id	sid	tid	token	lemma	pos
alces-443	1	1	4013.p65	4013.p65	NUM
alces-443	1	2	alces	alce	NOUN
alces-443	1	3	vol	vol	NOUN
alces-443	1	4	.	.	PROPN
alces-443	1	5	40	40	NUM
alces-443	1	6	,	,	PUNCT
alces-443	1	7	2004	2004	NUM
alces-443	1	8	moose	moose	NOUN
alces-443	1	9	genetics	genetic	NOUN
alces-443	1	10	hundertmark	hundertmark	NOUN
alces-443	1	11	and	and	CCONJ
alces-443	1	12	bowyer	bowyer	PROPN
alces-443	1	13	103	103	NUM
alces-443	1	14	genetics	genetic	NOUN
alces-443	1	15	,	,	PUNCT
alces-443	1	16	evolution	evolution	NOUN
alces-443	1	17	,	,	PUNCT
alces-443	1	18	and	and	CCONJ
alces-443	1	19	phylogeography	phylogeography	NOUN
alces-443	1	20	of	of	ADP
alces-443	1	21	moose	moose	PROPN
alces-443	1	22	kris	kris	PROPN
alces-443	1	23	j.	j.	PROPN
alces-443	1	24	hundertmark1	hundertmark1	PROPN
alces-443	1	25	and	and	CCONJ
alces-443	1	26	r.	r.	PROPN
alces-443	1	27	terry	terry	PROPN
alces-443	1	28	bowyer2	bowyer2	VERB
alces-443	2	1	1institute	1institute	NUM
alces-443	2	2	of	of	ADP
alces-443	2	3	arctic	arctic	ADJ
alces-443	2	4	biology	biology	NOUN
alces-443	2	5	and	and	CCONJ
alces-443	2	6	department	department	NOUN
alces-443	2	7	of	of	ADP
alces-443	2	8	biology	biology	NOUN
alces-443	2	9	and	and	CCONJ
alces-443	2	10	wildlife	wildlife	NOUN
alces-443	2	11	,	,	PUNCT
alces-443	2	12	university	university	NOUN
alces-443	2	13	of	of	ADP
alces-443	2	14	alaska	alaska	PROPN
alces-443	2	15	fairbanks	fairbanks	PROPN
alces-443	2	16	,	,	PUNCT
alces-443	2	17	fairbanks	fairbanks	PROPN
alces-443	2	18	,	,	PUNCT
alces-443	2	19	ak	ak	PROPN
alces-443	2	20	99775	99775	NUM
alces-443	2	21	,	,	PUNCT
alces-443	2	22	usa	usa	PROPN
alces-443	2	23	;	;	PUNCT
alces-443	2	24	2department	2department	NUM
alces-443	2	25	of	of	ADP
alces-443	2	26	biological	biological	ADJ
alces-443	2	27	sciences	science	NOUN
alces-443	2	28	,	,	PUNCT
alces-443	2	29	idaho	idaho	PROPN
alces-443	2	30	state	state	PROPN
alces-443	2	31	university	university	PROPN
alces-443	2	32	,	,	PUNCT
alces-443	2	33	pocatello	pocatello	PROPN
alces-443	2	34	,	,	PUNCT
alces-443	2	35	i	i	PROPN
alces-443	2	36	d	d	PROPN
alces-443	2	37	83209	83209	NUM
alces-443	2	38	,	,	PUNCT
alces-443	2	39	usa	usa	PROPN
alces-443	2	40	abstract	abstract	PROPN
alces-443	2	41	:	:	PUNCT
alces-443	2	42	early	early	ADJ
alces-443	2	43	studies	study	NOUN
alces-443	2	44	of	of	ADP
alces-443	2	45	genetic	genetic	ADJ
alces-443	2	46	variation	variation	NOUN
alces-443	2	47	in	in	ADP
alces-443	2	48	moose	moose	NOUN
alces-443	2	49	(	(	PUNCT
alces-443	2	50	alces	alce	NOUN
alces-443	2	51	alces	alce	NOUN
alces-443	2	52	)	)	PUNCT
alces-443	2	53	indicated	indicate	VERB
alces-443	2	54	little	little	ADJ
alces-443	2	55	variation	variation	NOUN
alces-443	2	56	.	.	PUNCT
alces-443	3	1	recent	recent	ADJ
alces-443	3	2	studies	study	NOUN
alces-443	3	3	have	have	AUX
alces-443	3	4	indicated	indicate	VERB
alces-443	3	5	higher	high	ADJ
alces-443	3	6	levels	level	NOUN
alces-443	3	7	of	of	ADP
alces-443	3	8	variation	variation	NOUN
alces-443	3	9	in	in	ADP
alces-443	3	10	nuclear	nuclear	ADJ
alces-443	3	11	markers	marker	NOUN
alces-443	3	12	;	;	PUNCT
alces-443	3	13	nonetheless	nonetheless	ADV
alces-443	3	14	,	,	PUNCT
alces-443	3	15	genetic	genetic	ADJ
alces-443	3	16	heterogeneity	heterogeneity	NOUN
alces-443	3	17	of	of	ADP
alces-443	3	18	moose	moose	NOUN
alces-443	3	19	is	be	AUX
alces-443	3	20	relatively	relatively	ADV
alces-443	3	21	low	low	ADJ
alces-443	3	22	compared	compare	VERB
alces-443	3	23	with	with	ADP
alces-443	3	24	other	other	ADJ
alces-443	3	25	mammals	mammal	NOUN
alces-443	3	26	.	.	PUNCT
alces-443	4	1	similarly	similarly	ADV
alces-443	4	2	,	,	PUNCT
alces-443	4	3	variation	variation	NOUN
alces-443	4	4	in	in	ADP
alces-443	4	5	mitochondrial	mitochondrial	ADJ
alces-443	4	6	dna	dna	NOUN
alces-443	4	7	of	of	ADP
alces-443	4	8	moose	moose	NOUN
alces-443	4	9	is	be	AUX
alces-443	4	10	limited	limit	VERB
alces-443	4	11	worldwide	worldwide	ADV
alces-443	4	12	,	,	PUNCT
alces-443	4	13	indicating	indicate	VERB
alces-443	4	14	low	low	ADJ
alces-443	4	15	historic	historic	ADJ
alces-443	4	16	effective	effective	ADJ
alces-443	4	17	population	population	NOUN
alces-443	4	18	size	size	NOUN
alces-443	4	19	and	and	CCONJ
alces-443	4	20	a	a	DET
alces-443	4	21	common	common	ADJ
alces-443	4	22	ancestry	ancestry	NOUN
alces-443	4	23	for	for	ADP
alces-443	4	24	moose	moose	NOUN
alces-443	4	25	within	within	ADP
alces-443	4	26	the	the	DET
alces-443	4	27	last	last	ADJ
alces-443	4	28	60,000	60,000	NUM
alces-443	4	29	years	year	NOUN
alces-443	4	30	.	.	PUNCT
alces-443	5	1	that	that	DET
alces-443	5	2	ancestor	ancestor	NOUN
alces-443	5	3	most	most	ADV
alces-443	5	4	likely	likely	ADV
alces-443	5	5	lived	live	VERB
alces-443	5	6	in	in	ADP
alces-443	5	7	central	central	ADJ
alces-443	5	8	asia	asia	PROPN
alces-443	5	9	.	.	PUNCT
alces-443	6	1	moose	moose	PROPN
alces-443	6	2	likely	likely	ADV
alces-443	6	3	exhibit	exhibit	VERB
alces-443	6	4	low	low	ADJ
alces-443	6	5	levels	level	NOUN
alces-443	6	6	of	of	ADP
alces-443	6	7	heterogeneity	heterogeneity	NOUN
alces-443	6	8	because	because	SCONJ
alces-443	6	9	of	of	ADP
alces-443	6	10	population	population	NOUN
alces-443	6	11	bottlenecks	bottleneck	NOUN
alces-443	6	12	in	in	ADP
alces-443	6	13	the	the	DET
alces-443	6	14	late	late	ADJ
alces-443	6	15	pleistocene	pleistocene	NOUN
alces-443	6	16	caused	cause	VERB
alces-443	6	17	by	by	ADP
alces-443	6	18	latitudinal	latitudinal	ADJ
alces-443	6	19	shifts	shift	NOUN
alces-443	6	20	in	in	ADP
alces-443	6	21	habitat	habitat	NOUN
alces-443	6	22	from	from	ADP
alces-443	6	23	recurrent	recurrent	ADJ
alces-443	6	24	climate	climate	NOUN
alces-443	6	25	reversals	reversal	NOUN
alces-443	6	26	.	.	PUNCT
alces-443	7	1	a	a	DET
alces-443	7	2	northward	northward	ADJ
alces-443	7	3	movement	movement	NOUN
alces-443	7	4	of	of	ADP
alces-443	7	5	boreal	boreal	NOUN
alces-443	7	6	forest	forest	NOUN
alces-443	7	7	associated	associate	VERB
alces-443	7	8	with	with	ADP
alces-443	7	9	the	the	DET
alces-443	7	10	end	end	NOUN
alces-443	7	11	of	of	ADP
alces-443	7	12	the	the	DET
alces-443	7	13	last	last	ADJ
alces-443	7	14	ice	ice	NOUN
alces-443	7	15	age	age	NOUN
alces-443	7	16	facilitated	facilitate	VERB
alces-443	7	17	the	the	DET
alces-443	7	18	northward	northward	ADJ
alces-443	7	19	advance	advance	NOUN
alces-443	7	20	of	of	ADP
alces-443	7	21	asian	asian	ADJ
alces-443	7	22	populations	population	NOUN
alces-443	7	23	and	and	CCONJ
alces-443	7	24	colonization	colonization	NOUN
alces-443	7	25	of	of	ADP
alces-443	7	26	the	the	DET
alces-443	7	27	new	new	ADJ
alces-443	7	28	world	world	NOUN
alces-443	7	29	,	,	PUNCT
alces-443	7	30	which	which	PRON
alces-443	7	31	occurred	occur	VERB
alces-443	7	32	as	as	ADP
alces-443	7	33	a	a	DET
alces-443	7	34	single	single	ADJ
alces-443	7	35	entry	entry	NOUN
alces-443	7	36	by	by	ADP
alces-443	7	37	relatively	relatively	ADV
alces-443	7	38	few	few	ADJ
alces-443	7	39	moose	moose	NOUN
alces-443	7	40	immediately	immediately	ADV
alces-443	7	41	prior	prior	ADV
alces-443	7	42	to	to	ADP
alces-443	7	43	the	the	DET
alces-443	7	44	last	last	ADJ
alces-443	7	45	flooding	flooding	NOUN
alces-443	7	46	of	of	ADP
alces-443	7	47	the	the	DET
alces-443	7	48	bering	bering	NOUN
alces-443	7	49	land	land	NOUN
alces-443	7	50	bridge	bridge	NOUN
alces-443	7	51	.	.	PUNCT
alces-443	8	1	despite	despite	SCONJ
alces-443	8	2	suffering	suffer	VERB
alces-443	8	3	serial	serial	ADJ
alces-443	8	4	population	population	NOUN
alces-443	8	5	bottlenecks	bottleneck	NOUN
alces-443	8	6	historically	historically	ADV
alces-443	8	7	,	,	PUNCT
alces-443	8	8	moose	moose	NOUN
alces-443	8	9	have	have	AUX
alces-443	8	10	exhibited	exhibit	VERB
alces-443	8	11	a	a	DET
alces-443	8	12	notable	notable	ADJ
alces-443	8	13	ability	ability	NOUN
alces-443	8	14	to	to	PART
alces-443	8	15	adapt	adapt	VERB
alces-443	8	16	to	to	ADP
alces-443	8	17	a	a	DET
alces-443	8	18	changing	change	VERB
alces-443	8	19	environment	environment	NOUN
alces-443	8	20	,	,	PUNCT
alces-443	8	21	indicating	indicate	VERB
alces-443	8	22	that	that	SCONJ
alces-443	8	23	limited	limited	ADJ
alces-443	8	24	neutral	neutral	ADJ
alces-443	8	25	genetic	genetic	ADJ
alces-443	8	26	variation	variation	NOUN
alces-443	8	27	may	may	AUX
alces-443	8	28	not	not	PART
alces-443	8	29	indicate	indicate	VERB
alces-443	8	30	limited	limited	ADJ
alces-443	8	31	adaptive	adaptive	ADJ
alces-443	8	32	genetic	genetic	ADJ
alces-443	8	33	variation	variation	NOUN
alces-443	8	34	.	.	PUNCT
alces-443	9	1	we	we	PRON
alces-443	9	2	conclude	conclude	VERB
alces-443	9	3	that	that	SCONJ
alces-443	9	4	morphological	morphological	ADJ
alces-443	9	5	variation	variation	NOUN
alces-443	9	6	among	among	ADP
alces-443	9	7	moose	moose	NOUN
alces-443	9	8	worldwide	worldwide	ADV
alces-443	9	9	occurred	occur	VERB
alces-443	9	10	within	within	ADP
alces-443	9	11	a	a	DET
alces-443	9	12	few	few	ADJ
alces-443	9	13	thousand	thousand	NUM
alces-443	9	14	years	year	NOUN
alces-443	9	15	and	and	CCONJ
alces-443	9	16	indicates	indicate	VERB
alces-443	9	17	that	that	PRON
alces-443	9	18	moose	moose	NOUN
alces-443	9	19	underwent	underwent	NOUN
alces-443	9	20	episodes	episode	NOUN
alces-443	9	21	of	of	ADP
alces-443	9	22	rapid	rapid	ADJ
alces-443	9	23	and	and	CCONJ
alces-443	9	24	occasionally	occasionally	ADV
alces-443	9	25	convergent	convergent	ADJ
alces-443	9	26	evolution	evolution	NOUN
alces-443	9	27	.	.	PUNCT
alces-443	10	1	genetic	genetic	ADJ
alces-443	10	2	change	change	NOUN
alces-443	10	3	in	in	ADP
alces-443	10	4	moose	moose	NOUN
alces-443	10	5	populations	population	NOUN
alces-443	10	6	over	over	ADP
alces-443	10	7	very	very	ADV
alces-443	10	8	short	short	ADJ
alces-443	10	9	time	time	NOUN
alces-443	10	10	scales	scale	NOUN
alces-443	10	11	(	(	PUNCT
alces-443	10	12	tens	ten	NOUN
alces-443	10	13	or	or	CCONJ
alces-443	10	14	hundreds	hundred	NOUN
alces-443	10	15	of	of	ADP
alces-443	10	16	years	year	NOUN
alces-443	10	17	)	)	PUNCT
alces-443	10	18	is	be	AUX
alces-443	10	19	possible	possible	ADJ
alces-443	10	20	under	under	ADP
alces-443	10	21	harvest	harvest	NOUN
alces-443	10	22	management	management	NOUN
alces-443	10	23	regimes	regime	NOUN
alces-443	10	24	and	and	CCONJ
alces-443	10	25	those	those	DET
alces-443	10	26	changes	change	NOUN
alces-443	10	27	may	may	AUX
alces-443	10	28	not	not	PART
alces-443	10	29	be	be	AUX
alces-443	10	30	beneficial	beneficial	ADJ
alces-443	10	31	to	to	AUX
alces-443	10	32	moose	moose	VERB
alces-443	10	33	in	in	ADP
alces-443	10	34	the	the	DET
alces-443	10	35	long	long	ADJ
alces-443	10	36	term	term	NOUN
alces-443	10	37	.	.	PUNCT
alces-443	11	1	modeling	model	VERB
alces-443	11	2	exercises	exercise	NOUN
alces-443	11	3	have	have	AUX
alces-443	11	4	demonstrated	demonstrate	VERB
alces-443	11	5	that	that	SCONJ
alces-443	11	6	harvest	harvest	NOUN
alces-443	11	7	strategies	strategy	NOUN
alces-443	11	8	can	can	AUX
alces-443	11	9	have	have	VERB
alces-443	11	10	negative	negative	ADJ
alces-443	11	11	consequences	consequence	NOUN
alces-443	11	12	on	on	ADP
alces-443	11	13	neutral	neutral	ADJ
alces-443	11	14	genetic	genetic	ADJ
alces-443	11	15	variation	variation	NOUN
alces-443	11	16	as	as	ADV
alces-443	11	17	well	well	ADV
alces-443	11	18	as	as	ADP
alces-443	11	19	alleles	allele	NOUN
alces-443	11	20	underpinning	underpin	VERB
alces-443	11	21	fitness	fitness	NOUN
alces-443	11	22	traits	trait	NOUN
alces-443	11	23	.	.	PUNCT
alces-443	12	1	biologists	biologist	NOUN
alces-443	12	2	should	should	AUX
alces-443	12	3	consider	consider	VERB
alces-443	12	4	such	such	ADJ
alces-443	12	5	outcomes	outcome	NOUN
alces-443	12	6	when	when	SCONJ
alces-443	12	7	evaluating	evaluate	VERB
alces-443	12	8	management	management	NOUN
alces-443	12	9	options	option	NOUN
alces-443	12	10	.	.	PUNCT
alces-443	13	1	alces	alce	NOUN
alces-443	13	2	vol	vol	NOUN
alces-443	13	3	.	.	PROPN
alces-443	14	1	40	40	NUM
alces-443	14	2	:	:	PUNCT
alces-443	14	3	103	103	NUM
alces-443	14	4	-	-	SYM
alces-443	14	5	122	122	NUM
alces-443	14	6	(	(	PUNCT
alces-443	14	7	2004	2004	NUM
alces-443	14	8	)	)	PUNCT
alces-443	14	9	key	key	ADJ
alces-443	14	10	words	word	NOUN
alces-443	14	11	:	:	PUNCT
alces-443	14	12	adaptation	adaptation	NOUN
alces-443	14	13	,	,	PUNCT
alces-443	14	14	alces	alce	NOUN
alces-443	14	15	,	,	PUNCT
alces-443	14	16	convergent	convergent	NOUN
alces-443	14	17	evolution	evolution	NOUN
alces-443	14	18	,	,	PUNCT
alces-443	14	19	moose	moose	NOUN
alces-443	14	20	,	,	PUNCT
alces-443	14	21	mtdna	mtdna	ADV
alces-443	14	22	,	,	PUNCT
alces-443	14	23	phylogeography	phylogeography	NOUN
alces-443	14	24	,	,	PUNCT
alces-443	14	25	pleistocene	pleistocene	NOUN
alces-443	14	26	,	,	PUNCT
alces-443	14	27	range	range	NOUN
alces-443	14	28	expansion	expansion	NOUN
alces-443	14	29	genetics	genetic	NOUN
alces-443	14	30	have	have	AUX
alces-443	14	31	long	long	ADV
alces-443	14	32	had	have	VERB
alces-443	14	33	a	a	DET
alces-443	14	34	central	central	ADJ
alces-443	14	35	role	role	NOUN
alces-443	14	36	in	in	ADP
alces-443	14	37	biological	biological	ADJ
alces-443	14	38	investigations	investigation	NOUN
alces-443	14	39	,	,	PUNCT
alces-443	14	40	and	and	CCONJ
alces-443	14	41	provide	provide	VERB
alces-443	14	42	analytical	analytical	ADJ
alces-443	14	43	tools	tool	NOUN
alces-443	14	44	that	that	PRON
alces-443	14	45	are	be	AUX
alces-443	14	46	applicable	applicable	ADJ
alces-443	14	47	across	across	ADP
alces-443	14	48	a	a	DET
alces-443	14	49	broad	broad	ADJ
alces-443	14	50	spectrum	spectrum	NOUN
alces-443	14	51	of	of	ADP
alces-443	14	52	investigation	investigation	NOUN
alces-443	14	53	.	.	PUNCT
alces-443	15	1	for	for	ADP
alces-443	15	2	instance	instance	NOUN
alces-443	15	3	,	,	PUNCT
alces-443	15	4	genetic	genetic	ADJ
alces-443	15	5	analysis	analysis	NOUN
alces-443	15	6	can	can	AUX
alces-443	15	7	provide	provide	VERB
alces-443	15	8	insights	insight	NOUN
alces-443	15	9	into	into	ADP
alces-443	15	10	such	such	ADJ
alces-443	15	11	diverse	diverse	ADJ
alces-443	15	12	investigations	investigation	NOUN
alces-443	15	13	as	as	ADP
alces-443	15	14	evolutionary	evolutionary	ADJ
alces-443	15	15	histories	history	NOUN
alces-443	15	16	of	of	ADP
alces-443	15	17	species	specie	NOUN
alces-443	15	18	(	(	PUNCT
alces-443	15	19	avise	avise	VERB
alces-443	15	20	et	et	PROPN
alces-443	15	21	al	al	PROPN
alces-443	15	22	.	.	PROPN
alces-443	15	23	1987	1987	NUM
alces-443	15	24	,	,	PUNCT
alces-443	15	25	2000	2000	NUM
alces-443	15	26	)	)	PUNCT
alces-443	15	27	,	,	PUNCT
alces-443	15	28	interactions	interaction	NOUN
alces-443	15	29	and	and	CCONJ
alces-443	15	30	relationships	relationship	NOUN
alces-443	15	31	among	among	ADP
alces-443	15	32	populations	population	NOUN
alces-443	15	33	(	(	PUNCT
alces-443	15	34	blundell	blundell	PROPN
alces-443	15	35	et	et	PROPN
alces-443	15	36	al	al	PROPN
alces-443	15	37	.	.	PROPN
alces-443	15	38	2002	2002	NUM
alces-443	15	39	)	)	PUNCT
alces-443	15	40	or	or	CCONJ
alces-443	15	41	individuals	individual	NOUN
alces-443	15	42	(	(	PUNCT
alces-443	15	43	quellar	quellar	ADJ
alces-443	15	44	et	et	PROPN
alces-443	15	45	al	al	PROPN
alces-443	15	46	.	.	PROPN
alces-443	15	47	1993	1993	NUM
alces-443	15	48	)	)	PUNCT
alces-443	15	49	,	,	PUNCT
alces-443	15	50	evaluation	evaluation	NOUN
alces-443	15	51	of	of	ADP
alces-443	15	52	the	the	DET
alces-443	15	53	success	success	NOUN
alces-443	15	54	of	of	ADP
alces-443	15	55	specific	specific	ADJ
alces-443	15	56	management	management	NOUN
alces-443	15	57	actions	action	NOUN
alces-443	15	58	(	(	PUNCT
alces-443	15	59	vernesi	vernesi	NOUN
alces-443	15	60	et	et	PROPN
alces-443	15	61	al	al	PROPN
alces-443	15	62	.	.	PROPN
alces-443	15	63	2002	2002	NUM
alces-443	15	64	)	)	PUNCT
alces-443	15	65	,	,	PUNCT
alces-443	15	66	population	population	NOUN
alces-443	15	67	and	and	CCONJ
alces-443	15	68	behavioral	behavioral	ADJ
alces-443	15	69	ecology	ecology	NOUN
alces-443	15	70	(	(	PUNCT
alces-443	15	71	scribner	scribner	NOUN
alces-443	15	72	and	and	CCONJ
alces-443	15	73	chesser	chesser	NOUN
alces-443	15	74	2001	2001	NUM
alces-443	15	75	)	)	PUNCT
alces-443	15	76	,	,	PUNCT
alces-443	15	77	and	and	CCONJ
alces-443	15	78	food	food	NOUN
alces-443	15	79	habits	habit	NOUN
alces-443	15	80	(	(	PUNCT
alces-443	15	81	symondson	symondson	NOUN
alces-443	15	82	2002	2002	NUM
alces-443	15	83	)	)	PUNCT
alces-443	15	84	.	.	PUNCT
alces-443	16	1	recent	recent	ADJ
alces-443	16	2	advances	advance	NOUN
alces-443	16	3	in	in	ADP
alces-443	16	4	collection	collection	NOUN
alces-443	16	5	and	and	CCONJ
alces-443	16	6	analysis	analysis	NOUN
alces-443	16	7	of	of	ADP
alces-443	16	8	genetic	genetic	ADJ
alces-443	16	9	data	datum	NOUN
alces-443	16	10	have	have	AUX
alces-443	16	11	facilitated	facilitate	VERB
alces-443	16	12	more	more	ADJ
alces-443	16	13	refined	refined	ADJ
alces-443	16	14	approaches	approach	NOUN
alces-443	16	15	to	to	ADP
alces-443	16	16	evolutionary	evolutionary	ADJ
alces-443	16	17	and	and	CCONJ
alces-443	16	18	population	population	NOUN
alces-443	16	19	genetic	genetic	ADJ
alces-443	16	20	questions	question	NOUN
alces-443	16	21	,	,	PUNCT
alces-443	16	22	and	and	CCONJ
alces-443	16	23	our	our	PRON
alces-443	16	24	understanding	understanding	NOUN
alces-443	16	25	of	of	ADP
alces-443	16	26	moose	moose	NOUN
alces-443	16	27	biology	biology	NOUN
alces-443	16	28	has	have	AUX
alces-443	16	29	benefited	benefit	VERB
alces-443	16	30	as	as	ADP
alces-443	16	31	a	a	DET
alces-443	16	32	result	result	NOUN
alces-443	16	33	of	of	ADP
alces-443	16	34	those	those	DET
alces-443	16	35	advances	advance	NOUN
alces-443	16	36	.	.	PUNCT
alces-443	17	1	evolution	evolution	NOUN
alces-443	17	2	and	and	CCONJ
alces-443	17	3	taxonomy	taxonomy	NOUN
alces-443	17	4	of	of	ADP
alces-443	17	5	moose	moose	NOUN
alces-443	17	6	(	(	PUNCT
alces-443	17	7	alces	alce	NOUN
alces-443	17	8	alces	alce	NOUN
alces-443	17	9	)	)	PUNCT
alces-443	17	10	have	have	AUX
alces-443	17	11	been	be	AUX
alces-443	17	12	reviewed	review	VERB
alces-443	17	13	previously	previously	ADV
alces-443	17	14	(	(	PUNCT
alces-443	17	15	peterson	peterson	PROPN
alces-443	17	16	1955	1955	NUM
alces-443	17	17	;	;	PUNCT
alces-443	17	18	groves	grove	NOUN
alces-443	17	19	and	and	CCONJ
alces-443	17	20	grubb	grubb	PROPN
alces-443	17	21	1987	1987	NUM
alces-443	17	22	;	;	PUNCT
alces-443	17	23	geist	geist	NOUN
alces-443	17	24	,	,	PUNCT
alces-443	17	25	1987a	1987a	NUM
alces-443	17	26	,	,	PUNCT
alces-443	17	27	b	b	NOUN
alces-443	17	28	,	,	PUNCT
alces-443	17	29	1998	1998	NUM
alces-443	17	30	;	;	PUNCT
alces-443	17	31	sher	sher	NOUN
alces-443	17	32	1987	1987	NUM
alces-443	17	33	;	;	PUNCT
alces-443	17	34	lister	lister	PROPN
alces-443	17	35	1993	1993	NUM
alces-443	17	36	;	;	PUNCT
alces-443	17	37	guthrie	guthrie	PROPN
alces-443	17	38	1995	1995	NUM
alces-443	17	39	;	;	PUNCT
alces-443	17	40	bubenik	bubenik	X
alces-443	17	41	1998	1998	NUM
alces-443	17	42	;	;	PUNCT
alces-443	17	43	bowyer	bowyer	PROPN
alces-443	17	44	et	et	PROPN
alces-443	17	45	al	al	PROPN
alces-443	17	46	.	.	PROPN
alces-443	17	47	2003	2003	NUM
alces-443	17	48	)	)	PUNCT
alces-443	17	49	and	and	CCONJ
alces-443	17	50	have	have	AUX
alces-443	17	51	encompassed	encompass	VERB
alces-443	17	52	aspects	aspect	NOUN
alces-443	17	53	of	of	ADP
alces-443	17	54	behavior	behavior	NOUN
alces-443	17	55	,	,	PUNCT
alces-443	17	56	morphology	morphology	NOUN
alces-443	17	57	,	,	PUNCT
alces-443	17	58	paleontology	paleontology	NOUN
alces-443	17	59	,	,	PUNCT
alces-443	17	60	and	and	CCONJ
alces-443	17	61	genetics	genetic	NOUN
alces-443	17	62	,	,	PUNCT
alces-443	17	63	but	but	CCONJ
alces-443	17	64	no	no	DET
alces-443	17	65	review	review	NOUN
alces-443	17	66	has	have	AUX
alces-443	17	67	dealt	deal	VERB
alces-443	17	68	specifihundertmark	specifihundertmark	NOUN
alces-443	17	69	and	and	CCONJ
alces-443	17	70	bowyer	bowyer	PROPN
alces-443	17	71	moose	moose	PROPN
alces-443	17	72	genetics	genetics	PROPN
alces-443	17	73	alces	alce	NOUN
alces-443	17	74	vol	vol	NOUN
alces-443	17	75	.	.	PROPN
alces-443	18	1	40	40	NUM
alces-443	18	2	,	,	PUNCT
alces-443	18	3	2004	2004	NUM
alces-443	18	4	104	104	NUM
alces-443	18	5	cally	cally	ADV
alces-443	18	6	with	with	ADP
alces-443	18	7	genetics	genetic	NOUN
alces-443	18	8	.	.	PUNCT
alces-443	19	1	in	in	ADP
alces-443	19	2	this	this	DET
alces-443	19	3	review	review	NOUN
alces-443	19	4	,	,	PUNCT
alces-443	19	5	our	our	PRON
alces-443	19	6	goal	goal	NOUN
alces-443	19	7	is	be	AUX
alces-443	19	8	to	to	PART
alces-443	19	9	provide	provide	VERB
alces-443	19	10	an	an	DET
alces-443	19	11	overview	overview	NOUN
alces-443	19	12	of	of	ADP
alces-443	19	13	older	old	ADJ
alces-443	19	14	studies	study	NOUN
alces-443	19	15	while	while	SCONJ
alces-443	19	16	focusing	focus	VERB
alces-443	19	17	on	on	ADP
alces-443	19	18	recent	recent	ADJ
alces-443	19	19	advances	advance	NOUN
alces-443	19	20	in	in	ADP
alces-443	19	21	genetics	genetic	NOUN
alces-443	19	22	and	and	CCONJ
alces-443	19	23	phylogeography	phylogeography	NOUN
alces-443	19	24	(	(	PUNCT
alces-443	19	25	see	see	VERB
alces-443	19	26	definition	definition	NOUN
alces-443	19	27	in	in	ADP
alces-443	19	28	appendix	appendix	NOUN
alces-443	19	29	1	1	NUM
alces-443	19	30	)	)	PUNCT
alces-443	19	31	of	of	ADP
alces-443	19	32	moose	moose	NOUN
alces-443	19	33	and	and	CCONJ
alces-443	19	34	the	the	DET
alces-443	19	35	insights	insight	NOUN
alces-443	19	36	they	they	PRON
alces-443	19	37	provide	provide	VERB
alces-443	19	38	.	.	PUNCT
alces-443	20	1	the	the	DET
alces-443	20	2	broad	broad	ADJ
alces-443	20	3	scope	scope	NOUN
alces-443	20	4	of	of	ADP
alces-443	20	5	genetic	genetic	ADJ
alces-443	20	6	and	and	CCONJ
alces-443	20	7	evolutionary	evolutionary	ADJ
alces-443	20	8	investigations	investigation	NOUN
alces-443	20	9	in	in	ADP
alces-443	20	10	species	specie	NOUN
alces-443	20	11	biology	biology	NOUN
alces-443	20	12	would	would	AUX
alces-443	20	13	make	make	VERB
alces-443	20	14	a	a	DET
alces-443	20	15	complete	complete	ADJ
alces-443	20	16	review	review	NOUN
alces-443	20	17	of	of	ADP
alces-443	20	18	all	all	DET
alces-443	20	19	studies	study	NOUN
alces-443	20	20	disjointed	disjoint	VERB
alces-443	20	21	.	.	PUNCT
alces-443	21	1	yet	yet	ADV
alces-443	21	2	,	,	PUNCT
alces-443	21	3	approximately	approximately	ADV
alces-443	21	4	half	half	NOUN
alces-443	21	5	of	of	ADP
alces-443	21	6	all	all	DET
alces-443	21	7	published	publish	VERB
alces-443	21	8	studies	study	NOUN
alces-443	21	9	of	of	ADP
alces-443	21	10	moose	moose	NOUN
alces-443	21	11	genetics	genetic	NOUN
alces-443	21	12	have	have	AUX
alces-443	21	13	been	be	AUX
alces-443	21	14	published	publish	VERB
alces-443	21	15	since	since	SCONJ
alces-443	21	16	the	the	DET
alces-443	21	17	comprehensive	comprehensive	ADJ
alces-443	21	18	treatise	treatise	NOUN
alces-443	21	19	“	"	PUNCT
alces-443	21	20	ecology	ecology	NOUN
alces-443	21	21	and	and	CCONJ
alces-443	21	22	management	management	NOUN
alces-443	21	23	of	of	ADP
alces-443	21	24	the	the	DET
alces-443	21	25	north	north	ADJ
alces-443	21	26	american	american	ADJ
alces-443	21	27	moose	moose	NOUN
alces-443	21	28	”	"	PUNCT
alces-443	21	29	(	(	PUNCT
alces-443	21	30	franzmann	franzmann	NOUN
alces-443	21	31	and	and	CCONJ
alces-443	21	32	schwartz	schwartz	PROPN
alces-443	21	33	1998	1998	NUM
alces-443	21	34	)	)	PUNCT
alces-443	21	35	was	be	AUX
alces-443	21	36	compiled	compile	VERB
alces-443	21	37	;	;	PUNCT
alces-443	21	38	thus	thus	ADV
alces-443	21	39	,	,	PUNCT
alces-443	21	40	we	we	PRON
alces-443	21	41	were	be	AUX
alces-443	21	42	compelled	compel	VERB
alces-443	21	43	to	to	PART
alces-443	21	44	present	present	VERB
alces-443	21	45	the	the	DET
alces-443	21	46	most	most	ADV
alces-443	21	47	complete	complete	ADJ
alces-443	21	48	review	review	NOUN
alces-443	21	49	possible	possible	ADJ
alces-443	21	50	.	.	PUNCT
alces-443	22	1	in	in	ADP
alces-443	22	2	an	an	DET
alces-443	22	3	effort	effort	NOUN
alces-443	22	4	to	to	PART
alces-443	22	5	present	present	VERB
alces-443	22	6	a	a	DET
alces-443	22	7	cogent	cogent	ADJ
alces-443	22	8	summary	summary	NOUN
alces-443	22	9	of	of	ADP
alces-443	22	10	all	all	DET
alces-443	22	11	relevant	relevant	ADJ
alces-443	22	12	studies	study	NOUN
alces-443	22	13	,	,	PUNCT
alces-443	22	14	we	we	PRON
alces-443	22	15	have	have	AUX
alces-443	22	16	divided	divide	VERB
alces-443	22	17	this	this	DET
alces-443	22	18	review	review	NOUN
alces-443	22	19	into	into	ADP
alces-443	22	20	3	3	NUM
alces-443	22	21	parts	part	NOUN
alces-443	22	22	:	:	PUNCT
alces-443	22	23	(	(	PUNCT
alces-443	22	24	1	1	X
alces-443	22	25	)	)	PUNCT
alces-443	22	26	assessing	assess	VERB
alces-443	22	27	genetic	genetic	ADJ
alces-443	22	28	diversity	diversity	NOUN
alces-443	22	29	,	,	PUNCT
alces-443	22	30	where	where	SCONJ
alces-443	22	31	we	we	PRON
alces-443	22	32	review	review	VERB
alces-443	22	33	the	the	DET
alces-443	22	34	different	different	ADJ
alces-443	22	35	types	type	NOUN
alces-443	22	36	of	of	ADP
alces-443	22	37	markers	marker	NOUN
alces-443	22	38	examined	examine	VERB
alces-443	22	39	in	in	ADP
alces-443	22	40	studies	study	NOUN
alces-443	22	41	of	of	ADP
alces-443	22	42	moose	moose	NOUN
alces-443	22	43	genetics	genetic	NOUN
alces-443	22	44	and	and	CCONJ
alces-443	22	45	the	the	DET
alces-443	22	46	conclusions	conclusion	NOUN
alces-443	22	47	drawn	draw	VERB
alces-443	22	48	from	from	ADP
alces-443	22	49	those	those	DET
alces-443	22	50	studies	study	NOUN
alces-443	22	51	;	;	PUNCT
alces-443	22	52	(	(	PUNCT
alces-443	22	53	2	2	X
alces-443	22	54	)	)	PUNCT
alces-443	22	55	moose	moose	NOUN
alces-443	22	56	evolution	evolution	NOUN
alces-443	22	57	and	and	CCONJ
alces-443	22	58	phylogeography	phylogeography	NOUN
alces-443	22	59	,	,	PUNCT
alces-443	22	60	where	where	SCONJ
alces-443	22	61	we	we	PRON
alces-443	22	62	examine	examine	VERB
alces-443	22	63	the	the	DET
alces-443	22	64	evolutionary	evolutionary	ADJ
alces-443	22	65	descent	descent	NOUN
alces-443	22	66	of	of	ADP
alces-443	22	67	moose	moose	NOUN
alces-443	22	68	and	and	CCONJ
alces-443	22	69	processes	process	NOUN
alces-443	22	70	that	that	PRON
alces-443	22	71	have	have	AUX
alces-443	22	72	shaped	shape	VERB
alces-443	22	73	the	the	DET
alces-443	22	74	genetic	genetic	ADJ
alces-443	22	75	variation	variation	NOUN
alces-443	22	76	and	and	CCONJ
alces-443	22	77	structure	structure	NOUN
alces-443	22	78	observed	observe	VERB
alces-443	22	79	today	today	NOUN
alces-443	22	80	;	;	PUNCT
alces-443	22	81	and	and	CCONJ
alces-443	22	82	(	(	PUNCT
alces-443	22	83	3	3	X
alces-443	22	84	)	)	PUNCT
alces-443	22	85	genetic	genetic	ADJ
alces-443	22	86	effects	effect	NOUN
alces-443	22	87	of	of	ADP
alces-443	22	88	harvest	harvest	NOUN
alces-443	22	89	,	,	PUNCT
alces-443	22	90	which	which	PRON
alces-443	22	91	reviews	review	VERB
alces-443	22	92	a	a	DET
alces-443	22	93	small	small	ADJ
alces-443	22	94	but	but	CCONJ
alces-443	22	95	important	important	ADJ
alces-443	22	96	body	body	NOUN
alces-443	22	97	of	of	ADP
alces-443	22	98	work	work	NOUN
alces-443	22	99	composed	compose	VERB
alces-443	22	100	of	of	ADP
alces-443	22	101	management	management	NOUN
alces-443	22	102	-	-	PUNCT
alces-443	22	103	based	base	VERB
alces-443	22	104	modeling	modeling	NOUN
alces-443	22	105	that	that	PRON
alces-443	22	106	examined	examine	VERB
alces-443	22	107	effects	effect	NOUN
alces-443	22	108	of	of	ADP
alces-443	22	109	various	various	ADJ
alces-443	22	110	harvest	harvest	NOUN
alces-443	22	111	regimes	regime	NOUN
alces-443	22	112	on	on	ADP
alces-443	22	113	population	population	NOUN
alces-443	22	114	and	and	CCONJ
alces-443	22	115	genetic	genetic	ADJ
alces-443	22	116	measures	measure	NOUN
alces-443	22	117	.	.	PUNCT
alces-443	23	1	assessing	assess	VERB
alces-443	23	2	genetic	genetic	ADJ
alces-443	23	3	diversity	diversity	NOUN
alces-443	23	4	one	one	NUM
alces-443	23	5	potential	potential	ADJ
alces-443	23	6	difficulty	difficulty	NOUN
alces-443	23	7	in	in	ADP
alces-443	23	8	discussing	discuss	VERB
alces-443	23	9	genetic	genetic	ADJ
alces-443	23	10	analyses	analysis	NOUN
alces-443	23	11	is	be	AUX
alces-443	23	12	the	the	DET
alces-443	23	13	use	use	NOUN
alces-443	23	14	of	of	ADP
alces-443	23	15	specialized	specialized	ADJ
alces-443	23	16	terminology	terminology	NOUN
alces-443	23	17	.	.	PUNCT
alces-443	24	1	to	to	PART
alces-443	24	2	avoid	avoid	VERB
alces-443	24	3	uncertainty	uncertainty	NOUN
alces-443	24	4	and	and	CCONJ
alces-443	24	5	enhance	enhance	VERB
alces-443	24	6	understanding	understanding	NOUN
alces-443	24	7	,	,	PUNCT
alces-443	24	8	we	we	PRON
alces-443	24	9	provide	provide	VERB
alces-443	24	10	a	a	DET
alces-443	24	11	brief	brief	ADJ
alces-443	24	12	glossary	glossary	NOUN
alces-443	24	13	of	of	ADP
alces-443	24	14	terms	term	NOUN
alces-443	24	15	used	use	VERB
alces-443	24	16	in	in	ADP
alces-443	24	17	this	this	DET
alces-443	24	18	review	review	NOUN
alces-443	24	19	(	(	PUNCT
alces-443	24	20	appendix	appendix	NOUN
alces-443	24	21	1	1	NUM
alces-443	24	22	)	)	PUNCT
alces-443	24	23	.	.	PUNCT
alces-443	25	1	terms	term	NOUN
alces-443	25	2	defined	define	VERB
alces-443	25	3	in	in	ADP
alces-443	25	4	the	the	DET
alces-443	25	5	glossary	glossary	NOUN
alces-443	25	6	are	be	AUX
alces-443	25	7	highlighted	highlight	VERB
alces-443	25	8	in	in	ADP
alces-443	25	9	bold	bold	ADJ
alces-443	25	10	in	in	ADP
alces-443	25	11	the	the	DET
alces-443	25	12	text	text	NOUN
alces-443	25	13	at	at	ADP
alces-443	25	14	their	their	PRON
alces-443	25	15	first	first	ADJ
alces-443	25	16	usage	usage	NOUN
alces-443	25	17	.	.	PUNCT
alces-443	26	1	the	the	DET
alces-443	26	2	allozyme	allozyme	NOUN
alces-443	26	3	era	era	NOUN
alces-443	26	4	first	first	ADJ
alces-443	26	5	reports	report	NOUN
alces-443	26	6	of	of	ADP
alces-443	26	7	genetic	genetic	ADJ
alces-443	26	8	investigations	investigation	NOUN
alces-443	26	9	of	of	ADP
alces-443	26	10	moose	moose	NOUN
alces-443	26	11	were	be	AUX
alces-443	26	12	published	publish	VERB
alces-443	26	13	by	by	ADP
alces-443	26	14	braend	braend	NOUN
alces-443	26	15	(	(	PUNCT
alces-443	26	16	1962	1962	NUM
alces-443	26	17	,	,	PUNCT
alces-443	26	18	cited	cite	VERB
alces-443	26	19	by	by	ADP
alces-443	26	20	gyllensten	gyllensten	PROPN
alces-443	26	21	et	et	PROPN
alces-443	26	22	al	al	PROPN
alces-443	26	23	.	.	PROPN
alces-443	26	24	1980	1980	NUM
alces-443	26	25	)	)	PUNCT
alces-443	26	26	,	,	PUNCT
alces-443	26	27	nadler	nadler	PROPN
alces-443	26	28	et	et	PROPN
alces-443	26	29	al	al	PROPN
alces-443	26	30	.	.	PROPN
alces-443	27	1	(	(	PUNCT
alces-443	27	2	1967	1967	NUM
alces-443	27	3	)	)	PUNCT
alces-443	27	4	,	,	PUNCT
alces-443	27	5	and	and	CCONJ
alces-443	27	6	shubin	shubin	PROPN
alces-443	27	7	(	(	PUNCT
alces-443	27	8	1969	1969	NUM
alces-443	27	9	,	,	PUNCT
alces-443	27	10	cited	cite	VERB
alces-443	27	11	by	by	ADP
alces-443	27	12	gyllensten	gyllensten	PROPN
alces-443	27	13	et	et	PROPN
alces-443	27	14	al	al	PROPN
alces-443	27	15	.	.	PROPN
alces-443	27	16	1980	1980	NUM
alces-443	27	17	)	)	PUNCT
alces-443	27	18	,	,	PUNCT
alces-443	27	19	wherein	wherein	SCONJ
alces-443	27	20	those	those	DET
alces-443	27	21	authors	author	NOUN
alces-443	27	22	examined	examine	VERB
alces-443	27	23	electrophoretic	electrophoretic	ADJ
alces-443	27	24	variation	variation	NOUN
alces-443	27	25	in	in	ADP
alces-443	27	26	proteins	protein	NOUN
alces-443	27	27	from	from	ADP
alces-443	27	28	blood	blood	NOUN
alces-443	27	29	serum	serum	NOUN
alces-443	27	30	;	;	PUNCT
alces-443	27	31	no	no	DET
alces-443	27	32	variation	variation	NOUN
alces-443	27	33	in	in	ADP
alces-443	27	34	those	those	DET
alces-443	27	35	genetic	genetic	ADJ
alces-443	27	36	markers	marker	NOUN
alces-443	27	37	occurred	occur	VERB
alces-443	27	38	in	in	ADP
alces-443	27	39	scandinavia	scandinavia	PROPN
alces-443	27	40	,	,	PUNCT
alces-443	27	41	north	north	PROPN
alces-443	27	42	america	america	PROPN
alces-443	27	43	,	,	PUNCT
alces-443	27	44	and	and	CCONJ
alces-443	27	45	central	central	ADJ
alces-443	27	46	russia	russia	PROPN
alces-443	27	47	,	,	PUNCT
alces-443	27	48	respectively	respectively	ADV
alces-443	27	49	.	.	PUNCT
alces-443	28	1	the	the	DET
alces-443	28	2	first	first	ADJ
alces-443	28	3	study	study	NOUN
alces-443	28	4	to	to	PART
alces-443	28	5	report	report	VERB
alces-443	28	6	genetic	genetic	ADJ
alces-443	28	7	variability	variability	NOUN
alces-443	28	8	was	be	AUX
alces-443	28	9	ryman	ryman	NOUN
alces-443	28	10	et	et	PROPN
alces-443	28	11	al	al	PROPN
alces-443	28	12	.	.	PROPN
alces-443	29	1	(	(	PUNCT
alces-443	29	2	1977	1977	NUM
alces-443	29	3	)	)	PUNCT
alces-443	29	4	,	,	PUNCT
alces-443	29	5	who	who	PRON
alces-443	29	6	examined	examine	VERB
alces-443	29	7	1,384	1,384	NUM
alces-443	29	8	moose	moose	NOUN
alces-443	29	9	from	from	ADP
alces-443	29	10	3	3	NUM
alces-443	29	11	areas	area	NOUN
alces-443	29	12	of	of	ADP
alces-443	29	13	sweden	sweden	PROPN
alces-443	29	14	for	for	ADP
alces-443	29	15	polymorphism	polymorphism	NOUN
alces-443	29	16	at	at	ADP
alces-443	29	17	23	23	NUM
alces-443	29	18	allozyme	allozyme	NOUN
alces-443	29	19	loci	locus	NOUN
alces-443	29	20	.	.	PUNCT
alces-443	30	1	that	that	DET
alces-443	30	2	study	study	NOUN
alces-443	30	3	reported	report	VERB
alces-443	30	4	only	only	ADV
alces-443	30	5	1	1	NUM
alces-443	30	6	locus	locus	NOUN
alces-443	30	7	to	to	PART
alces-443	30	8	be	be	AUX
alces-443	30	9	polymorphic	polymorphic	ADJ
alces-443	30	10	,	,	PUNCT
alces-443	30	11	however	however	ADV
alces-443	30	12	,	,	PUNCT
alces-443	30	13	and	and	CCONJ
alces-443	30	14	only	only	ADV
alces-443	30	15	in	in	ADP
alces-443	30	16	1	1	NUM
alces-443	30	17	region	region	NOUN
alces-443	30	18	.	.	PUNCT
alces-443	31	1	although	although	SCONJ
alces-443	31	2	they	they	PRON
alces-443	31	3	suspected	suspect	VERB
alces-443	31	4	that	that	SCONJ
alces-443	31	5	allele	allele	NOUN
alces-443	31	6	frequencies	frequency	NOUN
alces-443	31	7	varied	vary	VERB
alces-443	31	8	geographically	geographically	ADV
alces-443	31	9	within	within	ADP
alces-443	31	10	the	the	DET
alces-443	31	11	1	1	NUM
alces-443	31	12	variable	variable	NOUN
alces-443	31	13	region	region	NOUN
alces-443	31	14	,	,	PUNCT
alces-443	31	15	the	the	DET
alces-443	31	16	difference	difference	NOUN
alces-443	31	17	was	be	AUX
alces-443	31	18	not	not	PART
alces-443	31	19	statistically	statistically	ADV
alces-443	31	20	significant	significant	ADJ
alces-443	31	21	.	.	PUNCT
alces-443	32	1	those	those	DET
alces-443	32	2	authors	author	NOUN
alces-443	32	3	concluded	conclude	VERB
alces-443	32	4	that	that	SCONJ
alces-443	32	5	genetic	genetic	ADJ
alces-443	32	6	drift	drift	NOUN
alces-443	32	7	associated	associate	VERB
alces-443	32	8	with	with	ADP
alces-443	32	9	a	a	DET
alces-443	32	10	severe	severe	ADJ
alces-443	32	11	population	population	NOUN
alces-443	32	12	bottleneck	bottleneck	NOUN
alces-443	32	13	(	(	PUNCT
alces-443	32	14	reduction	reduction	NOUN
alces-443	32	15	in	in	ADP
alces-443	32	16	population	population	NOUN
alces-443	32	17	size	size	NOUN
alces-443	32	18	)	)	PUNCT
alces-443	32	19	in	in	ADP
alces-443	32	20	sweden	sweden	PROPN
alces-443	32	21	in	in	ADP
alces-443	32	22	the	the	DET
alces-443	32	23	19th	19th	ADJ
alces-443	32	24	century	century	NOUN
alces-443	32	25	was	be	AUX
alces-443	32	26	a	a	DET
alces-443	32	27	probable	probable	ADJ
alces-443	32	28	cause	cause	NOUN
alces-443	32	29	of	of	ADP
alces-443	32	30	the	the	DET
alces-443	32	31	observed	observe	VERB
alces-443	32	32	lack	lack	NOUN
alces-443	32	33	of	of	ADP
alces-443	32	34	diversity	diversity	NOUN
alces-443	32	35	.	.	PUNCT
alces-443	33	1	wilhelmson	wilhelmson	PROPN
alces-443	33	2	et	et	PROPN
alces-443	33	3	al	al	PROPN
alces-443	33	4	.	.	PROPN
alces-443	33	5	(	(	PUNCT
alces-443	33	6	1978	1978	NUM
alces-443	33	7	)	)	PUNCT
alces-443	33	8	,	,	PUNCT
alces-443	33	9	examining	examine	VERB
alces-443	33	10	variation	variation	NOUN
alces-443	33	11	in	in	ADP
alces-443	33	12	serum	serum	NOUN
alces-443	33	13	proteins	protein	NOUN
alces-443	33	14	,	,	PUNCT
alces-443	33	15	noted	note	VERB
alces-443	33	16	no	no	DET
alces-443	33	17	differences	difference	NOUN
alces-443	33	18	between	between	ADP
alces-443	33	19	canadian	canadian	ADJ
alces-443	33	20	and	and	CCONJ
alces-443	33	21	european	european	ADJ
alces-443	33	22	moose	moose	NOUN
alces-443	33	23	.	.	PUNCT
alces-443	34	1	from	from	ADP
alces-443	34	2	that	that	DET
alces-443	34	3	evidence	evidence	NOUN
alces-443	34	4	,	,	PUNCT
alces-443	34	5	they	they	PRON
alces-443	34	6	concluded	conclude	VERB
alces-443	34	7	that	that	SCONJ
alces-443	34	8	moose	moose	NOUN
alces-443	34	9	populations	population	NOUN
alces-443	34	10	on	on	ADP
alces-443	34	11	separate	separate	ADJ
alces-443	34	12	continents	continent	NOUN
alces-443	34	13	had	have	AUX
alces-443	34	14	not	not	PART
alces-443	34	15	undergone	undergo	VERB
alces-443	34	16	significant	significant	ADJ
alces-443	34	17	genetic	genetic	ADJ
alces-443	34	18	drift	drift	NOUN
alces-443	34	19	despite	despite	SCONJ
alces-443	34	20	being	be	AUX
alces-443	34	21	separated	separate	VERB
alces-443	34	22	for	for	ADP
alces-443	34	23	thousands	thousand	NOUN
alces-443	34	24	of	of	ADP
alces-443	34	25	years	year	NOUN
alces-443	34	26	,	,	PUNCT
alces-443	34	27	implying	imply	VERB
alces-443	34	28	that	that	SCONJ
alces-443	34	29	effective	effective	ADJ
alces-443	34	30	population	population	NOUN
alces-443	34	31	sizes	size	NOUN
alces-443	34	32	of	of	ADP
alces-443	34	33	moose	moose	NOUN
alces-443	34	34	populations	population	NOUN
alces-443	34	35	historically	historically	ADV
alces-443	34	36	had	have	AUX
alces-443	34	37	been	be	AUX
alces-443	34	38	large	large	ADJ
alces-443	34	39	.	.	PUNCT
alces-443	35	1	wilhelmson	wilhelmson	PROPN
alces-443	35	2	et	et	PROPN
alces-443	35	3	al	al	PROPN
alces-443	35	4	.	.	PROPN
alces-443	36	1	(	(	PUNCT
alces-443	36	2	1978	1978	NUM
alces-443	36	3	)	)	PUNCT
alces-443	36	4	also	also	ADV
alces-443	36	5	proposed	propose	VERB
alces-443	36	6	that	that	SCONJ
alces-443	36	7	historic	historic	ADJ
alces-443	36	8	population	population	NOUN
alces-443	36	9	bottlenecks	bottleneck	NOUN
alces-443	36	10	in	in	ADP
alces-443	36	11	sweden	sweden	PROPN
alces-443	36	12	had	have	AUX
alces-443	36	13	not	not	PART
alces-443	36	14	been	be	AUX
alces-443	36	15	severe	severe	ADJ
alces-443	36	16	enough	enough	ADV
alces-443	36	17	to	to	PART
alces-443	36	18	have	have	AUX
alces-443	36	19	had	have	VERB
alces-443	36	20	an	an	DET
alces-443	36	21	effect	effect	NOUN
alces-443	36	22	on	on	ADP
alces-443	36	23	genetic	genetic	ADJ
alces-443	36	24	diversity	diversity	NOUN
alces-443	36	25	of	of	ADP
alces-443	36	26	moose	moose	NOUN
alces-443	36	27	.	.	PUNCT
alces-443	37	1	gyllensten	gyllensten	VERB
alces-443	37	2	et	et	PROPN
alces-443	37	3	al	al	PROPN
alces-443	37	4	.	.	PROPN
alces-443	38	1	(	(	PUNCT
alces-443	38	2	1980	1980	NUM
alces-443	38	3	)	)	PUNCT
alces-443	38	4	conducted	conduct	VERB
alces-443	38	5	extensive	extensive	ADJ
alces-443	38	6	screening	screening	NOUN
alces-443	38	7	of	of	ADP
alces-443	38	8	a	a	DET
alces-443	38	9	transferrin	transferrin	NOUN
alces-443	38	10	locus	locus	NOUN
alces-443	38	11	from	from	ADP
alces-443	38	12	moose	moose	NOUN
alces-443	38	13	across	across	ADP
alces-443	38	14	fennoscandia	fennoscandia	NOUN
alces-443	38	15	and	and	CCONJ
alces-443	38	16	detected	detect	VERB
alces-443	38	17	a	a	DET
alces-443	38	18	single	single	ADJ
alces-443	38	19	polymorphism	polymorphism	NOUN
alces-443	38	20	occurring	occur	VERB
alces-443	38	21	in	in	ADP
alces-443	38	22	norway	norway	PROPN
alces-443	38	23	,	,	PUNCT
alces-443	38	24	sweden	sweden	PROPN
alces-443	38	25	,	,	PUNCT
alces-443	38	26	and	and	CCONJ
alces-443	38	27	finland	finland	PROPN
alces-443	38	28	.	.	PUNCT
alces-443	39	1	nonetheless	nonetheless	ADV
alces-443	39	2	,	,	PUNCT
alces-443	39	3	the	the	DET
alces-443	39	4	polymorphism	polymorphism	NOUN
alces-443	39	5	was	be	AUX
alces-443	39	6	present	present	ADJ
alces-443	39	7	in	in	ADP
alces-443	39	8	only	only	ADV
alces-443	39	9	6	6	NUM
alces-443	39	10	of	of	ADP
alces-443	39	11	16	16	NUM
alces-443	39	12	populations	population	NOUN
alces-443	39	13	and	and	CCONJ
alces-443	39	14	the	the	DET
alces-443	39	15	uncommon	uncommon	ADJ
alces-443	39	16	allele	allele	NOUN
alces-443	39	17	never	never	ADV
alces-443	39	18	exceeded	exceed	VERB
alces-443	39	19	6	6	NUM
alces-443	39	20	%	%	NOUN
alces-443	39	21	in	in	ADP
alces-443	39	22	any	any	DET
alces-443	39	23	population	population	NOUN
alces-443	39	24	.	.	PUNCT
alces-443	40	1	the	the	DET
alces-443	40	2	authors	author	NOUN
alces-443	40	3	presented	present	VERB
alces-443	40	4	differences	difference	NOUN
alces-443	40	5	in	in	ADP
alces-443	40	6	frequency	frequency	NOUN
alces-443	40	7	of	of	ADP
alces-443	40	8	the	the	DET
alces-443	40	9	uncommon	uncommon	ADJ
alces-443	40	10	allele	allele	NOUN
alces-443	40	11	as	as	ADP
alces-443	40	12	evidence	evidence	NOUN
alces-443	40	13	of	of	ADP
alces-443	40	14	differalces	differalce	NOUN
alces-443	40	15	vol	vol	NOUN
alces-443	40	16	.	.	PROPN
alces-443	40	17	40	40	NUM
alces-443	40	18	,	,	PUNCT
alces-443	40	19	2004	2004	NUM
alces-443	40	20	moose	moose	NOUN
alces-443	40	21	genetics	genetic	NOUN
alces-443	40	22	hundertmark	hundertmark	NOUN
alces-443	40	23	and	and	CCONJ
alces-443	40	24	bowyer	bowyer	PROPN
alces-443	40	25	105	105	NUM
alces-443	40	26	entiation	entiation	NOUN
alces-443	40	27	of	of	ADP
alces-443	40	28	populations	population	NOUN
alces-443	40	29	geographically	geographically	ADV
alces-443	40	30	,	,	PUNCT
alces-443	40	31	supporting	support	VERB
alces-443	40	32	observations	observation	NOUN
alces-443	40	33	of	of	ADP
alces-443	40	34	ryman	ryman	NOUN
alces-443	40	35	et	et	PROPN
alces-443	40	36	al	al	PROPN
alces-443	40	37	.	.	PROPN
alces-443	41	1	(	(	PUNCT
alces-443	41	2	1977	1977	NUM
alces-443	41	3	)	)	PUNCT
alces-443	41	4	.	.	PUNCT
alces-443	42	1	reliance	reliance	NOUN
alces-443	42	2	on	on	ADP
alces-443	42	3	the	the	DET
alces-443	42	4	occurrence	occurrence	NOUN
alces-443	42	5	of	of	ADP
alces-443	42	6	a	a	DET
alces-443	42	7	single	single	ADJ
alces-443	42	8	rare	rare	ADJ
alces-443	42	9	polymorphism	polymorphism	NOUN
alces-443	42	10	to	to	PART
alces-443	42	11	demonstrate	demonstrate	VERB
alces-443	42	12	population	population	NOUN
alces-443	42	13	subdivision	subdivision	NOUN
alces-443	42	14	,	,	PUNCT
alces-443	42	15	however	however	ADV
alces-443	42	16	,	,	PUNCT
alces-443	42	17	is	be	AUX
alces-443	42	18	tenuous	tenuous	ADJ
alces-443	42	19	at	at	ADV
alces-443	42	20	best	well	ADV
alces-443	42	21	.	.	PUNCT
alces-443	43	1	those	those	DET
alces-443	43	2	early	early	ADJ
alces-443	43	3	studies	study	NOUN
alces-443	43	4	and	and	CCONJ
alces-443	43	5	others	other	NOUN
alces-443	43	6	created	create	VERB
alces-443	43	7	an	an	DET
alces-443	43	8	impression	impression	NOUN
alces-443	43	9	among	among	ADP
alces-443	43	10	some	some	DET
alces-443	43	11	biologists	biologist	NOUN
alces-443	43	12	that	that	SCONJ
alces-443	43	13	certain	certain	ADJ
alces-443	43	14	species	specie	NOUN
alces-443	43	15	,	,	PUNCT
alces-443	43	16	including	include	VERB
alces-443	43	17	moose	moose	NOUN
alces-443	43	18	,	,	PUNCT
alces-443	43	19	possessed	possess	VERB
alces-443	43	20	little	little	ADJ
alces-443	43	21	genetic	genetic	ADJ
alces-443	43	22	variation	variation	NOUN
alces-443	43	23	across	across	ADP
alces-443	43	24	the	the	DET
alces-443	43	25	genome	genome	NOUN
alces-443	43	26	.	.	PUNCT
alces-443	44	1	hypotheses	hypothesis	NOUN
alces-443	44	2	explaining	explain	VERB
alces-443	44	3	this	this	PRON
alces-443	44	4	in	in	ADP
alces-443	44	5	evolutionary	evolutionary	ADJ
alces-443	44	6	terms	term	NOUN
alces-443	44	7	were	be	AUX
alces-443	44	8	proposed	propose	VERB
alces-443	44	9	.	.	PUNCT
alces-443	45	1	selander	seland	ADJ
alces-443	45	2	and	and	CCONJ
alces-443	45	3	kaufman	kaufman	NOUN
alces-443	45	4	(	(	PUNCT
alces-443	45	5	1973	1973	NUM
alces-443	45	6	)	)	PUNCT
alces-443	45	7	proposed	propose	VERB
alces-443	45	8	the	the	DET
alces-443	45	9	environmental	environmental	ADJ
alces-443	45	10	-	-	PUNCT
alces-443	45	11	grain	grain	NOUN
alces-443	45	12	hypothesis	hypothesis	NOUN
alces-443	45	13	,	,	PUNCT
alces-443	45	14	which	which	PRON
alces-443	45	15	stated	state	VERB
alces-443	45	16	that	that	SCONJ
alces-443	45	17	large	large	ADJ
alces-443	45	18	,	,	PUNCT
alces-443	45	19	highly	highly	ADV
alces-443	45	20	mobile	mobile	ADJ
alces-443	45	21	animals	animal	NOUN
alces-443	45	22	exhibited	exhibit	VERB
alces-443	45	23	less	less	ADJ
alces-443	45	24	genetic	genetic	ADJ
alces-443	45	25	variability	variability	NOUN
alces-443	45	26	than	than	ADP
alces-443	45	27	small	small	ADJ
alces-443	45	28	,	,	PUNCT
alces-443	45	29	sedentary	sedentary	ADJ
alces-443	45	30	species	specie	NOUN
alces-443	45	31	.	.	PUNCT
alces-443	46	1	that	that	DET
alces-443	46	2	hypothesis	hypothesis	NOUN
alces-443	46	3	further	far	ADV
alces-443	46	4	stated	state	VERB
alces-443	46	5	that	that	SCONJ
alces-443	46	6	highly	highly	ADV
alces-443	46	7	mobile	mobile	ADJ
alces-443	46	8	mammals	mammal	NOUN
alces-443	46	9	would	would	AUX
alces-443	46	10	exhibit	exhibit	VERB
alces-443	46	11	greater	great	ADJ
alces-443	46	12	homogeneity	homogeneity	NOUN
alces-443	46	13	across	across	ADP
alces-443	46	14	large	large	ADJ
alces-443	46	15	areas	area	NOUN
alces-443	46	16	than	than	ADP
alces-443	46	17	more	more	ADV
alces-443	46	18	sedentary	sedentary	ADJ
alces-443	46	19	forms	form	NOUN
alces-443	46	20	.	.	PUNCT
alces-443	47	1	other	other	ADJ
alces-443	47	2	hypotheses	hypothesis	NOUN
alces-443	47	3	proposed	propose	VERB
alces-443	47	4	that	that	SCONJ
alces-443	47	5	r	r	NOUN
alces-443	47	6	-	-	PUNCT
alces-443	47	7	strategists	strategist	NOUN
alces-443	47	8	were	be	AUX
alces-443	47	9	less	less	ADV
alces-443	47	10	variable	variable	ADJ
alces-443	47	11	than	than	ADP
alces-443	47	12	k	k	NOUN
alces-443	47	13	-	-	PUNCT
alces-443	47	14	strategists	strategist	NOUN
alces-443	47	15	(	(	PUNCT
alces-443	47	16	harrington	harrington	PROPN
alces-443	47	17	1985	1985	NUM
alces-443	47	18	)	)	PUNCT
alces-443	47	19	,	,	PUNCT
alces-443	47	20	genetic	genetic	ADJ
alces-443	47	21	heterogeneity	heterogeneity	NOUN
alces-443	47	22	was	be	AUX
alces-443	47	23	greater	great	ADJ
alces-443	47	24	in	in	ADP
alces-443	47	25	species	specie	NOUN
alces-443	47	26	inhabiting	inhabit	VERB
alces-443	47	27	broad	broad	ADJ
alces-443	47	28	arrays	array	NOUN
alces-443	47	29	of	of	ADP
alces-443	47	30	habitats	habitat	NOUN
alces-443	47	31	compared	compare	VERB
alces-443	47	32	with	with	ADP
alces-443	47	33	habitat	habitat	NOUN
alces-443	47	34	specialists	specialist	NOUN
alces-443	47	35	(	(	PUNCT
alces-443	47	36	nevo	nevo	PROPN
alces-443	47	37	1978	1978	NUM
alces-443	47	38	)	)	PUNCT
alces-443	47	39	,	,	PUNCT
alces-443	47	40	or	or	CCONJ
alces-443	47	41	that	that	SCONJ
alces-443	47	42	northern	northern	ADJ
alces-443	47	43	cervids	cervid	NOUN
alces-443	47	44	inhabiting	inhabit	VERB
alces-443	47	45	boreal	boreal	ADJ
alces-443	47	46	forests	forest	NOUN
alces-443	47	47	were	be	AUX
alces-443	47	48	less	less	ADV
alces-443	47	49	variable	variable	ADJ
alces-443	47	50	than	than	ADP
alces-443	47	51	their	their	PRON
alces-443	47	52	relatives	relative	NOUN
alces-443	47	53	to	to	ADP
alces-443	47	54	the	the	DET
alces-443	47	55	south	south	NOUN
alces-443	47	56	(	(	PUNCT
alces-443	47	57	smith	smith	PROPN
alces-443	47	58	et	et	PROPN
alces-443	47	59	al	al	PROPN
alces-443	47	60	.	.	PROPN
alces-443	47	61	1990	1990	NUM
alces-443	47	62	)	)	PUNCT
alces-443	47	63	.	.	PUNCT
alces-443	48	1	analyses	analysis	NOUN
alces-443	48	2	of	of	ADP
alces-443	48	3	23	23	NUM
alces-443	48	4	allozyme	allozyme	NOUN
alces-443	48	5	loci	locus	NOUN
alces-443	48	6	in	in	ADP
alces-443	48	7	>	>	X
alces-443	48	8	700	700	NUM
alces-443	48	9	individuals	individual	NOUN
alces-443	48	10	representing	represent	VERB
alces-443	48	11	18	18	NUM
alces-443	48	12	moose	moose	NOUN
alces-443	48	13	populations	population	NOUN
alces-443	48	14	in	in	ADP
alces-443	48	15	scandinavia	scandinavia	PROPN
alces-443	48	16	were	be	AUX
alces-443	48	17	required	require	VERB
alces-443	48	18	to	to	PART
alces-443	48	19	reveal	reveal	VERB
alces-443	48	20	extensive	extensive	ADJ
alces-443	48	21	genetic	genetic	ADJ
alces-443	48	22	variation	variation	NOUN
alces-443	48	23	in	in	ADP
alces-443	48	24	moose	moose	NOUN
alces-443	48	25	(	(	PUNCT
alces-443	48	26	ryman	ryman	NOUN
alces-443	48	27	et	et	PROPN
alces-443	48	28	al	al	PROPN
alces-443	48	29	.	.	PROPN
alces-443	48	30	1980	1980	NUM
alces-443	48	31	)	)	PUNCT
alces-443	48	32	.	.	PUNCT
alces-443	49	1	those	those	DET
alces-443	49	2	authors	author	NOUN
alces-443	49	3	refuted	refute	VERB
alces-443	49	4	the	the	DET
alces-443	49	5	environmental	environmental	ADJ
alces-443	49	6	-	-	PUNCT
alces-443	49	7	grain	grain	NOUN
alces-443	49	8	hypothesis	hypothesis	NOUN
alces-443	49	9	,	,	PUNCT
alces-443	49	10	concluding	conclude	VERB
alces-443	49	11	that	that	SCONJ
alces-443	49	12	large	large	ADJ
alces-443	49	13	mammals	mammal	NOUN
alces-443	49	14	in	in	ADP
alces-443	49	15	general	general	ADJ
alces-443	49	16	,	,	PUNCT
alces-443	49	17	and	and	CCONJ
alces-443	49	18	large	large	ADJ
alces-443	49	19	cervids	cervid	NOUN
alces-443	49	20	in	in	ADP
alces-443	49	21	particular	particular	ADJ
alces-443	49	22	,	,	PUNCT
alces-443	49	23	are	be	AUX
alces-443	49	24	not	not	PART
alces-443	49	25	naturally	naturally	ADV
alces-443	49	26	monomorphic	monomorphic	ADJ
alces-443	49	27	;	;	PUNCT
alces-443	49	28	previous	previous	ADJ
alces-443	49	29	studies	study	NOUN
alces-443	49	30	of	of	ADP
alces-443	49	31	moose	moose	NOUN
alces-443	49	32	had	have	AUX
alces-443	49	33	examined	examine	VERB
alces-443	49	34	too	too	ADV
alces-443	49	35	few	few	ADJ
alces-443	49	36	loci	locus	NOUN
alces-443	49	37	or	or	CCONJ
alces-443	49	38	individuals	individual	NOUN
alces-443	49	39	to	to	PART
alces-443	49	40	detect	detect	VERB
alces-443	49	41	variation	variation	NOUN
alces-443	49	42	.	.	PUNCT
alces-443	50	1	nonetheless	nonetheless	ADV
alces-443	50	2	,	,	PUNCT
alces-443	50	3	those	those	DET
alces-443	50	4	authors	author	NOUN
alces-443	50	5	noted	note	VERB
alces-443	50	6	that	that	SCONJ
alces-443	50	7	genetic	genetic	ADJ
alces-443	50	8	variability	variability	NOUN
alces-443	50	9	in	in	ADP
alces-443	50	10	moose	moose	NOUN
alces-443	50	11	was	be	AUX
alces-443	50	12	somewhat	somewhat	ADV
alces-443	50	13	less	less	ADJ
alces-443	50	14	than	than	ADP
alces-443	50	15	that	that	PRON
alces-443	50	16	observed	observe	VERB
alces-443	50	17	in	in	ADP
alces-443	50	18	many	many	ADJ
alces-443	50	19	other	other	ADJ
alces-443	50	20	species	specie	NOUN
alces-443	50	21	of	of	ADP
alces-443	50	22	mammals	mammal	NOUN
alces-443	50	23	(	(	PUNCT
alces-443	50	24	nevo	nevo	NOUN
alces-443	50	25	1978	1978	NUM
alces-443	50	26	)	)	PUNCT
alces-443	50	27	,	,	PUNCT
alces-443	50	28	but	but	CCONJ
alces-443	50	29	that	that	SCONJ
alces-443	50	30	genetic	genetic	ADJ
alces-443	50	31	drift	drift	NOUN
alces-443	50	32	due	due	ADP
alces-443	50	33	to	to	ADP
alces-443	50	34	small	small	ADJ
alces-443	50	35	historic	historic	ADJ
alces-443	50	36	population	population	NOUN
alces-443	50	37	size	size	NOUN
alces-443	50	38	was	be	AUX
alces-443	50	39	a	a	DET
alces-443	50	40	more	more	ADV
alces-443	50	41	likely	likely	ADJ
alces-443	50	42	explanation	explanation	NOUN
alces-443	50	43	than	than	ADP
alces-443	50	44	any	any	DET
alces-443	50	45	specific	specific	ADJ
alces-443	50	46	evolutionary	evolutionary	ADJ
alces-443	50	47	strategy	strategy	NOUN
alces-443	50	48	.	.	PUNCT
alces-443	51	1	thus	thus	ADV
alces-443	51	2	,	,	PUNCT
alces-443	51	3	genetic	genetic	ADJ
alces-443	51	4	drift	drift	NOUN
alces-443	51	5	was	be	AUX
alces-443	51	6	once	once	ADV
alces-443	51	7	again	again	ADV
alces-443	51	8	proposed	propose	VERB
alces-443	51	9	as	as	ADP
alces-443	51	10	being	be	AUX
alces-443	51	11	an	an	DET
alces-443	51	12	important	important	ADJ
alces-443	51	13	factor	factor	NOUN
alces-443	51	14	in	in	ADP
alces-443	51	15	determining	determine	VERB
alces-443	51	16	the	the	DET
alces-443	51	17	structure	structure	NOUN
alces-443	51	18	of	of	ADP
alces-443	51	19	genetic	genetic	ADJ
alces-443	51	20	diversity	diversity	NOUN
alces-443	51	21	.	.	PUNCT
alces-443	52	1	in	in	ADP
alces-443	52	2	another	another	DET
alces-443	52	3	comprehensive	comprehensive	ADJ
alces-443	52	4	study	study	NOUN
alces-443	52	5	,	,	PUNCT
alces-443	52	6	chesser	chesser	PROPN
alces-443	52	7	et	et	PROPN
alces-443	52	8	al	al	PROPN
alces-443	52	9	.	.	PROPN
alces-443	53	1	(	(	PUNCT
alces-443	53	2	1982	1982	NUM
alces-443	53	3	)	)	PUNCT
alces-443	53	4	examined	examine	VERB
alces-443	53	5	1,169	1,169	NUM
alces-443	53	6	individuals	individual	NOUN
alces-443	53	7	from	from	ADP
alces-443	53	8	4	4	NUM
alces-443	53	9	regions	region	NOUN
alces-443	53	10	in	in	ADP
alces-443	53	11	sweden	sweden	PROPN
alces-443	53	12	for	for	ADP
alces-443	53	13	a	a	DET
alces-443	53	14	single	single	ADJ
alces-443	53	15	polymorphic	polymorphic	ADJ
alces-443	53	16	locus	locus	NOUN
alces-443	53	17	and	and	CCONJ
alces-443	53	18	reported	report	VERB
alces-443	53	19	variation	variation	NOUN
alces-443	53	20	in	in	ADP
alces-443	53	21	allele	allele	NOUN
alces-443	53	22	frequencies	frequency	NOUN
alces-443	53	23	among	among	ADP
alces-443	53	24	those	those	DET
alces-443	53	25	regions	region	NOUN
alces-443	53	26	and	and	CCONJ
alces-443	53	27	,	,	PUNCT
alces-443	53	28	perhaps	perhaps	ADV
alces-443	53	29	more	more	ADV
alces-443	53	30	importantly	importantly	ADV
alces-443	53	31	,	,	PUNCT
alces-443	53	32	significant	significant	ADJ
alces-443	53	33	variation	variation	NOUN
alces-443	53	34	within	within	ADP
alces-443	53	35	1	1	NUM
alces-443	53	36	of	of	ADP
alces-443	53	37	those	those	DET
alces-443	53	38	regions	region	NOUN
alces-443	53	39	.	.	PUNCT
alces-443	54	1	detecting	detect	VERB
alces-443	54	2	variation	variation	NOUN
alces-443	54	3	at	at	ADP
alces-443	54	4	geographic	geographic	ADJ
alces-443	54	5	scales	scale	NOUN
alces-443	54	6	small	small	ADJ
alces-443	54	7	enough	enough	ADV
alces-443	54	8	to	to	PART
alces-443	54	9	be	be	AUX
alces-443	54	10	considered	consider	VERB
alces-443	54	11	within	within	ADP
alces-443	54	12	a	a	DET
alces-443	54	13	single	single	ADJ
alces-443	54	14	population	population	NOUN
alces-443	54	15	illustrated	illustrate	VERB
alces-443	54	16	that	that	DET
alces-443	54	17	structure	structure	NOUN
alces-443	54	18	of	of	ADP
alces-443	54	19	moose	moose	NOUN
alces-443	54	20	populations	population	NOUN
alces-443	54	21	existed	exist	VERB
alces-443	54	22	at	at	ADP
alces-443	54	23	scales	scale	NOUN
alces-443	54	24	smaller	small	ADJ
alces-443	54	25	than	than	SCONJ
alces-443	54	26	previously	previously	ADV
alces-443	54	27	imagined	imagine	VERB
alces-443	54	28	,	,	PUNCT
alces-443	54	29	and	and	CCONJ
alces-443	54	30	that	that	DET
alces-443	54	31	vagility	vagility	NOUN
alces-443	54	32	was	be	AUX
alces-443	54	33	not	not	PART
alces-443	54	34	inconsistent	inconsistent	ADJ
alces-443	54	35	with	with	ADP
alces-443	54	36	genetic	genetic	ADJ
alces-443	54	37	structuring	structuring	NOUN
alces-443	54	38	.	.	PUNCT
alces-443	55	1	the	the	DET
alces-443	55	2	most	most	ADV
alces-443	55	3	recent	recent	ADJ
alces-443	55	4	study	study	NOUN
alces-443	55	5	of	of	ADP
alces-443	55	6	allozyme	allozyme	NOUN
alces-443	55	7	variation	variation	NOUN
alces-443	55	8	in	in	ADP
alces-443	55	9	moose	moose	NOUN
alces-443	55	10	reported	report	VERB
alces-443	55	11	extensive	extensive	ADJ
alces-443	55	12	variation	variation	NOUN
alces-443	55	13	in	in	ADP
alces-443	55	14	a	a	DET
alces-443	55	15	moose	moose	NOUN
alces-443	55	16	population	population	NOUN
alces-443	55	17	in	in	ADP
alces-443	55	18	alaska	alaska	PROPN
alces-443	55	19	(	(	PUNCT
alces-443	55	20	hundertmark	hundertmark	PROPN
alces-443	55	21	et	et	PROPN
alces-443	55	22	al	al	PROPN
alces-443	55	23	.	.	PROPN
alces-443	55	24	1992	1992	NUM
alces-443	55	25	)	)	PUNCT
alces-443	55	26	.	.	PUNCT
alces-443	56	1	the	the	DET
alces-443	56	2	level	level	NOUN
alces-443	56	3	of	of	ADP
alces-443	56	4	genetic	genetic	ADJ
alces-443	56	5	diversity	diversity	NOUN
alces-443	56	6	observed	observe	VERB
alces-443	56	7	was	be	AUX
alces-443	56	8	greater	great	ADJ
alces-443	56	9	than	than	ADP
alces-443	56	10	that	that	PRON
alces-443	56	11	reported	report	VERB
alces-443	56	12	elsewhere	elsewhere	ADV
alces-443	56	13	in	in	ADP
alces-443	56	14	moose	moose	NOUN
alces-443	56	15	and	and	CCONJ
alces-443	56	16	was	be	AUX
alces-443	56	17	similar	similar	ADJ
alces-443	56	18	to	to	ADP
alces-443	56	19	levels	level	NOUN
alces-443	56	20	observed	observe	VERB
alces-443	56	21	in	in	ADP
alces-443	56	22	white	white	ADJ
alces-443	56	23	-	-	PUNCT
alces-443	56	24	tailed	tail	VERB
alces-443	56	25	deer	deer	NOUN
alces-443	56	26	,	,	PUNCT
alces-443	56	27	a	a	DET
alces-443	56	28	species	specie	NOUN
alces-443	56	29	known	know	VERB
alces-443	56	30	for	for	ADP
alces-443	56	31	extensive	extensive	ADJ
alces-443	56	32	allozyme	allozyme	NOUN
alces-443	56	33	diversity	diversity	NOUN
alces-443	56	34	(	(	PUNCT
alces-443	56	35	smith	smith	PROPN
alces-443	56	36	et	et	PROPN
alces-443	56	37	al	al	PROPN
alces-443	56	38	.	.	PROPN
alces-443	56	39	1984	1984	NUM
alces-443	56	40	)	)	PUNCT
alces-443	56	41	.	.	PUNCT
alces-443	57	1	hundertmark	hundertmark	PROPN
alces-443	57	2	et	et	PROPN
alces-443	57	3	al	al	PROPN
alces-443	57	4	.	.	PROPN
alces-443	58	1	(	(	PUNCT
alces-443	58	2	1992	1992	NUM
alces-443	58	3	)	)	PUNCT
alces-443	58	4	hypothesized	hypothesize	VERB
alces-443	58	5	that	that	SCONJ
alces-443	58	6	lesser	less	ADJ
alces-443	58	7	levels	level	NOUN
alces-443	58	8	of	of	ADP
alces-443	58	9	variability	variability	NOUN
alces-443	58	10	described	describe	VERB
alces-443	58	11	in	in	ADP
alces-443	58	12	moose	moose	NOUN
alces-443	58	13	from	from	ADP
alces-443	58	14	scandinavia	scandinavia	PROPN
alces-443	58	15	and	and	CCONJ
alces-443	58	16	other	other	ADJ
alces-443	58	17	regions	region	NOUN
alces-443	58	18	of	of	ADP
alces-443	58	19	north	north	PROPN
alces-443	58	20	america	america	PROPN
alces-443	58	21	were	be	AUX
alces-443	58	22	attributable	attributable	ADJ
alces-443	58	23	to	to	ADP
alces-443	58	24	glacial	glacial	ADJ
alces-443	58	25	history	history	NOUN
alces-443	58	26	.	.	PUNCT
alces-443	59	1	all	all	DET
alces-443	59	2	other	other	ADJ
alces-443	59	3	moose	moose	NOUN
alces-443	59	4	populations	population	NOUN
alces-443	59	5	studied	study	VERB
alces-443	59	6	to	to	ADP
alces-443	59	7	that	that	DET
alces-443	59	8	point	point	NOUN
alces-443	59	9	occurred	occur	VERB
alces-443	59	10	in	in	ADP
alces-443	59	11	previously	previously	ADV
alces-443	59	12	glaciated	glaciate	VERB
alces-443	59	13	terrain	terrain	NOUN
alces-443	59	14	that	that	PRON
alces-443	59	15	was	be	AUX
alces-443	59	16	colonized	colonize	VERB
alces-443	59	17	by	by	ADP
alces-443	59	18	moose	moose	NOUN
alces-443	59	19	after	after	ADP
alces-443	59	20	retreat	retreat	NOUN
alces-443	59	21	of	of	ADP
alces-443	59	22	pleistocene	pleistocene	NOUN
alces-443	59	23	ice	ice	NOUN
alces-443	59	24	sheets	sheet	NOUN
alces-443	59	25	.	.	PUNCT
alces-443	60	1	colonization	colonization	NOUN
alces-443	60	2	of	of	ADP
alces-443	60	3	previously	previously	ADV
alces-443	60	4	glaciated	glaciate	VERB
alces-443	60	5	areas	area	NOUN
alces-443	60	6	could	could	AUX
alces-443	60	7	have	have	AUX
alces-443	60	8	resulted	result	VERB
alces-443	60	9	in	in	ADP
alces-443	60	10	serial	serial	ADJ
alces-443	60	11	founder	founder	NOUN
alces-443	60	12	events	event	NOUN
alces-443	60	13	that	that	PRON
alces-443	60	14	reduced	reduce	VERB
alces-443	60	15	genetic	genetic	ADJ
alces-443	60	16	diversity	diversity	NOUN
alces-443	60	17	(	(	PUNCT
alces-443	60	18	sage	sage	NOUN
alces-443	60	19	and	and	CCONJ
alces-443	60	20	wolff	wolff	NOUN
alces-443	60	21	1986	1986	NUM
alces-443	60	22	)	)	PUNCT
alces-443	60	23	.	.	PUNCT
alces-443	61	1	hundertmark	hundertmark	PROPN
alces-443	61	2	et	et	PROPN
alces-443	61	3	al	al	PROPN
alces-443	61	4	.	.	PROPN
alces-443	62	1	(	(	PUNCT
alces-443	62	2	1992	1992	NUM
alces-443	62	3	)	)	PUNCT
alces-443	62	4	argued	argue	VERB
alces-443	62	5	that	that	SCONJ
alces-443	62	6	alaska	alaska	PROPN
alces-443	62	7	could	could	AUX
alces-443	62	8	have	have	AUX
alces-443	62	9	served	serve	VERB
alces-443	62	10	as	as	ADP
alces-443	62	11	a	a	DET
alces-443	62	12	refugium	refugium	NOUN
alces-443	62	13	for	for	ADP
alces-443	62	14	moose	moose	NOUN
alces-443	62	15	in	in	ADP
alces-443	62	16	which	which	PRON
alces-443	62	17	genetic	genetic	ADJ
alces-443	62	18	diversity	diversity	NOUN
alces-443	62	19	could	could	AUX
alces-443	62	20	have	have	AUX
alces-443	62	21	been	be	AUX
alces-443	62	22	maintained	maintain	VERB
alces-443	62	23	because	because	SCONJ
alces-443	62	24	of	of	ADP
alces-443	62	25	a	a	DET
alces-443	62	26	large	large	ADJ
alces-443	62	27	effective	effective	ADJ
alces-443	62	28	population	population	NOUN
alces-443	62	29	size	size	NOUN
alces-443	62	30	.	.	PUNCT
alces-443	63	1	assessing	assess	VERB
alces-443	63	2	variation	variation	NOUN
alces-443	63	3	at	at	ADP
alces-443	63	4	the	the	DET
alces-443	63	5	sequence	sequence	NOUN
alces-443	63	6	level	level	NOUN
alces-443	63	7	the	the	DET
alces-443	63	8	first	first	ADJ
alces-443	63	9	investigation	investigation	NOUN
alces-443	63	10	of	of	ADP
alces-443	63	11	highly	highly	ADV
alces-443	63	12	polymorphic	polymorphic	ADJ
alces-443	63	13	molecular	molecular	ADJ
alces-443	63	14	markers	marker	NOUN
alces-443	63	15	in	in	ADP
alces-443	63	16	moose	moose	NOUN
alces-443	63	17	documented	document	VERB
alces-443	63	18	4	4	NUM
alces-443	63	19	alleles	allele	NOUN
alces-443	63	20	in	in	ADP
alces-443	63	21	a	a	DET
alces-443	63	22	microsatellite	microsatellite	ADJ
alces-443	63	23	-	-	PUNCT
alces-443	63	24	like	like	ADJ
alces-443	63	25	hundertmark	hundertmark	NOUN
alces-443	63	26	and	and	CCONJ
alces-443	63	27	bowyer	bowyer	PROPN
alces-443	63	28	moose	moose	PROPN
alces-443	63	29	genetics	genetics	PROPN
alces-443	63	30	alces	alce	NOUN
alces-443	63	31	vol	vol	NOUN
alces-443	63	32	.	.	PROPN
alces-443	63	33	40	40	NUM
alces-443	63	34	,	,	PUNCT
alces-443	63	35	2004	2004	NUM
alces-443	63	36	106	106	NUM
alces-443	63	37	locus	locus	NOUN
alces-443	63	38	in	in	ADP
alces-443	63	39	17	17	NUM
alces-443	63	40	individuals	individual	NOUN
alces-443	63	41	from	from	ADP
alces-443	63	42	sweden	sweden	PROPN
alces-443	63	43	(	(	PUNCT
alces-443	63	44	ellegren	ellegren	PROPN
alces-443	63	45	et	et	PROPN
alces-443	63	46	al	al	PROPN
alces-443	63	47	.	.	PROPN
alces-443	63	48	1991	1991	NUM
alces-443	63	49	)	)	PUNCT
alces-443	63	50	.	.	PUNCT
alces-443	64	1	eight	eight	NUM
alces-443	64	2	genotypes	genotype	NOUN
alces-443	64	3	were	be	AUX
alces-443	64	4	reported	report	VERB
alces-443	64	5	,	,	PUNCT
alces-443	64	6	which	which	PRON
alces-443	64	7	represented	represent	VERB
alces-443	64	8	a	a	DET
alces-443	64	9	heretofore	heretofore	ADV
alces-443	64	10	unthinkable	unthinkable	ADJ
alces-443	64	11	level	level	NOUN
alces-443	64	12	of	of	ADP
alces-443	64	13	polymorphism	polymorphism	NOUN
alces-443	64	14	.	.	PUNCT
alces-443	65	1	those	those	DET
alces-443	65	2	authors	author	NOUN
alces-443	65	3	investigated	investigate	VERB
alces-443	65	4	the	the	DET
alces-443	65	5	inheritance	inheritance	NOUN
alces-443	65	6	of	of	ADP
alces-443	65	7	that	that	DET
alces-443	65	8	locus	locus	NOUN
alces-443	65	9	in	in	ADP
alces-443	65	10	a	a	DET
alces-443	65	11	2	2	NUM
alces-443	65	12	-	-	PUNCT
alces-443	65	13	generation	generation	NOUN
alces-443	65	14	pedigree	pedigree	NOUN
alces-443	65	15	and	and	CCONJ
alces-443	65	16	determined	determine	VERB
alces-443	65	17	that	that	SCONJ
alces-443	65	18	the	the	DET
alces-443	65	19	alleles	allele	NOUN
alces-443	65	20	exhibited	exhibit	VERB
alces-443	65	21	mendelian	mendelian	ADJ
alces-443	65	22	inheritance	inheritance	NOUN
alces-443	65	23	.	.	PUNCT
alces-443	66	1	that	that	DET
alces-443	66	2	study	study	NOUN
alces-443	66	3	and	and	CCONJ
alces-443	66	4	others	other	NOUN
alces-443	66	5	like	like	ADP
alces-443	66	6	it	it	PRON
alces-443	66	7	laid	lay	VERB
alces-443	66	8	the	the	DET
alces-443	66	9	foundation	foundation	NOUN
alces-443	66	10	for	for	ADP
alces-443	66	11	the	the	DET
alces-443	66	12	explosion	explosion	NOUN
alces-443	66	13	of	of	ADP
alces-443	66	14	interest	interest	NOUN
alces-443	66	15	in	in	ADP
alces-443	66	16	population	population	NOUN
alces-443	66	17	genetics	genetic	NOUN
alces-443	66	18	and	and	CCONJ
alces-443	66	19	phylogenetics	phylogenetic	NOUN
alces-443	66	20	based	base	VERB
alces-443	66	21	on	on	ADP
alces-443	66	22	molecular	molecular	ADJ
alces-443	66	23	markers	marker	NOUN
alces-443	66	24	and	and	CCONJ
alces-443	66	25	the	the	DET
alces-443	66	26	polymerase	polymerase	NOUN
alces-443	66	27	chain	chain	NOUN
alces-443	66	28	reaction	reaction	NOUN
alces-443	66	29	(	(	PUNCT
alces-443	66	30	pcr	pcr	PROPN
alces-443	66	31	;	;	PUNCT
alces-443	66	32	mullis	mullis	PROPN
alces-443	66	33	et	et	PROPN
alces-443	66	34	al	al	PROPN
alces-443	66	35	.	.	PROPN
alces-443	66	36	1986	1986	NUM
alces-443	66	37	)	)	PUNCT
alces-443	66	38	.	.	PUNCT
alces-443	67	1	the	the	DET
alces-443	67	2	advent	advent	NOUN
alces-443	67	3	of	of	ADP
alces-443	67	4	pcr	pcr	PROPN
alces-443	67	5	represented	represent	VERB
alces-443	67	6	one	one	NUM
alces-443	67	7	of	of	ADP
alces-443	67	8	the	the	DET
alces-443	67	9	truly	truly	ADV
alces-443	67	10	significant	significant	ADJ
alces-443	67	11	advances	advance	NOUN
alces-443	67	12	in	in	ADP
alces-443	67	13	the	the	DET
alces-443	67	14	history	history	NOUN
alces-443	67	15	of	of	ADP
alces-443	67	16	molecular	molecular	ADJ
alces-443	67	17	biology	biology	NOUN
alces-443	67	18	and	and	CCONJ
alces-443	67	19	genetics	genetic	NOUN
alces-443	67	20	.	.	PUNCT
alces-443	68	1	the	the	DET
alces-443	68	2	process	process	NOUN
alces-443	68	3	allows	allow	VERB
alces-443	68	4	in	in	ADP
alces-443	68	5	-	-	PUNCT
alces-443	68	6	vitro	vitro	X
alces-443	68	7	amplification	amplification	NOUN
alces-443	68	8	of	of	ADP
alces-443	68	9	dna	dna	NOUN
alces-443	68	10	from	from	ADP
alces-443	68	11	miniscule	miniscule	ADJ
alces-443	68	12	amounts	amount	NOUN
alces-443	68	13	of	of	ADP
alces-443	68	14	starting	start	VERB
alces-443	68	15	material	material	NOUN
alces-443	68	16	(	(	PUNCT
alces-443	68	17	theoretically	theoretically	ADV
alces-443	68	18	as	as	ADV
alces-443	68	19	little	little	ADJ
alces-443	68	20	as	as	ADP
alces-443	68	21	1	1	NUM
alces-443	68	22	molecule	molecule	NOUN
alces-443	68	23	of	of	ADP
alces-443	68	24	dna	dna	PROPN
alces-443	68	25	)	)	PUNCT
alces-443	68	26	to	to	PART
alces-443	68	27	provide	provide	VERB
alces-443	68	28	sufficient	sufficient	ADJ
alces-443	68	29	quantities	quantity	NOUN
alces-443	68	30	for	for	ADP
alces-443	68	31	analysis	analysis	NOUN
alces-443	68	32	.	.	PUNCT
alces-443	69	1	no	no	ADV
alces-443	69	2	longer	long	ADV
alces-443	69	3	were	be	AUX
alces-443	69	4	researchers	researcher	NOUN
alces-443	69	5	required	require	VERB
alces-443	69	6	to	to	PART
alces-443	69	7	sacrifice	sacrifice	VERB
alces-443	69	8	animals	animal	NOUN
alces-443	69	9	to	to	PART
alces-443	69	10	acquire	acquire	VERB
alces-443	69	11	sufficient	sufficient	ADJ
alces-443	69	12	quantities	quantity	NOUN
alces-443	69	13	and	and	CCONJ
alces-443	69	14	types	type	NOUN
alces-443	69	15	of	of	ADP
alces-443	69	16	tissues	tissue	NOUN
alces-443	69	17	for	for	ADP
alces-443	69	18	genetic	genetic	ADJ
alces-443	69	19	analyses	analysis	NOUN
alces-443	69	20	because	because	SCONJ
alces-443	69	21	any	any	DET
alces-443	69	22	nucleated	nucleated	ADJ
alces-443	69	23	cell	cell	NOUN
alces-443	69	24	held	hold	VERB
alces-443	69	25	the	the	DET
alces-443	69	26	complete	complete	ADJ
alces-443	69	27	genetic	genetic	ADJ
alces-443	69	28	complement	complement	NOUN
alces-443	69	29	of	of	ADP
alces-443	69	30	the	the	DET
alces-443	69	31	individual	individual	NOUN
alces-443	69	32	.	.	PUNCT
alces-443	70	1	pcr	pcr	PROPN
alces-443	70	2	offered	offer	VERB
alces-443	70	3	unsurpassed	unsurpassed	ADJ
alces-443	70	4	access	access	NOUN
alces-443	70	5	to	to	ADP
alces-443	70	6	the	the	DET
alces-443	70	7	genome	genome	NOUN
alces-443	70	8	,	,	PUNCT
alces-443	70	9	and	and	CCONJ
alces-443	70	10	researchers	researcher	NOUN
alces-443	70	11	soon	soon	ADV
alces-443	70	12	applied	apply	VERB
alces-443	70	13	that	that	PRON
alces-443	70	14	to	to	PART
alces-443	70	15	study	study	VERB
alces-443	70	16	genetics	genetic	NOUN
alces-443	70	17	of	of	ADP
alces-443	70	18	moose	moose	NOUN
alces-443	70	19	.	.	PUNCT
alces-443	71	1	mikko	mikko	PROPN
alces-443	71	2	and	and	CCONJ
alces-443	71	3	andersson	andersson	PROPN
alces-443	71	4	(	(	PUNCT
alces-443	71	5	1995	1995	NUM
alces-443	71	6	)	)	PUNCT
alces-443	71	7	conducted	conduct	VERB
alces-443	71	8	the	the	DET
alces-443	71	9	first	first	ADJ
alces-443	71	10	analysis	analysis	NOUN
alces-443	71	11	of	of	ADP
alces-443	71	12	functional	functional	ADJ
alces-443	71	13	loci	locus	NOUN
alces-443	71	14	in	in	ADP
alces-443	71	15	an	an	DET
alces-443	71	16	analysis	analysis	NOUN
alces-443	71	17	of	of	ADP
alces-443	71	18	variation	variation	NOUN
alces-443	71	19	in	in	ADP
alces-443	71	20	the	the	DET
alces-443	71	21	major	major	ADJ
alces-443	71	22	histocompatibility	histocompatibility	NOUN
alces-443	71	23	complex	complex	NOUN
alces-443	71	24	(	(	PUNCT
alces-443	71	25	mhc	mhc	PROPN
alces-443	71	26	)	)	PUNCT
alces-443	71	27	in	in	ADP
alces-443	71	28	moose	moose	NOUN
alces-443	71	29	from	from	ADP
alces-443	71	30	sweden	sweden	PROPN
alces-443	71	31	and	and	CCONJ
alces-443	71	32	canada	canada	PROPN
alces-443	71	33	.	.	PUNCT
alces-443	72	1	the	the	DET
alces-443	72	2	mhc	mhc	PROPN
alces-443	72	3	is	be	AUX
alces-443	72	4	a	a	DET
alces-443	72	5	family	family	NOUN
alces-443	72	6	of	of	ADP
alces-443	72	7	genes	gene	NOUN
alces-443	72	8	important	important	ADJ
alces-443	72	9	in	in	ADP
alces-443	72	10	immune	immune	ADJ
alces-443	72	11	system	system	NOUN
alces-443	72	12	function	function	NOUN
alces-443	72	13	,	,	PUNCT
alces-443	72	14	and	and	CCONJ
alces-443	72	15	low	low	ADJ
alces-443	72	16	levels	level	NOUN
alces-443	72	17	of	of	ADP
alces-443	72	18	diversity	diversity	NOUN
alces-443	72	19	of	of	ADP
alces-443	72	20	mhc	mhc	PROPN
alces-443	72	21	alleles	allele	NOUN
alces-443	72	22	have	have	AUX
alces-443	72	23	been	be	AUX
alces-443	72	24	interpreted	interpret	VERB
alces-443	72	25	as	as	ADP
alces-443	72	26	indicators	indicator	NOUN
alces-443	72	27	of	of	ADP
alces-443	72	28	lost	lose	VERB
alces-443	72	29	evolutionary	evolutionary	ADJ
alces-443	72	30	potential	potential	NOUN
alces-443	72	31	and	and	CCONJ
alces-443	72	32	increased	increase	VERB
alces-443	72	33	susceptibility	susceptibility	NOUN
alces-443	72	34	to	to	ADP
alces-443	72	35	pathogens	pathogen	NOUN
alces-443	72	36	(	(	PUNCT
alces-443	72	37	hedrick	hedrick	PROPN
alces-443	72	38	1994	1994	NUM
alces-443	72	39	)	)	PUNCT
alces-443	72	40	.	.	PUNCT
alces-443	73	1	mikko	mikko	PROPN
alces-443	73	2	and	and	CCONJ
alces-443	73	3	andersson	andersson	PROPN
alces-443	73	4	(	(	PUNCT
alces-443	73	5	1995	1995	NUM
alces-443	73	6	)	)	PUNCT
alces-443	73	7	noted	note	VERB
alces-443	73	8	very	very	ADV
alces-443	73	9	low	low	ADJ
alces-443	73	10	levels	level	NOUN
alces-443	73	11	of	of	ADP
alces-443	73	12	mhc	mhc	PROPN
alces-443	73	13	variation	variation	NOUN
alces-443	73	14	in	in	ADP
alces-443	73	15	both	both	CCONJ
alces-443	73	16	swedish	swedish	ADJ
alces-443	73	17	and	and	CCONJ
alces-443	73	18	canadian	canadian	ADJ
alces-443	73	19	moose	moose	NOUN
alces-443	73	20	.	.	PUNCT
alces-443	74	1	moreover	moreover	ADV
alces-443	74	2	,	,	PUNCT
alces-443	74	3	those	those	DET
alces-443	74	4	authors	author	NOUN
alces-443	74	5	documented	document	VERB
alces-443	74	6	similarity	similarity	NOUN
alces-443	74	7	among	among	ADP
alces-443	74	8	alleles	allele	NOUN
alces-443	74	9	between	between	ADP
alces-443	74	10	continents	continent	NOUN
alces-443	74	11	and	and	CCONJ
alces-443	74	12	inferred	infer	VERB
alces-443	74	13	the	the	DET
alces-443	74	14	existence	existence	NOUN
alces-443	74	15	of	of	ADP
alces-443	74	16	a	a	DET
alces-443	74	17	bottleneck	bottleneck	NOUN
alces-443	74	18	in	in	ADP
alces-443	74	19	an	an	DET
alces-443	74	20	ancient	ancient	ADJ
alces-443	74	21	moose	moose	NOUN
alces-443	74	22	lineage	lineage	NOUN
alces-443	74	23	prior	prior	ADV
alces-443	74	24	to	to	ADP
alces-443	74	25	divergence	divergence	NOUN
alces-443	74	26	of	of	ADP
alces-443	74	27	european	european	ADJ
alces-443	74	28	and	and	CCONJ
alces-443	74	29	canadian	canadian	ADJ
alces-443	74	30	lineages	lineage	NOUN
alces-443	74	31	.	.	PUNCT
alces-443	75	1	mikko	mikko	PROPN
alces-443	75	2	and	and	CCONJ
alces-443	75	3	andersson	andersson	PROPN
alces-443	75	4	(	(	PUNCT
alces-443	75	5	1995	1995	NUM
alces-443	75	6	)	)	PUNCT
alces-443	75	7	applied	apply	VERB
alces-443	75	8	a	a	DET
alces-443	75	9	molecular	molecular	ADJ
alces-443	75	10	clock	clock	NOUN
alces-443	75	11	to	to	ADP
alces-443	75	12	dna	dna	PROPN
alces-443	75	13	sequence	sequence	NOUN
alces-443	75	14	variation	variation	NOUN
alces-443	75	15	in	in	ADP
alces-443	75	16	the	the	DET
alces-443	75	17	control	control	NOUN
alces-443	75	18	region	region	NOUN
alces-443	75	19	of	of	ADP
alces-443	75	20	mitochondrial	mitochondrial	ADJ
alces-443	75	21	dna	dna	NOUN
alces-443	75	22	(	(	PUNCT
alces-443	75	23	mtdna	mtdna	NOUN
alces-443	75	24	)	)	PUNCT
alces-443	75	25	to	to	PART
alces-443	75	26	date	date	NOUN
alces-443	75	27	the	the	DET
alces-443	75	28	time	time	NOUN
alces-443	75	29	of	of	ADP
alces-443	75	30	divergence	divergence	NOUN
alces-443	75	31	of	of	ADP
alces-443	75	32	swedish	swedish	ADJ
alces-443	75	33	and	and	CCONJ
alces-443	75	34	canadian	canadian	ADJ
alces-443	75	35	moose	moose	NOUN
alces-443	75	36	,	,	PUNCT
alces-443	75	37	which	which	PRON
alces-443	75	38	they	they	PRON
alces-443	75	39	estimated	estimate	VERB
alces-443	75	40	at	at	ADP
alces-443	75	41	165,000	165,000	NUM
alces-443	75	42	-	-	SYM
alces-443	75	43	350,000	350,000	NUM
alces-443	75	44	years	year	NOUN
alces-443	75	45	ago	ago	ADV
alces-443	75	46	.	.	PUNCT
alces-443	76	1	the	the	DET
alces-443	76	2	time	time	NOUN
alces-443	76	3	since	since	SCONJ
alces-443	76	4	divergence	divergence	NOUN
alces-443	76	5	of	of	ADP
alces-443	76	6	european	european	ADJ
alces-443	76	7	moose	moose	NOUN
alces-443	76	8	also	also	ADV
alces-443	76	9	was	be	AUX
alces-443	76	10	analyzed	analyze	VERB
alces-443	76	11	by	by	ADP
alces-443	76	12	ellegren	ellegren	PROPN
alces-443	76	13	et	et	PROPN
alces-443	76	14	al	al	PROPN
alces-443	76	15	.	.	PROPN
alces-443	76	16	(	(	PUNCT
alces-443	76	17	1996	1996	NUM
alces-443	76	18	)	)	PUNCT
alces-443	76	19	.	.	PUNCT
alces-443	77	1	they	they	PRON
alces-443	77	2	assessed	assess	VERB
alces-443	77	3	variation	variation	NOUN
alces-443	77	4	in	in	ADP
alces-443	77	5	minisatellite	minisatellite	ADJ
alces-443	77	6	loci	locus	NOUN
alces-443	77	7	of	of	ADP
alces-443	77	8	swedish	swedish	ADJ
alces-443	77	9	moose	moose	NOUN
alces-443	77	10	and	and	CCONJ
alces-443	77	11	concluded	conclude	VERB
alces-443	77	12	that	that	SCONJ
alces-443	77	13	normal	normal	ADJ
alces-443	77	14	evolutionary	evolutionary	ADJ
alces-443	77	15	processes	process	NOUN
alces-443	77	16	could	could	AUX
alces-443	77	17	have	have	AUX
alces-443	77	18	generated	generate	VERB
alces-443	77	19	the	the	DET
alces-443	77	20	amount	amount	NOUN
alces-443	77	21	of	of	ADP
alces-443	77	22	variation	variation	NOUN
alces-443	77	23	observed	observe	VERB
alces-443	77	24	within	within	ADP
alces-443	77	25	10,000	10,000	NUM
alces-443	77	26	-	-	PUNCT
alces-443	77	27	50,000	50,000	NUM
alces-443	77	28	years	year	NOUN
alces-443	77	29	after	after	ADP
alces-443	77	30	a	a	DET
alces-443	77	31	severe	severe	ADJ
alces-443	77	32	bottleneck	bottleneck	NOUN
alces-443	77	33	.	.	PUNCT
alces-443	78	1	their	their	PRON
alces-443	78	2	implication	implication	NOUN
alces-443	78	3	,	,	PUNCT
alces-443	78	4	therefore	therefore	ADV
alces-443	78	5	,	,	PUNCT
alces-443	78	6	was	be	AUX
alces-443	78	7	that	that	SCONJ
alces-443	78	8	the	the	DET
alces-443	78	9	estimate	estimate	NOUN
alces-443	78	10	of	of	ADP
alces-443	78	11	divergence	divergence	NOUN
alces-443	78	12	provided	provide	VERB
alces-443	78	13	by	by	ADP
alces-443	78	14	mikko	mikko	PROPN
alces-443	78	15	and	and	CCONJ
alces-443	78	16	andersson	andersson	PROPN
alces-443	78	17	(	(	PUNCT
alces-443	78	18	1995	1995	NUM
alces-443	78	19	)	)	PUNCT
alces-443	78	20	was	be	AUX
alces-443	78	21	too	too	ADV
alces-443	78	22	old	old	ADJ
alces-443	78	23	by	by	ADP
alces-443	78	24	perhaps	perhaps	ADV
alces-443	78	25	1	1	NUM
alces-443	78	26	order	order	NOUN
alces-443	78	27	of	of	ADP
alces-443	78	28	magnitude	magnitude	NOUN
alces-443	78	29	.	.	PUNCT
alces-443	79	1	surprisingly	surprisingly	ADV
alces-443	79	2	,	,	PUNCT
alces-443	79	3	the	the	DET
alces-443	79	4	two	two	NUM
alces-443	79	5	data	data	NOUN
alces-443	79	6	sets	set	NOUN
alces-443	79	7	are	be	AUX
alces-443	79	8	not	not	PART
alces-443	79	9	inconsistent	inconsistent	ADJ
alces-443	79	10	;	;	PUNCT
alces-443	79	11	indeed	indeed	ADV
alces-443	79	12	,	,	PUNCT
alces-443	79	13	they	they	PRON
alces-443	79	14	show	show	VERB
alces-443	79	15	similar	similar	ADJ
alces-443	79	16	levels	level	NOUN
alces-443	79	17	of	of	ADP
alces-443	79	18	variation	variation	NOUN
alces-443	79	19	considering	consider	VERB
alces-443	79	20	differences	difference	NOUN
alces-443	79	21	in	in	ADP
alces-443	79	22	evolutionary	evolutionary	ADJ
alces-443	79	23	rates	rate	NOUN
alces-443	79	24	among	among	ADP
alces-443	79	25	marker	marker	NOUN
alces-443	79	26	types	type	NOUN
alces-443	79	27	.	.	PUNCT
alces-443	80	1	the	the	DET
alces-443	80	2	differences	difference	NOUN
alces-443	80	3	in	in	ADP
alces-443	80	4	estimates	estimate	NOUN
alces-443	80	5	for	for	ADP
alces-443	80	6	date	date	NOUN
alces-443	80	7	of	of	ADP
alces-443	80	8	divergence	divergence	NOUN
alces-443	80	9	relate	relate	VERB
alces-443	80	10	more	more	ADJ
alces-443	80	11	to	to	ADP
alces-443	80	12	the	the	DET
alces-443	80	13	evolutionary	evolutionary	ADJ
alces-443	80	14	rate	rate	NOUN
alces-443	80	15	estimates	estimate	NOUN
alces-443	80	16	used	use	VERB
alces-443	80	17	than	than	ADP
alces-443	80	18	to	to	ADP
alces-443	80	19	differences	difference	NOUN
alces-443	80	20	in	in	ADP
alces-443	80	21	genetic	genetic	ADJ
alces-443	80	22	variability	variability	NOUN
alces-443	80	23	.	.	PUNCT
alces-443	81	1	microsatellite	microsatellite	ADJ
alces-443	81	2	loci	locus	NOUN
alces-443	81	3	were	be	AUX
alces-443	81	4	first	first	ADV
alces-443	81	5	described	describe	VERB
alces-443	81	6	for	for	ADP
alces-443	81	7	moose	moose	NOUN
alces-443	81	8	by	by	ADP
alces-443	81	9	wilson	wilson	PROPN
alces-443	81	10	et	et	PROPN
alces-443	81	11	al	al	PROPN
alces-443	81	12	.	.	PROPN
alces-443	82	1	(	(	PUNCT
alces-443	82	2	1997	1997	NUM
alces-443	82	3	)	)	PUNCT
alces-443	82	4	and	and	CCONJ
alces-443	82	5	røed	røed	PROPN
alces-443	82	6	and	and	CCONJ
alces-443	82	7	midthjell	midthjell	PROPN
alces-443	82	8	(	(	PUNCT
alces-443	82	9	1998	1998	NUM
alces-443	82	10	)	)	PUNCT
alces-443	82	11	.	.	PUNCT
alces-443	83	1	broders	broder	NOUN
alces-443	83	2	et	et	PROPN
alces-443	83	3	al	al	PROPN
alces-443	83	4	.	.	PUNCT
alces-443	84	1	(	(	PUNCT
alces-443	84	2	1999	1999	NUM
alces-443	84	3	)	)	PUNCT
alces-443	84	4	demonstrated	demonstrate	VERB
alces-443	84	5	the	the	DET
alces-443	84	6	utility	utility	NOUN
alces-443	84	7	of	of	ADP
alces-443	84	8	microsatellites	microsatellite	NOUN
alces-443	84	9	for	for	ADP
alces-443	84	10	assessing	assess	VERB
alces-443	84	11	population	population	NOUN
alces-443	84	12	structure	structure	NOUN
alces-443	84	13	in	in	ADP
alces-443	84	14	moose	moose	NOUN
alces-443	84	15	by	by	ADP
alces-443	84	16	assessing	assess	VERB
alces-443	84	17	consequences	consequence	NOUN
alces-443	84	18	of	of	ADP
alces-443	84	19	founder	founder	NOUN
alces-443	84	20	events	event	NOUN
alces-443	84	21	in	in	ADP
alces-443	84	22	canada	canada	PROPN
alces-443	84	23	.	.	PUNCT
alces-443	85	1	heterozygosity	heterozygosity	NOUN
alces-443	85	2	in	in	ADP
alces-443	85	3	3	3	NUM
alces-443	85	4	populations	population	NOUN
alces-443	85	5	founded	found	VERB
alces-443	85	6	by	by	ADP
alces-443	85	7	few	few	ADJ
alces-443	85	8	individuals	individual	NOUN
alces-443	85	9	decreased	decrease	VERB
alces-443	85	10	from	from	ADP
alces-443	85	11	14	14	NUM
alces-443	85	12	-	-	SYM
alces-443	85	13	30	30	NUM
alces-443	85	14	%	%	NOUN
alces-443	85	15	compared	compare	VERB
alces-443	85	16	with	with	ADP
alces-443	85	17	the	the	DET
alces-443	85	18	source	source	NOUN
alces-443	85	19	population	population	NOUN
alces-443	85	20	.	.	PUNCT
alces-443	86	1	broders	broder	NOUN
alces-443	86	2	et	et	PROPN
alces-443	86	3	al	al	PROPN
alces-443	86	4	.	.	PUNCT
alces-443	87	1	(	(	PUNCT
alces-443	87	2	1999	1999	NUM
alces-443	87	3	)	)	PUNCT
alces-443	87	4	,	,	PUNCT
alces-443	87	5	in	in	ADP
alces-443	87	6	assessing	assess	VERB
alces-443	87	7	variability	variability	NOUN
alces-443	87	8	of	of	ADP
alces-443	87	9	moose	moose	NOUN
alces-443	87	10	on	on	ADP
alces-443	87	11	the	the	DET
alces-443	87	12	island	island	NOUN
alces-443	87	13	of	of	ADP
alces-443	87	14	newfoundland	newfoundland	PROPN
alces-443	87	15	,	,	PUNCT
alces-443	87	16	canada	canada	PROPN
alces-443	87	17	,	,	PUNCT
alces-443	87	18	demonstrated	demonstrate	VERB
alces-443	87	19	that	that	SCONJ
alces-443	87	20	2	2	NUM
alces-443	87	21	consecutive	consecutive	ADJ
alces-443	87	22	founder	founder	NOUN
alces-443	87	23	events	event	NOUN
alces-443	87	24	reduced	reduce	VERB
alces-443	87	25	heterozygosity	heterozygosity	NOUN
alces-443	87	26	by	by	ADP
alces-443	87	27	46	46	NUM
alces-443	87	28	%	%	NOUN
alces-443	87	29	.	.	PUNCT
alces-443	88	1	although	although	SCONJ
alces-443	88	2	they	they	PRON
alces-443	88	3	could	could	AUX
alces-443	88	4	not	not	PART
alces-443	88	5	discern	discern	VERB
alces-443	88	6	any	any	DET
alces-443	88	7	decrease	decrease	NOUN
alces-443	88	8	in	in	ADP
alces-443	88	9	fitness	fitness	NOUN
alces-443	88	10	as	as	ADP
alces-443	88	11	a	a	DET
alces-443	88	12	result	result	NOUN
alces-443	88	13	of	of	ADP
alces-443	88	14	the	the	DET
alces-443	88	15	decrease	decrease	NOUN
alces-443	88	16	in	in	ADP
alces-443	88	17	diversity	diversity	NOUN
alces-443	88	18	,	,	PUNCT
alces-443	88	19	the	the	DET
alces-443	88	20	authors	author	NOUN
alces-443	88	21	questioned	question	VERB
alces-443	88	22	the	the	DET
alces-443	88	23	long	long	ADJ
alces-443	88	24	-	-	PUNCT
alces-443	88	25	term	term	NOUN
alces-443	88	26	viability	viability	NOUN
alces-443	88	27	of	of	ADP
alces-443	88	28	those	those	DET
alces-443	88	29	moose	moose	NOUN
alces-443	88	30	populations	population	NOUN
alces-443	88	31	.	.	PUNCT
alces-443	89	1	nonetheless	nonetheless	ADV
alces-443	89	2	,	,	PUNCT
alces-443	89	3	levels	level	NOUN
alces-443	89	4	of	of	ADP
alces-443	89	5	diversity	diversity	NOUN
alces-443	89	6	in	in	ADP
alces-443	89	7	neutral	neutral	ADJ
alces-443	89	8	microsatellite	microsatellite	ADJ
alces-443	89	9	loci	locus	NOUN
alces-443	89	10	were	be	AUX
alces-443	89	11	not	not	PART
alces-443	89	12	indicative	indicative	ADJ
alces-443	89	13	of	of	ADP
alces-443	89	14	diversity	diversity	NOUN
alces-443	89	15	in	in	ADP
alces-443	89	16	functional	functional	ADJ
alces-443	89	17	loci	locus	NOUN
alces-443	89	18	in	in	ADP
alces-443	89	19	moose	moose	NOUN
alces-443	89	20	(	(	PUNCT
alces-443	89	21	wilson	wilson	PROPN
alces-443	89	22	et	et	PROPN
alces-443	89	23	al	al	PROPN
alces-443	89	24	.	.	PROPN
alces-443	89	25	2003	2003	NUM
alces-443	89	26	)	)	PUNCT
alces-443	89	27	.	.	PUNCT
alces-443	90	1	alces	alce	NOUN
alces-443	90	2	vol	vol	NOUN
alces-443	90	3	.	.	PROPN
alces-443	90	4	40	40	NUM
alces-443	90	5	,	,	PUNCT
alces-443	90	6	2004	2004	NUM
alces-443	90	7	moose	moose	NOUN
alces-443	90	8	genetics	genetic	NOUN
alces-443	90	9	hundertmark	hundertmark	NOUN
alces-443	90	10	and	and	CCONJ
alces-443	90	11	bowyer	bowyer	PROPN
alces-443	90	12	107	107	NUM
alces-443	90	13	the	the	DET
alces-443	90	14	future	future	ADJ
alces-443	90	15	microsatellites	microsatellite	NOUN
alces-443	90	16	have	have	AUX
alces-443	90	17	replaced	replace	VERB
alces-443	90	18	allozymes	allozyme	NOUN
alces-443	90	19	as	as	ADP
alces-443	90	20	the	the	DET
alces-443	90	21	most	most	ADV
alces-443	90	22	widely	widely	ADV
alces-443	90	23	used	use	VERB
alces-443	90	24	molecular	molecular	ADJ
alces-443	90	25	marker	marker	NOUN
alces-443	90	26	for	for	ADP
alces-443	90	27	assessing	assess	VERB
alces-443	90	28	nuclear	nuclear	ADJ
alces-443	90	29	genetic	genetic	ADJ
alces-443	90	30	diversity	diversity	NOUN
alces-443	90	31	,	,	PUNCT
alces-443	90	32	and	and	CCONJ
alces-443	90	33	will	will	AUX
alces-443	90	34	be	be	AUX
alces-443	90	35	the	the	DET
alces-443	90	36	choice	choice	NOUN
alces-443	90	37	of	of	ADP
alces-443	90	38	geneticists	geneticist	NOUN
alces-443	90	39	for	for	ADP
alces-443	90	40	the	the	DET
alces-443	90	41	foreseeable	foreseeable	ADJ
alces-443	90	42	future	future	NOUN
alces-443	90	43	(	(	PUNCT
alces-443	90	44	bruford	bruford	NOUN
alces-443	90	45	et	et	PROPN
alces-443	90	46	al	al	PROPN
alces-443	90	47	.	.	PROPN
alces-443	90	48	1996	1996	NUM
alces-443	90	49	)	)	PUNCT
alces-443	90	50	.	.	PUNCT
alces-443	91	1	there	there	PRON
alces-443	91	2	are	be	VERB
alces-443	91	3	problems	problem	NOUN
alces-443	91	4	,	,	PUNCT
alces-443	91	5	however	however	ADV
alces-443	91	6	,	,	PUNCT
alces-443	91	7	with	with	ADP
alces-443	91	8	analysis	analysis	NOUN
alces-443	91	9	of	of	ADP
alces-443	91	10	microsatellites	microsatellite	NOUN
alces-443	91	11	because	because	SCONJ
alces-443	91	12	the	the	DET
alces-443	91	13	ways	way	NOUN
alces-443	91	14	in	in	ADP
alces-443	91	15	which	which	PRON
alces-443	91	16	they	they	PRON
alces-443	91	17	mutate	mutate	VERB
alces-443	91	18	into	into	ADP
alces-443	91	19	new	new	ADJ
alces-443	91	20	forms	form	NOUN
alces-443	91	21	are	be	AUX
alces-443	91	22	not	not	PART
alces-443	91	23	entirely	entirely	ADV
alces-443	91	24	understood	understand	VERB
alces-443	91	25	(	(	PUNCT
alces-443	91	26	hancock	hancock	PROPN
alces-443	91	27	1999	1999	NUM
alces-443	91	28	)	)	PUNCT
alces-443	91	29	.	.	PUNCT
alces-443	92	1	in	in	ADP
alces-443	92	2	the	the	DET
alces-443	92	3	future	future	NOUN
alces-443	92	4	,	,	PUNCT
alces-443	92	5	a	a	DET
alces-443	92	6	new	new	ADJ
alces-443	92	7	type	type	NOUN
alces-443	92	8	of	of	ADP
alces-443	92	9	analysis	analysis	NOUN
alces-443	92	10	called	call	VERB
alces-443	92	11	single	single	ADJ
alces-443	92	12	nucleotide	nucleotide	ADJ
alces-443	92	13	polymorphism	polymorphism	NOUN
alces-443	92	14	(	(	PUNCT
alces-443	92	15	snp	snp	ADJ
alces-443	92	16	,	,	PUNCT
alces-443	92	17	pronounced	pronounce	VERB
alces-443	92	18	“	"	PUNCT
alces-443	92	19	snip	snip	NOUN
alces-443	92	20	”	"	PUNCT
alces-443	92	21	)	)	PUNCT
alces-443	92	22	may	may	AUX
alces-443	92	23	replace	replace	VERB
alces-443	92	24	microsatellites	microsatellite	NOUN
alces-443	92	25	for	for	ADP
alces-443	92	26	some	some	DET
alces-443	92	27	applications	application	NOUN
alces-443	92	28	(	(	PUNCT
alces-443	92	29	fries	fry	NOUN
alces-443	92	30	and	and	CCONJ
alces-443	92	31	durstewitz	durstewitz	NOUN
alces-443	92	32	2001	2001	NUM
alces-443	92	33	,	,	PUNCT
alces-443	92	34	brumfield	brumfield	PROPN
alces-443	92	35	et	et	PROPN
alces-443	92	36	al	al	PROPN
alces-443	92	37	.	.	PROPN
alces-443	92	38	2003	2003	NUM
alces-443	92	39	)	)	PUNCT
alces-443	92	40	.	.	PUNCT
alces-443	93	1	this	this	DET
alces-443	93	2	new	new	ADJ
alces-443	93	3	technology	technology	NOUN
alces-443	93	4	allows	allow	VERB
alces-443	93	5	a	a	DET
alces-443	93	6	single	single	ADJ
alces-443	93	7	nucleotide	nucleotide	NOUN
alces-443	93	8	site	site	NOUN
alces-443	93	9	to	to	PART
alces-443	93	10	be	be	AUX
alces-443	93	11	queried	query	VERB
alces-443	93	12	for	for	ADP
alces-443	93	13	presence	presence	NOUN
alces-443	93	14	of	of	ADP
alces-443	93	15	a	a	DET
alces-443	93	16	particular	particular	ADJ
alces-443	93	17	nucleotide	nucleotide	NOUN
alces-443	93	18	and	and	CCONJ
alces-443	93	19	presence	presence	NOUN
alces-443	93	20	or	or	CCONJ
alces-443	93	21	absence	absence	NOUN
alces-443	93	22	can	can	AUX
alces-443	93	23	be	be	AUX
alces-443	93	24	converted	convert	VERB
alces-443	93	25	to	to	ADP
alces-443	93	26	a	a	DET
alces-443	93	27	binary	binary	PROPN
alces-443	93	28	code	code	NOUN
alces-443	93	29	.	.	PUNCT
alces-443	94	1	current	current	ADJ
alces-443	94	2	technology	technology	NOUN
alces-443	94	3	(	(	PUNCT
alces-443	94	4	so	so	ADV
alces-443	94	5	-	-	PUNCT
alces-443	94	6	called	call	VERB
alces-443	94	7	“	"	PUNCT
alces-443	94	8	real	real	ADJ
alces-443	94	9	-	-	PUNCT
alces-443	94	10	time	time	NOUN
alces-443	94	11	pcr	pcr	NOUN
alces-443	94	12	”	"	PUNCT
alces-443	94	13	and	and	CCONJ
alces-443	94	14	“	"	PUNCT
alces-443	94	15	dna	dna	PROPN
alces-443	94	16	chips	chip	NOUN
alces-443	94	17	”	"	PUNCT
alces-443	94	18	)	)	PUNCT
alces-443	94	19	allows	allow	VERB
alces-443	94	20	for	for	ADP
alces-443	94	21	fast	fast	ADJ
alces-443	94	22	and	and	CCONJ
alces-443	94	23	accurate	accurate	ADJ
alces-443	94	24	examination	examination	NOUN
alces-443	94	25	of	of	ADP
alces-443	94	26	many	many	ADJ
alces-443	94	27	individuals	individual	NOUN
alces-443	94	28	and	and	CCONJ
alces-443	94	29	snps	snps	NOUN
alces-443	94	30	,	,	PUNCT
alces-443	94	31	but	but	CCONJ
alces-443	94	32	we	we	PRON
alces-443	94	33	must	must	AUX
alces-443	94	34	await	await	VERB
alces-443	94	35	the	the	DET
alces-443	94	36	development	development	NOUN
alces-443	94	37	and	and	CCONJ
alces-443	94	38	characterization	characterization	NOUN
alces-443	94	39	of	of	ADP
alces-443	94	40	marker	marker	NOUN
alces-443	94	41	loci	locus	NOUN
alces-443	94	42	before	before	SCONJ
alces-443	94	43	broad	broad	ADJ
alces-443	94	44	application	application	NOUN
alces-443	94	45	of	of	ADP
alces-443	94	46	this	this	DET
alces-443	94	47	new	new	ADJ
alces-443	94	48	family	family	NOUN
alces-443	94	49	of	of	ADP
alces-443	94	50	molecular	molecular	ADJ
alces-443	94	51	markers	marker	NOUN
alces-443	94	52	can	can	AUX
alces-443	94	53	be	be	AUX
alces-443	94	54	considered	consider	VERB
alces-443	94	55	.	.	PUNCT
alces-443	95	1	moose	moose	PROPN
alces-443	95	2	evolution	evolution	NOUN
alces-443	95	3	and	and	CCONJ
alces-443	95	4	phylogeography	phylogeography	NOUN
alces-443	95	5	origins	origin	NOUN
alces-443	95	6	of	of	ADP
alces-443	95	7	modern	modern	ADJ
alces-443	95	8	moose	moose	NOUN
alces-443	95	9	moose	moose	NOUN
alces-443	95	10	(	(	PUNCT
alces-443	95	11	alces	alce	NOUN
alces-443	95	12	alces	alce	NOUN
alces-443	95	13	)	)	PUNCT
alces-443	95	14	are	be	AUX
alces-443	95	15	a	a	DET
alces-443	95	16	young	young	ADJ
alces-443	95	17	species	specie	NOUN
alces-443	95	18	in	in	ADP
alces-443	95	19	the	the	DET
alces-443	95	20	evolutionary	evolutionary	ADJ
alces-443	95	21	scheme	scheme	NOUN
alces-443	95	22	of	of	ADP
alces-443	95	23	large	large	ADJ
alces-443	95	24	mammals	mammal	NOUN
alces-443	95	25	.	.	PUNCT
alces-443	96	1	the	the	DET
alces-443	96	2	genus	genus	PROPN
alces-443	96	3	alces	alce	NOUN
alces-443	96	4	first	first	ADV
alces-443	96	5	appears	appear	VERB
alces-443	96	6	in	in	ADP
alces-443	96	7	the	the	DET
alces-443	96	8	fossil	fossil	NOUN
alces-443	96	9	record	record	NOUN
alces-443	96	10	2	2	NUM
alces-443	96	11	million	million	NUM
alces-443	96	12	years	year	NOUN
alces-443	96	13	ago	ago	ADV
alces-443	96	14	(	(	PUNCT
alces-443	96	15	thouveny	thouveny	NOUN
alces-443	96	16	and	and	CCONJ
alces-443	96	17	bonifay	bonifay	NOUN
alces-443	96	18	1984	1984	NUM
alces-443	96	19	)	)	PUNCT
alces-443	96	20	and	and	CCONJ
alces-443	96	21	fossils	fossil	VERB
alces-443	96	22	attributable	attributable	ADJ
alces-443	96	23	to	to	ADP
alces-443	96	24	a.	a.	NOUN
alces-443	96	25	alces	alce	NOUN
alces-443	96	26	are	be	AUX
alces-443	96	27	first	first	ADV
alces-443	96	28	recorded	record	VERB
alces-443	96	29	approximately	approximately	ADV
alces-443	96	30	100,000	100,000	NUM
alces-443	96	31	years	year	NOUN
alces-443	96	32	ago	ago	ADV
alces-443	96	33	(	(	PUNCT
alces-443	96	34	lister	lister	PROPN
alces-443	96	35	1993	1993	NUM
alces-443	96	36	)	)	PUNCT
alces-443	96	37	.	.	PUNCT
alces-443	97	1	those	those	DET
alces-443	97	2	dates	date	NOUN
alces-443	97	3	are	be	AUX
alces-443	97	4	very	very	ADV
alces-443	97	5	recent	recent	ADJ
alces-443	97	6	considering	consider	VERB
alces-443	97	7	that	that	SCONJ
alces-443	97	8	the	the	DET
alces-443	97	9	subfamily	subfamily	ADV
alces-443	97	10	odocoileinae	odocoileinae	ADJ
alces-443	97	11	,	,	PUNCT
alces-443	97	12	to	to	PART
alces-443	97	13	which	which	PRON
alces-443	97	14	moose	moose	NOUN
alces-443	97	15	belong	belong	VERB
alces-443	97	16	,	,	PUNCT
alces-443	97	17	diverged	diverge	VERB
alces-443	97	18	from	from	ADP
alces-443	97	19	other	other	ADJ
alces-443	97	20	deer	deer	NOUN
alces-443	97	21	lineages	lineage	NOUN
alces-443	97	22	9	9	NUM
alces-443	97	23	-	-	SYM
alces-443	97	24	12	12	NUM
alces-443	97	25	million	million	NUM
alces-443	97	26	years	year	NOUN
alces-443	97	27	ago	ago	ADV
alces-443	97	28	(	(	PUNCT
alces-443	97	29	miyamoto	miyamoto	NOUN
alces-443	97	30	et	et	PROPN
alces-443	97	31	al	al	PROPN
alces-443	97	32	.	.	PROPN
alces-443	97	33	1990	1990	NUM
alces-443	97	34	)	)	PUNCT
alces-443	97	35	.	.	PUNCT
alces-443	98	1	paleontological	paleontological	ADJ
alces-443	98	2	evidence	evidence	NOUN
alces-443	98	3	indicated	indicate	VERB
alces-443	98	4	europe	europe	PROPN
alces-443	98	5	as	as	ADP
alces-443	98	6	the	the	DET
alces-443	98	7	place	place	NOUN
alces-443	98	8	of	of	ADP
alces-443	98	9	origin	origin	NOUN
alces-443	98	10	of	of	ADP
alces-443	98	11	the	the	DET
alces-443	98	12	genus	genus	ADJ
alces-443	98	13	alces	alce	NOUN
alces-443	98	14	(	(	PUNCT
alces-443	98	15	lister	lister	PROPN
alces-443	98	16	1993	1993	NUM
alces-443	98	17	)	)	PUNCT
alces-443	98	18	.	.	PUNCT
alces-443	99	1	the	the	DET
alces-443	99	2	genus	genus	NOUN
alces-443	99	3	never	never	ADV
alces-443	99	4	was	be	AUX
alces-443	99	5	diverse	diverse	ADJ
alces-443	99	6	,	,	PUNCT
alces-443	99	7	with	with	ADP
alces-443	99	8	only	only	ADV
alces-443	99	9	one	one	NUM
alces-443	99	10	species	specie	NOUN
alces-443	99	11	present	present	ADJ
alces-443	99	12	in	in	ADP
alces-443	99	13	the	the	DET
alces-443	99	14	fossil	fossil	NOUN
alces-443	99	15	record	record	NOUN
alces-443	99	16	at	at	ADP
alces-443	99	17	any	any	DET
alces-443	99	18	particular	particular	ADJ
alces-443	99	19	time	time	NOUN
alces-443	99	20	.	.	PUNCT
alces-443	100	1	yet	yet	ADV
alces-443	100	2	,	,	PUNCT
alces-443	100	3	the	the	DET
alces-443	100	4	species	specie	NOUN
alces-443	100	5	assumed	assume	VERB
alces-443	100	6	to	to	PART
alces-443	100	7	be	be	AUX
alces-443	100	8	the	the	DET
alces-443	100	9	precursor	precursor	NOUN
alces-443	100	10	to	to	ADP
alces-443	100	11	a.	a.	NOUN
alces-443	100	12	alces	alce	NOUN
alces-443	100	13	,	,	PUNCT
alces-443	100	14	the	the	DET
alces-443	100	15	broad	broad	ADV
alces-443	100	16	-	-	PUNCT
alces-443	100	17	fronted	front	VERB
alces-443	100	18	moose	moose	NOUN
alces-443	100	19	(	(	PUNCT
alces-443	100	20	a.	a.	NOUN
alces-443	100	21	latifrons	latifron	NOUN
alces-443	100	22	)	)	PUNCT
alces-443	100	23	was	be	AUX
alces-443	100	24	distributed	distribute	VERB
alces-443	100	25	across	across	ADP
alces-443	100	26	eurasia	eurasia	PROPN
alces-443	100	27	and	and	CCONJ
alces-443	100	28	into	into	ADP
alces-443	100	29	northwestern	northwestern	PROPN
alces-443	100	30	north	north	PROPN
alces-443	100	31	america	america	PROPN
alces-443	100	32	for	for	ADP
alces-443	100	33	a	a	DET
alces-443	100	34	time	time	NOUN
alces-443	100	35	before	before	ADP
alces-443	100	36	becoming	become	VERB
alces-443	100	37	extinct	extinct	ADJ
alces-443	100	38	in	in	ADP
alces-443	100	39	beringia	beringia	NOUN
alces-443	100	40	at	at	ADP
alces-443	100	41	the	the	DET
alces-443	100	42	end	end	NOUN
alces-443	100	43	of	of	ADP
alces-443	100	44	the	the	DET
alces-443	100	45	pleistocene	pleistocene	NOUN
alces-443	100	46	(	(	PUNCT
alces-443	100	47	guthrie	guthrie	PROPN
alces-443	100	48	1995	1995	NUM
alces-443	100	49	)	)	PUNCT
alces-443	100	50	.	.	PUNCT
alces-443	101	1	thus	thus	ADV
alces-443	101	2	,	,	PUNCT
alces-443	101	3	the	the	DET
alces-443	101	4	widespread	widespread	ADJ
alces-443	101	5	distribution	distribution	NOUN
alces-443	101	6	of	of	ADP
alces-443	101	7	modern	modern	ADJ
alces-443	101	8	moose	moose	NOUN
alces-443	101	9	and	and	CCONJ
alces-443	101	10	its	its	PRON
alces-443	101	11	immediate	immediate	ADJ
alces-443	101	12	ancestor	ancestor	NOUN
alces-443	101	13	indicate	indicate	VERB
alces-443	101	14	a	a	DET
alces-443	101	15	degree	degree	NOUN
alces-443	101	16	of	of	ADP
alces-443	101	17	evolutionary	evolutionary	ADJ
alces-443	101	18	success	success	NOUN
alces-443	101	19	despite	despite	SCONJ
alces-443	101	20	a	a	DET
alces-443	101	21	paucity	paucity	NOUN
alces-443	101	22	of	of	ADP
alces-443	101	23	species	specie	NOUN
alces-443	101	24	diversity	diversity	NOUN
alces-443	101	25	.	.	PUNCT
alces-443	102	1	up	up	ADP
alces-443	102	2	to	to	PART
alces-443	102	3	8	8	NUM
alces-443	102	4	subspecies	subspecie	NOUN
alces-443	102	5	of	of	ADP
alces-443	102	6	moose	moose	NOUN
alces-443	102	7	are	be	AUX
alces-443	102	8	recognized	recognize	VERB
alces-443	102	9	worldwide	worldwide	ADV
alces-443	102	10	(	(	PUNCT
alces-443	102	11	fig	fig	NOUN
alces-443	102	12	.	.	NOUN
alces-443	102	13	1	1	NUM
alces-443	102	14	)	)	PUNCT
alces-443	102	15	;	;	PUNCT
alces-443	102	16	4	4	NUM
alces-443	102	17	in	in	ADP
alces-443	102	18	eurasia	eurasia	PROPN
alces-443	102	19	and	and	CCONJ
alces-443	102	20	4	4	NUM
alces-443	102	21	in	in	ADP
alces-443	102	22	north	north	PROPN
alces-443	102	23	america	america	PROPN
alces-443	102	24	(	(	PUNCT
alces-443	102	25	peterson	peterson	PROPN
alces-443	102	26	1955	1955	NUM
alces-443	102	27	)	)	PUNCT
alces-443	102	28	.	.	PUNCT
alces-443	103	1	that	that	DET
alces-443	103	2	number	number	NOUN
alces-443	103	3	is	be	AUX
alces-443	103	4	open	open	ADJ
alces-443	103	5	to	to	ADP
alces-443	103	6	question	question	NOUN
alces-443	103	7	,	,	PUNCT
alces-443	103	8	however	however	ADV
alces-443	103	9	.	.	PUNCT
alces-443	104	1	geist	geist	PROPN
alces-443	104	2	(	(	PUNCT
alces-443	104	3	1987a	1987a	NUM
alces-443	104	4	,	,	PUNCT
alces-443	104	5	1998	1998	NUM
alces-443	104	6	)	)	PUNCT
alces-443	104	7	contends	contend	VERB
alces-443	104	8	that	that	SCONJ
alces-443	104	9	there	there	PRON
alces-443	104	10	are	be	VERB
alces-443	104	11	2	2	NUM
alces-443	104	12	predominant	predominant	ADJ
alces-443	104	13	types	type	NOUN
alces-443	104	14	of	of	ADP
alces-443	104	15	moose	moose	NOUN
alces-443	104	16	in	in	ADP
alces-443	104	17	the	the	DET
alces-443	104	18	world	world	NOUN
alces-443	104	19	:	:	PUNCT
alces-443	104	20	american	american	PROPN
alces-443	104	21	and	and	CCONJ
alces-443	104	22	european	european	ADJ
alces-443	104	23	,	,	PUNCT
alces-443	104	24	following	follow	VERB
alces-443	104	25	the	the	DET
alces-443	104	26	convention	convention	NOUN
alces-443	104	27	of	of	ADP
alces-443	104	28	flerov	flerov	PROPN
alces-443	104	29	(	(	PUNCT
alces-443	104	30	1952	1952	NUM
alces-443	104	31	)	)	PUNCT
alces-443	104	32	.	.	PUNCT
alces-443	105	1	to	to	ADP
alces-443	105	2	the	the	DET
alces-443	105	3	former	former	ADJ
alces-443	105	4	type	type	NOUN
alces-443	105	5	he	he	PRON
alces-443	105	6	assigns	assign	VERB
alces-443	105	7	all	all	DET
alces-443	105	8	north	north	ADJ
alces-443	105	9	american	american	ADJ
alces-443	105	10	moose	moose	NOUN
alces-443	105	11	as	as	ADV
alces-443	105	12	well	well	ADV
alces-443	105	13	as	as	ADP
alces-443	105	14	eastern	eastern	ADJ
alces-443	105	15	asian	asian	ADJ
alces-443	105	16	subspecies	subspecie	NOUN
alces-443	105	17	a.	a.	NOUN
alces-443	105	18	a.	a.	NOUN
alces-443	105	19	burturlini	burturlini	PROPN
alces-443	105	20	and	and	CCONJ
alces-443	105	21	a.	a.	NOUN
alces-443	105	22	a.	a.	NOUN
alces-443	105	23	pfizenmayeri	pfizenmayeri	PROPN
alces-443	105	24	.	.	PUNCT
alces-443	106	1	he	he	PRON
alces-443	106	2	based	base	VERB
alces-443	106	3	his	his	PRON
alces-443	106	4	opinion	opinion	NOUN
alces-443	106	5	primarily	primarily	ADV
alces-443	106	6	on	on	ADP
alces-443	106	7	morphology	morphology	NOUN
alces-443	106	8	and	and	CCONJ
alces-443	106	9	noted	note	VERB
alces-443	106	10	similar	similar	ADJ
alces-443	106	11	geographic	geographic	ADJ
alces-443	106	12	divisions	division	NOUN
alces-443	106	13	in	in	ADP
alces-443	106	14	taxonomy	taxonomy	NOUN
alces-443	106	15	among	among	ADP
alces-443	106	16	reindeer	reindeer	NOUN
alces-443	106	17	and	and	CCONJ
alces-443	106	18	caribou	caribou	PROPN
alces-443	106	19	(	(	PUNCT
alces-443	106	20	rangifer	rangifer	VERB
alces-443	106	21	tarandus	tarandus	NOUN
alces-443	106	22	)	)	PUNCT
alces-443	106	23	and	and	CCONJ
alces-443	106	24	red	red	ADJ
alces-443	106	25	deer	deer	NOUN
alces-443	106	26	and	and	CCONJ
alces-443	106	27	north	north	ADJ
alces-443	106	28	american	american	PROPN
alces-443	106	29	elk	elk	PROPN
alces-443	106	30	(	(	PUNCT
alces-443	106	31	cervus	cervus	NOUN
alces-443	106	32	elaphus	elaphu	NOUN
alces-443	106	33	;	;	PUNCT
alces-443	106	34	geist	geist	NOUN
alces-443	106	35	1998	1998	NUM
alces-443	106	36	)	)	PUNCT
alces-443	106	37	.	.	PUNCT
alces-443	107	1	he	he	PRON
alces-443	107	2	further	far	ADV
alces-443	107	3	contended	contend	VERB
alces-443	107	4	that	that	SCONJ
alces-443	107	5	those	those	DET
alces-443	107	6	morphological	morphological	ADJ
alces-443	107	7	types	type	NOUN
alces-443	107	8	should	should	AUX
alces-443	107	9	correspond	correspond	VERB
alces-443	107	10	to	to	ADP
alces-443	107	11	subspecies	subspecie	NOUN
alces-443	107	12	designations	designation	NOUN
alces-443	107	13	.	.	PUNCT
alces-443	108	1	therefore	therefore	ADV
alces-443	108	2	,	,	PUNCT
alces-443	108	3	geist	geist	PROPN
alces-443	108	4	(	(	PUNCT
alces-443	108	5	1998	1998	NUM
alces-443	108	6	)	)	PUNCT
alces-443	108	7	recognized	recognize	VERB
alces-443	108	8	a.	a.	NOUN
alces-443	108	9	a.	a.	NOUN
alces-443	108	10	alces	alce	NOUN
alces-443	108	11	of	of	ADP
alces-443	108	12	europe	europe	PROPN
alces-443	108	13	and	and	CCONJ
alces-443	108	14	a.	a.	NOUN
alces-443	108	15	a.	a.	NOUN
alces-443	108	16	americana	americana	PROPN
alces-443	108	17	in	in	ADP
alces-443	108	18	eastern	eastern	PROPN
alces-443	108	19	asia	asia	PROPN
alces-443	108	20	and	and	CCONJ
alces-443	108	21	north	north	PROPN
alces-443	108	22	america	america	PROPN
alces-443	108	23	.	.	PUNCT
alces-443	109	1	he	he	PRON
alces-443	109	2	also	also	ADV
alces-443	109	3	suggested	suggest	VERB
alces-443	109	4	that	that	SCONJ
alces-443	109	5	a.	a.	NOUN
alces-443	109	6	a.	a.	NOUN
alces-443	109	7	americanus	americanus	PROPN
alces-443	109	8	has	have	AUX
alces-443	109	9	precedence	precedence	NOUN
alces-443	109	10	under	under	ADP
alces-443	109	11	nomenclatural	nomenclatural	ADJ
alces-443	109	12	conventions	convention	NOUN
alces-443	109	13	as	as	ADP
alces-443	109	14	the	the	DET
alces-443	109	15	proper	proper	ADJ
alces-443	109	16	name	name	NOUN
alces-443	109	17	for	for	ADP
alces-443	109	18	the	the	DET
alces-443	109	19	east	east	ADJ
alces-443	109	20	asian	asian	ADJ
alces-443	109	21	-	-	PUNCT
alces-443	109	22	north	north	NOUN
alces-443	109	23	american	american	ADJ
alces-443	109	24	subspecies	subspecie	NOUN
alces-443	109	25	.	.	PUNCT
alces-443	110	1	he	he	PRON
alces-443	110	2	referred	refer	VERB
alces-443	110	3	to	to	ADP
alces-443	110	4	a.	a.	PROPN
alces-443	110	5	a.	a.	NOUN
alces-443	110	6	cameloides	cameloide	NOUN
alces-443	110	7	in	in	ADP
alces-443	110	8	northern	northern	ADJ
alces-443	110	9	china	china	PROPN
alces-443	110	10	,	,	PUNCT
alces-443	110	11	mongolia	mongolia	PROPN
alces-443	110	12	,	,	PUNCT
alces-443	110	13	and	and	CCONJ
alces-443	110	14	southeastern	southeastern	ADJ
alces-443	110	15	russia	russia	PROPN
alces-443	110	16	as	as	ADP
alces-443	110	17	part	part	NOUN
alces-443	110	18	of	of	ADP
alces-443	110	19	a	a	DET
alces-443	110	20	primitive	primitive	ADJ
alces-443	110	21	fauna	fauna	NOUN
alces-443	110	22	native	native	ADJ
alces-443	110	23	to	to	ADP
alces-443	110	24	that	that	DET
alces-443	110	25	region	region	NOUN
alces-443	110	26	and	and	CCONJ
alces-443	110	27	recognized	recognize	VERB
alces-443	110	28	that	that	SCONJ
alces-443	110	29	subspecies	subspecie	NOUN
alces-443	110	30	as	as	ADP
alces-443	110	31	a	a	DET
alces-443	110	32	valid	valid	ADJ
alces-443	110	33	taxon	taxon	NOUN
alces-443	110	34	although	although	SCONJ
alces-443	110	35	he	he	PRON
alces-443	110	36	also	also	ADV
alces-443	110	37	refers	refer	VERB
alces-443	110	38	to	to	ADP
alces-443	110	39	it	it	PRON
alces-443	110	40	as	as	ADP
alces-443	110	41	an	an	DET
alces-443	110	42	americantype	americantype	PROPN
alces-443	110	43	moose	moose	NOUN
alces-443	110	44	(	(	PUNCT
alces-443	110	45	geist	geist	PROPN
alces-443	110	46	1998:230	1998:230	NUM
alces-443	110	47	)	)	PUNCT
alces-443	110	48	.	.	PUNCT
alces-443	111	1	the	the	DET
alces-443	111	2	2	2	NUM
alces-443	111	3	-	-	PUNCT
alces-443	111	4	types	type	NOUN
alces-443	111	5	hypothesis	hypothesis	NOUN
alces-443	111	6	is	be	AUX
alces-443	111	7	supported	support	VERB
alces-443	111	8	to	to	ADP
alces-443	111	9	some	some	DET
alces-443	111	10	degree	degree	NOUN
alces-443	111	11	by	by	ADP
alces-443	111	12	karyotype	karyotype	NOUN
alces-443	111	13	(	(	PUNCT
alces-443	111	14	boeskorov	boeskorov	PROPN
alces-443	111	15	1996	1996	NUM
alces-443	111	16	,	,	PUNCT
alces-443	111	17	1997	1997	NUM
alces-443	111	18	)	)	PUNCT
alces-443	111	19	and	and	CCONJ
alces-443	111	20	some	some	DET
alces-443	111	21	data	datum	NOUN
alces-443	111	22	on	on	ADP
alces-443	111	23	mtdna	mtdna	PROPN
alces-443	111	24	(	(	PUNCT
alces-443	111	25	mikko	mikko	PROPN
alces-443	111	26	hundertmark	hundertmark	PROPN
alces-443	111	27	and	and	CCONJ
alces-443	111	28	bowyer	bowyer	PROPN
alces-443	111	29	moose	moose	PROPN
alces-443	111	30	genetics	genetics	PROPN
alces-443	111	31	alces	alce	NOUN
alces-443	111	32	vol	vol	NOUN
alces-443	111	33	.	.	PROPN
alces-443	111	34	40	40	NUM
alces-443	111	35	,	,	PUNCT
alces-443	111	36	2004	2004	NUM
alces-443	111	37	108	108	NUM
alces-443	111	38	fig	fig	NOUN
alces-443	111	39	.	.	PUNCT
alces-443	112	1	1	1	X
alces-443	112	2	.	.	NUM
alces-443	112	3	approximate	approximate	ADJ
alces-443	112	4	ranges	range	NOUN
alces-443	112	5	of	of	ADP
alces-443	112	6	8	8	NUM
alces-443	112	7	subspecies	subspecie	NOUN
alces-443	112	8	of	of	ADP
alces-443	112	9	moose	moose	NOUN
alces-443	112	10	worldwide	worldwide	ADV
alces-443	112	11	.	.	PUNCT
alces-443	113	1	a.	a.	NOUN
alces-443	113	2	a.	a.	NOUN
alces-443	113	3	a.	a.	NOUN
alces-443	113	4	=	=	PUNCT
alces-443	113	5	a.	a.	NOUN
alces-443	113	6	a.	a.	NOUN
alces-443	113	7	alces	alces	PROPN
alces-443	113	8	,	,	PUNCT
alces-443	113	9	a.	a.	NOUN
alces-443	113	10	a.	a.	NOUN
alces-443	113	11	p.	p.	PROPN
alces-443	113	12	=	=	PUNCT
alces-443	113	13	a.	a.	NOUN
alces-443	113	14	a.	a.	NOUN
alces-443	113	15	pfizenmayeri	pfizenmayeri	PROPN
alces-443	113	16	,	,	PUNCT
alces-443	113	17	a.	a.	NOUN
alces-443	113	18	a.	a.	PROPN
alces-443	113	19	c.	c.	PROPN
alces-443	113	20	=	=	PUNCT
alces-443	113	21	a.	a.	NOUN
alces-443	113	22	a.	a.	NOUN
alces-443	113	23	cameloides	cameloide	NOUN
alces-443	113	24	,	,	PUNCT
alces-443	113	25	a.	a.	NOUN
alces-443	113	26	a.	a.	PROPN
alces-443	113	27	b.	b.	PROPN
alces-443	114	1	=	=	PUNCT
alces-443	114	2	a.	a.	PROPN
alces-443	114	3	a.	a.	PROPN
alces-443	114	4	burturlini	burturlini	PROPN
alces-443	114	5	,	,	PUNCT
alces-443	114	6	a.	a.	NOUN
alces-443	114	7	a.	a.	NOUN
alces-443	114	8	g.	g.	PROPN
alces-443	115	1	=	=	PUNCT
alces-443	116	1	a.	a.	NOUN
alces-443	116	2	a.	a.	NOUN
alces-443	116	3	gigas	gigas	PROPN
alces-443	116	4	,	,	PUNCT
alces-443	116	5	a.	a.	NOUN
alces-443	116	6	a.	a.	NOUN
alces-443	116	7	an	an	PROPN
alces-443	116	8	.	.	PUNCT
alces-443	117	1	=	=	PUNCT
alces-443	117	2	a.	a.	NOUN
alces-443	117	3	a.	a.	NOUN
alces-443	117	4	andersoni	andersoni	PROPN
alces-443	117	5	,	,	PUNCT
alces-443	117	6	a.	a.	PROPN
alces-443	117	7	a.	a.	PROPN
alces-443	117	8	s.	s.	PROPN
alces-443	118	1	=	=	PUNCT
alces-443	118	2	a.	a.	PROPN
alces-443	118	3	a.	a.	PROPN
alces-443	118	4	shirasi	shirasi	PROPN
alces-443	118	5	,	,	PUNCT
alces-443	118	6	a.	a.	PROPN
alces-443	118	7	a.	a.	NOUN
alces-443	118	8	am	am	PROPN
alces-443	118	9	.	.	PUNCT
alces-443	119	1	=	=	PUNCT
alces-443	119	2	a.	a.	NOUN
alces-443	119	3	a.	a.	NOUN
alces-443	119	4	americana	americana	PROPN
alces-443	119	5	,	,	PUNCT
alces-443	119	6	*	*	PUNCT
alces-443	119	7	=	=	PRON
alces-443	119	8	introduced	introduce	VERB
alces-443	119	9	population	population	NOUN
alces-443	119	10	in	in	ADP
alces-443	119	11	newfoundland	newfoundland	PROPN
alces-443	119	12	.	.	PUNCT
alces-443	120	1	and	and	CCONJ
alces-443	120	2	andersson	andersson	PROPN
alces-443	120	3	1995	1995	NUM
alces-443	120	4	)	)	PUNCT
alces-443	120	5	.	.	PUNCT
alces-443	121	1	most	most	ADJ
alces-443	121	2	eurasian	eurasian	ADJ
alces-443	121	3	moose	moose	NOUN
alces-443	121	4	have	have	VERB
alces-443	121	5	a	a	DET
alces-443	121	6	karyotype	karyotype	NOUN
alces-443	121	7	of	of	ADP
alces-443	121	8	2n	2n	NUM
alces-443	121	9	=	=	SYM
alces-443	121	10	68	68	NUM
alces-443	121	11	,	,	PUNCT
alces-443	121	12	whereas	whereas	SCONJ
alces-443	121	13	north	north	ADJ
alces-443	121	14	american	american	PROPN
alces-443	121	15	moose	moose	NOUN
alces-443	121	16	have	have	VERB
alces-443	121	17	2n	2n	NUM
alces-443	121	18	=	=	SYM
alces-443	121	19	70	70	NUM
alces-443	121	20	.	.	PUNCT
alces-443	122	1	that	that	DET
alces-443	122	2	difference	difference	NOUN
alces-443	122	3	derives	derive	VERB
alces-443	122	4	from	from	ADP
alces-443	122	5	a	a	DET
alces-443	122	6	robertsonian	robertsonian	ADJ
alces-443	122	7	translocation	translocation	NOUN
alces-443	122	8	of	of	ADP
alces-443	122	9	2	2	NUM
alces-443	122	10	acrocentric	acrocentric	ADJ
alces-443	122	11	chromosomes	chromosome	NOUN
alces-443	122	12	into	into	ADP
alces-443	122	13	a	a	DET
alces-443	122	14	single	single	ADJ
alces-443	122	15	metacentric	metacentric	ADJ
alces-443	122	16	chromosome	chromosome	NOUN
alces-443	122	17	or	or	CCONJ
alces-443	122	18	vice	vice	NOUN
alces-443	122	19	versa	versa	ADV
alces-443	122	20	.	.	PUNCT
alces-443	123	1	although	although	SCONJ
alces-443	123	2	that	that	DET
alces-443	123	3	chromosomal	chromosomal	ADJ
alces-443	123	4	polymorphism	polymorphism	NOUN
alces-443	123	5	originally	originally	ADV
alces-443	123	6	was	be	AUX
alces-443	123	7	thought	think	VERB
alces-443	123	8	to	to	PART
alces-443	123	9	separate	separate	VERB
alces-443	123	10	eurasian	eurasian	ADJ
alces-443	123	11	and	and	CCONJ
alces-443	123	12	north	north	ADJ
alces-443	123	13	american	american	ADJ
alces-443	123	14	moose	moose	NOUN
alces-443	123	15	(	(	PUNCT
alces-443	123	16	groves	grove	NOUN
alces-443	123	17	and	and	CCONJ
alces-443	123	18	grubb	grubb	PROPN
alces-443	123	19	1987	1987	NUM
alces-443	123	20	)	)	PUNCT
alces-443	123	21	,	,	PUNCT
alces-443	123	22	the	the	DET
alces-443	123	23	2n	2n	NUM
alces-443	123	24	=	=	SYM
alces-443	123	25	70	70	NUM
alces-443	123	26	form	form	NOUN
alces-443	123	27	was	be	AUX
alces-443	123	28	recently	recently	ADV
alces-443	123	29	discovered	discover	VERB
alces-443	123	30	in	in	ADP
alces-443	123	31	eastern	eastern	PROPN
alces-443	123	32	asia	asia	PROPN
alces-443	123	33	(	(	PUNCT
alces-443	123	34	boeskorov	boeskorov	PROPN
alces-443	123	35	1996	1996	NUM
alces-443	123	36	,	,	PUNCT
alces-443	123	37	1997	1997	NUM
alces-443	123	38	)	)	PUNCT
alces-443	123	39	.	.	PUNCT
alces-443	124	1	similarly	similarly	ADV
alces-443	124	2	,	,	PUNCT
alces-443	124	3	a	a	DET
alces-443	124	4	length	length	NOUN
alces-443	124	5	mutation	mutation	NOUN
alces-443	124	6	(	(	PUNCT
alces-443	124	7	insertion	insertion	NOUN
alces-443	124	8	-	-	PUNCT
alces-443	124	9	deletion	deletion	NOUN
alces-443	124	10	,	,	PUNCT
alces-443	124	11	or	or	CCONJ
alces-443	124	12	indel	indel	NOUN
alces-443	124	13	)	)	PUNCT
alces-443	124	14	within	within	ADP
alces-443	124	15	the	the	DET
alces-443	124	16	control	control	NOUN
alces-443	124	17	region	region	NOUN
alces-443	124	18	of	of	ADP
alces-443	124	19	mtdna	mtdna	NOUN
alces-443	124	20	originally	originally	ADV
alces-443	124	21	was	be	AUX
alces-443	124	22	described	describe	VERB
alces-443	124	23	as	as	ADP
alces-443	124	24	discriminating	discriminate	VERB
alces-443	124	25	between	between	ADP
alces-443	124	26	north	north	PROPN
alces-443	124	27	american	american	ADJ
alces-443	124	28	and	and	CCONJ
alces-443	124	29	european	european	ADJ
alces-443	124	30	moose	moose	NOUN
alces-443	124	31	(	(	PUNCT
alces-443	124	32	mikko	mikko	PROPN
alces-443	124	33	and	and	CCONJ
alces-443	124	34	andersson	andersson	PROPN
alces-443	124	35	1995	1995	NUM
alces-443	124	36	)	)	PUNCT
alces-443	124	37	,	,	PUNCT
alces-443	124	38	but	but	CCONJ
alces-443	124	39	subsequent	subsequent	ADJ
alces-443	124	40	investigations	investigation	NOUN
alces-443	124	41	documented	document	VERB
alces-443	124	42	that	that	SCONJ
alces-443	124	43	indel	indel	NOUN
alces-443	124	44	in	in	ADP
alces-443	124	45	moose	moose	NOUN
alces-443	124	46	from	from	ADP
alces-443	124	47	eastern	eastern	PROPN
alces-443	124	48	asia	asia	PROPN
alces-443	124	49	(	(	PUNCT
alces-443	124	50	hundertmark	hundertmark	NOUN
alces-443	124	51	et	et	PROPN
alces-443	124	52	al	al	PROPN
alces-443	124	53	.	.	PROPN
alces-443	124	54	2002b	2002b	NUM
alces-443	124	55	,	,	PUNCT
alces-443	124	56	udina	udina	NOUN
alces-443	124	57	et	et	PROPN
alces-443	124	58	al	al	PROPN
alces-443	124	59	.	.	PROPN
alces-443	124	60	2002	2002	NUM
alces-443	124	61	)	)	PUNCT
alces-443	124	62	.	.	PUNCT
alces-443	125	1	although	although	SCONJ
alces-443	125	2	the	the	DET
alces-443	125	3	precise	precise	ADJ
alces-443	125	4	geographic	geographic	ADJ
alces-443	125	5	distributions	distribution	NOUN
alces-443	125	6	of	of	ADP
alces-443	125	7	those	those	DET
alces-443	125	8	polymorphisms	polymorphism	NOUN
alces-443	125	9	in	in	ADP
alces-443	125	10	karyotype	karyotype	NOUN
alces-443	125	11	and	and	CCONJ
alces-443	125	12	mtdna	mtdna	NOUN
alces-443	125	13	alces	alce	NOUN
alces-443	125	14	vol	vol	NOUN
alces-443	125	15	.	.	PROPN
alces-443	125	16	40	40	NUM
alces-443	125	17	,	,	PUNCT
alces-443	125	18	2004	2004	NUM
alces-443	125	19	moose	moose	NOUN
alces-443	125	20	genetics	genetic	NOUN
alces-443	125	21	hundertmark	hundertmark	NOUN
alces-443	125	22	and	and	CCONJ
alces-443	125	23	bowyer	bowyer	PROPN
alces-443	125	24	109	109	NUM
alces-443	125	25	length	length	NOUN
alces-443	125	26	are	be	AUX
alces-443	125	27	not	not	PART
alces-443	125	28	well	well	ADV
alces-443	125	29	described	describe	VERB
alces-443	125	30	,	,	PUNCT
alces-443	125	31	they	they	PRON
alces-443	125	32	seem	seem	VERB
alces-443	125	33	to	to	PART
alces-443	125	34	correspond	correspond	VERB
alces-443	125	35	geographically	geographically	ADV
alces-443	125	36	with	with	ADP
alces-443	125	37	a	a	DET
alces-443	125	38	zone	zone	NOUN
alces-443	125	39	of	of	ADP
alces-443	125	40	intergradation	intergradation	NOUN
alces-443	125	41	in	in	ADP
alces-443	125	42	east	east	ADJ
alces-443	125	43	-	-	PUNCT
alces-443	125	44	central	central	PROPN
alces-443	125	45	asia	asia	PROPN
alces-443	125	46	,	,	PUNCT
alces-443	125	47	similar	similar	ADJ
alces-443	125	48	to	to	ADP
alces-443	125	49	that	that	PRON
alces-443	125	50	proposed	propose	VERB
alces-443	125	51	for	for	ADP
alces-443	125	52	american	american	ADJ
alces-443	125	53	and	and	CCONJ
alces-443	125	54	eurasian	eurasian	ADJ
alces-443	125	55	types	type	NOUN
alces-443	125	56	of	of	ADP
alces-443	125	57	moose	moose	NOUN
alces-443	125	58	(	(	PUNCT
alces-443	125	59	flerov	flerov	NOUN
alces-443	125	60	1952	1952	NUM
alces-443	125	61	,	,	PUNCT
alces-443	125	62	geist	geist	NOUN
alces-443	125	63	1998	1998	NUM
alces-443	125	64	)	)	PUNCT
alces-443	125	65	.	.	PUNCT
alces-443	126	1	more	more	ADJ
alces-443	126	2	work	work	NOUN
alces-443	126	3	is	be	AUX
alces-443	126	4	needed	need	VERB
alces-443	126	5	to	to	PART
alces-443	126	6	determine	determine	VERB
alces-443	126	7	the	the	DET
alces-443	126	8	extent	extent	NOUN
alces-443	126	9	of	of	ADP
alces-443	126	10	that	that	DET
alces-443	126	11	geographic	geographic	ADJ
alces-443	126	12	correspondence	correspondence	NOUN
alces-443	126	13	and	and	CCONJ
alces-443	126	14	to	to	PART
alces-443	126	15	determine	determine	VERB
alces-443	126	16	if	if	SCONJ
alces-443	126	17	it	it	PRON
alces-443	126	18	coincides	coincide	VERB
alces-443	126	19	with	with	ADP
alces-443	126	20	subspecies	subspecie	NOUN
alces-443	126	21	boundaries	boundary	NOUN
alces-443	126	22	.	.	PUNCT
alces-443	127	1	moreover	moreover	ADV
alces-443	127	2	,	,	PUNCT
alces-443	127	3	the	the	DET
alces-443	127	4	question	question	NOUN
alces-443	127	5	of	of	ADP
alces-443	127	6	reproductive	reproductive	ADJ
alces-443	127	7	viability	viability	NOUN
alces-443	127	8	of	of	ADP
alces-443	127	9	the	the	DET
alces-443	127	10	two	two	NUM
alces-443	127	11	chromosomal	chromosomal	ADJ
alces-443	127	12	races	race	NOUN
alces-443	127	13	must	must	AUX
alces-443	127	14	be	be	AUX
alces-443	127	15	addressed	address	VERB
alces-443	127	16	.	.	PUNCT
alces-443	128	1	indeed	indeed	ADV
alces-443	128	2	,	,	PUNCT
alces-443	128	3	boeskorov	boeskorov	X
alces-443	128	4	(	(	PUNCT
alces-443	128	5	1997	1997	NUM
alces-443	128	6	)	)	PUNCT
alces-443	128	7	has	have	AUX
alces-443	128	8	proposed	propose	VERB
alces-443	128	9	that	that	SCONJ
alces-443	128	10	the	the	DET
alces-443	128	11	chromosomal	chromosomal	ADJ
alces-443	128	12	races	race	NOUN
alces-443	128	13	are	be	AUX
alces-443	128	14	different	different	ADJ
alces-443	128	15	species	specie	NOUN
alces-443	128	16	and	and	CCONJ
alces-443	128	17	groves	grove	NOUN
alces-443	128	18	and	and	CCONJ
alces-443	128	19	grubb	grubb	PROPN
alces-443	128	20	(	(	PUNCT
alces-443	128	21	1987	1987	NUM
alces-443	128	22	)	)	PUNCT
alces-443	128	23	have	have	AUX
alces-443	128	24	identified	identify	VERB
alces-443	128	25	them	they	PRON
alces-443	128	26	as	as	ADP
alces-443	128	27	“	"	PUNCT
alces-443	128	28	semispecies	semispecie	NOUN
alces-443	128	29	.	.	PUNCT
alces-443	128	30	”	"	PUNCT
alces-443	129	1	we	we	PRON
alces-443	129	2	caution	caution	VERB
alces-443	129	3	,	,	PUNCT
alces-443	129	4	however	however	ADV
alces-443	129	5	,	,	PUNCT
alces-443	129	6	that	that	PRON
alces-443	129	7	chromosome	chromosome	NOUN
alces-443	129	8	numbers	number	NOUN
alces-443	129	9	may	may	AUX
alces-443	129	10	be	be	AUX
alces-443	129	11	a	a	DET
alces-443	129	12	poor	poor	ADJ
alces-443	129	13	designator	designator	NOUN
alces-443	129	14	of	of	ADP
alces-443	129	15	species	specie	NOUN
alces-443	129	16	among	among	ADP
alces-443	129	17	large	large	ADJ
alces-443	129	18	mammals	mammal	NOUN
alces-443	129	19	(	(	PUNCT
alces-443	129	20	bowyer	bowyer	PROPN
alces-443	129	21	et	et	PROPN
alces-443	129	22	al	al	PROPN
alces-443	129	23	.	.	PROPN
alces-443	129	24	2000	2000	NUM
alces-443	129	25	)	)	PUNCT
alces-443	129	26	.	.	PUNCT
alces-443	130	1	hundertmark	hundertmark	PROPN
alces-443	130	2	et	et	PROPN
alces-443	130	3	al	al	PROPN
alces-443	130	4	.	.	PROPN
alces-443	130	5	(	(	PUNCT
alces-443	130	6	2002b	2002b	NUM
alces-443	130	7	)	)	PUNCT
alces-443	130	8	tested	test	VERB
alces-443	130	9	the	the	DET
alces-443	130	10	2	2	NUM
alces-443	130	11	-	-	PUNCT
alces-443	130	12	types	type	NOUN
alces-443	130	13	hypothesis	hypothesis	NOUN
alces-443	130	14	by	by	ADP
alces-443	130	15	examining	examine	VERB
alces-443	130	16	the	the	DET
alces-443	130	17	distribution	distribution	NOUN
alces-443	130	18	of	of	ADP
alces-443	130	19	genetic	genetic	ADJ
alces-443	130	20	variance	variance	NOUN
alces-443	130	21	of	of	ADP
alces-443	130	22	mtdna	mtdna	NOUN
alces-443	130	23	among	among	ADP
alces-443	130	24	and	and	CCONJ
alces-443	130	25	within	within	ADP
alces-443	130	26	different	different	ADJ
alces-443	130	27	hypothetical	hypothetical	ADJ
alces-443	130	28	population	population	NOUN
alces-443	130	29	structures	structure	NOUN
alces-443	130	30	.	.	PUNCT
alces-443	131	1	those	those	DET
alces-443	131	2	authors	author	NOUN
alces-443	131	3	sampled	sample	VERB
alces-443	131	4	moose	moose	NOUN
alces-443	131	5	throughout	throughout	ADP
alces-443	131	6	their	their	PRON
alces-443	131	7	worldwide	worldwide	ADJ
alces-443	131	8	distribution	distribution	NOUN
alces-443	131	9	and	and	CCONJ
alces-443	131	10	arranged	arrange	VERB
alces-443	131	11	them	they	PRON
alces-443	131	12	into	into	ADP
alces-443	131	13	either	either	DET
alces-443	131	14	2	2	NUM
alces-443	131	15	groups	group	NOUN
alces-443	131	16	(	(	PUNCT
alces-443	131	17	corresponding	correspond	VERB
alces-443	131	18	to	to	ADP
alces-443	131	19	the	the	DET
alces-443	131	20	2	2	NUM
alces-443	131	21	-	-	PUNCT
alces-443	131	22	types	type	NOUN
alces-443	131	23	hypothesis	hypothesis	NOUN
alces-443	131	24	)	)	PUNCT
alces-443	131	25	or	or	CCONJ
alces-443	131	26	3	3	NUM
alces-443	131	27	groups	group	NOUN
alces-443	131	28	corresponding	correspond	VERB
alces-443	131	29	to	to	ADP
alces-443	131	30	continent	continent	NOUN
alces-443	131	31	of	of	ADP
alces-443	131	32	origin	origin	NOUN
alces-443	131	33	(	(	PUNCT
alces-443	131	34	asia	asia	PROPN
alces-443	131	35	,	,	PUNCT
alces-443	131	36	europe	europe	PROPN
alces-443	131	37	,	,	PUNCT
alces-443	131	38	and	and	CCONJ
alces-443	131	39	north	north	PROPN
alces-443	131	40	america	america	PROPN
alces-443	131	41	)	)	PUNCT
alces-443	131	42	.	.	PUNCT
alces-443	132	1	they	they	PRON
alces-443	132	2	then	then	ADV
alces-443	132	3	examined	examine	VERB
alces-443	132	4	the	the	DET
alces-443	132	5	distribution	distribution	NOUN
alces-443	132	6	of	of	ADP
alces-443	132	7	genetic	genetic	ADJ
alces-443	132	8	variance	variance	NOUN
alces-443	132	9	within	within	ADP
alces-443	132	10	and	and	CCONJ
alces-443	132	11	among	among	ADP
alces-443	132	12	or	or	CCONJ
alces-443	132	13	between	between	ADP
alces-443	132	14	groups	group	NOUN
alces-443	132	15	,	,	PUNCT
alces-443	132	16	predicting	predict	VERB
alces-443	132	17	that	that	SCONJ
alces-443	132	18	the	the	DET
alces-443	132	19	correct	correct	ADJ
alces-443	132	20	structure	structure	NOUN
alces-443	132	21	would	would	AUX
alces-443	132	22	minimize	minimize	VERB
alces-443	132	23	within	within	ADP
alces-443	132	24	-	-	PUNCT
alces-443	132	25	group	group	NOUN
alces-443	132	26	variation	variation	NOUN
alces-443	132	27	and	and	CCONJ
alces-443	132	28	maximize	maximize	VERB
alces-443	132	29	among	among	ADP
alces-443	132	30	-	-	PUNCT
alces-443	132	31	group	group	NOUN
alces-443	132	32	variation	variation	NOUN
alces-443	132	33	.	.	PUNCT
alces-443	133	1	percentage	percentage	NOUN
alces-443	133	2	of	of	ADP
alces-443	133	3	total	total	ADJ
alces-443	133	4	variation	variation	NOUN
alces-443	133	5	observed	observe	VERB
alces-443	133	6	among	among	ADP
alces-443	133	7	the	the	DET
alces-443	133	8	3	3	NUM
alces-443	133	9	groups	group	NOUN
alces-443	133	10	was	be	AUX
alces-443	133	11	slightly	slightly	ADV
alces-443	133	12	greater	great	ADJ
alces-443	133	13	than	than	ADP
alces-443	133	14	variation	variation	NOUN
alces-443	133	15	between	between	ADP
alces-443	133	16	the	the	DET
alces-443	133	17	2	2	NUM
alces-443	133	18	groups	group	NOUN
alces-443	133	19	(	(	PUNCT
alces-443	133	20	61.5	61.5	NUM
alces-443	133	21	%	%	NOUN
alces-443	133	22	vs.	vs.	ADP
alces-443	133	23	58.1	58.1	NUM
alces-443	133	24	%	%	NOUN
alces-443	133	25	)	)	PUNCT
alces-443	133	26	,	,	PUNCT
alces-443	133	27	and	and	CCONJ
alces-443	133	28	variation	variation	NOUN
alces-443	133	29	among	among	ADP
alces-443	133	30	populations	population	NOUN
alces-443	133	31	within	within	ADP
alces-443	133	32	groups	group	NOUN
alces-443	133	33	was	be	AUX
alces-443	133	34	minimized	minimize	VERB
alces-443	133	35	in	in	ADP
alces-443	133	36	the	the	DET
alces-443	133	37	3	3	NUM
alces-443	133	38	-	-	PUNCT
alces-443	133	39	group	group	NOUN
alces-443	133	40	comparison	comparison	NOUN
alces-443	133	41	(	(	PUNCT
alces-443	133	42	21.6	21.6	NUM
alces-443	133	43	%	%	NOUN
alces-443	133	44	vs.	vs.	ADP
alces-443	133	45	28.8	28.8	NUM
alces-443	133	46	%	%	NOUN
alces-443	133	47	in	in	ADP
alces-443	133	48	the	the	DET
alces-443	133	49	2	2	NUM
alces-443	133	50	-	-	PUNCT
alces-443	133	51	group	group	NOUN
alces-443	133	52	comparison	comparison	NOUN
alces-443	133	53	)	)	PUNCT
alces-443	133	54	.	.	PUNCT
alces-443	134	1	therefore	therefore	ADV
alces-443	134	2	,	,	PUNCT
alces-443	134	3	hundertmark	hundertmark	PROPN
alces-443	134	4	et	et	PROPN
alces-443	134	5	al	al	PROPN
alces-443	134	6	.	.	PROPN
alces-443	134	7	(	(	PUNCT
alces-443	134	8	2002b	2002b	NUM
alces-443	134	9	)	)	PUNCT
alces-443	134	10	concluded	conclude	VERB
alces-443	134	11	that	that	SCONJ
alces-443	134	12	mtdna	mtdna	PROPN
alces-443	134	13	data	data	PROPN
alces-443	134	14	provided	provide	VERB
alces-443	134	15	no	no	DET
alces-443	134	16	support	support	NOUN
alces-443	134	17	for	for	ADP
alces-443	134	18	a	a	DET
alces-443	134	19	2	2	NUM
alces-443	134	20	-	-	PUNCT
alces-443	134	21	type	type	NOUN
alces-443	134	22	over	over	ADP
alces-443	134	23	a	a	DET
alces-443	134	24	3type	3type	NUM
alces-443	134	25	hypothesis	hypothesis	NOUN
alces-443	134	26	.	.	PUNCT
alces-443	135	1	that	that	DET
alces-443	135	2	finding	finding	NOUN
alces-443	135	3	can	can	AUX
alces-443	135	4	be	be	AUX
alces-443	135	5	visualized	visualize	VERB
alces-443	135	6	by	by	ADP
alces-443	135	7	a	a	DET
alces-443	135	8	phylogenetic	phylogenetic	ADJ
alces-443	135	9	tree	tree	NOUN
alces-443	135	10	constructed	construct	VERB
alces-443	135	11	from	from	ADP
alces-443	135	12	haplotypes	haplotype	NOUN
alces-443	135	13	originally	originally	ADV
alces-443	135	14	reported	report	VERB
alces-443	135	15	by	by	ADP
alces-443	135	16	hundertmark	hundertmark	PROPN
alces-443	135	17	et	et	PROPN
alces-443	135	18	al	al	PROPN
alces-443	135	19	.	.	PROPN
alces-443	135	20	(	(	PUNCT
alces-443	135	21	2002b	2002b	NUM
alces-443	135	22	,	,	PUNCT
alces-443	135	23	2003	2003	NUM
alces-443	135	24	)	)	PUNCT
alces-443	135	25	,	,	PUNCT
alces-443	135	26	which	which	PRON
alces-443	135	27	shows	show	VERB
alces-443	135	28	north	north	ADJ
alces-443	135	29	american	american	ADJ
alces-443	135	30	moose	moose	NOUN
alces-443	135	31	as	as	ADP
alces-443	135	32	distinct	distinct	ADJ
alces-443	135	33	from	from	ADP
alces-443	135	34	both	both	CCONJ
alces-443	135	35	asian	asian	ADJ
alces-443	135	36	and	and	CCONJ
alces-443	135	37	european	european	ADJ
alces-443	135	38	forms	form	NOUN
alces-443	135	39	(	(	PUNCT
alces-443	135	40	fig	fig	NOUN
alces-443	135	41	.	.	PUNCT
alces-443	136	1	2	2	NUM
alces-443	136	2	)	)	PUNCT
alces-443	136	3	.	.	PUNCT
alces-443	137	1	ancient	ancient	ADJ
alces-443	137	2	bottlenecks	bottleneck	NOUN
alces-443	137	3	and	and	CCONJ
alces-443	137	4	the	the	DET
alces-443	137	5	mother	mother	NOUN
alces-443	137	6	of	of	ADP
alces-443	137	7	all	all	DET
alces-443	137	8	moose	moose	NOUN
alces-443	137	9	moose	moose	NOUN
alces-443	137	10	worldwide	worldwide	ADV
alces-443	137	11	exhibit	exhibit	VERB
alces-443	137	12	little	little	ADJ
alces-443	137	13	variation	variation	NOUN
alces-443	137	14	in	in	ADP
alces-443	137	15	a	a	DET
alces-443	137	16	fragment	fragment	NOUN
alces-443	137	17	of	of	ADP
alces-443	137	18	the	the	DET
alces-443	137	19	mitochondrial	mitochondrial	ADJ
alces-443	137	20	cytochrome	cytochrome	NOUN
alces-443	137	21	-	-	PUNCT
alces-443	137	22	b	b	NOUN
alces-443	137	23	gene	gene	NOUN
alces-443	137	24	(	(	PUNCT
alces-443	137	25	hundertmark	hundertmark	NOUN
alces-443	137	26	et	et	PROPN
alces-443	137	27	al	al	PROPN
alces-443	137	28	.	.	PROPN
alces-443	137	29	2002a	2002a	NUM
alces-443	137	30	)	)	PUNCT
alces-443	137	31	.	.	PUNCT
alces-443	138	1	cytochrome	cytochrome	PROPN
alces-443	138	2	b	b	PROPN
alces-443	138	3	is	be	AUX
alces-443	138	4	useful	useful	ADJ
alces-443	138	5	for	for	ADP
alces-443	138	6	constructing	construct	VERB
alces-443	138	7	mammalian	mammalian	ADJ
alces-443	138	8	phylogenies	phylogeny	NOUN
alces-443	138	9	(	(	PUNCT
alces-443	138	10	irwin	irwin	PROPN
alces-443	138	11	et	et	PROPN
alces-443	138	12	al	al	PROPN
alces-443	138	13	.	.	PROPN
alces-443	138	14	1991	1991	NUM
alces-443	138	15	)	)	PUNCT
alces-443	138	16	and	and	CCONJ
alces-443	138	17	a	a	DET
alces-443	138	18	paucity	paucity	NOUN
alces-443	138	19	of	of	ADP
alces-443	138	20	variation	variation	NOUN
alces-443	138	21	in	in	ADP
alces-443	138	22	moose	moose	NOUN
alces-443	138	23	indicated	indicate	VERB
alces-443	138	24	a	a	DET
alces-443	138	25	recent	recent	ADJ
alces-443	138	26	common	common	ADJ
alces-443	138	27	ancestry	ancestry	NOUN
alces-443	138	28	likely	likely	ADV
alces-443	138	29	due	due	ADP
alces-443	138	30	to	to	ADP
alces-443	138	31	a	a	DET
alces-443	138	32	severe	severe	ADJ
alces-443	138	33	bottleneck	bottleneck	NOUN
alces-443	138	34	that	that	PRON
alces-443	138	35	affected	affect	VERB
alces-443	138	36	all	all	DET
alces-443	138	37	extant	extant	ADJ
alces-443	138	38	lineages	lineage	NOUN
alces-443	138	39	(	(	PUNCT
alces-443	138	40	hundertmark	hundertmark	NOUN
alces-443	138	41	et	et	PROPN
alces-443	138	42	al	al	PROPN
alces-443	138	43	.	.	PROPN
alces-443	138	44	2002a	2002a	NUM
alces-443	138	45	)	)	PUNCT
alces-443	138	46	,	,	PUNCT
alces-443	138	47	which	which	PRON
alces-443	138	48	is	be	AUX
alces-443	138	49	in	in	ADP
alces-443	138	50	agreement	agreement	NOUN
alces-443	138	51	with	with	ADP
alces-443	138	52	the	the	DET
alces-443	138	53	findings	finding	NOUN
alces-443	138	54	of	of	ADP
alces-443	138	55	mikko	mikko	PROPN
alces-443	138	56	and	and	CCONJ
alces-443	138	57	andersson	andersson	PROPN
alces-443	138	58	(	(	PUNCT
alces-443	138	59	1995	1995	NUM
alces-443	138	60	)	)	PUNCT
alces-443	138	61	.	.	PUNCT
alces-443	139	1	analysis	analysis	NOUN
alces-443	139	2	of	of	ADP
alces-443	139	3	variation	variation	NOUN
alces-443	139	4	within	within	ADP
alces-443	139	5	the	the	DET
alces-443	139	6	mitochondrial	mitochondrial	ADJ
alces-443	139	7	control	control	NOUN
alces-443	139	8	region	region	NOUN
alces-443	139	9	,	,	PUNCT
alces-443	139	10	which	which	PRON
alces-443	139	11	evolves	evolve	VERB
alces-443	139	12	at	at	ADP
alces-443	139	13	a	a	DET
alces-443	139	14	much	much	ADV
alces-443	139	15	faster	fast	ADJ
alces-443	139	16	rate	rate	NOUN
alces-443	139	17	than	than	ADP
alces-443	139	18	cytochrome	cytochrome	PROPN
alces-443	139	19	b	b	PROPN
alces-443	139	20	(	(	PUNCT
alces-443	139	21	lopez	lopez	PROPN
alces-443	139	22	et	et	PROPN
alces-443	139	23	al	al	PROPN
alces-443	139	24	.	.	PROPN
alces-443	139	25	1997	1997	NUM
alces-443	139	26	)	)	PUNCT
alces-443	139	27	,	,	PUNCT
alces-443	139	28	was	be	AUX
alces-443	139	29	fig	fig	NOUN
alces-443	139	30	.	.	PUNCT
alces-443	140	1	2	2	X
alces-443	140	2	.	.	X
alces-443	140	3	unrooted	unrooted	ADJ
alces-443	140	4	phylogenetic	phylogenetic	ADJ
alces-443	140	5	tree	tree	NOUN
alces-443	140	6	of	of	ADP
alces-443	140	7	moose	moose	NOUN
alces-443	140	8	populations	population	NOUN
alces-443	140	9	and	and	CCONJ
alces-443	140	10	subspecies	subspecie	NOUN
alces-443	140	11	worldwide	worldwide	ADV
alces-443	140	12	,	,	PUNCT
alces-443	140	13	with	with	ADP
alces-443	140	14	the	the	DET
alces-443	140	15	exception	exception	NOUN
alces-443	140	16	of	of	ADP
alces-443	140	17	a.	a.	NOUN
alces-443	140	18	a.	a.	NOUN
alces-443	140	19	pfizenmayeri	pfizenmayeri	PROPN
alces-443	140	20	,	,	PUNCT
alces-443	140	21	using	use	VERB
alces-443	140	22	fst	fst	NOUN
alces-443	140	23	as	as	ADP
alces-443	140	24	the	the	DET
alces-443	140	25	distance	distance	NOUN
alces-443	140	26	measure	measure	NOUN
alces-443	140	27	[	[	X
alces-443	140	28	data	datum	NOUN
alces-443	140	29	from	from	ADP
alces-443	140	30	hundertmark	hundertmark	PROPN
alces-443	140	31	et	et	PROPN
alces-443	140	32	al	al	PROPN
alces-443	140	33	.	.	PROPN
alces-443	141	1	(	(	PUNCT
alces-443	141	2	2002b	2002b	NUM
alces-443	141	3	,	,	PUNCT
alces-443	141	4	2003	2003	NUM
alces-443	141	5	)	)	PUNCT
alces-443	141	6	]	]	PUNCT
alces-443	141	7	.	.	PUNCT
alces-443	142	1	note	note	VERB
alces-443	142	2	the	the	DET
alces-443	142	3	distinct	distinct	ADJ
alces-443	142	4	positions	position	NOUN
alces-443	142	5	of	of	ADP
alces-443	142	6	the	the	DET
alces-443	142	7	3	3	NUM
alces-443	142	8	eurasian	eurasian	ADJ
alces-443	142	9	subspecies	subspecie	NOUN
alces-443	142	10	and	and	CCONJ
alces-443	142	11	north	north	ADJ
alces-443	142	12	american	american	ADJ
alces-443	142	13	moose	moose	NOUN
alces-443	142	14	,	,	PUNCT
alces-443	142	15	which	which	PRON
alces-443	142	16	do	do	AUX
alces-443	142	17	not	not	PART
alces-443	142	18	support	support	VERB
alces-443	142	19	an	an	DET
alces-443	142	20	hypothesis	hypothesis	NOUN
alces-443	142	21	of	of	ADP
alces-443	142	22	2	2	NUM
alces-443	142	23	or	or	CCONJ
alces-443	142	24	3	3	NUM
alces-443	142	25	races	race	NOUN
alces-443	142	26	of	of	ADP
alces-443	142	27	moose	moose	NOUN
alces-443	142	28	worldwide	worldwide	ADV
alces-443	142	29	.	.	PUNCT
alces-443	143	1	hundertmark	hundertmark	PROPN
alces-443	143	2	and	and	CCONJ
alces-443	143	3	bowyer	bowyer	PROPN
alces-443	143	4	moose	moose	PROPN
alces-443	143	5	genetics	genetics	PROPN
alces-443	143	6	alces	alce	NOUN
alces-443	143	7	vol	vol	NOUN
alces-443	143	8	.	.	PROPN
alces-443	143	9	40	40	NUM
alces-443	143	10	,	,	PUNCT
alces-443	143	11	2004	2004	NUM
alces-443	143	12	110	110	NUM
alces-443	143	13	fig	fig	NOUN
alces-443	143	14	.	.	PUNCT
alces-443	144	1	3	3	X
alces-443	144	2	.	.	X
alces-443	144	3	a	a	DET
alces-443	144	4	phylogenetic	phylogenetic	ADJ
alces-443	144	5	tree	tree	NOUN
alces-443	144	6	of	of	ADP
alces-443	144	7	haplotypes	haplotype	NOUN
alces-443	144	8	of	of	ADP
alces-443	144	9	the	the	DET
alces-443	144	10	mitochondrial	mitochondrial	ADJ
alces-443	144	11	control	control	NOUN
alces-443	144	12	region	region	NOUN
alces-443	144	13	of	of	ADP
alces-443	144	14	moose	moose	NOUN
alces-443	144	15	.	.	PUNCT
alces-443	145	1	symbols	symbol	NOUN
alces-443	145	2	indicate	indicate	VERB
alces-443	145	3	region	region	NOUN
alces-443	145	4	of	of	ADP
alces-443	145	5	origin	origin	NOUN
alces-443	145	6	,	,	PUNCT
alces-443	145	7	with	with	ADP
alces-443	145	8	black	black	ADJ
alces-443	145	9	symbols	symbol	NOUN
alces-443	145	10	indicating	indicate	VERB
alces-443	145	11	asian	asian	ADJ
alces-443	145	12	origin	origin	NOUN
alces-443	145	13	.	.	PUNCT
alces-443	146	1	distinct	distinct	ADJ
alces-443	146	2	clades	clade	NOUN
alces-443	146	3	or	or	CCONJ
alces-443	146	4	phylogroups	phylogroup	NOUN
alces-443	146	5	are	be	AUX
alces-443	146	6	indicated	indicate	VERB
alces-443	146	7	on	on	ADP
alces-443	146	8	the	the	DET
alces-443	146	9	right	right	NOUN
alces-443	146	10	.	.	PUNCT
alces-443	147	1	from	from	ADP
alces-443	147	2	hundertmark	hundertmark	PROPN
alces-443	147	3	et	et	PROPN
alces-443	147	4	al	al	PROPN
alces-443	147	5	.	.	PROPN
alces-443	147	6	(	(	PUNCT
alces-443	147	7	2002b	2002b	NUM
alces-443	147	8	)	)	PUNCT
alces-443	147	9	.	.	PUNCT
alces-443	147	10	necessary	necessary	ADJ
alces-443	147	11	to	to	PART
alces-443	147	12	reveal	reveal	VERB
alces-443	147	13	significant	significant	ADJ
alces-443	147	14	levels	level	NOUN
alces-443	147	15	of	of	ADP
alces-443	147	16	variation	variation	NOUN
alces-443	147	17	in	in	ADP
alces-443	147	18	moose	moose	NOUN
alces-443	147	19	and	and	CCONJ
alces-443	147	20	subsequently	subsequently	ADV
alces-443	147	21	geographic	geographic	ADJ
alces-443	147	22	patterns	pattern	NOUN
alces-443	147	23	were	be	AUX
alces-443	147	24	revealed	reveal	VERB
alces-443	147	25	.	.	PUNCT
alces-443	148	1	control	control	NOUN
alces-443	148	2	region	region	NOUN
alces-443	148	3	haplotypes	haplotype	NOUN
alces-443	148	4	of	of	ADP
alces-443	148	5	moose	moose	NOUN
alces-443	148	6	can	can	AUX
alces-443	148	7	be	be	AUX
alces-443	148	8	divided	divide	VERB
alces-443	148	9	into	into	ADP
alces-443	148	10	3	3	NUM
alces-443	148	11	clades	clade	NOUN
alces-443	148	12	,	,	PUNCT
alces-443	148	13	or	or	CCONJ
alces-443	148	14	haplogroups	haplogroup	NOUN
alces-443	148	15	(	(	PUNCT
alces-443	148	16	fig	fig	NOUN
alces-443	148	17	.	.	PUNCT
alces-443	149	1	3	3	NUM
alces-443	149	2	)	)	PUNCT
alces-443	149	3	.	.	PUNCT
alces-443	150	1	the	the	DET
alces-443	150	2	basal	basal	ADJ
alces-443	150	3	haplogroup	haplogroup	NOUN
alces-443	150	4	(	(	PUNCT
alces-443	150	5	i.e.	i.e.	X
alces-443	150	6	,	,	PUNCT
alces-443	150	7	the	the	DET
alces-443	150	8	group	group	NOUN
alces-443	150	9	that	that	PRON
alces-443	150	10	diverges	diverge	VERB
alces-443	150	11	first	first	ADV
alces-443	150	12	from	from	ADP
alces-443	150	13	the	the	DET
alces-443	150	14	base	base	NOUN
alces-443	150	15	of	of	ADP
alces-443	150	16	the	the	DET
alces-443	150	17	tree	tree	NOUN
alces-443	150	18	)	)	PUNCT
alces-443	150	19	is	be	AUX
alces-443	150	20	entirely	entirely	ADV
alces-443	150	21	asian	asian	ADJ
alces-443	150	22	,	,	PUNCT
alces-443	150	23	which	which	PRON
alces-443	150	24	suggests	suggest	VERB
alces-443	150	25	that	that	SCONJ
alces-443	150	26	those	those	PRON
alces-443	150	27	are	be	AUX
alces-443	150	28	the	the	DET
alces-443	150	29	oldest	old	ADJ
alces-443	150	30	moose	moose	NOUN
alces-443	150	31	haplotypes	haplotype	NOUN
alces-443	150	32	.	.	PUNCT
alces-443	151	1	the	the	DET
alces-443	151	2	two	two	NUM
alces-443	151	3	other	other	ADJ
alces-443	151	4	haplogroups	haplogroup	NOUN
alces-443	151	5	are	be	AUX
alces-443	151	6	primarily	primarily	ADV
alces-443	151	7	european	european	ADJ
alces-443	151	8	and	and	CCONJ
alces-443	151	9	primarily	primarily	ADV
alces-443	151	10	north	north	NOUN
alces-443	151	11	american	american	ADJ
alces-443	151	12	,	,	PUNCT
alces-443	151	13	although	although	SCONJ
alces-443	151	14	some	some	DET
alces-443	151	15	asian	asian	ADJ
alces-443	151	16	haplotypes	haplotype	NOUN
alces-443	151	17	occur	occur	VERB
alces-443	151	18	in	in	ADP
alces-443	151	19	both	both	PRON
alces-443	151	20	.	.	PUNCT
alces-443	152	1	the	the	DET
alces-443	152	2	distribution	distribution	NOUN
alces-443	152	3	of	of	ADP
alces-443	152	4	haplogroups	haplogroup	NOUN
alces-443	152	5	on	on	ADP
alces-443	152	6	a	a	DET
alces-443	152	7	worldwide	worldwide	ADJ
alces-443	152	8	scale	scale	NOUN
alces-443	152	9	illustrates	illustrate	VERB
alces-443	152	10	the	the	DET
alces-443	152	11	age	age	NOUN
alces-443	152	12	of	of	ADP
alces-443	152	13	continental	continental	ADJ
alces-443	152	14	assemblages	assemblage	NOUN
alces-443	152	15	of	of	ADP
alces-443	152	16	haplotypes	haplotype	NOUN
alces-443	152	17	(	(	PUNCT
alces-443	152	18	fig	fig	NOUN
alces-443	152	19	.	.	PUNCT
alces-443	152	20	4	4	NUM
alces-443	152	21	)	)	PUNCT
alces-443	152	22	.	.	PUNCT
alces-443	153	1	the	the	DET
alces-443	153	2	yakutia	yakutia	PROPN
alces-443	153	3	area	area	NOUN
alces-443	153	4	contains	contain	VERB
alces-443	153	5	moose	moose	NOUN
alces-443	153	6	from	from	ADP
alces-443	153	7	all	all	DET
alces-443	153	8	3	3	NUM
alces-443	153	9	haplogroups	haplogroup	NOUN
alces-443	153	10	.	.	PUNCT
alces-443	154	1	the	the	DET
alces-443	154	2	russian	russian	ADJ
alces-443	154	3	far	far	PROPN
alces-443	154	4	east	east	PROPN
alces-443	154	5	contains	contain	VERB
alces-443	154	6	both	both	CCONJ
alces-443	154	7	european	european	ADJ
alces-443	154	8	and	and	CCONJ
alces-443	154	9	asian	asian	ADJ
alces-443	154	10	haplogroups	haplogroup	NOUN
alces-443	154	11	,	,	PUNCT
alces-443	154	12	but	but	CCONJ
alces-443	154	13	not	not	PART
alces-443	154	14	north	north	NOUN
alces-443	154	15	american	american	ADJ
alces-443	154	16	,	,	PUNCT
alces-443	154	17	and	and	CCONJ
alces-443	154	18	both	both	CCONJ
alces-443	154	19	europe	europe	PROPN
alces-443	154	20	and	and	CCONJ
alces-443	154	21	north	north	PROPN
alces-443	154	22	america	america	PROPN
alces-443	154	23	contain	contain	VERB
alces-443	154	24	only	only	ADV
alces-443	154	25	1	1	NUM
alces-443	154	26	haplogroup	haplogroup	NOUN
alces-443	154	27	each	each	PRON
alces-443	154	28	.	.	PUNCT
alces-443	155	1	therefore	therefore	ADV
alces-443	155	2	,	,	PUNCT
alces-443	155	3	yakutia	yakutia	PROPN
alces-443	155	4	can	can	AUX
alces-443	155	5	be	be	AUX
alces-443	155	6	identified	identify	VERB
alces-443	155	7	as	as	ADP
alces-443	155	8	the	the	DET
alces-443	155	9	area	area	NOUN
alces-443	155	10	from	from	ADP
alces-443	155	11	which	which	PRON
alces-443	155	12	all	all	DET
alces-443	155	13	extant	extant	ADJ
alces-443	155	14	moose	moose	NOUN
alces-443	155	15	lineages	lineage	NOUN
alces-443	155	16	were	be	AUX
alces-443	155	17	derived	derive	VERB
alces-443	155	18	,	,	PUNCT
alces-443	155	19	i.e.	i.e.	X
alces-443	155	20	,	,	PUNCT
alces-443	155	21	it	it	PRON
alces-443	155	22	is	be	AUX
alces-443	155	23	the	the	DET
alces-443	155	24	oldest	old	ADJ
alces-443	155	25	extant	extant	ADJ
alces-443	155	26	moose	moose	NOUN
alces-443	155	27	population	population	NOUN
alces-443	155	28	that	that	PRON
alces-443	155	29	has	have	AUX
alces-443	155	30	been	be	AUX
alces-443	155	31	sampled	sample	VERB
alces-443	155	32	.	.	PUNCT
alces-443	156	1	yakutia	yakutia	PROPN
alces-443	156	2	probably	probably	ADV
alces-443	156	3	was	be	AUX
alces-443	156	4	the	the	DET
alces-443	156	5	center	center	NOUN
alces-443	156	6	of	of	ADP
alces-443	156	7	a	a	DET
alces-443	156	8	single	single	ADJ
alces-443	156	9	moose	moose	NOUN
alces-443	156	10	population	population	NOUN
alces-443	156	11	during	during	ADP
alces-443	156	12	the	the	DET
alces-443	156	13	last	last	ADJ
alces-443	156	14	ice	ice	NOUN
alces-443	156	15	age	age	NOUN
alces-443	156	16	,	,	PUNCT
alces-443	156	17	or	or	CCONJ
alces-443	156	18	at	at	ADP
alces-443	156	19	least	least	ADJ
alces-443	156	20	was	be	AUX
alces-443	156	21	the	the	DET
alces-443	156	22	location	location	NOUN
alces-443	156	23	of	of	ADP
alces-443	156	24	the	the	DET
alces-443	156	25	only	only	ADJ
alces-443	156	26	population	population	NOUN
alces-443	156	27	to	to	PART
alces-443	156	28	provide	provide	VERB
alces-443	156	29	descendants	descendant	NOUN
alces-443	156	30	of	of	ADP
alces-443	156	31	modern	modern	ADJ
alces-443	156	32	moose	moose	NOUN
alces-443	156	33	.	.	PUNCT
alces-443	157	1	moose	moose	NOUN
alces-443	157	2	would	would	AUX
alces-443	157	3	have	have	AUX
alces-443	157	4	been	be	AUX
alces-443	157	5	restricted	restrict	VERB
alces-443	157	6	in	in	ADP
alces-443	157	7	their	their	PRON
alces-443	157	8	distribution	distribution	NOUN
alces-443	157	9	because	because	SCONJ
alces-443	157	10	the	the	DET
alces-443	157	11	cooler	cool	ADJ
alces-443	157	12	climate	climate	NOUN
alces-443	157	13	in	in	ADP
alces-443	157	14	asia	asia	PROPN
alces-443	157	15	at	at	ADP
alces-443	157	16	that	that	DET
alces-443	157	17	time	time	NOUN
alces-443	157	18	would	would	AUX
alces-443	157	19	have	have	AUX
alces-443	157	20	resulted	result	VERB
alces-443	157	21	in	in	ADP
alces-443	157	22	a	a	DET
alces-443	157	23	shift	shift	NOUN
alces-443	157	24	of	of	ADP
alces-443	157	25	boreal	boreal	ADJ
alces-443	157	26	forest	forest	NOUN
alces-443	157	27	habitat	habitat	NOUN
alces-443	157	28	to	to	ADP
alces-443	157	29	the	the	DET
alces-443	157	30	south	south	NOUN
alces-443	157	31	.	.	PUNCT
alces-443	158	1	that	that	DET
alces-443	158	2	habitat	habitat	NOUN
alces-443	158	3	could	could	AUX
alces-443	158	4	have	have	AUX
alces-443	158	5	shifted	shift	VERB
alces-443	158	6	only	only	ADV
alces-443	158	7	so	so	ADV
alces-443	158	8	far	far	ADV
alces-443	158	9	southward	southward	ADV
alces-443	158	10	,	,	PUNCT
alces-443	158	11	because	because	SCONJ
alces-443	158	12	of	of	ADP
alces-443	158	13	prominent	prominent	ADJ
alces-443	158	14	mountains	mountain	NOUN
alces-443	158	15	running	run	VERB
alces-443	158	16	east	east	NOUN
alces-443	158	17	-	-	PUNCT
alces-443	158	18	west	west	PROPN
alces-443	158	19	,	,	PUNCT
alces-443	158	20	which	which	PRON
alces-443	158	21	would	would	AUX
alces-443	158	22	have	have	AUX
alces-443	158	23	formed	form	VERB
alces-443	158	24	an	an	DET
alces-443	158	25	effective	effective	ADJ
alces-443	158	26	barrier	barrier	NOUN
alces-443	158	27	to	to	ADP
alces-443	158	28	further	further	ADJ
alces-443	158	29	movement	movement	NOUN
alces-443	158	30	to	to	ADP
alces-443	158	31	the	the	DET
alces-443	158	32	south	south	NOUN
alces-443	158	33	(	(	PUNCT
alces-443	158	34	hewitt	hewitt	PROPN
alces-443	158	35	1996	1996	NUM
alces-443	158	36	)	)	PUNCT
alces-443	158	37	.	.	PUNCT
alces-443	159	1	during	during	ADP
alces-443	159	2	the	the	DET
alces-443	159	3	last	last	ADJ
alces-443	159	4	ice	ice	NOUN
alces-443	159	5	age	age	NOUN
alces-443	159	6	,	,	PUNCT
alces-443	159	7	there	there	PRON
alces-443	159	8	were	be	VERB
alces-443	159	9	2	2	NUM
alces-443	159	10	periods	period	NOUN
alces-443	159	11	of	of	ADP
alces-443	159	12	maximum	maximum	ADJ
alces-443	159	13	glacial	glacial	ADJ
alces-443	159	14	advance	advance	NOUN
alces-443	159	15	,	,	PUNCT
alces-443	159	16	termed	term	VERB
alces-443	159	17	the	the	DET
alces-443	159	18	lower	low	ADJ
alces-443	159	19	and	and	CCONJ
alces-443	159	20	upper	upper	ADJ
alces-443	159	21	pleniglacials	pleniglacial	NOUN
alces-443	159	22	.	.	PUNCT
alces-443	160	1	those	those	DET
alces-443	160	2	episodes	episode	NOUN
alces-443	160	3	occurred	occur	VERB
alces-443	160	4	at	at	ADP
alces-443	160	5	approximately	approximately	ADV
alces-443	160	6	62,000	62,000	NUM
alces-443	160	7	and	and	CCONJ
alces-443	160	8	20,000	20,000	NUM
alces-443	160	9	years	year	NOUN
alces-443	160	10	ago	ago	ADV
alces-443	160	11	,	,	PUNCT
alces-443	160	12	respectively	respectively	ADV
alces-443	160	13	(	(	PUNCT
alces-443	160	14	fulton	fulton	PROPN
alces-443	160	15	et	et	PROPN
alces-443	160	16	al	al	PROPN
alces-443	160	17	.	.	PROPN
alces-443	160	18	1986	1986	NUM
alces-443	160	19	)	)	PUNCT
alces-443	160	20	.	.	PUNCT
alces-443	161	1	boreal	boreal	ADJ
alces-443	161	2	forest	forest	NOUN
alces-443	161	3	habitats	habitat	NOUN
alces-443	161	4	in	in	ADP
alces-443	161	5	asia	asia	PROPN
alces-443	161	6	would	would	AUX
alces-443	161	7	have	have	AUX
alces-443	161	8	shifted	shift	VERB
alces-443	161	9	to	to	ADP
alces-443	161	10	the	the	DET
alces-443	161	11	south	south	NOUN
alces-443	161	12	during	during	ADP
alces-443	161	13	those	those	PRON
alces-443	161	14	cooling	cool	VERB
alces-443	161	15	episodes	episode	NOUN
alces-443	161	16	and	and	CCONJ
alces-443	161	17	would	would	AUX
alces-443	161	18	have	have	AUX
alces-443	161	19	been	be	AUX
alces-443	161	20	compressed	compress	VERB
alces-443	161	21	against	against	ADP
alces-443	161	22	the	the	DET
alces-443	161	23	northern	northern	ADJ
alces-443	161	24	slopes	slope	NOUN
alces-443	161	25	of	of	ADP
alces-443	161	26	mounfig	mounfig	PROPN
alces-443	161	27	.	.	PUNCT
alces-443	162	1	4	4	X
alces-443	162	2	.	.	X
alces-443	162	3	distribution	distribution	NOUN
alces-443	162	4	of	of	ADP
alces-443	162	5	mitochondrial	mitochondrial	ADJ
alces-443	162	6	haplogroups	haplogroup	NOUN
alces-443	162	7	worldwide	worldwide	ADV
alces-443	162	8	.	.	PUNCT
alces-443	163	1	note	note	VERB
alces-443	163	2	that	that	PRON
alces-443	163	3	moose	moose	NOUN
alces-443	163	4	from	from	ADP
alces-443	163	5	the	the	DET
alces-443	163	6	yakutia	yakutia	PROPN
alces-443	163	7	area	area	NOUN
alces-443	163	8	have	have	VERB
alces-443	163	9	the	the	DET
alces-443	163	10	most	most	ADV
alces-443	163	11	diverse	diverse	ADJ
alces-443	163	12	composition	composition	NOUN
alces-443	163	13	and	and	CCONJ
alces-443	163	14	that	that	DET
alces-443	163	15	moose	moose	NOUN
alces-443	163	16	from	from	ADP
alces-443	163	17	north	north	PROPN
alces-443	163	18	america	america	PROPN
alces-443	163	19	do	do	AUX
alces-443	163	20	not	not	PART
alces-443	163	21	share	share	VERB
alces-443	163	22	haplogroups	haplogroup	NOUN
alces-443	163	23	with	with	ADP
alces-443	163	24	moose	moose	NOUN
alces-443	163	25	from	from	ADP
alces-443	163	26	russian	russian	PROPN
alces-443	163	27	far	far	PROPN
alces-443	163	28	east	east	PROPN
alces-443	163	29	.	.	PUNCT
alces-443	164	1	alces	alce	NOUN
alces-443	164	2	vol	vol	NOUN
alces-443	164	3	.	.	PROPN
alces-443	164	4	40	40	NUM
alces-443	164	5	,	,	PUNCT
alces-443	164	6	2004	2004	NUM
alces-443	164	7	moose	moose	NOUN
alces-443	164	8	genetics	genetic	NOUN
alces-443	164	9	hundertmark	hundertmark	NOUN
alces-443	164	10	and	and	CCONJ
alces-443	164	11	bowyer	bowyer	PROPN
alces-443	164	12	111	111	NUM
alces-443	164	13	59,000	59,000	NUM
alces-443	164	14	and	and	CCONJ
alces-443	164	15	14,000	14,000	NUM
alces-443	164	16	years	year	NOUN
alces-443	164	17	ago	ago	ADV
alces-443	164	18	.	.	PUNCT
alces-443	165	1	when	when	SCONJ
alces-443	165	2	expansion	expansion	NOUN
alces-443	165	3	times	time	NOUN
alces-443	165	4	of	of	ADP
alces-443	165	5	moose	moose	NOUN
alces-443	165	6	are	be	AUX
alces-443	165	7	overlaid	overlay	VERB
alces-443	165	8	on	on	ADP
alces-443	165	9	a	a	DET
alces-443	165	10	profile	profile	NOUN
alces-443	165	11	of	of	ADP
alces-443	165	12	global	global	ADJ
alces-443	165	13	temperature	temperature	NOUN
alces-443	165	14	change	change	NOUN
alces-443	165	15	for	for	ADP
alces-443	165	16	the	the	DET
alces-443	165	17	last	last	ADJ
alces-443	165	18	ice	ice	NOUN
alces-443	165	19	age	age	NOUN
alces-443	165	20	(	(	PUNCT
alces-443	165	21	jouzel	jouzel	PROPN
alces-443	165	22	et	et	PROPN
alces-443	165	23	al	al	PROPN
alces-443	165	24	.	.	PROPN
alces-443	165	25	1987	1987	NUM
alces-443	165	26	)	)	PUNCT
alces-443	165	27	,	,	PUNCT
alces-443	165	28	population	population	NOUN
alces-443	165	29	expansion	expansion	NOUN
alces-443	165	30	is	be	AUX
alces-443	165	31	correlated	correlate	VERB
alces-443	165	32	with	with	ADP
alces-443	165	33	warming	warm	VERB
alces-443	165	34	trends	trend	NOUN
alces-443	165	35	following	follow	VERB
alces-443	165	36	the	the	DET
alces-443	165	37	pleniglacials	pleniglacial	NOUN
alces-443	165	38	(	(	PUNCT
alces-443	165	39	fig	fig	NOUN
alces-443	165	40	.	.	PUNCT
alces-443	166	1	6	6	NUM
alces-443	166	2	)	)	PUNCT
alces-443	166	3	.	.	PUNCT
alces-443	167	1	consequently	consequently	ADV
alces-443	167	2	,	,	PUNCT
alces-443	167	3	the	the	DET
alces-443	167	4	evolution	evolution	NOUN
alces-443	167	5	and	and	CCONJ
alces-443	167	6	geographic	geographic	ADJ
alces-443	167	7	distribution	distribution	NOUN
alces-443	167	8	of	of	ADP
alces-443	167	9	moose	moose	NOUN
alces-443	167	10	seems	seem	VERB
alces-443	167	11	to	to	PART
alces-443	167	12	have	have	AUX
alces-443	167	13	been	be	AUX
alces-443	167	14	affected	affect	VERB
alces-443	167	15	substantially	substantially	ADV
alces-443	167	16	by	by	ADP
alces-443	167	17	climate	climate	NOUN
alces-443	167	18	change	change	NOUN
alces-443	167	19	,	,	PUNCT
alces-443	167	20	particularly	particularly	ADV
alces-443	167	21	climate	climate	NOUN
alces-443	167	22	reversals	reversal	NOUN
alces-443	167	23	associated	associate	VERB
alces-443	167	24	with	with	ADP
alces-443	167	25	the	the	DET
alces-443	167	26	late	late	ADJ
alces-443	167	27	pleistocene	pleistocene	NOUN
alces-443	167	28	and	and	CCONJ
alces-443	167	29	early	early	ADJ
alces-443	167	30	holocene	holocene	PROPN
alces-443	167	31	.	.	PUNCT
alces-443	168	1	coming	come	VERB
alces-443	168	2	to	to	ADP
alces-443	168	3	america	america	PROPN
alces-443	168	4	cronin	cronin	PROPN
alces-443	168	5	(	(	PUNCT
alces-443	168	6	1992	1992	NUM
alces-443	168	7	)	)	PUNCT
alces-443	168	8	analyzed	analyze	VERB
alces-443	168	9	subspecific	subspecific	ADJ
alces-443	168	10	variation	variation	NOUN
alces-443	168	11	in	in	ADP
alces-443	168	12	north	north	PROPN
alces-443	168	13	american	american	PROPN
alces-443	168	14	cervids	cervid	NOUN
alces-443	168	15	using	use	VERB
alces-443	168	16	restriction	restriction	NOUN
alces-443	168	17	fragment	fragment	NOUN
alces-443	168	18	length	length	NOUN
alces-443	168	19	polymorphisms	polymorphism	NOUN
alces-443	168	20	(	(	PUNCT
alces-443	168	21	rflp	rflp	PROPN
alces-443	168	22	)	)	PUNCT
alces-443	168	23	of	of	ADP
alces-443	168	24	mtdna	mtdna	PROPN
alces-443	168	25	.	.	PUNCT
alces-443	169	1	despite	despite	SCONJ
alces-443	169	2	analyzing	analyze	VERB
alces-443	169	3	32	32	NUM
alces-443	169	4	moose	moose	NOUN
alces-443	169	5	sampled	sample	VERB
alces-443	169	6	from	from	ADP
alces-443	169	7	different	different	ADJ
alces-443	169	8	regions	region	NOUN
alces-443	169	9	of	of	ADP
alces-443	169	10	north	north	PROPN
alces-443	169	11	america	america	PROPN
alces-443	169	12	,	,	PUNCT
alces-443	169	13	he	he	PRON
alces-443	169	14	documented	document	VERB
alces-443	169	15	no	no	DET
alces-443	169	16	variation	variation	NOUN
alces-443	169	17	among	among	ADP
alces-443	169	18	haplotypes	haplotype	NOUN
alces-443	169	19	from	from	ADP
alces-443	169	20	that	that	DET
alces-443	169	21	continent	continent	NOUN
alces-443	169	22	.	.	PUNCT
alces-443	170	1	in	in	ADP
alces-443	170	2	comparison	comparison	NOUN
alces-443	170	3	to	to	ADP
alces-443	170	4	other	other	ADJ
alces-443	170	5	north	north	ADJ
alces-443	170	6	american	american	ADJ
alces-443	170	7	cervids	cervid	NOUN
alces-443	170	8	,	,	PUNCT
alces-443	170	9	the	the	DET
alces-443	170	10	lack	lack	NOUN
alces-443	170	11	of	of	ADP
alces-443	170	12	variation	variation	NOUN
alces-443	170	13	among	among	ADP
alces-443	170	14	subspecies	subspecie	NOUN
alces-443	170	15	of	of	ADP
alces-443	170	16	moose	moose	NOUN
alces-443	170	17	was	be	AUX
alces-443	170	18	interpreted	interpret	VERB
alces-443	170	19	as	as	ADP
alces-443	170	20	an	an	DET
alces-443	170	21	indicafig	indicafig	NOUN
alces-443	170	22	.	.	PUNCT
alces-443	171	1	5	5	X
alces-443	171	2	.	.	X
alces-443	171	3	glacial	glacial	ADJ
alces-443	171	4	coverage	coverage	NOUN
alces-443	171	5	during	during	ADP
alces-443	171	6	the	the	DET
alces-443	171	7	last	last	ADJ
alces-443	171	8	glacial	glacial	ADJ
alces-443	171	9	maximum	maximum	NOUN
alces-443	171	10	,	,	PUNCT
alces-443	171	11	superimposed	superimpose	VERB
alces-443	171	12	on	on	ADP
alces-443	171	13	a	a	DET
alces-443	171	14	map	map	NOUN
alces-443	171	15	of	of	ADP
alces-443	171	16	presentday	presentday	PROPN
alces-443	171	17	sea	sea	NOUN
alces-443	171	18	level	level	NOUN
alces-443	171	19	.	.	PUNCT
alces-443	172	1	note	note	VERB
alces-443	172	2	that	that	SCONJ
alces-443	172	3	the	the	DET
alces-443	172	4	bering	bere	VERB
alces-443	172	5	land	land	NOUN
alces-443	172	6	bridge	bridge	NOUN
alces-443	172	7	between	between	ADP
alces-443	172	8	north	north	PROPN
alces-443	172	9	america	america	PROPN
alces-443	172	10	and	and	CCONJ
alces-443	172	11	asia	asia	PROPN
alces-443	172	12	would	would	AUX
alces-443	172	13	have	have	AUX
alces-443	172	14	been	be	AUX
alces-443	172	15	exposed	expose	VERB
alces-443	172	16	during	during	ADP
alces-443	172	17	the	the	DET
alces-443	172	18	glacial	glacial	ADJ
alces-443	172	19	maximum	maximum	NOUN
alces-443	172	20	due	due	ADP
alces-443	172	21	to	to	ADP
alces-443	172	22	lower	low	ADJ
alces-443	172	23	sea	sea	NOUN
alces-443	172	24	levels	level	NOUN
alces-443	172	25	.	.	PUNCT
alces-443	173	1	names	name	NOUN
alces-443	173	2	of	of	ADP
alces-443	173	3	major	major	ADJ
alces-443	173	4	ice	ice	NOUN
alces-443	173	5	sheets	sheet	NOUN
alces-443	173	6	are	be	AUX
alces-443	173	7	provided	provide	VERB
alces-443	173	8	.	.	PUNCT
alces-443	174	1	tain	tain	PROPN
alces-443	174	2	ranges	range	VERB
alces-443	174	3	.	.	PUNCT
alces-443	175	1	during	during	ADP
alces-443	175	2	subsequent	subsequent	ADJ
alces-443	175	3	warming	warming	NOUN
alces-443	175	4	intervals	interval	NOUN
alces-443	175	5	,	,	PUNCT
alces-443	175	6	moose	moose	NOUN
alces-443	175	7	habitat	habitat	NOUN
alces-443	175	8	would	would	AUX
alces-443	175	9	have	have	AUX
alces-443	175	10	spread	spread	VERB
alces-443	175	11	to	to	ADP
alces-443	175	12	the	the	DET
alces-443	175	13	north	north	NOUN
alces-443	175	14	,	,	PUNCT
alces-443	175	15	allowing	allow	VERB
alces-443	175	16	moose	moose	NOUN
alces-443	175	17	populations	population	NOUN
alces-443	175	18	to	to	PART
alces-443	175	19	expand	expand	VERB
alces-443	175	20	(	(	PUNCT
alces-443	175	21	guthrie	guthrie	PROPN
alces-443	175	22	1995	1995	NUM
alces-443	175	23	)	)	PUNCT
alces-443	175	24	.	.	PUNCT
alces-443	176	1	unlike	unlike	ADP
alces-443	176	2	north	north	PROPN
alces-443	176	3	america	america	PROPN
alces-443	176	4	,	,	PUNCT
alces-443	176	5	much	much	ADJ
alces-443	176	6	of	of	ADP
alces-443	176	7	eurasia	eurasia	PROPN
alces-443	176	8	was	be	AUX
alces-443	176	9	not	not	PART
alces-443	176	10	glaciated	glaciate	VERB
alces-443	176	11	during	during	ADP
alces-443	176	12	those	those	DET
alces-443	176	13	periods	period	NOUN
alces-443	176	14	(	(	PUNCT
alces-443	176	15	arkhipov	arkhipov	PROPN
alces-443	176	16	et	et	PROPN
alces-443	176	17	al	al	PROPN
alces-443	176	18	.	.	PROPN
alces-443	176	19	1986	1986	NUM
alces-443	176	20	)	)	PUNCT
alces-443	176	21	,	,	PUNCT
alces-443	176	22	providing	provide	VERB
alces-443	176	23	potential	potential	ADJ
alces-443	176	24	dispersal	dispersal	NOUN
alces-443	176	25	routes	route	NOUN
alces-443	176	26	across	across	ADP
alces-443	176	27	the	the	DET
alces-443	176	28	continent	continent	NOUN
alces-443	176	29	(	(	PUNCT
alces-443	176	30	fig	fig	NOUN
alces-443	176	31	.	.	PUNCT
alces-443	177	1	5	5	NUM
alces-443	177	2	)	)	PUNCT
alces-443	177	3	.	.	PUNCT
alces-443	178	1	the	the	DET
alces-443	178	2	process	process	NOUN
alces-443	178	3	of	of	ADP
alces-443	178	4	latitudinal	latitudinal	ADJ
alces-443	178	5	shifts	shift	NOUN
alces-443	178	6	of	of	ADP
alces-443	178	7	range	range	NOUN
alces-443	178	8	associated	associate	VERB
alces-443	178	9	with	with	ADP
alces-443	178	10	episodes	episode	NOUN
alces-443	178	11	of	of	ADP
alces-443	178	12	climate	climate	NOUN
alces-443	178	13	change	change	NOUN
alces-443	178	14	results	result	NOUN
alces-443	178	15	in	in	ADP
alces-443	178	16	decreases	decrease	NOUN
alces-443	178	17	of	of	ADP
alces-443	178	18	existing	exist	VERB
alces-443	178	19	genetic	genetic	ADJ
alces-443	178	20	diversity	diversity	NOUN
alces-443	178	21	(	(	PUNCT
alces-443	178	22	hewitt	hewitt	PROPN
alces-443	178	23	1996	1996	NUM
alces-443	178	24	)	)	PUNCT
alces-443	178	25	and	and	CCONJ
alces-443	178	26	leaves	leave	VERB
alces-443	178	27	characteristic	characteristic	ADJ
alces-443	178	28	signatures	signature	NOUN
alces-443	178	29	in	in	ADP
alces-443	178	30	the	the	DET
alces-443	178	31	genome	genome	NOUN
alces-443	178	32	of	of	ADP
alces-443	178	33	modern	modern	ADJ
alces-443	178	34	moose	moose	NOUN
alces-443	178	35	.	.	PUNCT
alces-443	179	1	moose	moose	PROPN
alces-443	179	2	in	in	ADP
alces-443	179	3	eurasia	eurasia	PROPN
alces-443	179	4	underwent	undergo	VERB
alces-443	179	5	2	2	NUM
alces-443	179	6	distinct	distinct	ADJ
alces-443	179	7	,	,	PUNCT
alces-443	179	8	recent	recent	ADJ
alces-443	179	9	population	population	NOUN
alces-443	179	10	expansions	expansion	NOUN
alces-443	179	11	(	(	PUNCT
alces-443	179	12	hundertmark	hundertmark	NOUN
alces-443	179	13	et	et	PROPN
alces-443	179	14	al	al	PROPN
alces-443	179	15	.	.	PROPN
alces-443	179	16	2002b	2002b	NUM
alces-443	179	17	)	)	PUNCT
alces-443	179	18	.	.	PUNCT
alces-443	180	1	any	any	DET
alces-443	180	2	other	other	ADJ
alces-443	180	3	historic	historic	ADJ
alces-443	180	4	population	population	NOUN
alces-443	180	5	processes	process	NOUN
alces-443	180	6	preceding	precede	VERB
alces-443	180	7	those	those	DET
alces-443	180	8	expansions	expansion	NOUN
alces-443	180	9	are	be	AUX
alces-443	180	10	not	not	PART
alces-443	180	11	detectable	detectable	ADJ
alces-443	180	12	because	because	SCONJ
alces-443	180	13	low	low	ADJ
alces-443	180	14	population	population	NOUN
alces-443	180	15	sizes	size	NOUN
alces-443	180	16	eliminated	eliminate	VERB
alces-443	180	17	much	much	ADJ
alces-443	180	18	of	of	ADP
alces-443	180	19	the	the	DET
alces-443	180	20	genetic	genetic	ADJ
alces-443	180	21	variation	variation	NOUN
alces-443	180	22	present	present	ADJ
alces-443	180	23	in	in	ADP
alces-443	180	24	the	the	DET
alces-443	180	25	pre	pre	ADJ
alces-443	180	26	-	-	ADJ
alces-443	180	27	bottleneck	bottleneck	ADJ
alces-443	180	28	populations	population	NOUN
alces-443	180	29	,	,	PUNCT
alces-443	180	30	and	and	CCONJ
alces-443	180	31	hence	hence	ADV
alces-443	180	32	no	no	DET
alces-443	180	33	signature	signature	NOUN
alces-443	180	34	from	from	ADP
alces-443	180	35	those	those	DET
alces-443	180	36	times	time	NOUN
alces-443	180	37	exists	exist	VERB
alces-443	180	38	in	in	ADP
alces-443	180	39	the	the	DET
alces-443	180	40	present	present	ADJ
alces-443	180	41	genome	genome	NOUN
alces-443	180	42	.	.	PUNCT
alces-443	181	1	by	by	ADP
alces-443	181	2	applying	apply	VERB
alces-443	181	3	a	a	DET
alces-443	181	4	molecular	molecular	ADJ
alces-443	181	5	clock	clock	NOUN
alces-443	181	6	to	to	ADP
alces-443	181	7	those	those	DET
alces-443	181	8	expansion	expansion	NOUN
alces-443	181	9	data	datum	NOUN
alces-443	181	10	,	,	PUNCT
alces-443	181	11	hundertmark	hundertmark	PROPN
alces-443	181	12	et	et	PROPN
alces-443	181	13	al	al	PROPN
alces-443	181	14	.	.	PROPN
alces-443	182	1	(	(	PUNCT
alces-443	182	2	2002b	2002b	NUM
alces-443	182	3	)	)	PUNCT
alces-443	182	4	estimated	estimate	VERB
alces-443	182	5	that	that	SCONJ
alces-443	182	6	the	the	DET
alces-443	182	7	expansions	expansion	NOUN
alces-443	182	8	occurred	occur	VERB
alces-443	182	9	approximately	approximately	ADV
alces-443	182	10	160	160	NUM
alces-443	182	11	140	140	NUM
alces-443	182	12	120	120	NUM
alces-443	182	13	100	100	NUM
alces-443	182	14	80	80	NUM
alces-443	182	15	60	60	NUM
alces-443	182	16	40	40	NUM
alces-443	182	17	20	20	NUM
alces-443	182	18	0	0	NUM
alces-443	182	19	2	2	NUM
alces-443	182	20	0	0	NUM
alces-443	183	1	-2	-2	INTJ
alces-443	183	2	-4	-4	INTJ
alces-443	184	1	-6	-6	INTJ
alces-443	185	1	-8	-8	INTJ
alces-443	185	2	-10	-10	PUNCT
alces-443	185	3	thousands	thousand	NOUN
alces-443	185	4	of	of	ADP
alces-443	185	5	years	year	NOUN
alces-443	185	6	ago	ago	ADV
alces-443	185	7	te	te	PROPN
alces-443	185	8	m	m	VERB
alces-443	185	9	p	p	NOUN
alces-443	185	10	e	e	X
alces-443	185	11	ra	ra	PROPN
alces-443	185	12	tu	tu	X
alces-443	185	13	re	re	PROPN
alces-443	185	14	d	d	NOUN
alces-443	185	15	i	i	PRON
alces-443	185	16	ff	ff	NUM
alces-443	185	17	e	e	X
alces-443	185	18	re	re	VERB
alces-443	185	19	nc	nc	PROPN
alces-443	185	20	e	e	PROPN
alces-443	185	21	(	(	PUNCT
alces-443	185	22	c	c	PROPN
alces-443	185	23	º	º	PROPN
alces-443	185	24	)	)	PUNCT
alces-443	185	25	fig	fig	NOUN
alces-443	185	26	.	.	PUNCT
alces-443	186	1	6	6	NUM
alces-443	186	2	.	.	X
alces-443	186	3	representation	representation	NOUN
alces-443	186	4	of	of	ADP
alces-443	186	5	mean	mean	ADJ
alces-443	186	6	global	global	ADJ
alces-443	186	7	temperatures	temperature	NOUN
alces-443	186	8	during	during	ADP
alces-443	186	9	the	the	DET
alces-443	186	10	last	last	ADJ
alces-443	186	11	160,000	160,000	NUM
alces-443	186	12	years	year	NOUN
alces-443	186	13	relative	relative	ADJ
alces-443	186	14	to	to	PART
alces-443	186	15	mean	mean	VERB
alces-443	186	16	temperature	temperature	NOUN
alces-443	186	17	in	in	ADP
alces-443	186	18	1900	1900	NUM
alces-443	186	19	.	.	PUNCT
alces-443	187	1	negative	negative	ADJ
alces-443	187	2	temperature	temperature	NOUN
alces-443	187	3	differences	difference	NOUN
alces-443	187	4	indicate	indicate	VERB
alces-443	187	5	periods	period	NOUN
alces-443	187	6	colder	cold	ADJ
alces-443	187	7	than	than	ADP
alces-443	187	8	today	today	NOUN
alces-443	187	9	.	.	PUNCT
alces-443	188	1	the	the	DET
alces-443	188	2	2	2	NUM
alces-443	188	3	glacial	glacial	ADJ
alces-443	188	4	maxima	maxima	NOUN
alces-443	188	5	of	of	ADP
alces-443	188	6	the	the	DET
alces-443	188	7	last	last	ADJ
alces-443	188	8	ice	ice	NOUN
alces-443	188	9	age	age	NOUN
alces-443	188	10	are	be	AUX
alces-443	188	11	indicated	indicate	VERB
alces-443	188	12	by	by	ADP
alces-443	188	13	arrows	arrow	NOUN
alces-443	188	14	.	.	PUNCT
alces-443	189	1	estimated	estimate	VERB
alces-443	189	2	dates	date	NOUN
alces-443	189	3	of	of	ADP
alces-443	189	4	moose	moose	NOUN
alces-443	189	5	population	population	NOUN
alces-443	189	6	expansion	expansion	NOUN
alces-443	189	7	are	be	AUX
alces-443	189	8	indicated	indicate	VERB
alces-443	189	9	by	by	ADP
alces-443	189	10	dashed	dash	VERB
alces-443	189	11	lines	line	NOUN
alces-443	189	12	and	and	CCONJ
alces-443	189	13	correspond	correspond	VERB
alces-443	189	14	to	to	ADP
alces-443	189	15	periods	period	NOUN
alces-443	189	16	of	of	ADP
alces-443	189	17	warming	warming	NOUN
alces-443	189	18	associated	associate	VERB
alces-443	189	19	with	with	ADP
alces-443	189	20	glacial	glacial	ADJ
alces-443	189	21	retreat	retreat	NOUN
alces-443	189	22	and	and	CCONJ
alces-443	189	23	northward	northward	ADJ
alces-443	189	24	advance	advance	NOUN
alces-443	189	25	of	of	ADP
alces-443	189	26	the	the	DET
alces-443	189	27	boreal	boreal	NOUN
alces-443	189	28	forest	forest	NOUN
alces-443	189	29	in	in	ADP
alces-443	189	30	eurasia	eurasia	PROPN
alces-443	189	31	.	.	PUNCT
alces-443	190	1	temperature	temperature	NOUN
alces-443	190	2	profile	profile	NOUN
alces-443	190	3	adapted	adapt	VERB
alces-443	190	4	from	from	ADP
alces-443	190	5	http://gcrio.ciesin.org/consequnces/winter96/article1-fig3.html	http://gcrio.ciesin.org/consequnces/winter96/article1-fig3.html	NOUN
alces-443	190	6	.	.	PUNCT
alces-443	191	1	hundertmark	hundertmark	PROPN
alces-443	191	2	and	and	CCONJ
alces-443	191	3	bowyer	bowyer	PROPN
alces-443	191	4	moose	moose	PROPN
alces-443	191	5	genetics	genetics	PROPN
alces-443	191	6	alces	alce	NOUN
alces-443	191	7	vol	vol	NOUN
alces-443	191	8	.	.	PROPN
alces-443	191	9	40	40	NUM
alces-443	191	10	,	,	PUNCT
alces-443	191	11	2004	2004	NUM
alces-443	191	12	112	112	NUM
alces-443	191	13	ity	ity	NOUN
alces-443	191	14	in	in	ADP
alces-443	191	15	composition	composition	NOUN
alces-443	191	16	of	of	ADP
alces-443	191	17	mtdna	mtdna	NOUN
alces-443	191	18	haplotypes	haplotype	NOUN
alces-443	191	19	.	.	PUNCT
alces-443	192	1	as	as	SCONJ
alces-443	192	2	noted	note	VERB
alces-443	192	3	previously	previously	ADV
alces-443	192	4	,	,	PUNCT
alces-443	192	5	north	north	ADJ
alces-443	192	6	american	american	ADJ
alces-443	192	7	subspecies	subspecie	NOUN
alces-443	192	8	are	be	AUX
alces-443	192	9	distinct	distinct	ADJ
alces-443	192	10	from	from	ADP
alces-443	192	11	european	european	ADJ
alces-443	192	12	and	and	CCONJ
alces-443	192	13	asian	asian	ADJ
alces-443	192	14	subspecies	subspecie	NOUN
alces-443	192	15	(	(	PUNCT
alces-443	192	16	fig	fig	NOUN
alces-443	192	17	.	.	PUNCT
alces-443	193	1	2	2	NUM
alces-443	193	2	)	)	PUNCT
alces-443	193	3	and	and	CCONJ
alces-443	193	4	none	none	NOUN
alces-443	193	5	of	of	ADP
alces-443	193	6	the	the	DET
alces-443	193	7	asian	asian	ADJ
alces-443	193	8	haplotypes	haplotype	NOUN
alces-443	193	9	in	in	ADP
alces-443	193	10	the	the	DET
alces-443	193	11	north	north	ADJ
alces-443	193	12	american	american	ADJ
alces-443	193	13	haplogroup	haplogroup	NOUN
alces-443	193	14	were	be	AUX
alces-443	193	15	found	find	VERB
alces-443	193	16	in	in	ADP
alces-443	193	17	the	the	DET
alces-443	193	18	russian	russian	ADJ
alces-443	193	19	far	far	PROPN
alces-443	193	20	east	east	NOUN
alces-443	193	21	.	.	PUNCT
alces-443	194	1	all	all	DET
alces-443	194	2	haplotypes	haplotype	NOUN
alces-443	194	3	found	find	VERB
alces-443	194	4	in	in	ADP
alces-443	194	5	the	the	DET
alces-443	194	6	russian	russian	ADJ
alces-443	194	7	far	far	ADV
alces-443	194	8	east	east	PROPN
alces-443	194	9	(	(	PUNCT
alces-443	194	10	magadan	magadan	PROPN
alces-443	194	11	oblast	oblast	PROPN
alces-443	194	12	)	)	PUNCT
alces-443	194	13	are	be	AUX
alces-443	194	14	restricted	restrict	VERB
alces-443	194	15	to	to	ADP
alces-443	194	16	haplogroup	haplogroup	NOUN
alces-443	194	17	2	2	NUM
alces-443	194	18	(	(	PUNCT
alces-443	194	19	fig	fig	NOUN
alces-443	194	20	.	.	PUNCT
alces-443	195	1	3	3	NUM
alces-443	195	2	)	)	PUNCT
alces-443	195	3	,	,	PUNCT
alces-443	195	4	which	which	PRON
alces-443	195	5	contains	contain	VERB
alces-443	195	6	all	all	DET
alces-443	195	7	european	european	ADJ
alces-443	195	8	haplotypes	haplotype	NOUN
alces-443	195	9	.	.	PUNCT
alces-443	196	1	a	a	DET
alces-443	196	2	likely	likely	ADJ
alces-443	196	3	colonization	colonization	NOUN
alces-443	196	4	scenario	scenario	NOUN
alces-443	196	5	entails	entail	VERB
alces-443	196	6	closely	closely	ADV
alces-443	196	7	related	relate	VERB
alces-443	196	8	moose	moose	NOUN
alces-443	196	9	from	from	ADP
alces-443	196	10	central	central	ADJ
alces-443	196	11	asia	asia	PROPN
alces-443	196	12	colonizing	colonize	VERB
alces-443	196	13	both	both	CCONJ
alces-443	196	14	europe	europe	PROPN
alces-443	196	15	and	and	CCONJ
alces-443	196	16	north	north	PROPN
alces-443	196	17	america	america	PROPN
alces-443	196	18	.	.	PUNCT
alces-443	197	1	lack	lack	NOUN
alces-443	197	2	of	of	ADP
alces-443	197	3	genetic	genetic	ADJ
alces-443	197	4	similarity	similarity	NOUN
alces-443	197	5	between	between	ADP
alces-443	197	6	moose	moose	NOUN
alces-443	197	7	in	in	ADP
alces-443	197	8	alaska	alaska	PROPN
alces-443	197	9	and	and	CCONJ
alces-443	197	10	the	the	DET
alces-443	197	11	russian	russian	ADJ
alces-443	197	12	far	far	ADV
alces-443	197	13	east	east	PROPN
alces-443	197	14	is	be	AUX
alces-443	197	15	inconsistent	inconsistent	ADJ
alces-443	197	16	with	with	ADP
alces-443	197	17	the	the	DET
alces-443	197	18	scenario	scenario	NOUN
alces-443	197	19	proposed	propose	VERB
alces-443	197	20	by	by	ADP
alces-443	197	21	hundertmark	hundertmark	PROPN
alces-443	197	22	et	et	PROPN
alces-443	197	23	al	al	PROPN
alces-443	197	24	.	.	PROPN
alces-443	197	25	(	(	PUNCT
alces-443	197	26	2002b	2002b	NUM
alces-443	197	27	)	)	PUNCT
alces-443	197	28	concerning	concern	VERB
alces-443	197	29	a	a	DET
alces-443	197	30	colonizing	colonizing	ADJ
alces-443	197	31	wave	wave	NOUN
alces-443	197	32	of	of	ADP
alces-443	197	33	moose	moose	NOUN
alces-443	197	34	traveling	travel	VERB
alces-443	197	35	from	from	ADP
alces-443	197	36	asia	asia	PROPN
alces-443	197	37	to	to	ADP
alces-443	197	38	north	north	PROPN
alces-443	197	39	america	america	PROPN
alces-443	197	40	through	through	ADP
alces-443	197	41	beringia	beringia	NOUN
alces-443	197	42	.	.	PUNCT
alces-443	198	1	a	a	DET
alces-443	198	2	single	single	ADJ
alces-443	198	3	wave	wave	NOUN
alces-443	198	4	of	of	ADP
alces-443	198	5	moose	moose	NOUN
alces-443	198	6	colonizing	colonize	VERB
alces-443	198	7	north	north	PROPN
alces-443	198	8	america	america	PROPN
alces-443	198	9	through	through	ADP
alces-443	198	10	beringia	beringia	NOUN
alces-443	198	11	would	would	AUX
alces-443	198	12	have	have	AUX
alces-443	198	13	left	leave	VERB
alces-443	198	14	genetically	genetically	ADV
alces-443	198	15	similar	similar	ADJ
alces-443	198	16	populations	population	NOUN
alces-443	198	17	on	on	ADP
alces-443	198	18	either	either	DET
alces-443	198	19	side	side	NOUN
alces-443	198	20	of	of	ADP
alces-443	198	21	the	the	DET
alces-443	198	22	bering	bering	NOUN
alces-443	198	23	strait	strait	NOUN
alces-443	198	24	after	after	SCONJ
alces-443	198	25	the	the	DET
alces-443	198	26	bering	bere	VERB
alces-443	198	27	land	land	NOUN
alces-443	198	28	bridge	bridge	NOUN
alces-443	198	29	flooded	flood	VERB
alces-443	198	30	.	.	PUNCT
alces-443	199	1	yet	yet	ADV
alces-443	199	2	,	,	PUNCT
alces-443	199	3	ages	age	NOUN
alces-443	199	4	of	of	ADP
alces-443	199	5	subfossil	subfossil	NOUN
alces-443	199	6	remains	remain	NOUN
alces-443	199	7	of	of	ADP
alces-443	199	8	moose	moose	NOUN
alces-443	199	9	from	from	ADP
alces-443	199	10	north	north	PROPN
alces-443	199	11	america	america	PROPN
alces-443	199	12	indicate	indicate	VERB
alces-443	199	13	clearly	clearly	ADV
alces-443	199	14	that	that	SCONJ
alces-443	199	15	a.	a.	NOUN
alces-443	199	16	alces	alce	NOUN
alces-443	199	17	was	be	AUX
alces-443	199	18	present	present	ADJ
alces-443	199	19	in	in	ADP
alces-443	199	20	alaska	alaska	PROPN
alces-443	199	21	prior	prior	ADV
alces-443	199	22	to	to	ADP
alces-443	199	23	anywhere	anywhere	ADV
alces-443	199	24	else	else	ADV
alces-443	199	25	on	on	ADP
alces-443	199	26	the	the	DET
alces-443	199	27	continent	continent	NOUN
alces-443	199	28	(	(	PUNCT
alces-443	199	29	guthrie	guthrie	PROPN
alces-443	199	30	1990	1990	NUM
alces-443	199	31	,	,	PUNCT
alces-443	199	32	hundertmark	hundertmark	NOUN
alces-443	199	33	et	et	PROPN
alces-443	199	34	al	al	PROPN
alces-443	199	35	.	.	PROPN
alces-443	199	36	2003	2003	NUM
alces-443	199	37	)	)	PUNCT
alces-443	199	38	,	,	PUNCT
alces-443	199	39	unequivocally	unequivocally	ADV
alces-443	199	40	supporting	support	VERB
alces-443	199	41	an	an	DET
alces-443	199	42	entry	entry	NOUN
alces-443	199	43	through	through	ADP
alces-443	199	44	beringia	beringia	NOUN
alces-443	199	45	.	.	PUNCT
alces-443	200	1	one	one	NUM
alces-443	200	2	hypothesis	hypothesis	NOUN
alces-443	200	3	that	that	PRON
alces-443	200	4	accounts	account	VERB
alces-443	200	5	for	for	ADP
alces-443	200	6	this	this	DET
alces-443	200	7	seeming	seem	VERB
alces-443	200	8	paradox	paradox	NOUN
alces-443	200	9	is	be	AUX
alces-443	200	10	that	that	SCONJ
alces-443	200	11	1	1	NUM
alces-443	200	12	or	or	CCONJ
alces-443	200	13	both	both	PRON
alces-443	200	14	of	of	ADP
alces-443	200	15	those	those	DET
alces-443	200	16	populations	population	NOUN
alces-443	200	17	underwent	undergo	VERB
alces-443	200	18	population	population	NOUN
alces-443	200	19	bottlenecks	bottleneck	NOUN
alces-443	200	20	shortly	shortly	ADV
alces-443	200	21	after	after	ADP
alces-443	200	22	the	the	DET
alces-443	200	23	colonization	colonization	NOUN
alces-443	200	24	of	of	ADP
alces-443	200	25	north	north	PROPN
alces-443	200	26	america	america	PROPN
alces-443	200	27	,	,	PUNCT
alces-443	200	28	and	and	CCONJ
alces-443	200	29	have	have	AUX
alces-443	200	30	only	only	ADV
alces-443	200	31	recently	recently	ADV
alces-443	200	32	reestablished	reestablish	VERB
alces-443	200	33	populations	population	NOUN
alces-443	200	34	in	in	ADP
alces-443	200	35	those	those	DET
alces-443	200	36	areas	area	NOUN
alces-443	200	37	,	,	PUNCT
alces-443	200	38	leading	lead	VERB
alces-443	200	39	to	to	ADP
alces-443	200	40	the	the	DET
alces-443	200	41	presence	presence	NOUN
alces-443	200	42	of	of	ADP
alces-443	200	43	different	different	ADJ
alces-443	200	44	genetic	genetic	ADJ
alces-443	200	45	lineages	lineage	NOUN
alces-443	200	46	in	in	ADP
alces-443	200	47	each	each	PRON
alces-443	200	48	.	.	PUNCT
alces-443	201	1	moose	moose	PROPN
alces-443	201	2	in	in	ADP
alces-443	201	3	the	the	DET
alces-443	201	4	russian	russian	ADJ
alces-443	201	5	far	far	PROPN
alces-443	201	6	east	east	NOUN
alces-443	201	7	show	show	NOUN
alces-443	201	8	evidence	evidence	NOUN
alces-443	201	9	of	of	ADP
alces-443	201	10	an	an	DET
alces-443	201	11	expansion	expansion	NOUN
alces-443	201	12	approximately	approximately	ADV
alces-443	201	13	1,200	1,200	NUM
alces-443	201	14	years	year	NOUN
alces-443	201	15	ago	ago	ADV
alces-443	201	16	(	(	PUNCT
alces-443	201	17	hundertmark	hundertmark	NOUN
alces-443	201	18	et	et	PROPN
alces-443	201	19	al	al	PROPN
alces-443	201	20	.	.	PROPN
alces-443	201	21	2002b	2002b	NUM
alces-443	201	22	)	)	PUNCT
alces-443	201	23	and	and	CCONJ
alces-443	201	24	continue	continue	VERB
alces-443	201	25	to	to	PART
alces-443	201	26	expand	expand	VERB
alces-443	201	27	their	their	PRON
alces-443	201	28	range	range	NOUN
alces-443	201	29	toward	toward	ADP
alces-443	201	30	the	the	DET
alces-443	201	31	bering	bering	NOUN
alces-443	201	32	sea	sea	NOUN
alces-443	201	33	(	(	PUNCT
alces-443	201	34	zheleznov	zheleznov	PROPN
alces-443	201	35	1993	1993	NUM
alces-443	201	36	)	)	PUNCT
alces-443	201	37	.	.	PUNCT
alces-443	202	1	similarly	similarly	ADV
alces-443	202	2	,	,	PUNCT
alces-443	202	3	alaskan	alaskan	ADJ
alces-443	202	4	moose	moose	NOUN
alces-443	202	5	show	show	VERB
alces-443	202	6	a	a	DET
alces-443	202	7	surprising	surprising	ADJ
alces-443	202	8	lack	lack	NOUN
alces-443	202	9	of	of	ADP
alces-443	202	10	mitochondrial	mitochondrial	ADJ
alces-443	202	11	diversity	diversity	NOUN
alces-443	202	12	compared	compare	VERB
alces-443	202	13	with	with	ADP
alces-443	202	14	moose	moose	NOUN
alces-443	202	15	elsewhere	elsewhere	ADV
alces-443	202	16	on	on	ADP
alces-443	202	17	the	the	DET
alces-443	202	18	continent	continent	NOUN
alces-443	202	19	(	(	PUNCT
alces-443	202	20	hundertmark	hundertmark	NOUN
alces-443	202	21	et	et	PROPN
alces-443	202	22	al	al	PROPN
alces-443	202	23	.	.	PROPN
alces-443	202	24	2003	2003	NUM
alces-443	202	25	)	)	PUNCT
alces-443	202	26	,	,	PUNCT
alces-443	202	27	which	which	PRON
alces-443	202	28	is	be	AUX
alces-443	202	29	tion	tion	NOUN
alces-443	202	30	of	of	ADP
alces-443	202	31	a	a	DET
alces-443	202	32	recent	recent	ADJ
alces-443	202	33	common	common	ADJ
alces-443	202	34	ancestry	ancestry	NOUN
alces-443	202	35	,	,	PUNCT
alces-443	202	36	consistent	consistent	ADJ
alces-443	202	37	with	with	ADP
alces-443	202	38	colonization	colonization	NOUN
alces-443	202	39	of	of	ADP
alces-443	202	40	the	the	DET
alces-443	202	41	continent	continent	NOUN
alces-443	202	42	after	after	ADP
alces-443	202	43	the	the	DET
alces-443	202	44	retreat	retreat	NOUN
alces-443	202	45	of	of	ADP
alces-443	202	46	ice	ice	NOUN
alces-443	202	47	sheets	sheet	NOUN
alces-443	202	48	from	from	ADP
alces-443	202	49	the	the	DET
alces-443	202	50	last	last	ADJ
alces-443	202	51	ice	ice	NOUN
alces-443	202	52	age	age	NOUN
alces-443	202	53	(	(	PUNCT
alces-443	202	54	cronin	cronin	PROPN
alces-443	202	55	1992	1992	NUM
alces-443	202	56	)	)	PUNCT
alces-443	202	57	.	.	PUNCT
alces-443	203	1	similarly	similarly	ADV
alces-443	203	2	,	,	PUNCT
alces-443	203	3	no	no	DET
alces-443	203	4	variation	variation	NOUN
alces-443	203	5	was	be	AUX
alces-443	203	6	detected	detect	VERB
alces-443	203	7	within	within	ADP
alces-443	203	8	a	a	DET
alces-443	203	9	fragment	fragment	NOUN
alces-443	203	10	of	of	ADP
alces-443	203	11	cytochrome	cytochrome	PROPN
alces-443	203	12	b	b	PROPN
alces-443	203	13	in	in	ADP
alces-443	203	14	north	north	ADJ
alces-443	203	15	american	american	ADJ
alces-443	203	16	moose	moose	NOUN
alces-443	203	17	compared	compare	VERB
alces-443	203	18	with	with	ADP
alces-443	203	19	slight	slight	ADJ
alces-443	203	20	variation	variation	NOUN
alces-443	203	21	in	in	ADP
alces-443	203	22	eurasia	eurasia	PROPN
alces-443	203	23	(	(	PUNCT
alces-443	203	24	hundertmark	hundertmark	PROPN
alces-443	203	25	et	et	PROPN
alces-443	203	26	al	al	PROPN
alces-443	203	27	.	.	PROPN
alces-443	203	28	2002a	2002a	NUM
alces-443	203	29	)	)	PUNCT
alces-443	203	30	.	.	PUNCT
alces-443	204	1	moose	moose	NOUN
alces-443	204	2	populations	population	NOUN
alces-443	204	3	in	in	ADP
alces-443	204	4	north	north	PROPN
alces-443	204	5	america	america	PROPN
alces-443	204	6	were	be	AUX
alces-443	204	7	established	establish	VERB
alces-443	204	8	as	as	ADP
alces-443	204	9	a	a	DET
alces-443	204	10	result	result	NOUN
alces-443	204	11	of	of	ADP
alces-443	204	12	a	a	DET
alces-443	204	13	single	single	ADJ
alces-443	204	14	entry	entry	NOUN
alces-443	204	15	into	into	ADP
alces-443	204	16	the	the	DET
alces-443	204	17	continent	continent	NOUN
alces-443	204	18	,	,	PUNCT
alces-443	204	19	and	and	CCONJ
alces-443	204	20	that	that	DET
alces-443	204	21	entry	entry	NOUN
alces-443	204	22	occurred	occur	VERB
alces-443	204	23	during	during	ADP
alces-443	204	24	the	the	DET
alces-443	204	25	population	population	NOUN
alces-443	204	26	expansion	expansion	NOUN
alces-443	204	27	of	of	ADP
alces-443	204	28	moose	moose	NOUN
alces-443	204	29	14,000	14,000	NUM
alces-443	204	30	years	year	NOUN
alces-443	204	31	ago	ago	ADV
alces-443	204	32	at	at	ADP
alces-443	204	33	the	the	DET
alces-443	204	34	end	end	NOUN
alces-443	204	35	of	of	ADP
alces-443	204	36	the	the	DET
alces-443	204	37	last	last	ADJ
alces-443	204	38	ice	ice	NOUN
alces-443	204	39	age	age	NOUN
alces-443	204	40	,	,	PUNCT
alces-443	204	41	just	just	ADV
alces-443	204	42	before	before	ADP
alces-443	204	43	the	the	DET
alces-443	204	44	closure	closure	NOUN
alces-443	204	45	of	of	ADP
alces-443	204	46	the	the	DET
alces-443	204	47	bering	bering	NOUN
alces-443	204	48	land	land	NOUN
alces-443	204	49	bridge	bridge	PROPN
alces-443	204	50	(	(	PUNCT
alces-443	204	51	guthrie	guthrie	PROPN
alces-443	204	52	1995	1995	NUM
alces-443	204	53	,	,	PUNCT
alces-443	204	54	hundertmark	hundertmark	NOUN
alces-443	204	55	et	et	PROPN
alces-443	204	56	al	al	PROPN
alces-443	204	57	.	.	PROPN
alces-443	204	58	2002b	2002b	NUM
alces-443	204	59	)	)	PUNCT
alces-443	204	60	.	.	PUNCT
alces-443	205	1	a	a	DET
alces-443	205	2	recent	recent	ADJ
alces-443	205	3	entry	entry	NOUN
alces-443	205	4	into	into	ADP
alces-443	205	5	north	north	PROPN
alces-443	205	6	america	america	PROPN
alces-443	205	7	is	be	AUX
alces-443	205	8	the	the	DET
alces-443	205	9	only	only	ADJ
alces-443	205	10	conclusion	conclusion	NOUN
alces-443	205	11	that	that	PRON
alces-443	205	12	is	be	AUX
alces-443	205	13	consistent	consistent	ADJ
alces-443	205	14	with	with	ADP
alces-443	205	15	limited	limited	ADJ
alces-443	205	16	variation	variation	NOUN
alces-443	205	17	in	in	ADP
alces-443	205	18	the	the	DET
alces-443	205	19	mtdna	mtdna	PROPN
alces-443	205	20	control	control	PROPN
alces-443	205	21	region	region	NOUN
alces-443	205	22	both	both	CCONJ
alces-443	205	23	within	within	ADP
alces-443	205	24	north	north	PROPN
alces-443	205	25	america	america	PROPN
alces-443	205	26	and	and	CCONJ
alces-443	205	27	between	between	ADP
alces-443	205	28	north	north	PROPN
alces-443	205	29	america	america	PROPN
alces-443	205	30	and	and	CCONJ
alces-443	205	31	eurasia	eurasia	PROPN
alces-443	205	32	.	.	PUNCT
alces-443	206	1	if	if	SCONJ
alces-443	206	2	moose	moose	NOUN
alces-443	206	3	had	have	AUX
alces-443	206	4	existed	exist	VERB
alces-443	206	5	in	in	ADP
alces-443	206	6	4	4	NUM
alces-443	206	7	separate	separate	ADJ
alces-443	206	8	refugia	refugia	NOUN
alces-443	206	9	in	in	ADP
alces-443	206	10	north	north	PROPN
alces-443	206	11	america	america	PROPN
alces-443	206	12	during	during	ADP
alces-443	206	13	the	the	DET
alces-443	206	14	last	last	ADJ
alces-443	206	15	ice	ice	NOUN
alces-443	206	16	age	age	NOUN
alces-443	206	17	,	,	PUNCT
alces-443	206	18	as	as	SCONJ
alces-443	206	19	suggested	suggest	VERB
alces-443	206	20	by	by	ADP
alces-443	206	21	peterson	peterson	PROPN
alces-443	206	22	(	(	PUNCT
alces-443	206	23	1955	1955	NUM
alces-443	206	24	)	)	PUNCT
alces-443	206	25	,	,	PUNCT
alces-443	206	26	or	or	CCONJ
alces-443	206	27	had	have	AUX
alces-443	206	28	entered	enter	VERB
alces-443	206	29	north	north	PROPN
alces-443	206	30	america	america	PROPN
alces-443	206	31	from	from	ADP
alces-443	206	32	asia	asia	PROPN
alces-443	206	33	more	more	ADV
alces-443	206	34	than	than	ADP
alces-443	206	35	once	once	ADV
alces-443	206	36	,	,	PUNCT
alces-443	206	37	a	a	DET
alces-443	206	38	more	more	ADV
alces-443	206	39	distant	distant	ADJ
alces-443	206	40	common	common	ADJ
alces-443	206	41	ancestor	ancestor	NOUN
alces-443	206	42	would	would	AUX
alces-443	206	43	be	be	AUX
alces-443	206	44	indicated	indicate	VERB
alces-443	206	45	.	.	PUNCT
alces-443	207	1	the	the	DET
alces-443	207	2	timing	timing	NOUN
alces-443	207	3	of	of	ADP
alces-443	207	4	the	the	DET
alces-443	207	5	entry	entry	NOUN
alces-443	207	6	is	be	AUX
alces-443	207	7	supported	support	VERB
alces-443	207	8	not	not	PART
alces-443	207	9	only	only	ADV
alces-443	207	10	by	by	ADP
alces-443	207	11	genetic	genetic	ADJ
alces-443	207	12	data	datum	NOUN
alces-443	207	13	but	but	CCONJ
alces-443	207	14	also	also	ADV
alces-443	207	15	by	by	ADP
alces-443	207	16	the	the	DET
alces-443	207	17	distribution	distribution	NOUN
alces-443	207	18	of	of	ADP
alces-443	207	19	suitable	suitable	ADJ
alces-443	207	20	moose	moose	NOUN
alces-443	207	21	habitat	habitat	NOUN
alces-443	207	22	at	at	ADP
alces-443	207	23	the	the	DET
alces-443	207	24	end	end	NOUN
alces-443	207	25	of	of	ADP
alces-443	207	26	the	the	DET
alces-443	207	27	last	last	ADJ
alces-443	207	28	ice	ice	NOUN
alces-443	207	29	age	age	NOUN
alces-443	207	30	.	.	PUNCT
alces-443	208	1	the	the	DET
alces-443	208	2	cold	cold	ADJ
alces-443	208	3	,	,	PUNCT
alces-443	208	4	dry	dry	ADJ
alces-443	208	5	grassland	grassland	NOUN
alces-443	208	6	habitat	habitat	NOUN
alces-443	208	7	that	that	PRON
alces-443	208	8	prevailed	prevail	VERB
alces-443	208	9	in	in	ADP
alces-443	208	10	beringia	beringia	NOUN
alces-443	208	11	for	for	ADP
alces-443	208	12	most	most	ADJ
alces-443	208	13	of	of	ADP
alces-443	208	14	the	the	DET
alces-443	208	15	last	last	ADJ
alces-443	208	16	ice	ice	NOUN
alces-443	208	17	age	age	NOUN
alces-443	208	18	was	be	AUX
alces-443	208	19	unsuitable	unsuitable	ADJ
alces-443	208	20	for	for	ADP
alces-443	208	21	moose	moose	NOUN
alces-443	208	22	and	and	CCONJ
alces-443	208	23	was	be	AUX
alces-443	208	24	replaced	replace	VERB
alces-443	208	25	by	by	ADP
alces-443	208	26	boreal	boreal	ADJ
alces-443	208	27	forest	forest	NOUN
alces-443	208	28	only	only	ADV
alces-443	208	29	within	within	ADP
alces-443	208	30	the	the	DET
alces-443	208	31	last	last	ADJ
alces-443	208	32	14,000	14,000	NUM
alces-443	208	33	years	year	NOUN
alces-443	208	34	(	(	PUNCT
alces-443	208	35	guthrie	guthrie	PROPN
alces-443	208	36	1995	1995	NUM
alces-443	208	37	)	)	PUNCT
alces-443	208	38	.	.	PUNCT
alces-443	209	1	evidence	evidence	NOUN
alces-443	209	2	from	from	ADP
alces-443	209	3	mtdna	mtdna	NOUN
alces-443	209	4	variation	variation	NOUN
alces-443	209	5	also	also	ADV
alces-443	209	6	indicates	indicate	VERB
alces-443	209	7	that	that	SCONJ
alces-443	209	8	north	north	PROPN
alces-443	209	9	american	american	ADJ
alces-443	209	10	moose	moose	NOUN
alces-443	209	11	did	do	AUX
alces-443	209	12	not	not	PART
alces-443	209	13	originate	originate	VERB
alces-443	209	14	in	in	ADP
alces-443	209	15	beringia	beringia	NOUN
alces-443	209	16	,	,	PUNCT
alces-443	209	17	as	as	SCONJ
alces-443	209	18	some	some	PRON
alces-443	209	19	have	have	AUX
alces-443	209	20	speculated	speculate	VERB
alces-443	209	21	(	(	PUNCT
alces-443	209	22	cronin	cronin	PROPN
alces-443	209	23	1992	1992	NUM
alces-443	209	24	,	,	PUNCT
alces-443	209	25	geist	geist	NOUN
alces-443	209	26	1998	1998	NUM
alces-443	209	27	)	)	PUNCT
alces-443	209	28	,	,	PUNCT
alces-443	209	29	or	or	CCONJ
alces-443	209	30	recolonize	recolonize	VERB
alces-443	209	31	the	the	DET
alces-443	209	32	russian	russian	ADJ
alces-443	209	33	far	far	ADV
alces-443	209	34	east	east	ADV
alces-443	209	35	from	from	ADP
alces-443	209	36	north	north	PROPN
alces-443	209	37	america	america	PROPN
alces-443	209	38	(	(	PUNCT
alces-443	209	39	coady	coady	PROPN
alces-443	209	40	1982	1982	NUM
alces-443	209	41	)	)	PUNCT
alces-443	209	42	.	.	PUNCT
alces-443	210	1	moose	moose	NOUN
alces-443	210	2	in	in	ADP
alces-443	210	3	north	north	PROPN
alces-443	210	4	america	america	PROPN
alces-443	210	5	are	be	AUX
alces-443	210	6	not	not	PART
alces-443	210	7	closely	closely	ADV
alces-443	210	8	related	relate	VERB
alces-443	210	9	to	to	ADP
alces-443	210	10	moose	moose	VERB
alces-443	210	11	on	on	ADP
alces-443	210	12	the	the	DET
alces-443	210	13	western	western	ADJ
alces-443	210	14	side	side	NOUN
alces-443	210	15	of	of	ADP
alces-443	210	16	the	the	DET
alces-443	210	17	bering	bering	NOUN
alces-443	210	18	strait	strait	NOUN
alces-443	210	19	(	(	PUNCT
alces-443	210	20	russian	russian	ADJ
alces-443	210	21	far	far	PROPN
alces-443	210	22	east	east	NOUN
alces-443	210	23	)	)	PUNCT
alces-443	210	24	.	.	PUNCT
alces-443	211	1	if	if	SCONJ
alces-443	211	2	those	those	DET
alces-443	211	3	moose	moose	NOUN
alces-443	211	4	were	be	AUX
alces-443	211	5	once	once	ADV
alces-443	211	6	part	part	NOUN
alces-443	211	7	of	of	ADP
alces-443	211	8	the	the	DET
alces-443	211	9	same	same	ADJ
alces-443	211	10	population	population	NOUN
alces-443	211	11	recently	recently	ADV
alces-443	211	12	separated	separate	VERB
alces-443	211	13	by	by	ADP
alces-443	211	14	the	the	DET
alces-443	211	15	flooding	flooding	NOUN
alces-443	211	16	of	of	ADP
alces-443	211	17	the	the	DET
alces-443	211	18	bering	bering	NOUN
alces-443	211	19	land	land	NOUN
alces-443	211	20	bridge	bridge	NOUN
alces-443	211	21	,	,	PUNCT
alces-443	211	22	we	we	PRON
alces-443	211	23	would	would	AUX
alces-443	211	24	still	still	ADV
alces-443	211	25	expect	expect	VERB
alces-443	211	26	to	to	PART
alces-443	211	27	find	find	VERB
alces-443	211	28	similaralces	similaralce	NOUN
alces-443	211	29	vol	vol	NOUN
alces-443	211	30	.	.	PROPN
alces-443	211	31	40	40	NUM
alces-443	211	32	,	,	PUNCT
alces-443	211	33	2004	2004	NUM
alces-443	211	34	moose	moose	NOUN
alces-443	211	35	genetics	genetic	NOUN
alces-443	211	36	hundertmark	hundertmark	NOUN
alces-443	211	37	and	and	CCONJ
alces-443	211	38	bowyer	bowyer	PROPN
alces-443	211	39	113	113	NUM
alces-443	211	40	indicative	indicative	ADJ
alces-443	211	41	of	of	ADP
alces-443	211	42	a	a	DET
alces-443	211	43	bottleneck	bottleneck	NOUN
alces-443	211	44	notwithstanding	notwithstanding	ADP
alces-443	211	45	the	the	DET
alces-443	211	46	moderate	moderate	ADJ
alces-443	211	47	levels	level	NOUN
alces-443	211	48	of	of	ADP
alces-443	211	49	allozyme	allozyme	NOUN
alces-443	211	50	diversity	diversity	NOUN
alces-443	211	51	reported	report	VERB
alces-443	211	52	for	for	ADP
alces-443	211	53	alaskan	alaskan	ADJ
alces-443	211	54	moose	moose	NOUN
alces-443	211	55	by	by	ADP
alces-443	211	56	hundertmark	hundertmark	PROPN
alces-443	211	57	et	et	PROPN
alces-443	211	58	al	al	PROPN
alces-443	211	59	.	.	PROPN
alces-443	212	1	(	(	PUNCT
alces-443	212	2	1992	1992	NUM
alces-443	212	3	)	)	PUNCT
alces-443	212	4	.	.	PUNCT
alces-443	213	1	diversity	diversity	NOUN
alces-443	213	2	of	of	ADP
alces-443	213	3	mtdna	mtdna	PROPN
alces-443	213	4	can	can	AUX
alces-443	213	5	be	be	AUX
alces-443	213	6	reduced	reduce	VERB
alces-443	213	7	to	to	ADP
alces-443	213	8	a	a	DET
alces-443	213	9	much	much	ADV
alces-443	213	10	greater	great	ADJ
alces-443	213	11	degree	degree	NOUN
alces-443	213	12	by	by	ADP
alces-443	213	13	a	a	DET
alces-443	213	14	bottleneck	bottleneck	NOUN
alces-443	213	15	than	than	ADP
alces-443	213	16	diversity	diversity	NOUN
alces-443	213	17	of	of	ADP
alces-443	213	18	nuclear	nuclear	ADJ
alces-443	213	19	dna	dna	PROPN
alces-443	213	20	because	because	SCONJ
alces-443	213	21	mtdna	mtdna	PROPN
alces-443	213	22	is	be	AUX
alces-443	213	23	4	4	NUM
alces-443	213	24	times	time	NOUN
alces-443	213	25	more	more	ADV
alces-443	213	26	sensitive	sensitive	ADJ
alces-443	213	27	to	to	ADP
alces-443	213	28	genetic	genetic	ADJ
alces-443	213	29	drift	drift	NOUN
alces-443	213	30	due	due	ADP
alces-443	213	31	to	to	ADP
alces-443	213	32	its	its	PRON
alces-443	213	33	haploid	haploid	NOUN
alces-443	213	34	,	,	PUNCT
alces-443	213	35	uniparental	uniparental	ADJ
alces-443	213	36	mode	mode	NOUN
alces-443	213	37	of	of	ADP
alces-443	213	38	inheritance	inheritance	NOUN
alces-443	213	39	(	(	PUNCT
alces-443	213	40	birky	birky	VERB
alces-443	213	41	et	et	PROPN
alces-443	213	42	al	al	PROPN
alces-443	213	43	.	.	PROPN
alces-443	213	44	1983	1983	NUM
alces-443	213	45	)	)	PUNCT
alces-443	213	46	.	.	PUNCT
alces-443	214	1	simultaneous	simultaneous	ADJ
alces-443	214	2	bottlenecks	bottleneck	NOUN
alces-443	214	3	in	in	ADP
alces-443	214	4	moose	moose	NOUN
alces-443	214	5	from	from	ADP
alces-443	214	6	both	both	DET
alces-443	214	7	sides	side	NOUN
alces-443	214	8	of	of	ADP
alces-443	214	9	the	the	DET
alces-443	214	10	bering	bering	NOUN
alces-443	214	11	strait	strait	NOUN
alces-443	214	12	suggest	suggest	VERB
alces-443	214	13	a	a	DET
alces-443	214	14	widespread	widespread	ADJ
alces-443	214	15	causal	causal	ADJ
alces-443	214	16	factor	factor	NOUN
alces-443	214	17	.	.	PUNCT
alces-443	215	1	recent	recent	ADJ
alces-443	215	2	studies	study	NOUN
alces-443	215	3	have	have	AUX
alces-443	215	4	found	find	VERB
alces-443	215	5	evidence	evidence	NOUN
alces-443	215	6	of	of	ADP
alces-443	215	7	significant	significant	ADJ
alces-443	215	8	biotic	biotic	ADJ
alces-443	215	9	effects	effect	NOUN
alces-443	215	10	of	of	ADP
alces-443	215	11	climate	climate	NOUN
alces-443	215	12	reversals	reversal	NOUN
alces-443	215	13	in	in	ADP
alces-443	215	14	beringia	beringia	NOUN
alces-443	215	15	after	after	ADP
alces-443	215	16	flooding	flooding	NOUN
alces-443	215	17	of	of	ADP
alces-443	215	18	the	the	DET
alces-443	215	19	bering	bering	NOUN
alces-443	215	20	land	land	NOUN
alces-443	215	21	bridge	bridge	PROPN
alces-443	215	22	(	(	PUNCT
alces-443	215	23	elias	elias	PROPN
alces-443	215	24	2000	2000	NUM
alces-443	215	25	,	,	PUNCT
alces-443	215	26	mason	mason	PROPN
alces-443	215	27	et	et	PROPN
alces-443	215	28	al	al	PROPN
alces-443	215	29	.	.	PROPN
alces-443	215	30	2001	2001	NUM
alces-443	215	31	,	,	PUNCT
alces-443	215	32	anderson	anderson	PROPN
alces-443	215	33	et	et	PROPN
alces-443	215	34	al	al	PROPN
alces-443	215	35	.	.	PROPN
alces-443	215	36	2002	2002	NUM
alces-443	215	37	)	)	PUNCT
alces-443	215	38	.	.	PUNCT
alces-443	216	1	those	those	DET
alces-443	216	2	effects	effect	NOUN
alces-443	216	3	offer	offer	VERB
alces-443	216	4	an	an	DET
alces-443	216	5	intriguing	intriguing	ADJ
alces-443	216	6	mechanism	mechanism	NOUN
alces-443	216	7	for	for	ADP
alces-443	216	8	bottlenecks	bottleneck	NOUN
alces-443	216	9	in	in	ADP
alces-443	216	10	beringian	beringian	ADJ
alces-443	216	11	moose	moose	NOUN
alces-443	216	12	populations	population	NOUN
alces-443	216	13	.	.	PUNCT
alces-443	217	1	the	the	DET
alces-443	217	2	greatest	great	ADJ
alces-443	217	3	variation	variation	NOUN
alces-443	217	4	in	in	ADP
alces-443	217	5	mtdna	mtdna	PROPN
alces-443	217	6	in	in	ADP
alces-443	217	7	north	north	PROPN
alces-443	217	8	american	american	PROPN
alces-443	217	9	moose	moose	PROPN
alces-443	217	10	occurs	occur	VERB
alces-443	217	11	within	within	ADP
alces-443	217	12	the	the	DET
alces-443	217	13	range	range	NOUN
alces-443	217	14	of	of	ADP
alces-443	217	15	a.	a.	PROPN
alces-443	217	16	a.	a.	NOUN
alces-443	217	17	andersoni	andersoni	PROPN
alces-443	217	18	(	(	PUNCT
alces-443	217	19	hundertmark	hundertmark	NOUN
alces-443	217	20	et	et	PROPN
alces-443	217	21	al	al	PROPN
alces-443	217	22	.	.	PROPN
alces-443	217	23	2003	2003	NUM
alces-443	217	24	)	)	PUNCT
alces-443	217	25	.	.	PUNCT
alces-443	218	1	alces	alce	NOUN
alces-443	218	2	a.	a.	PROPN
alces-443	218	3	shirasi	shirasi	PROPN
alces-443	218	4	from	from	ADP
alces-443	218	5	colorado	colorado	PROPN
alces-443	218	6	exhibited	exhibit	VERB
alces-443	218	7	no	no	DET
alces-443	218	8	diversity	diversity	NOUN
alces-443	218	9	and	and	CCONJ
alces-443	218	10	a.	a.	NOUN
alces-443	218	11	a.	a.	NOUN
alces-443	218	12	gigas	gigas	PROPN
alces-443	218	13	and	and	CCONJ
alces-443	218	14	a.	a.	NOUN
alces-443	218	15	a.	a.	PROPN
alces-443	218	16	americana	americana	PROPN
alces-443	218	17	exhibited	exhibit	VERB
alces-443	218	18	very	very	ADV
alces-443	218	19	little	little	ADJ
alces-443	218	20	diversity	diversity	NOUN
alces-443	218	21	.	.	PUNCT
alces-443	219	1	if	if	SCONJ
alces-443	219	2	the	the	DET
alces-443	219	3	paucity	paucity	NOUN
alces-443	219	4	of	of	ADP
alces-443	219	5	mitochondrial	mitochondrial	ADJ
alces-443	219	6	diversity	diversity	NOUN
alces-443	219	7	in	in	ADP
alces-443	219	8	alaska	alaska	PROPN
alces-443	219	9	is	be	AUX
alces-443	219	10	due	due	ADJ
alces-443	219	11	to	to	ADP
alces-443	219	12	a	a	DET
alces-443	219	13	bottleneck	bottleneck	NOUN
alces-443	219	14	and	and	CCONJ
alces-443	219	15	recent	recent	ADJ
alces-443	219	16	expansion	expansion	NOUN
alces-443	219	17	,	,	PUNCT
alces-443	219	18	those	those	DET
alces-443	219	19	data	datum	NOUN
alces-443	219	20	would	would	AUX
alces-443	219	21	be	be	AUX
alces-443	219	22	consistent	consistent	ADJ
alces-443	219	23	with	with	ADP
alces-443	219	24	the	the	DET
alces-443	219	25	serial	serial	ADJ
alces-443	219	26	-	-	PUNCT
alces-443	219	27	founder	founder	NOUN
alces-443	219	28	-	-	PUNCT
alces-443	219	29	events	event	NOUN
alces-443	219	30	hypothesis	hypothesis	NOUN
alces-443	219	31	of	of	ADP
alces-443	219	32	north	north	PROPN
alces-443	219	33	american	american	ADJ
alces-443	219	34	colonization	colonization	NOUN
alces-443	219	35	(	(	PUNCT
alces-443	219	36	hundertmark	hundertmark	NOUN
alces-443	219	37	et	et	PROPN
alces-443	219	38	al	al	PROPN
alces-443	219	39	.	.	PROPN
alces-443	219	40	1992	1992	NUM
alces-443	219	41	)	)	PUNCT
alces-443	219	42	.	.	PUNCT
alces-443	220	1	the	the	DET
alces-443	220	2	pattern	pattern	NOUN
alces-443	220	3	of	of	ADP
alces-443	220	4	colonization	colonization	NOUN
alces-443	220	5	of	of	ADP
alces-443	220	6	north	north	PROPN
alces-443	220	7	america	america	PROPN
alces-443	220	8	undoubtedly	undoubtedly	ADV
alces-443	220	9	was	be	AUX
alces-443	220	10	influenced	influence	VERB
alces-443	220	11	by	by	ADP
alces-443	220	12	the	the	DET
alces-443	220	13	retreating	retreat	VERB
alces-443	220	14	glaciers	glacier	NOUN
alces-443	220	15	and	and	CCONJ
alces-443	220	16	may	may	AUX
alces-443	220	17	have	have	AUX
alces-443	220	18	had	have	VERB
alces-443	220	19	some	some	DET
alces-443	220	20	effect	effect	NOUN
alces-443	220	21	on	on	ADP
alces-443	220	22	genetic	genetic	ADJ
alces-443	220	23	structure	structure	NOUN
alces-443	220	24	(	(	PUNCT
alces-443	220	25	hundertmark	hundertmark	NOUN
alces-443	220	26	et	et	PROPN
alces-443	220	27	al	al	PROPN
alces-443	220	28	.	.	PROPN
alces-443	220	29	2003	2003	NUM
alces-443	220	30	)	)	PUNCT
alces-443	220	31	.	.	PUNCT
alces-443	221	1	based	base	VERB
alces-443	221	2	on	on	ADP
alces-443	221	3	the	the	DET
alces-443	221	4	reconstruction	reconstruction	NOUN
alces-443	221	5	of	of	ADP
alces-443	221	6	the	the	DET
alces-443	221	7	retreat	retreat	NOUN
alces-443	221	8	of	of	ADP
alces-443	221	9	the	the	DET
alces-443	221	10	laurentide	laurentide	ADJ
alces-443	221	11	ice	ice	NOUN
alces-443	221	12	sheet	sheet	NOUN
alces-443	221	13	by	by	ADP
alces-443	221	14	dyke	dyke	NOUN
alces-443	221	15	and	and	CCONJ
alces-443	221	16	prest	pr	ADJ
alces-443	221	17	(	(	PUNCT
alces-443	221	18	1987	1987	NUM
alces-443	221	19	)	)	PUNCT
alces-443	221	20	,	,	PUNCT
alces-443	221	21	we	we	PRON
alces-443	221	22	offer	offer	VERB
alces-443	221	23	the	the	DET
alces-443	221	24	following	follow	VERB
alces-443	221	25	scenario	scenario	NOUN
alces-443	221	26	of	of	ADP
alces-443	221	27	colonization	colonization	NOUN
alces-443	221	28	.	.	PUNCT
alces-443	222	1	at	at	ADP
alces-443	222	2	the	the	DET
alces-443	222	3	last	last	ADJ
alces-443	222	4	glacial	glacial	ADJ
alces-443	222	5	maximum	maximum	NOUN
alces-443	222	6	,	,	PUNCT
alces-443	222	7	the	the	DET
alces-443	222	8	cordilleran	cordilleran	ADJ
alces-443	222	9	and	and	CCONJ
alces-443	222	10	laurentide	laurentide	NOUN
alces-443	222	11	ice	ice	NOUN
alces-443	222	12	sheets	sheet	NOUN
alces-443	222	13	created	create	VERB
alces-443	222	14	an	an	DET
alces-443	222	15	effective	effective	ADJ
alces-443	222	16	barrier	barrier	NOUN
alces-443	222	17	between	between	ADP
alces-443	222	18	eastern	eastern	ADJ
alces-443	222	19	beringia	beringia	NOUN
alces-443	222	20	(	(	PUNCT
alces-443	222	21	alaska	alaska	PROPN
alces-443	222	22	)	)	PUNCT
alces-443	222	23	and	and	CCONJ
alces-443	222	24	other	other	ADJ
alces-443	222	25	parts	part	NOUN
alces-443	222	26	of	of	ADP
alces-443	222	27	the	the	DET
alces-443	222	28	continent	continent	NOUN
alces-443	222	29	(	(	PUNCT
alces-443	222	30	fig	fig	NOUN
alces-443	222	31	.	.	PUNCT
alces-443	223	1	5	5	NUM
alces-443	223	2	)	)	PUNCT
alces-443	223	3	.	.	PUNCT
alces-443	224	1	as	as	SCONJ
alces-443	224	2	glaciers	glacier	NOUN
alces-443	224	3	retreated	retreat	VERB
alces-443	224	4	,	,	PUNCT
alces-443	224	5	a	a	DET
alces-443	224	6	corridor	corridor	NOUN
alces-443	224	7	opened	open	VERB
alces-443	224	8	on	on	ADP
alces-443	224	9	the	the	DET
alces-443	224	10	eastern	eastern	ADJ
alces-443	224	11	slopes	slope	NOUN
alces-443	224	12	of	of	ADP
alces-443	224	13	the	the	DET
alces-443	224	14	rocky	rocky	ADJ
alces-443	224	15	mountains	mountain	NOUN
alces-443	224	16	allowing	allow	VERB
alces-443	224	17	passage	passage	NOUN
alces-443	224	18	to	to	ADP
alces-443	224	19	the	the	DET
alces-443	224	20	south	south	NOUN
alces-443	224	21	.	.	PUNCT
alces-443	225	1	by	by	ADP
alces-443	225	2	10,000	10,000	NUM
alces-443	225	3	years	year	NOUN
alces-443	225	4	ago	ago	ADV
alces-443	225	5	,	,	PUNCT
alces-443	225	6	western	western	ADJ
alces-443	225	7	canada	canada	PROPN
alces-443	225	8	was	be	AUX
alces-443	225	9	ice	ice	NOUN
alces-443	225	10	-	-	PUNCT
alces-443	225	11	free	free	ADJ
alces-443	225	12	but	but	CCONJ
alces-443	225	13	central	central	ADJ
alces-443	225	14	and	and	CCONJ
alces-443	225	15	eastern	eastern	ADJ
alces-443	225	16	canada	canada	PROPN
alces-443	225	17	remained	remain	VERB
alces-443	225	18	covered	cover	VERB
alces-443	225	19	by	by	ADP
alces-443	225	20	the	the	DET
alces-443	225	21	laurentide	laurentide	PROPN
alces-443	225	22	ice	ice	NOUN
alces-443	225	23	sheet	sheet	NOUN
alces-443	225	24	and	and	CCONJ
alces-443	225	25	large	large	ADJ
alces-443	225	26	proglacial	proglacial	ADJ
alces-443	225	27	lakes	lake	NOUN
alces-443	225	28	(	(	PUNCT
alces-443	225	29	fig	fig	NOUN
alces-443	225	30	.	.	PUNCT
alces-443	226	1	7	7	NUM
alces-443	226	2	)	)	PUNCT
alces-443	226	3	.	.	PUNCT
alces-443	227	1	passage	passage	NOUN
alces-443	227	2	to	to	ADP
alces-443	227	3	eastern	eastern	PROPN
alces-443	227	4	canada	canada	PROPN
alces-443	227	5	north	north	ADV
alces-443	227	6	of	of	ADP
alces-443	227	7	the	the	DET
alces-443	227	8	great	great	ADJ
alces-443	227	9	lakes	lake	NOUN
alces-443	227	10	was	be	AUX
alces-443	227	11	impossible	impossible	ADJ
alces-443	227	12	at	at	ADP
alces-443	227	13	this	this	DET
alces-443	227	14	time	time	NOUN
alces-443	227	15	and	and	CCONJ
alces-443	227	16	the	the	DET
alces-443	227	17	only	only	ADJ
alces-443	227	18	dispersal	dispersal	NOUN
alces-443	227	19	corridor	corridor	NOUN
alces-443	227	20	was	be	AUX
alces-443	227	21	south	south	ADV
alces-443	227	22	of	of	ADP
alces-443	227	23	the	the	DET
alces-443	227	24	lakes	lake	NOUN
alces-443	227	25	.	.	PUNCT
alces-443	228	1	by	by	ADP
alces-443	228	2	8,400	8,400	NUM
alces-443	228	3	years	year	NOUN
alces-443	228	4	ago	ago	ADV
alces-443	228	5	,	,	PUNCT
alces-443	228	6	moose	moose	NOUN
alces-443	228	7	arriving	arrive	VERB
alces-443	228	8	in	in	ADP
alces-443	228	9	the	the	DET
alces-443	228	10	eastern	eastern	ADJ
alces-443	228	11	continent	continent	NOUN
alces-443	228	12	could	could	AUX
alces-443	228	13	have	have	AUX
alces-443	228	14	dispersed	disperse	VERB
alces-443	228	15	westward	westward	ADV
alces-443	228	16	north	north	ADV
alces-443	228	17	of	of	ADP
alces-443	228	18	the	the	DET
alces-443	228	19	lakes	lake	NOUN
alces-443	228	20	,	,	PUNCT
alces-443	228	21	skirting	skirt	VERB
alces-443	228	22	the	the	DET
alces-443	228	23	southern	southern	ADJ
alces-443	228	24	shores	shore	NOUN
alces-443	228	25	of	of	ADP
alces-443	228	26	the	the	DET
alces-443	228	27	proglacial	proglacial	ADJ
alces-443	228	28	lakes	lake	NOUN
alces-443	228	29	to	to	ADP
alces-443	228	30	the	the	DET
alces-443	228	31	north	north	NOUN
alces-443	228	32	,	,	PUNCT
alces-443	228	33	and	and	CCONJ
alces-443	228	34	come	come	VERB
alces-443	228	35	into	into	ADP
alces-443	228	36	secondary	secondary	ADJ
alces-443	228	37	contact	contact	NOUN
alces-443	228	38	with	with	ADP
alces-443	228	39	moose	moose	NOUN
alces-443	228	40	from	from	ADP
alces-443	228	41	the	the	DET
alces-443	228	42	west	west	NOUN
alces-443	228	43	once	once	SCONJ
alces-443	228	44	the	the	DET
alces-443	228	45	proglacial	proglacial	ADJ
alces-443	228	46	lakes	lake	NOUN
alces-443	228	47	receded	recede	VERB
alces-443	228	48	.	.	PUNCT
alces-443	229	1	moose	moose	NOUN
alces-443	229	2	in	in	ADP
alces-443	229	3	the	the	DET
alces-443	229	4	eastern	eastern	ADJ
alces-443	229	5	part	part	NOUN
alces-443	229	6	of	of	ADP
alces-443	229	7	the	the	DET
alces-443	229	8	continent	continent	NOUN
alces-443	229	9	(	(	PUNCT
alces-443	229	10	a.	a.	NOUN
alces-443	229	11	a.	a.	NOUN
alces-443	229	12	americana	americana	PROPN
alces-443	229	13	)	)	PUNCT
alces-443	229	14	would	would	AUX
alces-443	229	15	have	have	AUX
alces-443	229	16	been	be	AUX
alces-443	229	17	on	on	ADP
alces-443	229	18	the	the	DET
alces-443	229	19	end	end	NOUN
alces-443	229	20	of	of	ADP
alces-443	229	21	a	a	DET
alces-443	229	22	series	series	NOUN
alces-443	229	23	of	of	ADP
alces-443	229	24	founder	founder	NOUN
alces-443	229	25	events	event	NOUN
alces-443	229	26	,	,	PUNCT
alces-443	229	27	explaining	explain	VERB
alces-443	229	28	their	their	PRON
alces-443	229	29	low	low	ADJ
alces-443	229	30	mitochondrial	mitochondrial	ADJ
alces-443	229	31	variability	variability	NOUN
alces-443	229	32	and	and	CCONJ
alces-443	229	33	the	the	DET
alces-443	229	34	presence	presence	NOUN
alces-443	229	35	of	of	ADP
alces-443	229	36	a	a	DET
alces-443	229	37	contact	contact	NOUN
alces-443	229	38	zone	zone	NOUN
alces-443	229	39	between	between	ADP
alces-443	229	40	a.	a.	NOUN
alces-443	229	41	a.	a.	NOUN
alces-443	229	42	americana	americana	PROPN
alces-443	229	43	and	and	CCONJ
alces-443	229	44	the	the	DET
alces-443	229	45	much	much	ADV
alces-443	229	46	more	more	ADJ
alces-443	229	47	variable	variable	ADJ
alces-443	229	48	a.	a.	NOUN
alces-443	229	49	a.	a.	NOUN
alces-443	229	50	andersoni	andersoni	PROPN
alces-443	229	51	in	in	ADP
alces-443	229	52	ontario	ontario	PROPN
alces-443	229	53	between	between	ADP
alces-443	229	54	the	the	DET
alces-443	229	55	great	great	ADJ
alces-443	229	56	lakes	lake	NOUN
alces-443	229	57	and	and	CCONJ
alces-443	229	58	hudson	hudson	PROPN
alces-443	229	59	bay	bay	PROPN
alces-443	229	60	.	.	PUNCT
alces-443	230	1	morphological	morphological	ADJ
alces-443	230	2	adaptation	adaptation	NOUN
alces-443	230	3	moose	moose	NOUN
alces-443	230	4	of	of	ADP
alces-443	230	5	the	the	DET
alces-443	230	6	pleistocene	pleistocene	NOUN
alces-443	230	7	and	and	CCONJ
alces-443	230	8	those	those	PRON
alces-443	230	9	that	that	PRON
alces-443	230	10	entered	enter	VERB
alces-443	230	11	north	north	PROPN
alces-443	230	12	america	america	PROPN
alces-443	230	13	at	at	ADP
alces-443	230	14	the	the	DET
alces-443	230	15	beginning	beginning	NOUN
alces-443	230	16	of	of	ADP
alces-443	230	17	the	the	DET
alces-443	230	18	holocene	holocene	PROPN
alces-443	230	19	were	be	AUX
alces-443	230	20	significantly	significantly	ADV
alces-443	230	21	larger	large	ADJ
alces-443	230	22	than	than	ADP
alces-443	230	23	those	those	PRON
alces-443	230	24	living	live	VERB
alces-443	230	25	today	today	NOUN
alces-443	230	26	,	,	PUNCT
alces-443	230	27	a	a	DET
alces-443	230	28	trait	trait	NOUN
alces-443	230	29	shared	share	VERB
alces-443	230	30	with	with	ADP
alces-443	230	31	other	other	ADJ
alces-443	230	32	northern	northern	ADJ
alces-443	230	33	ruminants	ruminant	NOUN
alces-443	230	34	(	(	PUNCT
alces-443	230	35	guthrie	guthrie	PROPN
alces-443	230	36	1984	1984	NUM
alces-443	230	37	)	)	PUNCT
alces-443	230	38	.	.	PUNCT
alces-443	231	1	guthrie	guthrie	PROPN
alces-443	231	2	(	(	PUNCT
alces-443	231	3	1984	1984	NUM
alces-443	231	4	)	)	PUNCT
alces-443	231	5	proposed	propose	VERB
alces-443	231	6	that	that	SCONJ
alces-443	231	7	the	the	DET
alces-443	231	8	reduction	reduction	NOUN
alces-443	231	9	in	in	ADP
alces-443	231	10	body	body	NOUN
alces-443	231	11	size	size	NOUN
alces-443	231	12	was	be	AUX
alces-443	231	13	an	an	DET
alces-443	231	14	adaptation	adaptation	NOUN
alces-443	231	15	to	to	ADP
alces-443	231	16	changes	change	NOUN
alces-443	231	17	in	in	ADP
alces-443	231	18	seasonal	seasonal	ADJ
alces-443	231	19	forage	forage	NOUN
alces-443	231	20	availability	availability	NOUN
alces-443	231	21	that	that	PRON
alces-443	231	22	occurred	occur	VERB
alces-443	231	23	as	as	ADP
alces-443	231	24	a	a	DET
alces-443	231	25	result	result	NOUN
alces-443	231	26	of	of	ADP
alces-443	231	27	climate	climate	NOUN
alces-443	231	28	amelioration	amelioration	NOUN
alces-443	231	29	at	at	ADP
alces-443	231	30	the	the	DET
alces-443	231	31	end	end	NOUN
alces-443	231	32	of	of	ADP
alces-443	231	33	the	the	DET
alces-443	231	34	last	last	ADJ
alces-443	231	35	ice	ice	NOUN
alces-443	231	36	age	age	NOUN
alces-443	231	37	.	.	PUNCT
alces-443	232	1	the	the	DET
alces-443	232	2	ability	ability	NOUN
alces-443	232	3	of	of	ADP
alces-443	232	4	moose	moose	NOUN
alces-443	232	5	to	to	PART
alces-443	232	6	respond	respond	VERB
alces-443	232	7	to	to	ADP
alces-443	232	8	a	a	DET
alces-443	232	9	rapidly	rapidly	ADV
alces-443	232	10	changing	change	VERB
alces-443	232	11	environment	environment	NOUN
alces-443	232	12	belies	belie	VERB
alces-443	232	13	the	the	DET
alces-443	232	14	relatively	relatively	ADV
alces-443	232	15	low	low	ADJ
alces-443	232	16	levels	level	NOUN
alces-443	232	17	of	of	ADP
alces-443	232	18	genetic	genetic	ADJ
alces-443	232	19	variation	variation	NOUN
alces-443	232	20	documented	document	VERB
alces-443	232	21	by	by	ADP
alces-443	232	22	the	the	DET
alces-443	232	23	studies	study	NOUN
alces-443	232	24	we	we	PRON
alces-443	232	25	have	have	AUX
alces-443	232	26	reviewed	review	VERB
alces-443	232	27	and	and	CCONJ
alces-443	232	28	demonstrates	demonstrate	VERB
alces-443	232	29	that	that	SCONJ
alces-443	232	30	evolutionary	evolutionary	ADJ
alces-443	232	31	potential	potential	NOUN
alces-443	232	32	is	be	AUX
alces-443	232	33	not	not	PART
alces-443	232	34	easily	easily	ADV
alces-443	232	35	predicted	predict	VERB
alces-443	232	36	solely	solely	ADV
alces-443	232	37	by	by	ADP
alces-443	232	38	genetic	genetic	ADJ
alces-443	232	39	variability	variability	NOUN
alces-443	232	40	but	but	CCONJ
alces-443	232	41	ultimately	ultimately	ADV
alces-443	232	42	is	be	AUX
alces-443	232	43	determined	determine	VERB
alces-443	232	44	by	by	ADP
alces-443	232	45	the	the	DET
alces-443	232	46	presence	presence	NOUN
alces-443	232	47	of	of	ADP
alces-443	232	48	adaptive	adaptive	ADJ
alces-443	232	49	genetic	genetic	ADJ
alces-443	232	50	variation	variation	NOUN
alces-443	232	51	and	and	CCONJ
alces-443	232	52	heritability	heritability	NOUN
alces-443	232	53	of	of	ADP
alces-443	232	54	traits	trait	NOUN
alces-443	232	55	that	that	PRON
alces-443	232	56	improve	improve	VERB
alces-443	232	57	fitness	fitness	NOUN
alces-443	232	58	(	(	PUNCT
alces-443	232	59	lynch	lynch	PROPN
alces-443	232	60	1996	1996	NUM
alces-443	232	61	)	)	PUNCT
alces-443	232	62	.	.	PUNCT
alces-443	233	1	a	a	DET
alces-443	233	2	general	general	ADJ
alces-443	233	3	reduction	reduction	NOUN
alces-443	233	4	in	in	ADP
alces-443	233	5	body	body	NOUN
alces-443	233	6	size	size	NOUN
alces-443	233	7	is	be	AUX
alces-443	233	8	not	not	PART
alces-443	233	9	the	the	DET
alces-443	233	10	only	only	ADJ
alces-443	233	11	change	change	NOUN
alces-443	233	12	to	to	PART
alces-443	233	13	occur	occur	VERB
alces-443	233	14	since	since	SCONJ
alces-443	233	15	the	the	DET
alces-443	233	16	colonization	colonization	NOUN
alces-443	233	17	of	of	ADP
alces-443	233	18	north	north	PROPN
alces-443	233	19	america	america	PROPN
alces-443	233	20	.	.	PUNCT
alces-443	234	1	moose	moose	PROPN
alces-443	234	2	in	in	ADP
alces-443	234	3	hundertmark	hundertmark	PROPN
alces-443	234	4	and	and	CCONJ
alces-443	234	5	bowyer	bowyer	PROPN
alces-443	234	6	moose	moose	PROPN
alces-443	234	7	genetics	genetics	PROPN
alces-443	234	8	alces	alce	NOUN
alces-443	234	9	vol	vol	NOUN
alces-443	234	10	.	.	PROPN
alces-443	235	1	40	40	NUM
alces-443	235	2	,	,	PUNCT
alces-443	235	3	2004	2004	NUM
alces-443	235	4	114	114	NUM
alces-443	235	5	fig	fig	NOUN
alces-443	235	6	.	.	PUNCT
alces-443	236	1	7	7	X
alces-443	236	2	.	.	X
alces-443	236	3	coverage	coverage	NOUN
alces-443	236	4	of	of	ADP
alces-443	236	5	north	north	PROPN
alces-443	236	6	america	america	PROPN
alces-443	236	7	by	by	ADP
alces-443	236	8	the	the	DET
alces-443	236	9	cordilleran	cordilleran	ADJ
alces-443	236	10	and	and	CCONJ
alces-443	236	11	laurentide	laurentide	NOUN
alces-443	236	12	ice	ice	NOUN
alces-443	236	13	sheets	sheet	NOUN
alces-443	236	14	and	and	CCONJ
alces-443	236	15	proglacial	proglacial	ADJ
alces-443	236	16	lakes	lake	NOUN
alces-443	236	17	at	at	ADP
alces-443	236	18	14,000	14,000	NUM
alces-443	236	19	,	,	PUNCT
alces-443	236	20	10,000	10,000	NUM
alces-443	236	21	,	,	PUNCT
alces-443	236	22	and	and	CCONJ
alces-443	236	23	8,400	8,400	NUM
alces-443	236	24	years	year	NOUN
alces-443	236	25	ago	ago	ADV
alces-443	236	26	(	(	PUNCT
alces-443	236	27	adapted	adapt	VERB
alces-443	236	28	from	from	ADP
alces-443	236	29	dyke	dyke	NOUN
alces-443	236	30	and	and	CCONJ
alces-443	236	31	prest	pr	ADJ
alces-443	236	32	1987	1987	NUM
alces-443	236	33	)	)	PUNCT
alces-443	236	34	.	.	PUNCT
alces-443	237	1	north	north	PROPN
alces-443	237	2	america	america	PROPN
alces-443	237	3	exhibit	exhibit	VERB
alces-443	237	4	many	many	ADJ
alces-443	237	5	differences	difference	NOUN
alces-443	237	6	in	in	ADP
alces-443	237	7	behavior	behavior	NOUN
alces-443	237	8	and	and	CCONJ
alces-443	237	9	morphology	morphology	NOUN
alces-443	237	10	.	.	PUNCT
alces-443	238	1	alaskan	alaskan	ADJ
alces-443	238	2	moose	moose	NOUN
alces-443	238	3	are	be	AUX
alces-443	238	4	perhaps	perhaps	ADV
alces-443	238	5	the	the	DET
alces-443	238	6	most	most	ADV
alces-443	238	7	divergent	divergent	ADJ
alces-443	238	8	;	;	PUNCT
alces-443	238	9	they	they	PRON
alces-443	238	10	exhibit	exhibit	VERB
alces-443	238	11	a	a	DET
alces-443	238	12	degree	degree	NOUN
alces-443	238	13	of	of	ADP
alces-443	238	14	sociality	sociality	NOUN
alces-443	238	15	not	not	PART
alces-443	238	16	observed	observe	VERB
alces-443	238	17	elsewhere	elsewhere	ADV
alces-443	238	18	(	(	PUNCT
alces-443	238	19	molvar	molvar	NOUN
alces-443	238	20	and	and	CCONJ
alces-443	238	21	bowyer	bowyer	PROPN
alces-443	238	22	1994	1994	NUM
alces-443	238	23	)	)	PUNCT
alces-443	238	24	and	and	CCONJ
alces-443	238	25	have	have	VERB
alces-443	238	26	more	more	ADJ
alces-443	238	27	distinctive	distinctive	ADJ
alces-443	238	28	body	body	NOUN
alces-443	238	29	markings	marking	NOUN
alces-443	238	30	,	,	PUNCT
alces-443	238	31	also	also	ADV
alces-443	238	32	indicative	indicative	ADJ
alces-443	238	33	of	of	ADP
alces-443	238	34	increased	increase	VERB
alces-443	238	35	sociality	sociality	NOUN
alces-443	238	36	(	(	PUNCT
alces-443	238	37	bowyer	bowyer	PROPN
alces-443	238	38	et	et	PROPN
alces-443	238	39	al	al	PROPN
alces-443	238	40	.	.	PROPN
alces-443	238	41	1991	1991	NUM
alces-443	238	42	)	)	PUNCT
alces-443	238	43	.	.	PUNCT
alces-443	239	1	molvar	molvar	NOUN
alces-443	239	2	and	and	CCONJ
alces-443	239	3	bowyer	bowyer	PROPN
alces-443	239	4	(	(	PUNCT
alces-443	239	5	1994	1994	NUM
alces-443	239	6	)	)	PUNCT
alces-443	239	7	suggested	suggest	VERB
alces-443	239	8	that	that	SCONJ
alces-443	239	9	moose	moose	NOUN
alces-443	239	10	in	in	ADP
alces-443	239	11	alaska	alaska	PROPN
alces-443	239	12	have	have	AUX
alces-443	239	13	evolved	evolve	VERB
alces-443	239	14	sociality	sociality	NOUN
alces-443	239	15	recently	recently	ADV
alces-443	239	16	as	as	ADP
alces-443	239	17	a	a	DET
alces-443	239	18	response	response	NOUN
alces-443	239	19	to	to	ADP
alces-443	239	20	living	live	VERB
alces-443	239	21	in	in	ADP
alces-443	239	22	open	open	ADJ
alces-443	239	23	environments	environment	NOUN
alces-443	239	24	.	.	PUNCT
alces-443	240	1	adaptation	adaptation	NOUN
alces-443	240	2	to	to	PART
alces-443	240	3	open	open	VERB
alces-443	240	4	environments	environment	NOUN
alces-443	240	5	also	also	ADV
alces-443	240	6	applies	apply	VERB
alces-443	240	7	to	to	ADP
alces-443	240	8	their	their	PRON
alces-443	240	9	mating	mating	NOUN
alces-443	240	10	system	system	NOUN
alces-443	240	11	,	,	PUNCT
alces-443	240	12	which	which	PRON
alces-443	240	13	is	be	AUX
alces-443	240	14	harem	harem	PROPN
alces-443	240	15	-	-	PUNCT
alces-443	240	16	based	base	VERB
alces-443	240	17	.	.	PUNCT
alces-443	241	1	harem	harem	PROPN
alces-443	241	2	mating	mating	NOUN
alces-443	241	3	is	be	AUX
alces-443	241	4	adaptive	adaptive	ADJ
alces-443	241	5	in	in	ADP
alces-443	241	6	open	open	ADJ
alces-443	241	7	environments	environment	NOUN
alces-443	241	8	(	(	PUNCT
alces-443	241	9	hirth	hirth	NOUN
alces-443	241	10	1977	1977	NUM
alces-443	241	11	)	)	PUNCT
alces-443	241	12	where	where	SCONJ
alces-443	241	13	a	a	DET
alces-443	241	14	male	male	NOUN
alces-443	241	15	can	can	AUX
alces-443	241	16	protect	protect	VERB
alces-443	241	17	a	a	DET
alces-443	241	18	harem	harem	PROPN
alces-443	241	19	from	from	ADP
alces-443	241	20	competitors	competitor	NOUN
alces-443	241	21	.	.	PUNCT
alces-443	242	1	moose	moose	VERB
alces-443	242	2	elsewhere	elsewhere	ADV
alces-443	242	3	in	in	ADP
alces-443	242	4	north	north	PROPN
alces-443	242	5	america	america	PROPN
alces-443	242	6	exhibit	exhibit	VERB
alces-443	242	7	a	a	DET
alces-443	242	8	tending	tend	VERB
alces-443	242	9	-	-	PUNCT
alces-443	242	10	bond	bond	NOUN
alces-443	242	11	system	system	NOUN
alces-443	242	12	of	of	ADP
alces-443	242	13	mating	mating	NOUN
alces-443	242	14	,	,	PUNCT
alces-443	242	15	which	which	PRON
alces-443	242	16	is	be	AUX
alces-443	242	17	adaptive	adaptive	ADJ
alces-443	242	18	for	for	ADP
alces-443	242	19	forested	forest	VERB
alces-443	242	20	environments	environment	NOUN
alces-443	242	21	.	.	PUNCT
alces-443	243	1	moose	moose	PROPN
alces-443	243	2	in	in	ADP
alces-443	243	3	alaska	alaska	PROPN
alces-443	243	4	and	and	CCONJ
alces-443	243	5	siberia	siberia	PROPN
alces-443	243	6	exhibit	exhibit	VERB
alces-443	243	7	the	the	DET
alces-443	243	8	largest	large	ADJ
alces-443	243	9	body	body	NOUN
alces-443	243	10	size	size	NOUN
alces-443	243	11	of	of	ADP
alces-443	243	12	moose	moose	NOUN
alces-443	243	13	in	in	ADP
alces-443	243	14	north	north	PROPN
alces-443	243	15	america	america	PROPN
alces-443	243	16	and	and	CCONJ
alces-443	243	17	eurasia	eurasia	PROPN
alces-443	243	18	,	,	PUNCT
alces-443	243	19	respectively	respectively	ADV
alces-443	243	20	.	.	PUNCT
alces-443	244	1	the	the	DET
alces-443	244	2	similar	similar	ADJ
alces-443	244	3	appearance	appearance	NOUN
alces-443	244	4	of	of	ADP
alces-443	244	5	moose	moose	NOUN
alces-443	244	6	occurring	occur	VERB
alces-443	244	7	on	on	ADP
alces-443	244	8	either	either	DET
alces-443	244	9	side	side	NOUN
alces-443	244	10	of	of	ADP
alces-443	244	11	the	the	DET
alces-443	244	12	bering	bering	NOUN
alces-443	244	13	strait	strait	NOUN
alces-443	244	14	has	have	AUX
alces-443	244	15	caused	cause	VERB
alces-443	244	16	some	some	DET
alces-443	244	17	investigators	investigator	NOUN
alces-443	244	18	to	to	PART
alces-443	244	19	consider	consider	VERB
alces-443	244	20	them	they	PRON
alces-443	244	21	the	the	DET
alces-443	244	22	same	same	ADJ
alces-443	244	23	subspecies	subspecie	NOUN
alces-443	244	24	(	(	PUNCT
alces-443	244	25	e.g.	e.g.	ADV
alces-443	244	26	,	,	PUNCT
alces-443	244	27	telfer	telfer	NOUN
alces-443	244	28	1984	1984	NUM
alces-443	244	29	)	)	PUNCT
alces-443	244	30	.	.	PUNCT
alces-443	245	1	as	as	SCONJ
alces-443	245	2	moose	moose	NOUN
alces-443	245	3	from	from	ADP
alces-443	245	4	those	those	DET
alces-443	245	5	2	2	NUM
alces-443	245	6	regions	region	NOUN
alces-443	245	7	are	be	AUX
alces-443	245	8	not	not	PART
alces-443	245	9	closely	closely	ADV
alces-443	245	10	related	relate	VERB
alces-443	245	11	(	(	PUNCT
alces-443	245	12	hundertmark	hundertmark	NOUN
alces-443	245	13	et	et	PROPN
alces-443	245	14	al	al	PROPN
alces-443	245	15	.	.	PROPN
alces-443	245	16	2002a	2002a	NOUN
alces-443	245	17	,	,	PUNCT
alces-443	245	18	b	b	NOUN
alces-443	245	19	)	)	PUNCT
alces-443	245	20	,	,	PUNCT
alces-443	245	21	their	their	PRON
alces-443	245	22	similarity	similarity	NOUN
alces-443	245	23	in	in	ADP
alces-443	245	24	size	size	NOUN
alces-443	245	25	must	must	AUX
alces-443	245	26	result	result	VERB
alces-443	245	27	from	from	ADP
alces-443	245	28	convergent	convergent	NOUN
alces-443	245	29	evolution	evolution	NOUN
alces-443	245	30	.	.	PUNCT
alces-443	246	1	both	both	DET
alces-443	246	2	subspecies	subspecie	NOUN
alces-443	246	3	have	have	AUX
alces-443	246	4	adapted	adapt	VERB
alces-443	246	5	to	to	PART
alces-443	246	6	open	open	VERB
alces-443	246	7	,	,	PUNCT
alces-443	246	8	northern	northern	ADJ
alces-443	246	9	habitats	habitat	NOUN
alces-443	246	10	by	by	ADP
alces-443	246	11	increasing	increase	VERB
alces-443	246	12	body	body	NOUN
alces-443	246	13	size	size	NOUN
alces-443	246	14	.	.	PUNCT
alces-443	247	1	adaptation	adaptation	NOUN
alces-443	247	2	to	to	PART
alces-443	247	3	open	open	VERB
alces-443	247	4	habitats	habitat	NOUN
alces-443	247	5	was	be	AUX
alces-443	247	6	demonstrated	demonstrate	VERB
alces-443	247	7	with	with	ADP
alces-443	247	8	a	a	DET
alces-443	247	9	multivariate	multivariate	NOUN
alces-443	247	10	analysis	analysis	NOUN
alces-443	247	11	of	of	ADP
alces-443	247	12	antler	antler	ADJ
alces-443	247	13	size	size	NOUN
alces-443	247	14	among	among	ADP
alces-443	247	15	moose	moose	NOUN
alces-443	247	16	inhabiting	inhabit	VERB
alces-443	247	17	different	different	ADJ
alces-443	247	18	areas	area	NOUN
alces-443	247	19	and	and	CCONJ
alces-443	247	20	habitats	habitat	NOUN
alces-443	247	21	in	in	ADP
alces-443	247	22	alaska	alaska	PROPN
alces-443	247	23	(	(	PUNCT
alces-443	247	24	bowyer	bowyer	PROPN
alces-443	247	25	et	et	PROPN
alces-443	247	26	al	al	PROPN
alces-443	247	27	.	.	PROPN
alces-443	247	28	2002	2002	NUM
alces-443	247	29	)	)	PUNCT
alces-443	247	30	.	.	PUNCT
alces-443	248	1	those	those	DET
alces-443	248	2	moose	moose	NOUN
alces-443	248	3	inhabiting	inhabit	VERB
alces-443	248	4	open	open	ADJ
alces-443	248	5	habitats	habitat	NOUN
alces-443	248	6	(	(	PUNCT
alces-443	248	7	tundra	tundra	NOUN
alces-443	248	8	)	)	PUNCT
alces-443	248	9	tended	tend	VERB
alces-443	248	10	to	to	PART
alces-443	248	11	have	have	VERB
alces-443	248	12	larger	large	ADJ
alces-443	248	13	antlers	antler	NOUN
alces-443	248	14	overall	overall	ADV
alces-443	248	15	than	than	ADP
alces-443	248	16	those	those	PRON
alces-443	248	17	living	live	VERB
alces-443	248	18	in	in	ADP
alces-443	248	19	boreal	boreal	ADJ
alces-443	248	20	forest	forest	NOUN
alces-443	248	21	(	(	PUNCT
alces-443	248	22	taiga	taiga	NOUN
alces-443	248	23	)	)	PUNCT
alces-443	248	24	.	.	PUNCT
alces-443	249	1	similarly	similarly	ADV
alces-443	249	2	,	,	PUNCT
alces-443	249	3	moose	moose	NOUN
alces-443	249	4	occupying	occupy	VERB
alces-443	249	5	mountainous	mountainous	ADJ
alces-443	249	6	habitat	habitat	NOUN
alces-443	249	7	in	in	ADP
alces-443	249	8	the	the	DET
alces-443	249	9	southern	southern	ADJ
alces-443	249	10	portions	portion	NOUN
alces-443	249	11	of	of	ADP
alces-443	249	12	their	their	PRON
alces-443	249	13	range	range	NOUN
alces-443	249	14	in	in	ADP
alces-443	249	15	north	north	PROPN
alces-443	249	16	america	america	PROPN
alces-443	249	17	(	(	PUNCT
alces-443	249	18	a.	a.	NOUN
alces-443	249	19	a.	a.	PROPN
alces-443	249	20	shirasi	shirasi	PROPN
alces-443	249	21	)	)	PUNCT
alces-443	249	22	and	and	CCONJ
alces-443	249	23	asia	asia	PROPN
alces-443	249	24	(	(	PUNCT
alces-443	249	25	a.	a.	NOUN
alces-443	249	26	a.	a.	NOUN
alces-443	249	27	cameloides	cameloide	NOUN
alces-443	249	28	)	)	PUNCT
alces-443	249	29	are	be	AUX
alces-443	249	30	similar	similar	ADJ
alces-443	249	31	;	;	PUNCT
alces-443	249	32	exhibiting	exhibit	VERB
alces-443	249	33	small	small	ADJ
alces-443	249	34	body	body	NOUN
alces-443	249	35	and	and	CCONJ
alces-443	249	36	antler	antler	NOUN
alces-443	249	37	size	size	NOUN
alces-443	249	38	(	(	PUNCT
alces-443	249	39	geist	geist	NOUN
alces-443	249	40	1998	1998	NUM
alces-443	249	41	)	)	PUNCT
alces-443	249	42	.	.	PUNCT
alces-443	250	1	bubenik	bubenik	PROPN
alces-443	250	2	(	(	PUNCT
alces-443	250	3	1998	1998	NUM
alces-443	250	4	)	)	PUNCT
alces-443	250	5	explained	explain	VERB
alces-443	250	6	that	that	SCONJ
alces-443	250	7	similarity	similarity	NOUN
alces-443	250	8	by	by	ADP
alces-443	250	9	proposing	propose	VERB
alces-443	250	10	a	a	DET
alces-443	250	11	second	second	ADJ
alces-443	250	12	entry	entry	NOUN
alces-443	250	13	into	into	ADP
alces-443	250	14	north	north	PROPN
alces-443	250	15	america	america	PROPN
alces-443	250	16	by	by	ADP
alces-443	250	17	asian	asian	ADJ
alces-443	250	18	moose	moose	NOUN
alces-443	250	19	—	—	PUNCT
alces-443	250	20	an	an	DET
alces-443	250	21	entry	entry	NOUN
alces-443	250	22	that	that	PRON
alces-443	250	23	bypassed	bypass	VERB
alces-443	250	24	beringia	beringia	NOUN
alces-443	250	25	by	by	ADP
alces-443	250	26	traveling	travel	VERB
alces-443	250	27	along	along	ADP
alces-443	250	28	the	the	DET
alces-443	250	29	southern	southern	ADJ
alces-443	250	30	coast	coast	NOUN
alces-443	250	31	of	of	ADP
alces-443	250	32	alaska	alaska	PROPN
alces-443	250	33	.	.	PUNCT
alces-443	251	1	neither	neither	CCONJ
alces-443	251	2	genetic	genetic	ADJ
alces-443	251	3	nor	nor	CCONJ
alces-443	251	4	fossil	fossil	NOUN
alces-443	251	5	data	datum	NOUN
alces-443	251	6	support	support	NOUN
alces-443	251	7	that	that	DET
alces-443	251	8	hypothesis	hypothesis	NOUN
alces-443	251	9	(	(	PUNCT
alces-443	251	10	guthrie	guthrie	PROPN
alces-443	251	11	1990	1990	NUM
alces-443	251	12	,	,	PUNCT
alces-443	251	13	hundertmark	hundertmark	NOUN
alces-443	251	14	et	et	PROPN
alces-443	251	15	al	al	PROPN
alces-443	251	16	.	.	PROPN
alces-443	251	17	2002a	2002a	NOUN
alces-443	251	18	,	,	PUNCT
alces-443	251	19	b	b	NOUN
alces-443	251	20	,	,	PUNCT
alces-443	251	21	2003	2003	NUM
alces-443	251	22	)	)	PUNCT
alces-443	251	23	.	.	PUNCT
alces-443	252	1	genetic	genetic	ADJ
alces-443	252	2	effects	effect	NOUN
alces-443	252	3	of	of	ADP
alces-443	252	4	management	management	NOUN
alces-443	252	5	to	to	ADP
alces-443	252	6	this	this	DET
alces-443	252	7	point	point	NOUN
alces-443	252	8	we	we	PRON
alces-443	252	9	have	have	AUX
alces-443	252	10	discussed	discuss	VERB
alces-443	252	11	distribution	distribution	NOUN
alces-443	252	12	of	of	ADP
alces-443	252	13	genetic	genetic	ADJ
alces-443	252	14	and	and	CCONJ
alces-443	252	15	morphological	morphological	ADJ
alces-443	252	16	diversity	diversity	NOUN
alces-443	252	17	over	over	ADP
alces-443	252	18	space	space	NOUN
alces-443	252	19	and	and	CCONJ
alces-443	252	20	time	time	NOUN
alces-443	252	21	in	in	ADP
alces-443	252	22	the	the	DET
alces-443	252	23	context	context	NOUN
alces-443	252	24	of	of	ADP
alces-443	252	25	genetic	genetic	ADJ
alces-443	252	26	drift	drift	NOUN
alces-443	252	27	and	and	CCONJ
alces-443	252	28	selection	selection	NOUN
alces-443	252	29	for	for	ADP
alces-443	252	30	locally	locally	ADV
alces-443	252	31	adaptive	adaptive	ADJ
alces-443	252	32	traits	trait	NOUN
alces-443	252	33	.	.	PUNCT
alces-443	253	1	those	those	DET
alces-443	253	2	patterns	pattern	NOUN
alces-443	253	3	are	be	AUX
alces-443	253	4	developed	develop	VERB
alces-443	253	5	over	over	ADP
alces-443	253	6	relatively	relatively	ADV
alces-443	253	7	long	long	ADJ
alces-443	253	8	periods	period	NOUN
alces-443	253	9	of	of	ADP
alces-443	253	10	time	time	NOUN
alces-443	253	11	and	and	CCONJ
alces-443	253	12	are	be	AUX
alces-443	253	13	integral	integral	ADJ
alces-443	253	14	to	to	ADP
alces-443	253	15	the	the	DET
alces-443	253	16	process	process	NOUN
alces-443	253	17	of	of	ADP
alces-443	253	18	evolution	evolution	NOUN
alces-443	253	19	.	.	PUNCT
alces-443	254	1	genetic	genetic	ADJ
alces-443	254	2	alces	alce	NOUN
alces-443	254	3	vol	vol	NOUN
alces-443	254	4	.	.	PROPN
alces-443	255	1	40	40	NUM
alces-443	255	2	,	,	PUNCT
alces-443	255	3	2004	2004	NUM
alces-443	255	4	moose	moose	NOUN
alces-443	255	5	genetics	genetic	NOUN
alces-443	255	6	hundertmark	hundertmark	NOUN
alces-443	255	7	and	and	CCONJ
alces-443	255	8	bowyer	bowyer	PROPN
alces-443	255	9	115	115	PROPN
alces-443	255	10	change	change	NOUN
alces-443	255	11	also	also	ADV
alces-443	255	12	can	can	AUX
alces-443	255	13	occur	occur	VERB
alces-443	255	14	over	over	ADP
alces-443	255	15	short	short	ADJ
alces-443	255	16	periods	period	NOUN
alces-443	255	17	due	due	ADJ
alces-443	255	18	to	to	ADP
alces-443	255	19	human	human	ADJ
alces-443	255	20	influences	influence	NOUN
alces-443	255	21	,	,	PUNCT
alces-443	255	22	notably	notably	ADV
alces-443	255	23	harvest	harvest	NOUN
alces-443	255	24	management	management	NOUN
alces-443	255	25	,	,	PUNCT
alces-443	255	26	and	and	CCONJ
alces-443	255	27	those	those	DET
alces-443	255	28	changes	change	NOUN
alces-443	255	29	may	may	AUX
alces-443	255	30	have	have	VERB
alces-443	255	31	unintended	unintended	ADJ
alces-443	255	32	and	and	CCONJ
alces-443	255	33	undesirable	undesirable	ADJ
alces-443	255	34	consequences	consequence	NOUN
alces-443	255	35	on	on	ADP
alces-443	255	36	individuals	individual	NOUN
alces-443	255	37	and	and	CCONJ
alces-443	255	38	populations	population	NOUN
alces-443	255	39	(	(	PUNCT
alces-443	255	40	harris	harris	PROPN
alces-443	255	41	et	et	PROPN
alces-443	255	42	al	al	PROPN
alces-443	255	43	.	.	PROPN
alces-443	255	44	2002	2002	NUM
alces-443	255	45	)	)	PUNCT
alces-443	255	46	and	and	CCONJ
alces-443	255	47	therefore	therefore	ADV
alces-443	255	48	are	be	AUX
alces-443	255	49	important	important	ADJ
alces-443	255	50	to	to	PART
alces-443	255	51	recognize	recognize	VERB
alces-443	255	52	.	.	PUNCT
alces-443	256	1	although	although	SCONJ
alces-443	256	2	it	it	PRON
alces-443	256	3	might	might	AUX
alces-443	256	4	be	be	AUX
alces-443	256	5	assumed	assume	VERB
alces-443	256	6	that	that	SCONJ
alces-443	256	7	a	a	DET
alces-443	256	8	well	well	ADV
alces-443	256	9	-	-	PUNCT
alces-443	256	10	designed	design	VERB
alces-443	256	11	harvest	harvest	NOUN
alces-443	256	12	plan	plan	NOUN
alces-443	256	13	acts	act	VERB
alces-443	256	14	in	in	ADP
alces-443	256	15	a	a	DET
alces-443	256	16	random	random	ADJ
alces-443	256	17	fashion	fashion	NOUN
alces-443	256	18	on	on	ADP
alces-443	256	19	genetic	genetic	ADJ
alces-443	256	20	makeup	makeup	NOUN
alces-443	256	21	of	of	ADP
alces-443	256	22	individuals	individual	NOUN
alces-443	256	23	,	,	PUNCT
alces-443	256	24	in	in	ADP
alces-443	256	25	reality	reality	NOUN
alces-443	256	26	even	even	ADV
alces-443	256	27	a	a	DET
alces-443	256	28	managed	manage	VERB
alces-443	256	29	harvest	harvest	NOUN
alces-443	256	30	can	can	AUX
alces-443	256	31	be	be	AUX
alces-443	256	32	a	a	DET
alces-443	256	33	highly	highly	ADV
alces-443	256	34	selective	selective	ADJ
alces-443	256	35	force	force	NOUN
alces-443	256	36	with	with	ADP
alces-443	256	37	measurable	measurable	ADJ
alces-443	256	38	consequences	consequence	NOUN
alces-443	256	39	in	in	ADP
alces-443	256	40	just	just	ADV
alces-443	256	41	a	a	DET
alces-443	256	42	few	few	ADJ
alces-443	256	43	generations	generation	NOUN
alces-443	256	44	(	(	PUNCT
alces-443	256	45	coltman	coltman	NOUN
alces-443	256	46	et	et	PROPN
alces-443	256	47	al	al	PROPN
alces-443	256	48	.	.	PROPN
alces-443	256	49	2003	2003	NUM
alces-443	256	50	)	)	PUNCT
alces-443	256	51	.	.	PUNCT
alces-443	257	1	ryman	ryman	NOUN
alces-443	257	2	et	et	PROPN
alces-443	257	3	al	al	PROPN
alces-443	257	4	.	.	PROPN
alces-443	257	5	(	(	PUNCT
alces-443	257	6	1981	1981	NUM
alces-443	257	7	)	)	PUNCT
alces-443	257	8	modeled	model	VERB
alces-443	257	9	the	the	DET
alces-443	257	10	effect	effect	NOUN
alces-443	257	11	of	of	ADP
alces-443	257	12	different	different	ADJ
alces-443	257	13	harvest	harvest	NOUN
alces-443	257	14	strategies	strategy	NOUN
alces-443	257	15	on	on	ADP
alces-443	257	16	2	2	NUM
alces-443	257	17	factors	factor	NOUN
alces-443	257	18	critical	critical	ADJ
alces-443	257	19	in	in	ADP
alces-443	257	20	determining	determine	VERB
alces-443	257	21	the	the	DET
alces-443	257	22	extent	extent	NOUN
alces-443	257	23	of	of	ADP
alces-443	257	24	genetic	genetic	ADJ
alces-443	257	25	drift	drift	NOUN
alces-443	257	26	and	and	CCONJ
alces-443	257	27	inbreeding	inbreede	VERB
alces-443	257	28	in	in	ADP
alces-443	257	29	populations	population	NOUN
alces-443	257	30	:	:	PUNCT
alces-443	257	31	effective	effective	ADJ
alces-443	257	32	population	population	NOUN
alces-443	257	33	size	size	NOUN
alces-443	257	34	and	and	CCONJ
alces-443	257	35	generation	generation	NOUN
alces-443	257	36	interval	interval	NOUN
alces-443	257	37	,	,	PUNCT
alces-443	257	38	respectively	respectively	ADV
alces-443	257	39	.	.	PUNCT
alces-443	258	1	hunting	hunt	VERB
alces-443	258	2	strategies	strategy	NOUN
alces-443	258	3	were	be	AUX
alces-443	258	4	defined	define	VERB
alces-443	258	5	by	by	ADP
alces-443	258	6	different	different	ADJ
alces-443	258	7	probabilities	probability	NOUN
alces-443	258	8	of	of	ADP
alces-443	258	9	harvest	harvest	NOUN
alces-443	258	10	for	for	ADP
alces-443	258	11	both	both	DET
alces-443	258	12	juveniles	juvenile	NOUN
alces-443	258	13	and	and	CCONJ
alces-443	258	14	adult	adult	NOUN
alces-443	258	15	females	female	NOUN
alces-443	258	16	,	,	PUNCT
alces-443	258	17	and	and	CCONJ
alces-443	258	18	resulted	result	VERB
alces-443	258	19	in	in	ADP
alces-443	258	20	stable	stable	ADJ
alces-443	258	21	populations	population	NOUN
alces-443	258	22	with	with	ADP
alces-443	258	23	decreased	decrease	VERB
alces-443	258	24	generation	generation	NOUN
alces-443	258	25	intervals	interval	NOUN
alces-443	258	26	and	and	CCONJ
alces-443	258	27	effective	effective	ADJ
alces-443	258	28	sizes	size	NOUN
alces-443	258	29	compared	compare	VERB
alces-443	258	30	with	with	ADP
alces-443	258	31	unhunted	unhunted	ADJ
alces-443	258	32	populations	population	NOUN
alces-443	258	33	.	.	PUNCT
alces-443	259	1	moreover	moreover	ADV
alces-443	259	2	,	,	PUNCT
alces-443	259	3	temporal	temporal	ADJ
alces-443	259	4	changes	change	NOUN
alces-443	259	5	in	in	ADP
alces-443	259	6	genetic	genetic	ADJ
alces-443	259	7	diversity	diversity	NOUN
alces-443	259	8	differed	differ	VERB
alces-443	259	9	for	for	ADP
alces-443	259	10	different	different	ADJ
alces-443	259	11	harvest	harvest	NOUN
alces-443	259	12	strategies	strategy	NOUN
alces-443	259	13	but	but	CCONJ
alces-443	259	14	always	always	ADV
alces-443	259	15	decreased	decrease	VERB
alces-443	259	16	.	.	PUNCT
alces-443	260	1	thus	thus	ADV
alces-443	260	2	,	,	PUNCT
alces-443	260	3	ryman	ryman	NOUN
alces-443	260	4	et	et	PROPN
alces-443	260	5	al	al	PROPN
alces-443	260	6	.	.	PROPN
alces-443	260	7	(	(	PUNCT
alces-443	260	8	1981	1981	NUM
alces-443	260	9	)	)	PUNCT
alces-443	260	10	demonstrated	demonstrate	VERB
alces-443	260	11	that	that	SCONJ
alces-443	260	12	improperly	improperly	ADV
alces-443	260	13	designed	design	VERB
alces-443	260	14	harvest	harvest	NOUN
alces-443	260	15	regimes	regime	NOUN
alces-443	260	16	can	can	AUX
alces-443	260	17	affect	affect	VERB
alces-443	260	18	genetic	genetic	ADJ
alces-443	260	19	characteristics	characteristic	NOUN
alces-443	260	20	of	of	ADP
alces-443	260	21	populations	population	NOUN
alces-443	260	22	and	and	CCONJ
alces-443	260	23	by	by	ADP
alces-443	260	24	extension	extension	NOUN
alces-443	260	25	may	may	AUX
alces-443	260	26	have	have	VERB
alces-443	260	27	an	an	DET
alces-443	260	28	influence	influence	NOUN
alces-443	260	29	on	on	ADP
alces-443	260	30	evolutionary	evolutionary	ADJ
alces-443	260	31	potential	potential	NOUN
alces-443	260	32	.	.	PUNCT
alces-443	261	1	conversely	conversely	ADV
alces-443	261	2	,	,	PUNCT
alces-443	261	3	cronin	cronin	PROPN
alces-443	261	4	et	et	PROPN
alces-443	261	5	al	al	PROPN
alces-443	261	6	.	.	PROPN
alces-443	261	7	(	(	PUNCT
alces-443	261	8	2001	2001	NUM
alces-443	261	9	)	)	PUNCT
alces-443	261	10	detected	detect	VERB
alces-443	261	11	no	no	DET
alces-443	261	12	differences	difference	NOUN
alces-443	261	13	in	in	ADP
alces-443	261	14	numbers	number	NOUN
alces-443	261	15	of	of	ADP
alces-443	261	16	alleles	allele	NOUN
alces-443	261	17	or	or	CCONJ
alces-443	261	18	levels	level	NOUN
alces-443	261	19	of	of	ADP
alces-443	261	20	heterozygosity	heterozygosity	NOUN
alces-443	261	21	for	for	ADP
alces-443	261	22	5	5	NUM
alces-443	261	23	microsatellite	microsatellite	ADJ
alces-443	261	24	loci	locus	NOUN
alces-443	261	25	among	among	ADP
alces-443	261	26	3	3	NUM
alces-443	261	27	moose	moose	NOUN
alces-443	261	28	populations	population	NOUN
alces-443	261	29	in	in	ADP
alces-443	261	30	quebec	quebec	PROPN
alces-443	261	31	,	,	PUNCT
alces-443	261	32	1	1	NUM
alces-443	261	33	heavily	heavily	ADV
alces-443	261	34	hunted	hunt	VERB
alces-443	261	35	,	,	PUNCT
alces-443	261	36	1	1	NUM
alces-443	261	37	lightly	lightly	ADV
alces-443	261	38	hunted	hunt	VERB
alces-443	261	39	,	,	PUNCT
alces-443	261	40	and	and	CCONJ
alces-443	261	41	1	1	NUM
alces-443	261	42	not	not	PART
alces-443	261	43	hunted	hunt	VERB
alces-443	261	44	.	.	PUNCT
alces-443	262	1	another	another	DET
alces-443	262	2	critical	critical	ADJ
alces-443	262	3	factor	factor	NOUN
alces-443	262	4	in	in	ADP
alces-443	262	5	management	management	NOUN
alces-443	262	6	is	be	AUX
alces-443	262	7	the	the	DET
alces-443	262	8	effect	effect	NOUN
alces-443	262	9	of	of	ADP
alces-443	262	10	harvest	harvest	NOUN
alces-443	262	11	on	on	ADP
alces-443	262	12	genetic	genetic	ADJ
alces-443	262	13	loci	locus	NOUN
alces-443	262	14	underlying	underlie	VERB
alces-443	262	15	characters	character	NOUN
alces-443	262	16	having	have	VERB
alces-443	262	17	a	a	DET
alces-443	262	18	direct	direct	ADJ
alces-443	262	19	effect	effect	NOUN
alces-443	262	20	on	on	ADP
alces-443	262	21	reproductive	reproductive	ADJ
alces-443	262	22	fitness	fitness	NOUN
alces-443	262	23	,	,	PUNCT
alces-443	262	24	e.g.	e.g.	ADV
alces-443	262	25	,	,	PUNCT
alces-443	262	26	antler	antler	NOUN
alces-443	262	27	size	size	NOUN
alces-443	262	28	in	in	ADP
alces-443	262	29	moose	moose	NOUN
alces-443	262	30	.	.	PUNCT
alces-443	263	1	controlling	control	VERB
alces-443	263	2	harvest	harvest	NOUN
alces-443	263	3	by	by	ADP
alces-443	263	4	defining	define	VERB
alces-443	263	5	legal	legal	ADJ
alces-443	263	6	males	male	NOUN
alces-443	263	7	according	accord	VERB
alces-443	263	8	to	to	ADP
alces-443	263	9	antler	antler	NOUN
alces-443	263	10	size	size	NOUN
alces-443	263	11	is	be	AUX
alces-443	263	12	common	common	ADJ
alces-443	263	13	in	in	ADP
alces-443	263	14	management	management	NOUN
alces-443	263	15	of	of	ADP
alces-443	263	16	north	north	ADJ
alces-443	263	17	american	american	PROPN
alces-443	263	18	elk	elk	PROPN
alces-443	263	19	(	(	PUNCT
alces-443	263	20	thelen	thelen	VERB
alces-443	263	21	1991	1991	NUM
alces-443	263	22	)	)	PUNCT
alces-443	263	23	and	and	CCONJ
alces-443	263	24	is	be	AUX
alces-443	263	25	a	a	DET
alces-443	263	26	strategy	strategy	NOUN
alces-443	263	27	employed	employ	VERB
alces-443	263	28	in	in	ADP
alces-443	263	29	moose	moose	NOUN
alces-443	263	30	management	management	NOUN
alces-443	263	31	in	in	ADP
alces-443	263	32	british	british	PROPN
alces-443	263	33	columbia	columbia	PROPN
alces-443	263	34	,	,	PUNCT
alces-443	263	35	canada	canada	PROPN
alces-443	263	36	,	,	PUNCT
alces-443	263	37	and	and	CCONJ
alces-443	263	38	alaska	alaska	PROPN
alces-443	263	39	,	,	PUNCT
alces-443	263	40	usa	usa	PROPN
alces-443	263	41	(	(	PUNCT
alces-443	263	42	child	child	PROPN
alces-443	263	43	1983	1983	NUM
alces-443	263	44	,	,	PUNCT
alces-443	263	45	schwartz	schwartz	PROPN
alces-443	263	46	et	et	PROPN
alces-443	263	47	al	al	PROPN
alces-443	263	48	.	.	PROPN
alces-443	263	49	1992	1992	NUM
alces-443	263	50	)	)	PUNCT
alces-443	263	51	.	.	PUNCT
alces-443	264	1	in	in	ADP
alces-443	264	2	an	an	DET
alces-443	264	3	effort	effort	NOUN
alces-443	264	4	to	to	PART
alces-443	264	5	evaluate	evaluate	VERB
alces-443	264	6	genetic	genetic	ADJ
alces-443	264	7	effects	effect	NOUN
alces-443	264	8	of	of	ADP
alces-443	264	9	the	the	DET
alces-443	264	10	selective	selective	ADJ
alces-443	264	11	harvest	harvest	NOUN
alces-443	264	12	system	system	NOUN
alces-443	264	13	in	in	ADP
alces-443	264	14	alaska	alaska	PROPN
alces-443	264	15	,	,	PUNCT
alces-443	264	16	hundertmark	hundertmark	PROPN
alces-443	264	17	et	et	PROPN
alces-443	264	18	al	al	PROPN
alces-443	264	19	.	.	PROPN
alces-443	265	1	(	(	PUNCT
alces-443	265	2	1993	1993	NUM
alces-443	265	3	,	,	PUNCT
alces-443	265	4	1998	1998	NUM
alces-443	265	5	)	)	PUNCT
alces-443	265	6	modeled	model	VERB
alces-443	265	7	moose	moose	NOUN
alces-443	265	8	populations	population	NOUN
alces-443	265	9	subject	subject	ADJ
alces-443	265	10	to	to	ADP
alces-443	265	11	harvest	harvest	NOUN
alces-443	265	12	strategies	strategy	NOUN
alces-443	265	13	employing	employ	VERB
alces-443	265	14	different	different	ADJ
alces-443	265	15	definitions	definition	NOUN
alces-443	265	16	of	of	ADP
alces-443	265	17	legal	legal	ADJ
alces-443	265	18	males	male	NOUN
alces-443	265	19	.	.	PUNCT
alces-443	266	1	they	they	PRON
alces-443	266	2	concluded	conclude	VERB
alces-443	266	3	that	that	SCONJ
alces-443	266	4	selective	selective	ADJ
alces-443	266	5	harvest	harvest	NOUN
alces-443	266	6	systems	system	NOUN
alces-443	266	7	could	could	AUX
alces-443	266	8	result	result	VERB
alces-443	266	9	in	in	ADP
alces-443	266	10	allele	allele	NOUN
alces-443	266	11	frequency	frequency	NOUN
alces-443	266	12	changes	change	NOUN
alces-443	266	13	at	at	ADP
alces-443	266	14	loci	loci	NOUN
alces-443	266	15	coding	code	VERB
alces-443	266	16	for	for	ADP
alces-443	266	17	antler	antler	NOUN
alces-443	266	18	characteristics	characteristic	NOUN
alces-443	266	19	(	(	PUNCT
alces-443	266	20	hundertmark	hundertmark	NOUN
alces-443	266	21	et	et	PROPN
alces-443	266	22	al	al	PROPN
alces-443	266	23	.	.	PROPN
alces-443	266	24	1993	1993	NUM
alces-443	266	25	)	)	PUNCT
alces-443	266	26	and	and	CCONJ
alces-443	266	27	that	that	SCONJ
alces-443	266	28	the	the	DET
alces-443	266	29	position	position	NOUN
alces-443	266	30	of	of	ADP
alces-443	266	31	a	a	DET
alces-443	266	32	population	population	NOUN
alces-443	266	33	relative	relative	ADJ
alces-443	266	34	to	to	ADP
alces-443	266	35	nutritional	nutritional	ADJ
alces-443	266	36	carrying	carrying	NOUN
alces-443	266	37	capacity	capacity	NOUN
alces-443	266	38	of	of	ADP
alces-443	266	39	the	the	DET
alces-443	266	40	habitat	habitat	NOUN
alces-443	266	41	affected	affect	VERB
alces-443	266	42	the	the	DET
alces-443	266	43	rate	rate	NOUN
alces-443	266	44	of	of	ADP
alces-443	266	45	change	change	NOUN
alces-443	266	46	in	in	ADP
alces-443	266	47	allele	allele	NOUN
alces-443	266	48	frequencies	frequency	NOUN
alces-443	266	49	(	(	PUNCT
alces-443	266	50	hundertmark	hundertmark	NOUN
alces-443	266	51	et	et	PROPN
alces-443	266	52	al	al	PROPN
alces-443	266	53	.	.	PROPN
alces-443	266	54	1998	1998	NUM
alces-443	266	55	)	)	PUNCT
alces-443	266	56	.	.	PUNCT
alces-443	267	1	those	those	DET
alces-443	267	2	results	result	NOUN
alces-443	267	3	indicated	indicate	VERB
alces-443	267	4	that	that	SCONJ
alces-443	267	5	limiting	limit	VERB
alces-443	267	6	harvest	harvest	NOUN
alces-443	267	7	to	to	PART
alces-443	267	8	moose	moose	VERB
alces-443	267	9	with	with	ADP
alces-443	267	10	large	large	ADJ
alces-443	267	11	antlers	antler	NOUN
alces-443	267	12	could	could	AUX
alces-443	267	13	cause	cause	VERB
alces-443	267	14	a	a	DET
alces-443	267	15	genetically	genetically	ADV
alces-443	267	16	based	base	VERB
alces-443	267	17	decrease	decrease	NOUN
alces-443	267	18	in	in	ADP
alces-443	267	19	antler	antler	ADJ
alces-443	267	20	size	size	NOUN
alces-443	267	21	over	over	ADP
alces-443	267	22	time	time	NOUN
alces-443	267	23	.	.	PUNCT
alces-443	268	1	such	such	DET
alces-443	268	2	a	a	DET
alces-443	268	3	reduction	reduction	NOUN
alces-443	268	4	in	in	ADP
alces-443	268	5	adaptive	adaptive	ADJ
alces-443	268	6	genetic	genetic	ADJ
alces-443	268	7	variation	variation	NOUN
alces-443	268	8	runs	run	VERB
alces-443	268	9	counter	counter	ADV
alces-443	268	10	to	to	ADP
alces-443	268	11	general	general	ADJ
alces-443	268	12	conservation	conservation	NOUN
alces-443	268	13	goals	goal	NOUN
alces-443	268	14	(	(	PUNCT
alces-443	268	15	lynch	lynch	PROPN
alces-443	268	16	1996	1996	NUM
alces-443	268	17	)	)	PUNCT
alces-443	268	18	.	.	PUNCT
alces-443	269	1	a	a	DET
alces-443	269	2	stunning	stunning	ADJ
alces-443	269	3	example	example	NOUN
alces-443	269	4	of	of	ADP
alces-443	269	5	the	the	DET
alces-443	269	6	effect	effect	NOUN
alces-443	269	7	of	of	ADP
alces-443	269	8	harvest	harvest	NOUN
alces-443	269	9	on	on	ADP
alces-443	269	10	fitness	fitness	NOUN
alces-443	269	11	traits	trait	NOUN
alces-443	269	12	was	be	AUX
alces-443	269	13	recently	recently	ADV
alces-443	269	14	reported	report	VERB
alces-443	269	15	by	by	ADP
alces-443	269	16	coltman	coltman	PROPN
alces-443	269	17	et	et	PROPN
alces-443	269	18	al	al	PROPN
alces-443	269	19	.	.	PROPN
alces-443	270	1	(	(	PUNCT
alces-443	270	2	2003	2003	NUM
alces-443	270	3	)	)	PUNCT
alces-443	270	4	,	,	PUNCT
alces-443	270	5	who	who	PRON
alces-443	270	6	documented	document	VERB
alces-443	270	7	significant	significant	ADJ
alces-443	270	8	genetic	genetic	ADJ
alces-443	270	9	effects	effect	NOUN
alces-443	270	10	on	on	ADP
alces-443	270	11	horn	horn	NOUN
alces-443	270	12	size	size	NOUN
alces-443	270	13	and	and	CCONJ
alces-443	270	14	body	body	NOUN
alces-443	270	15	mass	mass	NOUN
alces-443	270	16	in	in	ADP
alces-443	270	17	bighorn	bighorn	NOUN
alces-443	270	18	sheep	sheep	NOUN
alces-443	270	19	(	(	PUNCT
alces-443	270	20	ovis	ovis	ADJ
alces-443	270	21	canadensis	canadensis	NOUN
alces-443	270	22	)	)	PUNCT
alces-443	270	23	as	as	ADP
alces-443	270	24	a	a	DET
alces-443	270	25	result	result	NOUN
alces-443	270	26	of	of	ADP
alces-443	270	27	selective	selective	ADJ
alces-443	270	28	harvest	harvest	NOUN
alces-443	270	29	of	of	ADP
alces-443	270	30	males	male	NOUN
alces-443	270	31	with	with	ADP
alces-443	270	32	large	large	ADJ
alces-443	270	33	horns	horn	NOUN
alces-443	270	34	.	.	PUNCT
alces-443	271	1	detecting	detect	VERB
alces-443	271	2	potential	potential	ADJ
alces-443	271	3	changes	change	NOUN
alces-443	271	4	in	in	ADP
alces-443	271	5	genetic	genetic	ADJ
alces-443	271	6	composition	composition	NOUN
alces-443	271	7	of	of	ADP
alces-443	271	8	moose	moose	NOUN
alces-443	271	9	as	as	SCONJ
alces-443	271	10	they	they	PRON
alces-443	271	11	respond	respond	VERB
alces-443	271	12	to	to	ADP
alces-443	271	13	various	various	ADJ
alces-443	271	14	anthropogenic	anthropogenic	ADJ
alces-443	271	15	influences	influence	NOUN
alces-443	271	16	,	,	PUNCT
alces-443	271	17	whether	whether	SCONJ
alces-443	271	18	related	relate	VERB
alces-443	271	19	to	to	ADP
alces-443	271	20	management	management	NOUN
alces-443	271	21	or	or	CCONJ
alces-443	271	22	to	to	ADP
alces-443	271	23	changes	change	NOUN
alces-443	271	24	in	in	ADP
alces-443	271	25	the	the	DET
alces-443	271	26	environment	environment	NOUN
alces-443	271	27	,	,	PUNCT
alces-443	271	28	is	be	AUX
alces-443	271	29	a	a	DET
alces-443	271	30	difficult	difficult	ADJ
alces-443	271	31	task	task	NOUN
alces-443	271	32	.	.	PUNCT
alces-443	272	1	nonetheless	nonetheless	ADV
alces-443	272	2	,	,	PUNCT
alces-443	272	3	it	it	PRON
alces-443	272	4	is	be	AUX
alces-443	272	5	an	an	DET
alces-443	272	6	important	important	ADJ
alces-443	272	7	area	area	NOUN
alces-443	272	8	of	of	ADP
alces-443	272	9	investigation	investigation	NOUN
alces-443	272	10	and	and	CCONJ
alces-443	272	11	deserves	deserve	VERB
alces-443	272	12	attention	attention	NOUN
alces-443	272	13	.	.	PUNCT
alces-443	273	1	modeling	model	VERB
alces-443	273	2	exercises	exercise	NOUN
alces-443	273	3	and	and	CCONJ
alces-443	273	4	studies	study	NOUN
alces-443	273	5	of	of	ADP
alces-443	273	6	genetic	genetic	ADJ
alces-443	273	7	variation	variation	NOUN
alces-443	273	8	have	have	AUX
alces-443	273	9	not	not	PART
alces-443	273	10	addressed	address	VERB
alces-443	273	11	interrelationships	interrelationship	NOUN
alces-443	273	12	of	of	ADP
alces-443	273	13	moose	moose	NOUN
alces-443	273	14	populations	population	NOUN
alces-443	273	15	at	at	ADP
alces-443	273	16	fine	fine	ADJ
alces-443	273	17	scales	scale	NOUN
alces-443	273	18	.	.	PUNCT
alces-443	274	1	the	the	DET
alces-443	274	2	effects	effect	NOUN
alces-443	274	3	of	of	ADP
alces-443	274	4	management	management	NOUN
alces-443	274	5	and	and	CCONJ
alces-443	274	6	habitat	habitat	NOUN
alces-443	274	7	alteration	alteration	NOUN
alces-443	274	8	on	on	ADP
alces-443	274	9	processes	process	NOUN
alces-443	274	10	involved	involve	VERB
alces-443	274	11	in	in	ADP
alces-443	274	12	maintenance	maintenance	NOUN
alces-443	274	13	of	of	ADP
alces-443	274	14	connectivity	connectivity	NOUN
alces-443	274	15	of	of	ADP
alces-443	274	16	moose	moose	NOUN
alces-443	274	17	populations	population	NOUN
alces-443	274	18	,	,	PUNCT
alces-443	274	19	such	such	ADJ
alces-443	274	20	as	as	ADP
alces-443	274	21	malemediated	malemediate	VERB
alces-443	274	22	gene	gene	NOUN
alces-443	274	23	flow	flow	NOUN
alces-443	274	24	via	via	ADP
alces-443	274	25	yearling	yearling	NOUN
alces-443	274	26	dispersal	dispersal	NOUN
alces-443	274	27	are	be	AUX
alces-443	274	28	extremely	extremely	ADV
alces-443	274	29	important	important	ADJ
alces-443	274	30	and	and	CCONJ
alces-443	274	31	should	should	AUX
alces-443	274	32	be	be	AUX
alces-443	274	33	described	describe	VERB
alces-443	274	34	.	.	PUNCT
alces-443	275	1	moose	moose	PROPN
alces-443	275	2	are	be	AUX
alces-443	275	3	highly	highly	ADV
alces-443	275	4	adaptable	adaptable	ADJ
alces-443	275	5	animals	animal	NOUN
alces-443	275	6	,	,	PUNCT
alces-443	275	7	but	but	CCONJ
alces-443	275	8	the	the	DET
alces-443	275	9	intensive	intensive	ADJ
alces-443	275	10	management	management	NOUN
alces-443	275	11	of	of	ADP
alces-443	275	12	moose	moose	NOUN
alces-443	275	13	hundertmark	hundertmark	NOUN
alces-443	275	14	and	and	CCONJ
alces-443	275	15	bowyer	bowyer	PROPN
alces-443	275	16	moose	moose	PROPN
alces-443	275	17	genetics	genetics	PROPN
alces-443	275	18	alces	alce	NOUN
alces-443	275	19	vol	vol	NOUN
alces-443	275	20	.	.	PROPN
alces-443	275	21	40	40	NUM
alces-443	275	22	,	,	PUNCT
alces-443	275	23	2004	2004	NUM
alces-443	275	24	116	116	NUM
alces-443	275	25	populations	population	NOUN
alces-443	275	26	and	and	CCONJ
alces-443	275	27	the	the	DET
alces-443	275	28	environmental	environmental	ADJ
alces-443	275	29	factors	factor	NOUN
alces-443	275	30	influencing	influence	VERB
alces-443	275	31	their	their	PRON
alces-443	275	32	habitat	habitat	NOUN
alces-443	275	33	(	(	PUNCT
alces-443	275	34	e.g.	e.g.	ADV
alces-443	275	35	,	,	PUNCT
alces-443	275	36	wildfire	wildfire	NOUN
alces-443	275	37	)	)	PUNCT
alces-443	275	38	may	may	AUX
alces-443	275	39	have	have	VERB
alces-443	275	40	unintended	unintended	ADJ
alces-443	275	41	and	and	CCONJ
alces-443	275	42	significant	significant	ADJ
alces-443	275	43	consequences	consequence	NOUN
alces-443	275	44	on	on	ADP
alces-443	275	45	the	the	DET
alces-443	275	46	moose	moose	NOUN
alces-443	275	47	genome	genome	NOUN
alces-443	275	48	through	through	ADP
alces-443	275	49	a	a	DET
alces-443	275	50	change	change	NOUN
alces-443	275	51	in	in	ADP
alces-443	275	52	selective	selective	ADJ
alces-443	275	53	forces	force	NOUN
alces-443	275	54	.	.	PUNCT
alces-443	276	1	proper	proper	ADJ
alces-443	276	2	conservation	conservation	NOUN
alces-443	276	3	of	of	ADP
alces-443	276	4	this	this	DET
alces-443	276	5	species	species	NOUN
alces-443	276	6	requires	require	VERB
alces-443	276	7	that	that	SCONJ
alces-443	276	8	we	we	PRON
alces-443	276	9	recognize	recognize	VERB
alces-443	276	10	and	and	CCONJ
alces-443	276	11	avoid	avoid	VERB
alces-443	276	12	that	that	DET
alces-443	276	13	possibility	possibility	NOUN
alces-443	276	14	.	.	PUNCT
alces-443	277	1	acknowledgements	acknowledgement	NOUN
alces-443	277	2	we	we	PRON
alces-443	277	3	thank	thank	VERB
alces-443	277	4	the	the	DET
alces-443	277	5	organizers	organizer	NOUN
alces-443	277	6	of	of	ADP
alces-443	277	7	the	the	DET
alces-443	277	8	5th	5th	ADJ
alces-443	277	9	international	international	ADJ
alces-443	277	10	moose	moose	NOUN
alces-443	277	11	symposium	symposium	NOUN
alces-443	277	12	for	for	ADP
alces-443	277	13	the	the	DET
alces-443	277	14	opportunity	opportunity	NOUN
alces-443	277	15	to	to	PART
alces-443	277	16	synthesize	synthesize	VERB
alces-443	277	17	and	and	CCONJ
alces-443	277	18	present	present	VERB
alces-443	277	19	this	this	DET
alces-443	277	20	information	information	NOUN
alces-443	277	21	.	.	PUNCT
alces-443	278	1	we	we	PRON
alces-443	278	2	wish	wish	VERB
alces-443	278	3	to	to	PART
alces-443	278	4	acknowledge	acknowledge	VERB
alces-443	278	5	funding	funding	NOUN
alces-443	278	6	support	support	NOUN
alces-443	278	7	from	from	ADP
alces-443	278	8	federal	federal	ADJ
alces-443	278	9	aid	aid	NOUN
alces-443	278	10	in	in	ADP
alces-443	278	11	wildlife	wildlife	NOUN
alces-443	278	12	restoration	restoration	NOUN
alces-443	278	13	through	through	ADP
alces-443	278	14	the	the	DET
alces-443	278	15	alaska	alaska	PROPN
alces-443	278	16	department	department	PROPN
alces-443	278	17	of	of	ADP
alces-443	278	18	fish	fish	NOUN
alces-443	278	19	and	and	CCONJ
alces-443	278	20	game	game	NOUN
alces-443	278	21	as	as	ADV
alces-443	278	22	well	well	ADV
alces-443	278	23	as	as	ADP
alces-443	278	24	the	the	DET
alces-443	278	25	institute	institute	NOUN
alces-443	278	26	of	of	ADP
alces-443	278	27	arctic	arctic	PROPN
alces-443	278	28	biology	biology	NOUN
alces-443	278	29	,	,	PUNCT
alces-443	278	30	university	university	PROPN
alces-443	278	31	of	of	ADP
alces-443	278	32	alaska	alaska	PROPN
alces-443	278	33	fairbanks	fairbanks	PROPN
alces-443	278	34	.	.	PUNCT
alces-443	279	1	s.	s.	PROPN
alces-443	279	2	côté	côté	PROPN
alces-443	279	3	and	and	CCONJ
alces-443	279	4	an	an	DET
alces-443	279	5	anonymous	anonymous	ADJ
alces-443	279	6	reviewer	reviewer	NOUN
alces-443	279	7	provided	provide	VERB
alces-443	279	8	constructive	constructive	ADJ
alces-443	279	9	comments	comment	NOUN
alces-443	279	10	for	for	ADP
alces-443	279	11	improving	improve	VERB
alces-443	279	12	the	the	DET
alces-443	279	13	manuscript	manuscript	NOUN
alces-443	279	14	.	.	PUNCT
alces-443	280	1	references	reference	NOUN
alces-443	280	2	anderson	anderson	PROPN
alces-443	280	3	,	,	PUNCT
alces-443	280	4	p.	p.	PROPN
alces-443	280	5	m.	m.	NOUN
alces-443	280	6	,	,	PUNCT
alces-443	280	7	a.	a.	NOUN
alces-443	280	8	v.	v.	ADP
alces-443	280	9	lozhkin	lozhkin	PROPN
alces-443	280	10	,	,	PUNCT
alces-443	280	11	and	and	CCONJ
alces-443	280	12	l.	l.	PROPN
alces-443	280	13	b.	b.	PROPN
alces-443	280	14	brubaker	brubaker	PROPN
alces-443	280	15	.	.	PUNCT
alces-443	280	16	2002	2002	NUM
alces-443	280	17	.	.	PUNCT
alces-443	281	1	implications	implication	NOUN
alces-443	281	2	of	of	ADP
alces-443	281	3	a	a	DET
alces-443	281	4	24,000	24,000	NUM
alces-443	281	5	-	-	PUNCT
alces-443	281	6	yr	yr	NOUN
alces-443	281	7	palynological	palynological	ADJ
alces-443	281	8	record	record	NOUN
alces-443	281	9	for	for	ADP
alces-443	281	10	a	a	DET
alces-443	281	11	younger	young	ADJ
alces-443	281	12	dryas	dryas	NOUN
alces-443	281	13	cooling	cool	VERB
alces-443	281	14	and	and	CCONJ
alces-443	281	15	for	for	ADP
alces-443	281	16	boreal	boreal	NOUN
alces-443	281	17	forest	forest	NOUN
alces-443	281	18	development	development	NOUN
alces-443	281	19	in	in	ADP
alces-443	281	20	northeastern	northeastern	ADJ
alces-443	281	21	siberia	siberia	PROPN
alces-443	281	22	.	.	PUNCT
alces-443	282	1	quaternary	quaternary	ADJ
alces-443	282	2	research	research	NOUN
alces-443	282	3	57:325333	57:325333	ADJ
alces-443	282	4	.	.	PUNCT
alces-443	283	1	arkhipov	arkhipov	PROPN
alces-443	283	2	,	,	PUNCT
alces-443	283	3	s.	s.	PROPN
alces-443	283	4	a.	a.	PROPN
alces-443	283	5	,	,	PUNCT
alces-443	283	6	v.	v.	PROPN
alces-443	283	7	g.	g.	PROPN
alces-443	283	8	bespaly	bespaly	PROPN
alces-443	283	9	,	,	PUNCT
alces-443	283	10	m.	m.	NOUN
alces-443	283	11	a.	a.	PROPN
alces-443	283	12	faustova	faustova	PROPN
alces-443	283	13	,	,	PUNCT
alces-443	283	14	o.	o.	PROPN
alces-443	283	15	y.	y.	PROPN
alces-443	283	16	glushkova	glushkova	PROPN
alces-443	283	17	,	,	PUNCT
alces-443	283	18	l.	l.	PROPN
alces-443	283	19	l.	l.	PROPN
alces-443	283	20	isaeva	isaeva	PROPN
alces-443	283	21	,	,	PUNCT
alces-443	283	22	and	and	CCONJ
alces-443	283	23	a.	a.	NOUN
alces-443	283	24	a.	a.	NOUN
alces-443	283	25	velichko	velichko	PROPN
alces-443	283	26	.	.	PUNCT
alces-443	284	1	1986	1986	NUM
alces-443	284	2	.	.	PUNCT
alces-443	285	1	ice	ice	NOUN
alces-443	285	2	sheet	sheet	NOUN
alces-443	285	3	reconstructions	reconstruction	NOUN
alces-443	285	4	.	.	PUNCT
alces-443	286	1	pages	page	NOUN
alces-443	286	2	269	269	NUM
alces-443	286	3	-	-	SYM
alces-443	286	4	292	292	NUM
alces-443	286	5	in	in	ADP
alces-443	286	6	v.	v.	ADP
alces-443	286	7	šibrava	šibrava	PROPN
alces-443	286	8	,	,	PUNCT
alces-443	286	9	d.	d.	PROPN
alces-443	286	10	q.	q.	PROPN
alces-443	286	11	bowen	bowen	PROPN
alces-443	286	12	,	,	PUNCT
alces-443	286	13	and	and	CCONJ
alces-443	286	14	g.	g.	PROPN
alces-443	286	15	m.	m.	PROPN
alces-443	286	16	richmond	richmond	PROPN
alces-443	286	17	,	,	PUNCT
alces-443	286	18	editors	editor	NOUN
alces-443	286	19	.	.	PUNCT
alces-443	287	1	quaternary	quaternary	ADJ
alces-443	287	2	glaciations	glaciation	NOUN
alces-443	287	3	in	in	ADP
alces-443	287	4	the	the	DET
alces-443	287	5	northern	northern	ADJ
alces-443	287	6	hemisphere	hemisphere	NOUN
alces-443	287	7	.	.	PUNCT
alces-443	288	1	pergamon	pergamon	PROPN
alces-443	288	2	press	press	PROPN
alces-443	288	3	,	,	PUNCT
alces-443	288	4	oxford	oxford	PROPN
alces-443	288	5	,	,	PUNCT
alces-443	288	6	u.k	u.k	PROPN
alces-443	288	7	.	.	PROPN
alces-443	288	8	avise	avise	PROPN
alces-443	288	9	,	,	PUNCT
alces-443	288	10	j.	j.	PROPN
alces-443	288	11	c.	c.	PROPN
alces-443	288	12	2000	2000	NUM
alces-443	288	13	.	.	PUNCT
alces-443	289	1	phylogeography	phylogeography	NOUN
alces-443	289	2	:	:	PUNCT
alces-443	289	3	the	the	DET
alces-443	289	4	history	history	NOUN
alces-443	289	5	and	and	CCONJ
alces-443	289	6	formation	formation	NOUN
alces-443	289	7	of	of	ADP
alces-443	289	8	species	specie	NOUN
alces-443	289	9	.	.	PUNCT
alces-443	290	1	harvard	harvard	PROPN
alces-443	290	2	university	university	PROPN
alces-443	290	3	press	press	PROPN
alces-443	290	4	,	,	PUNCT
alces-443	290	5	cambridge	cambridge	PROPN
alces-443	290	6	,	,	PUNCT
alces-443	290	7	massachusetts	massachusetts	PROPN
alces-443	290	8	,	,	PUNCT
alces-443	290	9	usa	usa	PROPN
alces-443	290	10	.	.	PUNCT
alces-443	291	1	_	_	PUNCT
alces-443	292	1	_	_	PUNCT
alces-443	293	1	_	_	PUNCT
alces-443	294	1	_	_	PUNCT
alces-443	295	1	_	_	PUNCT
alces-443	295	2	,	,	PUNCT
alces-443	295	3	j.	j.	PROPN
alces-443	295	4	arnold	arnold	PROPN
alces-443	295	5	,	,	PUNCT
alces-443	295	6	r.	r.	PROPN
alces-443	295	7	m.	m.	PROPN
alces-443	295	8	ball	ball	PROPN
alces-443	295	9	,	,	PUNCT
alces-443	295	10	jr	jr	PROPN
alces-443	295	11	.	.	PROPN
alces-443	295	12	,	,	PUNCT
alces-443	295	13	e.	e.	PROPN
alces-443	295	14	bermingham	bermingham	PROPN
alces-443	295	15	,	,	PUNCT
alces-443	295	16	t.	t.	PROPN
alces-443	295	17	lamb	lamb	PROPN
alces-443	295	18	,	,	PUNCT
alces-443	295	19	j.	j.	PROPN
alces-443	295	20	e.	e.	PROPN
alces-443	295	21	neigel	neigel	PROPN
alces-443	295	22	,	,	PUNCT
alces-443	295	23	c.	c.	PROPN
alces-443	295	24	a.	a.	PROPN
alces-443	295	25	reeb	reeb	PROPN
alces-443	295	26	,	,	PUNCT
alces-443	295	27	and	and	CCONJ
alces-443	295	28	n.	n.	PROPN
alces-443	295	29	c.	c.	PROPN
alces-443	295	30	saunders	saunders	PROPN
alces-443	295	31	.	.	PUNCT
alces-443	296	1	1987	1987	NUM
alces-443	296	2	.	.	PUNCT
alces-443	297	1	intraspecific	intraspecific	NOUN
alces-443	297	2	phylogeography	phylogeography	NOUN
alces-443	297	3	:	:	PUNCT
alces-443	297	4	the	the	DET
alces-443	297	5	mitochondrial	mitochondrial	ADJ
alces-443	297	6	dna	dna	PROPN
alces-443	297	7	bridge	bridge	NOUN
alces-443	297	8	between	between	ADP
alces-443	297	9	population	population	NOUN
alces-443	297	10	genetics	genetic	NOUN
alces-443	297	11	and	and	CCONJ
alces-443	297	12	systematics	systematic	NOUN
alces-443	297	13	.	.	PUNCT
alces-443	298	1	annual	annual	ADJ
alces-443	298	2	review	review	NOUN
alces-443	298	3	of	of	ADP
alces-443	298	4	ecology	ecology	NOUN
alces-443	298	5	and	and	CCONJ
alces-443	298	6	systematics	systematic	NOUN
alces-443	298	7	18:489	18:489	NUM
alces-443	298	8	-	-	SYM
alces-443	298	9	522	522	NUM
alces-443	298	10	.	.	PUNCT
alces-443	299	1	birky	birky	NOUN
alces-443	299	2	,	,	PUNCT
alces-443	299	3	c.	c.	PROPN
alces-443	299	4	w.	w.	PROPN
alces-443	299	5	,	,	PUNCT
alces-443	299	6	t.	t.	PROPN
alces-443	299	7	maruyama	maruyama	NOUN
alces-443	299	8	,	,	PUNCT
alces-443	299	9	and	and	CCONJ
alces-443	299	10	p.	p.	PROPN
alces-443	299	11	fuerst	fuerst	PROPN
alces-443	299	12	.	.	PUNCT
alces-443	300	1	1983	1983	NUM
alces-443	300	2	.	.	PUNCT
alces-443	301	1	an	an	DET
alces-443	301	2	approach	approach	NOUN
alces-443	301	3	to	to	ADP
alces-443	301	4	population	population	NOUN
alces-443	301	5	and	and	CCONJ
alces-443	301	6	evolutionary	evolutionary	ADJ
alces-443	301	7	genetic	genetic	ADJ
alces-443	301	8	theory	theory	NOUN
alces-443	301	9	for	for	ADP
alces-443	301	10	genes	gene	NOUN
alces-443	301	11	in	in	ADP
alces-443	301	12	mitochondria	mitochondria	NOUN
alces-443	301	13	and	and	CCONJ
alces-443	301	14	chloroplasts	chloroplast	NOUN
alces-443	301	15	,	,	PUNCT
alces-443	301	16	and	and	CCONJ
alces-443	301	17	some	some	DET
alces-443	301	18	results	result	NOUN
alces-443	301	19	.	.	PUNCT
alces-443	302	1	genetics	genetic	NOUN
alces-443	302	2	103:513	103:513	PROPN
alces-443	302	3	-	-	SYM
alces-443	302	4	527	527	NUM
alces-443	302	5	.	.	PUNCT
alces-443	303	1	blundell	blundell	PROPN
alces-443	303	2	,	,	PUNCT
alces-443	303	3	g.	g.	PROPN
alces-443	303	4	m.	m.	PROPN
alces-443	303	5	,	,	PUNCT
alces-443	303	6	m.	m.	PROPN
alces-443	303	7	ben	ben	PROPN
alces-443	303	8	-	-	PROPN
alces-443	303	9	david	david	PROPN
alces-443	303	10	,	,	PUNCT
alces-443	303	11	p.	p.	NOUN
alces-443	303	12	groves	grove	NOUN
alces-443	303	13	,	,	PUNCT
alces-443	303	14	r.	r.	PROPN
alces-443	303	15	t.	t.	PROPN
alces-443	303	16	bowyer	bowyer	PROPN
alces-443	303	17	,	,	PUNCT
alces-443	303	18	and	and	CCONJ
alces-443	303	19	e.	e.	PROPN
alces-443	303	20	geffen	geffen	PROPN
alces-443	303	21	.	.	PUNCT
alces-443	304	1	2002	2002	NUM
alces-443	304	2	.	.	PUNCT
alces-443	305	1	characteristics	characteristic	NOUN
alces-443	305	2	of	of	ADP
alces-443	305	3	sex	sex	NOUN
alces-443	305	4	-	-	PUNCT
alces-443	305	5	biased	bias	VERB
alces-443	305	6	dispersal	dispersal	NOUN
alces-443	305	7	and	and	CCONJ
alces-443	305	8	gene	gene	NOUN
alces-443	305	9	flow	flow	NOUN
alces-443	305	10	in	in	ADP
alces-443	305	11	coastal	coastal	ADJ
alces-443	305	12	river	river	NOUN
alces-443	305	13	otters	otter	NOUN
alces-443	305	14	:	:	PUNCT
alces-443	305	15	implications	implication	NOUN
alces-443	305	16	for	for	ADP
alces-443	305	17	natural	natural	ADJ
alces-443	305	18	recolonization	recolonization	NOUN
alces-443	305	19	of	of	ADP
alces-443	305	20	extirpated	extirpated	ADJ
alces-443	305	21	populations	population	NOUN
alces-443	305	22	.	.	PUNCT
alces-443	306	1	molecular	molecular	ADJ
alces-443	306	2	ecology	ecology	NOUN
alces-443	306	3	11:289	11:289	NUM
alces-443	306	4	-	-	SYM
alces-443	306	5	303	303	NUM
alces-443	306	6	.	.	PUNCT
alces-443	306	7	boeskorov	boeskorov	PROPN
alces-443	306	8	,	,	PUNCT
alces-443	306	9	g.	g.	PROPN
alces-443	306	10	g.	g.	PROPN
alces-443	306	11	1996	1996	NUM
alces-443	306	12	.	.	PUNCT
alces-443	307	1	karyotype	karyotype	NOUN
alces-443	307	2	of	of	ADP
alces-443	307	3	moose	moose	NOUN
alces-443	307	4	(	(	PUNCT
alces-443	307	5	alces	alces	PROPN
alces-443	307	6	alces	alce	NOUN
alces-443	307	7	l.	l.	PROPN
alces-443	307	8	)	)	PUNCT
alces-443	307	9	from	from	ADP
alces-443	307	10	northeastern	northeastern	ADJ
alces-443	307	11	asia	asia	PROPN
alces-443	307	12	.	.	PUNCT
alces-443	308	1	proceedings	proceeding	NOUN
alces-443	308	2	of	of	ADP
alces-443	308	3	the	the	DET
alces-443	308	4	russian	russian	PROPN
alces-443	308	5	academy	academy	PROPN
alces-443	308	6	of	of	ADP
alces-443	308	7	sciences	sciences	PROPN
alces-443	308	8	329:506508	329:506508	NUM
alces-443	308	9	.	.	PUNCT
alces-443	309	1	(	(	PUNCT
alces-443	309	2	in	in	ADP
alces-443	309	3	russian	russian	NOUN
alces-443	309	4	)	)	PUNCT
alces-443	309	5	.	.	PUNCT
alces-443	310	1	_	_	PUNCT
alces-443	311	1	_	_	PUNCT
alces-443	312	1	_	_	PUNCT
alces-443	313	1	_	_	PUNCT
alces-443	314	1	_	_	PUNCT
alces-443	314	2	.	.	PUNCT
alces-443	315	1	1997	1997	NUM
alces-443	315	2	.	.	PUNCT
alces-443	316	1	chromosomal	chromosomal	ADJ
alces-443	316	2	differences	difference	NOUN
alces-443	316	3	in	in	ADP
alces-443	316	4	moose	moose	NOUN
alces-443	316	5	.	.	PUNCT
alces-443	317	1	genetika	genetika	NOUN
alces-443	317	2	33:974	33:974	NUM
alces-443	317	3	-	-	SYM
alces-443	317	4	978	978	NUM
alces-443	317	5	.	.	PUNCT
alces-443	318	1	(	(	PUNCT
alces-443	318	2	in	in	ADP
alces-443	318	3	russian	russian	NOUN
alces-443	318	4	)	)	PUNCT
alces-443	318	5	.	.	PUNCT
alces-443	319	1	bowyer	bowyer	PROPN
alces-443	319	2	,	,	PUNCT
alces-443	319	3	r.	r.	PROPN
alces-443	319	4	t.	t.	PROPN
alces-443	319	5	,	,	PUNCT
alces-443	319	6	d.	d.	PROPN
alces-443	319	7	m.	m.	PROPN
alces-443	319	8	leslie	leslie	PROPN
alces-443	319	9	,	,	PUNCT
alces-443	319	10	jr	jr	PROPN
alces-443	319	11	.	.	PROPN
alces-443	319	12	,	,	PUNCT
alces-443	319	13	and	and	CCONJ
alces-443	319	14	j.	j.	PROPN
alces-443	319	15	l.	l.	PROPN
alces-443	319	16	rachlow	rachlow	PROPN
alces-443	319	17	.	.	PUNCT
alces-443	320	1	2000	2000	NUM
alces-443	320	2	.	.	PUNCT
alces-443	321	1	dall	dall	PROPN
alces-443	321	2	’s	’s	PART
alces-443	321	3	and	and	CCONJ
alces-443	321	4	stone	stone	NOUN
alces-443	321	5	’s	’s	PART
alces-443	321	6	sheep	sheep	NOUN
alces-443	321	7	.	.	PUNCT
alces-443	322	1	pages	page	NOUN
alces-443	322	2	491	491	NUM
alces-443	322	3	-	-	SYM
alces-443	322	4	516	516	NUM
alces-443	322	5	in	in	ADP
alces-443	322	6	s.	s.	PROPN
alces-443	322	7	demarais	demarais	PROPN
alces-443	322	8	and	and	CCONJ
alces-443	322	9	p.	p.	PROPN
alces-443	322	10	r.	r.	PROPN
alces-443	322	11	krausman	krausman	PROPN
alces-443	322	12	,	,	PUNCT
alces-443	322	13	editors	editor	NOUN
alces-443	322	14	.	.	PUNCT
alces-443	323	1	ecology	ecology	NOUN
alces-443	323	2	and	and	CCONJ
alces-443	323	3	management	management	NOUN
alces-443	323	4	of	of	ADP
alces-443	323	5	large	large	ADJ
alces-443	323	6	mammals	mammal	NOUN
alces-443	323	7	in	in	ADP
alces-443	323	8	north	north	PROPN
alces-443	323	9	america	america	PROPN
alces-443	323	10	.	.	PUNCT
alces-443	324	1	prentice	prentice	PROPN
alces-443	324	2	hall	hall	PROPN
alces-443	324	3	,	,	PUNCT
alces-443	324	4	upper	upper	ADJ
alces-443	324	5	saddle	saddle	NOUN
alces-443	324	6	river	river	NOUN
alces-443	324	7	,	,	PUNCT
alces-443	324	8	new	new	PROPN
alces-443	324	9	jersey	jersey	PROPN
alces-443	324	10	,	,	PUNCT
alces-443	324	11	usa	usa	PROPN
alces-443	324	12	.	.	PUNCT
alces-443	325	1	_	_	PUNCT
alces-443	326	1	_	_	PUNCT
alces-443	327	1	_	_	PUNCT
alces-443	328	1	_	_	PUNCT
alces-443	329	1	_	_	PUNCT
alces-443	329	2	,	,	PUNCT
alces-443	329	3	j.	j.	PROPN
alces-443	329	4	l.	l.	PROPN
alces-443	329	5	rachlow	rachlow	PROPN
alces-443	329	6	,	,	PUNCT
alces-443	329	7	v.	v.	PROPN
alces-443	329	8	van	van	PROPN
alces-443	329	9	ballenberghe	ballenberghe	NOUN
alces-443	329	10	,	,	PUNCT
alces-443	329	11	and	and	CCONJ
alces-443	329	12	r.	r.	PROPN
alces-443	329	13	d.	d.	PROPN
alces-443	329	14	guthrie	guthrie	PROPN
alces-443	329	15	.	.	PROPN
alces-443	329	16	1991	1991	NUM
alces-443	329	17	.	.	PUNCT
alces-443	330	1	evolution	evolution	NOUN
alces-443	330	2	of	of	ADP
alces-443	330	3	a	a	DET
alces-443	330	4	rump	rump	NOUN
alces-443	330	5	patch	patch	NOUN
alces-443	330	6	in	in	ADP
alces-443	330	7	moose	moose	NOUN
alces-443	330	8	:	:	PUNCT
alces-443	330	9	an	an	DET
alces-443	330	10	hypothesis	hypothesis	NOUN
alces-443	330	11	.	.	PUNCT
alces-443	331	1	alces	alce	NOUN
alces-443	331	2	27:12	27:12	NUM
alces-443	331	3	-	-	SYM
alces-443	331	4	23	23	NUM
alces-443	331	5	.	.	PUNCT
alces-443	332	1	_	_	PUNCT
alces-443	333	1	_	_	PUNCT
alces-443	334	1	_	_	PUNCT
alces-443	335	1	_	_	PUNCT
alces-443	336	1	_	_	PUNCT
alces-443	336	2	,	,	PUNCT
alces-443	336	3	k.	k.	PROPN
alces-443	336	4	m.	m.	PROPN
alces-443	336	5	stewart	stewart	PROPN
alces-443	336	6	,	,	PUNCT
alces-443	336	7	b.	b.	PROPN
alces-443	336	8	m.	m.	PROPN
alces-443	336	9	pierce	pierce	PROPN
alces-443	336	10	,	,	PUNCT
alces-443	336	11	k.	k.	PROPN
alces-443	336	12	j.	j.	PROPN
alces-443	336	13	hundertmark	hundertmark	PROPN
alces-443	336	14	,	,	PUNCT
alces-443	336	15	and	and	CCONJ
alces-443	336	16	w.	w.	PROPN
alces-443	336	17	c.	c.	PROPN
alces-443	336	18	gasaway	gasaway	PROPN
alces-443	336	19	.	.	PUNCT
alces-443	337	1	2002	2002	NUM
alces-443	337	2	.	.	PUNCT
alces-443	338	1	geographical	geographical	ADJ
alces-443	338	2	variation	variation	NOUN
alces-443	338	3	in	in	ADP
alces-443	338	4	antler	antler	NOUN
alces-443	338	5	morphology	morphology	NOUN
alces-443	338	6	of	of	ADP
alces-443	338	7	alaskan	alaskan	ADJ
alces-443	338	8	moose	moose	NOUN
alces-443	338	9	:	:	PUNCT
alces-443	338	10	putative	putative	ADJ
alces-443	338	11	effects	effect	NOUN
alces-443	338	12	of	of	ADP
alces-443	338	13	habitat	habitat	NOUN
alces-443	338	14	and	and	CCONJ
alces-443	338	15	genetics	genetic	NOUN
alces-443	338	16	.	.	PUNCT
alces-443	339	1	alces	alce	NOUN
alces-443	339	2	38	38	NUM
alces-443	339	3	:	:	SYM
alces-443	339	4	155	155	NUM
alces-443	339	5	-	-	SYM
alces-443	339	6	165	165	NUM
alces-443	339	7	.	.	PUNCT
alces-443	340	1	_	_	PUNCT
alces-443	341	1	_	_	PUNCT
alces-443	342	1	_	_	PUNCT
alces-443	343	1	_	_	PUNCT
alces-443	344	1	_	_	PUNCT
alces-443	344	2	,	,	PUNCT
alces-443	344	3	v.	v.	PROPN
alces-443	344	4	van	van	PROPN
alces-443	344	5	ballenberghe	ballenberghe	NOUN
alces-443	344	6	,	,	PUNCT
alces-443	344	7	and	and	CCONJ
alces-443	344	8	j.g	j.g	PROPN
alces-443	344	9	.	.	PROPN
alces-443	344	10	kie	kie	PROPN
alces-443	344	11	.	.	PUNCT
alces-443	345	1	2003	2003	NUM
alces-443	345	2	.	.	PUNCT
alces-443	346	1	moose	moose	NOUN
alces-443	346	2	(	(	PUNCT
alces-443	346	3	alces	alce	NOUN
alces-443	346	4	alces	alce	NOUN
alces-443	346	5	)	)	PUNCT
alces-443	346	6	.	.	PUNCT
alces-443	347	1	pages	page	NOUN
alces-443	347	2	931	931	NUM
alces-443	347	3	964	964	NUM
alces-443	347	4	in	in	ADP
alces-443	347	5	g.a	g.a	PROPN
alces-443	347	6	.	.	PROPN
alces-443	347	7	feldhamer	feldhamer	PROPN
alces-443	347	8	,	,	PUNCT
alces-443	347	9	b.	b.	PROPN
alces-443	347	10	thompson	thompson	PROPN
alces-443	347	11	,	,	PUNCT
alces-443	347	12	and	and	CCONJ
alces-443	347	13	j.	j.	PROPN
alces-443	347	14	chapman	chapman	PROPN
alces-443	347	15	,	,	PUNCT
alces-443	347	16	editors	editor	NOUN
alces-443	347	17	.	.	PUNCT
alces-443	348	1	wild	wild	ADJ
alces-443	348	2	mammals	mammal	NOUN
alces-443	348	3	of	of	ADP
alces-443	348	4	north	north	PROPN
alces-443	348	5	america	america	PROPN
alces-443	348	6	:	:	PUNCT
alces-443	348	7	biology	biology	NOUN
alces-443	348	8	,	,	PUNCT
alces-443	348	9	management	management	NOUN
alces-443	348	10	,	,	PUNCT
alces-443	348	11	and	and	CCONJ
alces-443	348	12	economics	economic	NOUN
alces-443	348	13	.	.	PUNCT
alces-443	349	1	secalces	secalce	NOUN
alces-443	349	2	vol	vol	NOUN
alces-443	349	3	.	.	PROPN
alces-443	349	4	40	40	NUM
alces-443	349	5	,	,	PUNCT
alces-443	349	6	2004	2004	NUM
alces-443	349	7	moose	moose	NOUN
alces-443	349	8	genetics	genetic	NOUN
alces-443	349	9	hundertmark	hundertmark	NOUN
alces-443	349	10	and	and	CCONJ
alces-443	349	11	bowyer	bowyer	PROPN
alces-443	349	12	117	117	NUM
alces-443	349	13	ond	ond	PROPN
alces-443	349	14	edition	edition	NOUN
alces-443	349	15	.	.	PUNCT
alces-443	350	1	the	the	DET
alces-443	350	2	johns	johns	PROPN
alces-443	350	3	hopkins	hopkins	PROPN
alces-443	350	4	university	university	PROPN
alces-443	350	5	press	press	PROPN
alces-443	350	6	,	,	PUNCT
alces-443	350	7	baltimore	baltimore	PROPN
alces-443	350	8	,	,	PUNCT
alces-443	350	9	maryland	maryland	PROPN
alces-443	350	10	,	,	PUNCT
alces-443	350	11	usa	usa	PROPN
alces-443	350	12	.	.	PROPN
alces-443	350	13	braend	braend	PROPN
alces-443	350	14	,	,	PUNCT
alces-443	350	15	m.	m.	NOUN
alces-443	350	16	1962	1962	NUM
alces-443	350	17	.	.	PUNCT
alces-443	351	1	studies	study	NOUN
alces-443	351	2	on	on	ADP
alces-443	351	3	blood	blood	NOUN
alces-443	351	4	and	and	CCONJ
alces-443	351	5	serum	serum	NOUN
alces-443	351	6	groups	group	NOUN
alces-443	351	7	in	in	ADP
alces-443	351	8	the	the	DET
alces-443	351	9	elk	elk	NOUN
alces-443	351	10	(	(	PUNCT
alces-443	351	11	alces	alce	NOUN
alces-443	351	12	alces	alce	NOUN
alces-443	351	13	)	)	PUNCT
alces-443	351	14	.	.	PUNCT
alces-443	352	1	new	new	PROPN
alces-443	352	2	york	york	PROPN
alces-443	352	3	academy	academy	PROPN
alces-443	352	4	of	of	ADP
alces-443	352	5	sciences	sciences	PROPN
alces-443	352	6	97:296305	97:296305	PROPN
alces-443	352	7	.	.	PUNCT
alces-443	353	1	broders	broder	NOUN
alces-443	353	2	,	,	PUNCT
alces-443	353	3	h.	h.	PROPN
alces-443	353	4	g.	g.	PROPN
alces-443	353	5	,	,	PUNCT
alces-443	353	6	s.	s.	PROPN
alces-443	353	7	p.	p.	PROPN
alces-443	353	8	mahoney	mahoney	PROPN
alces-443	353	9	,	,	PUNCT
alces-443	353	10	w.	w.	PROPN
alces-443	353	11	a.	a.	PROPN
alces-443	353	12	montevecchi	montevecchi	PROPN
alces-443	353	13	,	,	PUNCT
alces-443	353	14	and	and	CCONJ
alces-443	353	15	w.	w.	PROPN
alces-443	353	16	s.	s.	PROPN
alces-443	353	17	davidson	davidson	PROPN
alces-443	353	18	.	.	PUNCT
alces-443	353	19	1999	1999	NUM
alces-443	353	20	.	.	PUNCT
alces-443	354	1	population	population	NOUN
alces-443	354	2	genetic	genetic	ADJ
alces-443	354	3	structure	structure	NOUN
alces-443	354	4	and	and	CCONJ
alces-443	354	5	the	the	DET
alces-443	354	6	effect	effect	NOUN
alces-443	354	7	of	of	ADP
alces-443	354	8	founder	founder	NOUN
alces-443	354	9	events	event	NOUN
alces-443	354	10	on	on	ADP
alces-443	354	11	the	the	DET
alces-443	354	12	genetic	genetic	ADJ
alces-443	354	13	variability	variability	NOUN
alces-443	354	14	of	of	ADP
alces-443	354	15	moose	moose	NOUN
alces-443	354	16	,	,	PUNCT
alces-443	354	17	alces	alce	NOUN
alces-443	354	18	alces	alce	NOUN
alces-443	354	19	,	,	PUNCT
alces-443	354	20	in	in	ADP
alces-443	354	21	canada	canada	PROPN
alces-443	354	22	.	.	PUNCT
alces-443	355	1	molecular	molecular	ADJ
alces-443	355	2	ecology	ecology	NOUN
alces-443	355	3	8:1309	8:1309	PROPN
alces-443	355	4	-	-	SYM
alces-443	355	5	1315	1315	NUM
alces-443	355	6	.	.	PUNCT
alces-443	356	1	bruford	bruford	PROPN
alces-443	356	2	,	,	PUNCT
alces-443	356	3	m.	m.	PROPN
alces-443	356	4	w.	w.	PROPN
alces-443	356	5	,	,	PUNCT
alces-443	356	6	d.	d.	PROPN
alces-443	356	7	j.	j.	PROPN
alces-443	356	8	cheesman	cheesman	PROPN
alces-443	356	9	,	,	PUNCT
alces-443	356	10	t.	t.	PROPN
alces-443	356	11	coote	coote	PROPN
alces-443	356	12	,	,	PUNCT
alces-443	356	13	h.	h.	PROPN
alces-443	356	14	a.	a.	PROPN
alces-443	356	15	a.	a.	PROPN
alces-443	356	16	green	green	PROPN
alces-443	356	17	,	,	PUNCT
alces-443	356	18	s.	s.	PROPN
alces-443	356	19	a.	a.	NOUN
alces-443	356	20	haines	haines	PROPN
alces-443	356	21	,	,	PUNCT
alces-443	356	22	c.	c.	PROPN
alces-443	356	23	o’ryan	o’ryan	PROPN
alces-443	356	24	,	,	PUNCT
alces-443	356	25	and	and	CCONJ
alces-443	356	26	t.	t.	PROPN
alces-443	356	27	r.	r.	PROPN
alces-443	356	28	williams	williams	PROPN
alces-443	356	29	.	.	PUNCT
alces-443	356	30	1996	1996	NUM
alces-443	356	31	.	.	PUNCT
alces-443	357	1	microsatellites	microsatellite	NOUN
alces-443	357	2	and	and	CCONJ
alces-443	357	3	their	their	PRON
alces-443	357	4	application	application	NOUN
alces-443	357	5	to	to	ADP
alces-443	357	6	conservation	conservation	NOUN
alces-443	357	7	genetics	genetic	NOUN
alces-443	357	8	.	.	PUNCT
alces-443	358	1	pages	page	NOUN
alces-443	358	2	278	278	NUM
alces-443	358	3	-	-	SYM
alces-443	358	4	297	297	NUM
alces-443	358	5	in	in	ADP
alces-443	358	6	t.	t.	PROPN
alces-443	358	7	b.	b.	PROPN
alces-443	358	8	smith	smith	PROPN
alces-443	358	9	and	and	CCONJ
alces-443	358	10	r.	r.	PROPN
alces-443	358	11	k.	k.	PROPN
alces-443	358	12	wayne	wayne	PROPN
alces-443	358	13	,	,	PUNCT
alces-443	358	14	editors	editor	NOUN
alces-443	358	15	.	.	PUNCT
alces-443	359	1	molecular	molecular	ADJ
alces-443	359	2	genetic	genetic	ADJ
alces-443	359	3	approaches	approach	NOUN
alces-443	359	4	in	in	ADP
alces-443	359	5	conservation	conservation	NOUN
alces-443	359	6	.	.	PUNCT
alces-443	360	1	oxford	oxford	PROPN
alces-443	360	2	university	university	PROPN
alces-443	360	3	press	press	NOUN
alces-443	360	4	,	,	PUNCT
alces-443	360	5	oxford	oxford	PROPN
alces-443	360	6	,	,	PUNCT
alces-443	360	7	u.k	u.k	PROPN
alces-443	360	8	.	.	PROPN
alces-443	360	9	brumfield	brumfield	PROPN
alces-443	360	10	,	,	PUNCT
alces-443	360	11	r.	r.	PROPN
alces-443	360	12	t.	t.	PROPN
alces-443	360	13	,	,	PUNCT
alces-443	360	14	p.	p.	NOUN
alces-443	360	15	beerli	beerli	PROPN
alces-443	360	16	,	,	PUNCT
alces-443	360	17	d.	d.	PROPN
alces-443	360	18	a.	a.	PROPN
alces-443	360	19	nickerson	nickerson	PROPN
alces-443	360	20	,	,	PUNCT
alces-443	360	21	and	and	CCONJ
alces-443	360	22	s.	s.	PROPN
alces-443	360	23	v.	v.	PROPN
alces-443	360	24	edwards	edwards	PROPN
alces-443	360	25	.	.	PROPN
alces-443	360	26	2003	2003	NUM
alces-443	360	27	.	.	PUNCT
alces-443	361	1	the	the	DET
alces-443	361	2	utili	utili	PROPN
alces-443	361	3	ty	ty	INTJ
alces-443	361	4	of	of	ADP
alces-443	361	5	single	single	ADJ
alces-443	361	6	nucleotide	nucleotide	ADJ
alces-443	361	7	polymorphisms	polymorphism	NOUN
alces-443	361	8	in	in	ADP
alces-443	361	9	inferences	inference	NOUN
alces-443	361	10	of	of	ADP
alces-443	361	11	population	population	NOUN
alces-443	361	12	history	history	NOUN
alces-443	361	13	.	.	PUNCT
alces-443	362	1	trends	trend	NOUN
alces-443	362	2	in	in	ADP
alces-443	362	3	ecology	ecology	NOUN
alces-443	362	4	and	and	CCONJ
alces-443	362	5	evolution	evolution	NOUN
alces-443	362	6	18:249	18:249	NUM
alces-443	362	7	-	-	SYM
alces-443	362	8	256	256	NUM
alces-443	362	9	.	.	PUNCT
alces-443	362	10	bubenik	bubenik	NOUN
alces-443	362	11	,	,	PUNCT
alces-443	362	12	a.	a.	PROPN
alces-443	362	13	b.	b.	PROPN
alces-443	362	14	1998	1998	NUM
alces-443	362	15	.	.	PUNCT
alces-443	363	1	evolution	evolution	NOUN
alces-443	363	2	,	,	PUNCT
alces-443	363	3	taxonomy	taxonomy	NOUN
alces-443	363	4	and	and	CCONJ
alces-443	363	5	morphophysiology	morphophysiology	NOUN
alces-443	363	6	.	.	PUNCT
alces-443	364	1	pages	page	NOUN
alces-443	364	2	77	77	NUM
alces-443	364	3	-	-	SYM
alces-443	364	4	123	123	NUM
alces-443	364	5	in	in	ADP
alces-443	364	6	a.	a.	NOUN
alces-443	364	7	w.	w.	PROPN
alces-443	364	8	franzmann	franzmann	PROPN
alces-443	364	9	and	and	CCONJ
alces-443	364	10	c.	c.	PROPN
alces-443	364	11	c.	c.	PROPN
alces-443	364	12	schwartz	schwartz	PROPN
alces-443	364	13	,	,	PUNCT
alces-443	364	14	editors	editor	NOUN
alces-443	364	15	.	.	PUNCT
alces-443	365	1	ecology	ecology	NOUN
alces-443	365	2	and	and	CCONJ
alces-443	365	3	management	management	NOUN
alces-443	365	4	of	of	ADP
alces-443	365	5	the	the	DET
alces-443	365	6	north	north	ADJ
alces-443	365	7	american	american	ADJ
alces-443	365	8	moose	moose	NOUN
alces-443	365	9	.	.	PUNCT
alces-443	366	1	smithsonian	smithsonian	PROPN
alces-443	366	2	institution	institution	PROPN
alces-443	366	3	press	press	PROPN
alces-443	366	4	,	,	PUNCT
alces-443	366	5	washington	washington	PROPN
alces-443	366	6	,	,	PUNCT
alces-443	366	7	d.c	d.c	PROPN
alces-443	366	8	.	.	PROPN
alces-443	366	9	,	,	PUNCT
alces-443	366	10	usa	usa	PROPN
alces-443	366	11	.	.	PROPN
alces-443	366	12	chesser	chesser	PROPN
alces-443	366	13	,	,	PUNCT
alces-443	366	14	r.	r.	PROPN
alces-443	366	15	k.	k.	PROPN
alces-443	366	16	,	,	PUNCT
alces-443	366	17	c.	c.	PROPN
alces-443	366	18	reuterwall	reuterwall	NOUN
alces-443	366	19	,	,	PUNCT
alces-443	366	20	and	and	CCONJ
alces-443	366	21	n.	n.	PROPN
alces-443	366	22	ryman	ryman	PROPN
alces-443	366	23	.	.	PUNCT
alces-443	367	1	1982	1982	NUM
alces-443	367	2	.	.	PUNCT
alces-443	368	1	genetic	genetic	ADJ
alces-443	368	2	differentiation	differentiation	NOUN
alces-443	368	3	of	of	ADP
alces-443	368	4	scandinavian	scandinavian	ADJ
alces-443	368	5	moose	moose	NOUN
alces-443	368	6	alces	alce	NOUN
alces-443	368	7	alces	alce	NOUN
alces-443	368	8	populations	population	NOUN
alces-443	368	9	over	over	ADP
alces-443	368	10	short	short	ADJ
alces-443	368	11	geographic	geographic	ADJ
alces-443	368	12	distances	distance	NOUN
alces-443	368	13	.	.	PUNCT
alces-443	369	1	oikos	oikos	PROPN
alces-443	369	2	39:125	39:125	NUM
alces-443	369	3	-	-	PUNCT
alces-443	369	4	130	130	NUM
alces-443	369	5	.	.	PUNCT
alces-443	369	6	child	child	NOUN
alces-443	369	7	,	,	PUNCT
alces-443	369	8	k.	k.	PROPN
alces-443	369	9	n.	n.	PROPN
alces-443	369	10	1983	1983	NUM
alces-443	369	11	.	.	PUNCT
alces-443	370	1	selective	selective	ADJ
alces-443	370	2	harvest	harvest	NOUN
alces-443	370	3	of	of	ADP
alces-443	370	4	moose	moose	NOUN
alces-443	370	5	in	in	ADP
alces-443	370	6	the	the	DET
alces-443	370	7	omineca	omineca	NOUN
alces-443	370	8	:	:	PUNCT
alces-443	370	9	some	some	DET
alces-443	370	10	preliminary	preliminary	ADJ
alces-443	370	11	results	result	NOUN
alces-443	370	12	.	.	PUNCT
alces-443	371	1	alces	alce	NOUN
alces-443	371	2	19:162	19:162	NUM
alces-443	371	3	-	-	SYM
alces-443	371	4	177	177	NUM
alces-443	371	5	.	.	PUNCT
alces-443	372	1	coady	coady	PROPN
alces-443	372	2	,	,	PUNCT
alces-443	372	3	j.	j.	PROPN
alces-443	372	4	w.	w.	PROPN
alces-443	372	5	1982	1982	NUM
alces-443	372	6	.	.	PUNCT
alces-443	373	1	moose	moose	PROPN
alces-443	373	2	.	.	PUNCT
alces-443	374	1	pages	page	NOUN
alces-443	374	2	902	902	NUM
alces-443	374	3	-	-	SYM
alces-443	374	4	922	922	NUM
alces-443	374	5	in	in	ADP
alces-443	374	6	j.	j.	PROPN
alces-443	374	7	a.	a.	PROPN
alces-443	374	8	chapman	chapman	PROPN
alces-443	374	9	and	and	CCONJ
alces-443	374	10	g.	g.	PROPN
alces-443	374	11	a.	a.	PROPN
alces-443	374	12	feldhamer	feldhamer	PROPN
alces-443	374	13	,	,	PUNCT
alces-443	374	14	editors	editor	NOUN
alces-443	374	15	.	.	PUNCT
alces-443	375	1	wild	wild	ADJ
alces-443	375	2	mammals	mammal	NOUN
alces-443	375	3	of	of	ADP
alces-443	375	4	north	north	PROPN
alces-443	375	5	america	america	PROPN
alces-443	375	6	:	:	PUNCT
alces-443	375	7	biology	biology	NOUN
alces-443	375	8	,	,	PUNCT
alces-443	375	9	management	management	NOUN
alces-443	375	10	,	,	PUNCT
alces-443	375	11	and	and	CCONJ
alces-443	375	12	economics	economic	NOUN
alces-443	375	13	.	.	PUNCT
alces-443	376	1	johns	johns	PROPN
alces-443	376	2	hopkins	hopkins	PROPN
alces-443	376	3	university	university	PROPN
alces-443	376	4	press	press	PROPN
alces-443	376	5	,	,	PUNCT
alces-443	376	6	baltimore	baltimore	PROPN
alces-443	376	7	,	,	PUNCT
alces-443	376	8	maryland	maryland	PROPN
alces-443	376	9	,	,	PUNCT
alces-443	376	10	usa	usa	PROPN
alces-443	376	11	.	.	PROPN
alces-443	376	12	coltman	coltman	PROPN
alces-443	376	13	,	,	PUNCT
alces-443	376	14	d.	d.	PROPN
alces-443	376	15	w.	w.	PROPN
alces-443	376	16	,	,	PUNCT
alces-443	376	17	p.	p.	PROPN
alces-443	376	18	o’donoghue	o’donoghue	PROPN
alces-443	376	19	,	,	PUNCT
alces-443	376	20	j.	j.	PROPN
alces-443	376	21	t.	t.	PROPN
alces-443	376	22	jorgenson	jorgenson	PROPN
alces-443	376	23	,	,	PUNCT
alces-443	376	24	j.	j.	PROPN
alces-443	376	25	t.	t.	PROPN
alces-443	376	26	hogg	hogg	PROPN
alces-443	376	27	,	,	PUNCT
alces-443	376	28	c.	c.	PROPN
alces-443	376	29	strobeck	strobeck	NOUN
alces-443	376	30	,	,	PUNCT
alces-443	376	31	and	and	CCONJ
alces-443	376	32	m.	m.	NOUN
alces-443	376	33	festa	festa	NOUN
alces-443	376	34	-	-	NOUN
alces-443	376	35	bianchet	bianchet	NOUN
alces-443	376	36	.	.	PUNCT
alces-443	377	1	2003	2003	NUM
alces-443	377	2	.	.	PUNCT
alces-443	378	1	undesirable	undesirable	ADJ
alces-443	378	2	evolutionary	evolutionary	ADJ
alces-443	378	3	consequences	consequence	NOUN
alces-443	378	4	of	of	ADP
alces-443	378	5	trophy	trophy	NOUN
alces-443	378	6	hunting	hunting	NOUN
alces-443	378	7	.	.	PUNCT
alces-443	379	1	nature	nature	NOUN
alces-443	379	2	426:655	426:655	NOUN
alces-443	379	3	-	-	PUNCT
alces-443	379	4	658	658	NUM
alces-443	379	5	.	.	PUNCT
alces-443	380	1	cronin	cronin	PROPN
alces-443	380	2	,	,	PUNCT
alces-443	380	3	m.	m.	NOUN
alces-443	380	4	a.	a.	PROPN
alces-443	380	5	1992	1992	NUM
alces-443	380	6	.	.	PUNCT
alces-443	381	1	intraspecific	intraspecific	NOUN
alces-443	381	2	variation	variation	NOUN
alces-443	381	3	in	in	ADP
alces-443	381	4	mitochondrial	mitochondrial	ADJ
alces-443	381	5	dna	dna	NOUN
alces-443	381	6	of	of	ADP
alces-443	381	7	north	north	ADJ
alces-443	381	8	american	american	PROPN
alces-443	381	9	cervids	cervid	NOUN
alces-443	381	10	.	.	PUNCT
alces-443	382	1	journal	journal	PROPN
alces-443	382	2	of	of	ADP
alces-443	382	3	mammalogy	mammalogy	NOUN
alces-443	382	4	73:70	73:70	NUM
alces-443	382	5	-	-	SYM
alces-443	382	6	82	82	NUM
alces-443	382	7	.	.	PUNCT
alces-443	383	1	_	_	PUNCT
alces-443	384	1	_	_	PUNCT
alces-443	385	1	_	_	PUNCT
alces-443	386	1	_	_	PUNCT
alces-443	387	1	_	_	PUNCT
alces-443	387	2	,	,	PUNCT
alces-443	387	3	j.	j.	PROPN
alces-443	387	4	c.	c.	PROPN
alces-443	387	5	patton	patton	PROPN
alces-443	387	6	,	,	PUNCT
alces-443	387	7	r.	r.	PROPN
alces-443	387	8	courtois	courtois	PROPN
alces-443	387	9	,	,	PUNCT
alces-443	387	10	and	and	CCONJ
alces-443	387	11	m.	m.	NOUN
alces-443	387	12	crête	crête	PROPN
alces-443	387	13	.	.	PUNCT
alces-443	388	1	2001	2001	NUM
alces-443	388	2	.	.	PUNCT
alces-443	389	1	genetic	genetic	ADJ
alces-443	389	2	variation	variation	NOUN
alces-443	389	3	of	of	ADP
alces-443	389	4	microsatellite	microsatellite	ADJ
alces-443	389	5	dna	dna	NOUN
alces-443	389	6	in	in	ADP
alces-443	389	7	moose	moose	NOUN
alces-443	389	8	in	in	ADP
alces-443	389	9	québec	québec	PROPN
alces-443	389	10	.	.	PUNCT
alces-443	390	1	alces	alce	NOUN
alces-443	390	2	37:175	37:175	NUM
alces-443	390	3	-	-	SYM
alces-443	390	4	187	187	NUM
alces-443	390	5	.	.	PUNCT
alces-443	391	1	dyke	dyke	PROPN
alces-443	391	2	,	,	PUNCT
alces-443	391	3	a.	a.	NOUN
alces-443	391	4	s.	s.	PROPN
alces-443	391	5	,	,	PUNCT
alces-443	391	6	and	and	CCONJ
alces-443	391	7	v.	v.	ADP
alces-443	391	8	k.	k.	PROPN
alces-443	391	9	prest	prest	PROPN
alces-443	391	10	.	.	PUNCT
alces-443	392	1	1987	1987	NUM
alces-443	392	2	.	.	PUNCT
alces-443	393	1	late	late	ADJ
alces-443	393	2	wisconsinan	wisconsinan	PROPN
alces-443	393	3	and	and	CCONJ
alces-443	393	4	holocene	holocene	PROPN
alces-443	393	5	history	history	NOUN
alces-443	393	6	of	of	ADP
alces-443	393	7	the	the	DET
alces-443	393	8	laurentide	laurentide	ADJ
alces-443	393	9	ice	ice	NOUN
alces-443	393	10	sheet	sheet	NOUN
alces-443	393	11	.	.	PUNCT
alces-443	394	1	geographie	geographie	PROPN
alces-443	394	2	physique	physique	PROPN
alces-443	394	3	et	et	PROPN
alces-443	394	4	quaternaire	quaternaire	VERB
alces-443	394	5	41:237	41:237	NUM
alces-443	394	6	-	-	SYM
alces-443	394	7	263	263	NUM
alces-443	394	8	.	.	PUNCT
alces-443	395	1	elias	elias	PROPN
alces-443	395	2	,	,	PUNCT
alces-443	395	3	s.	s.	PROPN
alces-443	395	4	a.	a.	PROPN
alces-443	395	5	2000	2000	NUM
alces-443	395	6	.	.	PUNCT
alces-443	396	1	late	late	ADJ
alces-443	396	2	pleistocene	pleistocene	PROPN
alces-443	396	3	climates	climate	NOUN
alces-443	396	4	of	of	ADP
alces-443	396	5	beringia	beringia	NOUN
alces-443	396	6	,	,	PUNCT
alces-443	396	7	based	base	VERB
alces-443	396	8	on	on	ADP
alces-443	396	9	analysis	analysis	NOUN
alces-443	396	10	of	of	ADP
alces-443	396	11	fossil	fossil	ADJ
alces-443	396	12	beetles	beetle	NOUN
alces-443	396	13	.	.	PUNCT
alces-443	397	1	quaternary	quaternary	ADJ
alces-443	397	2	research	research	NOUN
alces-443	397	3	53:229	53:229	NUM
alces-443	397	4	-	-	SYM
alces-443	397	5	235	235	NUM
alces-443	397	6	.	.	PUNCT
alces-443	398	1	ellegren	ellegren	PROPN
alces-443	398	2	,	,	PUNCT
alces-443	398	3	h.	h.	PROPN
alces-443	398	4	,	,	PUNCT
alces-443	398	5	l.	l.	PROPN
alces-443	398	6	andersson	andersson	PROPN
alces-443	398	7	,	,	PUNCT
alces-443	398	8	and	and	CCONJ
alces-443	398	9	k.	k.	PROPN
alces-443	398	10	wallin	wallin	PROPN
alces-443	398	11	.	.	PUNCT
alces-443	399	1	1991	1991	NUM
alces-443	399	2	.	.	PUNCT
alces-443	400	1	dna	dna	PROPN
alces-443	400	2	polymorphism	polymorphism	NOUN
alces-443	400	3	in	in	ADP
alces-443	400	4	moose	moose	NOUN
alces-443	400	5	(	(	PUNCT
alces-443	400	6	alces	alce	NOUN
alces-443	400	7	alces	alce	NOUN
alces-443	400	8	)	)	PUNCT
alces-443	400	9	revealed	reveal	VERB
alces-443	400	10	by	by	ADP
alces-443	400	11	polynucleotide	polynucleotide	ADJ
alces-443	400	12	probe	probe	NOUN
alces-443	400	13	(	(	PUNCT
alces-443	400	14	tc)n	tc)n	PROPN
alces-443	400	15	.	.	PROPN
alces-443	400	16	journal	journal	PROPN
alces-443	400	17	of	of	ADP
alces-443	400	18	heredity	heredity	NOUN
alces-443	400	19	82:429	82:429	PROPN
alces-443	400	20	-	-	PUNCT
alces-443	400	21	431	431	NUM
alces-443	400	22	.	.	PUNCT
alces-443	401	1	_	_	PUNCT
alces-443	402	1	_	_	PUNCT
alces-443	403	1	_	_	PUNCT
alces-443	404	1	_	_	PUNCT
alces-443	405	1	_	_	PUNCT
alces-443	405	2	,	,	PUNCT
alces-443	405	3	s.	s.	PROPN
alces-443	405	4	mikko	mikko	PROPN
alces-443	405	5	,	,	PUNCT
alces-443	405	6	k.	k.	PROPN
alces-443	405	7	wallin	wallin	PROPN
alces-443	405	8	,	,	PUNCT
alces-443	405	9	and	and	CCONJ
alces-443	405	10	l.	l.	PROPN
alces-443	405	11	andersson	andersson	PROPN
alces-443	405	12	.	.	PUNCT
alces-443	405	13	1996	1996	NUM
alces-443	405	14	.	.	PUNCT
alces-443	406	1	limited	limited	ADJ
alces-443	406	2	polymorphism	polymorphism	NOUN
alces-443	406	3	at	at	ADP
alces-443	406	4	major	major	ADJ
alces-443	406	5	histocompatibility	histocompatibility	NOUN
alces-443	406	6	complex	complex	NOUN
alces-443	406	7	(	(	PUNCT
alces-443	406	8	mhc	mhc	PROPN
alces-443	406	9	)	)	PUNCT
alces-443	406	10	loci	loci	NOUN
alces-443	406	11	in	in	ADP
alces-443	406	12	the	the	DET
alces-443	406	13	swedish	swedish	PROPN
alces-443	406	14	moose	moose	PROPN
alces-443	406	15	a.	a.	NOUN
alces-443	406	16	alces	alce	NOUN
alces-443	406	17	.	.	PUNCT
alces-443	407	1	molecular	molecular	ADJ
alces-443	407	2	ecology	ecology	NOUN
alces-443	407	3	5:3	5:3	NUM
alces-443	407	4	-	-	SYM
alces-443	407	5	9	9	NUM
alces-443	407	6	.	.	PUNCT
alces-443	407	7	flerov	flerov	PROPN
alces-443	407	8	,	,	PUNCT
alces-443	407	9	k.	k.	PROPN
alces-443	407	10	k.	k.	PROPN
alces-443	407	11	1952	1952	NUM
alces-443	407	12	.	.	PUNCT
alces-443	408	1	fauna	fauna	NOUN
alces-443	408	2	of	of	ADP
alces-443	408	3	the	the	DET
alces-443	408	4	ussr	ussr	ADJ
alces-443	408	5	:	:	PUNCT
alces-443	408	6	mammals	mammal	NOUN
alces-443	408	7	.	.	PUNCT
alces-443	409	1	vol	vol	NOUN
alces-443	409	2	.	.	NOUN
alces-443	409	3	1	1	NUM
alces-443	409	4	no	no	DET
alces-443	409	5	2	2	NUM
alces-443	409	6	.	.	PUNCT
alces-443	409	7	musk	musk	ADJ
alces-443	409	8	deer	deer	NOUN
alces-443	409	9	and	and	CCONJ
alces-443	409	10	deer	deer	NOUN
alces-443	409	11	.	.	PUNCT
alces-443	410	1	the	the	DET
alces-443	410	2	academy	academy	PROPN
alces-443	410	3	of	of	ADP
alces-443	410	4	sciences	sciences	PROPN
alces-443	410	5	of	of	ADP
alces-443	410	6	the	the	DET
alces-443	410	7	ussr	ussr	PROPN
alces-443	410	8	,	,	PUNCT
alces-443	410	9	leningrad	leningrad	PROPN
alces-443	410	10	.	.	PUNCT
alces-443	411	1	(	(	PUNCT
alces-443	411	2	english	english	ADJ
alces-443	411	3	translation	translation	NOUN
alces-443	411	4	by	by	ADP
alces-443	411	5	the	the	DET
alces-443	411	6	israel	israel	PROPN
alces-443	411	7	program	program	NOUN
alces-443	411	8	for	for	ADP
alces-443	411	9	scientific	scientific	ADJ
alces-443	411	10	translation	translation	NOUN
alces-443	411	11	,	,	PUNCT
alces-443	411	12	1960	1960	NUM
alces-443	411	13	,	,	PUNCT
alces-443	411	14	s.	s.	PROPN
alces-443	411	15	monson	monson	PROPN
alces-443	411	16	,	,	PUNCT
alces-443	411	17	jerusalem	jerusalem	PROPN
alces-443	411	18	)	)	PUNCT
alces-443	411	19	.	.	PUNCT
alces-443	412	1	franzmann	franzmann	PROPN
alces-443	412	2	,	,	PUNCT
alces-443	412	3	a.	a.	PROPN
alces-443	412	4	w.	w.	PROPN
alces-443	412	5	,	,	PUNCT
alces-443	412	6	and	and	CCONJ
alces-443	412	7	c.	c.	PROPN
alces-443	412	8	c.	c.	PROPN
alces-443	412	9	schwartz	schwartz	PROPN
alces-443	412	10	,	,	PUNCT
alces-443	412	11	editors	editor	NOUN
alces-443	412	12	.	.	PUNCT
alces-443	413	1	1998	1998	NUM
alces-443	413	2	.	.	PUNCT
alces-443	414	1	ecology	ecology	NOUN
alces-443	414	2	and	and	CCONJ
alces-443	414	3	management	management	NOUN
alces-443	414	4	of	of	ADP
alces-443	414	5	the	the	DET
alces-443	414	6	north	north	ADJ
alces-443	414	7	american	american	ADJ
alces-443	414	8	moose	moose	NOUN
alces-443	414	9	.	.	PUNCT
alces-443	415	1	smithsonian	smithsonian	PROPN
alces-443	415	2	institution	institution	PROPN
alces-443	415	3	press	press	PROPN
alces-443	415	4	,	,	PUNCT
alces-443	415	5	washhundertmark	washhundertmark	NOUN
alces-443	415	6	and	and	CCONJ
alces-443	415	7	bowyer	bowyer	PROPN
alces-443	415	8	moose	moose	PROPN
alces-443	415	9	genetics	genetics	PROPN
alces-443	415	10	alces	alce	NOUN
alces-443	415	11	vol	vol	NOUN
alces-443	415	12	.	.	PROPN
alces-443	415	13	40	40	NUM
alces-443	415	14	,	,	PUNCT
alces-443	415	15	2004	2004	NUM
alces-443	415	16	118	118	NUM
alces-443	415	17	ington	ington	NOUN
alces-443	415	18	,	,	PUNCT
alces-443	415	19	d.c	d.c	PROPN
alces-443	415	20	.	.	PROPN
alces-443	415	21	,	,	PUNCT
alces-443	415	22	usa	usa	PROPN
alces-443	415	23	.	.	PROPN
alces-443	415	24	fries	fries	PROPN
alces-443	415	25	,	,	PUNCT
alces-443	415	26	r.	r.	PROPN
alces-443	415	27	,	,	PUNCT
alces-443	415	28	and	and	CCONJ
alces-443	415	29	g.	g.	PROPN
alces-443	415	30	durstewitz	durstewitz	PROPN
alces-443	415	31	.	.	PUNCT
alces-443	416	1	2001	2001	NUM
alces-443	416	2	.	.	PUNCT
alces-443	417	1	digitaldna	digitaldna	PROPN
alces-443	417	2	signatures	signature	VERB
alces-443	417	3	:	:	PUNCT
alces-443	417	4	snps	snps	NOUN
alces-443	417	5	for	for	ADP
alces-443	417	6	animal	animal	NOUN
alces-443	417	7	tagging	tagging	NOUN
alces-443	417	8	.	.	PUNCT
alces-443	418	1	nature	nature	NOUN
alces-443	418	2	biotechnology	biotechnology	PROPN
alces-443	418	3	19:508	19:508	PROPN
alces-443	418	4	.	.	PUNCT
alces-443	419	1	fulton	fulton	PROPN
alces-443	419	2	,	,	PUNCT
alces-443	419	3	r.	r.	PROPN
alces-443	419	4	j.	j.	PROPN
alces-443	419	5	,	,	PUNCT
alces-443	419	6	m.	m.	PROPN
alces-443	419	7	m.	m.	PROPN
alces-443	419	8	fenton	fenton	PROPN
alces-443	419	9	,	,	PUNCT
alces-443	419	10	and	and	CCONJ
alces-443	419	11	n.	n.	PROPN
alces-443	419	12	w.	w.	PROPN
alces-443	419	13	rutter	rutter	PROPN
alces-443	419	14	.	.	PUNCT
alces-443	420	1	1986	1986	NUM
alces-443	420	2	.	.	PUNCT
alces-443	421	1	summary	summary	NOUN
alces-443	421	2	of	of	ADP
alces-443	421	3	quaternary	quaternary	ADJ
alces-443	421	4	stratigraphy	stratigraphy	NOUN
alces-443	421	5	and	and	CCONJ
alces-443	421	6	history	history	NOUN
alces-443	421	7	,	,	PUNCT
alces-443	421	8	western	western	PROPN
alces-443	421	9	canada	canada	PROPN
alces-443	421	10	.	.	PUNCT
alces-443	422	1	pages	page	NOUN
alces-443	422	2	229	229	NUM
alces-443	422	3	-	-	SYM
alces-443	422	4	242	242	NUM
alces-443	422	5	in	in	ADP
alces-443	422	6	v.	v.	ADP
alces-443	422	7	šibrava	šibrava	PROPN
alces-443	422	8	,	,	PUNCT
alces-443	422	9	d.	d.	PROPN
alces-443	422	10	q.	q.	PROPN
alces-443	422	11	bowen	bowen	PROPN
alces-443	422	12	,	,	PUNCT
alces-443	422	13	and	and	CCONJ
alces-443	422	14	g.	g.	PROPN
alces-443	422	15	m.	m.	PROPN
alces-443	422	16	richmond	richmond	PROPN
alces-443	422	17	,	,	PUNCT
alces-443	422	18	editors	editor	NOUN
alces-443	422	19	.	.	PUNCT
alces-443	423	1	quaternary	quaternary	ADJ
alces-443	423	2	glaciations	glaciation	NOUN
alces-443	423	3	in	in	ADP
alces-443	423	4	the	the	DET
alces-443	423	5	northern	northern	ADJ
alces-443	423	6	hemisphere	hemisphere	NOUN
alces-443	423	7	.	.	PUNCT
alces-443	424	1	pergamon	pergamon	PROPN
alces-443	424	2	press	press	PROPN
alces-443	424	3	,	,	PUNCT
alces-443	424	4	oxford	oxford	PROPN
alces-443	424	5	,	,	PUNCT
alces-443	424	6	u.k	u.k	PROPN
alces-443	424	7	.	.	PROPN
alces-443	424	8	geist	geist	PROPN
alces-443	424	9	,	,	PUNCT
alces-443	424	10	v.	v.	ADP
alces-443	424	11	1987a	1987a	NUM
alces-443	424	12	.	.	PUNCT
alces-443	425	1	on	on	ADP
alces-443	425	2	the	the	DET
alces-443	425	3	evolution	evolution	NOUN
alces-443	425	4	and	and	CCONJ
alces-443	425	5	adaptations	adaptation	NOUN
alces-443	425	6	of	of	ADP
alces-443	425	7	alces	alce	NOUN
alces-443	425	8	.	.	PUNCT
alces-443	426	1	swedish	swedish	ADJ
alces-443	426	2	wildlife	wildlife	NOUN
alces-443	426	3	research	research	NOUN
alces-443	426	4	supplement	supplement	VERB
alces-443	426	5	1:11	1:11	NUM
alces-443	426	6	-	-	SYM
alces-443	426	7	23	23	NUM
alces-443	426	8	.	.	PUNCT
alces-443	427	1	_	_	PUNCT
alces-443	428	1	_	_	PUNCT
alces-443	429	1	_	_	PUNCT
alces-443	430	1	_	_	PUNCT
alces-443	431	1	_	_	PUNCT
alces-443	431	2	.	.	PUNCT
alces-443	432	1	1987b	1987b	PROPN
alces-443	432	2	.	.	PUNCT
alces-443	433	1	on	on	ADP
alces-443	433	2	speciation	speciation	NOUN
alces-443	433	3	in	in	ADP
alces-443	433	4	ice	ice	NOUN
alces-443	433	5	age	age	NOUN
alces-443	433	6	mammals	mammal	NOUN
alces-443	433	7	,	,	PUNCT
alces-443	433	8	with	with	ADP
alces-443	433	9	special	special	ADJ
alces-443	433	10	reference	reference	NOUN
alces-443	433	11	to	to	ADP
alces-443	433	12	cervids	cervid	NOUN
alces-443	433	13	and	and	CCONJ
alces-443	433	14	caprids	caprid	NOUN
alces-443	433	15	.	.	PUNCT
alces-443	434	1	canadian	canadian	ADJ
alces-443	434	2	journal	journal	PROPN
alces-443	434	3	of	of	ADP
alces-443	434	4	zoology	zoology	NOUN
alces-443	434	5	65:1067	65:1067	PROPN
alces-443	434	6	-	-	PUNCT
alces-443	434	7	1084	1084	NUM
alces-443	434	8	.	.	PUNCT
alces-443	435	1	_	_	PUNCT
alces-443	436	1	_	_	PUNCT
alces-443	437	1	_	_	PUNCT
alces-443	438	1	_	_	PUNCT
alces-443	439	1	_	_	PUNCT
alces-443	439	2	.	.	PUNCT
alces-443	440	1	1998	1998	NUM
alces-443	440	2	.	.	PUNCT
alces-443	441	1	deer	deer	NOUN
alces-443	441	2	of	of	ADP
alces-443	441	3	the	the	DET
alces-443	441	4	world	world	NOUN
alces-443	441	5	:	:	PUNCT
alces-443	441	6	their	their	PRON
alces-443	441	7	evolution	evolution	NOUN
alces-443	441	8	,	,	PUNCT
alces-443	441	9	behavior	behavior	NOUN
alces-443	441	10	,	,	PUNCT
alces-443	441	11	and	and	CCONJ
alces-443	441	12	ecology	ecology	NOUN
alces-443	441	13	.	.	PUNCT
alces-443	442	1	stackpole	stackpole	PROPN
alces-443	442	2	books	books	PROPN
alces-443	442	3	,	,	PUNCT
alces-443	442	4	mechanicsburg	mechanicsburg	PROPN
alces-443	442	5	,	,	PUNCT
alces-443	442	6	pennsylvania	pennsylvania	PROPN
alces-443	442	7	,	,	PUNCT
alces-443	442	8	usa	usa	PROPN
alces-443	442	9	.	.	PROPN
alces-443	442	10	groves	grove	NOUN
alces-443	442	11	,	,	PUNCT
alces-443	442	12	c.	c.	PROPN
alces-443	442	13	p.	p.	PROPN
alces-443	442	14	,	,	PUNCT
alces-443	442	15	and	and	CCONJ
alces-443	442	16	p.	p.	PROPN
alces-443	442	17	grubb	grubb	PROPN
alces-443	442	18	.	.	PUNCT
alces-443	443	1	1987	1987	NUM
alces-443	443	2	.	.	PUNCT
alces-443	444	1	relationships	relationship	NOUN
alces-443	444	2	of	of	ADP
alces-443	444	3	living	live	VERB
alces-443	444	4	deer	deer	NOUN
alces-443	444	5	.	.	PUNCT
alces-443	445	1	pages	page	NOUN
alces-443	445	2	3	3	NUM
alces-443	445	3	-	-	SYM
alces-443	445	4	59	59	NUM
alces-443	445	5	in	in	ADP
alces-443	445	6	c.	c.	PROPN
alces-443	445	7	m.	m.	PROPN
alces-443	445	8	wemmer	wemmer	PROPN
alces-443	445	9	,	,	PUNCT
alces-443	445	10	editor	editor	NOUN
alces-443	445	11	.	.	PUNCT
alces-443	446	1	biology	biology	NOUN
alces-443	446	2	and	and	CCONJ
alces-443	446	3	management	management	NOUN
alces-443	446	4	of	of	ADP
alces-443	446	5	the	the	DET
alces-443	446	6	cervidae	cervidae	NOUN
alces-443	446	7	.	.	PUNCT
alces-443	447	1	smithsonian	smithsonian	PROPN
alces-443	447	2	institution	institution	PROPN
alces-443	447	3	press	press	PROPN
alces-443	447	4	,	,	PUNCT
alces-443	447	5	washington	washington	PROPN
alces-443	447	6	,	,	PUNCT
alces-443	447	7	d.c	d.c	PROPN
alces-443	447	8	.	.	PROPN
alces-443	447	9	,	,	PUNCT
alces-443	447	10	usa	usa	PROPN
alces-443	447	11	.	.	PROPN
alces-443	447	12	guthrie	guthrie	PROPN
alces-443	447	13	,	,	PUNCT
alces-443	447	14	r.	r.	PROPN
alces-443	447	15	d.	d.	PROPN
alces-443	447	16	1984	1984	NUM
alces-443	447	17	.	.	PUNCT
alces-443	448	1	alaska	alaska	PROPN
alces-443	448	2	megabucks	megabuck	NOUN
alces-443	448	3	,	,	PUNCT
alces-443	448	4	megabulls	megabull	NOUN
alces-443	448	5	,	,	PUNCT
alces-443	448	6	and	and	CCONJ
alces-443	448	7	megarams	megaram	NOUN
alces-443	448	8	:	:	PUNCT
alces-443	448	9	the	the	DET
alces-443	448	10	issue	issue	NOUN
alces-443	448	11	of	of	ADP
alces-443	448	12	pleistocene	pleistocene	NOUN
alces-443	448	13	gigantism	gigantism	NOUN
alces-443	448	14	.	.	PUNCT
alces-443	449	1	special	special	ADJ
alces-443	449	2	publications	publication	NOUN
alces-443	449	3	of	of	ADP
alces-443	449	4	the	the	DET
alces-443	449	5	carnegie	carnegie	NOUN
alces-443	449	6	museum	museum	PROPN
alces-443	449	7	of	of	ADP
alces-443	449	8	natural	natural	ADJ
alces-443	449	9	history	history	NOUN
alces-443	449	10	8:482	8:482	NUM
alces-443	449	11	-	-	SYM
alces-443	449	12	509	509	NUM
alces-443	449	13	.	.	PUNCT
alces-443	450	1	_	_	PUNCT
alces-443	451	1	_	_	PUNCT
alces-443	452	1	_	_	PUNCT
alces-443	453	1	_	_	PUNCT
alces-443	454	1	_	_	PUNCT
alces-443	454	2	.	.	PUNCT
alces-443	455	1	1990	1990	NUM
alces-443	455	2	.	.	PUNCT
alces-443	456	1	new	new	ADJ
alces-443	456	2	dates	date	NOUN
alces-443	456	3	in	in	ADP
alces-443	456	4	alaskan	alaskan	ADJ
alces-443	456	5	quaternary	quaternary	ADJ
alces-443	456	6	moose	moose	NOUN
alces-443	456	7	,	,	PUNCT
alces-443	456	8	cervalces	cervalce	NOUN
alces-443	456	9	-	-	PUNCT
alces-443	456	10	alces	alce	NOUN
alces-443	456	11	–	–	PUNCT
alces-443	456	12	archaeological	archaeological	ADJ
alces-443	456	13	,	,	PUNCT
alces-443	456	14	evolutionary	evolutionary	ADJ
alces-443	456	15	and	and	CCONJ
alces-443	456	16	ecological	ecological	ADJ
alces-443	456	17	implications	implication	NOUN
alces-443	456	18	.	.	PUNCT
alces-443	457	1	current	current	ADJ
alces-443	457	2	research	research	NOUN
alces-443	457	3	in	in	ADP
alces-443	457	4	the	the	DET
alces-443	457	5	pleistocene	pleistocene	NOUN
alces-443	457	6	7:111	7:111	PROPN
alces-443	457	7	-	-	SYM
alces-443	457	8	112	112	NUM
alces-443	457	9	.	.	PUNCT
alces-443	458	1	_	_	PUNCT
alces-443	459	1	_	_	PUNCT
alces-443	460	1	_	_	PUNCT
alces-443	461	1	_	_	PUNCT
alces-443	462	1	_	_	PUNCT
alces-443	462	2	.	.	PUNCT
alces-443	463	1	1995	1995	NUM
alces-443	463	2	.	.	PUNCT
alces-443	464	1	mammalian	mammalian	ADJ
alces-443	464	2	evolution	evolution	NOUN
alces-443	464	3	in	in	ADP
alces-443	464	4	response	response	NOUN
alces-443	464	5	to	to	ADP
alces-443	464	6	the	the	DET
alces-443	464	7	pleistocene	pleistocene	NOUN
alces-443	464	8	-	-	PUNCT
alces-443	464	9	holocene	holocene	PROPN
alces-443	464	10	transition	transition	NOUN
alces-443	464	11	and	and	CCONJ
alces-443	464	12	the	the	DET
alces-443	464	13	break	break	NOUN
alces-443	464	14	-	-	PUNCT
alces-443	464	15	up	up	NOUN
alces-443	464	16	of	of	ADP
alces-443	464	17	the	the	DET
alces-443	464	18	mammoth	mammoth	ADJ
alces-443	464	19	steppe	steppe	NOUN
alces-443	464	20	:	:	PUNCT
alces-443	464	21	two	two	NUM
alces-443	464	22	case	case	NOUN
alces-443	464	23	studies	study	NOUN
alces-443	464	24	.	.	PUNCT
alces-443	465	1	acta	acta	PROPN
alces-443	465	2	zoologica	zoologica	PROPN
alces-443	465	3	cracoviensia	cracoviensia	PROPN
alces-443	465	4	38:139	38:139	PROPN
alces-443	465	5	-	-	PUNCT
alces-443	465	6	154	154	NUM
alces-443	465	7	.	.	PUNCT
alces-443	466	1	gyllensten	gyllensten	VERB
alces-443	466	2	,	,	PUNCT
alces-443	466	3	u.	u.	PROPN
alces-443	466	4	,	,	PUNCT
alces-443	466	5	c.	c.	NOUN
alces-443	466	6	reuterwall	reuterwall	NOUN
alces-443	466	7	,	,	PUNCT
alces-443	466	8	n.	n.	PROPN
alces-443	466	9	ryman	ryman	PROPN
alces-443	466	10	,	,	PUNCT
alces-443	466	11	and	and	CCONJ
alces-443	466	12	g.	g.	PROPN
alces-443	466	13	stahl	stahl	PROPN
alces-443	466	14	.	.	PUNCT
alces-443	467	1	1980	1980	NUM
alces-443	467	2	.	.	PUNCT
alces-443	468	1	geographical	geographical	ADJ
alces-443	468	2	variation	variation	NOUN
alces-443	468	3	in	in	ADP
alces-443	468	4	transferrin	transferrin	ADJ
alces-443	468	5	allele	allele	NOUN
alces-443	468	6	frequencies	frequency	NOUN
alces-443	468	7	in	in	ADP
alces-443	468	8	three	three	NUM
alces-443	468	9	deer	deer	NOUN
alces-443	468	10	species	specie	NOUN
alces-443	468	11	from	from	ADP
alces-443	468	12	scandinavia	scandinavia	PROPN
alces-443	468	13	.	.	PUNCT
alces-443	469	1	hereditas	hereditas	PROPN
alces-443	469	2	92:237	92:237	NUM
alces-443	469	3	-	-	SYM
alces-443	469	4	241	241	NUM
alces-443	469	5	.	.	PUNCT
alces-443	470	1	hancock	hancock	PROPN
alces-443	470	2	,	,	PUNCT
alces-443	470	3	j.	j.	PROPN
alces-443	470	4	m.	m.	PROPN
alces-443	470	5	1999	1999	NUM
alces-443	470	6	.	.	PUNCT
alces-443	471	1	microsatellites	microsatellite	NOUN
alces-443	471	2	and	and	CCONJ
alces-443	471	3	other	other	ADJ
alces-443	471	4	simple	simple	ADJ
alces-443	471	5	sequences	sequence	NOUN
alces-443	471	6	:	:	PUNCT
alces-443	471	7	genomic	genomic	ADJ
alces-443	471	8	context	context	NOUN
alces-443	471	9	and	and	CCONJ
alces-443	471	10	mutational	mutational	ADJ
alces-443	471	11	mechanisms	mechanism	NOUN
alces-443	471	12	.	.	PUNCT
alces-443	472	1	pages	page	NOUN
alces-443	472	2	1	1	NUM
alces-443	472	3	-	-	SYM
alces-443	472	4	9	9	NUM
alces-443	472	5	in	in	ADP
alces-443	472	6	d.	d.	PROPN
alces-443	472	7	b.	b.	PROPN
alces-443	472	8	goldstin	goldstin	PROPN
alces-443	472	9	and	and	CCONJ
alces-443	472	10	c.	c.	PROPN
alces-443	472	11	schlötterer	schlötterer	PROPN
alces-443	472	12	,	,	PUNCT
alces-443	472	13	editors	editor	NOUN
alces-443	472	14	.	.	PUNCT
alces-443	473	1	microsatellites	microsatellite	NOUN
alces-443	473	2	:	:	PUNCT
alces-443	473	3	evolution	evolution	NOUN
alces-443	473	4	and	and	CCONJ
alces-443	473	5	applications	application	NOUN
alces-443	473	6	.	.	PUNCT
alces-443	474	1	oxford	oxford	PROPN
alces-443	474	2	university	university	PROPN
alces-443	474	3	press	press	NOUN
alces-443	474	4	,	,	PUNCT
alces-443	474	5	oxford	oxford	PROPN
alces-443	474	6	,	,	PUNCT
alces-443	474	7	u.k	u.k	PROPN
alces-443	474	8	.	.	PROPN
alces-443	474	9	harrington	harrington	PROPN
alces-443	474	10	,	,	PUNCT
alces-443	474	11	r.	r.	PROPN
alces-443	474	12	1985	1985	NUM
alces-443	474	13	.	.	PUNCT
alces-443	475	1	evolution	evolution	NOUN
alces-443	475	2	and	and	CCONJ
alces-443	475	3	distribution	distribution	NOUN
alces-443	475	4	of	of	ADP
alces-443	475	5	the	the	DET
alces-443	475	6	cervidae	cervidae	NOUN
alces-443	475	7	.	.	PUNCT
alces-443	476	1	biology	biology	NOUN
alces-443	476	2	of	of	ADP
alces-443	476	3	deer	deer	NOUN
alces-443	476	4	production	production	NOUN
alces-443	476	5	.	.	PUNCT
alces-443	477	1	the	the	DET
alces-443	477	2	royal	royal	ADJ
alces-443	477	3	society	society	NOUN
alces-443	477	4	of	of	ADP
alces-443	477	5	new	new	PROPN
alces-443	477	6	zealand	zealand	PROPN
alces-443	477	7	,	,	PUNCT
alces-443	477	8	bulletin	bulletin	NOUN
alces-443	477	9	22:3	22:3	NUM
alces-443	477	10	-	-	SYM
alces-443	477	11	11	11	NUM
alces-443	477	12	.	.	PUNCT
alces-443	478	1	harris	harris	PROPN
alces-443	478	2	,	,	PUNCT
alces-443	478	3	r.	r.	PROPN
alces-443	478	4	b.	b.	PROPN
alces-443	478	5	,	,	PUNCT
alces-443	478	6	w.	w.	PROPN
alces-443	478	7	a.	a.	PROPN
alces-443	478	8	wall	wall	PROPN
alces-443	478	9	,	,	PUNCT
alces-443	478	10	and	and	CCONJ
alces-443	478	11	f.	f.	PROPN
alces-443	478	12	w.	w.	PROPN
alces-443	478	13	allendorf	allendorf	PROPN
alces-443	478	14	.	.	PUNCT
alces-443	479	1	2002	2002	NUM
alces-443	479	2	.	.	PUNCT
alces-443	480	1	genetic	genetic	ADJ
alces-443	480	2	consequences	consequence	NOUN
alces-443	480	3	of	of	ADP
alces-443	480	4	hunting	hunt	VERB
alces-443	480	5	:	:	PUNCT
alces-443	480	6	what	what	PRON
alces-443	480	7	do	do	AUX
alces-443	480	8	we	we	PRON
alces-443	480	9	know	know	VERB
alces-443	480	10	and	and	CCONJ
alces-443	480	11	what	what	PRON
alces-443	480	12	should	should	AUX
alces-443	480	13	we	we	PRON
alces-443	480	14	do	do	VERB
alces-443	480	15	?	?	PUNCT
alces-443	481	1	wildlife	wildlife	NOUN
alces-443	481	2	society	society	NOUN
alces-443	481	3	bulletin	bulletin	NOUN
alces-443	481	4	30:634	30:634	NUM
alces-443	481	5	-	-	PUNCT
alces-443	481	6	643	643	NUM
alces-443	481	7	.	.	PUNCT
alces-443	482	1	hedrick	hedrick	PROPN
alces-443	482	2	,	,	PUNCT
alces-443	482	3	p.	p.	PROPN
alces-443	482	4	w.	w.	PROPN
alces-443	482	5	1994	1994	NUM
alces-443	482	6	.	.	PUNCT
alces-443	483	1	evolutionary	evolutionary	ADJ
alces-443	483	2	genetics	genetic	NOUN
alces-443	483	3	of	of	ADP
alces-443	483	4	the	the	DET
alces-443	483	5	major	major	ADJ
alces-443	483	6	histocompatibility	histocompatibility	NOUN
alces-443	483	7	complex	complex	NOUN
alces-443	483	8	.	.	PUNCT
alces-443	484	1	american	american	PROPN
alces-443	484	2	naturalist	naturalist	PROPN
alces-443	484	3	143:945964	143:945964	NUM
alces-443	484	4	.	.	PUNCT
alces-443	485	1	hewitt	hewitt	PROPN
alces-443	485	2	,	,	PUNCT
alces-443	485	3	g.	g.	PROPN
alces-443	485	4	m.	m.	PROPN
alces-443	485	5	1996	1996	NUM
alces-443	485	6	.	.	PUNCT
alces-443	486	1	some	some	DET
alces-443	486	2	genetic	genetic	ADJ
alces-443	486	3	consequences	consequence	NOUN
alces-443	486	4	of	of	ADP
alces-443	486	5	ice	ice	NOUN
alces-443	486	6	ages	age	NOUN
alces-443	486	7	,	,	PUNCT
alces-443	486	8	and	and	CCONJ
alces-443	486	9	their	their	PRON
alces-443	486	10	role	role	NOUN
alces-443	486	11	in	in	ADP
alces-443	486	12	divergence	divergence	NOUN
alces-443	486	13	and	and	CCONJ
alces-443	486	14	speciation	speciation	NOUN
alces-443	486	15	.	.	PUNCT
alces-443	487	1	biological	biological	ADJ
alces-443	487	2	journal	journal	NOUN
alces-443	487	3	of	of	ADP
alces-443	487	4	the	the	DET
alces-443	487	5	linnaen	linnaen	PROPN
alces-443	487	6	society	society	PROPN
alces-443	487	7	58:247276	58:247276	PROPN
alces-443	487	8	.	.	PUNCT
alces-443	488	1	hirth	hirth	NOUN
alces-443	488	2	,	,	PUNCT
alces-443	488	3	d.	d.	PROPN
alces-443	488	4	h.	h.	PROPN
alces-443	488	5	1977	1977	NUM
alces-443	488	6	.	.	PUNCT
alces-443	489	1	social	social	ADJ
alces-443	489	2	behavior	behavior	NOUN
alces-443	489	3	of	of	ADP
alces-443	489	4	white	white	ADJ
alces-443	489	5	-	-	PUNCT
alces-443	489	6	tailed	tail	VERB
alces-443	489	7	deer	deer	NOUN
alces-443	489	8	in	in	ADP
alces-443	489	9	relation	relation	NOUN
alces-443	489	10	to	to	ADP
alces-443	489	11	habitat	habitat	NOUN
alces-443	489	12	.	.	PUNCT
alces-443	490	1	wildlife	wildlife	NOUN
alces-443	490	2	monographs	monograph	NOUN
alces-443	490	3	53	53	NUM
alces-443	490	4	.	.	PUNCT
alces-443	491	1	hundertmark	hundertmark	PROPN
alces-443	491	2	,	,	PUNCT
alces-443	491	3	k.	k.	PROPN
alces-443	491	4	j.	j.	PROPN
alces-443	491	5	,	,	PUNCT
alces-443	491	6	r.	r.	PROPN
alces-443	491	7	t.	t.	PROPN
alces-443	491	8	bowyer	bowyer	PROPN
alces-443	491	9	,	,	PUNCT
alces-443	491	10	g.	g.	PROPN
alces-443	491	11	f.	f.	PROPN
alces-443	491	12	shields	shields	PROPN
alces-443	491	13	,	,	PUNCT
alces-443	491	14	and	and	CCONJ
alces-443	491	15	c.	c.	PROPN
alces-443	491	16	c.	c.	PROPN
alces-443	491	17	schwartz	schwartz	PROPN
alces-443	491	18	.	.	PUNCT
alces-443	492	1	2003	2003	NUM
alces-443	492	2	.	.	PUNCT
alces-443	493	1	mitochondrial	mitochondrial	ADJ
alces-443	493	2	phylogeography	phylogeography	NOUN
alces-443	493	3	of	of	ADP
alces-443	493	4	moose	moose	NOUN
alces-443	493	5	(	(	PUNCT
alces-443	493	6	alces	alce	NOUN
alces-443	493	7	alces	alce	NOUN
alces-443	493	8	)	)	PUNCT
alces-443	493	9	in	in	ADP
alces-443	493	10	north	north	PROPN
alces-443	493	11	america	america	PROPN
alces-443	493	12	.	.	PUNCT
alces-443	494	1	journal	journal	PROPN
alces-443	494	2	of	of	ADP
alces-443	494	3	mammalogy	mammalogy	NOUN
alces-443	494	4	84:718	84:718	NUM
alces-443	494	5	-	-	SYM
alces-443	494	6	728	728	NUM
alces-443	494	7	.	.	PUNCT
alces-443	495	1	_	_	PUNCT
alces-443	496	1	_	_	PUNCT
alces-443	497	1	_	_	PUNCT
alces-443	498	1	_	_	PUNCT
alces-443	499	1	_	_	PUNCT
alces-443	499	2	,	,	PUNCT
alces-443	499	3	p.	p.	PROPN
alces-443	499	4	e.	e.	PROPN
alces-443	499	5	johns	johns	PROPN
alces-443	499	6	,	,	PUNCT
alces-443	499	7	and	and	CCONJ
alces-443	499	8	m.	m.	PROPN
alces-443	499	9	h.	h.	PROPN
alces-443	499	10	smith	smith	PROPN
alces-443	499	11	.	.	PROPN
alces-443	500	1	1992	1992	NUM
alces-443	500	2	.	.	PUNCT
alces-443	501	1	genetic	genetic	ADJ
alces-443	501	2	diversity	diversity	NOUN
alces-443	501	3	of	of	ADP
alces-443	501	4	moose	moose	NOUN
alces-443	501	5	from	from	ADP
alces-443	501	6	the	the	DET
alces-443	501	7	kenai	kenai	PROPN
alces-443	501	8	peninsula	peninsula	PROPN
alces-443	501	9	,	,	PUNCT
alces-443	501	10	alaska	alaska	PROPN
alces-443	501	11	.	.	PUNCT
alces-443	501	12	alces	alce	VERB
alces-443	501	13	28:15	28:15	NUM
alces-443	501	14	-	-	SYM
alces-443	501	15	20	20	NUM
alces-443	501	16	.	.	PUNCT
alces-443	502	1	_	_	PUNCT
alces-443	503	1	_	_	PUNCT
alces-443	504	1	_	_	PUNCT
alces-443	505	1	_	_	PUNCT
alces-443	506	1	_	_	PUNCT
alces-443	506	2	,	,	PUNCT
alces-443	506	3	g.	g.	PROPN
alces-443	506	4	f.	f.	PROPN
alces-443	506	5	shields	shields	PROPN
alces-443	506	6	,	,	PUNCT
alces-443	506	7	r.	r.	PROPN
alces-443	506	8	t.	t.	PROPN
alces-443	506	9	bowyer	bowyer	PROPN
alces-443	506	10	,	,	PUNCT
alces-443	506	11	and	and	CCONJ
alces-443	506	12	c.	c.	PROPN
alces-443	506	13	c.	c.	PROPN
alces-443	506	14	schwartz	schwartz	PROPN
alces-443	506	15	.	.	PUNCT
alces-443	506	16	2002a	2002a	PROPN
alces-443	506	17	.	.	PUNCT
alces-443	507	1	genetic	genetic	ADJ
alces-443	507	2	relationships	relationship	NOUN
alces-443	507	3	deduced	deduce	VERB
alces-443	507	4	from	from	ADP
alces-443	507	5	cytochrome	cytochrome	NOUN
alces-443	507	6	-	-	PUNCT
alces-443	507	7	b	b	NOUN
alces-443	507	8	sequences	sequence	NOUN
alces-443	507	9	among	among	ADP
alces-443	507	10	moose	moose	NOUN
alces-443	507	11	.	.	PUNCT
alces-443	508	1	alces	alce	NOUN
alces-443	508	2	38:113122	38:113122	PROPN
alces-443	508	3	.	.	PUNCT
alces-443	508	4	alces	alce	NOUN
alces-443	508	5	vol	vol	NOUN
alces-443	508	6	.	.	PROPN
alces-443	508	7	40	40	NUM
alces-443	508	8	,	,	PUNCT
alces-443	508	9	2004	2004	NUM
alces-443	508	10	moose	moose	NOUN
alces-443	508	11	genetics	genetic	NOUN
alces-443	508	12	hundertmark	hundertmark	NOUN
alces-443	508	13	and	and	CCONJ
alces-443	508	14	bowyer	bowyer	PROPN
alces-443	509	1	119	119	NUM
alces-443	510	1	_	_	PUNCT
alces-443	511	1	_	_	PUNCT
alces-443	512	1	_	_	PUNCT
alces-443	513	1	_	_	PUNCT
alces-443	514	1	_	_	PUNCT
alces-443	514	2	,	,	PUNCT
alces-443	515	1	_	_	PUNCT
alces-443	516	1	_	_	PUNCT
alces-443	517	1	_	_	PUNCT
alces-443	518	1	_	_	PUNCT
alces-443	519	1	_	_	PUNCT
alces-443	519	2	,	,	PUNCT
alces-443	519	3	i.	i.	PROPN
alces-443	519	4	g.	g.	PROPN
alces-443	519	5	udina	udina	PROPN
alces-443	519	6	,	,	PUNCT
alces-443	519	7	r.	r.	PROPN
alces-443	519	8	t.	t.	PROPN
alces-443	519	9	bowyer	bowyer	PROPN
alces-443	519	10	,	,	PUNCT
alces-443	519	11	a.	a.	NOUN
alces-443	519	12	a.	a.	NOUN
alces-443	519	13	danilkin	danilkin	PROPN
alces-443	519	14	,	,	PUNCT
alces-443	519	15	and	and	CCONJ
alces-443	519	16	c.	c.	PROPN
alces-443	519	17	c.	c.	PROPN
alces-443	519	18	schwartz	schwartz	PROPN
alces-443	519	19	.	.	PUNCT
alces-443	520	1	2002b	2002b	NUM
alces-443	520	2	.	.	PUNCT
alces-443	521	1	mitochondrial	mitochondrial	ADJ
alces-443	521	2	phylogeography	phylogeography	NOUN
alces-443	521	3	of	of	ADP
alces-443	521	4	moose	moose	NOUN
alces-443	521	5	(	(	PUNCT
alces-443	521	6	alces	alce	NOUN
alces-443	521	7	alces	alce	NOUN
alces-443	521	8	):	):	PUNCT
alces-443	521	9	late	late	ADJ
alces-443	521	10	pleistocene	pleistocene	PROPN
alces-443	521	11	divergence	divergence	NOUN
alces-443	521	12	and	and	CCONJ
alces-443	521	13	population	population	NOUN
alces-443	521	14	expansion	expansion	NOUN
alces-443	521	15	.	.	PUNCT
alces-443	522	1	molecular	molecular	ADJ
alces-443	522	2	phylogenetics	phylogenetic	NOUN
alces-443	522	3	and	and	CCONJ
alces-443	522	4	evolution	evolution	NOUN
alces-443	522	5	22:375	22:375	NUM
alces-443	522	6	-	-	SYM
alces-443	522	7	387	387	NUM
alces-443	522	8	.	.	PUNCT
alces-443	523	1	_	_	PUNCT
alces-443	524	1	_	_	PUNCT
alces-443	525	1	_	_	PUNCT
alces-443	526	1	_	_	PUNCT
alces-443	527	1	_	_	PUNCT
alces-443	527	2	,	,	PUNCT
alces-443	527	3	t.	t.	PROPN
alces-443	527	4	h.	h.	PROPN
alces-443	527	5	thelen	thelen	PROPN
alces-443	527	6	,	,	PUNCT
alces-443	527	7	and	and	CCONJ
alces-443	527	8	r.	r.	PROPN
alces-443	527	9	t.	t.	PROPN
alces-443	527	10	bowyer	bowyer	PROPN
alces-443	527	11	.	.	PUNCT
alces-443	528	1	1998	1998	NUM
alces-443	528	2	.	.	PUNCT
alces-443	529	1	effects	effect	NOUN
alces-443	529	2	of	of	ADP
alces-443	529	3	population	population	NOUN
alces-443	529	4	density	density	NOUN
alces-443	529	5	and	and	CCONJ
alces-443	529	6	selective	selective	ADJ
alces-443	529	7	harvest	harvest	NOUN
alces-443	529	8	on	on	ADP
alces-443	529	9	antler	antler	NOUN
alces-443	529	10	phenotype	phenotype	NOUN
alces-443	529	11	in	in	ADP
alces-443	529	12	simulated	simulated	ADJ
alces-443	529	13	moose	moose	NOUN
alces-443	529	14	populations	population	NOUN
alces-443	529	15	.	.	PUNCT
alces-443	530	1	alces	alce	NOUN
alces-443	530	2	34:375	34:375	NUM
alces-443	530	3	-	-	SYM
alces-443	530	4	383	383	NUM
alces-443	530	5	.	.	PUNCT
alces-443	531	1	_	_	PUNCT
alces-443	532	1	_	_	PUNCT
alces-443	533	1	_	_	PUNCT
alces-443	534	1	_	_	PUNCT
alces-443	535	1	_	_	PUNCT
alces-443	535	2	,	,	PUNCT
alces-443	536	1	_	_	PUNCT
alces-443	537	1	_	_	PUNCT
alces-443	538	1	_	_	PUNCT
alces-443	539	1	_	_	PUNCT
alces-443	540	1	_	_	PUNCT
alces-443	540	2	,	,	PUNCT
alces-443	540	3	and	and	CCONJ
alces-443	540	4	c.	c.	PROPN
alces-443	540	5	c.	c.	PROPN
alces-443	540	6	schwartz	schwartz	PROPN
alces-443	540	7	.	.	PUNCT
alces-443	541	1	1993	1993	NUM
alces-443	541	2	.	.	PUNCT
alces-443	542	1	genetic	genetic	ADJ
alces-443	542	2	and	and	CCONJ
alces-443	542	3	population	population	NOUN
alces-443	542	4	effects	effect	NOUN
alces-443	542	5	of	of	ADP
alces-443	542	6	selective	selective	ADJ
alces-443	542	7	harvest	harvest	NOUN
alces-443	542	8	systems	system	NOUN
alces-443	542	9	in	in	ADP
alces-443	542	10	moose	moose	NOUN
alces-443	542	11	:	:	PUNCT
alces-443	542	12	a	a	DET
alces-443	542	13	modeling	modeling	NOUN
alces-443	542	14	approach	approach	NOUN
alces-443	542	15	.	.	PUNCT
alces-443	543	1	alces	alce	NOUN
alces-443	543	2	29:225	29:225	NUM
alces-443	543	3	-	-	SYM
alces-443	543	4	234	234	NUM
alces-443	543	5	.	.	PUNCT
alces-443	544	1	irwin	irwin	PROPN
alces-443	544	2	,	,	PUNCT
alces-443	544	3	d.	d.	PROPN
alces-443	544	4	m.	m.	PROPN
alces-443	544	5	,	,	PUNCT
alces-443	544	6	t.	t.	PROPN
alces-443	544	7	d.	d.	PROPN
alces-443	544	8	kocher	kocher	PROPN
alces-443	544	9	,	,	PUNCT
alces-443	544	10	and	and	CCONJ
alces-443	544	11	a.	a.	PROPN
alces-443	544	12	c.	c.	PROPN
alces-443	544	13	wilson	wilson	PROPN
alces-443	544	14	.	.	PUNCT
alces-443	545	1	1991	1991	NUM
alces-443	545	2	.	.	PUNCT
alces-443	546	1	evolution	evolution	NOUN
alces-443	546	2	of	of	ADP
alces-443	546	3	the	the	DET
alces-443	546	4	cytochrome	cytochrome	PROPN
alces-443	546	5	b	b	PROPN
alces-443	546	6	gene	gene	NOUN
alces-443	546	7	in	in	ADP
alces-443	546	8	mammals	mammal	NOUN
alces-443	546	9	.	.	PUNCT
alces-443	547	1	journal	journal	NOUN
alces-443	547	2	of	of	ADP
alces-443	547	3	molecular	molecular	ADJ
alces-443	547	4	evolution	evolution	NOUN
alces-443	547	5	.	.	PUNCT
alces-443	548	1	32	32	NUM
alces-443	548	2	:	:	SYM
alces-443	548	3	128	128	NUM
alces-443	548	4	144	144	NUM
alces-443	548	5	.	.	PUNCT
alces-443	549	1	jouzel	jouzel	PROPN
alces-443	549	2	,	,	PUNCT
alces-443	549	3	j.	j.	PROPN
alces-443	549	4	,	,	PUNCT
alces-443	549	5	c.	c.	PROPN
alces-443	549	6	lorius	lorius	PROPN
alces-443	549	7	,	,	PUNCT
alces-443	549	8	j.	j.	PROPN
alces-443	549	9	r.	r.	PROPN
alces-443	549	10	petit	petit	PROPN
alces-443	549	11	,	,	PUNCT
alces-443	549	12	c.	c.	PROPN
alces-443	549	13	genthon	genthon	PROPN
alces-443	549	14	,	,	PUNCT
alces-443	549	15	n.	n.	PROPN
alces-443	549	16	i.	i.	PROPN
alces-443	549	17	barkov	barkov	PROPN
alces-443	549	18	,	,	PUNCT
alces-443	549	19	v.	v.	ADP
alces-443	549	20	m.	m.	PROPN
alces-443	549	21	kotlyakov	kotlyakov	PROPN
alces-443	549	22	,	,	PUNCT
alces-443	549	23	and	and	CCONJ
alces-443	550	1	v.	v.	ADP
alces-443	550	2	m.	m.	NOUN
alces-443	550	3	petrov	petrov	PROPN
alces-443	550	4	.	.	PROPN
alces-443	550	5	1987	1987	NUM
alces-443	550	6	.	.	PUNCT
alces-443	551	1	vostok	vostok	PROPN
alces-443	551	2	ice	ice	NOUN
alces-443	551	3	core	core	NOUN
alces-443	551	4	:	:	PUNCT
alces-443	551	5	a	a	DET
alces-443	551	6	continuous	continuous	ADJ
alces-443	551	7	isotope	isotope	NOUN
alces-443	551	8	temperature	temperature	NOUN
alces-443	551	9	record	record	NOUN
alces-443	551	10	over	over	ADP
alces-443	551	11	the	the	DET
alces-443	551	12	last	last	ADJ
alces-443	551	13	climate	climate	NOUN
alces-443	551	14	cycle	cycle	NOUN
alces-443	551	15	(	(	PUNCT
alces-443	551	16	160,000	160,000	NUM
alces-443	551	17	years	year	NOUN
alces-443	551	18	)	)	PUNCT
alces-443	551	19	.	.	PUNCT
alces-443	552	1	nature	nature	NOUN
alces-443	553	1	329:403	329:403	PROPN
alces-443	553	2	-	-	PUNCT
alces-443	553	3	408	408	NUM
alces-443	553	4	.	.	PUNCT
alces-443	554	1	lister	lister	PROPN
alces-443	554	2	,	,	PUNCT
alces-443	554	3	a.	a.	NOUN
alces-443	554	4	m.	m.	NOUN
alces-443	554	5	1993	1993	NUM
alces-443	554	6	.	.	PUNCT
alces-443	555	1	evolution	evolution	NOUN
alces-443	555	2	of	of	ADP
alces-443	555	3	mammoths	mammoth	NOUN
alces-443	555	4	and	and	CCONJ
alces-443	555	5	moose	moose	NOUN
alces-443	555	6	:	:	PUNCT
alces-443	555	7	the	the	DET
alces-443	555	8	holarctic	holarctic	ADJ
alces-443	555	9	perspective	perspective	NOUN
alces-443	555	10	.	.	PUNCT
alces-443	556	1	pages	page	NOUN
alces-443	556	2	178	178	NUM
alces-443	556	3	-	-	SYM
alces-443	556	4	204	204	NUM
alces-443	556	5	in	in	ADP
alces-443	556	6	a.	a.	PROPN
alces-443	556	7	d.	d.	PROPN
alces-443	556	8	barnosky	barnosky	PROPN
alces-443	556	9	,	,	PUNCT
alces-443	556	10	editor	editor	NOUN
alces-443	556	11	.	.	PUNCT
alces-443	557	1	quaternary	quaternary	ADJ
alces-443	557	2	mammals	mammal	NOUN
alces-443	557	3	of	of	ADP
alces-443	557	4	north	north	PROPN
alces-443	557	5	america	america	PROPN
alces-443	557	6	.	.	PUNCT
alces-443	558	1	cambridge	cambridge	PROPN
alces-443	558	2	university	university	PROPN
alces-443	558	3	press	press	PROPN
alces-443	558	4	,	,	PUNCT
alces-443	558	5	cambridge	cambridge	PROPN
alces-443	558	6	,	,	PUNCT
alces-443	558	7	u.k	u.k	PROPN
alces-443	558	8	.	.	PROPN
alces-443	558	9	lopez	lopez	PROPN
alces-443	558	10	,	,	PUNCT
alces-443	558	11	j.	j.	PROPN
alces-443	558	12	v.	v.	PROPN
alces-443	558	13	,	,	PUNCT
alces-443	558	14	m.	m.	NOUN
alces-443	558	15	culver	culver	PROPN
alces-443	558	16	,	,	PUNCT
alces-443	558	17	j.	j.	PROPN
alces-443	558	18	c.	c.	PROPN
alces-443	558	19	stephens	stephens	PROPN
alces-443	558	20	,	,	PUNCT
alces-443	558	21	w.	w.	PROPN
alces-443	558	22	e.	e.	PROPN
alces-443	558	23	johnson	johnson	PROPN
alces-443	558	24	,	,	PUNCT
alces-443	558	25	and	and	CCONJ
alces-443	558	26	s.	s.	PROPN
alces-443	558	27	j.	j.	PROPN
alces-443	558	28	o’brien	o’brien	PROPN
alces-443	558	29	.	.	PUNCT
alces-443	559	1	1997	1997	NUM
alces-443	559	2	.	.	PUNCT
alces-443	560	1	rates	rate	NOUN
alces-443	560	2	of	of	ADP
alces-443	560	3	nuclear	nuclear	ADJ
alces-443	560	4	and	and	CCONJ
alces-443	560	5	cytoplasmic	cytoplasmic	ADJ
alces-443	560	6	mitochondrial	mitochondrial	ADJ
alces-443	560	7	dna	dna	NOUN
alces-443	560	8	sequence	sequence	NOUN
alces-443	560	9	divergence	divergence	NOUN
alces-443	560	10	in	in	ADP
alces-443	560	11	mammals	mammal	NOUN
alces-443	560	12	.	.	PUNCT
alces-443	561	1	molecular	molecular	ADJ
alces-443	561	2	biology	biology	NOUN
alces-443	561	3	and	and	CCONJ
alces-443	561	4	evolution	evolution	NOUN
alces-443	561	5	14:277	14:277	PROPN
alces-443	561	6	-	-	PUNCT
alces-443	561	7	286	286	NUM
alces-443	561	8	.	.	PUNCT
alces-443	562	1	lynch	lynch	PROPN
alces-443	562	2	,	,	PUNCT
alces-443	562	3	m.	m.	NOUN
alces-443	562	4	1996	1996	NUM
alces-443	562	5	.	.	PUNCT
alces-443	563	1	a	a	DET
alces-443	563	2	quantitative	quantitative	ADJ
alces-443	563	3	-	-	PUNCT
alces-443	563	4	genetic	genetic	ADJ
alces-443	563	5	perspective	perspective	NOUN
alces-443	563	6	on	on	ADP
alces-443	563	7	conservation	conservation	NOUN
alces-443	563	8	issues	issue	NOUN
alces-443	563	9	.	.	PUNCT
alces-443	564	1	pages	page	NOUN
alces-443	564	2	471	471	NUM
alces-443	564	3	-	-	SYM
alces-443	564	4	501	501	NUM
alces-443	564	5	in	in	ADP
alces-443	564	6	j.	j.	PROPN
alces-443	564	7	c.	c.	PROPN
alces-443	564	8	avise	avise	PROPN
alces-443	564	9	and	and	CCONJ
alces-443	564	10	j.	j.	PROPN
alces-443	564	11	l.	l.	PROPN
alces-443	564	12	hamrick	hamrick	PROPN
alces-443	564	13	,	,	PUNCT
alces-443	564	14	editors	editor	NOUN
alces-443	564	15	.	.	PUNCT
alces-443	565	1	conservation	conservation	NOUN
alces-443	565	2	genetics	genetic	NOUN
alces-443	565	3	:	:	PUNCT
alces-443	565	4	case	case	NOUN
alces-443	565	5	studies	study	NOUN
alces-443	565	6	from	from	ADP
alces-443	565	7	nature	nature	NOUN
alces-443	565	8	.	.	PUNCT
alces-443	566	1	chapman	chapman	PROPN
alces-443	566	2	&	&	CCONJ
alces-443	566	3	hall	hall	PROPN
alces-443	566	4	,	,	PUNCT
alces-443	566	5	new	new	PROPN
alces-443	566	6	york	york	PROPN
alces-443	566	7	,	,	PUNCT
alces-443	566	8	new	new	PROPN
alces-443	566	9	york	york	PROPN
alces-443	566	10	,	,	PUNCT
alces-443	566	11	usa	usa	PROPN
alces-443	566	12	.	.	PROPN
alces-443	566	13	mason	mason	PROPN
alces-443	566	14	,	,	PUNCT
alces-443	566	15	o.	o.	PROPN
alces-443	566	16	k.	k.	PROPN
alces-443	566	17	,	,	PUNCT
alces-443	566	18	p.	p.	NOUN
alces-443	566	19	m.	m.	NOUN
alces-443	566	20	bowers	bower	NOUN
alces-443	566	21	,	,	PUNCT
alces-443	566	22	and	and	CCONJ
alces-443	566	23	d.	d.	PROPN
alces-443	566	24	m.	m.	PROPN
alces-443	566	25	hopkins	hopkins	PROPN
alces-443	566	26	.	.	PROPN
alces-443	566	27	2001	2001	NUM
alces-443	566	28	.	.	PUNCT
alces-443	567	1	the	the	DET
alces-443	567	2	early	early	ADJ
alces-443	567	3	milankovitch	milankovitch	NOUN
alces-443	567	4	thermal	thermal	ADJ
alces-443	567	5	maximum	maximum	NOUN
alces-443	567	6	and	and	CCONJ
alces-443	567	7	humans	human	NOUN
alces-443	567	8	:	:	PUNCT
alces-443	567	9	adverse	adverse	ADJ
alces-443	567	10	conditions	condition	NOUN
alces-443	567	11	for	for	ADP
alces-443	567	12	the	the	DET
alces-443	567	13	denali	denali	PROPN
alces-443	567	14	complex	complex	NOUN
alces-443	567	15	of	of	ADP
alces-443	567	16	eastern	eastern	ADJ
alces-443	567	17	beringia	beringia	NOUN
alces-443	567	18	.	.	PUNCT
alces-443	568	1	quaternary	quaternary	ADJ
alces-443	568	2	science	science	NOUN
alces-443	568	3	reviews	review	NOUN
alces-443	568	4	20:525	20:525	NUM
alces-443	568	5	-	-	PUNCT
alces-443	568	6	548	548	NUM
alces-443	568	7	.	.	PUNCT
alces-443	569	1	mikko	mikko	PROPN
alces-443	569	2	,	,	PUNCT
alces-443	569	3	s.	s.	PROPN
alces-443	569	4	,	,	PUNCT
alces-443	569	5	and	and	CCONJ
alces-443	569	6	l.	l.	PROPN
alces-443	569	7	andersson	andersson	PROPN
alces-443	569	8	.	.	PUNCT
alces-443	569	9	1995	1995	NUM
alces-443	569	10	.	.	PUNCT
alces-443	570	1	low	low	ADJ
alces-443	570	2	major	major	ADJ
alces-443	570	3	histocompatibility	histocompatibility	NOUN
alces-443	570	4	complex	complex	ADJ
alces-443	570	5	class	class	NOUN
alces-443	570	6	ii	ii	NOUN
alces-443	570	7	diversity	diversity	NOUN
alces-443	570	8	in	in	ADP
alces-443	570	9	european	european	ADJ
alces-443	570	10	and	and	CCONJ
alces-443	570	11	north	north	ADJ
alces-443	570	12	american	american	ADJ
alces-443	570	13	moose	moose	NOUN
alces-443	570	14	.	.	PUNCT
alces-443	571	1	proceedings	proceeding	NOUN
alces-443	571	2	of	of	ADP
alces-443	571	3	the	the	DET
alces-443	571	4	national	national	PROPN
alces-443	571	5	academy	academy	PROPN
alces-443	571	6	of	of	ADP
alces-443	571	7	sciences	sciences	PROPN
alces-443	571	8	,	,	PUNCT
alces-443	571	9	usa	usa	PROPN
alces-443	571	10	92:4259	92:4259	PROPN
alces-443	571	11	-	-	PUNCT
alces-443	571	12	4263	4263	NUM
alces-443	571	13	.	.	PUNCT
alces-443	572	1	miyamoto	miyamoto	NOUN
alces-443	572	2	,	,	PUNCT
alces-443	572	3	m.	m.	NOUN
alces-443	572	4	m.	m.	NOUN
alces-443	572	5	,	,	PUNCT
alces-443	572	6	f.	f.	PROPN
alces-443	572	7	kraus	kraus	PROPN
alces-443	572	8	,	,	PUNCT
alces-443	572	9	and	and	CCONJ
alces-443	572	10	o.	o.	PROPN
alces-443	572	11	a.	a.	PROPN
alces-443	572	12	ryder	ryder	PROPN
alces-443	572	13	.	.	PUNCT
alces-443	573	1	1990	1990	NUM
alces-443	573	2	.	.	PUNCT
alces-443	574	1	phylogeny	phylogeny	NOUN
alces-443	574	2	and	and	CCONJ
alces-443	574	3	evolution	evolution	NOUN
alces-443	574	4	of	of	ADP
alces-443	574	5	antlered	antlere	VERB
alces-443	574	6	deer	deer	NOUN
alces-443	574	7	determined	determine	VERB
alces-443	574	8	from	from	ADP
alces-443	574	9	mitochondrial	mitochondrial	ADJ
alces-443	574	10	dna	dna	NOUN
alces-443	574	11	sequences	sequence	NOUN
alces-443	574	12	.	.	PUNCT
alces-443	575	1	proceedings	proceeding	NOUN
alces-443	575	2	of	of	ADP
alces-443	575	3	the	the	DET
alces-443	575	4	national	national	PROPN
alces-443	575	5	academy	academy	PROPN
alces-443	575	6	of	of	ADP
alces-443	575	7	sciences	sciences	PROPN
alces-443	575	8	,	,	PUNCT
alces-443	575	9	usa	usa	PROPN
alces-443	575	10	87:6127	87:6127	NUM
alces-443	575	11	-	-	SYM
alces-443	575	12	6131	6131	NUM
alces-443	575	13	.	.	PUNCT
alces-443	576	1	molvar	molvar	PROPN
alces-443	576	2	,	,	PUNCT
alces-443	576	3	e.	e.	PROPN
alces-443	576	4	m.	m.	PROPN
alces-443	576	5	,	,	PUNCT
alces-443	576	6	and	and	CCONJ
alces-443	576	7	r.	r.	PROPN
alces-443	576	8	t.	t.	PROPN
alces-443	576	9	bowyer	bowyer	PROPN
alces-443	576	10	.	.	PROPN
alces-443	576	11	1994	1994	NUM
alces-443	576	12	.	.	PUNCT
alces-443	577	1	costs	cost	NOUN
alces-443	577	2	and	and	CCONJ
alces-443	577	3	benefits	benefit	NOUN
alces-443	577	4	of	of	ADP
alces-443	577	5	group	group	NOUN
alces-443	577	6	living	live	VERB
alces-443	577	7	in	in	ADP
alces-443	577	8	a	a	DET
alces-443	577	9	recently	recently	ADV
alces-443	577	10	social	social	ADJ
alces-443	577	11	ungulate	ungulate	NOUN
alces-443	577	12	:	:	PUNCT
alces-443	577	13	the	the	DET
alces-443	577	14	alaskan	alaskan	ADJ
alces-443	577	15	moose	moose	NOUN
alces-443	577	16	.	.	PUNCT
alces-443	578	1	journal	journal	PROPN
alces-443	578	2	of	of	ADP
alces-443	578	3	mammalogy	mammalogy	NOUN
alces-443	578	4	75:621630	75:621630	NOUN
alces-443	578	5	.	.	PUNCT
alces-443	579	1	mullis	mullis	PROPN
alces-443	579	2	,	,	PUNCT
alces-443	579	3	k.	k.	PROPN
alces-443	579	4	,	,	PUNCT
alces-443	579	5	f.	f.	PROPN
alces-443	579	6	faloona	faloona	PROPN
alces-443	579	7	,	,	PUNCT
alces-443	579	8	s.	s.	PROPN
alces-443	579	9	scharf	scharf	PROPN
alces-443	579	10	,	,	PUNCT
alces-443	579	11	r.	r.	PROPN
alces-443	579	12	sakai	sakai	PROPN
alces-443	579	13	,	,	PUNCT
alces-443	579	14	g.	g.	PROPN
alces-443	579	15	horn	horn	PROPN
alces-443	579	16	,	,	PUNCT
alces-443	579	17	and	and	CCONJ
alces-443	579	18	h.	h.	PROPN
alces-443	579	19	erlich	erlich	PROPN
alces-443	579	20	.	.	PUNCT
alces-443	580	1	1986	1986	NUM
alces-443	580	2	.	.	PUNCT
alces-443	581	1	specific	specific	ADJ
alces-443	581	2	enzymatic	enzymatic	ADJ
alces-443	581	3	amplification	amplification	NOUN
alces-443	581	4	of	of	ADP
alces-443	581	5	dna	dna	PROPN
alces-443	581	6	in	in	ADP
alces-443	581	7	vitro	vitro	NOUN
alces-443	581	8	:	:	PUNCT
alces-443	581	9	the	the	DET
alces-443	581	10	polymerase	polymerase	NOUN
alces-443	581	11	chain	chain	NOUN
alces-443	581	12	reaction	reaction	NOUN
alces-443	581	13	.	.	PUNCT
alces-443	582	1	cold	cold	ADJ
alces-443	582	2	spring	spring	NOUN
alces-443	582	3	harbor	harbor	NOUN
alces-443	582	4	symposia	symposia	NOUN
alces-443	582	5	in	in	ADP
alces-443	582	6	quantitative	quantitative	ADJ
alces-443	582	7	biology	biology	NOUN
alces-443	582	8	51:263	51:263	NUM
alces-443	582	9	-	-	SYM
alces-443	582	10	273	273	NUM
alces-443	582	11	.	.	PUNCT
alces-443	583	1	nadler	nadler	PROPN
alces-443	583	2	,	,	PUNCT
alces-443	583	3	c.	c.	PROPN
alces-443	583	4	f.	f.	PROPN
alces-443	583	5	,	,	PUNCT
alces-443	583	6	c.	c.	PROPN
alces-443	583	7	e.	e.	PROPN
alces-443	583	8	hughes	hughes	PROPN
alces-443	583	9	,	,	PUNCT
alces-443	583	10	k.	k.	PROPN
alces-443	583	11	e.	e.	PROPN
alces-443	583	12	harris	harris	PROPN
alces-443	583	13	,	,	PUNCT
alces-443	583	14	and	and	CCONJ
alces-443	583	15	n.	n.	PROPN
alces-443	583	16	w.	w.	PROPN
alces-443	583	17	adler	adler	PROPN
alces-443	583	18	.	.	PUNCT
alces-443	584	1	1967	1967	NUM
alces-443	584	2	.	.	PUNCT
alces-443	585	1	electrophoresis	electrophoresis	NOUN
alces-443	585	2	of	of	ADP
alces-443	585	3	the	the	DET
alces-443	585	4	serum	serum	NOUN
alces-443	585	5	proteins	protein	NOUN
alces-443	585	6	and	and	CCONJ
alces-443	585	7	transferrins	transferrin	NOUN
alces-443	585	8	of	of	ADP
alces-443	585	9	alces	alces	PROPN
alces-443	585	10	alces	alce	NOUN
alces-443	585	11	(	(	PUNCT
alces-443	585	12	elk	elk	NOUN
alces-443	585	13	)	)	PUNCT
alces-443	585	14	,	,	PUNCT
alces-443	585	15	rangifer	rangifer	VERB
alces-443	585	16	tarandus	tarandus	NOUN
alces-443	585	17	(	(	PUNCT
alces-443	585	18	reindeer	reindeer	NOUN
alces-443	585	19	)	)	PUNCT
alces-443	585	20	,	,	PUNCT
alces-443	585	21	and	and	CCONJ
alces-443	585	22	ovis	ovis	PROPN
alces-443	585	23	dalli	dalli	NOUN
alces-443	585	24	(	(	PUNCT
alces-443	585	25	dall	dall	NOUN
alces-443	585	26	sheep	sheep	NOUN
alces-443	585	27	)	)	PUNCT
alces-443	585	28	from	from	ADP
alces-443	585	29	north	north	PROPN
alces-443	585	30	america	america	PROPN
alces-443	585	31	.	.	PUNCT
alces-443	586	1	comparative	comparative	ADJ
alces-443	586	2	biochemistry	biochemistry	NOUN
alces-443	586	3	and	and	CCONJ
alces-443	586	4	physiology	physiology	NOUN
alces-443	586	5	23:149157	23:149157	NUM
alces-443	586	6	.	.	PUNCT
alces-443	587	1	nevo	nevo	PROPN
alces-443	587	2	,	,	PUNCT
alces-443	587	3	e.	e.	PROPN
alces-443	587	4	1978	1978	NUM
alces-443	587	5	.	.	PUNCT
alces-443	588	1	genetic	genetic	ADJ
alces-443	588	2	variation	variation	NOUN
alces-443	588	3	in	in	ADP
alces-443	588	4	natural	natural	ADJ
alces-443	588	5	populations	population	NOUN
alces-443	588	6	:	:	PUNCT
alces-443	588	7	patterns	pattern	NOUN
alces-443	588	8	and	and	CCONJ
alces-443	588	9	theory	theory	NOUN
alces-443	588	10	.	.	PUNCT
alces-443	589	1	theoretical	theoretical	ADJ
alces-443	589	2	population	population	NOUN
alces-443	589	3	biology	biology	NOUN
alces-443	589	4	13:121	13:121	NUM
alces-443	589	5	-	-	SYM
alces-443	589	6	177	177	NUM
alces-443	589	7	.	.	PUNCT
alces-443	590	1	peterson	peterson	PROPN
alces-443	590	2	,	,	PUNCT
alces-443	590	3	r.	r.	PROPN
alces-443	590	4	l.	l.	PROPN
alces-443	590	5	1955	1955	NUM
alces-443	590	6	.	.	PUNCT
alces-443	591	1	north	north	ADJ
alces-443	591	2	american	american	PROPN
alces-443	591	3	moose	moose	NOUN
alces-443	591	4	.	.	PUNCT
alces-443	592	1	university	university	PROPN
alces-443	592	2	of	of	ADP
alces-443	592	3	toronto	toronto	PROPN
alces-443	592	4	press	press	PROPN
alces-443	592	5	,	,	PUNCT
alces-443	592	6	toronto	toronto	PROPN
alces-443	592	7	,	,	PUNCT
alces-443	592	8	ontario	ontario	PROPN
alces-443	592	9	,	,	PUNCT
alces-443	592	10	canada	canada	PROPN
alces-443	592	11	.	.	PUNCT
alces-443	593	1	quellar	quellar	PROPN
alces-443	593	2	,	,	PUNCT
alces-443	593	3	d.	d.	PROPN
alces-443	593	4	c.	c.	PROPN
alces-443	593	5	,	,	PUNCT
alces-443	593	6	j.	j.	PROPN
alces-443	593	7	e.	e.	PROPN
alces-443	593	8	strassmann	strassmann	PROPN
alces-443	593	9	,	,	PUNCT
alces-443	593	10	and	and	CCONJ
alces-443	593	11	c.	c.	PROPN
alces-443	593	12	r.	r.	PROPN
alces-443	593	13	hughes	hughes	PROPN
alces-443	593	14	.	.	PUNCT
alces-443	594	1	1993	1993	NUM
alces-443	594	2	.	.	PUNCT
alces-443	595	1	microsatellites	microsatellite	NOUN
alces-443	595	2	and	and	CCONJ
alces-443	595	3	kinship	kinship	NOUN
alces-443	595	4	.	.	PUNCT
alces-443	596	1	trends	trend	NOUN
alces-443	596	2	in	in	ADP
alces-443	596	3	ecology	ecology	NOUN
alces-443	596	4	and	and	CCONJ
alces-443	596	5	evolution	evolution	NOUN
alces-443	596	6	8:285	8:285	NUM
alces-443	596	7	-	-	SYM
alces-443	596	8	288	288	NUM
alces-443	596	9	.	.	PUNCT
alces-443	597	1	hundertmark	hundertmark	PROPN
alces-443	597	2	and	and	CCONJ
alces-443	597	3	bowyer	bowyer	PROPN
alces-443	597	4	moose	moose	PROPN
alces-443	597	5	genetics	genetics	PROPN
alces-443	597	6	alces	alce	NOUN
alces-443	597	7	vol	vol	NOUN
alces-443	597	8	.	.	PROPN
alces-443	597	9	40	40	NUM
alces-443	597	10	,	,	PUNCT
alces-443	597	11	2004	2004	NUM
alces-443	597	12	120	120	NUM
alces-443	597	13	røed	røed	NOUN
alces-443	597	14	,	,	PUNCT
alces-443	597	15	k.	k.	PROPN
alces-443	597	16	h.	h.	PROPN
alces-443	597	17	,	,	PUNCT
alces-443	597	18	and	and	CCONJ
alces-443	597	19	l.	l.	PROPN
alces-443	597	20	midthjell	midthjell	PROPN
alces-443	597	21	.	.	PUNCT
alces-443	598	1	1998	1998	NUM
alces-443	598	2	.	.	PUNCT
alces-443	599	1	microsatellites	microsatellite	NOUN
alces-443	599	2	in	in	ADP
alces-443	599	3	reindeer	reindeer	NOUN
alces-443	599	4	,	,	PUNCT
alces-443	599	5	rangifer	rangifer	VERB
alces-443	599	6	tarandus	tarandus	NOUN
alces-443	599	7	,	,	PUNCT
alces-443	599	8	and	and	CCONJ
alces-443	599	9	their	their	PRON
alces-443	599	10	use	use	NOUN
alces-443	599	11	in	in	ADP
alces-443	599	12	other	other	ADJ
alces-443	599	13	cervids	cervid	NOUN
alces-443	599	14	.	.	PUNCT
alces-443	600	1	molecular	molecular	ADJ
alces-443	600	2	ecology	ecology	NOUN
alces-443	600	3	7:1773	7:1773	PROPN
alces-443	600	4	-	-	PUNCT
alces-443	600	5	1776	1776	NUM
alces-443	600	6	.	.	PUNCT
alces-443	601	1	ryman	ryman	NOUN
alces-443	601	2	,	,	PUNCT
alces-443	601	3	n.	n.	PROPN
alces-443	601	4	,	,	PUNCT
alces-443	601	5	r.	r.	PROPN
alces-443	601	6	baccus	baccus	PROPN
alces-443	601	7	,	,	PUNCT
alces-443	601	8	c.	c.	PROPN
alces-443	601	9	reuterwall	reuterwall	NOUN
alces-443	601	10	,	,	PUNCT
alces-443	601	11	and	and	CCONJ
alces-443	601	12	m.	m.	PROPN
alces-443	601	13	h.	h.	PROPN
alces-443	601	14	smith	smith	PROPN
alces-443	601	15	.	.	PROPN
alces-443	601	16	1981	1981	NUM
alces-443	601	17	.	.	PUNCT
alces-443	602	1	effective	effective	ADJ
alces-443	602	2	population	population	NOUN
alces-443	602	3	size	size	NOUN
alces-443	602	4	,	,	PUNCT
alces-443	602	5	generation	generation	NOUN
alces-443	602	6	interval	interval	NOUN
alces-443	602	7	,	,	PUNCT
alces-443	602	8	and	and	CCONJ
alces-443	602	9	potential	potential	ADJ
alces-443	602	10	loss	loss	NOUN
alces-443	602	11	of	of	ADP
alces-443	602	12	genetic	genetic	ADJ
alces-443	602	13	variability	variability	NOUN
alces-443	602	14	in	in	ADP
alces-443	602	15	game	game	NOUN
alces-443	602	16	species	specie	NOUN
alces-443	602	17	under	under	ADP
alces-443	602	18	different	different	ADJ
alces-443	602	19	hunting	hunt	VERB
alces-443	602	20	regimes	regime	NOUN
alces-443	602	21	.	.	PUNCT
alces-443	603	1	oikos	oikos	PROPN
alces-443	603	2	36:257	36:257	PROPN
alces-443	603	3	-	-	SYM
alces-443	603	4	266	266	NUM
alces-443	603	5	.	.	PUNCT
alces-443	604	1	_	_	PUNCT
alces-443	605	1	_	_	PUNCT
alces-443	606	1	_	_	PUNCT
alces-443	607	1	_	_	PUNCT
alces-443	608	1	_	_	PUNCT
alces-443	608	2	,	,	PUNCT
alces-443	608	3	g.	g.	PROPN
alces-443	608	4	beckman	beckman	PROPN
alces-443	608	5	,	,	PUNCT
alces-443	608	6	g.	g.	PROPN
alces-443	608	7	bruun	bruun	PROPN
alces-443	608	8	-	-	PUNCT
alces-443	608	9	petersen	petersen	PROPN
alces-443	608	10	,	,	PUNCT
alces-443	608	11	and	and	CCONJ
alces-443	608	12	c.	c.	PROPN
alces-443	608	13	reuterwall	reuterwall	NOUN
alces-443	608	14	.	.	PUNCT
alces-443	609	1	1977	1977	NUM
alces-443	609	2	.	.	PUNCT
alces-443	610	1	variability	variability	NOUN
alces-443	610	2	of	of	ADP
alces-443	610	3	red	red	ADJ
alces-443	610	4	cell	cell	NOUN
alces-443	610	5	enzymes	enzyme	NOUN
alces-443	610	6	and	and	CCONJ
alces-443	610	7	genetic	genetic	ADJ
alces-443	610	8	implications	implication	NOUN
alces-443	610	9	of	of	ADP
alces-443	610	10	management	management	NOUN
alces-443	610	11	policies	policy	NOUN
alces-443	610	12	in	in	ADP
alces-443	610	13	scandinavian	scandinavian	ADJ
alces-443	610	14	moose	moose	NOUN
alces-443	610	15	(	(	PUNCT
alces-443	610	16	alces	alce	NOUN
alces-443	610	17	alces	alce	NOUN
alces-443	610	18	)	)	PUNCT
alces-443	610	19	.	.	PUNCT
alces-443	611	1	hereditas	hereditas	PROPN
alces-443	611	2	85:157	85:157	NUM
alces-443	611	3	-	-	SYM
alces-443	611	4	162	162	NUM
alces-443	611	5	.	.	PUNCT
alces-443	612	1	_	_	PUNCT
alces-443	613	1	_	_	PUNCT
alces-443	614	1	_	_	PUNCT
alces-443	615	1	_	_	PUNCT
alces-443	616	1	_	_	PUNCT
alces-443	616	2	,	,	PUNCT
alces-443	616	3	c.	c.	NOUN
alces-443	616	4	reuterwall	reuterwall	NOUN
alces-443	616	5	,	,	PUNCT
alces-443	616	6	k.	k.	PROPN
alces-443	616	7	nygren	nygren	PROPN
alces-443	616	8	,	,	PUNCT
alces-443	616	9	and	and	CCONJ
alces-443	616	10	t.	t.	PROPN
alces-443	616	11	nygren	nygren	PROPN
alces-443	616	12	.	.	PUNCT
alces-443	617	1	1980	1980	NUM
alces-443	617	2	.	.	PUNCT
alces-443	618	1	genetic	genetic	ADJ
alces-443	618	2	variation	variation	NOUN
alces-443	618	3	and	and	CCONJ
alces-443	618	4	differentiation	differentiation	NOUN
alces-443	618	5	in	in	ADP
alces-443	618	6	scandinavian	scandinavian	ADJ
alces-443	618	7	moose	moose	NOUN
alces-443	618	8	(	(	PUNCT
alces-443	618	9	alces	alce	NOUN
alces-443	618	10	alces	alce	NOUN
alces-443	618	11	):	):	PUNCT
alces-443	618	12	are	be	AUX
alces-443	618	13	large	large	ADJ
alces-443	618	14	mammals	mammal	NOUN
alces-443	618	15	monomorphic	monomorphic	ADJ
alces-443	618	16	.	.	PUNCT
alces-443	619	1	evolution	evolution	NOUN
alces-443	619	2	34:1037	34:1037	NOUN
alces-443	619	3	-	-	SYM
alces-443	619	4	1049	1049	NUM
alces-443	619	5	.	.	PUNCT
alces-443	620	1	sage	sage	NOUN
alces-443	620	2	,	,	PUNCT
alces-443	620	3	r.	r.	PROPN
alces-443	620	4	d.	d.	PROPN
alces-443	620	5	,	,	PUNCT
alces-443	620	6	and	and	CCONJ
alces-443	620	7	j.	j.	PROPN
alces-443	620	8	o.	o.	PROPN
alces-443	620	9	wolff	wolff	PROPN
alces-443	620	10	.	.	PUNCT
alces-443	621	1	1986	1986	NUM
alces-443	621	2	.	.	PUNCT
alces-443	622	1	pleistocene	pleistocene	NOUN
alces-443	622	2	glaciations	glaciation	NOUN
alces-443	622	3	,	,	PUNCT
alces-443	622	4	fluctuating	fluctuate	VERB
alces-443	622	5	ranges	range	NOUN
alces-443	622	6	,	,	PUNCT
alces-443	622	7	and	and	CCONJ
alces-443	622	8	low	low	ADJ
alces-443	622	9	genetic	genetic	ADJ
alces-443	622	10	variability	variability	NOUN
alces-443	622	11	in	in	ADP
alces-443	622	12	a	a	DET
alces-443	622	13	large	large	ADJ
alces-443	622	14	mammal	mammal	NOUN
alces-443	622	15	(	(	PUNCT
alces-443	622	16	ovis	ovis	ADJ
alces-443	622	17	dalli	dalli	NOUN
alces-443	622	18	)	)	PUNCT
alces-443	622	19	.	.	PUNCT
alces-443	623	1	evolution	evolution	NOUN
alces-443	623	2	40	40	NUM
alces-443	623	3	:	:	SYM
alces-443	623	4	1092	1092	NUM
alces-443	623	5	1095	1095	NUM
alces-443	623	6	.	.	PUNCT
alces-443	624	1	schwartz	schwartz	PROPN
alces-443	624	2	,	,	PUNCT
alces-443	624	3	c.	c.	PROPN
alces-443	624	4	c.	c.	PROPN
alces-443	624	5	,	,	PUNCT
alces-443	624	6	k.	k.	PROPN
alces-443	624	7	j.	j.	PROPN
alces-443	624	8	hundertmark	hundertmark	PROPN
alces-443	624	9	,	,	PUNCT
alces-443	624	10	and	and	CCONJ
alces-443	624	11	t.	t.	PROPN
alces-443	624	12	h.	h.	PROPN
alces-443	624	13	spraker	spraker	PROPN
alces-443	624	14	.	.	PUNCT
alces-443	625	1	1992	1992	NUM
alces-443	625	2	.	.	PUNCT
alces-443	626	1	an	an	DET
alces-443	626	2	evaluation	evaluation	NOUN
alces-443	626	3	of	of	ADP
alces-443	626	4	selective	selective	ADJ
alces-443	626	5	bull	bull	NOUN
alces-443	626	6	moose	moose	NOUN
alces-443	626	7	harvest	harvest	NOUN
alces-443	626	8	on	on	ADP
alces-443	626	9	the	the	DET
alces-443	626	10	kenai	kenai	PROPN
alces-443	626	11	peninsula	peninsula	PROPN
alces-443	626	12	,	,	PUNCT
alces-443	626	13	alaska	alaska	PROPN
alces-443	626	14	.	.	PUNCT
alces-443	626	15	alces	alce	VERB
alces-443	626	16	28:1	28:1	NUM
alces-443	626	17	-	-	SYM
alces-443	626	18	13	13	NUM
alces-443	626	19	.	.	PUNCT
alces-443	627	1	scribner	scribner	PROPN
alces-443	627	2	,	,	PUNCT
alces-443	627	3	k.	k.	PROPN
alces-443	627	4	t.	t.	PROPN
alces-443	627	5	,	,	PUNCT
alces-443	627	6	and	and	CCONJ
alces-443	627	7	r.	r.	PROPN
alces-443	627	8	k.	k.	PROPN
alces-443	627	9	chesser	chesser	PROPN
alces-443	627	10	.	.	PUNCT
alces-443	628	1	2001	2001	NUM
alces-443	628	2	.	.	PUNCT
alces-443	629	1	group	group	NOUN
alces-443	629	2	-	-	PUNCT
alces-443	629	3	structured	structure	VERB
alces-443	629	4	genetic	genetic	ADJ
alces-443	629	5	models	model	NOUN
alces-443	629	6	in	in	ADP
alces-443	629	7	analysis	analysis	NOUN
alces-443	629	8	of	of	ADP
alces-443	629	9	the	the	DET
alces-443	629	10	population	population	NOUN
alces-443	629	11	and	and	CCONJ
alces-443	629	12	behavioral	behavioral	ADJ
alces-443	629	13	ecology	ecology	NOUN
alces-443	629	14	of	of	ADP
alces-443	629	15	poikilothermic	poikilothermic	ADJ
alces-443	629	16	vertebrates	vertebrate	NOUN
alces-443	629	17	.	.	PUNCT
alces-443	630	1	journal	journal	NOUN
alces-443	630	2	of	of	ADP
alces-443	630	3	heredity	heredity	NOUN
alces-443	630	4	92:180	92:180	NUM
alces-443	630	5	-	-	SYM
alces-443	630	6	189	189	NUM
alces-443	630	7	.	.	PUNCT
alces-443	630	8	selander	selander	PROPN
alces-443	630	9	,	,	PUNCT
alces-443	630	10	r.	r.	PROPN
alces-443	630	11	k.	k.	PROPN
alces-443	630	12	,	,	PUNCT
alces-443	630	13	and	and	CCONJ
alces-443	630	14	d.	d.	PROPN
alces-443	630	15	w.	w.	PROPN
alces-443	630	16	kaufman	kaufman	PROPN
alces-443	630	17	.	.	PUNCT
alces-443	631	1	1973	1973	NUM
alces-443	631	2	.	.	PUNCT
alces-443	632	1	genic	genic	ADJ
alces-443	632	2	variability	variability	NOUN
alces-443	632	3	and	and	CCONJ
alces-443	632	4	strategies	strategy	NOUN
alces-443	632	5	of	of	ADP
alces-443	632	6	adaptation	adaptation	NOUN
alces-443	632	7	in	in	ADP
alces-443	632	8	animals	animal	NOUN
alces-443	632	9	.	.	PUNCT
alces-443	633	1	proceedings	proceeding	NOUN
alces-443	633	2	of	of	ADP
alces-443	633	3	the	the	DET
alces-443	633	4	national	national	PROPN
alces-443	633	5	academy	academy	PROPN
alces-443	633	6	of	of	ADP
alces-443	633	7	sciences	sciences	PROPN
alces-443	633	8	of	of	ADP
alces-443	633	9	the	the	DET
alces-443	633	10	usa	usa	PROPN
alces-443	633	11	70:1875	70:1875	PROPN
alces-443	633	12	-	-	PROPN
alces-443	633	13	1877	1877	NUM
alces-443	633	14	.	.	PUNCT
alces-443	634	1	sher	sher	PROPN
alces-443	634	2	,	,	PUNCT
alces-443	634	3	a.	a.	NOUN
alces-443	634	4	v.	v.	ADP
alces-443	634	5	1987	1987	NUM
alces-443	634	6	.	.	PUNCT
alces-443	635	1	history	history	NOUN
alces-443	635	2	and	and	CCONJ
alces-443	635	3	evolution	evolution	NOUN
alces-443	635	4	of	of	ADP
alces-443	635	5	moose	moose	NOUN
alces-443	635	6	.	.	PUNCT
alces-443	636	1	swedish	swedish	ADJ
alces-443	636	2	wildlife	wildlife	NOUN
alces-443	636	3	research	research	NOUN
alces-443	636	4	supplement	supplement	NOUN
alces-443	636	5	1:71	1:71	NUM
alces-443	636	6	-	-	SYM
alces-443	636	7	97	97	NUM
alces-443	636	8	.	.	PUNCT
alces-443	637	1	shubin	shubin	PROPN
alces-443	637	2	,	,	PUNCT
alces-443	637	3	p.	p.	PROPN
alces-443	637	4	n.	n.	PROPN
alces-443	637	5	1969	1969	NUM
alces-443	637	6	.	.	PUNCT
alces-443	638	1	the	the	DET
alces-443	638	2	genetics	genetic	NOUN
alces-443	638	3	of	of	ADP
alces-443	638	4	transferrins	transferrin	NOUN
alces-443	638	5	in	in	ADP
alces-443	638	6	the	the	DET
alces-443	638	7	reindeer	reindeer	NOUN
alces-443	638	8	and	and	CCONJ
alces-443	638	9	in	in	ADP
alces-443	638	10	the	the	DET
alces-443	638	11	european	european	PROPN
alces-443	638	12	elk	elk	PROPN
alces-443	638	13	.	.	PROPN
alces-443	638	14	genetika	genetika	NOUN
alces-443	638	15	5:37	5:37	NUM
alces-443	638	16	-	-	SYM
alces-443	638	17	41	41	NUM
alces-443	638	18	.	.	PUNCT
alces-443	639	1	(	(	PUNCT
alces-443	639	2	in	in	ADP
alces-443	639	3	russian	russian	NOUN
alces-443	639	4	)	)	PUNCT
alces-443	639	5	.	.	PUNCT
alces-443	640	1	smith	smith	PROPN
alces-443	640	2	,	,	PUNCT
alces-443	640	3	m.	m.	PROPN
alces-443	640	4	h.	h.	PROPN
alces-443	640	5	,	,	PUNCT
alces-443	640	6	r.	r.	PROPN
alces-443	640	7	baccus	baccus	PROPN
alces-443	640	8	,	,	PUNCT
alces-443	640	9	h.	h.	PROPN
alces-443	640	10	o.	o.	PROPN
alces-443	640	11	hillestad	hillestad	PROPN
alces-443	640	12	,	,	PUNCT
alces-443	640	13	and	and	CCONJ
alces-443	640	14	m.	m.	PROPN
alces-443	640	15	n.	n.	PROPN
alces-443	640	16	manlove	manlove	PROPN
alces-443	640	17	.	.	PUNCT
alces-443	640	18	1984	1984	NUM
alces-443	640	19	.	.	PUNCT
alces-443	641	1	population	population	NOUN
alces-443	641	2	genetics	genetic	NOUN
alces-443	641	3	.	.	PUNCT
alces-443	642	1	pages	page	NOUN
alces-443	642	2	119	119	NUM
alces-443	642	3	-	-	SYM
alces-443	642	4	128	128	NUM
alces-443	642	5	in	in	ADP
alces-443	642	6	l.	l.	PROPN
alces-443	642	7	k.	k.	PROPN
alces-443	642	8	halls	halls	PROPN
alces-443	642	9	,	,	PUNCT
alces-443	642	10	editor	editor	NOUN
alces-443	642	11	.	.	PUNCT
alces-443	643	1	white	white	ADJ
alces-443	643	2	-	-	PUNCT
alces-443	643	3	tailed	tail	VERB
alces-443	643	4	deer	deer	NOUN
alces-443	643	5	:	:	PUNCT
alces-443	643	6	ecology	ecology	NOUN
alces-443	643	7	and	and	CCONJ
alces-443	643	8	management	management	NOUN
alces-443	643	9	.	.	PUNCT
alces-443	644	1	stackpole	stackpole	PROPN
alces-443	644	2	books	books	PROPN
alces-443	644	3	,	,	PUNCT
alces-443	644	4	harrisburg	harrisburg	PROPN
alces-443	644	5	,	,	PUNCT
alces-443	644	6	pennsylvania	pennsylvania	PROPN
alces-443	644	7	,	,	PUNCT
alces-443	644	8	usa	usa	PROPN
alces-443	644	9	.	.	PUNCT
alces-443	645	1	_	_	PUNCT
alces-443	646	1	_	_	PUNCT
alces-443	647	1	_	_	PUNCT
alces-443	648	1	_	_	PUNCT
alces-443	649	1	_	_	PUNCT
alces-443	649	2	,	,	PUNCT
alces-443	649	3	k.	k.	PROPN
alces-443	649	4	t.	t.	PROPN
alces-443	649	5	scribner	scribner	PROPN
alces-443	649	6	,	,	PUNCT
alces-443	649	7	l.	l.	PROPN
alces-443	649	8	h.	h.	PROPN
alces-443	649	9	carpenter	carpenter	PROPN
alces-443	649	10	,	,	PUNCT
alces-443	649	11	and	and	CCONJ
alces-443	649	12	r.	r.	PROPN
alces-443	649	13	h.	h.	PROPN
alces-443	649	14	garrott	garrott	PROPN
alces-443	649	15	.	.	PUNCT
alces-443	650	1	1990	1990	NUM
alces-443	650	2	.	.	PUNCT
alces-443	651	1	genetic	genetic	ADJ
alces-443	651	2	characteristics	characteristic	NOUN
alces-443	651	3	of	of	ADP
alces-443	651	4	colorado	colorado	PROPN
alces-443	651	5	mule	mule	PROPN
alces-443	651	6	deer	deer	NOUN
alces-443	651	7	(	(	PUNCT
alces-443	651	8	odocoileus	odocoileus	ADJ
alces-443	651	9	hemionus	hemionus	PROPN
alces-443	651	10	)	)	PUNCT
alces-443	651	11	and	and	CCONJ
alces-443	651	12	comparisons	comparison	NOUN
alces-443	651	13	with	with	ADP
alces-443	651	14	other	other	ADJ
alces-443	651	15	cervids	cervid	NOUN
alces-443	651	16	.	.	PUNCT
alces-443	652	1	southwestern	southwestern	ADJ
alces-443	652	2	naturalist	naturalist	NOUN
alces-443	652	3	35	35	NUM
alces-443	652	4	:	:	SYM
alces-443	652	5	1	1	NUM
alces-443	652	6	8	8	NUM
alces-443	652	7	.	.	PUNCT
alces-443	653	1	symondson	symondson	PROPN
alces-443	653	2	,	,	PUNCT
alces-443	653	3	w.	w.	PROPN
alces-443	653	4	o.	o.	PROPN
alces-443	653	5	2002	2002	NUM
alces-443	653	6	.	.	PUNCT
alces-443	654	1	molecular	molecular	ADJ
alces-443	654	2	identification	identification	NOUN
alces-443	654	3	of	of	ADP
alces-443	654	4	prey	prey	NOUN
alces-443	654	5	in	in	ADP
alces-443	654	6	predator	predator	NOUN
alces-443	654	7	diets	diet	NOUN
alces-443	654	8	.	.	PUNCT
alces-443	655	1	molecular	molecular	ADJ
alces-443	655	2	ecology	ecology	NOUN
alces-443	655	3	11:627	11:627	NUM
alces-443	655	4	-	-	SYM
alces-443	655	5	641	641	NUM
alces-443	655	6	.	.	PUNCT
alces-443	655	7	telfer	telfer	PROPN
alces-443	655	8	,	,	PUNCT
alces-443	655	9	e.	e.	PROPN
alces-443	655	10	s.	s.	PROPN
alces-443	655	11	1984	1984	NUM
alces-443	655	12	.	.	PUNCT
alces-443	656	1	circumpolar	circumpolar	ADJ
alces-443	656	2	distribution	distribution	NOUN
alces-443	656	3	and	and	CCONJ
alces-443	656	4	habitat	habitat	NOUN
alces-443	656	5	requirements	requirement	NOUN
alces-443	656	6	of	of	ADP
alces-443	656	7	moose	moose	NOUN
alces-443	656	8	(	(	PUNCT
alces-443	656	9	alces	alce	NOUN
alces-443	656	10	alces	alce	NOUN
alces-443	656	11	)	)	PUNCT
alces-443	656	12	.	.	PUNCT
alces-443	657	1	pages	page	NOUN
alces-443	657	2	145	145	NUM
alces-443	657	3	-	-	SYM
alces-443	657	4	182	182	NUM
alces-443	657	5	in	in	ADP
alces-443	657	6	r.	r.	PROPN
alces-443	657	7	olson	olson	PROPN
alces-443	657	8	,	,	PUNCT
alces-443	657	9	f.	f.	PROPN
alces-443	657	10	geddes	geddes	PROPN
alces-443	657	11	,	,	PUNCT
alces-443	657	12	and	and	CCONJ
alces-443	657	13	r.	r.	PROPN
alces-443	657	14	hastings	hastings	PROPN
alces-443	657	15	,	,	PUNCT
alces-443	657	16	editors	editor	NOUN
alces-443	657	17	.	.	PUNCT
alces-443	658	1	northern	northern	ADJ
alces-443	658	2	ecology	ecology	NOUN
alces-443	658	3	and	and	CCONJ
alces-443	658	4	resource	resource	NOUN
alces-443	658	5	management	management	NOUN
alces-443	658	6	.	.	PUNCT
alces-443	659	1	university	university	NOUN
alces-443	659	2	of	of	ADP
alces-443	659	3	alberta	alberta	PROPN
alces-443	659	4	press	press	PROPN
alces-443	659	5	,	,	PUNCT
alces-443	659	6	edmonton	edmonton	PROPN
alces-443	659	7	,	,	PUNCT
alces-443	659	8	canada	canada	PROPN
alces-443	659	9	.	.	PUNCT
alces-443	660	1	thelen	thelen	VERB
alces-443	660	2	,	,	PUNCT
alces-443	660	3	t.	t.	PROPN
alces-443	660	4	h.	h.	PROPN
alces-443	660	5	1991	1991	NUM
alces-443	660	6	.	.	PUNCT
alces-443	661	1	effects	effect	NOUN
alces-443	661	2	of	of	ADP
alces-443	661	3	harvest	harvest	NOUN
alces-443	661	4	on	on	ADP
alces-443	661	5	antlers	antler	NOUN
alces-443	661	6	of	of	ADP
alces-443	661	7	simulated	simulated	ADJ
alces-443	661	8	populations	population	NOUN
alces-443	661	9	of	of	ADP
alces-443	661	10	elk	elk	PROPN
alces-443	661	11	.	.	PROPN
alces-443	661	12	journal	journal	PROPN
alces-443	661	13	of	of	ADP
alces-443	661	14	wildlife	wildlife	NOUN
alces-443	661	15	management	management	NOUN
alces-443	661	16	55:243249	55:243249	NUM
alces-443	661	17	.	.	PUNCT
alces-443	661	18	thouveny	thouveny	NOUN
alces-443	661	19	,	,	PUNCT
alces-443	661	20	n.	n.	NOUN
alces-443	661	21	,	,	PUNCT
alces-443	661	22	and	and	CCONJ
alces-443	661	23	e.	e.	PROPN
alces-443	661	24	bonifay	bonifay	PROPN
alces-443	661	25	.	.	PUNCT
alces-443	662	1	1984	1984	NUM
alces-443	662	2	.	.	PUNCT
alces-443	663	1	new	new	ADJ
alces-443	663	2	chronological	chronological	ADJ
alces-443	663	3	data	datum	NOUN
alces-443	663	4	on	on	ADP
alces-443	663	5	european	european	ADJ
alces-443	663	6	plio	plio	NOUN
alces-443	663	7	-	-	PUNCT
alces-443	663	8	pleistocene	pleistocene	NOUN
alces-443	663	9	faunas	fauna	NOUN
alces-443	663	10	and	and	CCONJ
alces-443	663	11	hominid	hominid	PROPN
alces-443	663	12	occupation	occupation	NOUN
alces-443	663	13	sites	site	NOUN
alces-443	663	14	.	.	PUNCT
alces-443	664	1	nature	nature	NOUN
alces-443	664	2	308:355	308:355	PROPN
alces-443	664	3	-	-	PUNCT
alces-443	664	4	358	358	NUM
alces-443	664	5	.	.	PUNCT
alces-443	665	1	udina	udina	PROPN
alces-443	665	2	,	,	PUNCT
alces-443	665	3	i.	i.	PROPN
alces-443	665	4	g.	g.	PROPN
alces-443	665	5	,	,	PUNCT
alces-443	665	6	a.	a.	NOUN
alces-443	665	7	a.	a.	NOUN
alces-443	665	8	danilkin	danilkin	PROPN
alces-443	665	9	,	,	PUNCT
alces-443	665	10	and	and	CCONJ
alces-443	665	11	g.	g.	PROPN
alces-443	665	12	g.	g.	PROPN
alces-443	665	13	boeskorov	boeskorov	PROPN
alces-443	665	14	.	.	PUNCT
alces-443	666	1	2002	2002	NUM
alces-443	666	2	.	.	PUNCT
alces-443	667	1	genetic	genetic	ADJ
alces-443	667	2	diversity	diversity	NOUN
alces-443	667	3	of	of	ADP
alces-443	667	4	moose	moose	NOUN
alces-443	667	5	(	(	PUNCT
alces-443	667	6	alces	alces	PROPN
alces-443	667	7	alces	alce	NOUN
alces-443	667	8	l.	l.	PROPN
alces-443	667	9	)	)	PUNCT
alces-443	667	10	in	in	ADP
alces-443	667	11	eurasia	eurasia	PROPN
alces-443	667	12	.	.	PUNCT
alces-443	668	1	genetika	genetika	PROPN
alces-443	668	2	38:951	38:951	NUM
alces-443	668	3	-	-	SYM
alces-443	668	4	957	957	NUM
alces-443	668	5	.	.	PUNCT
alces-443	669	1	(	(	PUNCT
alces-443	669	2	in	in	ADP
alces-443	669	3	russian	russian	NOUN
alces-443	669	4	)	)	PUNCT
alces-443	669	5	.	.	PUNCT
alces-443	670	1	vernesi	vernesi	PROPN
alces-443	670	2	,	,	PUNCT
alces-443	670	3	c.	c.	PROPN
alces-443	670	4	,	,	PUNCT
alces-443	670	5	e.	e.	PROPN
alces-443	670	6	pecchiolo	pecchiolo	PROPN
alces-443	670	7	,	,	PUNCT
alces-443	670	8	d.	d.	PROPN
alces-443	670	9	caramelli	caramelli	PROPN
alces-443	670	10	,	,	PUNCT
alces-443	670	11	r.	r.	PROPN
alces-443	670	12	tiedemann	tiedemann	PROPN
alces-443	670	13	,	,	PUNCT
alces-443	670	14	e.	e.	PROPN
alces-443	670	15	randi	randi	PROPN
alces-443	670	16	,	,	PUNCT
alces-443	670	17	and	and	CCONJ
alces-443	670	18	g.	g.	PROPN
alces-443	670	19	bertorelle	bertorelle	PROPN
alces-443	670	20	.	.	PUNCT
alces-443	671	1	2002	2002	NUM
alces-443	671	2	.	.	PUNCT
alces-443	672	1	the	the	DET
alces-443	672	2	genetic	genetic	ADJ
alces-443	672	3	structure	structure	NOUN
alces-443	672	4	of	of	ADP
alces-443	672	5	natural	natural	ADJ
alces-443	672	6	and	and	CCONJ
alces-443	672	7	reintroduced	reintroduce	VERB
alces-443	672	8	roe	roe	NOUN
alces-443	672	9	deer	deer	NOUN
alces-443	672	10	(	(	PUNCT
alces-443	672	11	capreolus	capreolus	ADJ
alces-443	672	12	capreolus	capreolus	ADJ
alces-443	672	13	)	)	PUNCT
alces-443	672	14	populations	population	NOUN
alces-443	672	15	in	in	ADP
alces-443	672	16	the	the	DET
alces-443	672	17	alps	alps	PROPN
alces-443	672	18	and	and	CCONJ
alces-443	672	19	central	central	PROPN
alces-443	672	20	italy	italy	PROPN
alces-443	672	21	,	,	PUNCT
alces-443	672	22	with	with	ADP
alces-443	672	23	reference	reference	NOUN
alces-443	672	24	to	to	ADP
alces-443	672	25	the	the	DET
alces-443	672	26	mitochondrial	mitochondrial	ADJ
alces-443	672	27	dna	dna	NOUN
alces-443	672	28	phylogeography	phylogeography	NOUN
alces-443	672	29	of	of	ADP
alces-443	672	30	europe	europe	PROPN
alces-443	672	31	.	.	PUNCT
alces-443	673	1	molecular	molecular	ADJ
alces-443	673	2	ecology	ecology	NOUN
alces-443	673	3	11:1285	11:1285	NUM
alces-443	673	4	-	-	SYM
alces-443	673	5	1297	1297	NUM
alces-443	673	6	.	.	PUNCT
alces-443	673	7	wilhelmson	wilhelmson	PROPN
alces-443	673	8	,	,	PUNCT
alces-443	673	9	m.	m.	NOUN
alces-443	673	10	,	,	PUNCT
alces-443	673	11	r.	r.	PROPN
alces-443	673	12	k.	k.	PROPN
alces-443	673	13	juneja	juneja	PROPN
alces-443	673	14	,	,	PUNCT
alces-443	673	15	and	and	CCONJ
alces-443	673	16	s.	s.	PROPN
alces-443	673	17	bengtsson	bengtsson	PROPN
alces-443	673	18	.	.	PUNCT
alces-443	674	1	1978	1978	NUM
alces-443	674	2	.	.	PUNCT
alces-443	675	1	lack	lack	NOUN
alces-443	675	2	of	of	ADP
alces-443	675	3	polymorphism	polymorphism	NOUN
alces-443	675	4	in	in	ADP
alces-443	675	5	certain	certain	ADJ
alces-443	675	6	blood	blood	NOUN
alces-443	675	7	proteins	protein	NOUN
alces-443	675	8	and	and	CCONJ
alces-443	675	9	enalces	enalce	NOUN
alces-443	675	10	vol	vol	NOUN
alces-443	675	11	.	.	PROPN
alces-443	675	12	40	40	NUM
alces-443	675	13	,	,	PUNCT
alces-443	675	14	2004	2004	NUM
alces-443	675	15	moose	moose	NOUN
alces-443	675	16	genetics	genetic	NOUN
alces-443	675	17	hundertmark	hundertmark	NOUN
alces-443	675	18	and	and	CCONJ
alces-443	675	19	bowyer	bowyer	PROPN
alces-443	675	20	121	121	NUM
alces-443	675	21	zymes	zyme	NOUN
alces-443	675	22	of	of	ADP
alces-443	675	23	european	european	ADJ
alces-443	675	24	and	and	CCONJ
alces-443	675	25	canadian	canadian	ADJ
alces-443	675	26	moose	moose	NOUN
alces-443	675	27	.	.	PUNCT
alces-443	676	1	naturaliste	naturaliste	PROPN
alces-443	676	2	canadien	canadien	PROPN
alces-443	676	3	105:445449	105:445449	PROPN
alces-443	676	4	.	.	PUNCT
alces-443	677	1	wilson	wilson	PROPN
alces-443	677	2	,	,	PUNCT
alces-443	677	3	g.	g.	PROPN
alces-443	677	4	a.	a.	PROPN
alces-443	677	5	,	,	PUNCT
alces-443	677	6	c.	c.	PROPN
alces-443	677	7	strobeck	strobeck	PROPN
alces-443	677	8	,	,	PUNCT
alces-443	677	9	l.	l.	PROPN
alces-443	677	10	wu	wu	PROPN
alces-443	677	11	,	,	PUNCT
alces-443	677	12	and	and	CCONJ
alces-443	677	13	j.	j.	PROPN
alces-443	677	14	w.	w.	PROPN
alces-443	677	15	coffin	coffin	PROPN
alces-443	677	16	.	.	PUNCT
alces-443	678	1	1997	1997	NUM
alces-443	678	2	.	.	PUNCT
alces-443	679	1	characterization	characterization	NOUN
alces-443	679	2	of	of	ADP
alces-443	679	3	microsatellite	microsatellite	ADJ
alces-443	679	4	loci	locus	NOUN
alces-443	679	5	in	in	ADP
alces-443	679	6	caribou	caribou	PROPN
alces-443	679	7	rangifer	rangifer	PROPN
alces-443	679	8	tarandus	tarandus	PROPN
alces-443	679	9	,	,	PUNCT
alces-443	679	10	and	and	CCONJ
alces-443	679	11	their	their	PRON
alces-443	679	12	use	use	NOUN
alces-443	679	13	in	in	ADP
alces-443	679	14	other	other	ADJ
alces-443	679	15	artiodactyls	artiodactyls	NOUN
alces-443	679	16	.	.	PUNCT
alces-443	680	1	molecular	molecular	ADJ
alces-443	680	2	ecology	ecology	NOUN
alces-443	680	3	6:697699	6:697699	PROPN
alces-443	680	4	.	.	PUNCT
alces-443	680	5	wilson	wilson	PROPN
alces-443	680	6	,	,	PUNCT
alces-443	680	7	p.	p.	PROPN
alces-443	680	8	j.	j.	PROPN
alces-443	680	9	,	,	PUNCT
alces-443	680	10	s.	s.	PROPN
alces-443	680	11	grewal	grewal	PROPN
alces-443	680	12	,	,	PUNCT
alces-443	680	13	a.	a.	NOUN
alces-443	680	14	rodgers	rodgers	PROPN
alces-443	680	15	,	,	PUNCT
alces-443	680	16	r.	r.	PROPN
alces-443	680	17	rempel	rempel	PROPN
alces-443	680	18	,	,	PUNCT
alces-443	680	19	j.	j.	PROPN
alces-443	680	20	saquet	saquet	PROPN
alces-443	680	21	,	,	PUNCT
alces-443	680	22	h.	h.	PROPN
alces-443	680	23	hristienko	hristienko	PROPN
alces-443	680	24	,	,	PUNCT
alces-443	680	25	f.	f.	PROPN
alces-443	680	26	burrows	burrows	PROPN
alces-443	680	27	,	,	PUNCT
alces-443	680	28	r.	r.	PROPN
alces-443	680	29	peterson	peterson	PROPN
alces-443	680	30	,	,	PUNCT
alces-443	680	31	and	and	CCONJ
alces-443	680	32	b.	b.	PROPN
alces-443	680	33	n.	n.	PROPN
alces-443	680	34	white	white	PROPN
alces-443	680	35	.	.	PUNCT
alces-443	681	1	2003	2003	NUM
alces-443	681	2	.	.	PUNCT
alces-443	682	1	genetic	genetic	ADJ
alces-443	682	2	variation	variation	NOUN
alces-443	682	3	and	and	CCONJ
alces-443	682	4	population	population	NOUN
alces-443	682	5	structure	structure	NOUN
alces-443	682	6	of	of	ADP
alces-443	682	7	moose	moose	NOUN
alces-443	682	8	(	(	PUNCT
alces-443	682	9	alces	alce	NOUN
alces-443	682	10	alces	alce	NOUN
alces-443	682	11	)	)	PUNCT
alces-443	682	12	at	at	ADP
alces-443	682	13	neutral	neutral	ADJ
alces-443	682	14	and	and	CCONJ
alces-443	682	15	functional	functional	ADJ
alces-443	682	16	dna	dna	PROPN
alces-443	682	17	loci	loci	PROPN
alces-443	682	18	.	.	PUNCT
alces-443	683	1	canadian	canadian	ADJ
alces-443	683	2	journal	journal	PROPN
alces-443	683	3	of	of	ADP
alces-443	683	4	zoology	zoology	NOUN
alces-443	683	5	81:670	81:670	NOUN
alces-443	683	6	-	-	SYM
alces-443	683	7	683	683	NUM
alces-443	683	8	.	.	PUNCT
alces-443	684	1	zheleznov	zheleznov	NOUN
alces-443	684	2	,	,	PUNCT
alces-443	684	3	n.	n.	PROPN
alces-443	684	4	k.	k.	PROPN
alces-443	684	5	1993	1993	NUM
alces-443	684	6	.	.	PUNCT
alces-443	685	1	historic	historic	ADJ
alces-443	685	2	and	and	CCONJ
alces-443	685	3	current	current	ADJ
alces-443	685	4	distribution	distribution	NOUN
alces-443	685	5	of	of	ADP
alces-443	685	6	moose	moose	NOUN
alces-443	685	7	in	in	ADP
alces-443	685	8	the	the	DET
alces-443	685	9	northeast	northeast	ADJ
alces-443	685	10	ussr	ussr	PROPN
alces-443	685	11	.	.	PUNCT
alces-443	686	1	alces	alce	VERB
alces-443	686	2	29:213	29:213	NUM
alces-443	686	3	-	-	SYM
alces-443	686	4	218	218	NUM
alces-443	686	5	.	.	PUNCT
alces-443	687	1	hundertmark	hundertmark	PROPN
alces-443	687	2	and	and	CCONJ
alces-443	687	3	bowyer	bowyer	PROPN
alces-443	687	4	moose	moose	PROPN
alces-443	687	5	genetics	genetics	PROPN
alces-443	687	6	alces	alce	NOUN
alces-443	687	7	vol	vol	NOUN
alces-443	687	8	.	.	PROPN
alces-443	687	9	40	40	NUM
alces-443	687	10	,	,	PUNCT
alces-443	687	11	2004	2004	NUM
alces-443	687	12	122	122	NUM
alces-443	687	13	term	term	NOUN
alces-443	687	14	definition	definition	NOUN
alces-443	687	15	adaptive	adaptive	ADJ
alces-443	687	16	genetic	genetic	ADJ
alces-443	687	17	variation	variation	NOUN
alces-443	687	18	genotypic	genotypic	NOUN
alces-443	687	19	variation	variation	NOUN
alces-443	687	20	at	at	ADP
alces-443	687	21	loci	locus	NOUN
alces-443	687	22	that	that	SCONJ
alces-443	687	23	control	control	NOUN
alces-443	687	24	traits	trait	NOUN
alces-443	687	25	important	important	ADJ
alces-443	687	26	to	to	ADP
alces-443	687	27	fitness	fitness	NOUN
alces-443	687	28	,	,	PUNCT
alces-443	687	29	such	such	ADJ
alces-443	687	30	as	as	ADP
alces-443	687	31	morphology	morphology	NOUN
alces-443	687	32	,	,	PUNCT
alces-443	687	33	physiology	physiology	NOUN
alces-443	687	34	,	,	PUNCT
alces-443	687	35	and	and	CCONJ
alces-443	687	36	behavior	behavior	NOUN
alces-443	687	37	.	.	PUNCT
alces-443	688	1	allozyme	allozyme	NOUN
alces-443	688	2	a	a	DET
alces-443	688	3	gene	gene	NOUN
alces-443	688	4	product	product	NOUN
alces-443	688	5	(	(	PUNCT
alces-443	688	6	protein	protein	NOUN
alces-443	688	7	)	)	PUNCT
alces-443	688	8	that	that	PRON
alces-443	688	9	is	be	AUX
alces-443	688	10	distinguished	distinguish	VERB
alces-443	688	11	by	by	ADP
alces-443	688	12	its	its	PRON
alces-443	688	13	migratory	migratory	ADJ
alces-443	688	14	characteristics	characteristic	NOUN
alces-443	688	15	in	in	ADP
alces-443	688	16	a	a	DET
alces-443	688	17	gel	gel	NOUN
alces-443	688	18	exposed	expose	VERB
alces-443	688	19	to	to	ADP
alces-443	688	20	an	an	DET
alces-443	688	21	electric	electric	ADJ
alces-443	688	22	field	field	NOUN
alces-443	688	23	(	(	PUNCT
alces-443	688	24	electrophoresis	electrophoresis	NOUN
alces-443	688	25	)	)	PUNCT
alces-443	688	26	.	.	PUNCT
alces-443	689	1	differences	difference	NOUN
alces-443	689	2	among	among	ADP
alces-443	689	3	alleles	allele	NOUN
alces-443	689	4	(	(	PUNCT
alces-443	689	5	different	different	ADJ
alces-443	689	6	variants	variant	NOUN
alces-443	689	7	of	of	ADP
alces-443	689	8	the	the	DET
alces-443	689	9	same	same	ADJ
alces-443	689	10	gene	gene	NOUN
alces-443	689	11	)	)	PUNCT
alces-443	689	12	at	at	ADP
alces-443	689	13	allozyme	allozyme	NOUN
alces-443	689	14	loci	locus	NOUN
alces-443	689	15	relate	relate	VERB
alces-443	689	16	to	to	ADP
alces-443	689	17	amino	amino	NOUN
alces-443	689	18	acid	acid	NOUN
alces-443	689	19	composition	composition	NOUN
alces-443	689	20	and	and	CCONJ
alces-443	689	21	secondary	secondary	ADJ
alces-443	689	22	structure	structure	NOUN
alces-443	689	23	of	of	ADP
alces-443	689	24	the	the	DET
alces-443	689	25	protein	protein	NOUN
alces-443	689	26	.	.	PUNCT
alces-443	690	1	only	only	ADV
alces-443	690	2	mutations	mutation	NOUN
alces-443	690	3	that	that	PRON
alces-443	690	4	create	create	VERB
alces-443	690	5	proteins	protein	NOUN
alces-443	690	6	with	with	ADP
alces-443	690	7	different	different	ADJ
alces-443	690	8	migratory	migratory	NOUN
alces-443	690	9	characteristics	characteristic	NOUN
alces-443	690	10	are	be	AUX
alces-443	690	11	detectable	detectable	ADJ
alces-443	690	12	.	.	PUNCT
alces-443	691	1	control	control	NOUN
alces-443	691	2	region	region	PROPN
alces-443	691	3	a	a	DET
alces-443	691	4	portion	portion	NOUN
alces-443	691	5	of	of	ADP
alces-443	691	6	mtdna	mtdna	NOUN
alces-443	691	7	that	that	PRON
alces-443	691	8	evolves	evolves	AUX
alces-443	691	9	(	(	PUNCT
alces-443	691	10	incorporates	incorporate	VERB
alces-443	691	11	mutations	mutation	NOUN
alces-443	691	12	)	)	PUNCT
alces-443	691	13	at	at	ADP
alces-443	691	14	a	a	DET
alces-443	691	15	very	very	ADV
alces-443	691	16	fast	fast	ADJ
alces-443	691	17	rate	rate	NOUN
alces-443	691	18	,	,	PUNCT
alces-443	691	19	which	which	PRON
alces-443	691	20	makes	make	VERB
alces-443	691	21	it	it	PRON
alces-443	691	22	a	a	DET
alces-443	691	23	valuable	valuable	ADJ
alces-443	691	24	marker	marker	NOUN
alces-443	691	25	for	for	ADP
alces-443	691	26	examining	examine	VERB
alces-443	691	27	intraspecific	intraspecific	NOUN
alces-443	691	28	genetic	genetic	ADJ
alces-443	691	29	variation	variation	NOUN
alces-443	691	30	.	.	PUNCT
alces-443	692	1	occasionally	occasionally	ADV
alces-443	692	2	referred	refer	VERB
alces-443	692	3	to	to	ADP
alces-443	692	4	as	as	ADP
alces-443	692	5	the	the	DET
alces-443	692	6	d	d	NOUN
alces-443	692	7	-	-	PUNCT
alces-443	692	8	loop	loop	NOUN
alces-443	692	9	.	.	PUNCT
alces-443	693	1	dna	dna	PROPN
alces-443	693	2	sequence	sequence	NOUN
alces-443	693	3	the	the	DET
alces-443	693	4	ultimate	ultimate	ADJ
alces-443	693	5	level	level	NOUN
alces-443	693	6	of	of	ADP
alces-443	693	7	analysis	analysis	NOUN
alces-443	693	8	of	of	ADP
alces-443	693	9	genetic	genetic	ADJ
alces-443	693	10	material	material	NOUN
alces-443	693	11	.	.	PUNCT
alces-443	694	1	this	this	DET
alces-443	694	2	technique	technique	NOUN
alces-443	694	3	deduces	deduce	VERB
alces-443	694	4	the	the	DET
alces-443	694	5	identity	identity	NOUN
alces-443	694	6	and	and	CCONJ
alces-443	694	7	order	order	NOUN
alces-443	694	8	of	of	ADP
alces-443	694	9	nucleotides	nucleotide	NOUN
alces-443	694	10	in	in	ADP
alces-443	694	11	a	a	DET
alces-443	694	12	fragment	fragment	NOUN
alces-443	694	13	of	of	ADP
alces-443	694	14	dna	dna	PROPN
alces-443	694	15	.	.	PUNCT
alces-443	695	1	mutations	mutation	NOUN
alces-443	695	2	(	(	PUNCT
alces-443	695	3	nucleotide	nucleotide	NOUN
alces-443	695	4	substitutions	substitution	NOUN
alces-443	695	5	)	)	PUNCT
alces-443	695	6	are	be	AUX
alces-443	695	7	detectable	detectable	ADJ
alces-443	695	8	whether	whether	SCONJ
alces-443	695	9	or	or	CCONJ
alces-443	695	10	not	not	PART
alces-443	695	11	they	they	PRON
alces-443	695	12	create	create	VERB
alces-443	695	13	different	different	ADJ
alces-443	695	14	gene	gene	NOUN
alces-443	695	15	products	product	NOUN
alces-443	695	16	.	.	PUNCT
alces-443	696	1	effective	effective	ADJ
alces-443	696	2	population	population	NOUN
alces-443	696	3	size	size	NOUN
alces-443	696	4	the	the	DET
alces-443	696	5	size	size	NOUN
alces-443	696	6	of	of	ADP
alces-443	696	7	a	a	DET
alces-443	696	8	standardized	standardized	ADJ
alces-443	696	9	population	population	NOUN
alces-443	696	10	that	that	PRON
alces-443	696	11	has	have	VERB
alces-443	696	12	the	the	DET
alces-443	696	13	same	same	ADJ
alces-443	696	14	degree	degree	NOUN
alces-443	696	15	of	of	ADP
alces-443	696	16	genetic	genetic	ADJ
alces-443	696	17	drift	drift	NOUN
alces-443	696	18	as	as	SCONJ
alces-443	696	19	the	the	DET
alces-443	696	20	population	population	NOUN
alces-443	696	21	being	be	AUX
alces-443	696	22	studied	study	VERB
alces-443	696	23	.	.	PUNCT
alces-443	697	1	an	an	DET
alces-443	697	2	ideal	ideal	ADJ
alces-443	697	3	population	population	NOUN
alces-443	697	4	is	be	AUX
alces-443	697	5	a	a	DET
alces-443	697	6	closed	closed	ADJ
alces-443	697	7	population	population	NOUN
alces-443	697	8	of	of	ADP
alces-443	697	9	constant	constant	ADJ
alces-443	697	10	size	size	NOUN
alces-443	697	11	with	with	ADP
alces-443	697	12	non	non	ADJ
alces-443	697	13	-	-	ADJ
alces-443	697	14	overlapping	overlapping	ADJ
alces-443	697	15	generations	generation	NOUN
alces-443	697	16	and	and	CCONJ
alces-443	697	17	no	no	DET
alces-443	697	18	variance	variance	NOUN
alces-443	697	19	in	in	ADP
alces-443	697	20	reproductive	reproductive	ADJ
alces-443	697	21	success	success	NOUN
alces-443	697	22	.	.	PUNCT
alces-443	698	1	the	the	PRON
alces-443	698	2	smaller	small	ADJ
alces-443	698	3	the	the	DET
alces-443	698	4	effective	effective	ADJ
alces-443	698	5	size	size	NOUN
alces-443	698	6	of	of	ADP
alces-443	698	7	a	a	DET
alces-443	698	8	population	population	NOUN
alces-443	698	9	,	,	PUNCT
alces-443	698	10	the	the	PRON
alces-443	698	11	faster	fast	ADV
alces-443	698	12	it	it	PRON
alces-443	698	13	will	will	AUX
alces-443	698	14	lose	lose	VERB
alces-443	698	15	genetic	genetic	ADJ
alces-443	698	16	diversity	diversity	NOUN
alces-443	698	17	through	through	ADP
alces-443	698	18	drift	drift	NOUN
alces-443	698	19	regardless	regardless	ADV
alces-443	698	20	of	of	ADP
alces-443	698	21	actual	actual	ADJ
alces-443	698	22	population	population	NOUN
alces-443	698	23	size	size	NOUN
alces-443	698	24	.	.	PUNCT
alces-443	699	1	effective	effective	ADJ
alces-443	699	2	population	population	NOUN
alces-443	699	3	size	size	NOUN
alces-443	699	4	(	(	PUNCT
alces-443	699	5	ne	ne	NOUN
alces-443	699	6	)	)	PUNCT
alces-443	699	7	is	be	AUX
alces-443	699	8	almost	almost	ADV
alces-443	699	9	always	always	ADV
alces-443	699	10	a	a	DET
alces-443	699	11	fraction	fraction	NOUN
alces-443	699	12	of	of	ADP
alces-443	699	13	the	the	DET
alces-443	699	14	true	true	ADJ
alces-443	699	15	population	population	NOUN
alces-443	699	16	size	size	NOUN
alces-443	699	17	(	(	PUNCT
alces-443	699	18	n	n	CCONJ
alces-443	699	19	)	)	PUNCT
alces-443	699	20	.	.	PUNCT
alces-443	700	1	generation	generation	NOUN
alces-443	700	2	interval	interval	NOUN
alces-443	700	3	mean	mean	VERB
alces-443	700	4	age	age	NOUN
alces-443	700	5	of	of	ADP
alces-443	700	6	all	all	DET
alces-443	700	7	parents	parent	NOUN
alces-443	700	8	.	.	PUNCT
alces-443	701	1	genetic	genetic	ADJ
alces-443	701	2	drift	drift	NOUN
alces-443	701	3	changes	change	NOUN
alces-443	701	4	in	in	ADP
alces-443	701	5	allele	allele	NOUN
alces-443	701	6	frequencies	frequency	NOUN
alces-443	701	7	across	across	ADP
alces-443	701	8	generations	generation	NOUN
alces-443	701	9	due	due	ADP
alces-443	701	10	to	to	ADP
alces-443	701	11	random	random	ADJ
alces-443	701	12	sampling	sampling	NOUN
alces-443	701	13	error	error	NOUN
alces-443	701	14	associated	associate	VERB
alces-443	701	15	with	with	ADP
alces-443	701	16	less	less	ADJ
alces-443	701	17	than	than	ADP
alces-443	701	18	infinite	infinite	ADJ
alces-443	701	19	population	population	NOUN
alces-443	701	20	size	size	NOUN
alces-443	701	21	.	.	PUNCT
alces-443	702	1	haplotype	haplotype	VERB
alces-443	702	2	the	the	DET
alces-443	702	3	haploid	haploid	NOUN
alces-443	702	4	equivalent	equivalent	NOUN
alces-443	702	5	of	of	ADP
alces-443	702	6	genotype	genotype	NOUN
alces-443	702	7	.	.	PUNCT
alces-443	703	1	genetic	genetic	ADJ
alces-443	703	2	type	type	NOUN
alces-443	703	3	of	of	ADP
alces-443	703	4	an	an	DET
alces-443	703	5	individual	individual	NOUN
alces-443	703	6	when	when	SCONJ
alces-443	703	7	haploid	haploid	ADJ
alces-443	703	8	dna	dna	NOUN
alces-443	703	9	,	,	PUNCT
alces-443	703	10	such	such	ADJ
alces-443	703	11	as	as	ADP
alces-443	703	12	mtdna	mtdna	PROPN
alces-443	703	13	,	,	PUNCT
alces-443	703	14	is	be	AUX
alces-443	703	15	analyzed	analyze	VERB
alces-443	703	16	.	.	PUNCT
alces-443	704	1	heritability	heritability	NOUN
alces-443	704	2	the	the	DET
alces-443	704	3	proportion	proportion	NOUN
alces-443	704	4	of	of	ADP
alces-443	704	5	variance	variance	NOUN
alces-443	704	6	in	in	ADP
alces-443	704	7	the	the	DET
alces-443	704	8	expression	expression	NOUN
alces-443	704	9	of	of	ADP
alces-443	704	10	a	a	DET
alces-443	704	11	trait	trait	NOUN
alces-443	704	12	,	,	PUNCT
alces-443	704	13	such	such	ADJ
alces-443	704	14	as	as	ADP
alces-443	704	15	antler	antler	NOUN
alces-443	704	16	size	size	NOUN
alces-443	704	17	or	or	CCONJ
alces-443	704	18	body	body	NOUN
alces-443	704	19	size	size	NOUN
alces-443	704	20	,	,	PUNCT
alces-443	704	21	that	that	PRON
alces-443	704	22	is	be	AUX
alces-443	704	23	due	due	ADJ
alces-443	704	24	to	to	ADP
alces-443	704	25	genetic	genetic	ADJ
alces-443	704	26	effects	effect	NOUN
alces-443	704	27	(	(	PUNCT
alces-443	704	28	as	as	SCONJ
alces-443	704	29	opposed	oppose	VERB
alces-443	704	30	to	to	ADP
alces-443	704	31	environmental	environmental	ADJ
alces-443	704	32	effects	effect	NOUN
alces-443	704	33	)	)	PUNCT
alces-443	704	34	,	,	PUNCT
alces-443	704	35	i.e.	i.e.	X
alces-443	704	36	,	,	PUNCT
alces-443	704	37	the	the	DET
alces-443	704	38	degree	degree	NOUN
alces-443	704	39	to	to	PART
alces-443	704	40	which	which	PRON
alces-443	704	41	a	a	DET
alces-443	704	42	trait	trait	NOUN
alces-443	704	43	can	can	AUX
alces-443	704	44	be	be	AUX
alces-443	704	45	passed	pass	VERB
alces-443	704	46	on	on	ADP
alces-443	704	47	to	to	ADP
alces-443	704	48	the	the	DET
alces-443	704	49	next	next	ADJ
alces-443	704	50	generation	generation	NOUN
alces-443	704	51	.	.	PUNCT
alces-443	705	1	microsatellite	microsatellite	ADJ
alces-443	705	2	segments	segment	NOUN
alces-443	705	3	of	of	ADP
alces-443	705	4	dna	dna	PROPN
alces-443	705	5	composed	compose	VERB
alces-443	705	6	of	of	ADP
alces-443	705	7	sequence	sequence	NOUN
alces-443	705	8	units	unit	NOUN
alces-443	705	9	varying	vary	VERB
alces-443	705	10	from	from	ADP
alces-443	705	11	2	2	NUM
alces-443	705	12	-	-	SYM
alces-443	705	13	4	4	NUM
alces-443	705	14	nucleotides	nucleotide	NOUN
alces-443	705	15	in	in	ADP
alces-443	705	16	length	length	NOUN
alces-443	705	17	that	that	PRON
alces-443	705	18	are	be	AUX
alces-443	705	19	tandemly	tandemly	ADV
alces-443	705	20	repeated	repeat	VERB
alces-443	705	21	.	.	PUNCT
alces-443	706	1	size	size	NOUN
alces-443	706	2	variation	variation	NOUN
alces-443	706	3	(	(	PUNCT
alces-443	706	4	number	number	NOUN
alces-443	706	5	of	of	ADP
alces-443	706	6	repeated	repeat	VERB
alces-443	706	7	units	unit	NOUN
alces-443	706	8	)	)	PUNCT
alces-443	706	9	defines	define	VERB
alces-443	706	10	alleles	allele	NOUN
alces-443	706	11	.	.	PUNCT
alces-443	707	1	microsatellites	microsatellite	NOUN
alces-443	707	2	are	be	AUX
alces-443	707	3	mendelian	mendelian	ADJ
alces-443	707	4	in	in	ADP
alces-443	707	5	their	their	PRON
alces-443	707	6	inheritance	inheritance	NOUN
alces-443	707	7	,	,	PUNCT
alces-443	707	8	i.e.	i.e.	X
alces-443	707	9	,	,	PUNCT
alces-443	707	10	they	they	PRON
alces-443	707	11	are	be	AUX
alces-443	707	12	diploid	diploid	ADJ
alces-443	707	13	,	,	PUNCT
alces-443	707	14	specific	specific	ADJ
alces-443	707	15	to	to	ADP
alces-443	707	16	a	a	DET
alces-443	707	17	site	site	NOUN
alces-443	707	18	on	on	ADP
alces-443	707	19	a	a	DET
alces-443	707	20	chromosome	chromosome	NOUN
alces-443	707	21	,	,	PUNCT
alces-443	707	22	and	and	CCONJ
alces-443	707	23	occur	occur	VERB
alces-443	707	24	in	in	ADP
alces-443	707	25	either	either	CCONJ
alces-443	707	26	a	a	DET
alces-443	707	27	homozygous	homozygous	ADJ
alces-443	707	28	or	or	CCONJ
alces-443	707	29	heterozygous	heterozygous	ADJ
alces-443	707	30	genotype	genotype	NOUN
alces-443	707	31	.	.	PUNCT
alces-443	708	1	minisatellite	minisatellite	NOUN
alces-443	708	2	similar	similar	ADJ
alces-443	708	3	to	to	ADP
alces-443	708	4	microsatellites	microsatellite	NOUN
alces-443	708	5	except	except	SCONJ
alces-443	708	6	that	that	SCONJ
alces-443	708	7	the	the	DET
alces-443	708	8	repeat	repeat	NOUN
alces-443	708	9	units	unit	NOUN
alces-443	708	10	vary	vary	VERB
alces-443	708	11	from	from	ADP
alces-443	708	12	16	16	NUM
alces-443	708	13	-	-	SYM
alces-443	708	14	64	64	NUM
alces-443	708	15	nucleotides	nucleotide	NOUN
alces-443	708	16	in	in	ADP
alces-443	708	17	length	length	NOUN
alces-443	708	18	and	and	CCONJ
alces-443	708	19	occur	occur	VERB
alces-443	708	20	at	at	ADP
alces-443	708	21	many	many	ADJ
alces-443	708	22	sites	site	NOUN
alces-443	708	23	throughout	throughout	ADP
alces-443	708	24	the	the	DET
alces-443	708	25	chromosomes	chromosome	NOUN
alces-443	708	26	,	,	PUNCT
alces-443	708	27	thus	thus	ADV
alces-443	708	28	exhibiting	exhibit	VERB
alces-443	708	29	more	more	ADJ
alces-443	708	30	than	than	ADP
alces-443	708	31	2	2	NUM
alces-443	708	32	alleles	allele	NOUN
alces-443	708	33	per	per	ADP
alces-443	708	34	individual	individual	NOUN
alces-443	708	35	and	and	CCONJ
alces-443	708	36	creating	create	VERB
alces-443	708	37	complex	complex	ADJ
alces-443	708	38	genotypes	genotype	NOUN
alces-443	708	39	of	of	ADP
alces-443	708	40	gel	gel	NOUN
alces-443	708	41	banding	banding	NOUN
alces-443	708	42	patterns	pattern	NOUN
alces-443	708	43	that	that	PRON
alces-443	708	44	resemble	resemble	VERB
alces-443	708	45	bar	bar	NOUN
alces-443	708	46	codes	code	NOUN
alces-443	708	47	.	.	PUNCT
alces-443	709	1	this	this	PRON
alces-443	709	2	is	be	AUX
alces-443	709	3	the	the	DET
alces-443	709	4	technique	technique	NOUN
alces-443	709	5	that	that	PRON
alces-443	709	6	pioneered	pioneer	VERB
alces-443	709	7	genetic	genetic	ADJ
alces-443	709	8	fingerprinting	fingerprinting	NOUN
alces-443	709	9	.	.	PUNCT
alces-443	710	1	mitochondrial	mitochondrial	ADJ
alces-443	710	2	dna	dna	PROPN
alces-443	710	3	(	(	PUNCT
alces-443	710	4	mtdna	mtdna	NOUN
alces-443	710	5	)	)	PUNCT
alces-443	710	6	a	a	DET
alces-443	710	7	circular	circular	ADJ
alces-443	710	8	dna	dna	NOUN
alces-443	710	9	molecule	molecule	NOUN
alces-443	710	10	occurring	occur	VERB
alces-443	710	11	in	in	ADP
alces-443	710	12	the	the	DET
alces-443	710	13	mitochondrion	mitochondrion	NOUN
alces-443	710	14	.	.	PUNCT
alces-443	711	1	unlike	unlike	ADP
alces-443	711	2	chromosomes	chromosome	NOUN
alces-443	711	3	,	,	PUNCT
alces-443	711	4	which	which	PRON
alces-443	711	5	occur	occur	VERB
alces-443	711	6	in	in	ADP
alces-443	711	7	pairs	pair	NOUN
alces-443	711	8	(	(	PUNCT
alces-443	711	9	diploid	diploid	ADJ
alces-443	711	10	)	)	PUNCT
alces-443	711	11	,	,	PUNCT
alces-443	711	12	mtdna	mtdna	PROPN
alces-443	711	13	occurs	occur	VERB
alces-443	711	14	as	as	ADP
alces-443	711	15	a	a	DET
alces-443	711	16	single	single	ADJ
alces-443	711	17	copy	copy	NOUN
alces-443	711	18	(	(	PUNCT
alces-443	711	19	haploid	haploid	NOUN
alces-443	711	20	)	)	PUNCT
alces-443	711	21	because	because	SCONJ
alces-443	711	22	it	it	PRON
alces-443	711	23	is	be	AUX
alces-443	711	24	inherited	inherit	VERB
alces-443	711	25	only	only	ADV
alces-443	711	26	from	from	ADP
alces-443	711	27	the	the	DET
alces-443	711	28	maternal	maternal	ADJ
alces-443	711	29	line	line	NOUN
alces-443	711	30	.	.	PUNCT
alces-443	712	1	therefore	therefore	ADV
alces-443	712	2	,	,	PUNCT
alces-443	712	3	it	it	PRON
alces-443	712	4	is	be	AUX
alces-443	712	5	not	not	PART
alces-443	712	6	subject	subject	ADJ
alces-443	712	7	to	to	ADP
alces-443	712	8	recombination	recombination	NOUN
alces-443	712	9	and	and	CCONJ
alces-443	712	10	changes	change	NOUN
alces-443	712	11	only	only	ADV
alces-443	712	12	via	via	ADP
alces-443	712	13	mutation	mutation	NOUN
alces-443	712	14	.	.	PUNCT
alces-443	713	1	that	that	DET
alces-443	713	2	property	property	NOUN
alces-443	713	3	makes	make	VERB
alces-443	713	4	it	it	PRON
alces-443	713	5	particularly	particularly	ADV
alces-443	713	6	useful	useful	ADJ
alces-443	713	7	for	for	ADP
alces-443	713	8	tracing	trace	VERB
alces-443	713	9	lineages	lineage	NOUN
alces-443	713	10	through	through	ADP
alces-443	713	11	time	time	NOUN
alces-443	713	12	.	.	PUNCT
alces-443	714	1	molecular	molecular	ADJ
alces-443	714	2	clock	clock	VERB
alces-443	714	3	the	the	DET
alces-443	714	4	assumption	assumption	NOUN
alces-443	714	5	that	that	SCONJ
alces-443	714	6	the	the	DET
alces-443	714	7	average	average	ADJ
alces-443	714	8	rate	rate	NOUN
alces-443	714	9	of	of	ADP
alces-443	714	10	mutation	mutation	NOUN
alces-443	714	11	for	for	ADP
alces-443	714	12	a	a	DET
alces-443	714	13	particular	particular	ADJ
alces-443	714	14	dna	dna	NOUN
alces-443	714	15	sequence	sequence	NOUN
alces-443	714	16	is	be	AUX
alces-443	714	17	constant	constant	ADJ
alces-443	714	18	over	over	ADP
alces-443	714	19	evolutionary	evolutionary	ADJ
alces-443	714	20	time	time	NOUN
alces-443	714	21	.	.	PUNCT
alces-443	715	1	if	if	SCONJ
alces-443	715	2	a	a	DET
alces-443	715	3	molecular	molecular	ADJ
alces-443	715	4	clock	clock	NOUN
alces-443	715	5	can	can	AUX
alces-443	715	6	be	be	AUX
alces-443	715	7	assumed	assume	VERB
alces-443	715	8	,	,	PUNCT
alces-443	715	9	the	the	DET
alces-443	715	10	amount	amount	NOUN
alces-443	715	11	of	of	ADP
alces-443	715	12	genetic	genetic	ADJ
alces-443	715	13	divergence	divergence	NOUN
alces-443	715	14	between	between	ADP
alces-443	715	15	populations	population	NOUN
alces-443	715	16	or	or	CCONJ
alces-443	715	17	species	specie	NOUN
alces-443	715	18	can	can	AUX
alces-443	715	19	be	be	AUX
alces-443	715	20	converted	convert	VERB
alces-443	715	21	to	to	ADP
alces-443	715	22	time	time	NOUN
alces-443	715	23	since	since	SCONJ
alces-443	715	24	divergence	divergence	NOUN
alces-443	715	25	.	.	PUNCT
alces-443	716	1	phylogeography	phylogeography	NOUN
alces-443	716	2	the	the	DET
alces-443	716	3	study	study	NOUN
alces-443	716	4	of	of	ADP
alces-443	716	5	genetic	genetic	ADJ
alces-443	716	6	lineages	lineage	NOUN
alces-443	716	7	in	in	ADP
alces-443	716	8	a	a	DET
alces-443	716	9	spatial	spatial	ADJ
alces-443	716	10	and	and	CCONJ
alces-443	716	11	temporal	temporal	ADJ
alces-443	716	12	context	context	NOUN
alces-443	716	13	,	,	PUNCT
alces-443	716	14	revealing	reveal	VERB
alces-443	716	15	historic	historic	ADJ
alces-443	716	16	population	population	NOUN
alces-443	716	17	processes	process	NOUN
alces-443	716	18	and	and	CCONJ
alces-443	716	19	evolutionary	evolutionary	ADJ
alces-443	716	20	histories	history	NOUN
alces-443	716	21	.	.	PUNCT
alces-443	717	1	rflp	rflp	PROPN
alces-443	717	2	restriction	restriction	NOUN
alces-443	717	3	fragment	fragment	NOUN
alces-443	717	4	length	length	NOUN
alces-443	717	5	polymorphism	polymorphism	NOUN
alces-443	717	6	–	–	PUNCT
alces-443	717	7	a	a	DET
alces-443	717	8	section	section	NOUN
alces-443	717	9	of	of	ADP
alces-443	717	10	dna	dna	NOUN
alces-443	717	11	of	of	ADP
alces-443	717	12	known	know	VERB
alces-443	717	13	length	length	NOUN
alces-443	717	14	is	be	AUX
alces-443	717	15	digested	digest	VERB
alces-443	717	16	with	with	ADP
alces-443	717	17	restriction	restriction	NOUN
alces-443	717	18	enzymes	enzyme	NOUN
alces-443	717	19	(	(	PUNCT
alces-443	717	20	endonucleases	endonuclease	NOUN
alces-443	717	21	)	)	PUNCT
alces-443	717	22	,	,	PUNCT
alces-443	717	23	which	which	PRON
alces-443	717	24	cleave	cleave	VERB
alces-443	717	25	dna	dna	PROPN
alces-443	717	26	at	at	ADP
alces-443	717	27	sites	site	NOUN
alces-443	717	28	with	with	ADP
alces-443	717	29	specific	specific	ADJ
alces-443	717	30	target	target	NOUN
alces-443	717	31	sequences	sequence	NOUN
alces-443	717	32	of	of	ADP
alces-443	717	33	4	4	NUM
alces-443	717	34	-	-	SYM
alces-443	717	35	6	6	NUM
alces-443	717	36	nucleotides	nucleotide	NOUN
alces-443	717	37	.	.	PUNCT
alces-443	718	1	the	the	DET
alces-443	718	2	digested	digest	VERB
alces-443	718	3	fragments	fragment	NOUN
alces-443	718	4	are	be	AUX
alces-443	718	5	separated	separate	VERB
alces-443	718	6	by	by	ADP
alces-443	718	7	length	length	NOUN
alces-443	718	8	(	(	PUNCT
alces-443	718	9	number	number	NOUN
alces-443	718	10	of	of	ADP
alces-443	718	11	nucleotides	nucleotide	NOUN
alces-443	718	12	)	)	PUNCT
alces-443	718	13	using	use	VERB
alces-443	718	14	gel	gel	NOUN
alces-443	718	15	electrophoresis	electrophoresis	NOUN
alces-443	718	16	and	and	CCONJ
alces-443	718	17	haplotypes	haplotype	NOUN
alces-443	718	18	are	be	AUX
alces-443	718	19	characterized	characterize	VERB
alces-443	718	20	by	by	ADP
alces-443	718	21	numbers	number	NOUN
alces-443	718	22	and	and	CCONJ
alces-443	718	23	sizes	size	NOUN
alces-443	718	24	of	of	ADP
alces-443	718	25	fragments	fragment	NOUN
alces-443	718	26	.	.	PUNCT
alces-443	719	1	appendix	appendix	NOUN
alces-443	719	2	1	1	NUM
alces-443	719	3	.	.	PUNCT
alces-443	720	1	glossary	glossary	NOUN
alces-443	720	2	of	of	ADP
alces-443	720	3	specialized	specialized	ADJ
alces-443	720	4	terms	term	NOUN
alces-443	720	5	used	use	VERB
alces-443	720	6	in	in	ADP
alces-443	720	7	this	this	DET
alces-443	720	8	review	review	NOUN
alces-443	720	9	.	.	PUNCT