.	.	PUNCT
alces-297	41	1	moose	moose	PROPN
alces-297	41	2	were	be	AUX
alces-297	41	3	probably	probably	ADV
alces-297	41	4	the	the	DET
alces-297	41	5	alces	alce	NOUN
alces-297	41	6	vol	vol	NOUN
alces-297	41	7	.	.	PROPN
alces-297	41	8	57	57	NUM
alces-297	41	9	,	,	PUNCT
alces-297	41	10	2021	2021	NUM
alces-297	41	11	modelling	model	VERB
alces-297	41	12	sustained	sustain	VERB
alces-297	41	13	yields	yield	NOUN
alces-297	41	14	for	for	ADP
alces-297	41	15	moose	moose	NOUN
alces-297	41	16	–	–	PUNCT
alces-297	41	17	hatter	hatter	NOUN
alces-297	41	18	.	.	PUNCT
alces-297	42	1	115	115	NUM
alces-297	42	2	predominant	predominant	ADJ
alces-297	42	3	ungulate	ungulate	ADJ
alces-297	42	4	prey	prey	NOUN
alces-297	42	5	for	for	ADP
alces-297	42	6	wolves	wolf	NOUN
alces-297	42	7	,	,	PUNCT
alces-297	42	8	black	black	ADJ
alces-297	42	9	bears	bear	NOUN
alces-297	42	10	,	,	PUNCT
alces-297	42	11	and	and	CCONJ
alces-297	42	12	grizzly	grizzly	NOUN
alces-297	42	13	bears	bear	NOUN
alces-297	42	14	because	because	SCONJ
alces-297	42	15	other	other	ADJ
alces-297	42	16	ungulates	ungulate	NOUN
alces-297	42	17	were	be	AUX
alces-297	42	18	rare	rare	ADJ
alces-297	42	19	,	,	PUNCT
alces-297	42	20	but	but	CCONJ
alces-297	42	21	included	include	VERB
alces-297	42	22	mule	mule	ADJ
alces-297	42	23	deer	deer	NOUN
alces-297	42	24	(	(	PUNCT
alces-297	42	25	odocoileus	odocoileus	ADJ
alces-297	42	26	hemionus	hemionus	PROPN
alces-297	42	27	)	)	PUNCT
alces-297	42	28	,	,	PUNCT
alces-297	42	29	white	white	ADJ
alces-297	42	30	-	-	PUNCT
alces-297	42	31	tailed	tail	VERB
alces-297	42	32	deer	deer	NOUN
alces-297	42	33	(	(	PUNCT
alces-297	42	34	o.	o.	NOUN
alces-297	42	35	virginianus	virginianus	PROPN
alces-297	42	36	)	)	PUNCT
alces-297	42	37	,	,	PUNCT
alces-297	42	38	elk	elk	PROPN
alces-297	42	39	(	(	PUNCT
alces-297	42	40	cervus	cervus	NOUN
alces-297	42	41	canadensis	canadensis	NOUN
alces-297	42	42	)	)	PUNCT
alces-297	42	43	,	,	PUNCT
alces-297	42	44	and	and	CCONJ
alces-297	42	45	caribou	caribou	NOUN
alces-297	42	46	(	(	PUNCT
alces-297	42	47	rangifer	rangifer	VERB
alces-297	42	48	tarandus	tarandus	NOUN
alces-297	42	49	)	)	PUNCT
alces-297	42	50	(	(	PUNCT
alces-297	42	51	heard	hear	VERB
alces-297	42	52	et	et	PROPN
alces-297	42	53	al	al	PROPN
alces-297	42	54	.	.	PROPN
alces-297	42	55	1999	1999	NUM
alces-297	42	56	)	)	PUNCT
alces-297	42	57	.	.	PUNCT
alces-297	43	1	the	the	DET
alces-297	43	2	moose	moose	NOUN
alces-297	43	3	density	density	NOUN
alces-297	43	4	estimated	estimate	VERB
alces-297	43	5	in	in	ADP
alces-297	43	6	1998	1998	NUM
alces-297	43	7	was	be	AUX
alces-297	43	8	1320	1320	NUM
alces-297	43	9	moose/1000	moose/1000	NOUN
alces-297	43	10	km2	km2	X
alces-297	43	11	from	from	ADP
alces-297	43	12	a	a	DET
alces-297	43	13	stratified	stratified	ADJ
alces-297	43	14	random	random	ADJ
alces-297	43	15	block	block	NOUN
alces-297	43	16	survey	survey	NOUN
alces-297	43	17	when	when	SCONJ
alces-297	43	18	predator	predator	NOUN
alces-297	43	19	densities	density	NOUN
alces-297	43	20	were	be	AUX
alces-297	43	21	~12.5	~12.5	ADJ
alces-297	43	22	wolves/1000	wolves/1000	PROPN
alces-297	43	23	km2	km2	PROPN
alces-297	43	24	,	,	PUNCT
alces-297	43	25	20	20	NUM
alces-297	43	26	grizzly	grizzly	NOUN
alces-297	43	27	bears/1000	bears/1000	PROPN
alces-297	43	28	km2	km2	PROPN
alces-297	43	29	,	,	PUNCT
alces-297	43	30	and	and	CCONJ
alces-297	43	31	211	211	NUM
alces-297	43	32	black	black	ADJ
alces-297	43	33	bears/1000	bears/1000	NOUN
alces-297	43	34	km2	km2	PROPN
alces-297	43	35	(	(	PUNCT
alces-297	43	36	heard	hear	VERB
alces-297	43	37	et	et	PROPN
alces-297	43	38	al	al	PROPN
alces-297	43	39	.	.	PROPN
alces-297	43	40	1999	1999	NUM
alces-297	43	41	)	)	PUNCT
alces-297	43	42	.	.	PUNCT
alces-297	44	1	since	since	SCONJ
alces-297	44	2	1980	1980	NUM
alces-297	44	3	,	,	PUNCT
alces-297	44	4	selective	selective	ADJ
alces-297	44	5	fall	fall	NOUN
alces-297	44	6	harvests	harvest	NOUN
alces-297	44	7	have	have	AUX
alces-297	44	8	governed	govern	VERB
alces-297	44	9	moose	moose	NOUN
alces-297	44	10	hunting	hunt	VERB
alces-297	44	11	through	through	ADP
alces-297	44	12	a	a	DET
alces-297	44	13	combination	combination	NOUN
alces-297	44	14	of	of	ADP
alces-297	44	15	differential	differential	NOUN
alces-297	44	16	licencing	licencing	NOUN
alces-297	44	17	for	for	ADP
alces-297	44	18	bulls	bull	NOUN
alces-297	44	19	and	and	CCONJ
alces-297	44	20	cows	cow	NOUN
alces-297	44	21	and	and	CCONJ
alces-297	44	22	open	open	ADJ
alces-297	44	23	seasons	season	NOUN
alces-297	44	24	on	on	ADP
alces-297	44	25	2	2	NUM
alces-297	44	26	-	-	PUNCT
alces-297	44	27	point	point	NOUN
alces-297	44	28	bulls	bull	NOUN
alces-297	44	29	and	and	CCONJ
alces-297	44	30	calves	calf	NOUN
alces-297	44	31	(	(	PUNCT
alces-297	44	32	child	child	NOUN
alces-297	44	33	1983	1983	NUM
alces-297	44	34	)	)	PUNCT
alces-297	44	35	.	.	PUNCT
alces-297	45	1	from	from	ADP
alces-297	45	2	1991	1991	NUM
alces-297	45	3	to	to	ADP
alces-297	45	4	1998	1998	NUM
alces-297	45	5	,	,	PUNCT
alces-297	45	6	licenced	licence	VERB
alces-297	45	7	hunters	hunter	NOUN
alces-297	45	8	harvested	harvest	VERB
alces-297	45	9	an	an	DET
alces-297	45	10	average	average	NOUN
alces-297	45	11	of	of	ADP
alces-297	45	12	16	16	NUM
alces-297	45	13	%	%	NOUN
alces-297	45	14	of	of	ADP
alces-297	45	15	the	the	DET
alces-297	45	16	bulls	bull	NOUN
alces-297	45	17	,	,	PUNCT
alces-297	45	18	2	2	NUM
alces-297	45	19	%	%	NOUN
alces-297	45	20	of	of	ADP
alces-297	45	21	cows	cow	NOUN
alces-297	45	22	,	,	PUNCT
alces-297	45	23	and	and	CCONJ
alces-297	45	24	9	9	NUM
alces-297	45	25	%	%	NOUN
alces-297	45	26	of	of	ADP
alces-297	45	27	calves	calf	NOUN
alces-297	45	28	for	for	ADP
alces-297	45	29	a	a	DET
alces-297	45	30	harvest	harvest	NOUN
alces-297	45	31	rate	rate	NOUN
alces-297	45	32	of	of	ADP
alces-297	45	33	7	7	NUM
alces-297	45	34	%	%	NOUN
alces-297	45	35	of	of	ADP
alces-297	45	36	the	the	DET
alces-297	45	37	pre	pre	ADJ
alces-297	45	38	-	-	ADJ
alces-297	45	39	hunt	hunt	ADJ
alces-297	45	40	population	population	NOUN
alces-297	45	41	(	(	PUNCT
alces-297	45	42	heard	hear	VERB
alces-297	45	43	et	et	PROPN
alces-297	45	44	al	al	PROPN
alces-297	45	45	.	.	PROPN
alces-297	45	46	1999	1999	NUM
alces-297	45	47	)	)	PUNCT
alces-297	45	48	.	.	PUNCT
alces-297	46	1	the	the	DET
alces-297	46	2	first	first	ADJ
alces-297	46	3	nations	nation	NOUN
alces-297	46	4	harvest	harvest	NOUN
alces-297	46	5	was	be	AUX
alces-297	46	6	unknown	unknown	ADJ
alces-297	46	7	,	,	PUNCT
alces-297	46	8	but	but	CCONJ
alces-297	46	9	likely	likely	ADV
alces-297	46	10	less	less	ADJ
alces-297	46	11	than	than	ADP
alces-297	46	12	the	the	DET
alces-297	46	13	licenced	licenced	ADJ
alces-297	46	14	harvest	harvest	NOUN
alces-297	46	15	(	(	PUNCT
alces-297	46	16	heard	hear	VERB
alces-297	46	17	,	,	PUNCT
alces-297	46	18	pers	per	NOUN
alces-297	46	19	.	.	PUNCT
alces-297	47	1	comm	comm	NOUN
alces-297	47	2	.	.	PUNCT
alces-297	47	3	)	)	PUNCT
alces-297	47	4	.	.	PUNCT
alces-297	48	1	moose	moose	NOUN
alces-297	48	2	densities	density	NOUN
alces-297	48	3	and	and	CCONJ
alces-297	48	4	composition	composition	NOUN
alces-297	48	5	,	,	PUNCT
alces-297	48	6	harvests	harvest	NOUN
alces-297	48	7	,	,	PUNCT
alces-297	48	8	and	and	CCONJ
alces-297	48	9	hunter	hunter	NOUN
alces-297	48	10	success	success	NOUN
alces-297	48	11	rates	rate	NOUN
alces-297	48	12	appeared	appear	VERB
alces-297	48	13	to	to	PART
alces-297	48	14	be	be	AUX
alces-297	48	15	relatively	relatively	ADV
alces-297	48	16	stable	stable	ADJ
alces-297	48	17	from	from	ADP
alces-297	48	18	1987	1987	NUM
alces-297	48	19	to	to	ADP
alces-297	48	20	1998	1998	NUM
alces-297	48	21	.	.	PUNCT
alces-297	49	1	there	there	PRON
alces-297	49	2	was	be	VERB
alces-297	49	3	no	no	DET
alces-297	49	4	trend	trend	NOUN
alces-297	49	5	in	in	ADP
alces-297	49	6	the	the	DET
alces-297	49	7	number	number	NOUN
alces-297	49	8	of	of	ADP
alces-297	49	9	wolves	wolf	NOUN
alces-297	49	10	shot	shoot	VERB
alces-297	49	11	by	by	ADP
alces-297	49	12	licensed	licensed	ADJ
alces-297	49	13	hunters	hunter	NOUN
alces-297	49	14	or	or	CCONJ
alces-297	49	15	conservation	conservation	NOUN
alces-297	49	16	officers	officer	NOUN
alces-297	49	17	during	during	ADP
alces-297	49	18	this	this	DET
alces-297	49	19	period	period	NOUN
alces-297	49	20	implying	imply	VERB
alces-297	49	21	that	that	SCONJ
alces-297	49	22	wolf	wolf	NOUN
alces-297	49	23	densities	density	NOUN
alces-297	49	24	were	be	AUX
alces-297	49	25	also	also	ADV
alces-297	49	26	stable	stable	ADJ
alces-297	49	27	.	.	PUNCT
alces-297	50	1	heard	hear	VERB
alces-297	50	2	et	et	PROPN
alces-297	50	3	al	al	PROPN
alces-297	50	4	.	.	PUNCT
alces-297	51	1	(	(	PUNCT
alces-297	51	2	1999	1999	NUM
alces-297	51	3	)	)	PUNCT
alces-297	51	4	suggested	suggest	VERB
alces-297	51	5	that	that	SCONJ
alces-297	51	6	the	the	DET
alces-297	51	7	elevated	elevated	ADJ
alces-297	51	8	moose	moose	NOUN
alces-297	51	9	density	density	NOUN
alces-297	51	10	was	be	AUX
alces-297	51	11	possible	possible	ADJ
alces-297	51	12	because	because	SCONJ
alces-297	51	13	wolf	wolf	PROPN
alces-297	51	14	removals	removal	NOUN
alces-297	51	15	(	(	PUNCT
alces-297	51	16	humancaused	humancause	VERB
alces-297	51	17	wolf	wolf	PROPN
alces-297	51	18	mortality	mortality	NOUN
alces-297	51	19	)	)	PUNCT
alces-297	51	20	were	be	AUX
alces-297	51	21	high	high	ADJ
alces-297	51	22	due	due	ADP
alces-297	51	23	to	to	ADP
alces-297	51	24	the	the	DET
alces-297	51	25	combined	combine	VERB
alces-297	51	26	effects	effect	NOUN
alces-297	51	27	of	of	ADP
alces-297	51	28	hunting	hunting	NOUN
alces-297	51	29	,	,	PUNCT
alces-297	51	30	trapping	trap	VERB
alces-297	51	31	,	,	PUNCT
alces-297	51	32	and	and	CCONJ
alces-297	51	33	killing	kill	VERB
alces-297	51	34	wolves	wolf	NOUN
alces-297	51	35	to	to	PART
alces-297	51	36	protect	protect	VERB
alces-297	51	37	livestock	livestock	NOUN
alces-297	51	38	.	.	PUNCT
alces-297	52	1	grizzly	grizzly	NOUN
alces-297	52	2	bears	bear	NOUN
alces-297	52	3	were	be	AUX
alces-297	52	4	lightly	lightly	ADV
alces-297	52	5	hunted	hunt	VERB
alces-297	52	6	and	and	CCONJ
alces-297	52	7	black	black	ADJ
alces-297	52	8	bears	bear	NOUN
alces-297	52	9	appeared	appear	VERB
alces-297	52	10	to	to	PART
alces-297	52	11	be	be	AUX
alces-297	52	12	moderately	moderately	ADV
alces-297	52	13	exploited	exploit	VERB
alces-297	52	14	(	(	PUNCT
alces-297	52	15	heard	hear	VERB
alces-297	52	16	et	et	PROPN
alces-297	52	17	al	al	PROPN
alces-297	52	18	.	.	PROPN
alces-297	52	19	1997	1997	NUM
alces-297	52	20	)	)	PUNCT
alces-297	52	21	.	.	PUNCT
alces-297	53	1	further	further	ADJ
alces-297	53	2	details	detail	NOUN
alces-297	53	3	of	of	ADP
alces-297	53	4	the	the	DET
alces-297	53	5	pgsa	pgsa	NOUN
alces-297	53	6	are	be	AUX
alces-297	53	7	provided	provide	VERB
alces-297	53	8	by	by	ADP
alces-297	53	9	heard	hear	VERB
alces-297	53	10	et	et	PROPN
alces-297	53	11	al	al	PROPN
alces-297	53	12	.	.	PROPN
alces-297	54	1	(	(	PUNCT
alces-297	54	2	1997	1997	NUM
alces-297	54	3	,	,	PUNCT
alces-297	54	4	1999	1999	NUM
alces-297	54	5	)	)	PUNCT
alces-297	54	6	.	.	PUNCT
alces-297	55	1	methods	method	NOUN
alces-297	55	2	model	model	PROPN
alces-297	55	3	structure	structure	NOUN
alces-297	55	4	moose	moose	VERB
alces-297	55	5	the	the	DET
alces-297	55	6	population	population	NOUN
alces-297	55	7	was	be	AUX
alces-297	55	8	stage	stage	NOUN
alces-297	55	9	-	-	PUNCT
alces-297	55	10	structured	structure	VERB
alces-297	55	11	as	as	ADP
alces-297	55	12	bulls	bull	NOUN
alces-297	55	13	(	(	PUNCT
alces-297	55	14	≥	≥	NOUN
alces-297	55	15	1	1	NUM
alces-297	55	16	year	year	NOUN
alces-297	55	17	-	-	PUNCT
alces-297	55	18	old	old	ADJ
alces-297	55	19	males	male	NOUN
alces-297	55	20	)	)	PUNCT
alces-297	55	21	,	,	PUNCT
alces-297	55	22	cows	cow	NOUN
alces-297	55	23	(	(	PUNCT
alces-297	55	24	≥	≥	NUM
alces-297	55	25	1	1	NUM
alces-297	55	26	yearold	yearold	ADJ
alces-297	55	27	females	female	NOUN
alces-297	55	28	)	)	PUNCT
alces-297	55	29	,	,	PUNCT
alces-297	55	30	and	and	CCONJ
alces-297	55	31	calves	calf	NOUN
alces-297	55	32	(	(	PUNCT
alces-297	55	33	<	<	X
alces-297	55	34	1year	1year	NUM
alces-297	55	35	-	-	ADJ
alces-297	55	36	old	old	ADJ
alces-297	55	37	with	with	ADP
alces-297	55	38	50:50	50:50	NUM
alces-297	55	39	sex	sex	NOUN
alces-297	55	40	ratio	ratio	NOUN
alces-297	55	41	)	)	PUNCT
alces-297	55	42	.	.	PUNCT
alces-297	56	1	density	density	NOUN
alces-297	56	2	dependence	dependence	NOUN
alces-297	56	3	in	in	ADP
alces-297	56	4	reproduction	reproduction	NOUN
alces-297	56	5	and	and	CCONJ
alces-297	56	6	survival	survival	NOUN
alces-297	56	7	were	be	AUX
alces-297	56	8	modelled	model	VERB
alces-297	56	9	fig	fig	NOUN
alces-297	56	10	.	.	PUNCT
alces-297	57	1	1	1	X
alces-297	57	2	.	.	X
alces-297	57	3	location	location	NOUN
alces-297	57	4	of	of	ADP
alces-297	57	5	the	the	DET
alces-297	57	6	prince	prince	PROPN
alces-297	57	7	george	george	PROPN
alces-297	57	8	study	study	PROPN
alces-297	57	9	area	area	NOUN
alces-297	57	10	(	(	PUNCT
alces-297	57	11	pgsa	pgsa	NOUN
alces-297	57	12	)	)	PUNCT
alces-297	57	13	in	in	ADP
alces-297	57	14	central	central	ADJ
alces-297	57	15	british	british	PROPN
alces-297	57	16	columbia	columbia	PROPN
alces-297	57	17	,	,	PUNCT
alces-297	57	18	canada	canada	PROPN
alces-297	57	19	.	.	PUNCT
alces-297	58	1	the	the	DET
alces-297	58	2	pgsa	pgsa	NOUN
alces-297	58	3	includes	include	VERB
alces-297	58	4	all	all	PRON
alces-297	58	5	of	of	ADP
alces-297	58	6	wildlife	wildlife	NOUN
alces-297	58	7	management	management	NOUN
alces-297	58	8	units	unit	NOUN
alces-297	58	9	(	(	PUNCT
alces-297	58	10	mu	mu	NOUN
alces-297	58	11	)	)	PUNCT
alces-297	58	12	707–715	707–715	NUM
alces-297	58	13	,	,	PUNCT
alces-297	58	14	and	and	CCONJ
alces-297	58	15	the	the	DET
alces-297	58	16	lower	low	ADJ
alces-297	58	17	portions	portion	NOUN
alces-297	58	18	of	of	ADP
alces-297	58	19	716	716	NUM
alces-297	58	20	and	and	CCONJ
alces-297	58	21	724	724	NUM
alces-297	58	22	.	.	PUNCT
alces-297	59	1	modelling	model	VERB
alces-297	59	2	sustained	sustained	ADJ
alces-297	59	3	yields	yield	NOUN
alces-297	59	4	for	for	ADP
alces-297	59	5	moose	moose	NOUN
alces-297	59	6	–	–	PUNCT
alces-297	59	7	hatter	hatter	NOUN
alces-297	59	8	.	.	PUNCT
alces-297	60	1	alces	alce	NOUN
alces-297	60	2	vol	vol	NOUN
alces-297	60	3	.	.	PROPN
alces-297	60	4	57	57	NUM
alces-297	60	5	,	,	PUNCT
alces-297	60	6	2021	2021	NUM
alces-297	60	7	116	116	NUM
alces-297	60	8	followed	follow	VERB
alces-297	60	9	the	the	DET
alces-297	60	10	approach	approach	NOUN
alces-297	60	11	outlined	outline	VERB
alces-297	60	12	by	by	ADP
alces-297	60	13	mcnay	mcnay	PROPN
alces-297	60	14	and	and	CCONJ
alces-297	60	15	delong	delong	PROPN
alces-297	60	16	(	(	PUNCT
alces-297	60	17	1998	1998	NUM
alces-297	60	18	)	)	PUNCT
alces-297	60	19	and	and	CCONJ
alces-297	60	20	ranged	range	VERB
alces-297	60	21	from	from	ADP
alces-297	60	22	51	51	NUM
alces-297	60	23	to	to	PART
alces-297	60	24	85	85	NUM
alces-297	60	25	calves/100	calves/100	NOUN
alces-297	60	26	cows	cow	NOUN
alces-297	60	27	at	at	ADP
alces-297	60	28	birth	birth	NOUN
alces-297	60	29	,	,	PUNCT
alces-297	60	30	15–39	15–39	NUM
alces-297	60	31	%	%	NOUN
alces-297	60	32	summer	summer	NOUN
alces-297	60	33	calf	calf	NOUN
alces-297	60	34	mortality	mortality	NOUN
alces-297	60	35	,	,	PUNCT
alces-297	60	36	10–12	10–12	NUM
alces-297	60	37	%	%	NOUN
alces-297	60	38	winter	winter	NOUN
alces-297	60	39	calf	calf	NOUN
alces-297	60	40	mortality	mortality	NOUN
alces-297	60	41	,	,	PUNCT
alces-297	60	42	2–12	2–12	ADP
alces-297	60	43	%	%	NOUN
alces-297	60	44	winter	winter	NOUN
alces-297	60	45	cow	cow	NOUN
alces-297	60	46	mortality	mortality	NOUN
alces-297	60	47	,	,	PUNCT
alces-297	60	48	and	and	CCONJ
alces-297	60	49	4–12	4–12	NOUN
alces-297	60	50	%	%	NOUN
alces-297	60	51	winter	winter	NOUN
alces-297	60	52	bull	bull	NOUN
alces-297	60	53	mortality	mortality	NOUN
alces-297	60	54	.	.	PUNCT
alces-297	61	1	all	all	DET
alces-297	61	2	density	density	NOUN
alces-297	61	3	dependent	dependent	ADJ
alces-297	61	4	responses	response	NOUN
alces-297	61	5	started	start	VERB
alces-297	61	6	at	at	ADP
alces-297	61	7	~	~	PUNCT
alces-297	61	8	65	65	NUM
alces-297	61	9	%	%	NOUN
alces-297	61	10	of	of	ADP
alces-297	61	11	k	k	PROPN
alces-297	61	12	and	and	CCONJ
alces-297	61	13	generally	generally	ADV
alces-297	61	14	increased	increase	VERB
alces-297	61	15	in	in	ADP
alces-297	61	16	a	a	DET
alces-297	61	17	linear	linear	NOUN
alces-297	61	18	manner	manner	NOUN
alces-297	61	19	;	;	PUNCT
alces-297	61	20	k	k	PROPN
alces-297	61	21	was	be	AUX
alces-297	61	22	set	set	VERB
alces-297	61	23	to	to	ADP
alces-297	61	24	2000	2000	NUM
alces-297	61	25	moose/1000	moose/1000	NOUN
alces-297	61	26	km2	km2	PROPN
alces-297	61	27	(	(	PUNCT
alces-297	61	28	crête	crête	PROPN
alces-297	61	29	1987	1987	NUM
alces-297	61	30	,	,	PUNCT
alces-297	61	31	heard	hear	VERB
alces-297	61	32	et	et	PROPN
alces-297	61	33	al	al	PROPN
alces-297	61	34	.	.	PROPN
alces-297	61	35	1999	1999	NUM
alces-297	61	36	)	)	PUNCT
alces-297	61	37	.	.	PUNCT
alces-297	62	1	summer	summer	NOUN
alces-297	62	2	adult	adult	NOUN
alces-297	62	3	mortality	mortality	NOUN
alces-297	62	4	(	(	PUNCT
alces-297	62	5	2	2	NUM
alces-297	62	6	%	%	NOUN
alces-297	62	7	)	)	PUNCT
alces-297	62	8	was	be	AUX
alces-297	62	9	assumed	assume	VERB
alces-297	62	10	to	to	PART
alces-297	62	11	be	be	AUX
alces-297	62	12	density	density	NOUN
alces-297	62	13	independent	independent	ADJ
alces-297	62	14	.	.	PUNCT
alces-297	63	1	the	the	DET
alces-297	63	2	modelled	model	VERB
alces-297	63	3	population	population	NOUN
alces-297	63	4	was	be	AUX
alces-297	63	5	censused	census	VERB
alces-297	63	6	3	3	NUM
alces-297	63	7	times	time	NOUN
alces-297	63	8	per	per	ADP
alces-297	63	9	year	year	NOUN
alces-297	63	10	including	include	VERB
alces-297	63	11	the	the	DET
alces-297	63	12	post	post	NOUN
alces-297	63	13	-	-	NOUN
alces-297	63	14	hunt	hunt	ADJ
alces-297	63	15	,	,	PUNCT
alces-297	63	16	post	post	ADJ
alces-297	63	17	-	-	ADJ
alces-297	63	18	calving	calving	ADJ
alces-297	63	19	,	,	PUNCT
alces-297	63	20	and	and	CCONJ
alces-297	63	21	pre	pre	ADJ
alces-297	63	22	-	-	ADJ
alces-297	63	23	hunt	hunt	ADJ
alces-297	63	24	periods	period	NOUN
alces-297	63	25	.	.	PUNCT
alces-297	64	1	wounding	wound	VERB
alces-297	64	2	loss	loss	NOUN
alces-297	64	3	was	be	AUX
alces-297	64	4	set	set	VERB
alces-297	64	5	to	to	ADP
alces-297	64	6	15	15	NUM
alces-297	64	7	%	%	NOUN
alces-297	64	8	of	of	ADP
alces-297	64	9	the	the	DET
alces-297	64	10	licenced	licenced	ADJ
alces-297	64	11	harvest	harvest	NOUN
alces-297	64	12	(	(	PUNCT
alces-297	64	13	kuzyk	kuzyk	NOUN
alces-297	64	14	et	et	PROPN
alces-297	64	15	al	al	PROPN
alces-297	64	16	.	.	PROPN
alces-297	64	17	2018	2018	NUM
alces-297	64	18	)	)	PUNCT
alces-297	64	19	.	.	PUNCT
alces-297	65	1	first	first	ADJ
alces-297	65	2	nations	nation	NOUN
alces-297	65	3	harvest	harvest	NOUN
alces-297	65	4	was	be	AUX
alces-297	65	5	assumed	assume	VERB
alces-297	65	6	to	to	PART
alces-297	65	7	be	be	AUX
alces-297	65	8	unselective	unselective	ADJ
alces-297	65	9	and	and	CCONJ
alces-297	65	10	occur	occur	VERB
alces-297	65	11	during	during	ADP
alces-297	65	12	winter	winter	NOUN
alces-297	65	13	.	.	PUNCT
alces-297	66	1	the	the	DET
alces-297	66	2	maximum	maximum	ADJ
alces-297	66	3	harvest	harvest	NOUN
alces-297	66	4	rate	rate	NOUN
alces-297	66	5	by	by	ADP
alces-297	66	6	first	first	ADJ
alces-297	66	7	nations	nation	NOUN
alces-297	66	8	was	be	AUX
alces-297	66	9	presumed	presume	VERB
alces-297	66	10	to	to	PART
alces-297	66	11	be	be	AUX
alces-297	66	12	7	7	NUM
alces-297	66	13	%	%	NOUN
alces-297	66	14	of	of	ADP
alces-297	66	15	the	the	DET
alces-297	66	16	post	post	ADJ
alces-297	66	17	-	-	ADJ
alces-297	66	18	hunt	hunt	ADJ
alces-297	66	19	moose	moose	NOUN
alces-297	66	20	population	population	NOUN
alces-297	66	21	at	at	ADP
alces-297	66	22	k	k	PROPN
alces-297	66	23	,	,	PUNCT
alces-297	66	24	declining	decline	VERB
alces-297	66	25	linearly	linearly	ADV
alces-297	66	26	to	to	ADP
alces-297	66	27	0	0	NUM
alces-297	66	28	%	%	NOUN
alces-297	66	29	at	at	ADP
alces-297	66	30	100	100	NUM
alces-297	66	31	moose/1000	moose/1000	NOUN
alces-297	66	32	km2	km2	PROPN
alces-297	66	33	.	.	PUNCT
alces-297	67	1	where	where	SCONJ
alces-297	67	2	not	not	PART
alces-297	67	3	specified	specify	VERB
alces-297	67	4	,	,	PUNCT
alces-297	67	5	harvest	harvest	NOUN
alces-297	67	6	rates	rate	NOUN
alces-297	67	7	were	be	AUX
alces-297	67	8	based	base	VERB
alces-297	67	9	on	on	ADP
alces-297	67	10	the	the	DET
alces-297	67	11	pre	pre	ADJ
alces-297	67	12	-	-	ADJ
alces-297	67	13	hunt	hunt	ADJ
alces-297	67	14	moose	moose	NOUN
alces-297	67	15	numbers	number	NOUN
alces-297	67	16	,	,	PUNCT
alces-297	67	17	while	while	SCONJ
alces-297	67	18	moose	moose	NOUN
alces-297	67	19	densities	density	NOUN
alces-297	67	20	and	and	CCONJ
alces-297	67	21	sex	sex	NOUN
alces-297	67	22	-	-	PUNCT
alces-297	67	23	age	age	NOUN
alces-297	67	24	composition	composition	NOUN
alces-297	67	25	apply	apply	VERB
alces-297	67	26	to	to	ADP
alces-297	67	27	the	the	DET
alces-297	67	28	post	post	ADJ
alces-297	67	29	-	-	ADJ
alces-297	67	30	hunt	hunt	ADJ
alces-297	67	31	period	period	NOUN
alces-297	67	32	.	.	PUNCT
alces-297	68	1	wolves	wolf	NOUN
alces-297	68	2	changes	change	VERB
alces-297	68	3	in	in	ADP
alces-297	68	4	wolf	wolf	PROPN
alces-297	68	5	density	density	NOUN
alces-297	68	6	were	be	AUX
alces-297	68	7	modelled	model	VERB
alces-297	68	8	as	as	ADP
alces-297	68	9	a	a	DET
alces-297	68	10	ratio	ratio	NOUN
alces-297	68	11	-	-	PUNCT
alces-297	68	12	dependent	dependent	ADJ
alces-297	68	13	numerical	numerical	ADJ
alces-297	68	14	response	response	NOUN
alces-297	68	15	:	:	PUNCT
alces-297	68	16	w	w	PROPN
alces-297	68	17	w	w	PROPN
alces-297	68	18	rw	rw	PROPN
alces-297	68	19	w	w	PROPN
alces-297	68	20	amt	amt	PROPN
alces-297	68	21	t	t	PROPN
alces-297	68	22	t	t	PROPN
alces-297	68	23	t	t	PROPN
alces-297	68	24	t	t	PROPN
alces-297	68	25	=	=	PUNCT
alces-297	69	1	+	+	CCONJ
alces-297	70	1	−	−	NUM
alces-297	70	2			NOUN
alces-297	70	3			NOUN
alces-297	70	4			NOUN
alces-297	70	5			NOUN
alces-297	70	6			PUNCT
alces-297	71	1	−	−	NUM
alces-297	71	2	−	−	NOUN
alces-297	72	1	−	−	NOUN
alces-297	72	2	−	−	NUM
alces-297	72	3	1	1	NUM
alces-297	72	4	1	1	NUM
alces-297	72	5	1	1	NUM
alces-297	72	6	1	1	NUM
alces-297	72	7	1	1	NUM
alces-297	72	8	(	(	PUNCT
alces-297	72	9	eberhardt	eberhardt	PROPN
alces-297	72	10	1997	1997	NUM
alces-297	72	11	,	,	PUNCT
alces-297	72	12	hatter	hatter	NOUN
alces-297	72	13	2019	2019	NUM
alces-297	72	14	)	)	PUNCT
alces-297	72	15	where	where	SCONJ
alces-297	72	16	wt	wt	NOUN
alces-297	72	17	denotes	denote	NOUN
alces-297	72	18	fall	fall	VERB
alces-297	72	19	wolf	wolf	PROPN
alces-297	72	20	density	density	NOUN
alces-297	72	21	at	at	ADP
alces-297	72	22	time	time	NOUN
alces-297	72	23	t	t	PROPN
alces-297	72	24	,	,	PUNCT
alces-297	72	25	r	r	NOUN
alces-297	72	26	is	be	AUX
alces-297	72	27	the	the	DET
alces-297	72	28	maximal	maximal	ADJ
alces-297	72	29	rate	rate	NOUN
alces-297	72	30	of	of	ADP
alces-297	72	31	increase	increase	NOUN
alces-297	72	32	for	for	ADP
alces-297	72	33	wolves	wolf	NOUN
alces-297	72	34	(	(	PUNCT
alces-297	72	35	r	r	NOUN
alces-297	72	36	=	=	SYM
alces-297	72	37	0.46	0.46	NUM
alces-297	72	38	,	,	PUNCT
alces-297	72	39	keith	keith	PROPN
alces-297	72	40	1983	1983	NUM
alces-297	72	41	)	)	PUNCT
alces-297	72	42	,	,	PUNCT
alces-297	72	43	a	a	PRON
alces-297	72	44	is	be	AUX
alces-297	72	45	the	the	DET
alces-297	72	46	equilibrium	equilibrium	NOUN
alces-297	72	47	ratio	ratio	NOUN
alces-297	72	48	of	of	ADP
alces-297	72	49	wolves	wolf	NOUN
alces-297	72	50	:	:	PUNCT
alces-297	72	51	moose	moose	NOUN
alces-297	72	52	,	,	PUNCT
alces-297	72	53	and	and	CCONJ
alces-297	72	54	mt	mt	PROPN
alces-297	72	55	is	be	AUX
alces-297	72	56	moose	moose	NOUN
alces-297	72	57	density	density	NOUN
alces-297	72	58	.	.	PUNCT
alces-297	73	1	eberhardt	eberhardt	PROPN
alces-297	73	2	et	et	PROPN
alces-297	73	3	al	al	PROPN
alces-297	73	4	.	.	PROPN
alces-297	74	1	(	(	PUNCT
alces-297	74	2	2003	2003	NUM
alces-297	74	3	)	)	PUNCT
alces-297	74	4	estimated	estimate	VERB
alces-297	74	5	a	a	DET
alces-297	74	6	=	=	SYM
alces-297	74	7	0.049	0.049	NUM
alces-297	74	8	by	by	ADP
alces-297	74	9	linear	linear	ADJ
alces-297	74	10	regression	regression	NOUN
alces-297	74	11	of	of	ADP
alces-297	74	12	the	the	DET
alces-297	74	13	observed	observe	VERB
alces-297	74	14	finite	finite	ADJ
alces-297	74	15	rate	rate	NOUN
alces-297	74	16	of	of	ADP
alces-297	74	17	increase	increase	NOUN
alces-297	74	18	of	of	ADP
alces-297	74	19	wolves	wolf	NOUN
alces-297	74	20	(	(	PUNCT
alces-297	74	21	λ	λ	NOUN
alces-297	74	22	)	)	PUNCT
alces-297	74	23	against	against	ADP
alces-297	74	24	the	the	DET
alces-297	74	25	wolf	wolf	NOUN
alces-297	74	26	:	:	PUNCT
alces-297	74	27	moose	moose	NOUN
alces-297	74	28	ratio	ratio	NOUN
alces-297	74	29	.	.	PUNCT
alces-297	75	1	i	i	PRON
alces-297	75	2	added	add	VERB
alces-297	75	3	6	6	NUM
alces-297	75	4	additional	additional	ADJ
alces-297	75	5	data	datum	NOUN
alces-297	75	6	points	point	NOUN
alces-297	75	7	(	(	PUNCT
alces-297	75	8	appendix	appendix	NOUN
alces-297	75	9	1	1	NUM
alces-297	75	10	)	)	PUNCT
alces-297	75	11	and	and	CCONJ
alces-297	75	12	estimated	estimate	VERB
alces-297	75	13	the	the	DET
alces-297	75	14	relationship	relationship	NOUN
alces-297	75	15	as	as	ADP
alces-297	75	16	:	:	PUNCT
alces-297	75	17	y	y	PROPN
alces-297	75	18	e1.534	e1.534	PROPN
alces-297	75	19	x10.72=	x10.72=	ADJ
alces-297	75	20	−	−	PROPN
alces-297	75	21	where	where	SCONJ
alces-297	75	22	y	y	PROPN
alces-297	75	23	is	be	AUX
alces-297	75	24	the	the	DET
alces-297	75	25	observed	observed	ADJ
alces-297	75	26	λ	λ	NOUN
alces-297	75	27	for	for	ADP
alces-297	75	28	wolves	wolf	NOUN
alces-297	75	29	,	,	PUNCT
alces-297	75	30	x	x	PUNCT
alces-297	75	31	is	be	AUX
alces-297	75	32	wolves	wolf	NOUN
alces-297	75	33	/	/	SYM
alces-297	75	34	moose	moose	NOUN
alces-297	75	35	,	,	PUNCT
alces-297	75	36	and	and	CCONJ
alces-297	75	37	a	a	DET
alces-297	75	38	=	=	SYM
alces-297	75	39	0.040	0.040	NUM
alces-297	75	40	(	(	PUNCT
alces-297	75	41	25	25	NUM
alces-297	75	42	moose/	moose/	NUM
alces-297	75	43	wolf	wolf	NOUN
alces-297	75	44	,	,	PUNCT
alces-297	75	45	95	95	NUM
alces-297	75	46	%	%	NOUN
alces-297	75	47	ci	ci	NOUN
alces-297	75	48	:	:	PUNCT
alces-297	75	49	20–29	20–29	NUM
alces-297	75	50	)	)	PUNCT
alces-297	75	51	when	when	SCONJ
alces-297	75	52	y	y	PROPN
alces-297	75	53	=	=	PROPN
alces-297	75	54	1.0	1.0	NUM
alces-297	75	55	.	.	PUNCT
alces-297	76	1	spring	spring	NOUN
alces-297	76	2	wolf	wolf	NOUN
alces-297	76	3	numbers	number	NOUN
alces-297	76	4	were	be	AUX
alces-297	76	5	estimated	estimate	VERB
alces-297	76	6	by	by	ADP
alces-297	76	7	subtracting	subtract	VERB
alces-297	76	8	the	the	DET
alces-297	76	9	annual	annual	ADJ
alces-297	76	10	wolf	wolf	NOUN
alces-297	76	11	removal	removal	NOUN
alces-297	76	12	from	from	ADP
alces-297	76	13	the	the	DET
alces-297	76	14	fall	fall	NOUN
alces-297	76	15	population	population	NOUN
alces-297	76	16	.	.	PUNCT
alces-297	77	1	the	the	DET
alces-297	77	2	wolf	wolf	PROPN
alces-297	77	3	removal	removal	NOUN
alces-297	77	4	rate	rate	NOUN
alces-297	77	5	included	include	VERB
alces-297	77	6	all	all	DET
alces-297	77	7	human	human	NOUN
alces-297	77	8	-	-	PUNCT
alces-297	77	9	caused	cause	VERB
alces-297	77	10	mortality	mortality	NOUN
alces-297	77	11	.	.	PUNCT
alces-297	78	1	several	several	ADJ
alces-297	78	2	models	model	NOUN
alces-297	78	3	have	have	AUX
alces-297	78	4	been	be	AUX
alces-297	78	5	proposed	propose	VERB
alces-297	78	6	for	for	ADP
alces-297	78	7	estimating	estimate	VERB
alces-297	78	8	the	the	DET
alces-297	78	9	annual	annual	ADJ
alces-297	78	10	kill	kill	NOUN
alces-297	78	11	rate	rate	NOUN
alces-297	78	12	of	of	ADP
alces-297	78	13	moose	moose	NOUN
alces-297	78	14	by	by	ADP
alces-297	78	15	wolves	wolf	NOUN
alces-297	78	16	.	.	PUNCT
alces-297	79	1	messier	messy	ADJ
alces-297	79	2	(	(	PUNCT
alces-297	79	3	1994	1994	NUM
alces-297	79	4	)	)	PUNCT
alces-297	79	5	found	find	VERB
alces-297	79	6	that	that	SCONJ
alces-297	79	7	winter	winter	NOUN
alces-297	79	8	kill	kill	NOUN
alces-297	79	9	rates	rate	NOUN
alces-297	79	10	were	be	AUX
alces-297	79	11	reduced	reduce	VERB
alces-297	79	12	at	at	ADP
alces-297	79	13	low	low	ADJ
alces-297	79	14	moose	moose	NOUN
alces-297	79	15	densities	density	NOUN
alces-297	79	16	,	,	PUNCT
alces-297	79	17	and	and	CCONJ
alces-297	79	18	proposed	propose	VERB
alces-297	79	19	a	a	DET
alces-297	79	20	type	type	NOUN
alces-297	79	21	ii	ii	NOUN
alces-297	79	22	functional	functional	ADJ
alces-297	79	23	response	response	NOUN
alces-297	79	24	.	.	PUNCT
alces-297	80	1	hayes	hayes	PROPN
alces-297	80	2	and	and	CCONJ
alces-297	80	3	harestad	harestad	PROPN
alces-297	80	4	(	(	PUNCT
alces-297	80	5	2000	2000	NUM
alces-297	80	6	)	)	PUNCT
alces-297	80	7	modified	modify	VERB
alces-297	80	8	the	the	DET
alces-297	80	9	parameter	parameter	NOUN
alces-297	80	10	estimates	estimate	NOUN
alces-297	80	11	for	for	ADP
alces-297	80	12	the	the	DET
alces-297	80	13	type	type	NOUN
alces-297	80	14	ii	ii	NOUN
alces-297	80	15	functional	functional	ADJ
alces-297	80	16	response	response	NOUN
alces-297	80	17	based	base	VERB
alces-297	80	18	on	on	ADP
alces-297	80	19	kill	kill	NOUN
alces-297	80	20	rates	rate	NOUN
alces-297	80	21	from	from	ADP
alces-297	80	22	the	the	DET
alces-297	80	23	yukon	yukon	PROPN
alces-297	80	24	,	,	PUNCT
alces-297	80	25	while	while	SCONJ
alces-297	80	26	eberhardt	eberhardt	PROPN
alces-297	80	27	(	(	PUNCT
alces-297	80	28	1997	1997	NUM
alces-297	80	29	)	)	PUNCT
alces-297	80	30	suggested	suggest	VERB
alces-297	80	31	kill	kill	NOUN
alces-297	80	32	rates	rate	NOUN
alces-297	80	33	were	be	AUX
alces-297	80	34	density	density	NOUN
alces-297	80	35	independent	independent	ADJ
alces-297	80	36	and	and	CCONJ
alces-297	80	37	averaged	average	VERB
alces-297	80	38	2.1	2.1	NUM
alces-297	80	39	kills	kill	NOUN
alces-297	80	40	/	/	SYM
alces-297	80	41	wolf/100	wolf/100	NOUN
alces-297	80	42	days	day	NOUN
alces-297	80	43	in	in	ADP
alces-297	80	44	winter	winter	NOUN
alces-297	80	45	.	.	PUNCT
alces-297	81	1	lake	lake	PROPN
alces-297	81	2	et	et	PROPN
alces-297	81	3	al	al	PROPN
alces-297	81	4	.	.	PROPN
alces-297	82	1	(	(	PUNCT
alces-297	82	2	2013	2013	NUM
alces-297	82	3	)	)	PUNCT
alces-297	82	4	concluded	conclude	VERB
alces-297	82	5	that	that	SCONJ
alces-297	82	6	kill	kill	NOUN
alces-297	82	7	rates	rate	NOUN
alces-297	82	8	were	be	AUX
alces-297	82	9	also	also	ADV
alces-297	82	10	density	density	NOUN
alces-297	82	11	independent	independent	ADJ
alces-297	82	12	in	in	ADP
alces-297	82	13	alaska	alaska	PROPN
alces-297	82	14	and	and	CCONJ
alces-297	82	15	yukon	yukon	PROPN
alces-297	82	16	.	.	PUNCT
alces-297	83	1	serrouya	serrouya	PROPN
alces-297	83	2	et	et	PROPN
alces-297	83	3	al	al	PROPN
alces-297	83	4	.	.	PROPN
alces-297	83	5	(	(	PUNCT
alces-297	83	6	2015	2015	NUM
alces-297	83	7	)	)	PUNCT
alces-297	83	8	,	,	PUNCT
alces-297	83	9	however	however	ADV
alces-297	83	10	,	,	PUNCT
alces-297	83	11	found	find	VERB
alces-297	83	12	that	that	SCONJ
alces-297	83	13	messier	messy	ADJ
alces-297	83	14	’s	’s	PART
alces-297	83	15	type	type	NOUN
alces-297	83	16	ii	ii	PROPN
alces-297	83	17	response	response	NOUN
alces-297	83	18	provided	provide	VERB
alces-297	83	19	the	the	DET
alces-297	83	20	best	good	ADJ
alces-297	83	21	fit	fit	NOUN
alces-297	83	22	among	among	ADP
alces-297	83	23	competing	compete	VERB
alces-297	83	24	models	model	NOUN
alces-297	83	25	for	for	ADP
alces-297	83	26	a	a	DET
alces-297	83	27	wolf	wolf	NOUN
alces-297	83	28	-	-	PUNCT
alces-297	83	29	moose	moose	NOUN
alces-297	83	30	-	-	PUNCT
alces-297	83	31	caribou	caribou	NOUN
alces-297	83	32	system	system	NOUN
alces-297	83	33	in	in	ADP
alces-297	83	34	south	south	ADJ
alces-297	83	35	-	-	PUNCT
alces-297	83	36	eastern	eastern	ADJ
alces-297	83	37	bc	bc	PROPN
alces-297	83	38	.	.	PUNCT
alces-297	84	1	i	i	PRON
alces-297	84	2	used	use	VERB
alces-297	84	3	messier	messy	ADJ
alces-297	84	4	’s	’s	PART
alces-297	84	5	(	(	PUNCT
alces-297	84	6	1994	1994	NUM
alces-297	84	7	)	)	PUNCT
alces-297	84	8	functional	functional	ADJ
alces-297	84	9	response	response	NOUN
alces-297	84	10	:	:	PUNCT
alces-297	84	11	y	y	NOUN
alces-297	84	12	x	x	PUNCT
alces-297	84	13	x	x	PUNCT
alces-297	84	14	3.36	3.36	NUM
alces-297	84	15	0.46	0.46	NUM
alces-297	84	16	=	=	PUNCT
alces-297	85	1	+	+	CCONJ
alces-297	85	2	where	where	SCONJ
alces-297	85	3	y	y	PROPN
alces-297	85	4	is	be	AUX
alces-297	85	5	the	the	DET
alces-297	85	6	number	number	NOUN
alces-297	85	7	of	of	ADP
alces-297	85	8	moose	moose	NOUN
alces-297	85	9	killed	kill	VERB
alces-297	85	10	per	per	ADP
alces-297	85	11	wolf	wolf	PROPN
alces-297	85	12	per	per	ADP
alces-297	85	13	100	100	NUM
alces-297	85	14	days	day	NOUN
alces-297	85	15	in	in	ADP
alces-297	85	16	winter	winter	NOUN
alces-297	85	17	,	,	PUNCT
alces-297	85	18	and	and	CCONJ
alces-297	85	19	x	x	X
alces-297	85	20	is	be	AUX
alces-297	85	21	the	the	DET
alces-297	85	22	number	number	NOUN
alces-297	85	23	of	of	ADP
alces-297	85	24	moose	moose	NOUN
alces-297	85	25	/	/	SYM
alces-297	85	26	km2	km2	PROPN
alces-297	85	27	,	,	PUNCT
alces-297	85	28	but	but	CCONJ
alces-297	85	29	capped	cap	VERB
alces-297	85	30	the	the	DET
alces-297	85	31	maximum	maximum	ADJ
alces-297	85	32	killing	killing	NOUN
alces-297	85	33	rate	rate	NOUN
alces-297	85	34	at	at	ADP
alces-297	85	35	2.1	2.1	NUM
alces-297	85	36	as	as	SCONJ
alces-297	85	37	suggested	suggest	VERB
alces-297	85	38	by	by	ADP
alces-297	85	39	eberhardt	eberhardt	PROPN
alces-297	85	40	(	(	PUNCT
alces-297	85	41	1997	1997	NUM
alces-297	85	42	)	)	PUNCT
alces-297	85	43	.	.	PUNCT
alces-297	86	1	following	follow	VERB
alces-297	86	2	eberhardt	eberhardt	PROPN
alces-297	86	3	(	(	PUNCT
alces-297	86	4	1997	1997	NUM
alces-297	86	5	)	)	PUNCT
alces-297	86	6	,	,	PUNCT
alces-297	86	7	i	i	PRON
alces-297	86	8	assumed	assume	VERB
alces-297	86	9	that	that	SCONJ
alces-297	86	10	summer	summer	NOUN
alces-297	86	11	kill	kill	NOUN
alces-297	86	12	rates	rate	NOUN
alces-297	86	13	were	be	AUX
alces-297	86	14	75	75	NUM
alces-297	86	15	%	%	NOUN
alces-297	86	16	of	of	ADP
alces-297	86	17	the	the	DET
alces-297	86	18	winter	winter	NOUN
alces-297	86	19	rate	rate	NOUN
alces-297	86	20	during	during	ADP
alces-297	86	21	5	5	NUM
alces-297	86	22	months	month	NOUN
alces-297	86	23	of	of	ADP
alces-297	86	24	summer	summer	NOUN
alces-297	86	25	(	(	PUNCT
alces-297	86	26	june	june	PROPN
alces-297	86	27	–	–	PUNCT
alces-297	86	28	october	october	PROPN
alces-297	86	29	)	)	PUNCT
alces-297	86	30	.	.	PUNCT
alces-297	87	1	i	i	PRON
alces-297	87	2	used	use	VERB
alces-297	87	3	fall	fall	NOUN
alces-297	87	4	wolf	wolf	PROPN
alces-297	87	5	densities	density	NOUN
alces-297	87	6	for	for	ADP
alces-297	87	7	winter	winter	NOUN
alces-297	87	8	predation	predation	NOUN
alces-297	87	9	and	and	CCONJ
alces-297	87	10	spring	spring	NOUN
alces-297	87	11	wolf	wolf	NOUN
alces-297	87	12	densities	density	NOUN
alces-297	87	13	for	for	ADP
alces-297	87	14	summer	summer	NOUN
alces-297	87	15	predation	predation	NOUN
alces-297	87	16	.	.	PUNCT
alces-297	88	1	wolf	wolf	PROPN
alces-297	88	2	kills	kills	PROPN
alces-297	88	3	were	be	AUX
alces-297	88	4	apportioned	apportion	VERB
alces-297	88	5	among	among	ADP
alces-297	88	6	the	the	DET
alces-297	88	7	bulls	bull	NOUN
alces-297	88	8	,	,	PUNCT
alces-297	88	9	cows	cow	NOUN
alces-297	88	10	,	,	PUNCT
alces-297	88	11	and	and	CCONJ
alces-297	88	12	calves	calf	NOUN
alces-297	88	13	by	by	ADP
alces-297	88	14	assuming	assume	VERB
alces-297	88	15	the	the	DET
alces-297	88	16	relative	relative	ADJ
alces-297	88	17	vulnerability	vulnerability	NOUN
alces-297	88	18	of	of	ADP
alces-297	88	19	calves	calf	NOUN
alces-297	88	20	was	be	AUX
alces-297	88	21	10	10	NUM
alces-297	88	22	-	-	ADJ
alces-297	88	23	fold	fold	VERB
alces-297	88	24	greater	great	ADJ
alces-297	88	25	alces	alce	NOUN
alces-297	88	26	vol	vol	NOUN
alces-297	88	27	.	.	PROPN
alces-297	88	28	57	57	NUM
alces-297	88	29	,	,	PUNCT
alces-297	88	30	2021	2021	NUM
alces-297	88	31	modelling	model	VERB
alces-297	88	32	sustained	sustain	VERB
alces-297	88	33	yields	yield	NOUN
alces-297	88	34	for	for	ADP
alces-297	88	35	moose	moose	NOUN
alces-297	88	36	–	–	PUNCT
alces-297	88	37	hatter	hatter	NOUN
alces-297	88	38	.	.	PUNCT
alces-297	89	1	117	117	NUM
alces-297	89	2	than	than	ADP
alces-297	89	3	adults	adult	NOUN
alces-297	89	4	during	during	ADP
alces-297	89	5	summer	summer	NOUN
alces-297	89	6	and	and	CCONJ
alces-297	89	7	2	2	NUM
alces-297	89	8	-	-	NOUN
alces-297	89	9	fold	fold	VERB
alces-297	89	10	greater	great	ADJ
alces-297	89	11	during	during	ADP
alces-297	89	12	winter	winter	NOUN
alces-297	89	13	.	.	PUNCT
alces-297	90	1	the	the	DET
alces-297	90	2	high	high	ADJ
alces-297	90	3	calf	calf	NOUN
alces-297	90	4	vulnerability	vulnerability	NOUN
alces-297	90	5	reflected	reflect	VERB
alces-297	90	6	the	the	DET
alces-297	90	7	preference	preference	NOUN
alces-297	90	8	by	by	ADP
alces-297	90	9	wolves	wolf	NOUN
alces-297	90	10	for	for	ADP
alces-297	90	11	calves	calf	NOUN
alces-297	90	12	(	(	PUNCT
alces-297	90	13	peterson	peterson	NOUN
alces-297	90	14	et	et	PROPN
alces-297	90	15	al	al	PROPN
alces-297	90	16	.	.	PROPN
alces-297	90	17	1984	1984	NUM
alces-297	90	18	,	,	PUNCT
alces-297	90	19	sand	sand	NOUN
alces-297	90	20	et	et	NOUN
alces-297	90	21	al	al	PROPN
alces-297	90	22	.	.	PROPN
alces-297	90	23	2008	2008	NUM
alces-297	90	24	)	)	PUNCT
alces-297	90	25	.	.	PUNCT
alces-297	91	1	calculations	calculation	NOUN
alces-297	91	2	for	for	ADP
alces-297	91	3	relative	relative	ADJ
alces-297	91	4	vulnerability	vulnerability	NOUN
alces-297	91	5	followed	follow	VERB
alces-297	91	6	mcnay	mcnay	PROPN
alces-297	91	7	and	and	CCONJ
alces-297	91	8	delong	delong	PROPN
alces-297	91	9	(	(	PUNCT
alces-297	91	10	1998	1998	NUM
alces-297	91	11	)	)	PUNCT
alces-297	91	12	.	.	PUNCT
alces-297	92	1	bears	bear	VERB
alces-297	92	2	ballard	ballard	PROPN
alces-297	92	3	(	(	PUNCT
alces-297	92	4	1992	1992	NUM
alces-297	92	5	)	)	PUNCT
alces-297	92	6	reported	report	VERB
alces-297	92	7	that	that	SCONJ
alces-297	92	8	calf	calf	NOUN
alces-297	92	9	mortality	mortality	NOUN
alces-297	92	10	rates	rate	NOUN
alces-297	92	11	due	due	ADP
alces-297	92	12	to	to	ADP
alces-297	92	13	grizzly	grizzly	NOUN
alces-297	92	14	bear	bear	NOUN
alces-297	92	15	predation	predation	NOUN
alces-297	92	16	ranged	range	VERB
alces-297	92	17	from	from	ADP
alces-297	92	18	3	3	NUM
alces-297	92	19	to	to	ADP
alces-297	92	20	52	52	NUM
alces-297	92	21	%	%	NOUN
alces-297	92	22	and	and	CCONJ
alces-297	92	23	predation	predation	NOUN
alces-297	92	24	rates	rate	NOUN
alces-297	92	25	were	be	AUX
alces-297	92	26	independent	independent	ADJ
alces-297	92	27	of	of	ADP
alces-297	92	28	moose	moose	NOUN
alces-297	92	29	density	density	NOUN
alces-297	92	30	;	;	PUNCT
alces-297	92	31	mortality	mortality	NOUN
alces-297	92	32	rates	rate	NOUN
alces-297	92	33	from	from	ADP
alces-297	92	34	black	black	ADJ
alces-297	92	35	bears	bear	NOUN
alces-297	92	36	predation	predation	NOUN
alces-297	92	37	ranged	range	VERB
alces-297	92	38	from	from	ADP
alces-297	92	39	2	2	NUM
alces-297	92	40	to	to	ADP
alces-297	92	41	50	50	NUM
alces-297	92	42	%	%	NOUN
alces-297	92	43	.	.	PUNCT
alces-297	93	1	the	the	DET
alces-297	93	2	predation	predation	NOUN
alces-297	93	3	rates	rate	NOUN
alces-297	93	4	of	of	ADP
alces-297	93	5	grizzly	grizzly	NOUN
alces-297	93	6	and	and	CCONJ
alces-297	93	7	black	black	ADJ
alces-297	93	8	bears	bear	NOUN
alces-297	93	9	on	on	ADP
alces-297	93	10	moose	moose	NOUN
alces-297	93	11	calves	calf	NOUN
alces-297	93	12	in	in	ADP
alces-297	93	13	the	the	DET
alces-297	93	14	pgsa	pgsa	NOUN
alces-297	93	15	were	be	AUX
alces-297	93	16	unknown	unknown	ADJ
alces-297	93	17	.	.	PUNCT
alces-297	94	1	rea	rea	PROPN
alces-297	94	2	et	et	PROPN
alces-297	94	3	al	al	PROPN
alces-297	94	4	.	.	PROPN
alces-297	95	1	(	(	PUNCT
alces-297	95	2	2019	2019	NUM
alces-297	95	3	)	)	PUNCT
alces-297	95	4	found	find	VERB
alces-297	95	5	that	that	SCONJ
alces-297	95	6	only	only	ADV
alces-297	95	7	2	2	NUM
alces-297	95	8	%	%	NOUN
alces-297	95	9	of	of	ADP
alces-297	95	10	bear	bear	NOUN
alces-297	95	11	scats	scat	NOUN
alces-297	95	12	collected	collect	VERB
alces-297	95	13	in	in	ADP
alces-297	95	14	the	the	DET
alces-297	95	15	pgsa	pgsa	NOUN
alces-297	95	16	during	during	ADP
alces-297	95	17	spring	spring	NOUN
alces-297	95	18	and	and	CCONJ
alces-297	95	19	summer	summer	NOUN
alces-297	95	20	(	(	PUNCT
alces-297	95	21	n	n	NOUN
alces-297	95	22	=	=	SYM
alces-297	95	23	1381	1381	NUM
alces-297	95	24	)	)	PUNCT
alces-297	95	25	contained	contain	VERB
alces-297	95	26	moose	moose	NOUN
alces-297	95	27	calf	calf	NOUN
alces-297	95	28	hair	hair	NOUN
alces-297	95	29	;	;	PUNCT
alces-297	95	30	however	however	ADV
alces-297	95	31	,	,	PUNCT
alces-297	95	32	they	they	PRON
alces-297	95	33	noted	note	VERB
alces-297	95	34	that	that	SCONJ
alces-297	95	35	even	even	ADV
alces-297	95	36	this	this	DET
alces-297	95	37	low	low	ADJ
alces-297	95	38	percentage	percentage	NOUN
alces-297	95	39	could	could	AUX
alces-297	95	40	result	result	VERB
alces-297	95	41	in	in	ADP
alces-297	95	42	substantial	substantial	ADJ
alces-297	95	43	calf	calf	NOUN
alces-297	95	44	mortality	mortality	NOUN
alces-297	95	45	.	.	PUNCT
alces-297	96	1	i	i	PRON
alces-297	96	2	assumed	assume	VERB
alces-297	96	3	a	a	DET
alces-297	96	4	combined	combined	ADJ
alces-297	96	5	annual	annual	ADJ
alces-297	96	6	calf	calf	NOUN
alces-297	96	7	predation	predation	NOUN
alces-297	96	8	rate	rate	NOUN
alces-297	96	9	by	by	ADP
alces-297	96	10	grizzly	grizzly	NOUN
alces-297	96	11	and	and	CCONJ
alces-297	96	12	black	black	ADJ
alces-297	96	13	bears	bear	NOUN
alces-297	96	14	of	of	ADP
alces-297	96	15	35	35	NUM
alces-297	96	16	%	%	NOUN
alces-297	96	17	,	,	PUNCT
alces-297	96	18	that	that	DET
alces-297	96	19	predation	predation	NOUN
alces-297	96	20	was	be	AUX
alces-297	96	21	density	density	NOUN
alces-297	96	22	independent	independent	ADJ
alces-297	96	23	,	,	PUNCT
alces-297	96	24	and	and	CCONJ
alces-297	96	25	that	that	SCONJ
alces-297	96	26	these	these	DET
alces-297	96	27	kill	kill	NOUN
alces-297	96	28	rates	rate	NOUN
alces-297	96	29	were	be	AUX
alces-297	96	30	additive	additive	ADJ
alces-297	96	31	to	to	ADP
alces-297	96	32	other	other	ADJ
alces-297	96	33	forms	form	NOUN
alces-297	96	34	of	of	ADP
alces-297	96	35	mortality	mortality	NOUN
alces-297	96	36	.	.	PUNCT
alces-297	97	1	adult	adult	NOUN
alces-297	97	2	grizzly	grizzly	NOUN
alces-297	97	3	bears	bear	NOUN
alces-297	97	4	have	have	AUX
alces-297	97	5	been	be	AUX
alces-297	97	6	reported	report	VERB
alces-297	97	7	to	to	PART
alces-297	97	8	kill	kill	VERB
alces-297	97	9	an	an	DET
alces-297	97	10	average	average	NOUN
alces-297	97	11	of	of	ADP
alces-297	97	12	0.5–2.2	0.5–2.2	X
alces-297	97	13	adult	adult	NOUN
alces-297	97	14	moose	moose	NOUN
alces-297	97	15	annually	annually	ADV
alces-297	97	16	(	(	PUNCT
alces-297	97	17	dahle	dahle	PROPN
alces-297	97	18	et	et	PROPN
alces-297	97	19	al	al	PROPN
alces-297	97	20	.	.	PROPN
alces-297	97	21	2013	2013	NUM
alces-297	97	22	)	)	PUNCT
alces-297	97	23	,	,	PUNCT
alces-297	97	24	while	while	SCONJ
alces-297	97	25	black	black	ADJ
alces-297	97	26	bears	bear	NOUN
alces-297	97	27	rarely	rarely	ADV
alces-297	97	28	kill	kill	VERB
alces-297	97	29	adult	adult	NOUN
alces-297	97	30	moose	moose	NOUN
alces-297	97	31	(	(	PUNCT
alces-297	97	32	ballad	ballad	ADJ
alces-297	97	33	1992	1992	NUM
alces-297	97	34	)	)	PUNCT
alces-297	97	35	.	.	PUNCT
alces-297	98	1	i	i	PRON
alces-297	98	2	assumed	assume	VERB
alces-297	98	3	that	that	SCONJ
alces-297	98	4	the	the	DET
alces-297	98	5	modelled	model	VERB
alces-297	98	6	2	2	NUM
alces-297	98	7	%	%	NOUN
alces-297	98	8	summer	summer	NOUN
alces-297	98	9	mortality	mortality	NOUN
alces-297	98	10	rate	rate	NOUN
alces-297	98	11	for	for	ADP
alces-297	98	12	adults	adult	NOUN
alces-297	98	13	included	include	VERB
alces-297	98	14	kills	kill	NOUN
alces-297	98	15	by	by	ADP
alces-297	98	16	bears	bear	NOUN
alces-297	98	17	.	.	PUNCT
alces-297	99	1	i	i	PRON
alces-297	99	2	did	do	AUX
alces-297	99	3	not	not	PART
alces-297	99	4	consider	consider	VERB
alces-297	99	5	the	the	DET
alces-297	99	6	effects	effect	NOUN
alces-297	99	7	of	of	ADP
alces-297	99	8	bear	bear	NOUN
alces-297	99	9	hunting	hunting	NOUN
alces-297	99	10	on	on	ADP
alces-297	99	11	moose	moose	NOUN
alces-297	99	12	population	population	NOUN
alces-297	99	13	dynamics	dynamic	NOUN
alces-297	99	14	as	as	SCONJ
alces-297	99	15	it	it	PRON
alces-297	99	16	would	would	AUX
alces-297	99	17	have	have	AUX
alces-297	99	18	made	make	VERB
alces-297	99	19	the	the	DET
alces-297	99	20	model	model	NOUN
alces-297	99	21	more	more	ADV
alces-297	99	22	complex	complex	ADJ
alces-297	99	23	and	and	CCONJ
alces-297	99	24	further	far	ADV
alces-297	99	25	increased	increase	VERB
alces-297	99	26	uncertainty	uncertainty	NOUN
alces-297	99	27	in	in	ADP
alces-297	99	28	the	the	DET
alces-297	99	29	model	model	NOUN
alces-297	99	30	projections	projection	NOUN
alces-297	99	31	.	.	PUNCT
alces-297	100	1	model	model	NOUN
alces-297	100	2	evaluation	evaluation	NOUN
alces-297	100	3	i	i	PRON
alces-297	100	4	evaluated	evaluate	VERB
alces-297	100	5	the	the	DET
alces-297	100	6	model	model	NOUN
alces-297	100	7	by	by	ADP
alces-297	100	8	assessing	assess	VERB
alces-297	100	9	how	how	SCONJ
alces-297	100	10	well	well	ADV
alces-297	100	11	the	the	DET
alces-297	100	12	modelled	model	VERB
alces-297	100	13	moose	moose	NOUN
alces-297	100	14	and	and	CCONJ
alces-297	100	15	wolf	wolf	NOUN
alces-297	100	16	parameters	parameter	NOUN
alces-297	100	17	compared	compare	VERB
alces-297	100	18	to	to	ADP
alces-297	100	19	the	the	DET
alces-297	100	20	observed	observed	ADJ
alces-297	100	21	parameters	parameter	NOUN
alces-297	100	22	in	in	ADP
alces-297	100	23	the	the	DET
alces-297	100	24	pgsa	pgsa	NOUN
alces-297	100	25	when	when	SCONJ
alces-297	100	26	16	16	NUM
alces-297	100	27	%	%	NOUN
alces-297	100	28	bulls	bull	NOUN
alces-297	100	29	,	,	PUNCT
alces-297	100	30	2	2	NUM
alces-297	100	31	%	%	NOUN
alces-297	100	32	cows	cow	NOUN
alces-297	100	33	,	,	PUNCT
alces-297	100	34	and	and	CCONJ
alces-297	100	35	9	9	NUM
alces-297	100	36	%	%	NOUN
alces-297	100	37	calves	calf	NOUN
alces-297	100	38	were	be	AUX
alces-297	100	39	harvested	harvest	VERB
alces-297	100	40	annually	annually	ADV
alces-297	100	41	.	.	PUNCT
alces-297	101	1	model	model	NOUN
alces-297	101	2	fitting	fitting	ADJ
alces-297	101	3	was	be	AUX
alces-297	101	4	primarily	primarily	ADV
alces-297	101	5	achieved	achieve	VERB
alces-297	101	6	by	by	ADP
alces-297	101	7	adjusting	adjust	VERB
alces-297	101	8	the	the	DET
alces-297	101	9	wolf	wolf	NOUN
alces-297	101	10	removal	removal	NOUN
alces-297	101	11	rate	rate	NOUN
alces-297	101	12	until	until	SCONJ
alces-297	101	13	the	the	DET
alces-297	101	14	modelled	model	VERB
alces-297	101	15	parameters	parameter	NOUN
alces-297	101	16	closely	closely	ADV
alces-297	101	17	matched	match	VERB
alces-297	101	18	the	the	DET
alces-297	101	19	observed	observed	ADJ
alces-297	101	20	parameters	parameter	NOUN
alces-297	101	21	.	.	PUNCT
alces-297	102	1	i	i	PRON
alces-297	102	2	ran	run	VERB
alces-297	102	3	each	each	DET
alces-297	102	4	simulation	simulation	NOUN
alces-297	102	5	for	for	ADP
alces-297	102	6	100	100	NUM
alces-297	102	7	years	year	NOUN
alces-297	102	8	to	to	PART
alces-297	102	9	remove	remove	VERB
alces-297	102	10	transient	transient	ADJ
alces-297	102	11	predator	predator	NOUN
alces-297	102	12	-	-	PUNCT
alces-297	102	13	prey	prey	NOUN
alces-297	102	14	dynamics	dynamic	NOUN
alces-297	102	15	and	and	CCONJ
alces-297	102	16	to	to	PART
alces-297	102	17	ensure	ensure	VERB
alces-297	102	18	that	that	DET
alces-297	102	19	moose	moose	NOUN
alces-297	102	20	,	,	PUNCT
alces-297	102	21	wolf	wolf	NOUN
alces-297	102	22	,	,	PUNCT
alces-297	102	23	and	and	CCONJ
alces-297	102	24	harvest	harvest	NOUN
alces-297	102	25	densities	density	NOUN
alces-297	102	26	were	be	AUX
alces-297	102	27	stable	stable	ADJ
alces-297	102	28	.	.	PUNCT
alces-297	103	1	i	i	PRON
alces-297	103	2	also	also	ADV
alces-297	103	3	compared	compare	VERB
alces-297	103	4	modelled	model	VERB
alces-297	103	5	estimates	estimate	NOUN
alces-297	103	6	of	of	ADP
alces-297	103	7	moose	moose	NOUN
alces-297	103	8	calf	calf	NOUN
alces-297	103	9	mortality	mortality	NOUN
alces-297	103	10	rates	rate	NOUN
alces-297	103	11	and	and	CCONJ
alces-297	103	12	total	total	ADJ
alces-297	103	13	annual	annual	ADJ
alces-297	103	14	mortality	mortality	NOUN
alces-297	103	15	rates	rate	NOUN
alces-297	103	16	with	with	ADP
alces-297	103	17	those	those	PRON
alces-297	103	18	from	from	ADP
alces-297	103	19	field	field	NOUN
alces-297	103	20	studies	study	NOUN
alces-297	103	21	in	in	ADP
alces-297	103	22	alaska	alaska	PROPN
alces-297	103	23	and	and	CCONJ
alces-297	103	24	yukon	yukon	PROPN
alces-297	103	25	summarized	summarize	VERB
alces-297	103	26	by	by	ADP
alces-297	103	27	boertje	boertje	PROPN
alces-297	103	28	et	et	PROPN
alces-297	103	29	al	al	PROPN
alces-297	103	30	.	.	PUNCT
alces-297	103	31	(	(	PUNCT
alces-297	103	32	2009	2009	NUM
alces-297	103	33	)	)	PUNCT
alces-297	103	34	.	.	PUNCT
alces-297	104	1	additionally	additionally	ADV
alces-297	104	2	,	,	PUNCT
alces-297	104	3	i	i	PRON
alces-297	104	4	tested	test	VERB
alces-297	104	5	for	for	ADP
alces-297	104	6	consistency	consistency	NOUN
alces-297	104	7	between	between	ADP
alces-297	104	8	modelled	model	VERB
alces-297	104	9	estimates	estimate	NOUN
alces-297	104	10	of	of	ADP
alces-297	104	11	wolf	wolf	PROPN
alces-297	104	12	density	density	NOUN
alces-297	104	13	with	with	ADP
alces-297	104	14	low	low	ADJ
alces-297	104	15	removal	removal	NOUN
alces-297	104	16	rates	rate	NOUN
alces-297	104	17	(	(	PUNCT
alces-297	104	18	0–15	0–15	PROPN
alces-297	104	19	%	%	NOUN
alces-297	104	20	)	)	PUNCT
alces-297	104	21	and	and	CCONJ
alces-297	104	22	those	those	PRON
alces-297	104	23	generated	generate	VERB
alces-297	104	24	from	from	ADP
alces-297	104	25	a	a	DET
alces-297	104	26	prey	prey	NOUN
alces-297	104	27	biomass	biomass	NOUN
alces-297	104	28	regression	regression	NOUN
alces-297	104	29	model	model	NOUN
alces-297	104	30	,	,	PUNCT
alces-297	104	31	where	where	SCONJ
alces-297	104	32	wolf	wolf	PROPN
alces-297	104	33	densities	density	NOUN
alces-297	104	34	were	be	AUX
alces-297	104	35	estimated	estimate	VERB
alces-297	104	36	from	from	ADP
alces-297	104	37	an	an	DET
alces-297	104	38	ungulate	ungulate	ADJ
alces-297	104	39	biomass	biomass	NOUN
alces-297	104	40	index	index	NOUN
alces-297	104	41	at	at	ADP
alces-297	104	42	the	the	DET
alces-297	104	43	regional	regional	ADJ
alces-297	104	44	level	level	NOUN
alces-297	104	45	in	in	ADP
alces-297	104	46	bc	bc	PROPN
alces-297	104	47	(	(	PUNCT
alces-297	104	48	kuzyk	kuzyk	X
alces-297	104	49	and	and	CCONJ
alces-297	104	50	hatter	hatter	NOUN
alces-297	104	51	2014	2014	NUM
alces-297	104	52	)	)	PUNCT
alces-297	104	53	.	.	PUNCT
alces-297	105	1	as	as	SCONJ
alces-297	105	2	calf	calf	NOUN
alces-297	105	3	predation	predation	NOUN
alces-297	105	4	rates	rate	NOUN
alces-297	105	5	by	by	ADP
alces-297	105	6	bears	bear	NOUN
alces-297	105	7	were	be	AUX
alces-297	105	8	unknown	unknown	ADJ
alces-297	105	9	,	,	PUNCT
alces-297	105	10	but	but	CCONJ
alces-297	105	11	could	could	AUX
alces-297	105	12	be	be	AUX
alces-297	105	13	substantial	substantial	ADJ
alces-297	105	14	,	,	PUNCT
alces-297	105	15	i	i	PRON
alces-297	105	16	evaluated	evaluate	VERB
alces-297	105	17	the	the	DET
alces-297	105	18	sensitivity	sensitivity	NOUN
alces-297	105	19	of	of	ADP
alces-297	105	20	different	different	ADJ
alces-297	105	21	predation	predation	NOUN
alces-297	105	22	rates	rate	NOUN
alces-297	105	23	on	on	ADP
alces-297	105	24	the	the	DET
alces-297	105	25	modelled	model	VERB
alces-297	105	26	parameters	parameter	NOUN
alces-297	105	27	.	.	PUNCT
alces-297	106	1	i	i	PRON
alces-297	106	2	considered	consider	VERB
alces-297	106	3	bear	bear	NOUN
alces-297	106	4	predation	predation	NOUN
alces-297	106	5	rates	rate	NOUN
alces-297	106	6	ranging	range	VERB
alces-297	106	7	from	from	ADP
alces-297	106	8	10	10	NUM
alces-297	106	9	to	to	PART
alces-297	106	10	50	50	NUM
alces-297	106	11	%	%	NOUN
alces-297	106	12	in	in	ADP
alces-297	106	13	increments	increment	NOUN
alces-297	106	14	of	of	ADP
alces-297	106	15	5	5	NUM
alces-297	106	16	%	%	NOUN
alces-297	106	17	.	.	PUNCT
alces-297	107	1	for	for	ADP
alces-297	107	2	each	each	DET
alces-297	107	3	predation	predation	NOUN
alces-297	107	4	rate	rate	NOUN
alces-297	107	5	,	,	PUNCT
alces-297	107	6	i	i	PRON
alces-297	107	7	used	use	VERB
alces-297	107	8	the	the	DET
alces-297	107	9	optimization	optimization	NOUN
alces-297	107	10	tool	tool	NOUN
alces-297	107	11	solver	solver	VERB
alces-297	107	12	in	in	ADP
alces-297	107	13	microsoft	microsoft	PROPN
alces-297	107	14	excel	excel	PROPN
alces-297	107	15	(	(	PUNCT
alces-297	107	16	microsoft	microsoft	PROPN
alces-297	107	17	,	,	PUNCT
alces-297	107	18	redmond	redmond	PROPN
alces-297	107	19	,	,	PUNCT
alces-297	107	20	washington	washington	PROPN
alces-297	107	21	,	,	PUNCT
alces-297	107	22	usa	usa	PROPN
alces-297	107	23	)	)	PUNCT
alces-297	107	24	to	to	PART
alces-297	107	25	minimize	minimize	VERB
alces-297	107	26	the	the	DET
alces-297	107	27	sums	sum	NOUN
alces-297	107	28	of	of	ADP
alces-297	107	29	squared	square	VERB
alces-297	107	30	differences	difference	NOUN
alces-297	107	31	between	between	ADP
alces-297	107	32	the	the	DET
alces-297	107	33	observed	observe	VERB
alces-297	107	34	and	and	CCONJ
alces-297	107	35	modelled	model	VERB
alces-297	107	36	parameters	parameter	NOUN
alces-297	107	37	(	(	PUNCT
alces-297	107	38	hilborn	hilborn	ADJ
alces-297	107	39	and	and	CCONJ
alces-297	107	40	mangel	mangel	NOUN
alces-297	107	41	1997	1997	NUM
alces-297	107	42	)	)	PUNCT
alces-297	107	43	by	by	ADP
alces-297	107	44	iteratively	iteratively	ADV
alces-297	107	45	changing	change	VERB
alces-297	107	46	the	the	DET
alces-297	107	47	wolf	wolf	NOUN
alces-297	107	48	removal	removal	NOUN
alces-297	107	49	rate	rate	NOUN
alces-297	107	50	.	.	PUNCT
alces-297	108	1	the	the	DET
alces-297	108	2	adjusted	adjust	VERB
alces-297	108	3	parameters	parameter	NOUN
alces-297	108	4	included	include	VERB
alces-297	108	5	moose	moose	NOUN
alces-297	108	6	density	density	NOUN
alces-297	108	7	,	,	PUNCT
alces-297	108	8	wolf	wolf	PROPN
alces-297	108	9	density	density	NOUN
alces-297	108	10	,	,	PUNCT
alces-297	108	11	moose	moose	NOUN
alces-297	108	12	harvest	harvest	NOUN
alces-297	108	13	density	density	NOUN
alces-297	108	14	,	,	PUNCT
alces-297	108	15	and	and	CCONJ
alces-297	108	16	moose	moose	NOUN
alces-297	108	17	sex	sex	NOUN
alces-297	108	18	:	:	PUNCT
alces-297	108	19	age	age	NOUN
alces-297	108	20	ratios	ratio	NOUN
alces-297	108	21	.	.	PUNCT
alces-297	109	1	sustained	sustain	VERB
alces-297	109	2	yield	yield	NOUN
alces-297	109	3	curve	curve	NOUN
alces-297	109	4	sys	sys	NOUN
alces-297	109	5	for	for	ADP
alces-297	109	6	moose	moose	NOUN
alces-297	109	7	in	in	ADP
alces-297	109	8	the	the	DET
alces-297	109	9	pgsa	pgsa	NOUN
alces-297	109	10	were	be	AUX
alces-297	109	11	simulated	simulate	VERB
alces-297	109	12	by	by	ADP
alces-297	109	13	increasing	increase	VERB
alces-297	109	14	annual	annual	ADJ
alces-297	109	15	wolf	wolf	NOUN
alces-297	109	16	removal	removal	NOUN
alces-297	109	17	rates	rate	NOUN
alces-297	109	18	from	from	ADP
alces-297	109	19	0	0	NUM
alces-297	109	20	to	to	PART
alces-297	109	21	40	40	NUM
alces-297	109	22	%	%	NOUN
alces-297	109	23	in	in	ADP
alces-297	109	24	increments	increment	NOUN
alces-297	109	25	of	of	ADP
alces-297	109	26	1	1	NUM
alces-297	109	27	%	%	NOUN
alces-297	109	28	while	while	SCONJ
alces-297	109	29	maintaining	maintain	VERB
alces-297	109	30	constant	constant	ADJ
alces-297	109	31	moose	moose	NOUN
alces-297	109	32	harvest	harvest	NOUN
alces-297	109	33	rates	rate	NOUN
alces-297	109	34	for	for	ADP
alces-297	109	35	bulls	bull	NOUN
alces-297	109	36	(	(	PUNCT
alces-297	109	37	16	16	NUM
alces-297	109	38	%	%	NOUN
alces-297	109	39	)	)	PUNCT
alces-297	109	40	,	,	PUNCT
alces-297	109	41	cows	cow	NOUN
alces-297	109	42	(	(	PUNCT
alces-297	109	43	2	2	NUM
alces-297	109	44	%	%	NOUN
alces-297	109	45	)	)	PUNCT
alces-297	109	46	,	,	PUNCT
alces-297	109	47	and	and	CCONJ
alces-297	109	48	calves	calf	NOUN
alces-297	109	49	(	(	PUNCT
alces-297	109	50	9	9	NUM
alces-297	109	51	%	%	NOUN
alces-297	109	52	)	)	PUNCT
alces-297	109	53	.	.	PUNCT
alces-297	110	1	the	the	DET
alces-297	110	2	sy	sy	PROPN
alces-297	110	3	was	be	AUX
alces-297	110	4	the	the	DET
alces-297	110	5	moose	moose	NOUN
alces-297	110	6	harvest	harvest	NOUN
alces-297	110	7	in	in	ADP
alces-297	110	8	the	the	DET
alces-297	110	9	100th	100th	ADJ
alces-297	110	10	year	year	NOUN
alces-297	110	11	of	of	ADP
alces-297	110	12	the	the	DET
alces-297	110	13	simulation	simulation	NOUN
alces-297	110	14	.	.	PUNCT
alces-297	111	1	sy	sy	PROPN
alces-297	111	2	curves	curve	NOUN
alces-297	111	3	were	be	AUX
alces-297	111	4	generated	generate	VERB
alces-297	111	5	for	for	ADP
alces-297	111	6	harvest	harvest	NOUN
alces-297	111	7	density	density	NOUN
alces-297	111	8	and	and	CCONJ
alces-297	111	9	wolf	wolf	NOUN
alces-297	111	10	removal	removal	NOUN
alces-297	111	11	rates	rate	NOUN
alces-297	111	12	,	,	PUNCT
alces-297	111	13	harvest	harvest	NOUN
alces-297	111	14	density	density	NOUN
alces-297	111	15	and	and	CCONJ
alces-297	111	16	wolf	wolf	PROPN
alces-297	111	17	density	density	NOUN
alces-297	111	18	,	,	PUNCT
alces-297	111	19	and	and	CCONJ
alces-297	111	20	harvest	harvest	NOUN
alces-297	111	21	density	density	NOUN
alces-297	111	22	and	and	CCONJ
alces-297	111	23	moose	moose	NOUN
alces-297	111	24	density	density	NOUN
alces-297	111	25	.	.	PUNCT
alces-297	112	1	modelling	model	VERB
alces-297	112	2	sustained	sustained	ADJ
alces-297	112	3	yields	yield	NOUN
alces-297	112	4	for	for	ADP
alces-297	112	5	moose	moose	NOUN
alces-297	112	6	–	–	PUNCT
alces-297	112	7	hatter	hatter	NOUN
alces-297	112	8	.	.	PUNCT
alces-297	113	1	alces	alce	NOUN
alces-297	113	2	vol	vol	NOUN
alces-297	113	3	.	.	PROPN
alces-297	113	4	57	57	NUM
alces-297	113	5	,	,	PUNCT
alces-297	113	6	2021	2021	NUM
alces-297	113	7	118	118	NUM
alces-297	114	1	i	i	PRON
alces-297	114	2	evaluated	evaluate	VERB
alces-297	114	3	the	the	DET
alces-297	114	4	modelled	model	VERB
alces-297	114	5	yield	yield	NOUN
alces-297	114	6	-	-	PUNCT
alces-297	114	7	density	density	NOUN
alces-297	114	8	curve	curve	NOUN
alces-297	114	9	by	by	ADP
alces-297	114	10	comparing	compare	VERB
alces-297	114	11	it	it	PRON
alces-297	114	12	to	to	ADP
alces-297	114	13	the	the	DET
alces-297	114	14	yield	yield	NOUN
alces-297	114	15	-	-	PUNCT
alces-297	114	16	density	density	NOUN
alces-297	114	17	curve	curve	NOUN
alces-297	114	18	developed	develop	VERB
alces-297	114	19	by	by	ADP
alces-297	114	20	hatter	hatter	NOUN
alces-297	114	21	(	(	PUNCT
alces-297	114	22	1999	1999	NUM
alces-297	114	23	)	)	PUNCT
alces-297	114	24	for	for	ADP
alces-297	114	25	stable	stable	ADJ
alces-297	114	26	moose	moose	NOUN
alces-297	114	27	populations	population	NOUN
alces-297	114	28	in	in	ADP
alces-297	114	29	north	north	NOUN
alces-297	114	30	-	-	PUNCT
alces-297	114	31	central	central	ADJ
alces-297	114	32	bc	bc	PROPN
alces-297	114	33	.	.	PUNCT
alces-297	115	1	optimal	optimal	ADJ
alces-297	115	2	harvests	harvest	NOUN
alces-297	115	3	optimal	optimal	ADJ
alces-297	115	4	harvests	harvest	NOUN
alces-297	115	5	were	be	AUX
alces-297	115	6	determined	determine	VERB
alces-297	115	7	by	by	ADP
alces-297	115	8	iteratively	iteratively	ADV
alces-297	115	9	changing	change	VERB
alces-297	115	10	the	the	DET
alces-297	115	11	harvest	harvest	NOUN
alces-297	115	12	rate	rate	NOUN
alces-297	115	13	for	for	ADP
alces-297	115	14	bulls	bull	NOUN
alces-297	115	15	,	,	PUNCT
alces-297	115	16	cows	cow	NOUN
alces-297	115	17	,	,	PUNCT
alces-297	115	18	and	and	CCONJ
alces-297	115	19	calves	calf	NOUN
alces-297	115	20	with	with	ADP
alces-297	115	21	solver	solver	NOUN
alces-297	115	22	until	until	SCONJ
alces-297	115	23	the	the	DET
alces-297	115	24	maximum	maximum	ADJ
alces-297	115	25	total	total	ADJ
alces-297	115	26	harvest	harvest	NOUN
alces-297	115	27	was	be	AUX
alces-297	115	28	achieved	achieve	VERB
alces-297	115	29	,	,	PUNCT
alces-297	115	30	and	and	CCONJ
alces-297	115	31	moose	moose	NOUN
alces-297	115	32	and	and	CCONJ
alces-297	115	33	wolf	wolf	NOUN
alces-297	115	34	populations	population	NOUN
alces-297	115	35	were	be	AUX
alces-297	115	36	stable	stable	ADJ
alces-297	115	37	.	.	PUNCT
alces-297	116	1	a	a	DET
alces-297	116	2	similar	similar	ADJ
alces-297	116	3	procedure	procedure	NOUN
alces-297	116	4	was	be	AUX
alces-297	116	5	used	use	VERB
alces-297	116	6	for	for	ADP
alces-297	116	7	optimizing	optimize	VERB
alces-297	116	8	bull	bull	NOUN
alces-297	116	9	and	and	CCONJ
alces-297	116	10	calf	calf	NOUN
alces-297	116	11	harvests	harvest	NOUN
alces-297	116	12	with	with	ADP
alces-297	116	13	cows	cow	NOUN
alces-297	116	14	harvested	harvest	VERB
alces-297	116	15	at	at	ADP
alces-297	116	16	2	2	NUM
alces-297	116	17	%	%	NOUN
alces-297	116	18	,	,	PUNCT
alces-297	116	19	and	and	CCONJ
alces-297	116	20	for	for	ADP
alces-297	116	21	bull	bull	NOUN
alces-297	116	22	harvests	harvest	NOUN
alces-297	116	23	with	with	ADP
alces-297	116	24	cows	cow	NOUN
alces-297	116	25	and	and	CCONJ
alces-297	116	26	calves	calf	NOUN
alces-297	116	27	unharvested	unharvested	ADJ
alces-297	116	28	.	.	PUNCT
alces-297	117	1	crête	crête	PROPN
alces-297	117	2	et	et	PROPN
alces-297	117	3	al	al	PROPN
alces-297	117	4	.	.	PROPN
alces-297	117	5	(	(	PUNCT
alces-297	117	6	1981	1981	NUM
alces-297	117	7	)	)	PUNCT
alces-297	117	8	recommended	recommend	VERB
alces-297	117	9	that	that	SCONJ
alces-297	117	10	in	in	ADP
alces-297	117	11	order	order	NOUN
alces-297	117	12	to	to	PART
alces-297	117	13	optimize	optimize	VERB
alces-297	117	14	moose	moose	NOUN
alces-297	117	15	harvest	harvest	NOUN
alces-297	117	16	in	in	ADP
alces-297	117	17	south	south	ADJ
alces-297	117	18	-	-	PUNCT
alces-297	117	19	western	western	ADJ
alces-297	117	20	quebec	quebec	PROPN
alces-297	117	21	,	,	PUNCT
alces-297	117	22	at	at	ADV
alces-297	117	23	least	least	ADV
alces-297	117	24	40	40	NUM
alces-297	117	25	%	%	NOUN
alces-297	117	26	bulls	bull	NOUN
alces-297	117	27	should	should	AUX
alces-297	117	28	be	be	AUX
alces-297	117	29	retained	retain	VERB
alces-297	117	30	among	among	ADP
alces-297	117	31	adults	adult	NOUN
alces-297	117	32	to	to	PART
alces-297	117	33	ensure	ensure	VERB
alces-297	117	34	that	that	SCONJ
alces-297	117	35	sex	sex	NOUN
alces-297	117	36	ratio	ratio	NOUN
alces-297	117	37	-	-	PUNCT
alces-297	117	38	dependent	dependent	ADJ
alces-297	117	39	fertilization	fertilization	NOUN
alces-297	117	40	was	be	AUX
alces-297	117	41	not	not	PART
alces-297	117	42	adversely	adversely	ADV
alces-297	117	43	affected	affect	VERB
alces-297	117	44	.	.	PUNCT
alces-297	118	1	the	the	DET
alces-297	118	2	provincial	provincial	ADJ
alces-297	118	3	moose	moose	NOUN
alces-297	118	4	harvest	harvest	NOUN
alces-297	118	5	management	management	NOUN
alces-297	118	6	procedure	procedure	NOUN
alces-297	118	7	for	for	ADP
alces-297	118	8	bc	bc	PROPN
alces-297	118	9	(	(	PUNCT
alces-297	118	10	flnro	flnro	ADJ
alces-297	118	11	2013	2013	NUM
alces-297	118	12	)	)	PUNCT
alces-297	118	13	states	state	VERB
alces-297	118	14	that	that	SCONJ
alces-297	118	15	the	the	DET
alces-297	118	16	lower	low	ADJ
alces-297	118	17	range	range	NOUN
alces-297	118	18	of	of	ADP
alces-297	118	19	the	the	DET
alces-297	118	20	post	post	ADJ
alces-297	118	21	-	-	ADJ
alces-297	118	22	hunt	hunt	ADJ
alces-297	118	23	adult	adult	NOUN
alces-297	118	24	sex	sex	NOUN
alces-297	118	25	ratio	ratio	NOUN
alces-297	118	26	should	should	AUX
alces-297	118	27	not	not	PART
alces-297	118	28	fall	fall	VERB
alces-297	118	29	below	below	ADP
alces-297	118	30	30	30	NUM
alces-297	118	31	bulls:100	bulls:100	NOUN
alces-297	118	32	cows	cow	NOUN
alces-297	118	33	,	,	PUNCT
alces-297	118	34	or	or	CCONJ
alces-297	118	35	50	50	NUM
alces-297	118	36	bulls:100	bulls:100	NOUN
alces-297	118	37	cows	cow	NOUN
alces-297	118	38	in	in	ADP
alces-297	118	39	low	low	ADJ
alces-297	118	40	density	density	NOUN
alces-297	118	41	(	(	PUNCT
alces-297	118	42	≤	≤	ADV
alces-297	118	43	200/1000	200/1000	NUM
alces-297	118	44	km2	km2	NOUN
alces-297	118	45	)	)	PUNCT
alces-297	118	46	moose	moose	NOUN
alces-297	118	47	populations	population	NOUN
alces-297	118	48	.	.	PUNCT
alces-297	119	1	therefore	therefore	ADV
alces-297	119	2	,	,	PUNCT
alces-297	119	3	i	i	PRON
alces-297	119	4	added	add	VERB
alces-297	119	5	a	a	DET
alces-297	119	6	sex	sex	NOUN
alces-297	119	7	ratio	ratio	NOUN
alces-297	119	8	constraint	constraint	NOUN
alces-297	119	9	when	when	SCONJ
alces-297	119	10	optimizing	optimize	VERB
alces-297	119	11	moose	moose	NOUN
alces-297	119	12	harvests	harvest	NOUN
alces-297	119	13	with	with	ADP
alces-297	119	14	solver	solver	NOUN
alces-297	119	15	to	to	PART
alces-297	119	16	ensure	ensure	VERB
alces-297	119	17	the	the	DET
alces-297	119	18	adult	adult	NOUN
alces-297	119	19	sex	sex	NOUN
alces-297	119	20	ratio	ratio	NOUN
alces-297	119	21	was	be	AUX
alces-297	119	22	maintained	maintain	VERB
alces-297	119	23	at	at	ADP
alces-297	119	24	either	either	CCONJ
alces-297	119	25	≥	≥	NUM
alces-297	119	26	30	30	NUM
alces-297	119	27	bulls:100	bulls:100	NOUN
alces-297	119	28	cows	cow	NOUN
alces-297	119	29	or	or	CCONJ
alces-297	119	30	≥	≥	NUM
alces-297	119	31	50	50	NUM
alces-297	119	32	bulls:100	bulls:100	PROPN
alces-297	119	33	cows	cow	NOUN
alces-297	119	34	depending	depend	VERB
alces-297	119	35	on	on	ADP
alces-297	119	36	moose	moose	NOUN
alces-297	119	37	density	density	NOUN
alces-297	119	38	.	.	PUNCT
alces-297	120	1	results	result	VERB
alces-297	120	2	the	the	DET
alces-297	120	3	modelled	model	VERB
alces-297	120	4	moose	moose	NOUN
alces-297	120	5	population	population	NOUN
alces-297	120	6	,	,	PUNCT
alces-297	120	7	in	in	ADP
alces-297	120	8	the	the	DET
alces-297	120	9	absence	absence	NOUN
alces-297	120	10	of	of	ADP
alces-297	120	11	hunting	hunting	NOUN
alces-297	120	12	or	or	CCONJ
alces-297	120	13	predation	predation	NOUN
alces-297	120	14	,	,	PUNCT
alces-297	120	15	stabilized	stabilize	VERB
alces-297	120	16	at	at	ADP
alces-297	120	17	2000	2000	NUM
alces-297	120	18	moose/1000	moose/1000	NOUN
alces-297	120	19	km2	km2	NOUN
alces-297	120	20	,	,	PUNCT
alces-297	120	21	with	with	ADP
alces-297	120	22	100	100	NUM
alces-297	120	23	bulls/100	bulls/100	PROPN
alces-297	120	24	cows	cow	NOUN
alces-297	120	25	and	and	CCONJ
alces-297	120	26	32	32	NUM
alces-297	120	27	calves/100	calves/100	NOUN
alces-297	120	28	cows	cow	NOUN
alces-297	120	29	(	(	PUNCT
alces-297	120	30	table	table	NOUN
alces-297	120	31	1	1	NUM
alces-297	120	32	)	)	PUNCT
alces-297	120	33	.	.	PUNCT
alces-297	121	1	the	the	DET
alces-297	121	2	finite	finite	ADJ
alces-297	121	3	rate	rate	NOUN
alces-297	121	4	of	of	ADP
alces-297	121	5	increase	increase	NOUN
alces-297	121	6	below	below	ADP
alces-297	121	7	population	population	NOUN
alces-297	121	8	levels	level	NOUN
alces-297	121	9	where	where	SCONJ
alces-297	121	10	density	density	NOUN
alces-297	121	11	dependent	dependent	ADJ
alces-297	121	12	effects	effect	NOUN
alces-297	121	13	became	become	VERB
alces-297	121	14	operative	operative	ADJ
alces-297	121	15	was	be	AUX
alces-297	121	16	λ	λ	X
alces-297	121	17	~	~	PUNCT
alces-297	121	18	1.3	1.3	NUM
alces-297	121	19	.	.	PUNCT
alces-297	122	1	the	the	DET
alces-297	122	2	stable	stable	ADJ
alces-297	122	3	moose	moose	NOUN
alces-297	122	4	density	density	NOUN
alces-297	122	5	was	be	AUX
alces-297	122	6	9	9	NUM
alces-297	122	7	%	%	NOUN
alces-297	122	8	lower	low	ADJ
alces-297	122	9	with	with	ADP
alces-297	122	10	bears	bear	NOUN
alces-297	122	11	only	only	ADV
alces-297	122	12	(	(	PUNCT
alces-297	122	13	wolves	wolf	NOUN
alces-297	122	14	and	and	CCONJ
alces-297	122	15	hunting	hunt	VERB
alces-297	122	16	absent	absent	ADJ
alces-297	122	17	)	)	PUNCT
alces-297	122	18	,	,	PUNCT
alces-297	122	19	and	and	CCONJ
alces-297	122	20	72	72	NUM
alces-297	122	21	%	%	NOUN
alces-297	122	22	lower	low	ADJ
alces-297	122	23	with	with	ADP
alces-297	122	24	wolves	wolf	NOUN
alces-297	122	25	only	only	ADV
alces-297	122	26	(	(	PUNCT
alces-297	122	27	bears	bear	NOUN
alces-297	122	28	and	and	CCONJ
alces-297	122	29	hunting	hunt	VERB
alces-297	122	30	absent	absent	ADJ
alces-297	122	31	)	)	PUNCT
alces-297	122	32	.	.	PUNCT
alces-297	123	1	with	with	ADP
alces-297	123	2	wolves	wolf	NOUN
alces-297	123	3	and	and	CCONJ
alces-297	123	4	bears	bear	NOUN
alces-297	123	5	(	(	PUNCT
alces-297	123	6	no	no	DET
alces-297	123	7	hunting	hunting	NOUN
alces-297	123	8	)	)	PUNCT
alces-297	123	9	,	,	PUNCT
alces-297	123	10	the	the	DET
alces-297	123	11	stabilizing	stabilize	VERB
alces-297	123	12	density	density	NOUN
alces-297	123	13	was	be	AUX
alces-297	123	14	almost	almost	ADV
alces-297	123	15	90	90	NUM
alces-297	123	16	%	%	NOUN
alces-297	123	17	lower	low	ADJ
alces-297	123	18	,	,	PUNCT
alces-297	123	19	and	and	CCONJ
alces-297	123	20	consisted	consist	VERB
alces-297	123	21	of	of	ADP
alces-297	123	22	87	87	NUM
alces-297	123	23	bulls/100	bulls/100	PROPN
alces-297	123	24	cows	cow	NOUN
alces-297	123	25	and	and	CCONJ
alces-297	123	26	35	35	NUM
alces-297	123	27	calves/100	calves/100	NOUN
alces-297	123	28	cows	cow	NOUN
alces-297	123	29	.	.	PUNCT
alces-297	124	1	the	the	DET
alces-297	124	2	estimated	estimate	VERB
alces-297	124	3	parameters	parameter	NOUN
alces-297	124	4	from	from	ADP
alces-297	124	5	the	the	DET
alces-297	124	6	predatorprey	predatorprey	PROPN
alces-297	124	7	model	model	NOUN
alces-297	124	8	compared	compare	VERB
alces-297	124	9	favourably	favourably	ADV
alces-297	124	10	with	with	ADP
alces-297	124	11	the	the	DET
alces-297	124	12	observed	observed	ADJ
alces-297	124	13	parameters	parameter	NOUN
alces-297	124	14	for	for	ADP
alces-297	124	15	the	the	DET
alces-297	124	16	pgsa	pgsa	NOUN
alces-297	124	17	during	during	ADP
alces-297	124	18	the	the	DET
alces-297	124	19	study	study	NOUN
alces-297	124	20	period	period	NOUN
alces-297	124	21	when	when	SCONJ
alces-297	124	22	the	the	DET
alces-297	124	23	wolf	wolf	PROPN
alces-297	124	24	removal	removal	NOUN
alces-297	124	25	rate	rate	NOUN
alces-297	124	26	was	be	AUX
alces-297	124	27	31	31	NUM
alces-297	124	28	%	%	NOUN
alces-297	124	29	.	.	PUNCT
alces-297	125	1	both	both	CCONJ
alces-297	125	2	the	the	DET
alces-297	125	3	modelled	model	VERB
alces-297	125	4	and	and	CCONJ
alces-297	125	5	observed	observe	VERB
alces-297	125	6	wolf	wolf	PROPN
alces-297	125	7	densities	density	NOUN
alces-297	125	8	were	be	AUX
alces-297	125	9	12.5	12.5	NUM
alces-297	125	10	wolves/1000	wolves/1000	NOUN
alces-297	125	11	km2	km2	PROPN
alces-297	125	12	.	.	PUNCT
alces-297	126	1	the	the	DET
alces-297	126	2	modelled	model	VERB
alces-297	126	3	moose	moose	NOUN
alces-297	126	4	density	density	NOUN
alces-297	126	5	was	be	AUX
alces-297	126	6	1310/1000	1310/1000	NUM
alces-297	126	7	km2	km2	NOUN
alces-297	126	8	(	(	PUNCT
alces-297	126	9	observed	observe	VERB
alces-297	126	10	:	:	PUNCT
alces-297	126	11	1320/1000	1320/1000	NUM
alces-297	126	12	km2	km2	NOUN
alces-297	126	13	)	)	PUNCT
alces-297	126	14	and	and	CCONJ
alces-297	126	15	the	the	DET
alces-297	126	16	harvest	harvest	NOUN
alces-297	126	17	density	density	NOUN
alces-297	126	18	was	be	AUX
alces-297	126	19	100/1000	100/1000	NUM
alces-297	126	20	km2	km2	X
alces-297	126	21	(	(	PUNCT
alces-297	126	22	observed	observe	VERB
alces-297	126	23	:	:	PUNCT
alces-297	126	24	99/1000	99/1000	NUM
alces-297	126	25	km2	km2	NOUN
alces-297	126	26	)	)	PUNCT
alces-297	126	27	,	,	PUNCT
alces-297	126	28	or	or	CCONJ
alces-297	126	29	7	7	NUM
alces-297	126	30	%	%	NOUN
alces-297	126	31	of	of	ADP
alces-297	126	32	the	the	DET
alces-297	126	33	pre	pre	ADJ
alces-297	126	34	-	-	ADJ
alces-297	126	35	hunt	hunt	ADJ
alces-297	126	36	moose	moose	NOUN
alces-297	126	37	density	density	NOUN
alces-297	126	38	=	=	SYM
alces-297	126	39	99/	99/	NUM
alces-297	126	40	(	(	PUNCT
alces-297	126	41	99	99	NUM
alces-297	126	42	+	+	NOUN
alces-297	126	43	1320	1320	NUM
alces-297	126	44	)	)	PUNCT
alces-297	126	45	.	.	PUNCT
alces-297	127	1	the	the	DET
alces-297	127	2	modelled	model	VERB
alces-297	127	3	adult	adult	NOUN
alces-297	127	4	sex	sex	NOUN
alces-297	127	5	ratio	ratio	NOUN
alces-297	127	6	(	(	PUNCT
alces-297	127	7	40	40	NUM
alces-297	127	8	bulls:100	bulls:100	PROPN
alces-297	127	9	cows	cow	NOUN
alces-297	127	10	)	)	PUNCT
alces-297	127	11	was	be	AUX
alces-297	127	12	lower	low	ADJ
alces-297	127	13	than	than	ADP
alces-297	127	14	the	the	DET
alces-297	127	15	observed	observed	ADJ
alces-297	127	16	ratio	ratio	NOUN
alces-297	127	17	(	(	PUNCT
alces-297	127	18	46	46	NUM
alces-297	127	19	bulls:100	bulls:100	PROPN
alces-297	127	20	cows	cow	NOUN
alces-297	127	21	)	)	PUNCT
alces-297	127	22	,	,	PUNCT
alces-297	127	23	while	while	SCONJ
alces-297	127	24	calf	calf	NOUN
alces-297	127	25	:	:	PUNCT
alces-297	127	26	cow	cow	NOUN
alces-297	127	27	ratios	ratio	NOUN
alces-297	127	28	were	be	AUX
alces-297	127	29	similar	similar	ADJ
alces-297	127	30	(	(	PUNCT
alces-297	127	31	modelled	model	VERB
alces-297	127	32	=	=	SYM
alces-297	127	33	38	38	NUM
alces-297	127	34	calves:100	calves:100	NOUN
alces-297	127	35	cows	cow	NOUN
alces-297	127	36	,	,	PUNCT
alces-297	127	37	observed	observe	VERB
alces-297	127	38	=	=	PUNCT
alces-297	127	39	41	41	NUM
alces-297	127	40	calves:100	calves:100	NOUN
alces-297	127	41	cows	cow	NOUN
alces-297	127	42	)	)	PUNCT
alces-297	127	43	.	.	PUNCT
alces-297	128	1	the	the	DET
alces-297	128	2	modelled	model	VERB
alces-297	128	3	and	and	CCONJ
alces-297	128	4	observed	observed	ADJ
alces-297	128	5	moose	moose	NOUN
alces-297	128	6	:	:	PUNCT
alces-297	128	7	wolf	wolf	PROPN
alces-297	128	8	ratios	ratio	NOUN
alces-297	128	9	were	be	AUX
alces-297	128	10	the	the	DET
alces-297	128	11	same	same	ADJ
alces-297	128	12	(	(	PUNCT
alces-297	128	13	105:1	105:1	NUM
alces-297	128	14	)	)	PUNCT
alces-297	128	15	.	.	PUNCT
alces-297	129	1	modelled	model	VERB
alces-297	129	2	summer	summer	NOUN
alces-297	129	3	moose	moose	NOUN
alces-297	129	4	calf	calf	NOUN
alces-297	129	5	mortality	mortality	NOUN
alces-297	129	6	rates	rate	NOUN
alces-297	129	7	from	from	ADP
alces-297	129	8	wolves	wolf	NOUN
alces-297	129	9	and	and	CCONJ
alces-297	129	10	bears	bear	NOUN
alces-297	129	11	,	,	PUNCT
alces-297	129	12	as	as	ADV
alces-297	129	13	well	well	ADV
alces-297	129	14	as	as	ADP
alces-297	129	15	total	total	ADJ
alces-297	129	16	mortality	mortality	NOUN
alces-297	129	17	rates	rate	NOUN
alces-297	129	18	,	,	PUNCT
alces-297	129	19	fell	fall	VERB
alces-297	129	20	within	within	ADP
alces-297	129	21	the	the	DET
alces-297	129	22	range	range	NOUN
alces-297	129	23	reported	report	VERB
alces-297	129	24	from	from	ADP
alces-297	129	25	studies	study	NOUN
alces-297	129	26	in	in	ADP
alces-297	129	27	alaska	alaska	PROPN
alces-297	129	28	and	and	CCONJ
alces-297	129	29	yukon	yukon	PROPN
alces-297	129	30	(	(	PUNCT
alces-297	129	31	table	table	NOUN
alces-297	129	32	2a	2a	NUM
alces-297	129	33	)	)	PUNCT
alces-297	129	34	.	.	PUNCT
alces-297	130	1	the	the	DET
alces-297	130	2	combined	combine	VERB
alces-297	130	3	predation	predation	NOUN
alces-297	130	4	rates	rate	NOUN
alces-297	130	5	from	from	ADP
alces-297	130	6	wolves	wolf	NOUN
alces-297	130	7	table	table	NOUN
alces-297	130	8	1	1	NUM
alces-297	130	9	.	.	PUNCT
alces-297	130	10	comparison	comparison	NOUN
alces-297	130	11	of	of	ADP
alces-297	130	12	modelled	model	VERB
alces-297	130	13	moose	moose	NOUN
alces-297	130	14	density	density	NOUN
alces-297	130	15	,	,	PUNCT
alces-297	130	16	bull	bull	NOUN
alces-297	130	17	:	:	PUNCT
alces-297	130	18	cow	cow	NOUN
alces-297	130	19	ratios	ratio	NOUN
alces-297	130	20	and	and	CCONJ
alces-297	130	21	calf	calf	NOUN
alces-297	130	22	:	:	PUNCT
alces-297	130	23	cow	cow	NOUN
alces-297	130	24	ratios	ratio	NOUN
alces-297	130	25	for	for	ADP
alces-297	130	26	the	the	DET
alces-297	130	27	pgsa	pgsa	NOUN
alces-297	130	28	without	without	ADP
alces-297	130	29	hunting	hunting	NOUN
alces-297	130	30	or	or	CCONJ
alces-297	130	31	predation	predation	NOUN
alces-297	130	32	,	,	PUNCT
alces-297	130	33	with	with	ADP
alces-297	130	34	bear	bear	NOUN
alces-297	130	35	predation	predation	NOUN
alces-297	130	36	only	only	ADV
alces-297	130	37	,	,	PUNCT
alces-297	130	38	wolf	wolf	PROPN
alces-297	130	39	predation	predation	NOUN
alces-297	130	40	only	only	ADV
alces-297	130	41	,	,	PUNCT
alces-297	130	42	and	and	CCONJ
alces-297	130	43	wolf	wolf	PROPN
alces-297	130	44	and	and	CCONJ
alces-297	130	45	bear	bear	VERB
alces-297	130	46	predation	predation	NOUN
alces-297	130	47	,	,	PUNCT
alces-297	130	48	central	central	PROPN
alces-297	130	49	british	british	PROPN
alces-297	130	50	columbia	columbia	PROPN
alces-297	130	51	,	,	PUNCT
alces-297	130	52	canada	canada	PROPN
alces-297	130	53	.	.	PUNCT
alces-297	131	1	treatment	treatment	NOUN
alces-297	131	2	moose/1000	moose/1000	PROPN
alces-297	131	3	km2	km2	PROPN
alces-297	131	4	bulls:100	bulls:100	PROPN
alces-297	131	5	cows	cow	NOUN
alces-297	131	6	calves:100	calves:100	NOUN
alces-297	131	7	cows	cow	NOUN
alces-297	131	8	no	no	DET
alces-297	131	9	hunting	hunting	NOUN
alces-297	131	10	or	or	CCONJ
alces-297	131	11	predation	predation	NOUN
alces-297	131	12	2000	2000	NUM
alces-297	131	13	100	100	NUM
alces-297	131	14	32	32	NUM
alces-297	131	15	bear	bear	NOUN
alces-297	131	16	predation	predation	NOUN
alces-297	131	17	only	only	ADV
alces-297	131	18	1828	1828	NUM
alces-297	131	19	100	100	NUM
alces-297	131	20	22	22	NUM
alces-297	131	21	wolf	wolf	PROPN
alces-297	131	22	predation	predation	NOUN
alces-297	131	23	only	only	ADV
alces-297	131	24	567	567	NUM
alces-297	131	25	90	90	NUM
alces-297	131	26	58	58	NUM
alces-297	131	27	wolf	wolf	NOUN
alces-297	131	28	and	and	CCONJ
alces-297	131	29	bear	bear	VERB
alces-297	131	30	predation	predation	NOUN
alces-297	131	31	218	218	NUM
alces-297	131	32	87	87	NUM
alces-297	131	33	35	35	NUM
alces-297	131	34	alces	alce	NOUN
alces-297	131	35	vol	vol	NOUN
alces-297	131	36	.	.	PROPN
alces-297	132	1	57	57	NUM
alces-297	132	2	,	,	PUNCT
alces-297	132	3	2021	2021	NUM
alces-297	132	4	modelling	model	VERB
alces-297	132	5	sustained	sustain	VERB
alces-297	132	6	yields	yield	NOUN
alces-297	132	7	for	for	ADP
alces-297	132	8	moose	moose	NOUN
alces-297	132	9	–	–	PUNCT
alces-297	132	10	hatter	hatter	NOUN
alces-297	132	11	.	.	PUNCT
alces-297	133	1	119	119	NUM
alces-297	133	2	and	and	CCONJ
alces-297	133	3	bears	bear	NOUN
alces-297	133	4	,	,	PUNCT
alces-297	133	5	however	however	ADV
alces-297	133	6	,	,	PUNCT
alces-297	133	7	were	be	AUX
alces-297	133	8	slightly	slightly	ADV
alces-297	133	9	lower	low	ADJ
alces-297	133	10	.	.	PUNCT
alces-297	134	1	annual	annual	ADJ
alces-297	134	2	wolf	wolf	NOUN
alces-297	134	3	and	and	CCONJ
alces-297	134	4	bear	bear	VERB
alces-297	134	5	predation	predation	NOUN
alces-297	134	6	rates	rate	NOUN
alces-297	134	7	on	on	ADP
alces-297	134	8	the	the	DET
alces-297	134	9	post	post	ADJ
alces-297	134	10	-	-	ADJ
alces-297	134	11	calving	calving	ADJ
alces-297	134	12	population	population	NOUN
alces-297	134	13	,	,	PUNCT
alces-297	134	14	as	as	ADV
alces-297	134	15	well	well	ADV
alces-297	134	16	as	as	ADP
alces-297	134	17	the	the	DET
alces-297	134	18	total	total	ADJ
alces-297	134	19	mortality	mortality	NOUN
alces-297	134	20	rate	rate	NOUN
alces-297	134	21	,	,	PUNCT
alces-297	134	22	also	also	ADV
alces-297	134	23	fell	fall	VERB
alces-297	134	24	within	within	ADP
alces-297	134	25	the	the	DET
alces-297	134	26	range	range	NOUN
alces-297	134	27	from	from	ADP
alces-297	134	28	alaska	alaska	PROPN
alces-297	134	29	and	and	CCONJ
alces-297	134	30	yukon	yukon	PROPN
alces-297	134	31	(	(	PUNCT
alces-297	134	32	table	table	NOUN
alces-297	134	33	2b	2b	NOUN
alces-297	134	34	)	)	PUNCT
alces-297	134	35	.	.	PUNCT
alces-297	135	1	modelled	model	VERB
alces-297	135	2	estimates	estimate	NOUN
alces-297	135	3	of	of	ADP
alces-297	135	4	wolf	wolf	PROPN
alces-297	135	5	density	density	NOUN
alces-297	135	6	with	with	ADP
alces-297	135	7	wolf	wolf	PROPN
alces-297	135	8	removal	removal	NOUN
alces-297	135	9	rates	rate	NOUN
alces-297	135	10	≤	≤	ADV
alces-297	135	11	15	15	NUM
alces-297	135	12	%	%	NOUN
alces-297	135	13	(	(	PUNCT
alces-297	135	14	x̄	x̄	NOUN
alces-297	135	15	=	=	SYM
alces-297	135	16	7	7	NUM
alces-297	135	17	wolves/1000	wolves/1000	PROPN
alces-297	135	18	km2	km2	NOUN
alces-297	135	19	,	,	PUNCT
alces-297	135	20	range	range	NOUN
alces-297	135	21	=	=	SYM
alces-297	135	22	6–7	6–7	NUM
alces-297	135	23	)	)	PUNCT
alces-297	135	24	were	be	AUX
alces-297	135	25	similar	similar	ADJ
alces-297	135	26	to	to	ADP
alces-297	135	27	those	those	PRON
alces-297	135	28	from	from	ADP
alces-297	135	29	the	the	DET
alces-297	135	30	ungulate	ungulate	ADJ
alces-297	135	31	biomass	biomass	NOUN
alces-297	135	32	regression	regression	NOUN
alces-297	135	33	(	(	PUNCT
alces-297	135	34	x̄	x̄	NOUN
alces-297	135	35	=	=	SYM
alces-297	135	36	6	6	NUM
alces-297	135	37	wolves/1000	wolves/1000	PROPN
alces-297	135	38	km2	km2	NOUN
alces-297	135	39	,	,	PUNCT
alces-297	135	40	range	range	NOUN
alces-297	135	41	=	=	SYM
alces-297	135	42	5–8	5–8	NUM
alces-297	135	43	)	)	PUNCT
alces-297	135	44	.	.	PUNCT
alces-297	136	1	the	the	DET
alces-297	136	2	modelled	model	VERB
alces-297	136	3	parameters	parameter	NOUN
alces-297	136	4	were	be	AUX
alces-297	136	5	sensitive	sensitive	ADJ
alces-297	136	6	to	to	ADP
alces-297	136	7	different	different	ADJ
alces-297	136	8	bear	bear	NOUN
alces-297	136	9	predation	predation	NOUN
alces-297	136	10	rates	rate	NOUN
alces-297	136	11	on	on	ADP
alces-297	136	12	moose	moose	NOUN
alces-297	136	13	calves	calf	NOUN
alces-297	136	14	(	(	PUNCT
alces-297	136	15	table	table	NOUN
alces-297	136	16	3	3	NUM
alces-297	136	17	)	)	PUNCT
alces-297	136	18	.	.	PUNCT
alces-297	137	1	moose	moose	PROPN
alces-297	137	2	densities	density	NOUN
alces-297	137	3	,	,	PUNCT
alces-297	137	4	wolf	wolf	PROPN
alces-297	137	5	densities	density	NOUN
alces-297	137	6	,	,	PUNCT
alces-297	137	7	sex	sex	NOUN
alces-297	137	8	/	/	SYM
alces-297	137	9	age	age	NOUN
alces-297	137	10	ratios	ratio	NOUN
alces-297	137	11	,	,	PUNCT
alces-297	137	12	and	and	CCONJ
alces-297	137	13	moose	moose	NOUN
alces-297	137	14	harvests	harvest	NOUN
alces-297	137	15	declined	decline	VERB
alces-297	137	16	as	as	SCONJ
alces-297	137	17	bear	bear	NOUN
alces-297	137	18	predation	predation	NOUN
alces-297	137	19	rates	rate	NOUN
alces-297	137	20	increased	increase	VERB
alces-297	137	21	.	.	PUNCT
alces-297	138	1	wolf	wolf	PROPN
alces-297	138	2	removal	removal	NOUN
alces-297	138	3	rates	rate	NOUN
alces-297	138	4	and	and	CCONJ
alces-297	138	5	the	the	DET
alces-297	138	6	moose	moose	NOUN
alces-297	138	7	:	:	PUNCT
alces-297	138	8	wolf	wolf	PROPN
alces-297	138	9	ratio	ratio	NOUN
alces-297	138	10	increased	increase	VERB
alces-297	138	11	as	as	SCONJ
alces-297	138	12	bear	bear	NOUN
alces-297	138	13	predation	predation	NOUN
alces-297	138	14	rates	rate	NOUN
alces-297	138	15	increased	increase	VERB
alces-297	138	16	.	.	PUNCT
alces-297	139	1	sys	sys	PROPN
alces-297	139	2	varied	varied	ADJ
alces-297	139	3	from	from	ADP
alces-297	139	4	94	94	NUM
alces-297	139	5	moose/1000	moose/1000	NOUN
alces-297	139	6	km2	km2	PROPN
alces-297	139	7	with	with	ADP
alces-297	139	8	50	50	NUM
alces-297	139	9	%	%	NOUN
alces-297	139	10	bear	bear	NOUN
alces-297	139	11	predation	predation	NOUN
alces-297	139	12	to	to	ADP
alces-297	139	13	119	119	NUM
alces-297	139	14	moose/1000	moose/1000	NOUN
alces-297	139	15	km2	km2	PROPN
alces-297	139	16	with	with	ADP
alces-297	139	17	10	10	NUM
alces-297	139	18	%	%	NOUN
alces-297	139	19	bear	bear	NOUN
alces-297	139	20	predation	predation	NOUN
alces-297	139	21	.	.	PUNCT
alces-297	140	1	sys	sys	PROPN
alces-297	140	2	for	for	ADP
alces-297	140	3	moose	moose	NOUN
alces-297	140	4	displayed	display	VERB
alces-297	140	5	an	an	DET
alces-297	140	6	s	s	NOUN
alces-297	140	7	-	-	PUNCT
alces-297	140	8	shape	shape	NOUN
alces-297	140	9	curve	curve	NOUN
alces-297	140	10	with	with	ADP
alces-297	140	11	increasing	increase	VERB
alces-297	140	12	wolf	wolf	PROPN
alces-297	140	13	removal	removal	NOUN
alces-297	140	14	rates	rate	NOUN
alces-297	140	15	(	(	PUNCT
alces-297	140	16	fig	fig	NOUN
alces-297	140	17	.	.	PUNCT
alces-297	140	18	2a	2a	NUM
alces-297	140	19	)	)	PUNCT
alces-297	140	20	.	.	PUNCT
alces-297	141	1	sys	sys	PROPN
alces-297	141	2	gradually	gradually	ADV
alces-297	141	3	increased	increase	VERB
alces-297	141	4	when	when	SCONJ
alces-297	141	5	wolf	wolf	PROPN
alces-297	141	6	removal	removal	NOUN
alces-297	141	7	rates	rate	NOUN
alces-297	141	8	rose	rise	VERB
alces-297	141	9	from	from	ADP
alces-297	141	10	0	0	NUM
alces-297	141	11	to	to	PART
alces-297	141	12	15	15	NUM
alces-297	141	13	%	%	NOUN
alces-297	141	14	,	,	PUNCT
alces-297	141	15	while	while	SCONJ
alces-297	141	16	removal	removal	NOUN
alces-297	141	17	rates	rate	NOUN
alces-297	141	18	between	between	ADP
alces-297	141	19	27	27	NUM
alces-297	141	20	and	and	CCONJ
alces-297	141	21	32	32	NUM
alces-297	141	22	%	%	NOUN
alces-297	141	23	greatly	greatly	ADV
alces-297	141	24	increased	increase	VERB
alces-297	141	25	sys	sy	NOUN
alces-297	141	26	;	;	PUNCT
alces-297	141	27	removal	removal	NOUN
alces-297	141	28	rates	rate	NOUN
alces-297	141	29	>	>	X
alces-297	141	30	31	31	NUM
alces-297	141	31	%	%	NOUN
alces-297	141	32	only	only	ADV
alces-297	141	33	slightly	slightly	ADV
alces-297	141	34	increased	increase	VERB
alces-297	141	35	sys	sys	PROPN
alces-297	141	36	.	.	PUNCT
alces-297	142	1	wolves	wolf	NOUN
alces-297	142	2	sustained	sustain	VERB
alces-297	142	3	removal	removal	NOUN
alces-297	142	4	rates	rate	NOUN
alces-297	142	5	up	up	ADP
alces-297	142	6	to	to	PART
alces-297	142	7	31	31	NUM
alces-297	142	8	%	%	NOUN
alces-297	142	9	,	,	PUNCT
alces-297	142	10	declined	decline	VERB
alces-297	142	11	rapidly	rapidly	ADV
alces-297	142	12	when	when	SCONJ
alces-297	142	13	removals	removal	NOUN
alces-297	142	14	were	be	AUX
alces-297	142	15	31–38	31–38	NUM
alces-297	142	16	%	%	NOUN
alces-297	142	17	,	,	PUNCT
alces-297	142	18	and	and	CCONJ
alces-297	142	19	were	be	AUX
alces-297	142	20	eliminated	eliminate	VERB
alces-297	142	21	with	with	ADP
alces-297	142	22	40	40	NUM
alces-297	142	23	%	%	NOUN
alces-297	142	24	annual	annual	ADJ
alces-297	142	25	removal	removal	NOUN
alces-297	142	26	(	(	PUNCT
alces-297	142	27	fig	fig	NOUN
alces-297	142	28	.	.	PUNCT
alces-297	142	29	2b	2b	NUM
alces-297	142	30	)	)	PUNCT
alces-297	142	31	.	.	PUNCT
alces-297	143	1	sys	sys	PROPN
alces-297	143	2	for	for	ADP
alces-297	143	3	moose	moose	NOUN
alces-297	143	4	based	base	VERB
alces-297	143	5	on	on	ADP
alces-297	143	6	wolf	wolf	PROPN
alces-297	143	7	densities	density	NOUN
alces-297	143	8	displayed	display	VERB
alces-297	143	9	a	a	DET
alces-297	143	10	hook	hook	NOUN
alces-297	143	11	-	-	PUNCT
alces-297	143	12	shape	shape	NOUN
alces-297	143	13	curve	curve	NOUN
alces-297	143	14	(	(	PUNCT
alces-297	143	15	fig	fig	NOUN
alces-297	143	16	.	.	PUNCT
alces-297	143	17	2c	2c	NUM
alces-297	143	18	)	)	PUNCT
alces-297	143	19	.	.	PUNCT
alces-297	144	1	wolf	wolf	PROPN
alces-297	144	2	densities	density	NOUN
alces-297	144	3	were	be	AUX
alces-297	144	4	6/1000	6/1000	NUM
alces-297	144	5	km2	km2	NOUN
alces-297	144	6	when	when	SCONJ
alces-297	144	7	wolf	wolf	PROPN
alces-297	144	8	removals	removal	NOUN
alces-297	144	9	were	be	AUX
alces-297	144	10	0	0	NUM
alces-297	144	11	%	%	NOUN
alces-297	144	12	.	.	PUNCT
alces-297	145	1	moose	moose	NOUN
alces-297	145	2	densities	density	NOUN
alces-297	145	3	(	(	PUNCT
alces-297	145	4	154/1000	154/1000	NUM
alces-297	145	5	km2	km2	NOUN
alces-297	145	6	)	)	PUNCT
alces-297	145	7	and	and	CCONJ
alces-297	145	8	harvest	harvest	NOUN
alces-297	145	9	densities	density	NOUN
alces-297	145	10	(	(	PUNCT
alces-297	145	11	12/1000	12/1000	NUM
alces-297	145	12	km2	km2	NOUN
alces-297	145	13	)	)	PUNCT
alces-297	145	14	were	be	AUX
alces-297	145	15	also	also	ADV
alces-297	145	16	low	low	ADJ
alces-297	145	17	at	at	ADP
alces-297	145	18	this	this	DET
alces-297	145	19	wolf	wolf	PROPN
alces-297	145	20	density	density	NOUN
alces-297	145	21	.	.	PUNCT
alces-297	146	1	as	as	SCONJ
alces-297	146	2	wolf	wolf	PROPN
alces-297	146	3	removal	removal	NOUN
alces-297	146	4	rates	rate	NOUN
alces-297	146	5	increased	increase	VERB
alces-297	146	6	,	,	PUNCT
alces-297	146	7	moose	moose	NOUN
alces-297	146	8	numbers	number	NOUN
alces-297	146	9	and	and	CCONJ
alces-297	146	10	harvest	harvest	NOUN
alces-297	146	11	also	also	ADV
alces-297	146	12	increased	increase	VERB
alces-297	146	13	.	.	PUNCT
alces-297	147	1	wolves	wolf	NOUN
alces-297	147	2	responded	respond	VERB
alces-297	147	3	to	to	ADP
alces-297	147	4	the	the	DET
alces-297	147	5	moose	moose	NOUN
alces-297	147	6	increase	increase	NOUN
alces-297	147	7	due	due	ADP
alces-297	147	8	to	to	ADP
alces-297	147	9	the	the	DET
alces-297	147	10	ratio	ratio	NOUN
alces-297	147	11	dependent	dependent	ADJ
alces-297	147	12	numerical	numerical	ADJ
alces-297	147	13	response	response	NOUN
alces-297	147	14	,	,	PUNCT
alces-297	147	15	and	and	CCONJ
alces-297	147	16	wolf	wolf	PROPN
alces-297	147	17	densities	density	NOUN
alces-297	147	18	rose	rise	VERB
alces-297	147	19	until	until	SCONJ
alces-297	147	20	removal	removal	NOUN
alces-297	147	21	rates	rate	NOUN
alces-297	147	22	table	table	VERB
alces-297	147	23	2	2	NUM
alces-297	147	24	.	.	PUNCT
alces-297	147	25	comparison	comparison	NOUN
alces-297	147	26	of	of	ADP
alces-297	147	27	modelled	model	VERB
alces-297	147	28	moose	moose	NOUN
alces-297	147	29	mortality	mortality	NOUN
alces-297	147	30	rates	rate	NOUN
alces-297	147	31	for	for	ADP
alces-297	147	32	the	the	DET
alces-297	147	33	pgsa	pgsa	NOUN
alces-297	147	34	in	in	ADP
alces-297	147	35	central	central	ADJ
alces-297	147	36	british	british	PROPN
alces-297	147	37	columbia	columbia	PROPN
alces-297	147	38	,	,	PUNCT
alces-297	147	39	canada	canada	PROPN
alces-297	147	40	with	with	ADP
alces-297	147	41	studies	study	NOUN
alces-297	147	42	in	in	ADP
alces-297	147	43	alaska	alaska	PROPN
alces-297	147	44	,	,	PUNCT
alces-297	147	45	usa	usa	PROPN
alces-297	147	46	and	and	CCONJ
alces-297	147	47	yukon	yukon	PROPN
alces-297	147	48	,	,	PUNCT
alces-297	147	49	canada	canada	PROPN
alces-297	147	50	.	.	PUNCT
alces-297	148	1	harvest	harvest	NOUN
alces-297	148	2	rates	rate	NOUN
alces-297	148	3	were	be	AUX
alces-297	148	4	16	16	NUM
alces-297	148	5	%	%	NOUN
alces-297	148	6	bulls	bull	NOUN
alces-297	148	7	,	,	PUNCT
alces-297	148	8	2	2	NUM
alces-297	148	9	%	%	NOUN
alces-297	148	10	cows	cow	NOUN
alces-297	148	11	,	,	PUNCT
alces-297	148	12	and	and	CCONJ
alces-297	148	13	9	9	NUM
alces-297	148	14	%	%	NOUN
alces-297	148	15	calves	calf	NOUN
alces-297	148	16	.	.	PUNCT
alces-297	149	1	a.	a.	NOUN
alces-297	149	2	summer	summer	NOUN
alces-297	149	3	mortality	mortality	NOUN
alces-297	149	4	rates	rate	NOUN
alces-297	149	5	(	(	PUNCT
alces-297	149	6	%	%	INTJ
alces-297	149	7	)	)	PUNCT
alces-297	149	8	on	on	ADP
alces-297	149	9	calves	calf	NOUN
alces-297	149	10	.	.	PUNCT
alces-297	150	1	area	area	NOUN
alces-297	150	2	,	,	PUNCT
alces-297	150	3	treatment	treatment	NOUN
alces-297	150	4	wolves	wolf	NOUN
alces-297	150	5	bears	bear	VERB
alces-297	150	6	predation	predation	NOUN
alces-297	150	7	other	other	ADJ
alces-297	150	8	total	total	ADJ
alces-297	150	9	pgsa	pgsa	NOUN
alces-297	150	10	,	,	PUNCT
alces-297	150	11	30	30	NUM
alces-297	150	12	%	%	NOUN
alces-297	150	13	wolf	wolf	NOUN
alces-297	150	14	removal	removal	NOUN
alces-297	150	15	3	3	NUM
alces-297	150	16	35	35	NUM
alces-297	150	17	38	38	NUM
alces-297	150	18	15	15	NUM
alces-297	150	19	53	53	NUM
alces-297	150	20	pgsa	pgsa	NOUN
alces-297	150	21	,	,	PUNCT
alces-297	150	22	20	20	NUM
alces-297	150	23	%	%	NOUN
alces-297	150	24	wolf	wolf	NOUN
alces-297	150	25	removal	removal	NOUN
alces-297	150	26	5	5	NUM
alces-297	150	27	35	35	NUM
alces-297	150	28	40	40	NUM
alces-297	150	29	15	15	NUM
alces-297	150	30	55	55	NUM
alces-297	150	31	pgsa	pgsa	NOUN
alces-297	150	32	,	,	PUNCT
alces-297	150	33	10	10	NUM
alces-297	150	34	%	%	NOUN
alces-297	150	35	wolf	wolf	NOUN
alces-297	150	36	removal	removal	NOUN
alces-297	150	37	6	6	NUM
alces-297	150	38	35	35	NUM
alces-297	150	39	41	41	NUM
alces-297	150	40	15	15	NUM
alces-297	150	41	56	56	NUM
alces-297	150	42	alaska	alaska	PROPN
alces-297	150	43	/	/	SYM
alces-297	150	44	yukon	yukon	PROPN
alces-297	150	45	,	,	PUNCT
alces-297	150	46	min.1	min.1	PROPN
alces-297	150	47	2	2	NUM
alces-297	150	48	25	25	NUM
alces-297	150	49	45	45	NUM
alces-297	150	50	2	2	NUM
alces-297	150	51	47	47	NUM
alces-297	150	52	alaska	alaska	PROPN
alces-297	150	53	/	/	SYM
alces-297	150	54	yukon	yukon	PROPN
alces-297	150	55	,	,	PUNCT
alces-297	150	56	max.1	max.1	VERB
alces-297	150	57	25	25	NUM
alces-297	150	58	67	67	NUM
alces-297	150	59	72	72	NUM
alces-297	150	60	15	15	NUM
alces-297	150	61	80	80	NUM
alces-297	150	62	1minimum	1minimum	NUM
alces-297	150	63	(	(	PUNCT
alces-297	150	64	min	min	NOUN
alces-297	150	65	.	.	PROPN
alces-297	150	66	)	)	PUNCT
alces-297	151	1	and	and	CCONJ
alces-297	151	2	maximum	maximum	ADJ
alces-297	151	3	(	(	PUNCT
alces-297	151	4	max	max	PROPN
alces-297	151	5	.	.	PUNCT
alces-297	151	6	)	)	PUNCT
alces-297	151	7	estimates	estimate	NOUN
alces-297	151	8	of	of	ADP
alces-297	151	9	collared	collared	ADJ
alces-297	151	10	moose	moose	NOUN
alces-297	151	11	calves	calf	NOUN
alces-297	151	12	killed	kill	VERB
alces-297	151	13	among	among	ADP
alces-297	151	14	8	8	NUM
alces-297	151	15	radiotelemetry	radiotelemetry	NOUN
alces-297	151	16	studies	study	NOUN
alces-297	151	17	in	in	ADP
alces-297	151	18	alaska	alaska	PROPN
alces-297	151	19	and	and	CCONJ
alces-297	151	20	yukon	yukon	PROPN
alces-297	151	21	(	(	PUNCT
alces-297	151	22	from	from	ADP
alces-297	151	23	boertje	boertje	PROPN
alces-297	151	24	et	et	PROPN
alces-297	151	25	al	al	PROPN
alces-297	151	26	.	.	PROPN
alces-297	151	27	2009	2009	NUM
alces-297	151	28	)	)	PUNCT
alces-297	151	29	.	.	PUNCT
alces-297	152	1	b.	b.	PROPN
alces-297	152	2	annual	annual	ADJ
alces-297	152	3	mortality	mortality	NOUN
alces-297	152	4	rates	rate	NOUN
alces-297	152	5	(	(	PUNCT
alces-297	152	6	%	%	INTJ
alces-297	152	7	)	)	PUNCT
alces-297	152	8	on	on	ADP
alces-297	152	9	total	total	ADJ
alces-297	152	10	post	post	ADJ
alces-297	152	11	-	-	ADJ
alces-297	152	12	calving	calving	ADJ
alces-297	152	13	moose	moose	NOUN
alces-297	152	14	population.1	population.1	NOUN
alces-297	152	15	area	area	NOUN
alces-297	152	16	,	,	PUNCT
alces-297	152	17	treatment	treatment	NOUN
alces-297	152	18	wolves	wolf	NOUN
alces-297	152	19	bears	bear	VERB
alces-297	152	20	other	other	ADJ
alces-297	152	21	hunting	hunt	VERB
alces-297	152	22	total	total	ADJ
alces-297	152	23	pgsa	pgsa	NOUN
alces-297	152	24	,	,	PUNCT
alces-297	152	25	30	30	NUM
alces-297	152	26	%	%	NOUN
alces-297	152	27	wolf	wolf	NOUN
alces-297	152	28	removal	removal	NOUN
alces-297	152	29	5	5	NUM
alces-297	152	30	13	13	NUM
alces-297	152	31	10	10	NUM
alces-297	152	32	9	9	NUM
alces-297	152	33	37	37	NUM
alces-297	152	34	pgsa	pgsa	NOUN
alces-297	152	35	,	,	PUNCT
alces-297	152	36	20	20	NUM
alces-297	152	37	%	%	NOUN
alces-297	152	38	wolf	wolf	NOUN
alces-297	152	39	removal	removal	NOUN
alces-297	152	40	7	7	NUM
alces-297	152	41	13	13	NUM
alces-297	152	42	10	10	NUM
alces-297	152	43	7	7	NUM
alces-297	152	44	37	37	NUM
alces-297	152	45	pgsa	pgsa	NOUN
alces-297	152	46	,	,	PUNCT
alces-297	152	47	10	10	NUM
alces-297	152	48	%	%	NOUN
alces-297	152	49	wolf	wolf	NOUN
alces-297	152	50	removal	removal	NOUN
alces-297	152	51	8	8	NUM
alces-297	152	52	13	13	NUM
alces-297	152	53	10	10	NUM
alces-297	152	54	7	7	NUM
alces-297	152	55	37	37	NUM
alces-297	152	56	alaska	alaska	PROPN
alces-297	152	57	/	/	SYM
alces-297	152	58	yukon	yukon	PROPN
alces-297	152	59	,	,	PUNCT
alces-297	152	60	min.2	min.2	PROPN
alces-297	152	61	8	8	NUM
alces-297	152	62	9	9	NUM
alces-297	152	63	1	1	NUM
alces-297	152	64	2	2	NUM
alces-297	152	65	27	27	NUM
alces-297	152	66	alaska	alaska	PROPN
alces-297	152	67	/	/	SYM
alces-297	152	68	yukon	yukon	PROPN
alces-297	152	69	,	,	PUNCT
alces-297	152	70	max.2	max.2	PROPN
alces-297	152	71	15	15	NUM
alces-297	152	72	27	27	NUM
alces-297	152	73	6	6	NUM
alces-297	152	74	6	6	NUM
alces-297	152	75	47	47	NUM
alces-297	152	76	1the	1the	NUM
alces-297	152	77	annual	annual	ADJ
alces-297	152	78	mortality	mortality	NOUN
alces-297	152	79	rate	rate	NOUN
alces-297	152	80	of	of	ADP
alces-297	152	81	the	the	DET
alces-297	152	82	post	post	ADJ
alces-297	152	83	-	-	ADJ
alces-297	152	84	calving	calving	ADJ
alces-297	152	85	population	population	NOUN
alces-297	152	86	=	=	PUNCT
alces-297	153	1	no	no	INTJ
alces-297	153	2	.	.	PUNCT
alces-297	153	3	of	of	ADP
alces-297	153	4	moose	moose	NOUN
alces-297	153	5	deaths	death	NOUN
alces-297	153	6	(	(	PUNCT
alces-297	153	7	including	include	VERB
alces-297	153	8	calves)/number	calves)/number	NOUN
alces-297	153	9	of	of	ADP
alces-297	153	10	moose	moose	NOUN
alces-297	153	11	alive	alive	ADJ
alces-297	153	12	after	after	SCONJ
alces-297	153	13	all	all	DET
alces-297	153	14	calves	calf	NOUN
alces-297	153	15	were	be	AUX
alces-297	153	16	born	bear	VERB
alces-297	153	17	.	.	PUNCT
alces-297	154	1	2minimum	2minimum	NUM
alces-297	154	2	(	(	PUNCT
alces-297	154	3	min	min	NOUN
alces-297	154	4	.	.	PROPN
alces-297	154	5	)	)	PUNCT
alces-297	155	1	and	and	CCONJ
alces-297	155	2	maximum	maximum	ADJ
alces-297	155	3	(	(	PUNCT
alces-297	155	4	max	max	PROPN
alces-297	155	5	.	.	PUNCT
alces-297	155	6	)	)	PUNCT
alces-297	155	7	estimates	estimate	NOUN
alces-297	155	8	of	of	ADP
alces-297	155	9	annual	annual	ADJ
alces-297	155	10	predation	predation	NOUN
alces-297	155	11	rates	rate	NOUN
alces-297	155	12	and	and	CCONJ
alces-297	155	13	mortality	mortality	NOUN
alces-297	155	14	rates	rate	NOUN
alces-297	155	15	among	among	ADP
alces-297	155	16	4	4	NUM
alces-297	155	17	postcalving	postcalve	VERB
alces-297	155	18	moose	moose	NOUN
alces-297	155	19	populations	population	NOUN
alces-297	155	20	during	during	ADP
alces-297	155	21	radiotelemetry	radiotelemetry	NOUN
alces-297	155	22	studies	study	NOUN
alces-297	155	23	in	in	ADP
alces-297	155	24	alaska	alaska	PROPN
alces-297	155	25	and	and	CCONJ
alces-297	155	26	yukon	yukon	PROPN
alces-297	155	27	(	(	PUNCT
alces-297	155	28	from	from	ADP
alces-297	155	29	boertje	boertje	PROPN
alces-297	155	30	et	et	PROPN
alces-297	155	31	al	al	PROPN
alces-297	155	32	.	.	PROPN
alces-297	155	33	2009	2009	NUM
alces-297	155	34	)	)	PUNCT
alces-297	155	35	.	.	PUNCT
alces-297	156	1	modelling	model	VERB
alces-297	156	2	sustained	sustained	ADJ
alces-297	156	3	yields	yield	NOUN
alces-297	156	4	for	for	ADP
alces-297	156	5	moose	moose	NOUN
alces-297	156	6	–	–	PUNCT
alces-297	156	7	hatter	hatter	NOUN
alces-297	156	8	.	.	PUNCT
alces-297	157	1	alces	alce	NOUN
alces-297	157	2	vol	vol	NOUN
alces-297	157	3	.	.	PROPN
alces-297	157	4	57	57	NUM
alces-297	157	5	,	,	PUNCT
alces-297	157	6	2021	2021	NUM
alces-297	157	7	120	120	NUM
alces-297	157	8	reached	reach	VERB
alces-297	157	9	31	31	NUM
alces-297	157	10	%	%	NOUN
alces-297	157	11	.	.	PUNCT
alces-297	158	1	wolves	wolf	NOUN
alces-297	158	2	were	be	AUX
alces-297	158	3	not	not	PART
alces-297	158	4	able	able	ADJ
alces-297	158	5	to	to	PART
alces-297	158	6	compensate	compensate	VERB
alces-297	158	7	for	for	ADP
alces-297	158	8	removal	removal	NOUN
alces-297	158	9	rates	rate	NOUN
alces-297	158	10	above	above	ADP
alces-297	158	11	31	31	NUM
alces-297	158	12	%	%	NOUN
alces-297	158	13	and	and	CCONJ
alces-297	158	14	densities	density	NOUN
alces-297	158	15	declined	decline	VERB
alces-297	158	16	.	.	PUNCT
alces-297	159	1	moose	moose	NOUN
alces-297	159	2	densities	density	NOUN
alces-297	159	3	and	and	CCONJ
alces-297	159	4	harvest	harvest	NOUN
alces-297	159	5	continued	continue	VERB
alces-297	159	6	to	to	PART
alces-297	159	7	increase	increase	VERB
alces-297	159	8	as	as	SCONJ
alces-297	159	9	wolf	wolf	NOUN
alces-297	159	10	densities	density	NOUN
alces-297	159	11	declined	decline	VERB
alces-297	159	12	,	,	PUNCT
alces-297	159	13	but	but	CCONJ
alces-297	159	14	at	at	ADP
alces-297	159	15	a	a	DET
alces-297	159	16	much	much	ADV
alces-297	159	17	slower	slow	ADJ
alces-297	159	18	rate	rate	NOUN
alces-297	159	19	due	due	ADP
alces-297	159	20	to	to	ADP
alces-297	159	21	density	density	NOUN
alces-297	159	22	dependent	dependent	ADJ
alces-297	159	23	declines	decline	NOUN
alces-297	159	24	in	in	ADP
alces-297	159	25	moose	moose	NOUN
alces-297	159	26	reproduction	reproduction	NOUN
alces-297	159	27	and	and	CCONJ
alces-297	159	28	survival	survival	NOUN
alces-297	159	29	.	.	PUNCT
alces-297	160	1	although	although	SCONJ
alces-297	160	2	wolf	wolf	PROPN
alces-297	160	3	densities	density	NOUN
alces-297	160	4	from	from	ADP
alces-297	160	5	6	6	NUM
alces-297	160	6	to	to	PART
alces-297	160	7	12/1000	12/1000	NUM
alces-297	160	8	km2	km2	NOUN
alces-297	160	9	were	be	AUX
alces-297	160	10	table	table	NOUN
alces-297	160	11	3	3	NUM
alces-297	160	12	.	.	X
alces-297	160	13	sensitivity	sensitivity	NOUN
alces-297	160	14	analysis	analysis	NOUN
alces-297	160	15	of	of	ADP
alces-297	160	16	different	different	ADJ
alces-297	160	17	bear	bear	NOUN
alces-297	160	18	predation	predation	NOUN
alces-297	160	19	rates	rate	NOUN
alces-297	160	20	on	on	ADP
alces-297	160	21	modelled	model	VERB
alces-297	160	22	wolf	wolf	NOUN
alces-297	160	23	,	,	PUNCT
alces-297	160	24	moose	moose	NOUN
alces-297	160	25	and	and	CCONJ
alces-297	160	26	harvest	harvest	NOUN
alces-297	160	27	parameters	parameter	NOUN
alces-297	160	28	for	for	ADP
alces-297	160	29	the	the	DET
alces-297	160	30	pgsa	pgsa	NOUN
alces-297	160	31	in	in	ADP
alces-297	160	32	central	central	ADJ
alces-297	160	33	british	british	PROPN
alces-297	160	34	columbia	columbia	PROPN
alces-297	160	35	,	,	PUNCT
alces-297	160	36	canada	canada	PROPN
alces-297	160	37	.	.	PUNCT
alces-297	161	1	harvest	harvest	NOUN
alces-297	161	2	rates	rate	NOUN
alces-297	161	3	were	be	AUX
alces-297	161	4	16	16	NUM
alces-297	161	5	%	%	NOUN
alces-297	161	6	bulls	bull	NOUN
alces-297	161	7	,	,	PUNCT
alces-297	161	8	2	2	NUM
alces-297	161	9	%	%	NOUN
alces-297	161	10	cows	cow	NOUN
alces-297	161	11	and	and	CCONJ
alces-297	161	12	9	9	NUM
alces-297	161	13	%	%	NOUN
alces-297	161	14	calves	calf	NOUN
alces-297	161	15	.	.	PUNCT
alces-297	162	1	the	the	DET
alces-297	162	2	sums	sum	NOUN
alces-297	162	3	of	of	ADP
alces-297	162	4	squares	square	NOUN
alces-297	162	5	fit	fit	ADJ
alces-297	162	6	is	be	AUX
alces-297	162	7	the	the	DET
alces-297	162	8	minimum	minimum	ADJ
alces-297	162	9	sum	sum	NOUN
alces-297	162	10	of	of	ADP
alces-297	162	11	squared	square	VERB
alces-297	162	12	differences	difference	NOUN
alces-297	162	13	between	between	ADP
alces-297	162	14	the	the	DET
alces-297	162	15	observed	observe	VERB
alces-297	162	16	and	and	CCONJ
alces-297	162	17	modelled	model	VERB
alces-297	162	18	moose	moose	NOUN
alces-297	162	19	density	density	NOUN
alces-297	162	20	,	,	PUNCT
alces-297	162	21	wolf	wolf	PROPN
alces-297	162	22	density	density	NOUN
alces-297	162	23	,	,	PUNCT
alces-297	162	24	harvest	harvest	NOUN
alces-297	162	25	density	density	NOUN
alces-297	162	26	,	,	PUNCT
alces-297	162	27	and	and	CCONJ
alces-297	162	28	bull	bull	NOUN
alces-297	162	29	:	:	PUNCT
alces-297	162	30	cow	cow	NOUN
alces-297	162	31	and	and	CCONJ
alces-297	162	32	calf	calf	NOUN
alces-297	162	33	:	:	PUNCT
alces-297	162	34	cow	cow	NOUN
alces-297	162	35	ratios	ratio	NOUN
alces-297	162	36	.	.	PUNCT
alces-297	163	1	bear	bear	PROPN
alces-297	163	2	predation	predation	NOUN
alces-297	163	3	rate	rate	NOUN
alces-297	163	4	(	(	PUNCT
alces-297	163	5	%	%	NOUN
alces-297	163	6	)	)	PUNCT
alces-297	163	7	10	10	NUM
alces-297	163	8	15	15	NUM
alces-297	163	9	20	20	NUM
alces-297	163	10	25	25	NUM
alces-297	163	11	30	30	NUM
alces-297	163	12	35	35	NUM
alces-297	163	13	40	40	NUM
alces-297	163	14	45	45	NUM
alces-297	163	15	50	50	NUM
alces-297	163	16	wolf	wolf	NOUN
alces-297	163	17	removal	removal	NOUN
alces-297	163	18	rate	rate	NOUN
alces-297	163	19	(	(	PUNCT
alces-297	163	20	%	%	INTJ
alces-297	163	21	)	)	PUNCT
alces-297	163	22	21	21	NUM
alces-297	163	23	23	23	NUM
alces-297	163	24	25	25	NUM
alces-297	163	25	27	27	NUM
alces-297	163	26	29	29	NUM
alces-297	163	27	31	31	NUM
alces-297	163	28	33	33	NUM
alces-297	163	29	35	35	NUM
alces-297	163	30	37	37	NUM
alces-297	163	31	moose	moose	NOUN
alces-297	163	32	harvest	harvest	NOUN
alces-297	163	33	rate	rate	NOUN
alces-297	163	34	(	(	PUNCT
alces-297	163	35	%	%	NOUN
alces-297	163	36	)	)	PUNCT
alces-297	163	37	8	8	NUM
alces-297	163	38	8	8	NUM
alces-297	163	39	8	8	NUM
alces-297	163	40	7	7	NUM
alces-297	163	41	7	7	NUM
alces-297	163	42	7	7	NUM
alces-297	163	43	7	7	NUM
alces-297	163	44	7	7	NUM
alces-297	163	45	7	7	NUM
alces-297	163	46	harvest/1000	harvest/1000	NOUN
alces-297	163	47	km2	km2	PROPN
alces-297	163	48	119	119	NUM
alces-297	163	49	117	117	NUM
alces-297	163	50	116	116	NUM
alces-297	163	51	115	115	NUM
alces-297	163	52	103	103	NUM
alces-297	163	53	101	101	NUM
alces-297	163	54	99	99	NUM
alces-297	163	55	97	97	NUM
alces-297	163	56	94	94	NUM
alces-297	163	57	wolves/1000	wolves/1000	PROPN
alces-297	163	58	km2	km2	PROPN
alces-297	163	59	31	31	NUM
alces-297	163	60	28	28	NUM
alces-297	163	61	24	24	NUM
alces-297	163	62	21	21	NUM
alces-297	163	63	16	16	NUM
alces-297	163	64	12	12	NUM
alces-297	163	65	9	9	NUM
alces-297	163	66	6	6	NUM
alces-297	163	67	2	2	NUM
alces-297	163	68	moose/1000	moose/1000	NOUN
alces-297	163	69	km2	km2	PROPN
alces-297	163	70	1400	1400	NUM
alces-297	163	71	1400	1400	NUM
alces-297	163	72	1400	1400	NUM
alces-297	163	73	1400	1400	NUM
alces-297	163	74	1319	1319	NUM
alces-297	163	75	1319	1319	NUM
alces-297	163	76	1319	1319	NUM
alces-297	163	77	1320	1320	NUM
alces-297	163	78	1320	1320	NUM
alces-297	163	79	moose	moose	NOUN
alces-297	163	80	/	/	SYM
alces-297	163	81	wolf	wolf	PROPN
alces-297	163	82	45	45	NUM
alces-297	163	83	51	51	NUM
alces-297	163	84	57	57	NUM
alces-297	163	85	67	67	NUM
alces-297	163	86	84	84	NUM
alces-297	163	87	106	106	NUM
alces-297	163	88	145	145	NUM
alces-297	163	89	238	238	NUM
alces-297	163	90	711	711	NUM
alces-297	163	91	bulls/100	bulls/100	PROPN
alces-297	163	92	cows	cow	NOUN
alces-297	163	93	49	49	NUM
alces-297	163	94	48	48	NUM
alces-297	163	95	47	47	NUM
alces-297	163	96	46	46	NUM
alces-297	163	97	41	41	NUM
alces-297	163	98	40	40	NUM
alces-297	163	99	38	38	NUM
alces-297	163	100	36	36	NUM
alces-297	163	101	34	34	NUM
alces-297	163	102	calves/100	calves/100	NOUN
alces-297	163	103	cows	cow	VERB
alces-297	163	104	60	60	NUM
alces-297	163	105	56	56	NUM
alces-297	163	106	53	53	NUM
alces-297	163	107	49	49	NUM
alces-297	163	108	41	41	NUM
alces-297	163	109	38	38	NUM
alces-297	163	110	34	34	NUM
alces-297	163	111	31	31	NUM
alces-297	163	112	28	28	NUM
alces-297	163	113	sums	sum	NOUN
alces-297	163	114	of	of	ADP
alces-297	163	115	squares	square	NOUN
alces-297	163	116	fit	fit	VERB
alces-297	163	117	7817	7817	NUM
alces-297	163	118	7526	7526	NUM
alces-297	163	119	7271	7271	NUM
alces-297	163	120	7061	7061	NUM
alces-297	164	1	156	156	NUM
alces-297	164	2	136	136	NUM
alces-297	164	3	171	171	NUM
alces-297	164	4	269	269	NUM
alces-297	164	5	441	441	NUM
alces-297	164	6	fig	fig	NOUN
alces-297	164	7	.	.	PUNCT
alces-297	165	1	2	2	X
alces-297	165	2	.	.	X
alces-297	165	3	relationship	relationship	NOUN
alces-297	165	4	between	between	ADP
alces-297	165	5	modelled	model	VERB
alces-297	165	6	moose	moose	NOUN
alces-297	165	7	and	and	CCONJ
alces-297	165	8	wolf	wolf	NOUN
alces-297	165	9	parameters	parameter	NOUN
alces-297	165	10	and	and	CCONJ
alces-297	165	11	sys	sys	PROPN
alces-297	165	12	(	(	PUNCT
alces-297	165	13	moose	moose	NOUN
alces-297	165	14	harvest	harvest	NOUN
alces-297	165	15	density	density	NOUN
alces-297	165	16	)	)	PUNCT
alces-297	165	17	for	for	ADP
alces-297	165	18	the	the	DET
alces-297	165	19	pgsa	pgsa	NOUN
alces-297	165	20	in	in	ADP
alces-297	165	21	central	central	ADJ
alces-297	165	22	british	british	PROPN
alces-297	165	23	columbia	columbia	PROPN
alces-297	165	24	,	,	PUNCT
alces-297	165	25	canada	canada	PROPN
alces-297	165	26	with	with	ADP
alces-297	165	27	annual	annual	ADJ
alces-297	165	28	harvests	harvest	NOUN
alces-297	165	29	of	of	ADP
alces-297	165	30	16	16	NUM
alces-297	165	31	%	%	NOUN
alces-297	165	32	bulls	bull	NOUN
alces-297	165	33	,	,	PUNCT
alces-297	165	34	2	2	NUM
alces-297	165	35	%	%	NOUN
alces-297	165	36	cows	cow	NOUN
alces-297	165	37	and	and	CCONJ
alces-297	165	38	9	9	NUM
alces-297	165	39	%	%	NOUN
alces-297	165	40	calves	calf	NOUN
alces-297	165	41	.	.	PUNCT
alces-297	166	1	each	each	DET
alces-297	166	2	data	data	NOUN
alces-297	166	3	point	point	NOUN
alces-297	166	4	(	(	PUNCT
alces-297	166	5	open	open	ADJ
alces-297	166	6	circle	circle	NOUN
alces-297	166	7	)	)	PUNCT
alces-297	166	8	was	be	AUX
alces-297	166	9	generated	generate	VERB
alces-297	166	10	by	by	ADP
alces-297	166	11	modelling	model	VERB
alces-297	166	12	predator	predator	NOUN
alces-297	166	13	-	-	PUNCT
alces-297	166	14	prey	prey	NOUN
alces-297	166	15	dynamics	dynamic	NOUN
alces-297	166	16	for	for	ADP
alces-297	166	17	100	100	NUM
alces-297	166	18	years	year	NOUN
alces-297	166	19	with	with	ADP
alces-297	166	20	wolf	wolf	PROPN
alces-297	166	21	removal	removal	NOUN
alces-297	166	22	rates	rate	NOUN
alces-297	166	23	ranging	range	VERB
alces-297	166	24	from	from	ADP
alces-297	166	25	0	0	NUM
alces-297	166	26	to	to	PART
alces-297	166	27	40	40	NUM
alces-297	166	28	%	%	NOUN
alces-297	166	29	.	.	PUNCT
alces-297	167	1	(	(	PUNCT
alces-297	167	2	a	a	X
alces-297	167	3	)	)	PUNCT
alces-297	167	4	relationship	relationship	NOUN
alces-297	167	5	between	between	ADP
alces-297	167	6	sys	sys	NOUN
alces-297	167	7	and	and	CCONJ
alces-297	167	8	wolf	wolf	PROPN
alces-297	167	9	removal	removal	NOUN
alces-297	167	10	rate	rate	NOUN
alces-297	167	11	;	;	PUNCT
alces-297	167	12	(	(	PUNCT
alces-297	167	13	b	b	X
alces-297	167	14	)	)	PUNCT
alces-297	167	15	relationship	relationship	NOUN
alces-297	167	16	between	between	ADP
alces-297	167	17	wolf	wolf	PROPN
alces-297	167	18	density	density	NOUN
alces-297	167	19	and	and	CCONJ
alces-297	167	20	wolf	wolf	PROPN
alces-297	167	21	removal	removal	NOUN
alces-297	167	22	rate	rate	NOUN
alces-297	167	23	;	;	PUNCT
alces-297	167	24	(	(	PUNCT
alces-297	167	25	c	c	X
alces-297	167	26	)	)	PUNCT
alces-297	167	27	relationship	relationship	NOUN
alces-297	167	28	between	between	ADP
alces-297	167	29	sys	sys	NOUN
alces-297	167	30	and	and	CCONJ
alces-297	167	31	wolf	wolf	PROPN
alces-297	167	32	density	density	NOUN
alces-297	167	33	;	;	PUNCT
alces-297	167	34	and	and	CCONJ
alces-297	167	35	(	(	PUNCT
alces-297	167	36	d	d	X
alces-297	167	37	)	)	PUNCT
alces-297	167	38	sy	sy	PROPN
alces-297	167	39	(	(	PUNCT
alces-297	167	40	yield	yield	NOUN
alces-297	167	41	-	-	PUNCT
alces-297	167	42	density	density	NOUN
alces-297	167	43	)	)	PUNCT
alces-297	167	44	curve	curve	NOUN
alces-297	167	45	for	for	ADP
alces-297	167	46	moose	moose	NOUN
alces-297	167	47	harvest	harvest	NOUN
alces-297	167	48	density	density	NOUN
alces-297	167	49	and	and	CCONJ
alces-297	167	50	moose	moose	NOUN
alces-297	167	51	density	density	NOUN
alces-297	167	52	.	.	PUNCT
alces-297	168	1	alces	alce	NOUN
alces-297	168	2	vol	vol	NOUN
alces-297	168	3	.	.	PROPN
alces-297	168	4	57	57	NUM
alces-297	168	5	,	,	PUNCT
alces-297	168	6	2021	2021	NUM
alces-297	168	7	modelling	model	VERB
alces-297	168	8	sustained	sustain	VERB
alces-297	168	9	yields	yield	NOUN
alces-297	168	10	for	for	ADP
alces-297	168	11	moose	moose	NOUN
alces-297	168	12	–	–	PUNCT
alces-297	168	13	hatter	hatter	NOUN
alces-297	168	14	.	.	PUNCT
alces-297	169	1	121	121	NUM
alces-297	169	2	associated	associate	VERB
alces-297	169	3	with	with	ADP
alces-297	169	4	both	both	CCONJ
alces-297	169	5	low	low	ADJ
alces-297	169	6	and	and	CCONJ
alces-297	169	7	high	high	ADJ
alces-297	169	8	sys	sys	PROPN
alces-297	169	9	,	,	PUNCT
alces-297	169	10	wolf	wolf	PROPN
alces-297	169	11	removal	removal	NOUN
alces-297	169	12	rates	rate	NOUN
alces-297	169	13	and	and	CCONJ
alces-297	169	14	moose	moose	NOUN
alces-297	169	15	population	population	NOUN
alces-297	169	16	dynamics	dynamic	NOUN
alces-297	169	17	varied	vary	VERB
alces-297	169	18	greatly	greatly	ADV
alces-297	169	19	within	within	ADP
alces-297	169	20	this	this	DET
alces-297	169	21	range	range	NOUN
alces-297	169	22	.	.	PUNCT
alces-297	170	1	the	the	DET
alces-297	170	2	sy	sy	NOUN
alces-297	170	3	or	or	CCONJ
alces-297	170	4	yield	yield	NOUN
alces-297	170	5	-	-	PUNCT
alces-297	170	6	density	density	NOUN
alces-297	170	7	curve	curve	NOUN
alces-297	170	8	was	be	AUX
alces-297	170	9	linear	linear	ADJ
alces-297	170	10	over	over	ADP
alces-297	170	11	the	the	DET
alces-297	170	12	range	range	NOUN
alces-297	170	13	in	in	ADP
alces-297	170	14	moose	moose	NOUN
alces-297	170	15	density	density	NOUN
alces-297	170	16	(	(	PUNCT
alces-297	170	17	fig	fig	NOUN
alces-297	170	18	.	.	PUNCT
alces-297	171	1	2d	2d	NOUN
alces-297	171	2	)	)	PUNCT
alces-297	171	3	,	,	PUNCT
alces-297	171	4	as	as	SCONJ
alces-297	171	5	harvest	harvest	NOUN
alces-297	171	6	rates	rate	NOUN
alces-297	171	7	were	be	AUX
alces-297	171	8	constant	constant	ADJ
alces-297	171	9	.	.	PUNCT
alces-297	172	1	sys	sys	PROPN
alces-297	172	2	ranged	range	VERB
alces-297	172	3	from	from	ADP
alces-297	172	4	12	12	NUM
alces-297	172	5	kills/1000	kills/1000	NOUN
alces-297	172	6	km2	km2	PROPN
alces-297	172	7	at	at	ADP
alces-297	172	8	154	154	NUM
alces-297	172	9	moose/1000	moose/1000	NOUN
alces-297	172	10	km2	km2	X
alces-297	172	11	to	to	ADP
alces-297	172	12	121	121	NUM
alces-297	172	13	kills/1000	kills/1000	NOUN
alces-297	172	14	km2	km2	PROPN
alces-297	172	15	at	at	ADP
alces-297	172	16	1571	1571	NUM
alces-297	172	17	moose/1000	moose/1000	PROPN
alces-297	172	18	km2	km2	PROPN
alces-297	172	19	.	.	PUNCT
alces-297	173	1	the	the	DET
alces-297	173	2	yield	yield	NOUN
alces-297	173	3	-	-	PUNCT
alces-297	173	4	density	density	NOUN
alces-297	173	5	curve	curve	NOUN
alces-297	173	6	for	for	ADP
alces-297	173	7	the	the	DET
alces-297	173	8	pgsa	pgsa	NOUN
alces-297	173	9	was	be	AUX
alces-297	173	10	similar	similar	ADJ
alces-297	173	11	to	to	ADP
alces-297	173	12	the	the	DET
alces-297	173	13	preliminary	preliminary	ADJ
alces-297	173	14	yield	yield	NOUN
alces-297	173	15	-	-	PUNCT
alces-297	173	16	density	density	NOUN
alces-297	173	17	curve	curve	NOUN
alces-297	173	18	for	for	ADP
alces-297	173	19	moose	moose	NOUN
alces-297	173	20	in	in	ADP
alces-297	173	21	north	north	NOUN
alces-297	173	22	-	-	PUNCT
alces-297	173	23	central	central	ADJ
alces-297	173	24	bc	bc	PROPN
alces-297	173	25	(	(	PUNCT
alces-297	173	26	fig	fig	NOUN
alces-297	173	27	.	.	PUNCT
alces-297	174	1	3	3	NUM
alces-297	174	2	)	)	PUNCT
alces-297	174	3	.	.	PUNCT
alces-297	175	1	optimal	optimal	ADJ
alces-297	175	2	harvests	harvest	NOUN
alces-297	175	3	of	of	ADP
alces-297	175	4	bulls	bull	NOUN
alces-297	175	5	,	,	PUNCT
alces-297	175	6	cows	cow	NOUN
alces-297	175	7	,	,	PUNCT
alces-297	175	8	and	and	CCONJ
alces-297	175	9	calves	calf	NOUN
alces-297	175	10	(	(	PUNCT
alces-297	175	11	“	"	PUNCT
alces-297	175	12	fully	fully	ADV
alces-297	175	13	optimized	optimize	VERB
alces-297	175	14	harvest	harvest	NOUN
alces-297	175	15	rates	rate	NOUN
alces-297	175	16	,	,	PUNCT
alces-297	175	17	”	"	PUNCT
alces-297	175	18	table	table	NOUN
alces-297	175	19	4a	4a	NOUN
alces-297	175	20	)	)	PUNCT
alces-297	175	21	,	,	PUNCT
alces-297	175	22	optimal	optimal	ADJ
alces-297	175	23	bull	bull	NOUN
alces-297	175	24	and	and	CCONJ
alces-297	175	25	calf	calf	NOUN
alces-297	175	26	harvests	harvest	NOUN
alces-297	175	27	with	with	ADP
alces-297	175	28	a	a	DET
alces-297	175	29	cow	cow	NOUN
alces-297	175	30	harvest	harvest	NOUN
alces-297	175	31	of	of	ADP
alces-297	175	32	2	2	NUM
alces-297	175	33	%	%	NOUN
alces-297	175	34	(	(	PUNCT
alces-297	175	35	“	"	PUNCT
alces-297	175	36	optimized	optimize	VERB
alces-297	175	37	bull	bull	NOUN
alces-297	175	38	and	and	CCONJ
alces-297	175	39	calf	calf	NOUN
alces-297	175	40	harvest	harvest	NOUN
alces-297	175	41	rates	rate	NOUN
alces-297	175	42	,	,	PUNCT
alces-297	175	43	”	"	PUNCT
alces-297	175	44	table	table	NOUN
alces-297	175	45	4b	4b	PROPN
alces-297	175	46	)	)	PUNCT
alces-297	175	47	,	,	PUNCT
alces-297	175	48	and	and	CCONJ
alces-297	175	49	optimal	optimal	ADJ
alces-297	175	50	harvest	harvest	NOUN
alces-297	175	51	of	of	ADP
alces-297	175	52	bulls	bull	NOUN
alces-297	175	53	in	in	ADP
alces-297	175	54	bull	bull	NOUN
alces-297	175	55	-	-	PUNCT
alces-297	175	56	only	only	ADV
alces-297	175	57	seasons	season	NOUN
alces-297	175	58	(	(	PUNCT
alces-297	175	59	“	"	PUNCT
alces-297	175	60	optimized	optimize	VERB
alces-297	175	61	bull	bull	NOUN
alces-297	175	62	harvest	harvest	NOUN
alces-297	175	63	rates	rate	NOUN
alces-297	175	64	,	,	PUNCT
alces-297	175	65	”	"	PUNCT
alces-297	175	66	table	table	NOUN
alces-297	175	67	4c	4c	NOUN
alces-297	175	68	)	)	PUNCT
alces-297	175	69	increased	increase	VERB
alces-297	175	70	with	with	ADP
alces-297	175	71	increasing	increase	VERB
alces-297	175	72	wolf	wolf	PROPN
alces-297	175	73	removal	removal	NOUN
alces-297	175	74	rates	rate	NOUN
alces-297	175	75	.	.	PUNCT
alces-297	176	1	optimized	optimize	VERB
alces-297	176	2	bull	bull	NOUN
alces-297	176	3	and	and	CCONJ
alces-297	176	4	calf	calf	NOUN
alces-297	176	5	harvest	harvest	NOUN
alces-297	176	6	rates	rate	NOUN
alces-297	176	7	produced	produce	VERB
alces-297	176	8	only	only	ADV
alces-297	176	9	slightly	slightly	ADV
alces-297	176	10	lower	low	ADJ
alces-297	176	11	yields	yield	NOUN
alces-297	176	12	than	than	ADP
alces-297	176	13	fully	fully	ADV
alces-297	176	14	optimized	optimize	VERB
alces-297	176	15	harvest	harvest	NOUN
alces-297	176	16	rates	rate	NOUN
alces-297	176	17	.	.	PUNCT
alces-297	177	1	fully	fully	ADV
alces-297	177	2	optimized	optimize	VERB
alces-297	177	3	harvest	harvest	NOUN
alces-297	177	4	rates	rate	NOUN
alces-297	177	5	and	and	CCONJ
alces-297	177	6	optimized	optimize	VERB
alces-297	177	7	bull	bull	NOUN
alces-297	177	8	harvest	harvest	NOUN
alces-297	177	9	rates	rate	NOUN
alces-297	177	10	produced	produce	VERB
alces-297	177	11	equivalent	equivalent	ADJ
alces-297	177	12	yields	yield	NOUN
alces-297	177	13	when	when	SCONJ
alces-297	177	14	wolf	wolf	PROPN
alces-297	177	15	removal	removal	NOUN
alces-297	177	16	rates	rate	NOUN
alces-297	177	17	were	be	AUX
alces-297	177	18	low	low	ADJ
alces-297	177	19	(	(	PUNCT
alces-297	177	20	0–10	0–10	NOUN
alces-297	177	21	%	%	NOUN
alces-297	177	22	)	)	PUNCT
alces-297	177	23	.	.	PUNCT
alces-297	178	1	this	this	PRON
alces-297	178	2	was	be	AUX
alces-297	178	3	because	because	SCONJ
alces-297	178	4	fully	fully	ADV
alces-297	178	5	optimized	optimize	VERB
alces-297	178	6	harvests	harvest	NOUN
alces-297	178	7	reduced	reduce	VERB
alces-297	178	8	moose	moose	NOUN
alces-297	178	9	densities	density	NOUN
alces-297	178	10	to	to	ADP
alces-297	178	11	<	<	X
alces-297	178	12	200/1000	200/1000	NUM
alces-297	178	13	km2	km2	X
alces-297	178	14	which	which	PRON
alces-297	178	15	required	require	VERB
alces-297	178	16	≥	≥	NOUN
alces-297	178	17	50	50	NUM
alces-297	178	18	bulls/100	bulls/100	PROPN
alces-297	178	19	cows	cow	NOUN
alces-297	178	20	to	to	PART
alces-297	178	21	comply	comply	VERB
alces-297	178	22	with	with	ADP
alces-297	178	23	bc	bc	PROPN
alces-297	178	24	’s	’s	PART
alces-297	178	25	harvest	harvest	NOUN
alces-297	178	26	management	management	NOUN
alces-297	178	27	procedure	procedure	NOUN
alces-297	178	28	,	,	PUNCT
alces-297	178	29	while	while	SCONJ
alces-297	178	30	optimized	optimize	VERB
alces-297	178	31	bull	bull	NOUN
alces-297	178	32	harvests	harvest	NOUN
alces-297	178	33	maintained	maintain	VERB
alces-297	178	34	densities	density	NOUN
alces-297	178	35	>	>	X
alces-297	178	36	200/1000	200/1000	NUM
alces-297	178	37	km2	km2	X
alces-297	178	38	where	where	SCONJ
alces-297	178	39	only	only	ADV
alces-297	178	40	≥	≥	X
alces-297	178	41	30	30	NUM
alces-297	178	42	bulls/100	bulls/100	PROPN
alces-297	178	43	cows	cow	NOUN
alces-297	178	44	were	be	AUX
alces-297	178	45	required	require	VERB
alces-297	178	46	by	by	ADP
alces-297	178	47	the	the	DET
alces-297	178	48	procedure	procedure	NOUN
alces-297	178	49	.	.	PUNCT
alces-297	179	1	in	in	ADP
alces-297	179	2	all	all	DET
alces-297	179	3	cases	case	NOUN
alces-297	179	4	,	,	PUNCT
alces-297	179	5	the	the	DET
alces-297	179	6	greatest	great	ADJ
alces-297	179	7	harvests	harvest	NOUN
alces-297	179	8	were	be	AUX
alces-297	179	9	achieved	achieve	VERB
alces-297	179	10	when	when	SCONJ
alces-297	179	11	adult	adult	NOUN
alces-297	179	12	sex	sex	NOUN
alces-297	179	13	ratios	ratio	NOUN
alces-297	179	14	were	be	AUX
alces-297	179	15	maintained	maintain	VERB
alces-297	179	16	at	at	ADP
alces-297	179	17	the	the	DET
alces-297	179	18	minimum	minimum	ADJ
alces-297	179	19	bull	bull	NOUN
alces-297	179	20	:	:	PUNCT
alces-297	179	21	cow	cow	NOUN
alces-297	179	22	ratio	ratio	NOUN
alces-297	179	23	objective	objective	NOUN
alces-297	179	24	.	.	PUNCT
alces-297	180	1	calf	calf	NOUN
alces-297	180	2	:	:	PUNCT
alces-297	180	3	cow	cow	NOUN
alces-297	180	4	ratios	ratio	NOUN
alces-297	180	5	were	be	AUX
alces-297	180	6	variable	variable	ADJ
alces-297	180	7	and	and	CCONJ
alces-297	180	8	reflected	reflect	VERB
alces-297	180	9	wolf	wolf	PROPN
alces-297	180	10	removal	removal	NOUN
alces-297	180	11	rates	rate	NOUN
alces-297	180	12	,	,	PUNCT
alces-297	180	13	calf	calf	NOUN
alces-297	180	14	harvest	harvest	NOUN
alces-297	180	15	rates	rate	NOUN
alces-297	180	16	,	,	PUNCT
alces-297	180	17	and	and	CCONJ
alces-297	180	18	density	density	NOUN
alces-297	180	19	dependence	dependence	NOUN
alces-297	180	20	.	.	PUNCT
alces-297	181	1	fully	fully	ADV
alces-297	181	2	optimized	optimize	VERB
alces-297	181	3	harvest	harvest	NOUN
alces-297	181	4	rates	rate	NOUN
alces-297	181	5	with	with	ADP
alces-297	181	6	low	low	ADJ
alces-297	181	7	wolf	wolf	NOUN
alces-297	181	8	removals	removal	NOUN
alces-297	181	9	(	(	PUNCT
alces-297	181	10	0–10	0–10	NOUN
alces-297	181	11	%	%	NOUN
alces-297	181	12	)	)	PUNCT
alces-297	181	13	were	be	AUX
alces-297	181	14	6	6	NUM
alces-297	181	15	%	%	NOUN
alces-297	181	16	for	for	ADP
alces-297	181	17	bulls	bull	NOUN
alces-297	181	18	,	,	PUNCT
alces-297	181	19	0	0	NUM
alces-297	181	20	%	%	NOUN
alces-297	181	21	for	for	ADP
alces-297	181	22	cows	cow	NOUN
alces-297	181	23	,	,	PUNCT
alces-297	181	24	and	and	CCONJ
alces-297	181	25	28–30	28–30	NUM
alces-297	181	26	%	%	NOUN
alces-297	181	27	for	for	ADP
alces-297	181	28	calves	calf	NOUN
alces-297	181	29	.	.	PUNCT
alces-297	182	1	with	with	ADP
alces-297	182	2	moderate	moderate	ADJ
alces-297	182	3	to	to	PART
alces-297	182	4	high	high	ADJ
alces-297	182	5	wolf	wolf	NOUN
alces-297	182	6	removals	removal	NOUN
alces-297	182	7	(	(	PUNCT
alces-297	182	8	20–35	20–35	NUM
alces-297	182	9	%	%	NOUN
alces-297	182	10	)	)	PUNCT
alces-297	182	11	,	,	PUNCT
alces-297	182	12	optimal	optimal	ADJ
alces-297	182	13	harvest	harvest	NOUN
alces-297	182	14	rates	rate	NOUN
alces-297	182	15	were	be	AUX
alces-297	182	16	17–21	17–21	NUM
alces-297	182	17	%	%	NOUN
alces-297	182	18	for	for	ADP
alces-297	182	19	bulls	bull	NOUN
alces-297	182	20	,	,	PUNCT
alces-297	182	21	0–0.2	0–0.2	NOUN
alces-297	182	22	%	%	NOUN
alces-297	182	23	for	for	ADP
alces-297	182	24	cows	cow	NOUN
alces-297	182	25	,	,	PUNCT
alces-297	182	26	and	and	CCONJ
alces-297	182	27	15–34	15–34	NUM
alces-297	182	28	%	%	NOUN
alces-297	182	29	for	for	ADP
alces-297	182	30	calves	calf	NOUN
alces-297	182	31	.	.	PUNCT
alces-297	183	1	optimal	optimal	ADJ
alces-297	183	2	harvest	harvest	NOUN
alces-297	183	3	rates	rate	NOUN
alces-297	183	4	,	,	PUNCT
alces-297	183	5	when	when	SCONJ
alces-297	183	6	wolves	wolf	NOUN
alces-297	183	7	were	be	AUX
alces-297	183	8	absent	absent	ADJ
alces-297	183	9	,	,	PUNCT
alces-297	183	10	were	be	AUX
alces-297	183	11	16	16	NUM
alces-297	183	12	%	%	NOUN
alces-297	183	13	for	for	ADP
alces-297	183	14	bulls	bull	NOUN
alces-297	183	15	,	,	PUNCT
alces-297	183	16	0.2	0.2	NUM
alces-297	183	17	%	%	NOUN
alces-297	183	18	for	for	ADP
alces-297	183	19	cows	cow	NOUN
alces-297	183	20	,	,	PUNCT
alces-297	183	21	and	and	CCONJ
alces-297	183	22	43	43	NUM
alces-297	183	23	%	%	NOUN
alces-297	183	24	for	for	ADP
alces-297	183	25	calves	calf	NOUN
alces-297	183	26	.	.	PUNCT
alces-297	184	1	moose	moose	PROPN
alces-297	184	2	harvest	harvest	NOUN
alces-297	184	3	rates	rate	NOUN
alces-297	184	4	ranged	range	VERB
alces-297	184	5	from	from	ADP
alces-297	184	6	8	8	NUM
alces-297	184	7	to	to	PART
alces-297	184	8	15	15	NUM
alces-297	184	9	%	%	NOUN
alces-297	184	10	and	and	CCONJ
alces-297	184	11	were	be	AUX
alces-297	184	12	highest	high	ADJ
alces-297	184	13	when	when	SCONJ
alces-297	184	14	wolves	wolf	NOUN
alces-297	184	15	were	be	AUX
alces-297	184	16	absent	absent	ADJ
alces-297	184	17	.	.	PUNCT
alces-297	185	1	y	y	NOUN
alces-297	185	2	=	=	SYM
alces-297	186	1	4e-05x2	4e-05x2	NUM
alces-297	187	1	+	+	NUM
alces-297	187	2	0.049x	0.049x	ADJ
alces-297	187	3	r²	r²	NOUN
alces-297	187	4	=	=	NOUN
alces-297	187	5	0.86	0.86	NUM
alces-297	187	6	0	0	NUM
alces-297	187	7	10	10	NUM
alces-297	187	8	20	20	NUM
alces-297	187	9	30	30	NUM
alces-297	187	10	40	40	NUM
alces-297	187	11	50	50	NUM
alces-297	187	12	60	60	NUM
alces-297	187	13	70	70	NUM
alces-297	187	14	80	80	NUM
alces-297	187	15	0	0	NUM
alces-297	187	16	100	100	NUM
alces-297	187	17	200	200	NUM
alces-297	187	18	300	300	NUM
alces-297	187	19	400	400	NUM
alces-297	187	20	500	500	NUM
alces-297	187	21	600	600	NUM
alces-297	187	22	700	700	NUM
alces-297	187	23	800	800	NUM
alces-297	187	24	ha	ha	INTJ
alces-297	187	25	rv	rv	NOUN
alces-297	187	26	es	es	PROPN
alces-297	187	27	t/	t/	PRON
alces-297	187	28	10	10	NUM
alces-297	187	29	00	00	NUM
alces-297	188	1	k	k	NUM
alces-297	188	2	m	m	VERB
alces-297	188	3	2	2	NUM
alces-297	188	4	moose/1000	moose/1000	NOUN
alces-297	188	5	km2	km2	PROPN
alces-297	188	6	fig	fig	NOUN
alces-297	188	7	.	.	PUNCT
alces-297	189	1	3	3	X
alces-297	189	2	.	.	X
alces-297	189	3	yield	yield	NOUN
alces-297	189	4	-	-	PUNCT
alces-297	189	5	density	density	NOUN
alces-297	189	6	curves	curve	NOUN
alces-297	189	7	for	for	ADP
alces-297	189	8	moose	moose	NOUN
alces-297	189	9	in	in	ADP
alces-297	189	10	central	central	ADJ
alces-297	189	11	british	british	PROPN
alces-297	189	12	columbia	columbia	PROPN
alces-297	189	13	,	,	PUNCT
alces-297	189	14	canada	canada	PROPN
alces-297	189	15	.	.	PUNCT
alces-297	190	1	the	the	DET
alces-297	190	2	solid	solid	ADJ
alces-297	190	3	line	line	NOUN
alces-297	190	4	is	be	AUX
alces-297	190	5	the	the	DET
alces-297	190	6	modelled	model	VERB
alces-297	190	7	yield	yield	NOUN
alces-297	190	8	-	-	PUNCT
alces-297	190	9	density	density	NOUN
alces-297	190	10	curve	curve	NOUN
alces-297	190	11	for	for	ADP
alces-297	190	12	moose	moose	NOUN
alces-297	190	13	in	in	ADP
alces-297	190	14	the	the	DET
alces-297	190	15	pgsa	pgsa	NOUN
alces-297	190	16	with	with	ADP
alces-297	190	17	a	a	DET
alces-297	190	18	harvest	harvest	NOUN
alces-297	190	19	of	of	ADP
alces-297	190	20	16	16	NUM
alces-297	190	21	%	%	NOUN
alces-297	190	22	bulls	bull	NOUN
alces-297	190	23	,	,	PUNCT
alces-297	190	24	2	2	NUM
alces-297	190	25	%	%	NOUN
alces-297	190	26	cows	cow	NOUN
alces-297	190	27	and	and	CCONJ
alces-297	190	28	9	9	NUM
alces-297	190	29	%	%	NOUN
alces-297	190	30	calves	calf	NOUN
alces-297	190	31	.	.	PUNCT
alces-297	191	1	the	the	DET
alces-297	191	2	open	open	ADJ
alces-297	191	3	circles	circle	NOUN
alces-297	191	4	are	be	AUX
alces-297	191	5	the	the	DET
alces-297	191	6	observed	observe	VERB
alces-297	191	7	moose	moose	NOUN
alces-297	191	8	harvest	harvest	NOUN
alces-297	191	9	and	and	CCONJ
alces-297	191	10	density	density	NOUN
alces-297	191	11	estimates	estimate	NOUN
alces-297	191	12	from	from	ADP
alces-297	191	13	13	13	NUM
alces-297	191	14	game	game	NOUN
alces-297	191	15	management	management	NOUN
alces-297	191	16	zones	zone	NOUN
alces-297	191	17	in	in	ADP
alces-297	191	18	north	north	NOUN
alces-297	191	19	-	-	PUNCT
alces-297	191	20	central	central	ADJ
alces-297	191	21	british	british	PROPN
alces-297	191	22	columbia	columbia	PROPN
alces-297	191	23	with	with	ADP
alces-297	191	24	stable	stable	ADJ
alces-297	191	25	moose	moose	NOUN
alces-297	191	26	populations	population	NOUN
alces-297	191	27	(	(	PUNCT
alces-297	191	28	from	from	ADP
alces-297	191	29	hatter	hatter	NOUN
alces-297	191	30	1999	1999	NUM
alces-297	191	31	)	)	PUNCT
alces-297	191	32	,	,	PUNCT
alces-297	191	33	and	and	CCONJ
alces-297	191	34	the	the	DET
alces-297	191	35	dashed	dash	VERB
alces-297	191	36	line	line	NOUN
alces-297	191	37	is	be	AUX
alces-297	191	38	the	the	DET
alces-297	191	39	fitted	fit	VERB
alces-297	191	40	regression	regression	NOUN
alces-297	191	41	.	.	PUNCT
alces-297	192	1	modelling	model	VERB
alces-297	192	2	sustained	sustained	ADJ
alces-297	192	3	yields	yield	NOUN
alces-297	192	4	for	for	ADP
alces-297	192	5	moose	moose	NOUN
alces-297	192	6	–	–	PUNCT
alces-297	192	7	hatter	hatter	NOUN
alces-297	192	8	.	.	PUNCT
alces-297	193	1	alces	alce	NOUN
alces-297	193	2	vol	vol	NOUN
alces-297	193	3	.	.	PROPN
alces-297	193	4	57	57	NUM
alces-297	193	5	,	,	PUNCT
alces-297	193	6	2021	2021	NUM
alces-297	193	7	122	122	NUM
alces-297	193	8	discussion	discussion	NOUN
alces-297	193	9	i	i	PRON
alces-297	193	10	used	use	VERB
alces-297	193	11	a	a	DET
alces-297	193	12	predator	predator	NOUN
alces-297	193	13	-	-	PUNCT
alces-297	193	14	prey	prey	NOUN
alces-297	193	15	model	model	NOUN
alces-297	193	16	to	to	PART
alces-297	193	17	investigate	investigate	VERB
alces-297	193	18	how	how	SCONJ
alces-297	193	19	predation	predation	NOUN
alces-297	193	20	may	may	AUX
alces-297	193	21	affect	affect	VERB
alces-297	193	22	sys	sys	NOUN
alces-297	193	23	and	and	CCONJ
alces-297	193	24	optimal	optimal	ADJ
alces-297	193	25	harvest	harvest	NOUN
alces-297	193	26	strategies	strategy	NOUN
alces-297	193	27	for	for	ADP
alces-297	193	28	moose	moose	NOUN
alces-297	193	29	in	in	ADP
alces-297	193	30	central	central	ADJ
alces-297	193	31	bc	bc	PROPN
alces-297	193	32	.	.	PUNCT
alces-297	194	1	i	i	PRON
alces-297	194	2	used	use	VERB
alces-297	194	3	a	a	DET
alces-297	194	4	modified	modify	VERB
alces-297	194	5	type	type	NOUN
alces-297	194	6	ii	ii	NOUN
alces-297	194	7	functional	functional	ADJ
alces-297	194	8	response	response	NOUN
alces-297	194	9	for	for	ADP
alces-297	194	10	wolves	wolf	NOUN
alces-297	194	11	(	(	PUNCT
alces-297	194	12	messier	messy	ADJ
alces-297	194	13	1994	1994	NUM
alces-297	194	14	)	)	PUNCT
alces-297	194	15	,	,	PUNCT
alces-297	194	16	a	a	DET
alces-297	194	17	ratio	ratio	NOUN
alces-297	194	18	dependent	dependent	ADJ
alces-297	194	19	wolf	wolf	PROPN
alces-297	194	20	numerical	numerical	ADJ
alces-297	194	21	response	response	NOUN
alces-297	194	22	(	(	PUNCT
alces-297	194	23	eberhardt	eberhardt	PROPN
alces-297	194	24	1997	1997	NUM
alces-297	194	25	)	)	PUNCT
alces-297	194	26	,	,	PUNCT
alces-297	194	27	and	and	CCONJ
alces-297	194	28	a	a	DET
alces-297	194	29	density	density	NOUN
alces-297	194	30	independent	independent	ADJ
alces-297	194	31	calf	calf	NOUN
alces-297	194	32	predation	predation	NOUN
alces-297	194	33	rate	rate	NOUN
alces-297	194	34	by	by	ADP
alces-297	194	35	bears	bear	NOUN
alces-297	194	36	.	.	PUNCT
alces-297	195	1	the	the	DET
alces-297	195	2	modelled	model	VERB
alces-297	195	3	predator	predator	NOUN
alces-297	195	4	-	-	PUNCT
alces-297	195	5	prey	prey	NOUN
alces-297	195	6	parameters	parameter	NOUN
alces-297	195	7	closely	closely	ADV
alces-297	195	8	matched	match	VERB
alces-297	195	9	the	the	DET
alces-297	195	10	observed	observe	VERB
alces-297	195	11	parameters	parameter	NOUN
alces-297	195	12	from	from	ADP
alces-297	195	13	the	the	DET
alces-297	195	14	pgsa	pgsa	NOUN
alces-297	195	15	when	when	SCONJ
alces-297	195	16	the	the	DET
alces-297	195	17	wolf	wolf	PROPN
alces-297	195	18	removal	removal	NOUN
alces-297	195	19	rate	rate	NOUN
alces-297	195	20	was	be	AUX
alces-297	195	21	31	31	NUM
alces-297	195	22	%	%	NOUN
alces-297	195	23	.	.	PUNCT
alces-297	196	1	modelled	model	VERB
alces-297	196	2	moose	moose	NOUN
alces-297	196	3	calf	calf	NOUN
alces-297	196	4	and	and	CCONJ
alces-297	196	5	annual	annual	ADJ
alces-297	196	6	mortality	mortality	NOUN
alces-297	196	7	rates	rate	NOUN
alces-297	196	8	were	be	AUX
alces-297	196	9	generally	generally	ADV
alces-297	196	10	consistent	consistent	ADJ
alces-297	196	11	with	with	ADP
alces-297	196	12	alaska	alaska	PROPN
alces-297	196	13	and	and	CCONJ
alces-297	196	14	yukon	yukon	PROPN
alces-297	196	15	studies	study	NOUN
alces-297	196	16	table	table	VERB
alces-297	196	17	4	4	NUM
alces-297	196	18	.	.	PUNCT
alces-297	196	19	optimal	optimal	ADJ
alces-297	196	20	harvest	harvest	NOUN
alces-297	196	21	rates	rate	NOUN
alces-297	196	22	based	base	VERB
alces-297	196	23	on	on	ADP
alces-297	196	24	moose	moose	NOUN
alces-297	196	25	and	and	CCONJ
alces-297	196	26	wolf	wolf	PROPN
alces-297	196	27	population	population	NOUN
alces-297	196	28	parameters	parameter	NOUN
alces-297	196	29	using	use	VERB
alces-297	196	30	wolf	wolf	PROPN
alces-297	196	31	removal	removal	NOUN
alces-297	196	32	rates	rate	NOUN
alces-297	196	33	from	from	ADP
alces-297	196	34	0	0	NUM
alces-297	196	35	to	to	PART
alces-297	196	36	40	40	NUM
alces-297	196	37	%	%	NOUN
alces-297	196	38	for	for	ADP
alces-297	196	39	the	the	DET
alces-297	196	40	pgsa	pgsa	NOUN
alces-297	196	41	in	in	ADP
alces-297	196	42	central	central	ADJ
alces-297	196	43	british	british	PROPN
alces-297	196	44	columbia	columbia	PROPN
alces-297	196	45	,	,	PUNCT
alces-297	196	46	canada	canada	PROPN
alces-297	196	47	.	.	PUNCT
alces-297	197	1	a.	a.	PROPN
alces-297	197	2	optimized	optimize	VERB
alces-297	197	3	harvest	harvest	NOUN
alces-297	197	4	rates	rate	NOUN
alces-297	197	5	for	for	ADP
alces-297	197	6	bulls	bull	NOUN
alces-297	197	7	,	,	PUNCT
alces-297	197	8	cows	cow	NOUN
alces-297	197	9	and	and	CCONJ
alces-297	197	10	calves	calf	NOUN
alces-297	197	11	.	.	PUNCT
alces-297	198	1	wolf	wolf	PROPN
alces-297	198	2	removal	removal	NOUN
alces-297	198	3	rate	rate	NOUN
alces-297	198	4	(	(	PUNCT
alces-297	198	5	%	%	NOUN
alces-297	198	6	)	)	PUNCT
alces-297	198	7	0	0	NUM
alces-297	198	8	10	10	NUM
alces-297	198	9	20	20	NUM
alces-297	198	10	25	25	NUM
alces-297	198	11	30	30	NUM
alces-297	198	12	35	35	NUM
alces-297	198	13	40	40	NUM
alces-297	198	14	wolves/1000	wolves/1000	NOUN
alces-297	198	15	km2	km2	PROPN
alces-297	198	16	5	5	NUM
alces-297	198	17	5	5	NUM
alces-297	198	18	8	8	NUM
alces-297	198	19	10	10	NUM
alces-297	198	20	15	15	NUM
alces-297	198	21	5	5	NUM
alces-297	198	22	0	0	NUM
alces-297	198	23	moose/1000	moose/1000	NOUN
alces-297	198	24	km2	km2	PROPN
alces-297	198	25	120	120	NUM
alces-297	198	26	166	166	NUM
alces-297	198	27	363	363	NUM
alces-297	198	28	609	609	NUM
alces-297	198	29	1400	1400	NUM
alces-297	198	30	1400	1400	NUM
alces-297	198	31	1400	1400	NUM
alces-297	198	32	harvest/1000	harvest/1000	NOUN
alces-297	198	33	km2	km2	PROPN
alces-297	198	34	10	10	NUM
alces-297	198	35	14	14	NUM
alces-297	198	36	31	31	NUM
alces-297	198	37	48	48	NUM
alces-297	198	38	130	130	NUM
alces-297	198	39	206	206	NUM
alces-297	198	40	245	245	NUM
alces-297	198	41	bulls/100	bulls/100	PROPN
alces-297	198	42	cows	cow	VERB
alces-297	198	43	50	50	NUM
alces-297	198	44	50	50	NUM
alces-297	198	45	30	30	NUM
alces-297	198	46	30	30	NUM
alces-297	198	47	30	30	NUM
alces-297	198	48	30	30	NUM
alces-297	198	49	30	30	NUM
alces-297	198	50	calves/100	calves/100	NOUN
alces-297	198	51	cows	cow	VERB
alces-297	198	52	26	26	NUM
alces-297	198	53	26	26	NUM
alces-297	198	54	31	31	NUM
alces-297	198	55	33	33	NUM
alces-297	198	56	38	38	NUM
alces-297	198	57	29	29	NUM
alces-297	198	58	24	24	NUM
alces-297	198	59	bull	bull	NOUN
alces-297	198	60	harvest	harvest	NOUN
alces-297	198	61	rate	rate	NOUN
alces-297	198	62	(	(	PUNCT
alces-297	198	63	%	%	NOUN
alces-297	198	64	)	)	PUNCT
alces-297	198	65	6	6	NUM
alces-297	198	66	6	6	NUM
alces-297	198	67	17	17	NUM
alces-297	198	68	18	18	NUM
alces-297	198	69	21	21	NUM
alces-297	198	70	18	18	NUM
alces-297	198	71	16	16	NUM
alces-297	198	72	cow	cow	NOUN
alces-297	198	73	harvest	harvest	NOUN
alces-297	198	74	rate	rate	NOUN
alces-297	198	75	(	(	PUNCT
alces-297	198	76	%	%	NOUN
alces-297	198	77	)	)	PUNCT
alces-297	198	78	0	0	NUM
alces-297	198	79	0	0	NUM
alces-297	198	80	0	0	NUM
alces-297	198	81	0	0	NUM
alces-297	198	82	0.2	0.2	NUM
alces-297	198	83	0.2	0.2	NUM
alces-297	198	84	0.2	0.2	NUM
alces-297	198	85	calf	calf	NOUN
alces-297	198	86	harvest	harvest	NOUN
alces-297	198	87	rate	rate	NOUN
alces-297	198	88	(	(	PUNCT
alces-297	198	89	%	%	NOUN
alces-297	198	90	)	)	PUNCT
alces-297	198	91	30	30	NUM
alces-297	198	92	28	28	NUM
alces-297	198	93	19	19	NUM
alces-297	198	94	15	15	NUM
alces-297	198	95	15	15	NUM
alces-297	198	96	34	34	NUM
alces-297	198	97	43	43	NUM
alces-297	198	98	moose	moose	NOUN
alces-297	198	99	harvest	harvest	NOUN
alces-297	198	100	rate	rate	NOUN
alces-297	198	101	(	(	PUNCT
alces-297	198	102	%	%	NOUN
alces-297	198	103	)	)	PUNCT
alces-297	198	104	8	8	NUM
alces-297	198	105	8	8	NUM
alces-297	198	106	8	8	NUM
alces-297	198	107	7	7	NUM
alces-297	198	108	8	8	NUM
alces-297	198	109	13	13	NUM
alces-297	198	110	15	15	NUM
alces-297	198	111	b.	b.	NOUN
alces-297	198	112	optimized	optimize	VERB
alces-297	198	113	bull	bull	NOUN
alces-297	198	114	and	and	CCONJ
alces-297	198	115	calf	calf	NOUN
alces-297	198	116	harvest	harvest	NOUN
alces-297	198	117	rates	rate	NOUN
alces-297	198	118	with	with	ADP
alces-297	198	119	cow	cow	NOUN
alces-297	198	120	harvest	harvest	NOUN
alces-297	198	121	rate	rate	NOUN
alces-297	198	122	=	=	PUNCT
alces-297	198	123	2	2	NUM
alces-297	198	124	%	%	NOUN
alces-297	198	125	.	.	PUNCT
alces-297	199	1	wolf	wolf	PROPN
alces-297	199	2	removal	removal	NOUN
alces-297	199	3	rate	rate	NOUN
alces-297	199	4	(	(	PUNCT
alces-297	199	5	%	%	NOUN
alces-297	199	6	)	)	PUNCT
alces-297	199	7	0	0	NUM
alces-297	199	8	10	10	NUM
alces-297	199	9	20	20	NUM
alces-297	199	10	25	25	NUM
alces-297	199	11	30	30	NUM
alces-297	199	12	35	35	NUM
alces-297	199	13	40	40	NUM
alces-297	199	14	wolves/1000	wolves/1000	NOUN
alces-297	199	15	km2	km2	PROPN
alces-297	199	16	4	4	NUM
alces-297	199	17	5	5	NUM
alces-297	199	18	8	8	NUM
alces-297	199	19	10	10	NUM
alces-297	199	20	15	15	NUM
alces-297	199	21	5	5	NUM
alces-297	199	22	0	0	NUM
alces-297	199	23	moose/1000	moose/1000	NOUN
alces-297	199	24	km2	km2	PROPN
alces-297	199	25	100	100	NUM
alces-297	199	26	161	161	NUM
alces-297	199	27	350	350	NUM
alces-297	199	28	585	585	NUM
alces-297	199	29	1400	1400	NUM
alces-297	199	30	1400	1400	NUM
alces-297	199	31	1400	1400	NUM
alces-297	199	32	harvest/1000	harvest/1000	NOUN
alces-297	199	33	km2	km2	PROPN
alces-297	199	34	10	10	NUM
alces-297	199	35	13	13	NUM
alces-297	199	36	30	30	NUM
alces-297	199	37	46	46	NUM
alces-297	199	38	123	123	NUM
alces-297	199	39	200	200	NUM
alces-297	199	40	239	239	NUM
alces-297	199	41	bulls/100	bulls/100	PROPN
alces-297	199	42	cows	cow	VERB
alces-297	199	43	50	50	NUM
alces-297	199	44	50	50	NUM
alces-297	199	45	30	30	NUM
alces-297	199	46	30	30	NUM
alces-297	199	47	30	30	NUM
alces-297	199	48	30	30	NUM
alces-297	199	49	30	30	NUM
alces-297	199	50	calves/100	calves/100	NOUN
alces-297	199	51	cows	cow	NOUN
alces-297	199	52	28	28	NUM
alces-297	199	53	32	32	NUM
alces-297	199	54	36	36	NUM
alces-297	199	55	38	38	NUM
alces-297	199	56	43	43	NUM
alces-297	199	57	34	34	NUM
alces-297	199	58	29	29	NUM
alces-297	199	59	bull	bull	NOUN
alces-297	199	60	harvest	harvest	NOUN
alces-297	199	61	rate	rate	NOUN
alces-297	199	62	(	(	PUNCT
alces-297	199	63	%	%	NOUN
alces-297	199	64	)	)	PUNCT
alces-297	199	65	9	9	NUM
alces-297	199	66	10	10	NUM
alces-297	199	67	21	21	NUM
alces-297	199	68	22	22	NUM
alces-297	199	69	25	25	NUM
alces-297	199	70	22	22	NUM
alces-297	199	71	20	20	NUM
alces-297	199	72	cow	cow	NOUN
alces-297	199	73	harvest	harvest	NOUN
alces-297	199	74	rate	rate	NOUN
alces-297	199	75	(	(	PUNCT
alces-297	199	76	%	%	NOUN
alces-297	199	77	)	)	PUNCT
alces-297	199	78	2	2	NUM
alces-297	199	79	2	2	NUM
alces-297	199	80	2	2	NUM
alces-297	199	81	2	2	NUM
alces-297	199	82	2	2	NUM
alces-297	199	83	2	2	NUM
alces-297	199	84	2	2	NUM
alces-297	199	85	calf	calf	NOUN
alces-297	199	86	harvest	harvest	NOUN
alces-297	199	87	rate	rate	NOUN
alces-297	199	88	(	(	PUNCT
alces-297	199	89	%	%	NOUN
alces-297	199	90	)	)	PUNCT
alces-297	199	91	27	27	NUM
alces-297	199	92	18	18	NUM
alces-297	199	93	9	9	NUM
alces-297	199	94	6	6	NUM
alces-297	199	95	6	6	NUM
alces-297	199	96	26	26	NUM
alces-297	199	97	35	35	NUM
alces-297	199	98	moose	moose	NOUN
alces-297	199	99	harvest	harvest	NOUN
alces-297	199	100	rate	rate	NOUN
alces-297	199	101	(	(	PUNCT
alces-297	199	102	%	%	NOUN
alces-297	199	103	)	)	PUNCT
alces-297	199	104	9	9	NUM
alces-297	199	105	7	7	NUM
alces-297	199	106	8	8	NUM
alces-297	199	107	7	7	NUM
alces-297	199	108	8	8	NUM
alces-297	199	109	12	12	NUM
alces-297	199	110	14	14	NUM
alces-297	199	111	c.	c.	NOUN
alces-297	199	112	optimized	optimize	VERB
alces-297	199	113	bull	bull	NOUN
alces-297	199	114	harvest	harvest	NOUN
alces-297	199	115	rates	rate	NOUN
alces-297	199	116	with	with	ADP
alces-297	199	117	bull	bull	NOUN
alces-297	199	118	-	-	PUNCT
alces-297	199	119	only	only	ADJ
alces-297	199	120	hunting	hunting	NOUN
alces-297	199	121	.	.	PUNCT
alces-297	200	1	wolf	wolf	PROPN
alces-297	200	2	removal	removal	NOUN
alces-297	200	3	rate	rate	NOUN
alces-297	200	4	(	(	PUNCT
alces-297	200	5	%	%	NOUN
alces-297	200	6	)	)	PUNCT
alces-297	200	7	0	0	NUM
alces-297	200	8	10	10	NUM
alces-297	200	9	20	20	NUM
alces-297	200	10	25	25	NUM
alces-297	200	11	30	30	NUM
alces-297	200	12	35	35	NUM
alces-297	200	13	40	40	NUM
alces-297	200	14	wolves/1000	wolves/1000	NOUN
alces-297	200	15	km2	km2	PROPN
alces-297	200	16	9	9	NUM
alces-297	200	17	10	10	NUM
alces-297	200	18	13	13	NUM
alces-297	200	19	18	18	NUM
alces-297	200	20	16	16	NUM
alces-297	200	21	6	6	NUM
alces-297	200	22	0	0	NUM
alces-297	200	23	moose/1000	moose/1000	NOUN
alces-297	200	24	km2	km2	PROPN
alces-297	200	25	229	229	NUM
alces-297	200	26	319	319	NUM
alces-297	200	27	563	563	NUM
alces-297	200	28	1040	1040	NUM
alces-297	200	29	1490	1490	NUM
alces-297	200	30	1606	1606	NUM
alces-297	200	31	1662	1662	NUM
alces-297	200	32	harvest/1000	harvest/1000	NOUN
alces-297	200	33	km2	km2	PROPN
alces-297	200	34	10	10	NUM
alces-297	200	35	14	14	NUM
alces-297	200	36	26	26	NUM
alces-297	200	37	50	50	NUM
alces-297	200	38	79	79	NUM
alces-297	200	39	81	81	NUM
alces-297	200	40	81	81	NUM
alces-297	200	41	bulls/100	bulls/100	PROPN
alces-297	200	42	cows	cow	VERB
alces-297	200	43	30	30	NUM
alces-297	200	44	30	30	NUM
alces-297	200	45	30	30	NUM
alces-297	200	46	30	30	NUM
alces-297	200	47	30	30	NUM
alces-297	200	48	30	30	NUM
alces-297	200	49	30	30	NUM
alces-297	200	50	calves/100	calves/100	NOUN
alces-297	200	51	cows	cow	VERB
alces-297	200	52	36	36	NUM
alces-297	200	53	37	37	NUM
alces-297	200	54	38	38	NUM
alces-297	200	55	39	39	NUM
alces-297	200	56	40	40	NUM
alces-297	200	57	34	34	NUM
alces-297	200	58	31	31	NUM
alces-297	200	59	bull	bull	NOUN
alces-297	200	60	harvest	harvest	NOUN
alces-297	200	61	rate	rate	NOUN
alces-297	200	62	(	(	PUNCT
alces-297	200	63	%	%	NOUN
alces-297	200	64	)	)	PUNCT
alces-297	200	65	19	19	NUM
alces-297	200	66	20	20	NUM
alces-297	200	67	20	20	NUM
alces-297	200	68	21	21	NUM
alces-297	200	69	22	22	NUM
alces-297	200	70	21	21	NUM
alces-297	200	71	20	20	NUM
alces-297	200	72	cow	cow	NOUN
alces-297	200	73	harvest	harvest	NOUN
alces-297	200	74	rate	rate	NOUN
alces-297	200	75	(	(	PUNCT
alces-297	200	76	%	%	NOUN
alces-297	200	77	)	)	PUNCT
alces-297	200	78	0	0	NUM
alces-297	200	79	0	0	NUM
alces-297	200	80	0	0	NUM
alces-297	200	81	0	0	NUM
alces-297	200	82	0	0	NUM
alces-297	200	83	0	0	NUM
alces-297	200	84	0	0	NUM
alces-297	200	85	calf	calf	NOUN
alces-297	200	86	harvest	harvest	NOUN
alces-297	200	87	rate	rate	NOUN
alces-297	200	88	(	(	PUNCT
alces-297	200	89	%	%	NOUN
alces-297	200	90	)	)	PUNCT
alces-297	200	91	0	0	NUM
alces-297	200	92	0	0	NUM
alces-297	200	93	0	0	NUM
alces-297	200	94	0	0	NUM
alces-297	200	95	0	0	NUM
alces-297	200	96	0	0	NUM
alces-297	200	97	0	0	NUM
alces-297	200	98	moose	moose	NOUN
alces-297	200	99	harvest	harvest	NOUN
alces-297	200	100	rate	rate	NOUN
alces-297	200	101	(	(	PUNCT
alces-297	200	102	%	%	NOUN
alces-297	200	103	)	)	PUNCT
alces-297	200	104	4	4	NUM
alces-297	200	105	4	4	NUM
alces-297	200	106	4	4	NUM
alces-297	200	107	5	5	NUM
alces-297	200	108	5	5	NUM
alces-297	200	109	5	5	NUM
alces-297	200	110	5	5	NUM
alces-297	200	111	alces	alce	NOUN
alces-297	200	112	vol	vol	NOUN
alces-297	200	113	.	.	PROPN
alces-297	200	114	57	57	NUM
alces-297	200	115	,	,	PUNCT
alces-297	200	116	2021	2021	NUM
alces-297	200	117	modelling	model	VERB
alces-297	200	118	sustained	sustain	VERB
alces-297	200	119	yields	yield	NOUN
alces-297	200	120	for	for	ADP
alces-297	200	121	moose	moose	NOUN
alces-297	200	122	–	–	PUNCT
alces-297	200	123	hatter	hatter	NOUN
alces-297	200	124	.	.	PUNCT
alces-297	201	1	123	123	NUM
alces-297	201	2	(	(	PUNCT
alces-297	201	3	boertje	boertje	NOUN
alces-297	201	4	et	et	PROPN
alces-297	201	5	al	al	PROPN
alces-297	201	6	.	.	PROPN
alces-297	201	7	2009	2009	NUM
alces-297	201	8	)	)	PUNCT
alces-297	201	9	,	,	PUNCT
alces-297	201	10	and	and	CCONJ
alces-297	201	11	modelled	model	VERB
alces-297	201	12	wolf	wolf	PROPN
alces-297	201	13	densities	density	NOUN
alces-297	201	14	were	be	AUX
alces-297	201	15	similar	similar	ADJ
alces-297	201	16	to	to	ADP
alces-297	201	17	those	those	PRON
alces-297	201	18	generated	generate	VERB
alces-297	201	19	from	from	ADP
alces-297	201	20	a	a	DET
alces-297	201	21	biomass	biomass	NOUN
alces-297	201	22	regression	regression	NOUN
alces-297	201	23	model	model	NOUN
alces-297	201	24	.	.	PUNCT
alces-297	202	1	the	the	DET
alces-297	202	2	modelled	model	VERB
alces-297	202	3	maximum	maximum	ADJ
alces-297	202	4	rate	rate	NOUN
alces-297	202	5	of	of	ADP
alces-297	202	6	increase	increase	NOUN
alces-297	202	7	for	for	ADP
alces-297	202	8	moose	moose	NOUN
alces-297	202	9	(	(	PUNCT
alces-297	202	10	λ	λ	X
alces-297	202	11	~	~	PUNCT
alces-297	202	12	1.3	1.3	NUM
alces-297	202	13	)	)	PUNCT
alces-297	202	14	was	be	AUX
alces-297	202	15	similar	similar	ADJ
alces-297	202	16	to	to	ADP
alces-297	202	17	1.35	1.35	NUM
alces-297	202	18	estimated	estimate	VERB
alces-297	202	19	by	by	ADP
alces-297	202	20	eberhardt	eberhardt	PROPN
alces-297	202	21	(	(	PUNCT
alces-297	202	22	1997	1997	NUM
alces-297	202	23	)	)	PUNCT
alces-297	202	24	,	,	PUNCT
alces-297	202	25	and	and	CCONJ
alces-297	202	26	unhunted	unhunted	ADJ
alces-297	202	27	adult	adult	NOUN
alces-297	202	28	sex	sex	NOUN
alces-297	202	29	ratios	ratio	NOUN
alces-297	202	30	(	(	PUNCT
alces-297	202	31	87–100	87–100	PROPN
alces-297	202	32	bulls/100	bulls/100	PROPN
alces-297	202	33	cows	cow	NOUN
alces-297	202	34	)	)	PUNCT
alces-297	202	35	were	be	AUX
alces-297	202	36	near	near	ADP
alces-297	202	37	parity	parity	NOUN
alces-297	202	38	as	as	SCONJ
alces-297	202	39	documented	document	VERB
alces-297	202	40	from	from	ADP
alces-297	202	41	published	publish	VERB
alces-297	202	42	studies	study	NOUN
alces-297	202	43	of	of	ADP
alces-297	202	44	naturally	naturally	ADV
alces-297	202	45	fluctuating	fluctuate	VERB
alces-297	202	46	moose	moose	NOUN
alces-297	202	47	populations	population	NOUN
alces-297	202	48	(	(	PUNCT
alces-297	202	49	timmermann	timmermann	NOUN
alces-297	202	50	1992	1992	NUM
alces-297	202	51	)	)	PUNCT
alces-297	202	52	.	.	PUNCT
alces-297	203	1	the	the	DET
alces-297	203	2	yield	yield	NOUN
alces-297	203	3	-	-	PUNCT
alces-297	203	4	density	density	NOUN
alces-297	203	5	curve	curve	NOUN
alces-297	203	6	for	for	ADP
alces-297	203	7	the	the	DET
alces-297	203	8	pgsa	pgsa	NOUN
alces-297	203	9	was	be	AUX
alces-297	203	10	comparable	comparable	ADJ
alces-297	203	11	to	to	ADP
alces-297	203	12	the	the	DET
alces-297	203	13	yield	yield	NOUN
alces-297	203	14	-	-	PUNCT
alces-297	203	15	density	density	NOUN
alces-297	203	16	curve	curve	NOUN
alces-297	203	17	for	for	ADP
alces-297	203	18	moose	moose	NOUN
alces-297	203	19	in	in	ADP
alces-297	203	20	north	north	PROPN
alces-297	203	21	central	central	ADJ
alces-297	203	22	bc	bc	PROPN
alces-297	203	23	(	(	PUNCT
alces-297	203	24	hatter	hatter	NOUN
alces-297	203	25	1999	1999	NUM
alces-297	203	26	)	)	PUNCT
alces-297	203	27	suggesting	suggest	VERB
alces-297	203	28	that	that	SCONJ
alces-297	203	29	the	the	DET
alces-297	203	30	sy	sy	PROPN
alces-297	203	31	curve	curve	NOUN
alces-297	203	32	for	for	ADP
alces-297	203	33	the	the	DET
alces-297	203	34	pgsa	pgsa	NOUN
alces-297	203	35	may	may	AUX
alces-297	203	36	be	be	AUX
alces-297	203	37	broadly	broadly	ADV
alces-297	203	38	applicable	applicable	ADJ
alces-297	203	39	to	to	ADP
alces-297	203	40	other	other	ADJ
alces-297	203	41	moose	moose	NOUN
alces-297	203	42	populations	population	NOUN
alces-297	203	43	in	in	ADP
alces-297	203	44	bc	bc	PROPN
alces-297	203	45	.	.	PROPN
alces-297	204	1	optimal	optimal	ADJ
alces-297	204	2	harvests	harvest	NOUN
alces-297	204	3	included	include	VERB
alces-297	204	4	a	a	DET
alces-297	204	5	moderate	moderate	ADJ
alces-297	204	6	harvest	harvest	NOUN
alces-297	204	7	of	of	ADP
alces-297	204	8	bulls	bull	NOUN
alces-297	204	9	(	(	PUNCT
alces-297	204	10	16–21	16–21	NUM
alces-297	204	11	%	%	NOUN
alces-297	204	12	)	)	PUNCT
alces-297	204	13	,	,	PUNCT
alces-297	204	14	a	a	DET
alces-297	204	15	nil	nil	NOUN
alces-297	204	16	-	-	PUNCT
alces-297	204	17	to	to	ADP
alces-297	204	18	-	-	PUNCT
alces-297	204	19	very	very	ADV
alces-297	204	20	low	low	ADJ
alces-297	204	21	harvest	harvest	NOUN
alces-297	204	22	of	of	ADP
alces-297	204	23	cows	cow	NOUN
alces-297	204	24	(	(	PUNCT
alces-297	204	25	0–0.2	0–0.2	NOUN
alces-297	204	26	%	%	NOUN
alces-297	204	27	)	)	PUNCT
alces-297	204	28	,	,	PUNCT
alces-297	204	29	and	and	CCONJ
alces-297	204	30	moderate	moderate	ADJ
alces-297	204	31	-	-	PUNCT
alces-297	204	32	to	to	ADP
alces-297	204	33	-	-	PUNCT
alces-297	204	34	high	high	ADJ
alces-297	204	35	harvests	harvest	NOUN
alces-297	204	36	of	of	ADP
alces-297	204	37	calves	calf	NOUN
alces-297	204	38	(	(	PUNCT
alces-297	204	39	15–43	15–43	NUM
alces-297	204	40	%	%	NOUN
alces-297	204	41	)	)	PUNCT
alces-297	204	42	when	when	SCONJ
alces-297	204	43	wolf	wolf	PROPN
alces-297	204	44	removal	removal	NOUN
alces-297	204	45	rates	rate	NOUN
alces-297	204	46	were	be	AUX
alces-297	204	47	≥	≥	NUM
alces-297	204	48	20	20	NUM
alces-297	204	49	%	%	NOUN
alces-297	204	50	.	.	PUNCT
alces-297	205	1	these	these	DET
alces-297	205	2	results	result	NOUN
alces-297	205	3	support	support	VERB
alces-297	205	4	previous	previous	ADJ
alces-297	205	5	findings	finding	NOUN
alces-297	205	6	from	from	ADP
alces-297	205	7	nilsen	nilsen	PROPN
alces-297	205	8	et	et	PROPN
alces-297	205	9	al	al	PROPN
alces-297	205	10	.	.	PROPN
alces-297	206	1	(	(	PUNCT
alces-297	206	2	2005	2005	NUM
alces-297	206	3	)	)	PUNCT
alces-297	206	4	and	and	CCONJ
alces-297	206	5	xu	xu	PROPN
alces-297	206	6	and	and	CCONJ
alces-297	206	7	boyce	boyce	PROPN
alces-297	206	8	(	(	PUNCT
alces-297	206	9	2010	2010	NUM
alces-297	206	10	)	)	PUNCT
alces-297	206	11	who	who	PRON
alces-297	206	12	also	also	ADV
alces-297	206	13	noted	note	VERB
alces-297	206	14	that	that	SCONJ
alces-297	206	15	optimal	optimal	ADJ
alces-297	206	16	moose	moose	NOUN
alces-297	206	17	harvests	harvest	NOUN
alces-297	206	18	in	in	ADP
alces-297	206	19	the	the	DET
alces-297	206	20	presence	presence	NOUN
alces-297	206	21	of	of	ADP
alces-297	206	22	predators	predator	NOUN
alces-297	206	23	involved	involve	VERB
alces-297	206	24	a	a	DET
alces-297	206	25	high	high	ADJ
alces-297	206	26	harvest	harvest	NOUN
alces-297	206	27	of	of	ADP
alces-297	206	28	bulls	bull	NOUN
alces-297	206	29	and	and	CCONJ
alces-297	206	30	calves	calf	NOUN
alces-297	206	31	with	with	ADP
alces-297	206	32	a	a	DET
alces-297	206	33	minimal	minimal	ADJ
alces-297	206	34	cow	cow	NOUN
alces-297	206	35	harvest	harvest	NOUN
alces-297	206	36	.	.	PUNCT
alces-297	207	1	xu	xu	PROPN
alces-297	207	2	and	and	CCONJ
alces-297	207	3	boyce	boyce	PROPN
alces-297	207	4	(	(	PUNCT
alces-297	207	5	2010	2010	NUM
alces-297	207	6	)	)	PUNCT
alces-297	207	7	found	find	VERB
alces-297	207	8	that	that	SCONJ
alces-297	207	9	harvest	harvest	NOUN
alces-297	207	10	rates	rate	NOUN
alces-297	207	11	to	to	PART
alces-297	207	12	optimize	optimize	VERB
alces-297	207	13	moose	moose	NOUN
alces-297	207	14	yields	yield	NOUN
alces-297	207	15	in	in	ADP
alces-297	207	16	alberta	alberta	PROPN
alces-297	207	17	included	include	VERB
alces-297	207	18	40–45	40–45	NUM
alces-297	207	19	%	%	NOUN
alces-297	207	20	of	of	ADP
alces-297	207	21	bulls	bull	NOUN
alces-297	207	22	,	,	PUNCT
alces-297	207	23	0.1–5	0.1–5	PROPN
alces-297	207	24	%	%	NOUN
alces-297	207	25	of	of	ADP
alces-297	207	26	cows	cow	NOUN
alces-297	207	27	,	,	PUNCT
alces-297	207	28	and	and	CCONJ
alces-297	207	29	35–40	35–40	NUM
alces-297	207	30	%	%	NOUN
alces-297	207	31	of	of	ADP
alces-297	207	32	calves	calf	NOUN
alces-297	207	33	.	.	PUNCT
alces-297	208	1	the	the	DET
alces-297	208	2	lower	low	ADJ
alces-297	208	3	optimal	optimal	ADJ
alces-297	208	4	harvest	harvest	NOUN
alces-297	208	5	rates	rate	NOUN
alces-297	208	6	for	for	ADP
alces-297	208	7	bulls	bull	NOUN
alces-297	208	8	in	in	ADP
alces-297	208	9	this	this	DET
alces-297	208	10	study	study	NOUN
alces-297	208	11	were	be	AUX
alces-297	208	12	likely	likely	ADJ
alces-297	208	13	due	due	ADP
alces-297	208	14	to	to	ADP
alces-297	208	15	the	the	DET
alces-297	208	16	harvest	harvest	NOUN
alces-297	208	17	management	management	NOUN
alces-297	208	18	constraints	constraint	NOUN
alces-297	208	19	placed	place	VERB
alces-297	208	20	on	on	ADP
alces-297	208	21	adult	adult	NOUN
alces-297	208	22	sex	sex	NOUN
alces-297	208	23	ratios	ratio	NOUN
alces-297	208	24	(	(	PUNCT
alces-297	208	25	i.e.	i.e.	X
alces-297	208	26	,	,	PUNCT
alces-297	208	27	≥	≥	NUM
alces-297	208	28	30	30	NUM
alces-297	208	29	bulls:100	bulls:100	PROPN
alces-297	208	30	cows	cow	NOUN
alces-297	208	31	for	for	ADP
alces-297	208	32	>	>	SYM
alces-297	208	33	200	200	NUM
alces-297	208	34	moose/1000	moose/1000	NOUN
alces-297	208	35	km2	km2	NOUN
alces-297	208	36	,	,	PUNCT
alces-297	208	37	and	and	CCONJ
alces-297	208	38	≥	≥	NUM
alces-297	208	39	50	50	NUM
alces-297	208	40	bulls:100	bulls:100	PROPN
alces-297	208	41	cows	cow	NOUN
alces-297	208	42	for	for	ADP
alces-297	208	43	≤	≤	ADJ
alces-297	208	44	200/1000	200/1000	NUM
alces-297	208	45	km2	km2	NOUN
alces-297	208	46	)	)	PUNCT
alces-297	208	47	.	.	PUNCT
alces-297	209	1	optimal	optimal	ADJ
alces-297	209	2	yields	yield	NOUN
alces-297	209	3	of	of	ADP
alces-297	209	4	bulls	bull	NOUN
alces-297	209	5	and	and	CCONJ
alces-297	209	6	calves	calf	NOUN
alces-297	209	7	,	,	PUNCT
alces-297	209	8	with	with	ADP
alces-297	209	9	a	a	DET
alces-297	209	10	cow	cow	NOUN
alces-297	209	11	harvest	harvest	NOUN
alces-297	209	12	of	of	ADP
alces-297	209	13	2	2	NUM
alces-297	209	14	%	%	NOUN
alces-297	209	15	were	be	AUX
alces-297	209	16	only	only	ADV
alces-297	209	17	slightly	slightly	ADV
alces-297	209	18	lower	low	ADJ
alces-297	209	19	than	than	ADP
alces-297	209	20	fully	fully	ADV
alces-297	209	21	optimized	optimize	VERB
alces-297	209	22	yields	yield	NOUN
alces-297	209	23	.	.	PUNCT
alces-297	210	1	some	some	DET
alces-297	210	2	advantages	advantage	NOUN
alces-297	210	3	of	of	ADP
alces-297	210	4	harvesting	harvest	VERB
alces-297	210	5	a	a	DET
alces-297	210	6	greater	great	ADJ
alces-297	210	7	proportion	proportion	NOUN
alces-297	210	8	of	of	ADP
alces-297	210	9	cows	cow	NOUN
alces-297	210	10	(	(	PUNCT
alces-297	210	11	i.e.	i.e.	X
alces-297	210	12	,	,	PUNCT
alces-297	210	13	2	2	NUM
alces-297	210	14	%	%	NOUN
alces-297	210	15	vs.	vs.	ADP
alces-297	210	16	0.2	0.2	NUM
alces-297	210	17	%	%	NOUN
alces-297	210	18	)	)	PUNCT
alces-297	210	19	include	include	VERB
alces-297	210	20	an	an	DET
alces-297	210	21	increased	increase	VERB
alces-297	210	22	sample	sample	NOUN
alces-297	210	23	size	size	NOUN
alces-297	210	24	for	for	ADP
alces-297	210	25	monitoring	monitor	VERB
alces-297	210	26	moose	moose	NOUN
alces-297	210	27	reproduction	reproduction	NOUN
alces-297	210	28	and	and	CCONJ
alces-297	210	29	nutritional	nutritional	ADJ
alces-297	210	30	status	status	NOUN
alces-297	210	31	(	(	PUNCT
alces-297	210	32	heard	hear	VERB
alces-297	210	33	et	et	PROPN
alces-297	210	34	al	al	PROPN
alces-297	210	35	.	.	PROPN
alces-297	210	36	1997	1997	NUM
alces-297	210	37	,	,	PUNCT
alces-297	210	38	boertje	boertje	PROPN
alces-297	210	39	et	et	PROPN
alces-297	210	40	al	al	PROPN
alces-297	210	41	.	.	PROPN
alces-297	210	42	2007	2007	NUM
alces-297	210	43	)	)	PUNCT
alces-297	210	44	,	,	PUNCT
alces-297	210	45	recovery	recovery	NOUN
alces-297	210	46	of	of	ADP
alces-297	210	47	low	low	ADJ
alces-297	210	48	bull	bull	NOUN
alces-297	210	49	:	:	PUNCT
alces-297	210	50	cow	cow	NOUN
alces-297	210	51	ratios	ratio	NOUN
alces-297	210	52	(	(	PUNCT
alces-297	210	53	young	young	ADJ
alces-297	210	54	and	and	CCONJ
alces-297	210	55	boertje	boertje	ADJ
alces-297	210	56	2008	2008	NUM
alces-297	210	57	)	)	PUNCT
alces-297	210	58	,	,	PUNCT
alces-297	210	59	and	and	CCONJ
alces-297	210	60	more	more	ADJ
alces-297	210	61	options	option	NOUN
alces-297	210	62	to	to	PART
alces-297	210	63	intensify	intensify	VERB
alces-297	210	64	moose	moose	NOUN
alces-297	210	65	management	management	NOUN
alces-297	210	66	at	at	ADP
alces-297	210	67	high	high	ADJ
alces-297	210	68	density	density	NOUN
alces-297	210	69	(	(	PUNCT
alces-297	210	70	young	young	PROPN
alces-297	210	71	et	et	PROPN
alces-297	210	72	al	al	PROPN
alces-297	210	73	.	.	PROPN
alces-297	210	74	2006	2006	NUM
alces-297	210	75	)	)	PUNCT
alces-297	210	76	.	.	PUNCT
alces-297	211	1	bull	bull	NOUN
alces-297	211	2	-	-	PUNCT
alces-297	211	3	only	only	ADV
alces-297	211	4	hunting	hunting	NOUN
alces-297	211	5	,	,	PUNCT
alces-297	211	6	however	however	ADV
alces-297	211	7	,	,	PUNCT
alces-297	211	8	may	may	AUX
alces-297	211	9	be	be	AUX
alces-297	211	10	preferable	preferable	ADJ
alces-297	211	11	for	for	ADP
alces-297	211	12	low	low	ADJ
alces-297	211	13	density	density	NOUN
alces-297	211	14	moose	moose	NOUN
alces-297	211	15	populations	population	NOUN
alces-297	211	16	limited	limit	VERB
alces-297	211	17	by	by	ADP
alces-297	211	18	predation	predation	NOUN
alces-297	211	19	(	(	PUNCT
alces-297	211	20	van	van	PROPN
alces-297	211	21	ballenberghe	ballenberghe	NOUN
alces-297	211	22	and	and	CCONJ
alces-297	211	23	dart	dart	NOUN
alces-297	211	24	1982	1982	NUM
alces-297	211	25	,	,	PUNCT
alces-297	211	26	environment	environment	NOUN
alces-297	211	27	yukon	yukon	PROPN
alces-297	211	28	2016	2016	NUM
alces-297	211	29	)	)	PUNCT
alces-297	211	30	.	.	PUNCT
alces-297	212	1	i	i	PRON
alces-297	212	2	found	find	VERB
alces-297	212	3	that	that	SCONJ
alces-297	212	4	bullonly	bullonly	ADV
alces-297	212	5	harvests	harvest	NOUN
alces-297	212	6	with	with	ADP
alces-297	212	7	low	low	ADJ
alces-297	212	8	wolf	wolf	NOUN
alces-297	212	9	removal	removal	NOUN
alces-297	212	10	rates	rate	NOUN
alces-297	212	11	(	(	PUNCT
alces-297	212	12	0–10	0–10	NOUN
alces-297	212	13	%	%	NOUN
alces-297	212	14	)	)	PUNCT
alces-297	212	15	provided	provide	VERB
alces-297	212	16	equivalent	equivalent	ADJ
alces-297	212	17	yields	yield	NOUN
alces-297	212	18	to	to	PART
alces-297	212	19	selective	selective	ADJ
alces-297	212	20	harvests	harvest	NOUN
alces-297	212	21	while	while	SCONJ
alces-297	212	22	maintaining	maintain	VERB
alces-297	212	23	higher	high	ADJ
alces-297	212	24	moose	moose	NOUN
alces-297	212	25	and	and	CCONJ
alces-297	212	26	wolf	wolf	NOUN
alces-297	212	27	densities	density	NOUN
alces-297	212	28	.	.	PUNCT
alces-297	213	1	wolf	wolf	PROPN
alces-297	213	2	control	control	PROPN
alces-297	213	3	studies	study	NOUN
alces-297	213	4	in	in	ADP
alces-297	213	5	alaska	alaska	PROPN
alces-297	213	6	and	and	CCONJ
alces-297	213	7	yukon	yukon	PROPN
alces-297	213	8	indicated	indicate	VERB
alces-297	213	9	that	that	SCONJ
alces-297	213	10	wolf	wolf	PROPN
alces-297	213	11	reductions	reduction	NOUN
alces-297	213	12	within	within	ADP
alces-297	213	13	low	low	ADJ
alces-297	213	14	-	-	PUNCT
alces-297	213	15	density	density	NOUN
alces-297	213	16	,	,	PUNCT
alces-297	213	17	predator	predator	NOUN
alces-297	213	18	-	-	PUNCT
alces-297	213	19	limited	limit	VERB
alces-297	213	20	moose	moose	NOUN
alces-297	213	21	populations	population	NOUN
alces-297	213	22	led	lead	VERB
alces-297	213	23	to	to	ADP
alces-297	213	24	elevated	elevated	ADJ
alces-297	213	25	moose	moose	NOUN
alces-297	213	26	and	and	CCONJ
alces-297	213	27	harvest	harvest	NOUN
alces-297	213	28	densities	density	NOUN
alces-297	213	29	followed	follow	VERB
alces-297	213	30	by	by	ADP
alces-297	213	31	elevated	elevated	ADJ
alces-297	213	32	wolf	wolf	NOUN
alces-297	213	33	densities	density	NOUN
alces-297	213	34	that	that	PRON
alces-297	213	35	equalled	equal	VERB
alces-297	213	36	or	or	CCONJ
alces-297	213	37	exceeded	exceed	VERB
alces-297	213	38	pre	pre	ADJ
alces-297	213	39	-	-	ADJ
alces-297	213	40	control	control	ADJ
alces-297	213	41	levels	level	NOUN
alces-297	213	42	(	(	PUNCT
alces-297	213	43	gasaway	gasaway	NOUN
alces-297	213	44	et	et	PROPN
alces-297	213	45	al	al	PROPN
alces-297	213	46	.	.	PROPN
alces-297	213	47	1992	1992	NUM
alces-297	213	48	,	,	PUNCT
alces-297	213	49	boertje	boertje	PROPN
alces-297	213	50	et	et	PROPN
alces-297	213	51	al	al	PROPN
alces-297	213	52	.	.	PROPN
alces-297	213	53	1996	1996	NUM
alces-297	213	54	)	)	PUNCT
alces-297	213	55	.	.	PUNCT
alces-297	214	1	modelled	model	VERB
alces-297	214	2	moose	moose	NOUN
alces-297	214	3	densities	density	NOUN
alces-297	214	4	and	and	CCONJ
alces-297	214	5	harvests	harvest	NOUN
alces-297	214	6	similarly	similarly	ADV
alces-297	214	7	increased	increase	VERB
alces-297	214	8	with	with	ADP
alces-297	214	9	intensified	intensify	VERB
alces-297	214	10	wolf	wolf	PROPN
alces-297	214	11	removals	removal	NOUN
alces-297	214	12	due	due	ADP
alces-297	214	13	to	to	ADP
alces-297	214	14	the	the	DET
alces-297	214	15	ratio	ratio	NOUN
alces-297	214	16	-	-	PUNCT
alces-297	214	17	dependent	dependent	ADJ
alces-297	214	18	numerical	numerical	ADJ
alces-297	214	19	response	response	NOUN
alces-297	214	20	which	which	PRON
alces-297	214	21	enabled	enable	VERB
alces-297	214	22	wolf	wolf	NOUN
alces-297	214	23	densities	density	NOUN
alces-297	214	24	to	to	PART
alces-297	214	25	increase	increase	VERB
alces-297	214	26	.	.	PUNCT
alces-297	215	1	for	for	ADP
alces-297	215	2	example	example	NOUN
alces-297	215	3	,	,	PUNCT
alces-297	215	4	moose	moose	NOUN
alces-297	215	5	densities	density	NOUN
alces-297	215	6	were	be	AUX
alces-297	215	7	elevated	elevate	VERB
alces-297	215	8	from	from	ADP
alces-297	215	9	207	207	NUM
alces-297	215	10	to	to	ADP
alces-297	215	11	1103	1103	NUM
alces-297	215	12	moose/1000	moose/1000	NOUN
alces-297	215	13	km2	km2	NOUN
alces-297	215	14	,	,	PUNCT
alces-297	215	15	harvests	harvest	NOUN
alces-297	215	16	from	from	ADP
alces-297	215	17	16	16	NUM
alces-297	215	18	to	to	PART
alces-297	215	19	84	84	NUM
alces-297	215	20	moose/1000	moose/1000	NOUN
alces-297	215	21	km2	km2	NOUN
alces-297	215	22	,	,	PUNCT
alces-297	215	23	and	and	CCONJ
alces-297	215	24	wolf	wolf	PROPN
alces-297	215	25	densities	density	NOUN
alces-297	215	26	from	from	ADP
alces-297	215	27	7	7	NUM
alces-297	215	28	to	to	PART
alces-297	215	29	12	12	NUM
alces-297	215	30	wolves/1000	wolves/1000	NOUN
alces-297	215	31	km2	km2	PROPN
alces-297	215	32	when	when	SCONJ
alces-297	215	33	wolf	wolf	PROPN
alces-297	215	34	removal	removal	NOUN
alces-297	215	35	rates	rate	NOUN
alces-297	215	36	were	be	AUX
alces-297	215	37	increased	increase	VERB
alces-297	215	38	from	from	ADP
alces-297	215	39	10	10	NUM
alces-297	215	40	to	to	PART
alces-297	215	41	30	30	NUM
alces-297	215	42	%	%	NOUN
alces-297	215	43	.	.	PUNCT
alces-297	216	1	the	the	DET
alces-297	216	2	modelled	model	VERB
alces-297	216	3	wolf	wolf	PROPN
alces-297	216	4	population	population	NOUN
alces-297	216	5	was	be	AUX
alces-297	216	6	able	able	ADJ
alces-297	216	7	to	to	PART
alces-297	216	8	sustain	sustain	VERB
alces-297	216	9	removal	removal	NOUN
alces-297	216	10	rates	rate	NOUN
alces-297	216	11	up	up	ADP
alces-297	216	12	to	to	PART
alces-297	216	13	31	31	NUM
alces-297	216	14	%	%	NOUN
alces-297	216	15	which	which	PRON
alces-297	216	16	was	be	AUX
alces-297	216	17	consistent	consistent	ADJ
alces-297	216	18	with	with	ADP
alces-297	216	19	adams	adams	PROPN
alces-297	216	20	et	et	PROPN
alces-297	216	21	al	al	PROPN
alces-297	216	22	.	.	PROPN
alces-297	217	1	(	(	PUNCT
alces-297	217	2	2008	2008	NUM
alces-297	217	3	)	)	PUNCT
alces-297	217	4	who	who	PRON
alces-297	217	5	analysed	analyse	VERB
alces-297	217	6	information	information	NOUN
alces-297	217	7	from	from	ADP
alces-297	217	8	39	39	NUM
alces-297	217	9	north	north	ADJ
alces-297	217	10	american	american	ADJ
alces-297	217	11	wolf	wolf	NOUN
alces-297	217	12	populations	population	NOUN
alces-297	217	13	and	and	CCONJ
alces-297	217	14	determined	determine	VERB
alces-297	217	15	that	that	SCONJ
alces-297	217	16	populations	population	NOUN
alces-297	217	17	were	be	AUX
alces-297	217	18	able	able	ADJ
alces-297	217	19	to	to	PART
alces-297	217	20	compensate	compensate	VERB
alces-297	217	21	for	for	ADP
alces-297	217	22	removal	removal	NOUN
alces-297	217	23	rates	rate	NOUN
alces-297	217	24	up	up	ADV
alces-297	217	25	to	to	PART
alces-297	217	26	29	29	NUM
alces-297	217	27	%	%	NOUN
alces-297	217	28	.	.	PUNCT
alces-297	218	1	while	while	SCONJ
alces-297	218	2	the	the	DET
alces-297	218	3	predator	predator	NOUN
alces-297	218	4	-	-	PUNCT
alces-297	218	5	prey	prey	NOUN
alces-297	218	6	model	model	NOUN
alces-297	218	7	appeared	appear	VERB
alces-297	218	8	to	to	PART
alces-297	218	9	provide	provide	VERB
alces-297	218	10	a	a	DET
alces-297	218	11	reasonable	reasonable	ADJ
alces-297	218	12	portrait	portrait	NOUN
alces-297	218	13	of	of	ADP
alces-297	218	14	wolfmoose	wolfmoose	NOUN
alces-297	218	15	-	-	PUNCT
alces-297	218	16	bear	bear	NOUN
alces-297	218	17	dynamics	dynamic	NOUN
alces-297	218	18	in	in	ADP
alces-297	218	19	the	the	DET
alces-297	218	20	pgsa	pgsa	NOUN
alces-297	218	21	from	from	ADP
alces-297	218	22	1987	1987	NUM
alces-297	218	23	to	to	ADP
alces-297	218	24	1998	1998	NUM
alces-297	218	25	,	,	PUNCT
alces-297	218	26	increasing	increase	VERB
alces-297	218	27	model	model	NOUN
alces-297	218	28	realism	realism	NOUN
alces-297	218	29	would	would	AUX
alces-297	218	30	have	have	AUX
alces-297	218	31	helped	help	VERB
alces-297	218	32	to	to	PART
alces-297	218	33	validate	validate	VERB
alces-297	218	34	the	the	DET
alces-297	218	35	model	model	NOUN
alces-297	218	36	.	.	PUNCT
alces-297	219	1	several	several	ADJ
alces-297	219	2	studies	study	NOUN
alces-297	219	3	have	have	AUX
alces-297	219	4	shown	show	VERB
alces-297	219	5	that	that	SCONJ
alces-297	219	6	winter	winter	NOUN
alces-297	219	7	kill	kill	NOUN
alces-297	219	8	rates	rate	NOUN
alces-297	219	9	by	by	ADP
alces-297	219	10	individual	individual	ADJ
alces-297	219	11	wolves	wolf	NOUN
alces-297	219	12	are	be	AUX
alces-297	219	13	inversely	inversely	ADV
alces-297	219	14	related	relate	VERB
alces-297	219	15	to	to	ADP
alces-297	219	16	pack	pack	NOUN
alces-297	219	17	size	size	NOUN
alces-297	219	18	,	,	PUNCT
alces-297	219	19	and	and	CCONJ
alces-297	219	20	that	that	DET
alces-297	219	21	wolf	wolf	PROPN
alces-297	219	22	predation	predation	NOUN
alces-297	219	23	should	should	AUX
alces-297	219	24	be	be	AUX
alces-297	219	25	modelled	model	VERB
alces-297	219	26	by	by	ADP
alces-297	219	27	the	the	DET
alces-297	219	28	number	number	NOUN
alces-297	219	29	and	and	CCONJ
alces-297	219	30	size	size	NOUN
alces-297	219	31	of	of	ADP
alces-297	219	32	packs	pack	NOUN
alces-297	219	33	(	(	PUNCT
alces-297	219	34	ballard	ballard	PROPN
alces-297	219	35	et	et	PROPN
alces-297	219	36	al	al	PROPN
alces-297	219	37	.	.	PROPN
alces-297	219	38	1987	1987	NUM
alces-297	219	39	,	,	PUNCT
alces-297	219	40	mcnay	mcnay	PROPN
alces-297	219	41	and	and	CCONJ
alces-297	219	42	delong	delong	PROPN
alces-297	219	43	1998	1998	NUM
alces-297	219	44	,	,	PUNCT
alces-297	219	45	hayes	hayes	PROPN
alces-297	219	46	et	et	PROPN
alces-297	219	47	al	al	PROPN
alces-297	219	48	.	.	PROPN
alces-297	219	49	2000	2000	NUM
alces-297	219	50	)	)	PUNCT
alces-297	219	51	.	.	PUNCT
alces-297	220	1	adams	adams	PROPN
alces-297	220	2	et	et	PROPN
alces-297	220	3	al	al	PROPN
alces-297	220	4	.	.	PROPN
alces-297	221	1	(	(	PUNCT
alces-297	221	2	2008	2008	NUM
alces-297	221	3	)	)	PUNCT
alces-297	221	4	discussed	discuss	VERB
alces-297	221	5	how	how	SCONJ
alces-297	221	6	wolves	wolf	NOUN
alces-297	221	7	adjust	adjust	VERB
alces-297	221	8	dispersal	dispersal	NOUN
alces-297	221	9	rates	rate	NOUN
alces-297	221	10	as	as	ADP
alces-297	221	11	a	a	DET
alces-297	221	12	primary	primary	ADJ
alces-297	221	13	mechanism	mechanism	NOUN
alces-297	221	14	to	to	ADP
alces-297	221	15	modelling	model	VERB
alces-297	221	16	sustained	sustained	ADJ
alces-297	221	17	yields	yield	NOUN
alces-297	221	18	for	for	ADP
alces-297	221	19	moose	moose	NOUN
alces-297	221	20	–	–	PUNCT
alces-297	221	21	hatter	hatter	NOUN
alces-297	221	22	.	.	PUNCT
alces-297	222	1	alces	alce	NOUN
alces-297	222	2	vol	vol	NOUN
alces-297	222	3	.	.	PROPN
alces-297	223	1	57	57	NUM
alces-297	223	2	,	,	PUNCT
alces-297	223	3	2021	2021	NUM
alces-297	223	4	124	124	NUM
alces-297	223	5	compensate	compensate	NOUN
alces-297	223	6	for	for	ADP
alces-297	223	7	human	human	ADJ
alces-297	223	8	harvest	harvest	NOUN
alces-297	223	9	which	which	PRON
alces-297	223	10	also	also	ADV
alces-297	223	11	could	could	AUX
alces-297	223	12	be	be	AUX
alces-297	223	13	modelled	model	VERB
alces-297	223	14	.	.	PUNCT
alces-297	224	1	most	most	ADJ
alces-297	224	2	northern	northern	ADJ
alces-297	224	3	wolf	wolf	NOUN
alces-297	224	4	-	-	PUNCT
alces-297	224	5	ungulate	ungulate	ADJ
alces-297	224	6	studies	study	NOUN
alces-297	224	7	have	have	AUX
alces-297	224	8	identified	identify	VERB
alces-297	224	9	stochastic	stochastic	ADJ
alces-297	224	10	weather	weather	NOUN
alces-297	224	11	events	event	NOUN
alces-297	224	12	as	as	ADP
alces-297	224	13	a	a	DET
alces-297	224	14	significant	significant	ADJ
alces-297	224	15	component	component	NOUN
alces-297	224	16	in	in	ADP
alces-297	224	17	predator	predator	NOUN
alces-297	224	18	-	-	PUNCT
alces-297	224	19	prey	prey	NOUN
alces-297	224	20	systems	system	NOUN
alces-297	224	21	(	(	PUNCT
alces-297	224	22	gasaway	gasaway	NOUN
alces-297	224	23	et	et	PROPN
alces-297	224	24	al	al	PROPN
alces-297	224	25	.	.	PROPN
alces-297	224	26	1983	1983	NUM
alces-297	224	27	gasaway	gasaway	NOUN
alces-297	224	28	et	et	PROPN
alces-297	224	29	al	al	PROPN
alces-297	224	30	.	.	PROPN
alces-297	224	31	1992	1992	NUM
alces-297	224	32	,	,	PUNCT
alces-297	224	33	boertje	boertje	NOUN
alces-297	224	34	et	et	NOUN
alces-297	224	35	al.1996	al.1996	PROPN
alces-297	224	36	,	,	PUNCT
alces-297	224	37	2009	2009	NUM
alces-297	224	38	,	,	PUNCT
alces-297	224	39	ballard	ballard	NOUN
alces-297	224	40	and	and	CCONJ
alces-297	224	41	van	van	PROPN
alces-297	224	42	ballenberghe	ballenberghe	NOUN
alces-297	224	43	1998	1998	NUM
alces-297	224	44	,	,	PUNCT
alces-297	224	45	mcnay	mcnay	PROPN
alces-297	224	46	and	and	CCONJ
alces-297	224	47	delong	delong	PROPN
alces-297	224	48	1998	1998	NUM
alces-297	224	49	)	)	PUNCT
alces-297	224	50	.	.	PUNCT
alces-297	225	1	i	i	PRON
alces-297	225	2	did	do	AUX
alces-297	225	3	not	not	PART
alces-297	225	4	model	model	VERB
alces-297	225	5	stochastic	stochastic	ADJ
alces-297	225	6	predator	predator	NOUN
alces-297	225	7	-	-	PUNCT
alces-297	225	8	prey	prey	NOUN
alces-297	225	9	-	-	PUNCT
alces-297	225	10	weather	weather	NOUN
alces-297	225	11	interactions	interaction	NOUN
alces-297	225	12	because	because	SCONJ
alces-297	225	13	many	many	ADJ
alces-297	225	14	parameters	parameter	NOUN
alces-297	225	15	in	in	ADP
alces-297	225	16	predator	predator	NOUN
alces-297	225	17	-	-	PUNCT
alces-297	225	18	prey	prey	NOUN
alces-297	225	19	models	model	NOUN
alces-297	225	20	are	be	AUX
alces-297	225	21	influenced	influence	VERB
alces-297	225	22	by	by	ADP
alces-297	225	23	weather	weather	NOUN
alces-297	225	24	which	which	PRON
alces-297	225	25	greatly	greatly	ADV
alces-297	225	26	increases	increase	VERB
alces-297	225	27	model	model	NOUN
alces-297	225	28	complexity	complexity	NOUN
alces-297	225	29	,	,	PUNCT
alces-297	225	30	and	and	CCONJ
alces-297	225	31	because	because	SCONJ
alces-297	225	32	quantitative	quantitative	ADJ
alces-297	225	33	measures	measure	NOUN
alces-297	225	34	of	of	ADP
alces-297	225	35	imprecision	imprecision	NOUN
alces-297	225	36	(	(	PUNCT
alces-297	225	37	ses	ses	PROPN
alces-297	225	38	)	)	PUNCT
alces-297	225	39	were	be	AUX
alces-297	225	40	difficult	difficult	ADJ
alces-297	225	41	to	to	PART
alces-297	225	42	parameterize	parameterize	VERB
alces-297	225	43	.	.	PUNCT
alces-297	226	1	however	however	ADV
alces-297	226	2	,	,	PUNCT
alces-297	226	3	xu	xu	PROPN
alces-297	226	4	and	and	CCONJ
alces-297	226	5	boyce	boyce	PROPN
alces-297	226	6	(	(	PUNCT
alces-297	226	7	2010	2010	NUM
alces-297	226	8	)	)	PUNCT
alces-297	226	9	cautioned	caution	VERB
alces-297	226	10	that	that	SCONJ
alces-297	226	11	predation	predation	NOUN
alces-297	226	12	and	and	CCONJ
alces-297	226	13	stochastic	stochastic	ADJ
alces-297	226	14	weather	weather	NOUN
alces-297	226	15	events	event	NOUN
alces-297	226	16	can	can	AUX
alces-297	226	17	drive	drive	VERB
alces-297	226	18	moose	moose	NOUN
alces-297	226	19	populations	population	NOUN
alces-297	226	20	to	to	ADP
alces-297	226	21	low	low	ADJ
alces-297	226	22	levels	level	NOUN
alces-297	226	23	and	and	CCONJ
alces-297	226	24	negatively	negatively	ADV
alces-297	226	25	influence	influence	NOUN
alces-297	226	26	sys	sys	PROPN
alces-297	226	27	.	.	PUNCT
alces-297	227	1	a	a	DET
alces-297	227	2	growing	grow	VERB
alces-297	227	3	number	number	NOUN
alces-297	227	4	of	of	ADP
alces-297	227	5	studies	study	NOUN
alces-297	227	6	suggest	suggest	VERB
alces-297	227	7	ratio	ratio	NOUN
alces-297	227	8	-	-	PUNCT
alces-297	227	9	dependence	dependence	NOUN
alces-297	227	10	may	may	AUX
alces-297	227	11	be	be	AUX
alces-297	227	12	common	common	ADJ
alces-297	227	13	in	in	ADP
alces-297	227	14	wolf	wolf	NOUN
alces-297	227	15	-	-	PUNCT
alces-297	227	16	ungulate	ungulate	ADJ
alces-297	227	17	systems	system	NOUN
alces-297	227	18	(	(	PUNCT
alces-297	227	19	hebblewhite	hebblewhite	NOUN
alces-297	227	20	2013	2013	NUM
alces-297	227	21	)	)	PUNCT
alces-297	227	22	.	.	PUNCT
alces-297	228	1	i	i	PRON
alces-297	228	2	used	use	VERB
alces-297	228	3	a	a	DET
alces-297	228	4	ratio	ratio	NOUN
alces-297	228	5	-	-	PUNCT
alces-297	228	6	dependent	dependent	ADJ
alces-297	228	7	,	,	PUNCT
alces-297	228	8	wolf	wolf	PROPN
alces-297	228	9	numerical	numerical	ADJ
alces-297	228	10	response	response	NOUN
alces-297	228	11	(	(	PUNCT
alces-297	228	12	eberhardt	eberhardt	PROPN
alces-297	228	13	1997	1997	NUM
alces-297	228	14	)	)	PUNCT
alces-297	228	15	with	with	ADP
alces-297	228	16	an	an	DET
alces-297	228	17	equilibrium	equilibrium	NOUN
alces-297	228	18	ratio	ratio	NOUN
alces-297	228	19	of	of	ADP
alces-297	228	20	25	25	NUM
alces-297	228	21	moose	moose	NOUN
alces-297	228	22	/	/	SYM
alces-297	228	23	wolf	wolf	NOUN
alces-297	228	24	.	.	PUNCT
alces-297	229	1	studies	study	NOUN
alces-297	229	2	of	of	ADP
alces-297	229	3	moose	moose	NOUN
alces-297	229	4	:	:	PUNCT
alces-297	229	5	wolf	wolf	PROPN
alces-297	229	6	ratios	ratio	NOUN
alces-297	229	7	during	during	ADP
alces-297	229	8	winter	winter	NOUN
alces-297	229	9	suggest	suggest	VERB
alces-297	229	10	moose	moose	NOUN
alces-297	229	11	densities	density	NOUN
alces-297	229	12	may	may	AUX
alces-297	229	13	stabilize	stabilize	VERB
alces-297	229	14	with	with	ADP
alces-297	229	15	20–30	20–30	NUM
alces-297	229	16	moose	moose	NOUN
alces-297	229	17	/	/	SYM
alces-297	229	18	wolf	wolf	NOUN
alces-297	229	19	.	.	PUNCT
alces-297	230	1	gasaway	gasaway	PROPN
alces-297	230	2	et	et	PROPN
alces-297	230	3	al	al	PROPN
alces-297	230	4	.	.	PROPN
alces-297	230	5	(	(	PUNCT
alces-297	230	6	1983	1983	NUM
alces-297	230	7	)	)	PUNCT
alces-297	230	8	summarized	summarize	VERB
alces-297	230	9	these	these	DET
alces-297	230	10	studies	study	NOUN
alces-297	230	11	and	and	CCONJ
alces-297	230	12	identified	identify	VERB
alces-297	230	13	3	3	NUM
alces-297	230	14	general	general	ADJ
alces-297	230	15	categories	category	NOUN
alces-297	230	16	of	of	ADP
alces-297	230	17	moose	moose	NOUN
alces-297	230	18	/	/	SYM
alces-297	230	19	wolf	wolf	NOUN
alces-297	230	20	relationships	relationship	NOUN
alces-297	230	21	:	:	PUNCT
alces-297	230	22	predation	predation	NOUN
alces-297	230	23	was	be	AUX
alces-297	230	24	sufficient	sufficient	ADJ
alces-297	230	25	to	to	PART
alces-297	230	26	cause	cause	VERB
alces-297	230	27	a	a	DET
alces-297	230	28	decline	decline	NOUN
alces-297	230	29	in	in	ADP
alces-297	230	30	moose	moose	NOUN
alces-297	230	31	abundance	abundance	NOUN
alces-297	230	32	at	at	ADP
alces-297	230	33	<	<	X
alces-297	230	34	20	20	NUM
alces-297	230	35	moose/	moose/	NUM
alces-297	230	36	wolf	wolf	NOUN
alces-297	230	37	,	,	PUNCT
alces-297	230	38	appeared	appear	VERB
alces-297	230	39	to	to	PART
alces-297	230	40	control	control	NOUN
alces-297	230	41	moose	moose	NOUN
alces-297	230	42	numbers	number	NOUN
alces-297	230	43	at	at	ADP
alces-297	230	44	20–30	20–30	NUM
alces-297	230	45	moose	moose	NOUN
alces-297	230	46	/	/	SYM
alces-297	230	47	wolf	wolf	NOUN
alces-297	230	48	,	,	PUNCT
alces-297	230	49	and	and	CCONJ
alces-297	230	50	was	be	AUX
alces-297	230	51	insufficient	insufficient	ADJ
alces-297	230	52	to	to	PART
alces-297	230	53	limit	limit	VERB
alces-297	230	54	growth	growth	NOUN
alces-297	230	55	at	at	ADP
alces-297	230	56	>	>	X
alces-297	230	57	30	30	NUM
alces-297	230	58	moose	moose	NOUN
alces-297	230	59	/	/	SYM
alces-297	230	60	wolf	wolf	NOUN
alces-297	230	61	.	.	PUNCT
alces-297	231	1	person	person	NOUN
alces-297	231	2	et	et	PROPN
alces-297	231	3	al	al	PROPN
alces-297	231	4	.	.	PROPN
alces-297	232	1	(	(	PUNCT
alces-297	232	2	2001	2001	NUM
alces-297	232	3	)	)	PUNCT
alces-297	232	4	and	and	CCONJ
alces-297	232	5	bowyer	bowyer	PROPN
alces-297	232	6	et	et	PROPN
alces-297	232	7	al	al	PROPN
alces-297	232	8	.	.	PROPN
alces-297	233	1	(	(	PUNCT
alces-297	233	2	2013	2013	NUM
alces-297	233	3	)	)	PUNCT
alces-297	233	4	,	,	PUNCT
alces-297	233	5	however	however	ADV
alces-297	233	6	,	,	PUNCT
alces-297	233	7	cautioned	caution	VERB
alces-297	233	8	against	against	ADP
alces-297	233	9	even	even	ADV
alces-297	233	10	a	a	DET
alces-297	233	11	general	general	ADJ
alces-297	233	12	interpretation	interpretation	NOUN
alces-297	233	13	of	of	ADP
alces-297	233	14	such	such	ADJ
alces-297	233	15	ratios	ratio	NOUN
alces-297	233	16	for	for	ADP
alces-297	233	17	interpreting	interpret	VERB
alces-297	233	18	impacts	impact	NOUN
alces-297	233	19	of	of	ADP
alces-297	233	20	wolf	wolf	PROPN
alces-297	233	21	predation	predation	NOUN
alces-297	233	22	on	on	ADP
alces-297	233	23	moose	moose	NOUN
alces-297	233	24	.	.	PUNCT
alces-297	234	1	the	the	DET
alces-297	234	2	predator	predator	NOUN
alces-297	234	3	-	-	PUNCT
alces-297	234	4	prey	prey	NOUN
alces-297	234	5	model	model	NOUN
alces-297	234	6	was	be	AUX
alces-297	234	7	based	base	VERB
alces-297	234	8	on	on	ADP
alces-297	234	9	moose	moose	NOUN
alces-297	234	10	-	-	PUNCT
alces-297	234	11	wolf	wolf	NOUN
alces-297	234	12	-	-	PUNCT
alces-297	234	13	bear	bear	NOUN
alces-297	234	14	relationships	relationship	NOUN
alces-297	234	15	in	in	ADP
alces-297	234	16	the	the	DET
alces-297	234	17	pgsa	pgsa	NOUN
alces-297	234	18	prior	prior	ADV
alces-297	234	19	to	to	ADP
alces-297	234	20	the	the	DET
alces-297	234	21	moose	moose	NOUN
alces-297	234	22	population	population	NOUN
alces-297	234	23	decline	decline	NOUN
alces-297	234	24	in	in	ADP
alces-297	234	25	the	the	DET
alces-297	234	26	early	early	ADJ
alces-297	234	27	2000s	2000s	NUM
alces-297	234	28	.	.	PUNCT
alces-297	235	1	the	the	DET
alces-297	235	2	decline	decline	NOUN
alces-297	235	3	coincided	coincide	VERB
alces-297	235	4	with	with	ADP
alces-297	235	5	a	a	DET
alces-297	235	6	mountain	mountain	NOUN
alces-297	235	7	pine	pine	NOUN
alces-297	235	8	beetle	beetle	NOUN
alces-297	235	9	(	(	PUNCT
alces-297	235	10	dendroctonus	dendroctonus	NOUN
alces-297	235	11	ponderosae	ponderosae	NOUN
alces-297	235	12	)	)	PUNCT
alces-297	235	13	outbreak	outbreak	NOUN
alces-297	235	14	where	where	SCONJ
alces-297	235	15	habitat	habitat	NOUN
alces-297	235	16	changes	change	NOUN
alces-297	235	17	and	and	CCONJ
alces-297	235	18	increased	increase	VERB
alces-297	235	19	salvage	salvage	NOUN
alces-297	235	20	logging	logging	NOUN
alces-297	235	21	and	and	CCONJ
alces-297	235	22	road	road	NOUN
alces-297	235	23	building	building	NOUN
alces-297	235	24	may	may	AUX
alces-297	235	25	have	have	AUX
alces-297	235	26	resulted	result	VERB
alces-297	235	27	in	in	ADP
alces-297	235	28	greater	great	ADJ
alces-297	235	29	vulnerability	vulnerability	NOUN
alces-297	235	30	to	to	AUX
alces-297	235	31	moose	moose	VERB
alces-297	235	32	from	from	ADP
alces-297	235	33	human	human	ADJ
alces-297	235	34	harvest	harvest	NOUN
alces-297	235	35	and	and	CCONJ
alces-297	235	36	predation	predation	NOUN
alces-297	235	37	,	,	PUNCT
alces-297	235	38	while	while	SCONJ
alces-297	235	39	elevating	elevate	VERB
alces-297	235	40	nutritional	nutritional	ADJ
alces-297	235	41	constraints	constraint	NOUN
alces-297	235	42	and	and	CCONJ
alces-297	235	43	health	health	NOUN
alces-297	235	44	/	/	SYM
alces-297	235	45	disease	disease	NOUN
alces-297	235	46	concerns	concern	NOUN
alces-297	235	47	(	(	PUNCT
alces-297	235	48	kuzyk	kuzyk	VERB
alces-297	235	49	and	and	CCONJ
alces-297	235	50	heard	hear	VERB
alces-297	235	51	2014	2014	NUM
alces-297	235	52	)	)	PUNCT
alces-297	235	53	.	.	PUNCT
alces-297	236	1	while	while	SCONJ
alces-297	236	2	disease	disease	NOUN
alces-297	236	3	was	be	AUX
alces-297	236	4	not	not	PART
alces-297	236	5	considered	consider	VERB
alces-297	236	6	a	a	DET
alces-297	236	7	substantive	substantive	ADJ
alces-297	236	8	cause	cause	NOUN
alces-297	236	9	of	of	ADP
alces-297	236	10	moose	moose	NOUN
alces-297	236	11	mortality	mortality	NOUN
alces-297	236	12	in	in	ADP
alces-297	236	13	british	british	PROPN
alces-297	236	14	columbia	columbia	PROPN
alces-297	236	15	from	from	ADP
alces-297	236	16	2012	2012	NUM
alces-297	236	17	to	to	ADP
alces-297	236	18	2019	2019	NUM
alces-297	236	19	(	(	PUNCT
alces-297	236	20	kuzyk	kuzyk	VERB
alces-297	236	21	et	et	PROPN
alces-297	236	22	al	al	PROPN
alces-297	236	23	.	.	PROPN
alces-297	236	24	2019	2019	NUM
alces-297	236	25	)	)	PUNCT
alces-297	236	26	,	,	PUNCT
alces-297	236	27	it	it	PRON
alces-297	236	28	is	be	AUX
alces-297	236	29	reported	report	VERB
alces-297	236	30	as	as	ADP
alces-297	236	31	a	a	DET
alces-297	236	32	significant	significant	ADJ
alces-297	236	33	mortality	mortality	NOUN
alces-297	236	34	factor	factor	NOUN
alces-297	236	35	in	in	ADP
alces-297	236	36	certain	certain	ADJ
alces-297	236	37	north	north	ADJ
alces-297	236	38	american	american	ADJ
alces-297	236	39	moose	moose	NOUN
alces-297	236	40	populations	population	NOUN
alces-297	236	41	(	(	PUNCT
alces-297	236	42	murray	murray	PROPN
alces-297	236	43	et	et	PROPN
alces-297	236	44	al	al	PROPN
alces-297	236	45	.	.	PROPN
alces-297	236	46	2006	2006	NUM
alces-297	236	47	)	)	PUNCT
alces-297	236	48	.	.	PUNCT
alces-297	237	1	sys	sys	PROPN
alces-297	237	2	appear	appear	VERB
alces-297	237	3	to	to	PART
alces-297	237	4	have	have	AUX
alces-297	237	5	been	be	AUX
alces-297	237	6	impacted	impact	VERB
alces-297	237	7	from	from	ADP
alces-297	237	8	the	the	DET
alces-297	237	9	beetle	beetle	NOUN
alces-297	237	10	outbreak	outbreak	NOUN
alces-297	237	11	(	(	PUNCT
alces-297	237	12	kuzyk	kuzyk	NOUN
alces-297	237	13	et	et	PROPN
alces-297	237	14	al	al	PROPN
alces-297	237	15	.	.	PROPN
alces-297	237	16	2018	2018	NUM
alces-297	237	17	)	)	PUNCT
alces-297	237	18	and	and	CCONJ
alces-297	237	19	optimal	optimal	ADJ
alces-297	237	20	harvest	harvest	NOUN
alces-297	237	21	rates	rate	NOUN
alces-297	237	22	may	may	AUX
alces-297	237	23	now	now	ADV
alces-297	237	24	be	be	AUX
alces-297	237	25	considerably	considerably	ADV
alces-297	237	26	lower	low	ADJ
alces-297	237	27	than	than	SCONJ
alces-297	237	28	projected	project	VERB
alces-297	237	29	by	by	ADP
alces-297	237	30	the	the	DET
alces-297	237	31	predator	predator	NOUN
alces-297	237	32	-	-	PUNCT
alces-297	237	33	prey	prey	NOUN
alces-297	237	34	model	model	NOUN
alces-297	237	35	.	.	PUNCT
alces-297	238	1	further	further	ADJ
alces-297	238	2	work	work	NOUN
alces-297	238	3	on	on	ADP
alces-297	238	4	anthropogenic	anthropogenic	ADJ
alces-297	238	5	and	and	CCONJ
alces-297	238	6	environmental	environmental	ADJ
alces-297	238	7	factors	factor	NOUN
alces-297	238	8	affecting	affect	VERB
alces-297	238	9	moose	moose	NOUN
alces-297	238	10	population	population	NOUN
alces-297	238	11	dynamics	dynamic	NOUN
alces-297	238	12	could	could	AUX
alces-297	238	13	improve	improve	VERB
alces-297	238	14	and	and	CCONJ
alces-297	238	15	make	make	VERB
alces-297	238	16	the	the	DET
alces-297	238	17	model	model	NOUN
alces-297	238	18	more	more	ADV
alces-297	238	19	suitable	suitable	ADJ
alces-297	238	20	for	for	ADP
alces-297	238	21	contemporary	contemporary	ADJ
alces-297	238	22	conditions	condition	NOUN
alces-297	238	23	.	.	PUNCT
alces-297	239	1	management	management	NOUN
alces-297	239	2	implications	implication	NOUN
alces-297	239	3	and	and	CCONJ
alces-297	239	4	recommendations	recommendation	NOUN
alces-297	239	5	sy	sy	ADP
alces-297	239	6	curves	curve	NOUN
alces-297	239	7	assume	assume	VERB
alces-297	239	8	that	that	SCONJ
alces-297	239	9	moose	moose	NOUN
alces-297	239	10	populations	population	NOUN
alces-297	239	11	are	be	AUX
alces-297	239	12	stable	stable	ADJ
alces-297	239	13	,	,	PUNCT
alces-297	239	14	while	while	SCONJ
alces-297	239	15	most	most	ADJ
alces-297	239	16	populations	population	NOUN
alces-297	239	17	fluctuate	fluctuate	NOUN
alces-297	239	18	and	and	CCONJ
alces-297	239	19	rarely	rarely	ADV
alces-297	239	20	,	,	PUNCT
alces-297	239	21	if	if	SCONJ
alces-297	239	22	ever	ever	ADV
alces-297	239	23	,	,	PUNCT
alces-297	239	24	achieve	achieve	VERB
alces-297	239	25	a	a	DET
alces-297	239	26	stable	stable	ADJ
alces-297	239	27	equilibrium	equilibrium	NOUN
alces-297	239	28	(	(	PUNCT
alces-297	239	29	sæther	sæther	NOUN
alces-297	239	30	et	et	PROPN
alces-297	239	31	al	al	PROPN
alces-297	239	32	.	.	PROPN
alces-297	239	33	2001	2001	NUM
alces-297	239	34	)	)	PUNCT
alces-297	239	35	.	.	PUNCT
alces-297	240	1	thus	thus	ADV
alces-297	240	2	,	,	PUNCT
alces-297	240	3	the	the	DET
alces-297	240	4	main	main	ADJ
alces-297	240	5	value	value	NOUN
alces-297	240	6	of	of	ADP
alces-297	240	7	estimating	estimate	VERB
alces-297	240	8	sys	sys	NOUN
alces-297	240	9	was	be	AUX
alces-297	240	10	to	to	PART
alces-297	240	11	provide	provide	VERB
alces-297	240	12	an	an	DET
alces-297	240	13	expectation	expectation	NOUN
alces-297	240	14	of	of	ADP
alces-297	240	15	harvest	harvest	NOUN
alces-297	240	16	under	under	ADP
alces-297	240	17	longterm	longterm	PROPN
alces-297	240	18	,	,	PUNCT
alces-297	240	19	stable	stable	ADJ
alces-297	240	20	predator	predator	NOUN
alces-297	240	21	-	-	PUNCT
alces-297	240	22	prey	prey	NOUN
alces-297	240	23	interactions	interaction	NOUN
alces-297	240	24	in	in	ADP
alces-297	240	25	central	central	PROPN
alces-297	240	26	bc	bc	PROPN
alces-297	240	27	.	.	PUNCT
alces-297	241	1	the	the	DET
alces-297	241	2	principal	principal	ADJ
alces-297	241	3	value	value	NOUN
alces-297	241	4	of	of	ADP
alces-297	241	5	the	the	DET
alces-297	241	6	optimal	optimal	ADJ
alces-297	241	7	harvest	harvest	NOUN
alces-297	241	8	calculations	calculation	NOUN
alces-297	241	9	was	be	AUX
alces-297	241	10	to	to	PART
alces-297	241	11	identify	identify	VERB
alces-297	241	12	how	how	SCONJ
alces-297	241	13	various	various	ADJ
alces-297	241	14	sex	sex	NOUN
alces-297	241	15	/	/	SYM
alces-297	241	16	age	age	NOUN
alces-297	241	17	classes	class	NOUN
alces-297	241	18	should	should	AUX
alces-297	241	19	be	be	AUX
alces-297	241	20	harvested	harvest	VERB
alces-297	241	21	under	under	ADP
alces-297	241	22	different	different	ADJ
alces-297	241	23	levels	level	NOUN
alces-297	241	24	of	of	ADP
alces-297	241	25	predation	predation	NOUN
alces-297	241	26	.	.	PUNCT
alces-297	242	1	estimating	estimate	VERB
alces-297	242	2	sys	sys	NOUN
alces-297	242	3	from	from	ADP
alces-297	242	4	the	the	DET
alces-297	242	5	predator	predator	NOUN
alces-297	242	6	-	-	PUNCT
alces-297	242	7	prey	prey	NOUN
alces-297	242	8	model	model	NOUN
alces-297	242	9	was	be	AUX
alces-297	242	10	limited	limit	VERB
alces-297	242	11	by	by	ADP
alces-297	242	12	the	the	DET
alces-297	242	13	lack	lack	NOUN
alces-297	242	14	of	of	ADP
alces-297	242	15	data	datum	NOUN
alces-297	242	16	on	on	ADP
alces-297	242	17	first	first	ADJ
alces-297	242	18	nations	nation	NOUN
alces-297	242	19	harvest	harvest	NOUN
alces-297	242	20	,	,	PUNCT
alces-297	242	21	calf	calf	NOUN
alces-297	242	22	vulnerability	vulnerability	NOUN
alces-297	242	23	to	to	ADP
alces-297	242	24	predation	predation	NOUN
alces-297	242	25	during	during	ADP
alces-297	242	26	summer	summer	NOUN
alces-297	242	27	,	,	PUNCT
alces-297	242	28	and	and	CCONJ
alces-297	242	29	uncertainty	uncertainty	NOUN
alces-297	242	30	in	in	ADP
alces-297	242	31	the	the	DET
alces-297	242	32	wolf	wolf	ADJ
alces-297	242	33	functional	functional	ADJ
alces-297	242	34	and	and	CCONJ
alces-297	242	35	numerical	numerical	ADJ
alces-297	242	36	responses	response	NOUN
alces-297	242	37	.	.	PUNCT
alces-297	243	1	nonetheless	nonetheless	ADV
alces-297	243	2	,	,	PUNCT
alces-297	243	3	the	the	DET
alces-297	243	4	model	model	NOUN
alces-297	243	5	findings	finding	VERB
alces-297	243	6	that	that	PRON
alces-297	243	7	moose	moose	VERB
alces-297	243	8	harvests	harvest	NOUN
alces-297	243	9	should	should	AUX
alces-297	243	10	consist	consist	VERB
alces-297	243	11	primarily	primarily	ADV
alces-297	243	12	of	of	ADP
alces-297	243	13	bulls	bull	NOUN
alces-297	243	14	and	and	CCONJ
alces-297	243	15	calves	calf	NOUN
alces-297	243	16	with	with	ADP
alces-297	243	17	nil	nil	NOUN
alces-297	243	18	to	to	ADP
alces-297	243	19	very	very	ADV
alces-297	243	20	low	low	ADJ
alces-297	243	21	cow	cow	NOUN
alces-297	243	22	harvests	harvest	NOUN
alces-297	243	23	was	be	AUX
alces-297	243	24	consistent	consistent	ADJ
alces-297	243	25	with	with	ADP
alces-297	243	26	other	other	ADJ
alces-297	243	27	published	publish	VERB
alces-297	243	28	studies	study	NOUN
alces-297	243	29	,	,	PUNCT
alces-297	243	30	and	and	CCONJ
alces-297	243	31	suggest	suggest	VERB
alces-297	243	32	these	these	DET
alces-297	243	33	findings	finding	NOUN
alces-297	243	34	are	be	AUX
alces-297	243	35	broadly	broadly	ADV
alces-297	243	36	applicable	applicable	ADJ
alces-297	243	37	to	to	ADP
alces-297	243	38	moose	moose	NOUN
alces-297	243	39	-	-	PUNCT
alces-297	243	40	wolf	wolf	NOUN
alces-297	243	41	-	-	PUNCT
alces-297	243	42	bear	bear	NOUN
alces-297	243	43	systems	system	NOUN
alces-297	243	44	where	where	SCONJ
alces-297	243	45	alternate	alternate	ADJ
alces-297	243	46	prey	prey	NOUN
alces-297	243	47	are	be	AUX
alces-297	243	48	rare	rare	ADJ
alces-297	243	49	.	.	PUNCT
alces-297	244	1	alces	alce	NOUN
alces-297	244	2	vol	vol	NOUN
alces-297	244	3	.	.	PROPN
alces-297	244	4	57	57	NUM
alces-297	244	5	,	,	PUNCT
alces-297	244	6	2021	2021	NUM
alces-297	244	7	modelling	model	VERB
alces-297	244	8	sustained	sustain	VERB
alces-297	244	9	yields	yield	NOUN
alces-297	244	10	for	for	ADP
alces-297	244	11	moose	moose	NOUN
alces-297	244	12	–	–	PUNCT
alces-297	244	13	hatter	hatter	NOUN
alces-297	244	14	.	.	PUNCT
alces-297	245	1	125	125	NUM
alces-297	245	2	managers	manager	NOUN
alces-297	245	3	who	who	PRON
alces-297	245	4	wish	wish	VERB
alces-297	245	5	to	to	PART
alces-297	245	6	elevate	elevate	VERB
alces-297	245	7	cow	cow	NOUN
alces-297	245	8	harvests	harvest	NOUN
alces-297	245	9	in	in	ADP
alces-297	245	10	order	order	NOUN
alces-297	245	11	to	to	PART
alces-297	245	12	monitor	monitor	VERB
alces-297	245	13	moose	moose	NOUN
alces-297	245	14	reproduction	reproduction	NOUN
alces-297	245	15	and	and	CCONJ
alces-297	245	16	nutritional	nutritional	ADJ
alces-297	245	17	status	status	NOUN
alces-297	245	18	,	,	PUNCT
alces-297	245	19	or	or	CCONJ
alces-297	245	20	to	to	PART
alces-297	245	21	recover	recover	VERB
alces-297	245	22	low	low	ADJ
alces-297	245	23	bull	bull	NOUN
alces-297	245	24	:	:	PUNCT
alces-297	245	25	cow	cow	NOUN
alces-297	245	26	ratios	ratio	NOUN
alces-297	245	27	,	,	PUNCT
alces-297	245	28	should	should	AUX
alces-297	245	29	reduce	reduce	VERB
alces-297	245	30	calf	calf	NOUN
alces-297	245	31	harvests	harvest	NOUN
alces-297	245	32	to	to	PART
alces-297	245	33	compensate	compensate	VERB
alces-297	245	34	for	for	ADP
alces-297	245	35	the	the	DET
alces-297	245	36	increased	increase	VERB
alces-297	245	37	harvest	harvest	NOUN
alces-297	245	38	of	of	ADP
alces-297	245	39	cows	cow	NOUN
alces-297	245	40	.	.	PUNCT
alces-297	246	1	further	further	ADJ
alces-297	246	2	restrictions	restriction	NOUN
alces-297	246	3	such	such	ADJ
alces-297	246	4	as	as	ADP
alces-297	246	5	bull	bull	NOUN
alces-297	246	6	-	-	PUNCT
alces-297	246	7	only	only	ADV
alces-297	246	8	harvests	harvest	NOUN
alces-297	246	9	should	should	AUX
alces-297	246	10	be	be	AUX
alces-297	246	11	considered	consider	VERB
alces-297	246	12	for	for	ADP
alces-297	246	13	low	low	ADJ
alces-297	246	14	density	density	NOUN
alces-297	246	15	,	,	PUNCT
alces-297	246	16	predator	predator	NOUN
alces-297	246	17	-	-	PUNCT
alces-297	246	18	limited	limit	VERB
alces-297	246	19	moose	moose	NOUN
alces-297	246	20	populations	population	NOUN
alces-297	246	21	.	.	PUNCT
alces-297	247	1	when	when	SCONJ
alces-297	247	2	moose	moose	NOUN
alces-297	247	3	populations	population	NOUN
alces-297	247	4	are	be	AUX
alces-297	247	5	food	food	NOUN
alces-297	247	6	-	-	PUNCT
alces-297	247	7	limited	limited	ADJ
alces-297	247	8	,	,	PUNCT
alces-297	247	9	harvests	harvest	NOUN
alces-297	247	10	of	of	ADP
alces-297	247	11	bulls	bull	NOUN
alces-297	247	12	,	,	PUNCT
alces-297	247	13	cows	cow	NOUN
alces-297	247	14	,	,	PUNCT
alces-297	247	15	and	and	CCONJ
alces-297	247	16	calves	calf	NOUN
alces-297	247	17	should	should	AUX
alces-297	247	18	be	be	AUX
alces-297	247	19	elevated	elevate	VERB
alces-297	247	20	to	to	PART
alces-297	247	21	maintain	maintain	VERB
alces-297	247	22	populations	population	NOUN
alces-297	247	23	below	below	ADP
alces-297	247	24	k	k	PROPN
alces-297	247	25	and	and	CCONJ
alces-297	247	26	enhance	enhance	VERB
alces-297	247	27	yields	yield	NOUN
alces-297	247	28	.	.	PUNCT
alces-297	248	1	finally	finally	ADV
alces-297	248	2	,	,	PUNCT
alces-297	248	3	and	and	CCONJ
alces-297	248	4	most	most	ADV
alces-297	248	5	importantly	importantly	ADV
alces-297	248	6	,	,	PUNCT
alces-297	248	7	moose	moose	NOUN
alces-297	248	8	harvests	harvest	NOUN
alces-297	248	9	should	should	AUX
alces-297	248	10	be	be	AUX
alces-297	248	11	set	set	VERB
alces-297	248	12	below	below	ADP
alces-297	248	13	optimal	optimal	ADJ
alces-297	248	14	yields	yield	NOUN
alces-297	248	15	to	to	PART
alces-297	248	16	account	account	VERB
alces-297	248	17	for	for	ADP
alces-297	248	18	stochastic	stochastic	ADJ
alces-297	248	19	variation	variation	NOUN
alces-297	248	20	in	in	ADP
alces-297	248	21	predator	predator	NOUN
alces-297	248	22	-	-	PUNCT
alces-297	248	23	prey	prey	NOUN
alces-297	248	24	interactions	interaction	NOUN
alces-297	248	25	,	,	PUNCT
alces-297	248	26	changing	change	VERB
alces-297	248	27	environmental	environmental	ADJ
alces-297	248	28	conditions	condition	NOUN
alces-297	248	29	,	,	PUNCT
alces-297	248	30	and	and	CCONJ
alces-297	248	31	management	management	NOUN
alces-297	248	32	uncertainty	uncertainty	NOUN
alces-297	248	33	.	.	PUNCT
alces-297	249	1	acknowledgements	acknowledgement	NOUN
alces-297	249	2	d.	d.	PROPN
alces-297	249	3	heard	hear	VERB
alces-297	249	4	,	,	PUNCT
alces-297	249	5	m.	m.	PROPN
alces-297	249	6	anderson	anderson	PROPN
alces-297	249	7	,	,	PUNCT
alces-297	249	8	and	and	CCONJ
alces-297	249	9	m.	m.	NOUN
alces-297	249	10	scheideman	scheideman	NOUN
alces-297	249	11	kindly	kindly	ADV
alces-297	249	12	reviewed	review	VERB
alces-297	249	13	an	an	DET
alces-297	249	14	early	early	ADJ
alces-297	249	15	draft	draft	NOUN
alces-297	249	16	of	of	ADP
alces-297	249	17	the	the	DET
alces-297	249	18	manuscript	manuscript	NOUN
alces-297	249	19	and	and	CCONJ
alces-297	249	20	made	make	VERB
alces-297	249	21	numerous	numerous	ADJ
alces-297	249	22	helpful	helpful	ADJ
alces-297	249	23	and	and	CCONJ
alces-297	249	24	insightful	insightful	ADJ
alces-297	249	25	suggestions	suggestion	NOUN
alces-297	249	26	.	.	PUNCT
alces-297	250	1	i	i	PRON
alces-297	250	2	am	be	AUX
alces-297	250	3	also	also	ADV
alces-297	250	4	grateful	grateful	ADJ
alces-297	250	5	for	for	ADP
alces-297	250	6	the	the	DET
alces-297	250	7	constructive	constructive	ADJ
alces-297	250	8	reviews	review	NOUN
alces-297	250	9	from	from	ADP
alces-297	250	10	e.	e.	PROPN
alces-297	250	11	addison	addison	PROPN
alces-297	250	12	and	and	CCONJ
alces-297	250	13	two	two	NUM
alces-297	250	14	anonymous	anonymous	ADJ
alces-297	250	15	reviewers	reviewer	NOUN
alces-297	250	16	which	which	PRON
alces-297	250	17	further	far	ADV
alces-297	250	18	improved	improve	VERB
alces-297	250	19	this	this	DET
alces-297	250	20	manuscript	manuscript	NOUN
alces-297	250	21	.	.	PUNCT
alces-297	251	1	references	reference	NOUN
alces-297	251	2	adams	adams	PROPN
alces-297	251	3	,	,	PUNCT
alces-297	251	4	l.	l.	PROPN
alces-297	251	5	g.	g.	PROPN
alces-297	251	6	,	,	PUNCT
alces-297	251	7	r.	r.	PROPN
alces-297	251	8	o.	o.	PROPN
alces-297	251	9	stephenson	stephenson	PROPN
alces-297	251	10	,	,	PUNCT
alces-297	251	11	b.	b.	PROPN
alces-297	251	12	w.	w.	PROPN
alces-297	251	13	dale	dale	PROPN
alces-297	251	14	,	,	PUNCT
alces-297	251	15	r.	r.	PROPN
alces-297	251	16	t.	t.	PROPN
alces-297	251	17	ahgook	ahgook	PROPN
alces-297	251	18	,	,	PUNCT
alces-297	251	19	and	and	CCONJ
alces-297	251	20	d.	d.	PROPN
alces-297	251	21	j.	j.	PROPN
alces-297	251	22	demma	demma	PROPN
alces-297	251	23	.	.	PUNCT
alces-297	252	1	2008	2008	NUM
alces-297	252	2	.	.	PUNCT
alces-297	253	1	population	population	NOUN
alces-297	253	2	dynamics	dynamic	NOUN
alces-297	253	3	and	and	CCONJ
alces-297	253	4	harvest	harvest	NOUN
alces-297	253	5	characteristics	characteristic	NOUN
alces-297	253	6	of	of	ADP
alces-297	253	7	wolves	wolf	NOUN
alces-297	253	8	in	in	ADP
alces-297	253	9	the	the	DET
alces-297	253	10	central	central	ADJ
alces-297	253	11	brooks	brooks	PROPN
alces-297	253	12	range	range	PROPN
alces-297	253	13	,	,	PUNCT
alces-297	253	14	alaska	alaska	PROPN
alces-297	253	15	.	.	PUNCT
alces-297	254	1	wildlife	wildlife	NOUN
alces-297	254	2	monographs	monograph	NOUN
alces-297	254	3	170(1	170(1	NUM
alces-297	254	4	):	):	PUNCT
alces-297	254	5	1–25	1–25	PROPN
alces-297	254	6	.	.	PUNCT
alces-297	255	1	doi	doi	NOUN
alces-297	255	2	:	:	PUNCT
alces-297	255	3	10.2193/2008	10.2193/2008	NUM
alces-297	255	4	-	-	PUNCT
alces-297	255	5	012	012	NUM
alces-297	255	6	ballard	ballard	PROPN
alces-297	255	7	,	,	PUNCT
alces-297	255	8	w.	w.	PROPN
alces-297	255	9	b.	b.	PROPN
alces-297	255	10	1992	1992	NUM
alces-297	255	11	.	.	PUNCT
alces-297	256	1	bear	bear	PROPN
alces-297	256	2	predation	predation	NOUN
alces-297	256	3	on	on	ADP
alces-297	256	4	moose	moose	NOUN
alces-297	256	5	:	:	PUNCT
alces-297	256	6	a	a	DET
alces-297	256	7	review	review	NOUN
alces-297	256	8	of	of	ADP
alces-297	256	9	recent	recent	ADJ
alces-297	256	10	north	north	ADJ
alces-297	256	11	american	american	ADJ
alces-297	256	12	studies	study	NOUN
alces-297	256	13	and	and	CCONJ
alces-297	256	14	their	their	PRON
alces-297	256	15	management	management	NOUN
alces-297	256	16	implications	implication	NOUN
alces-297	256	17	.	.	PUNCT
alces-297	257	1	alces	alce	NOUN
alces-297	257	2	supplement	supplement	VERB
alces-297	257	3	1	1	NUM
alces-297	257	4	:	:	PUNCT
alces-297	257	5	162–176	162–176	NUM
alces-297	257	6	.	.	PUNCT
alces-297	258	1	_	_	PUNCT
alces-297	259	1	_	_	PUNCT
alces-297	260	1	_	_	PUNCT
alces-297	261	1	_	_	PUNCT
alces-297	262	1	_	_	PUNCT
alces-297	262	2	,	,	PUNCT
alces-297	262	3	and	and	CCONJ
alces-297	262	4	v.	v.	ADP
alces-297	262	5	van	van	PROPN
alces-297	262	6	ballenberghe	ballenberghe	NOUN
alces-297	262	7	.	.	PUNCT
alces-297	263	1	1998	1998	NUM
alces-297	263	2	.	.	PUNCT
alces-297	264	1	predator	predator	NOUN
alces-297	264	2	-	-	PUNCT
alces-297	264	3	prey	prey	NOUN
alces-297	264	4	relationships	relationship	NOUN
alces-297	264	5	.	.	PUNCT
alces-297	265	1	pages	page	NOUN
alces-297	265	2	247	247	NUM
alces-297	265	3	–	–	PUNCT
alces-297	265	4	273	273	NUM
alces-297	265	5	in	in	ADP
alces-297	265	6	a.	a.	NOUN
alces-297	265	7	w.	w.	PROPN
alces-297	265	8	franzmann	franzmann	PROPN
alces-297	265	9	and	and	CCONJ
alces-297	265	10	c.	c.	PROPN
alces-297	265	11	c.	c.	PROPN
alces-297	265	12	schwartz	schwartz	PROPN
alces-297	265	13	,	,	PUNCT
alces-297	265	14	editors	editor	NOUN
alces-297	265	15	.	.	PUNCT
alces-297	266	1	ecology	ecology	NOUN
alces-297	266	2	and	and	CCONJ
alces-297	266	3	management	management	NOUN
alces-297	266	4	of	of	ADP
alces-297	266	5	the	the	DET
alces-297	266	6	north	north	ADJ
alces-297	266	7	american	american	ADJ
alces-297	266	8	moose	moose	NOUN
alces-297	266	9	.	.	PUNCT
alces-297	267	1	smithsonian	smithsonian	PROPN
alces-297	267	2	institution	institution	PROPN
alces-297	267	3	press	press	PROPN
alces-297	267	4	,	,	PUNCT
alces-297	267	5	washington	washington	PROPN
alces-297	267	6	,	,	PUNCT
alces-297	267	7	dc	dc	PROPN
alces-297	267	8	,	,	PUNCT
alces-297	267	9	usa	usa	PROPN
alces-297	267	10	.	.	PUNCT
alces-297	268	1	_	_	PUNCT
alces-297	269	1	_	_	PUNCT
alces-297	270	1	_	_	PUNCT
alces-297	271	1	_	_	PUNCT
alces-297	272	1	_	_	PUNCT
alces-297	272	2	,	,	PUNCT
alces-297	272	3	j.	j.	PROPN
alces-297	272	4	s.	s.	PROPN
alces-297	272	5	whitman	whitman	PROPN
alces-297	272	6	,	,	PUNCT
alces-297	272	7	and	and	CCONJ
alces-297	272	8	c.	c.	PROPN
alces-297	272	9	l.	l.	PROPN
alces-297	272	10	gardner	gardner	PROPN
alces-297	272	11	.	.	PUNCT
alces-297	272	12	1987	1987	NUM
alces-297	272	13	.	.	PUNCT
alces-297	273	1	ecology	ecology	NOUN
alces-297	273	2	of	of	ADP
alces-297	273	3	an	an	DET
alces-297	273	4	exploited	exploit	VERB
alces-297	273	5	wolf	wolf	NOUN
alces-297	273	6	population	population	NOUN
alces-297	273	7	in	in	ADP
alces-297	273	8	south	south	ADJ
alces-297	273	9	-	-	PUNCT
alces-297	273	10	central	central	ADJ
alces-297	273	11	alaska	alaska	PROPN
alces-297	273	12	.	.	PUNCT
alces-297	274	1	wildlife	wildlife	NOUN
alces-297	274	2	monographs	monograph	NOUN
alces-297	274	3	98	98	NUM
alces-297	274	4	:	:	PUNCT
alces-297	274	5	3–54	3–54	NUM
alces-297	274	6	.	.	PUNCT
alces-297	275	1	bergerud	bergerud	NOUN
alces-297	275	2	,	,	PUNCT
alces-297	275	3	a.	a.	PROPN
alces-297	275	4	t.	t.	PROPN
alces-297	275	5	,	,	PUNCT
alces-297	275	6	w.	w.	PROPN
alces-297	275	7	wyett	wyett	PROPN
alces-297	275	8	,	,	PUNCT
alces-297	275	9	and	and	CCONJ
alces-297	275	10	b.	b.	PROPN
alces-297	275	11	snider	snider	PROPN
alces-297	275	12	.	.	PUNCT
alces-297	276	1	1983	1983	NUM
alces-297	276	2	.	.	PUNCT
alces-297	277	1	the	the	DET
alces-297	277	2	role	role	NOUN
alces-297	277	3	of	of	ADP
alces-297	277	4	wolf	wolf	PROPN
alces-297	277	5	predation	predation	NOUN
alces-297	277	6	in	in	ADP
alces-297	277	7	limiting	limit	VERB
alces-297	277	8	a	a	DET
alces-297	277	9	moose	moose	NOUN
alces-297	277	10	population	population	NOUN
alces-297	277	11	.	.	PUNCT
alces-297	278	1	journal	journal	PROPN
alces-297	278	2	of	of	ADP
alces-297	278	3	wildlife	wildlife	NOUN
alces-297	278	4	management	management	NOUN
alces-297	278	5	47	47	NUM
alces-297	278	6	:	:	PUNCT
alces-297	279	1	977–988	977–988	NUM
alces-297	279	2	.	.	PUNCT
alces-297	279	3	doi	doi	NOUN
alces-297	279	4	:	:	PUNCT
alces-297	279	5	10.2307/3808156	10.2307/3808156	NUM
alces-297	279	6	boertje	boertje	ADJ
alces-297	279	7	,	,	PUNCT
alces-297	279	8	r.	r.	PROPN
alces-297	279	9	d.	d.	PROPN
alces-297	279	10	,	,	PUNCT
alces-297	279	11	m.	m.	NOUN
alces-297	279	12	a.	a.	PROPN
alces-297	279	13	keech	keech	PROPN
alces-297	279	14	,	,	PUNCT
alces-297	279	15	d.	d.	PROPN
alces-297	279	16	d.	d.	PROPN
alces-297	279	17	young	young	PROPN
alces-297	279	18	,	,	PUNCT
alces-297	279	19	k.	k.	PROPN
alces-297	279	20	a.	a.	PROPN
alces-297	279	21	kellie	kellie	PROPN
alces-297	279	22	,	,	PUNCT
alces-297	279	23	and	and	CCONJ
alces-297	279	24	c.	c.	PROPN
alces-297	279	25	t.	t.	PROPN
alces-297	279	26	seaton	seaton	PROPN
alces-297	279	27	.	.	PUNCT
alces-297	280	1	2009	2009	NUM
alces-297	280	2	.	.	PUNCT
alces-297	281	1	managing	manage	VERB
alces-297	281	2	for	for	ADP
alces-297	281	3	elevated	elevated	ADJ
alces-297	281	4	yield	yield	NOUN
alces-297	281	5	of	of	ADP
alces-297	281	6	moose	moose	NOUN
alces-297	281	7	in	in	ADP
alces-297	281	8	interior	interior	ADJ
alces-297	281	9	alaska	alaska	PROPN
alces-297	281	10	.	.	PUNCT
alces-297	282	1	journal	journal	PROPN
alces-297	282	2	of	of	ADP
alces-297	282	3	wildlife	wildlife	NOUN
alces-297	282	4	management	management	NOUN
alces-297	282	5	73	73	NUM
alces-297	282	6	:	:	PUNCT
alces-297	283	1	314–327	314–327	NUM
alces-297	283	2	.	.	PUNCT
alces-297	283	3	doi	doi	NOUN
alces-297	283	4	:	:	PUNCT
alces-297	283	5	10.2193/	10.2193/	NUM
alces-297	283	6	2007	2007	NUM
alces-297	283	7	-	-	SYM
alces-297	283	8	591	591	NUM
alces-297	283	9	_	_	PUNCT
alces-297	284	1	_	_	PUNCT
alces-297	285	1	_	_	PUNCT
alces-297	286	1	_	_	PUNCT
alces-297	287	1	_	_	PUNCT
alces-297	287	2	,	,	PUNCT
alces-297	287	3	k.	k.	PROPN
alces-297	287	4	a.	a.	PROPN
alces-297	287	5	kellie	kellie	PROPN
alces-297	287	6	,	,	PUNCT
alces-297	287	7	c.	c.	PROPN
alces-297	287	8	t.	t.	PROPN
alces-297	287	9	seaton	seaton	PROPN
alces-297	287	10	,	,	PUNCT
alces-297	287	11	m.	m.	NOUN
alces-297	287	12	a.	a.	PROPN
alces-297	287	13	keech	keech	PROPN
alces-297	287	14	,	,	PUNCT
alces-297	287	15	d.	d.	PROPN
alces-297	287	16	d.	d.	PROPN
alces-297	287	17	young	young	PROPN
alces-297	287	18	,	,	PUNCT
alces-297	287	19	and	and	CCONJ
alces-297	287	20	b.	b.	PROPN
alces-297	287	21	w.	w.	PROPN
alces-297	287	22	dale	dale	PROPN
alces-297	287	23	,	,	PUNCT
alces-297	287	24	2007	2007	NUM
alces-297	287	25	.	.	PUNCT
alces-297	288	1	ranking	rank	VERB
alces-297	288	2	alaska	alaska	PROPN
alces-297	288	3	moose	moose	PROPN
alces-297	288	4	nutrition	nutrition	PROPN
alces-297	288	5	:	:	PUNCT
alces-297	288	6	signals	signal	NOUN
alces-297	288	7	to	to	PART
alces-297	288	8	begin	begin	VERB
alces-297	288	9	liberal	liberal	ADJ
alces-297	288	10	antlerless	antlerless	NOUN
alces-297	288	11	harvests	harvest	NOUN
alces-297	288	12	.	.	PUNCT
alces-297	289	1	journal	journal	NOUN
alces-297	289	2	of	of	ADP
alces-297	289	3	wildlife	wildlife	NOUN
alces-297	289	4	management	management	NOUN
alces-297	289	5	71	71	NUM
alces-297	289	6	:	:	PUNCT
alces-297	289	7	1494–1506	1494–1506	NUM
alces-297	289	8	.	.	PUNCT
alces-297	290	1	doi	doi	NOUN
alces-297	290	2	:	:	PUNCT
alces-297	290	3	10.2193/2006	10.2193/2006	NUM
alces-297	290	4	-	-	SYM
alces-297	290	5	159	159	NUM
alces-297	290	6	_	_	PUNCT
alces-297	291	1	_	_	PUNCT
alces-297	292	1	_	_	PUNCT
alces-297	293	1	_	_	PUNCT
alces-297	294	1	_	_	PUNCT
alces-297	294	2	,	,	PUNCT
alces-297	294	3	p.	p.	NOUN
alces-297	294	4	valkenburg	valkenburg	PROPN
alces-297	294	5	,	,	PUNCT
alces-297	294	6	and	and	CCONJ
alces-297	294	7	m.	m.	PROPN
alces-297	294	8	e.	e.	PROPN
alces-297	294	9	mcnay	mcnay	PROPN
alces-297	294	10	.	.	PUNCT
alces-297	295	1	1996	1996	NUM
alces-297	295	2	.	.	PUNCT
alces-297	296	1	increases	increase	NOUN
alces-297	296	2	in	in	ADP
alces-297	296	3	moose	moose	NOUN
alces-297	296	4	,	,	PUNCT
alces-297	296	5	caribou	caribou	NOUN
alces-297	296	6	and	and	CCONJ
alces-297	296	7	wolves	wolf	NOUN
alces-297	296	8	following	follow	VERB
alces-297	296	9	wolf	wolf	PROPN
alces-297	296	10	control	control	NOUN
alces-297	296	11	in	in	ADP
alces-297	296	12	alaska	alaska	PROPN
alces-297	296	13	.	.	PUNCT
alces-297	297	1	journal	journal	PROPN
alces-297	297	2	of	of	ADP
alces-297	297	3	wildlife	wildlife	NOUN
alces-297	297	4	management	management	NOUN
alces-297	297	5	80	80	NUM
alces-297	297	6	:	:	PUNCT
alces-297	297	7	474–489	474–489	NUM
alces-297	297	8	.	.	PUNCT
alces-297	298	1	doi	doi	PROPN
alces-297	298	2	:	:	PUNCT
alces-297	298	3	10.2307/3802065	10.2307/3802065	NUM
alces-297	298	4	bowyer	bowyer	PROPN
alces-297	298	5	,	,	PUNCT
alces-297	298	6	r.	r.	PROPN
alces-297	298	7	t.	t.	PROPN
alces-297	298	8	,	,	PUNCT
alces-297	298	9	j.	j.	PROPN
alces-297	298	10	g.	g.	PROPN
alces-297	298	11	kie	kie	PROPN
alces-297	298	12	,	,	PUNCT
alces-297	298	13	d.	d.	PROPN
alces-297	298	14	k.	k.	PROPN
alces-297	298	15	person	person	PROPN
alces-297	298	16	,	,	PUNCT
alces-297	298	17	and	and	CCONJ
alces-297	298	18	k.	k.	PROPN
alces-297	298	19	l.	l.	PROPN
alces-297	298	20	monteith	monteith	PROPN
alces-297	298	21	.	.	PUNCT
alces-297	299	1	2013	2013	NUM
alces-297	299	2	.	.	PUNCT
alces-297	300	1	metrics	metric	NOUN
alces-297	300	2	of	of	ADP
alces-297	300	3	predation	predation	NOUN
alces-297	300	4	:	:	PUNCT
alces-297	300	5	perils	peril	NOUN
alces-297	300	6	of	of	ADP
alces-297	300	7	predator	predator	NOUN
alces-297	300	8	-	-	PUNCT
alces-297	300	9	prey	prey	NOUN
alces-297	300	10	ratios	ratio	NOUN
alces-297	300	11	.	.	PUNCT
alces-297	301	1	acta	acta	PROPN
alces-297	301	2	theriologica	theriologica	PROPN
alces-297	301	3	58	58	NUM
alces-297	301	4	:	:	PUNCT
alces-297	302	1	329–340	329–340	NUM
alces-297	302	2	.	.	PUNCT
alces-297	302	3	doi	doi	NOUN
alces-297	302	4	:	:	PUNCT
alces-297	302	5	10.1007/	10.1007/	NUM
alces-297	302	6	s13364	s13364	NOUN
alces-297	302	7	-	-	PUNCT
alces-297	302	8	013	013	NUM
alces-297	302	9	-	-	PUNCT
alces-297	302	10	0133	0133	NUM
alces-297	302	11	-	-	SYM
alces-297	302	12	1	1	NUM
alces-297	302	13	caughley	caughley	NOUN
alces-297	302	14	,	,	PUNCT
alces-297	302	15	g.	g.	PROPN
alces-297	302	16	1976	1976	NUM
alces-297	302	17	.	.	PUNCT
alces-297	303	1	wildlife	wildlife	NOUN
alces-297	303	2	management	management	NOUN
alces-297	303	3	and	and	CCONJ
alces-297	303	4	the	the	DET
alces-297	303	5	dynamics	dynamic	NOUN
alces-297	303	6	of	of	ADP
alces-297	303	7	ungulate	ungulate	ADJ
alces-297	303	8	populations	population	NOUN
alces-297	303	9	.	.	PUNCT
alces-297	304	1	pages	page	NOUN
alces-297	304	2	183–246	183–246	NUM
alces-297	304	3	in	in	ADP
alces-297	304	4	t.	t.	PROPN
alces-297	304	5	h.	h.	PROPN
alces-297	304	6	coaker	coaker	PROPN
alces-297	304	7	,	,	PUNCT
alces-297	304	8	editor	editor	NOUN
alces-297	304	9	.	.	PUNCT
alces-297	305	1	applied	apply	VERB
alces-297	305	2	biology	biology	NOUN
alces-297	305	3	,	,	PUNCT
alces-297	305	4	vol	vol	NOUN
alces-297	305	5	.	.	NOUN
alces-297	305	6	1	1	NUM
alces-297	305	7	.	.	PUNCT
alces-297	305	8	academic	academic	ADJ
alces-297	305	9	press	press	PROPN
alces-297	305	10	,	,	PUNCT
alces-297	305	11	london	london	PROPN
alces-297	305	12	,	,	PUNCT
alces-297	305	13	england	england	PROPN
alces-297	305	14	.	.	PUNCT
alces-297	306	1	child	child	NOUN
alces-297	306	2	,	,	PUNCT
alces-297	306	3	k.	k.	PROPN
alces-297	306	4	1983	1983	NUM
alces-297	306	5	.	.	PUNCT
alces-297	307	1	selective	selective	ADJ
alces-297	307	2	harvest	harvest	NOUN
alces-297	307	3	of	of	ADP
alces-297	307	4	moose	moose	NOUN
alces-297	307	5	in	in	ADP
alces-297	307	6	the	the	DET
alces-297	307	7	omineca	omineca	NOUN
alces-297	307	8	:	:	PUNCT
alces-297	307	9	some	some	DET
alces-297	307	10	preliminary	preliminary	ADJ
alces-297	307	11	results	result	NOUN
alces-297	307	12	.	.	PUNCT
alces-297	308	1	alces	alce	NOUN
alces-297	308	2	19	19	NUM
alces-297	308	3	:	:	PUNCT
alces-297	308	4	162–177	162–177	NUM
alces-297	308	5	.	.	PUNCT
alces-297	309	1	crête	crête	PROPN
alces-297	309	2	,	,	PUNCT
alces-297	309	3	m.	m.	NOUN
alces-297	309	4	1987	1987	NUM
alces-297	309	5	.	.	PUNCT
alces-297	310	1	the	the	DET
alces-297	310	2	impact	impact	NOUN
alces-297	310	3	of	of	ADP
alces-297	310	4	sport	sport	NOUN
alces-297	310	5	hunting	hunting	NOUN
alces-297	310	6	on	on	ADP
alces-297	310	7	north	north	PROPN
alces-297	310	8	american	american	ADJ
alces-297	310	9	moose	moose	NOUN
alces-297	310	10	.	.	PUNCT
alces-297	311	1	swedish	swedish	ADJ
alces-297	311	2	wildlife	wildlife	NOUN
alces-297	311	3	research	research	NOUN
alces-297	311	4	supplement	supplement	NOUN
alces-297	311	5	1	1	NUM
alces-297	311	6	:	:	PUNCT
alces-297	311	7	553–563	553–563	NUM
alces-297	311	8	.	.	PUNCT
alces-297	312	1	_	_	PUNCT
alces-297	313	1	_	_	PUNCT
alces-297	314	1	_	_	PUNCT
alces-297	315	1	_	_	PUNCT
alces-297	316	1	_	_	PUNCT
alces-297	316	2	,	,	PUNCT
alces-297	316	3	r.	r.	PROPN
alces-297	316	4	j.	j.	PROPN
alces-297	316	5	taylor	taylor	PROPN
alces-297	316	6	,	,	PUNCT
alces-297	316	7	and	and	CCONJ
alces-297	316	8	p.a	p.a	PROPN
alces-297	316	9	.	.	PUNCT
alces-297	316	10	jordon	jordon	PROPN
alces-297	316	11	.	.	PUNCT
alces-297	317	1	1981	1981	NUM
alces-297	317	2	.	.	PUNCT
alces-297	318	1	optimization	optimization	NOUN
alces-297	318	2	of	of	ADP
alces-297	318	3	moose	moose	NOUN
alces-297	318	4	harvest	harvest	NOUN
alces-297	318	5	in	in	ADP
alces-297	318	6	southwestern	southwestern	ADJ
alces-297	318	7	quebec	quebec	PROPN
alces-297	318	8	.	.	PUNCT
alces-297	319	1	journal	journal	PROPN
alces-297	319	2	of	of	ADP
alces-297	319	3	wildlife	wildlife	NOUN
alces-297	319	4	management	management	NOUN
alces-297	319	5	45	45	NUM
alces-297	319	6	:	:	PUNCT
alces-297	320	1	598–611	598–611	NUM
alces-297	320	2	.	.	PUNCT
alces-297	321	1	doi	doi	NOUN
alces-297	321	2	:	:	PUNCT
alces-297	321	3	10.2307/	10.2307/	NUM
alces-297	321	4	3808693	3808693	NUM
alces-297	321	5	modelling	model	VERB
alces-297	321	6	sustained	sustain	VERB
alces-297	321	7	yields	yield	NOUN
alces-297	321	8	for	for	ADP
alces-297	321	9	moose	moose	NOUN
alces-297	321	10	–	–	PUNCT
alces-297	321	11	hatter	hatter	NOUN
alces-297	321	12	.	.	PUNCT
alces-297	322	1	alces	alce	NOUN
alces-297	322	2	vol	vol	NOUN
alces-297	322	3	.	.	PROPN
alces-297	322	4	57	57	NUM
alces-297	322	5	,	,	PUNCT
alces-297	322	6	2021	2021	NUM
alces-297	322	7	126	126	NUM
alces-297	322	8	dahle	dahle	NOUN
alces-297	322	9	,	,	PUNCT
alces-297	322	10	b.	b.	PROPN
alces-297	322	11	,	,	PUNCT
alces-297	322	12	k.	k.	PROPN
alces-297	322	13	wallin	wallin	PROPN
alces-297	322	14	,	,	PUNCT
alces-297	322	15	g.	g.	PROPN
alces-297	322	16	cederlund	cederlund	PROPN
alces-297	322	17	,	,	PUNCT
alces-297	322	18	i.-l	i.-l	NOUN
alces-297	322	19	.	.	PUNCT
alces-297	323	1	persson	persson	PROPN
alces-297	323	2	,	,	PUNCT
alces-297	323	3	l.	l.	PROPN
alces-297	323	4	s.	s.	PROPN
alces-297	323	5	selvaag	selvaag	PROPN
alces-297	323	6	,	,	PUNCT
alces-297	323	7	and	and	CCONJ
alces-297	323	8	j.	j.	PROPN
alces-297	323	9	e.	e.	PROPN
alces-297	323	10	swenson	swenson	PROPN
alces-297	323	11	.	.	PUNCT
alces-297	324	1	2013	2013	NUM
alces-297	324	2	.	.	PUNCT
alces-297	325	1	predation	predation	NOUN
alces-297	325	2	on	on	ADP
alces-297	325	3	adult	adult	NOUN
alces-297	325	4	moose	moose	NOUN
alces-297	325	5	alces	alce	NOUN
alces-297	325	6	alces	alce	NOUN
alces-297	325	7	by	by	ADP
alces-297	325	8	european	european	ADJ
alces-297	325	9	brown	brown	PROPN
alces-297	325	10	bears	bears	PROPN
alces-297	325	11	ursus	ursus	PROPN
alces-297	325	12	arctos	arctos	PROPN
alces-297	325	13	.	.	PUNCT
alces-297	326	1	wildlife	wildlife	NOUN
alces-297	326	2	biology	biology	NOUN
alces-297	326	3	19	19	NUM
alces-297	326	4	:	:	PUNCT
alces-297	326	5	165–169	165–169	NUM
alces-297	326	6	.	.	PUNCT
alces-297	326	7	doi	doi	NOUN
alces-297	326	8	:	:	PUNCT
alces-297	326	9	10.2981/10	10.2981/10	NUM
alces-297	326	10	-	-	SYM
alces-297	326	11	113	113	NUM
alces-297	326	12	eberhardt	eberhardt	PROPN
alces-297	326	13	,	,	PUNCT
alces-297	326	14	l.	l.	PROPN
alces-297	326	15	l.	l.	PROPN
alces-297	326	16	1997	1997	NUM
alces-297	326	17	.	.	PUNCT
alces-297	326	18	is	be	AUX
alces-297	326	19	wolf	wolf	PROPN
alces-297	326	20	predation	predation	NOUN
alces-297	326	21	ratio	ratio	NOUN
alces-297	326	22	-	-	PUNCT
alces-297	326	23	dependent	dependent	ADJ
alces-297	326	24	?	?	PUNCT
alces-297	327	1	canadian	canadian	ADJ
alces-297	327	2	journal	journal	NOUN
alces-297	327	3	of	of	ADP
alces-297	327	4	zoology	zoology	NOUN
alces-297	327	5	75	75	NUM
alces-297	327	6	:	:	SYM
alces-297	327	7	1940–1944	1940–1944	NUM
alces-297	327	8	.	.	PUNCT
alces-297	328	1	doi	doi	NOUN
alces-297	328	2	:	:	PUNCT
alces-297	328	3	10.1139/	10.1139/	NUM
alces-297	328	4	z97	z97	NOUN
alces-297	328	5	-	-	PUNCT
alces-297	328	6	824	824	NUM
alces-297	328	7	_	_	PUNCT
alces-297	329	1	_	_	PUNCT
alces-297	330	1	_	_	PUNCT
alces-297	331	1	_	_	PUNCT
alces-297	332	1	_	_	PUNCT
alces-297	332	2	,	,	PUNCT
alces-297	332	3	r.	r.	PROPN
alces-297	332	4	a.	a.	PROPN
alces-297	332	5	garrott	garrott	PROPN
alces-297	332	6	,	,	PUNCT
alces-297	332	7	d.	d.	PROPN
alces-297	332	8	w.	w.	PROPN
alces-297	332	9	smith	smith	PROPN
alces-297	332	10	,	,	PUNCT
alces-297	332	11	p.	p.	PROPN
alces-297	332	12	i.	i.	PROPN
alces-297	332	13	white	white	PROPN
alces-297	332	14	,	,	PUNCT
alces-297	332	15	and	and	CCONJ
alces-297	332	16	r.	r.	PROPN
alces-297	332	17	o.	o.	PROPN
alces-297	332	18	peterson	peterson	PROPN
alces-297	332	19	.	.	PROPN
alces-297	332	20	2003	2003	NUM
alces-297	332	21	.	.	PUNCT
alces-297	333	1	assessing	assess	VERB
alces-297	333	2	the	the	DET
alces-297	333	3	impact	impact	NOUN
alces-297	333	4	of	of	ADP
alces-297	333	5	wolves	wolf	NOUN
alces-297	333	6	on	on	ADP
alces-297	333	7	ungulate	ungulate	ADJ
alces-297	333	8	prey	prey	NOUN
alces-297	333	9	.	.	PUNCT
alces-297	334	1	ecological	ecological	ADJ
alces-297	334	2	applications	application	NOUN
alces-297	334	3	13	13	NUM
alces-297	334	4	:	:	PUNCT
alces-297	334	5	776–783	776–783	NUM
alces-297	334	6	.	.	PUNCT
alces-297	335	1	doi	doi	NOUN
alces-297	335	2	:	:	PUNCT
alces-297	335	3	10.1890/1051	10.1890/1051	NUM
alces-297	335	4	-	-	SYM
alces-297	335	5	0761	0761	NUM
alces-297	335	6	(	(	PUNCT
alces-297	336	1	2003)013	2003)013	NUM
alces-297	336	2	[	[	X
alces-297	336	3	0776	0776	NUM
alces-297	336	4	:	:	PUNCT
alces-297	336	5	atiowo	atiowo	NOUN
alces-297	336	6	]	]	X
alces-297	337	1	2.0.co;2	2.0.co;2	NUM
alces-297	337	2	_	_	PUNCT
alces-297	338	1	_	_	PUNCT
alces-297	339	1	_	_	PUNCT
alces-297	340	1	_	_	PUNCT
alces-297	341	1	_	_	PUNCT
alces-297	341	2	,	,	PUNCT
alces-297	341	3	and	and	CCONJ
alces-297	341	4	r.	r.	PROPN
alces-297	341	5	o.	o.	PROPN
alces-297	341	6	peterson	peterson	PROPN
alces-297	341	7	.	.	PROPN
alces-297	341	8	1999	1999	NUM
alces-297	341	9	.	.	PUNCT
alces-297	342	1	predicting	predict	VERB
alces-297	342	2	the	the	DET
alces-297	342	3	wolf	wolf	NOUN
alces-297	342	4	-	-	PUNCT
alces-297	342	5	prey	prey	NOUN
alces-297	342	6	equilibrium	equilibrium	NOUN
alces-297	342	7	point	point	NOUN
alces-297	342	8	.	.	PUNCT
alces-297	343	1	canadian	canadian	ADJ
alces-297	343	2	journal	journal	PROPN
alces-297	343	3	of	of	ADP
alces-297	343	4	zoology	zoology	NOUN
alces-297	343	5	77	77	NUM
alces-297	343	6	:	:	PUNCT
alces-297	343	7	494	494	NUM
alces-297	343	8	–	–	PUNCT
alces-297	343	9	498	498	NUM
alces-297	343	10	.	.	PUNCT
alces-297	344	1	doi	doi	NOUN
alces-297	344	2	:	:	PUNCT
alces-297	344	3	10.1139	10.1139	NUM
alces-297	344	4	/	/	SYM
alces-297	344	5	z98	z98	NOUN
alces-297	344	6	-	-	PUNCT
alces-297	344	7	240	240	NUM
alces-297	344	8	environment	environment	NOUN
alces-297	344	9	yukon	yukon	PROPN
alces-297	344	10	.	.	PUNCT
alces-297	345	1	2016	2016	NUM
alces-297	345	2	.	.	PUNCT
alces-297	346	1	science	science	NOUN
alces-297	346	2	-	-	PUNCT
alces-297	346	3	based	base	VERB
alces-297	346	4	guidelines	guideline	NOUN
alces-297	346	5	for	for	ADP
alces-297	346	6	management	management	NOUN
alces-297	346	7	of	of	ADP
alces-297	346	8	moose	moose	NOUN
alces-297	346	9	in	in	ADP
alces-297	346	10	yukon	yukon	PROPN
alces-297	346	11	.	.	PUNCT
alces-297	347	1	yukon	yukon	PROPN
alces-297	347	2	fish	fish	PROPN
alces-297	347	3	and	and	CCONJ
alces-297	347	4	wildlife	wildlife	NOUN
alces-297	347	5	branch	branch	NOUN
alces-297	347	6	report	report	NOUN
alces-297	347	7	mr-16–02	mr-16–02	PROPN
alces-297	347	8	.	.	PUNCT
alces-297	347	9	whitehorse	whitehorse	PROPN
alces-297	347	10	,	,	PUNCT
alces-297	347	11	yukon	yukon	PROPN
alces-297	347	12	,	,	PUNCT
alces-297	347	13	canada	canada	PROPN
alces-297	347	14	flnro	flnro	PROPN
alces-297	347	15	.	.	PUNCT
alces-297	348	1	provincial	provincial	ADJ
alces-297	348	2	moose	moose	NOUN
alces-297	348	3	management	management	NOUN
alces-297	348	4	team	team	NOUN
alces-297	348	5	.	.	PUNCT
alces-297	349	1	2013	2013	NUM
alces-297	349	2	.	.	PUNCT
alces-297	350	1	draft	draft	VERB
alces-297	350	2	provincial	provincial	ADJ
alces-297	350	3	framework	framework	NOUN
alces-297	350	4	for	for	ADP
alces-297	350	5	moose	moose	NOUN
alces-297	350	6	management	management	NOUN
alces-297	350	7	in	in	ADP
alces-297	350	8	british	british	PROPN
alces-297	350	9	columbia	columbia	PROPN
alces-297	350	10	.	.	PUNCT
alces-297	351	1	unpublished	unpublished	ADJ
alces-297	351	2	report	report	NOUN
alces-297	351	3	.	.	PUNCT
alces-297	352	1	ministry	ministry	NOUN
alces-297	352	2	of	of	ADP
alces-297	352	3	forests	forest	NOUN
alces-297	352	4	,	,	PUNCT
alces-297	352	5	lands	land	NOUN
alces-297	352	6	and	and	CCONJ
alces-297	352	7	natural	natural	ADJ
alces-297	352	8	resource	resource	NOUN
alces-297	352	9	operations	operation	NOUN
alces-297	352	10	,	,	PUNCT
alces-297	352	11	victoria	victoria	PROPN
alces-297	352	12	,	,	PUNCT
alces-297	352	13	british	british	PROPN
alces-297	352	14	columbia	columbia	PROPN
alces-297	352	15	,	,	PUNCT
alces-297	352	16	canada	canada	PROPN
alces-297	352	17	.	.	PUNCT
alces-297	353	1	gasaway	gasaway	PROPN
alces-297	353	2	,	,	PUNCT
alces-297	353	3	w.	w.	PROPN
alces-297	353	4	c.	c.	PROPN
alces-297	353	5	,	,	PUNCT
alces-297	353	6	r.	r.	PROPN
alces-297	353	7	d.	d.	PROPN
alces-297	353	8	boertje	boertje	PROPN
alces-297	353	9	,	,	PUNCT
alces-297	353	10	d.	d.	PROPN
alces-297	353	11	v.	v.	PROPN
alces-297	353	12	grangaard	grangaard	PROPN
alces-297	353	13	,	,	PUNCT
alces-297	353	14	d.	d.	PROPN
alces-297	353	15	g.	g.	PROPN
alces-297	353	16	kelleyhouse	kelleyhouse	PROPN
alces-297	353	17	,	,	PUNCT
alces-297	353	18	r.	r.	PROPN
alces-297	353	19	o.	o.	PROPN
alces-297	353	20	stephenson	stephenson	PROPN
alces-297	353	21	,	,	PUNCT
alces-297	353	22	and	and	CCONJ
alces-297	353	23	d.	d.	PROPN
alces-297	353	24	g.	g.	PROPN
alces-297	353	25	larson	larson	PROPN
alces-297	353	26	.	.	PROPN
alces-297	354	1	1992	1992	NUM
alces-297	354	2	.	.	PUNCT
alces-297	355	1	the	the	DET
alces-297	355	2	role	role	NOUN
alces-297	355	3	of	of	ADP
alces-297	355	4	predation	predation	NOUN
alces-297	355	5	in	in	ADP
alces-297	355	6	limiting	limit	VERB
alces-297	355	7	moose	moose	NOUN
alces-297	355	8	at	at	ADP
alces-297	355	9	low	low	ADJ
alces-297	355	10	densities	density	NOUN
alces-297	355	11	in	in	ADP
alces-297	355	12	alaska	alaska	PROPN
alces-297	355	13	and	and	CCONJ
alces-297	355	14	yukon	yukon	PROPN
alces-297	355	15	and	and	CCONJ
alces-297	355	16	implications	implication	NOUN
alces-297	355	17	for	for	ADP
alces-297	355	18	conservation	conservation	NOUN
alces-297	355	19	.	.	PUNCT
alces-297	356	1	wildlife	wildlife	NOUN
alces-297	356	2	monographs	monograph	NOUN
alces-297	356	3	120	120	NUM
alces-297	356	4	:	:	PUNCT
alces-297	356	5	1–59	1–59	NOUN
alces-297	356	6	.	.	PUNCT
alces-297	357	1	_	_	PUNCT
alces-297	358	1	_	_	PUNCT
alces-297	359	1	_	_	PUNCT
alces-297	360	1	_	_	PUNCT
alces-297	361	1	_	_	PUNCT
alces-297	361	2	,	,	PUNCT
alces-297	361	3	r.	r.	PROPN
alces-297	361	4	o.	o.	PROPN
alces-297	361	5	stephenson	stephenson	PROPN
alces-297	361	6	,	,	PUNCT
alces-297	361	7	j.	j.	PROPN
alces-297	361	8	l.	l.	PROPN
alces-297	361	9	davis	davis	PROPN
alces-297	361	10	,	,	PUNCT
alces-297	361	11	p.	p.	PROPN
alces-297	361	12	e.	e.	PROPN
alces-297	361	13	k.	k.	PROPN
alces-297	361	14	shepherd	shepherd	PROPN
alces-297	361	15	,	,	PUNCT
alces-297	361	16	and	and	CCONJ
alces-297	361	17	o.	o.	PROPN
alces-297	361	18	e.	e.	PROPN
alces-297	361	19	burris	burris	PROPN
alces-297	361	20	.	.	PUNCT
alces-297	362	1	1983	1983	NUM
alces-297	362	2	.	.	PUNCT
alces-297	363	1	interrelationships	interrelationship	NOUN
alces-297	363	2	of	of	ADP
alces-297	363	3	wolves	wolf	NOUN
alces-297	363	4	,	,	PUNCT
alces-297	363	5	prey	prey	NOUN
alces-297	363	6	and	and	CCONJ
alces-297	363	7	man	man	NOUN
alces-297	363	8	in	in	ADP
alces-297	363	9	interior	interior	ADJ
alces-297	363	10	alaska	alaska	PROPN
alces-297	363	11	.	.	PUNCT
alces-297	364	1	wildlife	wildlife	NOUN
alces-297	364	2	monographs	monograph	NOUN
alces-297	364	3	84	84	NUM
alces-297	364	4	:	:	PUNCT
alces-297	364	5	1–50	1–50	NOUN
alces-297	364	6	.	.	PUNCT
alces-297	365	1	hatter	hatter	NOUN
alces-297	365	2	,	,	PUNCT
alces-297	365	3	i.	i.	PROPN
alces-297	365	4	w.	w.	PROPN
alces-297	365	5	1999	1999	NUM
alces-297	365	6	.	.	PUNCT
alces-297	366	1	an	an	DET
alces-297	366	2	evaluation	evaluation	NOUN
alces-297	366	3	of	of	ADP
alces-297	366	4	moose	moose	NOUN
alces-297	366	5	harvest	harvest	NOUN
alces-297	366	6	management	management	NOUN
alces-297	366	7	in	in	ADP
alces-297	366	8	central	central	ADJ
alces-297	366	9	and	and	CCONJ
alces-297	366	10	northern	northern	ADJ
alces-297	366	11	british	british	PROPN
alces-297	366	12	columbia	columbia	PROPN
alces-297	366	13	.	.	PUNCT
alces-297	367	1	alces	alce	NOUN
alces-297	367	2	35	35	NUM
alces-297	367	3	:	:	PUNCT
alces-297	367	4	91–103	91–103	NUM
alces-297	367	5	.	.	PUNCT
alces-297	368	1	_	_	PUNCT
alces-297	369	1	_	_	PUNCT
alces-297	370	1	_	_	PUNCT
alces-297	371	1	_	_	PUNCT
alces-297	372	1	_	_	PUNCT
alces-297	372	2	.	.	PUNCT
alces-297	373	1	2019	2019	NUM
alces-297	373	2	.	.	PUNCT
alces-297	374	1	an	an	DET
alces-297	374	2	assessment	assessment	NOUN
alces-297	374	3	of	of	ADP
alces-297	374	4	eberhardt	eberhardt	PROPN
alces-297	374	5	’s	’s	PART
alces-297	374	6	ratio	ratio	NOUN
alces-297	374	7	-	-	PUNCT
alces-297	374	8	dependent	dependent	ADJ
alces-297	374	9	wolf	wolf	NOUN
alces-297	374	10	-	-	PUNCT
alces-297	374	11	ungulate	ungulate	ADJ
alces-297	374	12	model	model	NOUN
alces-297	374	13	.	.	PUNCT
alces-297	375	1	journal	journal	PROPN
alces-297	375	2	of	of	ADP
alces-297	375	3	wildlife	wildlife	NOUN
alces-297	375	4	and	and	CCONJ
alces-297	375	5	biodiversity	biodiversity	NOUN
alces-297	375	6	3	3	NUM
alces-297	375	7	:	:	PUNCT
alces-297	375	8	1–8	1–8	X
alces-297	375	9	.	.	PUNCT
alces-297	376	1	hayes	hayes	PROPN
alces-297	376	2	,	,	PUNCT
alces-297	376	3	r.	r.	PROPN
alces-297	376	4	d.	d.	PROPN
alces-297	376	5	,	,	PUNCT
alces-297	376	6	a.	a.	PROPN
alces-297	376	7	m.	m.	PROPN
alces-297	376	8	baer	baer	PROPN
alces-297	376	9	,	,	PUNCT
alces-297	376	10	u.	u.	PROPN
alces-297	376	11	wotschikowsky	wotschikowsky	PROPN
alces-297	376	12	,	,	PUNCT
alces-297	376	13	and	and	CCONJ
alces-297	376	14	a.	a.	PROPN
alces-297	376	15	s.	s.	PROPN
alces-297	376	16	harestad	harestad	PROPN
alces-297	376	17	.	.	PUNCT
alces-297	377	1	2000	2000	NUM
alces-297	377	2	.	.	PUNCT
alces-297	378	1	kill	kill	VERB
alces-297	378	2	rate	rate	NOUN
alces-297	378	3	by	by	ADP
alces-297	378	4	wolves	wolf	NOUN
alces-297	378	5	on	on	ADP
alces-297	378	6	moose	moose	NOUN
alces-297	378	7	in	in	ADP
alces-297	378	8	the	the	DET
alces-297	378	9	yukon	yukon	PROPN
alces-297	378	10	.	.	PUNCT
alces-297	379	1	canadian	canadian	PROPN
alces-297	379	2	journal	journal	PROPN
alces-297	379	3	of	of	ADP
alces-297	379	4	zoology	zoology	NOUN
alces-297	379	5	78	78	NUM
alces-297	379	6	:	:	PUNCT
alces-297	379	7	49–59	49–59	NOUN
alces-297	379	8	.	.	PUNCT
alces-297	380	1	doi	doi	NOUN
alces-297	380	2	:	:	PUNCT
alces-297	380	3	10.1139	10.1139	NUM
alces-297	380	4	/	/	SYM
alces-297	380	5	z99	z99	PROPN
alces-297	380	6	-	-	PUNCT
alces-297	380	7	187	187	NUM
alces-297	380	8	_	_	PUNCT
alces-297	381	1	_	_	PUNCT
alces-297	382	1	_	_	PUNCT
alces-297	383	1	_	_	PUNCT
alces-297	384	1	_	_	PUNCT
alces-297	384	2	,	,	PUNCT
alces-297	384	3	and	and	CCONJ
alces-297	384	4	a.	a.	PROPN
alces-297	384	5	s.	s.	PROPN
alces-297	384	6	harestad	harestad	PROPN
alces-297	384	7	.	.	PUNCT
alces-297	385	1	2000	2000	NUM
alces-297	385	2	.	.	PUNCT
alces-297	386	1	wolf	wolf	PROPN
alces-297	386	2	functional	functional	ADJ
alces-297	386	3	response	response	NOUN
alces-297	386	4	and	and	CCONJ
alces-297	386	5	regulation	regulation	NOUN
alces-297	386	6	of	of	ADP
alces-297	386	7	moose	moose	NOUN
alces-297	386	8	in	in	ADP
alces-297	386	9	the	the	DET
alces-297	386	10	yukon	yukon	PROPN
alces-297	386	11	.	.	PUNCT
alces-297	387	1	canadian	canadian	PROPN
alces-297	387	2	journal	journal	PROPN
alces-297	387	3	of	of	ADP
alces-297	387	4	zoology	zoology	NOUN
alces-297	387	5	78	78	NUM
alces-297	387	6	:	:	PUNCT
alces-297	387	7	60–66	60–66	NUM
alces-297	387	8	.	.	PUNCT
alces-297	388	1	doi	doi	NOUN
alces-297	388	2	:	:	PUNCT
alces-297	388	3	10.1139/	10.1139/	NUM
alces-297	388	4	z99	z99	NUM
alces-297	388	5	-	-	PUNCT
alces-297	388	6	188	188	NUM
alces-297	388	7	heard	hear	VERB
alces-297	388	8	,	,	PUNCT
alces-297	388	9	d.	d.	PROPN
alces-297	388	10	c.	c.	PROPN
alces-297	388	11	,	,	PUNCT
alces-297	388	12	s.	s.	PROPN
alces-297	388	13	barry	barry	PROPN
alces-297	388	14	,	,	PUNCT
alces-297	388	15	g.	g.	PROPN
alces-297	388	16	watts	watts	PROPN
alces-297	388	17	,	,	PUNCT
alces-297	388	18	and	and	CCONJ
alces-297	388	19	k.	k.	PROPN
alces-297	388	20	child	child	NOUN
alces-297	388	21	.	.	PUNCT
alces-297	389	1	1997	1997	NUM
alces-297	389	2	.	.	PUNCT
alces-297	390	1	fertility	fertility	NOUN
alces-297	390	2	of	of	ADP
alces-297	390	3	female	female	ADJ
alces-297	390	4	moose	moose	NOUN
alces-297	390	5	(	(	PUNCT
alces-297	390	6	alces	alce	NOUN
alces-297	390	7	alces	alce	NOUN
alces-297	390	8	)	)	PUNCT
alces-297	390	9	in	in	ADP
alces-297	390	10	relation	relation	NOUN
alces-297	390	11	to	to	ADP
alces-297	390	12	age	age	NOUN
alces-297	390	13	and	and	CCONJ
alces-297	390	14	body	body	NOUN
alces-297	390	15	composition	composition	NOUN
alces-297	390	16	.	.	PUNCT
alces-297	391	1	alces	alce	NOUN
alces-297	391	2	33	33	NUM
alces-297	391	3	:	:	PUNCT
alces-297	391	4	165–176	165–176	NUM
alces-297	391	5	.	.	PUNCT
alces-297	392	1	_	_	PUNCT
alces-297	393	1	_	_	PUNCT
alces-297	394	1	_	_	PUNCT
alces-297	395	1	_	_	PUNCT
alces-297	396	1	_	_	PUNCT
alces-297	396	2	,	,	PUNCT
alces-297	396	3	k.	k.	PROPN
alces-297	396	4	l.	l.	PROPN
alces-297	396	5	zimmerman	zimmerman	PROPN
alces-297	396	6	,	,	PUNCT
alces-297	396	7	g.	g.	PROPN
alces-297	396	8	s.	s.	PROPN
alces-297	396	9	watts	watts	PROPN
alces-297	396	10	,	,	PUNCT
alces-297	396	11	and	and	CCONJ
alces-297	396	12	s.	s.	PROPN
alces-297	396	13	p.	p.	PROPN
alces-297	396	14	barry	barry	PROPN
alces-297	396	15	.	.	PUNCT
alces-297	397	1	1999	1999	NUM
alces-297	397	2	.	.	PUNCT
alces-297	398	1	moose	moose	NOUN
alces-297	398	2	density	density	NOUN
alces-297	398	3	and	and	CCONJ
alces-297	398	4	composition	composition	NOUN
alces-297	398	5	around	around	ADP
alces-297	398	6	prince	prince	PROPN
alces-297	398	7	george	george	PROPN
alces-297	398	8	,	,	PUNCT
alces-297	398	9	british	british	PROPN
alces-297	398	10	columbia	columbia	PROPN
alces-297	398	11	,	,	PUNCT
alces-297	398	12	december	december	PROPN
alces-297	398	13	1998	1998	NUM
alces-297	398	14	.	.	PUNCT
alces-297	399	1	final	final	ADJ
alces-297	399	2	report	report	NOUN
alces-297	399	3	for	for	ADP
alces-297	399	4	common	common	ADJ
alces-297	399	5	land	land	NOUN
alces-297	399	6	information	information	NOUN
alces-297	399	7	base	base	NOUN
alces-297	399	8	.	.	PUNCT
alces-297	400	1	project	project	NOUN
alces-297	400	2	no	no	NOUN
alces-297	400	3	.	.	NOUN
alces-297	400	4	99004	99004	NUM
alces-297	400	5	.	.	PUNCT
alces-297	401	1	ministry	ministry	NOUN
alces-297	401	2	of	of	ADP
alces-297	401	3	forests	forest	NOUN
alces-297	401	4	,	,	PUNCT
alces-297	401	5	lands	land	NOUN
alces-297	401	6	and	and	CCONJ
alces-297	401	7	natural	natural	ADJ
alces-297	401	8	resource	resource	NOUN
alces-297	401	9	operations	operation	NOUN
alces-297	401	10	,	,	PUNCT
alces-297	401	11	victoria	victoria	PROPN
alces-297	401	12	,	,	PUNCT
alces-297	401	13	british	british	PROPN
alces-297	401	14	columbia	columbia	PROPN
alces-297	401	15	,	,	PUNCT
alces-297	401	16	canada	canada	PROPN
alces-297	401	17	.	.	PUNCT
alces-297	401	18	hebblewhite	hebblewhite	PROPN
alces-297	401	19	,	,	PUNCT
alces-297	401	20	m.	m.	NOUN
alces-297	401	21	2013	2013	NUM
alces-297	401	22	.	.	PUNCT
alces-297	402	1	consequences	consequence	NOUN
alces-297	402	2	of	of	ADP
alces-297	402	3	ratio	ratio	NOUN
alces-297	402	4	dependent	dependent	ADJ
alces-297	402	5	predation	predation	NOUN
alces-297	402	6	by	by	ADP
alces-297	402	7	wolves	wolf	NOUN
alces-297	402	8	for	for	ADP
alces-297	402	9	elk	elk	NOUN
alces-297	402	10	population	population	NOUN
alces-297	402	11	dynamics	dynamic	NOUN
alces-297	402	12	.	.	PUNCT
alces-297	403	1	population	population	NOUN
alces-297	403	2	ecology	ecology	NOUN
alces-297	403	3	55	55	NUM
alces-297	403	4	:	:	PUNCT
alces-297	404	1	511–522	511–522	NUM
alces-297	404	2	.	.	PUNCT
alces-297	404	3	doi	doi	NOUN
alces-297	404	4	:	:	PUNCT
alces-297	404	5	10.1007/	10.1007/	NUM
alces-297	404	6	s10144	s10144	NUM
alces-297	404	7	-	-	PUNCT
alces-297	404	8	013	013	NUM
alces-297	404	9	-	-	PUNCT
alces-297	404	10	0384	0384	NUM
alces-297	404	11	-	-	SYM
alces-297	404	12	3	3	NUM
alces-297	404	13	hilborn	hilborn	ADJ
alces-297	404	14	,	,	PUNCT
alces-297	404	15	r.	r.	NOUN
alces-297	404	16	,	,	PUNCT
alces-297	404	17	and	and	CCONJ
alces-297	404	18	m.	m.	NOUN
alces-297	404	19	mangel	mangel	PROPN
alces-297	404	20	.	.	PUNCT
alces-297	405	1	1997	1997	NUM
alces-297	405	2	.	.	PUNCT
alces-297	406	1	the	the	DET
alces-297	406	2	ecological	ecological	ADJ
alces-297	406	3	detective	detective	NOUN
alces-297	406	4	:	:	PUNCT
alces-297	406	5	confronting	confront	VERB
alces-297	406	6	models	model	NOUN
alces-297	406	7	with	with	ADP
alces-297	406	8	data	datum	NOUN
alces-297	406	9	.	.	PUNCT
alces-297	407	1	princeton	princeton	PROPN
alces-297	407	2	university	university	PROPN
alces-297	407	3	press	press	PROPN
alces-297	407	4	,	,	PUNCT
alces-297	407	5	princeton	princeton	PROPN
alces-297	407	6	,	,	PUNCT
alces-297	407	7	new	new	PROPN
alces-297	407	8	jersey	jersey	PROPN
alces-297	407	9	,	,	PUNCT
alces-297	407	10	usa	usa	PROPN
alces-297	407	11	.	.	PROPN
alces-297	407	12	keith	keith	PROPN
alces-297	407	13	,	,	PUNCT
alces-297	407	14	l.	l.	PROPN
alces-297	407	15	b.	b.	PROPN
alces-297	407	16	1983	1983	NUM
alces-297	407	17	.	.	PUNCT
alces-297	408	1	population	population	NOUN
alces-297	408	2	dynamics	dynamic	NOUN
alces-297	408	3	of	of	ADP
alces-297	408	4	wolves	wolf	NOUN
alces-297	408	5	.	.	PUNCT
alces-297	409	1	pages	page	NOUN
alces-297	409	2	66–77	66–77	NUM
alces-297	409	3	in	in	ADP
alces-297	409	4	l.	l.	PROPN
alces-297	409	5	n.	n.	PROPN
alces-297	409	6	carbyn	carbyn	PROPN
alces-297	409	7	,	,	PUNCT
alces-297	409	8	editor	editor	NOUN
alces-297	409	9	.	.	PUNCT
alces-297	410	1	wolves	wolf	NOUN
alces-297	410	2	in	in	ADP
alces-297	410	3	canada	canada	PROPN
alces-297	410	4	and	and	CCONJ
alces-297	410	5	alaska	alaska	PROPN
alces-297	410	6	:	:	PUNCT
alces-297	410	7	their	their	PRON
alces-297	410	8	status	status	NOUN
alces-297	410	9	,	,	PUNCT
alces-297	410	10	biology	biology	NOUN
alces-297	410	11	,	,	PUNCT
alces-297	410	12	and	and	CCONJ
alces-297	410	13	management	management	NOUN
alces-297	410	14	.	.	PUNCT
alces-297	411	1	canadian	canadian	ADJ
alces-297	411	2	wildlife	wildlife	NOUN
alces-297	411	3	service	service	NOUN
alces-297	411	4	report	report	NOUN
alces-297	411	5	series	series	PROPN
alces-297	411	6	45	45	NUM
alces-297	411	7	.	.	PUNCT
alces-297	412	1	ottawa	ottawa	PROPN
alces-297	412	2	,	,	PUNCT
alces-297	412	3	ontario	ontario	PROPN
alces-297	412	4	,	,	PUNCT
alces-297	412	5	canada	canada	PROPN
alces-297	412	6	.	.	PUNCT
alces-297	413	1	kuzyk	kuzyk	PROPN
alces-297	413	2	,	,	PUNCT
alces-297	413	3	g.	g.	PROPN
alces-297	413	4	,	,	PUNCT
alces-297	413	5	and	and	CCONJ
alces-297	413	6	i.	i.	PROPN
alces-297	413	7	hatter	hatter	PROPN
alces-297	413	8	.	.	PUNCT
alces-297	414	1	2014	2014	NUM
alces-297	414	2	.	.	PUNCT
alces-297	415	1	using	use	VERB
alces-297	415	2	ungulate	ungulate	ADJ
alces-297	415	3	biomass	biomass	NOUN
alces-297	415	4	to	to	PART
alces-297	415	5	estimate	estimate	VERB
alces-297	415	6	abundance	abundance	NOUN
alces-297	415	7	of	of	ADP
alces-297	415	8	wolves	wolf	NOUN
alces-297	415	9	in	in	ADP
alces-297	415	10	british	british	PROPN
alces-297	415	11	columbia	columbia	PROPN
alces-297	415	12	.	.	PUNCT
alces-297	416	1	wildlife	wildlife	NOUN
alces-297	416	2	society	society	NOUN
alces-297	416	3	bulletin	bulletin	NOUN
alces-297	416	4	38	38	NUM
alces-297	416	5	:	:	PUNCT
alces-297	416	6	878–883	878–883	NUM
alces-297	416	7	.	.	PUNCT
alces-297	417	1	doi	doi	NOUN
alces-297	417	2	:	:	PUNCT
alces-297	417	3	10	10	NUM
alces-297	417	4	.	.	X
alces-297	417	5	1002/	1002/	NUM
alces-297	417	6	wsb.475	wsb.475	NOUN
alces-297	417	7	alces	alce	NOUN
alces-297	417	8	vol	vol	NOUN
alces-297	417	9	.	.	PROPN
alces-297	417	10	57	57	NUM
alces-297	417	11	,	,	PUNCT
alces-297	417	12	2021	2021	NUM
alces-297	417	13	modelling	model	VERB
alces-297	417	14	sustained	sustain	VERB
alces-297	417	15	yields	yield	NOUN
alces-297	417	16	for	for	ADP
alces-297	417	17	moose	moose	NOUN
alces-297	417	18	–	–	PUNCT
alces-297	417	19	hatter	hatter	NOUN
alces-297	417	20	.	.	PUNCT
alces-297	418	1	127	127	NUM
alces-297	419	1	_	_	PUNCT
alces-297	420	1	_	_	PUNCT
alces-297	421	1	_	_	PUNCT
alces-297	422	1	_	_	PUNCT
alces-297	422	2	_	_	PUNCT
alces-297	422	3	,	,	PUNCT
alces-297	422	4	and	and	CCONJ
alces-297	422	5	d.	d.	PROPN
alces-297	422	6	heard	hear	VERB
alces-297	422	7	.	.	PUNCT
alces-297	423	1	2014	2014	NUM
alces-297	423	2	.	.	PUNCT
alces-297	424	1	research	research	NOUN
alces-297	424	2	design	design	NOUN
alces-297	424	3	to	to	PART
alces-297	424	4	determine	determine	VERB
alces-297	424	5	factors	factor	NOUN
alces-297	424	6	affecting	affect	VERB
alces-297	424	7	moose	moose	NOUN
alces-297	424	8	population	population	NOUN
alces-297	424	9	change	change	NOUN
alces-297	424	10	in	in	ADP
alces-297	424	11	british	british	PROPN
alces-297	424	12	columbia	columbia	PROPN
alces-297	424	13	:	:	PUNCT
alces-297	424	14	testing	test	VERB
alces-297	424	15	the	the	DET
alces-297	424	16	landscape	landscape	NOUN
alces-297	424	17	change	change	NOUN
alces-297	424	18	hypothesis	hypothesis	NOUN
alces-297	424	19	.	.	PUNCT
alces-297	425	1	wildlife	wildlife	NOUN
alces-297	425	2	bulletin	bulletin	NOUN
alces-297	425	3	no	no	INTJ
alces-297	425	4	.	.	PUNCT
alces-297	426	1	b-126	b-126	NOUN
alces-297	426	2	.	.	PUNCT
alces-297	427	1	british	british	PROPN
alces-297	427	2	columbia	columbia	PROPN
alces-297	427	3	ministry	ministry	PROPN
alces-297	427	4	of	of	ADP
alces-297	427	5	forests	forest	NOUN
alces-297	427	6	,	,	PUNCT
alces-297	427	7	lands	land	NOUN
alces-297	427	8	and	and	CCONJ
alces-297	427	9	natural	natural	ADJ
alces-297	427	10	resource	resource	NOUN
alces-297	427	11	operations	operation	NOUN
alces-297	427	12	,	,	PUNCT
alces-297	427	13	victoria	victoria	PROPN
alces-297	427	14	,	,	PUNCT
alces-297	427	15	british	british	PROPN
alces-297	427	16	columbia	columbia	PROPN
alces-297	427	17	,	,	PUNCT
alces-297	427	18	canada	canada	PROPN
alces-297	427	19	.	.	PUNCT
alces-297	428	1	_	_	PUNCT
alces-297	429	1	_	_	PUNCT
alces-297	430	1	_	_	PUNCT
alces-297	431	1	_	_	PUNCT
alces-297	432	1	_	_	PUNCT
alces-297	432	2	,	,	PUNCT
alces-297	433	1	_	_	PUNCT
alces-297	434	1	_	_	PUNCT
alces-297	435	1	_	_	PUNCT
alces-297	436	1	_	_	PUNCT
alces-297	437	1	_	_	PUNCT
alces-297	437	2	,	,	PUNCT
alces-297	437	3	s.	s.	PROPN
alces-297	437	4	marshall	marshall	PROPN
alces-297	437	5	,	,	PUNCT
alces-297	437	6	c.	c.	PROPN
alces-297	437	7	procter	procter	PROPN
alces-297	437	8	,	,	PUNCT
alces-297	437	9	b.	b.	PROPN
alces-297	437	10	cadsand	cadsand	PROPN
alces-297	437	11	,	,	PUNCT
alces-297	437	12	d.	d.	PROPN
alces-297	437	13	lirette	lirette	PROPN
alces-297	437	14	,	,	PUNCT
alces-297	437	15	h.	h.	PROPN
alces-297	437	16	schindler	schindler	PROPN
alces-297	437	17	,	,	PUNCT
alces-297	437	18	m.	m.	PROPN
alces-297	437	19	bridger	bridger	PROPN
alces-297	437	20	,	,	PUNCT
alces-297	437	21	p.	p.	PROPN
alces-297	437	22	stent	stent	NOUN
alces-297	437	23	,	,	PUNCT
alces-297	437	24	a.	a.	NOUN
alces-297	437	25	walker	walker	PROPN
alces-297	437	26	,	,	PUNCT
alces-297	437	27	and	and	CCONJ
alces-297	437	28	m.	m.	NOUN
alces-297	437	29	klaczek	klaczek	PROPN
alces-297	437	30	.	.	PROPN
alces-297	438	1	2018	2018	NUM
alces-297	438	2	.	.	PUNCT
alces-297	439	1	moose	moose	NOUN
alces-297	439	2	population	population	NOUN
alces-297	439	3	dynamics	dynamic	NOUN
alces-297	439	4	during	during	ADP
alces-297	439	5	20	20	NUM
alces-297	439	6	years	year	NOUN
alces-297	439	7	of	of	ADP
alces-297	439	8	declining	decline	VERB
alces-297	439	9	harvest	harvest	NOUN
alces-297	439	10	in	in	ADP
alces-297	439	11	british	british	PROPN
alces-297	439	12	columbia	columbia	PROPN
alces-297	439	13	.	.	PUNCT
alces-297	440	1	alces	alce	NOUN
alces-297	440	2	54	54	NUM
alces-297	440	3	:	:	PUNCT
alces-297	440	4	101–119	101–119	NUM
alces-297	440	5	.	.	PUNCT
alces-297	441	1	_	_	PUNCT
alces-297	442	1	_	_	PUNCT
alces-297	443	1	_	_	PUNCT
alces-297	444	1	_	_	PUNCT
alces-297	445	1	_	_	PUNCT
alces-297	445	2	,	,	PUNCT
alces-297	445	3	c.	c.	PROPN
alces-297	445	4	procter	procter	PROPN
alces-297	445	5	,	,	PUNCT
alces-297	445	6	s.	s.	PROPN
alces-297	445	7	marshall	marshall	PROPN
alces-297	445	8	,	,	PUNCT
alces-297	445	9	h.	h.	PROPN
alces-297	445	10	schindler	schindler	PROPN
alces-297	445	11	,	,	PUNCT
alces-297	445	12	h.	h.	PROPN
alces-297	445	13	schwantje	schwantje	PROPN
alces-297	445	14	,	,	PUNCT
alces-297	445	15	m.	m.	NOUN
alces-297	445	16	scheideman	scheideman	NOUN
alces-297	445	17	,	,	PUNCT
alces-297	445	18	and	and	CCONJ
alces-297	445	19	d.	d.	PROPN
alces-297	445	20	hodder	hodder	PROPN
alces-297	445	21	.	.	PUNCT
alces-297	446	1	2019	2019	NUM
alces-297	446	2	.	.	PUNCT
alces-297	447	1	factors	factor	NOUN
alces-297	447	2	affecting	affect	VERB
alces-297	447	3	moose	moose	NOUN
alces-297	447	4	population	population	NOUN
alces-297	447	5	declines	decline	VERB
alces-297	447	6	in	in	ADP
alces-297	447	7	british	british	PROPN
alces-297	447	8	columbia	columbia	PROPN
alces-297	447	9	.	.	PUNCT
alces-297	448	1	2019	2019	NUM
alces-297	448	2	progress	progress	NOUN
alces-297	448	3	report	report	NOUN
alces-297	448	4	:	:	PUNCT
alces-297	448	5	february	february	PROPN
alces-297	448	6	2012	2012	NUM
alces-297	448	7	–	–	PUNCT
alces-297	448	8	may	may	PROPN
alces-297	448	9	2019	2019	NUM
alces-297	448	10	.	.	PUNCT
alces-297	449	1	wildlife	wildlife	NOUN
alces-297	449	2	working	working	NOUN
alces-297	449	3	report	report	NOUN
alces-297	449	4	no	no	INTJ
alces-297	449	5	.	.	PUNCT
alces-297	450	1	wr	wr	PROPN
alces-297	450	2	127	127	NUM
alces-297	450	3	.	.	PUNCT
alces-297	451	1	british	british	PROPN
alces-297	451	2	columbia	columbia	PROPN
alces-297	451	3	ministry	ministry	PROPN
alces-297	451	4	of	of	ADP
alces-297	451	5	forests	forest	NOUN
alces-297	451	6	,	,	PUNCT
alces-297	451	7	lands	land	NOUN
alces-297	451	8	and	and	CCONJ
alces-297	451	9	natural	natural	ADJ
alces-297	451	10	resource	resource	NOUN
alces-297	451	11	operations	operation	NOUN
alces-297	451	12	,	,	PUNCT
alces-297	451	13	victoria	victoria	PROPN
alces-297	451	14	,	,	PUNCT
alces-297	451	15	british	british	PROPN
alces-297	451	16	columbia	columbia	PROPN
alces-297	451	17	,	,	PUNCT
alces-297	451	18	canada	canada	PROPN
alces-297	451	19	.	.	PUNCT
alces-297	451	20	lake	lake	PROPN
alces-297	451	21	,	,	PUNCT
alces-297	451	22	b.	b.	PROPN
alces-297	451	23	c.	c.	PROPN
alces-297	451	24	,	,	PUNCT
alces-297	451	25	m.	m.	PROPN
alces-297	451	26	r.	r.	PROPN
alces-297	451	27	bertram	bertram	PROPN
alces-297	451	28	,	,	PUNCT
alces-297	451	29	n.	n.	PROPN
alces-297	451	30	guldager	guldager	PROPN
alces-297	451	31	,	,	PUNCT
alces-297	451	32	j.	j.	PROPN
alces-297	451	33	r.	r.	PROPN
alces-297	451	34	caikoski	caikoski	PROPN
alces-297	451	35	,	,	PUNCT
alces-297	451	36	and	and	CCONJ
alces-297	451	37	r.	r.	PROPN
alces-297	451	38	o.	o.	PROPN
alces-297	451	39	stephenson	stephenson	PROPN
alces-297	451	40	.	.	PUNCT
alces-297	452	1	2013	2013	NUM
alces-297	452	2	.	.	PUNCT
alces-297	453	1	wolf	wolf	PROPN
alces-297	453	2	kill	kill	VERB
alces-297	453	3	rates	rate	NOUN
alces-297	453	4	across	across	ADP
alces-297	453	5	winter	winter	NOUN
alces-297	453	6	in	in	ADP
alces-297	453	7	a	a	DET
alces-297	453	8	low	low	ADJ
alces-297	453	9	-	-	PUNCT
alces-297	453	10	density	density	NOUN
alces-297	453	11	moose	moose	NOUN
alces-297	453	12	system	system	NOUN
alces-297	453	13	in	in	ADP
alces-297	453	14	alaska	alaska	PROPN
alces-297	453	15	.	.	PUNCT
alces-297	454	1	journal	journal	PROPN
alces-297	454	2	of	of	ADP
alces-297	454	3	wildlife	wildlife	NOUN
alces-297	454	4	management	management	NOUN
alces-297	454	5	77	77	NUM
alces-297	454	6	:	:	PUNCT
alces-297	454	7	1512–1522	1512–1522	NUM
alces-297	454	8	.	.	PUNCT
alces-297	455	1	doi	doi	NOUN
alces-297	455	2	:	:	PUNCT
alces-297	455	3	10.1002	10.1002	NUM
alces-297	455	4	/	/	SYM
alces-297	455	5	jwmg.603	jwmg.603	PROPN
alces-297	455	6	mcnay	mcnay	PROPN
alces-297	455	7	,	,	PUNCT
alces-297	455	8	m.	m.	PROPN
alces-297	455	9	e.	e.	PROPN
alces-297	455	10	,	,	PUNCT
alces-297	455	11	and	and	CCONJ
alces-297	455	12	r.a	r.a	PROPN
alces-297	455	13	.	.	PROPN
alces-297	455	14	delong	delong	PROPN
alces-297	455	15	.	.	PUNCT
alces-297	456	1	1998	1998	NUM
alces-297	456	2	.	.	PUNCT
alces-297	457	1	development	development	NOUN
alces-297	457	2	and	and	CCONJ
alces-297	457	3	testing	testing	NOUN
alces-297	457	4	of	of	ADP
alces-297	457	5	a	a	DET
alces-297	457	6	general	general	ADJ
alces-297	457	7	predator	predator	NOUN
alces-297	457	8	-	-	PUNCT
alces-297	457	9	prey	prey	NOUN
alces-297	457	10	computer	computer	NOUN
alces-297	457	11	model	model	NOUN
alces-297	457	12	for	for	ADP
alces-297	457	13	use	use	NOUN
alces-297	457	14	in	in	ADP
alces-297	457	15	making	make	VERB
alces-297	457	16	management	management	NOUN
alces-297	457	17	decisions	decision	NOUN
alces-297	457	18	.	.	PUNCT
alces-297	458	1	federal	federal	ADJ
alces-297	458	2	aid	aid	NOUN
alces-297	458	3	in	in	ADP
alces-297	458	4	wildlife	wildlife	NOUN
alces-297	458	5	restoration	restoration	NOUN
alces-297	458	6	,	,	PUNCT
alces-297	458	7	research	research	NOUN
alces-297	458	8	final	final	ADJ
alces-297	458	9	report	report	NOUN
alces-297	458	10	,	,	PUNCT
alces-297	458	11	grants	grant	NOUN
alces-297	458	12	w-24	w-24	PROPN
alces-297	458	13	-	-	PUNCT
alces-297	458	14	1	1	NUM
alces-297	458	15	and	and	CCONJ
alces-297	458	16	w-24	w-24	PROPN
alces-297	458	17	-	-	PUNCT
alces-297	458	18	5	5	NUM
alces-297	458	19	,	,	PUNCT
alces-297	458	20	study	study	VERB
alces-297	458	21	1.46	1.46	NUM
alces-297	458	22	.	.	PUNCT
alces-297	459	1	alaska	alaska	PROPN
alces-297	459	2	department	department	PROPN
alces-297	459	3	of	of	ADP
alces-297	459	4	fish	fish	NOUN
alces-297	459	5	and	and	CCONJ
alces-297	459	6	game	game	NOUN
alces-297	459	7	,	,	PUNCT
alces-297	459	8	juneau	juneau	PROPN
alces-297	459	9	,	,	PUNCT
alces-297	459	10	alaska	alaska	PROPN
alces-297	459	11	,	,	PUNCT
alces-297	459	12	usa	usa	PROPN
alces-297	459	13	.	.	PROPN
alces-297	459	14	messier	messy	ADJ
alces-297	459	15	,	,	PUNCT
alces-297	459	16	f.	f.	PROPN
alces-297	459	17	1994	1994	NUM
alces-297	459	18	.	.	PUNCT
alces-297	460	1	ungulate	ungulate	ADJ
alces-297	460	2	population	population	NOUN
alces-297	460	3	models	model	NOUN
alces-297	460	4	with	with	ADP
alces-297	460	5	predation	predation	NOUN
alces-297	460	6	:	:	PUNCT
alces-297	460	7	a	a	DET
alces-297	460	8	case	case	NOUN
alces-297	460	9	study	study	NOUN
alces-297	460	10	with	with	ADP
alces-297	460	11	the	the	DET
alces-297	460	12	north	north	ADJ
alces-297	460	13	american	american	ADJ
alces-297	460	14	moose	moose	NOUN
alces-297	460	15	.	.	PUNCT
alces-297	461	1	ecology	ecology	NOUN
alces-297	461	2	75	75	NUM
alces-297	461	3	:	:	PUNCT
alces-297	461	4	478–488	478–488	NUM
alces-297	461	5	.	.	PUNCT
alces-297	462	1	doi	doi	NOUN
alces-297	462	2	:	:	PUNCT
alces-297	462	3	10.2307/1939551	10.2307/1939551	NUM
alces-297	462	4	murray	murray	PROPN
alces-297	462	5	,	,	PUNCT
alces-297	462	6	d.	d.	PROPN
alces-297	462	7	l.	l.	PROPN
alces-297	462	8	,	,	PUNCT
alces-297	462	9	e.	e.	PROPN
alces-297	462	10	w.	w.	PROPN
alces-297	462	11	cox	cox	PROPN
alces-297	462	12	,	,	PUNCT
alces-297	462	13	w.	w.	PROPN
alces-297	462	14	b.	b.	PROPN
alces-297	462	15	ballard	ballard	PROPN
alces-297	462	16	,	,	PUNCT
alces-297	462	17	h.	h.	PROPN
alces-297	462	18	a.	a.	PROPN
alces-297	462	19	whitlaw	whitlaw	PROPN
alces-297	462	20	,	,	PUNCT
alces-297	462	21	m.	m.	PROPN
alces-297	462	22	s.	s.	PROPN
alces-297	462	23	lenarz	lenarz	PROPN
alces-297	462	24	,	,	PUNCT
alces-297	462	25	t.	t.	PROPN
alces-297	462	26	w.	w.	PROPN
alces-297	462	27	custer	custer	PROPN
alces-297	462	28	,	,	PUNCT
alces-297	462	29	t.	t.	PROPN
alces-297	462	30	barnett	barnett	PROPN
alces-297	462	31	,	,	PUNCT
alces-297	462	32	and	and	CCONJ
alces-297	462	33	t.	t.	PROPN
alces-297	462	34	k.	k.	PROPN
alces-297	462	35	fuller	fuller	PROPN
alces-297	462	36	.	.	PUNCT
alces-297	463	1	2006	2006	NUM
alces-297	463	2	.	.	PUNCT
alces-297	464	1	pathogens	pathogen	NOUN
alces-297	464	2	,	,	PUNCT
alces-297	464	3	nutritional	nutritional	ADJ
alces-297	464	4	deficiency	deficiency	NOUN
alces-297	464	5	,	,	PUNCT
alces-297	464	6	and	and	CCONJ
alces-297	464	7	climate	climate	NOUN
alces-297	464	8	influences	influence	NOUN
alces-297	464	9	on	on	ADP
alces-297	464	10	a	a	DET
alces-297	464	11	declining	decline	VERB
alces-297	464	12	moose	moose	NOUN
alces-297	464	13	population	population	NOUN
alces-297	464	14	.	.	PUNCT
alces-297	465	1	wildlife	wildlife	NOUN
alces-297	465	2	monographs	monograph	NOUN
alces-297	465	3	166	166	NUM
alces-297	465	4	:	:	PUNCT
alces-297	465	5	1–30	1–30	PROPN
alces-297	465	6	.	.	PUNCT
alces-297	466	1	doi	doi	PROPN
alces-297	466	2	:	:	PUNCT
alces-297	466	3	10.2193/	10.2193/	NUM
alces-297	466	4	0084	0084	NUM
alces-297	466	5	-	-	SYM
alces-297	466	6	0173	0173	NUM
alces-297	466	7	(	(	PUNCT
alces-297	466	8	2006	2006	NUM
alces-297	466	9	)	)	PUNCT
alces-297	467	1	166[1	166[1	NUM
alces-297	467	2	:	:	PUNCT
alces-297	467	3	pndaci	pndaci	NOUN
alces-297	467	4	]	]	PUNCT
alces-297	467	5	2.0.co;2	2.0.co;2	NUM
alces-297	467	6	nilsen	nilsen	PROPN
alces-297	467	7	,	,	PUNCT
alces-297	467	8	e.	e.	PROPN
alces-297	467	9	b.	b.	PROPN
alces-297	467	10	,	,	PUNCT
alces-297	467	11	t.	t.	PROPN
alces-297	467	12	pettersen	pettersen	PROPN
alces-297	467	13	,	,	PUNCT
alces-297	467	14	h.	h.	PROPN
alces-297	467	15	gundersen	gundersen	PROPN
alces-297	467	16	,	,	PUNCT
alces-297	467	17	j.	j.	PROPN
alces-297	467	18	m.	m.	PROPN
alces-297	467	19	milner	milner	PROPN
alces-297	467	20	,	,	PUNCT
alces-297	467	21	a.	a.	NOUN
alces-297	467	22	mysterud	mysterud	NOUN
alces-297	467	23	,	,	PUNCT
alces-297	467	24	e.	e.	PROPN
alces-297	467	25	j.	j.	PROPN
alces-297	467	26	solberg	solberg	PROPN
alces-297	467	27	,	,	PUNCT
alces-297	467	28	h.	h.	PROPN
alces-297	467	29	p.	p.	PROPN
alces-297	467	30	andreassen	andreassen	PROPN
alces-297	467	31	,	,	PUNCT
alces-297	467	32	and	and	CCONJ
alces-297	467	33	n.	n.	PROPN
alces-297	467	34	c.	c.	PROPN
alces-297	467	35	stenseth	stenseth	PROPN
alces-297	467	36	.	.	PUNCT
alces-297	468	1	2005	2005	NUM
alces-297	468	2	.	.	PUNCT
alces-297	469	1	moose	moose	NOUN
alces-297	469	2	harvesting	harvesting	NOUN
alces-297	469	3	strategies	strategy	NOUN
alces-297	469	4	in	in	ADP
alces-297	469	5	the	the	DET
alces-297	469	6	presence	presence	NOUN
alces-297	469	7	of	of	ADP
alces-297	469	8	wolves	wolf	NOUN
alces-297	469	9	.	.	PUNCT
alces-297	470	1	journal	journal	NOUN
alces-297	470	2	of	of	ADP
alces-297	470	3	applied	applied	ADJ
alces-297	470	4	ecology	ecology	NOUN
alces-297	470	5	42	42	NUM
alces-297	470	6	:	:	PUNCT
alces-297	471	1	389–399	389–399	NUM
alces-297	471	2	.	.	PUNCT
alces-297	471	3	doi	doi	NOUN
alces-297	471	4	:	:	PUNCT
alces-297	471	5	10.1111/	10.1111/	NUM
alces-297	471	6	j.13652664.2005	j.13652664.2005	PROPN
alces-297	471	7	.	.	PUNCT
alces-297	472	1	01018.x	01018.x	PROPN
alces-297	472	2	person	person	NOUN
alces-297	472	3	,	,	PUNCT
alces-297	472	4	d.	d.	PROPN
alces-297	472	5	k.	k.	PROPN
alces-297	472	6	,	,	PUNCT
alces-297	472	7	r.	r.	PROPN
alces-297	472	8	t.	t.	PROPN
alces-297	472	9	bowyer	bowyer	PROPN
alces-297	472	10	,	,	PUNCT
alces-297	472	11	and	and	CCONJ
alces-297	472	12	v.	v.	ADP
alces-297	472	13	van	van	PROPN
alces-297	472	14	ballenberge	ballenberge	NOUN
alces-297	472	15	.	.	PUNCT
alces-297	473	1	2001	2001	NUM
alces-297	473	2	.	.	PUNCT
alces-297	474	1	density	density	NOUN
alces-297	474	2	dependence	dependence	NOUN
alces-297	474	3	of	of	ADP
alces-297	474	4	ungulates	ungulate	NOUN
alces-297	474	5	and	and	CCONJ
alces-297	474	6	functional	functional	ADJ
alces-297	474	7	responses	response	NOUN
alces-297	474	8	of	of	ADP
alces-297	474	9	wolves	wolf	NOUN
alces-297	474	10	:	:	PUNCT
alces-297	474	11	effects	effect	NOUN
alces-297	474	12	on	on	ADP
alces-297	474	13	predator	predator	NOUN
alces-297	474	14	-	-	PUNCT
alces-297	474	15	prey	prey	NOUN
alces-297	474	16	ratios	ratio	NOUN
alces-297	474	17	.	.	PUNCT
alces-297	475	1	alces	alce	VERB
alces-297	475	2	37	37	NUM
alces-297	475	3	:	:	PUNCT
alces-297	475	4	253–273	253–273	NUM
alces-297	475	5	.	.	PUNCT
alces-297	475	6	peterson	peterson	PROPN
alces-297	475	7	,	,	PUNCT
alces-297	475	8	r.	r.	PROPN
alces-297	475	9	o.	o.	PROPN
alces-297	475	10	,	,	PUNCT
alces-297	475	11	j.	j.	PROPN
alces-297	475	12	d.	d.	PROPN
alces-297	475	13	woolington	woolington	PROPN
alces-297	475	14	,	,	PUNCT
alces-297	475	15	and	and	CCONJ
alces-297	475	16	t.	t.	PROPN
alces-297	475	17	n.	n.	PROPN
alces-297	475	18	bailey.1984	bailey.1984	PROPN
alces-297	475	19	.	.	PUNCT
alces-297	476	1	wolves	wolf	NOUN
alces-297	476	2	of	of	ADP
alces-297	476	3	the	the	DET
alces-297	476	4	kenai	kenai	PROPN
alces-297	476	5	peninsula	peninsula	PROPN
alces-297	476	6	,	,	PUNCT
alces-297	476	7	alaska	alaska	PROPN
alces-297	476	8	.	.	PUNCT
alces-297	477	1	wildlife	wildlife	NOUN
alces-297	477	2	monographs	monograph	NOUN
alces-297	477	3	88	88	NUM
alces-297	477	4	:	:	PUNCT
alces-297	477	5	1–52	1–52	PROPN
alces-297	477	6	.	.	PUNCT
alces-297	478	1	rea	rea	PROPN
alces-297	478	2	,	,	PUNCT
alces-297	478	3	r.v	r.v	PROPN
alces-297	478	4	.	.	PROPN
alces-297	478	5	,	,	PUNCT
alces-297	478	6	l.	l.	PROPN
alces-297	478	7	ajala	ajala	PROPN
alces-297	478	8	-	-	PUNCT
alces-297	478	9	batista	batista	PROPN
alces-297	478	10	,	,	PUNCT
alces-297	478	11	d.	d.	PROPN
alces-297	478	12	a.	a.	PROPN
alces-297	478	13	aitken	aitken	PROPN
alces-297	478	14	,	,	PUNCT
alces-297	478	15	k.	k.	PROPN
alces-297	478	16	n.	n.	PROPN
alces-297	478	17	child	child	PROPN
alces-297	478	18	,	,	PUNCT
alces-297	478	19	n.	n.	PROPN
alces-297	478	20	thompson	thompson	PROPN
alces-297	478	21	,	,	PUNCT
alces-297	478	22	and	and	CCONJ
alces-297	478	23	d.	d.	PROPN
alces-297	478	24	p.	p.	PROPN
alces-297	478	25	hodder	hodder	PROPN
alces-297	478	26	.	.	PUNCT
alces-297	479	1	2019	2019	NUM
alces-297	479	2	.	.	PUNCT
alces-297	480	1	scat	scat	NOUN
alces-297	480	2	analysis	analysis	NOUN
alces-297	480	3	as	as	ADP
alces-297	480	4	a	a	DET
alces-297	480	5	preliminary	preliminary	ADJ
alces-297	480	6	assessment	assessment	NOUN
alces-297	480	7	of	of	ADP
alces-297	480	8	moose	moose	NOUN
alces-297	480	9	(	(	PUNCT
alces-297	480	10	alces	alces	PROPN
alces-297	480	11	alces	alce	NOUN
alces-297	480	12	andersoni	andersoni	NOUN
alces-297	480	13	)	)	PUNCT
alces-297	480	14	calf	calf	NOUN
alces-297	480	15	consumption	consumption	NOUN
alces-297	480	16	by	by	ADP
alces-297	480	17	bears	bear	NOUN
alces-297	480	18	(	(	PUNCT
alces-297	480	19	ursus	ursus	PROPN
alces-297	480	20	spp	spp	PROPN
alces-297	480	21	.	.	PUNCT
alces-297	480	22	)	)	PUNCT
alces-297	481	1	in	in	ADP
alces-297	481	2	north	north	NOUN
alces-297	481	3	-	-	PUNCT
alces-297	481	4	central	central	ADJ
alces-297	481	5	british	british	PROPN
alces-297	481	6	columbia	columbia	PROPN
alces-297	481	7	.	.	PUNCT
alces-297	482	1	animal	animal	NOUN
alces-297	482	2	biodiversity	biodiversity	NOUN
alces-297	482	3	and	and	CCONJ
alces-297	482	4	conservation	conservation	NOUN
alces-297	482	5	42	42	NUM
alces-297	482	6	:	:	PUNCT
alces-297	482	7	369–377	369–377	NUM
alces-297	482	8	.	.	PUNCT
alces-297	483	1	doi	doi	NOUN
alces-297	483	2	:	:	PUNCT
alces-297	483	3	10.32800/	10.32800/	NUM
alces-297	483	4	abc.2019.42.0369	abc.2019.42.0369	ADP
alces-297	483	5	sand	sand	NOUN
alces-297	483	6	,	,	PUNCT
alces-297	483	7	h.	h.	PROPN
alces-297	483	8	,	,	PUNCT
alces-297	483	9	p.	p.	PROPN
alces-297	483	10	wabakken	wabakken	PROPN
alces-297	483	11	,	,	PUNCT
alces-297	483	12	b.	b.	PROPN
alces-297	483	13	zimmermann	zimmermann	PROPN
alces-297	483	14	,	,	PUNCT
alces-297	483	15	o.	o.	PROPN
alces-297	483	16	johansson	johansson	PROPN
alces-297	483	17	,	,	PUNCT
alces-297	483	18	h.	h.	PROPN
alces-297	483	19	c.	c.	PROPN
alces-297	483	20	pedersen	pedersen	PROPN
alces-297	483	21	,	,	PUNCT
alces-297	483	22	and	and	CCONJ
alces-297	483	23	o.	o.	PROPN
alces-297	483	24	liberg	liberg	PROPN
alces-297	483	25	.	.	PUNCT
alces-297	484	1	2008	2008	NUM
alces-297	484	2	.	.	PUNCT
alces-297	485	1	summer	summer	NOUN
alces-297	485	2	kill	kill	NOUN
alces-297	485	3	rates	rate	NOUN
alces-297	485	4	and	and	CCONJ
alces-297	485	5	predation	predation	NOUN
alces-297	485	6	pattern	pattern	NOUN
alces-297	485	7	in	in	ADP
alces-297	485	8	a	a	DET
alces-297	485	9	wolf	wolf	ADJ
alces-297	485	10	–	–	PUNCT
alces-297	485	11	moose	moose	NOUN
alces-297	485	12	system	system	NOUN
alces-297	485	13	:	:	PUNCT
alces-297	485	14	can	can	AUX
alces-297	485	15	we	we	PRON
alces-297	485	16	rely	rely	VERB
alces-297	485	17	on	on	ADP
alces-297	485	18	winter	winter	NOUN
alces-297	485	19	estimates	estimate	NOUN
alces-297	485	20	?	?	PUNCT
alces-297	486	1	oecologia	oecologia	ADJ
alces-297	486	2	156	156	NUM
alces-297	486	3	:	:	PUNCT
alces-297	487	1	53–64	53–64	NUM
alces-297	487	2	.	.	PUNCT
alces-297	487	3	doi	doi	PROPN
alces-297	487	4	:	:	PUNCT
alces-297	487	5	10.1007/	10.1007/	NUM
alces-297	487	6	s00442	s00442	PROPN
alces-297	487	7	-	-	PUNCT
alces-297	487	8	008	008	NUM
alces-297	487	9	-	-	PUNCT
alces-297	487	10	0969	0969	NUM
alces-297	487	11	-	-	SYM
alces-297	487	12	2	2	NUM
alces-297	487	13	sæther	sæther	NOUN
alces-297	487	14	,	,	PUNCT
alces-297	487	15	b.-e	b.-e	PROPN
alces-297	487	16	.	.	PROPN
alces-297	487	17	,	,	PUNCT
alces-297	487	18	s.	s.	PROPN
alces-297	487	19	engen	engen	PROPN
alces-297	487	20	,	,	PUNCT
alces-297	487	21	and	and	CCONJ
alces-297	487	22	e.	e.	PROPN
alces-297	487	23	j.	j.	PROPN
alces-297	487	24	solberg	solberg	PROPN
alces-297	487	25	.	.	PUNCT
alces-297	488	1	2001	2001	NUM
alces-297	488	2	.	.	PUNCT
alces-297	489	1	optimal	optimal	ADJ
alces-297	489	2	harvest	harvest	NOUN
alces-297	489	3	of	of	ADP
alces-297	489	4	age	age	NOUN
alces-297	489	5	-	-	PUNCT
alces-297	489	6	structured	structure	VERB
alces-297	489	7	populations	population	NOUN
alces-297	489	8	of	of	ADP
alces-297	489	9	moose	moose	NOUN
alces-297	489	10	alces	alce	NOUN
alces-297	489	11	alces	alce	NOUN
alces-297	489	12	in	in	ADP
alces-297	489	13	a	a	DET
alces-297	489	14	fluctuating	fluctuate	VERB
alces-297	489	15	environment	environment	NOUN
alces-297	489	16	.	.	PUNCT
alces-297	490	1	wildlife	wildlife	NOUN
alces-297	490	2	biology	biology	NOUN
alces-297	490	3	7	7	NUM
alces-297	490	4	:	:	SYM
alces-297	490	5	171–179	171–179	NUM
alces-297	490	6	.	.	PUNCT
alces-297	491	1	doi	doi	NOUN
alces-297	491	2	:	:	PUNCT
alces-297	491	3	10.2981	10.2981	NUM
alces-297	491	4	/	/	SYM
alces-297	491	5	wlb.2001.021	wlb.2001.021	PROPN
alces-297	491	6	serrouya	serrouya	PROPN
alces-297	491	7	,	,	PUNCT
alces-297	491	8	r.	r.	PROPN
alces-297	491	9	,	,	PUNCT
alces-297	491	10	b.	b.	PROPN
alces-297	491	11	n.	n.	PROPN
alces-297	491	12	mclellan	mclellan	PROPN
alces-297	491	13	,	,	PUNCT
alces-297	491	14	and	and	CCONJ
alces-297	491	15	s.	s.	PROPN
alces-297	491	16	boutin	boutin	PROPN
alces-297	491	17	.	.	PUNCT
alces-297	492	1	2015	2015	NUM
alces-297	492	2	.	.	PUNCT
alces-297	493	1	testing	test	VERB
alces-297	493	2	predator	predator	NOUN
alces-297	493	3	-	-	PUNCT
alces-297	493	4	prey	prey	NOUN
alces-297	493	5	theory	theory	NOUN
alces-297	493	6	using	use	VERB
alces-297	493	7	broad	broad	ADJ
alces-297	493	8	-	-	PUNCT
alces-297	493	9	scale	scale	NOUN
alces-297	493	10	manipulations	manipulation	NOUN
alces-297	493	11	and	and	CCONJ
alces-297	493	12	independent	independent	ADJ
alces-297	493	13	validation	validation	NOUN
alces-297	493	14	.	.	PUNCT
alces-297	494	1	journal	journal	NOUN
alces-297	494	2	of	of	ADP
alces-297	494	3	animal	animal	NOUN
alces-297	494	4	ecology	ecology	NOUN
alces-297	494	5	84	84	NUM
alces-297	494	6	:	:	PUNCT
alces-297	494	7	1600–1609	1600–1609	NUM
alces-297	494	8	.	.	PUNCT
alces-297	495	1	doi	doi	NOUN
alces-297	495	2	:	:	PUNCT
alces-297	495	3	10	10	NUM
alces-297	495	4	.	.	PUNCT
alces-297	496	1	1111/1365	1111/1365	NUM
alces-297	496	2	-	-	SYM
alces-297	496	3	2656.12413	2656.12413	NUM
alces-297	496	4	modelling	model	VERB
alces-297	496	5	sustained	sustain	VERB
alces-297	496	6	yields	yield	NOUN
alces-297	496	7	for	for	ADP
alces-297	496	8	moose	moose	NOUN
alces-297	496	9	–	–	PUNCT
alces-297	496	10	hatter	hatter	NOUN
alces-297	496	11	.	.	PUNCT
alces-297	497	1	alces	alce	NOUN
alces-297	497	2	vol	vol	NOUN
alces-297	497	3	.	.	PROPN
alces-297	497	4	57	57	NUM
alces-297	497	5	,	,	PUNCT
alces-297	497	6	2021	2021	NUM
alces-297	497	7	128	128	NUM
alces-297	497	8	sinclair	sinclair	NOUN
alces-297	497	9	,	,	PUNCT
alces-297	497	10	a.	a.	PROPN
alces-297	497	11	r.	r.	PROPN
alces-297	497	12	e.	e.	PROPN
alces-297	497	13	,	,	PUNCT
alces-297	497	14	j.	j.	PROPN
alces-297	497	15	m.	m.	PROPN
alces-297	497	16	fryxell	fryxell	PROPN
alces-297	497	17	,	,	PUNCT
alces-297	497	18	and	and	CCONJ
alces-297	497	19	g.	g.	PROPN
alces-297	497	20	caughley	caughley	PROPN
alces-297	497	21	.	.	PUNCT
alces-297	498	1	2006	2006	NUM
alces-297	498	2	.	.	PUNCT
alces-297	499	1	wildlife	wildlife	NOUN
alces-297	499	2	ecology	ecology	NOUN
alces-297	499	3	,	,	PUNCT
alces-297	499	4	conservation	conservation	NOUN
alces-297	499	5	,	,	PUNCT
alces-297	499	6	and	and	CCONJ
alces-297	499	7	management	management	NOUN
alces-297	499	8	,	,	PUNCT
alces-297	499	9	second	second	ADJ
alces-297	499	10	edition	edition	NOUN
alces-297	499	11	.	.	PUNCT
alces-297	500	1	blackwell	blackwell	PROPN
alces-297	500	2	publishing	publishing	PROPN
alces-297	500	3	,	,	PUNCT
alces-297	500	4	malden	malden	PROPN
alces-297	500	5	,	,	PUNCT
alces-297	500	6	massachusetts	massachusetts	PROPN
alces-297	500	7	,	,	PUNCT
alces-297	500	8	usa	usa	PROPN
alces-297	500	9	.	.	PROPN
alces-297	500	10	timmermann	timmermann	PROPN
alces-297	500	11	,	,	PUNCT
alces-297	500	12	t.	t.	PROPN
alces-297	500	13	1992	1992	NUM
alces-297	500	14	.	.	PUNCT
alces-297	501	1	moose	moose	PROPN
alces-297	501	2	sociobiology	sociobiology	NOUN
alces-297	501	3	and	and	CCONJ
alces-297	501	4	implications	implication	NOUN
alces-297	501	5	for	for	ADP
alces-297	501	6	harvest	harvest	NOUN
alces-297	501	7	.	.	PUNCT
alces-297	502	1	alces	alce	NOUN
alces-297	502	2	28	28	NUM
alces-297	502	3	:	:	PUNCT
alces-297	502	4	59–77	59–77	NUM
alces-297	502	5	.	.	PUNCT
alces-297	503	1	van	van	PROPN
alces-297	503	2	ballenberghe	ballenberghe	NOUN
alces-297	503	3	,	,	PUNCT
alces-297	503	4	v.	v.	ADV
alces-297	503	5	,	,	PUNCT
alces-297	503	6	and	and	CCONJ
alces-297	503	7	j.	j.	PROPN
alces-297	503	8	dart	dart	PROPN
alces-297	503	9	.	.	PUNCT
alces-297	504	1	1982	1982	NUM
alces-297	504	2	.	.	PUNCT
alces-297	505	1	harvest	harvest	NOUN
alces-297	505	2	yields	yield	NOUN
alces-297	505	3	from	from	ADP
alces-297	505	4	moose	moose	NOUN
alces-297	505	5	populations	population	NOUN
alces-297	505	6	subject	subject	ADJ
alces-297	505	7	to	to	ADP
alces-297	505	8	wolf	wolf	NOUN
alces-297	505	9	and	and	CCONJ
alces-297	505	10	bear	bear	VERB
alces-297	505	11	predation	predation	NOUN
alces-297	505	12	.	.	PUNCT
alces-297	506	1	alces	alce	NOUN
alces-297	506	2	18	18	NUM
alces-297	506	3	:	:	PUNCT
alces-297	506	4	258–275	258–275	NUM
alces-297	506	5	.	.	PUNCT
alces-297	507	1	_	_	PUNCT
alces-297	508	1	_	_	PUNCT
alces-297	509	1	_	_	PUNCT
alces-297	510	1	_	_	PUNCT
alces-297	510	2	_	_	PUNCT
alces-297	510	3	,	,	PUNCT
alces-297	510	4	and	and	CCONJ
alces-297	510	5	w.	w.	PROPN
alces-297	510	6	b.	b.	PROPN
alces-297	510	7	ballard	ballard	PROPN
alces-297	510	8	.	.	PUNCT
alces-297	510	9	1994	1994	NUM
alces-297	510	10	.	.	PUNCT
alces-297	511	1	limitation	limitation	NOUN
alces-297	511	2	and	and	CCONJ
alces-297	511	3	regulation	regulation	NOUN
alces-297	511	4	of	of	ADP
alces-297	511	5	moose	moose	NOUN
alces-297	511	6	populations	population	NOUN
alces-297	511	7	:	:	PUNCT
alces-297	511	8	the	the	DET
alces-297	511	9	role	role	NOUN
alces-297	511	10	of	of	ADP
alces-297	511	11	predation	predation	NOUN
alces-297	511	12	.	.	PUNCT
alces-297	512	1	canadian	canadian	ADJ
alces-297	512	2	journal	journal	PROPN
alces-297	512	3	of	of	ADP
alces-297	512	4	zoology	zoology	NOUN
alces-297	512	5	72	72	NUM
alces-297	512	6	:	:	PUNCT
alces-297	512	7	2071–2077	2071–2077	NUM
alces-297	512	8	.	.	PUNCT
alces-297	513	1	doi	doi	NOUN
alces-297	513	2	:	:	PUNCT
alces-297	513	3	10.1139/	10.1139/	NUM
alces-297	513	4	z94	z94	NUM
alces-297	513	5	-	-	PUNCT
alces-297	513	6	277	277	NUM
alces-297	513	7	xu	xu	PROPN
alces-297	513	8	,	,	PUNCT
alces-297	513	9	c.	c.	PROPN
alces-297	513	10	,	,	PUNCT
alces-297	513	11	and	and	CCONJ
alces-297	513	12	m.	m.	PROPN
alces-297	513	13	s.	s.	PROPN
alces-297	513	14	boyce	boyce	PROPN
alces-297	513	15	.	.	PUNCT
alces-297	514	1	2010	2010	NUM
alces-297	514	2	.	.	PUNCT
alces-297	515	1	optimal	optimal	ADJ
alces-297	515	2	harvesting	harvesting	NOUN
alces-297	515	3	of	of	ADP
alces-297	515	4	moose	moose	NOUN
alces-297	515	5	in	in	ADP
alces-297	515	6	alberta	alberta	PROPN
alces-297	515	7	.	.	PUNCT
alces-297	516	1	alces	alce	VERB
alces-297	516	2	46	46	NUM
alces-297	516	3	:	:	PUNCT
alces-297	516	4	15–35	15–35	NUM
alces-297	516	5	.	.	PUNCT
alces-297	517	1	young	young	PROPN
alces-297	517	2	,	,	PUNCT
alces-297	517	3	d.	d.	PROPN
alces-297	517	4	d.	d.	PROPN
alces-297	517	5	,	,	PUNCT
alces-297	517	6	jr	jr	PROPN
alces-297	517	7	.	.	PROPN
alces-297	517	8	,	,	PUNCT
alces-297	517	9	and	and	CCONJ
alces-297	517	10	r.	r.	PROPN
alces-297	517	11	d.	d.	PROPN
alces-297	517	12	boertje	boertje	PROPN
alces-297	517	13	.	.	PUNCT
alces-297	518	1	2008	2008	NUM
alces-297	518	2	.	.	PUNCT
alces-297	519	1	recovery	recovery	NOUN
alces-297	519	2	of	of	ADP
alces-297	519	3	low	low	ADJ
alces-297	519	4	bull	bull	NOUN
alces-297	519	5	:	:	PUNCT
alces-297	519	6	cow	cow	NOUN
alces-297	519	7	ratios	ratio	NOUN
alces-297	519	8	of	of	ADP
alces-297	519	9	moose	moose	NOUN
alces-297	519	10	in	in	ADP
alces-297	519	11	interior	interior	ADJ
alces-297	519	12	alaska	alaska	PROPN
alces-297	519	13	.	.	PUNCT
alces-297	520	1	alces	alce	NOUN
alces-297	520	2	44	44	NUM
alces-297	520	3	:	:	PUNCT
alces-297	520	4	65–71	65–71	NUM
alces-297	520	5	.	.	PUNCT
alces-297	521	1	_	_	PUNCT
alces-297	522	1	_	_	PUNCT
alces-297	523	1	_	_	PUNCT
alces-297	524	1	_	_	PUNCT
alces-297	525	1	_	_	PUNCT
alces-297	525	2	,	,	PUNCT
alces-297	526	1	_	_	PUNCT
alces-297	527	1	_	_	PUNCT
alces-297	528	1	_	_	PUNCT
alces-297	529	1	_	_	PUNCT
alces-297	530	1	_	_	PUNCT
alces-297	530	2	,	,	PUNCT
alces-297	530	3	t.	t.	PROPN
alces-297	530	4	seaton	seaton	PROPN
alces-297	530	5	,	,	PUNCT
alces-297	530	6	and	and	CCONJ
alces-297	530	7	k.	k.	PROPN
alces-297	530	8	a.	a.	PROPN
alces-297	530	9	kellie	kellie	PROPN
alces-297	530	10	.	.	PUNCT
alces-297	531	1	2006	2006	NUM
alces-297	531	2	.	.	PUNCT
alces-297	532	1	intensive	intensive	ADJ
alces-297	532	2	management	management	NOUN
alces-297	532	3	of	of	ADP
alces-297	532	4	moose	moose	NOUN
alces-297	532	5	at	at	ADP
alces-297	532	6	high	high	ADJ
alces-297	532	7	density	density	NOUN
alces-297	532	8	:	:	PUNCT
alces-297	532	9	impediments	impediment	NOUN
alces-297	532	10	,	,	PUNCT
alces-297	532	11	achievements	achievement	NOUN
alces-297	532	12	,	,	PUNCT
alces-297	532	13	and	and	CCONJ
alces-297	532	14	recommendations	recommendation	NOUN
alces-297	532	15	.	.	PUNCT
alces-297	533	1	alces	alce	NOUN
alces-297	533	2	42	42	NUM
alces-297	533	3	:	:	PUNCT
alces-297	533	4	41–48	41–48	NUM
alces-297	533	5	.	.	PUNCT
alces-297	534	1	alces	alce	NOUN
alces-297	534	2	vol	vol	NOUN
alces-297	534	3	.	.	PROPN
alces-297	534	4	57	57	NUM
alces-297	534	5	,	,	PUNCT
alces-297	534	6	2021	2021	NUM
alces-297	534	7	modelling	model	VERB
alces-297	534	8	sustained	sustain	VERB
alces-297	534	9	yields	yield	NOUN
alces-297	534	10	for	for	ADP
alces-297	534	11	moose	moose	NOUN
alces-297	534	12	–	–	PUNCT
alces-297	534	13	hatter	hatter	NOUN
alces-297	534	14	.	.	PUNCT
alces-297	535	1	129	129	NUM
alces-297	535	2	appendix	appendix	NOUN
alces-297	535	3	1	1	NUM
alces-297	535	4	.	.	X
alces-297	535	5	relationship	relationship	NOUN
alces-297	535	6	between	between	ADP
alces-297	535	7	wolf	wolf	PROPN
alces-297	535	8	finite	finite	ADJ
alces-297	535	9	rate	rate	NOUN
alces-297	535	10	of	of	ADP
alces-297	535	11	increase	increase	NOUN
alces-297	535	12	(	(	PUNCT
alces-297	535	13	λ	λ	NOUN
alces-297	535	14	)	)	PUNCT
alces-297	535	15	and	and	CCONJ
alces-297	535	16	the	the	DET
alces-297	535	17	wolves	wolf	NOUN
alces-297	535	18	:	:	PUNCT
alces-297	535	19	moose	moose	NOUN
alces-297	535	20	ratio	ratio	NOUN
alces-297	535	21	from	from	ADP
alces-297	535	22	17	17	NUM
alces-297	535	23	north	north	ADJ
alces-297	535	24	american	american	ADJ
alces-297	535	25	wolf	wolf	PROPN
alces-297	535	26	populations	population	NOUN
alces-297	535	27	.	.	PUNCT
alces-297	536	1	y	y	NOUN
alces-297	536	2	=	=	SYM
alces-297	536	3	1.534e-10.77x	1.534e-10.77x	NUM
alces-297	536	4	r²	r²	NOUN
alces-297	536	5	=	=	NOUN
alces-297	536	6	0.78	0.78	NUM
alces-297	536	7	0	0	NUM
alces-297	536	8	0.2	0.2	NUM
alces-297	536	9	0.4	0.4	NUM
alces-297	536	10	0.6	0.6	NUM
alces-297	536	11	0.8	0.8	NUM
alces-297	536	12	1.0	1.0	NUM
alces-297	536	13	1.2	1.2	NUM
alces-297	536	14	1.4	1.4	NUM
alces-297	536	15	1.6	1.6	NUM
alces-297	536	16	0	0	NUM
alces-297	536	17	0.02	0.02	NUM
alces-297	536	18	0.04	0.04	NUM
alces-297	536	19	0.06	0.06	NUM
alces-297	536	20	0.08	0.08	NUM
alces-297	536	21	0.1	0.1	NUM
alces-297	536	22	w	w	PROPN
alces-297	536	23	ol	ol	PROPN
alces-297	536	24	f	f	PROPN
alces-297	536	25	fi	fi	PROPN
alces-297	536	26	ni	ni	PROPN
alces-297	536	27	te	te	PROPN
alces-297	536	28	ra	ra	PROPN
alces-297	536	29	te	te	INTJ
alces-297	536	30	o	o	X
alces-297	537	1	f	f	X
alces-297	537	2	i	i	PRON
alces-297	537	3	nc	nc	PROPN
alces-297	537	4	re	re	VERB
alces-297	537	5	as	as	ADP
alces-297	537	6	e	e	PROPN
alces-297	537	7	,	,	PUNCT
alces-297	537	8	λ	λ	NOUN
alces-297	537	9	wolves	wolf	NOUN
alces-297	537	10	/	/	SYM
alces-297	537	11	moose	moose	NOUN
alces-297	537	12	location	location	NOUN
alces-297	537	13	wolves	wolf	NOUN
alces-297	537	14	/	/	SYM
alces-297	537	15	moose1	moose1	NOUN
alces-297	537	16	wolf	wolf	PROPN
alces-297	537	17	λ	λ	PROPN
alces-297	537	18	reference	reference	NOUN
alces-297	537	19	alaska	alaska	PROPN
alces-297	537	20	0.0083	0.0083	NUM
alces-297	537	21	1.340	1.340	NUM
alces-297	537	22	keith	keith	PROPN
alces-297	537	23	1983	1983	NUM
alces-297	537	24	alberta	alberta	PROPN
alces-297	538	1	0.0085	0.0085	NUM
alces-297	538	2	1.460	1.460	NUM
alces-297	538	3	keith	keith	PROPN
alces-297	538	4	1983	1983	NUM
alces-297	538	5	michigan	michigan	PROPN
alces-297	538	6	0.0135	0.0135	NUM
alces-297	538	7	1.390	1.390	NUM
alces-297	538	8	keith	keith	PROPN
alces-297	538	9	1983	1983	NUM
alces-297	538	10	minnesota	minnesota	PROPN
alces-297	538	11	0.0148	0.0148	NUM
alces-297	538	12	1.310	1.310	NUM
alces-297	538	13	keith	keith	PROPN
alces-297	538	14	1983	1983	NUM
alces-297	538	15	denali	denali	PROPN
alces-297	538	16	park	park	NOUN
alces-297	538	17	0.0159	0.0159	NUM
alces-297	538	18	1.280	1.280	NUM
alces-297	538	19	mcnay	mcnay	NOUN
alces-297	538	20	and	and	CCONJ
alces-297	538	21	delong	delong	PROPN
alces-297	538	22	19982	19982	NUM
alces-297	538	23	kenai	kenai	PROPN
alces-297	538	24	,	,	PUNCT
alces-297	538	25	alaska	alaska	PROPN
alces-297	538	26	0.0174	0.0174	NUM
alces-297	538	27	1.180	1.180	NUM
alces-297	538	28	eberhardt	eberhardt	PROPN
alces-297	538	29	and	and	CCONJ
alces-297	538	30	peterson	peterson	PROPN
alces-297	538	31	1999	1999	NUM
alces-297	538	32	interior	interior	PROPN
alces-297	538	33	alaska	alaska	NOUN
alces-297	538	34	0.0183	0.0183	NUM
alces-297	538	35	1.280	1.280	NUM
alces-297	538	36	mcnay	mcnay	NOUN
alces-297	538	37	and	and	CCONJ
alces-297	538	38	delong	delong	PROPN
alces-297	538	39	19982	19982	NUM
alces-297	538	40	denali	denali	PROPN
alces-297	538	41	park	park	NOUN
alces-297	538	42	0.0260	0.0260	NUM
alces-297	538	43	0.900	0.900	NUM
alces-297	538	44	mcnay	mcnay	NOUN
alces-297	538	45	and	and	CCONJ
alces-297	538	46	delong	delong	PROPN
alces-297	538	47	19982	19982	NUM
alces-297	538	48	alberta	alberta	PROPN
alces-297	538	49	0.0306	0.0306	NUM
alces-297	538	50	1.210	1.210	NUM
alces-297	538	51	keith	keith	PROPN
alces-297	538	52	1983	1983	NUM
alces-297	538	53	michigan	michigan	NOUN
alces-297	538	54	0.0310	0.0310	NUM
alces-297	538	55	1.150	1.150	NUM
alces-297	538	56	keith	keith	PROPN
alces-297	538	57	1983	1983	NUM
alces-297	538	58	nelchina	nelchina	PROPN
alces-297	538	59	,	,	PUNCT
alces-297	538	60	alaska	alaska	PROPN
alces-297	538	61	0.0313	0.0313	NUM
alces-297	538	62	1.280	1.280	NUM
alces-297	538	63	eberhardt	eberhardt	PROPN
alces-297	538	64	and	and	CCONJ
alces-297	538	65	peterson	peterson	PROPN
alces-297	538	66	1999	1999	NUM
alces-297	538	67	ontario	ontario	NOUN
alces-297	538	68	0.0336	0.0336	NUM
alces-297	538	69	1.200	1.200	NUM
alces-297	538	70	keith	keith	PROPN
alces-297	538	71	1983	1983	NUM
alces-297	538	72	nc	nc	PROPN
alces-297	538	73	minnesota	minnesota	PROPN
alces-297	538	74	0.0373	0.0373	NUM
alces-297	538	75	1.070	1.070	NUM
alces-297	538	76	eberhardt	eberhardt	PROPN
alces-297	538	77	and	and	CCONJ
alces-297	538	78	peterson	peterson	PROPN
alces-297	538	79	1999	1999	NUM
alces-297	538	80	isle	isle	PROPN
alces-297	538	81	royale	royale	PROPN
alces-297	538	82	0.0376	0.0376	PROPN
alces-297	538	83	1.004	1.004	NUM
alces-297	538	84	https://isleroyalewolf.org/2	https://isleroyalewolf.org/2	PROPN
alces-297	538	85	isle	isle	NOUN
alces-297	538	86	royale	royale	NOUN
alces-297	538	87	0.0382	0.0382	NUM
alces-297	538	88	0.820	0.820	NUM
alces-297	538	89	mcnay	mcnay	NOUN
alces-297	538	90	and	and	CCONJ
alces-297	538	91	delong	delong	PROPN
alces-297	538	92	19982	19982	NUM
alces-297	538	93	quebec	quebec	PROPN
alces-297	538	94	0.0400	0.0400	NUM
alces-297	538	95	1.080	1.080	NUM
alces-297	538	96	eberhardt	eberhardt	PROPN
alces-297	538	97	and	and	CCONJ
alces-297	538	98	peterson	peterson	PROPN
alces-297	538	99	1999	1999	NUM
alces-297	538	100	interior	interior	PROPN
alces-297	538	101	alaska	alaska	PROPN
alces-297	538	102	0.0870	0.0870	PROPN
alces-297	538	103	0.590	0.590	NUM
alces-297	538	104	mcnay	mcnay	PROPN
alces-297	538	105	and	and	CCONJ
alces-297	538	106	delong	delong	PROPN
alces-297	538	107	19982	19982	NUM
alces-297	538	108	1wolves	1wolves	PROPN
alces-297	538	109	per	per	ADP
alces-297	538	110	moose	moose	NOUN
alces-297	538	111	biomass	biomass	NOUN
alces-297	538	112	equivalent	equivalent	NOUN
alces-297	538	113	(	(	PUNCT
alces-297	538	114	keith	keith	PROPN
alces-297	538	115	1983	1983	NUM
alces-297	538	116	)	)	PUNCT
alces-297	538	117	.	.	PUNCT
alces-297	539	1	2additional	2additional	NUM
alces-297	539	2	data	datum	NOUN
alces-297	539	3	used	use	VERB
alces-297	539	4	in	in	ADP
alces-297	539	5	this	this	DET
alces-297	539	6	study	study	NOUN
alces-297	539	7	to	to	PART
alces-297	539	8	estimate	estimate	VERB
alces-297	539	9	the	the	DET
alces-297	539	10	wolves	wolf	NOUN
alces-297	539	11	:	:	PUNCT
alces-297	539	12	moose	moose	NOUN
alces-297	539	13	equilibrium	equilibrium	NOUN
alces-297	539	14	ratio	ratio	NOUN
alces-297	539	15	.	.	PUNCT
alces-297	540	1	https://isleroyalewolf.org/	https://isleroyalewolf.org/	PRON