id	sid	tid	token	lemma	pos
alces-35	1	1	alces	alce	NOUN
alces-35	1	2	vol	vol	NOUN
alces-35	1	3	.	.	PROPN
alces-35	1	4	44	44	NUM
alces-35	1	5	,	,	PUNCT
alces-35	1	6	2008	2008	NUM
alces-35	1	7	härkönen	härkönen	NOUN
alces-35	1	8	et	et	PROPN
alces-35	1	9	al.moose	al.moose	X
alces-35	1	10	and	and	CCONJ
alces-35	1	11	aspen	aspen	VERB
alces-35	1	12	regeneration	regeneration	NOUN
alces-35	1	13	in	in	ADP
alces-35	1	14	finland	finland	PROPN
alces-35	1	15	31	31	NUM
alces-35	1	16	does	do	AUX
alces-35	1	17	moose	moose	NOUN
alces-35	1	18	browsing	browse	VERB
alces-35	1	19	threaten	threaten	ADJ
alces-35	1	20	european	european	ADJ
alces-35	1	21	aspen	aspen	PROPN
alces-35	1	22	regeneration	regeneration	NOUN
alces-35	1	23	in	in	ADP
alces-35	1	24	koli	koli	PROPN
alces-35	1	25	national	national	PROPN
alces-35	1	26	park	park	PROPN
alces-35	1	27	,	,	PUNCT
alces-35	1	28	finland	finland	PROPN
alces-35	1	29	?	?	PUNCT
alces-35	2	1	sauli	sauli	PROPN
alces-35	2	2	härkönen1	härkönen1	PROPN
alces-35	2	3	,	,	PUNCT
alces-35	2	4	kalle	kalle	PROPN
alces-35	2	5	eerikäinen1	eerikäinen1	PROPN
alces-35	2	6	,	,	PUNCT
alces-35	2	7	riikka	riikka	NOUN
alces-35	2	8	lähteenmäki2	lähteenmäki2	NOUN
alces-35	2	9	,	,	PUNCT
alces-35	2	10	and	and	CCONJ
alces-35	2	11	risto	risto	NOUN
alces-35	2	12	heikkilä3	heikkilä3	PROPN
alces-35	3	1	1finnish	1finnish	NUM
alces-35	3	2	forest	forest	NOUN
alces-35	3	3	research	research	NOUN
alces-35	3	4	institute	institute	PROPN
alces-35	3	5	,	,	PUNCT
alces-35	3	6	joensuu	joensuu	PROPN
alces-35	3	7	research	research	PROPN
alces-35	3	8	unit	unit	NOUN
alces-35	3	9	,	,	PUNCT
alces-35	3	10	p.o	p.o	PROPN
alces-35	3	11	.	.	PROPN
alces-35	3	12	box	box	PROPN
alces-35	3	13	68	68	NUM
alces-35	3	14	,	,	PUNCT
alces-35	3	15	fi-80101	fi-80101	NOUN
alces-35	3	16	joensuu	joensuu	NOUN
alces-35	3	17	,	,	PUNCT
alces-35	3	18	finland	finland	PROPN
alces-35	3	19	,	,	PUNCT
alces-35	3	20	email	email	NOUN
alces-35	3	21	:	:	PUNCT
alces-35	3	22	sauli.harkonen@metla.fi	sauli.harkonen@metla.fi	ADV
alces-35	3	23	;	;	PUNCT
alces-35	3	24	2university	2university	NUM
alces-35	3	25	of	of	ADP
alces-35	3	26	joensuu	joensuu	NOUN
alces-35	3	27	,	,	PUNCT
alces-35	3	28	faculty	faculty	NOUN
alces-35	3	29	of	of	ADP
alces-35	3	30	forest	forest	NOUN
alces-35	3	31	sciences	sciences	PROPN
alces-35	3	32	,	,	PUNCT
alces-35	3	33	p.o	p.o	PROPN
alces-35	3	34	.	.	PROPN
alces-35	3	35	box	box	PROPN
alces-35	3	36	111	111	NUM
alces-35	3	37	,	,	PUNCT
alces-35	3	38	fi-80101	fi-80101	NOUN
alces-35	3	39	joensuu	joensuu	NOUN
alces-35	3	40	,	,	PUNCT
alces-35	3	41	finland	finland	PROPN
alces-35	3	42	;	;	PUNCT
alces-35	3	43	3finnish	3finnish	NUM
alces-35	3	44	forest	forest	NOUN
alces-35	3	45	research	research	PROPN
alces-35	3	46	institute	institute	PROPN
alces-35	3	47	,	,	PUNCT
alces-35	3	48	vantaa	vantaa	PROPN
alces-35	3	49	research	research	NOUN
alces-35	3	50	unit	unit	NOUN
alces-35	3	51	,	,	PUNCT
alces-35	3	52	p.o	p.o	PROPN
alces-35	3	53	.	.	PROPN
alces-35	3	54	box	box	PROPN
alces-35	3	55	18	18	NUM
alces-35	3	56	,	,	PUNCT
alces-35	3	57	fi-01301	fi-01301	NOUN
alces-35	3	58	vantaa	vantaa	NOUN
alces-35	3	59	,	,	PUNCT
alces-35	3	60	finland	finland	PROPN
alces-35	3	61	abstract	abstract	NOUN
alces-35	3	62	:	:	PUNCT
alces-35	3	63	large	large	ADJ
alces-35	3	64	european	european	PROPN
alces-35	3	65	aspen	aspen	PROPN
alces-35	3	66	(	(	PUNCT
alces-35	3	67	populus	populus	PROPN
alces-35	3	68	tremula	tremula	NOUN
alces-35	3	69	)	)	PUNCT
alces-35	3	70	trees	tree	NOUN
alces-35	3	71	host	host	VERB
alces-35	3	72	hundreds	hundred	NOUN
alces-35	3	73	of	of	ADP
alces-35	3	74	species	specie	NOUN
alces-35	3	75	of	of	ADP
alces-35	3	76	which	which	PRON
alces-35	3	77	many	many	ADJ
alces-35	3	78	are	be	AUX
alces-35	3	79	threatened	threaten	VERB
alces-35	3	80	species	specie	NOUN
alces-35	3	81	of	of	ADP
alces-35	3	82	conifer	conifer	NOUN
alces-35	3	83	-	-	PUNCT
alces-35	3	84	dominated	dominate	VERB
alces-35	3	85	,	,	PUNCT
alces-35	3	86	old	old	ADJ
alces-35	3	87	-	-	PUNCT
alces-35	3	88	growth	growth	NOUN
alces-35	3	89	boreal	boreal	NOUN
alces-35	3	90	forests	forest	NOUN
alces-35	3	91	.	.	PUNCT
alces-35	4	1	aspen	aspen	PROPN
alces-35	4	2	is	be	AUX
alces-35	4	3	also	also	ADV
alces-35	4	4	one	one	NUM
alces-35	4	5	of	of	ADP
alces-35	4	6	the	the	DET
alces-35	4	7	deciduous	deciduous	ADJ
alces-35	4	8	tree	tree	NOUN
alces-35	4	9	species	specie	NOUN
alces-35	4	10	most	most	ADV
alces-35	4	11	intensively	intensively	ADV
alces-35	4	12	used	use	VERB
alces-35	4	13	by	by	ADP
alces-35	4	14	moose	moose	NOUN
alces-35	4	15	(	(	PUNCT
alces-35	4	16	alces	alce	NOUN
alces-35	4	17	alces	alce	NOUN
alces-35	4	18	)	)	PUNCT
alces-35	4	19	in	in	ADP
alces-35	4	20	finland	finland	PROPN
alces-35	4	21	.	.	PUNCT
alces-35	5	1	in	in	ADP
alces-35	5	2	conservation	conservation	NOUN
alces-35	5	3	areas	area	NOUN
alces-35	5	4	aspen	aspen	VERB
alces-35	5	5	regeneration	regeneration	NOUN
alces-35	5	6	is	be	AUX
alces-35	5	7	facilitated	facilitate	VERB
alces-35	5	8	by	by	ADP
alces-35	5	9	large	large	ADJ
alces-35	5	10	-	-	PUNCT
alces-35	5	11	scale	scale	NOUN
alces-35	5	12	disturbances	disturbance	NOUN
alces-35	5	13	,	,	PUNCT
alces-35	5	14	especially	especially	ADV
alces-35	5	15	fires	fire	NOUN
alces-35	5	16	and	and	CCONJ
alces-35	5	17	windstorms	windstorm	NOUN
alces-35	5	18	,	,	PUNCT
alces-35	5	19	and	and	CCONJ
alces-35	5	20	also	also	ADV
alces-35	5	21	by	by	ADP
alces-35	5	22	mortality	mortality	NOUN
alces-35	5	23	of	of	ADP
alces-35	5	24	individual	individual	ADJ
alces-35	5	25	trees	tree	NOUN
alces-35	5	26	and	and	CCONJ
alces-35	5	27	small	small	ADJ
alces-35	5	28	-	-	PUNCT
alces-35	5	29	scale	scale	NOUN
alces-35	5	30	disturbances	disturbance	NOUN
alces-35	5	31	that	that	PRON
alces-35	5	32	create	create	VERB
alces-35	5	33	small	small	ADJ
alces-35	5	34	openings	opening	NOUN
alces-35	5	35	.	.	PUNCT
alces-35	6	1	these	these	DET
alces-35	6	2	aggregated	aggregate	VERB
alces-35	6	3	patches	patch	NOUN
alces-35	6	4	of	of	ADP
alces-35	6	5	young	young	ADJ
alces-35	6	6	aspens	aspen	NOUN
alces-35	6	7	provide	provide	VERB
alces-35	6	8	high	high	ADJ
alces-35	6	9	quality	quality	NOUN
alces-35	6	10	feeding	feeding	NOUN
alces-35	6	11	sites	site	NOUN
alces-35	6	12	for	for	ADP
alces-35	6	13	moose	moose	NOUN
alces-35	6	14	.	.	PUNCT
alces-35	7	1	in	in	ADP
alces-35	7	2	finland	finland	PROPN
alces-35	7	3	,	,	PUNCT
alces-35	7	4	it	it	PRON
alces-35	7	5	has	have	AUX
alces-35	7	6	been	be	AUX
alces-35	7	7	hypothesized	hypothesize	VERB
alces-35	7	8	that	that	SCONJ
alces-35	7	9	intense	intense	ADJ
alces-35	7	10	browsing	browsing	NOUN
alces-35	7	11	pressure	pressure	NOUN
alces-35	7	12	by	by	ADP
alces-35	7	13	moose	moose	NOUN
alces-35	7	14	on	on	ADP
alces-35	7	15	aspen	aspen	PROPN
alces-35	7	16	may	may	AUX
alces-35	7	17	prevent	prevent	VERB
alces-35	7	18	new	new	ADJ
alces-35	7	19	aspen	aspen	NOUN
alces-35	7	20	cohorts	cohort	NOUN
alces-35	7	21	from	from	ADP
alces-35	7	22	emerging	emerge	VERB
alces-35	7	23	,	,	PUNCT
alces-35	7	24	and	and	CCONJ
alces-35	7	25	thus	thus	ADV
alces-35	7	26	endanger	endanger	VERB
alces-35	7	27	the	the	DET
alces-35	7	28	spatio	spatio	PROPN
alces-35	7	29	-	-	PUNCT
alces-35	7	30	temporal	temporal	ADJ
alces-35	7	31	continuum	continuum	NOUN
alces-35	7	32	of	of	ADP
alces-35	7	33	aspen	aspen	ADJ
alces-35	7	34	occurrence	occurrence	NOUN
alces-35	7	35	in	in	ADP
alces-35	7	36	the	the	DET
alces-35	7	37	long	long	ADJ
alces-35	7	38	term	term	NOUN
alces-35	7	39	.	.	PUNCT
alces-35	8	1	the	the	DET
alces-35	8	2	aim	aim	NOUN
alces-35	8	3	of	of	ADP
alces-35	8	4	this	this	DET
alces-35	8	5	study	study	NOUN
alces-35	8	6	was	be	AUX
alces-35	8	7	to	to	PART
alces-35	8	8	analyze	analyze	VERB
alces-35	8	9	the	the	DET
alces-35	8	10	influence	influence	NOUN
alces-35	8	11	of	of	ADP
alces-35	8	12	moose	moose	NOUN
alces-35	8	13	browsing	browse	VERB
alces-35	8	14	on	on	ADP
alces-35	8	15	the	the	DET
alces-35	8	16	regeneration	regeneration	NOUN
alces-35	8	17	of	of	ADP
alces-35	8	18	aspen	aspen	NOUN
alces-35	8	19	in	in	ADP
alces-35	8	20	koli	koli	PROPN
alces-35	8	21	national	national	PROPN
alces-35	8	22	park	park	PROPN
alces-35	8	23	in	in	ADP
alces-35	8	24	eastern	eastern	PROPN
alces-35	8	25	finland	finland	PROPN
alces-35	8	26	at	at	ADP
alces-35	8	27	2	2	NUM
alces-35	8	28	different	different	ADJ
alces-35	8	29	spatial	spatial	ADJ
alces-35	8	30	scales	scale	NOUN
alces-35	8	31	,	,	PUNCT
alces-35	8	32	the	the	DET
alces-35	8	33	landscape	landscape	NOUN
alces-35	8	34	level	level	NOUN
alces-35	8	35	and	and	CCONJ
alces-35	8	36	stand	stand	VERB
alces-35	8	37	level	level	NOUN
alces-35	8	38	.	.	PUNCT
alces-35	9	1	our	our	PRON
alces-35	9	2	results	result	NOUN
alces-35	9	3	indicated	indicate	VERB
alces-35	9	4	that	that	SCONJ
alces-35	9	5	moose	moose	NOUN
alces-35	9	6	browsing	browse	VERB
alces-35	9	7	on	on	ADP
alces-35	9	8	aspen	aspen	NOUN
alces-35	9	9	has	have	AUX
alces-35	9	10	been	be	AUX
alces-35	9	11	very	very	ADV
alces-35	9	12	intense	intense	ADJ
alces-35	9	13	in	in	ADP
alces-35	9	14	the	the	DET
alces-35	9	15	area	area	NOUN
alces-35	9	16	.	.	PUNCT
alces-35	10	1	at	at	ADP
alces-35	10	2	the	the	DET
alces-35	10	3	landscape	landscape	NOUN
alces-35	10	4	level	level	NOUN
alces-35	10	5	,	,	PUNCT
alces-35	10	6	moose	moose	NOUN
alces-35	10	7	damaged	damage	VERB
alces-35	10	8	(	(	PUNCT
alces-35	10	9	twig	twig	NOUN
alces-35	10	10	-	-	PUNCT
alces-35	10	11	browsing	browsing	NOUN
alces-35	10	12	,	,	PUNCT
alces-35	10	13	stem	stem	NOUN
alces-35	10	14	breakage	breakage	NOUN
alces-35	10	15	,	,	PUNCT
alces-35	10	16	or	or	CCONJ
alces-35	10	17	bark	bark	NOUN
alces-35	10	18	stripping	strip	VERB
alces-35	10	19	)	)	PUNCT
alces-35	10	20	96	96	NUM
alces-35	10	21	%	%	NOUN
alces-35	10	22	of	of	ADP
alces-35	10	23	aspens	aspen	NOUN
alces-35	10	24	in	in	ADP
alces-35	10	25	the	the	DET
alces-35	10	26	southern	southern	ADJ
alces-35	10	27	area	area	NOUN
alces-35	10	28	and	and	CCONJ
alces-35	10	29	62	62	NUM
alces-35	10	30	%	%	NOUN
alces-35	10	31	in	in	ADP
alces-35	10	32	the	the	DET
alces-35	10	33	northern	northern	ADJ
alces-35	10	34	area	area	NOUN
alces-35	10	35	of	of	ADP
alces-35	10	36	the	the	DET
alces-35	10	37	park	park	NOUN
alces-35	10	38	.	.	PUNCT
alces-35	11	1	in	in	ADP
alces-35	11	2	addition	addition	NOUN
alces-35	11	3	,	,	PUNCT
alces-35	11	4	23	23	NUM
alces-35	11	5	%	%	NOUN
alces-35	11	6	of	of	ADP
alces-35	11	7	the	the	DET
alces-35	11	8	damaged	damage	VERB
alces-35	11	9	aspens	aspen	NOUN
alces-35	11	10	(	(	PUNCT
alces-35	11	11	all	all	PRON
alces-35	11	12	<	<	X
alces-35	11	13	5	5	NUM
alces-35	11	14	m	m	NOUN
alces-35	11	15	)	)	PUNCT
alces-35	11	16	were	be	AUX
alces-35	11	17	dead	dead	ADJ
alces-35	11	18	in	in	ADP
alces-35	11	19	the	the	DET
alces-35	11	20	southern	southern	ADJ
alces-35	11	21	area	area	NOUN
alces-35	11	22	.	.	PUNCT
alces-35	12	1	according	accord	VERB
alces-35	12	2	to	to	ADP
alces-35	12	3	counts	count	NOUN
alces-35	12	4	of	of	ADP
alces-35	12	5	fecal	fecal	ADJ
alces-35	12	6	pellet	pellet	NOUN
alces-35	12	7	groups	group	NOUN
alces-35	12	8	,	,	PUNCT
alces-35	12	9	moose	moose	NOUN
alces-35	12	10	activity	activity	NOUN
alces-35	12	11	was	be	AUX
alces-35	12	12	higher	high	ADJ
alces-35	12	13	in	in	ADP
alces-35	12	14	the	the	DET
alces-35	12	15	southern	southern	ADJ
alces-35	12	16	area	area	NOUN
alces-35	12	17	than	than	ADP
alces-35	12	18	the	the	DET
alces-35	12	19	northern	northern	ADJ
alces-35	12	20	area	area	NOUN
alces-35	12	21	.	.	PUNCT
alces-35	13	1	at	at	ADP
alces-35	13	2	the	the	DET
alces-35	13	3	stand	stand	NOUN
alces-35	13	4	level	level	NOUN
alces-35	13	5	,	,	PUNCT
alces-35	13	6	on	on	ADP
alces-35	13	7	average	average	ADJ
alces-35	13	8	,	,	PUNCT
alces-35	13	9	79	79	NUM
alces-35	13	10	%	%	NOUN
alces-35	13	11	of	of	ADP
alces-35	13	12	the	the	DET
alces-35	13	13	aspens	aspen	NOUN
alces-35	13	14	in	in	ADP
alces-35	13	15	the	the	DET
alces-35	13	16	southern	southern	ADJ
alces-35	13	17	area	area	NOUN
alces-35	13	18	and	and	CCONJ
alces-35	13	19	73	73	NUM
alces-35	13	20	%	%	NOUN
alces-35	13	21	in	in	ADP
alces-35	13	22	the	the	DET
alces-35	13	23	northern	northern	ADJ
alces-35	13	24	area	area	NOUN
alces-35	13	25	were	be	AUX
alces-35	13	26	damaged	damage	VERB
alces-35	13	27	.	.	PUNCT
alces-35	14	1	the	the	DET
alces-35	14	2	proportion	proportion	NOUN
alces-35	14	3	of	of	ADP
alces-35	14	4	dead	dead	ADJ
alces-35	14	5	aspens	aspen	NOUN
alces-35	14	6	(	(	PUNCT
alces-35	14	7	35	35	NUM
alces-35	14	8	%	%	NOUN
alces-35	14	9	)	)	PUNCT
alces-35	14	10	was	be	AUX
alces-35	14	11	highest	high	ADJ
alces-35	14	12	in	in	ADP
alces-35	14	13	stands	stand	NOUN
alces-35	14	14	in	in	ADP
alces-35	14	15	height	height	NOUN
alces-35	14	16	category	category	NOUN
alces-35	14	17	of	of	ADP
alces-35	14	18	5	5	NUM
alces-35	14	19	-	-	SYM
alces-35	14	20	15	15	NUM
alces-35	14	21	m.	m.	NOUN
alces-35	14	22	aspen	aspen	PROPN
alces-35	14	23	density	density	NOUN
alces-35	14	24	declined	decline	VERB
alces-35	14	25	from	from	ADP
alces-35	14	26	young	young	ADJ
alces-35	14	27	to	to	ADP
alces-35	14	28	old	old	ADJ
alces-35	14	29	stands	stand	NOUN
alces-35	14	30	in	in	ADP
alces-35	14	31	both	both	DET
alces-35	14	32	areas	area	NOUN
alces-35	14	33	.	.	PUNCT
alces-35	15	1	bark	bark	NOUN
alces-35	15	2	stripping	strip	VERB
alces-35	15	3	was	be	AUX
alces-35	15	4	relatively	relatively	ADV
alces-35	15	5	common	common	ADJ
alces-35	15	6	in	in	ADP
alces-35	15	7	the	the	DET
alces-35	15	8	height	height	NOUN
alces-35	15	9	category	category	NOUN
alces-35	15	10	of	of	ADP
alces-35	15	11	5	5	NUM
alces-35	15	12	-	-	SYM
alces-35	15	13	15	15	NUM
alces-35	15	14	m	m	NOUN
alces-35	15	15	over	over	ADP
alces-35	15	16	the	the	DET
alces-35	15	17	whole	whole	ADJ
alces-35	15	18	area	area	NOUN
alces-35	15	19	.	.	PUNCT
alces-35	16	1	we	we	PRON
alces-35	16	2	concluded	conclude	VERB
alces-35	16	3	that	that	SCONJ
alces-35	16	4	the	the	DET
alces-35	16	5	current	current	ADJ
alces-35	16	6	browsing	browse	VERB
alces-35	16	7	pressure	pressure	NOUN
alces-35	16	8	retards	retard	VERB
alces-35	16	9	the	the	DET
alces-35	16	10	height	height	NOUN
alces-35	16	11	development	development	NOUN
alces-35	16	12	of	of	ADP
alces-35	16	13	young	young	ADJ
alces-35	16	14	aspens	aspen	NOUN
alces-35	16	15	because	because	SCONJ
alces-35	16	16	of	of	ADP
alces-35	16	17	the	the	DET
alces-35	16	18	repeated	repeat	VERB
alces-35	16	19	break	break	NOUN
alces-35	16	20	-	-	PUNCT
alces-35	16	21	off	off	NOUN
alces-35	16	22	of	of	ADP
alces-35	16	23	main	main	ADJ
alces-35	16	24	stems	stem	NOUN
alces-35	16	25	and	and	CCONJ
alces-35	16	26	leader	leader	NOUN
alces-35	16	27	shoots	shoot	NOUN
alces-35	16	28	.	.	PUNCT
alces-35	17	1	although	although	SCONJ
alces-35	17	2	occurrence	occurrence	NOUN
alces-35	17	3	of	of	ADP
alces-35	17	4	aspen	aspen	NOUN
alces-35	17	5	may	may	AUX
alces-35	17	6	decline	decline	VERB
alces-35	17	7	due	due	ADJ
alces-35	17	8	to	to	ADP
alces-35	17	9	high	high	ADJ
alces-35	17	10	browsing	browse	VERB
alces-35	17	11	pressure	pressure	NOUN
alces-35	17	12	by	by	ADP
alces-35	17	13	moose	moose	NOUN
alces-35	17	14	,	,	PUNCT
alces-35	17	15	the	the	DET
alces-35	17	16	majority	majority	NOUN
alces-35	17	17	of	of	ADP
alces-35	17	18	aspens	aspen	NOUN
alces-35	17	19	have	have	VERB
alces-35	17	20	excellent	excellent	ADJ
alces-35	17	21	tolerance	tolerance	NOUN
alces-35	17	22	to	to	PART
alces-35	17	23	heavy	heavy	ADJ
alces-35	17	24	and	and	CCONJ
alces-35	17	25	repeated	repeat	VERB
alces-35	17	26	browsing	browsing	NOUN
alces-35	17	27	.	.	PUNCT
alces-35	18	1	hence	hence	ADV
alces-35	18	2	,	,	PUNCT
alces-35	18	3	a	a	DET
alces-35	18	4	high	high	ADJ
alces-35	18	5	proportion	proportion	NOUN
alces-35	18	6	of	of	ADP
alces-35	18	7	aspens	aspen	NOUN
alces-35	18	8	may	may	AUX
alces-35	18	9	reach	reach	VERB
alces-35	18	10	maturity	maturity	NOUN
alces-35	18	11	,	,	PUNCT
alces-35	18	12	and	and	CCONJ
alces-35	18	13	thus	thus	ADV
alces-35	18	14	maintain	maintain	VERB
alces-35	18	15	the	the	DET
alces-35	18	16	spatio	spatio	PROPN
alces-35	18	17	-	-	PUNCT
alces-35	18	18	temporal	temporal	ADJ
alces-35	18	19	continuum	continuum	NOUN
alces-35	18	20	of	of	ADP
alces-35	18	21	aspen	aspen	ADJ
alces-35	18	22	occurrence	occurrence	NOUN
alces-35	18	23	at	at	ADP
alces-35	18	24	a	a	DET
alces-35	18	25	level	level	NOUN
alces-35	18	26	that	that	PRON
alces-35	18	27	contributes	contribute	VERB
alces-35	18	28	to	to	ADP
alces-35	18	29	its	its	PRON
alces-35	18	30	role	role	NOUN
alces-35	18	31	in	in	ADP
alces-35	18	32	community	community	NOUN
alces-35	18	33	dynamics	dynamic	NOUN
alces-35	18	34	and	and	CCONJ
alces-35	18	35	local	local	ADJ
alces-35	18	36	and	and	CCONJ
alces-35	18	37	regional	regional	ADJ
alces-35	18	38	biodiversity	biodiversity	NOUN
alces-35	18	39	.	.	PUNCT
alces-35	19	1	alces	alce	NOUN
alces-35	19	2	vol	vol	NOUN
alces-35	19	3	.	.	PROPN
alces-35	20	1	44	44	NUM
alces-35	20	2	:	:	PUNCT
alces-35	20	3	31	31	NUM
alces-35	20	4	-	-	SYM
alces-35	20	5	40	40	NUM
alces-35	20	6	(	(	PUNCT
alces-35	20	7	2008	2008	NUM
alces-35	20	8	)	)	PUNCT
alces-35	20	9	key	key	ADJ
alces-35	20	10	words	word	NOUN
alces-35	20	11	:	:	PUNCT
alces-35	20	12	alces	alce	NOUN
alces-35	20	13	alces	alce	NOUN
alces-35	20	14	,	,	PUNCT
alces-35	20	15	biodiversity	biodiversity	NOUN
alces-35	20	16	,	,	PUNCT
alces-35	20	17	browsing	browse	VERB
alces-35	20	18	,	,	PUNCT
alces-35	20	19	european	european	PROPN
alces-35	20	20	aspen	aspen	PROPN
alces-35	20	21	,	,	PUNCT
alces-35	20	22	moose	moose	NOUN
alces-35	20	23	,	,	PUNCT
alces-35	20	24	populus	populus	PROPN
alces-35	20	25	tremula	tremula	NOUN
alces-35	20	26	,	,	PUNCT
alces-35	20	27	regeneration	regeneration	NOUN
alces-35	20	28	browsing	browse	VERB
alces-35	20	29	by	by	ADP
alces-35	20	30	cervids	cervid	NOUN
alces-35	20	31	on	on	ADP
alces-35	20	32	tree	tree	NOUN
alces-35	20	33	seedlings	seedling	NOUN
alces-35	20	34	and	and	CCONJ
alces-35	20	35	saplings	sapling	NOUN
alces-35	20	36	has	have	AUX
alces-35	20	37	been	be	AUX
alces-35	20	38	regarded	regard	VERB
alces-35	20	39	as	as	ADP
alces-35	20	40	a	a	DET
alces-35	20	41	pivotal	pivotal	ADJ
alces-35	20	42	issue	issue	NOUN
alces-35	20	43	affecting	affect	VERB
alces-35	20	44	the	the	DET
alces-35	20	45	success	success	NOUN
alces-35	20	46	of	of	ADP
alces-35	20	47	forest	forest	NOUN
alces-35	20	48	regeneration	regeneration	NOUN
alces-35	20	49	in	in	ADP
alces-35	20	50	managed	manage	VERB
alces-35	20	51	commercial	commercial	ADJ
alces-35	20	52	forests	forest	NOUN
alces-35	20	53	and	and	CCONJ
alces-35	20	54	in	in	ADP
alces-35	20	55	nature	nature	NOUN
alces-35	20	56	conservation	conservation	NOUN
alces-35	20	57	areas	area	NOUN
alces-35	20	58	across	across	ADP
alces-35	20	59	europe	europe	PROPN
alces-35	20	60	(	(	PUNCT
alces-35	20	61	e.g.	e.g.	ADV
alces-35	20	62	,	,	PUNCT
alces-35	20	63	gill	gill	NOUN
alces-35	20	64	1992	1992	NUM
alces-35	20	65	,	,	PUNCT
alces-35	20	66	angelstam	angelstam	VERB
alces-35	20	67	et	et	PROPN
alces-35	20	68	al	al	PROPN
alces-35	20	69	.	.	PROPN
alces-35	20	70	2000	2000	NUM
alces-35	20	71	,	,	PUNCT
alces-35	20	72	heikkilä	heikkilä	VERB
alces-35	20	73	et	et	PROPN
alces-35	20	74	al	al	PROPN
alces-35	20	75	.	.	PROPN
alces-35	20	76	2003	2003	NUM
alces-35	20	77	,	,	PUNCT
alces-35	20	78	edenius	edenius	NOUN
alces-35	20	79	and	and	CCONJ
alces-35	20	80	ericsson	ericsson	NOUN
alces-35	20	81	2007	2007	NUM
alces-35	20	82	)	)	PUNCT
alces-35	20	83	and	and	CCONJ
alces-35	20	84	in	in	ADP
alces-35	20	85	northern	northern	PROPN
alces-35	20	86	america	america	PROPN
alces-35	20	87	(	(	PUNCT
alces-35	20	88	e.g.	e.g.	ADV
alces-35	20	89	,	,	PUNCT
alces-35	20	90	baker	baker	PROPN
alces-35	20	91	et	et	PROPN
alces-35	20	92	al	al	PROPN
alces-35	20	93	.	.	PROPN
alces-35	20	94	1997	1997	NUM
alces-35	20	95	,	,	PUNCT
alces-35	20	96	kay	kay	PROPN
alces-35	20	97	1997	1997	NUM
alces-35	20	98	,	,	PUNCT
alces-35	20	99	rooney	rooney	NOUN
alces-35	20	100	and	and	CCONJ
alces-35	20	101	waller	waller	PROPN
alces-35	20	102	2003	2003	NUM
alces-35	20	103	,	,	PUNCT
alces-35	20	104	mclaren	mclaren	PROPN
alces-35	20	105	et	et	PROPN
alces-35	20	106	al	al	PROPN
alces-35	20	107	.	.	PROPN
alces-35	20	108	2004	2004	NUM
alces-35	20	109	)	)	PUNCT
alces-35	20	110	.	.	PUNCT
alces-35	21	1	in	in	ADP
alces-35	21	2	finland	finland	PROPN
alces-35	21	3	,	,	PUNCT
alces-35	21	4	moose	moose	X
alces-35	21	5	(	(	PUNCT
alces-35	21	6	alces	alce	NOUN
alces-35	21	7	alces	alce	NOUN
alces-35	21	8	)	)	PUNCT
alces-35	21	9	density	density	NOUN
alces-35	21	10	has	have	AUX
alces-35	21	11	increased	increase	VERB
alces-35	21	12	continuously	continuously	ADV
alces-35	21	13	over	over	ADP
alces-35	21	14	the	the	DET
alces-35	21	15	last	last	ADJ
alces-35	21	16	few	few	ADJ
alces-35	21	17	decades	decade	NOUN
alces-35	21	18	(	(	PUNCT
alces-35	21	19	finnish	finnish	PROPN
alces-35	21	20	forest	forest	PROPN
alces-35	21	21	research	research	PROPN
alces-35	21	22	institute	institute	PROPN
alces-35	21	23	2006	2006	NUM
alces-35	21	24	)	)	PUNCT
alces-35	21	25	,	,	PUNCT
alces-35	21	26	which	which	PRON
alces-35	21	27	has	have	AUX
alces-35	21	28	led	lead	VERB
alces-35	21	29	to	to	ADP
alces-35	21	30	intense	intense	ADJ
alces-35	21	31	public	public	ADJ
alces-35	21	32	debate	debate	NOUN
alces-35	21	33	on	on	ADP
alces-35	21	34	the	the	DET
alces-35	21	35	need	need	NOUN
alces-35	21	36	to	to	PART
alces-35	21	37	search	search	VERB
alces-35	21	38	for	for	ADP
alces-35	21	39	balance	balance	NOUN
alces-35	21	40	between	between	ADP
alces-35	21	41	moose	moose	NOUN
alces-35	21	42	and	and	CCONJ
alces-35	21	43	forest	forest	NOUN
alces-35	21	44	management	management	NOUN
alces-35	21	45	,	,	PUNCT
alces-35	21	46	as	as	ADV
alces-35	21	47	well	well	ADV
alces-35	21	48	as	as	ADP
alces-35	21	49	nature	nature	NOUN
alces-35	21	50	conservation	conservation	NOUN
alces-35	21	51	.	.	PUNCT
alces-35	22	1	large	large	ADJ
alces-35	22	2	european	european	PROPN
alces-35	22	3	aspen	aspen	PROPN
alces-35	22	4	(	(	PUNCT
alces-35	22	5	populus	populus	PROPN
alces-35	22	6	tremula	tremula	NOUN
alces-35	22	7	)	)	PUNCT
alces-35	22	8	trees	tree	NOUN
alces-35	22	9	(	(	PUNCT
alces-35	22	10	i.e.	i.e.	X
alces-35	22	11	,	,	PUNCT
alces-35	22	12	living	living	NOUN
alces-35	22	13	and	and	CCONJ
alces-35	22	14	decaying	decay	VERB
alces-35	22	15	dead	dead	ADJ
alces-35	22	16	aspens	aspen	NOUN
alces-35	22	17	)	)	PUNCT
alces-35	22	18	moose	moose	NOUN
alces-35	22	19	and	and	CCONJ
alces-35	22	20	aspen	aspen	NOUN
alces-35	22	21	regeneration	regeneration	NOUN
alces-35	22	22	in	in	ADP
alces-35	22	23	finland	finland	PROPN
alces-35	22	24	härkönen	härkönen	PROPN
alces-35	22	25	et	et	PROPN
alces-35	22	26	al	al	PROPN
alces-35	22	27	.	.	PROPN
alces-35	23	1	alces	alce	NOUN
alces-35	23	2	vol	vol	NOUN
alces-35	23	3	.	.	PROPN
alces-35	23	4	44	44	NUM
alces-35	23	5	,	,	PUNCT
alces-35	24	1	2008	2008	NUM
alces-35	24	2	32	32	NUM
alces-35	24	3	host	host	NOUN
alces-35	24	4	hundreds	hundred	NOUN
alces-35	24	5	of	of	ADP
alces-35	24	6	herbivorous	herbivorous	ADJ
alces-35	24	7	and	and	CCONJ
alces-35	24	8	saproxylic	saproxylic	ADJ
alces-35	24	9	invertebrates	invertebrate	NOUN
alces-35	24	10	,	,	PUNCT
alces-35	24	11	polypore	polypore	NOUN
alces-35	24	12	fungi	fungi	PROPN
alces-35	24	13	,	,	PUNCT
alces-35	24	14	and	and	CCONJ
alces-35	24	15	epiphytic	epiphytic	ADJ
alces-35	24	16	lichens	lichen	NOUN
alces-35	24	17	of	of	ADP
alces-35	24	18	which	which	PRON
alces-35	24	19	many	many	ADJ
alces-35	24	20	are	be	AUX
alces-35	24	21	threatened	threaten	VERB
alces-35	24	22	species	specie	NOUN
alces-35	24	23	of	of	ADP
alces-35	24	24	conifer	conifer	NOUN
alces-35	24	25	-	-	PUNCT
alces-35	24	26	dominated	dominate	VERB
alces-35	24	27	,	,	PUNCT
alces-35	24	28	old	old	ADJ
alces-35	24	29	-	-	PUNCT
alces-35	24	30	growth	growth	NOUN
alces-35	24	31	boreal	boreal	NOUN
alces-35	24	32	forests	forest	NOUN
alces-35	24	33	(	(	PUNCT
alces-35	24	34	kouki	kouki	PROPN
alces-35	24	35	et	et	PROPN
alces-35	24	36	al	al	PROPN
alces-35	24	37	.	.	PROPN
alces-35	24	38	2004	2004	NUM
alces-35	24	39	)	)	PUNCT
alces-35	24	40	.	.	PUNCT
alces-35	25	1	it	it	PRON
alces-35	25	2	has	have	AUX
alces-35	25	3	been	be	AUX
alces-35	25	4	estimated	estimate	VERB
alces-35	25	5	that	that	SCONJ
alces-35	25	6	there	there	PRON
alces-35	25	7	are	be	VERB
alces-35	25	8	at	at	ADV
alces-35	25	9	least	least	ADJ
alces-35	25	10	150	150	NUM
alces-35	25	11	specialist	specialist	NOUN
alces-35	25	12	species	specie	NOUN
alces-35	25	13	that	that	PRON
alces-35	25	14	are	be	AUX
alces-35	25	15	entirely	entirely	ADV
alces-35	25	16	dependent	dependent	ADJ
alces-35	25	17	on	on	ADP
alces-35	25	18	aspen	aspen	NOUN
alces-35	25	19	in	in	ADP
alces-35	25	20	fennoscandia	fennoscandia	NOUN
alces-35	25	21	(	(	PUNCT
alces-35	25	22	siitonen	siitonen	NOUN
alces-35	25	23	1999	1999	NUM
alces-35	25	24	,	,	PUNCT
alces-35	25	25	kouki	kouki	PROPN
alces-35	25	26	et	et	PROPN
alces-35	25	27	al	al	PROPN
alces-35	25	28	.	.	PROPN
alces-35	25	29	2004	2004	NUM
alces-35	25	30	)	)	PUNCT
alces-35	25	31	.	.	PUNCT
alces-35	26	1	aspen	aspen	PROPN
alces-35	26	2	is	be	AUX
alces-35	26	3	also	also	ADV
alces-35	26	4	one	one	NUM
alces-35	26	5	of	of	ADP
alces-35	26	6	the	the	DET
alces-35	26	7	deciduous	deciduous	ADJ
alces-35	26	8	tree	tree	NOUN
alces-35	26	9	species	specie	NOUN
alces-35	26	10	most	most	ADV
alces-35	26	11	intensively	intensively	ADV
alces-35	26	12	used	use	VERB
alces-35	26	13	by	by	ADP
alces-35	26	14	large	large	ADJ
alces-35	26	15	herbivores	herbivore	NOUN
alces-35	26	16	,	,	PUNCT
alces-35	26	17	and	and	CCONJ
alces-35	26	18	ranks	rank	VERB
alces-35	26	19	among	among	ADP
alces-35	26	20	the	the	DET
alces-35	26	21	highly	highly	ADV
alces-35	26	22	preferred	preferred	ADJ
alces-35	26	23	woody	woody	NOUN
alces-35	26	24	plants	plant	NOUN
alces-35	26	25	in	in	ADP
alces-35	26	26	the	the	DET
alces-35	26	27	diet	diet	NOUN
alces-35	26	28	of	of	ADP
alces-35	26	29	moose	moose	NOUN
alces-35	26	30	(	(	PUNCT
alces-35	26	31	bergström	bergström	NOUN
alces-35	26	32	and	and	CCONJ
alces-35	26	33	hjeljord	hjeljord	NOUN
alces-35	26	34	1987	1987	NUM
alces-35	26	35	)	)	PUNCT
alces-35	26	36	.	.	PUNCT
alces-35	27	1	due	due	ADP
alces-35	27	2	to	to	ADP
alces-35	27	3	increasing	increase	VERB
alces-35	27	4	awareness	awareness	NOUN
alces-35	27	5	about	about	ADP
alces-35	27	6	explicit	explicit	ADJ
alces-35	27	7	ecological	ecological	ADJ
alces-35	27	8	values	value	NOUN
alces-35	27	9	of	of	ADP
alces-35	27	10	aspen	aspen	NOUN
alces-35	27	11	on	on	ADP
alces-35	27	12	forest	forest	NOUN
alces-35	27	13	biodiversity	biodiversity	NOUN
alces-35	27	14	,	,	PUNCT
alces-35	27	15	aspen	aspen	PROPN
alces-35	27	16	is	be	AUX
alces-35	27	17	receiving	receive	VERB
alces-35	27	18	more	more	ADJ
alces-35	27	19	consideration	consideration	NOUN
alces-35	27	20	in	in	ADP
alces-35	27	21	management	management	NOUN
alces-35	27	22	planning	planning	NOUN
alces-35	27	23	of	of	ADP
alces-35	27	24	commercial	commercial	ADJ
alces-35	27	25	forests	forest	NOUN
alces-35	27	26	,	,	PUNCT
alces-35	27	27	specifically	specifically	ADV
alces-35	27	28	in	in	ADP
alces-35	27	29	the	the	DET
alces-35	27	30	restoration	restoration	NOUN
alces-35	27	31	planning	planning	NOUN
alces-35	27	32	of	of	ADP
alces-35	27	33	formerly	formerly	ADV
alces-35	27	34	managed	manage	VERB
alces-35	27	35	forests	forest	NOUN
alces-35	27	36	within	within	ADP
alces-35	27	37	conservation	conservation	NOUN
alces-35	27	38	areas	area	NOUN
alces-35	27	39	.	.	PUNCT
alces-35	28	1	it	it	PRON
alces-35	28	2	has	have	AUX
alces-35	28	3	been	be	AUX
alces-35	28	4	estimated	estimate	VERB
alces-35	28	5	that	that	SCONJ
alces-35	28	6	only	only	ADV
alces-35	28	7	0.3	0.3	NUM
alces-35	28	8	%	%	NOUN
alces-35	28	9	of	of	ADP
alces-35	28	10	the	the	DET
alces-35	28	11	forest	forest	NOUN
alces-35	28	12	land	land	NOUN
alces-35	28	13	in	in	ADP
alces-35	28	14	finland	finland	PROPN
alces-35	28	15	is	be	AUX
alces-35	28	16	covered	cover	VERB
alces-35	28	17	by	by	ADP
alces-35	28	18	aspendominated	aspendominated	ADJ
alces-35	28	19	stands	stand	NOUN
alces-35	28	20	(	(	PUNCT
alces-35	28	21	finnish	finnish	PROPN
alces-35	28	22	forest	forest	PROPN
alces-35	28	23	research	research	PROPN
alces-35	28	24	institute	institute	PROPN
alces-35	28	25	2006	2006	NUM
alces-35	28	26	)	)	PUNCT
alces-35	28	27	.	.	PUNCT
alces-35	29	1	however	however	ADV
alces-35	29	2	,	,	PUNCT
alces-35	29	3	young	young	ADJ
alces-35	29	4	mixed	mixed	ADJ
alces-35	29	5	species	specie	NOUN
alces-35	29	6	stands	stand	VERB
alces-35	29	7	with	with	ADP
alces-35	29	8	aspen	aspen	NOUN
alces-35	29	9	trees	tree	NOUN
alces-35	29	10	are	be	AUX
alces-35	29	11	relatively	relatively	ADV
alces-35	29	12	common	common	ADJ
alces-35	29	13	,	,	PUNCT
alces-35	29	14	whereas	whereas	SCONJ
alces-35	29	15	densities	density	NOUN
alces-35	29	16	of	of	ADP
alces-35	29	17	large	large	ADJ
alces-35	29	18	aspens	aspen	NOUN
alces-35	29	19	are	be	AUX
alces-35	29	20	very	very	ADV
alces-35	29	21	low	low	ADJ
alces-35	29	22	in	in	ADP
alces-35	29	23	older	old	ADJ
alces-35	29	24	forests	forest	NOUN
alces-35	29	25	.	.	PUNCT
alces-35	30	1	according	accord	VERB
alces-35	30	2	to	to	ADP
alces-35	30	3	the	the	DET
alces-35	30	4	national	national	PROPN
alces-35	30	5	forest	forest	PROPN
alces-35	30	6	inventory	inventory	NOUN
alces-35	30	7	(	(	PUNCT
alces-35	30	8	nfi	nfi	NOUN
alces-35	30	9	)	)	PUNCT
alces-35	30	10	of	of	ADP
alces-35	30	11	finland	finland	PROPN
alces-35	30	12	,	,	PUNCT
alces-35	30	13	there	there	PRON
alces-35	30	14	are	be	VERB
alces-35	30	15	only	only	ADV
alces-35	30	16	0.35	0.35	NUM
alces-35	30	17	aspens	aspen	NOUN
alces-35	30	18	/	/	SYM
alces-35	30	19	ha	ha	INTJ
alces-35	30	20	with	with	ADP
alces-35	30	21	diameter	diameter	NOUN
alces-35	30	22	at	at	ADP
alces-35	30	23	breast	breast	NOUN
alces-35	30	24	height	height	NOUN
alces-35	30	25	>	>	PROPN
alces-35	30	26	30	30	NUM
alces-35	30	27	cm	cm	NOUN
alces-35	30	28	(	(	PUNCT
alces-35	30	29	finnish	finnish	PROPN
alces-35	30	30	forest	forest	PROPN
alces-35	30	31	research	research	PROPN
alces-35	30	32	institute	institute	PROPN
alces-35	30	33	2006	2006	NUM
alces-35	30	34	)	)	PUNCT
alces-35	30	35	.	.	PUNCT
alces-35	31	1	aspen	aspen	PROPN
alces-35	31	2	can	can	AUX
alces-35	31	3	regenerate	regenerate	VERB
alces-35	31	4	sexually	sexually	ADV
alces-35	31	5	by	by	ADP
alces-35	31	6	seed	seed	NOUN
alces-35	31	7	and	and	CCONJ
alces-35	31	8	asexually	asexually	ADV
alces-35	31	9	by	by	ADP
alces-35	31	10	root	root	NOUN
alces-35	31	11	suckers	sucker	NOUN
alces-35	31	12	.	.	PUNCT
alces-35	32	1	however	however	ADV
alces-35	32	2	,	,	PUNCT
alces-35	32	3	in	in	ADP
alces-35	32	4	the	the	DET
alces-35	32	5	finnish	finnish	ADJ
alces-35	32	6	landscape	landscape	NOUN
alces-35	32	7	asexual	asexual	ADJ
alces-35	32	8	type	type	NOUN
alces-35	32	9	of	of	ADP
alces-35	32	10	reproduction	reproduction	NOUN
alces-35	32	11	is	be	AUX
alces-35	32	12	more	more	ADV
alces-35	32	13	common	common	ADJ
alces-35	32	14	and	and	CCONJ
alces-35	32	15	in	in	ADP
alces-35	32	16	most	most	ADJ
alces-35	32	17	regeneration	regeneration	NOUN
alces-35	32	18	sites	site	NOUN
alces-35	32	19	practically	practically	ADV
alces-35	32	20	all	all	DET
alces-35	32	21	seedlings	seedling	NOUN
alces-35	32	22	originate	originate	VERB
alces-35	32	23	from	from	ADP
alces-35	32	24	root	root	NOUN
alces-35	32	25	suckers	sucker	NOUN
alces-35	32	26	(	(	PUNCT
alces-35	32	27	tikka	tikka	NOUN
alces-35	32	28	1955	1955	NUM
alces-35	32	29	)	)	PUNCT
alces-35	32	30	.	.	PUNCT
alces-35	33	1	some	some	DET
alces-35	33	2	disturbance	disturbance	NOUN
alces-35	33	3	is	be	AUX
alces-35	33	4	needed	need	VERB
alces-35	33	5	to	to	PART
alces-35	33	6	start	start	VERB
alces-35	33	7	aspen	aspen	NOUN
alces-35	33	8	regeneration	regeneration	NOUN
alces-35	33	9	.	.	PUNCT
alces-35	34	1	in	in	ADP
alces-35	34	2	natural	natural	ADJ
alces-35	34	3	forests	forest	NOUN
alces-35	34	4	,	,	PUNCT
alces-35	34	5	aspen	aspen	NOUN
alces-35	34	6	regeneration	regeneration	NOUN
alces-35	34	7	is	be	AUX
alces-35	34	8	facilitated	facilitate	VERB
alces-35	34	9	by	by	ADP
alces-35	34	10	large	large	ADJ
alces-35	34	11	-	-	PUNCT
alces-35	34	12	scale	scale	NOUN
alces-35	34	13	disturbances	disturbance	NOUN
alces-35	34	14	,	,	PUNCT
alces-35	34	15	especially	especially	ADV
alces-35	34	16	fires	fire	NOUN
alces-35	34	17	and	and	CCONJ
alces-35	34	18	windstorms	windstorm	NOUN
alces-35	34	19	,	,	PUNCT
alces-35	34	20	but	but	CCONJ
alces-35	34	21	also	also	ADV
alces-35	34	22	by	by	ADP
alces-35	34	23	the	the	DET
alces-35	34	24	death	death	NOUN
alces-35	34	25	of	of	ADP
alces-35	34	26	individual	individual	ADJ
alces-35	34	27	trees	tree	NOUN
alces-35	34	28	and	and	CCONJ
alces-35	34	29	small	small	ADJ
alces-35	34	30	-	-	PUNCT
alces-35	34	31	scale	scale	NOUN
alces-35	34	32	disturbances	disturbance	NOUN
alces-35	34	33	that	that	PRON
alces-35	34	34	create	create	VERB
alces-35	34	35	small	small	ADJ
alces-35	34	36	openings	opening	NOUN
alces-35	34	37	(	(	PUNCT
alces-35	34	38	syrjänen	syrjänen	NOUN
alces-35	34	39	et	et	PROPN
alces-35	34	40	al	al	PROPN
alces-35	34	41	.	.	PROPN
alces-35	34	42	1994	1994	NUM
alces-35	34	43	,	,	PUNCT
alces-35	34	44	cumming	cumme	VERB
alces-35	34	45	et	et	PROPN
alces-35	34	46	al	al	PROPN
alces-35	34	47	.	.	PROPN
alces-35	34	48	2000	2000	NUM
alces-35	34	49	)	)	PUNCT
alces-35	34	50	.	.	PUNCT
alces-35	35	1	these	these	PRON
alces-35	35	2	aggregated	aggregate	VERB
alces-35	35	3	patches	patch	NOUN
alces-35	35	4	of	of	ADP
alces-35	35	5	young	young	ADJ
alces-35	35	6	aspens	aspen	NOUN
alces-35	35	7	provide	provide	VERB
alces-35	35	8	high	high	ADJ
alces-35	35	9	quality	quality	NOUN
alces-35	35	10	feeding	feeding	NOUN
alces-35	35	11	sites	site	NOUN
alces-35	35	12	for	for	ADP
alces-35	35	13	moose	moose	NOUN
alces-35	35	14	.	.	PUNCT
alces-35	36	1	in	in	ADP
alces-35	36	2	finland	finland	PROPN
alces-35	36	3	,	,	PUNCT
alces-35	36	4	it	it	PRON
alces-35	36	5	has	have	AUX
alces-35	36	6	been	be	AUX
alces-35	36	7	hypothesized	hypothesize	VERB
alces-35	36	8	that	that	SCONJ
alces-35	36	9	intense	intense	ADJ
alces-35	36	10	browsing	browsing	NOUN
alces-35	36	11	pressure	pressure	NOUN
alces-35	36	12	by	by	ADP
alces-35	36	13	moose	moose	NOUN
alces-35	36	14	on	on	ADP
alces-35	36	15	preferred	preferred	ADJ
alces-35	36	16	aspens	aspen	NOUN
alces-35	36	17	may	may	AUX
alces-35	36	18	prevent	prevent	VERB
alces-35	36	19	new	new	ADJ
alces-35	36	20	aspen	aspen	NOUN
alces-35	36	21	cohorts	cohort	NOUN
alces-35	36	22	from	from	ADP
alces-35	36	23	emerging	emerge	VERB
alces-35	36	24	(	(	PUNCT
alces-35	36	25	kouki	kouki	PROPN
alces-35	36	26	et	et	PROPN
alces-35	36	27	al	al	PROPN
alces-35	36	28	.	.	PROPN
alces-35	36	29	2004	2004	NUM
alces-35	36	30	)	)	PUNCT
alces-35	36	31	,	,	PUNCT
alces-35	36	32	thus	thus	ADV
alces-35	36	33	,	,	PUNCT
alces-35	36	34	endangering	endanger	VERB
alces-35	36	35	the	the	DET
alces-35	36	36	spatio	spatio	NOUN
alces-35	36	37	-	-	PUNCT
alces-35	36	38	temporal	temporal	ADJ
alces-35	36	39	continuum	continuum	NOUN
alces-35	36	40	of	of	ADP
alces-35	36	41	aspen	aspen	ADJ
alces-35	36	42	occurrence	occurrence	NOUN
alces-35	36	43	.	.	PUNCT
alces-35	37	1	even	even	ADV
alces-35	37	2	in	in	ADP
alces-35	37	3	conservation	conservation	NOUN
alces-35	37	4	areas	area	NOUN
alces-35	37	5	this	this	PRON
alces-35	37	6	may	may	AUX
alces-35	37	7	result	result	VERB
alces-35	37	8	in	in	ADP
alces-35	37	9	a	a	DET
alces-35	37	10	situation	situation	NOUN
alces-35	37	11	where	where	SCONJ
alces-35	37	12	aspen	aspen	NOUN
alces-35	37	13	-	-	PUNCT
alces-35	37	14	associated	associate	VERB
alces-35	37	15	species	specie	NOUN
alces-35	37	16	may	may	AUX
alces-35	37	17	disappear	disappear	VERB
alces-35	37	18	both	both	CCONJ
alces-35	37	19	locally	locally	ADV
alces-35	37	20	and	and	CCONJ
alces-35	37	21	regionally	regionally	ADV
alces-35	37	22	.	.	PUNCT
alces-35	38	1	the	the	DET
alces-35	38	2	aim	aim	NOUN
alces-35	38	3	of	of	ADP
alces-35	38	4	this	this	DET
alces-35	38	5	study	study	NOUN
alces-35	38	6	was	be	AUX
alces-35	38	7	to	to	PART
alces-35	38	8	analyze	analyze	VERB
alces-35	38	9	the	the	DET
alces-35	38	10	influence	influence	NOUN
alces-35	38	11	of	of	ADP
alces-35	38	12	moose	moose	NOUN
alces-35	38	13	browsing	browse	VERB
alces-35	38	14	on	on	ADP
alces-35	38	15	the	the	DET
alces-35	38	16	regeneration	regeneration	NOUN
alces-35	38	17	of	of	ADP
alces-35	38	18	aspen	aspen	NOUN
alces-35	38	19	in	in	ADP
alces-35	38	20	a	a	DET
alces-35	38	21	conservation	conservation	NOUN
alces-35	38	22	area	area	NOUN
alces-35	38	23	.	.	PUNCT
alces-35	39	1	specifically	specifically	ADV
alces-35	39	2	,	,	PUNCT
alces-35	39	3	we	we	PRON
alces-35	39	4	assessed	assess	VERB
alces-35	39	5	past	past	NOUN
alces-35	39	6	and	and	CCONJ
alces-35	39	7	present	present	ADJ
alces-35	39	8	moose	moose	NOUN
alces-35	39	9	browsing	browse	VERB
alces-35	39	10	pressure	pressure	NOUN
alces-35	39	11	on	on	ADP
alces-35	39	12	aspen	aspen	NOUN
alces-35	39	13	at	at	ADP
alces-35	39	14	the	the	DET
alces-35	39	15	landscape	landscape	NOUN
alces-35	39	16	and	and	CCONJ
alces-35	39	17	stand	stand	VERB
alces-35	39	18	levels	level	NOUN
alces-35	39	19	within	within	ADP
alces-35	39	20	koli	koli	PROPN
alces-35	39	21	national	national	PROPN
alces-35	39	22	park	park	PROPN
alces-35	39	23	in	in	ADP
alces-35	39	24	eastern	eastern	PROPN
alces-35	39	25	finland	finland	PROPN
alces-35	39	26	.	.	PUNCT
alces-35	40	1	the	the	DET
alces-35	40	2	results	result	NOUN
alces-35	40	3	are	be	AUX
alces-35	40	4	related	relate	VERB
alces-35	40	5	to	to	ADP
alces-35	40	6	the	the	DET
alces-35	40	7	regeneration	regeneration	NOUN
alces-35	40	8	success	success	NOUN
alces-35	40	9	of	of	ADP
alces-35	40	10	aspen	aspen	NOUN
alces-35	40	11	in	in	ADP
alces-35	40	12	forests	forest	NOUN
alces-35	40	13	under	under	ADP
alces-35	40	14	high	high	ADJ
alces-35	40	15	browsing	browse	VERB
alces-35	40	16	pressure	pressure	NOUN
alces-35	40	17	by	by	ADP
alces-35	40	18	moose	moose	NOUN
alces-35	40	19	.	.	PUNCT
alces-35	41	1	study	study	NOUN
alces-35	41	2	area	area	NOUN
alces-35	41	3	the	the	DET
alces-35	41	4	study	study	NOUN
alces-35	41	5	occurred	occur	VERB
alces-35	41	6	in	in	ADP
alces-35	41	7	the	the	DET
alces-35	41	8	koli	koli	PROPN
alces-35	41	9	national	national	PROPN
alces-35	41	10	park	park	PROPN
alces-35	41	11	in	in	ADP
alces-35	41	12	eastern	eastern	PROPN
alces-35	41	13	finland	finland	PROPN
alces-35	41	14	(	(	PUNCT
alces-35	41	15	fig	fig	NOUN
alces-35	41	16	.	.	PUNCT
alces-35	42	1	1	1	NUM
alces-35	42	2	)	)	PUNCT
alces-35	42	3	.	.	PUNCT
alces-35	43	1	the	the	DET
alces-35	43	2	central	central	PROPN
alces-35	43	3	hill	hill	NOUN
alces-35	43	4	sites	site	NOUN
alces-35	43	5	of	of	ADP
alces-35	43	6	koli	koli	PROPN
alces-35	43	7	became	become	VERB
alces-35	43	8	state	state	NOUN
alces-35	43	9	-	-	PUNCT
alces-35	43	10	owned	own	VERB
alces-35	43	11	in	in	ADP
alces-35	43	12	1907	1907	NUM
alces-35	43	13	,	,	PUNCT
alces-35	43	14	and	and	CCONJ
alces-35	43	15	from	from	ADP
alces-35	43	16	1924	1924	NUM
alces-35	43	17	-	-	SYM
alces-35	43	18	2007	2007	NUM
alces-35	43	19	the	the	DET
alces-35	43	20	area	area	NOUN
alces-35	43	21	was	be	AUX
alces-35	43	22	administered	administer	VERB
alces-35	43	23	by	by	ADP
alces-35	43	24	the	the	DET
alces-35	43	25	finnish	finnish	PROPN
alces-35	43	26	forest	forest	PROPN
alces-35	43	27	research	research	PROPN
alces-35	43	28	institute	institute	PROPN
alces-35	43	29	.	.	PUNCT
alces-35	44	1	koli	koli	PROPN
alces-35	44	2	national	national	PROPN
alces-35	44	3	park	park	PROPN
alces-35	44	4	was	be	AUX
alces-35	44	5	established	establish	VERB
alces-35	44	6	by	by	ADP
alces-35	44	7	law	law	NOUN
alces-35	44	8	in	in	ADP
alces-35	44	9	1991	1991	NUM
alces-35	44	10	and	and	CCONJ
alces-35	44	11	covered	cover	VERB
alces-35	44	12	a	a	DET
alces-35	44	13	total	total	ADJ
alces-35	44	14	area	area	NOUN
alces-35	44	15	of	of	ADP
alces-35	44	16	1,135	1,135	NUM
alces-35	44	17	ha	ha	INTJ
alces-35	44	18	;	;	PUNCT
alces-35	44	19	it	it	PRON
alces-35	44	20	has	have	AUX
alces-35	44	21	since	since	ADV
alces-35	44	22	been	be	AUX
alces-35	44	23	enlarged	enlarge	VERB
alces-35	44	24	to	to	ADP
alces-35	44	25	about	about	ADP
alces-35	44	26	3,000	3,000	NUM
alces-35	44	27	ha	ha	INTJ
alces-35	44	28	.	.	PUNCT
alces-35	45	1	as	as	ADP
alces-35	45	2	of	of	ADP
alces-35	45	3	2008	2008	NUM
alces-35	45	4	,	,	PUNCT
alces-35	45	5	the	the	DET
alces-35	45	6	finnish	finnish	ADJ
alces-35	45	7	forest	forest	NOUN
alces-35	45	8	and	and	CCONJ
alces-35	45	9	park	park	NOUN
alces-35	45	10	service	service	NOUN
alces-35	45	11	is	be	AUX
alces-35	45	12	responsible	responsible	ADJ
alces-35	45	13	for	for	ADP
alces-35	45	14	administration	administration	NOUN
alces-35	45	15	of	of	ADP
alces-35	45	16	koli	koli	PROPN
alces-35	45	17	national	national	PROPN
alces-35	45	18	park	park	PROPN
alces-35	45	19	.	.	PUNCT
alces-35	46	1	the	the	DET
alces-35	46	2	purpose	purpose	NOUN
alces-35	46	3	of	of	ADP
alces-35	46	4	conserving	conserve	VERB
alces-35	46	5	the	the	DET
alces-35	46	6	area	area	NOUN
alces-35	46	7	by	by	ADP
alces-35	46	8	law	law	NOUN
alces-35	46	9	was	be	AUX
alces-35	46	10	to	to	PART
alces-35	46	11	ensure	ensure	VERB
alces-35	46	12	the	the	DET
alces-35	46	13	preservation	preservation	NOUN
alces-35	46	14	of	of	ADP
alces-35	46	15	koli	koli	PROPN
alces-35	46	16	’s	’s	PART
alces-35	46	17	heritage	heritage	PROPN
alces-35	46	18	landscape	landscape	NOUN
alces-35	46	19	and	and	CCONJ
alces-35	46	20	the	the	DET
alces-35	46	21	old	old	ADJ
alces-35	46	22	-	-	PUNCT
alces-35	46	23	growth	growth	NOUN
alces-35	46	24	forests	forest	NOUN
alces-35	46	25	of	of	ADP
alces-35	46	26	the	the	DET
alces-35	46	27	koli	koli	PROPN
alces-35	46	28	highlands	highlands	PROPN
alces-35	46	29	,	,	PUNCT
alces-35	46	30	as	as	ADV
alces-35	46	31	well	well	ADV
alces-35	46	32	as	as	ADP
alces-35	46	33	maintain	maintain	VERB
alces-35	46	34	the	the	DET
alces-35	46	35	plant	plant	NOUN
alces-35	46	36	communities	community	NOUN
alces-35	46	37	created	create	VERB
alces-35	46	38	in	in	ADP
alces-35	46	39	the	the	DET
alces-35	46	40	past	past	NOUN
alces-35	46	41	by	by	ADP
alces-35	46	42	swidden	swidden	ADJ
alces-35	46	43	cultivation	cultivation	NOUN
alces-35	46	44	(	(	PUNCT
alces-35	46	45	i.e.	i.e.	X
alces-35	46	46	,	,	PUNCT
alces-35	46	47	clearing	clearing	NOUN
alces-35	46	48	of	of	ADP
alces-35	46	49	land	land	NOUN
alces-35	46	50	for	for	ADP
alces-35	46	51	cultivation	cultivation	NOUN
alces-35	46	52	by	by	ADP
alces-35	46	53	slashing	slash	VERB
alces-35	46	54	and	and	CCONJ
alces-35	46	55	burning	burn	VERB
alces-35	46	56	the	the	DET
alces-35	46	57	forest	forest	NOUN
alces-35	46	58	vegetation	vegetation	NOUN
alces-35	46	59	cover	cover	NOUN
alces-35	46	60	)	)	PUNCT
alces-35	46	61	.	.	PUNCT
alces-35	47	1	other	other	ADJ
alces-35	47	2	aims	aim	NOUN
alces-35	47	3	were	be	AUX
alces-35	47	4	the	the	DET
alces-35	47	5	promotion	promotion	NOUN
alces-35	47	6	of	of	ADP
alces-35	47	7	environmental	environmental	ADJ
alces-35	47	8	research	research	NOUN
alces-35	47	9	,	,	PUNCT
alces-35	47	10	education	education	NOUN
alces-35	47	11	,	,	PUNCT
alces-35	47	12	and	and	CCONJ
alces-35	47	13	nature	nature	NOUN
alces-35	47	14	recreation	recreation	NOUN
alces-35	47	15	in	in	ADP
alces-35	47	16	the	the	DET
alces-35	47	17	area	area	NOUN
alces-35	47	18	.	.	PUNCT
alces-35	48	1	the	the	DET
alces-35	48	2	park	park	NOUN
alces-35	48	3	area	area	NOUN
alces-35	48	4	is	be	AUX
alces-35	48	5	characterized	characterize	VERB
alces-35	48	6	by	by	ADP
alces-35	48	7	a	a	DET
alces-35	48	8	highly	highly	ADV
alces-35	48	9	variable	variable	ADJ
alces-35	48	10	landscape	landscape	NOUN
alces-35	48	11	and	and	CCONJ
alces-35	48	12	topography	topography	NOUN
alces-35	48	13	where	where	SCONJ
alces-35	48	14	the	the	DET
alces-35	48	15	altitude	altitude	NOUN
alces-35	48	16	varies	vary	VERB
alces-35	48	17	from	from	ADP
alces-35	48	18	95	95	NUM
alces-35	48	19	-	-	SYM
alces-35	48	20	347	347	NUM
alces-35	48	21	m	m	NOUN
alces-35	48	22	above	above	ADP
alces-35	48	23	sea	sea	NOUN
alces-35	48	24	level	level	NOUN
alces-35	48	25	.	.	PUNCT
alces-35	49	1	the	the	DET
alces-35	49	2	soils	soil	NOUN
alces-35	49	3	are	be	AUX
alces-35	49	4	fertile	fertile	ADJ
alces-35	49	5	,	,	PUNCT
alces-35	49	6	especially	especially	ADV
alces-35	49	7	at	at	ADP
alces-35	49	8	lower	low	ADJ
alces-35	49	9	altitudes	altitude	NOUN
alces-35	49	10	,	,	PUNCT
alces-35	49	11	because	because	SCONJ
alces-35	49	12	the	the	DET
alces-35	49	13	crumbling	crumble	VERB
alces-35	49	14	substance	substance	NOUN
alces-35	49	15	originating	originate	VERB
alces-35	49	16	from	from	ADP
alces-35	49	17	the	the	DET
alces-35	49	18	calciferous	calciferous	ADJ
alces-35	49	19	bedrock	bedrock	NOUN
alces-35	49	20	(	(	PUNCT
alces-35	49	21	diabase	diabase	NOUN
alces-35	49	22	-	-	PUNCT
alces-35	49	23	rich	rich	ADJ
alces-35	49	24	or	or	CCONJ
alces-35	49	25	siliceous	siliceous	ADJ
alces-35	49	26	rocks	rock	NOUN
alces-35	49	27	,	,	PUNCT
alces-35	49	28	i.e.	i.e.	X
alces-35	49	29	,	,	PUNCT
alces-35	49	30	granitegneisses	granitegneisse	NOUN
alces-35	49	31	and	and	CCONJ
alces-35	49	32	quartzites	quartzite	NOUN
alces-35	49	33	;	;	PUNCT
alces-35	49	34	piirainen	piirainen	NOUN
alces-35	49	35	et	et	PROPN
alces-35	49	36	al	al	PROPN
alces-35	49	37	.	.	PROPN
alces-35	49	38	1974	1974	NUM
alces-35	49	39	)	)	PUNCT
alces-35	49	40	alces	alce	NOUN
alces-35	49	41	vol	vol	NOUN
alces-35	49	42	.	.	PROPN
alces-35	49	43	44	44	NUM
alces-35	49	44	,	,	PUNCT
alces-35	49	45	2008	2008	NUM
alces-35	49	46	härkönen	härkönen	NOUN
alces-35	49	47	et	et	PROPN
alces-35	49	48	al.moose	al.moose	X
alces-35	49	49	and	and	CCONJ
alces-35	49	50	aspen	aspen	VERB
alces-35	49	51	regeneration	regeneration	NOUN
alces-35	49	52	in	in	ADP
alces-35	49	53	finland	finland	PROPN
alces-35	49	54	33	33	NUM
alces-35	49	55	flow	flow	NOUN
alces-35	49	56	downhill	downhill	ADV
alces-35	49	57	with	with	ADP
alces-35	49	58	rainwater	rainwater	NOUN
alces-35	49	59	and	and	CCONJ
alces-35	49	60	streams	stream	NOUN
alces-35	49	61	of	of	ADP
alces-35	49	62	melting	melt	VERB
alces-35	49	63	snow	snow	NOUN
alces-35	49	64	.	.	PUNCT
alces-35	50	1	the	the	DET
alces-35	50	2	study	study	NOUN
alces-35	50	3	area	area	NOUN
alces-35	50	4	belongs	belong	VERB
alces-35	50	5	to	to	ADP
alces-35	50	6	the	the	DET
alces-35	50	7	border	border	NOUN
alces-35	50	8	area	area	NOUN
alces-35	50	9	between	between	ADP
alces-35	50	10	the	the	DET
alces-35	50	11	southern	southern	ADJ
alces-35	50	12	and	and	CCONJ
alces-35	50	13	middle	middle	ADJ
alces-35	50	14	boreal	boreal	NOUN
alces-35	50	15	forest	forest	NOUN
alces-35	50	16	vegetation	vegetation	NOUN
alces-35	50	17	zone	zone	NOUN
alces-35	50	18	(	(	PUNCT
alces-35	50	19	see	see	VERB
alces-35	50	20	kalliola	kalliola	PROPN
alces-35	50	21	1973	1973	NUM
alces-35	50	22	)	)	PUNCT
alces-35	50	23	(	(	PUNCT
alces-35	50	24	fig	fig	NOUN
alces-35	50	25	.	.	PUNCT
alces-35	51	1	1	1	NUM
alces-35	51	2	)	)	PUNCT
alces-35	51	3	.	.	PUNCT
alces-35	52	1	the	the	DET
alces-35	52	2	dominant	dominant	ADJ
alces-35	52	3	tree	tree	NOUN
alces-35	52	4	species	specie	NOUN
alces-35	52	5	are	be	AUX
alces-35	52	6	norway	norway	NOUN
alces-35	52	7	spruce	spruce	NOUN
alces-35	52	8	(	(	PUNCT
alces-35	52	9	picea	picea	NOUN
alces-35	52	10	abies	abie	NOUN
alces-35	52	11	)	)	PUNCT
alces-35	52	12	and	and	CCONJ
alces-35	52	13	scots	scot	NOUN
alces-35	52	14	pine	pine	NOUN
alces-35	52	15	(	(	PUNCT
alces-35	52	16	pinus	pinus	NOUN
alces-35	52	17	sylvestris	sylvestris	NOUN
alces-35	52	18	)	)	PUNCT
alces-35	52	19	.	.	PUNCT
alces-35	53	1	other	other	ADJ
alces-35	53	2	tree	tree	NOUN
alces-35	53	3	species	specie	NOUN
alces-35	53	4	such	such	ADJ
alces-35	53	5	as	as	ADP
alces-35	53	6	birches	birch	NOUN
alces-35	53	7	(	(	PUNCT
alces-35	53	8	betula	betula	PROPN
alces-35	53	9	pendula	pendula	PROPN
alces-35	53	10	and	and	CCONJ
alces-35	53	11	b.	b.	PROPN
alces-35	53	12	pubescens	pubescen	NOUN
alces-35	53	13	)	)	PUNCT
alces-35	53	14	,	,	PUNCT
alces-35	53	15	aspen	aspen	PROPN
alces-35	53	16	,	,	PUNCT
alces-35	53	17	rowan	rowan	PROPN
alces-35	53	18	(	(	PUNCT
alces-35	53	19	sorbus	sorbus	PROPN
alces-35	53	20	aucuparia	aucuparia	PROPN
alces-35	53	21	)	)	PUNCT
alces-35	53	22	,	,	PUNCT
alces-35	53	23	and	and	CCONJ
alces-35	53	24	alder	alder	NOUN
alces-35	53	25	(	(	PUNCT
alces-35	53	26	alnus	alnus	PROPN
alces-35	53	27	incana	incana	NOUN
alces-35	53	28	)	)	PUNCT
alces-35	53	29	occur	occur	VERB
alces-35	53	30	patchily	patchily	ADV
alces-35	53	31	within	within	ADP
alces-35	53	32	the	the	DET
alces-35	53	33	area	area	NOUN
alces-35	53	34	.	.	PUNCT
alces-35	54	1	the	the	DET
alces-35	54	2	forests	forest	NOUN
alces-35	54	3	outside	outside	ADP
alces-35	54	4	the	the	DET
alces-35	54	5	park	park	NOUN
alces-35	54	6	are	be	AUX
alces-35	54	7	intensively	intensively	ADV
alces-35	54	8	managed	manage	VERB
alces-35	54	9	and	and	CCONJ
alces-35	54	10	fragmented	fragment	VERB
alces-35	54	11	by	by	ADP
alces-35	54	12	lakes	lake	NOUN
alces-35	54	13	,	,	PUNCT
alces-35	54	14	mires	mire	NOUN
alces-35	54	15	,	,	PUNCT
alces-35	54	16	cultivated	cultivate	VERB
alces-35	54	17	land	land	NOUN
alces-35	54	18	,	,	PUNCT
alces-35	54	19	and	and	CCONJ
alces-35	54	20	small	small	ADJ
alces-35	54	21	villages	village	NOUN
alces-35	54	22	.	.	PUNCT
alces-35	55	1	hunting	hunting	NOUN
alces-35	55	2	of	of	ADP
alces-35	55	3	moose	moose	NOUN
alces-35	55	4	is	be	AUX
alces-35	55	5	strictly	strictly	ADV
alces-35	55	6	prohibited	prohibit	VERB
alces-35	55	7	in	in	ADP
alces-35	55	8	koli	koli	PROPN
alces-35	55	9	national	national	PROPN
alces-35	55	10	park	park	PROPN
alces-35	55	11	.	.	PUNCT
alces-35	56	1	the	the	DET
alces-35	56	2	northern	northern	PROPN
alces-35	56	3	karelia	karelia	PROPN
alces-35	56	4	game	game	PROPN
alces-35	56	5	management	management	PROPN
alces-35	56	6	district	district	NOUN
alces-35	56	7	estimated	estimate	VERB
alces-35	56	8	the	the	DET
alces-35	56	9	density	density	NOUN
alces-35	56	10	of	of	ADP
alces-35	56	11	moose	moose	NOUN
alces-35	56	12	outside	outside	ADP
alces-35	56	13	the	the	DET
alces-35	56	14	park	park	NOUN
alces-35	56	15	,	,	PUNCT
alces-35	56	16	after	after	ADP
alces-35	56	17	the	the	DET
alces-35	56	18	hunting	hunting	NOUN
alces-35	56	19	season	season	NOUN
alces-35	56	20	,	,	PUNCT
alces-35	56	21	as	as	ADP
alces-35	56	22	0.3	0.3	NUM
alces-35	56	23	-	-	SYM
alces-35	56	24	0.6	0.6	NUM
alces-35	56	25	moose	moose	NOUN
alces-35	56	26	/	/	SYM
alces-35	56	27	km2	km2	NOUN
alces-35	56	28	in	in	ADP
alces-35	56	29	2002	2002	NUM
alces-35	56	30	-	-	SYM
alces-35	56	31	2006	2006	NUM
alces-35	56	32	.	.	PUNCT
alces-35	57	1	local	local	ADJ
alces-35	57	2	hunters	hunter	NOUN
alces-35	57	3	estimated	estimate	VERB
alces-35	57	4	by	by	ADP
alces-35	57	5	snow	snow	NOUN
alces-35	57	6	tracking	tracking	NOUN
alces-35	57	7	that	that	SCONJ
alces-35	57	8	there	there	PRON
alces-35	57	9	were	be	VERB
alces-35	57	10	about	about	ADV
alces-35	57	11	0.75	0.75	NUM
alces-35	57	12	moose	moose	NOUN
alces-35	57	13	/	/	SYM
alces-35	57	14	km2	km2	NOUN
alces-35	57	15	within	within	ADP
alces-35	57	16	the	the	DET
alces-35	57	17	park	park	NOUN
alces-35	57	18	in	in	ADP
alces-35	57	19	november	november	PROPN
alces-35	57	20	2006	2006	NUM
alces-35	57	21	,	,	PUNCT
alces-35	57	22	and	and	CCONJ
alces-35	57	23	that	that	SCONJ
alces-35	57	24	this	this	DET
alces-35	57	25	density	density	NOUN
alces-35	57	26	remained	remain	VERB
alces-35	57	27	about	about	ADV
alces-35	57	28	the	the	DET
alces-35	57	29	same	same	ADJ
alces-35	57	30	in	in	ADP
alces-35	57	31	2002	2002	NUM
alces-35	57	32	-	-	SYM
alces-35	57	33	2006	2006	NUM
alces-35	57	34	.	.	PUNCT
alces-35	58	1	moose	moose	NOUN
alces-35	58	2	density	density	NOUN
alces-35	58	3	increases	increase	NOUN
alces-35	58	4	in	in	ADP
alces-35	58	5	the	the	DET
alces-35	58	6	park	park	NOUN
alces-35	58	7	during	during	ADP
alces-35	58	8	hunting	hunt	VERB
alces-35	58	9	season	season	NOUN
alces-35	58	10	(	(	PUNCT
alces-35	58	11	starting	start	VERB
alces-35	58	12	the	the	DET
alces-35	58	13	last	last	ADJ
alces-35	58	14	saturday	saturday	NOUN
alces-35	58	15	of	of	ADP
alces-35	58	16	september	september	PROPN
alces-35	58	17	)	)	PUNCT
alces-35	58	18	,	,	PUNCT
alces-35	58	19	when	when	SCONJ
alces-35	58	20	they	they	PRON
alces-35	58	21	escape	escape	VERB
alces-35	58	22	hunting	hunt	VERB
alces-35	58	23	pressure	pressure	NOUN
alces-35	58	24	from	from	ADP
alces-35	58	25	adjacent	adjacent	ADJ
alces-35	58	26	private	private	ADJ
alces-35	58	27	forestland	forestland	NOUN
alces-35	58	28	,	,	PUNCT
alces-35	58	29	until	until	ADP
alces-35	58	30	spring	spring	NOUN
alces-35	58	31	when	when	SCONJ
alces-35	58	32	they	they	PRON
alces-35	58	33	disperse	disperse	VERB
alces-35	58	34	back	back	ADV
alces-35	58	35	to	to	ADP
alces-35	58	36	their	their	PRON
alces-35	58	37	traditional	traditional	ADJ
alces-35	58	38	summer	summer	NOUN
alces-35	58	39	range	range	NOUN
alces-35	58	40	.	.	PUNCT
alces-35	59	1	methods	method	NOUN
alces-35	59	2	moose	moose	VERB
alces-35	59	3	browsing	browse	VERB
alces-35	59	4	on	on	ADP
alces-35	59	5	aspen	aspen	PROPN
alces-35	59	6	was	be	AUX
alces-35	59	7	investigated	investigate	VERB
alces-35	59	8	in	in	ADP
alces-35	59	9	summer	summer	NOUN
alces-35	59	10	2006	2006	NUM
alces-35	59	11	at	at	ADP
alces-35	59	12	2	2	NUM
alces-35	59	13	spatial	spatial	ADJ
alces-35	59	14	scales	scale	NOUN
alces-35	59	15	,	,	PUNCT
alces-35	59	16	the	the	DET
alces-35	59	17	landscape	landscape	NOUN
alces-35	59	18	level	level	NOUN
alces-35	59	19	and	and	CCONJ
alces-35	59	20	the	the	DET
alces-35	59	21	stand	stand	NOUN
alces-35	59	22	level	level	NOUN
alces-35	59	23	.	.	PUNCT
alces-35	60	1	the	the	DET
alces-35	60	2	park	park	NOUN
alces-35	60	3	was	be	AUX
alces-35	60	4	divided	divide	VERB
alces-35	60	5	into	into	ADP
alces-35	60	6	northern	northern	ADJ
alces-35	60	7	(	(	PUNCT
alces-35	60	8	no	no	NOUN
alces-35	60	9	)	)	PUNCT
alces-35	60	10	and	and	CCONJ
alces-35	60	11	southern	southern	ADJ
alces-35	60	12	(	(	PUNCT
alces-35	60	13	so	so	ADV
alces-35	60	14	)	)	PUNCT
alces-35	60	15	areas	area	NOUN
alces-35	60	16	at	at	ADP
alces-35	60	17	both	both	DET
alces-35	60	18	scales	scale	NOUN
alces-35	60	19	.	.	PUNCT
alces-35	61	1	this	this	PRON
alces-35	61	2	was	be	AUX
alces-35	61	3	reasonable	reasonable	ADJ
alces-35	61	4	because	because	SCONJ
alces-35	61	5	the	the	DET
alces-35	61	6	no	no	DET
alces-35	61	7	area	area	NOUN
alces-35	61	8	was	be	AUX
alces-35	61	9	characterized	characterize	VERB
alces-35	61	10	by	by	ADP
alces-35	61	11	representative	representative	ADJ
alces-35	61	12	old	old	ADJ
alces-35	61	13	-	-	PUNCT
alces-35	61	14	growth	growth	NOUN
alces-35	61	15	forests	forest	NOUN
alces-35	61	16	close	close	ADJ
alces-35	61	17	to	to	ADP
alces-35	61	18	the	the	DET
alces-35	61	19	natural	natural	ADJ
alces-35	61	20	stage	stage	NOUN
alces-35	61	21	,	,	PUNCT
alces-35	61	22	whereas	whereas	SCONJ
alces-35	61	23	,	,	PUNCT
alces-35	61	24	in	in	ADP
alces-35	61	25	the	the	DET
alces-35	61	26	so	so	ADJ
alces-35	61	27	area	area	NOUN
alces-35	61	28	signs	sign	NOUN
alces-35	61	29	of	of	ADP
alces-35	61	30	forestry	forestry	NOUN
alces-35	61	31	practices	practice	NOUN
alces-35	61	32	were	be	AUX
alces-35	61	33	still	still	ADV
alces-35	61	34	clearly	clearly	ADV
alces-35	61	35	visible	visible	ADJ
alces-35	61	36	both	both	PRON
alces-35	61	37	in	in	ADP
alces-35	61	38	the	the	DET
alces-35	61	39	landscape	landscape	NOUN
alces-35	61	40	and	and	CCONJ
alces-35	61	41	forest	forest	NOUN
alces-35	61	42	age	age	NOUN
alces-35	61	43	structure	structure	NOUN
alces-35	61	44	.	.	PUNCT
alces-35	62	1	at	at	ADP
alces-35	62	2	the	the	DET
alces-35	62	3	landscape	landscape	NOUN
alces-35	62	4	level	level	NOUN
alces-35	62	5	,	,	PUNCT
alces-35	62	6	data	datum	NOUN
alces-35	62	7	were	be	AUX
alces-35	62	8	collected	collect	VERB
alces-35	62	9	from	from	ADP
alces-35	62	10	systematically	systematically	ADV
alces-35	62	11	located	locate	VERB
alces-35	62	12	circular	circular	ADJ
alces-35	62	13	sample	sample	NOUN
alces-35	62	14	plots	plot	NOUN
alces-35	62	15	;	;	PUNCT
alces-35	62	16	132	132	NUM
alces-35	62	17	plots	plot	NOUN
alces-35	62	18	were	be	AUX
alces-35	62	19	located	locate	VERB
alces-35	62	20	in	in	ADP
alces-35	62	21	the	the	DET
alces-35	62	22	no	no	DET
alces-35	62	23	area	area	NOUN
alces-35	62	24	and	and	CCONJ
alces-35	62	25	141	141	NUM
alces-35	62	26	plots	plot	NOUN
alces-35	62	27	in	in	ADP
alces-35	62	28	the	the	DET
alces-35	62	29	so	so	ADJ
alces-35	62	30	area	area	NOUN
alces-35	62	31	.	.	PUNCT
alces-35	63	1	the	the	DET
alces-35	63	2	plot	plot	NOUN
alces-35	63	3	size	size	NOUN
alces-35	63	4	was	be	AUX
alces-35	63	5	113	113	NUM
alces-35	63	6	m2	m2	NOUN
alces-35	63	7	,	,	PUNCT
alces-35	63	8	and	and	CCONJ
alces-35	63	9	plots	plot	NOUN
alces-35	63	10	were	be	AUX
alces-35	63	11	placed	place	VERB
alces-35	63	12	on	on	ADP
alces-35	63	13	a	a	DET
alces-35	63	14	systematic	systematic	ADJ
alces-35	63	15	300	300	NUM
alces-35	63	16	m	m	NOUN
alces-35	63	17	x	x	PUNCT
alces-35	63	18	300	300	NUM
alces-35	63	19	m	m	NOUN
alces-35	63	20	grid	grid	ADJ
alces-35	63	21	.	.	PUNCT
alces-35	64	1	in	in	ADP
alces-35	64	2	addition	addition	NOUN
alces-35	64	3	,	,	PUNCT
alces-35	64	4	the	the	DET
alces-35	64	5	stand	stand	NOUN
alces-35	64	6	development	development	NOUN
alces-35	64	7	class	class	NOUN
alces-35	64	8	was	be	AUX
alces-35	64	9	determined	determine	VERB
alces-35	64	10	at	at	ADP
alces-35	64	11	each	each	DET
alces-35	64	12	plot	plot	NOUN
alces-35	64	13	.	.	PUNCT
alces-35	65	1	aspens	aspen	NOUN
alces-35	65	2	were	be	AUX
alces-35	65	3	also	also	ADV
alces-35	65	4	pooled	pool	VERB
alces-35	65	5	into	into	ADP
alces-35	65	6	three	three	NUM
alces-35	65	7	height	height	NOUN
alces-35	65	8	categories	category	NOUN
alces-35	65	9	(	(	PUNCT
alces-35	65	10	i	i	NOUN
alces-35	65	11	:	:	PUNCT
alces-35	65	12	<	<	X
alces-35	65	13	5	5	NUM
alces-35	65	14	m	m	PROPN
alces-35	65	15	,	,	PUNCT
alces-35	65	16	ii	ii	PROPN
alces-35	65	17	:	:	PUNCT
alces-35	65	18	5	5	NUM
alces-35	65	19	-	-	SYM
alces-35	65	20	15	15	NUM
alces-35	65	21	m	m	NOUN
alces-35	65	22	,	,	PUNCT
alces-35	65	23	and	and	CCONJ
alces-35	65	24	iii	iii	X
alces-35	65	25	:	:	PUNCT
alces-35	65	26	>	>	X
alces-35	65	27	15	15	NUM
alces-35	65	28	m	m	NOUN
alces-35	65	29	)	)	PUNCT
alces-35	65	30	.	.	PUNCT
alces-35	66	1	at	at	ADP
alces-35	66	2	the	the	DET
alces-35	66	3	stand	stand	NOUN
alces-35	66	4	level	level	NOUN
alces-35	66	5	,	,	PUNCT
alces-35	66	6	all	all	DET
alces-35	66	7	potential	potential	NOUN
alces-35	66	8	stands	stand	VERB
alces-35	66	9	where	where	SCONJ
alces-35	66	10	aspen	aspen	PROPN
alces-35	66	11	was	be	AUX
alces-35	66	12	dominant	dominant	ADJ
alces-35	66	13	or	or	CCONJ
alces-35	66	14	was	be	AUX
alces-35	66	15	important	important	ADJ
alces-35	66	16	in	in	ADP
alces-35	66	17	the	the	DET
alces-35	66	18	mixture	mixture	NOUN
alces-35	66	19	of	of	ADP
alces-35	66	20	tree	tree	NOUN
alces-35	66	21	species	specie	NOUN
alces-35	66	22	were	be	AUX
alces-35	66	23	identified	identify	VERB
alces-35	66	24	from	from	ADP
alces-35	66	25	aerial	aerial	ADJ
alces-35	66	26	photographs	photograph	NOUN
alces-35	66	27	.	.	PUNCT
alces-35	67	1	when	when	SCONJ
alces-35	67	2	verified	verify	VERB
alces-35	67	3	in	in	ADP
alces-35	67	4	the	the	DET
alces-35	67	5	field	field	NOUN
alces-35	67	6	,	,	PUNCT
alces-35	67	7	the	the	DET
alces-35	67	8	stand	stand	NOUN
alces-35	67	9	was	be	AUX
alces-35	67	10	included	include	VERB
alces-35	67	11	in	in	ADP
alces-35	67	12	the	the	DET
alces-35	67	13	sampling	sample	VERB
alces-35	67	14	procedure	procedure	NOUN
alces-35	67	15	,	,	PUNCT
alces-35	67	16	after	after	ADP
alces-35	67	17	which	which	PRON
alces-35	67	18	5	5	NUM
alces-35	67	19	young	young	ADJ
alces-35	67	20	(	(	PUNCT
alces-35	67	21	<	<	X
alces-35	67	22	5	5	NUM
alces-35	67	23	m	m	NOUN
alces-35	67	24	in	in	ADP
alces-35	67	25	height	height	NOUN
alces-35	67	26	)	)	PUNCT
alces-35	67	27	,	,	PUNCT
alces-35	67	28	5	5	NUM
alces-35	67	29	middle	middle	NOUN
alces-35	67	30	-	-	PUNCT
alces-35	67	31	aged	aged	ADJ
alces-35	67	32	(	(	PUNCT
alces-35	67	33	5	5	NUM
alces-35	67	34	-	-	SYM
alces-35	67	35	15	15	NUM
alces-35	67	36	m	m	NOUN
alces-35	67	37	in	in	ADP
alces-35	67	38	height	height	NOUN
alces-35	67	39	)	)	PUNCT
alces-35	67	40	,	,	PUNCT
alces-35	67	41	and	and	CCONJ
alces-35	67	42	5	5	NUM
alces-35	67	43	old	old	ADJ
alces-35	67	44	(	(	PUNCT
alces-35	67	45	>	>	X
alces-35	67	46	15	15	NUM
alces-35	67	47	m	m	NOUN
alces-35	67	48	in	in	ADP
alces-35	67	49	height	height	NOUN
alces-35	67	50	)	)	PUNCT
alces-35	67	51	aspen	aspen	PROPN
alces-35	67	52	-	-	PUNCT
alces-35	67	53	rich	rich	ADJ
alces-35	67	54	stands	stand	NOUN
alces-35	67	55	were	be	AUX
alces-35	67	56	randomly	randomly	ADV
alces-35	67	57	selected	select	VERB
alces-35	67	58	both	both	PRON
alces-35	67	59	in	in	ADP
alces-35	67	60	the	the	DET
alces-35	67	61	no	no	NOUN
alces-35	67	62	and	and	CCONJ
alces-35	67	63	so	so	ADV
alces-35	67	64	areas	area	NOUN
alces-35	67	65	to	to	PART
alces-35	67	66	ensure	ensure	VERB
alces-35	67	67	sufficient	sufficient	ADJ
alces-35	67	68	variation	variation	NOUN
alces-35	67	69	of	of	ADP
alces-35	67	70	stand	stand	NOUN
alces-35	67	71	structures	structure	NOUN
alces-35	67	72	in	in	ADP
alces-35	67	73	the	the	DET
alces-35	67	74	data	datum	NOUN
alces-35	67	75	(	(	PUNCT
alces-35	67	76	ericsson	ericsson	PROPN
alces-35	67	77	et	et	PROPN
alces-35	67	78	al	al	PROPN
alces-35	67	79	.	.	PROPN
alces-35	67	80	2001	2001	NUM
alces-35	67	81	)	)	PUNCT
alces-35	67	82	.	.	PUNCT
alces-35	68	1	three	three	NUM
alces-35	68	2	circular	circular	ADJ
alces-35	68	3	plots	plot	NOUN
alces-35	68	4	of	of	ADP
alces-35	68	5	113	113	NUM
alces-35	68	6	m2	m2	NOUN
alces-35	68	7	were	be	AUX
alces-35	68	8	systematically	systematically	ADV
alces-35	68	9	located	locate	VERB
alces-35	68	10	within	within	ADP
alces-35	68	11	each	each	DET
alces-35	68	12	sampled	sample	VERB
alces-35	68	13	stand	stand	NOUN
alces-35	68	14	,	,	PUNCT
alces-35	68	15	with	with	ADP
alces-35	68	16	the	the	DET
alces-35	68	17	first	first	ADJ
alces-35	68	18	plot	plot	NOUN
alces-35	68	19	placed	place	VERB
alces-35	68	20	in	in	ADP
alces-35	68	21	the	the	DET
alces-35	68	22	center	center	NOUN
alces-35	68	23	of	of	ADP
alces-35	68	24	the	the	DET
alces-35	68	25	stand	stand	NOUN
alces-35	68	26	,	,	PUNCT
alces-35	68	27	fig	fig	NOUN
alces-35	68	28	.	.	PUNCT
alces-35	69	1	1	1	X
alces-35	69	2	.	.	X
alces-35	69	3	location	location	NOUN
alces-35	69	4	of	of	ADP
alces-35	69	5	the	the	DET
alces-35	69	6	koli	koli	PROPN
alces-35	69	7	national	national	PROPN
alces-35	69	8	park	park	PROPN
alces-35	69	9	in	in	ADP
alces-35	69	10	eastern	eastern	PROPN
alces-35	69	11	finland	finland	PROPN
alces-35	69	12	.	.	PUNCT
alces-35	70	1	forest	forest	NOUN
alces-35	70	2	vegetation	vegetation	NOUN
alces-35	70	3	zones	zone	NOUN
alces-35	70	4	after	after	ADP
alces-35	70	5	kalela	kalela	PROPN
alces-35	70	6	(	(	PUNCT
alces-35	70	7	1970	1970	NUM
alces-35	70	8	in	in	ADP
alces-35	70	9	kalliola	kalliola	PROPN
alces-35	70	10	1973	1973	NUM
alces-35	70	11	):	):	PUNCT
alces-35	70	12	south	south	PROPN
alces-35	70	13	finland	finland	PROPN
alces-35	70	14	(	(	PUNCT
alces-35	70	15	1=	1=	X
alces-35	70	16	hemiboreal	hemiboreal	NOUN
alces-35	70	17	and	and	CCONJ
alces-35	70	18	2	2	NUM
alces-35	70	19	=	=	ADJ
alces-35	70	20	southern	southern	ADJ
alces-35	70	21	boreal	boreal	NOUN
alces-35	70	22	)	)	PUNCT
alces-35	70	23	,	,	PUNCT
alces-35	70	24	pohjanmaakainuu	pohjanmaakainuu	PROPN
alces-35	70	25	(	(	PUNCT
alces-35	70	26	3	3	NUM
alces-35	70	27	=	=	SYM
alces-35	70	28	middle	middle	ADJ
alces-35	70	29	boreal	boreal	NOUN
alces-35	70	30	)	)	PUNCT
alces-35	70	31	,	,	PUNCT
alces-35	70	32	and	and	CCONJ
alces-35	70	33	peräpohjola	peräpohjola	PROPN
alces-35	70	34	and	and	CCONJ
alces-35	70	35	metsä	metsä	NOUN
alces-35	70	36	-	-	PUNCT
alces-35	70	37	lappi	lappi	PROPN
alces-35	70	38	(	(	PUNCT
alces-35	70	39	4	4	NUM
alces-35	70	40	=	=	SYM
alces-35	70	41	northern	northern	ADJ
alces-35	70	42	boreal	boreal	NOUN
alces-35	70	43	)	)	PUNCT
alces-35	70	44	.	.	PUNCT
alces-35	71	1	moose	moose	NOUN
alces-35	71	2	and	and	CCONJ
alces-35	71	3	aspen	aspen	NOUN
alces-35	71	4	regeneration	regeneration	NOUN
alces-35	71	5	in	in	ADP
alces-35	71	6	finland	finland	PROPN
alces-35	71	7	härkönen	härkönen	PROPN
alces-35	71	8	et	et	PROPN
alces-35	71	9	al	al	PROPN
alces-35	71	10	.	.	PROPN
alces-35	72	1	alces	alce	NOUN
alces-35	72	2	vol	vol	NOUN
alces-35	72	3	.	.	PROPN
alces-35	72	4	44	44	NUM
alces-35	72	5	,	,	PUNCT
alces-35	72	6	2008	2008	NUM
alces-35	72	7	34	34	NUM
alces-35	72	8	and	and	CCONJ
alces-35	72	9	the	the	DET
alces-35	72	10	other	other	ADJ
alces-35	72	11	2	2	NUM
alces-35	72	12	placed	place	VERB
alces-35	72	13	20	20	NUM
alces-35	72	14	m	m	NOUN
alces-35	72	15	north	north	NOUN
alces-35	72	16	,	,	PUNCT
alces-35	72	17	south	south	NOUN
alces-35	72	18	,	,	PUNCT
alces-35	72	19	west	west	NOUN
alces-35	72	20	,	,	PUNCT
alces-35	72	21	or	or	CCONJ
alces-35	72	22	east	east	NOUN
alces-35	72	23	from	from	ADP
alces-35	72	24	the	the	DET
alces-35	72	25	center	center	NOUN
alces-35	72	26	plot	plot	NOUN
alces-35	72	27	depending	depend	VERB
alces-35	72	28	on	on	ADP
alces-35	72	29	the	the	DET
alces-35	72	30	shape	shape	NOUN
alces-35	72	31	of	of	ADP
alces-35	72	32	the	the	DET
alces-35	72	33	stand	stand	NOUN
alces-35	72	34	.	.	PUNCT
alces-35	73	1	the	the	DET
alces-35	73	2	condition	condition	NOUN
alces-35	73	3	of	of	ADP
alces-35	73	4	each	each	DET
alces-35	73	5	aspen	aspen	NOUN
alces-35	73	6	was	be	AUX
alces-35	73	7	classified	classify	VERB
alces-35	73	8	as	as	ADP
alces-35	73	9	undamaged	undamaged	ADJ
alces-35	73	10	or	or	CCONJ
alces-35	73	11	damaged	damage	VERB
alces-35	73	12	,	,	PUNCT
alces-35	73	13	and	and	CCONJ
alces-35	73	14	alive	alive	ADJ
alces-35	73	15	or	or	CCONJ
alces-35	73	16	dead	dead	ADJ
alces-35	73	17	.	.	PUNCT
alces-35	74	1	each	each	DET
alces-35	74	2	measured	measure	VERB
alces-35	74	3	aspen	aspen	NOUN
alces-35	74	4	was	be	AUX
alces-35	74	5	investigated	investigate	VERB
alces-35	74	6	for	for	ADP
alces-35	74	7	signs	sign	NOUN
alces-35	74	8	of	of	ADP
alces-35	74	9	fresh	fresh	ADJ
alces-35	74	10	and	and	CCONJ
alces-35	74	11	older	old	ADJ
alces-35	74	12	moose	moose	NOUN
alces-35	74	13	browsing	browse	VERB
alces-35	74	14	.	.	PUNCT
alces-35	75	1	fresh	fresh	ADJ
alces-35	75	2	browsing	browsing	NOUN
alces-35	75	3	meant	mean	VERB
alces-35	75	4	that	that	SCONJ
alces-35	75	5	browsing	browse	VERB
alces-35	75	6	had	have	AUX
alces-35	75	7	occurred	occur	VERB
alces-35	75	8	during	during	ADP
alces-35	75	9	winter	winter	NOUN
alces-35	75	10	2005	2005	NUM
alces-35	75	11	-	-	SYM
alces-35	75	12	2006	2006	NUM
alces-35	75	13	,	,	PUNCT
alces-35	75	14	and	and	CCONJ
alces-35	75	15	was	be	AUX
alces-35	75	16	distinguished	distinguish	VERB
alces-35	75	17	from	from	ADP
alces-35	75	18	older	old	ADJ
alces-35	75	19	browsing	browsing	NOUN
alces-35	75	20	by	by	ADP
alces-35	75	21	its	its	PRON
alces-35	75	22	white	white	ADJ
alces-35	75	23	color	color	NOUN
alces-35	75	24	at	at	ADP
alces-35	75	25	the	the	DET
alces-35	75	26	browsing	browse	VERB
alces-35	75	27	point	point	NOUN
alces-35	75	28	.	.	PUNCT
alces-35	76	1	moose	moose	NOUN
alces-35	76	2	browsing	browse	VERB
alces-35	76	3	was	be	AUX
alces-35	76	4	divided	divide	VERB
alces-35	76	5	into	into	ADP
alces-35	76	6	3	3	NUM
alces-35	76	7	damage	damage	NOUN
alces-35	76	8	categories	category	NOUN
alces-35	76	9	:	:	PUNCT
alces-35	77	1	twigbrowsing	twigbrowse	VERB
alces-35	77	2	,	,	PUNCT
alces-35	77	3	stem	stem	NOUN
alces-35	77	4	breakage	breakage	NOUN
alces-35	77	5	,	,	PUNCT
alces-35	77	6	and	and	CCONJ
alces-35	77	7	bark	bark	NOUN
alces-35	77	8	stripping	strip	VERB
alces-35	77	9	.	.	PUNCT
alces-35	78	1	the	the	DET
alces-35	78	2	total	total	ADJ
alces-35	78	3	tree	tree	NOUN
alces-35	78	4	height	height	NOUN
alces-35	78	5	was	be	AUX
alces-35	78	6	measured	measure	VERB
alces-35	78	7	of	of	ADP
alces-35	78	8	all	all	DET
alces-35	78	9	small	small	ADJ
alces-35	78	10	aspens	aspen	NOUN
alces-35	78	11	(	(	PUNCT
alces-35	78	12	i.e.	i.e.	X
alces-35	78	13	,	,	PUNCT
alces-35	78	14	height	height	NOUN
alces-35	78	15	0.5	0.5	NUM
alces-35	78	16	-	-	SYM
alces-35	78	17	4.0	4.0	NUM
alces-35	78	18	m	m	NOUN
alces-35	78	19	)	)	PUNCT
alces-35	78	20	.	.	PUNCT
alces-35	79	1	for	for	ADP
alces-35	79	2	aspens	aspen	NOUN
alces-35	79	3	taller	tall	ADJ
alces-35	79	4	than	than	ADP
alces-35	79	5	4	4	NUM
alces-35	79	6	m	m	NOUN
alces-35	79	7	,	,	PUNCT
alces-35	79	8	the	the	DET
alces-35	79	9	height	height	NOUN
alces-35	79	10	was	be	AUX
alces-35	79	11	estimated	estimate	VERB
alces-35	79	12	to	to	ADP
alces-35	79	13	the	the	DET
alces-35	79	14	nearest	near	ADJ
alces-35	79	15	0.5	0.5	NUM
alces-35	79	16	-	-	PUNCT
alces-35	79	17	m	m	NOUN
alces-35	79	18	class	class	NOUN
alces-35	79	19	.	.	PUNCT
alces-35	80	1	in	in	ADP
alces-35	80	2	order	order	NOUN
alces-35	80	3	to	to	PART
alces-35	80	4	estimate	estimate	VERB
alces-35	80	5	the	the	DET
alces-35	80	6	moose	moose	NOUN
alces-35	80	7	activity	activity	NOUN
alces-35	80	8	in	in	ADP
alces-35	80	9	the	the	DET
alces-35	80	10	area	area	NOUN
alces-35	80	11	,	,	PUNCT
alces-35	80	12	the	the	DET
alces-35	80	13	number	number	NOUN
alces-35	80	14	of	of	ADP
alces-35	80	15	fecal	fecal	ADJ
alces-35	80	16	pellet	pellet	NOUN
alces-35	80	17	groups	group	NOUN
alces-35	80	18	(	(	PUNCT
alces-35	80	19	1	1	NUM
alces-35	80	20	group	group	NOUN
alces-35	80	21	≥20	≥20	NOUN
alces-35	80	22	pellets	pellet	NOUN
alces-35	80	23	)	)	PUNCT
alces-35	80	24	was	be	AUX
alces-35	80	25	counted	count	VERB
alces-35	80	26	in	in	ADP
alces-35	80	27	each	each	DET
alces-35	80	28	plot	plot	NOUN
alces-35	80	29	.	.	PUNCT
alces-35	81	1	only	only	ADV
alces-35	81	2	pellet	pellet	NOUN
alces-35	81	3	groups	group	NOUN
alces-35	81	4	deposited	deposit	VERB
alces-35	81	5	during	during	ADP
alces-35	81	6	the	the	DET
alces-35	81	7	winter	winter	NOUN
alces-35	81	8	of	of	ADP
alces-35	81	9	2005	2005	NUM
alces-35	81	10	-	-	SYM
alces-35	81	11	2006	2006	NUM
alces-35	81	12	(	(	PUNCT
alces-35	81	13	i.e.	i.e.	X
alces-35	81	14	,	,	PUNCT
alces-35	81	15	those	those	PRON
alces-35	81	16	on	on	ADP
alces-35	81	17	top	top	NOUN
alces-35	81	18	of	of	ADP
alces-35	81	19	the	the	DET
alces-35	81	20	previous	previous	ADJ
alces-35	81	21	year	year	NOUN
alces-35	81	22	’s	’s	PART
alces-35	81	23	leaf	leaf	NOUN
alces-35	81	24	litter	litter	NOUN
alces-35	81	25	)	)	PUNCT
alces-35	81	26	were	be	AUX
alces-35	81	27	counted	count	VERB
alces-35	81	28	.	.	PUNCT
alces-35	82	1	all	all	DET
alces-35	82	2	statistical	statistical	ADJ
alces-35	82	3	analyses	analysis	NOUN
alces-35	82	4	were	be	AUX
alces-35	82	5	performed	perform	VERB
alces-35	82	6	with	with	ADP
alces-35	82	7	spss	spss	PROPN
alces-35	82	8	package	package	NOUN
alces-35	82	9	.	.	PUNCT
alces-35	83	1	nonparametric	nonparametric	NOUN
alces-35	83	2	tests	test	NOUN
alces-35	83	3	were	be	AUX
alces-35	83	4	employed	employ	VERB
alces-35	83	5	because	because	SCONJ
alces-35	83	6	none	none	NOUN
alces-35	83	7	of	of	ADP
alces-35	83	8	the	the	DET
alces-35	83	9	variables	variable	NOUN
alces-35	83	10	had	have	VERB
alces-35	83	11	normal	normal	ADJ
alces-35	83	12	distributions	distribution	NOUN
alces-35	83	13	.	.	PUNCT
alces-35	84	1	results	result	VERB
alces-35	84	2	landscape	landscape	NOUN
alces-35	84	3	level	level	VERB
alces-35	84	4	the	the	DET
alces-35	84	5	density	density	NOUN
alces-35	84	6	of	of	ADP
alces-35	84	7	aspen	aspen	NOUN
alces-35	84	8	was	be	AUX
alces-35	84	9	three	three	NUM
alces-35	84	10	-	-	ADJ
alces-35	84	11	fold	fold	VERB
alces-35	84	12	higher	high	ADJ
alces-35	84	13	in	in	ADP
alces-35	84	14	the	the	DET
alces-35	84	15	so	so	ADJ
alces-35	84	16	area	area	NOUN
alces-35	84	17	(	(	PUNCT
alces-35	84	18	152	152	NUM
alces-35	84	19	aspens	aspen	NOUN
alces-35	84	20	/	/	SYM
alces-35	84	21	ha	ha	NOUN
alces-35	84	22	±	±	NUM
alces-35	84	23	47	47	NUM
alces-35	84	24	se	se	PROPN
alces-35	84	25	)	)	PUNCT
alces-35	84	26	than	than	ADP
alces-35	84	27	in	in	ADP
alces-35	84	28	the	the	DET
alces-35	84	29	no	no	DET
alces-35	84	30	area	area	NOUN
alces-35	84	31	(	(	PUNCT
alces-35	84	32	48	48	NUM
alces-35	84	33	aspens	aspen	NOUN
alces-35	84	34	/	/	SYM
alces-35	84	35	ha	ha	PROPN
alces-35	84	36	±	±	NUM
alces-35	84	37	12	12	NUM
alces-35	84	38	se	se	ADJ
alces-35	84	39	)	)	PUNCT
alces-35	84	40	,	,	PUNCT
alces-35	84	41	but	but	CCONJ
alces-35	84	42	the	the	DET
alces-35	84	43	difference	difference	NOUN
alces-35	84	44	was	be	AUX
alces-35	84	45	not	not	PART
alces-35	84	46	significant	significant	ADJ
alces-35	84	47	(	(	PUNCT
alces-35	84	48	mannwhitney	mannwhitney	NOUN
alces-35	84	49	u	u	NOUN
alces-35	84	50	=	=	PROPN
alces-35	84	51	8759.5	8759.5	NUM
alces-35	84	52	,	,	PUNCT
alces-35	84	53	p	p	NOUN
alces-35	84	54	=	=	NOUN
alces-35	84	55	0.24	0.24	NUM
alces-35	84	56	)	)	PUNCT
alces-35	84	57	.	.	PUNCT
alces-35	85	1	the	the	DET
alces-35	85	2	number	number	NOUN
alces-35	85	3	of	of	ADP
alces-35	85	4	fecal	fecal	ADJ
alces-35	85	5	pellet	pellet	NOUN
alces-35	85	6	groups	group	NOUN
alces-35	85	7	indicated	indicate	VERB
alces-35	85	8	that	that	SCONJ
alces-35	85	9	moose	moose	NOUN
alces-35	85	10	used	use	VERB
alces-35	85	11	the	the	DET
alces-35	85	12	so	so	ADJ
alces-35	85	13	area	area	NOUN
alces-35	85	14	(	(	PUNCT
alces-35	85	15	32	32	NUM
alces-35	85	16	pellet	pellet	NOUN
alces-35	85	17	group	group	NOUN
alces-35	85	18	/	/	SYM
alces-35	85	19	ha	ha	X
alces-35	85	20	±	±	NUM
alces-35	85	21	6	6	NUM
alces-35	85	22	se	se	ADV
alces-35	85	23	)	)	PUNCT
alces-35	85	24	more	more	ADJ
alces-35	85	25	than	than	ADP
alces-35	85	26	the	the	DET
alces-35	85	27	no	no	DET
alces-35	85	28	area	area	NOUN
alces-35	85	29	(	(	PUNCT
alces-35	85	30	2	2	NUM
alces-35	85	31	pellet	pellet	NOUN
alces-35	85	32	groups	group	NOUN
alces-35	85	33	/	/	SYM
alces-35	85	34	ha	ha	X
alces-35	85	35	±	±	NUM
alces-35	85	36	1.5	1.5	NUM
alces-35	85	37	se	se	X
alces-35	85	38	)	)	PUNCT
alces-35	85	39	(	(	PUNCT
alces-35	85	40	mann	mann	PROPN
alces-35	85	41	-	-	PUNCT
alces-35	85	42	whitney	whitney	PROPN
alces-35	85	43	u	u	NOUN
alces-35	85	44	=	=	PROPN
alces-35	85	45	7401.0	7401.0	NUM
alces-35	85	46	,	,	PUNCT
alces-35	85	47	p	p	X
alces-35	85	48	=	=	NOUN
alces-35	85	49	0.000	0.000	NUM
alces-35	85	50	)	)	PUNCT
alces-35	85	51	.	.	PUNCT
alces-35	86	1	there	there	PRON
alces-35	86	2	was	be	VERB
alces-35	86	3	no	no	DET
alces-35	86	4	difference	difference	NOUN
alces-35	86	5	in	in	ADP
alces-35	86	6	mean	mean	PROPN
alces-35	86	7	aspen	aspen	ADJ
alces-35	86	8	height	height	NOUN
alces-35	86	9	between	between	ADP
alces-35	86	10	the	the	DET
alces-35	86	11	two	two	NUM
alces-35	86	12	areas	area	NOUN
alces-35	86	13	(	(	PUNCT
alces-35	86	14	so	so	ADV
alces-35	86	15	:	:	PUNCT
alces-35	86	16	9.7	9.7	NUM
alces-35	86	17	m	m	NOUN
alces-35	86	18	±	±	NUM
alces-35	86	19	1.4	1.4	NUM
alces-35	86	20	se	se	X
alces-35	86	21	vs.	vs.	ADP
alces-35	86	22	no	no	NOUN
alces-35	86	23	:	:	PUNCT
alces-35	86	24	10.0	10.0	NUM
alces-35	86	25	m	m	NOUN
alces-35	86	26	±	±	NUM
alces-35	86	27	2.4	2.4	NUM
alces-35	86	28	se	se	X
alces-35	86	29	,	,	PUNCT
alces-35	86	30	mann	mann	PROPN
alces-35	86	31	-	-	PUNCT
alces-35	86	32	whitney	whitney	PROPN
alces-35	86	33	u	u	NOUN
alces-35	86	34	=	=	PROPN
alces-35	86	35	267.5	267.5	NUM
alces-35	86	36	,	,	PUNCT
alces-35	86	37	p	p	NOUN
alces-35	86	38	=	=	NOUN
alces-35	86	39	0.36	0.36	NUM
alces-35	86	40	)	)	PUNCT
alces-35	86	41	.	.	PUNCT
alces-35	87	1	moose	moose	PROPN
alces-35	87	2	browsed	browse	VERB
alces-35	87	3	aspens	aspen	NOUN
alces-35	87	4	in	in	ADP
alces-35	87	5	the	the	DET
alces-35	87	6	so	so	ADV
alces-35	87	7	more	more	ADV
alces-35	87	8	often	often	ADV
alces-35	87	9	than	than	ADP
alces-35	87	10	in	in	ADP
alces-35	87	11	the	the	DET
alces-35	87	12	no	no	DET
alces-35	87	13	area	area	NOUN
alces-35	87	14	.	.	PUNCT
alces-35	88	1	in	in	ADP
alces-35	88	2	total	total	ADJ
alces-35	88	3	,	,	PUNCT
alces-35	88	4	96	96	NUM
alces-35	88	5	%	%	NOUN
alces-35	88	6	of	of	ADP
alces-35	88	7	aspens	aspen	NOUN
alces-35	88	8	were	be	AUX
alces-35	88	9	damaged	damage	VERB
alces-35	88	10	in	in	ADP
alces-35	88	11	the	the	DET
alces-35	88	12	so	so	ADJ
alces-35	88	13	area	area	NOUN
alces-35	88	14	and	and	CCONJ
alces-35	88	15	62	62	NUM
alces-35	88	16	%	%	NOUN
alces-35	88	17	in	in	ADP
alces-35	88	18	the	the	DET
alces-35	88	19	no	no	DET
alces-35	88	20	area	area	NOUN
alces-35	88	21	;	;	PUNCT
alces-35	88	22	23	23	NUM
alces-35	88	23	%	%	NOUN
alces-35	88	24	of	of	ADP
alces-35	88	25	damaged	damage	VERB
alces-35	88	26	aspens	aspen	NOUN
alces-35	88	27	were	be	AUX
alces-35	88	28	dead	dead	ADJ
alces-35	88	29	in	in	ADP
alces-35	88	30	the	the	DET
alces-35	88	31	so	so	ADJ
alces-35	88	32	area	area	NOUN
alces-35	88	33	,	,	PUNCT
alces-35	88	34	all	all	PRON
alces-35	88	35	shorter	short	ADJ
alces-35	88	36	than	than	ADP
alces-35	88	37	5	5	NUM
alces-35	88	38	m.	m.	NOUN
alces-35	88	39	only	only	ADV
alces-35	88	40	3	3	NUM
alces-35	88	41	%	%	NOUN
alces-35	88	42	of	of	ADP
alces-35	88	43	the	the	DET
alces-35	88	44	damaged	damage	VERB
alces-35	88	45	aspens	aspen	NOUN
alces-35	88	46	were	be	AUX
alces-35	88	47	dead	dead	ADJ
alces-35	88	48	in	in	ADP
alces-35	88	49	the	the	DET
alces-35	88	50	no	no	DET
alces-35	88	51	area	area	NOUN
alces-35	88	52	.	.	PUNCT
alces-35	89	1	repeated	repeat	VERB
alces-35	89	2	stem	stem	NOUN
alces-35	89	3	breakage	breakage	NOUN
alces-35	89	4	was	be	AUX
alces-35	89	5	very	very	ADV
alces-35	89	6	common	common	ADJ
alces-35	89	7	.	.	PUNCT
alces-35	90	1	the	the	DET
alces-35	90	2	mean	mean	ADJ
alces-35	90	3	number	number	NOUN
alces-35	90	4	of	of	ADP
alces-35	90	5	stem	stem	NOUN
alces-35	90	6	breakages	breakage	NOUN
alces-35	90	7	per	per	ADP
alces-35	90	8	aspen	aspen	NOUN
alces-35	90	9	did	do	AUX
alces-35	90	10	not	not	PART
alces-35	90	11	differ	differ	VERB
alces-35	90	12	between	between	ADP
alces-35	90	13	the	the	DET
alces-35	90	14	two	two	NUM
alces-35	90	15	areas	area	NOUN
alces-35	90	16	(	(	PUNCT
alces-35	90	17	so	so	ADV
alces-35	90	18	:	:	PUNCT
alces-35	90	19	2.7	2.7	NUM
alces-35	90	20	times	time	NOUN
alces-35	90	21	/	/	SYM
alces-35	90	22	aspen	aspen	PROPN
alces-35	90	23	±	±	NUM
alces-35	90	24	0.3	0.3	NUM
alces-35	90	25	se	se	X
alces-35	90	26	vs.	vs.	ADP
alces-35	90	27	no	no	INTJ
alces-35	90	28	:	:	PUNCT
alces-35	90	29	2.2	2.2	NUM
alces-35	90	30	times	time	NOUN
alces-35	90	31	/	/	SYM
alces-35	90	32	aspen	aspen	PROPN
alces-35	90	33	±	±	NUM
alces-35	90	34	0.5	0.5	NUM
alces-35	90	35	se	se	X
alces-35	90	36	,	,	PUNCT
alces-35	90	37	mann	mann	PROPN
alces-35	90	38	-	-	PUNCT
alces-35	90	39	whitney	whitney	PROPN
alces-35	90	40	u	u	NOUN
alces-35	90	41	=	=	PROPN
alces-35	90	42	62.5	62.5	NUM
alces-35	90	43	,	,	PUNCT
alces-35	90	44	p	p	NOUN
alces-35	90	45	=	=	NOUN
alces-35	90	46	0.12	0.12	NUM
alces-35	90	47	)	)	PUNCT
alces-35	90	48	.	.	PUNCT
alces-35	91	1	in	in	ADP
alces-35	91	2	order	order	NOUN
alces-35	91	3	to	to	PART
alces-35	91	4	illustrate	illustrate	VERB
alces-35	91	5	the	the	DET
alces-35	91	6	effect	effect	NOUN
alces-35	91	7	of	of	ADP
alces-35	91	8	aspen	aspen	ADJ
alces-35	91	9	height	height	NOUN
alces-35	91	10	on	on	ADP
alces-35	91	11	moose	moose	NOUN
alces-35	91	12	browsing	browsing	NOUN
alces-35	91	13	,	,	PUNCT
alces-35	91	14	a	a	DET
alces-35	91	15	closer	close	ADJ
alces-35	91	16	examination	examination	NOUN
alces-35	91	17	of	of	ADP
alces-35	91	18	moose	moose	NOUN
alces-35	91	19	browsing	browse	VERB
alces-35	91	20	in	in	ADP
alces-35	91	21	three	three	NUM
alces-35	91	22	aspen	aspen	ADJ
alces-35	91	23	height	height	NOUN
alces-35	91	24	categories	category	NOUN
alces-35	91	25	was	be	AUX
alces-35	91	26	performed	perform	VERB
alces-35	91	27	(	(	PUNCT
alces-35	91	28	fig	fig	NOUN
alces-35	91	29	.	.	PUNCT
alces-35	92	1	2	2	NUM
alces-35	92	2	)	)	PUNCT
alces-35	92	3	.	.	PUNCT
alces-35	93	1	there	there	PRON
alces-35	93	2	was	be	VERB
alces-35	93	3	a	a	DET
alces-35	93	4	clear	clear	ADJ
alces-35	93	5	tendency	tendency	NOUN
alces-35	93	6	that	that	SCONJ
alces-35	93	7	aspens	aspen	NOUN
alces-35	93	8	in	in	ADP
alces-35	93	9	height	height	NOUN
alces-35	93	10	categories	category	NOUN
alces-35	93	11	i	i	PRON
alces-35	93	12	and	and	CCONJ
alces-35	93	13	ii	ii	PROPN
alces-35	93	14	were	be	AUX
alces-35	93	15	damaged	damage	VERB
alces-35	93	16	more	more	ADV
alces-35	93	17	often	often	ADV
alces-35	93	18	(	(	PUNCT
alces-35	93	19	twig	twig	NOUN
alces-35	93	20	-	-	PUNCT
alces-35	93	21	browsing	browse	VERB
alces-35	93	22	and	and	CCONJ
alces-35	93	23	stem	stem	NOUN
alces-35	93	24	breakage	breakage	NOUN
alces-35	93	25	)	)	PUNCT
alces-35	93	26	than	than	ADP
alces-35	93	27	aspens	aspen	NOUN
alces-35	93	28	in	in	ADP
alces-35	93	29	height	height	NOUN
alces-35	93	30	category	category	PROPN
alces-35	93	31	iii	iii	PROPN
alces-35	93	32	.	.	PUNCT
alces-35	94	1	bark	bark	NOUN
alces-35	94	2	stripping	strip	VERB
alces-35	94	3	was	be	AUX
alces-35	94	4	relatively	relatively	ADV
alces-35	94	5	common	common	ADJ
alces-35	94	6	in	in	ADP
alces-35	94	7	height	height	NOUN
alces-35	94	8	category	category	NOUN
alces-35	94	9	ii	ii	PROPN
alces-35	94	10	in	in	ADP
alces-35	94	11	both	both	DET
alces-35	94	12	areas	area	NOUN
alces-35	94	13	.	.	PUNCT
alces-35	95	1	in	in	ADP
alces-35	95	2	height	height	NOUN
alces-35	95	3	category	category	NOUN
alces-35	95	4	i	i	PRON
alces-35	95	5	,	,	PUNCT
alces-35	95	6	all	all	DET
alces-35	95	7	aspens	aspen	NOUN
alces-35	95	8	in	in	ADP
alces-35	95	9	the	the	DET
alces-35	95	10	so	so	ADJ
alces-35	95	11	area	area	NOUN
alces-35	95	12	had	have	VERB
alces-35	95	13	signs	sign	NOUN
alces-35	95	14	of	of	ADP
alces-35	95	15	old	old	ADJ
alces-35	95	16	damage	damage	NOUN
alces-35	95	17	.	.	PUNCT
alces-35	96	1	the	the	DET
alces-35	96	2	percentage	percentage	NOUN
alces-35	96	3	of	of	ADP
alces-35	96	4	new	new	ADJ
alces-35	96	5	damage	damage	NOUN
alces-35	96	6	showed	show	VERB
alces-35	96	7	0	0	NUM
alces-35	96	8	20	20	NUM
alces-35	96	9	40	40	NUM
alces-35	96	10	60	60	NUM
alces-35	96	11	80	80	NUM
alces-35	96	12	100	100	NUM
alces-35	97	1	so	so	CCONJ
alces-35	97	2	<	<	X
alces-35	97	3	5	5	NUM
alces-35	97	4	m	m	NOUN
alces-35	97	5	so	so	ADV
alces-35	97	6	5	5	NUM
alces-35	97	7	-	-	SYM
alces-35	97	8	15	15	NUM
alces-35	97	9	m	m	NOUN
alces-35	97	10	so	so	ADV
alces-35	97	11	>	>	X
alces-35	97	12	15	15	NUM
alces-35	97	13	m	m	NOUN
alces-35	97	14	no	no	PRON
alces-35	97	15	<	<	X
alces-35	97	16	5	5	NUM
alces-35	97	17	m	m	NOUN
alces-35	97	18	no	no	DET
alces-35	97	19	5	5	NUM
alces-35	97	20	-	-	SYM
alces-35	97	21	15	15	NUM
alces-35	97	22	m	m	NOUN
alces-35	97	23	no	no	INTJ
alces-35	97	24	>	>	X
alces-35	97	25	15	15	NUM
alces-35	97	26	m	m	NOUN
alces-35	97	27	%	%	NOUN
alces-35	97	28	broken	break	VERB
alces-35	97	29	stem	stem	NOUN
alces-35	97	30	new	new	ADJ
alces-35	97	31	broken	break	VERB
alces-35	97	32	stem	stem	NOUN
alces-35	97	33	old	old	ADJ
alces-35	97	34	broken	break	VERB
alces-35	97	35	stem	stem	NOUN
alces-35	97	36	0	0	NUM
alces-35	97	37	20	20	NUM
alces-35	97	38	40	40	NUM
alces-35	97	39	60	60	NUM
alces-35	97	40	80	80	NUM
alces-35	97	41	100	100	NUM
alces-35	98	1	so	so	CCONJ
alces-35	98	2	<	<	X
alces-35	98	3	5	5	NUM
alces-35	98	4	m	m	NOUN
alces-35	98	5	so	so	ADV
alces-35	98	6	5	5	NUM
alces-35	98	7	-	-	SYM
alces-35	98	8	15	15	NUM
alces-35	98	9	m	m	NOUN
alces-35	98	10	so	so	ADV
alces-35	98	11	>	>	X
alces-35	98	12	15	15	NUM
alces-35	98	13	m	m	NOUN
alces-35	98	14	no	no	PRON
alces-35	98	15	<	<	X
alces-35	98	16	5	5	NUM
alces-35	98	17	m	m	NOUN
alces-35	98	18	no	no	DET
alces-35	98	19	5	5	NUM
alces-35	98	20	-	-	SYM
alces-35	98	21	15	15	NUM
alces-35	98	22	m	m	NOUN
alces-35	98	23	no	no	INTJ
alces-35	98	24	>	>	X
alces-35	98	25	15	15	NUM
alces-35	98	26	m	m	NOUN
alces-35	98	27	%	%	NOUN
alces-35	98	28	twig	twig	NOUN
alces-35	98	29	-	-	PUNCT
alces-35	98	30	browsing	browse	VERB
alces-35	98	31	twig	twig	NOUN
alces-35	98	32	-	-	PUNCT
alces-35	98	33	browsing	browse	VERB
alces-35	98	34	,	,	PUNCT
alces-35	98	35	new	new	ADJ
alces-35	98	36	twig	twig	NOUN
alces-35	98	37	-	-	PUNCT
alces-35	98	38	browsing	browse	VERB
alces-35	98	39	,	,	PUNCT
alces-35	98	40	old	old	ADJ
alces-35	98	41	0	0	NUM
alces-35	98	42	20	20	NUM
alces-35	98	43	40	40	NUM
alces-35	98	44	60	60	NUM
alces-35	98	45	80	80	NUM
alces-35	98	46	100	100	NUM
alces-35	99	1	so	so	CCONJ
alces-35	99	2	<	<	X
alces-35	99	3	5	5	NUM
alces-35	99	4	m	m	NOUN
alces-35	99	5	so	so	ADV
alces-35	99	6	5	5	NUM
alces-35	99	7	-	-	SYM
alces-35	99	8	15	15	NUM
alces-35	99	9	m	m	NOUN
alces-35	99	10	so	so	ADV
alces-35	99	11	>	>	X
alces-35	99	12	15	15	NUM
alces-35	99	13	m	m	NOUN
alces-35	99	14	no	no	PRON
alces-35	99	15	<	<	X
alces-35	99	16	5	5	NUM
alces-35	99	17	m	m	NOUN
alces-35	99	18	no	no	DET
alces-35	99	19	5	5	NUM
alces-35	99	20	-	-	SYM
alces-35	99	21	15	15	NUM
alces-35	99	22	m	m	NOUN
alces-35	99	23	no	no	INTJ
alces-35	99	24	>	>	X
alces-35	99	25	15	15	NUM
alces-35	99	26	m	m	NOUN
alces-35	99	27	%	%	NOUN
alces-35	99	28	bark	bark	NOUN
alces-35	99	29	stripping	strip	VERB
alces-35	99	30	new	new	ADJ
alces-35	99	31	bark	bark	NOUN
alces-35	99	32	stripping	strip	VERB
alces-35	99	33	old	old	ADJ
alces-35	99	34	bark	bark	NOUN
alces-35	99	35	stripping	strip	VERB
alces-35	99	36	0	0	NUM
alces-35	99	37	20	20	NUM
alces-35	99	38	40	40	NUM
alces-35	99	39	60	60	NUM
alces-35	99	40	80	80	NUM
alces-35	99	41	100	100	NUM
alces-35	99	42	so	so	CCONJ
alces-35	99	43	<	<	X
alces-35	99	44	5	5	NUM
alces-35	99	45	m	m	NOUN
alces-35	99	46	so	so	ADV
alces-35	99	47	5	5	NUM
alces-35	99	48	-	-	SYM
alces-35	99	49	15	15	NUM
alces-35	99	50	m	m	NOUN
alces-35	99	51	so	so	ADV
alces-35	99	52	>	>	X
alces-35	99	53	15	15	NUM
alces-35	99	54	m	m	NOUN
alces-35	99	55	no	no	PRON
alces-35	99	56	<	<	X
alces-35	99	57	5	5	NUM
alces-35	99	58	m	m	NOUN
alces-35	99	59	no	no	DET
alces-35	99	60	5	5	NUM
alces-35	99	61	-	-	SYM
alces-35	99	62	15	15	NUM
alces-35	99	63	m	m	NOUN
alces-35	99	64	no	no	INTJ
alces-35	99	65	>	>	SYM
alces-35	99	66	15	15	NUM
alces-35	99	67	m	m	NOUN
alces-35	99	68	%	%	NOUN
alces-35	99	69	damaged	damage	VERB
alces-35	99	70	new	new	ADJ
alces-35	99	71	damage	damage	NOUN
alces-35	99	72	old	old	ADJ
alces-35	99	73	damage	damage	NOUN
alces-35	99	74	fig	fig	NOUN
alces-35	99	75	.	.	PUNCT
alces-35	100	1	2	2	X
alces-35	100	2	.	.	X
alces-35	100	3	proportions	proportion	NOUN
alces-35	100	4	of	of	ADP
alces-35	100	5	different	different	ADJ
alces-35	100	6	damage	damage	NOUN
alces-35	100	7	types	type	NOUN
alces-35	100	8	in	in	ADP
alces-35	100	9	different	different	ADJ
alces-35	100	10	aspen	aspen	NOUN
alces-35	100	11	height	height	NOUN
alces-35	100	12	categories	category	NOUN
alces-35	100	13	at	at	ADP
alces-35	100	14	the	the	DET
alces-35	100	15	landscape	landscape	NOUN
alces-35	100	16	level	level	NOUN
alces-35	100	17	in	in	ADP
alces-35	100	18	koli	koli	PROPN
alces-35	100	19	national	national	PROPN
alces-35	100	20	park	park	PROPN
alces-35	100	21	,	,	PUNCT
alces-35	100	22	finland	finland	PROPN
alces-35	100	23	(	(	PUNCT
alces-35	100	24	so	so	ADV
alces-35	100	25	=	=	SYM
alces-35	100	26	southern	southern	ADJ
alces-35	100	27	area	area	NOUN
alces-35	100	28	;	;	PUNCT
alces-35	100	29	no	no	DET
alces-35	100	30	=	=	SYM
alces-35	100	31	northern	northern	ADJ
alces-35	100	32	area	area	NOUN
alces-35	100	33	)	)	PUNCT
alces-35	100	34	.	.	PUNCT
alces-35	101	1	the	the	DET
alces-35	101	2	sum	sum	NOUN
alces-35	101	3	of	of	ADP
alces-35	101	4	percentages	percentage	NOUN
alces-35	101	5	of	of	ADP
alces-35	101	6	new	new	ADJ
alces-35	101	7	and	and	CCONJ
alces-35	101	8	old	old	ADJ
alces-35	101	9	damage	damage	NOUN
alces-35	101	10	may	may	AUX
alces-35	101	11	exceed	exceed	VERB
alces-35	101	12	the	the	DET
alces-35	101	13	percentage	percentage	NOUN
alces-35	101	14	of	of	ADP
alces-35	101	15	total	total	ADJ
alces-35	101	16	damage	damage	NOUN
alces-35	101	17	within	within	ADP
alces-35	101	18	a	a	DET
alces-35	101	19	height	height	NOUN
alces-35	101	20	category	category	NOUN
alces-35	101	21	due	due	ADP
alces-35	101	22	to	to	ADP
alces-35	101	23	repeated	repeat	VERB
alces-35	101	24	browsing	browsing	NOUN
alces-35	101	25	of	of	ADP
alces-35	101	26	the	the	DET
alces-35	101	27	same	same	ADJ
alces-35	101	28	stems	stem	NOUN
alces-35	101	29	.	.	PUNCT
alces-35	102	1	alces	alce	NOUN
alces-35	102	2	vol	vol	NOUN
alces-35	102	3	.	.	PROPN
alces-35	102	4	44	44	NUM
alces-35	102	5	,	,	PUNCT
alces-35	102	6	2008	2008	NUM
alces-35	102	7	härkönen	härkönen	NOUN
alces-35	102	8	et	et	PROPN
alces-35	102	9	al.moose	al.moose	X
alces-35	102	10	and	and	CCONJ
alces-35	102	11	aspen	aspen	VERB
alces-35	102	12	regeneration	regeneration	NOUN
alces-35	102	13	in	in	ADP
alces-35	102	14	finland	finland	PROPN
alces-35	102	15	35	35	NUM
alces-35	103	1	that	that	PRON
alces-35	103	2	re	re	VERB
alces-35	103	3	-	-	VERB
alces-35	103	4	browsing	browse	VERB
alces-35	103	5	hampered	hamper	VERB
alces-35	103	6	young	young	ADJ
alces-35	103	7	trees	tree	NOUN
alces-35	103	8	most	most	ADV
alces-35	103	9	.	.	PUNCT
alces-35	104	1	the	the	DET
alces-35	104	2	proportion	proportion	NOUN
alces-35	104	3	of	of	ADP
alces-35	104	4	young	young	ADJ
alces-35	104	5	aspens	aspen	NOUN
alces-35	104	6	(	(	PUNCT
alces-35	104	7	i.e.	i.e.	X
alces-35	104	8	,	,	PUNCT
alces-35	104	9	height	height	NOUN
alces-35	104	10	category	category	NOUN
alces-35	104	11	i	i	PRON
alces-35	104	12	)	)	PUNCT
alces-35	104	13	decreased	decrease	VERB
alces-35	104	14	from	from	ADP
alces-35	104	15	45	45	NUM
alces-35	104	16	%	%	NOUN
alces-35	104	17	in	in	ADP
alces-35	104	18	advanced	advanced	ADJ
alces-35	104	19	seedling	seedling	NOUN
alces-35	104	20	stands	stand	VERB
alces-35	104	21	to	to	ADP
alces-35	104	22	17	17	NUM
alces-35	104	23	%	%	NOUN
alces-35	104	24	in	in	ADP
alces-35	104	25	mature	mature	ADJ
alces-35	104	26	stands	stand	NOUN
alces-35	104	27	indicating	indicate	VERB
alces-35	104	28	that	that	SCONJ
alces-35	104	29	regeneration	regeneration	NOUN
alces-35	104	30	of	of	ADP
alces-35	104	31	aspen	aspen	NOUN
alces-35	104	32	,	,	PUNCT
alces-35	104	33	to	to	ADP
alces-35	104	34	some	some	DET
alces-35	104	35	extent	extent	NOUN
alces-35	104	36	,	,	PUNCT
alces-35	104	37	occurs	occur	VERB
alces-35	104	38	in	in	ADP
alces-35	104	39	closed	close	VERB
alces-35	104	40	older	old	ADJ
alces-35	104	41	stands	stand	NOUN
alces-35	104	42	.	.	PUNCT
alces-35	105	1	stand	stand	VERB
alces-35	105	2	level	level	NOUN
alces-35	105	3	aspen	aspen	NOUN
alces-35	105	4	density	density	NOUN
alces-35	105	5	declined	decline	VERB
alces-35	105	6	from	from	ADP
alces-35	105	7	young	young	ADJ
alces-35	105	8	to	to	ADP
alces-35	105	9	old	old	ADJ
alces-35	105	10	stands	stand	NOUN
alces-35	105	11	in	in	ADP
alces-35	105	12	both	both	DET
alces-35	105	13	areas	area	NOUN
alces-35	105	14	(	(	PUNCT
alces-35	105	15	table	table	NOUN
alces-35	105	16	1	1	NUM
alces-35	105	17	)	)	PUNCT
alces-35	105	18	.	.	PUNCT
alces-35	106	1	moose	moose	NOUN
alces-35	106	2	seemingly	seemingly	ADV
alces-35	106	3	used	use	VERB
alces-35	106	4	stands	stand	VERB
alces-35	106	5	relatively	relatively	ADV
alces-35	106	6	little	little	ADJ
alces-35	106	7	as	as	SCONJ
alces-35	106	8	the	the	DET
alces-35	106	9	number	number	NOUN
alces-35	106	10	of	of	ADP
alces-35	106	11	fecal	fecal	ADJ
alces-35	106	12	pellet	pellet	NOUN
alces-35	106	13	groups	group	NOUN
alces-35	106	14	was	be	AUX
alces-35	106	15	very	very	ADV
alces-35	106	16	low	low	ADJ
alces-35	106	17	.	.	PUNCT
alces-35	107	1	the	the	DET
alces-35	107	2	mean	mean	ADJ
alces-35	107	3	height	height	NOUN
alces-35	107	4	was	be	AUX
alces-35	107	5	similar	similar	ADJ
alces-35	107	6	in	in	ADP
alces-35	107	7	same	same	ADJ
alces-35	107	8	aged	aged	ADJ
alces-35	107	9	stands	stand	VERB
alces-35	107	10	in	in	ADP
alces-35	107	11	both	both	DET
alces-35	107	12	areas	area	NOUN
alces-35	107	13	(	(	PUNCT
alces-35	107	14	table	table	NOUN
alces-35	107	15	1	1	NUM
alces-35	107	16	)	)	PUNCT
alces-35	107	17	.	.	PUNCT
alces-35	108	1	based	base	VERB
alces-35	108	2	on	on	ADP
alces-35	108	3	damage	damage	NOUN
alces-35	108	4	type	type	NOUN
alces-35	108	5	,	,	PUNCT
alces-35	108	6	browsing	browse	VERB
alces-35	108	7	pressure	pressure	NOUN
alces-35	108	8	by	by	ADP
alces-35	108	9	moose	moose	NOUN
alces-35	108	10	was	be	AUX
alces-35	108	11	very	very	ADV
alces-35	108	12	similar	similar	ADJ
alces-35	108	13	in	in	ADP
alces-35	108	14	both	both	DET
alces-35	108	15	areas	area	NOUN
alces-35	108	16	(	(	PUNCT
alces-35	108	17	fig	fig	NOUN
alces-35	108	18	.	.	PUNCT
alces-35	109	1	3	3	NUM
alces-35	109	2	)	)	PUNCT
alces-35	109	3	.	.	PUNCT
alces-35	110	1	on	on	ADP
alces-35	110	2	average	average	ADJ
alces-35	110	3	,	,	PUNCT
alces-35	110	4	79	79	NUM
alces-35	110	5	%	%	NOUN
alces-35	110	6	of	of	ADP
alces-35	110	7	the	the	DET
alces-35	110	8	aspens	aspen	NOUN
alces-35	110	9	in	in	ADP
alces-35	110	10	the	the	DET
alces-35	110	11	so	so	ADJ
alces-35	110	12	area	area	NOUN
alces-35	110	13	were	be	AUX
alces-35	110	14	damaged	damage	VERB
alces-35	110	15	,	,	PUNCT
alces-35	110	16	as	as	SCONJ
alces-35	110	17	compared	compare	VERB
alces-35	110	18	to	to	ADP
alces-35	110	19	73	73	NUM
alces-35	110	20	%	%	NOUN
alces-35	110	21	in	in	ADP
alces-35	110	22	the	the	DET
alces-35	110	23	no	no	DET
alces-35	110	24	area	area	NOUN
alces-35	110	25	.	.	PUNCT
alces-35	111	1	the	the	DET
alces-35	111	2	proportions	proportion	NOUN
alces-35	111	3	of	of	ADP
alces-35	111	4	new	new	ADJ
alces-35	111	5	damage	damage	NOUN
alces-35	111	6	showed	show	VERB
alces-35	111	7	that	that	SCONJ
alces-35	111	8	re	re	VERB
alces-35	111	9	-	-	VERB
alces-35	111	10	browsing	browse	VERB
alces-35	111	11	occurred	occur	VERB
alces-35	111	12	predominantly	predominantly	ADV
alces-35	111	13	in	in	ADP
alces-35	111	14	young	young	ADJ
alces-35	111	15	stands	stand	NOUN
alces-35	111	16	.	.	PUNCT
alces-35	112	1	the	the	DET
alces-35	112	2	proportion	proportion	NOUN
alces-35	112	3	of	of	ADP
alces-35	112	4	dead	dead	ADJ
alces-35	112	5	aspens	aspen	NOUN
alces-35	112	6	was	be	AUX
alces-35	112	7	relatively	relatively	ADV
alces-35	112	8	high	high	ADJ
alces-35	112	9	in	in	ADP
alces-35	112	10	middle	middle	NOUN
alces-35	112	11	-	-	PUNCT
alces-35	112	12	aged	aged	ADJ
alces-35	112	13	stands	stand	VERB
alces-35	112	14	in	in	ADP
alces-35	112	15	the	the	DET
alces-35	112	16	so	so	ADV
alces-35	112	17	and	and	CCONJ
alces-35	112	18	no	no	DET
alces-35	112	19	areas	area	NOUN
alces-35	112	20	(	(	PUNCT
alces-35	112	21	fig	fig	NOUN
alces-35	112	22	.	.	PUNCT
alces-35	112	23	4	4	NUM
alces-35	112	24	)	)	PUNCT
alces-35	112	25	.	.	PUNCT
alces-35	113	1	the	the	DET
alces-35	113	2	mean	mean	ADJ
alces-35	113	3	number	number	NOUN
alces-35	113	4	of	of	ADP
alces-35	113	5	stem	stem	NOUN
alces-35	113	6	breakages	breakage	NOUN
alces-35	113	7	per	per	ADP
alces-35	113	8	aspen	aspen	NOUN
alces-35	113	9	did	do	AUX
alces-35	113	10	not	not	PART
alces-35	113	11	differ	differ	VERB
alces-35	113	12	between	between	ADP
alces-35	113	13	the	the	DET
alces-35	113	14	two	two	NUM
alces-35	113	15	areas	area	NOUN
alces-35	113	16	(	(	PUNCT
alces-35	113	17	so	so	ADV
alces-35	113	18	:	:	PUNCT
alces-35	113	19	2.3	2.3	NUM
alces-35	113	20	times/	times/	NUM
alces-35	113	21	aspen	aspen	PROPN
alces-35	113	22	±	±	NUM
alces-35	113	23	0.3	0.3	NUM
alces-35	113	24	se	se	X
alces-35	113	25	vs.	vs.	ADP
alces-35	113	26	no	no	INTJ
alces-35	113	27	:	:	PUNCT
alces-35	113	28	2.0	2.0	NUM
alces-35	113	29	times	time	NOUN
alces-35	113	30	/	/	SYM
alces-35	113	31	aspen	aspen	PROPN
alces-35	113	32	±	±	NUM
alces-35	113	33	0.2	0.2	NUM
alces-35	113	34	se	se	X
alces-35	113	35	,	,	PUNCT
alces-35	113	36	mann	mann	PROPN
alces-35	113	37	-	-	PUNCT
alces-35	113	38	whitney	whitney	PROPN
alces-35	113	39	u	u	NOUN
alces-35	113	40	=	=	PROPN
alces-35	113	41	48.0	48.0	NUM
alces-35	113	42	,	,	PUNCT
alces-35	113	43	p	p	X
alces-35	113	44	=	=	NOUN
alces-35	113	45	0.43	0.43	NUM
alces-35	113	46	)	)	PUNCT
alces-35	113	47	.	.	PUNCT
alces-35	114	1	so	so	ADV
alces-35	114	2	<	<	X
alces-35	114	3	5	5	NUM
alces-35	114	4	m	m	NOUN
alces-35	114	5	so	so	ADV
alces-35	114	6	5	5	NUM
alces-35	114	7	-	-	SYM
alces-35	114	8	15	15	NUM
alces-35	114	9	m	m	NOUN
alces-35	114	10	so	so	ADV
alces-35	114	11	>	>	SYM
alces-35	114	12	15	15	NUM
alces-35	114	13	m	m	NOUN
alces-35	114	14	no	no	PRON
alces-35	114	15	<	<	X
alces-35	114	16	5	5	NUM
alces-35	114	17	m	m	NOUN
alces-35	114	18	no	no	DET
alces-35	114	19	5	5	NUM
alces-35	114	20	-	-	SYM
alces-35	114	21	15	15	NUM
alces-35	114	22	m	m	NOUN
alces-35	114	23	no	no	INTJ
alces-35	114	24	>	>	SYM
alces-35	114	25	15	15	NUM
alces-35	114	26	m	m	NOUN
alces-35	114	27	aspen	aspen	ADJ
alces-35	114	28	density	density	NOUN
alces-35	114	29	4,458	4,458	NUM
alces-35	114	30	±	±	NUM
alces-35	114	31	725	725	NUM
alces-35	114	32	1,987	1,987	NUM
alces-35	114	33	±	±	NUM
alces-35	114	34	253	253	NUM
alces-35	114	35	383	383	NUM
alces-35	114	36	±	±	NUM
alces-35	114	37	13	13	NUM
alces-35	114	38	2,371	2,371	NUM
alces-35	114	39	±	±	NUM
alces-35	114	40	343	343	NUM
alces-35	114	41	2,306	2,306	NUM
alces-35	114	42	±	±	NUM
alces-35	114	43	710	710	NUM
alces-35	114	44	896	896	NUM
alces-35	114	45	±	±	NUM
alces-35	114	46	151	151	NUM
alces-35	114	47	aspen	aspen	NOUN
alces-35	114	48	height	height	NOUN
alces-35	114	49	1.4	1.4	NUM
alces-35	114	50	±	±	NUM
alces-35	114	51	0.1	0.1	NUM
alces-35	114	52	6	6	NUM
alces-35	114	53	±	±	NUM
alces-35	114	54	0.6	0.6	NUM
alces-35	114	55	20.1	20.1	NUM
alces-35	114	56	±	±	NUM
alces-35	114	57	1.5	1.5	NUM
alces-35	114	58	1.4	1.4	NUM
alces-35	114	59	±	±	NUM
alces-35	114	60	0.2	0.2	NUM
alces-35	114	61	4.7	4.7	NUM
alces-35	114	62	±	±	NUM
alces-35	114	63	0.4	0.4	NUM
alces-35	114	64	17.6	17.6	NUM
alces-35	114	65	±	±	NUM
alces-35	114	66	1.8	1.8	NUM
alces-35	114	67	pellet	pellet	NOUN
alces-35	114	68	groups	group	NOUN
alces-35	114	69	12	12	NUM
alces-35	114	70	±	±	NUM
alces-35	114	71	7	7	NUM
alces-35	114	72	12	12	NUM
alces-35	114	73	±	±	NUM
alces-35	114	74	7	7	NUM
alces-35	114	75	table	table	NOUN
alces-35	114	76	1	1	NUM
alces-35	114	77	.	.	PUNCT
alces-35	115	1	aspen	aspen	ADJ
alces-35	115	2	density	density	NOUN
alces-35	115	3	(	(	PUNCT
alces-35	115	4	trees	trees	PROPN
alces-35	115	5	/	/	SYM
alces-35	115	6	ha	ha	NOUN
alces-35	115	7	)	)	PUNCT
alces-35	115	8	and	and	CCONJ
alces-35	115	9	height	height	NOUN
alces-35	115	10	(	(	PUNCT
alces-35	115	11	m	m	NOUN
alces-35	115	12	)	)	PUNCT
alces-35	115	13	,	,	PUNCT
alces-35	115	14	and	and	CCONJ
alces-35	115	15	number	number	NOUN
alces-35	115	16	of	of	ADP
alces-35	115	17	fecal	fecal	ADJ
alces-35	115	18	pellet	pellet	NOUN
alces-35	115	19	groups	group	NOUN
alces-35	115	20	(	(	PUNCT
alces-35	115	21	/ha	/ha	NOUN
alces-35	115	22	)	)	PUNCT
alces-35	115	23	in	in	ADP
alces-35	115	24	young	young	ADJ
alces-35	115	25	(	(	PUNCT
alces-35	115	26	<	<	X
alces-35	115	27	5	5	NUM
alces-35	115	28	m	m	NOUN
alces-35	115	29	in	in	ADP
alces-35	115	30	height	height	NOUN
alces-35	115	31	)	)	PUNCT
alces-35	115	32	,	,	PUNCT
alces-35	115	33	middle	middle	NOUN
alces-35	115	34	-	-	PUNCT
alces-35	115	35	aged	aged	ADJ
alces-35	115	36	(	(	PUNCT
alces-35	115	37	5	5	NUM
alces-35	115	38	-	-	SYM
alces-35	115	39	15	15	NUM
alces-35	115	40	m	m	NOUN
alces-35	115	41	in	in	ADP
alces-35	115	42	height	height	NOUN
alces-35	115	43	)	)	PUNCT
alces-35	115	44	,	,	PUNCT
alces-35	115	45	and	and	CCONJ
alces-35	115	46	old	old	ADJ
alces-35	115	47	(	(	PUNCT
alces-35	115	48	>	>	X
alces-35	115	49	15	15	NUM
alces-35	115	50	m	m	NOUN
alces-35	115	51	in	in	ADP
alces-35	115	52	height	height	NOUN
alces-35	115	53	)	)	PUNCT
alces-35	115	54	aspen	aspen	PROPN
alces-35	115	55	-	-	PUNCT
alces-35	115	56	rich	rich	ADJ
alces-35	115	57	stands	stand	VERB
alces-35	115	58	at	at	ADP
alces-35	115	59	the	the	DET
alces-35	115	60	stand	stand	NOUN
alces-35	115	61	level	level	NOUN
alces-35	115	62	in	in	ADP
alces-35	115	63	aspen	aspen	NOUN
alces-35	115	64	-	-	PUNCT
alces-35	115	65	rich	rich	ADJ
alces-35	115	66	forest	forest	NOUN
alces-35	115	67	stands	stand	VERB
alces-35	115	68	in	in	ADP
alces-35	115	69	koli	koli	PROPN
alces-35	115	70	national	national	PROPN
alces-35	115	71	park	park	PROPN
alces-35	115	72	,	,	PUNCT
alces-35	115	73	finland	finland	PROPN
alces-35	115	74	.	.	PUNCT
alces-35	116	1	means	mean	NOUN
alces-35	116	2	are	be	AUX
alces-35	116	3	given	give	VERB
alces-35	116	4	with	with	ADP
alces-35	116	5	their	their	PRON
alces-35	116	6	standard	standard	ADJ
alces-35	116	7	errors	error	NOUN
alces-35	116	8	.	.	PUNCT
alces-35	117	1	note	note	VERB
alces-35	117	2	:	:	PUNCT
alces-35	118	1	so	so	ADV
alces-35	118	2	=	=	SYM
alces-35	118	3	southern	southern	ADJ
alces-35	118	4	area	area	NOUN
alces-35	118	5	of	of	ADP
alces-35	118	6	koli	koli	PROPN
alces-35	118	7	national	national	PROPN
alces-35	118	8	park	park	PROPN
alces-35	118	9	;	;	PUNCT
alces-35	118	10	no	no	DET
alces-35	118	11	=	=	SYM
alces-35	118	12	northern	northern	ADJ
alces-35	118	13	area	area	NOUN
alces-35	118	14	of	of	ADP
alces-35	118	15	koli	koli	PROPN
alces-35	118	16	national	national	PROPN
alces-35	118	17	park	park	PROPN
alces-35	118	18	.	.	PUNCT
alces-35	119	1	0	0	NUM
alces-35	120	1	20	20	NUM
alces-35	120	2	40	40	NUM
alces-35	120	3	60	60	NUM
alces-35	120	4	80	80	NUM
alces-35	120	5	100	100	NUM
alces-35	121	1	so	so	CCONJ
alces-35	121	2	<	<	X
alces-35	121	3	5	5	NUM
alces-35	121	4	m	m	NOUN
alces-35	121	5	so	so	ADV
alces-35	121	6	5	5	NUM
alces-35	121	7	-	-	SYM
alces-35	121	8	15	15	NUM
alces-35	121	9	m	m	NOUN
alces-35	121	10	so	so	ADV
alces-35	121	11	>	>	X
alces-35	121	12	15	15	NUM
alces-35	121	13	m	m	NOUN
alces-35	121	14	no	no	PRON
alces-35	121	15	<	<	X
alces-35	121	16	5	5	NUM
alces-35	121	17	m	m	NOUN
alces-35	121	18	no	no	DET
alces-35	121	19	5	5	NUM
alces-35	121	20	-	-	SYM
alces-35	121	21	15	15	NUM
alces-35	121	22	m	m	NOUN
alces-35	121	23	no	no	INTJ
alces-35	121	24	>	>	X
alces-35	121	25	15	15	NUM
alces-35	121	26	m	m	NOUN
alces-35	121	27	%	%	NOUN
alces-35	121	28	broken	break	VERB
alces-35	121	29	stem	stem	NOUN
alces-35	121	30	new	new	ADJ
alces-35	121	31	broken	break	VERB
alces-35	121	32	stem	stem	NOUN
alces-35	121	33	old	old	ADJ
alces-35	121	34	broken	break	VERB
alces-35	121	35	stem	stem	NOUN
alces-35	121	36	0	0	NUM
alces-35	121	37	20	20	NUM
alces-35	121	38	40	40	NUM
alces-35	121	39	60	60	NUM
alces-35	121	40	80	80	NUM
alces-35	121	41	100	100	NUM
alces-35	122	1	so	so	CCONJ
alces-35	122	2	<	<	X
alces-35	122	3	5	5	NUM
alces-35	122	4	m	m	NOUN
alces-35	122	5	so	so	ADV
alces-35	122	6	5	5	NUM
alces-35	122	7	-	-	SYM
alces-35	122	8	15	15	NUM
alces-35	122	9	m	m	NOUN
alces-35	122	10	so	so	ADV
alces-35	122	11	>	>	X
alces-35	122	12	15	15	NUM
alces-35	122	13	m	m	NOUN
alces-35	122	14	no	no	PRON
alces-35	122	15	<	<	X
alces-35	122	16	5	5	NUM
alces-35	122	17	m	m	NOUN
alces-35	122	18	no	no	DET
alces-35	122	19	5	5	NUM
alces-35	122	20	-	-	SYM
alces-35	122	21	15	15	NUM
alces-35	122	22	m	m	NOUN
alces-35	122	23	no	no	INTJ
alces-35	122	24	>	>	X
alces-35	122	25	15	15	NUM
alces-35	122	26	m	m	NOUN
alces-35	122	27	%	%	NOUN
alces-35	122	28	twig	twig	NOUN
alces-35	122	29	-	-	PUNCT
alces-35	122	30	browsing	browse	VERB
alces-35	122	31	twig	twig	NOUN
alces-35	122	32	-	-	PUNCT
alces-35	122	33	browsing	browse	VERB
alces-35	122	34	,	,	PUNCT
alces-35	122	35	new	new	ADJ
alces-35	122	36	twig	twig	NOUN
alces-35	122	37	-	-	PUNCT
alces-35	122	38	browsing	browse	VERB
alces-35	122	39	,	,	PUNCT
alces-35	122	40	old	old	ADJ
alces-35	122	41	0	0	NUM
alces-35	122	42	20	20	NUM
alces-35	122	43	40	40	NUM
alces-35	122	44	60	60	NUM
alces-35	122	45	80	80	NUM
alces-35	122	46	100	100	NUM
alces-35	123	1	so	so	CCONJ
alces-35	123	2	<	<	X
alces-35	123	3	5	5	NUM
alces-35	123	4	m	m	NOUN
alces-35	123	5	so	so	ADV
alces-35	123	6	5	5	NUM
alces-35	123	7	-	-	SYM
alces-35	123	8	15	15	NUM
alces-35	123	9	m	m	NOUN
alces-35	123	10	so	so	ADV
alces-35	123	11	>	>	X
alces-35	123	12	15	15	NUM
alces-35	123	13	m	m	NOUN
alces-35	123	14	no	no	PRON
alces-35	123	15	<	<	X
alces-35	123	16	5	5	NUM
alces-35	123	17	m	m	NOUN
alces-35	123	18	no	no	DET
alces-35	123	19	5	5	NUM
alces-35	123	20	-	-	SYM
alces-35	123	21	15	15	NUM
alces-35	123	22	m	m	NOUN
alces-35	123	23	no	no	INTJ
alces-35	123	24	>	>	X
alces-35	123	25	15	15	NUM
alces-35	123	26	m	m	NOUN
alces-35	123	27	%	%	NOUN
alces-35	123	28	bark	bark	NOUN
alces-35	123	29	stripping	strip	VERB
alces-35	123	30	new	new	ADJ
alces-35	123	31	bark	bark	NOUN
alces-35	123	32	stripping	strip	VERB
alces-35	123	33	old	old	ADJ
alces-35	123	34	bark	bark	NOUN
alces-35	123	35	stripping	strip	VERB
alces-35	123	36	0	0	NUM
alces-35	123	37	20	20	NUM
alces-35	123	38	40	40	NUM
alces-35	123	39	60	60	NUM
alces-35	123	40	80	80	NUM
alces-35	123	41	100	100	NUM
alces-35	123	42	so	so	CCONJ
alces-35	123	43	<	<	X
alces-35	123	44	5	5	NUM
alces-35	123	45	m	m	NOUN
alces-35	123	46	so	so	ADV
alces-35	123	47	5	5	NUM
alces-35	123	48	-	-	SYM
alces-35	123	49	15	15	NUM
alces-35	123	50	m	m	NOUN
alces-35	123	51	so	so	ADV
alces-35	123	52	>	>	X
alces-35	123	53	15	15	NUM
alces-35	123	54	m	m	NOUN
alces-35	123	55	no	no	PRON
alces-35	123	56	<	<	X
alces-35	123	57	5	5	NUM
alces-35	123	58	m	m	NOUN
alces-35	123	59	no	no	DET
alces-35	123	60	5	5	NUM
alces-35	123	61	-	-	SYM
alces-35	123	62	15	15	NUM
alces-35	123	63	m	m	NOUN
alces-35	123	64	no	no	INTJ
alces-35	123	65	>	>	SYM
alces-35	123	66	15	15	NUM
alces-35	123	67	m	m	NOUN
alces-35	123	68	%	%	NOUN
alces-35	123	69	damaged	damage	VERB
alces-35	123	70	new	new	ADJ
alces-35	123	71	damage	damage	NOUN
alces-35	123	72	old	old	ADJ
alces-35	123	73	damage	damage	NOUN
alces-35	123	74	fig	fig	NOUN
alces-35	123	75	.	.	PUNCT
alces-35	124	1	3	3	X
alces-35	124	2	.	.	X
alces-35	124	3	proportions	proportion	NOUN
alces-35	124	4	of	of	ADP
alces-35	124	5	different	different	ADJ
alces-35	124	6	damage	damage	NOUN
alces-35	124	7	types	type	NOUN
alces-35	124	8	in	in	ADP
alces-35	124	9	young	young	ADJ
alces-35	124	10	(	(	PUNCT
alces-35	124	11	<	<	X
alces-35	124	12	5	5	NUM
alces-35	124	13	m	m	NOUN
alces-35	124	14	in	in	ADP
alces-35	124	15	height	height	NOUN
alces-35	124	16	)	)	PUNCT
alces-35	124	17	,	,	PUNCT
alces-35	124	18	middle	middle	NOUN
alces-35	124	19	-	-	PUNCT
alces-35	124	20	aged	aged	ADJ
alces-35	124	21	(	(	PUNCT
alces-35	124	22	5	5	NUM
alces-35	124	23	-	-	SYM
alces-35	124	24	15	15	NUM
alces-35	124	25	m	m	NOUN
alces-35	124	26	in	in	ADP
alces-35	124	27	height	height	NOUN
alces-35	124	28	)	)	PUNCT
alces-35	124	29	,	,	PUNCT
alces-35	124	30	and	and	CCONJ
alces-35	124	31	old	old	ADJ
alces-35	124	32	(	(	PUNCT
alces-35	124	33	>	>	X
alces-35	124	34	15	15	NUM
alces-35	124	35	m	m	NOUN
alces-35	124	36	in	in	ADP
alces-35	124	37	height	height	NOUN
alces-35	124	38	)	)	PUNCT
alces-35	124	39	aspen	aspen	PROPN
alces-35	124	40	-	-	PUNCT
alces-35	124	41	rich	rich	ADJ
alces-35	124	42	stands	stand	VERB
alces-35	124	43	at	at	ADP
alces-35	124	44	the	the	DET
alces-35	124	45	stand	stand	NOUN
alces-35	124	46	level	level	NOUN
alces-35	124	47	in	in	ADP
alces-35	124	48	koli	koli	PROPN
alces-35	124	49	national	national	PROPN
alces-35	124	50	park	park	PROPN
alces-35	124	51	,	,	PUNCT
alces-35	124	52	finland	finland	PROPN
alces-35	124	53	(	(	PUNCT
alces-35	124	54	so	so	ADV
alces-35	124	55	=	=	SYM
alces-35	124	56	southern	southern	ADJ
alces-35	124	57	area	area	NOUN
alces-35	124	58	;	;	PUNCT
alces-35	124	59	no	no	DET
alces-35	124	60	=	=	SYM
alces-35	124	61	northern	northern	ADJ
alces-35	124	62	area	area	NOUN
alces-35	124	63	)	)	PUNCT
alces-35	124	64	.	.	PUNCT
alces-35	125	1	the	the	DET
alces-35	125	2	sum	sum	NOUN
alces-35	125	3	of	of	ADP
alces-35	125	4	percentages	percentage	NOUN
alces-35	125	5	of	of	ADP
alces-35	125	6	new	new	ADJ
alces-35	125	7	and	and	CCONJ
alces-35	125	8	old	old	ADJ
alces-35	125	9	damage	damage	NOUN
alces-35	125	10	may	may	AUX
alces-35	125	11	exceed	exceed	VERB
alces-35	125	12	the	the	DET
alces-35	125	13	percentage	percentage	NOUN
alces-35	125	14	of	of	ADP
alces-35	125	15	total	total	ADJ
alces-35	125	16	damage	damage	NOUN
alces-35	125	17	within	within	ADP
alces-35	125	18	a	a	DET
alces-35	125	19	stand	stand	NOUN
alces-35	125	20	type	type	NOUN
alces-35	125	21	due	due	ADP
alces-35	125	22	to	to	ADP
alces-35	125	23	repeated	repeat	VERB
alces-35	125	24	browsing	browsing	NOUN
alces-35	125	25	of	of	ADP
alces-35	125	26	the	the	DET
alces-35	125	27	same	same	ADJ
alces-35	125	28	stems	stem	NOUN
alces-35	125	29	.	.	PUNCT
alces-35	126	1	0	0	NUM
alces-35	126	2	10	10	NUM
alces-35	126	3	20	20	NUM
alces-35	126	4	30	30	NUM
alces-35	126	5	40	40	NUM
alces-35	127	1	so	so	ADV
alces-35	127	2	<	<	X
alces-35	127	3	5	5	NUM
alces-35	127	4	m	m	NOUN
alces-35	127	5	so	so	ADV
alces-35	127	6	5	5	NUM
alces-35	127	7	-1	-1	SYM
alces-35	127	8	5	5	NUM
alces-35	127	9	m	m	NOUN
alces-35	127	10	so	so	ADV
alces-35	127	11	>	>	SYM
alces-35	127	12	1	1	NUM
alces-35	127	13	5	5	NUM
alces-35	127	14	m	m	NOUN
alces-35	127	15	n	n	ADV
alces-35	127	16	o	o	X
alces-35	127	17	<	<	X
alces-35	127	18	5	5	NUM
alces-35	127	19	m	m	NOUN
alces-35	127	20	n	n	NUM
alces-35	127	21	o	o	NOUN
alces-35	127	22	5	5	NUM
alces-35	127	23	-1	-1	SYM
alces-35	127	24	5	5	NUM
alces-35	127	25	m	m	NOUN
alces-35	127	26	n	n	ADV
alces-35	127	27	o	o	NOUN
alces-35	127	28	>	>	X
alces-35	127	29	1	1	NUM
alces-35	127	30	5	5	NUM
alces-35	127	31	m	m	NOUN
alces-35	127	32	%	%	NOUN
alces-35	127	33	fig	fig	NOUN
alces-35	127	34	.	.	PUNCT
alces-35	128	1	4	4	X
alces-35	128	2	.	.	X
alces-35	128	3	proportions	proportion	NOUN
alces-35	128	4	of	of	ADP
alces-35	128	5	dead	dead	ADJ
alces-35	128	6	aspens	aspen	NOUN
alces-35	128	7	in	in	ADP
alces-35	128	8	young	young	ADJ
alces-35	128	9	(	(	PUNCT
alces-35	128	10	<	<	X
alces-35	128	11	5	5	NUM
alces-35	128	12	m	m	NOUN
alces-35	128	13	in	in	ADP
alces-35	128	14	height	height	NOUN
alces-35	128	15	)	)	PUNCT
alces-35	128	16	,	,	PUNCT
alces-35	128	17	middle	middle	NOUN
alces-35	128	18	-	-	PUNCT
alces-35	128	19	aged	aged	ADJ
alces-35	128	20	(	(	PUNCT
alces-35	128	21	5	5	NUM
alces-35	128	22	-	-	SYM
alces-35	128	23	15	15	NUM
alces-35	128	24	m	m	NOUN
alces-35	128	25	in	in	ADP
alces-35	128	26	height	height	NOUN
alces-35	128	27	)	)	PUNCT
alces-35	128	28	,	,	PUNCT
alces-35	128	29	and	and	CCONJ
alces-35	128	30	old	old	ADJ
alces-35	128	31	(	(	PUNCT
alces-35	128	32	>	>	X
alces-35	128	33	15	15	NUM
alces-35	128	34	m	m	NOUN
alces-35	128	35	in	in	ADP
alces-35	128	36	height	height	NOUN
alces-35	128	37	)	)	PUNCT
alces-35	128	38	aspen	aspen	PROPN
alces-35	128	39	-	-	PUNCT
alces-35	128	40	rich	rich	ADJ
alces-35	128	41	stands	stand	VERB
alces-35	128	42	at	at	ADP
alces-35	128	43	the	the	DET
alces-35	128	44	stand	stand	NOUN
alces-35	128	45	level	level	NOUN
alces-35	128	46	in	in	ADP
alces-35	128	47	koli	koli	PROPN
alces-35	128	48	national	national	PROPN
alces-35	128	49	park	park	PROPN
alces-35	128	50	,	,	PUNCT
alces-35	128	51	finland	finland	PROPN
alces-35	128	52	(	(	PUNCT
alces-35	128	53	so	so	ADV
alces-35	128	54	=	=	SYM
alces-35	128	55	southern	southern	ADJ
alces-35	128	56	area	area	NOUN
alces-35	128	57	;	;	PUNCT
alces-35	128	58	no	no	DET
alces-35	128	59	=	=	SYM
alces-35	128	60	northern	northern	ADJ
alces-35	128	61	area	area	NOUN
alces-35	128	62	)	)	PUNCT
alces-35	128	63	.	.	PUNCT
alces-35	129	1	moose	moose	NOUN
alces-35	129	2	and	and	CCONJ
alces-35	129	3	aspen	aspen	NOUN
alces-35	129	4	regeneration	regeneration	NOUN
alces-35	129	5	in	in	ADP
alces-35	129	6	finland	finland	PROPN
alces-35	129	7	härkönen	härkönen	PROPN
alces-35	129	8	et	et	PROPN
alces-35	129	9	al	al	PROPN
alces-35	129	10	.	.	PROPN
alces-35	130	1	alces	alce	NOUN
alces-35	130	2	vol	vol	NOUN
alces-35	130	3	.	.	PROPN
alces-35	130	4	44	44	NUM
alces-35	130	5	,	,	PUNCT
alces-35	130	6	2008	2008	NUM
alces-35	130	7	36	36	NUM
alces-35	130	8	discussion	discussion	NOUN
alces-35	130	9	results	result	NOUN
alces-35	130	10	clearly	clearly	ADV
alces-35	130	11	showed	show	VERB
alces-35	130	12	that	that	SCONJ
alces-35	130	13	the	the	DET
alces-35	130	14	browsing	browse	VERB
alces-35	130	15	pressure	pressure	NOUN
alces-35	130	16	by	by	ADP
alces-35	130	17	moose	moose	NOUN
alces-35	130	18	on	on	ADP
alces-35	130	19	aspen	aspen	PROPN
alces-35	130	20	has	have	AUX
alces-35	130	21	been	be	AUX
alces-35	130	22	very	very	ADV
alces-35	130	23	intense	intense	ADJ
alces-35	130	24	in	in	ADP
alces-35	130	25	koli	koli	PROPN
alces-35	130	26	national	national	PROPN
alces-35	130	27	park	park	PROPN
alces-35	130	28	.	.	PUNCT
alces-35	131	1	at	at	ADP
alces-35	131	2	the	the	DET
alces-35	131	3	landscape	landscape	NOUN
alces-35	131	4	level	level	NOUN
alces-35	131	5	,	,	PUNCT
alces-35	131	6	moose	moose	NOUN
alces-35	131	7	had	have	AUX
alces-35	131	8	damaged	damage	VERB
alces-35	131	9	(	(	PUNCT
alces-35	131	10	twigbrowsing	twigbrowse	VERB
alces-35	131	11	,	,	PUNCT
alces-35	131	12	stem	stem	NOUN
alces-35	131	13	breakage	breakage	NOUN
alces-35	131	14	,	,	PUNCT
alces-35	131	15	or	or	CCONJ
alces-35	131	16	bark	bark	NOUN
alces-35	131	17	stripping	strip	VERB
alces-35	131	18	)	)	PUNCT
alces-35	131	19	96	96	NUM
alces-35	131	20	%	%	NOUN
alces-35	131	21	of	of	ADP
alces-35	131	22	the	the	DET
alces-35	131	23	aspens	aspen	NOUN
alces-35	131	24	in	in	ADP
alces-35	131	25	the	the	DET
alces-35	131	26	so	so	ADJ
alces-35	131	27	area	area	NOUN
alces-35	131	28	and	and	CCONJ
alces-35	131	29	62	62	NUM
alces-35	131	30	%	%	NOUN
alces-35	131	31	in	in	ADP
alces-35	131	32	the	the	DET
alces-35	131	33	no	no	DET
alces-35	131	34	area	area	NOUN
alces-35	131	35	,	,	PUNCT
alces-35	131	36	and	and	CCONJ
alces-35	131	37	the	the	DET
alces-35	131	38	moose	moose	NOUN
alces-35	131	39	damage	damage	NOUN
alces-35	131	40	at	at	ADP
alces-35	131	41	the	the	DET
alces-35	131	42	stand	stand	NOUN
alces-35	131	43	level	level	NOUN
alces-35	131	44	was	be	AUX
alces-35	131	45	79	79	NUM
alces-35	131	46	and	and	CCONJ
alces-35	131	47	73	73	NUM
alces-35	131	48	%	%	NOUN
alces-35	131	49	in	in	ADP
alces-35	131	50	the	the	DET
alces-35	131	51	no	no	NOUN
alces-35	131	52	and	and	CCONJ
alces-35	131	53	so	so	ADV
alces-35	131	54	areas	area	NOUN
alces-35	131	55	,	,	PUNCT
alces-35	131	56	respectively	respectively	ADV
alces-35	131	57	.	.	PUNCT
alces-35	132	1	these	these	DET
alces-35	132	2	damage	damage	NOUN
alces-35	132	3	levels	level	NOUN
alces-35	132	4	are	be	AUX
alces-35	132	5	more	more	ADJ
alces-35	132	6	than	than	ADP
alces-35	132	7	four	four	NUM
alces-35	132	8	-	-	ADJ
alces-35	132	9	fold	fold	VERB
alces-35	132	10	higher	high	ADJ
alces-35	132	11	than	than	ADP
alces-35	132	12	frequencies	frequency	NOUN
alces-35	132	13	observed	observe	VERB
alces-35	132	14	in	in	ADP
alces-35	132	15	old	old	ADJ
alces-35	132	16	-	-	PUNCT
alces-35	132	17	growth	growth	NOUN
alces-35	132	18	forests	forest	NOUN
alces-35	132	19	in	in	ADP
alces-35	132	20	eastern	eastern	PROPN
alces-35	132	21	finland	finland	PROPN
alces-35	132	22	.	.	PUNCT
alces-35	133	1	latva	latva	PROPN
alces-35	133	2	-	-	PUNCT
alces-35	133	3	karjanmaa	karjanmaa	VERB
alces-35	133	4	et	et	PROPN
alces-35	133	5	al	al	PROPN
alces-35	133	6	.	.	PROPN
alces-35	134	1	(	(	PUNCT
alces-35	134	2	2007	2007	NUM
alces-35	134	3	)	)	PUNCT
alces-35	134	4	reported	report	VERB
alces-35	134	5	that	that	SCONJ
alces-35	134	6	the	the	DET
alces-35	134	7	damage	damage	NOUN
alces-35	134	8	proportions	proportion	NOUN
alces-35	134	9	at	at	ADP
alces-35	134	10	the	the	DET
alces-35	134	11	stand	stand	NOUN
alces-35	134	12	level	level	NOUN
alces-35	134	13	varied	varied	ADJ
alces-35	134	14	from	from	ADP
alces-35	134	15	28	28	NUM
alces-35	134	16	-	-	SYM
alces-35	134	17	44	44	NUM
alces-35	134	18	%	%	NOUN
alces-35	134	19	in	in	ADP
alces-35	134	20	managed	manage	VERB
alces-35	134	21	forests	forest	NOUN
alces-35	134	22	in	in	ADP
alces-35	134	23	finland	finland	PROPN
alces-35	134	24	;	;	PUNCT
alces-35	134	25	similarly	similarly	ADV
alces-35	134	26	,	,	PUNCT
alces-35	134	27	härkönen	härkönen	PROPN
alces-35	134	28	(	(	PUNCT
alces-35	134	29	1998	1998	NUM
alces-35	134	30	)	)	PUNCT
alces-35	134	31	found	find	VERB
alces-35	134	32	them	they	PRON
alces-35	134	33	to	to	PART
alces-35	134	34	vary	vary	VERB
alces-35	134	35	from	from	ADP
alces-35	134	36	35	35	NUM
alces-35	134	37	-	-	SYM
alces-35	134	38	50	50	NUM
alces-35	134	39	%	%	NOUN
alces-35	134	40	.	.	PUNCT
alces-35	135	1	the	the	DET
alces-35	135	2	proportion	proportion	NOUN
alces-35	135	3	of	of	ADP
alces-35	135	4	dead	dead	ADJ
alces-35	135	5	aspen	aspen	NOUN
alces-35	135	6	further	far	ADV
alces-35	135	7	indicated	indicate	VERB
alces-35	135	8	that	that	SCONJ
alces-35	135	9	browsing	browse	VERB
alces-35	135	10	pressure	pressure	NOUN
alces-35	135	11	caused	cause	VERB
alces-35	135	12	by	by	ADP
alces-35	135	13	moose	moose	NOUN
alces-35	135	14	was	be	AUX
alces-35	135	15	high	high	ADJ
alces-35	135	16	.	.	PUNCT
alces-35	136	1	also	also	ADV
alces-35	136	2	,	,	PUNCT
alces-35	136	3	at	at	ADP
alces-35	136	4	the	the	DET
alces-35	136	5	landscape	landscape	NOUN
alces-35	136	6	level	level	NOUN
alces-35	136	7	all	all	DET
alces-35	136	8	aspens	aspen	NOUN
alces-35	136	9	in	in	ADP
alces-35	136	10	height	height	NOUN
alces-35	136	11	category	category	NOUN
alces-35	136	12	i	i	PRON
alces-35	136	13	in	in	ADP
alces-35	136	14	the	the	DET
alces-35	136	15	so	so	ADJ
alces-35	136	16	area	area	NOUN
alces-35	136	17	had	have	VERB
alces-35	136	18	signs	sign	NOUN
alces-35	136	19	of	of	ADP
alces-35	136	20	old	old	ADJ
alces-35	136	21	browsing	browsing	NOUN
alces-35	136	22	.	.	PUNCT
alces-35	137	1	bark	bark	NOUN
alces-35	137	2	stripping	strip	VERB
alces-35	137	3	in	in	ADP
alces-35	137	4	height	height	NOUN
alces-35	137	5	category	category	NOUN
alces-35	137	6	ii	ii	PROPN
alces-35	137	7	was	be	AUX
alces-35	137	8	relatively	relatively	ADV
alces-35	137	9	common	common	ADJ
alces-35	137	10	in	in	ADP
alces-35	137	11	the	the	DET
alces-35	137	12	so	so	ADV
alces-35	137	13	and	and	CCONJ
alces-35	137	14	no	no	DET
alces-35	137	15	areas	area	NOUN
alces-35	137	16	indicating	indicate	VERB
alces-35	137	17	that	that	SCONJ
alces-35	137	18	relatively	relatively	ADV
alces-35	137	19	large	large	ADJ
alces-35	137	20	aspens	aspen	NOUN
alces-35	137	21	are	be	AUX
alces-35	137	22	also	also	ADV
alces-35	137	23	susceptible	susceptible	ADJ
alces-35	137	24	to	to	ADP
alces-35	137	25	moose	moose	NOUN
alces-35	137	26	damage	damage	NOUN
alces-35	137	27	.	.	PUNCT
alces-35	138	1	at	at	ADP
alces-35	138	2	the	the	DET
alces-35	138	3	landscape	landscape	NOUN
alces-35	138	4	level	level	NOUN
alces-35	138	5	,	,	PUNCT
alces-35	138	6	23	23	NUM
alces-35	138	7	%	%	NOUN
alces-35	138	8	of	of	ADP
alces-35	138	9	the	the	DET
alces-35	138	10	damaged	damage	VERB
alces-35	138	11	aspens	aspen	NOUN
alces-35	138	12	(	(	PUNCT
alces-35	138	13	all	all	PRON
alces-35	138	14	in	in	ADP
alces-35	138	15	height	height	NOUN
alces-35	138	16	category	category	NOUN
alces-35	138	17	i	i	NOUN
alces-35	138	18	)	)	PUNCT
alces-35	138	19	were	be	AUX
alces-35	138	20	dead	dead	ADJ
alces-35	138	21	in	in	ADP
alces-35	138	22	the	the	DET
alces-35	138	23	so	so	ADJ
alces-35	138	24	area	area	NOUN
alces-35	138	25	.	.	PUNCT
alces-35	139	1	at	at	ADP
alces-35	139	2	the	the	DET
alces-35	139	3	stand	stand	NOUN
alces-35	139	4	level	level	NOUN
alces-35	139	5	,	,	PUNCT
alces-35	139	6	the	the	DET
alces-35	139	7	proportion	proportion	NOUN
alces-35	139	8	(	(	PUNCT
alces-35	139	9	35	35	NUM
alces-35	139	10	%	%	NOUN
alces-35	139	11	)	)	PUNCT
alces-35	139	12	of	of	ADP
alces-35	139	13	the	the	DET
alces-35	139	14	dead	dead	ADJ
alces-35	139	15	aspens	aspen	NOUN
alces-35	139	16	was	be	AUX
alces-35	139	17	highest	high	ADJ
alces-35	139	18	in	in	ADP
alces-35	139	19	the	the	DET
alces-35	139	20	middle	middle	NOUN
alces-35	139	21	-	-	PUNCT
alces-35	139	22	aged	aged	ADJ
alces-35	139	23	stands	stand	NOUN
alces-35	139	24	.	.	PUNCT
alces-35	140	1	these	these	DET
alces-35	140	2	results	result	NOUN
alces-35	140	3	could	could	AUX
alces-35	140	4	be	be	AUX
alces-35	140	5	explained	explain	VERB
alces-35	140	6	by	by	ADP
alces-35	140	7	cumulative	cumulative	ADJ
alces-35	140	8	stem	stem	NOUN
alces-35	140	9	breaking	break	VERB
alces-35	140	10	and	and	CCONJ
alces-35	140	11	browsing	browse	VERB
alces-35	140	12	which	which	PRON
alces-35	140	13	is	be	AUX
alces-35	140	14	concentrated	concentrate	VERB
alces-35	140	15	on	on	ADP
alces-35	140	16	leader	leader	NOUN
alces-35	140	17	shoots	shoot	NOUN
alces-35	140	18	and	and	CCONJ
alces-35	140	19	side	side	NOUN
alces-35	140	20	twigs	twig	NOUN
alces-35	140	21	of	of	ADP
alces-35	140	22	young	young	ADJ
alces-35	140	23	aspens	aspen	NOUN
alces-35	140	24	year	year	NOUN
alces-35	140	25	after	after	ADP
alces-35	140	26	year	year	NOUN
alces-35	140	27	,	,	PUNCT
alces-35	140	28	weakening	weaken	VERB
alces-35	140	29	the	the	DET
alces-35	140	30	ability	ability	NOUN
alces-35	140	31	of	of	ADP
alces-35	140	32	aspen	aspen	NOUN
alces-35	140	33	to	to	PART
alces-35	140	34	withstand	withstand	VERB
alces-35	140	35	sustained	sustain	VERB
alces-35	140	36	moose	moose	NOUN
alces-35	140	37	browsing	browse	VERB
alces-35	140	38	.	.	PUNCT
alces-35	141	1	in	in	ADP
alces-35	141	2	the	the	DET
alces-35	141	3	case	case	NOUN
alces-35	141	4	of	of	ADP
alces-35	141	5	managed	manage	VERB
alces-35	141	6	forests	forest	NOUN
alces-35	141	7	,	,	PUNCT
alces-35	141	8	edenius	edenius	NOUN
alces-35	141	9	et	et	PROPN
alces-35	141	10	al	al	PROPN
alces-35	141	11	.	.	PROPN
alces-35	142	1	(	(	PUNCT
alces-35	142	2	2002	2002	NUM
alces-35	142	3	)	)	PUNCT
alces-35	142	4	reported	report	VERB
alces-35	142	5	that	that	SCONJ
alces-35	142	6	aspens	aspen	NOUN
alces-35	142	7	growing	grow	VERB
alces-35	142	8	alone	alone	ADV
alces-35	142	9	are	be	AUX
alces-35	142	10	more	more	ADV
alces-35	142	11	susceptible	susceptible	ADJ
alces-35	142	12	to	to	ADP
alces-35	142	13	moose	moose	NOUN
alces-35	142	14	browsing	browse	VERB
alces-35	142	15	than	than	SCONJ
alces-35	142	16	aspens	aspen	NOUN
alces-35	142	17	aggregated	aggregate	VERB
alces-35	142	18	within	within	ADP
alces-35	142	19	stands	stand	NOUN
alces-35	142	20	.	.	PUNCT
alces-35	143	1	hence	hence	ADV
alces-35	143	2	,	,	PUNCT
alces-35	143	3	a	a	DET
alces-35	143	4	single	single	ADJ
alces-35	143	5	aspen	aspen	NOUN
alces-35	143	6	has	have	VERB
alces-35	143	7	the	the	DET
alces-35	143	8	best	good	ADJ
alces-35	143	9	chance	chance	NOUN
alces-35	143	10	of	of	ADP
alces-35	143	11	escaping	escape	VERB
alces-35	143	12	browsing	browse	VERB
alces-35	143	13	in	in	ADP
alces-35	143	14	a	a	DET
alces-35	143	15	stand	stand	NOUN
alces-35	143	16	with	with	ADP
alces-35	143	17	a	a	DET
alces-35	143	18	high	high	ADJ
alces-35	143	19	aspen	aspen	NOUN
alces-35	143	20	density	density	NOUN
alces-35	143	21	(	(	PUNCT
alces-35	143	22	ericsson	ericsson	PROPN
alces-35	143	23	et	et	PROPN
alces-35	143	24	al	al	PROPN
alces-35	143	25	.	.	PROPN
alces-35	143	26	2001	2001	NUM
alces-35	143	27	)	)	PUNCT
alces-35	143	28	.	.	PUNCT
alces-35	144	1	this	this	DET
alces-35	144	2	observation	observation	NOUN
alces-35	144	3	corresponds	correspond	VERB
alces-35	144	4	to	to	ADP
alces-35	144	5	our	our	PRON
alces-35	144	6	findings	finding	NOUN
alces-35	144	7	because	because	SCONJ
alces-35	144	8	overall	overall	ADJ
alces-35	144	9	proportion	proportion	NOUN
alces-35	144	10	of	of	ADP
alces-35	144	11	damaged	damage	VERB
alces-35	144	12	aspens	aspen	NOUN
alces-35	144	13	was	be	AUX
alces-35	144	14	higher	high	ADJ
alces-35	144	15	at	at	ADP
alces-35	144	16	the	the	DET
alces-35	144	17	landscape	landscape	NOUN
alces-35	144	18	level	level	NOUN
alces-35	144	19	(	(	PUNCT
alces-35	144	20	low	low	ADJ
alces-35	144	21	aspen	aspen	ADJ
alces-35	144	22	densities	density	NOUN
alces-35	144	23	)	)	PUNCT
alces-35	144	24	than	than	ADP
alces-35	144	25	in	in	ADP
alces-35	144	26	the	the	DET
alces-35	144	27	stand	stand	NOUN
alces-35	144	28	level	level	NOUN
alces-35	144	29	(	(	PUNCT
alces-35	144	30	high	high	ADJ
alces-35	144	31	aspen	aspen	ADJ
alces-35	144	32	densities	density	NOUN
alces-35	144	33	)	)	PUNCT
alces-35	144	34	.	.	PUNCT
alces-35	145	1	thus	thus	ADV
alces-35	145	2	,	,	PUNCT
alces-35	145	3	moose	moose	NOUN
alces-35	145	4	may	may	AUX
alces-35	145	5	reinforce	reinforce	VERB
alces-35	145	6	the	the	DET
alces-35	145	7	spatially	spatially	ADV
alces-35	145	8	aggregated	aggregate	VERB
alces-35	145	9	distribution	distribution	NOUN
alces-35	145	10	of	of	ADP
alces-35	145	11	aspen	aspen	NOUN
alces-35	145	12	both	both	PRON
alces-35	145	13	in	in	ADP
alces-35	145	14	managed	manage	VERB
alces-35	145	15	and	and	CCONJ
alces-35	145	16	protected	protect	VERB
alces-35	145	17	boreal	boreal	ADJ
alces-35	145	18	forest	forest	NOUN
alces-35	145	19	landscapes	landscape	NOUN
alces-35	145	20	.	.	PUNCT
alces-35	146	1	that	that	DET
alces-35	146	2	aspen	aspen	ADJ
alces-35	146	3	density	density	NOUN
alces-35	146	4	decreased	decrease	VERB
alces-35	146	5	from	from	ADP
alces-35	146	6	young	young	ADJ
alces-35	146	7	to	to	ADP
alces-35	146	8	old	old	ADJ
alces-35	146	9	stands	stand	NOUN
alces-35	146	10	,	,	PUNCT
alces-35	146	11	that	that	SCONJ
alces-35	146	12	low	low	ADJ
alces-35	146	13	aspen	aspen	NOUN
alces-35	146	14	density	density	NOUN
alces-35	146	15	occurred	occur	VERB
alces-35	146	16	in	in	ADP
alces-35	146	17	old	old	ADJ
alces-35	146	18	stands	stand	NOUN
alces-35	146	19	at	at	ADP
alces-35	146	20	the	the	DET
alces-35	146	21	stand	stand	NOUN
alces-35	146	22	level	level	NOUN
alces-35	146	23	,	,	PUNCT
alces-35	146	24	and	and	CCONJ
alces-35	146	25	that	that	SCONJ
alces-35	146	26	low	low	ADJ
alces-35	146	27	overall	overall	ADJ
alces-35	146	28	aspen	aspen	ADJ
alces-35	146	29	density	density	NOUN
alces-35	146	30	occurred	occur	VERB
alces-35	146	31	at	at	ADP
alces-35	146	32	the	the	DET
alces-35	146	33	landscape	landscape	NOUN
alces-35	146	34	level	level	NOUN
alces-35	146	35	were	be	AUX
alces-35	146	36	all	all	PRON
alces-35	146	37	expected	expect	VERB
alces-35	146	38	results	result	NOUN
alces-35	146	39	.	.	PUNCT
alces-35	147	1	however	however	ADV
alces-35	147	2	,	,	PUNCT
alces-35	147	3	the	the	DET
alces-35	147	4	observed	observed	ADJ
alces-35	147	5	densities	density	NOUN
alces-35	147	6	were	be	AUX
alces-35	147	7	relatively	relatively	ADV
alces-35	147	8	high	high	ADJ
alces-35	147	9	in	in	ADP
alces-35	147	10	comparison	comparison	NOUN
alces-35	147	11	to	to	ADP
alces-35	147	12	those	those	PRON
alces-35	147	13	found	find	VERB
alces-35	147	14	at	at	ADP
alces-35	147	15	other	other	ADJ
alces-35	147	16	conservation	conservation	NOUN
alces-35	147	17	areas	area	NOUN
alces-35	147	18	in	in	ADP
alces-35	147	19	eastern	eastern	PROPN
alces-35	147	20	finland	finland	PROPN
alces-35	147	21	where	where	SCONJ
alces-35	147	22	aspen	aspen	ADJ
alces-35	147	23	densities	density	NOUN
alces-35	147	24	have	have	AUX
alces-35	147	25	been	be	AUX
alces-35	147	26	as	as	ADV
alces-35	147	27	low	low	ADJ
alces-35	147	28	as	as	ADP
alces-35	147	29	1	1	NUM
alces-35	147	30	tree	tree	NOUN
alces-35	147	31	/	/	SYM
alces-35	147	32	hectare	hectare	NOUN
alces-35	147	33	(	(	PUNCT
alces-35	147	34	kouki	kouki	PROPN
alces-35	147	35	et	et	PROPN
alces-35	147	36	al	al	PROPN
alces-35	147	37	.	.	PROPN
alces-35	147	38	2004	2004	NUM
alces-35	147	39	)	)	PUNCT
alces-35	147	40	.	.	PUNCT
alces-35	148	1	nfi	nfi	PROPN
alces-35	148	2	results	result	NOUN
alces-35	148	3	also	also	ADV
alces-35	148	4	indicate	indicate	VERB
alces-35	148	5	the	the	DET
alces-35	148	6	relative	relative	ADJ
alces-35	148	7	rareness	rareness	NOUN
alces-35	148	8	of	of	ADP
alces-35	148	9	mature	mature	ADJ
alces-35	148	10	aspen	aspen	NOUN
alces-35	148	11	trees	tree	NOUN
alces-35	148	12	in	in	ADP
alces-35	148	13	finnish	finnish	ADJ
alces-35	148	14	forests	forest	NOUN
alces-35	148	15	(	(	PUNCT
alces-35	148	16	finnish	finnish	PROPN
alces-35	148	17	forest	forest	PROPN
alces-35	148	18	research	research	PROPN
alces-35	148	19	institute	institute	PROPN
alces-35	148	20	2006	2006	NUM
alces-35	148	21	)	)	PUNCT
alces-35	148	22	.	.	PUNCT
alces-35	149	1	in	in	ADP
alces-35	149	2	young	young	ADJ
alces-35	149	3	stands	stand	NOUN
alces-35	149	4	,	,	PUNCT
alces-35	149	5	aspen	aspen	ADJ
alces-35	149	6	densities	density	NOUN
alces-35	149	7	(	(	PUNCT
alces-35	149	8	2,300	2,300	NUM
alces-35	149	9	-	-	SYM
alces-35	149	10	4,500	4,500	NUM
alces-35	149	11	trees	tree	NOUN
alces-35	149	12	/	/	SYM
alces-35	149	13	ha	ha	CCONJ
alces-35	149	14	)	)	PUNCT
alces-35	149	15	were	be	AUX
alces-35	149	16	comparable	comparable	ADJ
alces-35	149	17	to	to	ADP
alces-35	149	18	those	those	PRON
alces-35	149	19	in	in	ADP
alces-35	149	20	conservation	conservation	NOUN
alces-35	149	21	areas	area	NOUN
alces-35	149	22	measured	measure	VERB
alces-35	149	23	by	by	ADP
alces-35	149	24	heikkilä	heikkilä	PROPN
alces-35	149	25	et	et	PROPN
alces-35	149	26	al	al	PROPN
alces-35	149	27	.	.	PROPN
alces-35	150	1	(	(	PUNCT
alces-35	150	2	2003	2003	NUM
alces-35	150	3	)	)	PUNCT
alces-35	150	4	.	.	PUNCT
alces-35	151	1	stand	stand	VERB
alces-35	151	2	level	level	NOUN
alces-35	151	3	volumes	volume	NOUN
alces-35	151	4	of	of	ADP
alces-35	151	5	aspen	aspen	PROPN
alces-35	151	6	were	be	AUX
alces-35	151	7	not	not	PART
alces-35	151	8	of	of	ADP
alces-35	151	9	primary	primary	ADJ
alces-35	151	10	interest	interest	NOUN
alces-35	151	11	in	in	ADP
alces-35	151	12	this	this	DET
alces-35	151	13	study	study	NOUN
alces-35	151	14	and	and	CCONJ
alces-35	151	15	they	they	PRON
alces-35	151	16	were	be	AUX
alces-35	151	17	not	not	PART
alces-35	151	18	calculated	calculate	VERB
alces-35	151	19	,	,	PUNCT
alces-35	151	20	but	but	CCONJ
alces-35	151	21	it	it	PRON
alces-35	151	22	has	have	AUX
alces-35	151	23	been	be	AUX
alces-35	151	24	estimated	estimate	VERB
alces-35	151	25	that	that	SCONJ
alces-35	151	26	an	an	DET
alces-35	151	27	average	average	ADJ
alces-35	151	28	volume	volume	NOUN
alces-35	151	29	of	of	ADP
alces-35	151	30	5	5	NUM
alces-35	151	31	-	-	SYM
alces-35	151	32	20	20	NUM
alces-35	151	33	m3	m3	PROPN
alces-35	151	34	/	/	SYM
alces-35	151	35	ha	ha	PROPN
alces-35	151	36	is	be	AUX
alces-35	151	37	typical	typical	ADJ
alces-35	151	38	for	for	ADP
alces-35	151	39	aspen	aspen	NOUN
alces-35	151	40	in	in	ADP
alces-35	151	41	norway	norway	PROPN
alces-35	151	42	spruce	spruce	NOUN
alces-35	151	43	-	-	PUNCT
alces-35	151	44	dominated	dominate	VERB
alces-35	151	45	,	,	PUNCT
alces-35	151	46	old	old	ADJ
alces-35	151	47	-	-	PUNCT
alces-35	151	48	growth	growth	NOUN
alces-35	151	49	forests	forest	NOUN
alces-35	151	50	in	in	ADP
alces-35	151	51	southern	southern	ADJ
alces-35	151	52	and	and	CCONJ
alces-35	151	53	middle	middle	ADJ
alces-35	151	54	boreal	boreal	ADJ
alces-35	151	55	fennoscandia	fennoscandia	NOUN
alces-35	151	56	(	(	PUNCT
alces-35	151	57	latva	latva	NOUN
alces-35	151	58	-	-	PUNCT
alces-35	151	59	karjanmaa	karjanmaa	VERB
alces-35	151	60	et	et	PROPN
alces-35	151	61	al	al	PROPN
alces-35	151	62	.	.	PROPN
alces-35	151	63	2007	2007	NUM
alces-35	151	64	)	)	PUNCT
alces-35	151	65	.	.	PUNCT
alces-35	152	1	pellet	pellet	NOUN
alces-35	152	2	group	group	NOUN
alces-35	152	3	counts	count	NOUN
alces-35	152	4	were	be	AUX
alces-35	152	5	used	use	VERB
alces-35	152	6	to	to	PART
alces-35	152	7	estimate	estimate	VERB
alces-35	152	8	moose	moose	NOUN
alces-35	152	9	activity	activity	NOUN
alces-35	152	10	in	in	ADP
alces-35	152	11	the	the	DET
alces-35	152	12	area	area	NOUN
alces-35	152	13	.	.	PUNCT
alces-35	153	1	according	accord	VERB
alces-35	153	2	to	to	ADP
alces-35	153	3	neff	neff	PROPN
alces-35	153	4	(	(	PUNCT
alces-35	153	5	1968	1968	NUM
alces-35	153	6	)	)	PUNCT
alces-35	153	7	,	,	PUNCT
alces-35	153	8	the	the	DET
alces-35	153	9	method	method	NOUN
alces-35	153	10	can	can	AUX
alces-35	153	11	provide	provide	VERB
alces-35	153	12	reliable	reliable	ADJ
alces-35	153	13	data	datum	NOUN
alces-35	153	14	under	under	ADP
alces-35	153	15	most	most	ADJ
alces-35	153	16	field	field	NOUN
alces-35	153	17	conditions	condition	NOUN
alces-35	153	18	.	.	PUNCT
alces-35	154	1	in	in	ADP
alces-35	154	2	this	this	DET
alces-35	154	3	study	study	NOUN
alces-35	154	4	,	,	PUNCT
alces-35	154	5	the	the	DET
alces-35	154	6	observed	observe	VERB
alces-35	154	7	numbers	number	NOUN
alces-35	154	8	of	of	ADP
alces-35	154	9	pellet	pellet	NOUN
alces-35	154	10	groups	group	NOUN
alces-35	154	11	were	be	AUX
alces-35	154	12	surprisingly	surprisingly	ADV
alces-35	154	13	low	low	ADJ
alces-35	154	14	when	when	SCONJ
alces-35	154	15	compared	compare	VERB
alces-35	154	16	to	to	ADP
alces-35	154	17	moose	moose	NOUN
alces-35	154	18	browsing	browse	VERB
alces-35	154	19	pressure	pressure	NOUN
alces-35	154	20	on	on	ADP
alces-35	154	21	aspens	aspen	NOUN
alces-35	154	22	and	and	CCONJ
alces-35	154	23	the	the	DET
alces-35	154	24	estimated	estimate	VERB
alces-35	154	25	moose	moose	NOUN
alces-35	154	26	density	density	NOUN
alces-35	154	27	in	in	ADP
alces-35	154	28	the	the	DET
alces-35	154	29	area	area	NOUN
alces-35	154	30	,	,	PUNCT
alces-35	154	31	and	and	CCONJ
alces-35	154	32	the	the	DET
alces-35	154	33	numbers	number	NOUN
alces-35	154	34	reported	report	VERB
alces-35	154	35	from	from	ADP
alces-35	154	36	managed	manage	VERB
alces-35	154	37	scots	scot	NOUN
alces-35	154	38	pine	pine	NOUN
alces-35	154	39	-	-	PUNCT
alces-35	154	40	dominated	dominate	VERB
alces-35	154	41	forests	forest	NOUN
alces-35	154	42	in	in	ADP
alces-35	154	43	finland	finland	PROPN
alces-35	154	44	(	(	PUNCT
alces-35	154	45	heikkilä	heikkilä	NOUN
alces-35	154	46	and	and	CCONJ
alces-35	154	47	härkönen	härkönen	NOUN
alces-35	154	48	1998	1998	NUM
alces-35	154	49	)	)	PUNCT
alces-35	154	50	.	.	PUNCT
alces-35	155	1	this	this	PRON
alces-35	155	2	may	may	AUX
alces-35	155	3	be	be	AUX
alces-35	155	4	explained	explain	VERB
alces-35	155	5	by	by	ADP
alces-35	155	6	overestimation	overestimation	NOUN
alces-35	155	7	of	of	ADP
alces-35	155	8	moose	moose	NOUN
alces-35	155	9	density	density	NOUN
alces-35	155	10	in	in	ADP
alces-35	155	11	the	the	DET
alces-35	155	12	area	area	NOUN
alces-35	155	13	or	or	CCONJ
alces-35	155	14	by	by	ADP
alces-35	155	15	difficulties	difficulty	NOUN
alces-35	155	16	in	in	ADP
alces-35	155	17	separating	separate	VERB
alces-35	155	18	old	old	ADJ
alces-35	155	19	pellet	pellet	NOUN
alces-35	155	20	groups	group	NOUN
alces-35	155	21	from	from	ADP
alces-35	155	22	new	new	ADJ
alces-35	155	23	pellet	pellet	NOUN
alces-35	155	24	groups	group	NOUN
alces-35	155	25	during	during	ADP
alces-35	155	26	fieldwork	fieldwork	NOUN
alces-35	155	27	(	(	PUNCT
alces-35	155	28	neff	neff	PROPN
alces-35	155	29	1968	1968	NUM
alces-35	155	30	,	,	PUNCT
alces-35	155	31	härkönen	härkönen	NOUN
alces-35	155	32	and	and	CCONJ
alces-35	155	33	heikkilä	heikkilä	NOUN
alces-35	155	34	1999	1999	NUM
alces-35	155	35	)	)	PUNCT
alces-35	155	36	.	.	PUNCT
alces-35	156	1	however	however	ADV
alces-35	156	2	,	,	PUNCT
alces-35	156	3	the	the	DET
alces-35	156	4	pellet	pellet	NOUN
alces-35	156	5	group	group	NOUN
alces-35	156	6	counts	count	NOUN
alces-35	156	7	were	be	AUX
alces-35	156	8	in	in	ADP
alces-35	156	9	line	line	NOUN
alces-35	156	10	with	with	ADP
alces-35	156	11	browsing	browse	VERB
alces-35	156	12	intensity	intensity	NOUN
alces-35	156	13	at	at	ADP
alces-35	156	14	the	the	DET
alces-35	156	15	landscape	landscape	NOUN
alces-35	156	16	level	level	NOUN
alces-35	156	17	as	as	SCONJ
alces-35	156	18	there	there	PRON
alces-35	156	19	were	be	VERB
alces-35	156	20	more	more	ADJ
alces-35	156	21	pellet	pellet	NOUN
alces-35	156	22	groups	group	NOUN
alces-35	156	23	in	in	ADP
alces-35	156	24	the	the	DET
alces-35	156	25	so	so	ADV
alces-35	156	26	than	than	ADP
alces-35	156	27	no	no	DET
alces-35	156	28	area	area	NOUN
alces-35	156	29	.	.	PUNCT
alces-35	157	1	also	also	ADV
alces-35	157	2	,	,	PUNCT
alces-35	157	3	the	the	DET
alces-35	157	4	proportion	proportion	NOUN
alces-35	157	5	of	of	ADP
alces-35	157	6	damaged	damage	VERB
alces-35	157	7	aspens	aspen	NOUN
alces-35	157	8	was	be	AUX
alces-35	157	9	higher	high	ADJ
alces-35	157	10	in	in	ADP
alces-35	157	11	the	the	DET
alces-35	157	12	so	so	ADJ
alces-35	157	13	area	area	NOUN
alces-35	157	14	.	.	PUNCT
alces-35	158	1	at	at	ADP
alces-35	158	2	the	the	DET
alces-35	158	3	stand	stand	NOUN
alces-35	158	4	level	level	NOUN
alces-35	158	5	,	,	PUNCT
alces-35	158	6	the	the	DET
alces-35	158	7	number	number	NOUN
alces-35	158	8	alces	alce	VERB
alces-35	158	9	vol	vol	NOUN
alces-35	158	10	.	.	PROPN
alces-35	158	11	44	44	NUM
alces-35	158	12	,	,	PUNCT
alces-35	158	13	2008	2008	NUM
alces-35	158	14	härkönen	härkönen	NOUN
alces-35	158	15	et	et	PROPN
alces-35	158	16	al.moose	al.moose	X
alces-35	158	17	and	and	CCONJ
alces-35	158	18	aspen	aspen	VERB
alces-35	158	19	regeneration	regeneration	NOUN
alces-35	158	20	in	in	ADP
alces-35	158	21	finland	finland	PROPN
alces-35	158	22	37	37	NUM
alces-35	158	23	of	of	ADP
alces-35	158	24	pellet	pellet	NOUN
alces-35	158	25	groups	group	NOUN
alces-35	158	26	was	be	AUX
alces-35	158	27	very	very	ADV
alces-35	158	28	low	low	ADJ
alces-35	158	29	,	,	PUNCT
alces-35	158	30	whereas	whereas	SCONJ
alces-35	158	31	the	the	DET
alces-35	158	32	browsing	browse	VERB
alces-35	158	33	intensity	intensity	NOUN
alces-35	158	34	was	be	AUX
alces-35	158	35	relatively	relatively	ADV
alces-35	158	36	high	high	ADJ
alces-35	158	37	.	.	PUNCT
alces-35	159	1	this	this	DET
alces-35	159	2	weak	weak	ADJ
alces-35	159	3	relationship	relationship	NOUN
alces-35	159	4	between	between	ADP
alces-35	159	5	browsing	browse	VERB
alces-35	159	6	incidence	incidence	NOUN
alces-35	159	7	on	on	ADP
alces-35	159	8	aspen	aspen	NOUN
alces-35	159	9	and	and	CCONJ
alces-35	159	10	pellet	pellet	NOUN
alces-35	159	11	counts	count	NOUN
alces-35	159	12	was	be	AUX
alces-35	159	13	also	also	ADV
alces-35	159	14	observed	observe	VERB
alces-35	159	15	in	in	ADP
alces-35	159	16	sweden	sweden	PROPN
alces-35	159	17	(	(	PUNCT
alces-35	159	18	edenius	edenius	NOUN
alces-35	159	19	and	and	CCONJ
alces-35	159	20	ericsson	ericsson	NOUN
alces-35	159	21	2007	2007	NUM
alces-35	159	22	)	)	PUNCT
alces-35	159	23	.	.	PUNCT
alces-35	160	1	it	it	PRON
alces-35	160	2	is	be	AUX
alces-35	160	3	evident	evident	ADJ
alces-35	160	4	that	that	SCONJ
alces-35	160	5	norway	norway	PROPN
alces-35	160	6	spruce	spruce	NOUN
alces-35	160	7	-	-	PUNCT
alces-35	160	8	dominated	dominate	VERB
alces-35	160	9	forest	forest	NOUN
alces-35	160	10	sites	site	NOUN
alces-35	160	11	,	,	PUNCT
alces-35	160	12	which	which	PRON
alces-35	160	13	are	be	AUX
alces-35	160	14	common	common	ADJ
alces-35	160	15	in	in	ADP
alces-35	160	16	koli	koli	PROPN
alces-35	160	17	national	national	PROPN
alces-35	160	18	park	park	PROPN
alces-35	160	19	,	,	PUNCT
alces-35	160	20	are	be	AUX
alces-35	160	21	less	less	ADV
alces-35	160	22	favorable	favorable	ADJ
alces-35	160	23	for	for	ADP
alces-35	160	24	moose	moose	NOUN
alces-35	160	25	because	because	SCONJ
alces-35	160	26	overall	overall	ADJ
alces-35	160	27	food	food	NOUN
alces-35	160	28	availability	availability	NOUN
alces-35	160	29	is	be	AUX
alces-35	160	30	low	low	ADJ
alces-35	160	31	in	in	ADP
alces-35	160	32	these	these	DET
alces-35	160	33	closed	close	VERB
alces-35	160	34	middle	middle	NOUN
alces-35	160	35	-	-	PUNCT
alces-35	160	36	aged	aged	ADJ
alces-35	160	37	or	or	CCONJ
alces-35	160	38	older	old	ADJ
alces-35	160	39	stands	stand	NOUN
alces-35	160	40	.	.	PUNCT
alces-35	161	1	in	in	ADP
alces-35	161	2	this	this	DET
alces-35	161	3	sense	sense	NOUN
alces-35	161	4	,	,	PUNCT
alces-35	161	5	small	small	ADJ
alces-35	161	6	aggregated	aggregate	VERB
alces-35	161	7	patches	patch	NOUN
alces-35	161	8	of	of	ADP
alces-35	161	9	young	young	ADJ
alces-35	161	10	aspens	aspen	NOUN
alces-35	161	11	may	may	AUX
alces-35	161	12	provide	provide	VERB
alces-35	161	13	so	so	ADV
alces-35	161	14	little	little	ADJ
alces-35	161	15	food	food	NOUN
alces-35	161	16	that	that	PRON
alces-35	161	17	moose	moose	VERB
alces-35	161	18	only	only	ADV
alces-35	161	19	visit	visit	VERB
alces-35	161	20	them	they	PRON
alces-35	161	21	briefly	briefly	ADV
alces-35	161	22	during	during	ADP
alces-35	161	23	a	a	DET
alces-35	161	24	foraging	forage	VERB
alces-35	161	25	bout	bout	NOUN
alces-35	161	26	.	.	PUNCT
alces-35	162	1	the	the	DET
alces-35	162	2	impact	impact	NOUN
alces-35	162	3	of	of	ADP
alces-35	162	4	ungulate	ungulate	ADJ
alces-35	162	5	populations	population	NOUN
alces-35	162	6	on	on	ADP
alces-35	162	7	regeneration	regeneration	NOUN
alces-35	162	8	of	of	ADP
alces-35	162	9	aspen	aspen	NOUN
alces-35	162	10	has	have	AUX
alces-35	162	11	been	be	AUX
alces-35	162	12	questioned	question	VERB
alces-35	162	13	globally	globally	ADV
alces-35	162	14	,	,	PUNCT
alces-35	162	15	for	for	ADP
alces-35	162	16	example	example	NOUN
alces-35	162	17	,	,	PUNCT
alces-35	162	18	in	in	ADP
alces-35	162	19	the	the	DET
alces-35	162	20	rocky	rocky	ADJ
alces-35	162	21	mountain	mountain	NOUN
alces-35	162	22	region	region	NOUN
alces-35	162	23	in	in	ADP
alces-35	162	24	western	western	PROPN
alces-35	162	25	north	north	PROPN
alces-35	162	26	america	america	PROPN
alces-35	162	27	(	(	PUNCT
alces-35	162	28	kay	kay	PROPN
alces-35	162	29	1997	1997	NUM
alces-35	162	30	,	,	PUNCT
alces-35	162	31	suzuki	suzuki	PROPN
alces-35	162	32	et	et	PROPN
alces-35	162	33	al	al	PROPN
alces-35	162	34	.	.	PROPN
alces-35	162	35	1999	1999	NUM
alces-35	162	36	)	)	PUNCT
alces-35	162	37	and	and	CCONJ
alces-35	162	38	in	in	ADP
alces-35	162	39	finland	finland	PROPN
alces-35	162	40	(	(	PUNCT
alces-35	162	41	kouki	kouki	PROPN
alces-35	162	42	et	et	PROPN
alces-35	162	43	al	al	PROPN
alces-35	162	44	.	.	PROPN
alces-35	162	45	2004	2004	NUM
alces-35	162	46	)	)	PUNCT
alces-35	162	47	.	.	PUNCT
alces-35	163	1	however	however	ADV
alces-35	163	2	,	,	PUNCT
alces-35	163	3	moose	moose	NOUN
alces-35	163	4	or	or	CCONJ
alces-35	163	5	deer	deer	NOUN
alces-35	163	6	browsing	browse	VERB
alces-35	163	7	is	be	AUX
alces-35	163	8	not	not	PART
alces-35	163	9	the	the	DET
alces-35	163	10	only	only	ADJ
alces-35	163	11	factor	factor	NOUN
alces-35	163	12	behind	behind	ADP
alces-35	163	13	the	the	DET
alces-35	163	14	poor	poor	ADJ
alces-35	163	15	recruitment	recruitment	NOUN
alces-35	163	16	of	of	ADP
alces-35	163	17	young	young	ADJ
alces-35	163	18	aspen	aspen	NOUN
alces-35	163	19	cohorts	cohort	NOUN
alces-35	163	20	in	in	ADP
alces-35	163	21	conservation	conservation	NOUN
alces-35	163	22	areas	area	NOUN
alces-35	163	23	.	.	PUNCT
alces-35	164	1	absence	absence	NOUN
alces-35	164	2	of	of	ADP
alces-35	164	3	large	large	ADJ
alces-35	164	4	scale	scale	NOUN
alces-35	164	5	disturbances	disturbance	NOUN
alces-35	164	6	(	(	PUNCT
alces-35	164	7	e.g.	e.g.	ADV
alces-35	164	8	,	,	PUNCT
alces-35	164	9	fires	fire	NOUN
alces-35	164	10	and	and	CCONJ
alces-35	164	11	windstorms	windstorm	NOUN
alces-35	164	12	)	)	PUNCT
alces-35	164	13	that	that	PRON
alces-35	164	14	create	create	VERB
alces-35	164	15	forest	forest	NOUN
alces-35	164	16	openings	opening	NOUN
alces-35	164	17	in	in	ADP
alces-35	164	18	conservation	conservation	NOUN
alces-35	164	19	areas	area	NOUN
alces-35	164	20	strongly	strongly	ADV
alces-35	164	21	hinder	hinder	VERB
alces-35	164	22	the	the	DET
alces-35	164	23	possibility	possibility	NOUN
alces-35	164	24	of	of	ADP
alces-35	164	25	successful	successful	ADJ
alces-35	164	26	regeneration	regeneration	NOUN
alces-35	164	27	(	(	PUNCT
alces-35	164	28	latva	latva	NOUN
alces-35	164	29	-	-	PUNCT
alces-35	164	30	karjanmaa	karjanmaa	VERB
alces-35	164	31	et	et	PROPN
alces-35	164	32	al	al	PROPN
alces-35	164	33	.	.	PROPN
alces-35	164	34	2007	2007	NUM
alces-35	164	35	)	)	PUNCT
alces-35	164	36	because	because	SCONJ
alces-35	164	37	some	some	DET
alces-35	164	38	disturbance	disturbance	NOUN
alces-35	164	39	is	be	AUX
alces-35	164	40	needed	need	VERB
alces-35	164	41	before	before	SCONJ
alces-35	164	42	aspen	aspen	NOUN
alces-35	164	43	regeneration	regeneration	NOUN
alces-35	164	44	starts	start	VERB
alces-35	164	45	from	from	ADP
alces-35	164	46	root	root	NOUN
alces-35	164	47	suckers	sucker	NOUN
alces-35	164	48	.	.	PUNCT
alces-35	165	1	in	in	ADP
alces-35	165	2	addition	addition	NOUN
alces-35	165	3	,	,	PUNCT
alces-35	165	4	young	young	ADJ
alces-35	165	5	aspen	aspen	NOUN
alces-35	165	6	seedlings	seedling	NOUN
alces-35	165	7	and	and	CCONJ
alces-35	165	8	saplings	sapling	NOUN
alces-35	165	9	are	be	AUX
alces-35	165	10	browsed	browse	VERB
alces-35	165	11	by	by	ADP
alces-35	165	12	voles	vole	NOUN
alces-35	165	13	,	,	PUNCT
alces-35	165	14	mountain	mountain	NOUN
alces-35	165	15	hare	hare	NOUN
alces-35	165	16	(	(	PUNCT
alces-35	165	17	lepus	lepus	PROPN
alces-35	165	18	timidus	timidus	PROPN
alces-35	165	19	)	)	PUNCT
alces-35	165	20	,	,	PUNCT
alces-35	165	21	and	and	CCONJ
alces-35	165	22	beavers	beaver	NOUN
alces-35	165	23	(	(	PUNCT
alces-35	165	24	castor	castor	NOUN
alces-35	165	25	spp	spp	NOUN
alces-35	165	26	.	.	PUNCT
alces-35	165	27	)	)	PUNCT
alces-35	165	28	.	.	PUNCT
alces-35	166	1	the	the	DET
alces-35	166	2	impact	impact	NOUN
alces-35	166	3	of	of	ADP
alces-35	166	4	these	these	DET
alces-35	166	5	browsers	browser	NOUN
alces-35	166	6	on	on	ADP
alces-35	166	7	aspen	aspen	NOUN
alces-35	166	8	mortality	mortality	NOUN
alces-35	166	9	might	might	AUX
alces-35	166	10	have	have	AUX
alces-35	166	11	been	be	AUX
alces-35	166	12	underestimated	underestimate	VERB
alces-35	166	13	in	in	ADP
alces-35	166	14	comparison	comparison	NOUN
alces-35	166	15	to	to	ADP
alces-35	166	16	moose	moose	NOUN
alces-35	166	17	browsing	browse	VERB
alces-35	166	18	.	.	PUNCT
alces-35	167	1	disease	disease	NOUN
alces-35	167	2	,	,	PUNCT
alces-35	167	3	including	include	VERB
alces-35	167	4	“	"	PUNCT
alces-35	167	5	black	black	ADJ
alces-35	167	6	shoot	shoot	NOUN
alces-35	167	7	blight	blight	NOUN
alces-35	167	8	”	"	PUNCT
alces-35	167	9	which	which	PRON
alces-35	167	10	is	be	AUX
alces-35	167	11	caused	cause	VERB
alces-35	167	12	by	by	ADP
alces-35	167	13	a	a	DET
alces-35	167	14	fungus	fungus	NOUN
alces-35	167	15	(	(	PUNCT
alces-35	167	16	venturia	venturia	PROPN
alces-35	167	17	tremulae	tremulae	NOUN
alces-35	167	18	)	)	PUNCT
alces-35	167	19	and	and	CCONJ
alces-35	167	20	kills	kill	VERB
alces-35	167	21	the	the	DET
alces-35	167	22	leader	leader	NOUN
alces-35	167	23	shoot	shoot	NOUN
alces-35	167	24	and	and	CCONJ
alces-35	167	25	leaves	leave	NOUN
alces-35	167	26	,	,	PUNCT
alces-35	167	27	may	may	AUX
alces-35	167	28	stunt	stunt	VERB
alces-35	167	29	the	the	DET
alces-35	167	30	height	height	NOUN
alces-35	167	31	development	development	NOUN
alces-35	167	32	of	of	ADP
alces-35	167	33	young	young	ADJ
alces-35	167	34	aspens	aspen	NOUN
alces-35	167	35	in	in	ADP
alces-35	167	36	conservation	conservation	NOUN
alces-35	167	37	areas	area	NOUN
alces-35	167	38	(	(	PUNCT
alces-35	167	39	heikkilä	heikkilä	VERB
alces-35	167	40	et	et	PROPN
alces-35	167	41	al	al	PROPN
alces-35	167	42	.	.	PROPN
alces-35	167	43	2003	2003	NUM
alces-35	167	44	)	)	PUNCT
alces-35	167	45	.	.	PUNCT
alces-35	168	1	this	this	DET
alces-35	168	2	disease	disease	NOUN
alces-35	168	3	causes	cause	VERB
alces-35	168	4	infected	infected	ADJ
alces-35	168	5	aspens	aspen	NOUN
alces-35	168	6	to	to	PART
alces-35	168	7	be	be	AUX
alces-35	168	8	available	available	ADJ
alces-35	168	9	longer	long	ADV
alces-35	168	10	as	as	ADP
alces-35	168	11	potential	potential	ADJ
alces-35	168	12	forage	forage	NOUN
alces-35	168	13	for	for	ADP
alces-35	168	14	moose	moose	NOUN
alces-35	168	15	or	or	CCONJ
alces-35	168	16	other	other	ADJ
alces-35	168	17	browsers	browser	NOUN
alces-35	168	18	,	,	PUNCT
alces-35	168	19	thereby	thereby	ADV
alces-35	168	20	increasing	increase	VERB
alces-35	168	21	the	the	DET
alces-35	168	22	risk	risk	NOUN
alces-35	168	23	that	that	PRON
alces-35	168	24	damaged	damage	VERB
alces-35	168	25	aspens	aspen	NOUN
alces-35	168	26	die	die	VERB
alces-35	168	27	due	due	ADJ
alces-35	168	28	to	to	ADP
alces-35	168	29	repetitive	repetitive	ADJ
alces-35	168	30	browsing	browsing	NOUN
alces-35	168	31	.	.	PUNCT
alces-35	169	1	in	in	ADP
alces-35	169	2	addition	addition	NOUN
alces-35	169	3	,	,	PUNCT
alces-35	169	4	young	young	ADJ
alces-35	169	5	aspen	aspen	NOUN
alces-35	169	6	seedlings	seedling	NOUN
alces-35	169	7	may	may	AUX
alces-35	169	8	also	also	ADV
alces-35	169	9	die	die	VERB
alces-35	169	10	because	because	SCONJ
alces-35	169	11	they	they	PRON
alces-35	169	12	can	can	AUX
alces-35	169	13	not	not	PART
alces-35	169	14	tolerate	tolerate	VERB
alces-35	169	15	local	local	ADJ
alces-35	169	16	climate	climate	NOUN
alces-35	169	17	and	and	CCONJ
alces-35	169	18	soil	soil	NOUN
alces-35	169	19	conditions	condition	NOUN
alces-35	169	20	(	(	PUNCT
alces-35	169	21	romme	romme	PROPN
alces-35	169	22	et	et	PROPN
alces-35	169	23	al	al	PROPN
alces-35	169	24	.	.	PROPN
alces-35	169	25	2005	2005	NUM
alces-35	169	26	)	)	PUNCT
alces-35	169	27	.	.	PUNCT
alces-35	170	1	in	in	ADP
alces-35	170	2	managed	manage	VERB
alces-35	170	3	forests	forest	NOUN
alces-35	170	4	of	of	ADP
alces-35	170	5	finland	finland	PROPN
alces-35	170	6	,	,	PUNCT
alces-35	170	7	aspen	aspen	PROPN
alces-35	170	8	has	have	AUX
alces-35	170	9	been	be	AUX
alces-35	170	10	an	an	DET
alces-35	170	11	unwanted	unwanted	ADJ
alces-35	170	12	species	specie	NOUN
alces-35	170	13	,	,	PUNCT
alces-35	170	14	especially	especially	ADV
alces-35	170	15	during	during	ADP
alces-35	170	16	the	the	DET
alces-35	170	17	1970s	1970	NOUN
alces-35	170	18	and	and	CCONJ
alces-35	170	19	1980s	1980	NOUN
alces-35	170	20	,	,	PUNCT
alces-35	170	21	because	because	SCONJ
alces-35	170	22	of	of	ADP
alces-35	170	23	its	its	PRON
alces-35	170	24	low	low	ADJ
alces-35	170	25	economic	economic	ADJ
alces-35	170	26	value	value	NOUN
alces-35	170	27	,	,	PUNCT
alces-35	170	28	fast	fast	ADJ
alces-35	170	29	early	early	ADJ
alces-35	170	30	growth	growth	NOUN
alces-35	170	31	and	and	CCONJ
alces-35	170	32	high	high	ADJ
alces-35	170	33	competitive	competitive	ADJ
alces-35	170	34	ability	ability	NOUN
alces-35	170	35	with	with	ADP
alces-35	170	36	more	more	ADV
alces-35	170	37	valuable	valuable	ADJ
alces-35	170	38	tree	tree	NOUN
alces-35	170	39	species	specie	NOUN
alces-35	170	40	,	,	PUNCT
alces-35	170	41	and	and	CCONJ
alces-35	170	42	its	its	PRON
alces-35	170	43	inclination	inclination	NOUN
alces-35	170	44	to	to	PART
alces-35	170	45	host	host	NOUN
alces-35	170	46	rust	rust	NOUN
alces-35	170	47	disease	disease	NOUN
alces-35	170	48	(	(	PUNCT
alces-35	170	49	melampsora	melampsora	NOUN
alces-35	170	50	pinitorqua	pinitorqua	NOUN
alces-35	170	51	)	)	PUNCT
alces-35	170	52	that	that	PRON
alces-35	170	53	is	be	AUX
alces-35	170	54	harmful	harmful	ADJ
alces-35	170	55	to	to	ADP
alces-35	170	56	young	young	ADJ
alces-35	170	57	scots	scot	NOUN
alces-35	170	58	pine	pine	NOUN
alces-35	170	59	stands	stand	NOUN
alces-35	170	60	.	.	PUNCT
alces-35	171	1	for	for	ADP
alces-35	171	2	these	these	DET
alces-35	171	3	reasons	reason	NOUN
alces-35	171	4	foresters	forester	NOUN
alces-35	171	5	have	have	AUX
alces-35	171	6	tried	try	VERB
alces-35	171	7	to	to	PART
alces-35	171	8	control	control	VERB
alces-35	171	9	aspen	aspen	PROPN
alces-35	171	10	with	with	ADP
alces-35	171	11	silvicultural	silvicultural	ADJ
alces-35	171	12	cleanings	cleaning	NOUN
alces-35	171	13	in	in	ADP
alces-35	171	14	young	young	ADJ
alces-35	171	15	stands	stand	NOUN
alces-35	171	16	,	,	PUNCT
alces-35	171	17	and	and	CCONJ
alces-35	171	18	by	by	ADP
alces-35	171	19	notching	notch	VERB
alces-35	171	20	and	and	CCONJ
alces-35	171	21	girdling	girdle	VERB
alces-35	171	22	larger	large	ADJ
alces-35	171	23	individuals	individual	NOUN
alces-35	171	24	in	in	ADP
alces-35	171	25	mature	mature	ADJ
alces-35	171	26	forests	forest	NOUN
alces-35	171	27	before	before	ADP
alces-35	171	28	clear	clear	ADV
alces-35	171	29	-	-	PUNCT
alces-35	171	30	cutting	cutting	NOUN
alces-35	171	31	.	.	PUNCT
alces-35	172	1	considering	consider	VERB
alces-35	172	2	these	these	DET
alces-35	172	3	other	other	ADJ
alces-35	172	4	factors	factor	NOUN
alces-35	172	5	in	in	ADP
alces-35	172	6	protected	protect	VERB
alces-35	172	7	and	and	CCONJ
alces-35	172	8	managed	manage	VERB
alces-35	172	9	forests	forest	NOUN
alces-35	172	10	,	,	PUNCT
alces-35	172	11	it	it	PRON
alces-35	172	12	is	be	AUX
alces-35	172	13	evident	evident	ADJ
alces-35	172	14	as	as	ADP
alces-35	172	15	edenius	edenius	NOUN
alces-35	172	16	and	and	CCONJ
alces-35	172	17	ericsson	ericsson	PROPN
alces-35	172	18	(	(	PUNCT
alces-35	172	19	2007	2007	NUM
alces-35	172	20	)	)	PUNCT
alces-35	172	21	have	have	AUX
alces-35	172	22	presented	present	VERB
alces-35	172	23	,	,	PUNCT
alces-35	172	24	that	that	SCONJ
alces-35	172	25	it	it	PRON
alces-35	172	26	is	be	AUX
alces-35	172	27	not	not	PART
alces-35	172	28	possible	possible	ADJ
alces-35	172	29	to	to	PART
alces-35	172	30	increase	increase	VERB
alces-35	172	31	the	the	DET
alces-35	172	32	abundance	abundance	NOUN
alces-35	172	33	of	of	ADP
alces-35	172	34	aspen	aspen	NOUN
alces-35	172	35	only	only	ADV
alces-35	172	36	by	by	ADP
alces-35	172	37	adjusting	adjust	VERB
alces-35	172	38	ungulate	ungulate	ADJ
alces-35	172	39	browsing	browse	VERB
alces-35	172	40	levels	level	NOUN
alces-35	172	41	.	.	PUNCT
alces-35	173	1	similarly	similarly	ADV
alces-35	173	2	,	,	PUNCT
alces-35	173	3	romme	romme	PROPN
alces-35	173	4	et	et	PROPN
alces-35	173	5	al	al	PROPN
alces-35	173	6	.	.	PROPN
alces-35	173	7	(	(	PUNCT
alces-35	173	8	1995	1995	NUM
alces-35	173	9	)	)	PUNCT
alces-35	173	10	concluded	conclude	VERB
alces-35	173	11	that	that	SCONJ
alces-35	173	12	low	low	ADJ
alces-35	173	13	aspen	aspen	NOUN
alces-35	173	14	regeneration	regeneration	NOUN
alces-35	173	15	can	can	AUX
alces-35	173	16	not	not	PART
alces-35	173	17	be	be	AUX
alces-35	173	18	explained	explain	VERB
alces-35	173	19	by	by	ADP
alces-35	173	20	any	any	DET
alces-35	173	21	single	single	ADJ
alces-35	173	22	factor	factor	NOUN
alces-35	173	23	,	,	PUNCT
alces-35	173	24	but	but	CCONJ
alces-35	173	25	involves	involve	VERB
alces-35	173	26	a	a	DET
alces-35	173	27	complex	complex	ADJ
alces-35	173	28	interaction	interaction	NOUN
alces-35	173	29	of	of	ADP
alces-35	173	30	many	many	ADJ
alces-35	173	31	factors	factor	NOUN
alces-35	173	32	.	.	PUNCT
alces-35	174	1	aspen	aspen	NOUN
alces-35	174	2	regeneration	regeneration	NOUN
alces-35	174	3	can	can	AUX
alces-35	174	4	be	be	AUX
alces-35	174	5	influenced	influence	VERB
alces-35	174	6	by	by	ADP
alces-35	174	7	certain	certain	ADJ
alces-35	174	8	disturbance	disturbance	NOUN
alces-35	174	9	activities	activity	NOUN
alces-35	174	10	.	.	PUNCT
alces-35	175	1	it	it	PRON
alces-35	175	2	has	have	AUX
alces-35	175	3	been	be	AUX
alces-35	175	4	suggested	suggest	VERB
alces-35	175	5	that	that	SCONJ
alces-35	175	6	prescribed	prescribed	ADJ
alces-35	175	7	burning	burning	NOUN
alces-35	175	8	and	and	CCONJ
alces-35	175	9	manmade	manmade	NOUN
alces-35	175	10	gaps	gap	NOUN
alces-35	175	11	(	(	PUNCT
alces-35	175	12	latva	latva	NOUN
alces-35	175	13	-	-	PUNCT
alces-35	175	14	karjanmaa	karjanmaa	VERB
alces-35	175	15	et	et	PROPN
alces-35	175	16	al	al	PROPN
alces-35	175	17	.	.	PROPN
alces-35	175	18	2007	2007	NUM
alces-35	175	19	)	)	PUNCT
alces-35	175	20	will	will	AUX
alces-35	175	21	facilitate	facilitate	VERB
alces-35	175	22	aspen	aspen	NOUN
alces-35	175	23	regeneration	regeneration	NOUN
alces-35	175	24	.	.	PUNCT
alces-35	176	1	creating	create	VERB
alces-35	176	2	large	large	ADJ
alces-35	176	3	canopy	canopy	NOUN
alces-35	176	4	gaps	gap	NOUN
alces-35	176	5	in	in	ADP
alces-35	176	6	old	old	ADJ
alces-35	176	7	-	-	PUNCT
alces-35	176	8	growth	growth	NOUN
alces-35	176	9	forests	forest	NOUN
alces-35	176	10	by	by	ADP
alces-35	176	11	killing	kill	VERB
alces-35	176	12	trees	tree	NOUN
alces-35	176	13	around	around	ADP
alces-35	176	14	mature	mature	ADJ
alces-35	176	15	aspens	aspen	NOUN
alces-35	176	16	may	may	AUX
alces-35	176	17	also	also	ADV
alces-35	176	18	promote	promote	VERB
alces-35	176	19	aspen	aspen	NOUN
alces-35	176	20	regeneration	regeneration	NOUN
alces-35	176	21	.	.	PUNCT
alces-35	177	1	successful	successful	ADJ
alces-35	177	2	aspen	aspen	NOUN
alces-35	177	3	regeneration	regeneration	NOUN
alces-35	177	4	can	can	AUX
alces-35	177	5	be	be	AUX
alces-35	177	6	facilitated	facilitate	VERB
alces-35	177	7	by	by	ADP
alces-35	177	8	other	other	ADJ
alces-35	177	9	human	human	ADJ
alces-35	177	10	manipulations	manipulation	NOUN
alces-35	177	11	,	,	PUNCT
alces-35	177	12	for	for	ADP
alces-35	177	13	instance	instance	NOUN
alces-35	177	14	,	,	PUNCT
alces-35	177	15	protecting	protect	VERB
alces-35	177	16	aspen	aspen	ADJ
alces-35	177	17	recruitment	recruitment	NOUN
alces-35	177	18	physically	physically	ADV
alces-35	177	19	from	from	ADP
alces-35	177	20	browsing	browse	VERB
alces-35	177	21	(	(	PUNCT
alces-35	177	22	angelstam	angelstam	PROPN
alces-35	177	23	et	et	PROPN
alces-35	177	24	al	al	PROPN
alces-35	177	25	.	.	PROPN
alces-35	177	26	2000	2000	NUM
alces-35	177	27	,	,	PUNCT
alces-35	177	28	mclaren	mclaren	PROPN
alces-35	177	29	et	et	PROPN
alces-35	177	30	al	al	PROPN
alces-35	177	31	.	.	PROPN
alces-35	177	32	2004	2004	NUM
alces-35	177	33	,	,	PUNCT
alces-35	177	34	kaye	kaye	PROPN
alces-35	177	35	et	et	PROPN
alces-35	177	36	al	al	PROPN
alces-35	177	37	.	.	PROPN
alces-35	177	38	2005	2005	NUM
alces-35	177	39	,	,	PUNCT
alces-35	177	40	edenius	edenius	NOUN
alces-35	177	41	and	and	CCONJ
alces-35	177	42	ericsson	ericsson	NOUN
alces-35	177	43	2007	2007	NUM
alces-35	177	44	)	)	PUNCT
alces-35	177	45	.	.	PUNCT
alces-35	178	1	in	in	ADP
alces-35	178	2	managed	manage	VERB
alces-35	178	3	forests	forest	NOUN
alces-35	178	4	,	,	PUNCT
alces-35	178	5	young	young	ADJ
alces-35	178	6	aspens	aspen	NOUN
alces-35	178	7	could	could	AUX
alces-35	178	8	be	be	AUX
alces-35	178	9	spared	spare	VERB
alces-35	178	10	during	during	ADP
alces-35	178	11	silvicultural	silvicultural	ADJ
alces-35	178	12	cleanings	cleaning	NOUN
alces-35	178	13	and	and	CCONJ
alces-35	178	14	thinnings	thinning	NOUN
alces-35	178	15	,	,	PUNCT
alces-35	178	16	and	and	CCONJ
alces-35	178	17	mature	mature	ADJ
alces-35	178	18	aspens	aspen	NOUN
alces-35	178	19	could	could	AUX
alces-35	178	20	be	be	AUX
alces-35	178	21	left	leave	VERB
alces-35	178	22	aside	aside	ADV
alces-35	178	23	as	as	ADP
alces-35	178	24	retention	retention	NOUN
alces-35	178	25	trees	tree	NOUN
alces-35	178	26	during	during	ADP
alces-35	178	27	clear	clear	ADJ
alces-35	178	28	-	-	PUNCT
alces-35	178	29	cutting	cutting	NOUN
alces-35	178	30	.	.	PUNCT
alces-35	179	1	the	the	DET
alces-35	179	2	proportion	proportion	NOUN
alces-35	179	3	of	of	ADP
alces-35	179	4	young	young	ADJ
alces-35	179	5	aspens	aspen	NOUN
alces-35	179	6	(	(	PUNCT
alces-35	179	7	<	<	X
alces-35	179	8	5	5	NUM
alces-35	179	9	m	m	NOUN
alces-35	179	10	in	in	ADP
alces-35	179	11	height	height	NOUN
alces-35	179	12	)	)	PUNCT
alces-35	179	13	decreased	decrease	VERB
alces-35	179	14	from	from	ADP
alces-35	179	15	45	45	NUM
alces-35	179	16	%	%	NOUN
alces-35	179	17	in	in	ADP
alces-35	179	18	advanced	advanced	ADJ
alces-35	179	19	seedling	seedling	NOUN
alces-35	179	20	stands	stand	VERB
alces-35	179	21	to	to	ADP
alces-35	179	22	17	17	NUM
alces-35	179	23	%	%	NOUN
alces-35	179	24	in	in	ADP
alces-35	179	25	mature	mature	ADJ
alces-35	179	26	stands	stand	NOUN
alces-35	179	27	.	.	PUNCT
alces-35	180	1	however	however	ADV
alces-35	180	2	,	,	PUNCT
alces-35	180	3	this	this	PRON
alces-35	180	4	clearly	clearly	ADV
alces-35	180	5	indicates	indicate	VERB
alces-35	180	6	that	that	SCONJ
alces-35	180	7	the	the	DET
alces-35	180	8	regeneration	regeneration	NOUN
alces-35	180	9	of	of	ADP
alces-35	180	10	aspen	aspen	NOUN
alces-35	180	11	is	be	AUX
alces-35	180	12	,	,	PUNCT
alces-35	180	13	at	at	ADP
alces-35	180	14	least	least	ADJ
alces-35	180	15	to	to	ADP
alces-35	180	16	some	some	DET
alces-35	180	17	extent	extent	NOUN
alces-35	180	18	,	,	PUNCT
alces-35	180	19	occurring	occur	VERB
alces-35	180	20	in	in	ADP
alces-35	180	21	closed	close	VERB
alces-35	180	22	older	old	ADJ
alces-35	180	23	stands	stand	NOUN
alces-35	180	24	in	in	ADP
alces-35	180	25	the	the	DET
alces-35	180	26	study	study	NOUN
alces-35	180	27	area	area	NOUN
alces-35	180	28	.	.	PUNCT
alces-35	181	1	it	it	PRON
alces-35	181	2	can	can	AUX
alces-35	181	3	be	be	AUX
alces-35	181	4	concluded	conclude	VERB
alces-35	181	5	that	that	SCONJ
alces-35	181	6	the	the	DET
alces-35	181	7	present	present	NOUN
alces-35	181	8	browsing	browse	VERB
alces-35	181	9	pressure	pressure	NOUN
alces-35	181	10	by	by	ADP
alces-35	181	11	overabundant	overabundant	ADJ
alces-35	181	12	moose	moose	NOUN
alces-35	181	13	in	in	ADP
alces-35	181	14	moose	moose	NOUN
alces-35	181	15	and	and	CCONJ
alces-35	181	16	aspen	aspen	NOUN
alces-35	181	17	regeneration	regeneration	NOUN
alces-35	181	18	in	in	ADP
alces-35	181	19	finland	finland	PROPN
alces-35	181	20	härkönen	härkönen	PROPN
alces-35	181	21	et	et	PROPN
alces-35	181	22	al	al	PROPN
alces-35	181	23	.	.	PROPN
alces-35	182	1	alces	alce	NOUN
alces-35	182	2	vol	vol	NOUN
alces-35	182	3	.	.	PROPN
alces-35	182	4	44	44	NUM
alces-35	182	5	,	,	PUNCT
alces-35	182	6	2008	2008	NUM
alces-35	182	7	38	38	NUM
alces-35	182	8	koli	koli	PROPN
alces-35	182	9	national	national	PROPN
alces-35	182	10	park	park	PROPN
alces-35	182	11	not	not	PART
alces-35	182	12	only	only	ADV
alces-35	182	13	kills	kill	VERB
alces-35	182	14	some	some	DET
alces-35	182	15	proportion	proportion	NOUN
alces-35	182	16	of	of	ADP
alces-35	182	17	young	young	ADJ
alces-35	182	18	aspens	aspen	NOUN
alces-35	182	19	,	,	PUNCT
alces-35	182	20	but	but	CCONJ
alces-35	182	21	also	also	ADV
alces-35	182	22	retards	retard	VERB
alces-35	182	23	the	the	DET
alces-35	182	24	height	height	NOUN
alces-35	182	25	development	development	NOUN
alces-35	182	26	of	of	ADP
alces-35	182	27	aspens	aspen	NOUN
alces-35	182	28	because	because	SCONJ
alces-35	182	29	of	of	ADP
alces-35	182	30	the	the	DET
alces-35	182	31	repeated	repeat	VERB
alces-35	182	32	break	break	NOUN
alces-35	182	33	-	-	PUNCT
alces-35	182	34	off	off	NOUN
alces-35	182	35	of	of	ADP
alces-35	182	36	main	main	ADJ
alces-35	182	37	stems	stem	NOUN
alces-35	182	38	and	and	CCONJ
alces-35	182	39	leader	leader	NOUN
alces-35	182	40	shoots	shoot	NOUN
alces-35	182	41	(	(	PUNCT
alces-35	182	42	angelstam	angelstam	PROPN
alces-35	182	43	et	et	PROPN
alces-35	182	44	al	al	PROPN
alces-35	182	45	.	.	PROPN
alces-35	182	46	2000	2000	NUM
alces-35	182	47	,	,	PUNCT
alces-35	182	48	heikkilä	heikkilä	VERB
alces-35	182	49	et	et	PROPN
alces-35	182	50	al	al	PROPN
alces-35	182	51	.	.	PROPN
alces-35	182	52	2003	2003	NUM
alces-35	182	53	)	)	PUNCT
alces-35	182	54	.	.	PUNCT
alces-35	183	1	this	this	PRON
alces-35	183	2	is	be	AUX
alces-35	183	3	supported	support	VERB
alces-35	183	4	with	with	ADP
alces-35	183	5	the	the	DET
alces-35	183	6	observation	observation	NOUN
alces-35	183	7	that	that	SCONJ
alces-35	183	8	where	where	SCONJ
alces-35	183	9	population	population	NOUN
alces-35	183	10	density	density	NOUN
alces-35	183	11	is	be	AUX
alces-35	183	12	>	>	X
alces-35	183	13	5	5	NUM
alces-35	183	14	moose/	moose/	NUM
alces-35	183	15	1,000	1,000	NUM
alces-35	183	16	ha	ha	NOUN
alces-35	183	17	,	,	PUNCT
alces-35	183	18	moose	moose	NOUN
alces-35	183	19	browsing	browse	VERB
alces-35	183	20	depresses	depress	VERB
alces-35	183	21	the	the	DET
alces-35	183	22	growth	growth	NOUN
alces-35	183	23	of	of	ADP
alces-35	183	24	aspen	aspen	PROPN
alces-35	183	25	(	(	PUNCT
alces-35	183	26	abaturov	abaturov	ADJ
alces-35	183	27	and	and	CCONJ
alces-35	183	28	smirnov	smirnov	ADJ
alces-35	183	29	2002	2002	NUM
alces-35	183	30	)	)	PUNCT
alces-35	183	31	.	.	PUNCT
alces-35	184	1	at	at	ADP
alces-35	184	2	present	present	ADJ
alces-35	184	3	,	,	PUNCT
alces-35	184	4	it	it	PRON
alces-35	184	5	seems	seem	VERB
alces-35	184	6	that	that	SCONJ
alces-35	184	7	the	the	DET
alces-35	184	8	major	major	ADJ
alces-35	184	9	proportion	proportion	NOUN
alces-35	184	10	of	of	ADP
alces-35	184	11	young	young	ADJ
alces-35	184	12	aspens	aspen	NOUN
alces-35	184	13	in	in	ADP
alces-35	184	14	koli	koli	PROPN
alces-35	184	15	national	national	PROPN
alces-35	184	16	park	park	PROPN
alces-35	184	17	can	can	AUX
alces-35	184	18	tolerate	tolerate	VERB
alces-35	184	19	both	both	CCONJ
alces-35	184	20	heavy	heavy	ADJ
alces-35	184	21	and	and	CCONJ
alces-35	184	22	repeated	repeat	VERB
alces-35	184	23	browsing	browsing	NOUN
alces-35	184	24	without	without	ADP
alces-35	184	25	dying	die	VERB
alces-35	184	26	.	.	PUNCT
alces-35	185	1	however	however	ADV
alces-35	185	2	,	,	PUNCT
alces-35	185	3	the	the	DET
alces-35	185	4	persistence	persistence	NOUN
alces-35	185	5	of	of	ADP
alces-35	185	6	aspen	aspen	NOUN
alces-35	185	7	may	may	AUX
alces-35	185	8	lead	lead	VERB
alces-35	185	9	to	to	PART
alces-35	185	10	collapse	collapse	VERB
alces-35	185	11	if	if	SCONJ
alces-35	185	12	moose	moose	NOUN
alces-35	185	13	density	density	NOUN
alces-35	185	14	increases	increase	NOUN
alces-35	185	15	and	and	CCONJ
alces-35	185	16	stays	stay	VERB
alces-35	185	17	higher	high	ADJ
alces-35	185	18	than	than	ADP
alces-35	185	19	the	the	DET
alces-35	185	20	current	current	ADJ
alces-35	185	21	carrying	carrying	NOUN
alces-35	185	22	capacity	capacity	NOUN
alces-35	185	23	,	,	PUNCT
alces-35	185	24	which	which	PRON
alces-35	185	25	is	be	AUX
alces-35	185	26	an	an	DET
alces-35	185	27	unforeseeable	unforeseeable	ADJ
alces-35	185	28	scenario	scenario	NOUN
alces-35	185	29	.	.	PUNCT
alces-35	186	1	the	the	DET
alces-35	186	2	long	long	ADJ
alces-35	186	3	-	-	PUNCT
alces-35	186	4	term	term	NOUN
alces-35	186	5	existence	existence	NOUN
alces-35	186	6	,	,	PUNCT
alces-35	186	7	dynamics	dynamic	NOUN
alces-35	186	8	,	,	PUNCT
alces-35	186	9	and	and	CCONJ
alces-35	186	10	possibilities	possibility	NOUN
alces-35	186	11	for	for	ADP
alces-35	186	12	natural	natural	ADJ
alces-35	186	13	regeneration	regeneration	NOUN
alces-35	186	14	of	of	ADP
alces-35	186	15	aspen	aspen	NOUN
alces-35	186	16	in	in	ADP
alces-35	186	17	old	old	ADJ
alces-35	186	18	-	-	PUNCT
alces-35	186	19	growth	growth	NOUN
alces-35	186	20	forests	forest	NOUN
alces-35	186	21	in	in	ADP
alces-35	186	22	koli	koli	PROPN
alces-35	186	23	national	national	PROPN
alces-35	186	24	park	park	PROPN
alces-35	186	25	have	have	AUX
alces-35	186	26	been	be	AUX
alces-35	186	27	determined	determine	VERB
alces-35	186	28	by	by	ADP
alces-35	186	29	the	the	DET
alces-35	186	30	compartmentwise	compartmentwise	NOUN
alces-35	186	31	inventory	inventory	NOUN
alces-35	186	32	data	datum	NOUN
alces-35	186	33	available	available	ADJ
alces-35	186	34	for	for	ADP
alces-35	186	35	the	the	DET
alces-35	186	36	period	period	NOUN
alces-35	186	37	1910–2004	1910–2004	NUM
alces-35	186	38	(	(	PUNCT
alces-35	186	39	m.	m.	NOUN
alces-35	186	40	vehmas	vehma	NOUN
alces-35	186	41	et	et	PROPN
alces-35	186	42	al	al	PROPN
alces-35	186	43	.	.	PROPN
alces-35	186	44	,	,	PUNCT
alces-35	186	45	university	university	NOUN
alces-35	186	46	of	of	ADP
alces-35	186	47	joensuu	joensuu	NOUN
alces-35	186	48	,	,	PUNCT
alces-35	186	49	unpubl	unpubl	NOUN
alces-35	186	50	.	.	PUNCT
alces-35	187	1	data	datum	NOUN
alces-35	187	2	)	)	PUNCT
alces-35	187	3	.	.	PUNCT
alces-35	188	1	their	their	PRON
alces-35	188	2	results	result	NOUN
alces-35	188	3	indicated	indicate	VERB
alces-35	188	4	that	that	SCONJ
alces-35	188	5	aspen	aspen	PROPN
alces-35	188	6	can	can	AUX
alces-35	188	7	regenerate	regenerate	VERB
alces-35	188	8	naturally	naturally	ADV
alces-35	188	9	in	in	ADP
alces-35	188	10	old	old	ADJ
alces-35	188	11	-	-	PUNCT
alces-35	188	12	growth	growth	NOUN
alces-35	188	13	forests	forest	NOUN
alces-35	188	14	and	and	CCONJ
alces-35	188	15	that	that	SCONJ
alces-35	188	16	its	its	PRON
alces-35	188	17	existence	existence	NOUN
alces-35	188	18	is	be	AUX
alces-35	188	19	not	not	PART
alces-35	188	20	completely	completely	ADV
alces-35	188	21	dependent	dependent	ADJ
alces-35	188	22	on	on	ADP
alces-35	188	23	young	young	ADJ
alces-35	188	24	successional	successional	ADJ
alces-35	188	25	stages	stage	NOUN
alces-35	188	26	of	of	ADP
alces-35	188	27	forests	forest	NOUN
alces-35	188	28	,	,	PUNCT
alces-35	188	29	which	which	PRON
alces-35	188	30	corresponds	correspond	VERB
alces-35	188	31	to	to	ADP
alces-35	188	32	the	the	DET
alces-35	188	33	findings	finding	NOUN
alces-35	188	34	of	of	ADP
alces-35	188	35	this	this	DET
alces-35	188	36	study	study	NOUN
alces-35	188	37	.	.	PUNCT
alces-35	189	1	however	however	ADV
alces-35	189	2	,	,	PUNCT
alces-35	189	3	they	they	PRON
alces-35	189	4	assumed	assume	VERB
alces-35	189	5	that	that	SCONJ
alces-35	189	6	the	the	DET
alces-35	189	7	long	long	ADJ
alces-35	189	8	-	-	PUNCT
alces-35	189	9	term	term	NOUN
alces-35	189	10	ecological	ecological	ADJ
alces-35	189	11	continuity	continuity	NOUN
alces-35	189	12	may	may	AUX
alces-35	189	13	be	be	AUX
alces-35	189	14	threatened	threaten	VERB
alces-35	189	15	by	by	ADP
alces-35	189	16	increased	increase	VERB
alces-35	189	17	population	population	NOUN
alces-35	189	18	densities	density	NOUN
alces-35	189	19	of	of	ADP
alces-35	189	20	large	large	ADJ
alces-35	189	21	mammalian	mammalian	ADJ
alces-35	189	22	browsers	browser	NOUN
alces-35	189	23	such	such	ADJ
alces-35	189	24	as	as	ADP
alces-35	189	25	overabundant	overabundant	ADJ
alces-35	189	26	moose	moose	NOUN
alces-35	189	27	.	.	PUNCT
alces-35	190	1	even	even	ADV
alces-35	190	2	if	if	SCONJ
alces-35	190	3	their	their	PRON
alces-35	190	4	study	study	NOUN
alces-35	190	5	was	be	AUX
alces-35	190	6	not	not	PART
alces-35	190	7	focused	focus	VERB
alces-35	190	8	on	on	ADP
alces-35	190	9	identifying	identify	VERB
alces-35	190	10	factors	factor	NOUN
alces-35	190	11	related	relate	VERB
alces-35	190	12	to	to	ADP
alces-35	190	13	the	the	DET
alces-35	190	14	success	success	NOUN
alces-35	190	15	of	of	ADP
alces-35	190	16	aspen	aspen	PROPN
alces-35	190	17	regeneration	regeneration	NOUN
alces-35	190	18	,	,	PUNCT
alces-35	190	19	it	it	PRON
alces-35	190	20	concluded	conclude	VERB
alces-35	190	21	that	that	SCONJ
alces-35	190	22	the	the	DET
alces-35	190	23	issue	issue	NOUN
alces-35	190	24	between	between	ADP
alces-35	190	25	the	the	DET
alces-35	190	26	seemingly	seemingly	ADV
alces-35	190	27	increasing	increase	VERB
alces-35	190	28	population	population	NOUN
alces-35	190	29	densities	density	NOUN
alces-35	190	30	of	of	ADP
alces-35	190	31	mammalian	mammalian	ADJ
alces-35	190	32	browsers	browser	NOUN
alces-35	190	33	and	and	CCONJ
alces-35	190	34	success	success	NOUN
alces-35	190	35	of	of	ADP
alces-35	190	36	aspen	aspen	ADJ
alces-35	190	37	regeneration	regeneration	NOUN
alces-35	190	38	needs	need	NOUN
alces-35	190	39	to	to	PART
alces-35	190	40	be	be	AUX
alces-35	190	41	considered	consider	VERB
alces-35	190	42	in	in	ADP
alces-35	190	43	future	future	ADJ
alces-35	190	44	restoration	restoration	NOUN
alces-35	190	45	and	and	CCONJ
alces-35	190	46	management	management	NOUN
alces-35	190	47	measures	measure	NOUN
alces-35	190	48	planned	plan	VERB
alces-35	190	49	for	for	ADP
alces-35	190	50	koli	koli	PROPN
alces-35	190	51	national	national	PROPN
alces-35	190	52	park	park	PROPN
alces-35	190	53	.	.	PUNCT
alces-35	191	1	suzuki	suzuki	PROPN
alces-35	191	2	et	et	PROPN
alces-35	191	3	al	al	PROPN
alces-35	191	4	.	.	PROPN
alces-35	191	5	(	(	PUNCT
alces-35	191	6	1999	1999	NUM
alces-35	191	7	)	)	PUNCT
alces-35	191	8	concluded	conclude	VERB
alces-35	191	9	that	that	SCONJ
alces-35	191	10	elk	elk	PROPN
alces-35	191	11	(	(	PUNCT
alces-35	191	12	cervus	cervus	NOUN
alces-35	191	13	elaphus	elaphu	NOUN
alces-35	191	14	)	)	PUNCT
alces-35	191	15	browsing	browse	VERB
alces-35	191	16	in	in	ADP
alces-35	191	17	rocky	rocky	ADJ
alces-35	191	18	mountain	mountain	NOUN
alces-35	191	19	national	national	ADJ
alces-35	191	20	park	park	NOUN
alces-35	191	21	was	be	AUX
alces-35	191	22	not	not	PART
alces-35	191	23	causing	cause	VERB
alces-35	191	24	failure	failure	NOUN
alces-35	191	25	of	of	ADP
alces-35	191	26	aspen	aspen	ADJ
alces-35	191	27	regeneration	regeneration	NOUN
alces-35	191	28	at	at	ADP
alces-35	191	29	landscape	landscape	NOUN
alces-35	191	30	scales	scale	NOUN
alces-35	191	31	,	,	PUNCT
alces-35	191	32	whereas	whereas	SCONJ
alces-35	191	33	in	in	ADP
alces-35	191	34	local	local	ADJ
alces-35	191	35	areas	area	NOUN
alces-35	191	36	with	with	ADP
alces-35	191	37	the	the	DET
alces-35	191	38	highest	high	ADJ
alces-35	191	39	browsing	browse	VERB
alces-35	191	40	pressure	pressure	NOUN
alces-35	191	41	from	from	ADP
alces-35	191	42	elk	elk	PROPN
alces-35	191	43	,	,	PUNCT
alces-35	191	44	little	little	ADJ
alces-35	191	45	regeneration	regeneration	NOUN
alces-35	191	46	has	have	AUX
alces-35	191	47	occurred	occur	VERB
alces-35	191	48	in	in	ADP
alces-35	191	49	the	the	DET
alces-35	191	50	past	past	ADJ
alces-35	191	51	30	30	NUM
alces-35	191	52	years	year	NOUN
alces-35	191	53	.	.	PUNCT
alces-35	192	1	a	a	DET
alces-35	192	2	similar	similar	ADJ
alces-35	192	3	observation	observation	NOUN
alces-35	192	4	was	be	AUX
alces-35	192	5	made	make	VERB
alces-35	192	6	by	by	ADP
alces-35	192	7	heikkilä	heikkilä	NOUN
alces-35	192	8	et	et	PROPN
alces-35	192	9	al	al	PROPN
alces-35	192	10	.	.	PROPN
alces-35	193	1	(	(	PUNCT
alces-35	193	2	2003	2003	NUM
alces-35	193	3	)	)	PUNCT
alces-35	193	4	when	when	SCONJ
alces-35	193	5	evaluating	evaluate	VERB
alces-35	193	6	the	the	DET
alces-35	193	7	effects	effect	NOUN
alces-35	193	8	of	of	ADP
alces-35	193	9	moose	moose	NOUN
alces-35	193	10	browsing	browse	VERB
alces-35	193	11	on	on	ADP
alces-35	193	12	aspen	aspen	NOUN
alces-35	193	13	in	in	ADP
alces-35	193	14	finland	finland	PROPN
alces-35	193	15	,	,	PUNCT
alces-35	193	16	and	and	CCONJ
alces-35	193	17	in	in	ADP
alces-35	193	18	national	national	ADJ
alces-35	193	19	parks	park	NOUN
alces-35	193	20	in	in	ADP
alces-35	193	21	newfoundland	newfoundland	PROPN
alces-35	193	22	and	and	CCONJ
alces-35	193	23	labrador	labrador	PROPN
alces-35	193	24	(	(	PUNCT
alces-35	193	25	mclaren	mclaren	PROPN
alces-35	193	26	et	et	PROPN
alces-35	193	27	al	al	PROPN
alces-35	193	28	.	.	PROPN
alces-35	193	29	2004	2004	NUM
alces-35	193	30	)	)	PUNCT
alces-35	193	31	.	.	PUNCT
alces-35	194	1	our	our	PRON
alces-35	194	2	study	study	NOUN
alces-35	194	3	supports	support	VERB
alces-35	194	4	the	the	DET
alces-35	194	5	conclusion	conclusion	NOUN
alces-35	194	6	that	that	SCONJ
alces-35	194	7	moose	moose	NOUN
alces-35	194	8	browsing	browse	VERB
alces-35	194	9	will	will	AUX
alces-35	194	10	most	most	ADV
alces-35	194	11	likely	likely	ADV
alces-35	194	12	contribute	contribute	VERB
alces-35	194	13	to	to	ADP
alces-35	194	14	an	an	DET
alces-35	194	15	altered	altered	ADJ
alces-35	194	16	size	size	NOUN
alces-35	194	17	distribution	distribution	NOUN
alces-35	194	18	of	of	ADP
alces-35	194	19	aspen	aspen	NOUN
alces-35	194	20	in	in	ADP
alces-35	194	21	the	the	DET
alces-35	194	22	boreal	boreal	ADJ
alces-35	194	23	landscape	landscape	NOUN
alces-35	194	24	(	(	PUNCT
alces-35	194	25	kay	kay	NOUN
alces-35	194	26	1997	1997	NUM
alces-35	194	27	,	,	PUNCT
alces-35	194	28	suzuki	suzuki	PROPN
alces-35	194	29	et	et	PROPN
alces-35	194	30	al	al	PROPN
alces-35	194	31	.	.	PROPN
alces-35	194	32	1999	1999	NUM
alces-35	194	33	,	,	PUNCT
alces-35	194	34	ericsson	ericsson	PROPN
alces-35	194	35	et	et	PROPN
alces-35	194	36	al	al	PROPN
alces-35	194	37	.	.	PROPN
alces-35	194	38	2001	2001	NUM
alces-35	194	39	)	)	PUNCT
alces-35	194	40	.	.	PUNCT
alces-35	195	1	hence	hence	ADV
alces-35	195	2	,	,	PUNCT
alces-35	195	3	the	the	DET
alces-35	195	4	number	number	NOUN
alces-35	195	5	of	of	ADP
alces-35	195	6	aspens	aspen	NOUN
alces-35	195	7	in	in	ADP
alces-35	195	8	koli	koli	PROPN
alces-35	195	9	national	national	PROPN
alces-35	195	10	park	park	PROPN
alces-35	195	11	may	may	AUX
alces-35	195	12	decrease	decrease	VERB
alces-35	195	13	due	due	ADP
alces-35	195	14	to	to	ADP
alces-35	195	15	the	the	DET
alces-35	195	16	high	high	ADJ
alces-35	195	17	browsing	browse	VERB
alces-35	195	18	pressure	pressure	NOUN
alces-35	195	19	by	by	ADP
alces-35	195	20	moose	moose	NOUN
alces-35	195	21	,	,	PUNCT
alces-35	195	22	but	but	CCONJ
alces-35	195	23	many	many	ADJ
alces-35	195	24	aspens	aspen	NOUN
alces-35	195	25	may	may	AUX
alces-35	195	26	reach	reach	VERB
alces-35	195	27	maturity	maturity	NOUN
alces-35	195	28	once	once	ADV
alces-35	195	29	having	having	AUX
alces-35	195	30	passed	pass	VERB
alces-35	195	31	a	a	DET
alces-35	195	32	risky	risky	ADJ
alces-35	195	33	regeneration	regeneration	NOUN
alces-35	195	34	stage	stage	NOUN
alces-35	195	35	.	.	PUNCT
alces-35	196	1	this	this	PRON
alces-35	196	2	should	should	AUX
alces-35	196	3	guarantee	guarantee	VERB
alces-35	196	4	the	the	DET
alces-35	196	5	spatio	spatio	PROPN
alces-35	196	6	-	-	PUNCT
alces-35	196	7	temporal	temporal	ADJ
alces-35	196	8	continuum	continuum	NOUN
alces-35	196	9	of	of	ADP
alces-35	196	10	aspen	aspen	NOUN
alces-35	196	11	at	at	ADP
alces-35	196	12	a	a	DET
alces-35	196	13	level	level	NOUN
alces-35	196	14	where	where	SCONJ
alces-35	196	15	biodiversity	biodiversity	NOUN
alces-35	196	16	is	be	AUX
alces-35	196	17	maintained	maintain	VERB
alces-35	196	18	in	in	ADP
alces-35	196	19	the	the	DET
alces-35	196	20	conservation	conservation	NOUN
alces-35	196	21	area	area	NOUN
alces-35	196	22	.	.	PUNCT
alces-35	197	1	however	however	ADV
alces-35	197	2	,	,	PUNCT
alces-35	197	3	aspen	aspen	PROPN
alces-35	197	4	may	may	AUX
alces-35	197	5	not	not	PART
alces-35	197	6	necessarily	necessarily	ADV
alces-35	197	7	continue	continue	VERB
alces-35	197	8	to	to	PART
alces-35	197	9	exist	exist	VERB
alces-35	197	10	where	where	SCONJ
alces-35	197	11	foresters	forester	NOUN
alces-35	197	12	and	and	CCONJ
alces-35	197	13	conservationists	conservationist	NOUN
alces-35	197	14	desire	desire	NOUN
alces-35	197	15	(	(	PUNCT
alces-35	197	16	ericsson	ericsson	PROPN
alces-35	197	17	et	et	PROPN
alces-35	197	18	al	al	PROPN
alces-35	197	19	.	.	PROPN
alces-35	197	20	2001	2001	NUM
alces-35	197	21	)	)	PUNCT
alces-35	197	22	.	.	PUNCT
alces-35	198	1	acknowledgements	acknowledgement	NOUN
alces-35	198	2	we	we	PRON
alces-35	198	3	would	would	AUX
alces-35	198	4	like	like	VERB
alces-35	198	5	to	to	PART
alces-35	198	6	thank	thank	VERB
alces-35	198	7	mr	mr	PROPN
alces-35	198	8	.	.	PROPN
alces-35	198	9	lasse	lasse	PROPN
alces-35	198	10	lovén	lovén	PROPN
alces-35	198	11	(	(	PUNCT
alces-35	198	12	finnish	finnish	PROPN
alces-35	198	13	forest	forest	PROPN
alces-35	198	14	research	research	PROPN
alces-35	198	15	institute	institute	PROPN
alces-35	198	16	,	,	PUNCT
alces-35	198	17	koli	koli	PROPN
alces-35	198	18	national	national	PROPN
alces-35	198	19	park	park	PROPN
alces-35	198	20	)	)	PUNCT
alces-35	198	21	for	for	ADP
alces-35	198	22	providing	provide	VERB
alces-35	198	23	the	the	DET
alces-35	198	24	facilities	facility	NOUN
alces-35	198	25	during	during	ADP
alces-35	198	26	fieldwork	fieldwork	NOUN
alces-35	198	27	.	.	PUNCT
alces-35	199	1	we	we	PRON
alces-35	199	2	are	be	AUX
alces-35	199	3	also	also	ADV
alces-35	199	4	grateful	grateful	ADJ
alces-35	199	5	to	to	ADP
alces-35	199	6	mr	mr	PROPN
alces-35	199	7	.	.	PROPN
alces-35	199	8	mika	mika	PROPN
alces-35	199	9	venho	venho	PROPN
alces-35	199	10	for	for	ADP
alces-35	199	11	his	his	PRON
alces-35	199	12	help	help	NOUN
alces-35	199	13	in	in	ADP
alces-35	199	14	extracting	extract	VERB
alces-35	199	15	the	the	DET
alces-35	199	16	aspen	aspen	NOUN
alces-35	199	17	stand	stand	VERB
alces-35	199	18	information	information	NOUN
alces-35	199	19	from	from	ADP
alces-35	199	20	the	the	DET
alces-35	199	21	forest	forest	NOUN
alces-35	199	22	inventory	inventory	NOUN
alces-35	199	23	data	datum	NOUN
alces-35	199	24	of	of	ADP
alces-35	199	25	koli	koli	PROPN
alces-35	199	26	national	national	PROPN
alces-35	199	27	park	park	PROPN
alces-35	199	28	,	,	PUNCT
alces-35	199	29	and	and	CCONJ
alces-35	199	30	to	to	ADP
alces-35	199	31	2	2	NUM
alces-35	199	32	anonymous	anonymous	ADJ
alces-35	199	33	referees	referee	NOUN
alces-35	199	34	for	for	ADP
alces-35	199	35	their	their	PRON
alces-35	199	36	valuable	valuable	ADJ
alces-35	199	37	comments	comment	NOUN
alces-35	199	38	on	on	ADP
alces-35	199	39	the	the	DET
alces-35	199	40	manuscript	manuscript	NOUN
alces-35	199	41	.	.	PUNCT
alces-35	200	1	references	reference	NOUN
alces-35	200	2	abaturov	abaturov	VERB
alces-35	200	3	,	,	PUNCT
alces-35	200	4	b.	b.	PROPN
alces-35	200	5	d.	d.	PROPN
alces-35	200	6	,	,	PUNCT
alces-35	200	7	and	and	CCONJ
alces-35	200	8	k.	k.	PROPN
alces-35	200	9	a.	a.	PROPN
alces-35	200	10	smirnov	smirnov	PROPN
alces-35	200	11	.	.	PUNCT
alces-35	200	12	2002	2002	NUM
alces-35	200	13	.	.	PUNCT
alces-35	201	1	effects	effect	NOUN
alces-35	201	2	of	of	ADP
alces-35	201	3	moose	moose	NOUN
alces-35	201	4	population	population	NOUN
alces-35	201	5	density	density	NOUN
alces-35	201	6	on	on	ADP
alces-35	201	7	development	development	NOUN
alces-35	201	8	of	of	ADP
alces-35	201	9	forest	forest	NOUN
alces-35	201	10	stands	stand	VERB
alces-35	201	11	in	in	ADP
alces-35	201	12	central	central	ADJ
alces-35	201	13	european	european	PROPN
alces-35	201	14	russia	russia	PROPN
alces-35	201	15	.	.	PUNCT
alces-35	202	1	alces	alce	NOUN
alces-35	202	2	supplement	supplement	VERB
alces-35	202	3	2:1	2:1	NUM
alces-35	202	4	-	-	SYM
alces-35	202	5	5	5	NUM
alces-35	202	6	.	.	PUNCT
alces-35	202	7	angelstam	angelstam	PROPN
alces-35	202	8	,	,	PUNCT
alces-35	202	9	p.	p.	NOUN
alces-35	202	10	,	,	PUNCT
alces-35	202	11	p.-e	p.-e	NOUN
alces-35	202	12	.	.	PUNCT
alces-35	203	1	wikberg	wikberg	PROPN
alces-35	203	2	,	,	PUNCT
alces-35	203	3	p.	p.	PROPN
alces-35	203	4	danilov	danilov	PROPN
alces-35	203	5	,	,	PUNCT
alces-35	203	6	w.	w.	PROPN
alces-35	203	7	e.	e.	PROPN
alces-35	203	8	faber	faber	PROPN
alces-35	203	9	,	,	PUNCT
alces-35	203	10	and	and	CCONJ
alces-35	203	11	k.	k.	PROPN
alces-35	203	12	nygrén	nygrén	PROPN
alces-35	203	13	.	.	PUNCT
alces-35	204	1	2000	2000	NUM
alces-35	204	2	.	.	PUNCT
alces-35	205	1	effects	effect	NOUN
alces-35	205	2	of	of	ADP
alces-35	205	3	moose	moose	NOUN
alces-35	205	4	density	density	NOUN
alces-35	205	5	on	on	ADP
alces-35	205	6	timber	timber	NOUN
alces-35	205	7	quality	quality	NOUN
alces-35	205	8	and	and	CCONJ
alces-35	205	9	biodiversity	biodiversity	NOUN
alces-35	205	10	restoration	restoration	NOUN
alces-35	205	11	in	in	ADP
alces-35	205	12	sweden	sweden	PROPN
alces-35	205	13	,	,	PUNCT
alces-35	205	14	finland	finland	PROPN
alces-35	205	15	,	,	PUNCT
alces-35	205	16	and	and	CCONJ
alces-35	205	17	russian	russian	PROPN
alces-35	205	18	karelia	karelia	PROPN
alces-35	205	19	.	.	PUNCT
alces-35	206	1	alces	alces	PROPN
alces-35	206	2	36:133	36:133	NUM
alces-35	206	3	-	-	SYM
alces-35	206	4	145	145	NUM
alces-35	206	5	.	.	PUNCT
alces-35	207	1	baker	baker	PROPN
alces-35	207	2	,	,	PUNCT
alces-35	207	3	w.	w.	PROPN
alces-35	207	4	l.	l.	PROPN
alces-35	207	5	,	,	PUNCT
alces-35	207	6	j.	j.	PROPN
alces-35	207	7	a.	a.	PROPN
alces-35	207	8	munroe	munroe	PROPN
alces-35	207	9	,	,	PUNCT
alces-35	207	10	and	and	CCONJ
alces-35	207	11	a.	a.	PROPN
alces-35	207	12	e.	e.	PROPN
alces-35	207	13	hessl	hessl	PROPN
alces-35	207	14	.	.	PUNCT
alces-35	208	1	alces	alce	NOUN
alces-35	208	2	vol	vol	NOUN
alces-35	208	3	.	.	PROPN
alces-35	208	4	44	44	NUM
alces-35	208	5	,	,	PUNCT
alces-35	208	6	2008	2008	NUM
alces-35	208	7	härkönen	härkönen	NOUN
alces-35	208	8	et	et	PROPN
alces-35	208	9	al.moose	al.moose	X
alces-35	208	10	and	and	CCONJ
alces-35	208	11	aspen	aspen	VERB
alces-35	208	12	regeneration	regeneration	NOUN
alces-35	208	13	in	in	ADP
alces-35	208	14	finland	finland	PROPN
alces-35	208	15	39	39	NUM
alces-35	208	16	1997	1997	NUM
alces-35	208	17	.	.	PUNCT
alces-35	209	1	the	the	DET
alces-35	209	2	effects	effect	NOUN
alces-35	209	3	of	of	ADP
alces-35	209	4	elk	elk	NOUN
alces-35	209	5	on	on	ADP
alces-35	209	6	aspen	aspen	NOUN
alces-35	209	7	in	in	ADP
alces-35	209	8	the	the	DET
alces-35	209	9	winter	winter	NOUN
alces-35	209	10	range	range	NOUN
alces-35	209	11	in	in	ADP
alces-35	209	12	rocky	rocky	ADJ
alces-35	209	13	mountain	mountain	NOUN
alces-35	209	14	national	national	PROPN
alces-35	209	15	park	park	PROPN
alces-35	209	16	.	.	PUNCT
alces-35	210	1	ecography	ecography	NOUN
alces-35	210	2	20:155	20:155	PROPN
alces-35	210	3	-	-	SYM
alces-35	210	4	165	165	NUM
alces-35	210	5	.	.	PUNCT
alces-35	211	1	bergström	bergström	NOUN
alces-35	211	2	,	,	PUNCT
alces-35	211	3	r.	r.	PROPN
alces-35	211	4	,	,	PUNCT
alces-35	211	5	and	and	CCONJ
alces-35	211	6	o.	o.	PROPN
alces-35	211	7	hjeljord	hjeljord	PROPN
alces-35	211	8	.	.	PUNCT
alces-35	212	1	1987	1987	NUM
alces-35	212	2	.	.	PUNCT
alces-35	213	1	moose	moose	NOUN
alces-35	213	2	and	and	CCONJ
alces-35	213	3	vegetation	vegetation	NOUN
alces-35	213	4	interactions	interaction	NOUN
alces-35	213	5	in	in	ADP
alces-35	213	6	northwestern	northwestern	PROPN
alces-35	213	7	europe	europe	PROPN
alces-35	213	8	and	and	CCONJ
alces-35	213	9	poland	poland	PROPN
alces-35	213	10	.	.	PUNCT
alces-35	214	1	swedish	swedish	ADJ
alces-35	214	2	wildlife	wildlife	PROPN
alces-35	214	3	research	research	NOUN
alces-35	214	4	supplement	supplement	NOUN
alces-35	214	5	1:213	1:213	NUM
alces-35	214	6	-	-	SYM
alces-35	214	7	228	228	NUM
alces-35	214	8	.	.	PUNCT
alces-35	215	1	cumming	cumming	PROPN
alces-35	215	2	,	,	PUNCT
alces-35	215	3	s.	s.	PROPN
alces-35	215	4	g.	g.	PROPN
alces-35	215	5	,	,	PUNCT
alces-35	215	6	f.	f.	PROPN
alces-35	215	7	k.	k.	PROPN
alces-35	215	8	a.	a.	PROPN
alces-35	215	9	schmiegelow	schmiegelow	PROPN
alces-35	215	10	,	,	PUNCT
alces-35	215	11	and	and	CCONJ
alces-35	215	12	p.	p.	PROPN
alces-35	215	13	j.	j.	PROPN
alces-35	215	14	burton	burton	PROPN
alces-35	215	15	.	.	PUNCT
alces-35	216	1	2000	2000	NUM
alces-35	216	2	.	.	PUNCT
alces-35	217	1	gap	gap	NOUN
alces-35	217	2	dynamics	dynamic	NOUN
alces-35	217	3	in	in	ADP
alces-35	217	4	boreal	boreal	ADJ
alces-35	217	5	aspen	aspen	PROPN
alces-35	217	6	stands	stand	NOUN
alces-35	217	7	:	:	PUNCT
alces-35	217	8	is	be	AUX
alces-35	217	9	the	the	DET
alces-35	217	10	forest	forest	NOUN
alces-35	217	11	older	old	ADJ
alces-35	217	12	than	than	SCONJ
alces-35	217	13	we	we	PRON
alces-35	217	14	think	think	VERB
alces-35	217	15	?	?	PUNCT
alces-35	218	1	ecological	ecological	ADJ
alces-35	218	2	applications	application	NOUN
alces-35	218	3	10:744	10:744	NUM
alces-35	218	4	-	-	SYM
alces-35	218	5	759	759	NUM
alces-35	218	6	.	.	PUNCT
alces-35	219	1	edenius	edenius	NOUN
alces-35	219	2	,	,	PUNCT
alces-35	219	3	l.	l.	PROPN
alces-35	219	4	,	,	PUNCT
alces-35	219	5	and	and	CCONJ
alces-35	219	6	g.	g.	PROPN
alces-35	219	7	ericsson	ericsson	PROPN
alces-35	219	8	.	.	PUNCT
alces-35	220	1	2007	2007	NUM
alces-35	220	2	.	.	PUNCT
alces-35	221	1	aspen	aspen	ADJ
alces-35	221	2	demographics	demographic	NOUN
alces-35	221	3	in	in	ADP
alces-35	221	4	relation	relation	NOUN
alces-35	221	5	to	to	ADP
alces-35	221	6	spatial	spatial	ADJ
alces-35	221	7	context	context	NOUN
alces-35	221	8	and	and	CCONJ
alces-35	221	9	ungulate	ungulate	ADJ
alces-35	221	10	browsing	browsing	NOUN
alces-35	221	11	:	:	PUNCT
alces-35	221	12	implications	implication	NOUN
alces-35	221	13	for	for	ADP
alces-35	221	14	conservation	conservation	NOUN
alces-35	221	15	and	and	CCONJ
alces-35	221	16	forest	forest	NOUN
alces-35	221	17	management	management	NOUN
alces-35	221	18	.	.	PUNCT
alces-35	222	1	biological	biological	ADJ
alces-35	222	2	conservation	conservation	NOUN
alces-35	222	3	135:293	135:293	PROPN
alces-35	222	4	-	-	SYM
alces-35	222	5	301	301	NUM
alces-35	222	6	.	.	PUNCT
alces-35	223	1	_	_	PUNCT
alces-35	224	1	_	_	PUNCT
alces-35	225	1	_	_	PUNCT
alces-35	226	1	_	_	PUNCT
alces-35	227	1	_	_	PUNCT
alces-35	227	2	,	,	PUNCT
alces-35	228	1	_	_	PUNCT
alces-35	229	1	_	_	PUNCT
alces-35	230	1	_	_	PUNCT
alces-35	231	1	_	_	PUNCT
alces-35	232	1	_	_	PUNCT
alces-35	232	2	,	,	PUNCT
alces-35	232	3	and	and	CCONJ
alces-35	232	4	p.	p.	PROPN
alces-35	232	5	näslund	näslund	PROPN
alces-35	232	6	.	.	PUNCT
alces-35	233	1	2002	2002	NUM
alces-35	233	2	.	.	PUNCT
alces-35	234	1	selectivity	selectivity	NOUN
alces-35	234	2	by	by	ADP
alces-35	234	3	moose	moose	NOUN
alces-35	234	4	vs	vs	ADP
alces-35	234	5	spatial	spatial	ADJ
alces-35	234	6	distribution	distribution	NOUN
alces-35	234	7	of	of	ADP
alces-35	234	8	aspen	aspen	NOUN
alces-35	234	9	:	:	PUNCT
alces-35	234	10	a	a	DET
alces-35	234	11	natural	natural	ADJ
alces-35	234	12	experiment	experiment	NOUN
alces-35	234	13	.	.	PUNCT
alces-35	235	1	ecography	ecography	NOUN
alces-35	235	2	25:289	25:289	NUM
alces-35	235	3	-	-	SYM
alces-35	235	4	294	294	NUM
alces-35	235	5	.	.	PUNCT
alces-35	235	6	ericsson	ericsson	PROPN
alces-35	235	7	,	,	PUNCT
alces-35	235	8	g.	g.	PROPN
alces-35	235	9	,	,	PUNCT
alces-35	235	10	l.	l.	PROPN
alces-35	235	11	edenius	edenius	PROPN
alces-35	235	12	,	,	PUNCT
alces-35	235	13	and	and	CCONJ
alces-35	235	14	d.	d.	PROPN
alces-35	235	15	sundström	sundström	PROPN
alces-35	235	16	.	.	PUNCT
alces-35	236	1	2001	2001	NUM
alces-35	236	2	.	.	PUNCT
alces-35	237	1	factors	factor	NOUN
alces-35	237	2	affecting	affect	VERB
alces-35	237	3	browsing	browse	VERB
alces-35	237	4	by	by	ADP
alces-35	237	5	moose	moose	NOUN
alces-35	237	6	(	(	PUNCT
alces-35	237	7	alces	alces	PROPN
alces-35	237	8	alces	alce	NOUN
alces-35	237	9	l.	l.	PROPN
alces-35	237	10	)	)	PUNCT
alces-35	237	11	on	on	ADP
alces-35	237	12	european	european	PROPN
alces-35	237	13	aspen	aspen	PROPN
alces-35	237	14	(	(	PUNCT
alces-35	237	15	populus	populus	PROPN
alces-35	237	16	tremula	tremula	NOUN
alces-35	237	17	l.	l.	PROPN
alces-35	237	18	)	)	PUNCT
alces-35	237	19	in	in	ADP
alces-35	237	20	a	a	DET
alces-35	237	21	managed	manage	VERB
alces-35	237	22	boreal	boreal	ADJ
alces-35	237	23	landscape	landscape	NOUN
alces-35	237	24	.	.	PUNCT
alces-35	238	1	écoscience	écoscience	NOUN
alces-35	238	2	8:344	8:344	NOUN
alces-35	238	3	-	-	SYM
alces-35	238	4	349	349	NUM
alces-35	238	5	.	.	PUNCT
alces-35	239	1	finnish	finnish	PROPN
alces-35	239	2	forest	forest	PROPN
alces-35	239	3	research	research	PROPN
alces-35	239	4	institute	institute	PROPN
alces-35	239	5	.	.	PUNCT
alces-35	240	1	2006	2006	NUM
alces-35	240	2	.	.	PUNCT
alces-35	241	1	finnish	finnish	ADJ
alces-35	241	2	statistical	statistical	ADJ
alces-35	241	3	yearbook	yearbook	NOUN
alces-35	241	4	of	of	ADP
alces-35	241	5	forestry	forestry	NOUN
alces-35	241	6	.	.	PUNCT
alces-35	242	1	2006	2006	NUM
alces-35	242	2	.	.	PUNCT
alces-35	243	1	helsinki	helsinki	PROPN
alces-35	243	2	,	,	PUNCT
alces-35	243	3	finland	finland	PROPN
alces-35	243	4	.	.	PUNCT
alces-35	244	1	438	438	NUM
alces-35	244	2	pp	pp	ADJ
alces-35	244	3	.	.	PUNCT
alces-35	245	1	gill	gill	PROPN
alces-35	245	2	,	,	PUNCT
alces-35	245	3	r.	r.	PROPN
alces-35	245	4	m.	m.	PROPN
alces-35	245	5	a.	a.	PROPN
alces-35	245	6	1992	1992	NUM
alces-35	245	7	.	.	PUNCT
alces-35	246	1	a	a	DET
alces-35	246	2	review	review	NOUN
alces-35	246	3	of	of	ADP
alces-35	246	4	damage	damage	NOUN
alces-35	246	5	by	by	ADP
alces-35	246	6	mammals	mammal	NOUN
alces-35	246	7	in	in	ADP
alces-35	246	8	north	north	ADJ
alces-35	246	9	temperate	temperate	ADJ
alces-35	246	10	forests	forest	NOUN
alces-35	246	11	.	.	PUNCT
alces-35	247	1	3	3	X
alces-35	247	2	.	.	X
alces-35	247	3	impact	impact	NOUN
alces-35	247	4	on	on	ADP
alces-35	247	5	trees	tree	NOUN
alces-35	247	6	and	and	CCONJ
alces-35	247	7	forests	forest	NOUN
alces-35	247	8	.	.	PUNCT
alces-35	248	1	forestry	forestry	NOUN
alces-35	248	2	65:363	65:363	PROPN
alces-35	248	3	-	-	PUNCT
alces-35	248	4	388	388	NUM
alces-35	248	5	.	.	PUNCT
alces-35	249	1	härkönen	härkönen	PROPN
alces-35	249	2	.	.	PUNCT
alces-35	250	1	s.	s.	PROPN
alces-35	250	2	1998	1998	NUM
alces-35	250	3	.	.	PUNCT
alces-35	251	1	effects	effect	NOUN
alces-35	251	2	of	of	ADP
alces-35	251	3	silvicultural	silvicultural	ADJ
alces-35	251	4	cleaning	cleaning	NOUN
alces-35	251	5	in	in	ADP
alces-35	251	6	mixed	mixed	ADJ
alces-35	251	7	pine	pine	NOUN
alces-35	251	8	-	-	PUNCT
alces-35	251	9	deciduous	deciduous	ADJ
alces-35	251	10	stands	stand	VERB
alces-35	251	11	on	on	ADP
alces-35	251	12	moose	moose	NOUN
alces-35	251	13	damage	damage	NOUN
alces-35	251	14	to	to	ADP
alces-35	251	15	scots	scots	PROPN
alces-35	251	16	pine	pine	NOUN
alces-35	251	17	(	(	PUNCT
alces-35	251	18	pinus	pinus	NOUN
alces-35	251	19	sylvestris	sylvestris	NOUN
alces-35	251	20	)	)	PUNCT
alces-35	251	21	.	.	PUNCT
alces-35	252	1	scandinavian	scandinavian	ADJ
alces-35	252	2	journal	journal	PROPN
alces-35	252	3	of	of	ADP
alces-35	252	4	forest	forest	PROPN
alces-35	252	5	research	research	NOUN
alces-35	252	6	13:429	13:429	NUM
alces-35	252	7	-	-	SYM
alces-35	252	8	436	436	NUM
alces-35	252	9	.	.	PUNCT
alces-35	253	1	_	_	PUNCT
alces-35	254	1	_	_	PUNCT
alces-35	255	1	_	_	PUNCT
alces-35	256	1	_	_	PUNCT
alces-35	257	1	_	_	PUNCT
alces-35	257	2	,	,	PUNCT
alces-35	257	3	and	and	CCONJ
alces-35	257	4	r.	r.	PROPN
alces-35	257	5	heikkilä	heikkilä	PROPN
alces-35	257	6	.	.	PUNCT
alces-35	258	1	1999	1999	NUM
alces-35	258	2	.	.	PUNCT
alces-35	259	1	use	use	NOUN
alces-35	259	2	of	of	ADP
alces-35	259	3	pellet	pellet	NOUN
alces-35	259	4	group	group	NOUN
alces-35	259	5	counts	count	VERB
alces-35	259	6	in	in	ADP
alces-35	259	7	determining	determine	VERB
alces-35	259	8	density	density	NOUN
alces-35	259	9	and	and	CCONJ
alces-35	259	10	habitat	habitat	NOUN
alces-35	259	11	use	use	NOUN
alces-35	259	12	of	of	ADP
alces-35	259	13	moose	moose	NOUN
alces-35	259	14	alces	alce	NOUN
alces-35	259	15	alces	alce	NOUN
alces-35	259	16	in	in	ADP
alces-35	259	17	finland	finland	PROPN
alces-35	259	18	.	.	PUNCT
alces-35	260	1	wildlife	wildlife	NOUN
alces-35	260	2	biology	biology	NOUN
alces-35	260	3	5:233	5:233	PROPN
alces-35	260	4	-	-	SYM
alces-35	260	5	239	239	NUM
alces-35	260	6	.	.	PUNCT
alces-35	261	1	heikkilä	heikkilä	PROPN
alces-35	261	2	,	,	PUNCT
alces-35	261	3	r.	r.	PROPN
alces-35	261	4	,	,	PUNCT
alces-35	261	5	and	and	CCONJ
alces-35	261	6	s.	s.	PROPN
alces-35	261	7	härkönen	härkönen	PROPN
alces-35	261	8	.	.	PUNCT
alces-35	262	1	1998	1998	NUM
alces-35	262	2	.	.	PUNCT
alces-35	263	1	the	the	DET
alces-35	263	2	effects	effect	NOUN
alces-35	263	3	of	of	ADP
alces-35	263	4	salt	salt	NOUN
alces-35	263	5	stones	stone	NOUN
alces-35	263	6	on	on	ADP
alces-35	263	7	moose	moose	NOUN
alces-35	263	8	browsing	browse	VERB
alces-35	263	9	in	in	ADP
alces-35	263	10	managed	manage	VERB
alces-35	263	11	forests	forest	NOUN
alces-35	263	12	in	in	ADP
alces-35	263	13	finland	finland	PROPN
alces-35	263	14	.	.	PUNCT
alces-35	264	1	alces	alces	PROPN
alces-35	264	2	34:435	34:435	NUM
alces-35	264	3	-	-	PUNCT
alces-35	264	4	444	444	NUM
alces-35	264	5	.	.	PUNCT
alces-35	265	1	_	_	PUNCT
alces-35	266	1	_	_	PUNCT
alces-35	267	1	_	_	PUNCT
alces-35	268	1	_	_	PUNCT
alces-35	269	1	_	_	PUNCT
alces-35	269	2	,	,	PUNCT
alces-35	269	3	p.	p.	NOUN
alces-35	269	4	hokkanen	hokkanen	NOUN
alces-35	269	5	,	,	PUNCT
alces-35	269	6	m.	m.	NOUN
alces-35	269	7	kooiman	kooiman	PROPN
alces-35	269	8	,	,	PUNCT
alces-35	269	9	n.	n.	PROPN
alces-35	269	10	ayguney	ayguney	PROPN
alces-35	269	11	,	,	PUNCT
alces-35	269	12	and	and	CCONJ
alces-35	269	13	c.	c.	NOUN
alces-35	269	14	bassoulet	bassoulet	NOUN
alces-35	269	15	.	.	PUNCT
alces-35	270	1	2003	2003	NUM
alces-35	270	2	.	.	PUNCT
alces-35	271	1	the	the	DET
alces-35	271	2	impact	impact	NOUN
alces-35	271	3	of	of	ADP
alces-35	271	4	moose	moose	NOUN
alces-35	271	5	browsing	browse	VERB
alces-35	271	6	on	on	ADP
alces-35	271	7	tree	tree	NOUN
alces-35	271	8	species	specie	NOUN
alces-35	271	9	composition	composition	NOUN
alces-35	271	10	in	in	ADP
alces-35	271	11	finland	finland	PROPN
alces-35	271	12	.	.	PUNCT
alces-35	272	1	alces	alce	NOUN
alces-35	272	2	39:203	39:203	NUM
alces-35	272	3	-	-	SYM
alces-35	272	4	213	213	NUM
alces-35	272	5	.	.	PUNCT
alces-35	273	1	kalliola	kalliola	PROPN
alces-35	273	2	,	,	PUNCT
alces-35	273	3	r.	r.	PROPN
alces-35	273	4	1973	1973	NUM
alces-35	273	5	.	.	PUNCT
alces-35	274	1	suomen	suomen	PROPN
alces-35	274	2	kasvimaantiede	kasvimaantiede	PROPN
alces-35	274	3	.	.	PUNCT
alces-35	275	1	werner	werner	PROPN
alces-35	275	2	söderström	söderström	PROPN
alces-35	275	3	osakeyhtiö	osakeyhtiö	PROPN
alces-35	275	4	,	,	PUNCT
alces-35	275	5	porvoo	porvoo	NOUN
alces-35	275	6	,	,	PUNCT
alces-35	275	7	finland	finland	PROPN
alces-35	275	8	.	.	PUNCT
alces-35	276	1	308	308	NUM
alces-35	276	2	pp	pp	NOUN
alces-35	276	3	.	.	PUNCT
alces-35	277	1	(	(	PUNCT
alces-35	277	2	in	in	ADP
alces-35	277	3	finnish	finnish	ADJ
alces-35	277	4	)	)	PUNCT
alces-35	277	5	.	.	PUNCT
alces-35	278	1	kay	kay	PROPN
alces-35	278	2	,	,	PUNCT
alces-35	278	3	c.	c.	PROPN
alces-35	278	4	e.	e.	PROPN
alces-35	278	5	1997	1997	NUM
alces-35	278	6	.	.	PUNCT
alces-35	279	1	is	be	AUX
alces-35	279	2	aspen	aspen	NOUN
alces-35	279	3	doomed	doom	VERB
alces-35	279	4	?	?	PUNCT
alces-35	280	1	journal	journal	PROPN
alces-35	280	2	of	of	ADP
alces-35	280	3	forestry	forestry	PROPN
alces-35	280	4	95(5):4	95(5):4	PROPN
alces-35	280	5	-	-	PUNCT
alces-35	280	6	11	11	NUM
alces-35	280	7	.	.	PUNCT
alces-35	281	1	kaye	kaye	PROPN
alces-35	281	2	,	,	PUNCT
alces-35	281	3	m.	m.	PROPN
alces-35	281	4	w.	w.	PROPN
alces-35	281	5	,	,	PUNCT
alces-35	281	6	d.	d.	PROPN
alces-35	281	7	binkley	binkley	PROPN
alces-35	281	8	,	,	PUNCT
alces-35	281	9	and	and	CCONJ
alces-35	281	10	t.	t.	PROPN
alces-35	281	11	j.	j.	PROPN
alces-35	281	12	stohlgren	stohlgren	PROPN
alces-35	281	13	.	.	PUNCT
alces-35	282	1	2005	2005	NUM
alces-35	282	2	.	.	PUNCT
alces-35	283	1	effects	effect	NOUN
alces-35	283	2	of	of	ADP
alces-35	283	3	conifers	conifer	NOUN
alces-35	283	4	and	and	CCONJ
alces-35	283	5	elk	elk	NOUN
alces-35	283	6	browsing	browse	VERB
alces-35	283	7	on	on	ADP
alces-35	283	8	quaking	quake	VERB
alces-35	283	9	aspen	aspen	ADJ
alces-35	283	10	forests	forest	NOUN
alces-35	283	11	in	in	ADP
alces-35	283	12	the	the	DET
alces-35	283	13	central	central	ADJ
alces-35	283	14	rocky	rocky	ADJ
alces-35	283	15	mountains	mountain	NOUN
alces-35	283	16	,	,	PUNCT
alces-35	283	17	usa	usa	PROPN
alces-35	283	18	.	.	PROPN
alces-35	283	19	canadian	canadian	ADJ
alces-35	283	20	jour	jour	ADJ
alces-35	283	21	-	-	ADJ
alces-35	283	22	canadian	canadian	ADJ
alces-35	283	23	journal	journal	NOUN
alces-35	283	24	of	of	ADP
alces-35	283	25	forest	forest	PROPN
alces-35	283	26	research	research	NOUN
alces-35	283	27	15:1284	15:1284	NUM
alces-35	283	28	-	-	SYM
alces-35	283	29	1295	1295	NUM
alces-35	283	30	.	.	PUNCT
alces-35	284	1	kouki	kouki	PROPN
alces-35	284	2	,	,	PUNCT
alces-35	284	3	j.	j.	PROPN
alces-35	284	4	,	,	PUNCT
alces-35	284	5	k.	k.	PROPN
alces-35	284	6	arnold	arnold	PROPN
alces-35	284	7	,	,	PUNCT
alces-35	284	8	and	and	CCONJ
alces-35	284	9	p.	p.	PROPN
alces-35	284	10	martikainen	martikainen	NOUN
alces-35	284	11	.	.	PUNCT
alces-35	285	1	2004	2004	NUM
alces-35	285	2	.	.	PUNCT
alces-35	286	1	long	long	ADJ
alces-35	286	2	-	-	PUNCT
alces-35	286	3	term	term	NOUN
alces-35	286	4	persistence	persistence	NOUN
alces-35	286	5	of	of	ADP
alces-35	286	6	aspen	aspen	NOUN
alces-35	286	7	–	–	PUNCT
alces-35	286	8	a	a	DET
alces-35	286	9	key	key	ADJ
alces-35	286	10	host	host	NOUN
alces-35	286	11	for	for	ADP
alces-35	286	12	many	many	ADJ
alces-35	286	13	threatened	threaten	VERB
alces-35	286	14	species	specie	NOUN
alces-35	286	15	–	–	PUNCT
alces-35	286	16	is	be	AUX
alces-35	286	17	endangered	endanger	VERB
alces-35	286	18	in	in	ADP
alces-35	286	19	old	old	ADJ
alces-35	286	20	-	-	PUNCT
alces-35	286	21	growth	growth	NOUN
alces-35	286	22	conservation	conservation	NOUN
alces-35	286	23	areas	area	NOUN
alces-35	286	24	in	in	ADP
alces-35	286	25	finland	finland	PROPN
alces-35	286	26	.	.	PUNCT
alces-35	287	1	journal	journal	PROPN
alces-35	287	2	for	for	ADP
alces-35	287	3	nature	nature	NOUN
alces-35	287	4	conservation	conservation	NOUN
alces-35	287	5	12:41	12:41	NUM
alces-35	287	6	-	-	SYM
alces-35	287	7	52	52	NUM
alces-35	287	8	.	.	PUNCT
alces-35	288	1	latva	latva	NOUN
alces-35	288	2	-	-	PUNCT
alces-35	288	3	karjanmaa	karjanmaa	VERB
alces-35	288	4	,	,	PUNCT
alces-35	288	5	t.	t.	PROPN
alces-35	288	6	,	,	PUNCT
alces-35	288	7	r.	r.	PROPN
alces-35	288	8	penttilä	penttilä	PROPN
alces-35	288	9	,	,	PUNCT
alces-35	288	10	and	and	CCONJ
alces-35	288	11	j.	j.	PROPN
alces-35	288	12	siitonen	siitonen	PROPN
alces-35	288	13	.	.	PUNCT
alces-35	289	1	2007	2007	NUM
alces-35	289	2	.	.	PUNCT
alces-35	290	1	the	the	DET
alces-35	290	2	demographic	demographic	ADJ
alces-35	290	3	structure	structure	NOUN
alces-35	290	4	of	of	ADP
alces-35	290	5	european	european	PROPN
alces-35	290	6	aspen	aspen	PROPN
alces-35	290	7	(	(	PUNCT
alces-35	290	8	populus	populus	PROPN
alces-35	290	9	tremula	tremula	NOUN
alces-35	290	10	)	)	PUNCT
alces-35	290	11	populations	population	NOUN
alces-35	290	12	in	in	ADP
alces-35	290	13	managed	manage	VERB
alces-35	290	14	and	and	CCONJ
alces-35	290	15	oldgrowth	oldgrowth	VERB
alces-35	290	16	boreal	boreal	NOUN
alces-35	290	17	forests	forest	NOUN
alces-35	290	18	in	in	ADP
alces-35	290	19	eastern	eastern	PROPN
alces-35	290	20	finland	finland	PROPN
alces-35	290	21	.	.	PUNCT
alces-35	291	1	canadian	canadian	ADJ
alces-35	291	2	journal	journal	PROPN
alces-35	291	3	of	of	ADP
alces-35	291	4	forest	forest	PROPN
alces-35	291	5	research	research	PROPN
alces-35	291	6	37:1070	37:1070	NUM
alces-35	291	7	-	-	SYM
alces-35	291	8	1081	1081	NUM
alces-35	291	9	.	.	PUNCT
alces-35	292	1	mclaren	mclaren	PROPN
alces-35	292	2	,	,	PUNCT
alces-35	292	3	b.	b.	PROPN
alces-35	292	4	e.	e.	PROPN
alces-35	292	5	,	,	PUNCT
alces-35	292	6	b.	b.	PROPN
alces-35	292	7	a.	a.	PROPN
alces-35	292	8	roberts	roberts	PROPN
alces-35	292	9	,	,	PUNCT
alces-35	292	10	n.	n.	NOUN
alces-35	292	11	djanchékar	djanchékar	NOUN
alces-35	292	12	,	,	PUNCT
alces-35	292	13	and	and	CCONJ
alces-35	292	14	k.	k.	PROPN
alces-35	292	15	p.	p.	PROPN
alces-35	292	16	lewis	lewis	PROPN
alces-35	292	17	.	.	PUNCT
alces-35	292	18	2004	2004	NUM
alces-35	292	19	.	.	PUNCT
alces-35	293	1	effects	effect	NOUN
alces-35	293	2	of	of	ADP
alces-35	293	3	overabundant	overabundant	ADJ
alces-35	293	4	moose	moose	NOUN
alces-35	293	5	on	on	ADP
alces-35	293	6	the	the	DET
alces-35	293	7	newfoundland	newfoundland	PROPN
alces-35	293	8	landscape	landscape	NOUN
alces-35	293	9	.	.	PUNCT
alces-35	294	1	alces	alce	NOUN
alces-35	294	2	40:45	40:45	NUM
alces-35	294	3	-	-	SYM
alces-35	294	4	59	59	NUM
alces-35	294	5	.	.	PUNCT
alces-35	295	1	neff	neff	PROPN
alces-35	295	2	,	,	PUNCT
alces-35	295	3	d.	d.	PROPN
alces-35	295	4	j.	j.	PROPN
alces-35	295	5	1968	1968	NUM
alces-35	295	6	.	.	PUNCT
alces-35	296	1	the	the	DET
alces-35	296	2	pellet	pellet	NOUN
alces-35	296	3	-	-	PUNCT
alces-35	296	4	group	group	NOUN
alces-35	296	5	count	count	NOUN
alces-35	296	6	technique	technique	NOUN
alces-35	296	7	for	for	ADP
alces-35	296	8	big	big	ADJ
alces-35	296	9	game	game	NOUN
alces-35	296	10	trend	trend	NOUN
alces-35	296	11	,	,	PUNCT
alces-35	296	12	census	census	NOUN
alces-35	296	13	,	,	PUNCT
alces-35	296	14	and	and	CCONJ
alces-35	296	15	distribution	distribution	NOUN
alces-35	296	16	:	:	PUNCT
alces-35	296	17	a	a	DET
alces-35	296	18	review	review	NOUN
alces-35	296	19	.	.	PUNCT
alces-35	297	1	journal	journal	NOUN
alces-35	297	2	of	of	ADP
alces-35	297	3	wildlife	wildlife	NOUN
alces-35	297	4	management	management	NOUN
alces-35	297	5	32:597	32:597	NUM
alces-35	297	6	-	-	SYM
alces-35	297	7	614	614	NUM
alces-35	297	8	.	.	PUNCT
alces-35	298	1	piirainen	piirainen	ADJ
alces-35	298	2	,	,	PUNCT
alces-35	298	3	t.	t.	PROPN
alces-35	298	4	,	,	PUNCT
alces-35	298	5	m.	m.	NOUN
alces-35	298	6	honkamo	honkamo	NOUN
alces-35	298	7	,	,	PUNCT
alces-35	298	8	and	and	CCONJ
alces-35	298	9	s.	s.	PROPN
alces-35	298	10	rossi	rossi	PROPN
alces-35	298	11	.	.	PROPN
alces-35	299	1	1974	1974	NUM
alces-35	299	2	.	.	PUNCT
alces-35	300	1	a	a	DET
alces-35	300	2	preliminary	preliminary	ADJ
alces-35	300	3	report	report	NOUN
alces-35	300	4	of	of	ADP
alces-35	300	5	the	the	DET
alces-35	300	6	geology	geology	NOUN
alces-35	300	7	of	of	ADP
alces-35	300	8	the	the	DET
alces-35	300	9	koli	koli	PROPN
alces-35	300	10	area	area	PROPN
alces-35	300	11	.	.	PUNCT
alces-35	301	1	bulletin	bulletin	NOUN
alces-35	301	2	of	of	ADP
alces-35	301	3	the	the	DET
alces-35	301	4	geological	geological	ADJ
alces-35	301	5	society	society	NOUN
alces-35	301	6	of	of	ADP
alces-35	301	7	finland	finland	PROPN
alces-35	301	8	46:161	46:161	PROPN
alces-35	301	9	-	-	SYM
alces-35	301	10	166	166	NUM
alces-35	301	11	.	.	PUNCT
alces-35	302	1	romme	romme	PROPN
alces-35	302	2	,	,	PUNCT
alces-35	302	3	w.	w.	PROPN
alces-35	302	4	h.	h.	PROPN
alces-35	302	5	,	,	PUNCT
alces-35	302	6	m.	m.	PROPN
alces-35	302	7	g.	g.	PROPN
alces-35	302	8	turner	turner	PROPN
alces-35	302	9	,	,	PUNCT
alces-35	302	10	g.	g.	PROPN
alces-35	302	11	a.	a.	PROPN
alces-35	302	12	tuskan	tuskan	PROPN
alces-35	302	13	,	,	PUNCT
alces-35	302	14	and	and	CCONJ
alces-35	302	15	r.	r.	PROPN
alces-35	302	16	a.	a.	PROPN
alces-35	302	17	reed	reed	PROPN
alces-35	302	18	.	.	PUNCT
alces-35	303	1	2005	2005	NUM
alces-35	303	2	.	.	PUNCT
alces-35	304	1	establishment	establishment	NOUN
alces-35	304	2	,	,	PUNCT
alces-35	304	3	persistence	persistence	NOUN
alces-35	304	4	,	,	PUNCT
alces-35	304	5	and	and	CCONJ
alces-35	304	6	growth	growth	NOUN
alces-35	304	7	of	of	ADP
alces-35	304	8	aspen	aspen	PROPN
alces-35	304	9	(	(	PUNCT
alces-35	304	10	populus	populus	ADJ
alces-35	304	11	tremuloides	tremuloide	NOUN
alces-35	304	12	)	)	PUNCT
alces-35	304	13	seedlings	seedling	NOUN
alces-35	304	14	in	in	ADP
alces-35	304	15	yellowstone	yellowstone	PROPN
alces-35	304	16	national	national	PROPN
alces-35	304	17	park	park	PROPN
alces-35	304	18	.	.	PUNCT
alces-35	305	1	ecology	ecology	NOUN
alces-35	305	2	86:404	86:404	PROPN
alces-35	305	3	-	-	PUNCT
alces-35	305	4	418	418	NUM
alces-35	305	5	.	.	PUNCT
alces-35	306	1	_	_	PUNCT
alces-35	307	1	_	_	PUNCT
alces-35	308	1	_	_	PUNCT
alces-35	309	1	_	_	PUNCT
alces-35	310	1	_	_	PUNCT
alces-35	310	2	,	,	PUNCT
alces-35	311	1	_	_	PUNCT
alces-35	312	1	_	_	PUNCT
alces-35	313	1	_	_	PUNCT
alces-35	314	1	_	_	PUNCT
alces-35	315	1	_	_	PUNCT
alces-35	315	2	,	,	PUNCT
alces-35	315	3	l.	l.	PROPN
alces-35	315	4	l.	l.	PROPN
alces-35	315	5	wallace	wallace	PROPN
alces-35	315	6	,	,	PUNCT
alces-35	315	7	and	and	CCONJ
alces-35	315	8	j.	j.	PROPN
alces-35	315	9	s.	s.	PROPN
alces-35	315	10	walker	walker	PROPN
alces-35	315	11	.	.	PROPN
alces-35	315	12	1995	1995	NUM
alces-35	315	13	.	.	PUNCT
alces-35	316	1	aspen	aspen	PROPN
alces-35	316	2	,	,	PUNCT
alces-35	316	3	elk	elk	NOUN
alces-35	316	4	,	,	PUNCT
alces-35	316	5	and	and	CCONJ
alces-35	316	6	fire	fire	NOUN
alces-35	316	7	in	in	ADP
alces-35	316	8	moose	moose	NOUN
alces-35	316	9	and	and	CCONJ
alces-35	316	10	aspen	aspen	NOUN
alces-35	316	11	regeneration	regeneration	NOUN
alces-35	316	12	in	in	ADP
alces-35	316	13	finland	finland	PROPN
alces-35	316	14	härkönen	härkönen	PROPN
alces-35	316	15	et	et	PROPN
alces-35	316	16	al	al	PROPN
alces-35	316	17	.	.	PROPN
alces-35	317	1	alces	alce	NOUN
alces-35	317	2	vol	vol	NOUN
alces-35	317	3	.	.	PROPN
alces-35	317	4	44	44	NUM
alces-35	317	5	,	,	PUNCT
alces-35	317	6	2008	2008	NUM
alces-35	317	7	40	40	NUM
alces-35	317	8	northern	northern	ADJ
alces-35	317	9	yellowstone	yellowstone	PROPN
alces-35	317	10	park	park	NOUN
alces-35	317	11	.	.	PUNCT
alces-35	318	1	ecology	ecology	NOUN
alces-35	318	2	76:2097	76:2097	PROPN
alces-35	318	3	-	-	PROPN
alces-35	318	4	2106	2106	NUM
alces-35	318	5	.	.	PUNCT
alces-35	319	1	rooney	rooney	NOUN
alces-35	319	2	,	,	PUNCT
alces-35	319	3	t.	t.	PROPN
alces-35	319	4	p.	p.	PROPN
alces-35	319	5	,	,	PUNCT
alces-35	319	6	and	and	CCONJ
alces-35	319	7	d.	d.	PROPN
alces-35	319	8	m.	m.	PROPN
alces-35	319	9	waller	waller	PROPN
alces-35	319	10	.	.	PUNCT
alces-35	320	1	2003	2003	NUM
alces-35	320	2	.	.	PUNCT
alces-35	321	1	direct	direct	ADJ
alces-35	321	2	and	and	CCONJ
alces-35	321	3	indirect	indirect	ADJ
alces-35	321	4	effects	effect	NOUN
alces-35	321	5	of	of	ADP
alces-35	321	6	white	white	ADJ
alces-35	321	7	-	-	PUNCT
alces-35	321	8	tailed	tail	VERB
alces-35	321	9	deer	deer	NOUN
alces-35	321	10	in	in	ADP
alces-35	321	11	forest	forest	NOUN
alces-35	321	12	ecosystems	ecosystem	NOUN
alces-35	321	13	.	.	PUNCT
alces-35	322	1	forest	forest	NOUN
alces-35	322	2	ecology	ecology	NOUN
alces-35	322	3	and	and	CCONJ
alces-35	322	4	management	management	NOUN
alces-35	322	5	181:165	181:165	PROPN
alces-35	322	6	-	-	NOUN
alces-35	322	7	176	176	NUM
alces-35	322	8	.	.	PUNCT
alces-35	322	9	siitonen	siitonen	NOUN
alces-35	322	10	,	,	PUNCT
alces-35	322	11	j.	j.	PROPN
alces-35	322	12	1999	1999	NUM
alces-35	322	13	.	.	PUNCT
alces-35	323	1	haavan	haavan	PROPN
alces-35	323	2	merkitys	merkitys	PROPN
alces-35	323	3	metsäluonnon	metsäluonnon	PROPN
alces-35	323	4	monimuotoisuudelle	monimuotoisuudelle	PROPN
alces-35	323	5	.	.	PUNCT
alces-35	324	1	pages	page	NOUN
alces-35	324	2	71	71	NUM
alces-35	324	3	-	-	SYM
alces-35	324	4	82	82	NUM
alces-35	324	5	in	in	ADP
alces-35	324	6	j.	j.	PROPN
alces-35	324	7	hynynen	hynynen	PROPN
alces-35	324	8	and	and	CCONJ
alces-35	324	9	a.	a.	PROPN
alces-35	324	10	viherä	viherä	PROPN
alces-35	324	11	-	-	PUNCT
alces-35	324	12	aarnio	aarnio	PROPN
alces-35	324	13	(	(	PUNCT
alces-35	324	14	eds	ed	NOUN
alces-35	324	15	.	.	PUNCT
alces-35	324	16	)	)	PUNCT
alces-35	325	1	haapa	haapa	ADJ
alces-35	325	2	–	–	PUNCT
alces-35	325	3	monimuotoisuutta	monimuotoisuutta	NOUN
alces-35	325	4	metsään	metsään	NOUN
alces-35	325	5	ja	ja	PROPN
alces-35	325	6	metsätalouteen	metsätalouteen	PROPN
alces-35	325	7	.	.	PUNCT
alces-35	326	1	vantaan	vantaan	PROPN
alces-35	326	2	tutkimuskeskuksen	tutkimuskeskuksen	VERB
alces-35	326	3	tutkimuspäivä	tutkimuspäivä	ADJ
alces-35	326	4	tammisaaressa	tammisaaressa	NOUN
alces-35	326	5	.	.	PUNCT
alces-35	327	1	tammisaari	tammisaari	PROPN
alces-35	327	2	,	,	PUNCT
alces-35	327	3	finland	finland	PROPN
alces-35	327	4	,	,	PUNCT
alces-35	327	5	november	november	PROPN
alces-35	327	6	12	12	NUM
alces-35	327	7	,	,	PUNCT
alces-35	327	8	1998	1998	NUM
alces-35	327	9	.	.	PUNCT
alces-35	328	1	(	(	PUNCT
alces-35	328	2	in	in	ADP
alces-35	328	3	finnish	finnish	PROPN
alces-35	328	4	)	)	PUNCT
alces-35	328	5	.	.	PUNCT
alces-35	329	1	suzuki	suzuki	PROPN
alces-35	329	2	,	,	PUNCT
alces-35	329	3	k.	k.	PROPN
alces-35	329	4	,	,	PUNCT
alces-35	329	5	h.	h.	PROPN
alces-35	329	6	suzuki	suzuki	PROPN
alces-35	329	7	,	,	PUNCT
alces-35	329	8	d.	d.	PROPN
alces-35	329	9	binkley	binkley	PROPN
alces-35	329	10	,	,	PUNCT
alces-35	329	11	and	and	CCONJ
alces-35	329	12	t.	t.	PROPN
alces-35	329	13	j.	j.	PROPN
alces-35	329	14	stohlgren	stohlgren	PROPN
alces-35	329	15	.	.	PUNCT
alces-35	330	1	1999	1999	NUM
alces-35	330	2	.	.	PUNCT
alces-35	331	1	aspen	aspen	NOUN
alces-35	331	2	regeneration	regeneration	NOUN
alces-35	331	3	in	in	ADP
alces-35	331	4	the	the	DET
alces-35	331	5	colorado	colorado	NOUN
alces-35	331	6	front	front	ADJ
alces-35	331	7	range	range	NOUN
alces-35	331	8	:	:	PUNCT
alces-35	331	9	differences	difference	NOUN
alces-35	331	10	at	at	ADP
alces-35	331	11	local	local	ADJ
alces-35	331	12	and	and	CCONJ
alces-35	331	13	landscape	landscape	NOUN
alces-35	331	14	scales	scale	NOUN
alces-35	331	15	.	.	PUNCT
alces-35	332	1	landscape	landscape	NOUN
alces-35	332	2	ecology	ecology	NOUN
alces-35	332	3	14:231	14:231	PROPN
alces-35	332	4	-	-	SYM
alces-35	332	5	237	237	NUM
alces-35	332	6	.	.	PUNCT
alces-35	333	1	syrjänen	syrjänen	PROPN
alces-35	333	2	,	,	PUNCT
alces-35	333	3	k.	k.	PROPN
alces-35	333	4	,	,	PUNCT
alces-35	333	5	r.	r.	PROPN
alces-35	333	6	kalliola	kalliola	PROPN
alces-35	333	7	,	,	PUNCT
alces-35	333	8	a.	a.	NOUN
alces-35	333	9	puolasmaa	puolasmaa	NOUN
alces-35	333	10	,	,	PUNCT
alces-35	333	11	and	and	CCONJ
alces-35	333	12	j.	j.	PROPN
alces-35	333	13	mattsson	mattsson	PROPN
alces-35	333	14	.	.	PROPN
alces-35	333	15	1994	1994	NUM
alces-35	333	16	.	.	PUNCT
alces-35	334	1	landscape	landscape	NOUN
alces-35	334	2	structure	structure	NOUN
alces-35	334	3	and	and	CCONJ
alces-35	334	4	forest	forest	NOUN
alces-35	334	5	dynamics	dynamic	NOUN
alces-35	334	6	in	in	ADP
alces-35	334	7	subcontinental	subcontinental	ADJ
alces-35	334	8	russian	russian	ADJ
alces-35	334	9	european	european	ADJ
alces-35	334	10	taiga	taiga	NOUN
alces-35	334	11	.	.	PUNCT
alces-35	335	1	annales	annales	PROPN
alces-35	335	2	�	�	PROPN
alces-35	335	3	oo	oo	ADJ
alces-35	335	4	-	-	ADJ
alces-35	335	5	ian	ian	ADJ
alces-35	335	6	european	european	ADJ
alces-35	335	7	taiga	taiga	NOUN
alces-35	335	8	.	.	PUNCT
alces-35	336	1	annales	annales	PROPN
alces-35	336	2	�	�	PROPN
alces-35	336	3	oologici	oologici	PROPN
alces-35	336	4	fennici	fennici	PROPN
alces-35	336	5	31:19	31:19	NUM
alces-35	336	6	-	-	SYM
alces-35	336	7	34	34	NUM
alces-35	336	8	.	.	PUNCT
alces-35	336	9	tikka	tikka	NOUN
alces-35	336	10	,	,	PUNCT
alces-35	336	11	p.	p.	PROPN
alces-35	336	12	s.	s.	PROPN
alces-35	337	1	1955	1955	NUM
alces-35	337	2	.	.	PUNCT
alces-35	338	1	structure	structure	NOUN
alces-35	338	2	and	and	CCONJ
alces-35	338	3	quality	quality	NOUN
alces-35	338	4	of	of	ADP
alces-35	338	5	aspen	aspen	PROPN
alces-35	338	6	stands	stand	NOUN
alces-35	338	7	.	.	PUNCT
alces-35	339	1	i.	i.	PROPN
alces-35	339	2	structure	structure	PROPN
alces-35	339	3	.	.	PUNCT
alces-35	340	1	finnish	finnish	PROPN
alces-35	340	2	forest	forest	PROPN
alces-35	340	3	research	research	PROPN
alces-35	340	4	institute	institute	PROPN
alces-35	340	5	44(4):1	44(4):1	PROPN
alces-35	340	6	-	-	PUNCT
alces-35	340	7	33	33	NUM
alces-35	340	8	.	.	PUNCT
alces-35	341	1	(	(	PUNCT
alces-35	341	2	in	in	ADP
alces-35	341	3	finnish	finnish	PROPN
alces-35	341	4	with	with	ADP
alces-35	341	5	english	english	PROPN
alces-35	341	6	summary	summary	NOUN
alces-35	341	7	)	)	PUNCT
alces-35	341	8	.	.	PUNCT