id	sid	tid	token	lemma	pos
alces-343	1	1	4301.pdf	4301.pdf	PROPN
alces-343	1	2	alces	alce	NOUN
alces-343	1	3	vol	vol	NOUN
alces-343	1	4	.	.	PROPN
alces-343	1	5	43	43	NUM
alces-343	1	6	,	,	PUNCT
alces-343	1	7	2007	2007	NUM
alces-343	1	8	courtois	courtois	NOUN
alces-343	1	9	and	and	CCONJ
alces-343	1	10	ouellet	ouellet	NOUN
alces-343	1	11	impact	impact	NOUN
alces-343	1	12	of	of	ADP
alces-343	1	13	moose	moose	NOUN
alces-343	1	14	and	and	CCONJ
alces-343	1	15	wolves	wolf	NOUN
alces-343	1	16	on	on	ADP
alces-343	1	17	caribou	caribou	NOUN
alces-343	1	18	13	13	NUM
alces-343	1	19	modeling	model	VERB
alces-343	1	20	the	the	DET
alces-343	1	21	impact	impact	NOUN
alces-343	1	22	of	of	ADP
alces-343	1	23	moose	moose	NOUN
alces-343	1	24	and	and	CCONJ
alces-343	1	25	wolf	wolf	PROPN
alces-343	1	26	management	management	NOUN
alces-343	1	27	on	on	ADP
alces-343	1	28	persistence	persistence	NOUN
alces-343	1	29	of	of	ADP
alces-343	1	30	woodland	woodland	PROPN
alces-343	1	31	caribou	caribou	PROPN
alces-343	1	32	réhaume	réhaume	NOUN
alces-343	1	33	courtois1,2	courtois1,2	PROPN
alces-343	1	34	and	and	CCONJ
alces-343	1	35	jean	jean	PROPN
alces-343	1	36	-	-	PUNCT
alces-343	1	37	pierre	pierre	PROPN
alces-343	1	38	ouellet3	ouellet3	ADJ
alces-343	1	39	1ministère	1ministère	NUM
alces-343	1	40	des	de	NOUN
alces-343	1	41	ressources	ressource	NOUN
alces-343	1	42	naturelles	naturelles	X
alces-343	1	43	et	et	PROPN
alces-343	1	44	de	de	X
alces-343	1	45	la	la	PROPN
alces-343	1	46	faune	faune	PROPN
alces-343	1	47	,	,	PUNCT
alces-343	1	48	direction	direction	NOUN
alces-343	1	49	du	du	PROPN
alces-343	1	50	développement	développement	X
alces-343	1	51	de	de	PROPN
alces-343	1	52	la	la	PROPN
alces-343	1	53	faune	faune	PROPN
alces-343	1	54	,	,	PUNCT
alces-343	1	55	880	880	NUM
alces-343	1	56	,	,	PUNCT
alces-343	1	57	ch	ch	NOUN
alces-343	1	58	.	.	PROPN
alces-343	1	59	ste	ste	PROPN
alces-343	1	60	-	-	PUNCT
alces-343	1	61	foy	foy	PROPN
alces-343	1	62	,	,	PUNCT
alces-343	1	63	2e	2e	PROPN
alces-343	1	64	étage	étage	PROPN
alces-343	1	65	,	,	PUNCT
alces-343	1	66	québec	québec	PROPN
alces-343	1	67	,	,	PUNCT
alces-343	1	68	pq	pq	PROPN
alces-343	1	69	,	,	PUNCT
alces-343	1	70	canada	canada	PROPN
alces-343	1	71	g1s	g1s	PROPN
alces-343	1	72	4x4	4x4	NUM
alces-343	1	73	;	;	PUNCT
alces-343	1	74	2université	2université	NUM
alces-343	1	75	laval	laval	NOUN
alces-343	1	76	,	,	PUNCT
alces-343	1	77	département	département	X
alces-343	1	78	de	de	X
alces-343	1	79	foresterie	foresterie	PROPN
alces-343	1	80	et	et	PROPN
alces-343	1	81	de	de	X
alces-343	1	82	géomatique	géomatique	PROPN
alces-343	1	83	,	,	PUNCT
alces-343	1	84	cité	cité	NOUN
alces-343	1	85	universitaire	universitaire	NOUN
alces-343	1	86	,	,	PUNCT
alces-343	1	87	ste	ste	PROPN
alces-343	1	88	-	-	PUNCT
alces-343	1	89	foy	foy	PROPN
alces-343	1	90	,	,	PUNCT
alces-343	1	91	pq	pq	PROPN
alces-343	1	92	,	,	PUNCT
alces-343	1	93	canada	canada	PROPN
alces-343	1	94	g1k	g1k	PROPN
alces-343	1	95	7p4	7p4	NUM
alces-343	1	96	;	;	PUNCT
alces-343	1	97	3jean	3jean	NUM
alces-343	1	98	-	-	PUNCT
alces-343	1	99	pierre	pierre	NOUN
alces-343	1	100	ouellet	ouellet	NOUN
alces-343	1	101	,	,	PUNCT
alces-343	1	102	université	université	PROPN
alces-343	1	103	du	du	PROPN
alces-343	1	104	québec	québec	PROPN
alces-343	1	105	à	à	X
alces-343	1	106	rimouski	rimouski	NOUN
alces-343	1	107	,	,	PUNCT
alces-343	1	108	département	département	PROPN
alces-343	1	109	de	de	X
alces-343	1	110	biologie	biologie	PROPN
alces-343	1	111	,	,	PUNCT
alces-343	1	112	centre	centre	NOUN
alces-343	1	113	d’études	d’étude	VERB
alces-343	1	114	nordiques	nordique	NOUN
alces-343	1	115	,	,	PUNCT
alces-343	1	116	300	300	NUM
alces-343	1	117	,	,	PUNCT
alces-343	1	118	allée	allée	NOUN
alces-343	1	119	des	des	PROPN
alces-343	1	120	ursulines	ursuline	NOUN
alces-343	1	121	,	,	PUNCT
alces-343	1	122	rimouski	rimouski	NOUN
alces-343	1	123	,	,	PUNCT
alces-343	1	124	pq	pq	INTJ
alces-343	1	125	,	,	PUNCT
alces-343	1	126	canada	canada	PROPN
alces-343	2	1	g5l	g5l	PROPN
alces-343	2	2	3a1	3a1	NUM
alces-343	2	3	abstract	abstract	NOUN
alces-343	2	4	:	:	PUNCT
alces-343	2	5	limiting	limit	VERB
alces-343	2	6	factors	factor	NOUN
alces-343	2	7	of	of	ADP
alces-343	2	8	caribou	caribou	NOUN
alces-343	2	9	(	(	PUNCT
alces-343	2	10	rangifer	rangifer	VERB
alces-343	2	11	tarandus	tarandus	NOUN
alces-343	2	12	)	)	PUNCT
alces-343	2	13	populations	population	NOUN
alces-343	2	14	vary	vary	VERB
alces-343	2	15	regionally	regionally	ADV
alces-343	2	16	.	.	PUNCT
alces-343	3	1	in	in	ADP
alces-343	3	2	tundra	tundra	NOUN
alces-343	3	3	environments	environment	NOUN
alces-343	3	4	,	,	PUNCT
alces-343	3	5	this	this	DET
alces-343	3	6	species	specie	NOUN
alces-343	3	7	appears	appear	VERB
alces-343	3	8	to	to	PART
alces-343	3	9	be	be	AUX
alces-343	3	10	regulated	regulate	VERB
alces-343	3	11	by	by	ADP
alces-343	3	12	food	food	NOUN
alces-343	3	13	,	,	PUNCT
alces-343	3	14	either	either	CCONJ
alces-343	3	15	because	because	SCONJ
alces-343	3	16	wolves	wolf	NOUN
alces-343	3	17	(	(	PUNCT
alces-343	3	18	canis	canis	NOUN
alces-343	3	19	lupus	lupus	NOUN
alces-343	3	20	)	)	PUNCT
alces-343	3	21	are	be	AUX
alces-343	3	22	absent	absent	ADJ
alces-343	3	23	or	or	CCONJ
alces-343	3	24	because	because	SCONJ
alces-343	3	25	migration	migration	NOUN
alces-343	3	26	of	of	ADP
alces-343	3	27	caribou	caribou	NOUN
alces-343	3	28	allows	allow	VERB
alces-343	3	29	escape	escape	NOUN
alces-343	3	30	from	from	ADP
alces-343	3	31	predation	predation	NOUN
alces-343	3	32	during	during	ADP
alces-343	3	33	part	part	NOUN
alces-343	3	34	of	of	ADP
alces-343	3	35	the	the	DET
alces-343	3	36	year	year	NOUN
alces-343	3	37	.	.	PUNCT
alces-343	4	1	in	in	ADP
alces-343	4	2	the	the	DET
alces-343	4	3	boreal	boreal	NOUN
alces-343	4	4	forest	forest	NOUN
alces-343	4	5	,	,	PUNCT
alces-343	4	6	the	the	DET
alces-343	4	7	main	main	ADJ
alces-343	4	8	limiting	limit	VERB
alces-343	4	9	factors	factor	NOUN
alces-343	4	10	are	be	AUX
alces-343	4	11	hunting	hunt	VERB
alces-343	4	12	and	and	CCONJ
alces-343	4	13	predation	predation	NOUN
alces-343	4	14	but	but	CCONJ
alces-343	4	15	because	because	SCONJ
alces-343	4	16	of	of	ADP
alces-343	4	17	low	low	ADJ
alces-343	4	18	caribou	caribou	NOUN
alces-343	4	19	densities	density	NOUN
alces-343	4	20	,	,	PUNCT
alces-343	4	21	no	no	DET
alces-343	4	22	regulation	regulation	NOUN
alces-343	4	23	mechanism	mechanism	NOUN
alces-343	4	24	seems	seem	VERB
alces-343	4	25	to	to	PART
alces-343	4	26	exist	exist	VERB
alces-343	4	27	between	between	ADP
alces-343	4	28	caribou	caribou	NOUN
alces-343	4	29	and	and	CCONJ
alces-343	4	30	wolves	wolf	NOUN
alces-343	4	31	.	.	PUNCT
alces-343	5	1	moose	moose	NOUN
alces-343	5	2	(	(	PUNCT
alces-343	5	3	alces	alce	NOUN
alces-343	5	4	alces	alce	NOUN
alces-343	5	5	)	)	PUNCT
alces-343	5	6	is	be	AUX
alces-343	5	7	the	the	DET
alces-343	5	8	primary	primary	ADJ
alces-343	5	9	prey	prey	NOUN
alces-343	5	10	species	specie	NOUN
alces-343	5	11	of	of	ADP
alces-343	5	12	wolves	wolf	NOUN
alces-343	5	13	and	and	CCONJ
alces-343	5	14	consequently	consequently	ADV
alces-343	5	15	,	,	PUNCT
alces-343	5	16	if	if	SCONJ
alces-343	5	17	moose	moose	NOUN
alces-343	5	18	abundance	abundance	NOUN
alces-343	5	19	increases	increase	NOUN
alces-343	5	20	,	,	PUNCT
alces-343	5	21	wolves	wolf	NOUN
alces-343	5	22	should	should	AUX
alces-343	5	23	also	also	ADV
alces-343	5	24	increase	increase	VERB
alces-343	5	25	,	,	PUNCT
alces-343	5	26	independently	independently	ADV
alces-343	5	27	of	of	ADP
alces-343	5	28	the	the	DET
alces-343	5	29	caribou	caribou	NOUN
alces-343	5	30	population	population	NOUN
alces-343	5	31	.	.	PUNCT
alces-343	6	1	thus	thus	ADV
alces-343	6	2	,	,	PUNCT
alces-343	6	3	caribou	caribou	PROPN
alces-343	6	4	could	could	AUX
alces-343	6	5	experience	experience	VERB
alces-343	6	6	high	high	ADJ
alces-343	6	7	predation	predation	NOUN
alces-343	6	8	rates	rate	NOUN
alces-343	6	9	and	and	CCONJ
alces-343	6	10	be	be	AUX
alces-343	6	11	eliminated	eliminate	VERB
alces-343	6	12	in	in	ADP
alces-343	6	13	high	high	ADJ
alces-343	6	14	wolf	wolf	NOUN
alces-343	6	15	densities	density	NOUN
alces-343	6	16	.	.	PUNCT
alces-343	7	1	here	here	ADV
alces-343	7	2	we	we	PRON
alces-343	7	3	attempted	attempt	VERB
alces-343	7	4	to	to	PART
alces-343	7	5	identify	identify	VERB
alces-343	7	6	the	the	DET
alces-343	7	7	necessary	necessary	ADJ
alces-343	7	8	conditions	condition	NOUN
alces-343	7	9	to	to	PART
alces-343	7	10	maintain	maintain	VERB
alces-343	7	11	caribou	caribou	NOUN
alces-343	7	12	numbers	number	NOUN
alces-343	7	13	in	in	ADP
alces-343	7	14	the	the	DET
alces-343	7	15	presence	presence	NOUN
alces-343	7	16	of	of	ADP
alces-343	7	17	moose	moose	NOUN
alces-343	7	18	.	.	PUNCT
alces-343	8	1	to	to	PART
alces-343	8	2	do	do	VERB
alces-343	8	3	so	so	ADV
alces-343	8	4	,	,	PUNCT
alces-343	8	5	we	we	PRON
alces-343	8	6	built	build	VERB
alces-343	8	7	a	a	DET
alces-343	8	8	deterministic	deterministic	ADJ
alces-343	8	9	model	model	NOUN
alces-343	8	10	that	that	PRON
alces-343	8	11	simulated	simulate	VERB
alces-343	8	12	the	the	DET
alces-343	8	13	relationship	relationship	NOUN
alces-343	8	14	between	between	ADP
alces-343	8	15	a	a	DET
alces-343	8	16	caribou	caribou	NOUN
alces-343	8	17	population	population	NOUN
alces-343	8	18	regulated	regulate	VERB
alces-343	8	19	by	by	ADP
alces-343	8	20	food	food	NOUN
alces-343	8	21	competition	competition	NOUN
alces-343	8	22	and	and	CCONJ
alces-343	8	23	limited	limit	VERB
alces-343	8	24	by	by	ADP
alces-343	8	25	predation	predation	NOUN
alces-343	8	26	,	,	PUNCT
alces-343	8	27	a	a	DET
alces-343	8	28	moose	moose	NOUN
alces-343	8	29	population	population	NOUN
alces-343	8	30	regulated	regulate	VERB
alces-343	8	31	by	by	ADP
alces-343	8	32	predation	predation	NOUN
alces-343	8	33	,	,	PUNCT
alces-343	8	34	and	and	CCONJ
alces-343	8	35	a	a	DET
alces-343	8	36	wolf	wolf	ADJ
alces-343	8	37	population	population	NOUN
alces-343	8	38	,	,	PUNCT
alces-343	8	39	the	the	DET
alces-343	8	40	abundance	abundance	NOUN
alces-343	8	41	of	of	ADP
alces-343	8	42	which	which	PRON
alces-343	8	43	is	be	AUX
alces-343	8	44	determined	determine	VERB
alces-343	8	45	by	by	ADP
alces-343	8	46	moose	moose	NOUN
alces-343	8	47	abundance	abundance	NOUN
alces-343	8	48	.	.	PUNCT
alces-343	9	1	at	at	ADP
alces-343	9	2	current	current	ADJ
alces-343	9	3	hunting	hunting	NOUN
alces-343	9	4	rates	rate	NOUN
alces-343	9	5	for	for	ADP
alces-343	9	6	caribou	caribou	NOUN
alces-343	9	7	and	and	CCONJ
alces-343	9	8	moose	moose	NOUN
alces-343	9	9	in	in	ADP
alces-343	9	10	the	the	DET
alces-343	9	11	boreal	boreal	NOUN
alces-343	9	12	forest	forest	NOUN
alces-343	9	13	,	,	PUNCT
alces-343	9	14	and	and	CCONJ
alces-343	9	15	in	in	ADP
alces-343	9	16	the	the	DET
alces-343	9	17	absence	absence	NOUN
alces-343	9	18	of	of	ADP
alces-343	9	19	wolf	wolf	NOUN
alces-343	9	20	trapping	trapping	NOUN
alces-343	9	21	,	,	PUNCT
alces-343	9	22	the	the	DET
alces-343	9	23	model	model	NOUN
alces-343	9	24	predicted	predict	VERB
alces-343	9	25	that	that	SCONJ
alces-343	9	26	the	the	DET
alces-343	9	27	caribou	caribou	NOUN
alces-343	9	28	population	population	NOUN
alces-343	9	29	would	would	AUX
alces-343	9	30	be	be	AUX
alces-343	9	31	extirpated	extirpate	VERB
alces-343	9	32	in	in	ADP
alces-343	9	33	approximately	approximately	ADV
alces-343	9	34	100	100	NUM
alces-343	9	35	years	year	NOUN
alces-343	9	36	.	.	PUNCT
alces-343	10	1	wolf	wolf	NOUN
alces-343	10	2	trapping	trap	VERB
alces-343	10	3	was	be	AUX
alces-343	10	4	not	not	PART
alces-343	10	5	adequate	adequate	ADJ
alces-343	10	6	to	to	PART
alces-343	10	7	conserve	conserve	VERB
alces-343	10	8	the	the	DET
alces-343	10	9	caribou	caribou	NOUN
alces-343	10	10	population	population	NOUN
alces-343	10	11	unless	unless	SCONJ
alces-343	10	12	very	very	ADV
alces-343	10	13	intensive	intensive	ADJ
alces-343	10	14	control	control	NOUN
alces-343	10	15	was	be	AUX
alces-343	10	16	undertaken	undertake	VERB
alces-343	10	17	.	.	PUNCT
alces-343	11	1	in	in	ADP
alces-343	11	2	the	the	DET
alces-343	11	3	absence	absence	NOUN
alces-343	11	4	of	of	ADP
alces-343	11	5	trapping	trap	VERB
alces-343	11	6	,	,	PUNCT
alces-343	11	7	cessation	cessation	NOUN
alces-343	11	8	of	of	ADP
alces-343	11	9	caribou	caribou	NOUN
alces-343	11	10	hunting	hunting	NOUN
alces-343	11	11	allowed	allow	VERB
alces-343	11	12	a	a	DET
alces-343	11	13	3	3	ADJ
alces-343	11	14	-	-	ADJ
alces-343	11	15	fold	fold	ADJ
alces-343	11	16	increase	increase	NOUN
alces-343	11	17	in	in	ADP
alces-343	11	18	caribou	caribou	NOUN
alces-343	11	19	numbers	number	NOUN
alces-343	11	20	over	over	ADP
alces-343	11	21	the	the	DET
alces-343	11	22	long	long	ADJ
alces-343	11	23	term	term	NOUN
alces-343	11	24	,	,	PUNCT
alces-343	11	25	if	if	SCONJ
alces-343	11	26	the	the	DET
alces-343	11	27	moose	moose	NOUN
alces-343	11	28	population	population	NOUN
alces-343	11	29	remained	remain	VERB
alces-343	11	30	low	low	ADJ
alces-343	11	31	.	.	PUNCT
alces-343	12	1	according	accord	VERB
alces-343	12	2	to	to	ADP
alces-343	12	3	our	our	PRON
alces-343	12	4	model	model	NOUN
alces-343	12	5	,	,	PUNCT
alces-343	12	6	the	the	DET
alces-343	12	7	best	good	ADJ
alces-343	12	8	management	management	NOUN
alces-343	12	9	measure	measure	NOUN
alces-343	12	10	for	for	ADP
alces-343	12	11	caribou	caribou	NOUN
alces-343	12	12	consisted	consist	VERB
alces-343	12	13	of	of	ADP
alces-343	12	14	maintaining	maintain	VERB
alces-343	12	15	a	a	DET
alces-343	12	16	low	low	ADJ
alces-343	12	17	moose	moose	NOUN
alces-343	12	18	density	density	NOUN
alces-343	12	19	through	through	ADP
alces-343	12	20	appropriate	appropriate	ADJ
alces-343	12	21	population	population	NOUN
alces-343	12	22	and	and	CCONJ
alces-343	12	23	habitat	habitat	NOUN
alces-343	12	24	management	management	NOUN
alces-343	12	25	strategies	strategy	NOUN
alces-343	12	26	,	,	PUNCT
alces-343	12	27	which	which	PRON
alces-343	12	28	prevented	prevent	VERB
alces-343	12	29	expansion	expansion	NOUN
alces-343	12	30	of	of	ADP
alces-343	12	31	the	the	DET
alces-343	12	32	wolf	wolf	PROPN
alces-343	12	33	population	population	NOUN
alces-343	12	34	and	and	CCONJ
alces-343	12	35	limited	limited	ADJ
alces-343	12	36	predation	predation	NOUN
alces-343	12	37	on	on	ADP
alces-343	12	38	caribou	caribou	PROPN
alces-343	12	39	.	.	PUNCT
alces-343	13	1	alces	alce	NOUN
alces-343	13	2	vol	vol	NOUN
alces-343	13	3	.	.	PROPN
alces-343	14	1	43	43	NUM
alces-343	14	2	:	:	PUNCT
alces-343	14	3	13	13	NUM
alces-343	14	4	-	-	SYM
alces-343	14	5	27	27	NUM
alces-343	14	6	(	(	PUNCT
alces-343	14	7	2007	2007	NUM
alces-343	14	8	)	)	PUNCT
alces-343	14	9	key	key	ADJ
alces-343	14	10	words	word	NOUN
alces-343	14	11	:	:	PUNCT
alces-343	14	12	caribou	caribou	NOUN
alces-343	14	13	,	,	PUNCT
alces-343	14	14	hunting	hunting	NOUN
alces-343	14	15	,	,	PUNCT
alces-343	14	16	interactions	interaction	NOUN
alces-343	14	17	,	,	PUNCT
alces-343	14	18	moose	moose	NOUN
alces-343	14	19	,	,	PUNCT
alces-343	14	20	predation	predation	NOUN
alces-343	14	21	,	,	PUNCT
alces-343	14	22	simulation	simulation	NOUN
alces-343	14	23	,	,	PUNCT
alces-343	14	24	trapping	trap	VERB
alces-343	14	25	,	,	PUNCT
alces-343	14	26	wolf	wolf	NOUN
alces-343	14	27	the	the	DET
alces-343	14	28	hypothesis	hypothesis	NOUN
alces-343	14	29	of	of	ADP
alces-343	14	30	exploitation	exploitation	NOUN
alces-343	14	31	ecosystems	ecosystem	NOUN
alces-343	14	32	predicts	predict	VERB
alces-343	14	33	that	that	SCONJ
alces-343	14	34	the	the	DET
alces-343	14	35	number	number	NOUN
alces-343	14	36	of	of	ADP
alces-343	14	37	trophic	trophic	ADJ
alces-343	14	38	levels	level	NOUN
alces-343	14	39	and	and	CCONJ
alces-343	14	40	population	population	NOUN
alces-343	14	41	regulating	regulate	VERB
alces-343	14	42	factors	factor	NOUN
alces-343	14	43	depend	depend	VERB
alces-343	14	44	on	on	ADP
alces-343	14	45	the	the	DET
alces-343	14	46	primary	primary	ADJ
alces-343	14	47	productivity	productivity	NOUN
alces-343	14	48	of	of	ADP
alces-343	14	49	an	an	DET
alces-343	14	50	ecosystem	ecosystem	NOUN
alces-343	14	51	(	(	PUNCT
alces-343	14	52	oksanen	oksanen	NOUN
alces-343	14	53	et	et	PROPN
alces-343	14	54	al	al	PROPN
alces-343	14	55	.	.	PROPN
alces-343	14	56	1981	1981	NUM
alces-343	14	57	,	,	PUNCT
alces-343	14	58	oksanen	oksanen	NOUN
alces-343	14	59	and	and	CCONJ
alces-343	14	60	oksanen	oksanen	NOUN
alces-343	14	61	2000	2000	NUM
alces-343	14	62	)	)	PUNCT
alces-343	14	63	.	.	PUNCT
alces-343	15	1	in	in	ADP
alces-343	15	2	very	very	ADV
alces-343	15	3	poor	poor	ADJ
alces-343	15	4	environments	environment	NOUN
alces-343	15	5	(	(	PUNCT
alces-343	15	6	e.g.	e.g.	ADV
alces-343	15	7	,	,	PUNCT
alces-343	15	8	high	high	ADJ
alces-343	15	9	arctic	arctic	ADJ
alces-343	15	10	and	and	CCONJ
alces-343	15	11	deserts	desert	NOUN
alces-343	15	12	,	,	PUNCT
alces-343	15	13	where	where	SCONJ
alces-343	15	14	productivity	productivity	NOUN
alces-343	15	15	is	be	AUX
alces-343	15	16	<	<	X
alces-343	15	17	40	40	NUM
alces-343	15	18	g	g	NOUN
alces-343	15	19	/	/	SYM
alces-343	15	20	m2	m2	PROPN
alces-343	15	21	/	/	SYM
alces-343	15	22	year	year	NOUN
alces-343	15	23	)	)	PUNCT
alces-343	15	24	,	,	PUNCT
alces-343	15	25	ecosystems	ecosystem	NOUN
alces-343	15	26	consist	consist	VERB
alces-343	15	27	of	of	ADP
alces-343	15	28	vegetation	vegetation	NOUN
alces-343	15	29	communities	community	NOUN
alces-343	15	30	that	that	PRON
alces-343	15	31	are	be	AUX
alces-343	15	32	regulated	regulate	VERB
alces-343	15	33	by	by	ADP
alces-343	15	34	competition	competition	NOUN
alces-343	15	35	for	for	ADP
alces-343	15	36	resources	resource	NOUN
alces-343	15	37	.	.	PUNCT
alces-343	16	1	in	in	ADP
alces-343	16	2	poor	poor	ADJ
alces-343	16	3	environments	environment	NOUN
alces-343	16	4	(	(	PUNCT
alces-343	16	5	e.g.	e.g.	ADV
alces-343	16	6	,	,	PUNCT
alces-343	16	7	tundra	tundra	VERB
alces-343	16	8	:	:	PUNCT
alces-343	16	9	40	40	NUM
alces-343	16	10	–	–	SYM
alces-343	16	11	700	700	NUM
alces-343	16	12	g	g	NOUN
alces-343	16	13	/	/	SYM
alces-343	16	14	m2	m2	PROPN
alces-343	16	15	/	/	SYM
alces-343	16	16	year	year	NOUN
alces-343	16	17	)	)	PUNCT
alces-343	16	18	,	,	PUNCT
alces-343	16	19	plants	plant	NOUN
alces-343	16	20	are	be	AUX
alces-343	16	21	regulated	regulate	VERB
alces-343	16	22	by	by	ADP
alces-343	16	23	herbivores	herbivore	NOUN
alces-343	16	24	.	.	PUNCT
alces-343	17	1	in	in	ADP
alces-343	17	2	contrast	contrast	NOUN
alces-343	17	3	,	,	PUNCT
alces-343	17	4	more	more	ADV
alces-343	17	5	productive	productive	ADJ
alces-343	17	6	environments	environment	NOUN
alces-343	17	7	are	be	AUX
alces-343	17	8	made	make	VERB
alces-343	17	9	up	up	ADP
alces-343	17	10	of	of	ADP
alces-343	17	11	three	three	NUM
alces-343	17	12	trophic	trophic	ADJ
alces-343	17	13	levels	level	NOUN
alces-343	17	14	that	that	PRON
alces-343	17	15	are	be	AUX
alces-343	17	16	regulated	regulate	VERB
alces-343	17	17	from	from	ADP
alces-343	17	18	the	the	DET
alces-343	17	19	top	top	NOUN
alces-343	17	20	.	.	PUNCT
alces-343	18	1	here	here	ADV
alces-343	18	2	,	,	PUNCT
alces-343	18	3	herbivores	herbivore	NOUN
alces-343	18	4	are	be	AUX
alces-343	18	5	regulated	regulate	VERB
alces-343	18	6	by	by	ADP
alces-343	18	7	carnivores	carnivore	NOUN
alces-343	18	8	that	that	PRON
alces-343	18	9	have	have	VERB
alces-343	18	10	only	only	ADV
alces-343	18	11	a	a	DET
alces-343	18	12	limited	limited	ADJ
alces-343	18	13	impact	impact	NOUN
alces-343	18	14	on	on	ADP
alces-343	18	15	vegetation	vegetation	NOUN
alces-343	18	16	(	(	PUNCT
alces-343	18	17	oksanen	oksanen	NOUN
alces-343	18	18	1988	1988	NUM
alces-343	18	19	)	)	PUNCT
alces-343	18	20	,	,	PUNCT
alces-343	18	21	so	so	CCONJ
alces-343	18	22	both	both	DET
alces-343	18	23	carnivores	carnivore	NOUN
alces-343	18	24	and	and	CCONJ
alces-343	18	25	plants	plant	NOUN
alces-343	18	26	because	because	SCONJ
alces-343	18	27	regulating	regulate	VERB
alces-343	18	28	factors	factor	NOUN
alces-343	18	29	of	of	ADP
alces-343	18	30	caribou	caribou	NOUN
alces-343	18	31	(	(	PUNCT
alces-343	18	32	rangifer	rangifer	PROPN
alces-343	18	33	tarandus	tarandus	NOUN
alces-343	18	34	)	)	PUNCT
alces-343	18	35	vary	vary	VERB
alces-343	18	36	between	between	ADP
alces-343	18	37	ecotypes	ecotype	NOUN
alces-343	18	38	,	,	PUNCT
alces-343	18	39	the	the	DET
alces-343	18	40	exploitation	exploitation	NOUN
alces-343	18	41	ecosystem	ecosystem	NOUN
alces-343	18	42	model	model	NOUN
alces-343	18	43	seems	seem	VERB
alces-343	18	44	appropriate	appropriate	ADJ
alces-343	18	45	for	for	ADP
alces-343	18	46	prediction	prediction	NOUN
alces-343	18	47	of	of	ADP
alces-343	18	48	caribou	caribou	NOUN
alces-343	18	49	population	population	NOUN
alces-343	18	50	changes	change	NOUN
alces-343	18	51	.	.	PUNCT
alces-343	19	1	in	in	ADP
alces-343	19	2	tundra	tundra	NOUN
alces-343	19	3	environments	environment	NOUN
alces-343	19	4	,	,	PUNCT
alces-343	19	5	low	low	ADJ
alces-343	19	6	productivity	productivity	NOUN
alces-343	19	7	and	and	CCONJ
alces-343	19	8	low	low	ADJ
alces-343	19	9	carrying	carrying	NOUN
alces-343	19	10	capacity	capacity	NOUN
alces-343	19	11	conbiological	conbiological	ADJ
alces-343	19	12	communities	community	NOUN
alces-343	19	13	.	.	PUNCT
alces-343	20	1	predators	predator	NOUN
alces-343	20	2	are	be	AUX
alces-343	20	3	even	even	ADV
alces-343	20	4	absent	absent	ADJ
alces-343	20	5	on	on	ADP
alces-343	20	6	certain	certain	ADJ
alces-343	20	7	arctic	arctic	ADJ
alces-343	20	8	islands	island	NOUN
alces-343	20	9	(	(	PUNCT
alces-343	20	10	klein	klein	PROPN
alces-343	20	11	1968	1968	NUM
alces-343	20	12	,	,	PUNCT
alces-343	20	13	ouellet	ouellet	NOUN
alces-343	20	14	et	et	PROPN
alces-343	20	15	al	al	PROPN
alces-343	20	16	.	.	PROPN
alces-343	20	17	1996	1996	NUM
alces-343	20	18	)	)	PUNCT
alces-343	20	19	and	and	CCONJ
alces-343	20	20	in	in	ADP
alces-343	20	21	such	such	ADJ
alces-343	20	22	cases	case	NOUN
alces-343	20	23	,	,	PUNCT
alces-343	20	24	caribou	caribou	NOUN
alces-343	20	25	impact	impact	NOUN
alces-343	20	26	of	of	ADP
alces-343	20	27	moose	moose	NOUN
alces-343	20	28	and	and	CCONJ
alces-343	20	29	wolves	wolf	NOUN
alces-343	20	30	on	on	ADP
alces-343	20	31	caribou	caribou	NOUN
alces-343	20	32	–	–	PUNCT
alces-343	20	33	courtois	courtois	NOUN
alces-343	20	34	and	and	CCONJ
alces-343	20	35	ouellet	ouellet	PROPN
alces-343	20	36	alces	alces	PROPN
alces-343	20	37	vol	vol	NOUN
alces-343	20	38	.	.	PROPN
alces-343	21	1	43	43	NUM
alces-343	21	2	,	,	PUNCT
alces-343	21	3	2007	2007	NUM
alces-343	21	4	14	14	NUM
alces-343	21	5	populations	population	NOUN
alces-343	21	6	are	be	AUX
alces-343	21	7	regulated	regulate	VERB
alces-343	21	8	by	by	ADP
alces-343	21	9	competition	competition	NOUN
alces-343	21	10	for	for	ADP
alces-343	21	11	food	food	NOUN
alces-343	21	12	and	and	CCONJ
alces-343	21	13	population	population	NOUN
alces-343	21	14	changes	change	NOUN
alces-343	21	15	can	can	AUX
alces-343	21	16	be	be	AUX
alces-343	21	17	described	describe	VERB
alces-343	21	18	by	by	ADP
alces-343	21	19	a	a	DET
alces-343	21	20	logistic	logistic	ADJ
alces-343	21	21	model	model	NOUN
alces-343	21	22	(	(	PUNCT
alces-343	21	23	caughley	caughley	NOUN
alces-343	21	24	1977	1977	NUM
alces-343	21	25	)	)	PUNCT
alces-343	21	26	.	.	PUNCT
alces-343	22	1	net	net	ADJ
alces-343	22	2	population	population	NOUN
alces-343	22	3	growth	growth	NOUN
alces-343	22	4	rate	rate	NOUN
alces-343	22	5	varies	vary	VERB
alces-343	22	6	with	with	ADP
alces-343	22	7	density	density	NOUN
alces-343	22	8	:	:	PUNCT
alces-343	22	9	and	and	CCONJ
alces-343	22	10	then	then	ADV
alces-343	22	11	growth	growth	NOUN
alces-343	22	12	rate	rate	NOUN
alces-343	22	13	gradually	gradually	ADV
alces-343	22	14	slows	slow	VERB
alces-343	22	15	down	down	ADP
alces-343	22	16	as	as	ADP
alces-343	22	17	populations	population	NOUN
alces-343	22	18	approach	approach	NOUN
alces-343	22	19	carrying	carry	VERB
alces-343	22	20	capacity	capacity	NOUN
alces-343	22	21	.	.	PUNCT
alces-343	23	1	in	in	ADP
alces-343	23	2	such	such	ADJ
alces-343	23	3	environments	environment	NOUN
alces-343	23	4	,	,	PUNCT
alces-343	23	5	the	the	DET
alces-343	23	6	carrying	carry	VERB
alces-343	23	7	capacity	capacity	NOUN
alces-343	23	8	seems	seem	VERB
alces-343	23	9	to	to	PART
alces-343	23	10	be	be	AUX
alces-343	23	11	on	on	ADP
alces-343	23	12	the	the	DET
alces-343	23	13	order	order	NOUN
alces-343	23	14	of	of	ADP
alces-343	23	15	60	60	NUM
alces-343	23	16	–	–	SYM
alces-343	23	17	100	100	NUM
alces-343	23	18	caribou	caribou	NOUN
alces-343	23	19	per	per	ADP
alces-343	23	20	100	100	NUM
alces-343	23	21	km2	km2	NOUN
alces-343	23	22	,	,	PUNCT
alces-343	23	23	as	as	SCONJ
alces-343	23	24	observed	observe	VERB
alces-343	23	25	on	on	ADP
alces-343	23	26	coats	coat	NOUN
alces-343	23	27	island	island	NOUN
alces-343	23	28	in	in	ADP
alces-343	23	29	hudson	hudson	PROPN
alces-343	23	30	bay	bay	PROPN
alces-343	23	31	where	where	SCONJ
alces-343	23	32	caribou	caribou	PROPN
alces-343	23	33	have	have	AUX
alces-343	23	34	occurred	occur	VERB
alces-343	23	35	since	since	SCONJ
alces-343	23	36	at	at	ADP
alces-343	23	37	least	least	ADJ
alces-343	23	38	the	the	DET
alces-343	23	39	beginning	beginning	NOUN
alces-343	23	40	of	of	ADP
alces-343	23	41	the	the	DET
alces-343	23	42	20th	20th	ADJ
alces-343	23	43	century	century	NOUN
alces-343	23	44	(	(	PUNCT
alces-343	23	45	ouellet	ouellet	NOUN
alces-343	23	46	et	et	PROPN
alces-343	23	47	al	al	PROPN
alces-343	23	48	.	.	PROPN
alces-343	23	49	1996	1996	NUM
alces-343	23	50	)	)	PUNCT
alces-343	23	51	.	.	PUNCT
alces-343	24	1	massive	massive	ADJ
alces-343	24	2	mortalities	mortality	NOUN
alces-343	24	3	from	from	ADP
alces-343	24	4	starvation	starvation	NOUN
alces-343	24	5	sometimes	sometimes	ADV
alces-343	24	6	take	take	VERB
alces-343	24	7	place	place	NOUN
alces-343	24	8	overwinter	overwinter	NOUN
alces-343	24	9	due	due	ADP
alces-343	24	10	to	to	ADP
alces-343	24	11	food	food	NOUN
alces-343	24	12	overexploitation	overexploitation	NOUN
alces-343	24	13	or	or	CCONJ
alces-343	24	14	density	density	NOUN
alces-343	24	15	-	-	PUNCT
alces-343	24	16	independent	independent	ADJ
alces-343	24	17	factors	factor	NOUN
alces-343	24	18	,	,	PUNCT
alces-343	24	19	such	such	ADJ
alces-343	24	20	as	as	ADP
alces-343	24	21	climatic	climatic	ADJ
alces-343	24	22	conditions	condition	NOUN
alces-343	24	23	preventing	prevent	VERB
alces-343	24	24	access	access	NOUN
alces-343	24	25	to	to	ADP
alces-343	24	26	food	food	NOUN
alces-343	24	27	(	(	PUNCT
alces-343	24	28	e.g.	e.g.	ADV
alces-343	24	29	,	,	PUNCT
alces-343	24	30	ice	ice	NOUN
alces-343	24	31	crust	crust	NOUN
alces-343	24	32	;	;	PUNCT
alces-343	24	33	klein	klein	PROPN
alces-343	24	34	1968	1968	NUM
alces-343	24	35	,	,	PUNCT
alces-343	24	36	reimers	reimer	NOUN
alces-343	24	37	1982	1982	NUM
alces-343	24	38	)	)	PUNCT
alces-343	24	39	.	.	PUNCT
alces-343	25	1	in	in	ADP
alces-343	25	2	continental	continental	ADJ
alces-343	25	3	tundra	tundra	PROPN
alces-343	25	4	,	,	PUNCT
alces-343	25	5	caribou	caribou	NOUN
alces-343	25	6	populations	population	NOUN
alces-343	25	7	also	also	ADV
alces-343	25	8	seem	seem	VERB
alces-343	25	9	to	to	PART
alces-343	25	10	be	be	AUX
alces-343	25	11	regulated	regulate	VERB
alces-343	25	12	by	by	ADP
alces-343	25	13	food	food	NOUN
alces-343	25	14	.	.	PUNCT
alces-343	26	1	caribou	caribou	PROPN
alces-343	26	2	may	may	AUX
alces-343	26	3	undertake	undertake	VERB
alces-343	26	4	very	very	ADV
alces-343	26	5	long	long	ADJ
alces-343	26	6	migrations	migration	NOUN
alces-343	26	7	that	that	PRON
alces-343	26	8	allow	allow	VERB
alces-343	26	9	escape	escape	NOUN
alces-343	26	10	from	from	ADP
alces-343	26	11	predation	predation	NOUN
alces-343	26	12	most	most	ADJ
alces-343	26	13	of	of	ADP
alces-343	26	14	the	the	DET
alces-343	26	15	year	year	NOUN
alces-343	26	16	,	,	PUNCT
alces-343	26	17	partly	partly	ADV
alces-343	26	18	because	because	SCONJ
alces-343	26	19	wolf	wolf	PROPN
alces-343	26	20	(	(	PUNCT
alces-343	26	21	canis	canis	NOUN
alces-343	26	22	lupus	lupus	NOUN
alces-343	26	23	)	)	PUNCT
alces-343	26	24	packs	pack	NOUN
alces-343	26	25	are	be	AUX
alces-343	26	26	because	because	SCONJ
alces-343	26	27	wolf	wolf	NOUN
alces-343	26	28	packs	pack	NOUN
alces-343	26	29	can	can	AUX
alces-343	26	30	not	not	PART
alces-343	26	31	move	move	VERB
alces-343	26	32	away	away	ADV
alces-343	26	33	from	from	ADP
alces-343	26	34	a	a	DET
alces-343	26	35	den	den	NOUN
alces-343	26	36	site	site	NOUN
alces-343	26	37	when	when	SCONJ
alces-343	26	38	raising	raise	VERB
alces-343	26	39	pups	pup	NOUN
alces-343	26	40	during	during	ADP
alces-343	26	41	summer	summer	NOUN
alces-343	26	42	(	(	PUNCT
alces-343	26	43	bergerud	bergerud	NOUN
alces-343	26	44	1996	1996	NUM
alces-343	26	45	)	)	PUNCT
alces-343	26	46	.	.	PUNCT
alces-343	27	1	in	in	ADP
alces-343	27	2	tundra	tundra	NOUN
alces-343	27	3	environments	environment	NOUN
alces-343	27	4	,	,	PUNCT
alces-343	27	5	prepredation	prepredation	NOUN
alces-343	27	6	rate	rate	NOUN
alces-343	27	7	does	do	AUX
alces-343	27	8	not	not	PART
alces-343	27	9	increase	increase	VERB
alces-343	27	10	as	as	ADP
alces-343	27	11	a	a	DET
alces-343	27	12	function	function	NOUN
alces-343	27	13	of	of	ADP
alces-343	27	14	caribou	caribou	NOUN
alces-343	27	15	abundance	abundance	PROPN
alces-343	27	16	.	.	PUNCT
alces-343	28	1	caribou	caribou	NOUN
alces-343	28	2	numbers	number	NOUN
alces-343	28	3	can	can	AUX
alces-343	28	4	therefore	therefore	ADV
alces-343	28	5	increase	increase	VERB
alces-343	28	6	and	and	CCONJ
alces-343	28	7	,	,	PUNCT
alces-343	28	8	at	at	ADP
alces-343	28	9	high	high	ADJ
alces-343	28	10	density	density	NOUN
alces-343	28	11	,	,	PUNCT
alces-343	28	12	they	they	PRON
alces-343	28	13	can	can	AUX
alces-343	28	14	overexploit	overexploit	VERB
alces-343	28	15	food	food	NOUN
alces-343	28	16	available	available	ADJ
alces-343	28	17	during	during	ADP
alces-343	28	18	summer	summer	NOUN
alces-343	28	19	(	(	PUNCT
alces-343	28	20	bergerud	bergerud	NOUN
alces-343	28	21	1996	1996	NUM
alces-343	28	22	,	,	PUNCT
alces-343	28	23	crête	crête	ADJ
alces-343	28	24	and	and	CCONJ
alces-343	28	25	doucet	doucet	PROPN
alces-343	28	26	1998	1998	NUM
alces-343	28	27	)	)	PUNCT
alces-343	28	28	or	or	CCONJ
alces-343	28	29	winter	winter	NOUN
alces-343	28	30	(	(	PUNCT
alces-343	28	31	ouellet	ouellet	NOUN
alces-343	28	32	et	et	PROPN
alces-343	28	33	al	al	PROPN
alces-343	28	34	.	.	PROPN
alces-343	28	35	1994	1994	NUM
alces-343	28	36	,	,	PUNCT
alces-343	28	37	1996	1996	NUM
alces-343	28	38	,	,	PUNCT
alces-343	28	39	1997	1997	NUM
alces-343	28	40	)	)	PUNCT
alces-343	28	41	.	.	PUNCT
alces-343	29	1	as	as	SCONJ
alces-343	29	2	resources	resource	NOUN
alces-343	29	3	available	available	ADJ
alces-343	29	4	per	per	ADP
alces-343	29	5	animal	animal	NOUN
alces-343	29	6	diminish	diminish	NOUN
alces-343	29	7	with	with	ADP
alces-343	29	8	increasing	increase	VERB
alces-343	29	9	population	population	NOUN
alces-343	29	10	,	,	PUNCT
alces-343	29	11	a	a	DET
alces-343	29	12	decrease	decrease	NOUN
alces-343	29	13	in	in	ADP
alces-343	29	14	birth	birth	NOUN
alces-343	29	15	rate	rate	NOUN
alces-343	29	16	and	and	CCONJ
alces-343	29	17	an	an	DET
alces-343	29	18	increase	increase	NOUN
alces-343	29	19	in	in	ADP
alces-343	29	20	adult	adult	NOUN
alces-343	29	21	and	and	CCONJ
alces-343	29	22	calf	calf	NOUN
alces-343	29	23	mortality	mortality	NOUN
alces-343	29	24	are	be	AUX
alces-343	29	25	noted	note	VERB
alces-343	29	26	.	.	PUNCT
alces-343	30	1	densities	density	NOUN
alces-343	30	2	averaging	average	VERB
alces-343	30	3	60	60	NUM
alces-343	30	4	–	–	PUNCT
alces-343	30	5	110	110	NUM
alces-343	30	6	caribou	caribou	NOUN
alces-343	30	7	per	per	ADP
alces-343	30	8	100	100	NUM
alces-343	30	9	km2	km2	NOUN
alces-343	30	10	have	have	AUX
alces-343	30	11	been	be	AUX
alces-343	30	12	observed	observe	VERB
alces-343	30	13	in	in	ADP
alces-343	30	14	this	this	DET
alces-343	30	15	environment	environment	NOUN
alces-343	30	16	(	(	PUNCT
alces-343	30	17	messier	messier	X
alces-343	30	18	et	et	PROPN
alces-343	30	19	al	al	PROPN
alces-343	30	20	.	.	PROPN
alces-343	30	21	1988	1988	NUM
alces-343	30	22	,	,	PUNCT
alces-343	30	23	seip	seip	NOUN
alces-343	30	24	1991	1991	NUM
alces-343	30	25	)	)	PUNCT
alces-343	30	26	.	.	PUNCT
alces-343	31	1	using	use	VERB
alces-343	31	2	the	the	DET
alces-343	31	3	annual	annual	ADJ
alces-343	31	4	increase	increase	NOUN
alces-343	31	5	in	in	ADP
alces-343	31	6	lichen	lichen	NOUN
alces-343	31	7	biomass	biomass	NOUN
alces-343	31	8	,	,	PUNCT
alces-343	31	9	which	which	PRON
alces-343	31	10	is	be	AUX
alces-343	31	11	the	the	DET
alces-343	31	12	primary	primary	ADJ
alces-343	31	13	food	food	NOUN
alces-343	31	14	source	source	NOUN
alces-343	31	15	for	for	ADP
alces-343	31	16	caribou	caribou	PROPN
alces-343	31	17	(	(	PUNCT
alces-343	31	18	gauthier	gauthier	PROPN
alces-343	31	19	et	et	PROPN
alces-343	31	20	al	al	PROPN
alces-343	31	21	.	.	PROPN
alces-343	31	22	1989	1989	NUM
alces-343	31	23	)	)	PUNCT
alces-343	31	24	,	,	PUNCT
alces-343	31	25	along	along	ADP
alces-343	31	26	with	with	ADP
alces-343	31	27	losses	loss	NOUN
alces-343	31	28	caused	cause	VERB
alces-343	31	29	by	by	ADP
alces-343	31	30	animal	animal	NOUN
alces-343	31	31	trampling	trampling	NOUN
alces-343	31	32	,	,	PUNCT
alces-343	31	33	arsenault	arsenault	PROPN
alces-343	31	34	et	et	PROPN
alces-343	31	35	al	al	PROPN
alces-343	31	36	.	.	PROPN
alces-343	32	1	(	(	PUNCT
alces-343	32	2	1997	1997	NUM
alces-343	32	3	)	)	PUNCT
alces-343	32	4	have	have	AUX
alces-343	32	5	estimated	estimate	VERB
alces-343	32	6	the	the	DET
alces-343	32	7	carrying	carry	VERB
alces-343	32	8	capacity	capacity	NOUN
alces-343	32	9	of	of	ADP
alces-343	32	10	caribou	caribou	NOUN
alces-343	32	11	in	in	ADP
alces-343	32	12	tundra	tundra	NOUN
alces-343	32	13	at	at	ADP
alces-343	32	14	approximately	approximately	ADV
alces-343	32	15	20	20	NUM
alces-343	32	16	animals	animal	NOUN
alces-343	32	17	per	per	ADP
alces-343	32	18	100	100	NUM
alces-343	32	19	km2	km2	NOUN
alces-343	32	20	.	.	PUNCT
alces-343	33	1	in	in	ADP
alces-343	33	2	the	the	DET
alces-343	33	3	boreal	boreal	NOUN
alces-343	33	4	forest	forest	NOUN
alces-343	33	5	,	,	PUNCT
alces-343	33	6	the	the	DET
alces-343	33	7	carrying	carrying	NOUN
alces-343	33	8	capacity	capacity	NOUN
alces-343	33	9	for	for	ADP
alces-343	33	10	caribou	caribou	NOUN
alces-343	33	11	is	be	AUX
alces-343	33	12	not	not	PART
alces-343	33	13	precisely	precisely	ADV
alces-343	33	14	known	know	VERB
alces-343	33	15	.	.	PUNCT
alces-343	34	1	based	base	VERB
alces-343	34	2	on	on	ADP
alces-343	34	3	lichen	lichen	NOUN
alces-343	34	4	biomass	biomass	NOUN
alces-343	34	5	,	,	PUNCT
alces-343	34	6	crête	crête	ADJ
alces-343	34	7	and	and	CCONJ
alces-343	34	8	manseau	manseau	NOUN
alces-343	34	9	(	(	PUNCT
alces-343	34	10	1996	1996	NUM
alces-343	34	11	)	)	PUNCT
alces-343	34	12	estimated	estimate	VERB
alces-343	34	13	that	that	SCONJ
alces-343	34	14	carrying	carry	VERB
alces-343	34	15	capacity	capacity	NOUN
alces-343	34	16	should	should	AUX
alces-343	34	17	be	be	AUX
alces-343	34	18	at	at	ADP
alces-343	34	19	least	least	ADJ
alces-343	34	20	that	that	PRON
alces-343	34	21	observed	observe	VERB
alces-343	34	22	in	in	ADP
alces-343	34	23	tundra	tundra	NOUN
alces-343	34	24	,	,	PUNCT
alces-343	34	25	because	because	SCONJ
alces-343	34	26	alternative	alternative	ADJ
alces-343	34	27	food	food	NOUN
alces-343	34	28	sources	source	NOUN
alces-343	34	29	(	(	PUNCT
alces-343	34	30	such	such	ADJ
alces-343	34	31	as	as	ADP
alces-343	34	32	leaves	leave	NOUN
alces-343	34	33	,	,	PUNCT
alces-343	34	34	twigs	twig	NOUN
alces-343	34	35	,	,	PUNCT
alces-343	34	36	and	and	CCONJ
alces-343	34	37	deciduous	deciduous	ADJ
alces-343	34	38	shrubs	shrub	NOUN
alces-343	34	39	)	)	PUNCT
alces-343	34	40	are	be	AUX
alces-343	34	41	abundant	abundant	ADJ
alces-343	34	42	and	and	CCONJ
alces-343	34	43	the	the	DET
alces-343	34	44	climate	climate	NOUN
alces-343	34	45	is	be	AUX
alces-343	34	46	milder	mild	ADJ
alces-343	34	47	in	in	ADP
alces-343	34	48	that	that	DET
alces-343	34	49	environment	environment	NOUN
alces-343	34	50	.	.	PUNCT
alces-343	35	1	for	for	ADP
alces-343	35	2	example	example	NOUN
alces-343	35	3	,	,	PUNCT
alces-343	35	4	in	in	ADP
alces-343	35	5	east	east	NOUN
alces-343	35	6	-	-	PUNCT
alces-343	35	7	central	central	ADJ
alces-343	35	8	québec	québec	NOUN
alces-343	35	9	,	,	PUNCT
alces-343	35	10	carrying	carry	VERB
alces-343	35	11	capacity	capacity	NOUN
alces-343	35	12	based	base	VERB
alces-343	35	13	solely	solely	ADV
alces-343	35	14	on	on	ADP
alces-343	35	15	terrestrial	terrestrial	ADJ
alces-343	35	16	lichen	lichen	NOUN
alces-343	35	17	has	have	AUX
alces-343	35	18	recently	recently	ADV
alces-343	35	19	been	be	AUX
alces-343	35	20	estimated	estimate	VERB
alces-343	35	21	at	at	ADP
alces-343	35	22	4.1	4.1	NUM
alces-343	35	23	–	–	PUNCT
alces-343	35	24	7.7	7.7	NUM
alces-343	35	25	caribou	caribou	NOUN
alces-343	35	26	per	per	ADP
alces-343	35	27	100	100	NUM
alces-343	35	28	km2	km2	NOUN
alces-343	35	29	(	(	PUNCT
alces-343	35	30	courtois	courtois	PROPN
alces-343	35	31	2003	2003	NUM
alces-343	35	32	)	)	PUNCT
alces-343	35	33	.	.	PUNCT
alces-343	36	1	despite	despite	SCONJ
alces-343	36	2	a	a	DET
alces-343	36	3	relatively	relatively	ADV
alces-343	36	4	high	high	ADJ
alces-343	36	5	potential	potential	ADJ
alces-343	36	6	carrying	carrying	NOUN
alces-343	36	7	capacity	capacity	NOUN
alces-343	36	8	,	,	PUNCT
alces-343	36	9	woodland	woodland	NOUN
alces-343	36	10	caribou	caribou	NOUN
alces-343	36	11	populations	population	NOUN
alces-343	36	12	experience	experience	VERB
alces-343	36	13	very	very	ADV
alces-343	36	14	low	low	ADJ
alces-343	36	15	densities	density	NOUN
alces-343	36	16	of	of	ADP
alces-343	36	17	between	between	ADP
alces-343	36	18	1	1	NUM
alces-343	36	19	and	and	CCONJ
alces-343	36	20	3	3	NUM
alces-343	36	21	individuals	individual	NOUN
alces-343	36	22	per	per	ADP
alces-343	36	23	100	100	NUM
alces-343	36	24	km2	km2	NOUN
alces-343	36	25	(	(	PUNCT
alces-343	36	26	seip	seip	NOUN
alces-343	36	27	1991	1991	NUM
alces-343	36	28	,	,	PUNCT
alces-343	36	29	courtois	courtois	PROPN
alces-343	36	30	2003	2003	NUM
alces-343	36	31	)	)	PUNCT
alces-343	36	32	and	and	CCONJ
alces-343	36	33	most	most	ADJ
alces-343	36	34	are	be	AUX
alces-343	36	35	declining	decline	VERB
alces-343	36	36	in	in	ADP
alces-343	36	37	north	north	PROPN
alces-343	36	38	america	america	PROPN
alces-343	36	39	(	(	PUNCT
alces-343	36	40	mallory	mallory	PROPN
alces-343	36	41	and	and	CCONJ
alces-343	36	42	hillis	hillis	PROPN
alces-343	36	43	1998	1998	NUM
alces-343	36	44	)	)	PUNCT
alces-343	36	45	.	.	PUNCT
alces-343	37	1	those	those	DET
alces-343	37	2	observations	observation	NOUN
alces-343	37	3	suggest	suggest	VERB
alces-343	37	4	that	that	SCONJ
alces-343	37	5	woodland	woodland	PROPN
alces-343	37	6	caribou	caribou	NOUN
alces-343	37	7	populations	population	NOUN
alces-343	37	8	for	for	ADP
alces-343	37	9	winter	winter	NOUN
alces-343	37	10	food	food	NOUN
alces-343	37	11	.	.	PUNCT
alces-343	38	1	moreover	moreover	ADV
alces-343	38	2	,	,	PUNCT
alces-343	38	3	as	as	SCONJ
alces-343	38	4	caribou	caribou	PROPN
alces-343	38	5	food	food	NOUN
alces-343	38	6	habits	habit	NOUN
alces-343	38	7	are	be	AUX
alces-343	38	8	much	much	ADV
alces-343	38	9	less	less	ADV
alces-343	38	10	restrictive	restrictive	ADJ
alces-343	38	11	in	in	ADP
alces-343	38	12	summer	summer	NOUN
alces-343	38	13	,	,	PUNCT
alces-343	38	14	the	the	DET
alces-343	38	15	main	main	ADJ
alces-343	38	16	limiting	limit	VERB
alces-343	38	17	factors	factor	NOUN
alces-343	38	18	seem	seem	VERB
alces-343	38	19	to	to	PART
alces-343	38	20	be	be	AUX
alces-343	38	21	hunting	hunt	VERB
alces-343	38	22	and	and	CCONJ
alces-343	38	23	predation	predation	NOUN
alces-343	38	24	(	(	PUNCT
alces-343	38	25	stuart	stuart	PROPN
alces-343	38	26	-	-	PUNCT
alces-343	38	27	smith	smith	PROPN
alces-343	38	28	et	et	PROPN
alces-343	38	29	al	al	PROPN
alces-343	38	30	.	.	PROPN
alces-343	38	31	1997	1997	NUM
alces-343	38	32	,	,	PUNCT
alces-343	38	33	rettie	rettie	NOUN
alces-343	38	34	and	and	CCONJ
alces-343	38	35	messier	messy	ADJ
alces-343	38	36	1998	1998	NUM
alces-343	38	37	)	)	PUNCT
alces-343	38	38	.	.	PUNCT
alces-343	39	1	caribou	caribou	NOUN
alces-343	39	2	densities	density	NOUN
alces-343	39	3	in	in	ADP
alces-343	39	4	the	the	DET
alces-343	39	5	boreal	boreal	NOUN
alces-343	39	6	forest	forest	NOUN
alces-343	39	7	(	(	PUNCT
alces-343	39	8	1	1	NUM
alces-343	39	9	–	–	SYM
alces-343	39	10	3	3	NUM
alces-343	39	11	per	per	ADP
alces-343	39	12	100	100	NUM
alces-343	39	13	km2	km2	NUM
alces-343	39	14	;	;	PUNCT
alces-343	39	15	seip	seip	NOUN
alces-343	39	16	1991	1991	NUM
alces-343	39	17	,	,	PUNCT
alces-343	39	18	courtois	courtois	PROPN
alces-343	39	19	et	et	PROPN
alces-343	39	20	al	al	PROPN
alces-343	39	21	.	.	PROPN
alces-343	39	22	2003	2003	NUM
alces-343	39	23	)	)	PUNCT
alces-343	39	24	are	be	AUX
alces-343	39	25	typically	typically	ADV
alces-343	39	26	incapable	incapable	ADJ
alces-343	39	27	of	of	ADP
alces-343	39	28	supporting	support	VERB
alces-343	39	29	wolf	wolf	NOUN
alces-343	39	30	populations	population	NOUN
alces-343	39	31	.	.	PUNCT
alces-343	40	1	wolves	wolf	NOUN
alces-343	40	2	in	in	ADP
alces-343	40	3	the	the	DET
alces-343	40	4	boreal	boreal	NOUN
alces-343	40	5	forest	forest	NOUN
alces-343	40	6	therefore	therefore	ADV
alces-343	40	7	depend	depend	VERB
alces-343	40	8	on	on	ADP
alces-343	40	9	moose	moose	NOUN
alces-343	40	10	(	(	PUNCT
alces-343	40	11	alces	alce	NOUN
alces-343	40	12	alces	alce	NOUN
alces-343	40	13	)	)	PUNCT
alces-343	40	14	,	,	PUNCT
alces-343	40	15	a	a	DET
alces-343	40	16	larger	large	ADJ
alces-343	40	17	and	and	CCONJ
alces-343	40	18	more	more	ADV
alces-343	40	19	abundant	abundant	ADJ
alces-343	40	20	ungulate	ungulate	NOUN
alces-343	40	21	that	that	PRON
alces-343	40	22	generally	generally	ADV
alces-343	40	23	lives	live	VERB
alces-343	40	24	at	at	ADP
alces-343	40	25	densities	density	NOUN
alces-343	40	26	of	of	ADP
alces-343	40	27	between	between	ADP
alces-343	40	28	10	10	NUM
alces-343	40	29	and	and	CCONJ
alces-343	40	30	20	20	NUM
alces-343	40	31	individuals	individual	NOUN
alces-343	40	32	per	per	ADP
alces-343	40	33	100	100	NUM
alces-343	40	34	km2	km2	NOUN
alces-343	40	35	(	(	PUNCT
alces-343	40	36	messier	messy	ADJ
alces-343	40	37	1985	1985	NUM
alces-343	40	38	)	)	PUNCT
alces-343	40	39	.	.	PUNCT
alces-343	41	1	due	due	ADP
alces-343	41	2	to	to	ADP
alces-343	41	3	the	the	DET
alces-343	41	4	absence	absence	NOUN
alces-343	41	5	of	of	ADP
alces-343	41	6	a	a	DET
alces-343	41	7	regulating	regulate	VERB
alces-343	41	8	mechanism	mechanism	NOUN
alces-343	41	9	between	between	ADP
alces-343	41	10	caribou	caribou	NOUN
alces-343	41	11	and	and	CCONJ
alces-343	41	12	wolves	wolf	NOUN
alces-343	41	13	in	in	ADP
alces-343	41	14	the	the	DET
alces-343	41	15	boreal	boreal	NOUN
alces-343	41	16	forest	forest	NOUN
alces-343	41	17	(	(	PUNCT
alces-343	41	18	seip	seip	NOUN
alces-343	41	19	1991	1991	NUM
alces-343	41	20	)	)	PUNCT
alces-343	41	21	,	,	PUNCT
alces-343	41	22	an	an	DET
alces-343	41	23	increase	increase	NOUN
alces-343	41	24	in	in	ADP
alces-343	41	25	moose	moose	NOUN
alces-343	41	26	abundance	abundance	NOUN
alces-343	41	27	should	should	AUX
alces-343	41	28	provoke	provoke	VERB
alces-343	41	29	a	a	DET
alces-343	41	30	wolf	wolf	ADJ
alces-343	41	31	population	population	NOUN
alces-343	41	32	increase	increase	NOUN
alces-343	41	33	,	,	PUNCT
alces-343	41	34	independent	independent	ADJ
alces-343	41	35	of	of	ADP
alces-343	41	36	caribou	caribou	PROPN
alces-343	41	37	abundance	abundance	NOUN
alces-343	41	38	.	.	PUNCT
alces-343	42	1	in	in	ADP
alces-343	42	2	such	such	DET
alces-343	42	3	a	a	DET
alces-343	42	4	situation	situation	NOUN
alces-343	42	5	,	,	PUNCT
alces-343	42	6	more	more	ADV
alces-343	42	7	frequent	frequent	ADJ
alces-343	42	8	encounters	encounter	NOUN
alces-343	42	9	between	between	ADP
alces-343	42	10	wolf	wolf	PROPN
alces-343	42	11	and	and	CCONJ
alces-343	42	12	caribou	caribou	NOUN
alces-343	42	13	should	should	AUX
alces-343	42	14	be	be	AUX
alces-343	42	15	expected	expect	VERB
alces-343	42	16	leading	lead	VERB
alces-343	42	17	to	to	ADP
alces-343	42	18	an	an	DET
alces-343	42	19	increased	increase	VERB
alces-343	42	20	predation	predation	NOUN
alces-343	42	21	rate	rate	NOUN
alces-343	42	22	on	on	ADP
alces-343	42	23	caribou	caribou	NOUN
alces-343	42	24	that	that	PRON
alces-343	42	25	could	could	AUX
alces-343	42	26	decline	decline	VERB
alces-343	42	27	,	,	PUNCT
alces-343	42	28	eventually	eventually	ADV
alces-343	42	29	down	down	ADP
alces-343	42	30	to	to	ADP
alces-343	42	31	extirpation	extirpation	NOUN
alces-343	42	32	(	(	PUNCT
alces-343	42	33	seip	seip	NOUN
alces-343	42	34	1991	1991	NUM
alces-343	42	35	)	)	PUNCT
alces-343	42	36	.	.	PUNCT
alces-343	43	1	caribou	caribou	PROPN
alces-343	43	2	abundance	abundance	NOUN
alces-343	43	3	decreases	decrease	VERB
alces-343	43	4	when	when	SCONJ
alces-343	43	5	there	there	PRON
alces-343	43	6	are	be	VERB
alces-343	43	7	more	more	ADJ
alces-343	43	8	than	than	ADP
alces-343	43	9	0.60	0.60	NUM
alces-343	43	10	–	–	PUNCT
alces-343	43	11	0.65	0.65	NUM
alces-343	43	12	wolves	wolf	NOUN
alces-343	43	13	per	per	ADP
alces-343	43	14	100	100	NUM
alces-343	43	15	km2	km2	NOUN
alces-343	43	16	(	(	PUNCT
alces-343	43	17	bergerud	bergerud	NOUN
alces-343	43	18	and	and	CCONJ
alces-343	43	19	elliot	elliot	PROPN
alces-343	43	20	1986	1986	NUM
alces-343	43	21	,	,	PUNCT
alces-343	43	22	bergalces	bergalce	NOUN
alces-343	43	23	vol	vol	NOUN
alces-343	43	24	.	.	PROPN
alces-343	44	1	43	43	NUM
alces-343	44	2	,	,	PUNCT
alces-343	44	3	2007	2007	NUM
alces-343	44	4	courtois	courtois	NOUN
alces-343	44	5	and	and	CCONJ
alces-343	44	6	ouellet	ouellet	NOUN
alces-343	44	7	impact	impact	NOUN
alces-343	44	8	of	of	ADP
alces-343	44	9	moose	moose	NOUN
alces-343	44	10	and	and	CCONJ
alces-343	44	11	wolves	wolf	NOUN
alces-343	44	12	on	on	ADP
alces-343	44	13	caribou	caribou	NOUN
alces-343	44	14	15	15	NUM
alces-343	44	15	erud	erud	NOUN
alces-343	44	16	1996	1996	NUM
alces-343	44	17	)	)	PUNCT
alces-343	44	18	and	and	CCONJ
alces-343	44	19	caribou	caribou	NOUN
alces-343	44	20	populations	population	NOUN
alces-343	44	21	increase	increase	VERB
alces-343	44	22	when	when	SCONJ
alces-343	44	23	wolf	wolf	PROPN
alces-343	44	24	numbers	number	NOUN
alces-343	44	25	are	be	AUX
alces-343	44	26	controlled	control	VERB
alces-343	44	27	(	(	PUNCT
alces-343	44	28	boertje	boertje	X
alces-343	44	29	et	et	PROPN
alces-343	44	30	al	al	PROPN
alces-343	44	31	.	.	PROPN
alces-343	44	32	1996	1996	NUM
alces-343	44	33	,	,	PUNCT
alces-343	44	34	hayes	hayes	PROPN
alces-343	44	35	et	et	PROPN
alces-343	44	36	al	al	PROPN
alces-343	44	37	.	.	PROPN
alces-343	44	38	2003	2003	NUM
alces-343	44	39	)	)	PUNCT
alces-343	44	40	.	.	PUNCT
alces-343	45	1	the	the	DET
alces-343	45	2	necessity	necessity	NOUN
alces-343	45	3	for	for	SCONJ
alces-343	45	4	caribou	caribou	NOUN
alces-343	45	5	to	to	PART
alces-343	45	6	adopt	adopt	VERB
alces-343	45	7	avoidance	avoidance	NOUN
alces-343	45	8	strategies	strategy	NOUN
alces-343	45	9	of	of	ADP
alces-343	45	10	both	both	DET
alces-343	45	11	predators	predator	NOUN
alces-343	45	12	and	and	CCONJ
alces-343	45	13	other	other	ADJ
alces-343	45	14	ungulates	ungulate	NOUN
alces-343	45	15	in	in	ADP
alces-343	45	16	order	order	NOUN
alces-343	45	17	to	to	PART
alces-343	45	18	survive	survive	VERB
alces-343	45	19	is	be	AUX
alces-343	45	20	now	now	ADV
alces-343	45	21	recoget	recoget	VERB
alces-343	45	22	al	al	PROPN
alces-343	45	23	.	.	PROPN
alces-343	45	24	1984	1984	NUM
alces-343	45	25	;	;	PUNCT
alces-343	45	26	bergerud	bergerud	NOUN
alces-343	45	27	1985	1985	NUM
alces-343	45	28	,	,	PUNCT
alces-343	45	29	1996	1996	NUM
alces-343	45	30	;	;	PUNCT
alces-343	45	31	seip	seip	NOUN
alces-343	45	32	1991	1991	NUM
alces-343	45	33	,	,	PUNCT
alces-343	45	34	1992	1992	NUM
alces-343	45	35	;	;	PUNCT
alces-343	45	36	cumming	cumme	VERB
alces-343	45	37	et	et	PROPN
alces-343	45	38	al	al	PROPN
alces-343	45	39	.	.	PROPN
alces-343	45	40	1996	1996	NUM
alces-343	45	41	;	;	PUNCT
alces-343	45	42	stuart	stuart	PROPN
alces-343	45	43	-	-	PUNCT
alces-343	45	44	smith	smith	PROPN
alces-343	45	45	et	et	PROPN
alces-343	45	46	al	al	PROPN
alces-343	45	47	.	.	PROPN
alces-343	45	48	1997	1997	NUM
alces-343	45	49	;	;	PUNCT
alces-343	45	50	rettie	rettie	NOUN
alces-343	45	51	and	and	CCONJ
alces-343	45	52	messier	messy	ADJ
alces-343	45	53	1998	1998	NUM
alces-343	45	54	)	)	PUNCT
alces-343	45	55	.	.	PUNCT
alces-343	46	1	however	however	ADV
alces-343	46	2	,	,	PUNCT
alces-343	46	3	the	the	DET
alces-343	46	4	mechanisms	mechanism	NOUN
alces-343	46	5	underlying	underlie	VERB
alces-343	46	6	caribou	caribou	NOUN
alces-343	46	7	,	,	PUNCT
alces-343	46	8	moose	moose	NOUN
alces-343	46	9	,	,	PUNCT
alces-343	46	10	and	and	CCONJ
alces-343	46	11	wolf	wolf	NOUN
alces-343	46	12	interactions	interaction	NOUN
alces-343	46	13	still	still	ADV
alces-343	46	14	remain	remain	VERB
alces-343	46	15	obscure	obscure	ADJ
alces-343	46	16	.	.	PUNCT
alces-343	47	1	in	in	ADP
alces-343	47	2	this	this	DET
alces-343	47	3	study	study	NOUN
alces-343	47	4	,	,	PUNCT
alces-343	47	5	we	we	PRON
alces-343	47	6	built	build	VERB
alces-343	47	7	a	a	DET
alces-343	47	8	deterministic	deterministic	ADJ
alces-343	47	9	model	model	NOUN
alces-343	47	10	that	that	PRON
alces-343	47	11	mimicked	mimic	VERB
alces-343	47	12	the	the	DET
alces-343	47	13	relationships	relationship	NOUN
alces-343	47	14	between	between	ADP
alces-343	47	15	moose	moose	NOUN
alces-343	47	16	,	,	PUNCT
alces-343	47	17	wolves	wolf	NOUN
alces-343	47	18	,	,	PUNCT
alces-343	47	19	and	and	CCONJ
alces-343	47	20	caribou	caribou	NOUN
alces-343	47	21	and	and	CCONJ
alces-343	47	22	then	then	ADV
alces-343	47	23	simulated	simulate	VERB
alces-343	47	24	how	how	SCONJ
alces-343	47	25	variation	variation	NOUN
alces-343	47	26	in	in	ADP
alces-343	47	27	hunting	hunting	NOUN
alces-343	47	28	of	of	ADP
alces-343	47	29	caribou	caribou	PROPN
alces-343	47	30	and	and	CCONJ
alces-343	47	31	caribou	caribou	PROPN
alces-343	47	32	abundance	abundance	PROPN
alces-343	47	33	.	.	PUNCT
alces-343	48	1	model	model	NOUN
alces-343	48	2	results	result	NOUN
alces-343	48	3	should	should	AUX
alces-343	48	4	help	help	VERB
alces-343	48	5	to	to	PART
alces-343	48	6	determine	determine	VERB
alces-343	48	7	which	which	DET
alces-343	48	8	management	management	NOUN
alces-343	48	9	strategies	strategy	NOUN
alces-343	48	10	are	be	AUX
alces-343	48	11	most	most	ADV
alces-343	48	12	suitable	suitable	ADJ
alces-343	48	13	to	to	PART
alces-343	48	14	maintain	maintain	VERB
alces-343	48	15	equilibrium	equilibrium	NOUN
alces-343	48	16	between	between	ADP
alces-343	48	17	these	these	DET
alces-343	48	18	three	three	NUM
alces-343	48	19	species	specie	NOUN
alces-343	48	20	in	in	ADP
alces-343	48	21	order	order	NOUN
alces-343	48	22	to	to	PART
alces-343	48	23	help	help	VERB
alces-343	48	24	maintain	maintain	VERB
alces-343	48	25	caribou	caribou	NOUN
alces-343	48	26	,	,	PUNCT
alces-343	48	27	a	a	DET
alces-343	48	28	threatened	threaten	VERB
alces-343	48	29	species	specie	NOUN
alces-343	48	30	in	in	ADP
alces-343	48	31	the	the	DET
alces-343	48	32	boreal	boreal	NOUN
alces-343	48	33	forest	forest	NOUN
alces-343	48	34	.	.	PUNCT
alces-343	49	1	methods	method	NOUN
alces-343	49	2	the	the	DET
alces-343	49	3	model	model	NOUN
alces-343	49	4	the	the	DET
alces-343	49	5	model	model	NOUN
alces-343	49	6	simulates	simulate	NOUN
alces-343	49	7	changes	change	NOUN
alces-343	49	8	in	in	ADP
alces-343	49	9	caribou	caribou	PROPN
alces-343	49	10	,	,	PUNCT
alces-343	49	11	moose	moose	NOUN
alces-343	49	12	,	,	PUNCT
alces-343	49	13	and	and	CCONJ
alces-343	49	14	wolf	wolf	NOUN
alces-343	49	15	numbers	number	NOUN
alces-343	49	16	in	in	ADP
alces-343	49	17	a	a	DET
alces-343	49	18	1,000	1,000	NUM
alces-343	49	19	-	-	PUNCT
alces-343	49	20	km2	km2	NOUN
alces-343	49	21	study	study	NOUN
alces-343	49	22	area	area	NOUN
alces-343	49	23	of	of	ADP
alces-343	49	24	the	the	DET
alces-343	49	25	québec	québec	PROPN
alces-343	49	26	boreal	boreal	NOUN
alces-343	49	27	forest	forest	NOUN
alces-343	49	28	where	where	SCONJ
alces-343	49	29	the	the	DET
alces-343	49	30	three	three	NUM
alces-343	49	31	species	specie	NOUN
alces-343	49	32	live	live	VERB
alces-343	49	33	in	in	ADP
alces-343	49	34	sympatry	sympatry	NOUN
alces-343	49	35	.	.	PUNCT
alces-343	50	1	according	accord	VERB
alces-343	50	2	to	to	ADP
alces-343	50	3	the	the	DET
alces-343	50	4	model	model	NOUN
alces-343	50	5	,	,	PUNCT
alces-343	50	6	in	in	ADP
alces-343	50	7	the	the	DET
alces-343	50	8	absence	absence	NOUN
alces-343	50	9	of	of	ADP
alces-343	50	10	wolves	wolf	NOUN
alces-343	50	11	,	,	PUNCT
alces-343	50	12	the	the	DET
alces-343	50	13	moose	moose	NOUN
alces-343	50	14	and	and	CCONJ
alces-343	50	15	caribou	caribou	NOUN
alces-343	50	16	populations	population	NOUN
alces-343	50	17	are	be	AUX
alces-343	50	18	regulated	regulate	VERB
alces-343	50	19	by	by	ADP
alces-343	50	20	food	food	NOUN
alces-343	50	21	competition	competition	NOUN
alces-343	50	22	and	and	CCONJ
alces-343	50	23	follow	follow	VERB
alces-343	50	24	a	a	DET
alces-343	50	25	logistic	logistic	ADJ
alces-343	50	26	equation	equation	NOUN
alces-343	50	27	(	(	PUNCT
alces-343	50	28	nt+1	nt+1	X
alces-343	50	29	=	=	SYM
alces-343	50	30	nt	not	PART
alces-343	50	31	r	r	NOUN
alces-343	50	32	1	1	NUM
alces-343	50	33	nt	nt	PROPN
alces-343	50	34	/	/	SYM
alces-343	50	35	kcc	kcc	PROPN
alces-343	50	36	,	,	PUNCT
alces-343	50	37	where	where	SCONJ
alces-343	50	38	nt	not	PART
alces-343	50	39	and	and	CCONJ
alces-343	50	40	nt+1	nt+1	X
alces-343	50	41	=	=	VERB
alces-343	50	42	numbers	number	NOUN
alces-343	50	43	at	at	ADP
alces-343	50	44	time	time	NOUN
alces-343	50	45	t	t	PROPN
alces-343	50	46	and	and	CCONJ
alces-343	50	47	time	time	NOUN
alces-343	50	48	t+1	t+1	NUM
alces-343	50	49	,	,	PUNCT
alces-343	50	50	respectively	respectively	ADV
alces-343	50	51	,	,	PUNCT
alces-343	50	52	r	r	NOUN
alces-343	50	53	=	=	SYM
alces-343	50	54	maximum	maximum	ADJ
alces-343	50	55	population	population	NOUN
alces-343	50	56	growth	growth	NOUN
alces-343	50	57	rate	rate	NOUN
alces-343	50	58	,	,	PUNCT
alces-343	50	59	and	and	CCONJ
alces-343	50	60	kcc	kcc	PROPN
alces-343	50	61	=	=	SYM
alces-343	50	62	food	food	NOUN
alces-343	50	63	carrying	carry	VERB
alces-343	50	64	capacity	capacity	NOUN
alces-343	50	65	;	;	PUNCT
alces-343	50	66	caughley	caughley	NOUN
alces-343	50	67	1977	1977	NUM
alces-343	50	68	)	)	PUNCT
alces-343	50	69	.	.	PUNCT
alces-343	51	1	in	in	ADP
alces-343	51	2	the	the	DET
alces-343	51	3	presence	presence	NOUN
alces-343	51	4	of	of	ADP
alces-343	51	5	wolves	wolf	NOUN
alces-343	51	6	,	,	PUNCT
alces-343	51	7	the	the	DET
alces-343	51	8	moose	moose	NOUN
alces-343	51	9	population	population	NOUN
alces-343	51	10	is	be	AUX
alces-343	51	11	regulated	regulate	VERB
alces-343	51	12	by	by	ADP
alces-343	51	13	predation	predation	NOUN
alces-343	51	14	in	in	ADP
alces-343	51	15	accordance	accordance	NOUN
alces-343	51	16	with	with	ADP
alces-343	51	17	the	the	DET
alces-343	51	18	predation	predation	NOUN
alces-343	51	19	model	model	NOUN
alces-343	51	20	of	of	ADP
alces-343	51	21	messier	messy	ADJ
alces-343	51	22	(	(	PUNCT
alces-343	51	23	1994	1994	NUM
alces-343	51	24	)	)	PUNCT
alces-343	51	25	.	.	PUNCT
alces-343	52	1	wolf	wolf	PROPN
alces-343	52	2	numbers	number	NOUN
alces-343	52	3	are	be	AUX
alces-343	52	4	determined	determine	VERB
alces-343	52	5	by	by	ADP
alces-343	52	6	moose	moose	NOUN
alces-343	52	7	numbers	number	NOUN
alces-343	52	8	,	,	PUNCT
alces-343	52	9	as	as	SCONJ
alces-343	52	10	predicted	predict	VERB
alces-343	52	11	by	by	ADP
alces-343	52	12	the	the	DET
alces-343	52	13	michaelis	michaeli	NOUN
alces-343	52	14	-	-	PUNCT
alces-343	52	15	menten	menten	ADJ
alces-343	52	16	hyperbolic	hyperbolic	ADJ
alces-343	52	17	equation	equation	NOUN
alces-343	52	18	of	of	ADP
alces-343	52	19	messier	messy	ADJ
alces-343	52	20	(	(	PUNCT
alces-343	52	21	1994	1994	NUM
alces-343	52	22	)	)	PUNCT
alces-343	52	23	.	.	PUNCT
alces-343	53	1	in	in	ADP
alces-343	53	2	the	the	DET
alces-343	53	3	combined	combined	ADJ
alces-343	53	4	model	model	NOUN
alces-343	53	5	,	,	PUNCT
alces-343	53	6	where	where	SCONJ
alces-343	53	7	the	the	DET
alces-343	53	8	three	three	NUM
alces-343	53	9	species	specie	NOUN
alces-343	53	10	interact	interact	VERB
alces-343	53	11	,	,	PUNCT
alces-343	53	12	wolves	wolf	NOUN
alces-343	53	13	have	have	VERB
alces-343	53	14	access	access	NOUN
alces-343	53	15	to	to	ADP
alces-343	53	16	caribou	caribou	NOUN
alces-343	53	17	and	and	CCONJ
alces-343	53	18	carry	carry	VERB
alces-343	53	19	out	out	ADP
alces-343	53	20	density	density	NOUN
alces-343	53	21	-	-	PUNCT
alces-343	53	22	independent	independent	ADJ
alces-343	53	23	survival	survival	NOUN
alces-343	53	24	of	of	ADP
alces-343	53	25	calves	calf	NOUN
alces-343	53	26	and	and	CCONJ
alces-343	53	27	adults	adult	NOUN
alces-343	53	28	,	,	PUNCT
alces-343	53	29	as	as	SCONJ
alces-343	53	30	predicted	predict	VERB
alces-343	53	31	by	by	ADP
alces-343	53	32	the	the	DET
alces-343	53	33	bergerud	bergerud	NOUN
alces-343	53	34	and	and	CCONJ
alces-343	53	35	elliot	elliot	PROPN
alces-343	53	36	(	(	PUNCT
alces-343	53	37	1986	1986	NUM
alces-343	53	38	)	)	PUNCT
alces-343	53	39	model	model	NOUN
alces-343	53	40	.	.	PUNCT
alces-343	54	1	we	we	PRON
alces-343	54	2	completed	complete	VERB
alces-343	54	3	the	the	DET
alces-343	54	4	model	model	NOUN
alces-343	54	5	by	by	ADP
alces-343	54	6	adding	add	VERB
alces-343	54	7	management	management	NOUN
alces-343	54	8	parameters	parameter	NOUN
alces-343	54	9	,	,	PUNCT
alces-343	54	10	which	which	PRON
alces-343	54	11	allowed	allow	VERB
alces-343	54	12	changing	change	VERB
alces-343	54	13	the	the	DET
alces-343	54	14	moose	moose	NOUN
alces-343	54	15	and	and	CCONJ
alces-343	54	16	caribou	caribou	NOUN
alces-343	54	17	hunting	hunting	NOUN
alces-343	54	18	and	and	CCONJ
alces-343	54	19	wolf	wolf	NOUN
alces-343	54	20	trapping	trap	VERB
alces-343	54	21	mortalities	mortality	NOUN
alces-343	54	22	that	that	PRON
alces-343	54	23	were	be	AUX
alces-343	54	24	additive	additive	ADJ
alces-343	54	25	to	to	ADP
alces-343	54	26	natural	natural	ADJ
alces-343	54	27	mortality	mortality	NOUN
alces-343	54	28	.	.	PUNCT
alces-343	55	1	additional	additional	ADJ
alces-343	55	2	parameters	parameter	NOUN
alces-343	55	3	allowed	allow	VERB
alces-343	55	4	stochastic	stochastic	ADJ
alces-343	55	5	variation	variation	NOUN
alces-343	55	6	of	of	ADP
alces-343	55	7	moose	moose	NOUN
alces-343	55	8	and	and	CCONJ
alces-343	55	9	caribou	caribou	NOUN
alces-343	55	10	productivity	productivity	NOUN
alces-343	55	11	due	due	ADP
alces-343	55	12	to	to	ADP
alces-343	55	13	uncontrolled	uncontrolled	ADJ
alces-343	55	14	environmental	environmental	ADJ
alces-343	55	15	conditions	condition	NOUN
alces-343	55	16	(	(	PUNCT
alces-343	55	17	crête	crête	ADJ
alces-343	55	18	and	and	CCONJ
alces-343	55	19	courtois	courtois	NOUN
alces-343	55	20	1997	1997	NUM
alces-343	55	21	)	)	PUNCT
alces-343	55	22	.	.	PUNCT
alces-343	56	1	the	the	DET
alces-343	56	2	model	model	NOUN
alces-343	56	3	(	(	PUNCT
alces-343	56	4	appendix	appendix	VERB
alces-343	56	5	i	i	PRON
alces-343	56	6	)	)	PUNCT
alces-343	56	7	was	be	AUX
alces-343	56	8	elaborated	elaborate	VERB
alces-343	56	9	using	use	VERB
alces-343	56	10	stella	stella	PROPN
alces-343	56	11	4.0	4.0	NUM
alces-343	56	12	software	software	NOUN
alces-343	56	13	(	(	PUNCT
alces-343	56	14	isee	isee	NOUN
alces-343	56	15	systems	systems	PROPN
alces-343	56	16	,	,	PUNCT
alces-343	56	17	lebanon	lebanon	PROPN
alces-343	56	18	,	,	PUNCT
alces-343	56	19	nh	nh	PROPN
alces-343	56	20	,	,	PUNCT
alces-343	56	21	usa	usa	PROPN
alces-343	56	22	)	)	PUNCT
alces-343	56	23	.	.	PUNCT
alces-343	57	1	in	in	ADP
alces-343	57	2	summary	summary	NOUN
alces-343	57	3	,	,	PUNCT
alces-343	57	4	our	our	PRON
alces-343	57	5	combined	combined	ADJ
alces-343	57	6	model	model	NOUN
alces-343	57	7	is	be	AUX
alces-343	57	8	of	of	ADP
alces-343	57	9	predation	predation	NOUN
alces-343	57	10	or	or	CCONJ
alces-343	57	11	when	when	SCONJ
alces-343	57	12	predators	predator	NOUN
alces-343	57	13	are	be	AUX
alces-343	57	14	controlled	control	VERB
alces-343	57	15	,	,	PUNCT
alces-343	57	16	caribou	caribou	NOUN
alces-343	57	17	and	and	CCONJ
alces-343	57	18	moose	moose	NOUN
alces-343	57	19	populations	population	NOUN
alces-343	57	20	are	be	AUX
alces-343	57	21	regulated	regulate	VERB
alces-343	57	22	by	by	ADP
alces-343	57	23	competition	competition	NOUN
alces-343	57	24	for	for	ADP
alces-343	57	25	food	food	NOUN
alces-343	57	26	(	(	PUNCT
alces-343	57	27	messier	messy	ADJ
alces-343	57	28	1994	1994	NUM
alces-343	57	29	,	,	PUNCT
alces-343	57	30	crête	crête	ADJ
alces-343	57	31	and	and	CCONJ
alces-343	57	32	manseau	manseau	NOUN
alces-343	57	33	1996	1996	NUM
alces-343	57	34	)	)	PUNCT
alces-343	57	35	;	;	PUNCT
alces-343	57	36	(	(	PUNCT
alces-343	57	37	2	2	X
alces-343	57	38	)	)	PUNCT
alces-343	57	39	in	in	ADP
alces-343	57	40	the	the	DET
alces-343	57	41	presence	presence	NOUN
alces-343	57	42	of	of	ADP
alces-343	57	43	wolves	wolf	NOUN
alces-343	57	44	,	,	PUNCT
alces-343	57	45	the	the	DET
alces-343	57	46	moose	moose	NOUN
alces-343	57	47	population	population	NOUN
alces-343	57	48	is	be	AUX
alces-343	57	49	regulated	regulate	VERB
alces-343	57	50	by	by	ADP
alces-343	57	51	wolf	wolf	PROPN
alces-343	57	52	predation	predation	NOUN
alces-343	57	53	(	(	PUNCT
alces-343	57	54	messier	messy	ADJ
alces-343	57	55	1994	1994	NUM
alces-343	57	56	)	)	PUNCT
alces-343	57	57	;	;	PUNCT
alces-343	57	58	(	(	PUNCT
alces-343	57	59	3	3	X
alces-343	57	60	)	)	PUNCT
alces-343	57	61	wolf	wolf	NOUN
alces-343	57	62	numbers	number	NOUN
alces-343	57	63	are	be	AUX
alces-343	57	64	determined	determine	VERB
alces-343	57	65	by	by	ADP
alces-343	57	66	moose	moose	NOUN
alces-343	57	67	abundance	abundance	NOUN
alces-343	57	68	(	(	PUNCT
alces-343	57	69	messier	messy	ADJ
alces-343	57	70	1994	1994	NUM
alces-343	57	71	)	)	PUNCT
alces-343	57	72	,	,	PUNCT
alces-343	57	73	but	but	CCONJ
alces-343	57	74	there	there	PRON
alces-343	57	75	is	be	VERB
alces-343	57	76	no	no	DET
alces-343	57	77	dependance	dependance	NOUN
alces-343	57	78	of	of	ADP
alces-343	57	79	wolves	wolf	NOUN
alces-343	57	80	on	on	ADP
alces-343	57	81	caribou	caribou	NOUN
alces-343	57	82	numbers	number	NOUN
alces-343	57	83	(	(	PUNCT
alces-343	57	84	seip	seip	NOUN
alces-343	57	85	1991	1991	NUM
alces-343	57	86	)	)	PUNCT
alces-343	57	87	;	;	PUNCT
alces-343	57	88	(	(	PUNCT
alces-343	57	89	4	4	X
alces-343	57	90	)	)	PUNCT
alces-343	57	91	caribou	caribou	NOUN
alces-343	57	92	predation	predation	NOUN
alces-343	57	93	increases	increase	VERB
alces-343	57	94	non	non	ADJ
alces-343	57	95	-	-	ADJ
alces-343	57	96	linearly	linearly	ADV
alces-343	57	97	with	with	ADP
alces-343	57	98	wolf	wolf	PROPN
alces-343	57	99	abundance	abundance	PROPN
alces-343	57	100	(	(	PUNCT
alces-343	57	101	bergerud	bergerud	NOUN
alces-343	57	102	and	and	CCONJ
alces-343	57	103	elliot	elliot	PROPN
alces-343	57	104	1986	1986	NUM
alces-343	57	105	)	)	PUNCT
alces-343	57	106	;	;	PUNCT
alces-343	57	107	and	and	CCONJ
alces-343	57	108	(	(	PUNCT
alces-343	57	109	5	5	X
alces-343	57	110	)	)	PUNCT
alces-343	57	111	there	there	PRON
alces-343	57	112	is	be	VERB
alces-343	57	113	no	no	DET
alces-343	57	114	immigration	immigration	NOUN
alces-343	57	115	or	or	CCONJ
alces-343	57	116	emigration	emigration	NOUN
alces-343	57	117	in	in	ADP
alces-343	57	118	the	the	DET
alces-343	57	119	system	system	NOUN
alces-343	57	120	or	or	CCONJ
alces-343	57	121	these	these	DET
alces-343	57	122	two	two	NUM
alces-343	57	123	opposite	opposite	ADJ
alces-343	57	124	processes	process	NOUN
alces-343	57	125	are	be	AUX
alces-343	57	126	equal	equal	ADJ
alces-343	57	127	.	.	PUNCT
alces-343	58	1	caribou	caribou	NOUN
alces-343	58	2	population	population	NOUN
alces-343	58	3	parameters	parameter	NOUN
alces-343	58	4	due	due	ADP
alces-343	58	5	to	to	ADP
alces-343	58	6	the	the	DET
alces-343	58	7	paucity	paucity	NOUN
alces-343	58	8	of	of	ADP
alces-343	58	9	woodland	woodland	PROPN
alces-343	58	10	caribou	caribou	PROPN
alces-343	58	11	population	population	NOUN
alces-343	58	12	dynamics	dynamics	PROPN
alces-343	58	13	data	datum	NOUN
alces-343	58	14	,	,	PUNCT
alces-343	58	15	maximum	maximum	ADJ
alces-343	58	16	growth	growth	NOUN
alces-343	58	17	rate	rate	NOUN
alces-343	58	18	and	and	CCONJ
alces-343	58	19	food	food	NOUN
alces-343	58	20	carrying	carry	VERB
alces-343	58	21	capacity	capacity	NOUN
alces-343	58	22	were	be	AUX
alces-343	58	23	taken	take	VERB
alces-343	58	24	from	from	ADP
alces-343	58	25	published	publish	VERB
alces-343	58	26	data	datum	NOUN
alces-343	58	27	for	for	ADP
alces-343	58	28	barren	barren	NOUN
alces-343	58	29	-	-	PUNCT
alces-343	58	30	ground	ground	NOUN
alces-343	58	31	caribou	caribou	NOUN
alces-343	58	32	.	.	PUNCT
alces-343	59	1	maximum	maximum	ADJ
alces-343	59	2	population	population	NOUN
alces-343	59	3	growth	growth	NOUN
alces-343	59	4	rate	rate	NOUN
alces-343	59	5	of	of	ADP
alces-343	59	6	caribou	caribou	NOUN
alces-343	59	7	was	be	AUX
alces-343	59	8	based	base	VERB
alces-343	59	9	on	on	ADP
alces-343	59	10	observations	observation	NOUN
alces-343	59	11	of	of	ADP
alces-343	59	12	a	a	DET
alces-343	59	13	population	population	NOUN
alces-343	59	14	reintroduced	reintroduce	VERB
alces-343	59	15	on	on	ADP
alces-343	59	16	southampton	southampton	PROPN
alces-343	59	17	island	island	PROPN
alces-343	59	18	in	in	ADP
alces-343	59	19	hudson	hudson	PROPN
alces-343	59	20	bay	bay	PROPN
alces-343	59	21	(	(	PUNCT
alces-343	59	22	ouellet	ouellet	NOUN
alces-343	59	23	et	et	PROPN
alces-343	59	24	al	al	PROPN
alces-343	59	25	.	.	PROPN
alces-343	59	26	1996	1996	NUM
alces-343	59	27	,	,	PUNCT
alces-343	59	28	1997	1997	NUM
alces-343	59	29	)	)	PUNCT
alces-343	59	30	.	.	PUNCT
alces-343	60	1	in	in	ADP
alces-343	60	2	1991	1991	NUM
alces-343	60	3	,	,	PUNCT
alces-343	60	4	this	this	DET
alces-343	60	5	population	population	NOUN
alces-343	60	6	was	be	AUX
alces-343	60	7	estimated	estimate	VERB
alces-343	60	8	at	at	ADP
alces-343	60	9	13,700	13,700	NUM
alces-343	60	10	1,580	1,580	NUM
alces-343	60	11	adults	adult	NOUN
alces-343	60	12	,	,	PUNCT
alces-343	60	13	with	with	ADP
alces-343	60	14	an	an	DET
alces-343	60	15	annual	annual	ADJ
alces-343	60	16	growth	growth	NOUN
alces-343	60	17	rate	rate	NOUN
alces-343	60	18	(	(	PUNCT
alces-343	60	19	r	r	NOUN
alces-343	60	20	)	)	PUNCT
alces-343	60	21	of	of	ADP
alces-343	60	22	0.245	0.245	NUM
alces-343	60	23	since	since	SCONJ
alces-343	60	24	the	the	DET
alces-343	60	25	introduction	introduction	NOUN
alces-343	60	26	of	of	ADP
alces-343	60	27	38	38	NUM
alces-343	60	28	one	one	NUM
alces-343	60	29	-	-	PUNCT
alces-343	60	30	yearold	yearold	ADJ
alces-343	60	31	individuals	individual	NOUN
alces-343	60	32	in	in	ADP
alces-343	60	33	1967	1967	NUM
alces-343	60	34	(	(	PUNCT
alces-343	60	35	nt	not	PART
alces-343	60	36	=	=	PROPN
alces-343	60	37	n0	n0	NUM
alces-343	60	38	ert	ert	PROPN
alces-343	60	39	,	,	PUNCT
alces-343	60	40	hence	hence	ADV
alces-343	60	41	13,700	13,700	NUM
alces-343	60	42	=	=	SYM
alces-343	60	43	38	38	NUM
alces-343	60	44	er	er	INTJ
alces-343	60	45	24	24	NUM
alces-343	60	46	;	;	PUNCT
alces-343	60	47	where	where	SCONJ
alces-343	60	48	nt	not	PART
alces-343	60	49	=	=	SYM
alces-343	60	50	number	number	NOUN
alces-343	60	51	of	of	ADP
alces-343	60	52	caribou	caribou	NOUN
alces-343	60	53	at	at	ADP
alces-343	60	54	time	time	NOUN
alces-343	60	55	t	t	PROPN
alces-343	60	56	,	,	PUNCT
alces-343	60	57	n0	n0	ADJ
alces-343	60	58	=	=	PUNCT
alces-343	60	59	number	number	NOUN
alces-343	60	60	of	of	ADP
alces-343	60	61	caribou	caribou	NOUN
alces-343	60	62	in	in	ADP
alces-343	60	63	1967	1967	NUM
alces-343	60	64	,	,	PUNCT
alces-343	60	65	and	and	CCONJ
alces-343	60	66	t	t	NOUN
alces-343	60	67	=	=	PUNCT
alces-343	60	68	time	time	NOUN
alces-343	60	69	in	in	ADP
alces-343	60	70	years	year	NOUN
alces-343	60	71	)	)	PUNCT
alces-343	60	72	.	.	PUNCT
alces-343	61	1	no	no	DET
alces-343	61	2	decline	decline	NOUN
alces-343	61	3	in	in	ADP
alces-343	61	4	growth	growth	NOUN
alces-343	61	5	impact	impact	NOUN
alces-343	61	6	of	of	ADP
alces-343	61	7	moose	moose	NOUN
alces-343	61	8	and	and	CCONJ
alces-343	61	9	wolves	wolf	NOUN
alces-343	61	10	on	on	ADP
alces-343	61	11	caribou	caribou	NOUN
alces-343	61	12	–	–	PUNCT
alces-343	61	13	courtois	courtois	NOUN
alces-343	61	14	and	and	CCONJ
alces-343	61	15	ouellet	ouellet	PROPN
alces-343	61	16	alces	alces	PROPN
alces-343	61	17	vol	vol	NOUN
alces-343	61	18	.	.	PROPN
alces-343	61	19	43	43	NUM
alces-343	61	20	,	,	PUNCT
alces-343	61	21	2007	2007	NUM
alces-343	61	22	16	16	NUM
alces-343	61	23	rate	rate	NOUN
alces-343	61	24	was	be	AUX
alces-343	61	25	observed	observe	VERB
alces-343	61	26	over	over	ADP
alces-343	61	27	the	the	DET
alces-343	61	28	24	24	NUM
alces-343	61	29	years	year	NOUN
alces-343	61	30	,	,	PUNCT
alces-343	61	31	so	so	SCONJ
alces-343	61	32	this	this	DET
alces-343	61	33	value	value	NOUN
alces-343	61	34	of	of	ADP
alces-343	61	35	r	r	NOUN
alces-343	61	36	should	should	AUX
alces-343	61	37	be	be	AUX
alces-343	61	38	close	close	ADJ
alces-343	61	39	to	to	ADP
alces-343	61	40	the	the	DET
alces-343	61	41	maximum	maximum	NOUN
alces-343	61	42	attainable	attainable	ADJ
alces-343	61	43	by	by	ADP
alces-343	61	44	a	a	DET
alces-343	61	45	caribou	caribou	NOUN
alces-343	61	46	population	population	NOUN
alces-343	61	47	that	that	PRON
alces-343	61	48	is	be	AUX
alces-343	61	49	subject	subject	ADJ
alces-343	61	50	to	to	ADP
alces-343	61	51	negligible	negligible	ADJ
alces-343	61	52	hunting	hunting	NOUN
alces-343	61	53	(	(	PUNCT
alces-343	61	54	<	<	X
alces-343	61	55	1	1	NUM
alces-343	61	56	%	%	NOUN
alces-343	61	57	per	per	ADP
alces-343	61	58	year	year	NOUN
alces-343	61	59	)	)	PUNCT
alces-343	61	60	and	and	CCONJ
alces-343	61	61	lacks	lack	VERB
alces-343	61	62	wolf	wolf	PROPN
alces-343	61	63	predation	predation	NOUN
alces-343	61	64	.	.	PUNCT
alces-343	62	1	maximum	maximum	ADJ
alces-343	62	2	growth	growth	NOUN
alces-343	62	3	rate	rate	NOUN
alces-343	62	4	is	be	AUX
alces-343	62	5	comparable	comparable	ADJ
alces-343	62	6	to	to	ADP
alces-343	62	7	that	that	PRON
alces-343	62	8	reported	report	VERB
alces-343	62	9	for	for	ADP
alces-343	62	10	the	the	DET
alces-343	62	11	caribou	caribou	NOUN
alces-343	62	12	herd	herd	NOUN
alces-343	62	13	introduced	introduce	VERB
alces-343	62	14	on	on	ADP
alces-343	62	15	st	st	PROPN
alces-343	62	16	.	.	PROPN
alces-343	62	17	matthew	matthew	PROPN
alces-343	62	18	island	island	PROPN
alces-343	62	19	in	in	ADP
alces-343	62	20	the	the	DET
alces-343	62	21	bering	bering	NOUN
alces-343	62	22	sea	sea	NOUN
alces-343	62	23	(	(	PUNCT
alces-343	62	24	17	17	NUM
alces-343	62	25	–	–	SYM
alces-343	62	26	29	29	NUM
alces-343	62	27	%	%	NOUN
alces-343	62	28	;	;	PUNCT
alces-343	62	29	klein	klein	PROPN
alces-343	62	30	1968	1968	NUM
alces-343	62	31	)	)	PUNCT
alces-343	62	32	.	.	PUNCT
alces-343	63	1	carrying	carry	VERB
alces-343	63	2	capacity	capacity	NOUN
alces-343	63	3	based	base	VERB
alces-343	63	4	on	on	ADP
alces-343	63	5	availability	availability	NOUN
alces-343	63	6	of	of	ADP
alces-343	63	7	terrestrial	terrestrial	ADJ
alces-343	63	8	lichens	lichen	NOUN
alces-343	63	9	was	be	AUX
alces-343	63	10	estimated	estimate	VERB
alces-343	63	11	at	at	ADP
alces-343	63	12	20	20	NUM
alces-343	63	13	caribou	caribou	NOUN
alces-343	63	14	per	per	ADP
alces-343	63	15	100	100	NUM
alces-343	63	16	km2	km2	NOUN
alces-343	63	17	(	(	PUNCT
alces-343	63	18	crête	crête	ADJ
alces-343	63	19	and	and	CCONJ
alces-343	63	20	manseau	manseau	NOUN
alces-343	63	21	1996	1996	NUM
alces-343	63	22	,	,	PUNCT
alces-343	63	23	arsenault	arsenault	PROPN
alces-343	63	24	et	et	PROPN
alces-343	63	25	al	al	PROPN
alces-343	63	26	.	.	PROPN
alces-343	63	27	1997	1997	NUM
alces-343	63	28	)	)	PUNCT
alces-343	63	29	.	.	PUNCT
alces-343	64	1	caribou	caribou	NOUN
alces-343	64	2	can	can	AUX
alces-343	64	3	eat	eat	VERB
alces-343	64	4	various	various	ADJ
alces-343	64	5	items	item	NOUN
alces-343	64	6	(	(	PUNCT
alces-343	64	7	arboreal	arboreal	NOUN
alces-343	64	8	lichens	lichen	NOUN
alces-343	64	9	,	,	PUNCT
alces-343	64	10	deciduous	deciduous	ADJ
alces-343	64	11	leaves	leave	NOUN
alces-343	64	12	,	,	PUNCT
alces-343	64	13	sedges	sedge	NOUN
alces-343	64	14	,	,	PUNCT
alces-343	64	15	forbs	forb	NOUN
alces-343	64	16	,	,	PUNCT
alces-343	64	17	etc	etc	X
alces-343	64	18	.	.	X
alces-343	64	19	)	)	PUNCT
alces-343	64	20	,	,	PUNCT
alces-343	64	21	but	but	CCONJ
alces-343	64	22	these	these	PRON
alces-343	64	23	were	be	AUX
alces-343	64	24	not	not	PART
alces-343	64	25	included	include	VERB
alces-343	64	26	in	in	ADP
alces-343	64	27	the	the	DET
alces-343	64	28	carrying	carry	VERB
alces-343	64	29	capacity	capacity	NOUN
alces-343	64	30	estimates	estimate	NOUN
alces-343	64	31	to	to	PART
alces-343	64	32	avoid	avoid	VERB
alces-343	64	33	overestimation	overestimation	NOUN
alces-343	64	34	since	since	SCONJ
alces-343	64	35	winter	winter	NOUN
alces-343	64	36	diet	diet	NOUN
alces-343	64	37	is	be	AUX
alces-343	64	38	largely	largely	ADV
alces-343	64	39	dominated	dominate	VERB
alces-343	64	40	by	by	ADP
alces-343	64	41	terrestrial	terrestrial	ADJ
alces-343	64	42	lichens	lichen	NOUN
alces-343	64	43	(	(	PUNCT
alces-343	64	44	gauthier	gauthier	PROPN
alces-343	64	45	et	et	PROPN
alces-343	64	46	al	al	PROPN
alces-343	64	47	.	.	PROPN
alces-343	64	48	1989	1989	NUM
alces-343	64	49	)	)	PUNCT
alces-343	64	50	and	and	CCONJ
alces-343	64	51	this	this	DET
alces-343	64	52	season	season	NOUN
alces-343	64	53	is	be	AUX
alces-343	64	54	the	the	DET
alces-343	64	55	most	most	ADV
alces-343	64	56	likely	likely	ADJ
alces-343	64	57	to	to	PART
alces-343	64	58	be	be	AUX
alces-343	64	59	limiting	limit	VERB
alces-343	64	60	caribou	caribou	NOUN
alces-343	64	61	(	(	PUNCT
alces-343	64	62	klein	klein	PROPN
alces-343	64	63	1968	1968	NUM
alces-343	64	64	)	)	PUNCT
alces-343	64	65	.	.	PUNCT
alces-343	65	1	initial	initial	ADJ
alces-343	65	2	winter	winter	NOUN
alces-343	65	3	density	density	NOUN
alces-343	65	4	was	be	AUX
alces-343	65	5	set	set	VERB
alces-343	65	6	at	at	ADP
alces-343	65	7	1.63	1.63	NUM
alces-343	65	8	individuals	individual	NOUN
alces-343	65	9	per	per	ADP
alces-343	65	10	100	100	NUM
alces-343	65	11	km2	km2	NOUN
alces-343	65	12	,	,	PUNCT
alces-343	65	13	with	with	ADP
alces-343	65	14	16	16	NUM
alces-343	65	15	%	%	NOUN
alces-343	65	16	calves	calf	NOUN
alces-343	65	17	in	in	ADP
alces-343	65	18	the	the	DET
alces-343	65	19	population	population	NOUN
alces-343	65	20	,	,	PUNCT
alces-343	65	21	as	as	SCONJ
alces-343	65	22	noted	note	VERB
alces-343	65	23	in	in	ADP
alces-343	65	24	a	a	DET
alces-343	65	25	recent	recent	ADJ
alces-343	65	26	aerial	aerial	ADJ
alces-343	65	27	survey	survey	NOUN
alces-343	65	28	(	(	PUNCT
alces-343	65	29	courtois	courtois	PROPN
alces-343	65	30	et	et	PROPN
alces-343	65	31	al	al	PROPN
alces-343	65	32	.	.	PROPN
alces-343	65	33	2003	2003	NUM
alces-343	65	34	)	)	PUNCT
alces-343	65	35	.	.	PUNCT
alces-343	66	1	annual	annual	ADJ
alces-343	66	2	losses	loss	NOUN
alces-343	66	3	due	due	ADP
alces-343	66	4	to	to	ADP
alces-343	66	5	predation	predation	NOUN
alces-343	66	6	and	and	CCONJ
alces-343	66	7	hunting	hunting	NOUN
alces-343	66	8	(	(	PUNCT
alces-343	66	9	adults	adult	NOUN
alces-343	66	10	only	only	ADV
alces-343	66	11	)	)	PUNCT
alces-343	66	12	were	be	AUX
alces-343	66	13	initially	initially	ADV
alces-343	66	14	established	establish	VERB
alces-343	66	15	at	at	ADP
alces-343	66	16	3	3	NUM
alces-343	66	17	%	%	NOUN
alces-343	66	18	and	and	CCONJ
alces-343	66	19	8	8	NUM
alces-343	66	20	%	%	NOUN
alces-343	66	21	,	,	PUNCT
alces-343	66	22	respectively	respectively	ADV
alces-343	66	23	,	,	PUNCT
alces-343	66	24	as	as	SCONJ
alces-343	66	25	noted	note	VERB
alces-343	66	26	in	in	ADP
alces-343	66	27	central	central	ADJ
alces-343	66	28	québec	québec	NOUN
alces-343	66	29	(	(	PUNCT
alces-343	66	30	courtois	courtois	PROPN
alces-343	66	31	2003	2003	NUM
alces-343	66	32	)	)	PUNCT
alces-343	66	33	.	.	PUNCT
alces-343	67	1	we	we	PRON
alces-343	67	2	considered	consider	VERB
alces-343	67	3	that	that	SCONJ
alces-343	67	4	using	use	VERB
alces-343	67	5	empirical	empirical	ADJ
alces-343	67	6	data	datum	NOUN
alces-343	67	7	would	would	AUX
alces-343	67	8	allow	allow	VERB
alces-343	67	9	more	more	ADV
alces-343	67	10	realistic	realistic	ADJ
alces-343	67	11	simulations	simulation	NOUN
alces-343	67	12	for	for	ADP
alces-343	67	13	the	the	DET
alces-343	67	14	québec	québec	PROPN
alces-343	67	15	boreal	boreal	PROPN
alces-343	67	16	forest	forest	NOUN
alces-343	67	17	.	.	PUNCT
alces-343	68	1	moose	moose	PROPN
alces-343	68	2	population	population	NOUN
alces-343	68	3	parameters	parameter	NOUN
alces-343	68	4	maximum	maximum	ADJ
alces-343	68	5	annual	annual	ADJ
alces-343	68	6	growth	growth	NOUN
alces-343	68	7	rate	rate	NOUN
alces-343	68	8	of	of	ADP
alces-343	68	9	the	the	DET
alces-343	68	10	moose	moose	NOUN
alces-343	68	11	population	population	NOUN
alces-343	68	12	was	be	AUX
alces-343	68	13	estimated	estimate	VERB
alces-343	68	14	to	to	PART
alces-343	68	15	be	be	AUX
alces-343	68	16	25	25	NUM
alces-343	68	17	%	%	NOUN
alces-343	68	18	,	,	PUNCT
alces-343	68	19	based	base	VERB
alces-343	68	20	on	on	ADP
alces-343	68	21	observations	observation	NOUN
alces-343	68	22	in	in	ADP
alces-343	68	23	newfoundland	newfoundland	PROPN
alces-343	68	24	in	in	ADP
alces-343	68	25	the	the	DET
alces-343	68	26	absence	absence	NOUN
alces-343	68	27	of	of	ADP
alces-343	68	28	predation	predation	NOUN
alces-343	68	29	(	(	PUNCT
alces-343	68	30	fryxell	fryxell	NOUN
alces-343	68	31	et	et	PROPN
alces-343	68	32	al	al	PROPN
alces-343	68	33	.	.	PROPN
alces-343	68	34	1988	1988	NUM
alces-343	68	35	)	)	PUNCT
alces-343	68	36	.	.	PUNCT
alces-343	69	1	similar	similar	ADJ
alces-343	69	2	rates	rate	NOUN
alces-343	69	3	of	of	ADP
alces-343	69	4	increase	increase	NOUN
alces-343	69	5	were	be	AUX
alces-343	69	6	reported	report	VERB
alces-343	69	7	in	in	ADP
alces-343	69	8	south	south	ADJ
alces-343	69	9	-	-	PUNCT
alces-343	69	10	central	central	ADJ
alces-343	69	11	québec	québec	NOUN
alces-343	69	12	(	(	PUNCT
alces-343	69	13	23	23	NUM
alces-343	69	14	-	-	SYM
alces-343	69	15	24	24	NUM
alces-343	69	16	%	%	NOUN
alces-343	69	17	,	,	PUNCT
alces-343	69	18	laurian	laurian	PROPN
alces-343	69	19	et	et	PROPN
alces-343	69	20	al	al	PROPN
alces-343	69	21	.	.	PROPN
alces-343	69	22	2000	2000	NUM
alces-343	69	23	)	)	PUNCT
alces-343	69	24	.	.	PUNCT
alces-343	70	1	other	other	ADJ
alces-343	70	2	parameters	parameter	NOUN
alces-343	70	3	were	be	AUX
alces-343	70	4	taken	take	VERB
alces-343	70	5	from	from	ADP
alces-343	70	6	studies	study	NOUN
alces-343	70	7	carried	carry	VERB
alces-343	70	8	out	out	ADP
alces-343	70	9	in	in	ADP
alces-343	70	10	east	east	ADJ
alces-343	70	11	-	-	PUNCT
alces-343	70	12	central	central	ADJ
alces-343	70	13	québec	québec	NOUN
alces-343	70	14	where	where	SCONJ
alces-343	70	15	densities	density	NOUN
alces-343	70	16	are	be	AUX
alces-343	70	17	approximately	approximately	ADV
alces-343	70	18	3.0	3.0	NUM
alces-343	70	19	moose	moose	NOUN
alces-343	70	20	per	per	ADP
alces-343	70	21	100	100	NUM
alces-343	70	22	km2	km2	NOUN
alces-343	70	23	(	(	PUNCT
alces-343	70	24	gingras	gingras	X
alces-343	70	25	et	et	PROPN
alces-343	70	26	al.1989	al.1989	PROPN
alces-343	70	27	)	)	PUNCT
alces-343	70	28	.	.	PUNCT
alces-343	71	1	the	the	DET
alces-343	71	2	habitat	habitat	NOUN
alces-343	71	3	carrying	carry	VERB
alces-343	71	4	capacity	capacity	NOUN
alces-343	71	5	was	be	AUX
alces-343	71	6	estimated	estimate	VERB
alces-343	71	7	at	at	ADP
alces-343	71	8	84	84	NUM
alces-343	71	9	moose	moose	NOUN
alces-343	71	10	per	per	ADP
alces-343	71	11	100	100	NUM
alces-343	71	12	km2	km2	NOUN
alces-343	71	13	,	,	PUNCT
alces-343	71	14	based	base	VERB
alces-343	71	15	on	on	ADP
alces-343	71	16	annual	annual	ADJ
alces-343	71	17	production	production	NOUN
alces-343	71	18	of	of	ADP
alces-343	71	19	deciduous	deciduous	ADJ
alces-343	71	20	twigs	twig	NOUN
alces-343	71	21	available	available	ADJ
alces-343	71	22	during	during	ADP
alces-343	71	23	winter	winter	NOUN
alces-343	71	24	(	(	PUNCT
alces-343	71	25	courtois	courtois	PROPN
alces-343	71	26	et	et	PROPN
alces-343	71	27	al	al	PROPN
alces-343	71	28	.	.	PROPN
alces-343	71	29	1993	1993	NUM
alces-343	71	30	)	)	PUNCT
alces-343	71	31	.	.	PUNCT
alces-343	72	1	although	although	SCONJ
alces-343	72	2	this	this	DET
alces-343	72	3	estimate	estimate	NOUN
alces-343	72	4	may	may	AUX
alces-343	72	5	appear	appear	VERB
alces-343	72	6	low	low	ADJ
alces-343	72	7	,	,	PUNCT
alces-343	72	8	it	it	PRON
alces-343	72	9	is	be	AUX
alces-343	72	10	four	four	NUM
alces-343	72	11	times	time	NOUN
alces-343	72	12	higher	high	ADJ
alces-343	72	13	than	than	ADP
alces-343	72	14	the	the	DET
alces-343	72	15	minimum	minimum	ADJ
alces-343	72	16	density	density	NOUN
alces-343	72	17	required	require	VERB
alces-343	72	18	to	to	PART
alces-343	72	19	maintain	maintain	VERB
alces-343	72	20	viable	viable	ADJ
alces-343	72	21	wolf	wolf	NOUN
alces-343	72	22	populations	population	NOUN
alces-343	72	23	(	(	PUNCT
alces-343	72	24	20	20	NUM
alces-343	72	25	moose	moose	NOUN
alces-343	72	26	per	per	ADP
alces-343	72	27	100	100	NUM
alces-343	72	28	km2	km2	NOUN
alces-343	72	29	;	;	PUNCT
alces-343	72	30	messier	messy	ADJ
alces-343	72	31	1985	1985	NUM
alces-343	72	32	)	)	PUNCT
alces-343	72	33	.	.	PUNCT
alces-343	73	1	natural	natural	ADJ
alces-343	73	2	mortality	mortality	NOUN
alces-343	73	3	rate	rate	NOUN
alces-343	73	4	was	be	AUX
alces-343	73	5	set	set	VERB
alces-343	73	6	at	at	ADP
alces-343	73	7	9.2	9.2	NUM
alces-343	73	8	%	%	NOUN
alces-343	73	9	per	per	ADP
alces-343	73	10	year	year	NOUN
alces-343	73	11	(	(	PUNCT
alces-343	73	12	of	of	ADP
alces-343	73	13	which	which	PRON
alces-343	73	14	4.5	4.5	NUM
alces-343	73	15	%	%	NOUN
alces-343	73	16	was	be	AUX
alces-343	73	17	due	due	ADJ
alces-343	73	18	to	to	ADP
alces-343	73	19	causes	cause	NOUN
alces-343	73	20	other	other	ADJ
alces-343	73	21	than	than	ADP
alces-343	73	22	predation	predation	NOUN
alces-343	73	23	)	)	PUNCT
alces-343	73	24	while	while	SCONJ
alces-343	73	25	annual	annual	ADJ
alces-343	73	26	hunting	hunting	NOUN
alces-343	73	27	mortality	mortality	NOUN
alces-343	73	28	was	be	AUX
alces-343	73	29	estimated	estimate	VERB
alces-343	73	30	to	to	PART
alces-343	73	31	be	be	AUX
alces-343	73	32	9.0	9.0	NUM
alces-343	73	33	%	%	NOUN
alces-343	73	34	(	(	PUNCT
alces-343	73	35	courtois	courtois	PROPN
alces-343	73	36	et	et	PROPN
alces-343	73	37	al	al	PROPN
alces-343	73	38	.	.	PROPN
alces-343	73	39	1994b	1994b	NUM
alces-343	73	40	)	)	PUNCT
alces-343	73	41	.	.	PUNCT
alces-343	74	1	as	as	ADP
alces-343	74	2	for	for	ADP
alces-343	74	3	caribou	caribou	NOUN
alces-343	74	4	,	,	PUNCT
alces-343	74	5	these	these	DET
alces-343	74	6	estimates	estimate	NOUN
alces-343	74	7	were	be	AUX
alces-343	74	8	the	the	DET
alces-343	74	9	only	only	ADJ
alces-343	74	10	ones	one	NOUN
alces-343	74	11	available	available	ADJ
alces-343	74	12	for	for	ADP
alces-343	74	13	the	the	DET
alces-343	74	14	québec	québec	PROPN
alces-343	74	15	boreal	boreal	NOUN
alces-343	74	16	forest	forest	NOUN
alces-343	74	17	.	.	PUNCT
alces-343	75	1	additional	additional	ADJ
alces-343	75	2	simulations	simulation	NOUN
alces-343	75	3	involved	involve	VERB
alces-343	75	4	annual	annual	ADJ
alces-343	75	5	stochastic	stochastic	ADJ
alces-343	75	6	variation	variation	NOUN
alces-343	75	7	in	in	ADP
alces-343	75	8	birth	birth	NOUN
alces-343	75	9	rate	rate	NOUN
alces-343	75	10	in	in	ADP
alces-343	75	11	order	order	NOUN
alces-343	75	12	to	to	PART
alces-343	75	13	mimic	mimic	VERB
alces-343	75	14	productivity	productivity	NOUN
alces-343	75	15	changes	change	NOUN
alces-343	75	16	(	(	PUNCT
alces-343	75	17	0.56	0.56	NUM
alces-343	75	18	–	–	PUNCT
alces-343	75	19	1.00	1.00	NUM
alces-343	75	20	)	)	PUNCT
alces-343	75	21	that	that	PRON
alces-343	75	22	were	be	AUX
alces-343	75	23	caused	cause	VERB
alces-343	75	24	by	by	ADP
alces-343	75	25	variations	variation	NOUN
alces-343	75	26	in	in	ADP
alces-343	75	27	snowfall	snowfall	NOUN
alces-343	75	28	and	and	CCONJ
alces-343	75	29	summer	summer	NOUN
alces-343	75	30	temperature	temperature	NOUN
alces-343	75	31	(	(	PUNCT
alces-343	75	32	crête	crête	NOUN
alces-343	75	33	et	et	NOUN
alces-343	75	34	courtois	courtois	NOUN
alces-343	75	35	1997	1997	NUM
alces-343	75	36	)	)	PUNCT
alces-343	75	37	.	.	PUNCT
alces-343	76	1	it	it	PRON
alces-343	76	2	would	would	AUX
alces-343	76	3	have	have	AUX
alces-343	76	4	been	be	AUX
alces-343	76	5	possible	possible	ADJ
alces-343	76	6	to	to	PART
alces-343	76	7	use	use	VERB
alces-343	76	8	stochastic	stochastic	ADJ
alces-343	76	9	variability	variability	NOUN
alces-343	76	10	in	in	ADP
alces-343	76	11	all	all	DET
alces-343	76	12	parameters	parameter	NOUN
alces-343	76	13	but	but	CCONJ
alces-343	76	14	we	we	PRON
alces-343	76	15	preferred	prefer	VERB
alces-343	76	16	using	use	VERB
alces-343	76	17	actual	actual	ADJ
alces-343	76	18	estimates	estimate	NOUN
alces-343	76	19	to	to	PART
alces-343	76	20	allow	allow	VERB
alces-343	76	21	realistic	realistic	ADJ
alces-343	76	22	simulations	simulation	NOUN
alces-343	76	23	for	for	ADP
alces-343	76	24	the	the	DET
alces-343	76	25	study	study	NOUN
alces-343	76	26	site	site	NOUN
alces-343	76	27	.	.	PUNCT
alces-343	77	1	besides	besides	SCONJ
alces-343	77	2	,	,	PUNCT
alces-343	77	3	anthropogenic	anthropogenic	ADJ
alces-343	77	4	mortality	mortality	NOUN
alces-343	77	5	does	do	AUX
alces-343	77	6	not	not	PART
alces-343	77	7	vary	vary	VERB
alces-343	77	8	substantially	substantially	ADV
alces-343	77	9	from	from	ADP
alces-343	77	10	year	year	NOUN
alces-343	77	11	to	to	ADP
alces-343	77	12	year	year	NOUN
alces-343	77	13	because	because	SCONJ
alces-343	77	14	hunting	hunt	VERB
alces-343	77	15	regulation	regulation	NOUN
alces-343	77	16	remains	remain	VERB
alces-343	77	17	stable	stable	ADJ
alces-343	77	18	in	in	ADP
alces-343	77	19	the	the	DET
alces-343	77	20	area	area	NOUN
alces-343	77	21	.	.	PUNCT
alces-343	78	1	finally	finally	ADV
alces-343	78	2	,	,	PUNCT
alces-343	78	3	interpretation	interpretation	NOUN
alces-343	78	4	would	would	AUX
alces-343	78	5	have	have	VERB
alces-343	78	6	many	many	ADJ
alces-343	78	7	parameters	parameter	NOUN
alces-343	78	8	.	.	PUNCT
alces-343	79	1	wolf	wolf	PROPN
alces-343	79	2	population	population	NOUN
alces-343	79	3	estimation	estimation	NOUN
alces-343	79	4	our	our	PRON
alces-343	79	5	model	model	NOUN
alces-343	79	6	considers	consider	VERB
alces-343	79	7	wolf	wolf	PROPN
alces-343	79	8	population	population	NOUN
alces-343	79	9	size	size	NOUN
alces-343	79	10	to	to	PART
alces-343	79	11	be	be	AUX
alces-343	79	12	determined	determine	VERB
alces-343	79	13	by	by	ADP
alces-343	79	14	moose	moose	NOUN
alces-343	79	15	population	population	NOUN
alces-343	79	16	size	size	NOUN
alces-343	79	17	,	,	PUNCT
alces-343	79	18	according	accord	VERB
alces-343	79	19	to	to	ADP
alces-343	79	20	the	the	DET
alces-343	79	21	predation	predation	NOUN
alces-343	79	22	model	model	NOUN
alces-343	79	23	developed	develop	VERB
alces-343	79	24	by	by	ADP
alces-343	79	25	messier	messy	ADJ
alces-343	79	26	(	(	PUNCT
alces-343	79	27	1994	1994	NUM
alces-343	79	28	,	,	PUNCT
alces-343	79	29	1995	1995	NUM
alces-343	79	30	)	)	PUNCT
alces-343	79	31	from	from	ADP
alces-343	79	32	27	27	NUM
alces-343	79	33	north	north	ADJ
alces-343	79	34	american	american	ADJ
alces-343	79	35	studies	study	NOUN
alces-343	79	36	.	.	PUNCT
alces-343	80	1	this	this	DET
alces-343	80	2	author	author	NOUN
alces-343	80	3	demonstrated	demonstrate	VERB
alces-343	80	4	that	that	SCONJ
alces-343	80	5	it	it	PRON
alces-343	80	6	was	be	AUX
alces-343	80	7	possible	possible	ADJ
alces-343	80	8	to	to	PART
alces-343	80	9	predict	predict	VERB
alces-343	80	10	wolf	wolf	PROPN
alces-343	80	11	abundance	abundance	NOUN
alces-343	80	12	as	as	ADP
alces-343	80	13	a	a	DET
alces-343	80	14	function	function	NOUN
alces-343	80	15	of	of	ADP
alces-343	80	16	moose	moose	NOUN
alces-343	80	17	density	density	NOUN
alces-343	80	18	(	(	PUNCT
alces-343	80	19	numerical	numerical	ADJ
alces-343	80	20	response	response	NOUN
alces-343	80	21	:	:	PUNCT
alces-343	80	22	number	number	NOUN
alces-343	80	23	of	of	ADP
alces-343	80	24	wolves/1,000	wolves/1,000	PROPN
alces-343	80	25	km2	km2	PROPN
alces-343	80	26	=	=	PUNCT
alces-343	81	1	58.73	58.73	NUM
alces-343	81	2	[	[	X
alces-343	81	3	number	number	NOUN
alces-343	81	4	of	of	ADP
alces-343	81	5	moose	moose	NOUN
alces-343	81	6	/	/	SYM
alces-343	81	7	km2	km2	PROPN
alces-343	81	8	–	–	PUNCT
alces-343	81	9	0.031]/[0.76	0.031]/[0.76	NOUN
alces-343	81	10	+	+	CCONJ
alces-343	81	11	number	number	NOUN
alces-343	81	12	of	of	ADP
alces-343	81	13	moose	moose	NOUN
alces-343	81	14	/	/	SYM
alces-343	81	15	km2	km2	NOUN
alces-343	81	16	]	]	PUNCT
alces-343	81	17	,	,	PUNCT
alces-343	81	18	r2	r2	PROPN
alces-343	81	19	=	=	NOUN
alces-343	81	20	0.62	0.62	NUM
alces-343	81	21	)	)	PUNCT
alces-343	81	22	.	.	PUNCT
alces-343	82	1	the	the	DET
alces-343	82	2	number	number	NOUN
alces-343	82	3	of	of	ADP
alces-343	82	4	moose	moose	NOUN
alces-343	82	5	exponentially	exponentially	ADV
alces-343	82	6	as	as	ADP
alces-343	82	7	a	a	DET
alces-343	82	8	function	function	NOUN
alces-343	82	9	of	of	ADP
alces-343	82	10	prey	prey	PROPN
alces-343	82	11	density	density	NOUN
alces-343	82	12	then	then	ADV
alces-343	82	13	decelerates	decelerate	VERB
alces-343	82	14	and	and	CCONJ
alces-343	82	15	starts	start	VERB
alces-343	82	16	declining	decline	VERB
alces-343	82	17	at	at	ADP
alces-343	82	18	0.65	0.65	NUM
alces-343	82	19	moose	moose	NOUN
alces-343	82	20	/	/	SYM
alces-343	82	21	km2	km2	PROPN
alces-343	82	22	(	(	PUNCT
alces-343	82	23	functional	functional	ADJ
alces-343	82	24	response	response	NOUN
alces-343	82	25	:	:	PUNCT
alces-343	82	26	number	number	NOUN
alces-343	82	27	of	of	ADP
alces-343	82	28	moose	moose	NOUN
alces-343	82	29	killed	kill	VERB
alces-343	82	30	per	per	ADP
alces-343	82	31	wolf	wolf	PROPN
alces-343	82	32	per	per	ADP
alces-343	82	33	100	100	NUM
alces-343	82	34	days	day	NOUN
alces-343	82	35	=	=	SYM
alces-343	82	36	3.3	3.3	NUM
alces-343	82	37	number	number	NOUN
alces-343	82	38	of	of	ADP
alces-343	82	39	moose	moose	NOUN
alces-343	82	40	per	per	ADP
alces-343	82	41	km2/[0.46	km2/[0.46	NOUN
alces-343	82	42	+	+	CCONJ
alces-343	82	43	number	number	NOUN
alces-343	82	44	of	of	ADP
alces-343	82	45	moose	moose	NOUN
alces-343	82	46	per	per	ADP
alces-343	82	47	km2	km2	NOUN
alces-343	82	48	]	]	PUNCT
alces-343	82	49	,	,	PUNCT
alces-343	82	50	r2	r2	PROPN
alces-343	82	51	=	=	NOUN
alces-343	82	52	0.53	0.53	NUM
alces-343	82	53	;	;	PUNCT
alces-343	82	54	messier	messy	ADJ
alces-343	82	55	1994	1994	NUM
alces-343	82	56	)	)	PUNCT
alces-343	82	57	.	.	PUNCT
alces-343	83	1	in	in	ADP
alces-343	83	2	this	this	DET
alces-343	83	3	model	model	NOUN
alces-343	83	4	,	,	PUNCT
alces-343	83	5	the	the	DET
alces-343	83	6	total	total	ADJ
alces-343	83	7	response	response	NOUN
alces-343	83	8	is	be	AUX
alces-343	83	9	the	the	DET
alces-343	83	10	product	product	NOUN
alces-343	83	11	of	of	ADP
alces-343	83	12	numerical	numerical	ADJ
alces-343	83	13	and	and	CCONJ
alces-343	83	14	functional	functional	ADJ
alces-343	83	15	responses	response	NOUN
alces-343	83	16	.	.	PUNCT
alces-343	84	1	following	follow	VERB
alces-343	84	2	recommendations	recommendation	NOUN
alces-343	84	3	of	of	ADP
alces-343	84	4	messier	messy	ADJ
alces-343	84	5	(	(	PUNCT
alces-343	84	6	1994	1994	NUM
alces-343	84	7	)	)	PUNCT
alces-343	84	8	,	,	PUNCT
alces-343	84	9	a	a	DET
alces-343	84	10	correction	correction	NOUN
alces-343	84	11	factor	factor	NOUN
alces-343	84	12	of	of	ADP
alces-343	84	13	0.71	0.71	NUM
alces-343	84	14	was	be	AUX
alces-343	84	15	applied	apply	VERB
alces-343	84	16	to	to	ADP
alces-343	84	17	winter	winter	NOUN
alces-343	84	18	predation	predation	NOUN
alces-343	84	19	rates	rate	NOUN
alces-343	84	20	to	to	PART
alces-343	84	21	obtain	obtain	VERB
alces-343	84	22	annual	annual	ADJ
alces-343	84	23	rates	rate	NOUN
alces-343	84	24	and	and	CCONJ
alces-343	84	25	values	value	NOUN
alces-343	84	26	for	for	ADP
alces-343	84	27	100	100	NUM
alces-343	84	28	days	day	NOUN
alces-343	84	29	were	be	AUX
alces-343	84	30	converted	convert	VERB
alces-343	84	31	to	to	ADP
alces-343	84	32	alces	alce	NOUN
alces-343	84	33	vol	vol	NOUN
alces-343	84	34	.	.	PROPN
alces-343	85	1	43	43	NUM
alces-343	85	2	,	,	PUNCT
alces-343	85	3	2007	2007	NUM
alces-343	85	4	courtois	courtois	NOUN
alces-343	85	5	and	and	CCONJ
alces-343	85	6	ouellet	ouellet	NOUN
alces-343	85	7	impact	impact	NOUN
alces-343	85	8	of	of	ADP
alces-343	85	9	moose	moose	NOUN
alces-343	85	10	and	and	CCONJ
alces-343	85	11	wolves	wolf	NOUN
alces-343	85	12	on	on	ADP
alces-343	85	13	caribou	caribou	NOUN
alces-343	85	14	17	17	NUM
alces-343	85	15	annual	annual	ADJ
alces-343	85	16	values	value	NOUN
alces-343	85	17	.	.	PUNCT
alces-343	86	1	the	the	DET
alces-343	86	2	impact	impact	NOUN
alces-343	86	3	of	of	ADP
alces-343	86	4	wolves	wolf	NOUN
alces-343	86	5	on	on	ADP
alces-343	86	6	the	the	DET
alces-343	86	7	caribou	caribou	NOUN
alces-343	86	8	population	population	NOUN
alces-343	86	9	was	be	AUX
alces-343	86	10	estimated	estimate	VERB
alces-343	86	11	using	use	VERB
alces-343	86	12	the	the	DET
alces-343	86	13	bergerud	bergerud	NOUN
alces-343	86	14	and	and	CCONJ
alces-343	86	15	elliot	elliot	PROPN
alces-343	86	16	(	(	PUNCT
alces-343	86	17	1986	1986	NUM
alces-343	86	18	)	)	PUNCT
alces-343	86	19	density	density	NOUN
alces-343	86	20	-	-	PUNCT
alces-343	86	21	independent	independent	ADJ
alces-343	86	22	model	model	NOUN
alces-343	86	23	that	that	PRON
alces-343	86	24	was	be	AUX
alces-343	86	25	based	base	VERB
alces-343	86	26	on	on	ADP
alces-343	86	27	17	17	NUM
alces-343	86	28	north	north	ADJ
alces-343	86	29	american	american	ADJ
alces-343	86	30	studies	study	NOUN
alces-343	86	31	.	.	PUNCT
alces-343	87	1	we	we	PRON
alces-343	87	2	thus	thus	ADV
alces-343	87	3	predicted	predict	VERB
alces-343	87	4	annual	annual	ADJ
alces-343	87	5	recruitment	recruitment	NOUN
alces-343	87	6	rate	rate	NOUN
alces-343	87	7	(	(	PUNCT
alces-343	87	8	percentage	percentage	NOUN
alces-343	87	9	calves	calf	NOUN
alces-343	87	10	during	during	ADP
alces-343	87	11	winter	winter	NOUN
alces-343	87	12	=	=	SYM
alces-343	87	13	e(3.340	e(3.340	NUM
alces-343	87	14	-	-	PUNCT
alces-343	87	15	0.127	0.127	NUM
alces-343	87	16	wolves/1,000	wolves/1,000	ADJ
alces-343	87	17	km2	km2	PROPN
alces-343	87	18	)	)	PUNCT
alces-343	87	19	,	,	PUNCT
alces-343	87	20	r2	r2	NOUN
alces-343	87	21	=	=	NOUN
alces-343	87	22	0.69	0.69	NUM
alces-343	87	23	)	)	PUNCT
alces-343	87	24	and	and	CCONJ
alces-343	87	25	annual	annual	ADJ
alces-343	87	26	mortality	mortality	NOUN
alces-343	87	27	rate	rate	NOUN
alces-343	87	28	of	of	ADP
alces-343	87	29	adults	adult	NOUN
alces-343	87	30	(	(	PUNCT
alces-343	87	31	percentage	percentage	NOUN
alces-343	87	32	adult	adult	NOUN
alces-343	87	33	mortality	mortality	NOUN
alces-343	87	34	=	=	SYM
alces-343	88	1	4,766	4,766	NUM
alces-343	89	1	+	+	CCONJ
alces-343	89	2	(	(	PUNCT
alces-343	89	3	0.699	0.699	NUM
alces-343	89	4	wolves	wolf	NOUN
alces-343	89	5	per	per	ADP
alces-343	89	6	1,000	1,000	NUM
alces-343	89	7	km2)1,275	km2)1,275	PROPN
alces-343	89	8	,	,	PUNCT
alces-343	89	9	r2	r2	PROPN
alces-343	89	10	=	=	NOUN
alces-343	89	11	0.73	0.73	NUM
alces-343	89	12	)	)	PUNCT
alces-343	89	13	.	.	PUNCT
alces-343	90	1	simulations	simulation	NOUN
alces-343	90	2	parameters	parameter	NOUN
alces-343	90	3	and	and	CCONJ
alces-343	90	4	initial	initial	ADJ
alces-343	90	5	and	and	CCONJ
alces-343	90	6	calculated	calculated	ADJ
alces-343	90	7	variables	variable	NOUN
alces-343	90	8	are	be	AUX
alces-343	90	9	presented	present	VERB
alces-343	90	10	in	in	ADP
alces-343	90	11	appendix	appendix	PROPN
alces-343	90	12	ii	ii	PROPN
alces-343	90	13	.	.	PUNCT
alces-343	91	1	preliminary	preliminary	ADJ
alces-343	91	2	simulations	simulation	NOUN
alces-343	91	3	were	be	AUX
alces-343	91	4	performed	perform	VERB
alces-343	91	5	on	on	ADP
alces-343	91	6	caribou	caribou	PROPN
alces-343	91	7	and	and	CCONJ
alces-343	91	8	moose	moose	NOUN
alces-343	91	9	populations	population	NOUN
alces-343	91	10	separately	separately	ADV
alces-343	91	11	illustrated	illustrate	VERB
alces-343	91	12	the	the	DET
alces-343	91	13	trajectories	trajectory	NOUN
alces-343	91	14	of	of	ADP
alces-343	91	15	populations	population	NOUN
alces-343	91	16	that	that	PRON
alces-343	91	17	petition	petition	VERB
alces-343	91	18	for	for	ADP
alces-343	91	19	food	food	NOUN
alces-343	91	20	.	.	PUNCT
alces-343	92	1	in	in	ADP
alces-343	92	2	a	a	DET
alces-343	92	3	second	second	ADJ
alces-343	92	4	trial	trial	NOUN
alces-343	92	5	,	,	PUNCT
alces-343	92	6	we	we	PRON
alces-343	92	7	used	use	VERB
alces-343	92	8	the	the	DET
alces-343	92	9	moose	moose	NOUN
alces-343	92	10	-	-	PUNCT
alces-343	92	11	wolf	wolf	NOUN
alces-343	92	12	model	model	NOUN
alces-343	92	13	of	of	ADP
alces-343	92	14	messier	messy	ADJ
alces-343	92	15	(	(	PUNCT
alces-343	92	16	1994	1994	NUM
alces-343	92	17	)	)	PUNCT
alces-343	92	18	to	to	PART
alces-343	92	19	illustrate	illustrate	VERB
alces-343	92	20	the	the	DET
alces-343	92	21	reaction	reaction	NOUN
alces-343	92	22	of	of	ADP
alces-343	92	23	wolves	wolf	NOUN
alces-343	92	24	in	in	ADP
alces-343	92	25	the	the	DET
alces-343	92	26	presence	presence	NOUN
alces-343	92	27	of	of	ADP
alces-343	92	28	moose	moose	NOUN
alces-343	92	29	only	only	ADV
alces-343	92	30	.	.	PUNCT
alces-343	93	1	after	after	ADP
alces-343	93	2	these	these	DET
alces-343	93	3	preliminary	preliminary	ADJ
alces-343	93	4	trials	trial	NOUN
alces-343	93	5	,	,	PUNCT
alces-343	93	6	we	we	PRON
alces-343	93	7	ran	run	VERB
alces-343	93	8	the	the	DET
alces-343	93	9	combined	combined	ADJ
alces-343	93	10	model	model	NOUN
alces-343	93	11	using	use	VERB
alces-343	93	12	the	the	DET
alces-343	93	13	three	three	NUM
alces-343	93	14	species	specie	NOUN
alces-343	93	15	’	'	PUNCT
alces-343	93	16	interactions	interaction	NOUN
alces-343	93	17	to	to	PART
alces-343	93	18	investigate	investigate	VERB
alces-343	93	19	the	the	DET
alces-343	93	20	outcome	outcome	NOUN
alces-343	93	21	of	of	ADP
alces-343	93	22	a	a	DET
alces-343	93	23	limited	limited	ADJ
alces-343	93	24	number	number	NOUN
alces-343	93	25	of	of	ADP
alces-343	93	26	realistic	realistic	ADJ
alces-343	93	27	management	management	NOUN
alces-343	93	28	scenarios	scenario	NOUN
alces-343	93	29	over	over	ADP
alces-343	93	30	a	a	DET
alces-343	93	31	100	100	NUM
alces-343	93	32	-	-	PUNCT
alces-343	93	33	year	year	NOUN
alces-343	93	34	span	span	NOUN
alces-343	93	35	.	.	PUNCT
alces-343	94	1	four	four	NUM
alces-343	94	2	deterministic	deterministic	ADJ
alces-343	94	3	scenarios	scenario	NOUN
alces-343	94	4	were	be	AUX
alces-343	94	5	conducted	conduct	VERB
alces-343	94	6	.	.	PUNCT
alces-343	95	1	in	in	ADP
alces-343	95	2	trajectories	trajectory	NOUN
alces-343	95	3	under	under	ADP
alces-343	95	4	management	management	NOUN
alces-343	95	5	strategies	strategy	NOUN
alces-343	95	6	that	that	PRON
alces-343	95	7	had	have	AUX
alces-343	95	8	prevailed	prevail	VERB
alces-343	95	9	in	in	ADP
alces-343	95	10	central	central	ADJ
alces-343	95	11	québec	québec	NOUN
alces-343	95	12	until	until	ADP
alces-343	95	13	autumn	autumn	NOUN
alces-343	95	14	2000	2000	NUM
alces-343	95	15	(	(	PUNCT
alces-343	95	16	i.e.	i.e.	X
alces-343	95	17	,	,	PUNCT
alces-343	95	18	moose	moose	NOUN
alces-343	95	19	and	and	CCONJ
alces-343	95	20	caribou	caribou	NOUN
alces-343	95	21	hunting	hunting	NOUN
alces-343	95	22	at	at	ADP
alces-343	95	23	9	9	NUM
alces-343	95	24	%	%	NOUN
alces-343	95	25	and	and	CCONJ
alces-343	95	26	8	8	NUM
alces-343	95	27	%	%	NOUN
alces-343	95	28	,	,	PUNCT
alces-343	95	29	respectively	respectively	ADV
alces-343	95	30	,	,	PUNCT
alces-343	95	31	but	but	CCONJ
alces-343	95	32	no	no	DET
alces-343	95	33	wolf	wolf	NOUN
alces-343	95	34	trapping	trapping	NOUN
alces-343	95	35	)	)	PUNCT
alces-343	95	36	.	.	PUNCT
alces-343	96	1	then	then	ADV
alces-343	96	2	we	we	PRON
alces-343	96	3	successively	successively	ADV
alces-343	96	4	simulated	simulate	VERB
alces-343	96	5	effects	effect	NOUN
alces-343	96	6	of	of	ADP
alces-343	96	7	(	(	PUNCT
alces-343	96	8	2	2	X
alces-343	96	9	)	)	PUNCT
alces-343	96	10	a	a	DET
alces-343	96	11	caribou	caribou	NOUN
alces-343	96	12	hunting	hunt	VERB
alces-343	96	13	ban	ban	NOUN
alces-343	96	14	;	;	PUNCT
alces-343	96	15	(	(	PUNCT
alces-343	96	16	3	3	X
alces-343	96	17	)	)	PUNCT
alces-343	96	18	intensive	intensive	ADJ
alces-343	96	19	wolf	wolf	NOUN
alces-343	96	20	trapping	trap	VERB
alces-343	96	21	(	(	PUNCT
alces-343	96	22	at	at	ADP
alces-343	96	23	30	30	NUM
alces-343	96	24	%	%	NOUN
alces-343	96	25	;	;	PUNCT
alces-343	96	26	larivière	larivière	NOUN
alces-343	96	27	et	et	PROPN
alces-343	96	28	al	al	PROPN
alces-343	96	29	.	.	PROPN
alces-343	96	30	2000	2000	NUM
alces-343	96	31	)	)	PUNCT
alces-343	96	32	,	,	PUNCT
alces-343	96	33	and	and	CCONJ
alces-343	96	34	(	(	PUNCT
alces-343	96	35	4	4	X
alces-343	96	36	)	)	PUNCT
alces-343	96	37	moose	moose	NOUN
alces-343	96	38	hunting	hunting	NOUN
alces-343	96	39	increased	increase	VERB
alces-343	96	40	to	to	ADP
alces-343	96	41	15	15	NUM
alces-343	96	42	%	%	NOUN
alces-343	96	43	(	(	PUNCT
alces-343	96	44	in	in	ADP
alces-343	96	45	order	order	NOUN
alces-343	96	46	to	to	PART
alces-343	96	47	allow	allow	VERB
alces-343	96	48	only	only	ADV
alces-343	96	49	a	a	DET
alces-343	96	50	light	light	ADJ
alces-343	96	51	population	population	NOUN
alces-343	96	52	increase	increase	NOUN
alces-343	96	53	)	)	PUNCT
alces-343	96	54	without	without	ADP
alces-343	96	55	wolf	wolf	NOUN
alces-343	96	56	trapping	trapping	NOUN
alces-343	96	57	.	.	PUNCT
alces-343	97	1	finally	finally	ADV
alces-343	97	2	(	(	PUNCT
alces-343	97	3	5	5	NUM
alces-343	97	4	)	)	PUNCT
alces-343	97	5	,	,	PUNCT
alces-343	97	6	in	in	ADP
alces-343	97	7	order	order	NOUN
alces-343	97	8	to	to	PART
alces-343	97	9	experiment	experiment	VERB
alces-343	97	10	with	with	ADP
alces-343	97	11	the	the	DET
alces-343	97	12	effects	effect	NOUN
alces-343	97	13	of	of	ADP
alces-343	97	14	changes	change	NOUN
alces-343	97	15	not	not	PART
alces-343	97	16	controlled	control	VERB
alces-343	97	17	by	by	ADP
alces-343	97	18	harvest	harvest	NOUN
alces-343	97	19	regulation	regulation	NOUN
alces-343	97	20	,	,	PUNCT
alces-343	97	21	we	we	PRON
alces-343	97	22	included	include	VERB
alces-343	97	23	negative	negative	ADJ
alces-343	97	24	effects	effect	NOUN
alces-343	97	25	of	of	ADP
alces-343	97	26	stochastic	stochastic	ADJ
alces-343	97	27	environmental	environmental	ADJ
alces-343	97	28	variations	variation	NOUN
alces-343	97	29	on	on	ADP
alces-343	97	30	productivity	productivity	NOUN
alces-343	97	31	(	(	PUNCT
alces-343	97	32	crête	crête	ADJ
alces-343	97	33	and	and	CCONJ
alces-343	97	34	courtois	courtois	PROPN
alces-343	97	35	1997	1997	NUM
alces-343	97	36	)	)	PUNCT
alces-343	97	37	of	of	ADP
alces-343	97	38	moose	moose	NOUN
alces-343	97	39	(	(	PUNCT
alces-343	97	40	from	from	ADP
alces-343	97	41	0	0	NUM
alces-343	97	42	%	%	NOUN
alces-343	97	43	–	–	PUNCT
alces-343	97	44	40	40	NUM
alces-343	97	45	%	%	NOUN
alces-343	97	46	)	)	PUNCT
alces-343	97	47	and	and	CCONJ
alces-343	97	48	caribou	caribou	NOUN
alces-343	97	49	(	(	PUNCT
alces-343	97	50	20	20	NUM
alces-343	97	51	%	%	NOUN
alces-343	97	52	)	)	PUNCT
alces-343	97	53	,	,	PUNCT
alces-343	97	54	on	on	ADP
alces-343	97	55	scenarios	scenario	NOUN
alces-343	97	56	1	1	NUM
alces-343	97	57	and	and	CCONJ
alces-343	97	58	2	2	NUM
alces-343	97	59	.	.	NOUN
alces-343	97	60	results	result	VERB
alces-343	97	61	simple	simple	ADJ
alces-343	97	62	models	model	NOUN
alces-343	97	63	for	for	ADP
alces-343	97	64	caribou	caribou	NOUN
alces-343	97	65	,	,	PUNCT
alces-343	97	66	moose	moose	NOUN
alces-343	97	67	,	,	PUNCT
alces-343	97	68	and	and	CCONJ
alces-343	97	69	wolf	wolf	NOUN
alces-343	97	70	populations	population	NOUN
alces-343	97	71	in	in	ADP
alces-343	97	72	the	the	DET
alces-343	97	73	absence	absence	NOUN
alces-343	97	74	of	of	ADP
alces-343	97	75	hunting	hunting	NOUN
alces-343	97	76	and	and	CCONJ
alces-343	97	77	predation	predation	NOUN
alces-343	97	78	,	,	PUNCT
alces-343	97	79	caribou	caribou	PROPN
alces-343	97	80	reached	reach	VERB
alces-343	97	81	their	their	PRON
alces-343	97	82	carrying	carrying	NOUN
alces-343	97	83	capacity	capacity	NOUN
alces-343	97	84	(	(	PUNCT
alces-343	97	85	k	k	NOUN
alces-343	97	86	)	)	PUNCT
alces-343	97	87	in	in	ADP
alces-343	97	88	53	53	NUM
alces-343	97	89	years	year	NOUN
alces-343	97	90	,	,	PUNCT
alces-343	97	91	with	with	ADP
alces-343	97	92	numbers	number	NOUN
alces-343	97	93	increasing	increase	VERB
alces-343	97	94	from	from	ADP
alces-343	97	95	16	16	NUM
alces-343	97	96	to	to	ADP
alces-343	97	97	200	200	NUM
alces-343	97	98	individuals	individual	NOUN
alces-343	97	99	in	in	ADP
alces-343	97	100	the	the	DET
alces-343	97	101	1,000	1,000	NUM
alces-343	97	102	-	-	PUNCT
alces-343	97	103	km2	km2	NOUN
alces-343	97	104	site	site	NOUN
alces-343	97	105	.	.	PUNCT
alces-343	98	1	half	half	NOUN
alces-343	98	2	of	of	ADP
alces-343	98	3	the	the	DET
alces-343	98	4	carrying	carry	VERB
alces-343	98	5	capacity	capacity	NOUN
alces-343	98	6	(	(	PUNCT
alces-343	98	7	optimal	optimal	ADJ
alces-343	98	8	density	density	NOUN
alces-343	98	9	0.5k	0.5k	NOUN
alces-343	98	10	)	)	PUNCT
alces-343	98	11	was	be	AUX
alces-343	98	12	reached	reach	VERB
alces-343	98	13	in	in	ADP
alces-343	98	14	10	10	NUM
alces-343	98	15	years	year	NOUN
alces-343	98	16	,	,	PUNCT
alces-343	98	17	allowing	allow	VERB
alces-343	98	18	a	a	DET
alces-343	98	19	maximum	maximum	ADJ
alces-343	98	20	sustainable	sustainable	ADJ
alces-343	98	21	yield	yield	NOUN
alces-343	98	22	of	of	ADP
alces-343	98	23	12	12	NUM
alces-343	98	24	caribou	caribou	NOUN
alces-343	98	25	per	per	ADP
alces-343	98	26	year	year	NOUN
alces-343	98	27	in	in	ADP
alces-343	98	28	a	a	DET
alces-343	98	29	stable	stable	ADJ
alces-343	98	30	population	population	NOUN
alces-343	98	31	of	of	ADP
alces-343	98	32	100	100	NUM
alces-343	98	33	individuals	individual	NOUN
alces-343	98	34	.	.	PUNCT
alces-343	99	1	adding	add	VERB
alces-343	99	2	a	a	DET
alces-343	99	3	natural	natural	ADJ
alces-343	99	4	mortality	mortality	NOUN
alces-343	99	5	of	of	ADP
alces-343	99	6	3	3	NUM
alces-343	99	7	%	%	NOUN
alces-343	99	8	resulted	result	VERB
alces-343	99	9	in	in	ADP
alces-343	99	10	0.5	0.5	NUM
alces-343	99	11	k	k	NOUN
alces-343	99	12	being	be	AUX
alces-343	99	13	reached	reach	VERB
alces-343	99	14	in	in	ADP
alces-343	99	15	12	12	NUM
alces-343	99	16	years	year	NOUN
alces-343	99	17	,	,	PUNCT
alces-343	99	18	with	with	ADP
alces-343	99	19	a	a	DET
alces-343	99	20	maximum	maximum	ADJ
alces-343	99	21	sustainable	sustainable	ADJ
alces-343	99	22	yield	yield	NOUN
alces-343	99	23	of	of	ADP
alces-343	99	24	9	9	NUM
alces-343	99	25	individuals	individual	NOUN
alces-343	99	26	per	per	ADP
alces-343	99	27	year	year	NOUN
alces-343	99	28	.	.	PUNCT
alces-343	100	1	when	when	SCONJ
alces-343	100	2	an	an	DET
alces-343	100	3	additional	additional	ADJ
alces-343	100	4	anthropogenic	anthropogenic	ADJ
alces-343	100	5	harvest	harvest	NOUN
alces-343	100	6	of	of	ADP
alces-343	100	7	8	8	NUM
alces-343	100	8	%	%	NOUN
alces-343	100	9	was	be	AUX
alces-343	100	10	included	include	VERB
alces-343	100	11	,	,	PUNCT
alces-343	100	12	the	the	DET
alces-343	100	13	population	population	NOUN
alces-343	100	14	reached	reach	VERB
alces-343	100	15	0.5	0.5	NUM
alces-343	100	16	k	k	NOUN
alces-343	100	17	after	after	ADP
alces-343	100	18	28	28	NUM
alces-343	100	19	years	year	NOUN
alces-343	100	20	and	and	CCONJ
alces-343	100	21	did	do	AUX
alces-343	100	22	not	not	PART
alces-343	100	23	exceed	exceed	VERB
alces-343	100	24	112	112	NUM
alces-343	100	25	caribou	caribou	NOUN
alces-343	100	26	after	after	ADP
alces-343	100	27	100	100	NUM
alces-343	100	28	years	year	NOUN
alces-343	100	29	.	.	PUNCT
alces-343	101	1	in	in	ADP
alces-343	101	2	the	the	DET
alces-343	101	3	absence	absence	NOUN
alces-343	101	4	of	of	ADP
alces-343	101	5	hunting	hunting	NOUN
alces-343	101	6	and	and	CCONJ
alces-343	101	7	predation	predation	NOUN
alces-343	101	8	,	,	PUNCT
alces-343	101	9	the	the	DET
alces-343	101	10	30	30	NUM
alces-343	101	11	initial	initial	ADJ
alces-343	101	12	moose	moose	NOUN
alces-343	101	13	reached	reach	VERB
alces-343	101	14	their	their	PRON
alces-343	101	15	carrying	carry	VERB
alces-343	101	16	capacity	capacity	NOUN
alces-343	101	17	(	(	PUNCT
alces-343	101	18	840	840	NUM
alces-343	101	19	individuals	individual	NOUN
alces-343	101	20	)	)	PUNCT
alces-343	101	21	in	in	ADP
alces-343	101	22	62	62	NUM
alces-343	101	23	years	year	NOUN
alces-343	101	24	.	.	PUNCT
alces-343	102	1	the	the	DET
alces-343	102	2	maximum	maximum	ADJ
alces-343	102	3	sustainable	sustainable	ADJ
alces-343	102	4	yield	yield	NOUN
alces-343	102	5	(	(	PUNCT
alces-343	102	6	52	52	NUM
alces-343	102	7	moose	moose	NOUN
alces-343	102	8	)	)	PUNCT
alces-343	102	9	was	be	AUX
alces-343	102	10	attained	attain	VERB
alces-343	102	11	with	with	ADP
alces-343	102	12	a	a	DET
alces-343	102	13	population	population	NOUN
alces-343	102	14	of	of	ADP
alces-343	102	15	420	420	NUM
alces-343	102	16	moose	moose	NOUN
alces-343	102	17	(	(	PUNCT
alces-343	102	18	optimal	optimal	ADJ
alces-343	102	19	density	density	NOUN
alces-343	102	20	:	:	PUNCT
alces-343	102	21	42	42	NUM
alces-343	102	22	moose	moose	NOUN
alces-343	102	23	per	per	ADP
alces-343	102	24	100	100	NUM
alces-343	102	25	km2	km2	NOUN
alces-343	102	26	)	)	PUNCT
alces-343	102	27	in	in	ADP
alces-343	102	28	13	13	NUM
alces-343	102	29	years	year	NOUN
alces-343	102	30	.	.	PUNCT
alces-343	103	1	including	include	VERB
alces-343	103	2	a	a	DET
alces-343	103	3	natural	natural	ADJ
alces-343	103	4	mortality	mortality	NOUN
alces-343	103	5	of	of	ADP
alces-343	103	6	9	9	NUM
alces-343	103	7	%	%	NOUN
alces-343	103	8	resulted	result	VERB
alces-343	103	9	in	in	ADP
alces-343	103	10	0.5	0.5	NUM
alces-343	103	11	k	k	NOUN
alces-343	103	12	being	be	AUX
alces-343	103	13	reached	reach	VERB
alces-343	103	14	in	in	ADP
alces-343	103	15	25	25	NUM
alces-343	103	16	years	year	NOUN
alces-343	103	17	,	,	PUNCT
alces-343	103	18	but	but	CCONJ
alces-343	103	19	the	the	DET
alces-343	103	20	total	total	ADJ
alces-343	103	21	population	population	NOUN
alces-343	103	22	did	do	AUX
alces-343	103	23	not	not	PART
alces-343	103	24	exceed	exceed	VERB
alces-343	103	25	537	537	NUM
alces-343	103	26	moose	moose	NOUN
alces-343	103	27	after	after	ADP
alces-343	103	28	100	100	NUM
alces-343	103	29	years	year	NOUN
alces-343	103	30	.	.	PUNCT
alces-343	104	1	after	after	ADP
alces-343	104	2	adding	add	VERB
alces-343	104	3	another	another	DET
alces-343	104	4	9	9	NUM
alces-343	104	5	%	%	NOUN
alces-343	104	6	hunting	hunting	NOUN
alces-343	104	7	mortality	mortality	NOUN
alces-343	104	8	,	,	PUNCT
alces-343	104	9	the	the	DET
alces-343	104	10	population	population	NOUN
alces-343	104	11	comprised	comprise	VERB
alces-343	104	12	only	only	ADV
alces-343	104	13	234	234	NUM
alces-343	104	14	moose	moose	NOUN
alces-343	104	15	(	(	PUNCT
alces-343	104	16	0.28	0.28	NUM
alces-343	104	17	k	k	NOUN
alces-343	104	18	)	)	PUNCT
alces-343	104	19	after	after	ADP
alces-343	104	20	100	100	NUM
alces-343	104	21	years	year	NOUN
alces-343	104	22	.	.	PUNCT
alces-343	105	1	according	accord	VERB
alces-343	105	2	to	to	ADP
alces-343	105	3	the	the	DET
alces-343	105	4	predation	predation	NOUN
alces-343	105	5	model	model	NOUN
alces-343	105	6	(	(	PUNCT
alces-343	105	7	messier	messy	ADJ
alces-343	105	8	1994	1994	NUM
alces-343	105	9	)	)	PUNCT
alces-343	105	10	,	,	PUNCT
alces-343	105	11	the	the	DET
alces-343	105	12	wolf	wolf	PROPN
alces-343	105	13	population	population	NOUN
alces-343	105	14	was	be	AUX
alces-343	105	15	sustained	sustain	VERB
alces-343	105	16	by	by	ADP
alces-343	105	17	the	the	DET
alces-343	105	18	moose	moose	NOUN
alces-343	105	19	population	population	NOUN
alces-343	105	20	and	and	CCONJ
alces-343	105	21	also	also	ADV
alces-343	105	22	regulated	regulate	VERB
alces-343	105	23	moose	moose	NOUN
alces-343	105	24	numbers	number	NOUN
alces-343	105	25	.	.	PUNCT
alces-343	106	1	both	both	PRON
alces-343	106	2	wolf	wolf	PROPN
alces-343	106	3	abundance	abundance	PROPN
alces-343	106	4	and	and	CCONJ
alces-343	106	5	predation	predation	NOUN
alces-343	106	6	rate	rate	NOUN
alces-343	106	7	increased	increase	VERB
alces-343	106	8	with	with	ADP
alces-343	106	9	moose	moose	NOUN
alces-343	106	10	abundance	abundance	NOUN
alces-343	106	11	.	.	PUNCT
alces-343	107	1	without	without	ADP
alces-343	107	2	hunting	hunt	VERB
alces-343	107	3	,	,	PUNCT
alces-343	107	4	the	the	DET
alces-343	107	5	moose	moose	NOUN
alces-343	107	6	population	population	NOUN
alces-343	107	7	stabilized	stabilize	VERB
alces-343	107	8	at	at	ADP
alces-343	107	9	650	650	NUM
alces-343	107	10	individuals	individual	NOUN
alces-343	107	11	(	(	PUNCT
alces-343	107	12	65	65	NUM
alces-343	107	13	moose	moose	NOUN
alces-343	107	14	per	per	ADP
alces-343	107	15	100	100	NUM
alces-343	107	16	km2	km2	NUM
alces-343	107	17	)	)	PUNCT
alces-343	107	18	.	.	PUNCT
alces-343	108	1	at	at	ADP
alces-343	108	2	this	this	DET
alces-343	108	3	plateau	plateau	NOUN
alces-343	108	4	,	,	PUNCT
alces-343	108	5	predator	predator	NOUN
alces-343	108	6	density	density	NOUN
alces-343	108	7	reached	reach	VERB
alces-343	108	8	2.58	2.58	NUM
alces-343	108	9	wolves	wolf	NOUN
alces-343	108	10	per	per	ADP
alces-343	108	11	100	100	NUM
alces-343	108	12	km2	km2	NOUN
alces-343	108	13	,	,	PUNCT
alces-343	108	14	as	as	SCONJ
alces-343	108	15	predicted	predict	VERB
alces-343	108	16	by	by	ADP
alces-343	108	17	the	the	DET
alces-343	108	18	numerical	numerical	ADJ
alces-343	108	19	response	response	PROPN
alces-343	108	20	equation	equation	NOUN
alces-343	108	21	of	of	ADP
alces-343	108	22	messier	messy	ADJ
alces-343	108	23	(	(	PUNCT
alces-343	108	24	1994	1994	NUM
alces-343	108	25	)	)	PUNCT
alces-343	108	26	.	.	PUNCT
alces-343	109	1	impact	impact	NOUN
alces-343	109	2	of	of	ADP
alces-343	109	3	moose	moose	NOUN
alces-343	109	4	and	and	CCONJ
alces-343	109	5	wolves	wolf	NOUN
alces-343	109	6	on	on	ADP
alces-343	109	7	caribou	caribou	NOUN
alces-343	109	8	–	–	PUNCT
alces-343	109	9	courtois	courtois	NOUN
alces-343	109	10	and	and	CCONJ
alces-343	109	11	ouellet	ouellet	PROPN
alces-343	109	12	alces	alces	PROPN
alces-343	109	13	vol	vol	NOUN
alces-343	109	14	.	.	PROPN
alces-343	109	15	43	43	NUM
alces-343	109	16	,	,	PUNCT
alces-343	109	17	2007	2007	NUM
alces-343	109	18	18	18	NUM
alces-343	109	19	combined	combine	VERB
alces-343	109	20	model	model	NOUN
alces-343	109	21	incorporating	incorporate	VERB
alces-343	109	22	interactions	interaction	NOUN
alces-343	109	23	between	between	ADP
alces-343	109	24	moose	moose	NOUN
alces-343	109	25	,	,	PUNCT
alces-343	109	26	wolves	wolf	NOUN
alces-343	109	27	and	and	CCONJ
alces-343	109	28	caribou	caribou	NOUN
alces-343	109	29	scenario	scenario	NOUN
alces-343	109	30	1	1	NUM
alces-343	109	31	:	:	PUNCT
alces-343	109	32	moderate	moderate	ADJ
alces-343	109	33	moose	moose	NOUN
alces-343	109	34	(	(	PUNCT
alces-343	109	35	9	9	NUM
alces-343	109	36	%	%	NOUN
alces-343	109	37	)	)	PUNCT
alces-343	109	38	and	and	CCONJ
alces-343	109	39	caribou	caribou	NOUN
alces-343	109	40	(	(	PUNCT
alces-343	109	41	8	8	NUM
alces-343	109	42	%	%	NOUN
alces-343	109	43	)	)	PUNCT
alces-343	109	44	hunting	hunting	NOUN
alces-343	109	45	but	but	CCONJ
alces-343	109	46	no	no	DET
alces-343	109	47	wolf	wolf	NOUN
alces-343	109	48	trapping	trap	VERB
alces-343	109	49	.	.	PUNCT
alces-343	109	50	—	—	PUNCT
alces-343	109	51	when	when	SCONJ
alces-343	109	52	moose	moose	NOUN
alces-343	109	53	and	and	CCONJ
alces-343	109	54	caribou	caribou	NOUN
alces-343	109	55	were	be	AUX
alces-343	109	56	exploited	exploit	VERB
alces-343	109	57	at	at	ADP
alces-343	109	58	moderate	moderate	ADJ
alces-343	109	59	rates	rate	NOUN
alces-343	109	60	and	and	CCONJ
alces-343	109	61	in	in	ADP
alces-343	109	62	the	the	DET
alces-343	109	63	absence	absence	NOUN
alces-343	109	64	of	of	ADP
alces-343	109	65	wolf	wolf	NOUN
alces-343	109	66	trapping	trapping	NOUN
alces-343	109	67	,	,	PUNCT
alces-343	109	68	moose	moose	NOUN
alces-343	109	69	abundance	abundance	NOUN
alces-343	109	70	increased	increase	VERB
alces-343	109	71	from	from	ADP
alces-343	109	72	30	30	NUM
alces-343	109	73	to	to	ADP
alces-343	109	74	110	110	NUM
alces-343	109	75	individuals	individual	NOUN
alces-343	109	76	in	in	ADP
alces-343	109	77	55	55	NUM
alces-343	109	78	years	year	NOUN
alces-343	109	79	and	and	CCONJ
alces-343	109	80	then	then	ADV
alces-343	109	81	remained	remain	VERB
alces-343	109	82	stable	stable	ADJ
alces-343	109	83	(	(	PUNCT
alces-343	109	84	fig	fig	NOUN
alces-343	109	85	.	.	NOUN
alces-343	110	1	1	1	NUM
alces-343	110	2	)	)	PUNCT
alces-343	110	3	.	.	PUNCT
alces-343	111	1	simultaneously	simultaneously	ADV
alces-343	111	2	,	,	PUNCT
alces-343	111	3	wolves	wolf	NOUN
alces-343	111	4	increased	increase	VERB
alces-343	111	5	from	from	ADP
alces-343	111	6	0	0	NUM
alces-343	111	7	at	at	ADP
alces-343	111	8	the	the	DET
alces-343	111	9	start	start	NOUN
alces-343	111	10	of	of	ADP
alces-343	111	11	the	the	DET
alces-343	111	12	run	run	NOUN
alces-343	111	13	to	to	ADP
alces-343	111	14	5	5	NUM
alces-343	111	15	individuals	individual	NOUN
alces-343	111	16	after	after	ADP
alces-343	111	17	30	30	NUM
alces-343	111	18	years	year	NOUN
alces-343	111	19	only	only	ADV
alces-343	111	20	to	to	PART
alces-343	111	21	stabilize	stabilize	VERB
alces-343	111	22	afterwards	afterwards	ADV
alces-343	111	23	.	.	PUNCT
alces-343	112	1	moose	moose	PROPN
alces-343	112	2	predation	predation	NOUN
alces-343	112	3	rate	rate	NOUN
alces-343	112	4	reached	reach	VERB
alces-343	112	5	8	8	NUM
alces-343	112	6	%	%	NOUN
alces-343	112	7	per	per	ADP
alces-343	112	8	year	year	NOUN
alces-343	112	9	,	,	PUNCT
alces-343	112	10	which	which	PRON
alces-343	112	11	stabilized	stabilize	VERB
alces-343	112	12	the	the	DET
alces-343	112	13	moose	moose	ADJ
alces-343	112	14	-	-	PUNCT
alces-343	112	15	wolf	wolf	NOUN
alces-343	112	16	system	system	NOUN
alces-343	112	17	.	.	PUNCT
alces-343	113	1	caribou	caribou	PROPN
alces-343	113	2	abundance	abundance	PROPN
alces-343	113	3	increased	increase	VERB
alces-343	113	4	from	from	ADP
alces-343	113	5	16	16	NUM
alces-343	113	6	to	to	PART
alces-343	113	7	40	40	NUM
alces-343	113	8	individuals	individual	NOUN
alces-343	113	9	in	in	ADP
alces-343	113	10	14	14	NUM
alces-343	113	11	years	year	NOUN
alces-343	113	12	and	and	CCONJ
alces-343	113	13	then	then	ADV
alces-343	113	14	gradually	gradually	ADV
alces-343	113	15	declined	decline	VERB
alces-343	113	16	to	to	ADP
alces-343	113	17	quasi	quasi	NOUN
alces-343	113	18	-	-	NOUN
alces-343	113	19	extirpation	extirpation	NOUN
alces-343	113	20	in	in	ADP
alces-343	113	21	100	100	NUM
alces-343	113	22	years	year	NOUN
alces-343	113	23	(	(	PUNCT
alces-343	113	24	2	2	NUM
alces-343	113	25	caribou	caribou	NOUN
alces-343	113	26	remaining	remain	VERB
alces-343	113	27	)	)	PUNCT
alces-343	113	28	.	.	PUNCT
alces-343	114	1	the	the	DET
alces-343	114	2	caribou	caribou	NOUN
alces-343	114	3	decline	decline	NOUN
alces-343	114	4	initiated	initiate	VERB
alces-343	114	5	when	when	SCONJ
alces-343	114	6	moose	moose	NOUN
alces-343	114	7	and	and	CCONJ
alces-343	114	8	wolf	wolf	PROPN
alces-343	114	9	reached	reach	VERB
alces-343	114	10	76	76	NUM
alces-343	114	11	and	and	CCONJ
alces-343	114	12	32	32	NUM
alces-343	114	13	individuals	individual	NOUN
alces-343	114	14	(	(	PUNCT
alces-343	114	15	7.6	7.6	NUM
alces-343	114	16	and	and	CCONJ
alces-343	114	17	0.32	0.32	NUM
alces-343	114	18	individuals	individual	NOUN
alces-343	114	19	per	per	ADP
alces-343	114	20	100	100	NUM
alces-343	114	21	km2	km2	NUM
alces-343	114	22	)	)	PUNCT
alces-343	114	23	,	,	PUNCT
alces-343	114	24	respectively	respectively	ADV
alces-343	114	25	.	.	PUNCT
alces-343	115	1	scenario	scenario	NOUN
alces-343	115	2	2	2	NUM
alces-343	115	3	:	:	PUNCT
alces-343	115	4	ban	ban	NOUN
alces-343	115	5	on	on	ADP
alces-343	115	6	caribou	caribou	NOUN
alces-343	115	7	hunting	hunting	NOUN
alces-343	115	8	with	with	ADP
alces-343	115	9	9	9	NUM
alces-343	115	10	%	%	NOUN
alces-343	115	11	moose	moose	NOUN
alces-343	115	12	hunting	hunting	NOUN
alces-343	115	13	and	and	CCONJ
alces-343	115	14	no	no	DET
alces-343	115	15	wolf	wolf	NOUN
alces-343	115	16	trapping	trapping	NOUN
alces-343	115	17	.	.	PUNCT
alces-343	115	18	—	—	PUNCT
alces-343	115	19	the	the	DET
alces-343	115	20	moose	moose	NOUN
alces-343	115	21	and	and	CCONJ
alces-343	115	22	wolf	wolf	PROPN
alces-343	115	23	population	population	NOUN
alces-343	115	24	trajectories	trajectory	NOUN
alces-343	115	25	,	,	PUNCT
alces-343	115	26	since	since	SCONJ
alces-343	115	27	the	the	DET
alces-343	115	28	relevant	relevant	ADJ
alces-343	115	29	population	population	NOUN
alces-343	115	30	parameters	parameter	NOUN
alces-343	115	31	remained	remain	VERB
alces-343	115	32	the	the	DET
alces-343	115	33	same	same	ADJ
alces-343	115	34	(	(	PUNCT
alces-343	115	35	fig	fig	NOUN
alces-343	115	36	.	.	PUNCT
alces-343	115	37	2	2	NUM
alces-343	115	38	)	)	PUNCT
alces-343	115	39	.	.	PUNCT
alces-343	116	1	the	the	DET
alces-343	116	2	caribou	caribou	NOUN
alces-343	116	3	hunting	hunt	VERB
alces-343	116	4	ban	ban	NOUN
alces-343	116	5	allowed	allow	VERB
alces-343	116	6	the	the	DET
alces-343	116	7	caribou	caribou	NOUN
alces-343	116	8	population	population	NOUN
alces-343	116	9	to	to	PART
alces-343	116	10	increase	increase	VERB
alces-343	116	11	and	and	CCONJ
alces-343	116	12	reach	reach	VERB
alces-343	116	13	94	94	NUM
alces-343	116	14	individuals	individual	NOUN
alces-343	116	15	after	after	ADP
alces-343	116	16	22	22	NUM
alces-343	116	17	years	year	NOUN
alces-343	116	18	;	;	PUNCT
alces-343	116	19	however	however	ADV
alces-343	116	20	,	,	PUNCT
alces-343	116	21	despite	despite	SCONJ
alces-343	116	22	this	this	DET
alces-343	116	23	increase	increase	NOUN
alces-343	116	24	,	,	PUNCT
alces-343	116	25	the	the	DET
alces-343	116	26	caribou	caribou	NOUN
alces-343	116	27	population	population	NOUN
alces-343	116	28	started	start	VERB
alces-343	116	29	to	to	PART
alces-343	116	30	decline	decline	VERB
alces-343	116	31	as	as	ADV
alces-343	116	32	soon	soon	ADV
alces-343	116	33	as	as	SCONJ
alces-343	116	34	moose	moose	NOUN
alces-343	116	35	numbers	number	NOUN
alces-343	116	36	attained	attain	VERB
alces-343	116	37	95	95	NUM
alces-343	116	38	,	,	PUNCT
alces-343	116	39	thus	thus	ADV
alces-343	116	40	supporting	support	VERB
alces-343	116	41	4.5	4.5	NUM
alces-343	116	42	wolves	wolf	NOUN
alces-343	116	43	(	(	PUNCT
alces-343	116	44	9.5	9.5	NUM
alces-343	116	45	moose	moose	NOUN
alces-343	116	46	and	and	CCONJ
alces-343	116	47	0.45	0.45	NUM
alces-343	116	48	wolves	wolf	NOUN
alces-343	116	49	per	per	ADP
alces-343	116	50	100	100	NUM
alces-343	116	51	km2	km2	NUM
alces-343	116	52	)	)	PUNCT
alces-343	116	53	.	.	PUNCT
alces-343	117	1	caribou	caribou	NOUN
alces-343	117	2	were	be	AUX
alces-343	117	3	not	not	PART
alces-343	117	4	extirpated	extirpate	VERB
alces-343	117	5	and	and	CCONJ
alces-343	117	6	the	the	DET
alces-343	117	7	population	population	NOUN
alces-343	117	8	stabilized	stabilize	VERB
alces-343	117	9	at	at	ADP
alces-343	117	10	approximately	approximately	ADV
alces-343	117	11	64	64	NUM
alces-343	117	12	individuals	individual	NOUN
alces-343	117	13	(	(	PUNCT
alces-343	117	14	6.4	6.4	NUM
alces-343	117	15	per	per	ADP
alces-343	117	16	100	100	NUM
alces-343	117	17	km2	km2	NUM
alces-343	117	18	)	)	PUNCT
alces-343	117	19	.	.	PUNCT
alces-343	118	1	scenario	scenario	NOUN
alces-343	118	2	3	3	NUM
alces-343	118	3	:	:	PUNCT
alces-343	118	4	intensive	intensive	ADJ
alces-343	118	5	wolf	wolf	NOUN
alces-343	118	6	trapping	trap	VERB
alces-343	118	7	(	(	PUNCT
alces-343	118	8	30	30	NUM
alces-343	118	9	%	%	NOUN
alces-343	118	10	)	)	PUNCT
alces-343	118	11	with	with	ADP
alces-343	118	12	9	9	NUM
alces-343	118	13	%	%	NOUN
alces-343	118	14	moose	moose	NOUN
alces-343	118	15	and	and	CCONJ
alces-343	118	16	no	no	DET
alces-343	118	17	caribou	caribou	NOUN
alces-343	118	18	hunting	hunting	NOUN
alces-343	118	19	.	.	PUNCT
alces-343	118	20	—	—	PUNCT
alces-343	118	21	introducing	introduce	VERB
alces-343	118	22	intensive	intensive	ADJ
alces-343	118	23	trapping	trapping	NOUN
alces-343	118	24	of	of	ADP
alces-343	118	25	wolves	wolf	NOUN
alces-343	118	26	(	(	PUNCT
alces-343	118	27	30	30	NUM
alces-343	118	28	%	%	NOUN
alces-343	118	29	)	)	PUNCT
alces-343	118	30	to	to	ADP
alces-343	118	31	parameters	parameter	NOUN
alces-343	118	32	of	of	ADP
alces-343	118	33	the	the	DET
alces-343	118	34	previous	previous	ADJ
alces-343	118	35	simulation	simulation	NOUN
alces-343	118	36	did	do	AUX
alces-343	118	37	not	not	PART
alces-343	118	38	markedly	markedly	ADV
alces-343	118	39	improve	improve	VERB
alces-343	118	40	the	the	DET
alces-343	118	41	situation	situation	NOUN
alces-343	118	42	for	for	ADP
alces-343	118	43	caribou	caribou	NOUN
alces-343	118	44	.	.	PUNCT
alces-343	119	1	wolf	wolf	PROPN
alces-343	119	2	abundance	abundance	PROPN
alces-343	119	3	increased	increase	VERB
alces-343	119	4	steadily	steadily	ADV
alces-343	119	5	despite	despite	SCONJ
alces-343	119	6	exploitation	exploitation	NOUN
alces-343	119	7	,	,	PUNCT
alces-343	119	8	in	in	ADP
alces-343	119	9	parallel	parallel	NOUN
alces-343	119	10	with	with	ADP
alces-343	119	11	the	the	DET
alces-343	119	12	increase	increase	NOUN
alces-343	119	13	in	in	ADP
alces-343	119	14	moose	moose	NOUN
alces-343	119	15	abundance	abundance	NOUN
alces-343	119	16	(	(	PUNCT
alces-343	119	17	fig	fig	NOUN
alces-343	119	18	.	.	PUNCT
alces-343	120	1	3	3	NUM
alces-343	120	2	)	)	PUNCT
alces-343	120	3	.	.	PUNCT
alces-343	121	1	abundance	abundance	NOUN
alces-343	121	2	after	after	ADP
alces-343	121	3	100	100	NUM
alces-343	121	4	years	year	NOUN
alces-343	121	5	was	be	AUX
alces-343	121	6	5	5	NUM
alces-343	121	7	wolves	wolf	NOUN
alces-343	121	8	and	and	CCONJ
alces-343	121	9	141	141	NUM
alces-343	121	10	moose	moose	NOUN
alces-343	121	11	(	(	PUNCT
alces-343	121	12	0.5	0.5	NUM
alces-343	121	13	and	and	CCONJ
alces-343	121	14	14.7	14.7	NUM
alces-343	121	15	per	per	ADP
alces-343	121	16	100	100	NUM
alces-343	121	17	km2	km2	NOUN
alces-343	121	18	,	,	PUNCT
alces-343	121	19	respectively	respectively	ADV
alces-343	121	20	)	)	PUNCT
alces-343	121	21	.	.	PUNCT
alces-343	122	1	the	the	DET
alces-343	122	2	caribou	caribou	PROPN
alces-343	122	3	population	population	NOUN
alces-343	122	4	reached	reach	VERB
alces-343	122	5	109	109	NUM
alces-343	122	6	individuals	individual	NOUN
alces-343	122	7	after	after	ADP
alces-343	122	8	23	23	NUM
alces-343	122	9	years	year	NOUN
alces-343	122	10	,	,	PUNCT
alces-343	122	11	but	but	CCONJ
alces-343	122	12	then	then	ADV
alces-343	122	13	declined	decline	VERB
alces-343	122	14	and	and	CCONJ
alces-343	122	15	stabilized	stabilize	VERB
alces-343	122	16	at	at	ADP
alces-343	122	17	(	(	PUNCT
alces-343	122	18	75	75	NUM
alces-343	122	19	individuals	individual	NOUN
alces-343	122	20	7.5	7.5	NUM
alces-343	122	21	per	per	ADP
alces-343	122	22	100	100	NUM
alces-343	122	23	km2	km2	NUM
alces-343	122	24	)	)	PUNCT
alces-343	122	25	.	.	PUNCT
alces-343	123	1	scenario	scenario	NOUN
alces-343	123	2	4	4	NUM
alces-343	123	3	:	:	PUNCT
alces-343	123	4	increased	increase	VERB
alces-343	123	5	moose	moose	NOUN
alces-343	123	6	hunting	hunting	NOUN
alces-343	123	7	(	(	PUNCT
alces-343	123	8	15	15	NUM
alces-343	123	9	%	%	NOUN
alces-343	123	10	)	)	PUNCT
alces-343	123	11	with	with	ADP
alces-343	123	12	no	no	DET
alces-343	123	13	wolf	wolf	NOUN
alces-343	123	14	and	and	CCONJ
alces-343	123	15	caribou	caribou	NOUN
alces-343	123	16	harvesting	harvesting	NOUN
alces-343	123	17	.	.	PUNCT
alces-343	124	1	—	—	PUNCT
alces-343	124	2	intensifying	intensify	VERB
alces-343	124	3	moose	moose	NOUN
alces-343	124	4	harvest	harvest	NOUN
alces-343	124	5	to	to	ADP
alces-343	124	6	15	15	NUM
alces-343	124	7	%	%	NOUN
alces-343	124	8	but	but	CCONJ
alces-343	124	9	discontinuing	discontinue	VERB
alces-343	124	10	wolf	wolf	NOUN
alces-343	124	11	trapping	trapping	NOUN
alces-343	124	12	maintained	maintain	VERB
alces-343	124	13	the	the	DET
alces-343	124	14	moose	moose	NOUN
alces-343	124	15	population	population	NOUN
alces-343	124	16	at	at	ADP
alces-343	124	17	a	a	DET
alces-343	124	18	relatively	relatively	ADV
alces-343	124	19	low	low	ADJ
alces-343	124	20	level	level	NOUN
alces-343	124	21	of	of	ADP
alces-343	124	22	61	61	NUM
alces-343	124	23	individuals	individual	NOUN
alces-343	124	24	(	(	PUNCT
alces-343	124	25	6.1	6.1	NUM
alces-343	124	26	moose	moose	NOUN
alces-343	124	27	per100	per100	PROPN
alces-343	124	28	km2	km2	PROPN
alces-343	124	29	)	)	PUNCT
alces-343	124	30	,	,	PUNCT
alces-343	124	31	which	which	PRON
alces-343	124	32	directly	directly	ADV
alces-343	124	33	reduced	reduce	VERB
alces-343	124	34	the	the	DET
alces-343	124	35	wolf	wolf	ADJ
alces-343	124	36	population	population	NOUN
alces-343	124	37	to	to	ADP
alces-343	124	38	less	less	ADJ
alces-343	124	39	than	than	ADP
alces-343	124	40	2.2	2.2	NUM
alces-343	124	41	individuals	individual	NOUN
alces-343	124	42	(	(	PUNCT
alces-343	124	43	<	<	X
alces-343	124	44	0.22	0.22	NUM
alces-343	124	45	wolves	wolf	NOUN
alces-343	124	46	per	per	ADP
alces-343	124	47	100	100	NUM
alces-343	124	48	km2	km2	NOUN
alces-343	124	49	;	;	PUNCT
alces-343	124	50	fig	fig	NOUN
alces-343	124	51	.	.	PUNCT
alces-343	125	1	4	4	NUM
alces-343	125	2	)	)	PUNCT
alces-343	125	3	.	.	PUNCT
alces-343	126	1	this	this	DET
alces-343	126	2	management	management	NOUN
alces-343	126	3	strategy	strategy	NOUN
alces-343	126	4	was	be	AUX
alces-343	126	5	very	very	ADV
alces-343	126	6	favourable	favourable	ADJ
alces-343	126	7	for	for	ADP
alces-343	126	8	caribou	caribou	PROPN
alces-343	126	9	and	and	CCONJ
alces-343	126	10	the	the	DET
alces-343	126	11	population	population	NOUN
alces-343	126	12	reached	reach	VERB
alces-343	126	13	146	146	NUM
alces-343	126	14	individuals	individual	NOUN
alces-343	126	15	after	after	ADP
alces-343	126	16	30	30	NUM
alces-343	126	17	years	year	NOUN
alces-343	126	18	and	and	CCONJ
alces-343	126	19	then	then	ADV
alces-343	126	20	stabilized	stabilize	VERB
alces-343	126	21	at	at	ADP
alces-343	126	22	141	141	NUM
alces-343	126	23	caribou	caribou	NOUN
alces-343	126	24	(	(	PUNCT
alces-343	126	25	14.1	14.1	NUM
alces-343	126	26	per	per	ADP
alces-343	126	27	100	100	NUM
alces-343	126	28	km2	km2	NUM
alces-343	126	29	)	)	PUNCT
alces-343	126	30	.	.	PUNCT
alces-343	127	1	scenario	scenario	NOUN
alces-343	127	2	5	5	NUM
alces-343	127	3	:	:	PUNCT
alces-343	127	4	stochastic	stochastic	ADJ
alces-343	127	5	environmental	environmental	ADJ
alces-343	127	6	variations	variation	NOUN
alces-343	127	7	.	.	PUNCT
alces-343	127	8	—	—	PUNCT
alces-343	128	1	with	with	ADP
alces-343	128	2	random	random	ADJ
alces-343	128	3	annual	annual	ADJ
alces-343	128	4	variation	variation	NOUN
alces-343	128	5	in	in	ADP
alces-343	128	6	birth	birth	NOUN
alces-343	128	7	rates	rate	NOUN
alces-343	128	8	,	,	PUNCT
alces-343	128	9	each	each	DET
alces-343	128	10	simulation	simulation	NOUN
alces-343	128	11	followed	follow	VERB
alces-343	128	12	a	a	DET
alces-343	128	13	new	new	ADJ
alces-343	128	14	trajectory	trajectory	NOUN
alces-343	128	15	(	(	PUNCT
alces-343	128	16	fig	fig	NOUN
alces-343	128	17	.	.	PUNCT
alces-343	128	18	5	5	NUM
alces-343	128	19	)	)	PUNCT
alces-343	128	20	.	.	PUNCT
alces-343	129	1	stochastic	stochastic	ADJ
alces-343	129	2	variation	variation	NOUN
alces-343	129	3	0	0	NUM
alces-343	129	4	20	20	NUM
alces-343	129	5	40	40	NUM
alces-343	129	6	60	60	NUM
alces-343	129	7	80	80	NUM
alces-343	129	8	100	100	NUM
alces-343	129	9	120	120	NUM
alces-343	129	10	0	0	NUM
alces-343	129	11	10	10	NUM
alces-343	129	12	20	20	NUM
alces-343	129	13	30	30	NUM
alces-343	129	14	40	40	NUM
alces-343	129	15	50	50	NUM
alces-343	129	16	60	60	NUM
alces-343	129	17	70	70	NUM
alces-343	129	18	80	80	NUM
alces-343	129	19	90	90	NUM
alces-343	129	20	100	100	NUM
alces-343	129	21	time	time	NOUN
alces-343	129	22	(	(	PUNCT
alces-343	129	23	years	year	NOUN
alces-343	129	24	)	)	PUNCT
alces-343	129	25	n	n	CCONJ
alces-343	129	26	o	o	NOUN
alces-343	129	27	.i	.i	NOUN
alces-343	130	1	n	n	CCONJ
alces-343	130	2	d	d	NOUN
alces-343	130	3	iv	iv	X
alces-343	131	1	i	i	PROPN
alces-343	131	2	d	d	PROPN
alces-343	131	3	u	u	PROPN
alces-343	131	4	a	a	DET
alces-343	131	5	ls	ls	PROPN
alces-343	131	6	moose	moose	NOUN
alces-343	131	7	caribou	caribou	PROPN
alces-343	131	8	wolf	wolf	PROPN
alces-343	131	9	fig	fig	PROPN
alces-343	131	10	.	.	PUNCT
alces-343	132	1	1	1	NUM
alces-343	132	2	.	.	NUM
alces-343	132	3	moose	moose	NOUN
alces-343	132	4	,	,	PUNCT
alces-343	132	5	wolf	wolf	NOUN
alces-343	132	6	,	,	PUNCT
alces-343	132	7	and	and	CCONJ
alces-343	132	8	caribou	caribou	NOUN
alces-343	132	9	population	population	NOUN
alces-343	132	10	trajectories	trajectorie	VERB
alces-343	132	11	with	with	ADP
alces-343	132	12	moderate	moderate	ADJ
alces-343	132	13	moose	moose	NOUN
alces-343	132	14	(	(	PUNCT
alces-343	132	15	9	9	NUM
alces-343	132	16	%	%	NOUN
alces-343	132	17	per	per	ADP
alces-343	132	18	year	year	NOUN
alces-343	132	19	)	)	PUNCT
alces-343	132	20	and	and	CCONJ
alces-343	132	21	caribou	caribou	NOUN
alces-343	132	22	(	(	PUNCT
alces-343	132	23	8	8	NUM
alces-343	132	24	%	%	NOUN
alces-343	132	25	per	per	ADP
alces-343	132	26	year	year	NOUN
alces-343	132	27	)	)	PUNCT
alces-343	133	1	hunting	hunting	NOUN
alces-343	133	2	but	but	CCONJ
alces-343	133	3	no	no	DET
alces-343	133	4	wolf	wolf	NOUN
alces-343	133	5	trapping	trap	VERB
alces-343	133	6	(	(	PUNCT
alces-343	133	7	scenario	scenario	NOUN
alces-343	133	8	1	1	NUM
alces-343	133	9	)	)	PUNCT
alces-343	133	10	.	.	PUNCT
alces-343	134	1	0	0	NUM
alces-343	135	1	20	20	NUM
alces-343	135	2	40	40	NUM
alces-343	135	3	60	60	NUM
alces-343	135	4	80	80	NUM
alces-343	135	5	100	100	NUM
alces-343	135	6	120	120	NUM
alces-343	135	7	0	0	NUM
alces-343	135	8	10	10	NUM
alces-343	135	9	20	20	NUM
alces-343	135	10	30	30	NUM
alces-343	135	11	40	40	NUM
alces-343	135	12	50	50	NUM
alces-343	135	13	60	60	NUM
alces-343	135	14	70	70	NUM
alces-343	135	15	80	80	NUM
alces-343	135	16	90	90	NUM
alces-343	135	17	100	100	NUM
alces-343	135	18	time	time	NOUN
alces-343	135	19	(	(	PUNCT
alces-343	135	20	years	year	NOUN
alces-343	135	21	)	)	PUNCT
alces-343	135	22	n	n	CCONJ
alces-343	135	23	o	o	NOUN
alces-343	135	24	.i	.i	NOUN
alces-343	136	1	n	n	CCONJ
alces-343	136	2	d	d	NOUN
alces-343	136	3	iv	iv	X
alces-343	137	1	i	i	PROPN
alces-343	137	2	d	d	PROPN
alces-343	137	3	u	u	PROPN
alces-343	137	4	a	a	DET
alces-343	137	5	ls	ls	PROPN
alces-343	137	6	moose	moose	NOUN
alces-343	137	7	caribou	caribou	PROPN
alces-343	137	8	wolf	wolf	PROPN
alces-343	137	9	fig	fig	PROPN
alces-343	137	10	.	.	PUNCT
alces-343	138	1	2	2	NUM
alces-343	138	2	.	.	NUM
alces-343	138	3	moose	moose	NOUN
alces-343	138	4	,	,	PUNCT
alces-343	138	5	wolf	wolf	NOUN
alces-343	138	6	,	,	PUNCT
alces-343	138	7	and	and	CCONJ
alces-343	138	8	caribou	caribou	NOUN
alces-343	138	9	population	population	NOUN
alces-343	138	10	trajectories	trajectorie	VERB
alces-343	138	11	with	with	ADP
alces-343	138	12	moderate	moderate	ADJ
alces-343	138	13	moose	moose	NOUN
alces-343	138	14	hunting	hunting	NOUN
alces-343	138	15	(	(	PUNCT
alces-343	138	16	9	9	NUM
alces-343	138	17	%	%	NOUN
alces-343	138	18	per	per	ADP
alces-343	138	19	year	year	NOUN
alces-343	138	20	)	)	PUNCT
alces-343	139	1	but	but	CCONJ
alces-343	139	2	no	no	DET
alces-343	139	3	caribou	caribou	NOUN
alces-343	139	4	hunting	hunting	NOUN
alces-343	139	5	or	or	CCONJ
alces-343	139	6	wolf	wolf	NOUN
alces-343	139	7	trapping	trap	VERB
alces-343	139	8	(	(	PUNCT
alces-343	139	9	scenario	scenario	NOUN
alces-343	139	10	2	2	NUM
alces-343	139	11	)	)	PUNCT
alces-343	139	12	.	.	PUNCT
alces-343	140	1	alces	alce	NOUN
alces-343	140	2	vol	vol	NOUN
alces-343	140	3	.	.	PROPN
alces-343	141	1	43	43	NUM
alces-343	141	2	,	,	PUNCT
alces-343	141	3	2007	2007	NUM
alces-343	141	4	courtois	courtois	NOUN
alces-343	141	5	and	and	CCONJ
alces-343	141	6	ouellet	ouellet	NOUN
alces-343	141	7	impact	impact	NOUN
alces-343	141	8	of	of	ADP
alces-343	141	9	moose	moose	NOUN
alces-343	141	10	and	and	CCONJ
alces-343	141	11	wolves	wolf	NOUN
alces-343	141	12	on	on	ADP
alces-343	141	13	caribou	caribou	NOUN
alces-343	141	14	19	19	NUM
alces-343	141	15	tions	tion	NOUN
alces-343	141	16	in	in	ADP
alces-343	141	17	both	both	DET
alces-343	141	18	moose	moose	NOUN
alces-343	141	19	and	and	CCONJ
alces-343	141	20	wolf	wolf	NOUN
alces-343	141	21	numbers	number	NOUN
alces-343	141	22	,	,	PUNCT
alces-343	141	23	which	which	PRON
alces-343	141	24	always	always	ADV
alces-343	141	25	remained	remain	VERB
alces-343	141	26	lower	low	ADJ
alces-343	141	27	than	than	ADP
alces-343	141	28	in	in	ADP
alces-343	141	29	previous	previous	ADJ
alces-343	141	30	simulations	simulation	NOUN
alces-343	141	31	.	.	PUNCT
alces-343	142	1	these	these	DET
alces-343	142	2	variations	variation	NOUN
alces-343	142	3	had	have	VERB
alces-343	142	4	a	a	DET
alces-343	142	5	positive	positive	ADJ
alces-343	142	6	impact	impact	NOUN
alces-343	142	7	on	on	ADP
alces-343	142	8	caribou	caribou	NOUN
alces-343	142	9	numbers	number	NOUN
alces-343	142	10	.	.	PUNCT
alces-343	143	1	in	in	ADP
alces-343	143	2	the	the	DET
alces-343	143	3	presence	presence	NOUN
alces-343	143	4	of	of	ADP
alces-343	143	5	moderate	moderate	ADJ
alces-343	143	6	moose	moose	NOUN
alces-343	143	7	harvesting	harvesting	NOUN
alces-343	143	8	(	(	PUNCT
alces-343	143	9	9	9	NUM
alces-343	143	10	%	%	NOUN
alces-343	143	11	)	)	PUNCT
alces-343	144	1	but	but	CCONJ
alces-343	144	2	no	no	DET
alces-343	144	3	wolf	wolf	NOUN
alces-343	144	4	trapping	trapping	NOUN
alces-343	144	5	or	or	CCONJ
alces-343	144	6	caribou	caribou	NOUN
alces-343	144	7	hunting	hunting	NOUN
alces-343	144	8	(	(	PUNCT
alces-343	144	9	previous	previous	ADJ
alces-343	144	10	scenario	scenario	NOUN
alces-343	144	11	2	2	NUM
alces-343	144	12	)	)	PUNCT
alces-343	144	13	,	,	PUNCT
alces-343	144	14	the	the	DET
alces-343	144	15	caribou	caribou	NOUN
alces-343	144	16	population	population	NOUN
alces-343	144	17	reached	reach	VERB
alces-343	144	18	125	125	NUM
alces-343	144	19	-	-	SYM
alces-343	144	20	135	135	NUM
alces-343	144	21	individuals	individual	NOUN
alces-343	144	22	and	and	CCONJ
alces-343	144	23	stabilized	stabilize	VERB
alces-343	144	24	at	at	ADP
alces-343	144	25	this	this	DET
alces-343	144	26	level	level	NOUN
alces-343	144	27	(	(	PUNCT
alces-343	144	28	fig	fig	NOUN
alces-343	144	29	.	.	PUNCT
alces-343	145	1	5a	5a	NUM
alces-343	145	2	)	)	PUNCT
alces-343	145	3	.	.	PUNCT
alces-343	146	1	if	if	SCONJ
alces-343	146	2	caribou	caribou	NOUN
alces-343	146	3	hunting	hunting	NOUN
alces-343	146	4	was	be	AUX
alces-343	146	5	also	also	ADV
alces-343	146	6	introduced	introduce	VERB
alces-343	146	7	into	into	ADP
alces-343	146	8	this	this	DET
alces-343	146	9	model	model	NOUN
alces-343	146	10	(	(	PUNCT
alces-343	146	11	previous	previous	ADJ
alces-343	146	12	scenario	scenario	NOUN
alces-343	146	13	1	1	NUM
alces-343	146	14	)	)	PUNCT
alces-343	146	15	,	,	PUNCT
alces-343	146	16	the	the	DET
alces-343	146	17	caribou	caribou	NOUN
alces-343	146	18	population	population	NOUN
alces-343	146	19	stabilized	stabilize	VERB
alces-343	146	20	at	at	ADP
alces-343	146	21	approximately	approximately	ADV
alces-343	146	22	50	50	NUM
alces-343	146	23	–	–	PUNCT
alces-343	146	24	60	60	NUM
alces-343	146	25	individuals	individual	NOUN
alces-343	146	26	(	(	PUNCT
alces-343	146	27	fig	fig	NOUN
alces-343	146	28	.	.	PUNCT
alces-343	147	1	5b	5b	NUM
alces-343	147	2	)	)	PUNCT
alces-343	147	3	.	.	PUNCT
alces-343	148	1	when	when	SCONJ
alces-343	148	2	environmental	environmental	ADJ
alces-343	148	3	stochasticity	stochasticity	NOUN
alces-343	148	4	was	be	AUX
alces-343	148	5	allowed	allow	VERB
alces-343	148	6	to	to	PART
alces-343	148	7	increase	increase	VERB
alces-343	148	8	the	the	DET
alces-343	148	9	variation	variation	NOUN
alces-343	148	10	in	in	ADP
alces-343	148	11	caribou	caribou	NOUN
alces-343	148	12	birth	birth	NOUN
alces-343	148	13	rate	rate	NOUN
alces-343	148	14	in	in	ADP
alces-343	148	15	addition	addition	NOUN
alces-343	148	16	to	to	ADP
alces-343	148	17	the	the	DET
alces-343	148	18	moose	moose	NOUN
alces-343	148	19	birth	birth	NOUN
alces-343	148	20	rate	rate	NOUN
alces-343	148	21	,	,	PUNCT
alces-343	148	22	the	the	DET
alces-343	148	23	caribou	caribou	NOUN
alces-343	148	24	population	population	NOUN
alces-343	148	25	declined	decline	VERB
alces-343	148	26	to	to	ADP
alces-343	148	27	approximately	approximately	ADV
alces-343	148	28	35	35	NUM
alces-343	148	29	individuals	individual	NOUN
alces-343	148	30	(	(	PUNCT
alces-343	148	31	fig	fig	NOUN
alces-343	148	32	.	.	PUNCT
alces-343	148	33	5c	5c	NUM
alces-343	148	34	)	)	PUNCT
alces-343	148	35	.	.	PUNCT
alces-343	149	1	equilibrium	equilibrium	NOUN
alces-343	149	2	points	point	NOUN
alces-343	149	3	of	of	ADP
alces-343	149	4	the	the	DET
alces-343	149	5	combined	combine	VERB
alces-343	149	6	models	model	NOUN
alces-343	149	7	theoretical	theoretical	ADJ
alces-343	149	8	equilibrium	equilibrium	NOUN
alces-343	149	9	points	point	NOUN
alces-343	149	10	of	of	ADP
alces-343	149	11	the	the	DET
alces-343	149	12	simple	simple	ADJ
alces-343	149	13	moose	moose	NOUN
alces-343	149	14	-	-	PUNCT
alces-343	149	15	wolf	wolf	NOUN
alces-343	149	16	model	model	NOUN
alces-343	149	17	were	be	AUX
alces-343	149	18	65.0	65.0	NUM
alces-343	149	19	moose	moose	NOUN
alces-343	149	20	and	and	CCONJ
alces-343	149	21	2.58	2.58	NUM
alces-343	149	22	wolves	wolf	NOUN
alces-343	149	23	per	per	ADP
alces-343	149	24	100	100	NUM
alces-343	149	25	km2	km2	NOUN
alces-343	149	26	.	.	PUNCT
alces-343	150	1	in	in	ADP
alces-343	150	2	the	the	DET
alces-343	150	3	moosewolf	moosewolf	PROPN
alces-343	150	4	-	-	PUNCT
alces-343	150	5	caribou	caribou	PROPN
alces-343	150	6	model	model	NOUN
alces-343	150	7	,	,	PUNCT
alces-343	150	8	equilibrium	equilibrium	NOUN
alces-343	150	9	points	point	NOUN
alces-343	150	10	were	be	AUX
alces-343	150	11	much	much	ADV
alces-343	150	12	lower	low	ADJ
alces-343	150	13	than	than	ADP
alces-343	150	14	these	these	DET
alces-343	150	15	values	value	NOUN
alces-343	150	16	.	.	PUNCT
alces-343	151	1	in	in	ADP
alces-343	151	2	simulations	simulation	NOUN
alces-343	151	3	without	without	ADP
alces-343	151	4	stochastic	stochastic	ADJ
alces-343	151	5	environmental	environmental	ADJ
alces-343	151	6	variation	variation	NOUN
alces-343	151	7	,	,	PUNCT
alces-343	151	8	equilibrium	equilibrium	NOUN
alces-343	151	9	points	point	NOUN
alces-343	151	10	were	be	AUX
alces-343	151	11	approximately	approximately	ADV
alces-343	151	12	7	7	NUM
alces-343	151	13	–	–	PUNCT
alces-343	151	14	10	10	NUM
alces-343	151	15	0	0	NUM
alces-343	151	16	20	20	NUM
alces-343	151	17	40	40	NUM
alces-343	151	18	60	60	NUM
alces-343	151	19	80	80	NUM
alces-343	151	20	100	100	NUM
alces-343	151	21	120	120	NUM
alces-343	151	22	140	140	NUM
alces-343	151	23	160	160	NUM
alces-343	151	24	0	0	NUM
alces-343	151	25	10	10	NUM
alces-343	151	26	20	20	NUM
alces-343	151	27	30	30	NUM
alces-343	151	28	40	40	NUM
alces-343	151	29	50	50	NUM
alces-343	151	30	60	60	NUM
alces-343	151	31	70	70	NUM
alces-343	151	32	80	80	NUM
alces-343	151	33	90	90	NUM
alces-343	151	34	100	100	NUM
alces-343	151	35	time	time	NOUN
alces-343	151	36	(	(	PUNCT
alces-343	151	37	years	year	NOUN
alces-343	151	38	)	)	PUNCT
alces-343	151	39	n	n	CCONJ
alces-343	151	40	o	o	NOUN
alces-343	151	41	.i	.i	NOUN
alces-343	152	1	n	n	CCONJ
alces-343	152	2	d	d	NOUN
alces-343	152	3	iv	iv	X
alces-343	153	1	i	i	PROPN
alces-343	153	2	d	d	PROPN
alces-343	153	3	u	u	PROPN
alces-343	153	4	a	a	DET
alces-343	153	5	ls	ls	PROPN
alces-343	153	6	moose	moose	NOUN
alces-343	153	7	caribou	caribou	PROPN
alces-343	153	8	wolf	wolf	PROPN
alces-343	153	9	fig	fig	PROPN
alces-343	153	10	.	.	PUNCT
alces-343	154	1	3	3	X
alces-343	154	2	.	.	NUM
alces-343	154	3	moose	moose	NOUN
alces-343	154	4	,	,	PUNCT
alces-343	154	5	wolf	wolf	NOUN
alces-343	154	6	,	,	PUNCT
alces-343	154	7	and	and	CCONJ
alces-343	154	8	caribou	caribou	NOUN
alces-343	154	9	population	population	NOUN
alces-343	154	10	trajectories	trajectorie	VERB
alces-343	154	11	with	with	ADP
alces-343	154	12	moderate	moderate	ADJ
alces-343	154	13	moose	moose	NOUN
alces-343	154	14	hunting	hunting	NOUN
alces-343	154	15	(	(	PUNCT
alces-343	154	16	9	9	NUM
alces-343	154	17	%	%	NOUN
alces-343	154	18	per	per	ADP
alces-343	154	19	year	year	NOUN
alces-343	154	20	)	)	PUNCT
alces-343	154	21	and	and	CCONJ
alces-343	154	22	intensive	intensive	ADJ
alces-343	154	23	wolf	wolf	NOUN
alces-343	154	24	trapping	trap	VERB
alces-343	154	25	(	(	PUNCT
alces-343	154	26	30	30	NUM
alces-343	154	27	%	%	NOUN
alces-343	154	28	per	per	ADP
alces-343	154	29	year	year	NOUN
alces-343	154	30	)	)	PUNCT
alces-343	155	1	but	but	CCONJ
alces-343	155	2	no	no	DET
alces-343	155	3	caribou	caribou	NOUN
alces-343	155	4	hunting	hunting	NOUN
alces-343	155	5	(	(	PUNCT
alces-343	155	6	scenario	scenario	NOUN
alces-343	155	7	3	3	NUM
alces-343	155	8	)	)	PUNCT
alces-343	155	9	.	.	PUNCT
alces-343	156	1	0	0	NUM
alces-343	157	1	20	20	NUM
alces-343	157	2	40	40	NUM
alces-343	157	3	60	60	NUM
alces-343	157	4	80	80	NUM
alces-343	157	5	100	100	NUM
alces-343	157	6	120	120	NUM
alces-343	157	7	140	140	NUM
alces-343	157	8	160	160	NUM
alces-343	157	9	0	0	NUM
alces-343	157	10	10	10	NUM
alces-343	157	11	20	20	NUM
alces-343	157	12	30	30	NUM
alces-343	157	13	40	40	NUM
alces-343	157	14	50	50	NUM
alces-343	157	15	60	60	NUM
alces-343	157	16	70	70	NUM
alces-343	157	17	80	80	NUM
alces-343	157	18	90	90	NUM
alces-343	157	19	100	100	NUM
alces-343	157	20	time	time	NOUN
alces-343	157	21	(	(	PUNCT
alces-343	157	22	years	year	NOUN
alces-343	157	23	)	)	PUNCT
alces-343	157	24	n	n	CCONJ
alces-343	157	25	o	o	NOUN
alces-343	157	26	.i	.i	NOUN
alces-343	158	1	n	n	CCONJ
alces-343	158	2	d	d	NOUN
alces-343	158	3	iv	iv	X
alces-343	159	1	i	i	PROPN
alces-343	159	2	d	d	PROPN
alces-343	159	3	u	u	PROPN
alces-343	159	4	a	a	DET
alces-343	159	5	ls	ls	PROPN
alces-343	159	6	moose	moose	NOUN
alces-343	159	7	caribou	caribou	PROPN
alces-343	159	8	wolf	wolf	PROPN
alces-343	159	9	fig	fig	PROPN
alces-343	159	10	.	.	PUNCT
alces-343	160	1	4	4	NUM
alces-343	160	2	.	.	NUM
alces-343	160	3	moose	moose	NOUN
alces-343	160	4	,	,	PUNCT
alces-343	160	5	wolf	wolf	NOUN
alces-343	160	6	,	,	PUNCT
alces-343	160	7	and	and	CCONJ
alces-343	160	8	caribou	caribou	NOUN
alces-343	160	9	population	population	NOUN
alces-343	160	10	trajectories	trajectorie	VERB
alces-343	160	11	with	with	ADP
alces-343	160	12	moose	moose	NOUN
alces-343	160	13	hunting	hunting	NOUN
alces-343	160	14	set	set	VERB
alces-343	160	15	at	at	ADP
alces-343	160	16	15	15	NUM
alces-343	160	17	%	%	NOUN
alces-343	160	18	per	per	ADP
alces-343	160	19	year	year	NOUN
alces-343	160	20	but	but	CCONJ
alces-343	160	21	no	no	DET
alces-343	160	22	caribou	caribou	NOUN
alces-343	160	23	hunting	hunting	NOUN
alces-343	160	24	or	or	CCONJ
alces-343	160	25	wolf	wolf	NOUN
alces-343	160	26	trapping	trap	VERB
alces-343	160	27	(	(	PUNCT
alces-343	160	28	scenario	scenario	NOUN
alces-343	160	29	4	4	NUM
alces-343	160	30	)	)	PUNCT
alces-343	160	31	.	.	PUNCT
alces-343	161	1	a	a	PRON
alces-343	161	2	)	)	PUNCT
alces-343	161	3	0	0	NUM
alces-343	161	4	20	20	NUM
alces-343	161	5	40	40	NUM
alces-343	161	6	60	60	NUM
alces-343	161	7	80	80	NUM
alces-343	161	8	100	100	NUM
alces-343	161	9	120	120	NUM
alces-343	161	10	140	140	NUM
alces-343	161	11	160	160	NUM
alces-343	161	12	0	0	NUM
alces-343	161	13	20	20	NUM
alces-343	161	14	40	40	NUM
alces-343	161	15	60	60	NUM
alces-343	161	16	80	80	NUM
alces-343	161	17	100	100	NUM
alces-343	161	18	time	time	NOUN
alces-343	161	19	(	(	PUNCT
alces-343	161	20	years	year	NOUN
alces-343	161	21	)	)	PUNCT
alces-343	161	22	n	n	CCONJ
alces-343	161	23	o	o	NOUN
alces-343	161	24	.i	.i	NOUN
alces-343	162	1	n	n	CCONJ
alces-343	162	2	d	d	NOUN
alces-343	162	3	iv	iv	X
alces-343	163	1	i	i	PROPN
alces-343	163	2	d	d	PROPN
alces-343	163	3	u	u	PROPN
alces-343	163	4	a	a	DET
alces-343	163	5	ls	ls	ADJ
alces-343	163	6	_	_	PUNCT
alces-343	163	7	b	b	X
alces-343	163	8	)	)	PUNCT
alces-343	163	9	0	0	NUM
alces-343	163	10	20	20	NUM
alces-343	163	11	40	40	NUM
alces-343	163	12	60	60	NUM
alces-343	163	13	80	80	NUM
alces-343	163	14	100	100	NUM
alces-343	163	15	120	120	NUM
alces-343	163	16	140	140	NUM
alces-343	163	17	0	0	NUM
alces-343	163	18	20	20	NUM
alces-343	163	19	40	40	NUM
alces-343	163	20	60	60	NUM
alces-343	163	21	80	80	NUM
alces-343	163	22	100	100	NUM
alces-343	163	23	time	time	NOUN
alces-343	163	24	(	(	PUNCT
alces-343	163	25	years	year	NOUN
alces-343	163	26	)	)	PUNCT
alces-343	163	27	n	n	CCONJ
alces-343	163	28	o	o	NOUN
alces-343	163	29	.i	.i	NOUN
alces-343	164	1	n	n	CCONJ
alces-343	164	2	d	d	NOUN
alces-343	164	3	iv	iv	X
alces-343	165	1	i	i	PROPN
alces-343	165	2	d	d	PROPN
alces-343	165	3	u	u	PROPN
alces-343	165	4	a	a	DET
alces-343	165	5	ls	ls	ADJ
alces-343	165	6	_	_	PUNCT
alces-343	166	1	c	c	NOUN
alces-343	166	2	)	)	PUNCT
alces-343	166	3	0	0	NUM
alces-343	166	4	20	20	NUM
alces-343	166	5	40	40	NUM
alces-343	166	6	60	60	NUM
alces-343	166	7	80	80	NUM
alces-343	166	8	100	100	NUM
alces-343	166	9	120	120	NUM
alces-343	166	10	140	140	NUM
alces-343	166	11	0	0	NUM
alces-343	166	12	20	20	NUM
alces-343	166	13	40	40	NUM
alces-343	166	14	60	60	NUM
alces-343	166	15	80	80	NUM
alces-343	166	16	100	100	NUM
alces-343	166	17	time	time	NOUN
alces-343	166	18	(	(	PUNCT
alces-343	166	19	years	year	NOUN
alces-343	166	20	)	)	PUNCT
alces-343	166	21	n	n	CCONJ
alces-343	166	22	o	o	NOUN
alces-343	166	23	.i	.i	NOUN
alces-343	167	1	n	n	CCONJ
alces-343	167	2	d	d	NOUN
alces-343	167	3	iv	iv	X
alces-343	168	1	i	i	PROPN
alces-343	168	2	d	d	PROPN
alces-343	168	3	u	u	PROPN
alces-343	168	4	a	a	DET
alces-343	168	5	ls	ls	PROPN
alces-343	168	6	_	_	PROPN
alces-343	168	7	moose	moose	PROPN
alces-343	168	8	caribou	caribou	PROPN
alces-343	168	9	wolf	wolf	PROPN
alces-343	168	10	fig	fig	PROPN
alces-343	168	11	.	.	PUNCT
alces-343	169	1	5	5	NUM
alces-343	169	2	.	.	PUNCT
alces-343	169	3	typical	typical	ADJ
alces-343	169	4	moose	moose	NOUN
alces-343	169	5	,	,	PUNCT
alces-343	169	6	wolf	wolf	NOUN
alces-343	169	7	,	,	PUNCT
alces-343	169	8	and	and	CCONJ
alces-343	169	9	caribou	caribou	NOUN
alces-343	169	10	population	population	NOUN
alces-343	169	11	trajectories	trajectory	NOUN
alces-343	169	12	according	accord	VERB
alces-343	169	13	to	to	ADP
alces-343	169	14	different	different	ADJ
alces-343	169	15	management	management	NOUN
alces-343	169	16	strategies	strategy	NOUN
alces-343	169	17	and	and	CCONJ
alces-343	169	18	by	by	ADP
alces-343	169	19	including	include	VERB
alces-343	169	20	random	random	ADJ
alces-343	169	21	variations	variation	NOUN
alces-343	169	22	in	in	ADP
alces-343	169	23	moose	moose	NOUN
alces-343	169	24	(	(	PUNCT
alces-343	169	25	all	all	DET
alces-343	169	26	simulations	simulation	NOUN
alces-343	169	27	)	)	PUNCT
alces-343	169	28	and	and	CCONJ
alces-343	169	29	caribou	caribou	NOUN
alces-343	169	30	birth	birth	NOUN
alces-343	169	31	rates	rate	NOUN
alces-343	169	32	(	(	PUNCT
alces-343	169	33	last	last	ADJ
alces-343	169	34	simulation	simulation	NOUN
alces-343	169	35	)	)	PUNCT
alces-343	169	36	(	(	PUNCT
alces-343	169	37	scenario	scenario	NOUN
alces-343	169	38	5	5	NUM
alces-343	169	39	)	)	PUNCT
alces-343	169	40	.	.	PUNCT
alces-343	170	1	(	(	PUNCT
alces-343	170	2	a	a	X
alces-343	170	3	)	)	PUNCT
alces-343	170	4	moderate	moderate	ADJ
alces-343	170	5	moose	moose	NOUN
alces-343	170	6	hunting	hunting	NOUN
alces-343	170	7	(	(	PUNCT
alces-343	170	8	9	9	NUM
alces-343	170	9	%	%	NOUN
alces-343	170	10	per	per	ADP
alces-343	170	11	year	year	NOUN
alces-343	170	12	)	)	PUNCT
alces-343	170	13	,	,	PUNCT
alces-343	170	14	with	with	ADP
alces-343	170	15	no	no	DET
alces-343	170	16	caribou	caribou	NOUN
alces-343	170	17	hunting	hunting	NOUN
alces-343	170	18	or	or	CCONJ
alces-343	170	19	wolf	wolf	NOUN
alces-343	170	20	trapping	trapping	NOUN
alces-343	170	21	;	;	PUNCT
alces-343	170	22	moose	moose	NOUN
alces-343	170	23	birth	birth	NOUN
alces-343	170	24	rate	rate	NOUN
alces-343	170	25	varying	vary	VERB
alces-343	170	26	up	up	ADP
alces-343	170	27	to	to	PART
alces-343	170	28	40	40	NUM
alces-343	170	29	%	%	NOUN
alces-343	170	30	a	a	DET
alces-343	170	31	year	year	NOUN
alces-343	170	32	;	;	PUNCT
alces-343	170	33	(	(	PUNCT
alces-343	170	34	b	b	X
alces-343	170	35	)	)	PUNCT
alces-343	170	36	moderate	moderate	ADJ
alces-343	170	37	moose	moose	NOUN
alces-343	170	38	(	(	PUNCT
alces-343	170	39	9	9	NUM
alces-343	170	40	%	%	NOUN
alces-343	170	41	per	per	ADP
alces-343	170	42	year	year	NOUN
alces-343	170	43	)	)	PUNCT
alces-343	170	44	and	and	CCONJ
alces-343	170	45	caribou	caribou	NOUN
alces-343	170	46	(	(	PUNCT
alces-343	170	47	8	8	NUM
alces-343	170	48	%	%	NOUN
alces-343	170	49	per	per	ADP
alces-343	170	50	year	year	NOUN
alces-343	170	51	)	)	PUNCT
alces-343	170	52	hunting	hunting	NOUN
alces-343	170	53	with	with	ADP
alces-343	170	54	no	no	DET
alces-343	170	55	wolf	wolf	NOUN
alces-343	170	56	trapping	trapping	NOUN
alces-343	170	57	;	;	PUNCT
alces-343	170	58	moose	moose	NOUN
alces-343	170	59	birth	birth	NOUN
alces-343	170	60	rate	rate	NOUN
alces-343	170	61	varying	vary	VERB
alces-343	170	62	randomly	randomly	ADV
alces-343	170	63	(	(	PUNCT
alces-343	170	64	0	0	NUM
alces-343	170	65	–	–	PUNCT
alces-343	170	66	40	40	NUM
alces-343	170	67	%	%	NOUN
alces-343	170	68	)	)	PUNCT
alces-343	170	69	;	;	PUNCT
alces-343	170	70	(	(	PUNCT
alces-343	170	71	c	c	X
alces-343	170	72	)	)	PUNCT
alces-343	170	73	moderate	moderate	ADJ
alces-343	170	74	moose	moose	NOUN
alces-343	170	75	(	(	PUNCT
alces-343	170	76	9	9	NUM
alces-343	170	77	%	%	NOUN
alces-343	170	78	per	per	ADP
alces-343	170	79	year	year	NOUN
alces-343	170	80	)	)	PUNCT
alces-343	170	81	and	and	CCONJ
alces-343	170	82	caribou	caribou	NOUN
alces-343	170	83	(	(	PUNCT
alces-343	170	84	8	8	NUM
alces-343	170	85	%	%	NOUN
alces-343	170	86	per	per	ADP
alces-343	170	87	year	year	NOUN
alces-343	170	88	)	)	PUNCT
alces-343	170	89	hunting	hunting	NOUN
alces-343	170	90	with	with	ADP
alces-343	170	91	no	no	DET
alces-343	170	92	wolf	wolf	NOUN
alces-343	170	93	trapping	trapping	NOUN
alces-343	170	94	;	;	PUNCT
alces-343	170	95	both	both	DET
alces-343	170	96	moose	moose	NOUN
alces-343	170	97	and	and	CCONJ
alces-343	170	98	caribou	caribou	NOUN
alces-343	170	99	birth	birth	NOUN
alces-343	170	100	rates	rate	NOUN
alces-343	170	101	varying	vary	VERB
alces-343	170	102	randomly	randomly	ADV
alces-343	170	103	,	,	PUNCT
alces-343	170	104	between	between	ADP
alces-343	170	105	0	0	NUM
alces-343	170	106	and	and	CCONJ
alces-343	170	107	40	40	NUM
alces-343	170	108	%	%	NOUN
alces-343	170	109	and	and	CCONJ
alces-343	170	110	0	0	NUM
alces-343	170	111	and	and	CCONJ
alces-343	170	112	20	20	NUM
alces-343	170	113	%	%	NOUN
alces-343	170	114	,	,	PUNCT
alces-343	170	115	respectively	respectively	ADV
alces-343	170	116	.	.	PUNCT
alces-343	171	1	impact	impact	NOUN
alces-343	171	2	of	of	ADP
alces-343	171	3	moose	moose	NOUN
alces-343	171	4	and	and	CCONJ
alces-343	171	5	wolves	wolf	NOUN
alces-343	171	6	on	on	ADP
alces-343	171	7	caribou	caribou	NOUN
alces-343	171	8	–	–	PUNCT
alces-343	171	9	courtois	courtois	NOUN
alces-343	171	10	and	and	CCONJ
alces-343	171	11	ouellet	ouellet	PROPN
alces-343	171	12	alces	alces	PROPN
alces-343	171	13	vol	vol	NOUN
alces-343	171	14	.	.	PROPN
alces-343	172	1	43	43	NUM
alces-343	172	2	,	,	PUNCT
alces-343	172	3	2007	2007	NUM
alces-343	172	4	20	20	NUM
alces-343	172	5	moose	moose	NOUN
alces-343	172	6	,	,	PUNCT
alces-343	172	7	0.3	0.3	NUM
alces-343	172	8	–	–	PUNCT
alces-343	172	9	0.5	0.5	NUM
alces-343	172	10	wolves	wolf	NOUN
alces-343	172	11	,	,	PUNCT
alces-343	172	12	and	and	CCONJ
alces-343	172	13	3	3	NUM
alces-343	172	14	–	–	SYM
alces-343	172	15	14	14	NUM
alces-343	172	16	caribou	caribou	NOUN
alces-343	172	17	per	per	ADP
alces-343	172	18	100	100	NUM
alces-343	172	19	km2	km2	NOUN
alces-343	172	20	.	.	PUNCT
alces-343	173	1	in	in	ADP
alces-343	173	2	the	the	DET
alces-343	173	3	last	last	ADJ
alces-343	173	4	simulation	simulation	NOUN
alces-343	173	5	,	,	PUNCT
alces-343	173	6	which	which	PRON
alces-343	173	7	appears	appear	VERB
alces-343	173	8	a	a	DET
alces-343	173	9	plausible	plausible	ADJ
alces-343	173	10	situation	situation	NOUN
alces-343	173	11	in	in	ADP
alces-343	173	12	nature	nature	NOUN
alces-343	173	13	,	,	PUNCT
alces-343	173	14	populations	population	NOUN
alces-343	173	15	stabilized	stabilize	VERB
alces-343	173	16	at	at	ADP
alces-343	173	17	6	6	NUM
alces-343	173	18	–	–	SYM
alces-343	173	19	8	8	NUM
alces-343	173	20	moose	moose	NOUN
alces-343	173	21	,	,	PUNCT
alces-343	173	22	0.2	0.2	NUM
alces-343	173	23	–	–	PUNCT
alces-343	173	24	0.3	0.3	NUM
alces-343	173	25	wolves	wolf	NOUN
alces-343	173	26	,	,	PUNCT
alces-343	173	27	and	and	CCONJ
alces-343	173	28	3.5	3.5	NUM
alces-343	173	29	–	–	SYM
alces-343	173	30	4	4	NUM
alces-343	173	31	.0	.0	NUM
alces-343	173	32	caribou	caribou	NOUN
alces-343	173	33	per	per	ADP
alces-343	173	34	100	100	NUM
alces-343	173	35	km2	km2	NOUN
alces-343	173	36	.	.	PUNCT
alces-343	174	1	discussion	discussion	NOUN
alces-343	174	2	usefulness	usefulness	NOUN
alces-343	174	3	of	of	ADP
alces-343	174	4	the	the	DET
alces-343	174	5	combined	combined	ADJ
alces-343	174	6	model	model	NOUN
alces-343	174	7	in	in	ADP
alces-343	174	8	caribou	caribou	PROPN
alces-343	174	9	,	,	PUNCT
alces-343	174	10	moose	moose	NOUN
alces-343	174	11	,	,	PUNCT
alces-343	174	12	and	and	CCONJ
alces-343	174	13	wolf	wolf	PROPN
alces-343	174	14	management	management	NOUN
alces-343	174	15	our	our	PRON
alces-343	174	16	moose	moose	NOUN
alces-343	174	17	-	-	PUNCT
alces-343	174	18	wolf	wolf	NOUN
alces-343	174	19	-	-	PUNCT
alces-343	174	20	caribou	caribou	NOUN
alces-343	174	21	model	model	NOUN
alces-343	174	22	suggests	suggest	VERB
alces-343	174	23	interesting	interesting	ADJ
alces-343	174	24	alternatives	alternative	NOUN
alces-343	174	25	for	for	ADP
alces-343	174	26	the	the	DET
alces-343	174	27	management	management	NOUN
alces-343	174	28	of	of	ADP
alces-343	174	29	these	these	DET
alces-343	174	30	three	three	NUM
alces-343	174	31	species	specie	NOUN
alces-343	174	32	.	.	PUNCT
alces-343	175	1	seip	seip	NOUN
alces-343	175	2	(	(	PUNCT
alces-343	175	3	1991	1991	NUM
alces-343	175	4	,	,	PUNCT
alces-343	175	5	1992	1992	NUM
alces-343	175	6	)	)	PUNCT
alces-343	175	7	proposed	propose	VERB
alces-343	175	8	that	that	SCONJ
alces-343	175	9	predation	predation	NOUN
alces-343	175	10	could	could	AUX
alces-343	175	11	eliminate	eliminate	VERB
alces-343	175	12	woodland	woodland	NOUN
alces-343	175	13	caribou	caribou	NOUN
alces-343	175	14	whenever	whenever	SCONJ
alces-343	175	15	wolves	wolf	NOUN
alces-343	175	16	are	be	AUX
alces-343	175	17	sustained	sustain	VERB
alces-343	175	18	by	by	ADP
alces-343	175	19	another	another	DET
alces-343	175	20	species	specie	NOUN
alces-343	175	21	because	because	SCONJ
alces-343	175	22	there	there	PRON
alces-343	175	23	is	be	VERB
alces-343	175	24	no	no	DET
alces-343	175	25	retroactive	retroactive	ADJ
alces-343	175	26	mechanism	mechanism	NOUN
alces-343	175	27	that	that	PRON
alces-343	175	28	decreases	decrease	VERB
alces-343	175	29	the	the	DET
alces-343	175	30	impact	impact	NOUN
alces-343	175	31	of	of	ADP
alces-343	175	32	wolves	wolf	NOUN
alces-343	175	33	when	when	SCONJ
alces-343	175	34	the	the	DET
alces-343	175	35	caribou	caribou	NOUN
alces-343	175	36	population	population	NOUN
alces-343	175	37	declines	decline	VERB
alces-343	175	38	.	.	PUNCT
alces-343	176	1	our	our	PRON
alces-343	176	2	simulations	simulation	NOUN
alces-343	176	3	suggest	suggest	VERB
alces-343	176	4	that	that	SCONJ
alces-343	176	5	certain	certain	ADJ
alces-343	176	6	management	management	NOUN
alces-343	176	7	strategies	strategy	NOUN
alces-343	176	8	could	could	AUX
alces-343	176	9	help	help	VERB
alces-343	176	10	maintain	maintain	VERB
alces-343	176	11	caribou	caribou	NOUN
alces-343	176	12	numbers	number	NOUN
alces-343	176	13	by	by	ADP
alces-343	176	14	limiting	limit	VERB
alces-343	176	15	wolf	wolf	PROPN
alces-343	176	16	expansion	expansion	NOUN
alces-343	176	17	.	.	PUNCT
alces-343	177	1	wolf	wolf	PROPN
alces-343	177	2	control	control	NOUN
alces-343	177	3	could	could	AUX
alces-343	177	4	be	be	AUX
alces-343	177	5	considered	consider	VERB
alces-343	177	6	(	(	PUNCT
alces-343	177	7	bergerud	bergerud	NOUN
alces-343	177	8	and	and	CCONJ
alces-343	177	9	elliot	elliot	PROPN
alces-343	177	10	1986	1986	NUM
alces-343	177	11	,	,	PUNCT
alces-343	177	12	seip	seip	NOUN
alces-343	177	13	1991	1991	NUM
alces-343	177	14	)	)	PUNCT
alces-343	177	15	,	,	PUNCT
alces-343	177	16	but	but	CCONJ
alces-343	177	17	this	this	DET
alces-343	177	18	strategy	strategy	NOUN
alces-343	177	19	would	would	AUX
alces-343	177	20	only	only	ADV
alces-343	177	21	have	have	VERB
alces-343	177	22	a	a	DET
alces-343	177	23	minor	minor	ADJ
alces-343	177	24	impact	impact	NOUN
alces-343	177	25	on	on	ADP
alces-343	177	26	caribou	caribou	PROPN
alces-343	177	27	,	,	PUNCT
alces-343	177	28	unless	unless	SCONJ
alces-343	177	29	a	a	DET
alces-343	177	30	very	very	ADV
alces-343	177	31	intensive	intensive	ADJ
alces-343	177	32	control	control	NOUN
alces-343	177	33	is	be	AUX
alces-343	177	34	performed	perform	VERB
alces-343	177	35	.	.	PUNCT
alces-343	178	1	weclaw	weclaw	NOUN
alces-343	178	2	and	and	CCONJ
alces-343	178	3	hudson	hudson	PROPN
alces-343	178	4	(	(	PUNCT
alces-343	178	5	2004	2004	NUM
alces-343	178	6	)	)	PUNCT
alces-343	178	7	obtained	obtain	VERB
alces-343	178	8	similar	similar	ADJ
alces-343	178	9	results	result	NOUN
alces-343	178	10	using	use	VERB
alces-343	178	11	another	another	DET
alces-343	178	12	model	model	NOUN
alces-343	178	13	based	base	VERB
alces-343	178	14	on	on	ADP
alces-343	178	15	responses	response	NOUN
alces-343	178	16	of	of	ADP
alces-343	178	17	wolves	wolf	NOUN
alces-343	178	18	,	,	PUNCT
alces-343	178	19	moose	moose	NOUN
alces-343	178	20	,	,	PUNCT
alces-343	178	21	caribou	caribou	PROPN
alces-343	178	22	,	,	PUNCT
alces-343	178	23	lichens	lichen	NOUN
alces-343	178	24	,	,	PUNCT
alces-343	178	25	and	and	CCONJ
alces-343	178	26	vascular	vascular	ADJ
alces-343	178	27	plants	plant	NOUN
alces-343	178	28	to	to	ADP
alces-343	178	29	various	various	ADJ
alces-343	178	30	natural	natural	ADJ
alces-343	178	31	and	and	CCONJ
alces-343	178	32	anthropogenic	anthropogenic	ADJ
alces-343	178	33	factors	factor	NOUN
alces-343	178	34	.	.	PUNCT
alces-343	179	1	a	a	DET
alces-343	179	2	30	30	NUM
alces-343	179	3	%	%	NOUN
alces-343	179	4	harvest	harvest	NOUN
alces-343	179	5	rate	rate	NOUN
alces-343	179	6	exerted	exert	VERB
alces-343	179	7	by	by	ADP
alces-343	179	8	trappers	trapper	NOUN
alces-343	179	9	is	be	AUX
alces-343	179	10	not	not	PART
alces-343	179	11	high	high	ADJ
alces-343	179	12	birth	birth	NOUN
alces-343	179	13	rate	rate	NOUN
alces-343	179	14	,	,	PUNCT
alces-343	179	15	wolf	wolf	PROPN
alces-343	179	16	populations	population	NOUN
alces-343	179	17	can	can	AUX
alces-343	179	18	increase	increase	VERB
alces-343	179	19	quickly	quickly	ADV
alces-343	179	20	if	if	SCONJ
alces-343	179	21	moose	moose	NOUN
alces-343	179	22	abundance	abundance	NOUN
alces-343	179	23	is	be	AUX
alces-343	179	24	high	high	ADJ
alces-343	179	25	.	.	PUNCT
alces-343	180	1	cessation	cessation	NOUN
alces-343	180	2	of	of	ADP
alces-343	180	3	caribou	caribou	PROPN
alces-343	180	4	hunting	hunting	NOUN
alces-343	180	5	seems	seem	VERB
alces-343	180	6	more	more	ADJ
alces-343	180	7	of	of	ADP
alces-343	180	8	the	the	DET
alces-343	180	9	caribou	caribou	NOUN
alces-343	180	10	population	population	NOUN
alces-343	180	11	in	in	ADP
alces-343	180	12	the	the	DET
alces-343	180	13	long	long	ADJ
alces-343	180	14	term	term	NOUN
alces-343	180	15	if	if	SCONJ
alces-343	180	16	the	the	DET
alces-343	180	17	moose	moose	NOUN
alces-343	180	18	(	(	PUNCT
alces-343	180	19	and	and	CCONJ
alces-343	180	20	wolf	wolf	PROPN
alces-343	180	21	)	)	PUNCT
alces-343	180	22	population	population	NOUN
alces-343	180	23	does	do	AUX
alces-343	180	24	not	not	PART
alces-343	180	25	the	the	DET
alces-343	180	26	habitat	habitat	NOUN
alces-343	180	27	carrying	carry	VERB
alces-343	180	28	capacity	capacity	NOUN
alces-343	180	29	for	for	ADP
alces-343	180	30	moose	moose	NOUN
alces-343	180	31	is	be	AUX
alces-343	180	32	relatively	relatively	ADV
alces-343	180	33	low	low	ADJ
alces-343	180	34	.	.	PUNCT
alces-343	181	1	moose	moose	NOUN
alces-343	181	2	densities	density	NOUN
alces-343	181	3	are	be	AUX
alces-343	181	4	generally	generally	ADV
alces-343	181	5	higher	high	ADJ
alces-343	181	6	in	in	ADP
alces-343	181	7	disturbed	disturbed	ADJ
alces-343	181	8	habitats	habitat	NOUN
alces-343	181	9	and	and	CCONJ
alces-343	181	10	young	young	ADJ
alces-343	181	11	forests	forest	NOUN
alces-343	181	12	rich	rich	ADJ
alces-343	181	13	in	in	ADP
alces-343	181	14	deciduous	deciduous	ADJ
alces-343	181	15	browse	browse	NOUN
alces-343	181	16	than	than	ADP
alces-343	181	17	in	in	ADP
alces-343	181	18	mature	mature	ADJ
alces-343	181	19	spruce	spruce	NOUN
alces-343	181	20	forests	forest	NOUN
alces-343	181	21	(	(	PUNCT
alces-343	181	22	timmermann	timmermann	VERB
alces-343	181	23	and	and	CCONJ
alces-343	181	24	mcnicol	mcnicol	NOUN
alces-343	181	25	1988	1988	NUM
alces-343	181	26	)	)	PUNCT
alces-343	181	27	.	.	PUNCT
alces-343	182	1	moose	moose	NOUN
alces-343	182	2	expansion	expansion	NOUN
alces-343	182	3	could	could	AUX
alces-343	182	4	therefore	therefore	ADV
alces-343	182	5	be	be	AUX
alces-343	182	6	controlled	control	VERB
alces-343	182	7	by	by	ADP
alces-343	182	8	limiting	limit	VERB
alces-343	182	9	forest	forest	NOUN
alces-343	182	10	exploitation	exploitation	NOUN
alces-343	182	11	in	in	ADP
alces-343	182	12	sites	site	NOUN
alces-343	182	13	used	use	VERB
alces-343	182	14	by	by	ADP
alces-343	182	15	caribou	caribou	NOUN
alces-343	182	16	or	or	CCONJ
alces-343	182	17	by	by	ADP
alces-343	182	18	promoting	promote	VERB
alces-343	182	19	spruce	spruce	ADJ
alces-343	182	20	wolves	wolf	NOUN
alces-343	182	21	,	,	PUNCT
alces-343	182	22	our	our	PRON
alces-343	182	23	simulations	simulation	NOUN
alces-343	182	24	suggest	suggest	VERB
alces-343	182	25	that	that	SCONJ
alces-343	182	26	caribou	caribou	PROPN
alces-343	182	27	can	can	AUX
alces-343	182	28	be	be	AUX
alces-343	182	29	maintained	maintain	VERB
alces-343	182	30	in	in	ADP
alces-343	182	31	natural	natural	ADJ
alces-343	182	32	ecosystems	ecosystem	NOUN
alces-343	182	33	,	,	PUNCT
alces-343	182	34	at	at	ADP
alces-343	182	35	least	least	ADJ
alces-343	182	36	in	in	ADP
alces-343	182	37	the	the	DET
alces-343	182	38	absence	absence	NOUN
alces-343	182	39	of	of	ADP
alces-343	182	40	human	human	ADJ
alces-343	182	41	disturbance	disturbance	NOUN
alces-343	182	42	as	as	SCONJ
alces-343	182	43	suggested	suggest	VERB
alces-343	182	44	by	by	ADP
alces-343	182	45	weclaw	weclaw	NOUN
alces-343	182	46	and	and	CCONJ
alces-343	182	47	hudson	hudson	PROPN
alces-343	182	48	(	(	PUNCT
alces-343	182	49	2004	2004	NUM
alces-343	182	50	)	)	PUNCT
alces-343	182	51	.	.	PUNCT
alces-343	183	1	the	the	DET
alces-343	183	2	best	good	ADJ
alces-343	183	3	strategy	strategy	NOUN
alces-343	183	4	,	,	PUNCT
alces-343	183	5	however	however	ADV
alces-343	183	6	,	,	PUNCT
alces-343	183	7	would	would	AUX
alces-343	183	8	consist	consist	VERB
alces-343	183	9	of	of	ADP
alces-343	183	10	maintaining	maintain	VERB
alces-343	183	11	low	low	ADJ
alces-343	183	12	moose	moose	NOUN
alces-343	183	13	densities	density	NOUN
alces-343	183	14	through	through	ADP
alces-343	183	15	population	population	NOUN
alces-343	183	16	and	and	CCONJ
alces-343	183	17	habitat	habitat	NOUN
alces-343	183	18	management	management	NOUN
alces-343	183	19	.	.	PUNCT
alces-343	184	1	low	low	ADJ
alces-343	184	2	moose	moose	NOUN
alces-343	184	3	densities	density	NOUN
alces-343	184	4	imply	imply	VERB
alces-343	184	5	low	low	ADJ
alces-343	184	6	wolf	wolf	NOUN
alces-343	184	7	densities	density	NOUN
alces-343	184	8	and	and	CCONJ
alces-343	184	9	,	,	PUNCT
alces-343	184	10	therefore	therefore	ADV
alces-343	184	11	,	,	PUNCT
alces-343	184	12	low	low	ADJ
alces-343	184	13	predation	predation	NOUN
alces-343	184	14	rates	rate	NOUN
alces-343	184	15	on	on	ADP
alces-343	184	16	caribou	caribou	NOUN
alces-343	184	17	.	.	PUNCT
alces-343	185	1	increasing	increase	VERB
alces-343	185	2	the	the	DET
alces-343	185	3	harvest	harvest	NOUN
alces-343	185	4	of	of	ADP
alces-343	185	5	moose	moose	NOUN
alces-343	185	6	would	would	AUX
alces-343	185	7	be	be	AUX
alces-343	185	8	the	the	DET
alces-343	185	9	easiest	easy	ADJ
alces-343	185	10	and	and	CCONJ
alces-343	185	11	most	most	ADV
alces-343	185	12	convenient	convenient	ADJ
alces-343	185	13	management	management	NOUN
alces-343	185	14	measure	measure	NOUN
alces-343	185	15	to	to	PART
alces-343	185	16	adopt	adopt	VERB
alces-343	185	17	.	.	PUNCT
alces-343	186	1	moose	moose	PROPN
alces-343	186	2	hunting	hunting	NOUN
alces-343	186	3	is	be	AUX
alces-343	186	4	highly	highly	ADV
alces-343	186	5	popular	popular	ADJ
alces-343	186	6	.	.	PUNCT
alces-343	187	1	with	with	ADP
alces-343	187	2	a	a	DET
alces-343	187	3	3to	3to	ADJ
alces-343	187	4	4	4	NUM
alces-343	187	5	-	-	PUNCT
alces-343	187	6	week	week	NOUN
alces-343	187	7	hunting	hunting	NOUN
alces-343	187	8	season	season	NOUN
alces-343	187	9	,	,	PUNCT
alces-343	187	10	the	the	DET
alces-343	187	11	harvest	harvest	NOUN
alces-343	187	12	rate	rate	NOUN
alces-343	187	13	could	could	AUX
alces-343	187	14	reach	reach	VERB
alces-343	187	15	15	15	NUM
alces-343	187	16	%	%	NOUN
alces-343	187	17	if	if	SCONJ
alces-343	187	18	all	all	DET
alces-343	187	19	segments	segment	NOUN
alces-343	187	20	of	of	ADP
alces-343	187	21	the	the	DET
alces-343	187	22	population	population	NOUN
alces-343	187	23	(	(	PUNCT
alces-343	187	24	both	both	DET
alces-343	187	25	sexes	sex	NOUN
alces-343	187	26	of	of	ADP
alces-343	187	27	adults	adult	NOUN
alces-343	187	28	and	and	CCONJ
alces-343	187	29	calves	calf	NOUN
alces-343	187	30	)	)	PUNCT
alces-343	187	31	were	be	AUX
alces-343	187	32	targeted	target	VERB
alces-343	187	33	without	without	ADP
alces-343	187	34	setting	set	VERB
alces-343	187	35	a	a	DET
alces-343	187	36	limit	limit	NOUN
alces-343	187	37	on	on	ADP
alces-343	187	38	the	the	DET
alces-343	187	39	number	number	NOUN
alces-343	187	40	of	of	ADP
alces-343	187	41	hunting	hunting	NOUN
alces-343	187	42	permits	permit	NOUN
alces-343	187	43	issued	issue	VERB
alces-343	187	44	(	(	PUNCT
alces-343	187	45	courtois	courtois	PROPN
alces-343	187	46	et	et	PROPN
alces-343	187	47	al	al	PROPN
alces-343	187	48	.	.	PROPN
alces-343	187	49	1994a	1994a	NUM
alces-343	187	50	)	)	PUNCT
alces-343	187	51	.	.	PUNCT
alces-343	188	1	in	in	ADP
alces-343	188	2	such	such	DET
alces-343	188	3	a	a	DET
alces-343	188	4	situation	situation	NOUN
alces-343	188	5	,	,	PUNCT
alces-343	188	6	our	our	PRON
alces-343	188	7	simulation	simulation	NOUN
alces-343	188	8	suggests	suggest	VERB
alces-343	188	9	that	that	SCONJ
alces-343	188	10	moose	moose	NOUN
alces-343	188	11	density	density	NOUN
alces-343	188	12	would	would	AUX
alces-343	188	13	stabilize	stabilize	VERB
alces-343	188	14	at	at	ADP
alces-343	188	15	about	about	ADP
alces-343	188	16	than	than	ADP
alces-343	188	17	7	7	NUM
alces-343	188	18	–	–	SYM
alces-343	188	19	8	8	NUM
alces-343	188	20	individuals	individual	NOUN
alces-343	188	21	per	per	ADP
alces-343	188	22	100	100	NUM
alces-343	188	23	km2	km2	NOUN
alces-343	188	24	and	and	CCONJ
alces-343	188	25	wolf	wolf	PROPN
alces-343	188	26	density	density	NOUN
alces-343	188	27	at	at	ADP
alces-343	188	28	less	less	ADJ
alces-343	188	29	than	than	ADP
alces-343	188	30	0.2	0.2	NUM
alces-343	188	31	–	–	PUNCT
alces-343	188	32	0.3	0.3	NUM
alces-343	188	33	individuals	individual	NOUN
alces-343	188	34	per	per	ADP
alces-343	188	35	100	100	NUM
alces-343	188	36	km2	km2	NOUN
alces-343	188	37	.	.	PUNCT
alces-343	189	1	annual	annual	ADJ
alces-343	189	2	caribou	caribou	PROPN
alces-343	189	3	recruitment	recruitment	NOUN
alces-343	189	4	would	would	AUX
alces-343	189	5	then	then	ADV
alces-343	189	6	be	be	AUX
alces-343	189	7	approximately	approximately	ADV
alces-343	189	8	19	19	NUM
alces-343	189	9	–	–	PUNCT
alces-343	189	10	20	20	NUM
alces-343	189	11	%	%	NOUN
alces-343	189	12	and	and	CCONJ
alces-343	189	13	adult	adult	NOUN
alces-343	189	14	predation	predation	NOUN
alces-343	189	15	rate	rate	NOUN
alces-343	189	16	would	would	AUX
alces-343	189	17	be	be	AUX
alces-343	189	18	around	around	ADV
alces-343	189	19	7	7	NUM
alces-343	189	20	–	–	PUNCT
alces-343	189	21	8	8	NUM
alces-343	189	22	%	%	NOUN
alces-343	189	23	.	.	PUNCT
alces-343	190	1	a	a	DET
alces-343	190	2	population	population	NOUN
alces-343	190	3	increase	increase	NOUN
alces-343	190	4	in	in	ADP
alces-343	190	5	caribou	caribou	NOUN
alces-343	190	6	to	to	ADP
alces-343	190	7	3	3	NUM
alces-343	190	8	–	–	PUNCT
alces-343	190	9	14	14	NUM
alces-343	190	10	individuals	individual	NOUN
alces-343	190	11	per	per	ADP
alces-343	190	12	100	100	NUM
alces-343	190	13	km2	km2	NOUN
alces-343	190	14	could	could	AUX
alces-343	190	15	thus	thus	ADV
alces-343	190	16	be	be	AUX
alces-343	190	17	seen	see	VERB
alces-343	190	18	in	in	ADP
alces-343	190	19	several	several	ADJ
alces-343	190	20	decades	decade	NOUN
alces-343	190	21	.	.	PUNCT
alces-343	191	1	even	even	ADV
alces-343	191	2	if	if	SCONJ
alces-343	191	3	conservative	conservative	ADJ
alces-343	191	4	caribou	caribou	NOUN
alces-343	191	5	hunting	hunting	NOUN
alces-343	191	6	was	be	AUX
alces-343	191	7	maintained	maintain	VERB
alces-343	191	8	,	,	PUNCT
alces-343	191	9	a	a	DET
alces-343	191	10	doubling	doubling	NOUN
alces-343	191	11	or	or	CCONJ
alces-343	191	12	tripling	tripling	NOUN
alces-343	191	13	of	of	ADP
alces-343	191	14	the	the	DET
alces-343	191	15	population	population	NOUN
alces-343	191	16	could	could	AUX
alces-343	191	17	be	be	AUX
alces-343	191	18	observed	observe	VERB
alces-343	191	19	over	over	ADP
alces-343	191	20	the	the	DET
alces-343	191	21	long	long	ADJ
alces-343	191	22	term	term	NOUN
alces-343	191	23	,	,	PUNCT
alces-343	191	24	depending	depend	VERB
alces-343	191	25	on	on	ADP
alces-343	191	26	the	the	DET
alces-343	191	27	magnitude	magnitude	NOUN
alces-343	191	28	of	of	ADP
alces-343	191	29	variations	variation	NOUN
alces-343	191	30	due	due	ADP
alces-343	191	31	to	to	ADP
alces-343	191	32	environmental	environmental	ADJ
alces-343	191	33	factors	factor	NOUN
alces-343	191	34	.	.	PUNCT
alces-343	192	1	this	this	DET
alces-343	192	2	scenario	scenario	NOUN
alces-343	192	3	supports	support	VERB
alces-343	192	4	the	the	DET
alces-343	192	5	growing	grow	VERB
alces-343	192	6	evidence	evidence	NOUN
alces-343	192	7	that	that	PRON
alces-343	192	8	moose	moose	NOUN
alces-343	192	9	management	management	NOUN
alces-343	192	10	may	may	AUX
alces-343	192	11	be	be	AUX
alces-343	192	12	an	an	DET
alces-343	192	13	integrated	integrate	VERB
alces-343	192	14	aspect	aspect	NOUN
alces-343	192	15	of	of	ADP
alces-343	192	16	caribou	caribou	NOUN
alces-343	192	17	management	management	NOUN
alces-343	192	18	,	,	PUNCT
alces-343	192	19	as	as	SCONJ
alces-343	192	20	indicated	indicate	VERB
alces-343	192	21	by	by	ADP
alces-343	192	22	controlled	control	VERB
alces-343	192	23	experiments	experiment	NOUN
alces-343	192	24	in	in	ADP
alces-343	192	25	the	the	DET
alces-343	192	26	moose	moose	NOUN
alces-343	192	27	hunting	hunt	VERB
alces-343	192	28	rates	rate	NOUN
alces-343	192	29	higher	high	ADJ
alces-343	192	30	than	than	ADP
alces-343	192	31	15	15	NUM
alces-343	192	32	%	%	NOUN
alces-343	192	33	in	in	ADP
alces-343	192	34	order	order	NOUN
alces-343	192	35	to	to	PART
alces-343	192	36	maintain	maintain	VERB
alces-343	192	37	a	a	DET
alces-343	192	38	stable	stable	ADJ
alces-343	192	39	or	or	CCONJ
alces-343	192	40	slightly	slightly	ADV
alces-343	192	41	increasing	increase	VERB
alces-343	192	42	population	population	NOUN
alces-343	192	43	.	.	PUNCT
alces-343	193	1	management	management	NOUN
alces-343	193	2	that	that	PRON
alces-343	193	3	encourages	encourage	VERB
alces-343	193	4	declining	decline	VERB
alces-343	193	5	moose	moose	NOUN
alces-343	193	6	populations	population	NOUN
alces-343	193	7	will	will	AUX
alces-343	193	8	lead	lead	VERB
alces-343	193	9	to	to	ADP
alces-343	193	10	lower	low	ADJ
alces-343	193	11	hunting	hunt	VERB
alces-343	193	12	success	success	NOUN
alces-343	193	13	rate	rate	NOUN
alces-343	193	14	,	,	PUNCT
alces-343	193	15	which	which	PRON
alces-343	193	16	in	in	ADP
alces-343	193	17	the	the	DET
alces-343	193	18	long	long	ADJ
alces-343	193	19	term	term	NOUN
alces-343	193	20	may	may	AUX
alces-343	193	21	be	be	AUX
alces-343	193	22	a	a	DET
alces-343	193	23	less	less	ADV
alces-343	193	24	popular	popular	ADJ
alces-343	193	25	decision	decision	NOUN
alces-343	193	26	among	among	ADP
alces-343	193	27	hunters	hunter	NOUN
alces-343	193	28	.	.	PUNCT
alces-343	194	1	our	our	PRON
alces-343	194	2	simulations	simulation	NOUN
alces-343	194	3	seem	seem	VERB
alces-343	194	4	optimistic	optimistic	ADJ
alces-343	194	5	in	in	ADP
alces-343	194	6	comparison	comparison	NOUN
alces-343	194	7	with	with	ADP
alces-343	194	8	actual	actual	ADJ
alces-343	194	9	woodland	woodland	NOUN
alces-343	194	10	caribou	caribou	NOUN
alces-343	194	11	densities	density	NOUN
alces-343	194	12	(	(	PUNCT
alces-343	194	13	1	1	NUM
alces-343	194	14	–	–	SYM
alces-343	194	15	3	3	NUM
alces-343	194	16	individuals	individual	NOUN
alces-343	194	17	per	per	ADP
alces-343	194	18	100	100	NUM
alces-343	194	19	km2	km2	NOUN
alces-343	194	20	,	,	PUNCT
alces-343	194	21	seip	seip	NOUN
alces-343	194	22	1991	1991	NUM
alces-343	194	23	,	,	PUNCT
alces-343	194	24	courtois	courtois	PROPN
alces-343	194	25	2003	2003	NUM
alces-343	194	26	)	)	PUNCT
alces-343	194	27	;	;	PUNCT
alces-343	194	28	however	however	ADV
alces-343	194	29	,	,	PUNCT
alces-343	194	30	historical	historical	ADJ
alces-343	194	31	densities	density	NOUN
alces-343	194	32	alces	alce	NOUN
alces-343	194	33	vol	vol	NOUN
alces-343	194	34	.	.	PROPN
alces-343	195	1	43	43	NUM
alces-343	195	2	,	,	PUNCT
alces-343	195	3	2007	2007	NUM
alces-343	195	4	courtois	courtois	NOUN
alces-343	195	5	and	and	CCONJ
alces-343	195	6	ouellet	ouellet	NOUN
alces-343	195	7	impact	impact	NOUN
alces-343	195	8	of	of	ADP
alces-343	195	9	moose	moose	NOUN
alces-343	195	10	and	and	CCONJ
alces-343	195	11	wolves	wolf	NOUN
alces-343	195	12	on	on	ADP
alces-343	195	13	caribou	caribou	NOUN
alces-343	195	14	21	21	NUM
alces-343	195	15	were	be	AUX
alces-343	195	16	much	much	ADV
alces-343	195	17	higher	high	ADJ
alces-343	195	18	.	.	PUNCT
alces-343	196	1	brassard	brassard	PROPN
alces-343	196	2	(	(	PUNCT
alces-343	196	3	1967	1967	NUM
alces-343	196	4	)	)	PUNCT
alces-343	196	5	measured	measure	VERB
alces-343	196	6	a	a	DET
alces-343	196	7	density	density	NOUN
alces-343	196	8	of	of	ADP
alces-343	196	9	26.2	26.2	NUM
alces-343	196	10	caribou	caribou	NOUN
alces-343	196	11	per	per	ADP
alces-343	196	12	100	100	NUM
alces-343	196	13	km2	km2	NOUN
alces-343	196	14	in	in	ADP
alces-343	196	15	a	a	DET
alces-343	196	16	25,723	25,723	NUM
alces-343	196	17	km2	km2	NOUN
alces-343	196	18	study	study	NOUN
alces-343	196	19	site	site	NOUN
alces-343	196	20	in	in	ADP
alces-343	196	21	the	the	DET
alces-343	196	22	boreal	boreal	NOUN
alces-343	196	23	forest	forest	NOUN
alces-343	196	24	of	of	ADP
alces-343	196	25	the	the	DET
alces-343	196	26	québec	québec	PROPN
alces-343	196	27	north	north	PROPN
alces-343	196	28	shore	shore	PROPN
alces-343	196	29	.	.	PUNCT
alces-343	197	1	in	in	ADP
alces-343	197	2	labrador	labrador	NOUN
alces-343	197	3	,	,	PUNCT
alces-343	197	4	bergerud	bergerud	NOUN
alces-343	197	5	(	(	PUNCT
alces-343	197	6	1967	1967	NUM
alces-343	197	7	)	)	PUNCT
alces-343	197	8	estimated	estimate	VERB
alces-343	197	9	the	the	DET
alces-343	197	10	1958	1958	NUM
alces-343	197	11	density	density	NOUN
alces-343	197	12	of	of	ADP
alces-343	197	13	the	the	DET
alces-343	197	14	mealy	mealy	ADJ
alces-343	197	15	mountain	mountain	NOUN
alces-343	197	16	herd	herd	NOUN
alces-343	197	17	at	at	ADP
alces-343	197	18	7.9	7.9	NUM
alces-343	197	19	individuals	individual	NOUN
alces-343	197	20	per	per	ADP
alces-343	197	21	100	100	NUM
alces-343	197	22	km2	km2	NOUN
alces-343	197	23	over	over	ADP
alces-343	197	24	30,303	30,303	NUM
alces-343	197	25	km2	km2	NOUN
alces-343	197	26	.	.	PUNCT
alces-343	198	1	both	both	DET
alces-343	198	2	these	these	DET
alces-343	198	3	densities	density	NOUN
alces-343	198	4	rapidly	rapidly	ADV
alces-343	198	5	declined	decline	VERB
alces-343	198	6	following	follow	VERB
alces-343	198	7	intensive	intensive	ADJ
alces-343	198	8	hunting	hunting	NOUN
alces-343	198	9	during	during	ADP
alces-343	198	10	the	the	DET
alces-343	198	11	1960s	1960	NOUN
alces-343	198	12	and	and	CCONJ
alces-343	198	13	1970s	1970	NOUN
alces-343	198	14	following	follow	VERB
alces-343	198	15	the	the	DET
alces-343	198	16	invention	invention	NOUN
alces-343	198	17	of	of	ADP
alces-343	198	18	snowmobiles	snowmobile	NOUN
alces-343	198	19	that	that	PRON
alces-343	198	20	greatly	greatly	ADV
alces-343	198	21	facilitated	facilitate	VERB
alces-343	198	22	hunter	hunter	NOUN
alces-343	198	23	access	access	NOUN
alces-343	198	24	.	.	PUNCT
alces-343	199	1	similarly	similarly	ADV
alces-343	199	2	,	,	PUNCT
alces-343	199	3	caribou	caribou	NOUN
alces-343	199	4	density	density	NOUN
alces-343	199	5	was	be	AUX
alces-343	199	6	approximately	approximately	ADV
alces-343	199	7	4	4	NUM
alces-343	199	8	–	–	SYM
alces-343	199	9	5	5	NUM
alces-343	199	10	individuals	individual	NOUN
alces-343	199	11	per	per	ADP
alces-343	199	12	100	100	NUM
alces-343	199	13	km2	km2	NOUN
alces-343	199	14	in	in	ADP
alces-343	199	15	1992	1992	NUM
alces-343	199	16	in	in	ADP
alces-343	199	17	the	the	DET
alces-343	199	18	charlevoix	charlevoix	PROPN
alces-343	199	19	herd	herd	NOUN
alces-343	199	20	,	,	PUNCT
alces-343	199	21	north	north	ADV
alces-343	199	22	of	of	ADP
alces-343	199	23	québec	québec	PROPN
alces-343	199	24	city	city	PROPN
alces-343	199	25	,	,	PUNCT
alces-343	199	26	in	in	ADP
alces-343	199	27	the	the	DET
alces-343	199	28	absence	absence	NOUN
alces-343	199	29	of	of	ADP
alces-343	199	30	hunting	hunting	NOUN
alces-343	199	31	and	and	CCONJ
alces-343	199	32	restricted	restrict	VERB
alces-343	199	33	forest	forest	NOUN
alces-343	199	34	exploitation	exploitation	NOUN
alces-343	199	35	(	(	PUNCT
alces-343	199	36	sebbane	sebbane	NOUN
alces-343	199	37	2003	2003	NUM
alces-343	199	38	)	)	PUNCT
alces-343	199	39	.	.	PUNCT
alces-343	200	1	caribou	caribou	PROPN
alces-343	200	2	declines	decline	VERB
alces-343	200	3	followed	follow	VERB
alces-343	200	4	anthropogenic	anthropogenic	ADJ
alces-343	200	5	and	and	CCONJ
alces-343	200	6	natural	natural	ADJ
alces-343	200	7	habitat	habitat	NOUN
alces-343	200	8	disturbance	disturbance	NOUN
alces-343	200	9	in	in	ADP
alces-343	200	10	the	the	DET
alces-343	200	11	absence	absence	NOUN
alces-343	200	12	of	of	ADP
alces-343	200	13	hunting	hunting	NOUN
alces-343	200	14	,	,	PUNCT
alces-343	200	15	presumably	presumably	ADV
alces-343	200	16	due	due	ADP
alces-343	200	17	to	to	ADP
alces-343	200	18	increased	increase	VERB
alces-343	200	19	predation	predation	NOUN
alces-343	200	20	.	.	PUNCT
alces-343	201	1	controlled	control	VERB
alces-343	201	2	experiments	experiment	NOUN
alces-343	201	3	(	(	PUNCT
alces-343	201	4	boertje	boertje	NOUN
alces-343	201	5	et	et	PROPN
alces-343	201	6	al	al	PROPN
alces-343	201	7	.	.	PROPN
alces-343	201	8	1996	1996	NUM
alces-343	201	9	,	,	PUNCT
alces-343	201	10	hayes	hayes	PROPN
alces-343	201	11	et	et	PROPN
alces-343	201	12	al	al	PROPN
alces-343	201	13	.	.	PROPN
alces-343	201	14	2003	2003	NUM
alces-343	201	15	)	)	PUNCT
alces-343	201	16	and	and	CCONJ
alces-343	201	17	other	other	ADJ
alces-343	201	18	simulation	simulation	NOUN
alces-343	201	19	models	model	NOUN
alces-343	201	20	(	(	PUNCT
alces-343	201	21	weclaw	weclaw	NOUN
alces-343	201	22	and	and	CCONJ
alces-343	201	23	hudson	hudson	PROPN
alces-343	201	24	2004	2004	NUM
alces-343	201	25	)	)	PUNCT
alces-343	201	26	also	also	ADV
alces-343	201	27	suggest	suggest	VERB
alces-343	201	28	that	that	SCONJ
alces-343	201	29	caribou	caribou	NOUN
alces-343	201	30	densities	density	NOUN
alces-343	201	31	can	can	AUX
alces-343	201	32	attain	attain	VERB
alces-343	201	33	10	10	NUM
alces-343	201	34	individuals	individual	NOUN
alces-343	201	35	per	per	ADP
alces-343	201	36	100	100	NUM
alces-343	201	37	km2	km2	NOUN
alces-343	201	38	,	,	PUNCT
alces-343	201	39	depending	depend	VERB
alces-343	201	40	on	on	ADP
alces-343	201	41	the	the	DET
alces-343	201	42	extent	extent	NOUN
alces-343	201	43	of	of	ADP
alces-343	201	44	wolf	wolf	PROPN
alces-343	201	45	predation	predation	NOUN
alces-343	201	46	and	and	CCONJ
alces-343	201	47	hunting	hunting	NOUN
alces-343	201	48	-	-	PUNCT
alces-343	201	49	related	relate	VERB
alces-343	201	50	mortality	mortality	NOUN
alces-343	201	51	.	.	PUNCT
alces-343	202	1	effects	effect	NOUN
alces-343	202	2	of	of	ADP
alces-343	202	3	increase	increase	NOUN
alces-343	202	4	in	in	ADP
alces-343	202	5	caribou	caribou	NOUN
alces-343	202	6	biomass	biomass	NOUN
alces-343	202	7	we	we	PRON
alces-343	202	8	intentionally	intentionally	ADV
alces-343	202	9	developed	develop	VERB
alces-343	202	10	a	a	DET
alces-343	202	11	simple	simple	ADJ
alces-343	202	12	model	model	NOUN
alces-343	202	13	based	base	VERB
alces-343	202	14	on	on	ADP
alces-343	202	15	empirical	empirical	ADJ
alces-343	202	16	data	datum	NOUN
alces-343	202	17	from	from	ADP
alces-343	202	18	the	the	DET
alces-343	202	19	quéorder	quéorder	NOUN
alces-343	202	20	to	to	PART
alces-343	202	21	avoid	avoid	VERB
alces-343	202	22	including	include	VERB
alces-343	202	23	variables	variable	NOUN
alces-343	202	24	that	that	PRON
alces-343	202	25	were	be	AUX
alces-343	202	26	depended	depend	VERB
alces-343	202	27	exclusively	exclusively	ADV
alces-343	202	28	on	on	ADP
alces-343	202	29	moose	moose	NOUN
alces-343	202	30	density	density	NOUN
alces-343	202	31	.	.	PUNCT
alces-343	203	1	this	this	DET
alces-343	203	2	ence	ence	NOUN
alces-343	203	3	of	of	ADP
alces-343	203	4	ungulates	ungulate	NOUN
alces-343	203	5	is	be	AUX
alces-343	203	6	necessary	necessary	ADJ
alces-343	203	7	to	to	PART
alces-343	203	8	sustain	sustain	VERB
alces-343	203	9	wolf	wolf	NOUN
alces-343	203	10	populations	population	NOUN
alces-343	203	11	(	(	PUNCT
alces-343	203	12	messier	messy	ADJ
alces-343	203	13	1994	1994	NUM
alces-343	203	14	,	,	PUNCT
alces-343	203	15	1995	1995	NUM
alces-343	203	16	)	)	PUNCT
alces-343	203	17	and	and	CCONJ
alces-343	203	18	also	also	ADV
alces-343	203	19	caribou	caribou	NOUN
alces-343	203	20	density	density	NOUN
alces-343	203	21	was	be	AUX
alces-343	203	22	too	too	ADV
alces-343	203	23	low	low	ADJ
alces-343	203	24	to	to	PART
alces-343	203	25	constitute	constitute	VERB
alces-343	203	26	a	a	DET
alces-343	203	27	white	white	ADJ
alces-343	203	28	-	-	PUNCT
alces-343	203	29	tailed	tail	VERB
alces-343	203	30	deer	deer	NOUN
alces-343	203	31	,	,	PUNCT
alces-343	203	32	could	could	AUX
alces-343	203	33	be	be	AUX
alces-343	203	34	an	an	DET
alces-343	203	35	important	important	ADJ
alces-343	203	36	prey	prey	NOUN
alces-343	203	37	for	for	ADP
alces-343	203	38	wolves	wolf	NOUN
alces-343	203	39	during	during	ADP
alces-343	203	40	winter	winter	NOUN
alces-343	203	41	,	,	PUNCT
alces-343	203	42	but	but	CCONJ
alces-343	203	43	this	this	DET
alces-343	203	44	deer	deer	NOUN
alces-343	203	45	is	be	AUX
alces-343	203	46	not	not	PART
alces-343	203	47	found	find	VERB
alces-343	203	48	where	where	SCONJ
alces-343	203	49	caribou	caribou	NOUN
alces-343	203	50	live	live	VERB
alces-343	203	51	in	in	ADP
alces-343	203	52	the	the	DET
alces-343	203	53	northern	northern	ADJ
alces-343	203	54	boreal	boreal	NOUN
alces-343	203	55	forest	forest	NOUN
alces-343	203	56	.	.	PUNCT
alces-343	204	1	in	in	ADP
alces-343	204	2	sites	site	NOUN
alces-343	204	3	used	use	VERB
alces-343	204	4	by	by	ADP
alces-343	204	5	woodland	woodland	PROPN
alces-343	204	6	caribou	caribou	PROPN
alces-343	204	7	,	,	PUNCT
alces-343	204	8	for	for	ADP
alces-343	204	9	wolves	wolf	NOUN
alces-343	204	10	.	.	PUNCT
alces-343	205	1	although	although	SCONJ
alces-343	205	2	wolves	wolf	NOUN
alces-343	205	3	certainly	certainly	ADV
alces-343	205	4	consume	consume	VERB
alces-343	205	5	beavers	beaver	NOUN
alces-343	205	6	in	in	ADP
alces-343	205	7	summer	summer	NOUN
alces-343	205	8	,	,	PUNCT
alces-343	205	9	which	which	PRON
alces-343	205	10	presumably	presumably	ADV
alces-343	205	11	increases	increase	VERB
alces-343	205	12	pup	pup	ADJ
alces-343	205	13	survival	survival	NOUN
alces-343	205	14	,	,	PUNCT
alces-343	205	15	beavers	beaver	NOUN
alces-343	205	16	are	be	AUX
alces-343	205	17	not	not	PART
alces-343	205	18	readily	readily	ADV
alces-343	205	19	available	available	ADJ
alces-343	205	20	in	in	ADP
alces-343	205	21	winter	winter	NOUN
alces-343	205	22	because	because	SCONJ
alces-343	205	23	beaver	beaver	ADJ
alces-343	205	24	movements	movement	NOUN
alces-343	205	25	are	be	AUX
alces-343	205	26	mostly	mostly	ADV
alces-343	205	27	restricted	restrict	VERB
alces-343	205	28	to	to	ADP
alces-343	205	29	beneath	beneath	ADP
alces-343	205	30	the	the	DET
alces-343	205	31	ice	ice	NOUN
alces-343	205	32	where	where	SCONJ
alces-343	205	33	nevertheless	nevertheless	ADV
alces-343	205	34	,	,	PUNCT
alces-343	205	35	we	we	PRON
alces-343	205	36	tested	test	VERB
alces-343	205	37	the	the	DET
alces-343	205	38	effect	effect	NOUN
alces-343	205	39	of	of	ADP
alces-343	205	40	a	a	DET
alces-343	205	41	possible	possible	ADJ
alces-343	205	42	caribou	caribou	NOUN
alces-343	205	43	-	-	PUNCT
alces-343	205	44	wolf	wolf	NOUN
alces-343	205	45	retroaction	retroaction	NOUN
alces-343	205	46	mechanism	mechanism	NOUN
alces-343	205	47	.	.	PUNCT
alces-343	206	1	we	we	PRON
alces-343	206	2	the	the	DET
alces-343	206	3	carrying	carry	VERB
alces-343	206	4	capacity	capacity	NOUN
alces-343	206	5	of	of	ADP
alces-343	206	6	wolves	wolf	NOUN
alces-343	206	7	depended	depend	VERB
alces-343	206	8	on	on	ADP
alces-343	206	9	the	the	DET
alces-343	206	10	total	total	ADJ
alces-343	206	11	biomass	biomass	NOUN
alces-343	206	12	of	of	ADP
alces-343	206	13	available	available	ADJ
alces-343	206	14	ungulates	ungulate	NOUN
alces-343	206	15	(	(	PUNCT
alces-343	206	16	fuller	full	ADJ
alces-343	206	17	1989	1989	NUM
alces-343	206	18	)	)	PUNCT
alces-343	206	19	and	and	CCONJ
alces-343	206	20	we	we	PRON
alces-343	206	21	considered	consider	VERB
alces-343	206	22	that	that	SCONJ
alces-343	206	23	1	1	NUM
alces-343	206	24	caribou	caribou	NOUN
alces-343	206	25	was	be	AUX
alces-343	206	26	equivalent	equivalent	ADJ
alces-343	206	27	to	to	ADP
alces-343	206	28	0.29	0.29	NUM
alces-343	206	29	moose	moose	NOUN
alces-343	206	30	,	,	PUNCT
alces-343	206	31	based	base	VERB
alces-343	206	32	on	on	ADP
alces-343	206	33	respective	respective	ADJ
alces-343	206	34	masses	masse	NOUN
alces-343	206	35	.	.	PUNCT
alces-343	207	1	as	as	ADP
alces-343	207	2	a	a	DET
alces-343	207	3	result	result	NOUN
alces-343	207	4	,	,	PUNCT
alces-343	207	5	the	the	DET
alces-343	207	6	wolf	wolf	PROPN
alces-343	207	7	population	population	NOUN
alces-343	207	8	increased	increase	VERB
alces-343	207	9	by	by	ADP
alces-343	207	10	2	2	NUM
alces-343	207	11	%	%	NOUN
alces-343	207	12	compared	compare	VERB
alces-343	207	13	to	to	ADP
alces-343	207	14	scenario	scenario	NOUN
alces-343	207	15	2	2	NUM
alces-343	207	16	(	(	PUNCT
alces-343	207	17	harvest	harvest	NOUN
alces-343	207	18	rates	rate	NOUN
alces-343	207	19	:	:	PUNCT
alces-343	207	20	9	9	NUM
alces-343	207	21	%	%	NOUN
alces-343	207	22	,	,	PUNCT
alces-343	207	23	0	0	NUM
alces-343	207	24	%	%	NOUN
alces-343	207	25	,	,	PUNCT
alces-343	207	26	and	and	CCONJ
alces-343	207	27	0	0	NUM
alces-343	207	28	%	%	NOUN
alces-343	207	29	for	for	ADP
alces-343	207	30	moose	moose	NOUN
alces-343	207	31	,	,	PUNCT
alces-343	207	32	caribou	caribou	NOUN
alces-343	207	33	,	,	PUNCT
alces-343	207	34	and	and	CCONJ
alces-343	207	35	wolves	wolf	NOUN
alces-343	207	36	,	,	PUNCT
alces-343	207	37	respectively	respectively	ADV
alces-343	207	38	)	)	PUNCT
alces-343	207	39	,	,	PUNCT
alces-343	207	40	which	which	PRON
alces-343	207	41	created	create	VERB
alces-343	207	42	population	population	NOUN
alces-343	207	43	declines	decline	NOUN
alces-343	207	44	of	of	ADP
alces-343	207	45	14	14	NUM
alces-343	207	46	%	%	NOUN
alces-343	207	47	in	in	ADP
alces-343	207	48	moose	moose	NOUN
alces-343	207	49	and	and	CCONJ
alces-343	207	50	6	6	NUM
alces-343	207	51	%	%	NOUN
alces-343	207	52	in	in	ADP
alces-343	207	53	caribou	caribou	NOUN
alces-343	207	54	.	.	PUNCT
alces-343	208	1	assuming	assume	VERB
alces-343	208	2	that	that	SCONJ
alces-343	208	3	wolves	wolf	NOUN
alces-343	208	4	make	make	VERB
alces-343	208	5	hunting	hunt	VERB
alces-343	208	6	decisions	decision	NOUN
alces-343	208	7	based	base	VERB
alces-343	208	8	solely	solely	ADV
alces-343	208	9	on	on	ADP
alces-343	208	10	prey	prey	PROPN
alces-343	208	11	biomass	biomass	NOUN
alces-343	208	12	,	,	PUNCT
alces-343	208	13	not	not	PART
alces-343	208	14	risk	risk	NOUN
alces-343	208	15	associated	associate	VERB
alces-343	208	16	with	with	ADP
alces-343	208	17	their	their	PRON
alces-343	208	18	capture	capture	NOUN
alces-343	208	19	(	(	PUNCT
alces-343	208	20	which	which	PRON
alces-343	208	21	is	be	AUX
alces-343	208	22	lower	low	ADJ
alces-343	208	23	for	for	ADP
alces-343	208	24	caribou	caribou	NOUN
alces-343	208	25	than	than	ADP
alces-343	208	26	moose	moose	NOUN
alces-343	208	27	)	)	PUNCT
alces-343	208	28	,	,	PUNCT
alces-343	208	29	the	the	DET
alces-343	208	30	impact	impact	NOUN
alces-343	208	31	of	of	ADP
alces-343	208	32	an	an	DET
alces-343	208	33	increase	increase	NOUN
alces-343	208	34	in	in	ADP
alces-343	208	35	wolves	wolf	NOUN
alces-343	208	36	would	would	AUX
alces-343	208	37	probably	probably	ADV
alces-343	208	38	be	be	AUX
alces-343	208	39	greater	great	ADJ
alces-343	208	40	on	on	ADP
alces-343	208	41	moose	moose	NOUN
alces-343	208	42	than	than	ADP
alces-343	208	43	caribou	caribou	NOUN
alces-343	208	44	.	.	PUNCT
alces-343	209	1	seip	seip	NOUN
alces-343	209	2	(	(	PUNCT
alces-343	209	3	1991	1991	NUM
alces-343	209	4	)	)	PUNCT
alces-343	209	5	and	and	CCONJ
alces-343	209	6	bergerud	bergerud	NOUN
alces-343	209	7	(	(	PUNCT
alces-343	209	8	1996	1996	NUM
alces-343	209	9	)	)	PUNCT
alces-343	209	10	have	have	AUX
alces-343	209	11	suggested	suggest	VERB
alces-343	209	12	that	that	SCONJ
alces-343	209	13	caribou	caribou	NOUN
alces-343	209	14	could	could	AUX
alces-343	209	15	be	be	AUX
alces-343	209	16	extirpated	extirpate	VERB
alces-343	209	17	from	from	ADP
alces-343	209	18	an	an	DET
alces-343	209	19	area	area	NOUN
alces-343	209	20	in	in	ADP
alces-343	209	21	the	the	DET
alces-343	209	22	presence	presence	NOUN
alces-343	209	23	of	of	ADP
alces-343	209	24	high	high	ADJ
alces-343	209	25	moose	moose	NOUN
alces-343	209	26	densities	density	NOUN
alces-343	209	27	due	due	ADJ
alces-343	209	28	to	to	ADP
alces-343	209	29	increased	increase	VERB
alces-343	209	30	predation	predation	NOUN
alces-343	209	31	on	on	ADP
alces-343	209	32	caribou	caribou	NOUN
alces-343	209	33	from	from	ADP
alces-343	209	34	increased	increase	VERB
alces-343	209	35	wolf	wolf	NOUN
alces-343	209	36	densities	density	NOUN
alces-343	209	37	.	.	PUNCT
alces-343	210	1	in	in	ADP
alces-343	210	2	contrast	contrast	NOUN
alces-343	210	3	,	,	PUNCT
alces-343	210	4	hayes	hayes	PROPN
alces-343	210	5	et	et	PROPN
alces-343	210	6	al	al	PROPN
alces-343	210	7	.	.	PROPN
alces-343	210	8	(	(	PUNCT
alces-343	210	9	2003	2003	NUM
alces-343	210	10	)	)	PUNCT
alces-343	210	11	have	have	AUX
alces-343	210	12	suggested	suggest	VERB
alces-343	210	13	that	that	SCONJ
alces-343	210	14	reduced	reduce	VERB
alces-343	210	15	moose	moose	NOUN
alces-343	210	16	densities	density	NOUN
alces-343	210	17	would	would	AUX
alces-343	210	18	lead	lead	VERB
alces-343	210	19	to	to	ADP
alces-343	210	20	caribou	caribou	NOUN
alces-343	210	21	extirpation	extirpation	NOUN
alces-343	210	22	in	in	ADP
alces-343	210	23	areas	area	NOUN
alces-343	210	24	where	where	SCONJ
alces-343	210	25	wolves	wolf	NOUN
alces-343	210	26	,	,	PUNCT
alces-343	210	27	moose	moose	NOUN
alces-343	210	28	,	,	PUNCT
alces-343	210	29	and	and	CCONJ
alces-343	210	30	caribou	caribou	NOUN
alces-343	210	31	use	use	VERB
alces-343	210	32	similar	similar	ADJ
alces-343	210	33	habitats	habitat	NOUN
alces-343	210	34	in	in	ADP
alces-343	210	35	summer	summer	NOUN
alces-343	210	36	because	because	SCONJ
alces-343	210	37	in	in	ADP
alces-343	210	38	such	such	DET
alces-343	210	39	a	a	DET
alces-343	210	40	case	case	NOUN
alces-343	210	41	caribou	caribou	NOUN
alces-343	210	42	lose	lose	VERB
alces-343	210	43	the	the	DET
alces-343	210	44	advantage	advantage	NOUN
alces-343	210	45	of	of	ADP
alces-343	210	46	their	their	PRON
alces-343	210	47	spacing	space	VERB
alces-343	210	48	strategy	strategy	NOUN
alces-343	210	49	to	to	PART
alces-343	210	50	avoid	avoid	VERB
alces-343	210	51	predation	predation	NOUN
alces-343	210	52	.	.	PUNCT
alces-343	211	1	in	in	ADP
alces-343	211	2	the	the	DET
alces-343	211	3	model	model	NOUN
alces-343	211	4	including	include	VERB
alces-343	211	5	both	both	DET
alces-343	211	6	moose	moose	NOUN
alces-343	211	7	and	and	CCONJ
alces-343	211	8	caribou	caribou	NOUN
alces-343	211	9	biomass	biomass	NOUN
alces-343	211	10	to	to	PART
alces-343	211	11	support	support	VERB
alces-343	211	12	wolves	wolf	NOUN
alces-343	211	13	,	,	PUNCT
alces-343	211	14	increasing	increase	VERB
alces-343	211	15	the	the	DET
alces-343	211	16	moose	moose	NOUN
alces-343	211	17	harvest	harvest	NOUN
alces-343	211	18	rate	rate	NOUN
alces-343	211	19	to	to	ADP
alces-343	211	20	22	22	NUM
alces-343	211	21	%	%	NOUN
alces-343	211	22	led	lead	VERB
alces-343	211	23	to	to	ADP
alces-343	211	24	extirpation	extirpation	NOUN
alces-343	211	25	of	of	ADP
alces-343	211	26	this	this	DET
alces-343	211	27	species	specie	NOUN
alces-343	211	28	in	in	ADP
alces-343	211	29	100	100	NUM
alces-343	211	30	years	year	NOUN
alces-343	211	31	,	,	PUNCT
alces-343	211	32	while	while	SCONJ
alces-343	211	33	caribou	caribou	PROPN
alces-343	211	34	increased	increase	VERB
alces-343	211	35	to	to	ADP
alces-343	211	36	156	156	NUM
alces-343	211	37	individuals	individual	NOUN
alces-343	211	38	.	.	PUNCT
alces-343	212	1	despite	despite	SCONJ
alces-343	212	2	a	a	DET
alces-343	212	3	10	10	NUM
alces-343	212	4	-	-	ADJ
alces-343	212	5	fold	fold	ADJ
alces-343	212	6	increase	increase	NOUN
alces-343	212	7	,	,	PUNCT
alces-343	212	8	the	the	DET
alces-343	212	9	caribou	caribou	NOUN
alces-343	212	10	biomass	biomass	NOUN
alces-343	212	11	(	(	PUNCT
alces-343	212	12	4.5	4.5	NUM
alces-343	212	13	moose	moose	NOUN
alces-343	212	14	-	-	PUNCT
alces-343	212	15	equivalent	equivalent	NOUN
alces-343	212	16	per	per	ADP
alces-343	212	17	100	100	NUM
alces-343	212	18	km2	km2	NOUN
alces-343	212	19	a	a	DET
alces-343	212	20	viable	viable	ADJ
alces-343	212	21	wolf	wolf	NOUN
alces-343	212	22	population	population	NOUN
alces-343	212	23	that	that	PRON
alces-343	212	24	subsequently	subsequently	ADV
alces-343	212	25	declined	decline	VERB
alces-343	212	26	to	to	ADP
alces-343	212	27	1	1	NUM
alces-343	212	28	animal	animal	NOUN
alces-343	212	29	after	after	ADP
alces-343	212	30	100	100	NUM
alces-343	212	31	years	year	NOUN
alces-343	212	32	,	,	PUNCT
alces-343	212	33	suggesting	suggest	VERB
alces-343	212	34	that	that	SCONJ
alces-343	212	35	an	an	DET
alces-343	212	36	abundant	abundant	ADJ
alces-343	212	37	primary	primary	ADJ
alces-343	212	38	prey	prey	NOUN
alces-343	212	39	(	(	PUNCT
alces-343	212	40	moose	moose	NOUN
alces-343	212	41	or	or	CCONJ
alces-343	212	42	deer	deer	NOUN
alces-343	212	43	)	)	PUNCT
alces-343	212	44	is	be	AUX
alces-343	212	45	required	require	VERB
alces-343	212	46	to	to	PART
alces-343	212	47	maintain	maintain	VERB
alces-343	212	48	wolves	wolf	NOUN
alces-343	212	49	in	in	ADP
alces-343	212	50	the	the	DET
alces-343	212	51	boreal	boreal	NOUN
alces-343	212	52	forest	forest	NOUN
alces-343	212	53	.	.	PUNCT
alces-343	213	1	our	our	PRON
alces-343	213	2	model	model	NOUN
alces-343	213	3	implies	imply	VERB
alces-343	213	4	that	that	SCONJ
alces-343	213	5	natural	natural	ADJ
alces-343	213	6	and	and	CCONJ
alces-343	213	7	anthropogenic	anthropogenic	ADJ
alces-343	213	8	mortality	mortality	NOUN
alces-343	213	9	are	be	AUX
alces-343	213	10	additive	additive	ADJ
alces-343	213	11	.	.	PUNCT
alces-343	214	1	this	this	DET
alces-343	214	2	assumption	assumption	NOUN
alces-343	214	3	is	be	AUX
alces-343	214	4	probably	probably	ADV
alces-343	214	5	valid	valid	ADJ
alces-343	214	6	for	for	ADP
alces-343	214	7	ungulates	ungulate	NOUN
alces-343	214	8	,	,	PUNCT
alces-343	214	9	impact	impact	NOUN
alces-343	214	10	of	of	ADP
alces-343	214	11	moose	moose	NOUN
alces-343	214	12	and	and	CCONJ
alces-343	214	13	wolves	wolf	NOUN
alces-343	214	14	on	on	ADP
alces-343	214	15	caribou	caribou	NOUN
alces-343	214	16	–	–	PUNCT
alces-343	214	17	courtois	courtois	NOUN
alces-343	214	18	and	and	CCONJ
alces-343	214	19	ouellet	ouellet	PROPN
alces-343	214	20	alces	alces	PROPN
alces-343	214	21	vol	vol	NOUN
alces-343	214	22	.	.	PROPN
alces-343	215	1	43	43	NUM
alces-343	215	2	,	,	PUNCT
alces-343	215	3	2007	2007	NUM
alces-343	215	4	22	22	NUM
alces-343	216	1	but	but	CCONJ
alces-343	216	2	wolf	wolf	NOUN
alces-343	216	3	trapping	trap	VERB
alces-343	216	4	could	could	AUX
alces-343	216	5	be	be	AUX
alces-343	216	6	partly	partly	ADV
alces-343	216	7	compensatory	compensatory	ADJ
alces-343	216	8	(	(	PUNCT
alces-343	216	9	fuller	full	ADJ
alces-343	216	10	1989	1989	NUM
alces-343	216	11	)	)	PUNCT
alces-343	216	12	.	.	PUNCT
alces-343	217	1	harvesting	harvest	VERB
alces-343	217	2	wolves	wolf	NOUN
alces-343	217	3	would	would	AUX
alces-343	217	4	indeed	indeed	ADV
alces-343	217	5	leave	leave	VERB
alces-343	217	6	more	more	ADJ
alces-343	217	7	food	food	NOUN
alces-343	217	8	available	available	ADJ
alces-343	217	9	for	for	ADP
alces-343	217	10	each	each	DET
alces-343	217	11	surviving	survive	VERB
alces-343	217	12	wolf	wolf	NOUN
alces-343	217	13	,	,	PUNCT
alces-343	217	14	which	which	PRON
alces-343	217	15	could	could	AUX
alces-343	217	16	contribute	contribute	VERB
alces-343	217	17	to	to	ADP
alces-343	217	18	a	a	DET
alces-343	217	19	wolf	wolf	ADJ
alces-343	217	20	population	population	NOUN
alces-343	217	21	increase	increase	NOUN
alces-343	217	22	.	.	PUNCT
alces-343	218	1	we	we	PRON
alces-343	218	2	did	do	AUX
alces-343	218	3	not	not	PART
alces-343	218	4	have	have	VERB
alces-343	218	5	any	any	DET
alces-343	218	6	empirical	empirical	ADJ
alces-343	218	7	evidence	evidence	NOUN
alces-343	218	8	to	to	PART
alces-343	218	9	quantify	quantify	VERB
alces-343	218	10	this	this	DET
alces-343	218	11	possibility	possibility	NOUN
alces-343	218	12	,	,	PUNCT
alces-343	218	13	however	however	ADV
alces-343	218	14	.	.	PUNCT
alces-343	219	1	moreover	moreover	ADV
alces-343	219	2	,	,	PUNCT
alces-343	219	3	if	if	SCONJ
alces-343	219	4	trapping	trap	VERB
alces-343	219	5	is	be	AUX
alces-343	219	6	extensive	extensive	ADJ
alces-343	219	7	,	,	PUNCT
alces-343	219	8	the	the	DET
alces-343	219	9	reduced	reduce	VERB
alces-343	219	10	wolf	wolf	NOUN
alces-343	219	11	packs	pack	NOUN
alces-343	219	12	should	should	AUX
alces-343	219	13	be	be	AUX
alces-343	219	14	biomass	biomass	NOUN
alces-343	219	15	per	per	ADP
alces-343	219	16	animal	animal	NOUN
alces-343	219	17	.	.	PUNCT
alces-343	220	1	limits	limit	NOUN
alces-343	220	2	of	of	ADP
alces-343	220	3	the	the	DET
alces-343	220	4	model	model	NOUN
alces-343	220	5	simulation	simulation	NOUN
alces-343	220	6	results	result	NOUN
alces-343	220	7	are	be	AUX
alces-343	220	8	strongly	strongly	ADV
alces-343	220	9	dependent	dependent	ADJ
alces-343	220	10	on	on	ADP
alces-343	220	11	which	which	PRON
alces-343	220	12	combinations	combination	NOUN
alces-343	220	13	of	of	ADP
alces-343	220	14	parameters	parameter	NOUN
alces-343	220	15	are	be	AUX
alces-343	220	16	used	use	VERB
alces-343	220	17	and	and	CCONJ
alces-343	220	18	on	on	ADP
alces-343	220	19	the	the	DET
alces-343	220	20	accuracy	accuracy	NOUN
alces-343	220	21	of	of	ADP
alces-343	220	22	the	the	DET
alces-343	220	23	parameters	parameter	NOUN
alces-343	220	24	included	include	VERB
alces-343	220	25	.	.	PUNCT
alces-343	221	1	caribou	caribou	NOUN
alces-343	221	2	and	and	CCONJ
alces-343	221	3	moose	moose	NOUN
alces-343	221	4	densities	density	NOUN
alces-343	221	5	,	,	PUNCT
alces-343	221	6	food	food	NOUN
alces-343	221	7	carrying	carry	VERB
alces-343	221	8	capacity	capacity	NOUN
alces-343	221	9	,	,	PUNCT
alces-343	221	10	and	and	CCONJ
alces-343	221	11	hunting	hunt	VERB
alces-343	221	12	mortalities	mortality	NOUN
alces-343	221	13	were	be	AUX
alces-343	221	14	based	base	VERB
alces-343	221	15	on	on	ADP
alces-343	221	16	empirical	empirical	ADJ
alces-343	221	17	data	datum	NOUN
alces-343	221	18	from	from	ADP
alces-343	221	19	the	the	DET
alces-343	221	20	québec	québec	PROPN
alces-343	221	21	boreal	boreal	NOUN
alces-343	221	22	forest	forest	NOUN
alces-343	221	23	(	(	PUNCT
alces-343	221	24	courtois	courtois	PROPN
alces-343	221	25	2003	2003	NUM
alces-343	221	26	)	)	PUNCT
alces-343	221	27	,	,	PUNCT
alces-343	221	28	but	but	CCONJ
alces-343	221	29	wolf	wolf	PROPN
alces-343	221	30	densities	density	NOUN
alces-343	221	31	and	and	CCONJ
alces-343	221	32	their	their	PRON
alces-343	221	33	related	relate	VERB
alces-343	221	34	impact	impact	NOUN
alces-343	221	35	on	on	ADP
alces-343	221	36	caribou	caribou	NOUN
alces-343	221	37	numbers	number	NOUN
alces-343	221	38	could	could	AUX
alces-343	221	39	be	be	AUX
alces-343	221	40	greater	great	ADJ
alces-343	221	41	than	than	ADP
alces-343	221	42	those	those	PRON
alces-343	221	43	predicted	predict	VERB
alces-343	221	44	by	by	ADP
alces-343	221	45	our	our	PRON
alces-343	221	46	model	model	NOUN
alces-343	221	47	.	.	PUNCT
alces-343	222	1	larivière	larivière	VERB
alces-343	222	2	et	et	PROPN
alces-343	222	3	al	al	PROPN
alces-343	222	4	.	.	PROPN
alces-343	223	1	(	(	PUNCT
alces-343	223	2	2000	2000	NUM
alces-343	223	3	)	)	PUNCT
alces-343	223	4	used	use	VERB
alces-343	223	5	the	the	DET
alces-343	223	6	number	number	NOUN
alces-343	223	7	of	of	ADP
alces-343	223	8	wolf	wolf	PROPN
alces-343	223	9	howls	howls	AUX
alces-343	223	10	heard	hear	VERB
alces-343	223	11	by	by	ADP
alces-343	223	12	hunters	hunter	NOUN
alces-343	223	13	to	to	PART
alces-343	223	14	predict	predict	VERB
alces-343	223	15	wolf	wolf	PROPN
alces-343	223	16	density	density	PROPN
alces-343	223	17	.	.	PUNCT
alces-343	224	1	for	for	ADP
alces-343	224	2	the	the	DET
alces-343	224	3	port	port	NOUN
alces-343	224	4	-	-	PUNCT
alces-343	224	5	cartier	cartier	NOUN
alces-343	224	6	–	–	PUNCT
alces-343	224	7	sept	sept	PROPN
alces-343	224	8	-	-	PUNCT
alces-343	224	9	îles	îles	PROPN
alces-343	224	10	wildlife	wildlife	NOUN
alces-343	224	11	reserve	reserve	NOUN
alces-343	224	12	,	,	PUNCT
alces-343	224	13	which	which	PRON
alces-343	224	14	is	be	AUX
alces-343	224	15	located	locate	VERB
alces-343	224	16	in	in	ADP
alces-343	224	17	the	the	DET
alces-343	224	18	main	main	ADJ
alces-343	224	19	distribution	distribution	NOUN
alces-343	224	20	area	area	NOUN
alces-343	224	21	of	of	ADP
alces-343	224	22	woodland	woodland	PROPN
alces-343	224	23	caribou	caribou	PROPN
alces-343	224	24	,	,	PUNCT
alces-343	224	25	these	these	DET
alces-343	224	26	authors	author	NOUN
alces-343	224	27	estimated	estimate	VERB
alces-343	224	28	a	a	DET
alces-343	224	29	density	density	NOUN
alces-343	224	30	of	of	ADP
alces-343	224	31	0.85	0.85	NUM
alces-343	224	32	wolves	wolf	NOUN
alces-343	224	33	per	per	ADP
alces-343	224	34	100	100	NUM
alces-343	224	35	km2	km2	NOUN
alces-343	224	36	.	.	PUNCT
alces-343	225	1	using	use	VERB
alces-343	225	2	fuller	full	ADJ
alces-343	225	3	’s	’s	PART
alces-343	225	4	model	model	NOUN
alces-343	225	5	(	(	PUNCT
alces-343	225	6	1989	1989	NUM
alces-343	225	7	)	)	PUNCT
alces-343	225	8	,	,	PUNCT
alces-343	225	9	0.48	0.48	NUM
alces-343	225	10	wolves	wolf	NOUN
alces-343	225	11	per	per	ADP
alces-343	225	12	100	100	NUM
alces-343	225	13	km2	km2	NOUN
alces-343	225	14	would	would	AUX
alces-343	225	15	be	be	AUX
alces-343	225	16	obtained	obtain	VERB
alces-343	225	17	.	.	PUNCT
alces-343	226	1	the	the	DET
alces-343	226	2	model	model	NOUN
alces-343	226	3	we	we	PRON
alces-343	226	4	used	use	VERB
alces-343	226	5	(	(	PUNCT
alces-343	226	6	messier	messy	ADJ
alces-343	226	7	1994	1994	NUM
alces-343	226	8	)	)	PUNCT
alces-343	226	9	predicted	predict	VERB
alces-343	226	10	0.22	0.22	NUM
alces-343	226	11	wolves	wolf	NOUN
alces-343	226	12	per	per	ADP
alces-343	226	13	100	100	NUM
alces-343	226	14	km2	km2	NOUN
alces-343	226	15	for	for	ADP
alces-343	226	16	a	a	DET
alces-343	226	17	similar	similar	ADJ
alces-343	226	18	moose	moose	NOUN
alces-343	226	19	abundance	abundance	NOUN
alces-343	226	20	(	(	PUNCT
alces-343	226	21	6	6	NUM
alces-343	226	22	individuals	individual	NOUN
alces-343	226	23	per	per	ADP
alces-343	226	24	100	100	NUM
alces-343	226	25	km2	km2	NUM
alces-343	226	26	)	)	PUNCT
alces-343	226	27	,	,	PUNCT
alces-343	226	28	and	and	CCONJ
alces-343	226	29	maximum	maximum	ADJ
alces-343	226	30	density	density	NOUN
alces-343	226	31	of	of	ADP
alces-343	226	32	0.38	0.38	NUM
alces-343	226	33	wolves	wolf	NOUN
alces-343	226	34	per	per	ADP
alces-343	226	35	100	100	NUM
alces-343	226	36	km2	km2	NOUN
alces-343	226	37	at	at	ADP
alces-343	226	38	carrying	carry	VERB
alces-343	226	39	capacity	capacity	NOUN
alces-343	226	40	(	(	PUNCT
alces-343	226	41	84	84	NUM
alces-343	226	42	moose	moose	NOUN
alces-343	226	43	per	per	ADP
alces-343	226	44	100	100	NUM
alces-343	226	45	km2	km2	NUM
alces-343	226	46	)	)	PUNCT
alces-343	226	47	.	.	PUNCT
alces-343	227	1	only	only	ADV
alces-343	227	2	an	an	DET
alces-343	227	3	inventory	inventory	NOUN
alces-343	227	4	could	could	AUX
alces-343	227	5	provide	provide	VERB
alces-343	227	6	accurate	accurate	ADJ
alces-343	227	7	wolf	wolf	NOUN
alces-343	227	8	numbers	number	NOUN
alces-343	227	9	in	in	ADP
alces-343	227	10	the	the	DET
alces-343	227	11	study	study	NOUN
alces-343	227	12	area	area	NOUN
alces-343	227	13	.	.	PUNCT
alces-343	228	1	however	however	ADV
alces-343	228	2	,	,	PUNCT
alces-343	228	3	if	if	SCONJ
alces-343	228	4	the	the	DET
alces-343	228	5	wolf	wolf	NOUN
alces-343	228	6	densities	density	NOUN
alces-343	228	7	predicted	predict	VERB
alces-343	228	8	by	by	ADP
alces-343	228	9	larivière	larivière	PROPN
alces-343	228	10	et	et	PROPN
alces-343	228	11	al	al	PROPN
alces-343	228	12	.	.	PROPN
alces-343	228	13	(	(	PUNCT
alces-343	228	14	2000	2000	NUM
alces-343	228	15	)	)	PUNCT
alces-343	228	16	and	and	CCONJ
alces-343	228	17	fuller	full	ADJ
alces-343	228	18	(	(	PUNCT
alces-343	228	19	1989	1989	NUM
alces-343	228	20	)	)	PUNCT
alces-343	228	21	are	be	AUX
alces-343	228	22	correct	correct	ADJ
alces-343	228	23	,	,	PUNCT
alces-343	228	24	more	more	ADV
alces-343	228	25	restrictive	restrictive	ADJ
alces-343	228	26	measures	measure	NOUN
alces-343	228	27	would	would	AUX
alces-343	228	28	be	be	AUX
alces-343	228	29	needed	need	VERB
alces-343	228	30	to	to	PART
alces-343	228	31	conserve	conserve	VERB
alces-343	228	32	caribou	caribou	NOUN
alces-343	228	33	in	in	ADP
alces-343	228	34	the	the	DET
alces-343	228	35	presence	presence	NOUN
alces-343	228	36	of	of	ADP
alces-343	228	37	moose	moose	NOUN
alces-343	228	38	and	and	CCONJ
alces-343	228	39	wolves	wolf	NOUN
alces-343	228	40	.	.	PUNCT
alces-343	229	1	similarly	similarly	ADV
alces-343	229	2	,	,	PUNCT
alces-343	229	3	the	the	DET
alces-343	229	4	population	population	NOUN
alces-343	229	5	growth	growth	NOUN
alces-343	229	6	rate	rate	NOUN
alces-343	229	7	for	for	ADP
alces-343	229	8	both	both	PRON
alces-343	229	9	by	by	ADP
alces-343	229	10	certain	certain	ADJ
alces-343	229	11	environmental	environmental	ADJ
alces-343	229	12	variables	variable	NOUN
alces-343	229	13	,	,	PUNCT
alces-343	229	14	such	such	ADJ
alces-343	229	15	as	as	ADP
alces-343	229	16	snow	snow	NOUN
alces-343	229	17	conditions	condition	NOUN
alces-343	229	18	(	(	PUNCT
alces-343	229	19	schaefer	schaefer	NOUN
alces-343	229	20	and	and	CCONJ
alces-343	229	21	messier	messy	ADJ
alces-343	229	22	1991	1991	NUM
alces-343	229	23	)	)	PUNCT
alces-343	229	24	.	.	PUNCT
alces-343	230	1	therefore	therefore	ADV
alces-343	230	2	,	,	PUNCT
alces-343	230	3	simulations	simulation	NOUN
alces-343	230	4	that	that	PRON
alces-343	230	5	do	do	AUX
alces-343	230	6	not	not	PART
alces-343	230	7	include	include	VERB
alces-343	230	8	the	the	DET
alces-343	230	9	impact	impact	NOUN
alces-343	230	10	of	of	ADP
alces-343	230	11	such	such	ADJ
alces-343	230	12	stochastic	stochastic	ADJ
alces-343	230	13	limiting	limit	VERB
alces-343	230	14	factors	factor	NOUN
alces-343	230	15	are	be	AUX
alces-343	230	16	likely	likely	ADJ
alces-343	230	17	to	to	PART
alces-343	230	18	yield	yield	VERB
alces-343	230	19	optimistic	optimistic	ADJ
alces-343	230	20	results	result	NOUN
alces-343	230	21	.	.	PUNCT
alces-343	231	1	bergerud	bergerud	NOUN
alces-343	231	2	and	and	CCONJ
alces-343	231	3	elliot	elliot	PROPN
alces-343	231	4	(	(	PUNCT
alces-343	231	5	1986	1986	NUM
alces-343	231	6	)	)	PUNCT
alces-343	231	7	estimated	estimate	VERB
alces-343	231	8	that	that	SCONJ
alces-343	231	9	caribou	caribou	PROPN
alces-343	231	10	abundance	abundance	NOUN
alces-343	231	11	should	should	AUX
alces-343	231	12	decline	decline	VERB
alces-343	231	13	when	when	SCONJ
alces-343	231	14	wolf	wolf	PROPN
alces-343	231	15	density	density	NOUN
alces-343	231	16	exceeds	exceed	VERB
alces-343	231	17	0.65	0.65	NUM
alces-343	231	18	individuals	individual	NOUN
alces-343	231	19	per	per	ADP
alces-343	231	20	100	100	NUM
alces-343	231	21	km2	km2	NOUN
alces-343	231	22	.	.	PUNCT
alces-343	232	1	in	in	ADP
alces-343	232	2	our	our	PRON
alces-343	232	3	simulations	simulation	NOUN
alces-343	232	4	,	,	PUNCT
alces-343	232	5	caribou	caribou	NOUN
alces-343	232	6	numbers	number	NOUN
alces-343	232	7	started	start	VERB
alces-343	232	8	declining	decline	VERB
alces-343	232	9	as	as	ADV
alces-343	232	10	soon	soon	ADV
alces-343	232	11	as	as	SCONJ
alces-343	232	12	the	the	DET
alces-343	232	13	wolf	wolf	PROPN
alces-343	232	14	population	population	NOUN
alces-343	232	15	exceeded	exceed	VERB
alces-343	232	16	0.45	0.45	NUM
alces-343	232	17	individuals	individual	NOUN
alces-343	232	18	per	per	ADP
alces-343	232	19	100	100	NUM
alces-343	232	20	km2	km2	NOUN
alces-343	232	21	.	.	PUNCT
alces-343	233	1	caribou	caribou	PROPN
alces-343	233	2	declines	decline	NOUN
alces-343	233	3	always	always	ADV
alces-343	233	4	followed	follow	VERB
alces-343	233	5	an	an	DET
alces-343	233	6	expansion	expansion	NOUN
alces-343	233	7	phase	phase	NOUN
alces-343	233	8	,	,	PUNCT
alces-343	233	9	which	which	PRON
alces-343	233	10	means	mean	VERB
alces-343	233	11	that	that	SCONJ
alces-343	233	12	food	food	NOUN
alces-343	233	13	density	density	NOUN
alces-343	233	14	-	-	PUNCT
alces-343	233	15	dependent	dependent	ADJ
alces-343	233	16	mechanisms	mechanism	NOUN
alces-343	233	17	intervened	intervene	VERB
alces-343	233	18	when	when	SCONJ
alces-343	233	19	caribou	caribou	NOUN
alces-343	233	20	numbers	number	NOUN
alces-343	233	21	increased	increase	VERB
alces-343	233	22	and	and	CCONJ
alces-343	233	23	that	that	SCONJ
alces-343	233	24	these	these	DET
alces-343	233	25	mechanisms	mechanism	NOUN
alces-343	233	26	were	be	AUX
alces-343	233	27	additive	additive	ADJ
alces-343	233	28	to	to	ADP
alces-343	233	29	predation	predation	NOUN
alces-343	233	30	effects	effect	NOUN
alces-343	233	31	.	.	PUNCT
alces-343	234	1	the	the	DET
alces-343	234	2	carrying	carry	VERB
alces-343	234	3	capacity	capacity	NOUN
alces-343	234	4	that	that	PRON
alces-343	234	5	we	we	PRON
alces-343	234	6	selected	select	VERB
alces-343	234	7	(	(	PUNCT
alces-343	234	8	20	20	NUM
alces-343	234	9	caribou	caribou	NOUN
alces-343	234	10	per	per	ADP
alces-343	234	11	100	100	NUM
alces-343	234	12	km2	km2	NOUN
alces-343	234	13	,	,	PUNCT
alces-343	234	14	arsenault	arsenault	PROPN
alces-343	234	15	et	et	PROPN
alces-343	234	16	al	al	PROPN
alces-343	234	17	.	.	PROPN
alces-343	234	18	1997	1997	NUM
alces-343	234	19	)	)	PUNCT
alces-343	234	20	could	could	AUX
alces-343	234	21	also	also	ADV
alces-343	234	22	be	be	AUX
alces-343	234	23	too	too	ADV
alces-343	234	24	high	high	ADJ
alces-343	234	25	based	base	VERB
alces-343	234	26	on	on	ADP
alces-343	234	27	recent	recent	ADJ
alces-343	234	28	estimates	estimate	NOUN
alces-343	234	29	(	(	PUNCT
alces-343	234	30	4.1	4.1	NUM
alces-343	234	31	–	–	PUNCT
alces-343	234	32	7.7	7.7	NUM
alces-343	234	33	individuals	individual	NOUN
alces-343	234	34	per	per	ADP
alces-343	234	35	100	100	NUM
alces-343	234	36	km2	km2	NOUN
alces-343	234	37	,	,	PUNCT
alces-343	234	38	courtois	courtois	PROPN
alces-343	234	39	2003	2003	NUM
alces-343	234	40	)	)	PUNCT
alces-343	234	41	,	,	PUNCT
alces-343	234	42	and	and	CCONJ
alces-343	234	43	recent	recent	ADJ
alces-343	234	44	studies	study	NOUN
alces-343	234	45	suggest	suggest	VERB
alces-343	234	46	that	that	SCONJ
alces-343	234	47	caribou	caribou	PROPN
alces-343	234	48	avoid	avoid	VERB
alces-343	234	49	human	human	NOUN
alces-343	234	50	-	-	PUNCT
alces-343	234	51	related	relate	VERB
alces-343	234	52	infrastructures	infrastructure	NOUN
alces-343	234	53	leading	lead	VERB
alces-343	234	54	to	to	ADP
alces-343	234	55	loss	loss	NOUN
alces-343	234	56	of	of	ADP
alces-343	234	57	available	available	ADJ
alces-343	234	58	habitat	habitat	NOUN
alces-343	234	59	(	(	PUNCT
alces-343	234	60	see	see	VERB
alces-343	234	61	weclaw	weclaw	NOUN
alces-343	234	62	and	and	CCONJ
alces-343	234	63	hudson	hudson	PROPN
alces-343	234	64	2004	2004	NUM
alces-343	234	65	for	for	ADP
alces-343	234	66	a	a	DET
alces-343	234	67	review	review	NOUN
alces-343	234	68	)	)	PUNCT
alces-343	234	69	.	.	PUNCT
alces-343	235	1	black	black	ADJ
alces-343	235	2	bear	bear	NOUN
alces-343	235	3	(	(	PUNCT
alces-343	235	4	ursus	ursus	PROPN
alces-343	235	5	americanus	americanus	PROPN
alces-343	235	6	)	)	PUNCT
alces-343	235	7	are	be	AUX
alces-343	235	8	not	not	PART
alces-343	235	9	abundant	abundant	ADJ
alces-343	235	10	in	in	ADP
alces-343	235	11	prime	prime	ADJ
alces-343	235	12	caribou	caribou	PROPN
alces-343	235	13	habitat	habitat	NOUN
alces-343	235	14	,	,	PUNCT
alces-343	235	15	but	but	CCONJ
alces-343	235	16	this	this	DET
alces-343	235	17	species	specie	NOUN
alces-343	235	18	can	can	AUX
alces-343	235	19	exert	exert	VERB
alces-343	235	20	an	an	DET
alces-343	235	21	important	important	ADJ
alces-343	235	22	additional	additional	ADJ
alces-343	235	23	density	density	NOUN
alces-343	235	24	-	-	PUNCT
alces-343	235	25	independent	independent	ADJ
alces-343	235	26	pressure	pressure	NOUN
alces-343	235	27	on	on	ADP
alces-343	235	28	caribou	caribou	NOUN
alces-343	235	29	in	in	ADP
alces-343	235	30	some	some	DET
alces-343	235	31	areas	area	NOUN
alces-343	235	32	(	(	PUNCT
alces-343	235	33	ballard	ballard	PROPN
alces-343	235	34	1994	1994	NUM
alces-343	235	35	)	)	PUNCT
alces-343	235	36	,	,	PUNCT
alces-343	235	37	particularly	particularly	ADV
alces-343	235	38	the	the	DET
alces-343	235	39	southern	southern	ADJ
alces-343	235	40	region	region	NOUN
alces-343	235	41	of	of	ADP
alces-343	235	42	the	the	DET
alces-343	235	43	boreal	boreal	NOUN
alces-343	235	44	forest	forest	NOUN
alces-343	235	45	.	.	PUNCT
alces-343	236	1	therefore	therefore	ADV
alces-343	236	2	,	,	PUNCT
alces-343	236	3	the	the	DET
alces-343	236	4	model	model	NOUN
alces-343	236	5	may	may	AUX
alces-343	236	6	be	be	AUX
alces-343	236	7	inappropriate	inappropriate	ADJ
alces-343	236	8	in	in	ADP
alces-343	236	9	areas	area	NOUN
alces-343	236	10	with	with	ADP
alces-343	236	11	bears	bear	NOUN
alces-343	236	12	,	,	PUNCT
alces-343	236	13	cougars	cougar	NOUN
alces-343	236	14	,	,	PUNCT
alces-343	236	15	or	or	CCONJ
alces-343	236	16	other	other	ADJ
alces-343	236	17	predators	predator	NOUN
alces-343	236	18	.	.	PUNCT
alces-343	237	1	taken	take	VERB
alces-343	237	2	management	management	NOUN
alces-343	237	3	decisions	decision	NOUN
alces-343	237	4	.	.	PUNCT
alces-343	238	1	model	model	NOUN
alces-343	238	2	improvements	improvement	NOUN
alces-343	238	3	improvements	improvement	NOUN
alces-343	238	4	could	could	AUX
alces-343	238	5	be	be	AUX
alces-343	238	6	made	make	VERB
alces-343	238	7	to	to	ADP
alces-343	238	8	our	our	PRON
alces-343	238	9	model	model	NOUN
alces-343	238	10	.	.	PUNCT
alces-343	239	1	for	for	ADP
alces-343	239	2	example	example	NOUN
alces-343	239	3	,	,	PUNCT
alces-343	239	4	the	the	DET
alces-343	239	5	wolf	wolf	PROPN
alces-343	239	6	population	population	NOUN
alces-343	239	7	could	could	AUX
alces-343	239	8	be	be	AUX
alces-343	239	9	considered	consider	VERB
alces-343	239	10	as	as	ADP
alces-343	239	11	a	a	DET
alces-343	239	12	reservoir	reservoir	NOUN
alces-343	239	13	,	,	PUNCT
alces-343	239	14	rather	rather	ADV
alces-343	239	15	than	than	ADP
alces-343	239	16	as	as	ADP
alces-343	239	17	a	a	DET
alces-343	239	18	parameter	parameter	NOUN
alces-343	239	19	linked	link	VERB
alces-343	239	20	to	to	ADP
alces-343	239	21	the	the	DET
alces-343	239	22	moose	moose	NOUN
alces-343	239	23	population	population	NOUN
alces-343	239	24	.	.	PUNCT
alces-343	240	1	this	this	DET
alces-343	240	2	change	change	NOUN
alces-343	240	3	would	would	AUX
alces-343	240	4	allow	allow	VERB
alces-343	240	5	consideration	consideration	NOUN
alces-343	240	6	of	of	ADP
alces-343	240	7	other	other	ADJ
alces-343	240	8	variables	variable	NOUN
alces-343	240	9	that	that	PRON
alces-343	240	10	affect	affect	VERB
alces-343	240	11	wolves	wolf	NOUN
alces-343	240	12	,	,	PUNCT
alces-343	240	13	such	such	ADJ
alces-343	240	14	as	as	ADP
alces-343	240	15	the	the	DET
alces-343	240	16	presence	presence	NOUN
alces-343	240	17	of	of	ADP
alces-343	240	18	other	other	ADJ
alces-343	240	19	prey	prey	NOUN
alces-343	240	20	species	specie	NOUN
alces-343	240	21	or	or	CCONJ
alces-343	240	22	demographic	demographic	ADJ
alces-343	240	23	uncontrolled	uncontrolled	ADJ
alces-343	240	24	variables	variable	NOUN
alces-343	240	25	and	and	CCONJ
alces-343	240	26	measurement	measurement	NOUN
alces-343	240	27	errors	error	NOUN
alces-343	240	28	were	be	AUX
alces-343	240	29	responsible	responsible	ADJ
alces-343	240	30	for	for	ADP
alces-343	240	31	approximately	approximately	ADV
alces-343	240	32	40	40	NUM
alces-343	240	33	%	%	NOUN
alces-343	240	34	of	of	ADP
alces-343	240	35	the	the	DET
alces-343	240	36	variance	variance	NOUN
alces-343	240	37	in	in	ADP
alces-343	240	38	wolf	wolf	PROPN
alces-343	240	39	density	density	NOUN
alces-343	240	40	between	between	ADP
alces-343	240	41	studies	study	NOUN
alces-343	240	42	consulted	consult	VERB
alces-343	240	43	by	by	ADP
alces-343	240	44	messier	messy	ADJ
alces-343	240	45	(	(	PUNCT
alces-343	240	46	1994	1994	NUM
alces-343	240	47	)	)	PUNCT
alces-343	240	48	.	.	PUNCT
alces-343	241	1	the	the	DET
alces-343	241	2	regulation	regulation	NOUN
alces-343	241	3	mechanism	mechanism	NOUN
alces-343	241	4	between	between	ADP
alces-343	241	5	moose	moose	NOUN
alces-343	241	6	and	and	CCONJ
alces-343	241	7	wolves	wolf	NOUN
alces-343	241	8	in	in	ADP
alces-343	241	9	our	our	PRON
alces-343	241	10	alces	alce	NOUN
alces-343	241	11	vol	vol	NOUN
alces-343	241	12	.	.	PROPN
alces-343	241	13	43	43	NUM
alces-343	241	14	,	,	PUNCT
alces-343	241	15	2007	2007	NUM
alces-343	241	16	courtois	courtois	NOUN
alces-343	241	17	and	and	CCONJ
alces-343	241	18	ouellet	ouellet	NOUN
alces-343	241	19	impact	impact	NOUN
alces-343	241	20	of	of	ADP
alces-343	241	21	moose	moose	NOUN
alces-343	241	22	and	and	CCONJ
alces-343	241	23	wolves	wolf	NOUN
alces-343	241	24	on	on	ADP
alces-343	241	25	caribou	caribou	NOUN
alces-343	241	26	23	23	NUM
alces-343	241	27	model	model	NOUN
alces-343	241	28	,	,	PUNCT
alces-343	241	29	however	however	ADV
alces-343	241	30	,	,	PUNCT
alces-343	241	31	seems	seem	VERB
alces-343	241	32	more	more	ADV
alces-343	241	33	appropriate	appropriate	ADJ
alces-343	241	34	to	to	PART
alces-343	241	35	mimic	mimic	VERB
alces-343	241	36	natural	natural	ADJ
alces-343	241	37	predator	predator	NOUN
alces-343	241	38	-	-	PUNCT
alces-343	241	39	prey	prey	NOUN
alces-343	241	40	dynamics	dynamic	NOUN
alces-343	241	41	than	than	ADP
alces-343	241	42	a	a	DET
alces-343	241	43	priori	priori	ADJ
alces-343	241	44	parameters	parameter	NOUN
alces-343	241	45	.	.	PUNCT
alces-343	242	1	furthermore	furthermore	ADV
alces-343	242	2	,	,	PUNCT
alces-343	242	3	instead	instead	ADV
alces-343	242	4	of	of	ADP
alces-343	242	5	considering	consider	VERB
alces-343	242	6	ungulate	ungulate	ADJ
alces-343	242	7	populations	population	NOUN
alces-343	242	8	as	as	ADP
alces-343	242	9	homogeneous	homogeneous	ADJ
alces-343	242	10	reservoirs	reservoir	NOUN
alces-343	242	11	,	,	PUNCT
alces-343	242	12	we	we	PRON
alces-343	242	13	could	could	AUX
alces-343	242	14	categorize	categorize	VERB
alces-343	242	15	populations	population	NOUN
alces-343	242	16	by	by	ADP
alces-343	242	17	age	age	NOUN
alces-343	242	18	and	and	CCONJ
alces-343	242	19	only	only	ADV
alces-343	242	20	if	if	SCONJ
alces-343	242	21	proportions	proportion	NOUN
alces-343	242	22	of	of	ADP
alces-343	242	23	males	male	NOUN
alces-343	242	24	and	and	CCONJ
alces-343	242	25	females	female	NOUN
alces-343	242	26	vary	vary	VERB
alces-343	242	27	substantially	substantially	ADV
alces-343	242	28	.	.	PUNCT
alces-343	243	1	according	accord	VERB
alces-343	243	2	to	to	ADP
alces-343	243	3	sensitivity	sensitivity	NOUN
alces-343	243	4	analyses	analysis	NOUN
alces-343	243	5	performed	perform	VERB
alces-343	243	6	by	by	ADP
alces-343	243	7	fancy	fancy	ADJ
alces-343	243	8	et	et	PROPN
alces-343	243	9	al	al	PROPN
alces-343	243	10	.	.	PROPN
alces-343	244	1	(	(	PUNCT
alces-343	244	2	1994	1994	NUM
alces-343	244	3	)	)	PUNCT
alces-343	244	4	,	,	PUNCT
alces-343	244	5	moose	moose	NOUN
alces-343	244	6	and	and	CCONJ
alces-343	244	7	caribou	caribou	NOUN
alces-343	244	8	population	population	NOUN
alces-343	244	9	growth	growth	NOUN
alces-343	244	10	rates	rate	NOUN
alces-343	244	11	are	be	AUX
alces-343	244	12	not	not	PART
alces-343	244	13	highly	highly	ADV
alces-343	244	14	sensitive	sensitive	ADJ
alces-343	244	15	to	to	ADP
alces-343	244	16	age	age	NOUN
alces-343	244	17	structure	structure	NOUN
alces-343	244	18	,	,	PUNCT
alces-343	244	19	but	but	CCONJ
alces-343	244	20	rather	rather	ADV
alces-343	244	21	depend	depend	VERB
alces-343	244	22	on	on	ADP
alces-343	244	23	recruitment	recruitment	NOUN
alces-343	244	24	rate	rate	NOUN
alces-343	244	25	,	,	PUNCT
alces-343	244	26	which	which	PRON
alces-343	244	27	is	be	AUX
alces-343	244	28	strongly	strongly	ADV
alces-343	244	29	and	and	CCONJ
alces-343	244	30	their	their	PRON
alces-343	244	31	survival	survival	NOUN
alces-343	244	32	.	.	PUNCT
alces-343	245	1	more	more	ADV
alces-343	245	2	importantly	importantly	ADV
alces-343	245	3	,	,	PUNCT
alces-343	245	4	a	a	DET
alces-343	245	5	spatially	spatially	ADV
alces-343	245	6	explicit	explicit	ADJ
alces-343	245	7	model	model	NOUN
alces-343	245	8	would	would	AUX
alces-343	245	9	be	be	AUX
alces-343	245	10	particularly	particularly	ADV
alces-343	245	11	useful	useful	ADJ
alces-343	245	12	to	to	PART
alces-343	245	13	investigate	investigate	VERB
alces-343	245	14	the	the	DET
alces-343	245	15	possibility	possibility	NOUN
alces-343	245	16	of	of	ADP
alces-343	245	17	excluding	exclude	VERB
alces-343	245	18	wolves	wolf	NOUN
alces-343	245	19	from	from	ADP
alces-343	245	20	areas	area	NOUN
alces-343	245	21	used	use	VERB
alces-343	245	22	by	by	ADP
alces-343	245	23	caribou	caribou	PROPN
alces-343	245	24	,	,	PUNCT
alces-343	245	25	for	for	ADP
alces-343	245	26	example	example	NOUN
alces-343	245	27	through	through	ADP
alces-343	245	28	habitat	habitat	ADJ
alces-343	245	29	management	management	NOUN
alces-343	245	30	that	that	PRON
alces-343	245	31	renders	render	VERB
alces-343	245	32	areas	area	NOUN
alces-343	245	33	unsuitable	unsuitable	ADJ
alces-343	245	34	for	for	ADP
alces-343	245	35	moose	moose	NOUN
alces-343	245	36	.	.	PUNCT
alces-343	246	1	such	such	DET
alces-343	246	2	a	a	DET
alces-343	246	3	model	model	NOUN
alces-343	246	4	could	could	AUX
alces-343	246	5	be	be	AUX
alces-343	246	6	based	base	VERB
alces-343	246	7	on	on	ADP
alces-343	246	8	the	the	DET
alces-343	246	9	movement	movement	NOUN
alces-343	246	10	rate	rate	NOUN
alces-343	246	11	and	and	CCONJ
alces-343	246	12	direction	direction	NOUN
alces-343	246	13	of	of	ADP
alces-343	246	14	wolves	wolf	NOUN
alces-343	246	15	could	could	AUX
alces-343	246	16	help	help	VERB
alces-343	246	17	to	to	PART
alces-343	246	18	determine	determine	VERB
alces-343	246	19	the	the	DET
alces-343	246	20	minimal	minimal	ADJ
alces-343	246	21	size	size	NOUN
alces-343	246	22	of	of	ADP
alces-343	246	23	a	a	DET
alces-343	246	24	protected	protect	VERB
alces-343	246	25	area	area	NOUN
alces-343	246	26	for	for	ADP
alces-343	246	27	caribou	caribou	NOUN
alces-343	246	28	that	that	PRON
alces-343	246	29	would	would	AUX
alces-343	246	30	permit	permit	VERB
alces-343	246	31	avoidance	avoidance	NOUN
alces-343	246	32	of	of	ADP
alces-343	246	33	most	most	ADJ
alces-343	246	34	encounters	encounter	NOUN
alces-343	246	35	between	between	ADP
alces-343	246	36	wolves	wolf	NOUN
alces-343	246	37	and	and	CCONJ
alces-343	246	38	caribou	caribou	NOUN
alces-343	246	39	.	.	PUNCT
alces-343	247	1	acknowledgements	acknowledgement	NOUN
alces-343	247	2	we	we	PRON
alces-343	247	3	wish	wish	VERB
alces-343	247	4	to	to	PART
alces-343	247	5	thank	thank	VERB
alces-343	247	6	pierre	pierre	NOUN
alces-343	247	7	etcheverry	etcheverry	VERB
alces-343	247	8	for	for	ADP
alces-343	247	9	helpful	helpful	ADJ
alces-343	247	10	insights	insight	NOUN
alces-343	247	11	during	during	ADP
alces-343	247	12	elaboration	elaboration	NOUN
alces-343	247	13	of	of	ADP
alces-343	247	14	the	the	DET
alces-343	247	15	model	model	NOUN
alces-343	247	16	and	and	CCONJ
alces-343	247	17	selection	selection	NOUN
alces-343	247	18	of	of	ADP
alces-343	247	19	the	the	DET
alces-343	247	20	parameters	parameter	NOUN
alces-343	247	21	.	.	PUNCT
alces-343	248	1	we	we	PRON
alces-343	248	2	also	also	ADV
alces-343	248	3	address	address	VERB
alces-343	248	4	special	special	ADJ
alces-343	248	5	thanks	thank	NOUN
alces-343	248	6	to	to	ADP
alces-343	248	7	pierre	pierre	PROPN
alces-343	248	8	drapeau	drapeau	PROPN
alces-343	248	9	,	,	PUNCT
alces-343	248	10	robert	robert	PROPN
alces-343	248	11	hudson	hudson	PROPN
alces-343	248	12	,	,	PUNCT
alces-343	248	13	françois	françois	PROPN
alces-343	248	14	potvin	potvin	PROPN
alces-343	248	15	,	,	PUNCT
alces-343	248	16	jim	jim	PROPN
alces-343	248	17	schaefer	schaefer	PROPN
alces-343	248	18	,	,	PUNCT
alces-343	248	19	and	and	CCONJ
alces-343	248	20	two	two	NUM
alces-343	248	21	anonymous	anonymous	ADJ
alces-343	248	22	reviewers	reviewer	NOUN
alces-343	248	23	for	for	ADP
alces-343	248	24	their	their	PRON
alces-343	248	25	helpthe	helpthe	PRON
alces-343	248	26	ministère	ministère	PROPN
alces-343	248	27	des	des	PROPN
alces-343	248	28	ressources	ressource	NOUN
alces-343	248	29	naturelles	naturelle	NOUN
alces-343	248	30	et	et	PROPN
alces-343	248	31	de	de	X
alces-343	248	32	la	la	PROPN
alces-343	248	33	faune	faune	PROPN
alces-343	248	34	,	,	PUNCT
alces-343	248	35	the	the	DET
alces-343	248	36	association	association	NOUN
alces-343	248	37	des	des	PROPN
alces-343	248	38	manufacturiers	manufacturiers	PROPN
alces-343	248	39	de	de	X
alces-343	248	40	bois	bois	X
alces-343	248	41	de	de	PROPN
alces-343	248	42	sciage	sciage	PROPN
alces-343	248	43	du	du	PROPN
alces-343	248	44	québec	québec	PROPN
alces-343	248	45	,	,	PUNCT
alces-343	248	46	the	the	DET
alces-343	248	47	fondation	fondation	PROPN
alces-343	248	48	de	de	PROPN
alces-343	248	49	la	la	PROPN
alces-343	248	50	faune	faune	PROPN
alces-343	248	51	du	du	PROPN
alces-343	248	52	québec	québec	PROPN
alces-343	248	53	,	,	PUNCT
alces-343	248	54	abitibi	abitibi	NOUN
alces-343	248	55	-	-	PUNCT
alces-343	248	56	consolidated	consolidated	PROPN
alces-343	248	57	inc	inc	PROPN
alces-343	248	58	.	.	PROPN
alces-343	248	59	,	,	PUNCT
alces-343	248	60	and	and	CCONJ
alces-343	248	61	kruger	kruger	PROPN
alces-343	248	62	(	(	PUNCT
alces-343	248	63	scierie	scierie	NOUN
alces-343	248	64	-	-	PUNCT
alces-343	248	65	manic	manic	ADJ
alces-343	248	66	)	)	PUNCT
alces-343	248	67	.	.	PUNCT
alces-343	249	1	references	reference	NOUN
alces-343	249	2	arsenault	arsenault	PROPN
alces-343	249	3	,	,	PUNCT
alces-343	249	4	d.	d.	PROPN
alces-343	249	5	,	,	PUNCT
alces-343	249	6	n.	n.	PROPN
alces-343	249	7	villeneuve	villeneuve	PROPN
alces-343	249	8	,	,	PUNCT
alces-343	249	9	c.	c.	PROPN
alces-343	249	10	boismenu	boismenu	PROPN
alces-343	249	11	,	,	PUNCT
alces-343	249	12	y.	y.	PROPN
alces-343	249	13	leblanc	leblanc	PROPN
alces-343	249	14	,	,	PUNCT
alces-343	249	15	and	and	CCONJ
alces-343	249	16	j.	j.	PROPN
alces-343	249	17	deshaye	deshaye	PROPN
alces-343	249	18	.	.	PUNCT
alces-343	250	1	1997	1997	NUM
alces-343	250	2	.	.	PUNCT
alces-343	251	1	estimating	estimate	VERB
alces-343	251	2	lichen	lichen	NOUN
alces-343	251	3	biomass	biomass	NOUN
alces-343	251	4	and	and	CCONJ
alces-343	251	5	caribou	caribou	NOUN
alces-343	251	6	grazing	grazing	NOUN
alces-343	251	7	on	on	ADP
alces-343	251	8	the	the	DET
alces-343	251	9	winter	winter	NOUN
alces-343	251	10	grounds	ground	NOUN
alces-343	251	11	of	of	ADP
alces-343	251	12	northern	northern	ADJ
alces-343	251	13	québec	québec	NOUN
alces-343	251	14	:	:	PUNCT
alces-343	251	15	an	an	DET
alces-343	251	16	application	application	NOUN
alces-343	251	17	of	of	ADP
alces-343	251	18	fire	fire	NOUN
alces-343	251	19	history	history	NOUN
alces-343	251	20	and	and	CCONJ
alces-343	251	21	landsat	landsat	PROPN
alces-343	251	22	data	data	PROPN
alces-343	251	23	.	.	PUNCT
alces-343	252	1	journal	journal	PROPN
alces-343	252	2	of	of	ADP
alces-343	252	3	applied	applied	ADJ
alces-343	252	4	ecology	ecology	NOUN
alces-343	252	5	34:65	34:65	NUM
alces-343	252	6	-	-	SYM
alces-343	252	7	78	78	NUM
alces-343	252	8	.	.	PUNCT
alces-343	253	1	ballard	ballard	PROPN
alces-343	253	2	,	,	PUNCT
alces-343	253	3	w.	w.	PROPN
alces-343	253	4	b.	b.	PROPN
alces-343	253	5	1994	1994	NUM
alces-343	253	6	.	.	PUNCT
alces-343	254	1	effects	effect	NOUN
alces-343	254	2	of	of	ADP
alces-343	254	3	black	black	ADJ
alces-343	254	4	bear	bear	NOUN
alces-343	254	5	predation	predation	NOUN
alces-343	254	6	on	on	ADP
alces-343	254	7	caribou	caribou	PROPN
alces-343	254	8	–	–	PUNCT
alces-343	254	9	a	a	DET
alces-343	254	10	review	review	NOUN
alces-343	254	11	.	.	PUNCT
alces-343	255	1	alces	alce	NOUN
alces-343	255	2	30:25	30:25	NUM
alces-343	255	3	-	-	SYM
alces-343	255	4	35	35	NUM
alces-343	255	5	.	.	PUNCT
alces-343	256	1	banfield	banfield	PROPN
alces-343	256	2	,	,	PUNCT
alces-343	256	3	a.	a.	PROPN
alces-343	256	4	w.	w.	PROPN
alces-343	256	5	f.	f.	PROPN
alces-343	256	6	1974	1974	NUM
alces-343	256	7	.	.	PUNCT
alces-343	257	1	les	les	PROPN
alces-343	257	2	mammifères	mammifère	NOUN
alces-343	257	3	du	du	PROPN
alces-343	257	4	canada	canada	PROPN
alces-343	257	5	.	.	PUNCT
alces-343	258	1	les	les	PROPN
alces-343	258	2	presses	press	NOUN
alces-343	258	3	de	de	X
alces-343	258	4	l’université	l’université	NOUN
alces-343	258	5	laval	laval	NOUN
alces-343	258	6	,	,	PUNCT
alces-343	258	7	québec	québec	PROPN
alces-343	258	8	,	,	PUNCT
alces-343	258	9	québec	québec	PROPN
alces-343	258	10	,	,	PUNCT
alces-343	258	11	canada	canada	PROPN
alces-343	258	12	.	.	PUNCT
alces-343	259	1	bergerud	bergerud	PROPN
alces-343	259	2	,	,	PUNCT
alces-343	259	3	a.	a.	NOUN
alces-343	259	4	t.	t.	PROPN
alces-343	259	5	1967	1967	NUM
alces-343	259	6	.	.	PUNCT
alces-343	260	1	management	management	NOUN
alces-343	260	2	of	of	ADP
alces-343	260	3	labrador	labrador	PROPN
alces-343	260	4	caribou	caribou	PROPN
alces-343	260	5	.	.	PUNCT
alces-343	261	1	journal	journal	PROPN
alces-343	261	2	of	of	ADP
alces-343	261	3	wildlife	wildlife	NOUN
alces-343	261	4	management	management	NOUN
alces-343	261	5	31:621	31:621	NUM
alces-343	261	6	-	-	SYM
alces-343	261	7	635	635	NUM
alces-343	261	8	.	.	PUNCT
alces-343	262	1	_	_	PUNCT
alces-343	263	1	_	_	PUNCT
alces-343	264	1	_	_	PUNCT
alces-343	265	1	_	_	PUNCT
alces-343	266	1	_	_	PUNCT
alces-343	266	2	.	.	PUNCT
alces-343	267	1	1985	1985	NUM
alces-343	267	2	.	.	PUNCT
alces-343	268	1	antipredator	antipredator	ADJ
alces-343	268	2	strategies	strategy	NOUN
alces-343	268	3	of	of	ADP
alces-343	268	4	caribou	caribou	NOUN
alces-343	268	5	:	:	PUNCT
alces-343	268	6	dispersion	dispersion	NOUN
alces-343	268	7	along	along	ADP
alces-343	268	8	shorelines	shoreline	NOUN
alces-343	268	9	.	.	PUNCT
alces-343	269	1	canadian	canadian	ADJ
alces-343	269	2	journal	journal	PROPN
alces-343	269	3	of	of	ADP
alces-343	269	4	zoology	zoology	NOUN
alces-343	269	5	63:1324	63:1324	PROPN
alces-343	269	6	-	-	SYM
alces-343	269	7	1329	1329	NUM
alces-343	269	8	.	.	PUNCT
alces-343	270	1	_	_	PUNCT
alces-343	271	1	_	_	PUNCT
alces-343	272	1	_	_	PUNCT
alces-343	273	1	_	_	PUNCT
alces-343	274	1	_	_	PUNCT
alces-343	274	2	.	.	PUNCT
alces-343	275	1	1996	1996	NUM
alces-343	275	2	.	.	PUNCT
alces-343	276	1	evolving	evolve	VERB
alces-343	276	2	perspectives	perspective	NOUN
alces-343	276	3	on	on	ADP
alces-343	276	4	caribou	caribou	NOUN
alces-343	276	5	population	population	NOUN
alces-343	276	6	dynamics	dynamic	NOUN
alces-343	276	7	,	,	PUNCT
alces-343	276	8	have	have	AUX
alces-343	276	9	we	we	PRON
alces-343	276	10	got	get	VERB
alces-343	276	11	it	it	PRON
alces-343	276	12	right	right	ADV
alces-343	276	13	yet	yet	ADV
alces-343	276	14	?	?	PUNCT
alces-343	277	1	rangifer	rangifer	NOUN
alces-343	277	2	,	,	PUNCT
alces-343	277	3	special	special	ADJ
alces-343	277	4	issue	issue	NOUN
alces-343	277	5	9:95	9:95	NUM
alces-343	277	6	-	-	SYM
alces-343	277	7	116	116	NUM
alces-343	277	8	.	.	PUNCT
alces-343	278	1	_	_	PUNCT
alces-343	279	1	_	_	PUNCT
alces-343	280	1	_	_	PUNCT
alces-343	281	1	_	_	PUNCT
alces-343	281	2	_	_	PUNCT
alces-343	281	3	,	,	PUNCT
alces-343	281	4	and	and	CCONJ
alces-343	281	5	j.	j.	PROPN
alces-343	281	6	p.	p.	PROPN
alces-343	281	7	elliot	elliot	PROPN
alces-343	281	8	.	.	PROPN
alces-343	282	1	1986	1986	NUM
alces-343	282	2	.	.	PUNCT
alces-343	283	1	dynamics	dynamic	NOUN
alces-343	283	2	of	of	ADP
alces-343	283	3	caribou	caribou	NOUN
alces-343	283	4	and	and	CCONJ
alces-343	283	5	wolves	wolf	NOUN
alces-343	283	6	in	in	ADP
alces-343	283	7	northern	northern	ADJ
alces-343	283	8	british	british	PROPN
alces-343	283	9	columbia	columbia	PROPN
alces-343	283	10	.	.	PUNCT
alces-343	284	1	canadian	canadian	ADJ
alces-343	284	2	journal	journal	PROPN
alces-343	284	3	of	of	ADP
alces-343	284	4	zoology	zoology	NOUN
alces-343	284	5	64:1515	64:1515	NOUN
alces-343	284	6	-	-	PUNCT
alces-343	284	7	1529	1529	NUM
alces-343	284	8	.	.	PUNCT
alces-343	285	1	_	_	PUNCT
alces-343	286	1	_	_	PUNCT
alces-343	287	1	_	_	PUNCT
alces-343	288	1	_	_	PUNCT
alces-343	289	1	_	_	PUNCT
alces-343	289	2	,	,	PUNCT
alces-343	289	3	r.	r.	PROPN
alces-343	289	4	d.	d.	PROPN
alces-343	289	5	jaminchuk	jaminchuk	PROPN
alces-343	289	6	,	,	PUNCT
alces-343	289	7	and	and	CCONJ
alces-343	289	8	d.	d.	PROPN
alces-343	289	9	r.	r.	PROPN
alces-343	289	10	carruthers	carruthers	PROPN
alces-343	289	11	.	.	PUNCT
alces-343	289	12	1984	1984	NUM
alces-343	289	13	.	.	PUNCT
alces-343	290	1	the	the	DET
alces-343	290	2	buffalo	buffalo	NOUN
alces-343	290	3	of	of	ADP
alces-343	290	4	the	the	DET
alces-343	290	5	north	north	NOUN
alces-343	290	6	:	:	PUNCT
alces-343	290	7	woodland	woodland	NOUN
alces-343	290	8	caribou	caribou	NOUN
alces-343	290	9	at	at	ADP
alces-343	290	10	calving	calving	NOUN
alces-343	290	11	.	.	PUNCT
alces-343	291	1	animal	animal	NOUN
alces-343	291	2	behavior	behavior	NOUN
alces-343	291	3	39:360	39:360	NUM
alces-343	291	4	-	-	SYM
alces-343	291	5	368	368	NUM
alces-343	291	6	.	.	PUNCT
alces-343	292	1	boertje	boertje	PROPN
alces-343	292	2	,	,	PUNCT
alces-343	292	3	r.	r.	PROPN
alces-343	292	4	d.	d.	PROPN
alces-343	292	5	,	,	PUNCT
alces-343	292	6	p.	p.	PROPN
alces-343	292	7	valkenburg	valkenburg	PROPN
alces-343	292	8	,	,	PUNCT
alces-343	292	9	and	and	CCONJ
alces-343	292	10	m.	m.	PROPN
alces-343	292	11	e.	e.	PROPN
alces-343	292	12	mcnay	mcnay	PROPN
alces-343	292	13	.	.	PUNCT
alces-343	293	1	1996	1996	NUM
alces-343	293	2	.	.	PUNCT
alces-343	294	1	increases	increase	NOUN
alces-343	294	2	in	in	ADP
alces-343	294	3	moose	moose	NOUN
alces-343	294	4	,	,	PUNCT
alces-343	294	5	caribou	caribou	NOUN
alces-343	294	6	,	,	PUNCT
alces-343	294	7	and	and	CCONJ
alces-343	294	8	wolves	wolf	NOUN
alces-343	294	9	following	follow	VERB
alces-343	294	10	wolf	wolf	PROPN
alces-343	294	11	control	control	NOUN
alces-343	294	12	in	in	ADP
alces-343	294	13	alaska	alaska	PROPN
alces-343	294	14	.	.	PUNCT
alces-343	295	1	journal	journal	PROPN
alces-343	295	2	of	of	ADP
alces-343	295	3	wildlife	wildlife	NOUN
alces-343	295	4	management	management	NOUN
alces-343	295	5	60:474	60:474	PROPN
alces-343	295	6	-	-	PUNCT
alces-343	295	7	489	489	NUM
alces-343	295	8	.	.	PUNCT
alces-343	296	1	brassard	brassard	PROPN
alces-343	296	2	,	,	PUNCT
alces-343	296	3	j.	j.	PROPN
alces-343	296	4	m.	m.	PROPN
alces-343	296	5	1967	1967	NUM
alces-343	296	6	.	.	PUNCT
alces-343	297	1	inventaire	inventaire	PROPN
alces-343	297	2	aérien	aérien	PROPN
alces-343	297	3	des	des	PROPN
alces-343	297	4	ongulés	ongulés	PROPN
alces-343	297	5	sauvages	sauvages	X
alces-343	297	6	de	de	X
alces-343	297	7	la	la	X
alces-343	297	8	côte	côte	NOUN
alces-343	297	9	-	-	PUNCT
alces-343	297	10	nord	nord	PROPN
alces-343	297	11	et	et	PROPN
alces-343	297	12	identification	identification	PROPN
alces-343	297	13	des	des	PROPN
alces-343	297	14	aires	aires	PROPN
alces-343	297	15	d’hivernement	d’hivernement	PROPN
alces-343	297	16	en	en	PROPN
alces-343	297	17	fonction	fonction	PROPN
alces-343	297	18	des	des	X
alces-343	297	19	formes	formes	PROPN
alces-343	297	20	du	du	PROPN
alces-343	297	21	relief	relief	PROPN
alces-343	297	22	et	et	PROPN
alces-343	297	23	de	de	PROPN
alces-343	297	24	la	la	PROPN
alces-343	297	25	végétation	végétation	NOUN
alces-343	297	26	.	.	PUNCT
alces-343	298	1	service	service	PROPN
alces-343	298	2	de	de	PROPN
alces-343	298	3	la	la	PROPN
alces-343	298	4	faune	faune	PROPN
alces-343	298	5	.	.	PUNCT
alces-343	299	1	québec	québec	PROPN
alces-343	299	2	,	,	PUNCT
alces-343	299	3	québec	québec	PROPN
alces-343	299	4	,	,	PUNCT
alces-343	299	5	canada	canada	PROPN
alces-343	299	6	.	.	PUNCT
alces-343	300	1	caughley	caughley	PROPN
alces-343	300	2	,	,	PUNCT
alces-343	300	3	g.	g.	PROPN
alces-343	300	4	1977	1977	NUM
alces-343	300	5	.	.	PUNCT
alces-343	301	1	analysis	analysis	NOUN
alces-343	301	2	of	of	ADP
alces-343	301	3	vertebrate	vertebrate	ADJ
alces-343	301	4	populations	population	NOUN
alces-343	301	5	.	.	PUNCT
alces-343	302	1	john	john	PROPN
alces-343	302	2	wiley	wiley	PROPN
alces-343	302	3	and	and	CCONJ
alces-343	302	4	sons	son	NOUN
alces-343	302	5	,	,	PUNCT
alces-343	302	6	london	london	PROPN
alces-343	302	7	,	,	PUNCT
alces-343	302	8	u.k	u.k	PROPN
alces-343	302	9	.	.	PROPN
alces-343	302	10	courtois	courtois	PROPN
alces-343	302	11	,	,	PUNCT
alces-343	302	12	r.	r.	PROPN
alces-343	302	13	2003	2003	NUM
alces-343	302	14	.	.	PUNCT
alces-343	303	1	la	la	PROPN
alces-343	303	2	conservation	conservation	PROPN
alces-343	303	3	du	du	PROPN
alces-343	303	4	caribou	caribou	PROPN
alces-343	303	5	forestier	forestier	PROPN
alces-343	303	6	dans	dans	PROPN
alces-343	303	7	un	un	PROPN
alces-343	303	8	contexte	contexte	PROPN
alces-343	303	9	de	de	X
alces-343	303	10	perte	perte	PROPN
alces-343	303	11	d’habitat	d’habitat	X
alces-343	303	12	et	et	PROPN
alces-343	303	13	de	de	X
alces-343	303	14	fragmentation	fragmentation	PROPN
alces-343	303	15	du	du	PROPN
alces-343	303	16	milieu	milieu	PROPN
alces-343	303	17	.	.	PUNCT
alces-343	304	1	impact	impact	NOUN
alces-343	304	2	of	of	ADP
alces-343	304	3	moose	moose	NOUN
alces-343	304	4	and	and	CCONJ
alces-343	304	5	wolves	wolf	NOUN
alces-343	304	6	on	on	ADP
alces-343	304	7	caribou	caribou	NOUN
alces-343	304	8	–	–	PUNCT
alces-343	304	9	courtois	courtois	NOUN
alces-343	304	10	and	and	CCONJ
alces-343	304	11	ouellet	ouellet	PROPN
alces-343	304	12	alces	alces	PROPN
alces-343	304	13	vol	vol	NOUN
alces-343	304	14	.	.	PROPN
alces-343	305	1	43	43	NUM
alces-343	305	2	,	,	PUNCT
alces-343	305	3	2007	2007	NUM
alces-343	305	4	24	24	NUM
alces-343	305	5	ph.d	ph.d	PROPN
alces-343	305	6	.	.	PUNCT
alces-343	306	1	thesis	thesis	NOUN
alces-343	306	2	.	.	PUNCT
alces-343	307	1	université	université	PROPN
alces-343	307	2	du	du	PROPN
alces-343	307	3	québec	québec	PROPN
alces-343	307	4	à	à	X
alces-343	307	5	rimouski	rimouski	NOUN
alces-343	307	6	,	,	PUNCT
alces-343	307	7	rimouski	rimouski	ADJ
alces-343	307	8	,	,	PUNCT
alces-343	307	9	québec	québec	PROPN
alces-343	307	10	,	,	PUNCT
alces-343	307	11	canada	canada	PROPN
alces-343	307	12	.	.	PUNCT
alces-343	308	1	_	_	PUNCT
alces-343	309	1	_	_	PUNCT
alces-343	310	1	_	_	PUNCT
alces-343	311	1	_	_	PUNCT
alces-343	312	1	_	_	PUNCT
alces-343	312	2	,	,	PUNCT
alces-343	312	3	m.	m.	NOUN
alces-343	312	4	crête	crête	PROPN
alces-343	312	5	,	,	PUNCT
alces-343	312	6	and	and	CCONJ
alces-343	312	7	f.	f.	PROPN
alces-343	312	8	barnard	barnard	PROPN
alces-343	312	9	.	.	PUNCT
alces-343	313	1	1993	1993	NUM
alces-343	313	2	.	.	PUNCT
alces-343	314	1	productivité	productivité	PROPN
alces-343	314	2	et	et	PROPN
alces-343	314	3	dynamique	dynamique	PROPN
alces-343	314	4	d’une	d’une	VERB
alces-343	314	5	population	population	NOUN
alces-343	314	6	d’orignaux	d’orignaux	PUNCT
alces-343	314	7	du	du	X
alces-343	314	8	sud	sud	X
alces-343	314	9	de	de	X
alces-343	314	10	la	la	PROPN
alces-343	314	11	taïga	taïga	PROPN
alces-343	314	12	québécoise	québécoise	PROPN
alces-343	314	13	.	.	PUNCT
alces-343	315	1	ministère	ministère	PROPN
alces-343	315	2	du	du	PROPN
alces-343	315	3	loisir	loisir	PROPN
alces-343	315	4	,	,	PUNCT
alces-343	315	5	de	de	X
alces-343	315	6	la	la	X
alces-343	315	7	chasse	chasse	PROPN
alces-343	315	8	et	et	PROPN
alces-343	315	9	de	de	X
alces-343	315	10	la	la	X
alces-343	315	11	pêche	pêche	PROPN
alces-343	315	12	.	.	PUNCT
alces-343	316	1	québec	québec	PROPN
alces-343	316	2	,	,	PUNCT
alces-343	316	3	québec	québec	PROPN
alces-343	316	4	,	,	PUNCT
alces-343	316	5	canada	canada	PROPN
alces-343	316	6	.	.	PUNCT
alces-343	317	1	_	_	PUNCT
alces-343	318	1	_	_	PUNCT
alces-343	319	1	_	_	PUNCT
alces-343	320	1	_	_	PUNCT
alces-343	321	1	_	_	PUNCT
alces-343	321	2	,	,	PUNCT
alces-343	321	3	a.	a.	NOUN
alces-343	321	4	gingras	gingras	PROPN
alces-343	321	5	,	,	PUNCT
alces-343	321	6	c.	c.	PROPN
alces-343	321	7	dussault	dussault	PROPN
alces-343	321	8	,	,	PUNCT
alces-343	321	9	l.	l.	PROPN
alces-343	321	10	breton	breton	PROPN
alces-343	321	11	,	,	PUNCT
alces-343	321	12	and	and	CCONJ
alces-343	321	13	j.-p	j.-p	PROPN
alces-343	321	14	.	.	PUNCT
alces-343	322	1	ouellet	ouellet	NOUN
alces-343	322	2	.	.	PUNCT
alces-343	323	1	2003	2003	NUM
alces-343	323	2	.	.	PUNCT
alces-343	324	1	an	an	DET
alces-343	324	2	aerial	aerial	ADJ
alces-343	324	3	survey	survey	NOUN
alces-343	324	4	technique	technique	NOUN
alces-343	324	5	for	for	ADP
alces-343	324	6	the	the	DET
alces-343	324	7	forest	forest	NOUN
alces-343	324	8	-	-	PUNCT
alces-343	324	9	dwelling	dwell	VERB
alces-343	324	10	ecotype	ecotype	NOUN
alces-343	324	11	of	of	ADP
alces-343	324	12	woodland	woodland	PROPN
alces-343	324	13	caribou	caribou	PROPN
alces-343	324	14	,	,	PUNCT
alces-343	324	15	rangifer	rangifer	VERB
alces-343	324	16	tarandus	tarandus	PROPN
alces-343	324	17	caribou	caribou	PROPN
alces-343	324	18	.	.	PUNCT
alces-343	325	1	canadian	canadian	ADJ
alces-343	325	2	field	field	NOUN
alces-343	325	3	-	-	PUNCT
alces-343	325	4	naturalist	naturalist	NOUN
alces-343	325	5	117:546	117:546	NUM
alces-343	325	6	-	-	PUNCT
alces-343	325	7	554	554	NUM
alces-343	325	8	.	.	PUNCT
alces-343	326	1	_	_	PUNCT
alces-343	327	1	_	_	PUNCT
alces-343	328	1	_	_	PUNCT
alces-343	329	1	_	_	PUNCT
alces-343	330	1	_	_	PUNCT
alces-343	330	2	,	,	PUNCT
alces-343	330	3	y.leblanc	y.leblanc	NOUN
alces-343	330	4	,	,	PUNCT
alces-343	330	5	j.	j.	PROPN
alces-343	330	6	maltais	maltais	PROPN
alces-343	330	7	,	,	PUNCT
alces-343	330	8	andh.crépeau	andh.crépeau	PROPN
alces-343	330	9	.	.	PROPN
alces-343	330	10	1994a	1994a	NUM
alces-343	330	11	.	.	PUNCT
alces-343	331	1	québec	québec	PROPN
alces-343	331	2	moose	moose	NOUN
alces-343	331	3	aerial	aerial	ADJ
alces-343	331	4	surveys	survey	NOUN
alces-343	331	5	:	:	PUNCT
alces-343	331	6	methods	method	NOUN
alces-343	331	7	to	to	PART
alces-343	331	8	estimate	estimate	VERB
alces-343	331	9	population	population	NOUN
alces-343	331	10	characteristics	characteristic	NOUN
alces-343	331	11	and	and	CCONJ
alces-343	331	12	improved	improved	ADJ
alces-343	331	13	sampling	sampling	NOUN
alces-343	331	14	srategies	srategie	NOUN
alces-343	331	15	.	.	PUNCT
alces-343	332	1	alces	alce	NOUN
alces-343	332	2	30:159	30:159	NUM
alces-343	332	3	-	-	SYM
alces-343	332	4	171	171	NUM
alces-343	332	5	.	.	PUNCT
alces-343	333	1	_	_	PUNCT
alces-343	334	1	_	_	PUNCT
alces-343	335	1	_	_	PUNCT
alces-343	336	1	_	_	PUNCT
alces-343	337	1	_	_	PUNCT
alces-343	337	2	,	,	PUNCT
alces-343	337	3	d.	d.	PROPN
alces-343	337	4	sigouin	sigouin	PROPN
alces-343	337	5	,	,	PUNCT
alces-343	337	6	j.	j.	PROPN
alces-343	338	1	-p	-p	PROPN
alces-343	338	2	.	.	PUNCT
alces-343	339	1	ouellet	ouellet	PROPN
alces-343	339	2	,	,	PUNCT
alces-343	339	3	a.	a.	NOUN
alces-343	339	4	beaumont	beaumont	NOUN
alces-343	339	5	,	,	PUNCT
alces-343	339	6	and	and	CCONJ
alces-343	339	7	m.	m.	NOUN
alces-343	339	8	crête	crête	PROPN
alces-343	339	9	.	.	PUNCT
alces-343	340	1	1994b	1994b	NUM
alces-343	340	2	.	.	PUNCT
alces-343	341	1	mortalité	mortalité	NOUN
alces-343	341	2	naturelle	naturelle	PROPN
alces-343	341	3	et	et	NOUN
alces-343	341	4	d’origine	d’origine	PROPN
alces-343	341	5	anthropique	anthropique	NOUN
alces-343	341	6	de	de	X
alces-343	341	7	l’orignal	l’orignal	PROPN
alces-343	341	8	au	au	PROPN
alces-343	341	9	québec	québec	PROPN
alces-343	341	10	.	.	PUNCT
alces-343	342	1	ministère	ministère	PROPN
alces-343	342	2	de	de	PROPN
alces-343	342	3	l’environnement	l’environnement	PROPN
alces-343	342	4	et	et	PROPN
alces-343	342	5	de	de	PROPN
alces-343	342	6	la	la	X
alces-343	342	7	faune	faune	PROPN
alces-343	342	8	.	.	PUNCT
alces-343	343	1	québec	québec	PROPN
alces-343	343	2	,	,	PUNCT
alces-343	343	3	québec	québec	PROPN
alces-343	343	4	,	,	PUNCT
alces-343	343	5	canada	canada	PROPN
alces-343	343	6	.	.	PUNCT
alces-343	344	1	crête	crête	PROPN
alces-343	344	2	,	,	PUNCT
alces-343	344	3	m.	m.	NOUN
alces-343	344	4	,	,	PUNCT
alces-343	344	5	and	and	CCONJ
alces-343	344	6	r.	r.	PROPN
alces-343	344	7	courtois	courtois	PROPN
alces-343	344	8	.	.	PUNCT
alces-343	345	1	1997	1997	NUM
alces-343	345	2	.	.	PUNCT
alces-343	346	1	limiting	limit	VERB
alces-343	346	2	factors	factor	NOUN
alces-343	346	3	might	might	AUX
alces-343	346	4	obscure	obscure	VERB
alces-343	346	5	population	population	NOUN
alces-343	346	6	regulation	regulation	NOUN
alces-343	346	7	of	of	ADP
alces-343	346	8	moose	moose	NOUN
alces-343	346	9	(	(	PUNCT
alces-343	346	10	cervidae	cervidae	NOUN
alces-343	346	11	:	:	PUNCT
alces-343	346	12	alces	alce	NOUN
alces-343	346	13	alces	alce	NOUN
alces-343	346	14	)	)	PUNCT
alces-343	346	15	in	in	ADP
alces-343	346	16	unproductive	unproductive	ADJ
alces-343	346	17	forests	forest	NOUN
alces-343	346	18	.	.	PUNCT
alces-343	347	1	journal	journal	PROPN
alces-343	347	2	of	of	ADP
alces-343	347	3	zoology	zoology	NOUN
alces-343	347	4	242:765	242:765	PROPN
alces-343	347	5	-	-	PUNCT
alces-343	347	6	781	781	NUM
alces-343	347	7	.	.	PUNCT
alces-343	348	1	_	_	PUNCT
alces-343	349	1	_	_	PUNCT
alces-343	350	1	_	_	PUNCT
alces-343	351	1	_	_	PUNCT
alces-343	352	1	_	_	PUNCT
alces-343	352	2	,	,	PUNCT
alces-343	352	3	and	and	CCONJ
alces-343	352	4	j.	j.	PROPN
alces-343	352	5	doucet	doucet	PROPN
alces-343	352	6	.	.	PUNCT
alces-343	353	1	1998	1998	NUM
alces-343	353	2	.	.	PUNCT
alces-343	354	1	persistent	persistent	ADJ
alces-343	354	2	suppression	suppression	NOUN
alces-343	354	3	in	in	ADP
alces-343	354	4	dwarf	dwarf	NOUN
alces-343	354	5	birch	birch	NOUN
alces-343	354	6	after	after	ADP
alces-343	354	7	release	release	NOUN
alces-343	354	8	from	from	ADP
alces-343	354	9	heavy	heavy	ADJ
alces-343	354	10	summer	summer	NOUN
alces-343	354	11	browsing	browse	VERB
alces-343	354	12	by	by	ADP
alces-343	354	13	caribou	caribou	PROPN
alces-343	354	14	.	.	PUNCT
alces-343	355	1	arctic	arctic	PROPN
alces-343	355	2	and	and	CCONJ
alces-343	355	3	alpine	alpine	PROPN
alces-343	355	4	research	research	NOUN
alces-343	355	5	30:126	30:126	NUM
alces-343	355	6	-	-	SYM
alces-343	355	7	132	132	NUM
alces-343	355	8	.	.	PUNCT
alces-343	356	1	_	_	PUNCT
alces-343	357	1	_	_	PUNCT
alces-343	358	1	_	_	PUNCT
alces-343	359	1	_	_	PUNCT
alces-343	360	1	_	_	PUNCT
alces-343	360	2	,	,	PUNCT
alces-343	360	3	and	and	CCONJ
alces-343	360	4	m.	m.	NOUN
alces-343	360	5	manseau	manseau	PROPN
alces-343	360	6	.	.	PUNCT
alces-343	361	1	1996	1996	NUM
alces-343	361	2	.	.	PUNCT
alces-343	362	1	natural	natural	ADJ
alces-343	362	2	regulation	regulation	NOUN
alces-343	362	3	of	of	ADP
alces-343	362	4	cervidae	cervidae	NOUN
alces-343	362	5	along	along	ADP
alces-343	362	6	a	a	DET
alces-343	362	7	1000	1000	NUM
alces-343	362	8	km	km	NOUN
alces-343	362	9	latitudinal	latitudinal	ADJ
alces-343	362	10	gradient	gradient	NOUN
alces-343	362	11	:	:	PUNCT
alces-343	362	12	change	change	NOUN
alces-343	362	13	in	in	ADP
alces-343	362	14	trophic	trophic	ADJ
alces-343	362	15	dominance	dominance	NOUN
alces-343	362	16	.	.	PUNCT
alces-343	363	1	evolutionary	evolutionary	ADJ
alces-343	363	2	ecology	ecology	NOUN
alces-343	363	3	10:51	10:51	NUM
alces-343	363	4	-	-	SYM
alces-343	363	5	62	62	NUM
alces-343	363	6	.	.	PUNCT
alces-343	364	1	cumming	cumme	VERB
alces-343	364	2	,	,	PUNCT
alces-343	364	3	h.	h.	PROPN
alces-343	364	4	g.	g.	PROPN
alces-343	364	5	,	,	PUNCT
alces-343	364	6	d.	d.	PROPN
alces-343	364	7	b.	b.	PROPN
alces-343	364	8	beange	beange	PROPN
alces-343	364	9	,	,	PUNCT
alces-343	364	10	and	and	CCONJ
alces-343	364	11	g.	g.	PROPN
alces-343	364	12	lavoie	lavoie	PROPN
alces-343	364	13	.	.	PUNCT
alces-343	364	14	1996	1996	NUM
alces-343	364	15	.	.	PUNCT
alces-343	365	1	habitat	habitat	NOUN
alces-343	365	2	partitioning	partitioning	NOUN
alces-343	365	3	between	between	ADP
alces-343	365	4	woodland	woodland	PROPN
alces-343	365	5	caribou	caribou	PROPN
alces-343	365	6	and	and	CCONJ
alces-343	365	7	moose	moose	NOUN
alces-343	365	8	in	in	ADP
alces-343	365	9	ontario	ontario	PROPN
alces-343	365	10	:	:	PUNCT
alces-343	365	11	the	the	DET
alces-343	365	12	potential	potential	ADJ
alces-343	365	13	role	role	NOUN
alces-343	365	14	of	of	ADP
alces-343	365	15	shared	share	VERB
alces-343	365	16	predation	predation	NOUN
alces-343	365	17	risk	risk	NOUN
alces-343	365	18	.	.	PUNCT
alces-343	366	1	rangifer	rangifer	NOUN
alces-343	366	2	,	,	PUNCT
alces-343	366	3	special	special	ADJ
alces-343	366	4	issue	issue	NOUN
alces-343	366	5	9:81	9:81	NUM
alces-343	366	6	-	-	SYM
alces-343	366	7	94	94	NUM
alces-343	366	8	.	.	PUNCT
alces-343	367	1	fancy	fancy	ADJ
alces-343	367	2	,	,	PUNCT
alces-343	367	3	s.	s.	PROPN
alces-343	367	4	g.	g.	PROPN
alces-343	367	5	,	,	PUNCT
alces-343	367	6	k.	k.	PROPN
alces-343	367	7	r.	r.	PROPN
alces-343	367	8	whitten	whitten	PROPN
alces-343	367	9	,	,	PUNCT
alces-343	367	10	and	and	CCONJ
alces-343	367	11	d.	d.	PROPN
alces-343	367	12	e.	e.	PROPN
alces-343	367	13	russell	russell	PROPN
alces-343	367	14	.	.	PUNCT
alces-343	368	1	1994	1994	NUM
alces-343	368	2	.	.	PUNCT
alces-343	369	1	demography	demography	NOUN
alces-343	369	2	of	of	ADP
alces-343	369	3	the	the	DET
alces-343	369	4	porcupine	porcupine	NOUN
alces-343	369	5	caribou	caribou	PROPN
alces-343	369	6	herd	herd	NOUN
alces-343	369	7	,	,	PUNCT
alces-343	369	8	1983	1983	NUM
alces-343	369	9	-	-	SYM
alces-343	369	10	1992	1992	NUM
alces-343	369	11	.	.	PUNCT
alces-343	370	1	canadian	canadian	ADJ
alces-343	370	2	journal	journal	PROPN
alces-343	370	3	of	of	ADP
alces-343	370	4	zoology	zoology	NOUN
alces-343	370	5	.	.	PUNCT
alces-343	371	1	72:840	72:840	NUM
alces-343	371	2	-	-	SYM
alces-343	371	3	846	846	NUM
alces-343	371	4	.	.	PUNCT
alces-343	371	5	fryxell	fryxell	PROPN
alces-343	371	6	,	,	PUNCT
alces-343	371	7	j.	j.	PROPN
alces-343	371	8	m.	m.	PROPN
alces-343	371	9	,	,	PUNCT
alces-343	371	10	w.	w.	PROPN
alces-343	371	11	e.	e.	PROPN
alces-343	371	12	mercer	mercer	PROPN
alces-343	371	13	,	,	PUNCT
alces-343	371	14	and	and	CCONJ
alces-343	371	15	r.	r.	PROPN
alces-343	371	16	b	b	PROPN
alces-343	371	17	gellatel	gellatel	PROPN
alces-343	371	18	.	.	PUNCT
alces-343	371	19	1988	1988	NUM
alces-343	371	20	.	.	PUNCT
alces-343	372	1	population	population	NOUN
alces-343	372	2	dynamics	dynamic	NOUN
alces-343	372	3	of	of	ADP
alces-343	372	4	newfoundland	newfoundland	PROPN
alces-343	372	5	moose	moose	PROPN
alces-343	372	6	using	use	VERB
alces-343	372	7	cohort	cohort	NOUN
alces-343	372	8	analysis	analysis	NOUN
alces-343	372	9	.	.	PUNCT
alces-343	373	1	journal	journal	NOUN
alces-343	373	2	of	of	ADP
alces-343	373	3	wildlife	wildlife	NOUN
alces-343	373	4	management	management	NOUN
alces-343	373	5	52:14	52:14	NUM
alces-343	373	6	-	-	SYM
alces-343	373	7	21	21	NUM
alces-343	373	8	.	.	PUNCT
alces-343	374	1	fuller	full	ADJ
alces-343	374	2	,	,	PUNCT
alces-343	374	3	t.	t.	PROPN
alces-343	374	4	k.	k.	PROPN
alces-343	374	5	1989	1989	NUM
alces-343	374	6	.	.	PUNCT
alces-343	375	1	population	population	NOUN
alces-343	375	2	dynamics	dynamic	NOUN
alces-343	375	3	of	of	ADP
alces-343	375	4	wolves	wolf	NOUN
alces-343	375	5	in	in	ADP
alces-343	375	6	north	north	NOUN
alces-343	375	7	-	-	PUNCT
alces-343	375	8	central	central	ADJ
alces-343	375	9	minnesota	minnesota	NOUN
alces-343	375	10	.	.	PUNCT
alces-343	376	1	wildlife	wildlife	NOUN
alces-343	376	2	monographs	monograph	NOUN
alces-343	376	3	105	105	NUM
alces-343	376	4	.	.	PUNCT
alces-343	377	1	gauthier	gauthier	PROPN
alces-343	377	2	,	,	PUNCT
alces-343	377	3	l.	l.	PROPN
alces-343	377	4	,	,	PUNCT
alces-343	377	5	r.	r.	PROPN
alces-343	377	6	nault	nault	PROPN
alces-343	377	7	,	,	PUNCT
alces-343	377	8	and	and	CCONJ
alces-343	377	9	m.	m.	NOUN
alces-343	377	10	crête	crête	PROPN
alces-343	377	11	.	.	PUNCT
alces-343	377	12	1989	1989	NUM
alces-343	377	13	.	.	PUNCT
alces-343	378	1	variations	variation	NOUN
alces-343	378	2	saisonnières	saisonnière	VERB
alces-343	378	3	du	du	PROPN
alces-343	378	4	régime	régime	PROPN
alces-343	378	5	alimentaire	alimentaire	PROPN
alces-343	378	6	des	des	PROPN
alces-343	378	7	caribous	caribous	PROPN
alces-343	378	8	du	du	PROPN
alces-343	378	9	troupeau	troupeau	PROPN
alces-343	378	10	de	de	PROPN
alces-343	378	11	la	la	PROPN
alces-343	378	12	rivière	rivière	PROPN
alces-343	378	13	george	george	PROPN
alces-343	378	14	,	,	PUNCT
alces-343	378	15	québec	québec	PROPN
alces-343	378	16	nordique	nordique	PROPN
alces-343	378	17	.	.	PUNCT
alces-343	378	18	naturaliste	naturaliste	PROPN
alces-343	378	19	canadien	canadien	PROPN
alces-343	378	20	116:101	116:101	PROPN
alces-343	378	21	-	-	PUNCT
alces-343	378	22	112	112	NUM
alces-343	378	23	.	.	PUNCT
alces-343	379	1	gingras	gingras	PROPN
alces-343	379	2	,	,	PUNCT
alces-343	379	3	a.	a.	PROPN
alces-343	379	4	,	,	PUNCT
alces-343	379	5	r.	r.	PROPN
alces-343	379	6	audy	audy	PROPN
alces-343	379	7	,	,	PUNCT
alces-343	379	8	and	and	CCONJ
alces-343	379	9	r.	r.	PROPN
alces-343	379	10	courtois	courtois	PROPN
alces-343	379	11	.	.	PUNCT
alces-343	380	1	1989	1989	NUM
alces-343	380	2	.	.	PUNCT
alces-343	381	1	inventaire	inventaire	PROPN
alces-343	381	2	aérien	aérien	PROPN
alces-343	381	3	de	de	PROPN
alces-343	381	4	l’orignal	l’orignal	PROPN
alces-343	381	5	dans	dans	PROPN
alces-343	381	6	la	la	PROPN
alces-343	381	7	zone	zone	PROPN
alces-343	381	8	de	de	X
alces-343	381	9	chasse	chasse	PROPN
alces-343	381	10	19	19	NUM
alces-343	381	11	à	à	X
alces-343	381	12	l’hiver	l’hiver	X
alces-343	381	13	1987	1987	NUM
alces-343	381	14	-	-	SYM
alces-343	381	15	88	88	NUM
alces-343	381	16	.	.	PUNCT
alces-343	382	1	ministère	ministère	PROPN
alces-343	382	2	du	du	PROPN
alces-343	382	3	loisir	loisir	PROPN
alces-343	382	4	,	,	PUNCT
alces-343	382	5	de	de	X
alces-343	382	6	la	la	X
alces-343	382	7	chasse	chasse	PROPN
alces-343	382	8	et	et	PROPN
alces-343	382	9	de	de	X
alces-343	382	10	la	la	PROPN
alces-343	382	11	pêche	pêche	PROPN
alces-343	382	12	.	.	PUNCT
alces-343	383	1	sept	sept	PROPN
alces-343	383	2	-	-	PUNCT
alces-343	383	3	îles	île	NOUN
alces-343	383	4	,	,	PUNCT
alces-343	383	5	québec	québec	PROPN
alces-343	383	6	,	,	PUNCT
alces-343	383	7	canada	canada	PROPN
alces-343	383	8	.	.	PUNCT
alces-343	384	1	hayes	hayes	PROPN
alces-343	384	2	,	,	PUNCT
alces-343	384	3	r.	r.	PROPN
alces-343	384	4	d.	d.	PROPN
alces-343	384	5	,	,	PUNCT
alces-343	384	6	r.	r.	PROPN
alces-343	384	7	farnel	farnel	PROPN
alces-343	384	8	,	,	PUNCT
alces-343	384	9	r.	r.	PROPN
alces-343	384	10	m.	m.	PROPN
alces-343	384	11	p.	p.	PROPN
alces-343	384	12	ward	ward	PROPN
alces-343	384	13	,	,	PUNCT
alces-343	384	14	j.	j.	PROPN
alces-343	384	15	carey	carey	PROPN
alces-343	384	16	,	,	PUNCT
alces-343	384	17	m.	m.	NOUN
alces-343	384	18	dehn	dehn	PROPN
alces-343	384	19	,	,	PUNCT
alces-343	384	20	g.	g.	PROPN
alces-343	384	21	w.	w.	PROPN
alces-343	384	22	kuzyk	kuzyk	PROPN
alces-343	384	23	,	,	PUNCT
alces-343	384	24	a.	a.	NOUN
alces-343	384	25	m.baer	m.baer	NOUN
alces-343	384	26	,	,	PUNCT
alces-343	384	27	c.	c.	PROPN
alces-343	384	28	l.	l.	PROPN
alces-343	384	29	gardner	gardner	PROPN
alces-343	384	30	,	,	PUNCT
alces-343	384	31	and	and	CCONJ
alces-343	384	32	m.	m.	PROPN
alces-343	384	33	o’donoghue	o’donoghue	PROPN
alces-343	384	34	.	.	PROPN
alces-343	384	35	2003	2003	NUM
alces-343	384	36	.	.	PUNCT
alces-343	385	1	experimental	experimental	ADJ
alces-343	385	2	reduction	reduction	NOUN
alces-343	385	3	of	of	ADP
alces-343	385	4	wolves	wolf	NOUN
alces-343	385	5	in	in	ADP
alces-343	385	6	the	the	DET
alces-343	385	7	yukon	yukon	PROPN
alces-343	385	8	:	:	PUNCT
alces-343	385	9	ungulate	ungulate	ADJ
alces-343	385	10	responses	response	NOUN
alces-343	385	11	and	and	CCONJ
alces-343	385	12	management	management	NOUN
alces-343	385	13	implications	implication	NOUN
alces-343	385	14	.	.	PUNCT
alces-343	386	1	wildlife	wildlife	NOUN
alces-343	386	2	monographs	monograph	NOUN
alces-343	386	3	152	152	NUM
alces-343	386	4	.	.	PUNCT
alces-343	387	1	klein	klein	PROPN
alces-343	387	2	,	,	PUNCT
alces-343	387	3	d.	d.	PROPN
alces-343	387	4	r.	r.	PROPN
alces-343	387	5	1968	1968	NUM
alces-343	387	6	.	.	PUNCT
alces-343	388	1	the	the	DET
alces-343	388	2	introduction	introduction	NOUN
alces-343	388	3	,	,	PUNCT
alces-343	388	4	increase	increase	NOUN
alces-343	388	5	,	,	PUNCT
alces-343	388	6	and	and	CCONJ
alces-343	388	7	crash	crash	NOUN
alces-343	388	8	of	of	ADP
alces-343	388	9	reindeer	reindeer	NOUN
alces-343	388	10	on	on	ADP
alces-343	388	11	st	st	PROPN
alces-343	388	12	.	.	PROPN
alces-343	388	13	mattew	mattew	PROPN
alces-343	388	14	island	island	PROPN
alces-343	388	15	.	.	PUNCT
alces-343	389	1	journal	journal	NOUN
alces-343	389	2	of	of	ADP
alces-343	389	3	wildlife	wildlife	NOUN
alces-343	389	4	management	management	NOUN
alces-343	389	5	32:350	32:350	NUM
alces-343	389	6	-	-	SYM
alces-343	389	7	367	367	NUM
alces-343	389	8	.	.	PUNCT
alces-343	390	1	larivière	larivière	PROPN
alces-343	390	2	,	,	PUNCT
alces-343	390	3	s.	s.	PROPN
alces-343	390	4	,	,	PUNCT
alces-343	390	5	h.	h.	PROPN
alces-343	390	6	jolicoeur	jolicoeur	PROPN
alces-343	390	7	,	,	PUNCT
alces-343	390	8	and	and	CCONJ
alces-343	390	9	m.	m.	NOUN
alces-343	390	10	crête	crête	PROPN
alces-343	390	11	.	.	PUNCT
alces-343	391	1	2000	2000	NUM
alces-343	391	2	.	.	PUNCT
alces-343	392	1	status	status	NOUN
alces-343	392	2	and	and	CCONJ
alces-343	392	3	conservation	conservation	NOUN
alces-343	392	4	of	of	ADP
alces-343	392	5	the	the	DET
alces-343	392	6	gray	gray	ADJ
alces-343	392	7	wolf	wolf	NOUN
alces-343	392	8	(	(	PUNCT
alces-343	392	9	canis	canis	NOUN
alces-343	392	10	lupus	lupus	NOUN
alces-343	392	11	)	)	PUNCT
alces-343	392	12	in	in	ADP
alces-343	392	13	wildlife	wildlife	NOUN
alces-343	392	14	reserves	reserve	NOUN
alces-343	392	15	of	of	ADP
alces-343	392	16	québec	québec	PROPN
alces-343	392	17	.	.	PUNCT
alces-343	393	1	biological	biological	ADJ
alces-343	393	2	conservation	conservation	NOUN
alces-343	393	3	94:143	94:143	PROPN
alces-343	393	4	-	-	PUNCT
alces-343	393	5	151	151	NUM
alces-343	393	6	.	.	PUNCT
alces-343	394	1	laurian	laurian	PROPN
alces-343	394	2	,	,	PUNCT
alces-343	394	3	c.	c.	PROPN
alces-343	394	4	,	,	PUNCT
alces-343	394	5	j.	j.	PROPN
alces-343	395	1	-p	-p	PROPN
alces-343	395	2	.	.	PUNCT
alces-343	396	1	ouellet	ouellet	PROPN
alces-343	396	2	,	,	PUNCT
alces-343	396	3	r.	r.	PROPN
alces-343	396	4	courtois	courtois	PROPN
alces-343	396	5	,	,	PUNCT
alces-343	396	6	l.	l.	PROPN
alces-343	396	7	breton	breton	PROPN
alces-343	396	8	,	,	PUNCT
alces-343	396	9	and	and	CCONJ
alces-343	396	10	s.	s.	PROPN
alces-343	396	11	st	st	PROPN
alces-343	396	12	-	-	PUNCT
alces-343	396	13	onge	onge	NOUN
alces-343	396	14	.	.	PUNCT
alces-343	397	1	2000	2000	NUM
alces-343	397	2	.	.	PUNCT
alces-343	398	1	the	the	DET
alces-343	398	2	effects	effect	NOUN
alces-343	398	3	of	of	ADP
alces-343	398	4	intensive	intensive	ADJ
alces-343	398	5	harvesting	harvesting	NOUN
alces-343	398	6	on	on	ADP
alces-343	398	7	moose	moose	NOUN
alces-343	398	8	reproduction	reproduction	NOUN
alces-343	398	9	.	.	PUNCT
alces-343	399	1	journal	journal	PROPN
alces-343	399	2	of	of	ADP
alces-343	399	3	applied	apply	VERB
alces-343	399	4	ecology	ecology	NOUN
alces-343	399	5	37:515	37:515	NOUN
alces-343	399	6	-	-	SYM
alces-343	399	7	531	531	NUM
alces-343	399	8	.	.	PUNCT
alces-343	400	1	mallory	mallory	PROPN
alces-343	400	2	,	,	PUNCT
alces-343	400	3	f.	f.	PROPN
alces-343	400	4	f.	f.	PROPN
alces-343	400	5	,	,	PUNCT
alces-343	400	6	and	and	CCONJ
alces-343	400	7	t.	t.	PROPN
alces-343	400	8	l.	l.	PROPN
alces-343	400	9	hillis	hillis	PROPN
alces-343	400	10	.	.	PUNCT
alces-343	401	1	1998	1998	NUM
alces-343	401	2	.	.	PUNCT
alces-343	402	1	demographic	demographic	ADJ
alces-343	402	2	characteristics	characteristic	NOUN
alces-343	402	3	of	of	ADP
alces-343	402	4	circumpolar	circumpolar	ADJ
alces-343	402	5	caribou	caribou	NOUN
alces-343	402	6	populations	population	NOUN
alces-343	402	7	:	:	PUNCT
alces-343	402	8	ecotypes	ecotype	NOUN
alces-343	402	9	,	,	PUNCT
alces-343	402	10	ecological	ecological	ADJ
alces-343	402	11	constraints	constraint	NOUN
alces-343	402	12	/	/	SYM
alces-343	402	13	releases	release	NOUN
alces-343	402	14	,	,	PUNCT
alces-343	402	15	and	and	CCONJ
alces-343	402	16	population	population	NOUN
alces-343	402	17	dynamics	dynamic	NOUN
alces-343	402	18	.	.	PUNCT
alces-343	403	1	rangifer	rangifer	NOUN
alces-343	403	2	,	,	PUNCT
alces-343	403	3	special	special	ADJ
alces-343	403	4	issue	issue	NOUN
alces-343	403	5	10:49	10:49	NUM
alces-343	403	6	-	-	SYM
alces-343	403	7	60	60	NUM
alces-343	403	8	.	.	PUNCT
alces-343	404	1	messier	messy	ADJ
alces-343	404	2	,	,	PUNCT
alces-343	404	3	f.	f.	PROPN
alces-343	404	4	1985	1985	NUM
alces-343	404	5	.	.	PUNCT
alces-343	405	1	social	social	ADJ
alces-343	405	2	organization	organization	NOUN
alces-343	405	3	,	,	PUNCT
alces-343	405	4	spaalces	spaalce	NOUN
alces-343	405	5	vol	vol	NOUN
alces-343	405	6	.	.	PROPN
alces-343	406	1	43	43	NUM
alces-343	406	2	,	,	PUNCT
alces-343	406	3	2007	2007	NUM
alces-343	406	4	courtois	courtois	NOUN
alces-343	406	5	and	and	CCONJ
alces-343	406	6	ouellet	ouellet	NOUN
alces-343	406	7	impact	impact	NOUN
alces-343	406	8	of	of	ADP
alces-343	406	9	moose	moose	NOUN
alces-343	406	10	and	and	CCONJ
alces-343	406	11	wolves	wolf	NOUN
alces-343	406	12	on	on	ADP
alces-343	406	13	caribou	caribou	NOUN
alces-343	406	14	25	25	NUM
alces-343	406	15	tial	tial	ADJ
alces-343	406	16	distribution	distribution	NOUN
alces-343	406	17	and	and	CCONJ
alces-343	406	18	population	population	NOUN
alces-343	406	19	density	density	NOUN
alces-343	406	20	of	of	ADP
alces-343	406	21	wolves	wolf	NOUN
alces-343	406	22	in	in	ADP
alces-343	406	23	relation	relation	NOUN
alces-343	406	24	to	to	PART
alces-343	406	25	moose	moose	NOUN
alces-343	406	26	density	density	NOUN
alces-343	406	27	.	.	PUNCT
alces-343	407	1	canadian	canadian	ADJ
alces-343	407	2	journal	journal	PROPN
alces-343	407	3	of	of	ADP
alces-343	407	4	zoology	zoology	NOUN
alces-343	407	5	63:10681077	63:10681077	PUNCT
alces-343	407	6	.	.	PUNCT
alces-343	408	1	_	_	PUNCT
alces-343	409	1	_	_	PUNCT
alces-343	410	1	_	_	PUNCT
alces-343	411	1	_	_	PUNCT
alces-343	412	1	_	_	PUNCT
alces-343	412	2	.	.	PUNCT
alces-343	413	1	1994	1994	NUM
alces-343	413	2	.	.	PUNCT
alces-343	414	1	ungulate	ungulate	ADJ
alces-343	414	2	population	population	NOUN
alces-343	414	3	models	model	NOUN
alces-343	414	4	with	with	ADP
alces-343	414	5	predation	predation	NOUN
alces-343	414	6	:	:	PUNCT
alces-343	414	7	a	a	DET
alces-343	414	8	case	case	NOUN
alces-343	414	9	study	study	NOUN
alces-343	414	10	with	with	ADP
alces-343	414	11	north	north	PROPN
alces-343	414	12	american	american	ADJ
alces-343	414	13	moose	moose	NOUN
alces-343	414	14	.	.	PUNCT
alces-343	415	1	ecology	ecology	NOUN
alces-343	415	2	75:478	75:478	PROPN
alces-343	415	3	-	-	SYM
alces-343	415	4	488	488	NUM
alces-343	415	5	.	.	PUNCT
alces-343	416	1	_	_	PUNCT
alces-343	417	1	_	_	PUNCT
alces-343	418	1	_	_	PUNCT
alces-343	419	1	_	_	PUNCT
alces-343	420	1	_	_	PUNCT
alces-343	420	2	.	.	PUNCT
alces-343	421	1	1995	1995	NUM
alces-343	421	2	.	.	PUNCT
alces-343	422	1	trophic	trophic	ADJ
alces-343	422	2	interactions	interaction	NOUN
alces-343	422	3	in	in	ADP
alces-343	422	4	two	two	NUM
alces-343	422	5	northern	northern	ADJ
alces-343	422	6	wolf	wolf	NOUN
alces-343	422	7	-	-	PUNCT
alces-343	422	8	ungulate	ungulate	ADJ
alces-343	422	9	systems	system	NOUN
alces-343	422	10	.	.	PUNCT
alces-343	423	1	wildlife	wildlife	NOUN
alces-343	423	2	research	research	NOUN
alces-343	423	3	22:131	22:131	NUM
alces-343	423	4	-	-	SYM
alces-343	423	5	146	146	NUM
alces-343	423	6	.	.	PUNCT
alces-343	424	1	_	_	PUNCT
alces-343	425	1	_	_	PUNCT
alces-343	426	1	_	_	PUNCT
alces-343	427	1	_	_	PUNCT
alces-343	428	1	_	_	PUNCT
alces-343	428	2	,	,	PUNCT
alces-343	428	3	j.	j.	PROPN
alces-343	428	4	huot	huot	PROPN
alces-343	428	5	,	,	PUNCT
alces-343	428	6	d.	d.	PROPN
alces-343	428	7	le	le	PROPN
alces-343	428	8	henaff	henaff	PROPN
alces-343	428	9	,	,	PUNCT
alces-343	428	10	and	and	CCONJ
alces-343	428	11	s.	s.	PROPN
alces-343	428	12	luttich	luttich	PROPN
alces-343	428	13	.	.	PUNCT
alces-343	429	1	1988	1988	NUM
alces-343	429	2	.	.	PUNCT
alces-343	430	1	demography	demography	NOUN
alces-343	430	2	of	of	ADP
alces-343	430	3	the	the	DET
alces-343	430	4	george	george	PROPN
alces-343	430	5	river	river	PROPN
alces-343	430	6	caribou	caribou	PROPN
alces-343	430	7	herd	herd	NOUN
alces-343	430	8	:	:	PUNCT
alces-343	430	9	evidence	evidence	NOUN
alces-343	430	10	of	of	ADP
alces-343	430	11	population	population	NOUN
alces-343	430	12	regulation	regulation	NOUN
alces-343	430	13	by	by	ADP
alces-343	430	14	forage	forage	NOUN
alces-343	430	15	exploitation	exploitation	NOUN
alces-343	430	16	and	and	CCONJ
alces-343	430	17	range	range	NOUN
alces-343	430	18	expansion	expansion	NOUN
alces-343	430	19	.	.	PUNCT
alces-343	431	1	arctic	arctic	PROPN
alces-343	431	2	41:279	41:279	PROPN
alces-343	431	3	-	-	PUNCT
alces-343	431	4	287	287	NUM
alces-343	431	5	.	.	PUNCT
alces-343	431	6	oksanen	oksanen	PROPN
alces-343	431	7	,	,	PUNCT
alces-343	431	8	l.	l.	PROPN
alces-343	431	9	1988	1988	NUM
alces-343	431	10	.	.	PUNCT
alces-343	432	1	ecosystem	ecosystem	NOUN
alces-343	432	2	organization	organization	NOUN
alces-343	432	3	:	:	PUNCT
alces-343	432	4	mutualism	mutualism	NOUN
alces-343	432	5	and	and	CCONJ
alces-343	432	6	cybernetics	cybernetic	NOUN
alces-343	432	7	or	or	CCONJ
alces-343	432	8	plain	plain	ADJ
alces-343	432	9	darwinian	darwinian	ADJ
alces-343	432	10	struggle	struggle	NOUN
alces-343	432	11	for	for	ADP
alces-343	432	12	existence	existence	NOUN
alces-343	432	13	?	?	PUNCT
alces-343	433	1	the	the	DET
alces-343	433	2	american	american	PROPN
alces-343	433	3	naturalist	naturalist	NOUN
alces-343	433	4	131:424	131:424	PROPN
alces-343	433	5	-	-	PUNCT
alces-343	433	6	444	444	NUM
alces-343	433	7	.	.	PUNCT
alces-343	434	1	_	_	PUNCT
alces-343	435	1	_	_	PUNCT
alces-343	436	1	_	_	PUNCT
alces-343	437	1	_	_	PUNCT
alces-343	438	1	_	_	PUNCT
alces-343	438	2	,	,	PUNCT
alces-343	438	3	and	and	CCONJ
alces-343	438	4	t.	t.	PROPN
alces-343	438	5	oksanen	oksanen	NOUN
alces-343	438	6	.	.	PROPN
alces-343	438	7	2000	2000	NUM
alces-343	438	8	.	.	PUNCT
alces-343	439	1	the	the	DET
alces-343	439	2	logic	logic	NOUN
alces-343	439	3	and	and	CCONJ
alces-343	439	4	realism	realism	NOUN
alces-343	439	5	of	of	ADP
alces-343	439	6	the	the	DET
alces-343	439	7	hypothesis	hypothesis	NOUN
alces-343	439	8	of	of	ADP
alces-343	439	9	exploitation	exploitation	NOUN
alces-343	439	10	ecosystem	ecosystem	NOUN
alces-343	439	11	.	.	PUNCT
alces-343	440	1	the	the	DET
alces-343	440	2	american	american	PROPN
alces-343	440	3	naturalist	naturalist	PROPN
alces-343	440	4	155:703	155:703	PROPN
alces-343	440	5	-	-	SYM
alces-343	440	6	723	723	NUM
alces-343	440	7	.	.	PUNCT
alces-343	441	1	_	_	PUNCT
alces-343	442	1	_	_	PUNCT
alces-343	443	1	_	_	PUNCT
alces-343	444	1	_	_	PUNCT
alces-343	445	1	_	_	PUNCT
alces-343	445	2	,	,	PUNCT
alces-343	445	3	d.	d.	PROPN
alces-343	445	4	stephen	stephen	PROPN
alces-343	445	5	,	,	PUNCT
alces-343	445	6	j.	j.	PROPN
alces-343	445	7	a.	a.	PROPN
alces-343	445	8	fretwell	fretwell	PROPN
alces-343	445	9	,	,	PUNCT
alces-343	445	10	and	and	CCONJ
alces-343	445	11	n.	n.	PROPN
alces-343	445	12	pekka	pekka	PROPN
alces-343	445	13	.	.	PUNCT
alces-343	445	14	1981	1981	NUM
alces-343	445	15	.	.	PUNCT
alces-343	446	1	exploitation	exploitation	NOUN
alces-343	446	2	ecosystems	ecosystem	NOUN
alces-343	446	3	in	in	ADP
alces-343	446	4	gradients	gradient	NOUN
alces-343	446	5	of	of	ADP
alces-343	446	6	primary	primary	ADJ
alces-343	446	7	productivity	productivity	NOUN
alces-343	446	8	.	.	PUNCT
alces-343	447	1	the	the	DET
alces-343	447	2	american	american	PROPN
alces-343	447	3	naturalist	naturalist	PROPN
alces-343	447	4	118:240	118:240	PROPN
alces-343	447	5	-	-	PUNCT
alces-343	447	6	261	261	NUM
alces-343	447	7	.	.	PUNCT
alces-343	448	1	ouellet	ouellet	NOUN
alces-343	448	2	,	,	PUNCT
alces-343	448	3	j.-p	j.-p	PROPN
alces-343	448	4	.	.	PROPN
alces-343	448	5	,	,	PUNCT
alces-343	448	6	s.	s.	PROPN
alces-343	448	7	boutin	boutin	PROPN
alces-343	448	8	,	,	PUNCT
alces-343	448	9	and	and	CCONJ
alces-343	448	10	d.	d.	PROPN
alces-343	448	11	c.	c.	PROPN
alces-343	448	12	heard	hear	VERB
alces-343	448	13	.	.	PUNCT
alces-343	449	1	1994	1994	NUM
alces-343	449	2	.	.	PUNCT
alces-343	450	1	responses	response	NOUN
alces-343	450	2	to	to	ADP
alces-343	450	3	simulated	simulate	VERB
alces-343	450	4	grazing	grazing	NOUN
alces-343	450	5	and	and	CCONJ
alces-343	450	6	browsing	browsing	NOUN
alces-343	450	7	of	of	ADP
alces-343	450	8	vegetation	vegetation	NOUN
alces-343	450	9	available	available	ADJ
alces-343	450	10	to	to	ADP
alces-343	450	11	caribou	caribou	PROPN
alces-343	450	12	in	in	ADP
alces-343	450	13	the	the	DET
alces-343	450	14	arctic	arctic	PROPN
alces-343	450	15	.	.	PUNCT
alces-343	451	1	canadian	canadian	ADJ
alces-343	451	2	journal	journal	PROPN
alces-343	451	3	of	of	ADP
alces-343	451	4	zoology	zoology	NOUN
alces-343	451	5	72:1426	72:1426	PROPN
alces-343	451	6	-	-	SYM
alces-343	451	7	1435	1435	NUM
alces-343	451	8	.	.	PUNCT
alces-343	452	1	_	_	PUNCT
alces-343	453	1	_	_	PUNCT
alces-343	454	1	_	_	PUNCT
alces-343	455	1	_	_	PUNCT
alces-343	456	1	_	_	PUNCT
alces-343	456	2	,	,	PUNCT
alces-343	456	3	d.	d.	PROPN
alces-343	456	4	c.	c.	PROPN
alces-343	456	5	heard	hear	VERB
alces-343	456	6	,	,	PUNCT
alces-343	456	7	s.	s.	PROPN
alces-343	456	8	boutin	boutin	PROPN
alces-343	456	9	,	,	PUNCT
alces-343	456	10	and	and	CCONJ
alces-343	456	11	r.	r.	PROPN
alces-343	456	12	mulders	mulder	NOUN
alces-343	456	13	.	.	PUNCT
alces-343	457	1	1997	1997	NUM
alces-343	457	2	.	.	PUNCT
alces-343	458	1	a	a	DET
alces-343	458	2	comparison	comparison	NOUN
alces-343	458	3	of	of	ADP
alces-343	458	4	body	body	NOUN
alces-343	458	5	condition	condition	NOUN
alces-343	458	6	and	and	CCONJ
alces-343	458	7	reproduction	reproduction	NOUN
alces-343	458	8	of	of	ADP
alces-343	458	9	caribou	caribou	NOUN
alces-343	458	10	on	on	ADP
alces-343	458	11	two	two	NUM
alces-343	458	12	predator	predator	NOUN
alces-343	458	13	-	-	PUNCT
alces-343	458	14	free	free	ADJ
alces-343	458	15	arctic	arctic	ADJ
alces-343	458	16	islands	island	NOUN
alces-343	458	17	.	.	PUNCT
alces-343	459	1	canadian	canadian	ADJ
alces-343	459	2	journal	journal	PROPN
alces-343	459	3	of	of	ADP
alces-343	459	4	zoology	zoology	NOUN
alces-343	459	5	75:11	75:11	NUM
alces-343	459	6	-	-	SYM
alces-343	459	7	17	17	NUM
alces-343	459	8	.	.	PUNCT
alces-343	460	1	_	_	PUNCT
alces-343	461	1	_	_	PUNCT
alces-343	462	1	_	_	PUNCT
alces-343	463	1	_	_	PUNCT
alces-343	464	1	_	_	PUNCT
alces-343	464	2	,	,	PUNCT
alces-343	465	1	_	_	PUNCT
alces-343	466	1	_	_	PUNCT
alces-343	467	1	_	_	PUNCT
alces-343	468	1	_	_	PUNCT
alces-343	469	1	_	_	PUNCT
alces-343	469	2	,	,	PUNCT
alces-343	469	3	and	and	CCONJ
alces-343	469	4	r.	r.	PROPN
alces-343	469	5	mulder	mulder	PROPN
alces-343	469	6	.	.	PUNCT
alces-343	469	7	1996	1996	NUM
alces-343	469	8	.	.	PUNCT
alces-343	470	1	population	population	NOUN
alces-343	470	2	ecology	ecology	NOUN
alces-343	470	3	of	of	ADP
alces-343	470	4	caribou	caribou	NOUN
alces-343	470	5	populations	population	NOUN
alces-343	470	6	without	without	ADP
alces-343	470	7	predators	predator	NOUN
alces-343	470	8	:	:	PUNCT
alces-343	470	9	southampton	southampton	PROPN
alces-343	470	10	and	and	CCONJ
alces-343	470	11	coats	coat	VERB
alces-343	470	12	island	island	NOUN
alces-343	470	13	herds	herd	NOUN
alces-343	470	14	.	.	PUNCT
alces-343	471	1	rangifer	rangifer	NOUN
alces-343	471	2	,	,	PUNCT
alces-343	471	3	special	special	ADJ
alces-343	471	4	issue	issue	NOUN
alces-343	471	5	9:17	9:17	NUM
alces-343	471	6	-	-	SYM
alces-343	471	7	26	26	NUM
alces-343	471	8	.	.	PUNCT
alces-343	472	1	reimers	reimer	NOUN
alces-343	472	2	,	,	PUNCT
alces-343	472	3	e.	e.	PROPN
alces-343	472	4	1982	1982	NUM
alces-343	472	5	.	.	PUNCT
alces-343	473	1	winter	winter	NOUN
alces-343	473	2	mortality	mortality	NOUN
alces-343	473	3	and	and	CCONJ
alces-343	473	4	population	population	NOUN
alces-343	473	5	trends	trend	NOUN
alces-343	473	6	of	of	ADP
alces-343	473	7	reindeer	reindeer	NOUN
alces-343	473	8	on	on	ADP
alces-343	473	9	svalbard	svalbard	PROPN
alces-343	473	10	,	,	PUNCT
alces-343	473	11	norway	norway	PROPN
alces-343	473	12	.	.	PUNCT
alces-343	474	1	arctic	arctic	PROPN
alces-343	474	2	and	and	CCONJ
alces-343	474	3	alpine	alpine	PROPN
alces-343	474	4	research	research	NOUN
alces-343	474	5	14:295	14:295	NUM
alces-343	474	6	-	-	SYM
alces-343	474	7	300	300	NUM
alces-343	474	8	.	.	PUNCT
alces-343	474	9	rettie	rettie	NOUN
alces-343	474	10	,	,	PUNCT
alces-343	474	11	w.	w.	PROPN
alces-343	474	12	j.	j.	PROPN
alces-343	474	13	,	,	PUNCT
alces-343	474	14	and	and	CCONJ
alces-343	474	15	f.	f.	PROPN
alces-343	474	16	messier	messy	ADJ
alces-343	474	17	.	.	PUNCT
alces-343	474	18	1991	1991	NUM
alces-343	474	19	.	.	PUNCT
alces-343	475	1	the	the	DET
alces-343	475	2	implications	implication	NOUN
alces-343	475	3	of	of	ADP
alces-343	475	4	environmental	environmental	ADJ
alces-343	475	5	variability	variability	NOUN
alces-343	475	6	on	on	ADP
alces-343	475	7	caribou	caribou	NOUN
alces-343	475	8	demography	demography	NOUN
alces-343	475	9	:	:	PUNCT
alces-343	475	10	theoretical	theoretical	ADJ
alces-343	475	11	consideration	consideration	NOUN
alces-343	475	12	.	.	PUNCT
alces-343	476	1	rangifer	rangifer	NOUN
alces-343	476	2	,	,	PUNCT
alces-343	476	3	special	special	ADJ
alces-343	476	4	issue	issue	NOUN
alces-343	476	5	7:53	7:53	NUM
alces-343	476	6	-	-	SYM
alces-343	476	7	59	59	NUM
alces-343	476	8	.	.	PUNCT
alces-343	477	1	_	_	PUNCT
alces-343	478	1	_	_	PUNCT
alces-343	479	1	_	_	PUNCT
alces-343	480	1	_	_	PUNCT
alces-343	480	2	_	_	PUNCT
alces-343	480	3	,	,	PUNCT
alces-343	480	4	and	and	CCONJ
alces-343	480	5	_	_	PUNCT
alces-343	481	1	_	_	PUNCT
alces-343	482	1	_	_	PUNCT
alces-343	483	1	_	_	PUNCT
alces-343	484	1	_	_	PUNCT
alces-343	484	2	.	.	PUNCT
alces-343	485	1	1998	1998	NUM
alces-343	485	2	.	.	PUNCT
alces-343	486	1	dynamics	dynamic	NOUN
alces-343	486	2	of	of	ADP
alces-343	486	3	woodland	woodland	PROPN
alces-343	486	4	caribou	caribou	NOUN
alces-343	486	5	populations	population	NOUN
alces-343	486	6	at	at	ADP
alces-343	486	7	the	the	DET
alces-343	486	8	southern	southern	ADJ
alces-343	486	9	limit	limit	NOUN
alces-343	486	10	of	of	ADP
alces-343	486	11	their	their	PRON
alces-343	486	12	range	range	NOUN
alces-343	486	13	in	in	ADP
alces-343	486	14	saskatchewan	saskatchewan	PROPN
alces-343	486	15	.	.	PUNCT
alces-343	487	1	canadian	canadian	ADJ
alces-343	487	2	journal	journal	PROPN
alces-343	487	3	of	of	ADP
alces-343	487	4	zoology	zoology	NOUN
alces-343	487	5	76:251	76:251	PROPN
alces-343	487	6	-	-	SYM
alces-343	487	7	259	259	NUM
alces-343	487	8	.	.	PUNCT
alces-343	488	1	sebbane	sebbane	PROPN
alces-343	488	2	,	,	PUNCT
alces-343	488	3	a.	a.	PROPN
alces-343	488	4	,	,	PUNCT
alces-343	488	5	r.	r.	PROPN
alces-343	488	6	courtois	courtois	PROPN
alces-343	488	7	,	,	PUNCT
alces-343	488	8	s.	s.	PROPN
alces-343	488	9	st	st	PROPN
alces-343	488	10	-	-	PUNCT
alces-343	488	11	onge	onge	PROPN
alces-343	488	12	,	,	PUNCT
alces-343	488	13	l.	l.	PROPN
alces-343	488	14	breton	breton	PROPN
alces-343	488	15	,	,	PUNCT
alces-343	488	16	and	and	CCONJ
alces-343	488	17	p.	p.	NOUN
alces-343	488	18	-é	-é	NUM
alces-343	488	19	.	.	PUNCT
alces-343	489	1	lafleur	lafleur	PROPN
alces-343	489	2	.	.	PUNCT
alces-343	490	1	2003	2003	NUM
alces-343	490	2	.	.	PUNCT
alces-343	491	1	trente	trente	PROPN
alces-343	491	2	ans	ans	PROPN
alces-343	491	3	après	après	PROPN
alces-343	491	4	sa	sa	PROPN
alces-343	491	5	réintroduction	réintroduction	PROPN
alces-343	491	6	,	,	PUNCT
alces-343	491	7	quel	quel	PROPN
alces-343	491	8	est	est	PROPN
alces-343	491	9	l’avenir	l’avenir	PROPN
alces-343	491	10	du	du	PROPN
alces-343	491	11	caribou	caribou	PROPN
alces-343	491	12	de	de	PROPN
alces-343	491	13	charlevoix	charlevoix	PROPN
alces-343	491	14	?	?	PUNCT
alces-343	492	1	naturaliste	naturaliste	PROPN
alces-343	492	2	canadien	canadien	PROPN
alces-343	492	3	127:55	127:55	PROPN
alces-343	492	4	-	-	PUNCT
alces-343	492	5	62	62	NUM
alces-343	492	6	.	.	PUNCT
alces-343	493	1	seip	seip	PROPN
alces-343	493	2	,	,	PUNCT
alces-343	493	3	d.	d.	PROPN
alces-343	493	4	r.	r.	PROPN
alces-343	493	5	1991	1991	NUM
alces-343	493	6	.	.	PUNCT
alces-343	494	1	predation	predation	NOUN
alces-343	494	2	and	and	CCONJ
alces-343	494	3	caribou	caribou	NOUN
alces-343	494	4	populations	population	NOUN
alces-343	494	5	.	.	PUNCT
alces-343	495	1	rangifer	rangifer	NOUN
alces-343	495	2	,	,	PUNCT
alces-343	495	3	special	special	ADJ
alces-343	495	4	issue	issue	NOUN
alces-343	495	5	7:46	7:46	NUM
alces-343	495	6	-	-	SYM
alces-343	495	7	52	52	NUM
alces-343	495	8	.	.	PUNCT
alces-343	496	1	_	_	PUNCT
alces-343	497	1	_	_	PUNCT
alces-343	498	1	_	_	PUNCT
alces-343	499	1	_	_	PUNCT
alces-343	500	1	_	_	PUNCT
alces-343	500	2	.	.	PUNCT
alces-343	501	1	1992	1992	NUM
alces-343	501	2	.	.	PUNCT
alces-343	502	1	factors	factor	NOUN
alces-343	502	2	limiting	limit	VERB
alces-343	502	3	woodland	woodland	NOUN
alces-343	502	4	caribou	caribou	NOUN
alces-343	502	5	populations	population	NOUN
alces-343	502	6	and	and	CCONJ
alces-343	502	7	their	their	PRON
alces-343	502	8	interrelationships	interrelationship	NOUN
alces-343	502	9	with	with	ADP
alces-343	502	10	wolves	wolf	NOUN
alces-343	502	11	and	and	CCONJ
alces-343	502	12	moose	moose	NOUN
alces-343	502	13	in	in	ADP
alces-343	502	14	southeastern	southeastern	ADJ
alces-343	502	15	british	british	PROPN
alces-343	502	16	columbia	columbia	PROPN
alces-343	502	17	.	.	PUNCT
alces-343	503	1	canadian	canadian	ADJ
alces-343	503	2	journal	journal	PROPN
alces-343	503	3	of	of	ADP
alces-343	503	4	zoology	zoology	NOUN
alces-343	503	5	70:1494	70:1494	PROPN
alces-343	503	6	-	-	SYM
alces-343	503	7	1503	1503	NUM
alces-343	503	8	.	.	PUNCT
alces-343	504	1	stuart	stuart	PROPN
alces-343	504	2	-	-	PUNCT
alces-343	504	3	smith	smith	PROPN
alces-343	504	4	,	,	PUNCT
alces-343	504	5	a.	a.	PROPN
alces-343	504	6	k.	k.	PROPN
alces-343	504	7	,	,	PUNCT
alces-343	504	8	j.	j.	PROPN
alces-343	504	9	a.	a.	PROPN
alces-343	504	10	corey	corey	PROPN
alces-343	504	11	,	,	PUNCT
alces-343	504	12	s.	s.	PROPN
alces-343	504	13	boutin	boutin	PROPN
alces-343	504	14	,	,	PUNCT
alces-343	504	15	d.	d.	PROPN
alces-343	504	16	h.	h.	PROPN
alces-343	504	17	hebert	hebert	PROPN
alces-343	504	18	,	,	PUNCT
alces-343	504	19	and	and	CCONJ
alces-343	504	20	a.	a.	PROPN
alces-343	504	21	b.	b.	PROPN
alces-343	504	22	rippin	rippin	PROPN
alces-343	504	23	.	.	PUNCT
alces-343	505	1	1997	1997	NUM
alces-343	505	2	.	.	PUNCT
alces-343	506	1	woodland	woodland	PROPN
alces-343	506	2	caribou	caribou	PROPN
alces-343	506	3	relative	relative	ADJ
alces-343	506	4	to	to	ADP
alces-343	506	5	landscape	landscape	NOUN
alces-343	506	6	patterns	pattern	NOUN
alces-343	506	7	in	in	ADP
alces-343	506	8	northeastern	northeastern	ADJ
alces-343	506	9	alberta	alberta	PROPN
alces-343	506	10	.	.	PUNCT
alces-343	507	1	journal	journal	PROPN
alces-343	507	2	of	of	ADP
alces-343	507	3	wildlife	wildlife	NOUN
alces-343	507	4	management	management	NOUN
alces-343	507	5	61:622	61:622	NUM
alces-343	507	6	-	-	SYM
alces-343	507	7	633	633	NUM
alces-343	507	8	.	.	PUNCT
alces-343	508	1	timmermann	timmermann	PROPN
alces-343	508	2	,	,	PUNCT
alces-343	508	3	h.	h.	PROPN
alces-343	508	4	r.	r.	PROPN
alces-343	508	5	,	,	PUNCT
alces-343	508	6	and	and	CCONJ
alces-343	508	7	j.	j.	PROPN
alces-343	508	8	g	g	PROPN
alces-343	508	9	mcnicol	mcnicol	PROPN
alces-343	508	10	.	.	PUNCT
alces-343	509	1	1988	1988	NUM
alces-343	509	2	.	.	PUNCT
alces-343	510	1	moose	moose	PROPN
alces-343	510	2	habitat	habitat	NOUN
alces-343	510	3	needs	need	NOUN
alces-343	510	4	.	.	PUNCT
alces-343	511	1	the	the	DET
alces-343	511	2	forestry	forestry	PROPN
alces-343	511	3	chronicle	chronicle	PROPN
alces-343	511	4	64:238	64:238	PROPN
alces-343	511	5	-	-	SYM
alces-343	511	6	245	245	NUM
alces-343	511	7	.	.	PUNCT
alces-343	511	8	weclaw	weclaw	NOUN
alces-343	511	9	,	,	PUNCT
alces-343	511	10	p.	p.	NOUN
alces-343	511	11	,	,	PUNCT
alces-343	511	12	and	and	CCONJ
alces-343	511	13	r.	r.	PROPN
alces-343	511	14	j.	j.	PROPN
alces-343	511	15	hudson	hudson	PROPN
alces-343	511	16	.	.	PUNCT
alces-343	512	1	2004	2004	NUM
alces-343	512	2	.	.	PUNCT
alces-343	513	1	simulation	simulation	NOUN
alces-343	513	2	of	of	ADP
alces-343	513	3	conservation	conservation	NOUN
alces-343	513	4	and	and	CCONJ
alces-343	513	5	management	management	NOUN
alces-343	513	6	of	of	ADP
alces-343	513	7	woodland	woodland	PROPN
alces-343	513	8	caribou	caribou	PROPN
alces-343	513	9	.	.	PUNCT
alces-343	514	1	ecological	ecological	ADJ
alces-343	514	2	modeling	modeling	ADJ
alces-343	514	3	177:75	177:75	NUM
alces-343	514	4	-	-	SYM
alces-343	514	5	94	94	NUM
alces-343	514	6	.	.	PUNCT
alces-343	515	1	impact	impact	NOUN
alces-343	515	2	of	of	ADP
alces-343	515	3	moose	moose	NOUN
alces-343	515	4	and	and	CCONJ
alces-343	515	5	wolves	wolf	NOUN
alces-343	515	6	on	on	ADP
alces-343	515	7	caribou	caribou	NOUN
alces-343	515	8	–	–	PUNCT
alces-343	515	9	courtois	courtois	NOUN
alces-343	515	10	and	and	CCONJ
alces-343	515	11	ouellet	ouellet	PROPN
alces-343	515	12	alces	alces	PROPN
alces-343	515	13	vol	vol	NOUN
alces-343	515	14	.	.	PROPN
alces-343	516	1	43	43	NUM
alces-343	516	2	,	,	PUNCT
alces-343	516	3	2007	2007	NUM
alces-343	516	4	26	26	NUM
alces-343	516	5	appendix	appendix	NOUN
alces-343	516	6	i	i	PRON
alces-343	516	7	stella	stella	NOUN
alces-343	516	8	model	model	NOUN
alces-343	516	9	incorporating	incorporate	VERB
alces-343	516	10	moose	moose	NOUN
alces-343	516	11	,	,	PUNCT
alces-343	516	12	wolf	wolf	NOUN
alces-343	516	13	,	,	PUNCT
alces-343	516	14	and	and	CCONJ
alces-343	516	15	caribou	caribou	NOUN
alces-343	516	16	interactions	interaction	NOUN
alces-343	516	17	in	in	ADP
alces-343	516	18	the	the	DET
alces-343	516	19	boreal	boreal	NOUN
alces-343	516	20	forest	forest	NOUN
alces-343	516	21	.	.	PUNCT
alces-343	517	1	rectangles	rectangle	NOUN
alces-343	517	2	represent	represent	VERB
alces-343	517	3	reservoirs	reservoir	NOUN
alces-343	517	4	(	(	PUNCT
alces-343	517	5	populations	population	NOUN
alces-343	517	6	)	)	PUNCT
alces-343	517	7	,	,	PUNCT
alces-343	517	8	double	double	ADJ
alces-343	517	9	arrows	arrow	NOUN
alces-343	517	10	identify	identify	VERB
alces-343	517	11	sources	source	NOUN
alces-343	517	12	(	(	PUNCT
alces-343	517	13	births	birth	NOUN
alces-343	517	14	)	)	PUNCT
alces-343	517	15	and	and	CCONJ
alces-343	517	16	sinks	sink	NOUN
alces-343	517	17	tions	tion	NOUN
alces-343	517	18	tested	test	VERB
alces-343	517	19	the	the	DET
alces-343	517	20	impact	impact	NOUN
alces-343	517	21	of	of	ADP
alces-343	517	22	changes	change	NOUN
alces-343	517	23	in	in	ADP
alces-343	517	24	caribou	caribou	NOUN
alces-343	517	25	and	and	CCONJ
alces-343	517	26	moose	moose	NOUN
alces-343	517	27	hunting	hunt	VERB
alces-343	517	28	rates	rate	NOUN
alces-343	517	29	(	(	PUNCT
alces-343	517	30	caribouhuntingrate	caribouhuntingrate	NOUN
alces-343	517	31	and	and	CCONJ
alces-343	517	32	moosehuntingrate	moosehuntingrate	NOUN
alces-343	517	33	,	,	PUNCT
alces-343	517	34	respectively	respectively	ADV
alces-343	517	35	)	)	PUNCT
alces-343	517	36	,	,	PUNCT
alces-343	517	37	wolf	wolf	NOUN
alces-343	517	38	trapping	trap	VERB
alces-343	517	39	rate	rate	NOUN
alces-343	517	40	(	(	PUNCT
alces-343	517	41	wolftrappingrate	wolftrappingrate	NOUN
alces-343	517	42	)	)	PUNCT
alces-343	517	43	and	and	CCONJ
alces-343	517	44	changes	change	NOUN
alces-343	517	45	in	in	ADP
alces-343	517	46	caribou	caribou	NOUN
alces-343	517	47	and	and	CCONJ
alces-343	517	48	moose	moose	NOUN
alces-343	517	49	productivity	productivity	NOUN
alces-343	517	50	due	due	ADP
alces-343	517	51	to	to	ADP
alces-343	517	52	random	random	ADJ
alces-343	517	53	variations	variation	NOUN
alces-343	517	54	in	in	ADP
alces-343	517	55	environmental	environmental	ADJ
alces-343	517	56	conditions	condition	NOUN
alces-343	517	57	(	(	PUNCT
alces-343	517	58	caribourandomenvcdn	caribourandomenvcdn	NOUN
alces-343	517	59	and	and	CCONJ
alces-343	517	60	mooserandomenvcdn	mooserandomenvcdn	NOUN
alces-343	517	61	)	)	PUNCT
alces-343	517	62	on	on	ADP
alces-343	517	63	caribou	caribou	PROPN
alces-343	517	64	and	and	CCONJ
alces-343	517	65	moose	moose	VERB
alces-343	517	66	reproductive	reproductive	ADJ
alces-343	517	67	rates	rate	NOUN
alces-343	517	68	.	.	PUNCT
alces-343	518	1	other	other	ADJ
alces-343	518	2	parameters	parameter	NOUN
alces-343	518	3	and	and	CCONJ
alces-343	518	4	equations	equation	NOUN
alces-343	518	5	were	be	AUX
alces-343	518	6	obtained	obtain	VERB
alces-343	518	7	from	from	ADP
alces-343	518	8	the	the	DET
alces-343	518	9	literature	literature	NOUN
alces-343	518	10	(	(	PUNCT
alces-343	518	11	see	see	VERB
alces-343	518	12	methods	method	NOUN
alces-343	518	13	)	)	PUNCT
alces-343	518	14	.	.	PUNCT
alces-343	519	1	minprodcaribou	minprodcaribou	PROPN
alces-343	519	2	and	and	CCONJ
alces-343	519	3	minprodmoose	minprodmoose	VERB
alces-343	519	4	indicate	indicate	VERB
alces-343	519	5	the	the	DET
alces-343	519	6	minimum	minimum	ADJ
alces-343	519	7	productivity	productivity	NOUN
alces-343	519	8	of	of	ADP
alces-343	519	9	the	the	DET
alces-343	519	10	species	specie	NOUN
alces-343	519	11	under	under	ADP
alces-343	519	12	the	the	DET
alces-343	519	13	worst	bad	ADJ
alces-343	519	14	environmental	environmental	ADJ
alces-343	519	15	conditions	condition	NOUN
alces-343	519	16	.	.	PUNCT
alces-343	520	1	other	other	ADJ
alces-343	520	2	acronyms	acronym	NOUN
alces-343	520	3	are	be	AUX
alces-343	520	4	self	self	NOUN
alces-343	520	5	explanatory	explanatory	ADJ
alces-343	520	6	.	.	PUNCT
alces-343	521	1	alces	alce	NOUN
alces-343	521	2	vol	vol	NOUN
alces-343	521	3	.	.	PROPN
alces-343	522	1	43	43	NUM
alces-343	522	2	,	,	PUNCT
alces-343	522	3	2007	2007	NUM
alces-343	522	4	courtois	courtois	NOUN
alces-343	522	5	and	and	CCONJ
alces-343	522	6	ouellet	ouellet	NOUN
alces-343	522	7	impact	impact	NOUN
alces-343	522	8	of	of	ADP
alces-343	522	9	moose	moose	NOUN
alces-343	522	10	and	and	CCONJ
alces-343	522	11	wolves	wolf	NOUN
alces-343	522	12	on	on	ADP
alces-343	522	13	caribou	caribou	NOUN
alces-343	522	14	27	27	NUM
alces-343	522	15	appendix	appendix	PROPN
alces-343	522	16	ii	ii	NOUN
alces-343	522	17	parameters	parameter	NOUN
alces-343	522	18	and	and	CCONJ
alces-343	522	19	variables	variable	NOUN
alces-343	522	20	used	use	VERB
alces-343	522	21	in	in	ADP
alces-343	522	22	the	the	DET
alces-343	522	23	models	model	NOUN
alces-343	522	24	,	,	PUNCT
alces-343	522	25	values	value	NOUN
alces-343	522	26	employed	employ	VERB
alces-343	522	27	in	in	ADP
alces-343	522	28	the	the	DET
alces-343	522	29	simulations	simulation	NOUN
alces-343	522	30	,	,	PUNCT
alces-343	522	31	and	and	CCONJ
alces-343	522	32	data	datum	NOUN
alces-343	522	33	sources	source	NOUN
alces-343	522	34	.	.	PUNCT
alces-343	523	1	parameter	parameter	NOUN
alces-343	523	2	or	or	CCONJ
alces-343	523	3	variable	variable	ADJ
alces-343	523	4	values	value	NOUN
alces-343	523	5	employed	employ	VERB
alces-343	523	6	data	datum	NOUN
alces-343	523	7	source	source	NOUN
alces-343	523	8	moose	moose	NOUN
alces-343	523	9	habitat	habitat	NOUN
alces-343	523	10	carrying	carry	VERB
alces-343	523	11	capacity	capacity	NOUN
alces-343	523	12	840	840	NUM
alces-343	523	13	in	in	ADP
alces-343	523	14	the	the	DET
alces-343	523	15	study	study	NOUN
alces-343	523	16	area	area	NOUN
alces-343	523	17	(	(	PUNCT
alces-343	523	18	84	84	NUM
alces-343	523	19	per	per	ADP
alces-343	523	20	100	100	NUM
alces-343	523	21	km2	km2	NUM
alces-343	523	22	)	)	PUNCT
alces-343	523	23	courtois	courtois	PROPN
alces-343	523	24	et	et	PROPN
alces-343	523	25	al	al	PROPN
alces-343	523	26	.	.	PROPN
alces-343	524	1	(	(	PUNCT
alces-343	524	2	1993	1993	NUM
alces-343	524	3	)	)	PUNCT
alces-343	524	4	population	population	NOUN
alces-343	524	5	30	30	NUM
alces-343	524	6	in	in	ADP
alces-343	524	7	the	the	DET
alces-343	524	8	study	study	NOUN
alces-343	524	9	area	area	NOUN
alces-343	524	10	at	at	ADP
alces-343	524	11	time	time	NOUN
alces-343	524	12	0	0	NUM
alces-343	524	13	;	;	PUNCT
alces-343	524	14	calculated	calculate	VERB
alces-343	524	15	by	by	ADP
alces-343	524	16	the	the	DET
alces-343	524	17	model	model	NOUN
alces-343	524	18	afterwards	afterwards	ADV
alces-343	524	19	gingras	gingras	PROPN
alces-343	524	20	et	et	PROPN
alces-343	524	21	al	al	PROPN
alces-343	524	22	.	.	PROPN
alces-343	525	1	(	(	PUNCT
alces-343	525	2	1989	1989	NUM
alces-343	525	3	)	)	PUNCT
alces-343	525	4	maximum	maximum	ADJ
alces-343	525	5	growth	growth	NOUN
alces-343	525	6	rate	rate	NOUN
alces-343	525	7	25	25	NUM
alces-343	525	8	%	%	NOUN
alces-343	525	9	per	per	ADP
alces-343	525	10	year	year	NOUN
alces-343	525	11	fryxell	fryxell	NOUN
alces-343	525	12	et	et	PROPN
alces-343	525	13	al	al	PROPN
alces-343	525	14	.	.	PROPN
alces-343	526	1	(	(	PUNCT
alces-343	526	2	1988	1988	NUM
alces-343	526	3	)	)	PUNCT
alces-343	526	4	annual	annual	ADJ
alces-343	526	5	predation	predation	NOUN
alces-343	526	6	rate	rate	NOUN
alces-343	526	7	calculated	calculate	VERB
alces-343	526	8	from	from	ADP
alces-343	526	9	messier	messy	ADJ
alces-343	526	10	’s	’s	PART
alces-343	526	11	predation	predation	NOUN
alces-343	526	12	model	model	NOUN
alces-343	526	13	(	(	PUNCT
alces-343	526	14	depends	depend	VERB
alces-343	526	15	on	on	ADP
alces-343	526	16	moose	moose	NOUN
alces-343	526	17	density	density	NOUN
alces-343	526	18	)	)	PUNCT
alces-343	526	19	messier	messy	ADJ
alces-343	526	20	(	(	PUNCT
alces-343	526	21	1994	1994	NUM
alces-343	526	22	)	)	PUNCT
alces-343	526	23	annual	annual	ADJ
alces-343	526	24	hunting	hunting	NOUN
alces-343	526	25	rate	rate	NOUN
alces-343	526	26	9	9	NUM
alces-343	526	27	%	%	NOUN
alces-343	526	28	(	(	PUNCT
alces-343	526	29	simulations	simulation	NOUN
alces-343	526	30	1	1	NUM
alces-343	526	31	,	,	PUNCT
alces-343	526	32	2	2	NUM
alces-343	526	33	,	,	PUNCT
alces-343	526	34	3	3	NUM
alces-343	526	35	,	,	PUNCT
alces-343	526	36	5	5	NUM
alces-343	526	37	)	)	PUNCT
alces-343	526	38	or	or	CCONJ
alces-343	526	39	15	15	NUM
alces-343	526	40	%	%	NOUN
alces-343	526	41	(	(	PUNCT
alces-343	526	42	simulation	simulation	NOUN
alces-343	526	43	4	4	NUM
alces-343	526	44	)	)	PUNCT
alces-343	526	45	courtois	courtois	PROPN
alces-343	526	46	et	et	PROPN
alces-343	526	47	al	al	PROPN
alces-343	526	48	.	.	PROPN
alces-343	527	1	(	(	PUNCT
alces-343	527	2	1994b	1994b	NUM
alces-343	527	3	)	)	PUNCT
alces-343	527	4	annual	annual	ADJ
alces-343	527	5	rate	rate	NOUN
alces-343	527	6	of	of	ADP
alces-343	527	7	mortality	mortality	NOUN
alces-343	527	8	due	due	ADP
alces-343	527	9	to	to	ADP
alces-343	527	10	other	other	ADJ
alces-343	527	11	causes	cause	NOUN
alces-343	527	12	4.50	4.50	NUM
alces-343	527	13	%	%	NOUN
alces-343	527	14	courtois	courtois	NOUN
alces-343	527	15	et	et	PROPN
alces-343	527	16	al	al	PROPN
alces-343	527	17	.	.	PROPN
alces-343	528	1	(	(	PUNCT
alces-343	528	2	1994b	1994b	NUM
alces-343	528	3	)	)	PUNCT
alces-343	528	4	annual	annual	ADJ
alces-343	528	5	variability	variability	NOUN
alces-343	528	6	of	of	ADP
alces-343	528	7	recruitment	recruitment	NOUN
alces-343	528	8	0	0	NUM
alces-343	528	9	%	%	NOUN
alces-343	528	10	(	(	PUNCT
alces-343	528	11	simulations	simulation	NOUN
alces-343	528	12	1	1	NUM
alces-343	528	13	to	to	PART
alces-343	528	14	4	4	NUM
alces-343	528	15	)	)	PUNCT
alces-343	528	16	;	;	PUNCT
alces-343	528	17	random	random	ADJ
alces-343	528	18	variation	variation	NOUN
alces-343	528	19	between	between	ADP
alces-343	528	20	0	0	NUM
alces-343	528	21	and	and	CCONJ
alces-343	528	22	40	40	NUM
alces-343	528	23	%	%	NOUN
alces-343	528	24	(	(	PUNCT
alces-343	528	25	simulation	simulation	NOUN
alces-343	528	26	5	5	NUM
alces-343	528	27	)	)	PUNCT
alces-343	528	28	crête	crête	NOUN
alces-343	528	29	and	and	CCONJ
alces-343	528	30	courtois	courtois	PROPN
alces-343	528	31	(	(	PUNCT
alces-343	528	32	1997	1997	NUM
alces-343	528	33	)	)	PUNCT
alces-343	528	34	wolf	wolf	NOUN
alces-343	528	35	population	population	NOUN
alces-343	528	36	calculated	calculate	VERB
alces-343	528	37	from	from	ADP
alces-343	528	38	messier	messy	ADJ
alces-343	528	39	’s	’s	PART
alces-343	528	40	michaelismenten	michaelismenten	VERB
alces-343	528	41	hyperbolic	hyperbolic	ADJ
alces-343	528	42	equation	equation	NOUN
alces-343	528	43	(	(	PUNCT
alces-343	528	44	depends	depend	VERB
alces-343	528	45	on	on	ADP
alces-343	528	46	moose	moose	NOUN
alces-343	528	47	population	population	NOUN
alces-343	528	48	)	)	PUNCT
alces-343	528	49	messier	messy	ADJ
alces-343	528	50	(	(	PUNCT
alces-343	528	51	1994	1994	NUM
alces-343	528	52	)	)	PUNCT
alces-343	528	53	trapping	trap	VERB
alces-343	528	54	rate	rate	NOUN
alces-343	528	55	0	0	NUM
alces-343	528	56	%	%	NOUN
alces-343	528	57	(	(	PUNCT
alces-343	528	58	simulations	simulation	NOUN
alces-343	528	59	1	1	NUM
alces-343	528	60	,	,	PUNCT
alces-343	528	61	2	2	NUM
alces-343	528	62	,	,	PUNCT
alces-343	528	63	4	4	NUM
alces-343	528	64	,	,	PUNCT
alces-343	528	65	5	5	NUM
alces-343	528	66	)	)	PUNCT
alces-343	528	67	or	or	CCONJ
alces-343	528	68	30	30	NUM
alces-343	528	69	%	%	NOUN
alces-343	528	70	(	(	PUNCT
alces-343	528	71	simulation	simulation	NOUN
alces-343	528	72	3	3	NUM
alces-343	528	73	)	)	PUNCT
alces-343	528	74	larivière	larivière	NOUN
alces-343	528	75	et	et	PROPN
alces-343	528	76	al	al	PROPN
alces-343	528	77	.	.	PROPN
alces-343	529	1	(	(	PUNCT
alces-343	529	2	2000	2000	NUM
alces-343	529	3	)	)	PUNCT
alces-343	529	4	caribou	caribou	NOUN
alces-343	529	5	habitat	habitat	NOUN
alces-343	529	6	carrying	carry	VERB
alces-343	529	7	capacity	capacity	NOUN
alces-343	529	8	200	200	NUM
alces-343	529	9	in	in	ADP
alces-343	529	10	the	the	DET
alces-343	529	11	study	study	NOUN
alces-343	529	12	area	area	NOUN
alces-343	529	13	(	(	PUNCT
alces-343	529	14	20	20	NUM
alces-343	529	15	per	per	ADP
alces-343	529	16	100	100	NUM
alces-343	529	17	km2	km2	NUM
alces-343	529	18	)	)	PUNCT
alces-343	529	19	arsenault	arsenault	PROPN
alces-343	529	20	et	et	PROPN
alces-343	529	21	al	al	PROPN
alces-343	529	22	.	.	PROPN
alces-343	530	1	(	(	PUNCT
alces-343	530	2	1997	1997	NUM
alces-343	530	3	)	)	PUNCT
alces-343	530	4	population	population	NOUN
alces-343	530	5	163	163	NUM
alces-343	530	6	in	in	ADP
alces-343	530	7	the	the	DET
alces-343	530	8	study	study	NOUN
alces-343	530	9	area	area	NOUN
alces-343	530	10	at	at	ADP
alces-343	530	11	time	time	NOUN
alces-343	530	12	0	0	NUM
alces-343	530	13	;	;	PUNCT
alces-343	530	14	calculated	calculate	VERB
alces-343	530	15	by	by	ADP
alces-343	530	16	the	the	DET
alces-343	530	17	model	model	NOUN
alces-343	530	18	afterwards	afterwards	ADV
alces-343	530	19	courtois	courtois	PROPN
alces-343	530	20	et	et	PROPN
alces-343	530	21	al	al	PROPN
alces-343	530	22	.	.	PROPN
alces-343	531	1	(	(	PUNCT
alces-343	531	2	2003	2003	NUM
alces-343	531	3	)	)	PUNCT
alces-343	531	4	maximum	maximum	ADJ
alces-343	531	5	growth	growth	NOUN
alces-343	531	6	rate	rate	NOUN
alces-343	531	7	24.5	24.5	NUM
alces-343	531	8	%	%	NOUN
alces-343	531	9	per	per	ADP
alces-343	531	10	year	year	NOUN
alces-343	531	11	estimated	estimate	VERB
alces-343	531	12	from	from	ADP
alces-343	531	13	ouellet	ouellet	PROPN
alces-343	531	14	et	et	PROPN
alces-343	531	15	al	al	PROPN
alces-343	531	16	.	.	PROPN
alces-343	531	17	(	(	PUNCT
alces-343	531	18	1996	1996	NUM
alces-343	531	19	,	,	PUNCT
alces-343	531	20	1997	1997	NUM
alces-343	531	21	)	)	PUNCT
alces-343	531	22	data	datum	NOUN
alces-343	531	23	annual	annual	ADJ
alces-343	531	24	recruitment	recruitment	NOUN
alces-343	531	25	rate	rate	NOUN
alces-343	531	26	calculated	calculate	VERB
alces-343	531	27	from	from	ADP
alces-343	531	28	bergerud	bergerud	NOUN
alces-343	531	29	and	and	CCONJ
alces-343	531	30	elliot	elliot	PROPN
alces-343	531	31	’s	’s	PART
alces-343	531	32	model	model	NOUN
alces-343	531	33	(	(	PUNCT
alces-343	531	34	depends	depend	VERB
alces-343	531	35	on	on	ADP
alces-343	531	36	wolf	wolf	PROPN
alces-343	531	37	density	density	NOUN
alces-343	531	38	)	)	PUNCT
alces-343	531	39	bergerud	bergerud	NOUN
alces-343	531	40	and	and	CCONJ
alces-343	531	41	elliot	elliot	PROPN
alces-343	531	42	(	(	PUNCT
alces-343	531	43	1986	1986	NUM
alces-343	531	44	)	)	PUNCT
alces-343	531	45	annual	annual	ADJ
alces-343	531	46	rate	rate	NOUN
alces-343	531	47	of	of	ADP
alces-343	531	48	adult	adult	NOUN
alces-343	531	49	natural	natural	ADJ
alces-343	531	50	mortality	mortality	NOUN
alces-343	531	51	calculated	calculate	VERB
alces-343	531	52	from	from	ADP
alces-343	531	53	bergerud	bergerud	NOUN
alces-343	531	54	and	and	CCONJ
alces-343	531	55	elliot	elliot	PROPN
alces-343	531	56	’s	’s	PART
alces-343	531	57	model	model	NOUN
alces-343	531	58	(	(	PUNCT
alces-343	531	59	depends	depend	VERB
alces-343	531	60	on	on	ADP
alces-343	531	61	wolf	wolf	PROPN
alces-343	531	62	density	density	NOUN
alces-343	531	63	)	)	PUNCT
alces-343	531	64	bergerud	bergerud	NOUN
alces-343	531	65	and	and	CCONJ
alces-343	531	66	elliot	elliot	PROPN
alces-343	531	67	(	(	PUNCT
alces-343	531	68	1986	1986	NUM
alces-343	531	69	)	)	PUNCT
alces-343	531	70	annual	annual	ADJ
alces-343	531	71	hunting	hunting	NOUN
alces-343	531	72	rate	rate	NOUN
alces-343	531	73	9	9	NUM
alces-343	531	74	%	%	NOUN
alces-343	531	75	(	(	PUNCT
alces-343	531	76	simulations	simulation	NOUN
alces-343	531	77	1	1	NUM
alces-343	531	78	,	,	PUNCT
alces-343	531	79	2	2	NUM
alces-343	531	80	,	,	PUNCT
alces-343	531	81	3	3	NUM
alces-343	531	82	,	,	PUNCT
alces-343	531	83	5	5	NUM
alces-343	531	84	)	)	PUNCT
alces-343	531	85	or	or	CCONJ
alces-343	531	86	15	15	NUM
alces-343	531	87	%	%	NOUN
alces-343	531	88	(	(	PUNCT
alces-343	531	89	simulation	simulation	NOUN
alces-343	531	90	4	4	NUM
alces-343	531	91	)	)	PUNCT
alces-343	531	92	courtois	courtois	PROPN
alces-343	531	93	et	et	PROPN
alces-343	531	94	al	al	PROPN
alces-343	531	95	.	.	PROPN
alces-343	532	1	(	(	PUNCT
alces-343	532	2	2003	2003	NUM
alces-343	532	3	)	)	PUNCT
alces-343	532	4	annual	annual	ADJ
alces-343	532	5	variability	variability	NOUN
alces-343	532	6	of	of	ADP
alces-343	532	7	recruitment	recruitment	NOUN
alces-343	532	8	0	0	NUM
alces-343	532	9	%	%	NOUN
alces-343	532	10	(	(	PUNCT
alces-343	532	11	simulations	simulation	NOUN
alces-343	532	12	1	1	NUM
alces-343	532	13	to	to	PART
alces-343	532	14	4	4	NUM
alces-343	532	15	,	,	PUNCT
alces-343	532	16	5a	5a	NUM
alces-343	532	17	and	and	CCONJ
alces-343	532	18	5b	5b	NUM
alces-343	532	19	)	)	PUNCT
alces-343	532	20	;	;	PUNCT
alces-343	532	21	random	random	ADJ
alces-343	532	22	variation	variation	NOUN
alces-343	532	23	between	between	ADP
alces-343	532	24	0	0	NUM
alces-343	532	25	and	and	CCONJ
alces-343	532	26	20	20	NUM
alces-343	532	27	%	%	NOUN
alces-343	532	28	(	(	PUNCT
alces-343	532	29	simulation	simulation	NOUN
alces-343	532	30	5c	5c	NOUN
alces-343	532	31	)	)	PUNCT
alces-343	532	32	arbitrary	arbitrary	ADJ