id	sid	tid	token	lemma	pos
alces-483	1	1	alces39_215.pdf	alces39_215.pdf	PROPN
alces-483	1	2	alces	alce	NOUN
alces-483	1	3	vol	vol	NOUN
alces-483	1	4	.	.	PROPN
alces-483	1	5	39	39	NUM
alces-483	1	6	,	,	PUNCT
alces-483	1	7	2003	2003	NUM
alces-483	1	8	peterson	peterson	PROPN
alces-483	1	9	et	et	PROPN
alces-483	1	10	al	al	PROPN
alces-483	1	11	.	.	PROPN
alces-483	1	12	–	–	PUNCT
alces-483	1	13	space	space	NOUN
alces-483	1	14	and	and	CCONJ
alces-483	1	15	time	time	NOUN
alces-483	1	16	in	in	ADP
alces-483	1	17	predation	predation	NOUN
alces-483	1	18	dynamics	dynamic	NOUN
alces-483	1	19	215	215	NUM
alces-483	1	20	temporal	temporal	ADJ
alces-483	1	21	and	and	CCONJ
alces-483	1	22	spatial	spatial	ADJ
alces-483	1	23	aspects	aspect	NOUN
alces-483	1	24	of	of	ADP
alces-483	1	25	predator	predator	NOUN
alces-483	1	26	-	-	PUNCT
alces-483	1	27	prey	prey	NOUN
alces-483	1	28	dynamics	dynamic	NOUN
alces-483	1	29	rolf	rolf	PROPN
alces-483	1	30	o.	o.	PROPN
alces-483	1	31	peterson1	peterson1	PROPN
alces-483	1	32	,	,	PUNCT
alces-483	1	33	john	john	PROPN
alces-483	1	34	a.	a.	PROPN
alces-483	1	35	vucetich1	vucetich1	PROPN
alces-483	1	36	,	,	PUNCT
alces-483	1	37	richard	richard	PROPN
alces-483	1	38	e.	e.	PROPN
alces-483	1	39	page2	page2	PROPN
alces-483	1	40	,	,	PUNCT
alces-483	1	41	and	and	CCONJ
alces-483	1	42	anne	anne	PROPN
alces-483	1	43	chouinard3	chouinard3	PROPN
alces-483	2	1	1school	1school	NUM
alces-483	2	2	of	of	ADP
alces-483	2	3	forest	forest	NOUN
alces-483	2	4	resources	resource	NOUN
alces-483	2	5	and	and	CCONJ
alces-483	2	6	environmental	environmental	ADJ
alces-483	2	7	science	science	NOUN
alces-483	2	8	,	,	PUNCT
alces-483	2	9	michigan	michigan	PROPN
alces-483	2	10	technological	technological	PROPN
alces-483	2	11	university	university	PROPN
alces-483	2	12	,	,	PUNCT
alces-483	2	13	houghton	houghton	PROPN
alces-483	2	14	,	,	PUNCT
alces-483	2	15	mi	mi	PROPN
alces-483	2	16	49931	49931	NUM
alces-483	2	17	,	,	PUNCT
alces-483	2	18	usa	usa	PROPN
alces-483	2	19	;	;	PUNCT
alces-483	2	20	2page	2page	NUM
alces-483	2	21	and	and	CCONJ
alces-483	2	22	associates	associate	NOUN
alces-483	2	23	,	,	PUNCT
alces-483	2	24	3915	3915	NUM
alces-483	2	25	scolton	scolton	PROPN
alces-483	2	26	road	road	NOUN
alces-483	2	27	,	,	PUNCT
alces-483	2	28	victoria	victoria	PROPN
alces-483	2	29	,	,	PUNCT
alces-483	2	30	bc	bc	PROPN
alces-483	2	31	,	,	PUNCT
alces-483	2	32	canada	canada	PROPN
alces-483	2	33	v8n	v8n	PROPN
alces-483	2	34	4e1	4e1	NUM
alces-483	2	35	;	;	PUNCT
alces-483	2	36	3centre	3centre	NUM
alces-483	2	37	d’études	d’étude	NOUN
alces-483	2	38	nordiques	nordique	NOUN
alces-483	2	39	,	,	PUNCT
alces-483	2	40	université	université	PROPN
alces-483	2	41	laval	laval	PROPN
alces-483	2	42	,	,	PUNCT
alces-483	2	43	sainte	sainte	PROPN
alces-483	2	44	-	-	PUNCT
alces-483	2	45	foy	foy	PROPN
alces-483	2	46	,	,	PUNCT
alces-483	2	47	pq	pq	PROPN
alces-483	2	48	,	,	PUNCT
alces-483	2	49	canada	canada	PROPN
alces-483	2	50	g1k	g1k	PROPN
alces-483	3	1	7p4	7p4	NUM
alces-483	3	2	abstract	abstract	NOUN
alces-483	3	3	:	:	PUNCT
alces-483	3	4	ungulates	ungulate	NOUN
alces-483	3	5	are	be	AUX
alces-483	3	6	both	both	PRON
alces-483	3	7	major	major	ADJ
alces-483	3	8	consumers	consumer	NOUN
alces-483	3	9	of	of	ADP
alces-483	3	10	vegetation	vegetation	NOUN
alces-483	3	11	and	and	CCONJ
alces-483	3	12	are	be	AUX
alces-483	3	13	themselves	themselves	PRON
alces-483	3	14	consumed	consume	VERB
alces-483	3	15	by	by	ADP
alces-483	3	16	carnivores	carnivore	NOUN
alces-483	3	17	,	,	PUNCT
alces-483	3	18	so	so	ADV
alces-483	3	19	food	food	NOUN
alces-483	3	20	web	web	NOUN
alces-483	3	21	dynamics	dynamic	NOUN
alces-483	3	22	,	,	PUNCT
alces-483	3	23	both	both	DET
alces-483	3	24	top	top	NOUN
alces-483	3	25	-	-	PUNCT
alces-483	3	26	down	down	NOUN
alces-483	3	27	(	(	PUNCT
alces-483	3	28	predation	predation	NOUN
alces-483	3	29	)	)	PUNCT
alces-483	3	30	and	and	CCONJ
alces-483	3	31	bottom	bottom	NOUN
alces-483	3	32	-	-	PUNCT
alces-483	3	33	up	up	NOUN
alces-483	3	34	(	(	PUNCT
alces-483	3	35	food	food	NOUN
alces-483	3	36	and	and	CCONJ
alces-483	3	37	weather	weather	NOUN
alces-483	3	38	effects	effect	NOUN
alces-483	3	39	)	)	PUNCT
alces-483	3	40	,	,	PUNCT
alces-483	3	41	are	be	AUX
alces-483	3	42	prominent	prominent	ADJ
alces-483	3	43	in	in	ADP
alces-483	3	44	theoretical	theoretical	ADJ
alces-483	3	45	and	and	CCONJ
alces-483	3	46	applied	apply	VERB
alces-483	3	47	research	research	NOUN
alces-483	3	48	involving	involve	VERB
alces-483	3	49	ungulates	ungulate	NOUN
alces-483	3	50	.	.	PUNCT
alces-483	4	1	the	the	DET
alces-483	4	2	long	long	ADJ
alces-483	4	3	generation	generation	NOUN
alces-483	4	4	time	time	NOUN
alces-483	4	5	of	of	ADP
alces-483	4	6	ungulates	ungulate	NOUN
alces-483	4	7	induces	induce	VERB
alces-483	4	8	long	long	ADJ
alces-483	4	9	lags	lag	NOUN
alces-483	4	10	in	in	ADP
alces-483	4	11	population	population	NOUN
alces-483	4	12	responses	response	NOUN
alces-483	4	13	.	.	PUNCT
alces-483	5	1	over	over	ADP
alces-483	5	2	broad	broad	ADJ
alces-483	5	3	geographic	geographic	ADJ
alces-483	5	4	regions	region	NOUN
alces-483	5	5	,	,	PUNCT
alces-483	5	6	ungulates	ungulate	NOUN
alces-483	5	7	commonly	commonly	ADV
alces-483	5	8	achieve	achieve	VERB
alces-483	5	9	high	high	ADJ
alces-483	5	10	density	density	NOUN
alces-483	5	11	only	only	ADV
alces-483	5	12	when	when	SCONJ
alces-483	5	13	predation	predation	NOUN
alces-483	5	14	is	be	AUX
alces-483	5	15	relatively	relatively	ADV
alces-483	5	16	low	low	ADJ
alces-483	5	17	(	(	PUNCT
alces-483	5	18	<	<	X
alces-483	5	19	2	2	NUM
alces-483	5	20	species	specie	NOUN
alces-483	5	21	of	of	ADP
alces-483	5	22	predator	predator	NOUN
alces-483	5	23	)	)	PUNCT
alces-483	5	24	,	,	PUNCT
alces-483	5	25	suggesting	suggest	VERB
alces-483	5	26	that	that	SCONJ
alces-483	5	27	predation	predation	NOUN
alces-483	5	28	provides	provide	VERB
alces-483	5	29	a	a	DET
alces-483	5	30	pervasive	pervasive	ADJ
alces-483	5	31	limitation	limitation	NOUN
alces-483	5	32	of	of	ADP
alces-483	5	33	large	large	ADJ
alces-483	5	34	herbivores	herbivore	NOUN
alces-483	5	35	.	.	PUNCT
alces-483	6	1	ungulate	ungulate	ADJ
alces-483	6	2	stability	stability	NOUN
alces-483	6	3	is	be	AUX
alces-483	6	4	fundamentally	fundamentally	ADV
alces-483	6	5	a	a	DET
alces-483	6	6	trophic	trophic	ADJ
alces-483	6	7	-	-	PUNCT
alces-483	6	8	dynamics	dynamic	NOUN
alces-483	6	9	issue	issue	NOUN
alces-483	6	10	,	,	PUNCT
alces-483	6	11	usually	usually	ADV
alces-483	6	12	a	a	DET
alces-483	6	13	mix	mix	NOUN
alces-483	6	14	of	of	ADP
alces-483	6	15	top	top	ADJ
alces-483	6	16	-	-	PUNCT
alces-483	6	17	down	down	NOUN
alces-483	6	18	and	and	CCONJ
alces-483	6	19	bottom	bottom	ADJ
alces-483	6	20	-	-	PUNCT
alces-483	6	21	up	up	ADP
alces-483	6	22	influences	influence	NOUN
alces-483	6	23	.	.	PUNCT
alces-483	7	1	the	the	DET
alces-483	7	2	isle	isle	PROPN
alces-483	7	3	royale	royale	PROPN
alces-483	7	4	case	case	NOUN
alces-483	7	5	history	history	NOUN
alces-483	7	6	,	,	PUNCT
alces-483	7	7	spanning	span	VERB
alces-483	7	8	4	4	NUM
alces-483	7	9	decades	decade	NOUN
alces-483	7	10	,	,	PUNCT
alces-483	7	11	reveals	reveal	VERB
alces-483	7	12	a	a	DET
alces-483	7	13	wolf	wolf	NOUN
alces-483	7	14	-	-	PUNCT
alces-483	7	15	moose	moose	NOUN
alces-483	7	16	system	system	NOUN
alces-483	7	17	fluctuating	fluctuate	VERB
alces-483	7	18	at	at	ADP
alces-483	7	19	2	2	NUM
alces-483	7	20	-	-	PUNCT
alces-483	7	21	decade	decade	NOUN
alces-483	7	22	intervals	interval	NOUN
alces-483	7	23	with	with	ADP
alces-483	7	24	significant	significant	ADJ
alces-483	7	25	predation	predation	NOUN
alces-483	7	26	,	,	PUNCT
alces-483	7	27	food	food	NOUN
alces-483	7	28	,	,	PUNCT
alces-483	7	29	and	and	CCONJ
alces-483	7	30	weather	weather	NOUN
alces-483	7	31	effects	effect	NOUN
alces-483	7	32	on	on	ADP
alces-483	7	33	ungulates	ungulate	NOUN
alces-483	7	34	.	.	PUNCT
alces-483	8	1	after	after	ADP
alces-483	8	2	a	a	DET
alces-483	8	3	century	century	NOUN
alces-483	8	4	,	,	PUNCT
alces-483	8	5	an	an	DET
alces-483	8	6	equilibrium	equilibrium	NOUN
alces-483	8	7	between	between	ADP
alces-483	8	8	moose	moose	NOUN
alces-483	8	9	and	and	CCONJ
alces-483	8	10	forest	forest	NOUN
alces-483	8	11	vegetation	vegetation	NOUN
alces-483	8	12	has	have	AUX
alces-483	8	13	not	not	PART
alces-483	8	14	yet	yet	ADV
alces-483	8	15	been	be	AUX
alces-483	8	16	reached	reach	VERB
alces-483	8	17	,	,	PUNCT
alces-483	8	18	and	and	CCONJ
alces-483	8	19	a	a	DET
alces-483	8	20	historical	historical	ADJ
alces-483	8	21	context	context	NOUN
alces-483	8	22	seems	seem	VERB
alces-483	8	23	necessary	necessary	ADJ
alces-483	8	24	to	to	PART
alces-483	8	25	understand	understand	VERB
alces-483	8	26	trophic	trophic	ADJ
alces-483	8	27	relationships	relationship	NOUN
alces-483	8	28	.	.	PUNCT
alces-483	9	1	components	component	NOUN
alces-483	9	2	of	of	ADP
alces-483	9	3	predation	predation	NOUN
alces-483	9	4	compared	compare	VERB
alces-483	9	5	at	at	ADP
alces-483	9	6	large	large	ADJ
alces-483	9	7	spatial	spatial	ADJ
alces-483	9	8	scales	scale	NOUN
alces-483	9	9	reveal	reveal	VERB
alces-483	9	10	different	different	ADJ
alces-483	9	11	predator	predator	NOUN
alces-483	9	12	-	-	PUNCT
alces-483	9	13	prey	prey	NOUN
alces-483	9	14	patterns	pattern	NOUN
alces-483	9	15	than	than	ADP
alces-483	9	16	the	the	DET
alces-483	9	17	single	single	ADJ
alces-483	9	18	system	system	NOUN
alces-483	9	19	at	at	ADP
alces-483	9	20	isle	isle	PROPN
alces-483	9	21	royale	royale	PROPN
alces-483	9	22	,	,	PUNCT
alces-483	9	23	and	and	CCONJ
alces-483	9	24	analyses	analysis	NOUN
alces-483	9	25	involving	involve	VERB
alces-483	9	26	substitution	substitution	NOUN
alces-483	9	27	of	of	ADP
alces-483	9	28	space	space	NOUN
alces-483	9	29	for	for	ADP
alces-483	9	30	time	time	NOUN
alces-483	9	31	also	also	ADV
alces-483	9	32	run	run	VERB
alces-483	9	33	counter	counter	ADV
alces-483	9	34	to	to	ADP
alces-483	9	35	studies	study	NOUN
alces-483	9	36	of	of	ADP
alces-483	9	37	single	single	ADJ
alces-483	9	38	systems	system	NOUN
alces-483	9	39	.	.	PUNCT
alces-483	10	1	choice	choice	NOUN
alces-483	10	2	of	of	ADP
alces-483	10	3	spatial	spatial	ADJ
alces-483	10	4	and	and	CCONJ
alces-483	10	5	temporal	temporal	ADJ
alces-483	10	6	scales	scale	NOUN
alces-483	10	7	for	for	ADP
alces-483	10	8	field	field	NOUN
alces-483	10	9	studies	study	NOUN
alces-483	10	10	and	and	CCONJ
alces-483	10	11	meta	meta	NOUN
alces-483	10	12	-	-	PUNCT
alces-483	10	13	analyses	analysis	NOUN
alces-483	10	14	appear	appear	VERB
alces-483	10	15	to	to	PART
alces-483	10	16	have	have	VERB
alces-483	10	17	a	a	DET
alces-483	10	18	strong	strong	ADJ
alces-483	10	19	bearing	bearing	NOUN
alces-483	10	20	on	on	ADP
alces-483	10	21	the	the	DET
alces-483	10	22	results	result	NOUN
alces-483	10	23	and	and	CCONJ
alces-483	10	24	their	their	PRON
alces-483	10	25	interpretation	interpretation	NOUN
alces-483	10	26	.	.	PUNCT
alces-483	11	1	thus	thus	ADV
alces-483	11	2	temporal	temporal	ADJ
alces-483	11	3	and	and	CCONJ
alces-483	11	4	spatial	spatial	ADJ
alces-483	11	5	scales	scale	NOUN
alces-483	11	6	enter	enter	VERB
alces-483	11	7	influentially	influentially	ADV
alces-483	11	8	in	in	ADP
alces-483	11	9	the	the	DET
alces-483	11	10	actual	actual	ADJ
alces-483	11	11	dynamics	dynamic	NOUN
alces-483	11	12	of	of	ADP
alces-483	11	13	carnivore	carnivore	ADJ
alces-483	11	14	-	-	PUNCT
alces-483	11	15	ungulate	ungulate	ADJ
alces-483	11	16	interaction	interaction	NOUN
alces-483	11	17	as	as	ADV
alces-483	11	18	well	well	ADV
alces-483	11	19	problematically	problematically	ADV
alces-483	11	20	in	in	ADP
alces-483	11	21	our	our	PRON
alces-483	11	22	analyses	analysis	NOUN
alces-483	11	23	of	of	ADP
alces-483	11	24	them	they	PRON
alces-483	11	25	.	.	PUNCT
alces-483	12	1	alces	alce	NOUN
alces-483	12	2	vol	vol	NOUN
alces-483	12	3	.	.	PROPN
alces-483	12	4	39	39	NUM
alces-483	12	5	:	:	PUNCT
alces-483	12	6	215	215	NUM
alces-483	12	7	-	-	SYM
alces-483	12	8	232	232	NUM
alces-483	12	9	(	(	PUNCT
alces-483	12	10	2003	2003	NUM
alces-483	12	11	)	)	PUNCT
alces-483	12	12	key	key	ADJ
alces-483	12	13	words	word	NOUN
alces-483	12	14	:	:	PUNCT
alces-483	12	15	bear	bear	NOUN
alces-483	12	16	,	,	PUNCT
alces-483	12	17	bottom	bottom	NOUN
alces-483	12	18	-	-	PUNCT
alces-483	12	19	up	up	NOUN
alces-483	12	20	,	,	PUNCT
alces-483	12	21	herbivory	herbivory	NOUN
alces-483	12	22	,	,	PUNCT
alces-483	12	23	moose	moose	NOUN
alces-483	12	24	,	,	PUNCT
alces-483	12	25	population	population	NOUN
alces-483	12	26	,	,	PUNCT
alces-483	12	27	predation	predation	NOUN
alces-483	12	28	,	,	PUNCT
alces-483	12	29	scale	scale	NOUN
alces-483	12	30	,	,	PUNCT
alces-483	12	31	space	space	NOUN
alces-483	12	32	,	,	PUNCT
alces-483	12	33	time	time	NOUN
alces-483	12	34	,	,	PUNCT
alces-483	12	35	topdown	topdown	ADJ
alces-483	12	36	,	,	PUNCT
alces-483	12	37	trophic	trophic	ADJ
alces-483	12	38	,	,	PUNCT
alces-483	12	39	wolf	wolf	NOUN
alces-483	12	40	interactions	interaction	NOUN
alces-483	12	41	among	among	ADP
alces-483	12	42	trophic	trophic	ADJ
alces-483	12	43	levels	level	NOUN
alces-483	12	44	have	have	AUX
alces-483	12	45	been	be	AUX
alces-483	12	46	pervasive	pervasive	ADJ
alces-483	12	47	themes	theme	NOUN
alces-483	12	48	in	in	ADP
alces-483	12	49	animal	animal	NOUN
alces-483	12	50	ecology	ecology	NOUN
alces-483	12	51	since	since	SCONJ
alces-483	12	52	its	its	PRON
alces-483	12	53	inception	inception	NOUN
alces-483	12	54	(	(	PUNCT
alces-483	12	55	elton	elton	PROPN
alces-483	12	56	1927	1927	NUM
alces-483	12	57	)	)	PUNCT
alces-483	12	58	.	.	PUNCT
alces-483	13	1	ungulates	ungulate	NOUN
alces-483	13	2	are	be	AUX
alces-483	13	3	both	both	PRON
alces-483	13	4	major	major	ADJ
alces-483	13	5	consumers	consumer	NOUN
alces-483	13	6	of	of	ADP
alces-483	13	7	vegetation	vegetation	NOUN
alces-483	13	8	and	and	CCONJ
alces-483	13	9	are	be	AUX
alces-483	13	10	themselves	themselves	PRON
alces-483	13	11	consumed	consume	VERB
alces-483	13	12	by	by	ADP
alces-483	13	13	carnivores	carnivore	NOUN
alces-483	13	14	,	,	PUNCT
alces-483	13	15	so	so	SCONJ
alces-483	13	16	studies	study	NOUN
alces-483	13	17	of	of	ADP
alces-483	13	18	their	their	PRON
alces-483	13	19	population	population	NOUN
alces-483	13	20	dynamics	dynamic	NOUN
alces-483	13	21	should	should	AUX
alces-483	13	22	reveal	reveal	VERB
alces-483	13	23	much	much	ADJ
alces-483	13	24	about	about	ADP
alces-483	13	25	dominant	dominant	ADJ
alces-483	13	26	trophic	trophic	ADJ
alces-483	13	27	linkages	linkage	NOUN
alces-483	13	28	in	in	ADP
alces-483	13	29	terrestrial	terrestrial	ADJ
alces-483	13	30	systems	system	NOUN
alces-483	13	31	(	(	PUNCT
alces-483	13	32	schmitz	schmitz	PROPN
alces-483	13	33	et	et	PROPN
alces-483	13	34	al	al	PROPN
alces-483	13	35	.	.	PROPN
alces-483	13	36	2000	2000	NUM
alces-483	13	37	)	)	PUNCT
alces-483	13	38	.	.	PUNCT
alces-483	14	1	ungulates	ungulate	NOUN
alces-483	14	2	are	be	AUX
alces-483	14	3	also	also	ADV
alces-483	14	4	large	large	ADJ
alces-483	14	5	-	-	PUNCT
alces-483	14	6	bodied	bodied	ADJ
alces-483	14	7	organisms	organism	NOUN
alces-483	14	8	;	;	PUNCT
alces-483	14	9	they	they	PRON
alces-483	14	10	may	may	AUX
alces-483	14	11	forage	forage	VERB
alces-483	14	12	at	at	ADP
alces-483	14	13	local	local	ADJ
alces-483	14	14	scales	scale	NOUN
alces-483	14	15	(	(	PUNCT
alces-483	14	16	risenhoover	risenhoover	NOUN
alces-483	14	17	1987	1987	NUM
alces-483	14	18	,	,	PUNCT
alces-483	14	19	spalinger	spalinger	NOUN
alces-483	14	20	and	and	CCONJ
alces-483	14	21	hobbs	hobbs	PROPN
alces-483	14	22	1992	1992	NUM
alces-483	14	23	)	)	PUNCT
alces-483	14	24	,	,	PUNCT
alces-483	14	25	but	but	CCONJ
alces-483	14	26	they	they	PRON
alces-483	14	27	make	make	VERB
alces-483	14	28	decisions	decision	NOUN
alces-483	14	29	about	about	ADP
alces-483	14	30	movements	movement	NOUN
alces-483	14	31	that	that	PRON
alces-483	14	32	may	may	AUX
alces-483	14	33	cover	cover	VERB
alces-483	14	34	tens	ten	NOUN
alces-483	14	35	or	or	CCONJ
alces-483	14	36	even	even	ADV
alces-483	14	37	hundreds	hundred	NOUN
alces-483	14	38	of	of	ADP
alces-483	14	39	square	square	ADJ
alces-483	14	40	kilometers	kilometer	NOUN
alces-483	14	41	.	.	PUNCT
alces-483	15	1	their	their	PRON
alces-483	15	2	population	population	NOUN
alces-483	15	3	dynamics	dynamic	NOUN
alces-483	15	4	and	and	CCONJ
alces-483	15	5	ecological	ecological	ADJ
alces-483	15	6	influence	influence	NOUN
alces-483	15	7	may	may	AUX
alces-483	15	8	likewise	likewise	ADV
alces-483	15	9	reflect	reflect	VERB
alces-483	15	10	ecological	ecological	ADJ
alces-483	15	11	phenomena	phenomenon	NOUN
alces-483	15	12	that	that	PRON
alces-483	15	13	occur	occur	VERB
alces-483	15	14	at	at	ADP
alces-483	15	15	different	different	ADJ
alces-483	15	16	scales	scale	NOUN
alces-483	15	17	,	,	PUNCT
alces-483	15	18	e.g.	e.g.	ADV
alces-483	15	19	,	,	PUNCT
alces-483	15	20	vegetation	vegetation	NOUN
alces-483	15	21	patch	patch	NOUN
alces-483	15	22	dynamics	dynamic	NOUN
alces-483	15	23	and	and	CCONJ
alces-483	15	24	local	local	ADJ
alces-483	15	25	nutrient	nutrient	NOUN
alces-483	15	26	fluxes	flux	NOUN
alces-483	15	27	may	may	AUX
alces-483	15	28	be	be	AUX
alces-483	15	29	tracked	track	VERB
alces-483	15	30	by	by	ADP
alces-483	15	31	foraging	forage	VERB
alces-483	15	32	ungulates	ungulate	NOUN
alces-483	15	33	(	(	PUNCT
alces-483	15	34	white	white	NOUN
alces-483	15	35	1983	1983	NUM
alces-483	15	36	,	,	PUNCT
alces-483	15	37	pastor	pastor	PROPN
alces-483	15	38	et	et	PROPN
alces-483	15	39	al	al	PROPN
alces-483	15	40	.	.	PROPN
alces-483	15	41	1997	1997	NUM
alces-483	15	42	,	,	PUNCT
alces-483	15	43	etzenhouser	etzenhouser	NOUN
alces-483	15	44	et	et	PROPN
alces-483	15	45	al	al	PROPN
alces-483	15	46	.	.	PROPN
alces-483	15	47	1998	1998	NUM
alces-483	15	48	,	,	PUNCT
alces-483	15	49	shipley	shipley	PROPN
alces-483	15	50	et	et	PROPN
alces-483	15	51	al	al	PROPN
alces-483	15	52	.	.	PROPN
alces-483	15	53	1999	1999	NUM
alces-483	15	54	)	)	PUNCT
alces-483	15	55	,	,	PUNCT
alces-483	15	56	but	but	CCONJ
alces-483	15	57	their	their	PRON
alces-483	15	58	population	population	NOUN
alces-483	15	59	dynamics	dynamic	NOUN
alces-483	15	60	may	may	AUX
alces-483	15	61	reflect	reflect	VERB
alces-483	15	62	broad	broad	ADJ
alces-483	15	63	-	-	PUNCT
alces-483	15	64	scale	scale	NOUN
alces-483	15	65	forest	forest	NOUN
alces-483	15	66	successional	successional	ADJ
alces-483	15	67	patterns	pattern	NOUN
alces-483	15	68	or	or	CCONJ
alces-483	15	69	pervasive	pervasive	ADJ
alces-483	15	70	loss	loss	NOUN
alces-483	15	71	to	to	ADP
alces-483	15	72	predators	predator	NOUN
alces-483	15	73	with	with	ADP
alces-483	15	74	very	very	ADV
alces-483	15	75	large	large	ADJ
alces-483	15	76	home	home	NOUN
alces-483	15	77	ranges	range	NOUN
alces-483	15	78	(	(	PUNCT
alces-483	15	79	schwartz	schwartz	NOUN
alces-483	15	80	and	and	CCONJ
alces-483	15	81	franzmann	franzmann	NOUN
alces-483	15	82	1991	1991	NUM
alces-483	15	83	,	,	PUNCT
alces-483	15	84	gasaway	gasaway	NOUN
alces-483	15	85	et	et	PROPN
alces-483	15	86	al	al	PROPN
alces-483	15	87	.	.	PROPN
alces-483	15	88	1992	1992	NUM
alces-483	15	89	)	)	PUNCT
alces-483	15	90	.	.	PUNCT
alces-483	16	1	their	their	PRON
alces-483	16	2	large	large	ADJ
alces-483	16	3	body	body	NOUN
alces-483	16	4	size	size	NOUN
alces-483	16	5	also	also	ADV
alces-483	16	6	introduces	introduce	VERB
alces-483	16	7	potential	potential	ADJ
alces-483	16	8	complexity	complexity	NOUN
alces-483	16	9	to	to	ADP
alces-483	16	10	predatorprey	predatorprey	ADJ
alces-483	16	11	relationships	relationship	NOUN
alces-483	16	12	.	.	PUNCT
alces-483	17	1	ungulates	ungulate	NOUN
alces-483	17	2	may	may	AUX
alces-483	17	3	be	be	AUX
alces-483	17	4	larger	large	ADJ
alces-483	17	5	than	than	ADP
alces-483	17	6	sympatric	sympatric	ADJ
alces-483	17	7	predators	predator	NOUN
alces-483	17	8	,	,	PUNCT
alces-483	17	9	thus	thus	ADV
alces-483	17	10	difficult	difficult	ADJ
alces-483	17	11	to	to	PART
alces-483	17	12	kill	kill	VERB
alces-483	17	13	as	as	ADP
alces-483	17	14	adults	adult	NOUN
alces-483	17	15	but	but	CCONJ
alces-483	17	16	more	more	ADV
alces-483	17	17	easily	easily	ADV
alces-483	17	18	killed	kill	VERB
alces-483	17	19	as	as	ADP
alces-483	17	20	slowgrowing	slowgrowe	VERB
alces-483	17	21	juveniles	juvenile	NOUN
alces-483	17	22	(	(	PUNCT
alces-483	17	23	peterson	peterson	NOUN
alces-483	17	24	1977	1977	NUM
alces-483	17	25	)	)	PUNCT
alces-483	17	26	.	.	PUNCT
alces-483	18	1	ungulates	ungulate	NOUN
alces-483	18	2	are	be	AUX
alces-483	18	3	relatively	relatively	ADV
alces-483	18	4	long	long	ADV
alces-483	18	5	-	-	PUNCT
alces-483	18	6	lived	live	VERB
alces-483	18	7	,	,	PUNCT
alces-483	18	8	with	with	ADP
alces-483	18	9	individuspace	individuspace	NOUN
alces-483	18	10	and	and	CCONJ
alces-483	18	11	time	time	NOUN
alces-483	18	12	in	in	ADP
alces-483	18	13	predation	predation	NOUN
alces-483	18	14	dynamics	dynamic	NOUN
alces-483	18	15	–	–	PUNCT
alces-483	18	16	peterson	peterson	PROPN
alces-483	18	17	et	et	PROPN
alces-483	18	18	al	al	PROPN
alces-483	18	19	.	.	PROPN
alces-483	19	1	alces	alce	NOUN
alces-483	19	2	vol	vol	NOUN
alces-483	19	3	.	.	PROPN
alces-483	19	4	39	39	NUM
alces-483	19	5	,	,	PUNCT
alces-483	19	6	2003	2003	NUM
alces-483	19	7	216	216	NUM
alces-483	19	8	als	als	NOUN
alces-483	19	9	persisting	persist	VERB
alces-483	19	10	for	for	ADP
alces-483	19	11	many	many	ADJ
alces-483	19	12	annual	annual	ADJ
alces-483	19	13	cycles	cycle	NOUN
alces-483	19	14	of	of	ADP
alces-483	19	15	seasonal	seasonal	ADJ
alces-483	19	16	changes	change	NOUN
alces-483	19	17	(	(	PUNCT
alces-483	19	18	caughley	caughley	NOUN
alces-483	19	19	and	and	CCONJ
alces-483	19	20	krebs	krebs	NOUN
alces-483	19	21	1983	1983	NUM
alces-483	19	22	)	)	PUNCT
alces-483	19	23	,	,	PUNCT
alces-483	19	24	so	so	ADV
alces-483	19	25	ecological	ecological	ADJ
alces-483	19	26	lag	lag	NOUN
alces-483	19	27	effects	effect	NOUN
alces-483	19	28	relating	relate	VERB
alces-483	19	29	to	to	ADP
alces-483	19	30	maternal	maternal	ADJ
alces-483	19	31	effects	effect	NOUN
alces-483	19	32	and	and	CCONJ
alces-483	19	33	individual	individual	ADJ
alces-483	19	34	vigor	vigor	NOUN
alces-483	19	35	can	can	AUX
alces-483	19	36	be	be	AUX
alces-483	19	37	anticipated	anticipate	VERB
alces-483	19	38	(	(	PUNCT
alces-483	19	39	mech	mech	NOUN
alces-483	19	40	et	et	PROPN
alces-483	19	41	al	al	PROPN
alces-483	19	42	.	.	PROPN
alces-483	19	43	1991	1991	NUM
alces-483	19	44	)	)	PUNCT
alces-483	19	45	.	.	PUNCT
alces-483	20	1	here	here	ADV
alces-483	20	2	we	we	PRON
alces-483	20	3	will	will	AUX
alces-483	20	4	attempt	attempt	VERB
alces-483	20	5	to	to	PART
alces-483	20	6	assess	assess	VERB
alces-483	20	7	how	how	SCONJ
alces-483	20	8	predator	predator	NOUN
alces-483	20	9	-	-	PUNCT
alces-483	20	10	prey	prey	NOUN
alces-483	20	11	dynamics	dynamic	NOUN
alces-483	20	12	,	,	PUNCT
alces-483	20	13	and	and	CCONJ
alces-483	20	14	our	our	PRON
alces-483	20	15	understanding	understanding	NOUN
alces-483	20	16	of	of	ADP
alces-483	20	17	such	such	ADJ
alces-483	20	18	dynamics	dynamic	NOUN
alces-483	20	19	,	,	PUNCT
alces-483	20	20	depends	depend	VERB
alces-483	20	21	critically	critically	ADV
alces-483	20	22	on	on	ADP
alces-483	20	23	several	several	ADJ
alces-483	20	24	spatio	spatio	ADJ
alces-483	20	25	-	-	PUNCT
alces-483	20	26	temporal	temporal	ADJ
alces-483	20	27	scale	scale	NOUN
alces-483	20	28	issues	issue	NOUN
alces-483	20	29	,	,	PUNCT
alces-483	20	30	as	as	SCONJ
alces-483	20	31	illustrated	illustrate	VERB
alces-483	20	32	by	by	ADP
alces-483	20	33	the	the	DET
alces-483	20	34	wolf	wolf	NOUN
alces-483	20	35	-	-	PUNCT
alces-483	20	36	moose	moose	NOUN
alces-483	20	37	-	-	PUNCT
alces-483	20	38	fir	fir	NOUN
alces-483	20	39	system	system	NOUN
alces-483	20	40	at	at	ADP
alces-483	20	41	isle	isle	PROPN
alces-483	20	42	royale	royale	PROPN
alces-483	20	43	.	.	PUNCT
alces-483	21	1	top	top	ADJ
alces-483	21	2	-	-	PUNCT
alces-483	21	3	down	down	NOUN
alces-483	21	4	and	and	CCONJ
alces-483	21	5	bottom	bottom	VERB
alces-483	21	6	up	up	ADP
alces-483	21	7	the	the	DET
alces-483	21	8	seminal	seminal	ADJ
alces-483	21	9	essay	essay	NOUN
alces-483	21	10	by	by	ADP
alces-483	21	11	hairston	hairston	PROPN
alces-483	21	12	et	et	PROPN
alces-483	21	13	al	al	PROPN
alces-483	21	14	.	.	PROPN
alces-483	22	1	(	(	PUNCT
alces-483	22	2	1960	1960	NUM
alces-483	22	3	)	)	PUNCT
alces-483	22	4	provides	provide	VERB
alces-483	22	5	a	a	DET
alces-483	22	6	useful	useful	ADJ
alces-483	22	7	starting	starting	NOUN
alces-483	22	8	point	point	NOUN
alces-483	22	9	for	for	ADP
alces-483	22	10	“	"	PUNCT
alces-483	22	11	modern	modern	ADJ
alces-483	22	12	”	"	PUNCT
alces-483	22	13	consideration	consideration	NOUN
alces-483	22	14	of	of	ADP
alces-483	22	15	trophic	trophic	ADJ
alces-483	22	16	linkages	linkage	NOUN
alces-483	22	17	,	,	PUNCT
alces-483	22	18	by	by	ADP
alces-483	22	19	clearly	clearly	ADV
alces-483	22	20	laying	lay	VERB
alces-483	22	21	out	out	ADP
alces-483	22	22	concepts	concept	NOUN
alces-483	22	23	that	that	PRON
alces-483	22	24	were	be	AUX
alces-483	22	25	spawned	spawn	VERB
alces-483	22	26	earlier	early	ADV
alces-483	22	27	by	by	ADP
alces-483	22	28	elton	elton	PROPN
alces-483	22	29	(	(	PUNCT
alces-483	22	30	1927	1927	NUM
alces-483	22	31	)	)	PUNCT
alces-483	22	32	and	and	CCONJ
alces-483	22	33	lindemann	lindemann	PROPN
alces-483	22	34	(	(	PUNCT
alces-483	22	35	1942	1942	NUM
alces-483	22	36	)	)	PUNCT
alces-483	22	37	.	.	PUNCT
alces-483	23	1	paine	paine	NOUN
alces-483	23	2	(	(	PUNCT
alces-483	23	3	1980	1980	NUM
alces-483	23	4	)	)	PUNCT
alces-483	23	5	attached	attach	VERB
alces-483	23	6	the	the	DET
alces-483	23	7	name	name	NOUN
alces-483	23	8	“	"	PUNCT
alces-483	23	9	trophic	trophic	ADJ
alces-483	23	10	cascade	cascade	NOUN
alces-483	23	11	”	"	PUNCT
alces-483	23	12	to	to	ADP
alces-483	23	13	the	the	DET
alces-483	23	14	conceptual	conceptual	ADJ
alces-483	23	15	world	world	NOUN
alces-483	23	16	of	of	ADP
alces-483	23	17	hairston	hairston	PROPN
alces-483	23	18	et	et	PROPN
alces-483	23	19	al	al	PROPN
alces-483	23	20	.	.	PROPN
alces-483	24	1	(	(	PUNCT
alces-483	24	2	1960	1960	NUM
alces-483	24	3	)	)	PUNCT
alces-483	24	4	.	.	PUNCT
alces-483	25	1	models	model	NOUN
alces-483	25	2	by	by	ADP
alces-483	25	3	rosenzweig	rosenzweig	NOUN
alces-483	25	4	(	(	PUNCT
alces-483	25	5	1968	1968	NUM
alces-483	25	6	,	,	PUNCT
alces-483	25	7	1969	1969	NUM
alces-483	25	8	,	,	PUNCT
alces-483	25	9	1971	1971	NUM
alces-483	25	10	,	,	PUNCT
alces-483	25	11	1973	1973	NUM
alces-483	25	12	)	)	PUNCT
alces-483	25	13	,	,	PUNCT
alces-483	25	14	oksanen	oksanen	PROPN
alces-483	25	15	et	et	PROPN
alces-483	25	16	al	al	PROPN
alces-483	25	17	.	.	PROPN
alces-483	25	18	(	(	PUNCT
alces-483	25	19	1981	1981	NUM
alces-483	25	20	)	)	PUNCT
alces-483	25	21	,	,	PUNCT
alces-483	25	22	and	and	CCONJ
alces-483	25	23	oksanen	oksanen	PROPN
alces-483	25	24	(	(	PUNCT
alces-483	25	25	1983	1983	NUM
alces-483	25	26	)	)	PUNCT
alces-483	25	27	furthered	further	VERB
alces-483	25	28	understanding	understanding	NOUN
alces-483	25	29	of	of	ADP
alces-483	25	30	how	how	SCONJ
alces-483	25	31	length	length	NOUN
alces-483	25	32	of	of	ADP
alces-483	25	33	food	food	NOUN
alces-483	25	34	chains	chain	NOUN
alces-483	25	35	and	and	CCONJ
alces-483	25	36	variations	variation	NOUN
alces-483	25	37	in	in	ADP
alces-483	25	38	primary	primary	ADJ
alces-483	25	39	productivity	productivity	NOUN
alces-483	25	40	might	might	AUX
alces-483	25	41	alter	alter	VERB
alces-483	25	42	outcomes	outcome	NOUN
alces-483	25	43	(	(	PUNCT
alces-483	25	44	fretwell	fretwell	NOUN
alces-483	25	45	1987	1987	NUM
alces-483	25	46	)	)	PUNCT
alces-483	25	47	.	.	PUNCT
alces-483	26	1	opposing	oppose	VERB
alces-483	26	2	viewpoints	viewpoint	NOUN
alces-483	26	3	were	be	AUX
alces-483	26	4	numerous	numerous	ADJ
alces-483	26	5	,	,	PUNCT
alces-483	26	6	both	both	PRON
alces-483	26	7	then	then	ADV
alces-483	26	8	(	(	PUNCT
alces-483	26	9	murdoch	murdoch	PROPN
alces-483	26	10	1966	1966	NUM
alces-483	26	11	,	,	PUNCT
alces-483	26	12	ehrlich	ehrlich	NOUN
alces-483	26	13	and	and	CCONJ
alces-483	26	14	birch	birch	NOUN
alces-483	26	15	1967	1967	NUM
alces-483	26	16	)	)	PUNCT
alces-483	26	17	and	and	CCONJ
alces-483	26	18	now	now	ADV
alces-483	26	19	(	(	PUNCT
alces-483	26	20	paine	paine	NOUN
alces-483	26	21	2000	2000	NUM
alces-483	26	22	,	,	PUNCT
alces-483	26	23	polis	polis	PROPN
alces-483	26	24	et	et	PROPN
alces-483	26	25	al	al	PROPN
alces-483	26	26	.	.	PROPN
alces-483	26	27	2000	2000	NUM
alces-483	26	28	,	,	PUNCT
alces-483	26	29	schmitz	schmitz	PROPN
alces-483	26	30	et	et	PROPN
alces-483	26	31	al	al	PROPN
alces-483	26	32	.	.	PROPN
alces-483	26	33	2000	2000	NUM
alces-483	26	34	)	)	PUNCT
alces-483	26	35	.	.	PUNCT
alces-483	27	1	for	for	ADP
alces-483	27	2	the	the	DET
alces-483	27	3	most	most	ADJ
alces-483	27	4	part	part	NOUN
alces-483	27	5	,	,	PUNCT
alces-483	27	6	research	research	NOUN
alces-483	27	7	on	on	ADP
alces-483	27	8	ungulate	ungulate	ADJ
alces-483	27	9	population	population	NOUN
alces-483	27	10	dynamics	dynamic	NOUN
alces-483	27	11	has	have	AUX
alces-483	27	12	been	be	AUX
alces-483	27	13	a	a	DET
alces-483	27	14	separate	separate	ADJ
alces-483	27	15	but	but	CCONJ
alces-483	27	16	parallel	parallel	ADJ
alces-483	27	17	parade	parade	NOUN
alces-483	27	18	,	,	PUNCT
alces-483	27	19	highlighting	highlight	VERB
alces-483	27	20	the	the	DET
alces-483	27	21	respective	respective	ADJ
alces-483	27	22	roles	role	NOUN
alces-483	27	23	of	of	ADP
alces-483	27	24	density	density	NOUN
alces-483	27	25	-	-	PUNCT
alces-483	27	26	dependence	dependence	NOUN
alces-483	27	27	and	and	CCONJ
alces-483	27	28	predation	predation	NOUN
alces-483	27	29	(	(	PUNCT
alces-483	27	30	krebs	krebs	NOUN
alces-483	27	31	1995	1995	NUM
alces-483	27	32	)	)	PUNCT
alces-483	27	33	.	.	PUNCT
alces-483	28	1	ungulate	ungulate	ADJ
alces-483	28	2	researchers	researcher	NOUN
alces-483	28	3	seemed	seem	VERB
alces-483	28	4	“	"	PUNCT
alces-483	28	5	data	datum	NOUN
alces-483	28	6	-	-	PUNCT
alces-483	28	7	challenged	challenge	VERB
alces-483	28	8	”	"	PUNCT
alces-483	28	9	,	,	PUNCT
alces-483	28	10	with	with	ADP
alces-483	28	11	findings	finding	NOUN
alces-483	28	12	limited	limit	VERB
alces-483	28	13	by	by	ADP
alces-483	28	14	a	a	DET
alces-483	28	15	paucity	paucity	NOUN
alces-483	28	16	of	of	ADP
alces-483	28	17	experimental	experimental	ADJ
alces-483	28	18	studies	study	NOUN
alces-483	28	19	,	,	PUNCT
alces-483	28	20	the	the	DET
alces-483	28	21	slow	slow	ADJ
alces-483	28	22	temporal	temporal	ADJ
alces-483	28	23	scale	scale	NOUN
alces-483	28	24	of	of	ADP
alces-483	28	25	population	population	NOUN
alces-483	28	26	fluctuations	fluctuation	NOUN
alces-483	28	27	,	,	PUNCT
alces-483	28	28	and	and	CCONJ
alces-483	28	29	logistical	logistical	ADJ
alces-483	28	30	challenges	challenge	NOUN
alces-483	28	31	that	that	PRON
alces-483	28	32	have	have	AUX
alces-483	28	33	limited	limit	VERB
alces-483	28	34	the	the	DET
alces-483	28	35	spatial	spatial	ADJ
alces-483	28	36	and	and	CCONJ
alces-483	28	37	temporal	temporal	ADJ
alces-483	28	38	context	context	NOUN
alces-483	28	39	of	of	ADP
alces-483	28	40	research	research	NOUN
alces-483	28	41	.	.	PUNCT
alces-483	29	1	the	the	DET
alces-483	29	2	search	search	NOUN
alces-483	29	3	for	for	ADP
alces-483	29	4	general	general	ADJ
alces-483	29	5	patterns	pattern	NOUN
alces-483	29	6	in	in	ADP
alces-483	29	7	population	population	NOUN
alces-483	29	8	ecology	ecology	NOUN
alces-483	29	9	has	have	AUX
alces-483	29	10	not	not	PART
alces-483	29	11	been	be	AUX
alces-483	29	12	spearheaded	spearhead	VERB
alces-483	29	13	by	by	ADP
alces-483	29	14	large	large	ADJ
alces-483	29	15	-	-	PUNCT
alces-483	29	16	mammal	mammal	ADJ
alces-483	29	17	ecologists	ecologist	NOUN
alces-483	29	18	and	and	CCONJ
alces-483	29	19	,	,	PUNCT
alces-483	29	20	as	as	ADP
alces-483	29	21	a	a	DET
alces-483	29	22	result	result	NOUN
alces-483	29	23	,	,	PUNCT
alces-483	29	24	ungulate	ungulate	ADJ
alces-483	29	25	dynamics	dynamic	NOUN
alces-483	29	26	have	have	AUX
alces-483	29	27	usually	usually	ADV
alces-483	29	28	been	be	AUX
alces-483	29	29	understood	understand	VERB
alces-483	29	30	through	through	ADP
alces-483	29	31	a	a	DET
alces-483	29	32	theoretical	theoretical	ADJ
alces-483	29	33	looking	look	VERB
alces-483	29	34	-	-	PUNCT
alces-483	29	35	glass	glass	NOUN
alces-483	29	36	that	that	PRON
alces-483	29	37	was	be	AUX
alces-483	29	38	built	build	VERB
alces-483	29	39	to	to	PART
alces-483	29	40	view	view	VERB
alces-483	29	41	much	much	ADV
alces-483	29	42	smaller	small	ADJ
alces-483	29	43	animals	animal	NOUN
alces-483	29	44	,	,	PUNCT
alces-483	29	45	even	even	ADV
alces-483	29	46	invertebrates	invertebrate	NOUN
alces-483	29	47	(	(	PUNCT
alces-483	29	48	eberhardt	eberhardt	PROPN
alces-483	29	49	1997	1997	NUM
alces-483	29	50	)	)	PUNCT
alces-483	29	51	.	.	PUNCT
alces-483	30	1	or	or	CCONJ
alces-483	30	2	,	,	PUNCT
alces-483	30	3	more	more	ADV
alces-483	30	4	commonly	commonly	ADV
alces-483	30	5	,	,	PUNCT
alces-483	30	6	ungulate	ungulate	ADJ
alces-483	30	7	population	population	NOUN
alces-483	30	8	dynamics	dynamic	NOUN
alces-483	30	9	were	be	AUX
alces-483	30	10	viewed	view	VERB
alces-483	30	11	in	in	ADP
alces-483	30	12	a	a	DET
alces-483	30	13	game	game	NOUN
alces-483	30	14	management	management	NOUN
alces-483	30	15	context	context	NOUN
alces-483	30	16	built	build	VERB
alces-483	30	17	around	around	ADP
alces-483	30	18	the	the	DET
alces-483	30	19	dichotomy	dichotomy	NOUN
alces-483	30	20	that	that	SCONJ
alces-483	30	21	populations	population	NOUN
alces-483	30	22	were	be	AUX
alces-483	30	23	commonly	commonly	ADV
alces-483	30	24	limited	limit	VERB
alces-483	30	25	by	by	ADP
alces-483	30	26	food	food	NOUN
alces-483	30	27	or	or	CCONJ
alces-483	30	28	predators	predator	NOUN
alces-483	30	29	,	,	PUNCT
alces-483	30	30	with	with	ADP
alces-483	30	31	weather	weather	NOUN
alces-483	30	32	contributing	contribute	VERB
alces-483	30	33	to	to	ADP
alces-483	30	34	complexity	complexity	NOUN
alces-483	30	35	(	(	PUNCT
alces-483	30	36	bergerud	bergerud	NOUN
alces-483	30	37	1980	1980	NUM
alces-483	30	38	,	,	PUNCT
alces-483	30	39	sinclair	sinclair	NOUN
alces-483	30	40	1985	1985	NUM
alces-483	30	41	,	,	PUNCT
alces-483	30	42	mech	mech	NOUN
alces-483	30	43	et	et	PROPN
alces-483	30	44	al	al	PROPN
alces-483	30	45	.	.	PROPN
alces-483	30	46	1987	1987	NUM
alces-483	30	47	,	,	PUNCT
alces-483	30	48	fryxell	fryxell	NOUN
alces-483	30	49	and	and	CCONJ
alces-483	30	50	sinclair	sinclair	PROPN
alces-483	30	51	1988	1988	NUM
alces-483	30	52	,	,	PUNCT
alces-483	30	53	messier	messy	ADJ
alces-483	30	54	1991	1991	NUM
alces-483	30	55	,	,	PUNCT
alces-483	30	56	vanballenberghe	vanballenberghe	NOUN
alces-483	30	57	and	and	CCONJ
alces-483	30	58	ballard	ballard	PROPN
alces-483	30	59	1994	1994	NUM
alces-483	30	60	,	,	PUNCT
alces-483	30	61	ballard	ballard	NOUN
alces-483	30	62	and	and	CCONJ
alces-483	30	63	vanballenberghe	vanballenberghe	NOUN
alces-483	30	64	1998	1998	NUM
alces-483	30	65	)	)	PUNCT
alces-483	30	66	.	.	PUNCT
alces-483	31	1	the	the	DET
alces-483	31	2	food	food	NOUN
alces-483	31	3	or	or	CCONJ
alces-483	31	4	predation	predation	NOUN
alces-483	31	5	dichotomy	dichotomy	NOUN
alces-483	31	6	surfaces	surface	NOUN
alces-483	31	7	in	in	ADP
alces-483	31	8	the	the	DET
alces-483	31	9	recent	recent	ADJ
alces-483	31	10	ecological	ecological	ADJ
alces-483	31	11	literature	literature	NOUN
alces-483	31	12	as	as	ADP
alces-483	31	13	bottom	bottom	NOUN
alces-483	31	14	-	-	PUNCT
alces-483	31	15	up	up	NOUN
alces-483	31	16	or	or	CCONJ
alces-483	31	17	topdown	topdown	ADJ
alces-483	31	18	control	control	NOUN
alces-483	31	19	of	of	ADP
alces-483	31	20	food	food	NOUN
alces-483	31	21	webs	webs	NOUN
alces-483	31	22	(	(	PUNCT
alces-483	31	23	holt	holt	PROPN
alces-483	31	24	2000	2000	NUM
alces-483	31	25	,	,	PUNCT
alces-483	31	26	power	power	NOUN
alces-483	31	27	2000	2000	NUM
alces-483	31	28	)	)	PUNCT
alces-483	31	29	,	,	PUNCT
alces-483	31	30	respectively	respectively	ADV
alces-483	31	31	,	,	PUNCT
alces-483	31	32	and	and	CCONJ
alces-483	31	33	may	may	AUX
alces-483	31	34	be	be	AUX
alces-483	31	35	generalized	generalize	VERB
alces-483	31	36	and	and	CCONJ
alces-483	31	37	enlarged	enlarge	VERB
alces-483	31	38	into	into	ADP
alces-483	31	39	recent	recent	ADJ
alces-483	31	40	debate	debate	NOUN
alces-483	31	41	about	about	ADP
alces-483	31	42	ratio	ratio	NOUN
alces-483	31	43	-	-	PUNCT
alces-483	31	44	dependent	dependent	ADJ
alces-483	31	45	predation	predation	NOUN
alces-483	31	46	(	(	PUNCT
alces-483	31	47	arditi	arditi	NOUN
alces-483	31	48	and	and	CCONJ
alces-483	31	49	ginzburg	ginzburg	NOUN
alces-483	31	50	1989	1989	NUM
alces-483	31	51	)	)	PUNCT
alces-483	31	52	and	and	CCONJ
alces-483	31	53	trophic	trophic	ADJ
alces-483	31	54	cascades	cascade	NOUN
alces-483	31	55	(	(	PUNCT
alces-483	31	56	paine	paine	NOUN
alces-483	31	57	2000	2000	NUM
alces-483	31	58	,	,	PUNCT
alces-483	31	59	schmitz	schmitz	PROPN
alces-483	31	60	et	et	PROPN
alces-483	31	61	al	al	PROPN
alces-483	31	62	.	.	PROPN
alces-483	31	63	2000	2000	NUM
alces-483	31	64	)	)	PUNCT
alces-483	31	65	.	.	PUNCT
alces-483	32	1	ungulates	ungulate	NOUN
alces-483	32	2	and	and	CCONJ
alces-483	32	3	their	their	PRON
alces-483	32	4	predators	predator	NOUN
alces-483	32	5	in	in	ADP
alces-483	32	6	the	the	DET
alces-483	32	7	“	"	PUNCT
alces-483	32	8	real	real	ADJ
alces-483	32	9	”	"	PUNCT
alces-483	32	10	world	world	NOUN
alces-483	32	11	several	several	ADJ
alces-483	32	12	species	specie	NOUN
alces-483	32	13	of	of	ADP
alces-483	32	14	wild	wild	ADJ
alces-483	32	15	ursids	ursid	NOUN
alces-483	32	16	,	,	PUNCT
alces-483	32	17	felids	felid	NOUN
alces-483	32	18	,	,	PUNCT
alces-483	32	19	and	and	CCONJ
alces-483	32	20	canids	canids	PROPN
alces-483	32	21	together	together	ADV
alces-483	32	22	consitute	consitute	VERB
alces-483	32	23	the	the	DET
alces-483	32	24	predator	predator	NOUN
alces-483	32	25	fauna	fauna	NOUN
alces-483	32	26	for	for	ADP
alces-483	32	27	northern	northern	ADJ
alces-483	32	28	hemisphere	hemisphere	ADJ
alces-483	32	29	ungulates	ungulate	NOUN
alces-483	32	30	.	.	PUNCT
alces-483	33	1	by	by	ADP
alces-483	33	2	virtue	virtue	NOUN
alces-483	33	3	of	of	ADP
alces-483	33	4	their	their	PRON
alces-483	33	5	widespread	widespread	ADJ
alces-483	33	6	geographic	geographic	ADJ
alces-483	33	7	distribution	distribution	NOUN
alces-483	33	8	,	,	PUNCT
alces-483	33	9	group	group	NOUN
alces-483	33	10	-	-	PUNCT
alces-483	33	11	hunting	hunt	VERB
alces-483	33	12	nature	nature	NOUN
alces-483	33	13	,	,	PUNCT
alces-483	33	14	and	and	CCONJ
alces-483	33	15	yearround	yearround	VERB
alces-483	33	16	activity	activity	NOUN
alces-483	33	17	,	,	PUNCT
alces-483	33	18	we	we	PRON
alces-483	33	19	argue	argue	VERB
alces-483	33	20	that	that	SCONJ
alces-483	33	21	the	the	DET
alces-483	33	22	gray	gray	ADJ
alces-483	33	23	wolf	wolf	NOUN
alces-483	33	24	(	(	PUNCT
alces-483	33	25	canis	canis	NOUN
alces-483	33	26	lupus	lupus	NOUN
alces-483	33	27	)	)	PUNCT
alces-483	33	28	is	be	AUX
alces-483	33	29	the	the	DET
alces-483	33	30	most	most	ADV
alces-483	33	31	significant	significant	ADJ
alces-483	33	32	predator	predator	NOUN
alces-483	33	33	of	of	ADP
alces-483	33	34	ungulates	ungulate	NOUN
alces-483	33	35	in	in	ADP
alces-483	33	36	the	the	DET
alces-483	33	37	northern	northern	ADJ
alces-483	33	38	hemisphere	hemisphere	NOUN
alces-483	33	39	.	.	PUNCT
alces-483	34	1	where	where	SCONJ
alces-483	34	2	additional	additional	ADJ
alces-483	34	3	large	large	ADJ
alces-483	34	4	predators	predator	NOUN
alces-483	34	5	coexist	coexist	VERB
alces-483	34	6	with	with	ADP
alces-483	34	7	wolves	wolf	NOUN
alces-483	34	8	,	,	PUNCT
alces-483	34	9	along	along	ADP
alces-483	34	10	with	with	ADP
alces-483	34	11	human	human	ADJ
alces-483	34	12	hunters	hunter	NOUN
alces-483	34	13	,	,	PUNCT
alces-483	34	14	it	it	PRON
alces-483	34	15	is	be	AUX
alces-483	34	16	probably	probably	ADV
alces-483	34	17	reasonable	reasonable	ADJ
alces-483	34	18	to	to	PART
alces-483	34	19	assume	assume	VERB
alces-483	34	20	that	that	SCONJ
alces-483	34	21	predation	predation	NOUN
alces-483	34	22	by	by	ADP
alces-483	34	23	these	these	DET
alces-483	34	24	different	different	ADJ
alces-483	34	25	species	specie	NOUN
alces-483	34	26	is	be	AUX
alces-483	34	27	at	at	ADV
alces-483	34	28	least	least	ADJ
alces-483	34	29	partially	partially	ADV
alces-483	34	30	additive	additive	ADJ
alces-483	34	31	,	,	PUNCT
alces-483	34	32	thus	thus	ADV
alces-483	34	33	enlarging	enlarge	VERB
alces-483	34	34	the	the	DET
alces-483	34	35	ecological	ecological	ADJ
alces-483	34	36	influence	influence	NOUN
alces-483	34	37	of	of	ADP
alces-483	34	38	carnivores	carnivore	NOUN
alces-483	34	39	.	.	PUNCT
alces-483	35	1	it	it	PRON
alces-483	35	2	is	be	AUX
alces-483	35	3	also	also	ADV
alces-483	35	4	important	important	ADJ
alces-483	35	5	to	to	PART
alces-483	35	6	emphasize	emphasize	VERB
alces-483	35	7	that	that	SCONJ
alces-483	35	8	humans	human	NOUN
alces-483	35	9	continue	continue	VERB
alces-483	35	10	to	to	PART
alces-483	35	11	occupy	occupy	VERB
alces-483	35	12	an	an	DET
alces-483	35	13	ecological	ecological	ADJ
alces-483	35	14	niche	niche	NOUN
alces-483	35	15	as	as	ADP
alces-483	35	16	a	a	DET
alces-483	35	17	top	top	ADJ
alces-483	35	18	carnivore	carnivore	NOUN
alces-483	35	19	.	.	PUNCT
alces-483	36	1	as	as	ADP
alces-483	36	2	with	with	ADP
alces-483	36	3	other	other	ADJ
alces-483	36	4	large	large	ADJ
alces-483	36	5	predators	predator	NOUN
alces-483	36	6	,	,	PUNCT
alces-483	36	7	we	we	PRON
alces-483	36	8	have	have	VERB
alces-483	36	9	enormous	enormous	ADJ
alces-483	36	10	potential	potential	NOUN
alces-483	36	11	to	to	PART
alces-483	36	12	influence	influence	VERB
alces-483	36	13	ungulate	ungulate	ADJ
alces-483	36	14	dynamics	dynamic	NOUN
alces-483	36	15	through	through	ADP
alces-483	36	16	additive	additive	ADJ
alces-483	36	17	mortality	mortality	NOUN
alces-483	36	18	(	(	PUNCT
alces-483	36	19	crête	crête	ADJ
alces-483	36	20	1987	1987	NUM
alces-483	36	21	)	)	PUNCT
alces-483	36	22	,	,	PUNCT
alces-483	36	23	and	and	CCONJ
alces-483	36	24	certainly	certainly	ADV
alces-483	36	25	there	there	PRON
alces-483	36	26	is	be	VERB
alces-483	36	27	plenty	plenty	NOUN
alces-483	36	28	of	of	ADP
alces-483	36	29	historical	historical	ADJ
alces-483	36	30	evidence	evidence	NOUN
alces-483	36	31	that	that	PRON
alces-483	36	32	demonstrates	demonstrate	VERB
alces-483	36	33	our	our	PRON
alces-483	36	34	prowess	prowess	NOUN
alces-483	36	35	in	in	ADP
alces-483	36	36	excluding	exclude	VERB
alces-483	36	37	other	other	ADJ
alces-483	36	38	carnivores	carnivore	NOUN
alces-483	36	39	(	(	PUNCT
alces-483	36	40	hampton	hampton	PROPN
alces-483	36	41	1997	1997	NUM
alces-483	36	42	)	)	PUNCT
alces-483	36	43	.	.	PUNCT
alces-483	37	1	although	although	SCONJ
alces-483	37	2	we	we	PRON
alces-483	37	3	have	have	AUX
alces-483	37	4	instituted	institute	VERB
alces-483	37	5	regulations	regulation	NOUN
alces-483	37	6	to	to	PART
alces-483	37	7	control	control	VERB
alces-483	37	8	our	our	PRON
alces-483	37	9	alces	alce	NOUN
alces-483	37	10	vol	vol	NOUN
alces-483	37	11	.	.	PROPN
alces-483	37	12	39	39	NUM
alces-483	37	13	,	,	PUNCT
alces-483	37	14	2003	2003	NUM
alces-483	37	15	peterson	peterson	PROPN
alces-483	37	16	et	et	PROPN
alces-483	37	17	al	al	PROPN
alces-483	37	18	.	.	PROPN
alces-483	37	19	–	–	PUNCT
alces-483	37	20	space	space	NOUN
alces-483	37	21	and	and	CCONJ
alces-483	37	22	time	time	NOUN
alces-483	37	23	in	in	ADP
alces-483	37	24	predation	predation	NOUN
alces-483	37	25	dynamics	dynamic	VERB
alces-483	37	26	217	217	NUM
alces-483	37	27	harvest	harvest	NOUN
alces-483	37	28	,	,	PUNCT
alces-483	37	29	we	we	PRON
alces-483	37	30	have	have	AUX
alces-483	37	31	also	also	ADV
alces-483	37	32	increased	increase	VERB
alces-483	37	33	in	in	ADP
alces-483	37	34	number	number	NOUN
alces-483	37	35	ourselves	ourselves	PRON
alces-483	37	36	and	and	CCONJ
alces-483	37	37	,	,	PUNCT
alces-483	37	38	through	through	ADP
alces-483	37	39	technology	technology	NOUN
alces-483	37	40	,	,	PUNCT
alces-483	37	41	we	we	PRON
alces-483	37	42	have	have	AUX
alces-483	37	43	greatly	greatly	ADV
alces-483	37	44	increased	increase	VERB
alces-483	37	45	our	our	PRON
alces-483	37	46	ability	ability	NOUN
alces-483	37	47	to	to	PART
alces-483	37	48	exploit	exploit	VERB
alces-483	37	49	and	and	CCONJ
alces-483	37	50	overexploit	overexploit	ADJ
alces-483	37	51	.	.	PUNCT
alces-483	38	1	two	two	NUM
alces-483	38	2	studies	study	NOUN
alces-483	38	3	,	,	PUNCT
alces-483	38	4	on	on	ADP
alces-483	38	5	opposite	opposite	ADJ
alces-483	38	6	sides	side	NOUN
alces-483	38	7	of	of	ADP
alces-483	38	8	the	the	DET
alces-483	38	9	north	north	ADJ
alces-483	38	10	american	american	ADJ
alces-483	38	11	continent	continent	NOUN
alces-483	38	12	,	,	PUNCT
alces-483	38	13	illustrate	illustrate	VERB
alces-483	38	14	the	the	DET
alces-483	38	15	significance	significance	NOUN
alces-483	38	16	of	of	ADP
alces-483	38	17	predation	predation	NOUN
alces-483	38	18	in	in	ADP
alces-483	38	19	an	an	DET
alces-483	38	20	ecosystem	ecosystem	NOUN
alces-483	38	21	context	context	NOUN
alces-483	38	22	.	.	PUNCT
alces-483	39	1	gasaway	gasaway	ADV
alces-483	39	2	et	et	PROPN
alces-483	39	3	al	al	PROPN
alces-483	39	4	.	.	PROPN
alces-483	40	1	(	(	PUNCT
alces-483	40	2	1992	1992	NUM
alces-483	40	3	)	)	PUNCT
alces-483	40	4	compared	compare	VERB
alces-483	40	5	moose	moose	NOUN
alces-483	40	6	(	(	PUNCT
alces-483	40	7	alces	alce	NOUN
alces-483	40	8	alces	alce	NOUN
alces-483	40	9	)	)	PUNCT
alces-483	40	10	densities	density	NOUN
alces-483	40	11	in	in	ADP
alces-483	40	12	19	19	NUM
alces-483	40	13	study	study	NOUN
alces-483	40	14	areas	area	NOUN
alces-483	40	15	in	in	ADP
alces-483	40	16	the	the	DET
alces-483	40	17	yukon	yukon	PROPN
alces-483	40	18	and	and	CCONJ
alces-483	40	19	alaska	alaska	PROPN
alces-483	40	20	.	.	PUNCT
alces-483	41	1	the	the	DET
alces-483	41	2	combination	combination	NOUN
alces-483	41	3	of	of	ADP
alces-483	41	4	wolf	wolf	PROPN
alces-483	41	5	predation	predation	NOUN
alces-483	41	6	and	and	CCONJ
alces-483	41	7	bear	bear	PROPN
alces-483	41	8	(	(	PUNCT
alces-483	41	9	ursus	ursus	PROPN
alces-483	41	10	spp	spp	PROPN
alces-483	41	11	.	.	PUNCT
alces-483	41	12	)	)	PUNCT
alces-483	42	1	predation	predation	NOUN
alces-483	42	2	was	be	AUX
alces-483	42	3	sufficient	sufficient	ADJ
alces-483	42	4	to	to	PART
alces-483	42	5	reduce	reduce	VERB
alces-483	42	6	moose	moose	NOUN
alces-483	42	7	to	to	ADP
alces-483	42	8	a	a	DET
alces-483	42	9	level	level	NOUN
alces-483	42	10	far	far	ADV
alces-483	42	11	below	below	ADP
alces-483	42	12	“	"	PUNCT
alces-483	42	13	ecological	ecological	ADJ
alces-483	42	14	carrying	carrying	NOUN
alces-483	42	15	capacity	capacity	NOUN
alces-483	42	16	,	,	PUNCT
alces-483	42	17	”	"	PUNCT
alces-483	42	18	where	where	SCONJ
alces-483	42	19	density	density	NOUN
alces-483	42	20	-	-	PUNCT
alces-483	42	21	dependent	dependent	ADJ
alces-483	42	22	responses	response	NOUN
alces-483	42	23	to	to	ADP
alces-483	42	24	food	food	NOUN
alces-483	42	25	shortage	shortage	NOUN
alces-483	42	26	would	would	AUX
alces-483	42	27	be	be	AUX
alces-483	42	28	evident	evident	ADJ
alces-483	42	29	.	.	PUNCT
alces-483	43	1	this	this	PRON
alces-483	43	2	was	be	AUX
alces-483	43	3	true	true	ADJ
alces-483	43	4	even	even	ADV
alces-483	43	5	in	in	ADP
alces-483	43	6	denali	denali	PROPN
alces-483	43	7	national	national	PROPN
alces-483	43	8	park	park	PROPN
alces-483	43	9	,	,	PUNCT
alces-483	43	10	where	where	SCONJ
alces-483	43	11	moose	moose	NOUN
alces-483	43	12	are	be	AUX
alces-483	43	13	not	not	PART
alces-483	43	14	hunted	hunt	VERB
alces-483	43	15	by	by	ADP
alces-483	43	16	humans	human	NOUN
alces-483	43	17	.	.	PUNCT
alces-483	44	1	crête	crête	ADJ
alces-483	44	2	and	and	CCONJ
alces-483	44	3	manseau	manseau	NOUN
alces-483	44	4	(	(	PUNCT
alces-483	44	5	1996	1996	NUM
alces-483	44	6	)	)	PUNCT
alces-483	44	7	contrasted	contrast	VERB
alces-483	44	8	predictions	prediction	NOUN
alces-483	44	9	of	of	ADP
alces-483	44	10	prey	prey	NOUN
alces-483	44	11	-	-	PUNCT
alces-483	44	12	based	base	VERB
alces-483	44	13	trophic	trophic	ADJ
alces-483	44	14	dynamics	dynamic	NOUN
alces-483	44	15	models	model	NOUN
alces-483	44	16	(	(	PUNCT
alces-483	44	17	conforming	conform	VERB
alces-483	44	18	to	to	ADP
alces-483	44	19	ideas	idea	NOUN
alces-483	44	20	of	of	ADP
alces-483	44	21	hairston	hairston	PROPN
alces-483	44	22	et	et	PROPN
alces-483	44	23	al	al	PROPN
alces-483	44	24	.	.	PROPN
alces-483	44	25	1960	1960	NUM
alces-483	44	26	)	)	PUNCT
alces-483	44	27	with	with	ADP
alces-483	44	28	models	model	NOUN
alces-483	44	29	relying	rely	VERB
alces-483	44	30	on	on	ADP
alces-483	44	31	ratio	ratio	NOUN
alces-483	44	32	-	-	PUNCT
alces-483	44	33	dependent	dependent	ADJ
alces-483	44	34	predation	predation	NOUN
alces-483	44	35	,	,	PUNCT
alces-483	44	36	along	along	ADP
alces-483	44	37	a	a	DET
alces-483	44	38	latitudinal	latitudinal	ADJ
alces-483	44	39	productivity	productivity	NOUN
alces-483	44	40	gradient	gradient	NOUN
alces-483	44	41	(	(	PUNCT
alces-483	44	42	south	south	ADV
alces-483	44	43	to	to	ADP
alces-483	44	44	north	north	NOUN
alces-483	44	45	)	)	PUNCT
alces-483	44	46	in	in	ADP
alces-483	44	47	the	the	DET
alces-483	44	48	québec	québec	PROPN
alces-483	44	49	-	-	PUNCT
alces-483	44	50	labrador	labrador	NOUN
alces-483	44	51	peninsula	peninsula	NOUN
alces-483	44	52	.	.	PUNCT
alces-483	45	1	preybased	preybase	VERB
alces-483	45	2	models	model	NOUN
alces-483	45	3	predict	predict	VERB
alces-483	45	4	that	that	SCONJ
alces-483	45	5	a	a	DET
alces-483	45	6	change	change	NOUN
alces-483	45	7	at	at	ADP
alces-483	45	8	one	one	NUM
alces-483	45	9	trophic	trophic	ADJ
alces-483	45	10	level	level	NOUN
alces-483	45	11	will	will	AUX
alces-483	45	12	prompt	prompt	VERB
alces-483	45	13	changes	change	NOUN
alces-483	45	14	of	of	ADP
alces-483	45	15	alternating	alternate	VERB
alces-483	45	16	sign	sign	NOUN
alces-483	45	17	in	in	ADP
alces-483	45	18	successively	successively	ADV
alces-483	45	19	lower	low	ADJ
alces-483	45	20	trophic	trophic	ADJ
alces-483	45	21	levels	level	NOUN
alces-483	45	22	,	,	PUNCT
alces-483	45	23	while	while	SCONJ
alces-483	45	24	models	model	NOUN
alces-483	45	25	of	of	ADP
alces-483	45	26	ratio	ratio	NOUN
alces-483	45	27	-	-	PUNCT
alces-483	45	28	dependent	dependent	ADJ
alces-483	45	29	predation	predation	NOUN
alces-483	45	30	predict	predict	VERB
alces-483	45	31	that	that	SCONJ
alces-483	45	32	consumers	consumer	NOUN
alces-483	45	33	and	and	CCONJ
alces-483	45	34	their	their	PRON
alces-483	45	35	resources	resource	NOUN
alces-483	45	36	will	will	AUX
alces-483	45	37	increase	increase	VERB
alces-483	45	38	in	in	ADP
alces-483	45	39	parallel	parallel	NOUN
alces-483	45	40	as	as	ADP
alces-483	45	41	ecosystem	ecosystem	NOUN
alces-483	45	42	productivity	productivity	NOUN
alces-483	45	43	increases	increase	NOUN
alces-483	45	44	(	(	PUNCT
alces-483	45	45	arditi	arditi	NOUN
alces-483	45	46	and	and	CCONJ
alces-483	45	47	ginsburg	ginsburg	PROPN
alces-483	45	48	1989	1989	NUM
alces-483	45	49	)	)	PUNCT
alces-483	45	50	.	.	PUNCT
alces-483	46	1	crête	crête	PROPN
alces-483	46	2	and	and	CCONJ
alces-483	46	3	manseau	manseau	NOUN
alces-483	46	4	(	(	PUNCT
alces-483	46	5	1996	1996	NUM
alces-483	46	6	)	)	PUNCT
alces-483	46	7	found	find	VERB
alces-483	46	8	that	that	SCONJ
alces-483	46	9	tundra	tundra	NOUN
alces-483	46	10	areas	area	NOUN
alces-483	46	11	with	with	ADP
alces-483	46	12	inherently	inherently	ADV
alces-483	46	13	low	low	ADJ
alces-483	46	14	productivity	productivity	NOUN
alces-483	46	15	,	,	PUNCT
alces-483	46	16	where	where	SCONJ
alces-483	46	17	caribou	caribou	PROPN
alces-483	46	18	(	(	PUNCT
alces-483	46	19	rangifer	rangifer	PROPN
alces-483	46	20	tarandus	tarandus	NOUN
alces-483	46	21	)	)	PUNCT
alces-483	46	22	are	be	AUX
alces-483	46	23	the	the	DET
alces-483	46	24	only	only	ADJ
alces-483	46	25	ungulate	ungulate	ADJ
alces-483	46	26	present	present	NOUN
alces-483	46	27	,	,	PUNCT
alces-483	46	28	supported	support	VERB
alces-483	46	29	a	a	DET
alces-483	46	30	basic	basic	ADJ
alces-483	46	31	2	2	NUM
alces-483	46	32	-	-	PUNCT
alces-483	46	33	link	link	NOUN
alces-483	46	34	system	system	NOUN
alces-483	46	35	(	(	PUNCT
alces-483	46	36	caribou	caribou	NOUN
alces-483	46	37	and	and	CCONJ
alces-483	46	38	forage	forage	NOUN
alces-483	46	39	)	)	PUNCT
alces-483	46	40	and	and	CCONJ
alces-483	46	41	wolf	wolf	PROPN
alces-483	46	42	predation	predation	NOUN
alces-483	46	43	was	be	AUX
alces-483	46	44	unimportant	unimportant	ADJ
alces-483	46	45	.	.	PUNCT
alces-483	47	1	forage	forage	NOUN
alces-483	47	2	production	production	NOUN
alces-483	47	3	increased	increase	VERB
alces-483	47	4	within	within	ADP
alces-483	47	5	the	the	DET
alces-483	47	6	forested	forest	VERB
alces-483	47	7	zone	zone	NOUN
alces-483	47	8	where	where	SCONJ
alces-483	47	9	caribou	caribou	NOUN
alces-483	47	10	and	and	CCONJ
alces-483	47	11	moose	moose	PROPN
alces-483	47	12	supported	support	VERB
alces-483	47	13	wolf	wolf	NOUN
alces-483	47	14	populations	population	NOUN
alces-483	47	15	,	,	PUNCT
alces-483	47	16	but	but	CCONJ
alces-483	47	17	both	both	DET
alces-483	47	18	herbivores	herbivore	NOUN
alces-483	47	19	and	and	CCONJ
alces-483	47	20	carnivores	carnivore	NOUN
alces-483	47	21	were	be	AUX
alces-483	47	22	relatively	relatively	ADV
alces-483	47	23	scarce	scarce	ADJ
alces-483	47	24	.	.	PUNCT
alces-483	48	1	further	far	ADV
alces-483	48	2	south	south	ADV
alces-483	48	3	,	,	PUNCT
alces-483	48	4	just	just	ADV
alces-483	48	5	north	north	ADV
alces-483	48	6	of	of	ADP
alces-483	48	7	the	the	DET
alces-483	48	8	saint	saint	PROPN
alces-483	48	9	lawrence	lawrence	PROPN
alces-483	48	10	river	river	PROPN
alces-483	48	11	,	,	PUNCT
alces-483	48	12	forage	forage	NOUN
alces-483	48	13	production	production	NOUN
alces-483	48	14	was	be	AUX
alces-483	48	15	high	high	ADJ
alces-483	48	16	but	but	CCONJ
alces-483	48	17	predation	predation	NOUN
alces-483	48	18	by	by	ADP
alces-483	48	19	wolves	wolf	NOUN
alces-483	48	20	and	and	CCONJ
alces-483	48	21	black	black	ADJ
alces-483	48	22	bears	bear	NOUN
alces-483	48	23	(	(	PUNCT
alces-483	48	24	ursus	ursus	PROPN
alces-483	48	25	americanus	americanus	PROPN
alces-483	48	26	)	)	PUNCT
alces-483	48	27	was	be	AUX
alces-483	48	28	thought	think	VERB
alces-483	48	29	to	to	PART
alces-483	48	30	limit	limit	VERB
alces-483	48	31	moose	moose	NOUN
alces-483	48	32	density	density	NOUN
alces-483	48	33	.	.	PUNCT
alces-483	49	1	where	where	SCONJ
alces-483	49	2	wolves	wolf	NOUN
alces-483	49	3	were	be	AUX
alces-483	49	4	extirpated	extirpate	VERB
alces-483	49	5	on	on	ADP
alces-483	49	6	the	the	DET
alces-483	49	7	gaspé	gaspé	NOUN
alces-483	49	8	peninsula	peninsula	NOUN
alces-483	49	9	,	,	PUNCT
alces-483	49	10	moose	moose	PROPN
alces-483	49	11	were	be	AUX
alces-483	49	12	7	7	NUM
alces-483	49	13	times	time	NOUN
alces-483	49	14	more	more	ADV
alces-483	49	15	abundant	abundant	ADJ
alces-483	49	16	than	than	ADP
alces-483	49	17	in	in	ADP
alces-483	49	18	the	the	DET
alces-483	49	19	wolf	wolf	NOUN
alces-483	49	20	-	-	PUNCT
alces-483	49	21	inhabited	inhabit	VERB
alces-483	49	22	area	area	NOUN
alces-483	49	23	by	by	ADP
alces-483	49	24	the	the	DET
alces-483	49	25	saint	saint	PROPN
alces-483	49	26	lawrence	lawrence	PROPN
alces-483	49	27	river	river	PROPN
alces-483	49	28	,	,	PUNCT
alces-483	49	29	even	even	ADV
alces-483	49	30	though	though	SCONJ
alces-483	49	31	preferred	prefer	VERB
alces-483	49	32	-	-	PUNCT
alces-483	49	33	forage	forage	NOUN
alces-483	49	34	production	production	NOUN
alces-483	49	35	was	be	AUX
alces-483	49	36	relatively	relatively	ADV
alces-483	49	37	low	low	ADJ
alces-483	49	38	for	for	ADP
alces-483	49	39	that	that	DET
alces-483	49	40	latitude	latitude	NOUN
alces-483	49	41	.	.	PUNCT
alces-483	50	1	results	result	NOUN
alces-483	50	2	were	be	AUX
alces-483	50	3	interpreted	interpret	VERB
alces-483	50	4	as	as	ADP
alces-483	50	5	support	support	NOUN
alces-483	50	6	for	for	ADP
alces-483	50	7	the	the	DET
alces-483	50	8	socalled	socalled	ADJ
alces-483	50	9	trophic	trophic	ADJ
alces-483	50	10	-	-	PUNCT
alces-483	50	11	cascade	cascade	NOUN
alces-483	50	12	model	model	NOUN
alces-483	50	13	(	(	PUNCT
alces-483	50	14	polis	polis	PROPN
alces-483	50	15	et	et	PROPN
alces-483	50	16	al	al	PROPN
alces-483	50	17	.	.	PROPN
alces-483	50	18	2000	2000	NUM
alces-483	50	19	)	)	PUNCT
alces-483	50	20	.	.	PUNCT
alces-483	51	1	gasaway	gasaway	VERB
alces-483	51	2	et	et	PROPN
alces-483	51	3	al	al	PROPN
alces-483	51	4	.	.	PROPN
alces-483	52	1	(	(	PUNCT
alces-483	52	2	1992	1992	NUM
alces-483	52	3	)	)	PUNCT
alces-483	52	4	asserted	assert	VERB
alces-483	52	5	that	that	SCONJ
alces-483	52	6	each	each	DET
alces-483	52	7	additional	additional	ADJ
alces-483	52	8	large	large	ADJ
alces-483	52	9	carnivore	carnivore	ADJ
alces-483	52	10	species	specie	NOUN
alces-483	52	11	resulted	result	VERB
alces-483	52	12	in	in	ADP
alces-483	52	13	a	a	DET
alces-483	52	14	stepwise	stepwise	ADJ
alces-483	52	15	reduction	reduction	NOUN
alces-483	52	16	in	in	ADP
alces-483	52	17	moose	moose	NOUN
alces-483	52	18	density	density	NOUN
alces-483	52	19	.	.	PUNCT
alces-483	53	1	if	if	SCONJ
alces-483	53	2	one	one	PRON
alces-483	53	3	considers	consider	VERB
alces-483	53	4	the	the	DET
alces-483	53	5	wolf	wolf	NOUN
alces-483	53	6	,	,	PUNCT
alces-483	53	7	brown	brown	ADJ
alces-483	53	8	bear	bear	NOUN
alces-483	53	9	,	,	PUNCT
alces-483	53	10	black	black	ADJ
alces-483	53	11	bear	bear	NOUN
alces-483	53	12	,	,	PUNCT
alces-483	53	13	and	and	CCONJ
alces-483	53	14	human	human	NOUN
alces-483	53	15	as	as	ADP
alces-483	53	16	the	the	DET
alces-483	53	17	suite	suite	NOUN
alces-483	53	18	of	of	ADP
alces-483	53	19	potential	potential	ADJ
alces-483	53	20	predators	predator	NOUN
alces-483	53	21	of	of	ADP
alces-483	53	22	moose	moose	NOUN
alces-483	53	23	,	,	PUNCT
alces-483	53	24	we	we	PRON
alces-483	53	25	see	see	VERB
alces-483	53	26	a	a	DET
alces-483	53	27	general	general	ADJ
alces-483	53	28	pattern	pattern	NOUN
alces-483	53	29	of	of	ADP
alces-483	53	30	reduced	reduce	VERB
alces-483	53	31	moose	moose	NOUN
alces-483	53	32	density	density	NOUN
alces-483	53	33	with	with	ADP
alces-483	53	34	each	each	DET
alces-483	53	35	additional	additional	ADJ
alces-483	53	36	predator	predator	NOUN
alces-483	53	37	,	,	PUNCT
alces-483	53	38	at	at	ADP
alces-483	53	39	least	least	ADJ
alces-483	53	40	when	when	SCONJ
alces-483	53	41	large	large	ADJ
alces-483	53	42	areas	area	NOUN
alces-483	53	43	of	of	ADP
alces-483	53	44	contiguous	contiguous	ADJ
alces-483	53	45	habitat	habitat	NOUN
alces-483	53	46	are	be	AUX
alces-483	53	47	considered	consider	VERB
alces-483	53	48	(	(	PUNCT
alces-483	53	49	fig	fig	NOUN
alces-483	53	50	.	.	NOUN
alces-483	53	51	1	1	NUM
alces-483	53	52	)	)	PUNCT
alces-483	53	53	.	.	PUNCT
alces-483	54	1	gasaway	gasaway	VERB
alces-483	54	2	et	et	PROPN
alces-483	54	3	al	al	PROPN
alces-483	54	4	.	.	PROPN
alces-483	55	1	(	(	PUNCT
alces-483	55	2	1992	1992	NUM
alces-483	55	3	)	)	PUNCT
alces-483	55	4	compared	compare	VERB
alces-483	55	5	moose	moose	NOUN
alces-483	55	6	density	density	NOUN
alces-483	55	7	where	where	SCONJ
alces-483	55	8	predators	predator	NOUN
alces-483	55	9	were	be	AUX
alces-483	55	10	lightly	lightly	ADV
alces-483	55	11	exploited	exploit	VERB
alces-483	55	12	to	to	ADP
alces-483	55	13	those	those	PRON
alces-483	55	14	where	where	SCONJ
alces-483	55	15	predafig	predafig	NOUN
alces-483	55	16	.	.	PROPN
alces-483	56	1	1	1	NUM
alces-483	56	2	.	.	X
alces-483	56	3	ungulate	ungulate	ADJ
alces-483	56	4	density	density	NOUN
alces-483	56	5	is	be	AUX
alces-483	56	6	related	relate	VERB
alces-483	56	7	to	to	ADP
alces-483	56	8	number	number	NOUN
alces-483	56	9	of	of	ADP
alces-483	56	10	predator	predator	NOUN
alces-483	56	11	species	specie	NOUN
alces-483	56	12	in	in	ADP
alces-483	56	13	large	large	ADJ
alces-483	56	14	areas	area	NOUN
alces-483	56	15	of	of	ADP
alces-483	56	16	contiguous	contiguous	ADJ
alces-483	56	17	habitat	habitat	NOUN
alces-483	56	18	where	where	SCONJ
alces-483	56	19	they	they	PRON
alces-483	56	20	are	be	AUX
alces-483	56	21	principal	principal	ADJ
alces-483	56	22	prey	prey	NOUN
alces-483	56	23	species	specie	NOUN
alces-483	56	24	for	for	ADP
alces-483	56	25	black	black	ADJ
alces-483	56	26	bears	bear	NOUN
alces-483	56	27	,	,	PUNCT
alces-483	56	28	brown	brown	ADJ
alces-483	56	29	bears	bear	NOUN
alces-483	56	30	,	,	PUNCT
alces-483	56	31	gray	gray	ADJ
alces-483	56	32	wolves	wolf	NOUN
alces-483	56	33	,	,	PUNCT
alces-483	56	34	and	and	CCONJ
alces-483	56	35	humans	human	NOUN
alces-483	56	36	.	.	PUNCT
alces-483	57	1	moose	moose	PROPN
alces-483	57	2	,	,	PUNCT
alces-483	57	3	elk	elk	NOUN
alces-483	57	4	,	,	PUNCT
alces-483	57	5	and	and	CCONJ
alces-483	57	6	bison	bison	NOUN
alces-483	57	7	are	be	AUX
alces-483	57	8	indicated	indicate	VERB
alces-483	57	9	by	by	ADP
alces-483	57	10	unfilled	unfilled	ADJ
alces-483	57	11	circles	circle	NOUN
alces-483	57	12	,	,	PUNCT
alces-483	57	13	white	white	ADJ
alces-483	57	14	-	-	PUNCT
alces-483	57	15	tailed	tail	VERB
alces-483	57	16	deer	deer	NOUN
alces-483	57	17	are	be	AUX
alces-483	57	18	indicated	indicate	VERB
alces-483	57	19	by	by	ADP
alces-483	57	20	“	"	PUNCT
alces-483	57	21	x	x	NOUN
alces-483	57	22	”	"	PUNCT
alces-483	57	23	,	,	PUNCT
alces-483	57	24	and	and	CCONJ
alces-483	57	25	elk	elk	PROPN
alces-483	57	26	in	in	ADP
alces-483	57	27	yellowstone	yellowstone	PROPN
alces-483	57	28	’s	’s	PART
alces-483	57	29	northern	northern	ADJ
alces-483	57	30	range	range	NOUN
alces-483	57	31	(	(	PUNCT
alces-483	57	32	prior	prior	ADV
alces-483	57	33	to	to	ADP
alces-483	57	34	wolf	wolf	NOUN
alces-483	57	35	introduction	introduction	NOUN
alces-483	57	36	in	in	ADP
alces-483	57	37	1995	1995	NUM
alces-483	57	38	)	)	PUNCT
alces-483	57	39	are	be	AUX
alces-483	57	40	indicated	indicate	VERB
alces-483	57	41	by	by	ADP
alces-483	57	42	filled	fill	VERB
alces-483	57	43	circle	circle	NOUN
alces-483	57	44	.	.	PUNCT
alces-483	58	1	data	datum	NOUN
alces-483	58	2	points	point	NOUN
alces-483	58	3	provided	provide	VERB
alces-483	58	4	in	in	ADP
alces-483	58	5	appendix	appendix	NOUN
alces-483	58	6	1	1	NUM
alces-483	58	7	.	.	PUNCT
alces-483	58	8	space	space	NOUN
alces-483	58	9	and	and	CCONJ
alces-483	58	10	time	time	NOUN
alces-483	58	11	in	in	ADP
alces-483	58	12	predation	predation	NOUN
alces-483	58	13	dynamics	dynamic	NOUN
alces-483	58	14	–	–	PUNCT
alces-483	58	15	peterson	peterson	PROPN
alces-483	58	16	et	et	PROPN
alces-483	58	17	al	al	PROPN
alces-483	58	18	.	.	PROPN
alces-483	58	19	alces	alce	NOUN
alces-483	58	20	vol	vol	NOUN
alces-483	58	21	.	.	PROPN
alces-483	58	22	39	39	NUM
alces-483	58	23	,	,	PUNCT
alces-483	58	24	2003	2003	NUM
alces-483	58	25	218	218	NUM
alces-483	58	26	tors	tor	NOUN
alces-483	58	27	were	be	AUX
alces-483	58	28	reduced	reduce	VERB
alces-483	58	29	by	by	ADP
alces-483	58	30	humans	human	NOUN
alces-483	58	31	.	.	PUNCT
alces-483	59	1	the	the	DET
alces-483	59	2	latter	latter	ADJ
alces-483	59	3	areas	area	NOUN
alces-483	59	4	had	have	VERB
alces-483	59	5	an	an	DET
alces-483	59	6	average	average	ADJ
alces-483	59	7	moose	moose	NOUN
alces-483	59	8	density	density	NOUN
alces-483	59	9	of	of	ADP
alces-483	59	10	0.66	0.66	NUM
alces-483	59	11	moose	moose	NOUN
alces-483	59	12	/	/	SYM
alces-483	59	13	km2	km2	NOUN
alces-483	59	14	,	,	PUNCT
alces-483	59	15	while	while	SCONJ
alces-483	59	16	the	the	DET
alces-483	59	17	former	former	ADJ
alces-483	59	18	had	have	VERB
alces-483	59	19	0.15	0.15	NUM
alces-483	59	20	moose	moose	NOUN
alces-483	59	21	/	/	SYM
alces-483	59	22	km2	km2	NOUN
alces-483	59	23	.	.	PUNCT
alces-483	60	1	likewise	likewise	ADV
alces-483	60	2	,	,	PUNCT
alces-483	60	3	in	in	ADP
alces-483	60	4	the	the	DET
alces-483	60	5	absence	absence	NOUN
alces-483	60	6	of	of	ADP
alces-483	60	7	predators	predator	NOUN
alces-483	60	8	,	,	PUNCT
alces-483	60	9	caribou	caribou	NOUN
alces-483	60	10	exist	exist	VERB
alces-483	60	11	at	at	ADP
alces-483	60	12	densities	density	NOUN
alces-483	60	13	2	2	NUM
alces-483	60	14	orders	order	NOUN
alces-483	60	15	of	of	ADP
alces-483	60	16	magnitude	magnitude	NOUN
alces-483	60	17	higher	high	ADJ
alces-483	60	18	than	than	ADP
alces-483	60	19	when	when	SCONJ
alces-483	60	20	coexisting	coexist	VERB
alces-483	60	21	with	with	ADP
alces-483	60	22	multiple	multiple	ADJ
alces-483	60	23	species	specie	NOUN
alces-483	60	24	of	of	ADP
alces-483	60	25	carnivore	carnivore	NOUN
alces-483	60	26	(	(	PUNCT
alces-483	60	27	appendix	appendix	NOUN
alces-483	60	28	1	1	NUM
alces-483	60	29	)	)	PUNCT
alces-483	60	30	.	.	PUNCT
alces-483	61	1	caribou	caribou	NOUN
alces-483	61	2	employ	employ	VERB
alces-483	61	3	a	a	DET
alces-483	61	4	spacingout	spacingout	NOUN
alces-483	61	5	strategy	strategy	NOUN
alces-483	61	6	to	to	PART
alces-483	61	7	avoid	avoid	VERB
alces-483	61	8	wolf	wolf	PROPN
alces-483	61	9	predation	predation	NOUN
alces-483	61	10	,	,	PUNCT
alces-483	61	11	and	and	CCONJ
alces-483	61	12	so	so	ADV
alces-483	61	13	exist	exist	VERB
alces-483	61	14	at	at	ADP
alces-483	61	15	low	low	ADJ
alces-483	61	16	density	density	NOUN
alces-483	61	17	when	when	SCONJ
alces-483	61	18	coexisting	coexist	VERB
alces-483	61	19	with	with	ADP
alces-483	61	20	wolves	wolf	NOUN
alces-483	61	21	(	(	PUNCT
alces-483	61	22	bergerud	bergerud	NOUN
alces-483	61	23	1983a	1983a	NUM
alces-483	61	24	,	,	PUNCT
alces-483	61	25	b	b	NOUN
alces-483	61	26	)	)	PUNCT
alces-483	61	27	.	.	PUNCT
alces-483	62	1	where	where	SCONJ
alces-483	62	2	moose	moose	NOUN
alces-483	62	3	coexist	coexist	VERB
alces-483	62	4	with	with	ADP
alces-483	62	5	only	only	ADV
alces-483	62	6	one	one	NUM
alces-483	62	7	predator	predator	NOUN
alces-483	62	8	species	specie	NOUN
alces-483	62	9	,	,	PUNCT
alces-483	62	10	they	they	PRON
alces-483	62	11	can	can	AUX
alces-483	62	12	achieve	achieve	VERB
alces-483	62	13	high	high	ADJ
alces-483	62	14	population	population	NOUN
alces-483	62	15	density	density	NOUN
alces-483	62	16	(	(	PUNCT
alces-483	62	17	fig	fig	NOUN
alces-483	62	18	.	.	PUNCT
alces-483	63	1	1	1	NUM
alces-483	63	2	)	)	PUNCT
alces-483	63	3	.	.	PUNCT
alces-483	64	1	moose	moose	PROPN
alces-483	64	2	reach	reach	VERB
alces-483	64	3	densities	density	NOUN
alces-483	64	4	exceeding	exceed	VERB
alces-483	64	5	2	2	NUM
alces-483	64	6	/	/	SYM
alces-483	64	7	km2	km2	PROPN
alces-483	64	8	in	in	ADP
alces-483	64	9	sweden	sweden	PROPN
alces-483	64	10	where	where	SCONJ
alces-483	64	11	they	they	PRON
alces-483	64	12	face	face	VERB
alces-483	64	13	only	only	ADV
alces-483	64	14	human	human	ADJ
alces-483	64	15	hunters	hunter	NOUN
alces-483	64	16	,	,	PUNCT
alces-483	64	17	now	now	ADV
alces-483	64	18	carefully	carefully	ADV
alces-483	64	19	regulated	regulate	VERB
alces-483	64	20	.	.	PUNCT
alces-483	65	1	in	in	ADP
alces-483	65	2	a	a	DET
alces-483	65	3	park	park	NOUN
alces-483	65	4	with	with	ADP
alces-483	65	5	no	no	DET
alces-483	65	6	hunting	hunting	NOUN
alces-483	65	7	on	on	ADP
alces-483	65	8	the	the	DET
alces-483	65	9	gaspé	gaspé	NOUN
alces-483	65	10	peninsula	peninsula	NOUN
alces-483	65	11	in	in	ADP
alces-483	65	12	québec	québec	PROPN
alces-483	65	13	,	,	PUNCT
alces-483	65	14	and	and	CCONJ
alces-483	65	15	only	only	ADV
alces-483	65	16	black	black	ADJ
alces-483	65	17	bears	bear	NOUN
alces-483	65	18	as	as	ADP
alces-483	65	19	predators	predator	NOUN
alces-483	65	20	,	,	PUNCT
alces-483	65	21	moose	moose	NOUN
alces-483	65	22	density	density	NOUN
alces-483	65	23	is	be	AUX
alces-483	65	24	also	also	ADV
alces-483	65	25	>	>	X
alces-483	66	1	2	2	NUM
alces-483	66	2	/	/	SYM
alces-483	66	3	km2	km2	PROPN
alces-483	66	4	.	.	PUNCT
alces-483	66	5	moose	moose	PROPN
alces-483	66	6	exist	exist	VERB
alces-483	66	7	at	at	ADP
alces-483	66	8	a	a	DET
alces-483	66	9	comparably	comparably	ADV
alces-483	66	10	high	high	ADJ
alces-483	66	11	density	density	NOUN
alces-483	66	12	at	at	ADP
alces-483	66	13	isle	isle	PROPN
alces-483	66	14	royale	royale	PROPN
alces-483	66	15	,	,	PUNCT
alces-483	66	16	where	where	SCONJ
alces-483	66	17	unmanipulated	unmanipulated	ADJ
alces-483	66	18	wolves	wolf	NOUN
alces-483	66	19	are	be	AUX
alces-483	66	20	the	the	DET
alces-483	66	21	only	only	ADJ
alces-483	66	22	predator	predator	NOUN
alces-483	66	23	.	.	PUNCT
alces-483	67	1	add	add	VERB
alces-483	67	2	a	a	DET
alces-483	67	3	second	second	ADJ
alces-483	67	4	predator	predator	NOUN
alces-483	67	5	species	specie	NOUN
alces-483	67	6	to	to	ADP
alces-483	67	7	any	any	PRON
alces-483	67	8	of	of	ADP
alces-483	67	9	these	these	DET
alces-483	67	10	moose	moose	NOUN
alces-483	67	11	-	-	PUNCT
alces-483	67	12	dominated	dominate	VERB
alces-483	67	13	systems	system	NOUN
alces-483	67	14	and	and	CCONJ
alces-483	67	15	one	one	PRON
alces-483	67	16	should	should	AUX
alces-483	67	17	expect	expect	VERB
alces-483	67	18	a	a	DET
alces-483	67	19	decline	decline	NOUN
alces-483	67	20	in	in	ADP
alces-483	67	21	moose	moose	NOUN
alces-483	67	22	density	density	NOUN
alces-483	67	23	,	,	PUNCT
alces-483	67	24	a	a	DET
alces-483	67	25	prediction	prediction	NOUN
alces-483	67	26	borne	bear	VERB
alces-483	67	27	out	out	ADP
alces-483	67	28	in	in	ADP
alces-483	67	29	many	many	ADJ
alces-483	67	30	areas	area	NOUN
alces-483	67	31	(	(	PUNCT
alces-483	67	32	fig	fig	NOUN
alces-483	67	33	.	.	PUNCT
alces-483	67	34	1	1	NUM
alces-483	67	35	)	)	PUNCT
alces-483	67	36	.	.	PUNCT
alces-483	68	1	put	put	VERB
alces-483	68	2	bears	bear	NOUN
alces-483	68	3	,	,	PUNCT
alces-483	68	4	wolves	wolf	NOUN
alces-483	68	5	,	,	PUNCT
alces-483	68	6	and	and	CCONJ
alces-483	68	7	humans	human	NOUN
alces-483	68	8	together	together	ADV
alces-483	68	9	in	in	ADP
alces-483	68	10	the	the	DET
alces-483	68	11	same	same	ADJ
alces-483	68	12	area	area	NOUN
alces-483	68	13	,	,	PUNCT
alces-483	68	14	and	and	CCONJ
alces-483	68	15	only	only	ADV
alces-483	68	16	rarely	rarely	ADV
alces-483	68	17	would	would	AUX
alces-483	68	18	moose	moose	VERB
alces-483	68	19	density	density	NOUN
alces-483	68	20	exceed	exceed	VERB
alces-483	68	21	1	1	NUM
alces-483	68	22	/	/	SYM
alces-483	68	23	km2	km2	NOUN
alces-483	68	24	.	.	PUNCT
alces-483	69	1	with	with	ADP
alces-483	69	2	exceptionally	exceptionally	ADV
alces-483	69	3	favorable	favorable	ADJ
alces-483	69	4	,	,	PUNCT
alces-483	69	5	but	but	CCONJ
alces-483	69	6	transient	transient	ADJ
alces-483	69	7	,	,	PUNCT
alces-483	69	8	habitat	habitat	NOUN
alces-483	69	9	,	,	PUNCT
alces-483	69	10	moose	moose	NOUN
alces-483	69	11	reached	reach	VERB
alces-483	69	12	0.8	0.8	NUM
alces-483	69	13	/	/	SYM
alces-483	69	14	km2	km2	PROPN
alces-483	69	15	on	on	ADP
alces-483	69	16	alaska	alaska	PROPN
alces-483	69	17	’s	’s	PART
alces-483	69	18	kenai	kenai	PROPN
alces-483	69	19	peninsula	peninsula	PROPN
alces-483	69	20	,	,	PUNCT
alces-483	69	21	where	where	SCONJ
alces-483	69	22	a	a	DET
alces-483	69	23	hunted	hunted	ADJ
alces-483	69	24	moose	moose	NOUN
alces-483	69	25	population	population	NOUN
alces-483	69	26	was	be	AUX
alces-483	69	27	also	also	ADV
alces-483	69	28	preyed	preyed	ADJ
alces-483	69	29	on	on	ADP
alces-483	69	30	by	by	ADP
alces-483	69	31	wolves	wolf	NOUN
alces-483	69	32	and	and	CCONJ
alces-483	69	33	black	black	ADJ
alces-483	69	34	bears	bear	NOUN
alces-483	69	35	(	(	PUNCT
alces-483	69	36	bangs	bang	NOUN
alces-483	69	37	and	and	CCONJ
alces-483	69	38	bailey	bailey	NOUN
alces-483	69	39	1980	1980	NUM
alces-483	69	40	,	,	PUNCT
alces-483	69	41	peterson	peterson	PROPN
alces-483	69	42	et	et	PROPN
alces-483	69	43	al	al	PROPN
alces-483	69	44	.	.	PROPN
alces-483	69	45	1984	1984	NUM
alces-483	69	46	)	)	PUNCT
alces-483	69	47	.	.	PUNCT
alces-483	70	1	but	but	CCONJ
alces-483	70	2	across	across	ADP
alces-483	70	3	the	the	DET
alces-483	70	4	northern	northern	ADJ
alces-483	70	5	hemisphere	hemisphere	NOUN
alces-483	70	6	,	,	PUNCT
alces-483	70	7	where	where	SCONJ
alces-483	70	8	moose	moose	NOUN
alces-483	70	9	are	be	AUX
alces-483	70	10	typically	typically	ADV
alces-483	70	11	hunted	hunt	VERB
alces-483	70	12	by	by	ADP
alces-483	70	13	bears	bear	NOUN
alces-483	70	14	,	,	PUNCT
alces-483	70	15	wolves	wolf	NOUN
alces-483	70	16	,	,	PUNCT
alces-483	70	17	and	and	CCONJ
alces-483	70	18	humans	human	NOUN
alces-483	70	19	,	,	PUNCT
alces-483	70	20	their	their	PRON
alces-483	70	21	density	density	NOUN
alces-483	70	22	is	be	AUX
alces-483	70	23	usually	usually	ADV
alces-483	70	24	on	on	ADP
alces-483	70	25	the	the	DET
alces-483	70	26	order	order	NOUN
alces-483	70	27	of	of	ADP
alces-483	70	28	0.4	0.4	NUM
alces-483	70	29	/	/	SYM
alces-483	70	30	km2	km2	PROPN
alces-483	70	31	or	or	CCONJ
alces-483	70	32	less	less	ADJ
alces-483	70	33	.	.	PUNCT
alces-483	71	1	anything	anything	PRON
alces-483	71	2	that	that	PRON
alces-483	71	3	increases	increase	VERB
alces-483	71	4	the	the	DET
alces-483	71	5	reproductive	reproductive	ADJ
alces-483	71	6	potential	potential	NOUN
alces-483	71	7	of	of	ADP
alces-483	71	8	a	a	DET
alces-483	71	9	prey	prey	NOUN
alces-483	71	10	population	population	NOUN
alces-483	71	11	will	will	AUX
alces-483	71	12	tend	tend	VERB
alces-483	71	13	to	to	PART
alces-483	71	14	reduce	reduce	VERB
alces-483	71	15	the	the	DET
alces-483	71	16	impact	impact	NOUN
alces-483	71	17	of	of	ADP
alces-483	71	18	large	large	ADJ
alces-483	71	19	carnivores	carnivore	NOUN
alces-483	71	20	(	(	PUNCT
alces-483	71	21	seip	seip	NOUN
alces-483	71	22	1995	1995	NUM
alces-483	71	23	)	)	PUNCT
alces-483	71	24	.	.	PUNCT
alces-483	72	1	deer	deer	NOUN
alces-483	72	2	(	(	PUNCT
alces-483	72	3	odocoileus	odocoileus	ADJ
alces-483	72	4	virginianus	virginianus	PROPN
alces-483	72	5	)	)	PUNCT
alces-483	72	6	,	,	PUNCT
alces-483	72	7	for	for	ADP
alces-483	72	8	example	example	NOUN
alces-483	72	9	,	,	PUNCT
alces-483	72	10	should	should	AUX
alces-483	72	11	be	be	AUX
alces-483	72	12	able	able	ADJ
alces-483	72	13	to	to	PART
alces-483	72	14	coexist	coexist	VERB
alces-483	72	15	with	with	ADP
alces-483	72	16	wolves	wolf	NOUN
alces-483	72	17	at	at	ADP
alces-483	72	18	a	a	DET
alces-483	72	19	higher	high	ADJ
alces-483	72	20	density	density	NOUN
alces-483	72	21	than	than	ADP
alces-483	72	22	prey	prey	NOUN
alces-483	72	23	with	with	ADP
alces-483	72	24	lower	low	ADJ
alces-483	72	25	rates	rate	NOUN
alces-483	72	26	of	of	ADP
alces-483	72	27	increase	increase	NOUN
alces-483	72	28	(	(	PUNCT
alces-483	72	29	e.g.	e.g.	ADV
alces-483	72	30	,	,	PUNCT
alces-483	72	31	caribou	caribou	NOUN
alces-483	72	32	)	)	PUNCT
alces-483	72	33	.	.	PUNCT
alces-483	73	1	potentially	potentially	ADV
alces-483	73	2	,	,	PUNCT
alces-483	73	3	habitat	habitat	ADJ
alces-483	73	4	improvement	improvement	NOUN
alces-483	73	5	could	could	AUX
alces-483	73	6	accomplish	accomplish	VERB
alces-483	73	7	the	the	DET
alces-483	73	8	same	same	ADJ
alces-483	73	9	(	(	PUNCT
alces-483	73	10	orians	orian	NOUN
alces-483	73	11	et	et	PROPN
alces-483	73	12	al	al	PROPN
alces-483	73	13	.	.	PROPN
alces-483	73	14	1997	1997	NUM
alces-483	73	15	)	)	PUNCT
alces-483	73	16	.	.	PUNCT
alces-483	74	1	thus	thus	ADV
alces-483	74	2	,	,	PUNCT
alces-483	74	3	deer	deer	NOUN
alces-483	74	4	exist	exist	VERB
alces-483	74	5	at	at	ADP
alces-483	74	6	higher	high	ADJ
alces-483	74	7	density	density	NOUN
alces-483	74	8	in	in	ADP
alces-483	74	9	north	north	NOUN
alces-483	74	10	-	-	PUNCT
alces-483	74	11	central	central	ADJ
alces-483	74	12	minnesota	minnesota	NOUN
alces-483	74	13	,	,	PUNCT
alces-483	74	14	with	with	ADP
alces-483	74	15	intensive	intensive	ADJ
alces-483	74	16	forest	forest	NOUN
alces-483	74	17	management	management	NOUN
alces-483	74	18	,	,	PUNCT
alces-483	74	19	than	than	ADP
alces-483	74	20	in	in	ADP
alces-483	74	21	the	the	DET
alces-483	74	22	old	old	ADJ
alces-483	74	23	-	-	PUNCT
alces-483	74	24	growth	growth	NOUN
alces-483	74	25	forests	forest	NOUN
alces-483	74	26	of	of	ADP
alces-483	74	27	extreme	extreme	ADJ
alces-483	74	28	northern	northern	ADJ
alces-483	74	29	minnesota	minnesota	PROPN
alces-483	74	30	(	(	PUNCT
alces-483	74	31	nelson	nelson	PROPN
alces-483	74	32	and	and	CCONJ
alces-483	74	33	mech	mech	NOUN
alces-483	74	34	1986	1986	NUM
alces-483	74	35	,	,	PUNCT
alces-483	74	36	fuller	full	ADJ
alces-483	74	37	1989	1989	NUM
alces-483	74	38	)	)	PUNCT
alces-483	74	39	.	.	PUNCT
alces-483	75	1	additionally	additionally	ADV
alces-483	75	2	,	,	PUNCT
alces-483	75	3	the	the	DET
alces-483	75	4	presence	presence	NOUN
alces-483	75	5	of	of	ADP
alces-483	75	6	buffer	buffer	NOUN
alces-483	75	7	prey	prey	NOUN
alces-483	75	8	may	may	AUX
alces-483	75	9	modify	modify	VERB
alces-483	75	10	(	(	PUNCT
alces-483	75	11	increase	increase	VERB
alces-483	75	12	or	or	CCONJ
alces-483	75	13	decrease	decrease	NOUN
alces-483	75	14	)	)	PUNCT
alces-483	75	15	predation	predation	NOUN
alces-483	75	16	on	on	ADP
alces-483	75	17	a	a	DET
alces-483	75	18	single	single	ADJ
alces-483	75	19	prey	prey	NOUN
alces-483	75	20	species	specie	NOUN
alces-483	75	21	.	.	PUNCT
alces-483	76	1	the	the	DET
alces-483	76	2	mechanism	mechanism	NOUN
alces-483	76	3	by	by	ADP
alces-483	76	4	which	which	PRON
alces-483	76	5	predation	predation	NOUN
alces-483	76	6	limits	limit	VERB
alces-483	76	7	ungulate	ungulate	ADJ
alces-483	76	8	populations	population	NOUN
alces-483	76	9	is	be	AUX
alces-483	76	10	pervasive	pervasive	ADJ
alces-483	76	11	removal	removal	NOUN
alces-483	76	12	of	of	ADP
alces-483	76	13	pre	pre	ADJ
alces-483	76	14	-	-	ADJ
alces-483	76	15	reproductive	reproductive	ADJ
alces-483	76	16	juveniles	juvenile	NOUN
alces-483	76	17	(	(	PUNCT
alces-483	76	18	pimlott	pimlott	NOUN
alces-483	76	19	1967	1967	NUM
alces-483	76	20	)	)	PUNCT
alces-483	76	21	;	;	PUNCT
alces-483	76	22	where	where	SCONJ
alces-483	76	23	juveniles	juvenile	NOUN
alces-483	76	24	survive	survive	VERB
alces-483	76	25	poorly	poorly	ADV
alces-483	76	26	,	,	PUNCT
alces-483	76	27	populations	population	NOUN
alces-483	76	28	tend	tend	VERB
alces-483	76	29	to	to	PART
alces-483	76	30	decline	decline	VERB
alces-483	76	31	.	.	PUNCT
alces-483	77	1	at	at	ADP
alces-483	77	2	isle	isle	PROPN
alces-483	77	3	royale	royale	PROPN
alces-483	77	4	moose	moose	PROPN
alces-483	77	5	density	density	NOUN
alces-483	77	6	increased	increase	VERB
alces-483	77	7	during	during	ADP
alces-483	77	8	periods	period	NOUN
alces-483	77	9	when	when	SCONJ
alces-483	77	10	calf	calf	NOUN
alces-483	77	11	overwinter	overwinter	ADJ
alces-483	77	12	survival	survival	NOUN
alces-483	77	13	was	be	AUX
alces-483	77	14	~80	~80	NOUN
alces-483	77	15	%	%	NOUN
alces-483	77	16	,	,	PUNCT
alces-483	77	17	but	but	CCONJ
alces-483	77	18	declined	decline	VERB
alces-483	77	19	when	when	SCONJ
alces-483	77	20	survival	survival	NOUN
alces-483	77	21	was	be	AUX
alces-483	77	22	~50	~50	NUM
alces-483	77	23	%	%	NOUN
alces-483	77	24	(	(	PUNCT
alces-483	77	25	r.	r.	PROPN
alces-483	77	26	peterson	peterson	PROPN
alces-483	77	27	,	,	PUNCT
alces-483	77	28	unpublished	unpublished	ADJ
alces-483	77	29	data	datum	NOUN
alces-483	77	30	)	)	PUNCT
alces-483	77	31	.	.	PUNCT
alces-483	78	1	predator	predator	NOUN
alces-483	78	2	reduction	reduction	NOUN
alces-483	78	3	experiments	experiment	NOUN
alces-483	78	4	improved	improve	VERB
alces-483	78	5	moose	moose	NOUN
alces-483	78	6	calf	calf	NOUN
alces-483	78	7	survival	survival	NOUN
alces-483	78	8	about	about	ADV
alces-483	78	9	3	3	ADV
alces-483	78	10	-	-	NOUN
alces-483	78	11	fold	fold	ADJ
alces-483	78	12	and	and	CCONJ
alces-483	78	13	increased	increase	VERB
alces-483	78	14	finite	finite	ADJ
alces-483	78	15	rate	rate	NOUN
alces-483	78	16	of	of	ADP
alces-483	78	17	population	population	NOUN
alces-483	78	18	increase	increase	NOUN
alces-483	78	19	from	from	ADP
alces-483	78	20	1.0	1.0	NUM
alces-483	78	21	to	to	ADP
alces-483	78	22	2.3	2.3	NUM
alces-483	78	23	(	(	PUNCT
alces-483	78	24	gasaway	gasaway	NOUN
alces-483	78	25	et	et	PROPN
alces-483	78	26	al	al	PROPN
alces-483	78	27	.	.	PROPN
alces-483	78	28	1992	1992	NUM
alces-483	78	29	)	)	PUNCT
alces-483	78	30	.	.	PUNCT
alces-483	79	1	over	over	ADP
alces-483	79	2	the	the	DET
alces-483	79	3	past	past	ADJ
alces-483	79	4	25	25	NUM
alces-483	79	5	years	year	NOUN
alces-483	79	6	there	there	PRON
alces-483	79	7	have	have	AUX
alces-483	79	8	been	be	AUX
alces-483	79	9	numerous	numerous	ADJ
alces-483	79	10	studies	study	NOUN
alces-483	79	11	to	to	PART
alces-483	79	12	determine	determine	VERB
alces-483	79	13	the	the	DET
alces-483	79	14	extent	extent	NOUN
alces-483	79	15	and	and	CCONJ
alces-483	79	16	nature	nature	NOUN
alces-483	79	17	of	of	ADP
alces-483	79	18	juvenile	juvenile	ADJ
alces-483	79	19	ungulate	ungulate	ADJ
alces-483	79	20	mortality	mortality	NOUN
alces-483	79	21	(	(	PUNCT
alces-483	79	22	reviewed	review	VERB
alces-483	79	23	by	by	ADP
alces-483	79	24	ballard	ballard	PROPN
alces-483	79	25	and	and	CCONJ
alces-483	79	26	larsen	larsen	PROPN
alces-483	79	27	1987	1987	NUM
alces-483	79	28	,	,	PUNCT
alces-483	79	29	van	van	PROPN
alces-483	79	30	ballenberghe	ballenberghe	NOUN
alces-483	79	31	1987	1987	NUM
alces-483	79	32	,	,	PUNCT
alces-483	79	33	orians	orian	NOUN
alces-483	79	34	et	et	PROPN
alces-483	79	35	al	al	PROPN
alces-483	79	36	.	.	PROPN
alces-483	79	37	1997	1997	NUM
alces-483	79	38	)	)	PUNCT
alces-483	79	39	.	.	PUNCT
alces-483	80	1	in	in	ADP
alces-483	80	2	11	11	NUM
alces-483	80	3	studies	study	NOUN
alces-483	80	4	located	locate	VERB
alces-483	80	5	in	in	ADP
alces-483	80	6	9	9	NUM
alces-483	80	7	areas	area	NOUN
alces-483	80	8	of	of	ADP
alces-483	80	9	alaska	alaska	PROPN
alces-483	80	10	,	,	PUNCT
alces-483	80	11	yukon	yukon	PROPN
alces-483	80	12	,	,	PUNCT
alces-483	80	13	and	and	CCONJ
alces-483	80	14	british	british	PROPN
alces-483	80	15	columbia	columbia	PROPN
alces-483	80	16	where	where	SCONJ
alces-483	80	17	wolves	wolf	NOUN
alces-483	80	18	and	and	CCONJ
alces-483	80	19	bears	bear	NOUN
alces-483	80	20	existed	exist	VERB
alces-483	80	21	,	,	PUNCT
alces-483	80	22	radio	radio	NOUN
alces-483	80	23	-	-	PUNCT
alces-483	80	24	collars	collar	NOUN
alces-483	80	25	have	have	AUX
alces-483	80	26	been	be	AUX
alces-483	80	27	placed	place	VERB
alces-483	80	28	on	on	ADP
alces-483	80	29	623	623	NUM
alces-483	80	30	moose	moose	NOUN
alces-483	80	31	and	and	CCONJ
alces-483	80	32	462	462	NUM
alces-483	80	33	caribou	caribou	NOUN
alces-483	80	34	soon	soon	ADV
alces-483	80	35	after	after	ADP
alces-483	80	36	birth	birth	NOUN
alces-483	80	37	(	(	PUNCT
alces-483	80	38	orians	orian	NOUN
alces-483	80	39	et	et	PROPN
alces-483	80	40	al	al	PROPN
alces-483	80	41	.	.	PROPN
alces-483	80	42	1997	1997	NUM
alces-483	80	43	)	)	PUNCT
alces-483	80	44	.	.	PUNCT
alces-483	81	1	survival	survival	NOUN
alces-483	81	2	was	be	AUX
alces-483	81	3	monitored	monitor	VERB
alces-483	81	4	for	for	ADP
alces-483	81	5	variable	variable	ADJ
alces-483	81	6	periods	period	NOUN
alces-483	81	7	,	,	PUNCT
alces-483	81	8	from	from	ADP
alces-483	81	9	2	2	NUM
alces-483	81	10	months	month	NOUN
alces-483	81	11	to	to	ADP
alces-483	81	12	1	1	NUM
alces-483	81	13	year	year	NOUN
alces-483	81	14	,	,	PUNCT
alces-483	81	15	but	but	CCONJ
alces-483	81	16	clearly	clearly	ADV
alces-483	81	17	there	there	PRON
alces-483	81	18	was	be	VERB
alces-483	81	19	an	an	DET
alces-483	81	20	early	early	ADJ
alces-483	81	21	period	period	NOUN
alces-483	81	22	of	of	ADP
alces-483	81	23	greatest	great	ADJ
alces-483	81	24	risk	risk	NOUN
alces-483	81	25	during	during	ADP
alces-483	81	26	the	the	DET
alces-483	81	27	first	first	ADJ
alces-483	81	28	2	2	NUM
alces-483	81	29	months	month	NOUN
alces-483	81	30	of	of	ADP
alces-483	81	31	life	life	NOUN
alces-483	81	32	.	.	PUNCT
alces-483	82	1	for	for	ADP
alces-483	82	2	both	both	DET
alces-483	82	3	species	specie	NOUN
alces-483	82	4	,	,	PUNCT
alces-483	82	5	average	average	ADJ
alces-483	82	6	(	(	PUNCT
alces-483	82	7	mean	mean	VERB
alces-483	82	8	for	for	ADP
alces-483	82	9	all	all	DET
alces-483	82	10	studies	study	NOUN
alces-483	82	11	)	)	PUNCT
alces-483	82	12	survival	survival	NOUN
alces-483	82	13	rate	rate	NOUN
alces-483	82	14	of	of	ADP
alces-483	82	15	juveniles	juvenile	NOUN
alces-483	82	16	in	in	ADP
alces-483	82	17	their	their	PRON
alces-483	82	18	first	first	ADJ
alces-483	82	19	year	year	NOUN
alces-483	82	20	was	be	AUX
alces-483	82	21	only	only	ADV
alces-483	82	22	40	40	NUM
alces-483	82	23	%	%	NOUN
alces-483	82	24	.	.	PUNCT
alces-483	83	1	ballard	ballard	PROPN
alces-483	83	2	and	and	CCONJ
alces-483	83	3	larsen	larsen	PROPN
alces-483	83	4	(	(	PUNCT
alces-483	83	5	1987	1987	NUM
alces-483	83	6	)	)	PUNCT
alces-483	83	7	and	and	CCONJ
alces-483	83	8	van	van	PROPN
alces-483	83	9	ballenberghe	ballenberghe	NOUN
alces-483	83	10	(	(	PUNCT
alces-483	83	11	1987	1987	NUM
alces-483	83	12	)	)	PUNCT
alces-483	83	13	cited	cite	VERB
alces-483	83	14	several	several	ADJ
alces-483	83	15	studies	study	NOUN
alces-483	83	16	which	which	PRON
alces-483	83	17	indicated	indicate	VERB
alces-483	83	18	that	that	SCONJ
alces-483	83	19	predation	predation	NOUN
alces-483	83	20	on	on	ADP
alces-483	83	21	young	young	ADJ
alces-483	83	22	moose	moose	NOUN
alces-483	83	23	may	may	AUX
alces-483	83	24	account	account	VERB
alces-483	83	25	for	for	ADP
alces-483	83	26	79	79	NUM
alces-483	83	27	%	%	NOUN
alces-483	83	28	of	of	ADP
alces-483	83	29	neonate	neonate	NOUN
alces-483	83	30	deaths	death	NOUN
alces-483	83	31	;	;	PUNCT
alces-483	83	32	survival	survival	NOUN
alces-483	83	33	in	in	ADP
alces-483	83	34	the	the	DET
alces-483	83	35	first	first	ADJ
alces-483	83	36	8	8	NUM
alces-483	83	37	weeks	week	NOUN
alces-483	83	38	can	can	AUX
alces-483	83	39	be	be	AUX
alces-483	83	40	as	as	ADV
alces-483	83	41	low	low	ADJ
alces-483	83	42	as	as	ADP
alces-483	83	43	17	17	NUM
alces-483	83	44	%	%	NOUN
alces-483	83	45	.	.	PUNCT
alces-483	84	1	where	where	SCONJ
alces-483	84	2	moose	moose	NOUN
alces-483	84	3	were	be	AUX
alces-483	84	4	hunted	hunt	VERB
alces-483	84	5	in	in	ADP
alces-483	84	6	alaska	alaska	PROPN
alces-483	84	7	,	,	PUNCT
alces-483	84	8	losses	loss	NOUN
alces-483	84	9	to	to	ADP
alces-483	84	10	predation	predation	NOUN
alces-483	84	11	(	(	PUNCT
alces-483	84	12	31	31	NUM
alces-483	84	13	%	%	NOUN
alces-483	84	14	of	of	ADP
alces-483	84	15	postcalving	postcalve	VERB
alces-483	84	16	numbers	number	NOUN
alces-483	84	17	)	)	PUNCT
alces-483	84	18	alces	alce	NOUN
alces-483	84	19	vol	vol	NOUN
alces-483	84	20	.	.	PROPN
alces-483	84	21	39	39	NUM
alces-483	84	22	,	,	PUNCT
alces-483	84	23	2003	2003	NUM
alces-483	84	24	peterson	peterson	PROPN
alces-483	84	25	et	et	PROPN
alces-483	84	26	al	al	PROPN
alces-483	84	27	.	.	PROPN
alces-483	84	28	–	–	PUNCT
alces-483	84	29	space	space	NOUN
alces-483	84	30	and	and	CCONJ
alces-483	84	31	time	time	NOUN
alces-483	84	32	in	in	ADP
alces-483	84	33	predation	predation	NOUN
alces-483	84	34	dynamics	dynamic	NOUN
alces-483	84	35	219	219	NUM
alces-483	84	36	greatly	greatly	ADV
alces-483	84	37	exceeded	exceed	VERB
alces-483	84	38	loss	loss	NOUN
alces-483	84	39	to	to	ADP
alces-483	84	40	hunting	hunting	NOUN
alces-483	84	41	(	(	PUNCT
alces-483	84	42	1.5	1.5	NUM
alces-483	84	43	%	%	NOUN
alces-483	84	44	)	)	PUNCT
alces-483	84	45	and	and	CCONJ
alces-483	85	1	other	other	ADJ
alces-483	85	2	losses	loss	NOUN
alces-483	85	3	(	(	PUNCT
alces-483	85	4	6	6	NUM
alces-483	85	5	%	%	NOUN
alces-483	85	6	)	)	PUNCT
alces-483	85	7	(	(	PUNCT
alces-483	85	8	gasaway	gasaway	NOUN
alces-483	85	9	et	et	PROPN
alces-483	85	10	al	al	PROPN
alces-483	85	11	.	.	PROPN
alces-483	85	12	1992	1992	NUM
alces-483	85	13	)	)	PUNCT
alces-483	85	14	.	.	PUNCT
alces-483	86	1	in	in	ADP
alces-483	86	2	an	an	DET
alces-483	86	3	unhunted	unhunted	ADJ
alces-483	86	4	population	population	NOUN
alces-483	86	5	free	free	ADJ
alces-483	86	6	of	of	ADP
alces-483	86	7	bears	bear	NOUN
alces-483	86	8	and	and	CCONJ
alces-483	86	9	wolves	wolf	NOUN
alces-483	86	10	(	(	PUNCT
alces-483	86	11	rochester	rochester	PROPN
alces-483	86	12	,	,	PUNCT
alces-483	86	13	alberta	alberta	PROPN
alces-483	86	14	)	)	PUNCT
alces-483	86	15	,	,	PUNCT
alces-483	86	16	annual	annual	ADJ
alces-483	86	17	survival	survival	NOUN
alces-483	86	18	of	of	ADP
alces-483	86	19	moose	moose	NOUN
alces-483	86	20	calves	calf	NOUN
alces-483	86	21	was	be	AUX
alces-483	86	22	as	as	ADV
alces-483	86	23	high	high	ADJ
alces-483	86	24	as	as	ADP
alces-483	86	25	67	67	NUM
alces-483	86	26	%	%	NOUN
alces-483	86	27	,	,	PUNCT
alces-483	86	28	and	and	CCONJ
alces-483	86	29	41	41	NUM
alces-483	86	30	%	%	NOUN
alces-483	86	31	of	of	ADP
alces-483	86	32	calf	calf	NOUN
alces-483	86	33	/	/	SYM
alces-483	86	34	cow	cow	NOUN
alces-483	86	35	groups	group	NOUN
alces-483	86	36	in	in	ADP
alces-483	86	37	winter	winter	NOUN
alces-483	86	38	included	include	VERB
alces-483	86	39	twins	twin	NOUN
alces-483	86	40	(	(	PUNCT
alces-483	86	41	rolley	rolley	NOUN
alces-483	86	42	and	and	CCONJ
alces-483	86	43	keith	keith	PROPN
alces-483	86	44	1980	1980	NUM
alces-483	86	45	)	)	PUNCT
alces-483	86	46	.	.	PUNCT
alces-483	87	1	where	where	SCONJ
alces-483	87	2	2	2	NUM
alces-483	87	3	or	or	CCONJ
alces-483	87	4	more	more	ADJ
alces-483	87	5	species	specie	NOUN
alces-483	87	6	of	of	ADP
alces-483	87	7	large	large	ADJ
alces-483	87	8	carnivore	carnivore	NOUN
alces-483	87	9	were	be	AUX
alces-483	87	10	present	present	ADJ
alces-483	87	11	(	(	PUNCT
alces-483	87	12	8	8	NUM
alces-483	87	13	studies	study	NOUN
alces-483	87	14	with	with	ADP
alces-483	87	15	469	469	NUM
alces-483	87	16	moose	moose	NOUN
alces-483	87	17	calves	calf	NOUN
alces-483	87	18	total	total	NOUN
alces-483	87	19	)	)	PUNCT
alces-483	87	20	,	,	PUNCT
alces-483	87	21	average	average	ADJ
alces-483	87	22	survival	survival	NOUN
alces-483	87	23	to	to	ADP
alces-483	87	24	age	age	NOUN
alces-483	87	25	of	of	ADP
alces-483	87	26	6	6	NUM
alces-483	87	27	months	month	NOUN
alces-483	87	28	was	be	AUX
alces-483	87	29	30	30	NUM
alces-483	87	30	%	%	NOUN
alces-483	87	31	.	.	PUNCT
alces-483	88	1	in	in	ADP
alces-483	88	2	3	3	NUM
alces-483	88	3	study	study	NOUN
alces-483	88	4	areas	area	NOUN
alces-483	88	5	with	with	ADP
alces-483	88	6	0	0	NUM
alces-483	88	7	-	-	SYM
alces-483	88	8	1	1	NUM
alces-483	88	9	species	specie	NOUN
alces-483	88	10	of	of	ADP
alces-483	88	11	carnivore	carnivore	NOUN
alces-483	88	12	,	,	PUNCT
alces-483	88	13	average	average	ADJ
alces-483	88	14	6	6	NUM
alces-483	88	15	-	-	PUNCT
alces-483	88	16	months	month	NOUN
alces-483	88	17	survival	survival	NOUN
alces-483	88	18	of	of	ADP
alces-483	88	19	moose	moose	NOUN
alces-483	88	20	was	be	AUX
alces-483	88	21	67	67	NUM
alces-483	88	22	%	%	NOUN
alces-483	88	23	(	(	PUNCT
alces-483	88	24	3	3	NUM
alces-483	88	25	studies	study	NOUN
alces-483	88	26	with	with	ADP
alces-483	88	27	111	111	NUM
alces-483	88	28	calves	calf	NOUN
alces-483	88	29	)	)	PUNCT
alces-483	88	30	.	.	PUNCT
alces-483	89	1	most	most	ADJ
alces-483	89	2	mortality	mortality	NOUN
alces-483	89	3	occurred	occur	VERB
alces-483	89	4	within	within	ADP
alces-483	89	5	the	the	DET
alces-483	89	6	first	first	ADJ
alces-483	89	7	6	6	NUM
alces-483	89	8	weeks	week	NOUN
alces-483	89	9	of	of	ADP
alces-483	89	10	life	life	NOUN
alces-483	89	11	–	–	PUNCT
alces-483	89	12	black	black	ADJ
alces-483	89	13	bears	bear	NOUN
alces-483	89	14	being	be	AUX
alces-483	89	15	responsible	responsible	ADJ
alces-483	89	16	for	for	ADP
alces-483	89	17	2	2	NUM
alces-483	89	18	-	-	SYM
alces-483	89	19	50	50	NUM
alces-483	89	20	%	%	NOUN
alces-483	89	21	,	,	PUNCT
alces-483	89	22	grizzlies	grizzly	NOUN
alces-483	89	23	3	3	NUM
alces-483	89	24	-	-	SYM
alces-483	89	25	52	52	NUM
alces-483	89	26	%	%	NOUN
alces-483	89	27	,	,	PUNCT
alces-483	89	28	and	and	CCONJ
alces-483	89	29	wolves	wolf	NOUN
alces-483	89	30	2	2	NUM
alces-483	89	31	-	-	SYM
alces-483	89	32	18	18	NUM
alces-483	89	33	%	%	NOUN
alces-483	89	34	.	.	PUNCT
alces-483	90	1	if	if	SCONJ
alces-483	90	2	moose	moose	NOUN
alces-483	90	3	density	density	NOUN
alces-483	90	4	was	be	AUX
alces-483	90	5	high	high	ADJ
alces-483	90	6	,	,	PUNCT
alces-483	90	7	approaching	approach	VERB
alces-483	90	8	k	k	NOUN
alces-483	90	9	carrying	carry	VERB
alces-483	90	10	capacity	capacity	NOUN
alces-483	90	11	,	,	PUNCT
alces-483	90	12	predatorinduced	predatorinduce	VERB
alces-483	90	13	mortality	mortality	NOUN
alces-483	90	14	was	be	AUX
alces-483	90	15	considered	consider	VERB
alces-483	90	16	more	more	ADV
alces-483	90	17	likely	likely	ADJ
alces-483	90	18	to	to	PART
alces-483	90	19	be	be	AUX
alces-483	90	20	compensatory	compensatory	ADJ
alces-483	90	21	(	(	PUNCT
alces-483	90	22	orians	orian	NOUN
alces-483	90	23	et	et	PROPN
alces-483	90	24	al	al	PROPN
alces-483	90	25	.	.	PROPN
alces-483	90	26	1997	1997	NUM
alces-483	90	27	)	)	PUNCT
alces-483	90	28	.	.	PUNCT
alces-483	91	1	bear	bear	PROPN
alces-483	91	2	predation	predation	NOUN
alces-483	91	3	may	may	AUX
alces-483	91	4	be	be	AUX
alces-483	91	5	densityindependent	densityindependent	ADJ
alces-483	91	6	,	,	PUNCT
alces-483	91	7	but	but	CCONJ
alces-483	91	8	nevertheless	nevertheless	ADV
alces-483	91	9	a	a	DET
alces-483	91	10	significant	significant	ADJ
alces-483	91	11	mortality	mortality	NOUN
alces-483	91	12	factor	factor	NOUN
alces-483	91	13	in	in	ADP
alces-483	91	14	the	the	DET
alces-483	91	15	first	first	ADJ
alces-483	91	16	6	6	NUM
alces-483	91	17	months	month	NOUN
alces-483	91	18	of	of	ADP
alces-483	91	19	life	life	NOUN
alces-483	91	20	.	.	PUNCT
alces-483	92	1	on	on	ADP
alces-483	92	2	the	the	DET
alces-483	92	3	other	other	ADJ
alces-483	92	4	hand	hand	NOUN
alces-483	92	5	,	,	PUNCT
alces-483	92	6	wolf	wolf	PROPN
alces-483	92	7	predation	predation	NOUN
alces-483	92	8	has	have	VERB
alces-483	92	9	its	its	PRON
alces-483	92	10	greatest	great	ADJ
alces-483	92	11	impact	impact	NOUN
alces-483	92	12	in	in	ADP
alces-483	92	13	winter	winter	NOUN
alces-483	92	14	,	,	PUNCT
alces-483	92	15	when	when	SCONJ
alces-483	92	16	calves	calf	NOUN
alces-483	92	17	make	make	VERB
alces-483	92	18	up	up	ADP
alces-483	92	19	30	30	NUM
alces-483	92	20	-	-	SYM
alces-483	92	21	40	40	NUM
alces-483	92	22	%	%	NOUN
alces-483	92	23	of	of	ADP
alces-483	92	24	observed	observed	ADJ
alces-483	92	25	kills	kill	NOUN
alces-483	92	26	(	(	PUNCT
alces-483	92	27	peterson	peterson	PROPN
alces-483	92	28	1977	1977	NUM
alces-483	92	29	,	,	PUNCT
alces-483	92	30	ballard	ballard	PROPN
alces-483	92	31	et	et	PROPN
alces-483	92	32	al	al	PROPN
alces-483	92	33	.	.	PROPN
alces-483	92	34	1987	1987	NUM
alces-483	92	35	,	,	PUNCT
alces-483	92	36	mech	mech	NOUN
alces-483	92	37	et	et	PROPN
alces-483	92	38	al	al	PROPN
alces-483	92	39	.	.	PROPN
alces-483	92	40	1995	1995	NUM
alces-483	92	41	)	)	PUNCT
alces-483	92	42	.	.	PUNCT
alces-483	93	1	page	page	NOUN
alces-483	93	2	’s	’s	PART
alces-483	93	3	(	(	PUNCT
alces-483	93	4	1989	1989	NUM
alces-483	93	5	)	)	PUNCT
alces-483	93	6	analysis	analysis	NOUN
alces-483	93	7	of	of	ADP
alces-483	93	8	cohort	cohort	NOUN
alces-483	93	9	survival	survival	NOUN
alces-483	93	10	indicated	indicate	VERB
alces-483	93	11	that	that	SCONJ
alces-483	93	12	overwinter	overwinter	ADJ
alces-483	93	13	survival	survival	NOUN
alces-483	93	14	of	of	ADP
alces-483	93	15	moose	moose	NOUN
alces-483	93	16	calves	calf	NOUN
alces-483	93	17	ranged	range	VERB
alces-483	93	18	from	from	ADP
alces-483	93	19	only	only	ADV
alces-483	93	20	30	30	NUM
alces-483	93	21	%	%	NOUN
alces-483	93	22	to	to	ADP
alces-483	93	23	almost	almost	ADV
alces-483	93	24	100	100	NUM
alces-483	93	25	%	%	NOUN
alces-483	93	26	,	,	PUNCT
alces-483	93	27	depending	depend	VERB
alces-483	93	28	on	on	ADP
alces-483	93	29	wolf	wolf	PROPN
alces-483	93	30	density	density	NOUN
alces-483	93	31	and	and	CCONJ
alces-483	93	32	relative	relative	ADJ
alces-483	93	33	nutritional	nutritional	ADJ
alces-483	93	34	stress	stress	NOUN
alces-483	93	35	(	(	PUNCT
alces-483	93	36	caused	cause	VERB
alces-483	93	37	by	by	ADP
alces-483	93	38	deep	deep	ADJ
alces-483	93	39	-	-	PUNCT
alces-483	93	40	snow	snow	NOUN
alces-483	93	41	winters	winter	NOUN
alces-483	93	42	and	and	CCONJ
alces-483	93	43	high	high	ADJ
alces-483	93	44	population	population	NOUN
alces-483	93	45	density	density	NOUN
alces-483	93	46	)	)	PUNCT
alces-483	93	47	.	.	PUNCT
alces-483	94	1	predation	predation	NOUN
alces-483	94	2	may	may	AUX
alces-483	94	3	likewise	likewise	ADV
alces-483	94	4	reduce	reduce	VERB
alces-483	94	5	adult	adult	NOUN
alces-483	94	6	survivorship	survivorship	NOUN
alces-483	94	7	(	(	PUNCT
alces-483	94	8	van	van	PROPN
alces-483	94	9	ballenberghe	ballenberghe	NOUN
alces-483	94	10	and	and	CCONJ
alces-483	94	11	ballard	ballard	NOUN
alces-483	94	12	1998	1998	NUM
alces-483	94	13	)	)	PUNCT
alces-483	94	14	,	,	PUNCT
alces-483	94	15	but	but	CCONJ
alces-483	94	16	variance	variance	NOUN
alces-483	94	17	in	in	ADP
alces-483	94	18	adult	adult	NOUN
alces-483	94	19	survival	survival	NOUN
alces-483	94	20	is	be	AUX
alces-483	94	21	much	much	ADV
alces-483	94	22	less	less	ADJ
alces-483	94	23	than	than	ADP
alces-483	94	24	for	for	ADP
alces-483	94	25	juveniles	juvenile	NOUN
alces-483	94	26	.	.	PUNCT
alces-483	95	1	adult	adult	NOUN
alces-483	95	2	survival	survival	NOUN
alces-483	95	3	rates	rate	NOUN
alces-483	95	4	depend	depend	VERB
alces-483	95	5	heavily	heavily	ADV
alces-483	95	6	on	on	ADP
alces-483	95	7	the	the	DET
alces-483	95	8	intensity	intensity	NOUN
alces-483	95	9	of	of	ADP
alces-483	95	10	hunting	hunting	NOUN
alces-483	95	11	.	.	PUNCT
alces-483	96	1	peterson	peterson	NOUN
alces-483	96	2	(	(	PUNCT
alces-483	96	3	1977	1977	NUM
alces-483	96	4	)	)	PUNCT
alces-483	96	5	estimated	estimate	VERB
alces-483	96	6	that	that	SCONJ
alces-483	96	7	equilibrium	equilibrium	NOUN
alces-483	96	8	survival	survival	NOUN
alces-483	96	9	of	of	ADP
alces-483	96	10	an	an	DET
alces-483	96	11	unhunted	unhunted	ADJ
alces-483	96	12	moose	moose	NOUN
alces-483	96	13	population	population	NOUN
alces-483	96	14	was	be	AUX
alces-483	96	15	~87	~87	NOUN
alces-483	96	16	%	%	NOUN
alces-483	96	17	for	for	ADP
alces-483	96	18	yearlings	yearling	NOUN
alces-483	96	19	and	and	CCONJ
alces-483	96	20	adults	adult	NOUN
alces-483	96	21	;	;	PUNCT
alces-483	96	22	gasaway	gasaway	NOUN
alces-483	96	23	et	et	PROPN
alces-483	96	24	al	al	PROPN
alces-483	96	25	.	.	PROPN
alces-483	97	1	(	(	PUNCT
alces-483	97	2	1983	1983	NUM
alces-483	97	3	)	)	PUNCT
alces-483	97	4	reported	report	VERB
alces-483	97	5	that	that	SCONJ
alces-483	97	6	the	the	DET
alces-483	97	7	adult	adult	NOUN
alces-483	97	8	moose	moose	NOUN
alces-483	97	9	survival	survival	NOUN
alces-483	97	10	rate	rate	NOUN
alces-483	97	11	improved	improve	VERB
alces-483	97	12	from	from	ADP
alces-483	97	13	80	80	NUM
alces-483	97	14	%	%	NOUN
alces-483	97	15	to	to	PART
alces-483	97	16	94	94	NUM
alces-483	97	17	%	%	NOUN
alces-483	97	18	after	after	SCONJ
alces-483	97	19	wolf	wolf	NOUN
alces-483	97	20	densities	density	NOUN
alces-483	97	21	were	be	AUX
alces-483	97	22	reduced	reduce	VERB
alces-483	97	23	in	in	ADP
alces-483	97	24	the	the	DET
alces-483	97	25	tanana	tanana	PROPN
alces-483	97	26	flats	flat	NOUN
alces-483	97	27	,	,	PUNCT
alces-483	97	28	alaska	alaska	PROPN
alces-483	98	1	.	.	PUNCT
alces-483	98	2	messier	messy	ADJ
alces-483	98	3	(	(	PUNCT
alces-483	98	4	1994	1994	NUM
alces-483	98	5	)	)	PUNCT
alces-483	98	6	reviewed	review	VERB
alces-483	98	7	27	27	NUM
alces-483	98	8	studies	study	NOUN
alces-483	98	9	in	in	ADP
alces-483	98	10	which	which	PRON
alces-483	98	11	moose	moose	NOUN
alces-483	98	12	were	be	AUX
alces-483	98	13	the	the	DET
alces-483	98	14	primary	primary	ADJ
alces-483	98	15	prey	prey	NOUN
alces-483	98	16	,	,	PUNCT
alces-483	98	17	and	and	CCONJ
alces-483	98	18	from	from	ADP
alces-483	98	19	his	his	PRON
alces-483	98	20	analysis	analysis	NOUN
alces-483	98	21	he	he	PRON
alces-483	98	22	concluded	conclude	VERB
alces-483	98	23	that	that	SCONJ
alces-483	98	24	wolf	wolf	PROPN
alces-483	98	25	predation	predation	NOUN
alces-483	98	26	was	be	AUX
alces-483	98	27	density	density	NOUN
alces-483	98	28	-	-	PUNCT
alces-483	98	29	dependent	dependent	ADJ
alces-483	98	30	at	at	ADP
alces-483	98	31	the	the	DET
alces-483	98	32	low	low	ADJ
alces-483	98	33	range	range	NOUN
alces-483	98	34	of	of	ADP
alces-483	98	35	moose	moose	NOUN
alces-483	98	36	density	density	NOUN
alces-483	98	37	,	,	PUNCT
alces-483	98	38	therefore	therefore	ADV
alces-483	98	39	regulatory	regulatory	ADJ
alces-483	98	40	.	.	PUNCT
alces-483	99	1	he	he	PRON
alces-483	99	2	predicted	predict	VERB
alces-483	99	3	that	that	SCONJ
alces-483	99	4	moose	moose	NOUN
alces-483	99	5	would	would	AUX
alces-483	99	6	stabilize	stabilize	VERB
alces-483	99	7	at	at	ADP
alces-483	99	8	2	2	NUM
alces-483	99	9	moose	moose	NOUN
alces-483	99	10	/	/	SYM
alces-483	99	11	km2	km2	NOUN
alces-483	99	12	in	in	ADP
alces-483	99	13	the	the	DET
alces-483	99	14	absence	absence	NOUN
alces-483	99	15	of	of	ADP
alces-483	99	16	predators	predator	NOUN
alces-483	99	17	via	via	ADP
alces-483	99	18	density	density	NOUN
alces-483	99	19	-	-	PUNCT
alces-483	99	20	dependent	dependent	ADJ
alces-483	99	21	mechanisms	mechanism	NOUN
alces-483	99	22	,	,	PUNCT
alces-483	99	23	at	at	ADP
alces-483	99	24	1.3	1.3	NUM
alces-483	99	25	moose	moose	NOUN
alces-483	99	26	/	/	SYM
alces-483	99	27	km2	km2	NOUN
alces-483	99	28	in	in	ADP
alces-483	99	29	simple	simple	ADJ
alces-483	99	30	wolf	wolf	NOUN
alces-483	99	31	/	/	SYM
alces-483	99	32	moose	moose	NOUN
alces-483	99	33	systems	system	NOUN
alces-483	99	34	,	,	PUNCT
alces-483	99	35	and	and	CCONJ
alces-483	99	36	0.2	0.2	NUM
alces-483	99	37	-	-	PUNCT
alces-483	99	38	0.4	0.4	NUM
alces-483	99	39	moose	moose	NOUN
alces-483	99	40	/	/	SYM
alces-483	99	41	km2	km2	NOUN
alces-483	99	42	in	in	ADP
alces-483	99	43	a	a	DET
alces-483	99	44	single	single	ADJ
alces-483	99	45	stable	stable	ADJ
alces-483	99	46	-	-	PUNCT
alces-483	99	47	state	state	NOUN
alces-483	99	48	(	(	PUNCT
alces-483	99	49	low	low	ADJ
alces-483	99	50	density	density	NOUN
alces-483	99	51	)	)	PUNCT
alces-483	99	52	equilibrium	equilibrium	NOUN
alces-483	99	53	with	with	ADP
alces-483	99	54	multiple	multiple	ADJ
alces-483	99	55	predators	predator	NOUN
alces-483	99	56	.	.	PUNCT
alces-483	100	1	thomas	thomas	PROPN
alces-483	100	2	(	(	PUNCT
alces-483	100	3	1995	1995	NUM
alces-483	100	4	)	)	PUNCT
alces-483	100	5	asserted	assert	VERB
alces-483	100	6	that	that	SCONJ
alces-483	100	7	caribou	caribou	NOUN
alces-483	100	8	have	have	VERB
alces-483	100	9	no	no	DET
alces-483	100	10	intrinsic	intrinsic	ADJ
alces-483	100	11	(	(	PUNCT
alces-483	100	12	e.g.	e.g.	ADV
alces-483	100	13	,	,	PUNCT
alces-483	100	14	social	social	ADJ
alces-483	100	15	)	)	PUNCT
alces-483	100	16	population	population	NOUN
alces-483	100	17	limitations	limitation	NOUN
alces-483	100	18	.	.	PUNCT
alces-483	101	1	in	in	ADP
alces-483	101	2	his	his	PRON
alces-483	101	3	view	view	NOUN
alces-483	101	4	the	the	DET
alces-483	101	5	evolutionary	evolutionary	ADJ
alces-483	101	6	pressure	pressure	NOUN
alces-483	101	7	of	of	ADP
alces-483	101	8	wolf	wolf	PROPN
alces-483	101	9	predation	predation	NOUN
alces-483	101	10	pervades	pervade	VERB
alces-483	101	11	caribou	caribou	PROPN
alces-483	101	12	ecology	ecology	NOUN
alces-483	101	13	.	.	PUNCT
alces-483	102	1	the	the	DET
alces-483	102	2	suite	suite	NOUN
alces-483	102	3	of	of	ADP
alces-483	102	4	predators	predator	NOUN
alces-483	102	5	for	for	ADP
alces-483	102	6	caribou	caribou	NOUN
alces-483	102	7	is	be	AUX
alces-483	102	8	dominated	dominate	VERB
alces-483	102	9	by	by	ADP
alces-483	102	10	wolves	wolf	NOUN
alces-483	102	11	but	but	CCONJ
alces-483	102	12	also	also	ADV
alces-483	102	13	includes	include	VERB
alces-483	102	14	lynx	lynx	NOUN
alces-483	102	15	(	(	PUNCT
alces-483	102	16	lynx	lynx	NOUN
alces-483	102	17	canadensis	canadensis	NOUN
alces-483	102	18	)	)	PUNCT
alces-483	102	19	,	,	PUNCT
alces-483	102	20	coyote	coyote	NOUN
alces-483	102	21	(	(	PUNCT
alces-483	102	22	canis	canis	NOUN
alces-483	102	23	latrans	latran	NOUN
alces-483	102	24	)	)	PUNCT
alces-483	102	25	,	,	PUNCT
alces-483	102	26	brown	brown	ADJ
alces-483	102	27	bear	bear	NOUN
alces-483	102	28	,	,	PUNCT
alces-483	102	29	black	black	ADJ
alces-483	102	30	bear	bear	NOUN
alces-483	102	31	,	,	PUNCT
alces-483	102	32	cougar	cougar	NOUN
alces-483	102	33	(	(	PUNCT
alces-483	102	34	felis	felis	NOUN
alces-483	102	35	concolor	concolor	NOUN
alces-483	102	36	)	)	PUNCT
alces-483	102	37	,	,	PUNCT
alces-483	102	38	and	and	CCONJ
alces-483	102	39	wolverine	wolverine	NOUN
alces-483	102	40	(	(	PUNCT
alces-483	102	41	gulo	gulo	PROPN
alces-483	102	42	gulo	gulo	PROPN
alces-483	102	43	)	)	PUNCT
alces-483	102	44	.	.	PUNCT
alces-483	103	1	he	he	PRON
alces-483	103	2	stated	state	VERB
alces-483	103	3	that	that	SCONJ
alces-483	103	4	predators	predator	NOUN
alces-483	103	5	“	"	PUNCT
alces-483	103	6	can	can	AUX
alces-483	103	7	keep	keep	VERB
alces-483	103	8	caribou	caribou	NOUN
alces-483	103	9	populations	population	NOUN
alces-483	103	10	depressed	depress	VERB
alces-483	103	11	for	for	ADP
alces-483	103	12	long	long	ADJ
alces-483	103	13	periods	period	NOUN
alces-483	103	14	if	if	SCONJ
alces-483	103	15	alternate	alternate	ADJ
alces-483	103	16	prey	prey	NOUN
alces-483	103	17	are	be	AUX
alces-483	103	18	abundant	abundant	ADJ
alces-483	103	19	.	.	PUNCT
alces-483	103	20	”	"	PUNCT
alces-483	104	1	farnell	farnell	PROPN
alces-483	104	2	and	and	CCONJ
alces-483	104	3	mcdonald	mcdonald	PROPN
alces-483	104	4	(	(	PUNCT
alces-483	104	5	1988	1988	NUM
alces-483	104	6	)	)	PUNCT
alces-483	104	7	reported	report	VERB
alces-483	104	8	that	that	SCONJ
alces-483	104	9	recruitment	recruitment	NOUN
alces-483	104	10	of	of	ADP
alces-483	104	11	caribou	caribou	NOUN
alces-483	104	12	increased	increase	VERB
alces-483	104	13	113	113	NUM
alces-483	104	14	%	%	NOUN
alces-483	104	15	and	and	CCONJ
alces-483	104	16	adult	adult	NOUN
alces-483	104	17	mortality	mortality	NOUN
alces-483	104	18	declined	decline	VERB
alces-483	104	19	60	60	NUM
alces-483	104	20	%	%	NOUN
alces-483	104	21	in	in	ADP
alces-483	104	22	a	a	DET
alces-483	104	23	population	population	NOUN
alces-483	104	24	where	where	SCONJ
alces-483	104	25	wolf	wolf	NOUN
alces-483	104	26	numbers	number	NOUN
alces-483	104	27	were	be	AUX
alces-483	104	28	reduced	reduce	VERB
alces-483	104	29	80	80	NUM
alces-483	104	30	%	%	NOUN
alces-483	104	31	.	.	PUNCT
alces-483	105	1	thomas	thomas	PROPN
alces-483	105	2	(	(	PUNCT
alces-483	105	3	1995	1995	NUM
alces-483	105	4	)	)	PUNCT
alces-483	105	5	concluded	conclude	VERB
alces-483	105	6	that	that	SCONJ
alces-483	105	7	when	when	SCONJ
alces-483	105	8	predators	predator	NOUN
alces-483	105	9	are	be	AUX
alces-483	105	10	present	present	ADJ
alces-483	105	11	,	,	PUNCT
alces-483	105	12	average	average	ADJ
alces-483	105	13	natural	natural	ADJ
alces-483	105	14	mortality	mortality	NOUN
alces-483	105	15	of	of	ADP
alces-483	105	16	caribou	caribou	NOUN
alces-483	105	17	is	be	AUX
alces-483	105	18	approximately	approximately	ADV
alces-483	105	19	50	50	NUM
alces-483	105	20	%	%	NOUN
alces-483	105	21	in	in	ADP
alces-483	105	22	calves	calf	NOUN
alces-483	105	23	in	in	ADP
alces-483	105	24	forest	forest	NOUN
alces-483	105	25	-	-	PUNCT
alces-483	105	26	tundra	tundra	NOUN
alces-483	105	27	areas	area	NOUN
alces-483	105	28	and	and	CCONJ
alces-483	105	29	50	50	NUM
alces-483	105	30	-	-	SYM
alces-483	105	31	70	70	NUM
alces-483	105	32	%	%	NOUN
alces-483	105	33	in	in	ADP
alces-483	105	34	forestalpine	forestalpine	NOUN
alces-483	105	35	and	and	CCONJ
alces-483	105	36	forest	forest	NOUN
alces-483	105	37	-	-	PUNCT
alces-483	105	38	forest	forest	NOUN
alces-483	105	39	moose	moose	NOUN
alces-483	105	40	zones	zone	NOUN
alces-483	105	41	;	;	PUNCT
alces-483	105	42	annual	annual	ADJ
alces-483	105	43	mortality	mortality	NOUN
alces-483	105	44	was	be	AUX
alces-483	105	45	7	7	NUM
alces-483	105	46	-	-	SYM
alces-483	105	47	30	30	NUM
alces-483	105	48	%	%	NOUN
alces-483	105	49	in	in	ADP
alces-483	105	50	adults	adult	NOUN
alces-483	105	51	(	(	PUNCT
alces-483	105	52	bergerud	bergerud	NOUN
alces-483	105	53	1980	1980	NUM
alces-483	105	54	,	,	PUNCT
alces-483	105	55	farnell	farnell	PROPN
alces-483	105	56	and	and	CCONJ
alces-483	105	57	mcdonald	mcdonald	PROPN
alces-483	105	58	1988	1988	NUM
alces-483	105	59	,	,	PUNCT
alces-483	105	60	seip	seip	NOUN
alces-483	105	61	1992	1992	NUM
alces-483	105	62	)	)	PUNCT
alces-483	105	63	.	.	PUNCT
alces-483	106	1	in	in	ADP
alces-483	106	2	contrast	contrast	NOUN
alces-483	106	3	,	,	PUNCT
alces-483	106	4	annual	annual	ADJ
alces-483	106	5	mortality	mortality	NOUN
alces-483	106	6	approaches	approach	VERB
alces-483	106	7	zero	zero	NUM
alces-483	106	8	where	where	SCONJ
alces-483	106	9	wolves	wolf	NOUN
alces-483	106	10	are	be	AUX
alces-483	106	11	absent	absent	ADJ
alces-483	106	12	or	or	CCONJ
alces-483	106	13	rare	rare	ADJ
alces-483	106	14	.	.	PUNCT
alces-483	107	1	from	from	ADP
alces-483	107	2	his	his	PRON
alces-483	107	3	review	review	NOUN
alces-483	107	4	of	of	ADP
alces-483	107	5	the	the	DET
alces-483	107	6	literature	literature	NOUN
alces-483	107	7	,	,	PUNCT
alces-483	107	8	ranging	range	VERB
alces-483	107	9	from	from	ADP
alces-483	107	10	kelsall	kelsall	PROPN
alces-483	107	11	(	(	PUNCT
alces-483	107	12	1968	1968	NUM
alces-483	107	13	)	)	PUNCT
alces-483	107	14	to	to	ADP
alces-483	107	15	seip	seip	PROPN
alces-483	107	16	(	(	PUNCT
alces-483	107	17	1992	1992	NUM
alces-483	107	18	)	)	PUNCT
alces-483	107	19	,	,	PUNCT
alces-483	107	20	thomas	thomas	PROPN
alces-483	107	21	(	(	PUNCT
alces-483	107	22	1995	1995	NUM
alces-483	107	23	)	)	PUNCT
alces-483	107	24	concluded	conclude	VERB
alces-483	107	25	that	that	SCONJ
alces-483	107	26	(	(	PUNCT
alces-483	107	27	1	1	X
alces-483	107	28	)	)	PUNCT
alces-483	107	29	wolf	wolf	PROPN
alces-483	107	30	predation	predation	NOUN
alces-483	107	31	is	be	AUX
alces-483	107	32	the	the	DET
alces-483	107	33	major	major	ADJ
alces-483	107	34	direct	direct	ADJ
alces-483	107	35	cause	cause	NOUN
alces-483	107	36	of	of	ADP
alces-483	107	37	natural	natural	ADJ
alces-483	107	38	mortality	mortality	NOUN
alces-483	107	39	of	of	ADP
alces-483	107	40	calf	calf	NOUN
alces-483	107	41	and	and	CCONJ
alces-483	107	42	adult	adult	NOUN
alces-483	107	43	caribou	caribou	NOUN
alces-483	107	44	and	and	CCONJ
alces-483	107	45	(	(	PUNCT
alces-483	107	46	2	2	X
alces-483	107	47	)	)	PUNCT
alces-483	107	48	dense	dense	ADJ
alces-483	107	49	caribou	caribou	NOUN
alces-483	107	50	populations	population	NOUN
alces-483	107	51	occur	occur	VERB
alces-483	107	52	only	only	ADV
alces-483	107	53	in	in	ADP
alces-483	107	54	the	the	DET
alces-483	107	55	absence	absence	NOUN
alces-483	107	56	of	of	ADP
alces-483	107	57	wolves	wolf	NOUN
alces-483	107	58	.	.	PUNCT
alces-483	108	1	from	from	ADP
alces-483	108	2	studies	study	NOUN
alces-483	108	3	of	of	ADP
alces-483	108	4	deer	deer	NOUN
alces-483	108	5	in	in	ADP
alces-483	108	6	northern	northern	ADJ
alces-483	108	7	minnesota	minnesota	PROPN
alces-483	108	8	,	,	PUNCT
alces-483	108	9	mech	mech	NOUN
alces-483	108	10	et	et	PROPN
alces-483	108	11	al	al	PROPN
alces-483	108	12	.	.	PROPN
alces-483	109	1	(	(	PUNCT
alces-483	109	2	1987	1987	NUM
alces-483	109	3	)	)	PUNCT
alces-483	109	4	showed	show	VERB
alces-483	109	5	that	that	SCONJ
alces-483	109	6	deer	deer	NOUN
alces-483	109	7	fawn	fawn	NOUN
alces-483	109	8	abundance	abundance	NOUN
alces-483	109	9	was	be	AUX
alces-483	109	10	correlated	correlate	VERB
alces-483	109	11	not	not	PART
alces-483	109	12	with	with	ADP
alces-483	109	13	space	space	NOUN
alces-483	109	14	and	and	CCONJ
alces-483	109	15	time	time	NOUN
alces-483	109	16	in	in	ADP
alces-483	109	17	predation	predation	NOUN
alces-483	109	18	dynamics	dynamic	NOUN
alces-483	109	19	–	–	PUNCT
alces-483	109	20	peterson	peterson	PROPN
alces-483	109	21	et	et	PROPN
alces-483	109	22	al	al	PROPN
alces-483	109	23	.	.	PROPN
alces-483	109	24	alces	alce	NOUN
alces-483	109	25	vol	vol	NOUN
alces-483	109	26	.	.	PROPN
alces-483	109	27	39	39	NUM
alces-483	109	28	,	,	PUNCT
alces-483	109	29	2003	2003	NUM
alces-483	109	30	220	220	NUM
alces-483	109	31	wolf	wolf	NOUN
alces-483	109	32	density	density	NOUN
alces-483	109	33	but	but	CCONJ
alces-483	109	34	rather	rather	ADV
alces-483	109	35	the	the	DET
alces-483	109	36	cumulative	cumulative	ADJ
alces-483	109	37	severity	severity	NOUN
alces-483	109	38	of	of	ADP
alces-483	109	39	3	3	NUM
alces-483	109	40	previous	previous	ADJ
alces-483	109	41	winters	winter	NOUN
alces-483	109	42	.	.	PUNCT
alces-483	110	1	this	this	PRON
alces-483	110	2	does	do	AUX
alces-483	110	3	not	not	PART
alces-483	110	4	indicate	indicate	VERB
alces-483	110	5	that	that	SCONJ
alces-483	110	6	wolves	wolf	NOUN
alces-483	110	7	were	be	AUX
alces-483	110	8	unimportant	unimportant	ADJ
alces-483	110	9	in	in	ADP
alces-483	110	10	deer	deer	NOUN
alces-483	110	11	dynamics	dynamic	NOUN
alces-483	110	12	(	(	PUNCT
alces-483	110	13	cf	cf	NOUN
alces-483	110	14	.	.	PUNCT
alces-483	110	15	mech	mech	PROPN
alces-483	110	16	and	and	CCONJ
alces-483	110	17	karns	karn	NOUN
alces-483	110	18	1977	1977	NUM
alces-483	110	19	)	)	PUNCT
alces-483	110	20	,	,	PUNCT
alces-483	110	21	but	but	CCONJ
alces-483	110	22	might	might	AUX
alces-483	110	23	instead	instead	ADV
alces-483	110	24	mean	mean	VERB
alces-483	110	25	that	that	SCONJ
alces-483	110	26	the	the	DET
alces-483	110	27	dominant	dominant	ADJ
alces-483	110	28	variance	variance	NOUN
alces-483	110	29	in	in	ADP
alces-483	110	30	the	the	DET
alces-483	110	31	system	system	NOUN
alces-483	110	32	was	be	AUX
alces-483	110	33	winter	winter	NOUN
alces-483	110	34	severity	severity	NOUN
alces-483	110	35	(	(	PUNCT
alces-483	110	36	boyce	boyce	PROPN
alces-483	110	37	and	and	CCONJ
alces-483	110	38	anderson	anderson	PROPN
alces-483	110	39	1999	1999	NUM
alces-483	110	40	)	)	PUNCT
alces-483	110	41	.	.	PUNCT
alces-483	111	1	further	further	PROPN
alces-483	111	2	south	south	ADV
alces-483	111	3	in	in	ADP
alces-483	111	4	minnesota	minnesota	PROPN
alces-483	111	5	,	,	PUNCT
alces-483	111	6	where	where	SCONJ
alces-483	111	7	timber	timber	NOUN
alces-483	111	8	harvest	harvest	NOUN
alces-483	111	9	created	create	VERB
alces-483	111	10	optimum	optimum	ADJ
alces-483	111	11	deer	deer	NOUN
alces-483	111	12	habitat	habitat	NOUN
alces-483	111	13	,	,	PUNCT
alces-483	111	14	fuller	full	ADJ
alces-483	111	15	(	(	PUNCT
alces-483	111	16	1989	1989	NUM
alces-483	111	17	)	)	PUNCT
alces-483	111	18	found	find	VERB
alces-483	111	19	that	that	SCONJ
alces-483	111	20	hunting	hunt	VERB
alces-483	111	21	mortality	mortality	NOUN
alces-483	111	22	was	be	AUX
alces-483	111	23	more	more	ADV
alces-483	111	24	important	important	ADJ
alces-483	111	25	than	than	ADP
alces-483	111	26	wolf	wolf	PROPN
alces-483	111	27	predation	predation	NOUN
alces-483	111	28	in	in	ADP
alces-483	111	29	deer	deer	NOUN
alces-483	111	30	dynamics	dynamic	NOUN
alces-483	111	31	.	.	PUNCT
alces-483	112	1	resource	resource	NOUN
alces-483	112	2	abundance	abundance	NOUN
alces-483	112	3	may	may	AUX
alces-483	112	4	indeed	indeed	ADV
alces-483	112	5	modify	modify	VERB
alces-483	112	6	the	the	DET
alces-483	112	7	effect	effect	NOUN
alces-483	112	8	of	of	ADP
alces-483	112	9	predation	predation	NOUN
alces-483	112	10	through	through	ADP
alces-483	112	11	effects	effect	NOUN
alces-483	112	12	on	on	ADP
alces-483	112	13	reproduction	reproduction	NOUN
alces-483	112	14	and	and	CCONJ
alces-483	112	15	individual	individual	ADJ
alces-483	112	16	vigor	vigor	NOUN
alces-483	112	17	.	.	PUNCT
alces-483	113	1	wolves	wolf	NOUN
alces-483	113	2	and	and	CCONJ
alces-483	113	3	moose	moose	NOUN
alces-483	113	4	on	on	ADP
alces-483	113	5	isle	isle	PROPN
alces-483	113	6	royale	royale	PROPN
alces-483	113	7	:	:	PUNCT
alces-483	113	8	dynamics	dynamic	NOUN
alces-483	113	9	of	of	ADP
alces-483	113	10	a	a	DET
alces-483	113	11	simple	simple	ADJ
alces-483	113	12	system	system	NOUN
alces-483	113	13	a	a	DET
alces-483	113	14	perspective	perspective	NOUN
alces-483	113	15	on	on	ADP
alces-483	113	16	wolf	wolf	NOUN
alces-483	113	17	-	-	PUNCT
alces-483	113	18	moose	moose	NOUN
alces-483	113	19	relations	relation	NOUN
alces-483	113	20	in	in	ADP
alces-483	113	21	isle	isle	PROPN
alces-483	113	22	royale	royale	PROPN
alces-483	113	23	national	national	PROPN
alces-483	113	24	park	park	PROPN
alces-483	113	25	(	(	PUNCT
alces-483	113	26	544	544	NUM
alces-483	113	27	-	-	PUNCT
alces-483	113	28	km2	km2	NOUN
alces-483	113	29	island	island	NOUN
alces-483	113	30	in	in	ADP
alces-483	113	31	lake	lake	PROPN
alces-483	113	32	superior	superior	PROPN
alces-483	113	33	)	)	PUNCT
alces-483	113	34	is	be	AUX
alces-483	113	35	offered	offer	VERB
alces-483	113	36	,	,	PUNCT
alces-483	113	37	to	to	PART
alces-483	113	38	provide	provide	VERB
alces-483	113	39	an	an	DET
alces-483	113	40	update	update	NOUN
alces-483	113	41	on	on	ADP
alces-483	113	42	a	a	DET
alces-483	113	43	very	very	ADV
alces-483	113	44	dynamic	dynamic	ADJ
alces-483	113	45	case	case	NOUN
alces-483	113	46	history	history	NOUN
alces-483	113	47	and	and	CCONJ
alces-483	113	48	to	to	PART
alces-483	113	49	illustrate	illustrate	VERB
alces-483	113	50	temporal	temporal	ADJ
alces-483	113	51	and	and	CCONJ
alces-483	113	52	spatial	spatial	ADJ
alces-483	113	53	issues	issue	NOUN
alces-483	113	54	involved	involve	VERB
alces-483	113	55	in	in	ADP
alces-483	113	56	interpreting	interpret	VERB
alces-483	113	57	predator	predator	NOUN
alces-483	113	58	-	-	PUNCT
alces-483	113	59	prey	prey	NOUN
alces-483	113	60	interactions	interaction	NOUN
alces-483	113	61	.	.	PUNCT
alces-483	114	1	as	as	SCONJ
alces-483	114	2	moose	moose	NOUN
alces-483	114	3	and	and	CCONJ
alces-483	114	4	wolf	wolf	NOUN
alces-483	114	5	populations	population	NOUN
alces-483	114	6	have	have	AUX
alces-483	114	7	changed	change	VERB
alces-483	114	8	over	over	ADP
alces-483	114	9	the	the	DET
alces-483	114	10	past	past	ADJ
alces-483	114	11	40	40	NUM
alces-483	114	12	years	year	NOUN
alces-483	114	13	(	(	PUNCT
alces-483	114	14	fig	fig	NOUN
alces-483	114	15	.	.	NOUN
alces-483	115	1	2	2	X
alces-483	115	2	)	)	PUNCT
alces-483	115	3	there	there	PRON
alces-483	115	4	have	have	AUX
alces-483	115	5	been	be	AUX
alces-483	115	6	significant	significant	ADJ
alces-483	115	7	changes	change	NOUN
alces-483	115	8	in	in	ADP
alces-483	115	9	how	how	SCONJ
alces-483	115	10	the	the	DET
alces-483	115	11	system	system	NOUN
alces-483	115	12	has	have	AUX
alces-483	115	13	been	be	AUX
alces-483	115	14	interpreted	interpret	VERB
alces-483	115	15	by	by	ADP
alces-483	115	16	observers	observer	NOUN
alces-483	115	17	,	,	PUNCT
alces-483	115	18	and	and	CCONJ
alces-483	115	19	valuable	valuable	ADJ
alces-483	115	20	perspective	perspective	NOUN
alces-483	115	21	has	have	AUX
alces-483	115	22	also	also	ADV
alces-483	115	23	been	be	AUX
alces-483	115	24	provided	provide	VERB
alces-483	115	25	by	by	ADP
alces-483	115	26	studies	study	NOUN
alces-483	115	27	elsewhere	elsewhere	ADV
alces-483	115	28	.	.	PUNCT
alces-483	116	1	annual	annual	ADJ
alces-483	116	2	winter	winter	NOUN
alces-483	116	3	counts	count	NOUN
alces-483	116	4	of	of	ADP
alces-483	116	5	wolves	wolf	NOUN
alces-483	116	6	began	begin	VERB
alces-483	116	7	at	at	ADP
alces-483	116	8	isle	isle	PROPN
alces-483	116	9	royale	royale	PROPN
alces-483	116	10	about	about	ADV
alces-483	116	11	10	10	NUM
alces-483	116	12	years	year	NOUN
alces-483	116	13	after	after	SCONJ
alces-483	116	14	the	the	DET
alces-483	116	15	island	island	NOUN
alces-483	116	16	was	be	AUX
alces-483	116	17	colonized	colonize	VERB
alces-483	116	18	by	by	ADP
alces-483	116	19	wolves	wolf	NOUN
alces-483	116	20	in	in	ADP
alces-483	116	21	the	the	DET
alces-483	116	22	late	late	ADJ
alces-483	116	23	1940s	1940	NOUN
alces-483	116	24	.	.	PUNCT
alces-483	117	1	efforts	effort	NOUN
alces-483	117	2	were	be	AUX
alces-483	117	3	begun	begin	VERB
alces-483	117	4	to	to	PART
alces-483	117	5	estimate	estimate	VERB
alces-483	117	6	moose	moose	NOUN
alces-483	117	7	numbers	number	NOUN
alces-483	117	8	,	,	PUNCT
alces-483	117	9	with	with	ADP
alces-483	117	10	methods	method	NOUN
alces-483	117	11	steadily	steadily	ADV
alces-483	117	12	evolving	evolve	VERB
alces-483	117	13	into	into	ADP
alces-483	117	14	a	a	DET
alces-483	117	15	“	"	PUNCT
alces-483	117	16	gasaway	gasaway	NOUN
alces-483	117	17	-	-	PUNCT
alces-483	117	18	type	type	NOUN
alces-483	117	19	”	"	PUNCT
alces-483	117	20	stratified	stratify	VERB
alces-483	117	21	plot	plot	NOUN
alces-483	117	22	count	count	NOUN
alces-483	117	23	in	in	ADP
alces-483	117	24	which	which	PRON
alces-483	117	25	about	about	ADP
alces-483	117	26	17	17	NUM
alces-483	117	27	%	%	NOUN
alces-483	117	28	of	of	ADP
alces-483	117	29	the	the	DET
alces-483	117	30	land	land	NOUN
alces-483	117	31	area	area	NOUN
alces-483	117	32	is	be	AUX
alces-483	117	33	intensively	intensively	ADV
alces-483	117	34	searched	search	VERB
alces-483	117	35	from	from	ADP
alces-483	117	36	aircraft	aircraft	NOUN
alces-483	117	37	in	in	ADP
alces-483	117	38	winter	winter	NOUN
alces-483	117	39	.	.	PUNCT
alces-483	118	1	an	an	DET
alces-483	118	2	independent	independent	ADJ
alces-483	118	3	method	method	NOUN
alces-483	118	4	to	to	PART
alces-483	118	5	track	track	VERB
alces-483	118	6	historic	historic	ADJ
alces-483	118	7	change	change	NOUN
alces-483	118	8	in	in	ADP
alces-483	118	9	moose	moose	NOUN
alces-483	118	10	numbers	number	NOUN
alces-483	118	11	has	have	AUX
alces-483	118	12	been	be	AUX
alces-483	118	13	retrospective	retrospective	ADJ
alces-483	118	14	reconstruction	reconstruction	NOUN
alces-483	118	15	,	,	PUNCT
alces-483	118	16	based	base	VERB
alces-483	118	17	on	on	ADP
alces-483	118	18	recoveries	recovery	NOUN
alces-483	118	19	of	of	ADP
alces-483	118	20	approximately	approximately	ADV
alces-483	118	21	one	one	NUM
alces-483	118	22	-	-	PUNCT
alces-483	118	23	third	third	NOUN
alces-483	118	24	of	of	ADP
alces-483	118	25	the	the	DET
alces-483	118	26	moose	moose	NOUN
alces-483	118	27	after	after	ADP
alces-483	118	28	death	death	NOUN
alces-483	118	29	(	(	PUNCT
alces-483	118	30	page	page	NOUN
alces-483	118	31	1989	1989	NUM
alces-483	118	32	)	)	PUNCT
alces-483	118	33	.	.	PUNCT
alces-483	119	1	temporal	temporal	ADJ
alces-483	119	2	chronology	chronology	NOUN
alces-483	119	3	early	early	ADV
alces-483	119	4	in	in	ADP
alces-483	119	5	the	the	DET
alces-483	119	6	study	study	NOUN
alces-483	119	7	,	,	PUNCT
alces-483	119	8	in	in	ADP
alces-483	119	9	the	the	DET
alces-483	119	10	early	early	ADJ
alces-483	119	11	1960s	1960	NOUN
alces-483	119	12	,	,	PUNCT
alces-483	119	13	wolf	wolf	PROPN
alces-483	119	14	population	population	NOUN
alces-483	119	15	size	size	NOUN
alces-483	119	16	was	be	AUX
alces-483	119	17	stable	stable	ADJ
alces-483	119	18	and	and	CCONJ
alces-483	119	19	moose	moose	NOUN
alces-483	119	20	exhibited	exhibit	VERB
alces-483	119	21	a	a	DET
alces-483	119	22	high	high	ADJ
alces-483	119	23	twinning	twin	VERB
alces-483	119	24	rate	rate	NOUN
alces-483	119	25	and	and	CCONJ
alces-483	119	26	also	also	ADV
alces-483	119	27	seemed	seem	VERB
alces-483	119	28	relatively	relatively	ADV
alces-483	119	29	stable	stable	ADJ
alces-483	119	30	,	,	PUNCT
alces-483	119	31	so	so	ADV
alces-483	119	32	mech	mech	NOUN
alces-483	119	33	(	(	PUNCT
alces-483	119	34	1966	1966	NUM
alces-483	119	35	)	)	PUNCT
alces-483	119	36	suggested	suggest	VERB
alces-483	119	37	that	that	SCONJ
alces-483	119	38	wolf	wolf	PROPN
alces-483	119	39	predation	predation	NOUN
alces-483	119	40	was	be	AUX
alces-483	119	41	keeping	keep	VERB
alces-483	119	42	moose	moose	NOUN
alces-483	119	43	density	density	NOUN
alces-483	119	44	(	(	PUNCT
alces-483	119	45	about	about	ADV
alces-483	119	46	1	1	NUM
alces-483	119	47	/	/	SYM
alces-483	119	48	km2	km2	NOUN
alces-483	119	49	)	)	PUNCT
alces-483	119	50	below	below	ADP
alces-483	119	51	the	the	DET
alces-483	119	52	level	level	NOUN
alces-483	119	53	at	at	ADP
alces-483	119	54	which	which	PRON
alces-483	119	55	food	food	NOUN
alces-483	119	56	supply	supply	NOUN
alces-483	119	57	might	might	AUX
alces-483	119	58	be	be	AUX
alces-483	119	59	limiting	limit	VERB
alces-483	119	60	.	.	PUNCT
alces-483	120	1	by	by	ADP
alces-483	120	2	the	the	DET
alces-483	120	3	early	early	ADJ
alces-483	120	4	1970s	1970	NOUN
alces-483	120	5	,	,	PUNCT
alces-483	120	6	however	however	ADV
alces-483	120	7	,	,	PUNCT
alces-483	120	8	it	it	PRON
alces-483	120	9	was	be	AUX
alces-483	120	10	evident	evident	ADJ
alces-483	120	11	that	that	SCONJ
alces-483	120	12	moose	moose	NOUN
alces-483	120	13	had	have	AUX
alces-483	120	14	increased	increase	VERB
alces-483	120	15	in	in	ADP
alces-483	120	16	the	the	DET
alces-483	120	17	1960s	1960	NOUN
alces-483	120	18	(	(	PUNCT
alces-483	120	19	krefting	krefte	VERB
alces-483	120	20	1974	1974	NUM
alces-483	120	21	,	,	PUNCT
alces-483	120	22	peterson	peterson	NOUN
alces-483	120	23	1977	1977	NUM
alces-483	120	24	)	)	PUNCT
alces-483	120	25	,	,	PUNCT
alces-483	120	26	reaching	reach	VERB
alces-483	120	27	a	a	DET
alces-483	120	28	level	level	NOUN
alces-483	120	29	(	(	PUNCT
alces-483	120	30	almost	almost	ADV
alces-483	120	31	3	3	NUM
alces-483	120	32	/	/	SYM
alces-483	120	33	km2	km2	NOUN
alces-483	120	34	)	)	PUNCT
alces-483	120	35	in	in	ADP
alces-483	120	36	the	the	DET
alces-483	120	37	early	early	ADJ
alces-483	120	38	1970s	1970	NOUN
alces-483	120	39	where	where	SCONJ
alces-483	120	40	nutrition	nutrition	NOUN
alces-483	120	41	was	be	AUX
alces-483	120	42	poor	poor	ADJ
alces-483	120	43	,	,	PUNCT
alces-483	120	44	at	at	ADP
alces-483	120	45	least	least	ADJ
alces-483	120	46	during	during	ADP
alces-483	120	47	severe	severe	ADJ
alces-483	120	48	winters	winter	NOUN
alces-483	120	49	in	in	ADP
alces-483	120	50	1969	1969	NUM
alces-483	120	51	-	-	SYM
alces-483	120	52	1972	1972	NUM
alces-483	120	53	(	(	PUNCT
alces-483	120	54	peterson	peterson	NOUN
alces-483	120	55	1975	1975	NUM
alces-483	120	56	)	)	PUNCT
alces-483	120	57	.	.	PUNCT
alces-483	121	1	the	the	DET
alces-483	121	2	wolf	wolf	PROPN
alces-483	121	3	population	population	NOUN
alces-483	121	4	expanded	expand	VERB
alces-483	121	5	during	during	ADP
alces-483	121	6	1969	1969	NUM
alces-483	121	7	-	-	SYM
alces-483	121	8	1980	1980	NUM
alces-483	121	9	and	and	CCONJ
alces-483	121	10	moose	moose	NOUN
alces-483	121	11	density	density	NOUN
alces-483	121	12	declined	decline	VERB
alces-483	121	13	in	in	ADP
alces-483	121	14	1972	1972	NUM
alces-483	121	15	-	-	SYM
alces-483	121	16	1982	1982	NUM
alces-483	121	17	as	as	SCONJ
alces-483	121	18	wolf	wolf	PROPN
alces-483	121	19	predation	predation	NOUN
alces-483	121	20	intensified	intensify	VERB
alces-483	121	21	.	.	PUNCT
alces-483	122	1	wolves	wolf	NOUN
alces-483	122	2	briefly	briefly	ADV
alces-483	122	3	reached	reach	VERB
alces-483	122	4	in	in	ADP
alces-483	122	5	1980	1980	NUM
alces-483	122	6	the	the	DET
alces-483	122	7	highest	high	ADJ
alces-483	122	8	year	year	NOUN
alces-483	122	9	-	-	PUNCT
alces-483	122	10	round	round	NOUN
alces-483	122	11	density	density	NOUN
alces-483	122	12	documented	document	VERB
alces-483	122	13	for	for	ADP
alces-483	122	14	wolves	wolf	NOUN
alces-483	122	15	in	in	ADP
alces-483	122	16	nature	nature	NOUN
alces-483	122	17	up	up	ADP
alces-483	122	18	to	to	ADP
alces-483	122	19	that	that	DET
alces-483	122	20	time	time	NOUN
alces-483	122	21	(	(	PUNCT
alces-483	122	22	almost	almost	ADV
alces-483	122	23	0.1/	0.1/	PUNCT
alces-483	122	24	km2	km2	NOUN
alces-483	122	25	)	)	PUNCT
alces-483	122	26	.	.	PUNCT
alces-483	123	1	as	as	SCONJ
alces-483	123	2	wolf	wolf	PROPN
alces-483	123	3	numbers	number	NOUN
alces-483	123	4	grew	grow	VERB
alces-483	123	5	it	it	PRON
alces-483	123	6	was	be	AUX
alces-483	123	7	obvious	obvious	ADJ
alces-483	123	8	that	that	SCONJ
alces-483	123	9	calf	calf	NOUN
alces-483	123	10	survival	survival	NOUN
alces-483	123	11	was	be	AUX
alces-483	123	12	negatively	negatively	ADV
alces-483	123	13	affected	affect	VERB
alces-483	123	14	(	(	PUNCT
alces-483	123	15	fig	fig	NOUN
alces-483	123	16	.	.	PUNCT
alces-483	124	1	2	2	NUM
alces-483	124	2	)	)	PUNCT
alces-483	124	3	,	,	PUNCT
alces-483	124	4	prompting	prompt	VERB
alces-483	124	5	population	population	NOUN
alces-483	124	6	decline	decline	NOUN
alces-483	124	7	.	.	PUNCT
alces-483	125	1	taking	take	VERB
alces-483	125	2	stock	stock	NOUN
alces-483	125	3	of	of	ADP
alces-483	125	4	the	the	DET
alces-483	125	5	situation	situation	NOUN
alces-483	125	6	in	in	ADP
alces-483	125	7	the	the	DET
alces-483	125	8	mid-1970s	mid-1970	NOUN
alces-483	125	9	,	,	PUNCT
alces-483	125	10	peterson	peterson	NOUN
alces-483	125	11	(	(	PUNCT
alces-483	125	12	1977	1977	NUM
alces-483	125	13	)	)	PUNCT
alces-483	125	14	interpreted	interpret	VERB
alces-483	125	15	the	the	DET
alces-483	125	16	moose	moose	NOUN
alces-483	125	17	decline	decline	NOUN
alces-483	125	18	as	as	ADP
alces-483	125	19	a	a	DET
alces-483	125	20	response	response	NOUN
alces-483	125	21	to	to	ADP
alces-483	125	22	habitat	habitat	NOUN
alces-483	125	23	deterioration	deterioration	NOUN
alces-483	125	24	as	as	SCONJ
alces-483	125	25	post	post	ADJ
alces-483	125	26	-	-	ADJ
alces-483	125	27	fire	fire	ADJ
alces-483	125	28	successional	successional	ADJ
alces-483	125	29	forests	forest	NOUN
alces-483	125	30	(	(	PUNCT
alces-483	125	31	regenerating	regenerate	VERB
alces-483	125	32	after	after	ADP
alces-483	125	33	fires	fire	NOUN
alces-483	125	34	in	in	ADP
alces-483	125	35	1936	1936	NUM
alces-483	125	36	and	and	CCONJ
alces-483	125	37	1948	1948	NUM
alces-483	125	38	)	)	PUNCT
alces-483	125	39	matured	mature	VERB
alces-483	125	40	and	and	CCONJ
alces-483	125	41	moose	moose	NOUN
alces-483	125	42	became	become	VERB
alces-483	125	43	more	more	ADV
alces-483	125	44	dependent	dependent	ADJ
alces-483	125	45	on	on	ADP
alces-483	125	46	older	old	ADJ
alces-483	125	47	forests	forest	NOUN
alces-483	125	48	over	over	ADP
alces-483	125	49	100	100	NUM
alces-483	125	50	years	year	NOUN
alces-483	125	51	old	old	ADJ
alces-483	125	52	.	.	PUNCT
alces-483	126	1	peterson	peterson	NOUN
alces-483	126	2	(	(	PUNCT
alces-483	126	3	1977	1977	NUM
alces-483	126	4	)	)	PUNCT
alces-483	126	5	asserted	assert	VERB
alces-483	126	6	that	that	SCONJ
alces-483	126	7	density	density	NOUN
alces-483	126	8	-	-	PUNCT
alces-483	126	9	dependent	dependent	ADJ
alces-483	126	10	mechanisms	mechanism	NOUN
alces-483	126	11	prompted	prompt	VERB
alces-483	126	12	the	the	DET
alces-483	126	13	moose	moose	NOUN
alces-483	126	14	decline	decline	NOUN
alces-483	126	15	,	,	PUNCT
alces-483	126	16	although	although	SCONJ
alces-483	126	17	wolf	wolf	PROPN
alces-483	126	18	predation	predation	NOUN
alces-483	126	19	probably	probably	ADV
alces-483	126	20	accelerated	accelerate	VERB
alces-483	126	21	it	it	PRON
alces-483	126	22	.	.	PUNCT
alces-483	127	1	in	in	ADP
alces-483	127	2	short	short	ADJ
alces-483	127	3	,	,	PUNCT
alces-483	127	4	wolf	wolf	PROPN
alces-483	127	5	predation	predation	NOUN
alces-483	127	6	was	be	AUX
alces-483	127	7	considered	consider	VERB
alces-483	127	8	largely	largely	ADV
alces-483	127	9	compensatory	compensatory	ADJ
alces-483	127	10	,	,	PUNCT
alces-483	127	11	and	and	CCONJ
alces-483	127	12	not	not	PART
alces-483	127	13	ultimately	ultimately	ADV
alces-483	127	14	responsible	responsible	ADJ
alces-483	127	15	for	for	ADP
alces-483	127	16	the	the	DET
alces-483	127	17	moose	moose	NOUN
alces-483	127	18	decline	decline	NOUN
alces-483	127	19	.	.	PUNCT
alces-483	128	1	this	this	PRON
alces-483	128	2	was	be	AUX
alces-483	128	3	essentially	essentially	ADV
alces-483	128	4	a	a	DET
alces-483	128	5	bottom	bottom	ADJ
alces-483	128	6	-	-	PUNCT
alces-483	128	7	up	up	ADP
alces-483	128	8	interpretation	interpretation	NOUN
alces-483	128	9	,	,	PUNCT
alces-483	128	10	with	with	ADP
alces-483	128	11	the	the	DET
alces-483	128	12	logical	logical	ADJ
alces-483	128	13	prediction	prediction	NOUN
alces-483	128	14	being	be	AUX
alces-483	128	15	that	that	DET
alces-483	128	16	moose	moose	NOUN
alces-483	128	17	density	density	NOUN
alces-483	128	18	would	would	AUX
alces-483	128	19	stabilize	stabilize	VERB
alces-483	128	20	at	at	ADP
alces-483	128	21	a	a	DET
alces-483	128	22	new	new	ADJ
alces-483	128	23	,	,	PUNCT
alces-483	128	24	lower	low	ADJ
alces-483	128	25	level	level	NOUN
alces-483	128	26	dictated	dictate	VERB
alces-483	128	27	by	by	ADP
alces-483	128	28	habitat	habitat	NOUN
alces-483	128	29	,	,	PUNCT
alces-483	128	30	where	where	SCONJ
alces-483	128	31	it	it	PRON
alces-483	128	32	would	would	AUX
alces-483	128	33	remain	remain	VERB
alces-483	128	34	as	as	ADV
alces-483	128	35	long	long	ADV
alces-483	128	36	as	as	SCONJ
alces-483	128	37	new	new	ADJ
alces-483	128	38	forest	forest	NOUN
alces-483	128	39	disturbance	disturbance	NOUN
alces-483	128	40	did	do	AUX
alces-483	128	41	not	not	PART
alces-483	128	42	intervene	intervene	VERB
alces-483	128	43	.	.	PUNCT
alces-483	129	1	however	however	ADV
alces-483	129	2	,	,	PUNCT
alces-483	129	3	the	the	DET
alces-483	129	4	moose	moose	NOUN
alces-483	129	5	population	population	NOUN
alces-483	129	6	quickly	quickly	ADV
alces-483	129	7	grew	grow	VERB
alces-483	129	8	again	again	ADV
alces-483	129	9	after	after	SCONJ
alces-483	129	10	the	the	DET
alces-483	129	11	wolf	wolf	PROPN
alces-483	129	12	population	population	NOUN
alces-483	129	13	crashed	crash	VERB
alces-483	129	14	in	in	ADP
alces-483	129	15	1980	1980	NUM
alces-483	129	16	-	-	SYM
alces-483	129	17	1982	1982	NUM
alces-483	129	18	(	(	PUNCT
alces-483	129	19	coincident	coincident	NOUN
alces-483	129	20	with	with	ADP
alces-483	129	21	the	the	DET
alces-483	129	22	arrival	arrival	NOUN
alces-483	129	23	of	of	ADP
alces-483	129	24	canine	canine	NOUN
alces-483	129	25	parvovirus	parvovirus	NOUN
alces-483	129	26	;	;	PUNCT
alces-483	129	27	peterson	peterson	PROPN
alces-483	129	28	et	et	PROPN
alces-483	129	29	al	al	PROPN
alces-483	129	30	.	.	PROPN
alces-483	129	31	1998	1998	NUM
alces-483	129	32	)	)	PUNCT
alces-483	129	33	.	.	PUNCT
alces-483	130	1	the	the	DET
alces-483	130	2	1981	1981	NUM
alces-483	130	3	cohort	cohort	NOUN
alces-483	130	4	of	of	ADP
alces-483	130	5	moose	moose	NOUN
alces-483	130	6	calves	calf	NOUN
alces-483	130	7	,	,	PUNCT
alces-483	130	8	coincident	coincident	NOUN
alces-483	130	9	with	with	ADP
alces-483	130	10	the	the	DET
alces-483	130	11	wolf	wolf	NOUN
alces-483	130	12	decline	decline	NOUN
alces-483	130	13	,	,	PUNCT
alces-483	130	14	was	be	AUX
alces-483	130	15	proportionately	proportionately	ADV
alces-483	130	16	among	among	ADP
alces-483	130	17	the	the	DET
alces-483	130	18	largest	large	ADJ
alces-483	130	19	ever	ever	ADV
alces-483	130	20	seen	see	VERB
alces-483	130	21	at	at	ADP
alces-483	130	22	isle	isle	PROPN
alces-483	130	23	alces	alces	PROPN
alces-483	130	24	vol	vol	NOUN
alces-483	130	25	.	.	PROPN
alces-483	130	26	39	39	NUM
alces-483	130	27	,	,	PUNCT
alces-483	130	28	2003	2003	NUM
alces-483	130	29	peterson	peterson	PROPN
alces-483	130	30	et	et	PROPN
alces-483	130	31	al	al	PROPN
alces-483	130	32	.	.	PROPN
alces-483	130	33	–	–	PUNCT
alces-483	130	34	space	space	NOUN
alces-483	130	35	and	and	CCONJ
alces-483	130	36	time	time	NOUN
alces-483	130	37	in	in	ADP
alces-483	130	38	predation	predation	NOUN
alces-483	130	39	dynamics	dynamic	NOUN
alces-483	130	40	221	221	NUM
alces-483	130	41	royale	royale	NOUN
alces-483	130	42	,	,	PUNCT
alces-483	130	43	constituting	constitute	VERB
alces-483	130	44	about	about	ADP
alces-483	130	45	one	one	NUM
alces-483	130	46	-	-	PUNCT
alces-483	130	47	fifth	fifth	NOUN
alces-483	130	48	of	of	ADP
alces-483	130	49	the	the	DET
alces-483	130	50	moose	moose	NOUN
alces-483	130	51	population	population	NOUN
alces-483	130	52	in	in	ADP
alces-483	130	53	mid	mid	NOUN
alces-483	130	54	-	-	NOUN
alces-483	130	55	winter	winter	ADJ
alces-483	130	56	(	(	PUNCT
alces-483	130	57	fig	fig	NOUN
alces-483	130	58	.	.	PUNCT
alces-483	131	1	2	2	NUM
alces-483	131	2	)	)	PUNCT
alces-483	131	3	.	.	PUNCT
alces-483	132	1	peterson	peterson	NOUN
alces-483	132	2	and	and	CCONJ
alces-483	132	3	page	page	NOUN
alces-483	132	4	(	(	PUNCT
alces-483	132	5	1983	1983	NUM
alces-483	132	6	)	)	PUNCT
alces-483	132	7	acknowledged	acknowledge	VERB
alces-483	132	8	that	that	SCONJ
alces-483	132	9	wolf	wolf	PROPN
alces-483	132	10	predation	predation	NOUN
alces-483	132	11	,	,	PUNCT
alces-483	132	12	not	not	PART
alces-483	132	13	deficient	deficient	ADJ
alces-483	132	14	habitat	habitat	NOUN
alces-483	132	15	,	,	PUNCT
alces-483	132	16	had	have	AUX
alces-483	132	17	been	be	AUX
alces-483	132	18	limiting	limit	VERB
alces-483	132	19	moose	moose	NOUN
alces-483	132	20	population	population	NOUN
alces-483	132	21	growth	growth	NOUN
alces-483	132	22	–	–	PUNCT
alces-483	132	23	evidently	evidently	ADV
alces-483	132	24	predation	predation	NOUN
alces-483	132	25	loss	loss	NOUN
alces-483	132	26	was	be	AUX
alces-483	132	27	not	not	PART
alces-483	132	28	simply	simply	ADV
alces-483	132	29	compensatory	compensatory	ADJ
alces-483	132	30	.	.	PUNCT
alces-483	133	1	significantly	significantly	ADV
alces-483	133	2	,	,	PUNCT
alces-483	133	3	balsam	balsam	NOUN
alces-483	133	4	fir	fir	NOUN
alces-483	133	5	trees	tree	NOUN
alces-483	133	6	throughout	throughout	ADP
alces-483	133	7	the	the	DET
alces-483	133	8	winter	winter	NOUN
alces-483	133	9	range	range	NOUN
alces-483	133	10	of	of	ADP
alces-483	133	11	isle	isle	PROPN
alces-483	133	12	royale	royale	PROPN
alces-483	133	13	moose	moose	PROPN
alces-483	133	14	exhibited	exhibit	VERB
alces-483	133	15	lagged	lag	VERB
alces-483	133	16	oscillations	oscillation	NOUN
alces-483	133	17	in	in	ADP
alces-483	133	18	growth	growth	NOUN
alces-483	133	19	that	that	PRON
alces-483	133	20	mirrored	mirror	VERB
alces-483	133	21	the	the	DET
alces-483	133	22	inverse	inverse	NOUN
alces-483	133	23	of	of	ADP
alces-483	133	24	moose	moose	NOUN
alces-483	133	25	density	density	NOUN
alces-483	133	26	–	–	PUNCT
alces-483	133	27	when	when	SCONJ
alces-483	133	28	wolves	wolf	NOUN
alces-483	133	29	were	be	AUX
alces-483	133	30	high	high	ADJ
alces-483	133	31	,	,	PUNCT
alces-483	133	32	moose	moose	NOUN
alces-483	133	33	declined	decline	VERB
alces-483	133	34	,	,	PUNCT
alces-483	133	35	prompting	prompt	VERB
alces-483	133	36	growth	growth	NOUN
alces-483	133	37	of	of	ADP
alces-483	133	38	the	the	DET
alces-483	133	39	forest	forest	NOUN
alces-483	133	40	(	(	PUNCT
alces-483	133	41	mclaren	mclaren	PROPN
alces-483	133	42	and	and	CCONJ
alces-483	133	43	peterson	peterson	PROPN
alces-483	133	44	1994	1994	NUM
alces-483	133	45	)	)	PUNCT
alces-483	133	46	.	.	PUNCT
alces-483	134	1	throughout	throughout	ADP
alces-483	134	2	the	the	DET
alces-483	134	3	1980s	1980	NOUN
alces-483	134	4	and	and	CCONJ
alces-483	134	5	early	early	ADJ
alces-483	134	6	1990s	1990	NOUN
alces-483	134	7	,	,	PUNCT
alces-483	134	8	with	with	ADP
alces-483	134	9	low	low	ADJ
alces-483	134	10	wolf	wolf	NOUN
alces-483	134	11	numbers	number	NOUN
alces-483	134	12	,	,	PUNCT
alces-483	134	13	the	the	DET
alces-483	134	14	moose	moose	NOUN
alces-483	134	15	population	population	NOUN
alces-483	134	16	grew	grow	VERB
alces-483	134	17	almost	almost	ADV
alces-483	134	18	without	without	ADP
alces-483	134	19	interruption	interruption	NOUN
alces-483	134	20	(	(	PUNCT
alces-483	134	21	winter	winter	NOUN
alces-483	134	22	ticks	tick	NOUN
alces-483	134	23	were	be	AUX
alces-483	134	24	implicated	implicate	VERB
alces-483	134	25	in	in	ADP
alces-483	134	26	high	high	ADJ
alces-483	134	27	mortality	mortality	NOUN
alces-483	134	28	in	in	ADP
alces-483	134	29	1989	1989	NUM
alces-483	134	30	;	;	PUNCT
alces-483	134	31	delgiudice	delgiudice	NOUN
alces-483	134	32	et	et	PROPN
alces-483	134	33	al	al	PROPN
alces-483	134	34	.	.	PROPN
alces-483	134	35	1997	1997	NUM
alces-483	134	36	)	)	PUNCT
alces-483	134	37	.	.	PUNCT
alces-483	135	1	wolves	wolf	NOUN
alces-483	135	2	were	be	AUX
alces-483	135	3	themselves	themselves	PRON
alces-483	135	4	mysteriously	mysteriously	ADV
alces-483	135	5	maintained	maintain	VERB
alces-483	135	6	at	at	ADP
alces-483	135	7	a	a	DET
alces-483	135	8	low	low	ADJ
alces-483	135	9	level	level	NOUN
alces-483	135	10	by	by	ADP
alces-483	135	11	lingering	linger	VERB
alces-483	135	12	effects	effect	NOUN
alces-483	135	13	of	of	ADP
alces-483	135	14	disease	disease	NOUN
alces-483	135	15	,	,	PUNCT
alces-483	135	16	inbreeding	inbreede	VERB
alces-483	135	17	,	,	PUNCT
alces-483	135	18	stochastic	stochastic	ADJ
alces-483	135	19	demography	demography	NOUN
alces-483	135	20	,	,	PUNCT
alces-483	135	21	or	or	CCONJ
alces-483	135	22	some	some	DET
alces-483	135	23	combination	combination	NOUN
alces-483	135	24	of	of	ADP
alces-483	135	25	these	these	DET
alces-483	135	26	factors	factor	NOUN
alces-483	135	27	(	(	PUNCT
alces-483	135	28	peterson	peterson	PROPN
alces-483	135	29	et	et	PROPN
alces-483	135	30	al	al	PROPN
alces-483	135	31	.	.	PROPN
alces-483	135	32	1998	1998	NUM
alces-483	135	33	,	,	PUNCT
alces-483	135	34	peterson	peterson	NOUN
alces-483	135	35	1999	1999	NUM
alces-483	135	36	)	)	PUNCT
alces-483	135	37	.	.	PUNCT
alces-483	136	1	by	by	ADP
alces-483	136	2	1995	1995	NUM
alces-483	136	3	the	the	DET
alces-483	136	4	moose	moose	NOUN
alces-483	136	5	population	population	NOUN
alces-483	136	6	had	have	AUX
alces-483	136	7	grown	grow	VERB
alces-483	136	8	to	to	PART
alces-483	136	9	exceed	exceed	VERB
alces-483	136	10	4/	4/	NUM
alces-483	136	11	km2	km2	NOUN
alces-483	136	12	and	and	CCONJ
alces-483	136	13	there	there	PRON
alces-483	136	14	was	be	VERB
alces-483	136	15	ample	ample	ADJ
alces-483	136	16	evidence	evidence	NOUN
alces-483	136	17	of	of	ADP
alces-483	136	18	severe	severe	ADJ
alces-483	136	19	undernutrition	undernutrition	NOUN
alces-483	136	20	in	in	ADP
alces-483	136	21	winter	winter	NOUN
alces-483	136	22	.	.	PUNCT
alces-483	137	1	twin	twin	ADJ
alces-483	137	2	calves	calf	NOUN
alces-483	137	3	were	be	AUX
alces-483	137	4	rarely	rarely	ADV
alces-483	137	5	seen	see	VERB
alces-483	137	6	,	,	PUNCT
alces-483	137	7	and	and	CCONJ
alces-483	137	8	moose	moose	NOUN
alces-483	137	9	phenotype	phenotype	NOUN
alces-483	137	10	reflected	reflect	VERB
alces-483	137	11	food	food	NOUN
alces-483	137	12	shortage	shortage	NOUN
alces-483	137	13	(	(	PUNCT
alces-483	137	14	peterson	peterson	NOUN
alces-483	137	15	1995	1995	NUM
alces-483	137	16	)	)	PUNCT
alces-483	137	17	.	.	PUNCT
alces-483	138	1	although	although	SCONJ
alces-483	138	2	calves	calf	NOUN
alces-483	138	3	were	be	AUX
alces-483	138	4	growth	growth	NOUN
alces-483	138	5	-	-	PUNCT
alces-483	138	6	retarded	retard	VERB
alces-483	138	7	,	,	PUNCT
alces-483	138	8	most	most	ADV
alces-483	138	9	nevertheless	nevertheless	ADV
alces-483	138	10	survived	survive	VERB
alces-483	138	11	their	their	PRON
alces-483	138	12	first	first	ADJ
alces-483	138	13	winter	winter	NOUN
alces-483	138	14	to	to	PART
alces-483	138	15	live	live	VERB
alces-483	138	16	on	on	ADP
alces-483	138	17	as	as	ADP
alces-483	138	18	adults	adult	NOUN
alces-483	138	19	;	;	PUNCT
alces-483	138	20	density	density	NOUN
alces-483	138	21	-	-	PUNCT
alces-483	138	22	dependence	dependence	NOUN
alces-483	138	23	was	be	AUX
alces-483	138	24	reflected	reflect	VERB
alces-483	138	25	in	in	ADP
alces-483	138	26	moose	moose	NOUN
alces-483	138	27	morphology	morphology	NOUN
alces-483	138	28	but	but	CCONJ
alces-483	138	29	not	not	PART
alces-483	138	30	in	in	ADP
alces-483	138	31	population	population	NOUN
alces-483	138	32	dynamics	dynamic	NOUN
alces-483	138	33	.	.	PUNCT
alces-483	139	1	the	the	DET
alces-483	139	2	winter	winter	NOUN
alces-483	139	3	of	of	ADP
alces-483	139	4	1995	1995	NUM
alces-483	139	5	-	-	SYM
alces-483	139	6	1996	1996	NUM
alces-483	139	7	was	be	AUX
alces-483	139	8	the	the	DET
alces-483	139	9	most	most	ADV
alces-483	139	10	extreme	extreme	ADJ
alces-483	139	11	in	in	ADP
alces-483	139	12	a	a	DET
alces-483	139	13	century	century	NOUN
alces-483	139	14	(	(	PUNCT
alces-483	139	15	delgiudice	delgiudice	NOUN
alces-483	139	16	1998	1998	NUM
alces-483	139	17	,	,	PUNCT
alces-483	139	18	post	post	VERB
alces-483	139	19	et	et	PROPN
alces-483	139	20	al	al	PROPN
alces-483	139	21	.	.	PROPN
alces-483	139	22	1999	1999	NUM
alces-483	139	23	)	)	PUNCT
alces-483	139	24	,	,	PUNCT
alces-483	139	25	with	with	ADP
alces-483	139	26	early	early	ADJ
alces-483	139	27	winter	winter	NOUN
alces-483	139	28	storms	storm	NOUN
alces-483	139	29	,	,	PUNCT
alces-483	139	30	persistent	persistent	ADJ
alces-483	139	31	deep	deep	ADJ
alces-483	139	32	snowcover	snowcover	NOUN
alces-483	139	33	,	,	PUNCT
alces-483	139	34	and	and	CCONJ
alces-483	139	35	cold	cold	ADJ
alces-483	139	36	temperatures	temperature	NOUN
alces-483	139	37	that	that	PRON
alces-483	139	38	delayed	delay	VERB
alces-483	139	39	the	the	DET
alces-483	139	40	arrival	arrival	NOUN
alces-483	139	41	of	of	ADP
alces-483	139	42	spring	spring	NOUN
alces-483	139	43	by	by	ADP
alces-483	139	44	about	about	ADV
alces-483	139	45	6	6	NUM
alces-483	139	46	weeks	week	NOUN
alces-483	139	47	.	.	PUNCT
alces-483	140	1	moose	moose	PROPN
alces-483	140	2	began	begin	VERB
alces-483	140	3	dying	die	VERB
alces-483	140	4	of	of	ADP
alces-483	140	5	starvation	starvation	NOUN
alces-483	140	6	by	by	ADP
alces-483	140	7	february	february	PROPN
alces-483	140	8	1996	1996	NUM
alces-483	140	9	and	and	CCONJ
alces-483	140	10	about	about	ADV
alces-483	140	11	80	80	NUM
alces-483	140	12	%	%	NOUN
alces-483	140	13	of	of	ADP
alces-483	140	14	the	the	DET
alces-483	140	15	population	population	NOUN
alces-483	140	16	perished	perish	VERB
alces-483	140	17	in	in	ADP
alces-483	140	18	the	the	DET
alces-483	140	19	next	next	ADJ
alces-483	140	20	3	3	NUM
alces-483	140	21	months	month	NOUN
alces-483	140	22	,	,	PUNCT
alces-483	140	23	reducing	reduce	VERB
alces-483	140	24	moose	moose	NOUN
alces-483	140	25	density	density	NOUN
alces-483	140	26	once	once	ADV
alces-483	140	27	again	again	ADV
alces-483	140	28	to	to	ADP
alces-483	140	29	about	about	ADP
alces-483	140	30	1	1	NUM
alces-483	140	31	/	/	SYM
alces-483	140	32	km2	km2	NOUN
alces-483	140	33	.	.	PUNCT
alces-483	141	1	a	a	DET
alces-483	141	2	dieoff	dieoff	NOUN
alces-483	141	3	of	of	ADP
alces-483	141	4	this	this	DET
alces-483	141	5	scale	scale	NOUN
alces-483	141	6	also	also	ADV
alces-483	141	7	happened	happen	VERB
alces-483	141	8	on	on	ADP
alces-483	141	9	isle	isle	PROPN
alces-483	141	10	royale	royale	PROPN
alces-483	141	11	in	in	ADP
alces-483	141	12	1934	1934	NUM
alces-483	141	13	,	,	PUNCT
alces-483	141	14	following	follow	VERB
alces-483	141	15	the	the	DET
alces-483	141	16	initial	initial	ADJ
alces-483	141	17	irruption	irruption	NOUN
alces-483	141	18	of	of	ADP
alces-483	141	19	moose	moose	NOUN
alces-483	141	20	after	after	ADP
alces-483	141	21	colonization	colonization	NOUN
alces-483	141	22	(	(	PUNCT
alces-483	141	23	mech	mech	NOUN
alces-483	141	24	1966	1966	NUM
alces-483	141	25	)	)	PUNCT
alces-483	141	26	.	.	PUNCT
alces-483	142	1	spatial	spatial	ADJ
alces-483	142	2	heterogeneity	heterogeneity	NOUN
alces-483	142	3	while	while	SCONJ
alces-483	142	4	temporal	temporal	ADJ
alces-483	142	5	variation	variation	NOUN
alces-483	142	6	in	in	ADP
alces-483	142	7	the	the	DET
alces-483	142	8	isle	isle	PROPN
alces-483	142	9	royale	royale	PROPN
alces-483	142	10	chronology	chronology	NOUN
alces-483	142	11	is	be	AUX
alces-483	142	12	striking	striking	ADJ
alces-483	142	13	,	,	PUNCT
alces-483	142	14	there	there	PRON
alces-483	142	15	is	be	VERB
alces-483	142	16	also	also	ADV
alces-483	142	17	spatial	spatial	ADJ
alces-483	142	18	heterogeneity	heterogeneity	NOUN
alces-483	142	19	in	in	ADP
alces-483	142	20	this	this	DET
alces-483	142	21	ecosystem	ecosystem	NOUN
alces-483	142	22	.	.	PUNCT
alces-483	143	1	the	the	DET
alces-483	143	2	east	east	ADJ
alces-483	143	3	end	end	NOUN
alces-483	143	4	of	of	ADP
alces-483	143	5	the	the	DET
alces-483	143	6	island	island	NOUN
alces-483	143	7	,	,	PUNCT
alces-483	143	8	by	by	ADP
alces-483	143	9	virtue	virtue	NOUN
alces-483	143	10	of	of	ADP
alces-483	143	11	its	its	PRON
alces-483	143	12	glacial	glacial	ADJ
alces-483	143	13	history	history	NOUN
alces-483	143	14	,	,	PUNCT
alces-483	143	15	has	have	VERB
alces-483	143	16	thin	thin	ADJ
alces-483	143	17	soils	soil	NOUN
alces-483	143	18	and	and	CCONJ
alces-483	143	19	more	more	ADJ
alces-483	143	20	forest	forest	NOUN
alces-483	143	21	disturbance	disturbance	NOUN
alces-483	143	22	caused	cause	VERB
alces-483	143	23	by	by	ADP
alces-483	143	24	wind	wind	NOUN
alces-483	143	25	,	,	PUNCT
alces-483	143	26	resulting	result	VERB
alces-483	143	27	in	in	ADP
alces-483	143	28	more	more	ADJ
alces-483	143	29	light	light	NOUN
alces-483	143	30	reaching	reach	VERB
alces-483	143	31	the	the	DET
alces-483	143	32	forest	forest	NOUN
alces-483	143	33	floor	floor	NOUN
alces-483	143	34	(	(	PUNCT
alces-483	143	35	mclaren	mclaren	PROPN
alces-483	143	36	and	and	CCONJ
alces-483	143	37	janke	janke	PROPN
alces-483	143	38	1996	1996	NUM
alces-483	143	39	)	)	PUNCT
alces-483	143	40	.	.	PUNCT
alces-483	144	1	the	the	DET
alces-483	144	2	resulting	result	VERB
alces-483	144	3	higher	high	ADJ
alces-483	144	4	production	production	NOUN
alces-483	144	5	of	of	ADP
alces-483	144	6	moose	moose	NOUN
alces-483	144	7	forage	forage	NOUN
alces-483	144	8	at	at	ADP
alces-483	144	9	the	the	DET
alces-483	144	10	east	east	ADJ
alces-483	144	11	end	end	NOUN
alces-483	144	12	contrasts	contrast	VERB
alces-483	144	13	with	with	ADP
alces-483	144	14	conditions	condition	NOUN
alces-483	144	15	at	at	ADP
alces-483	144	16	the	the	DET
alces-483	144	17	west	west	PROPN
alces-483	144	18	end	end	NOUN
alces-483	144	19	,	,	PUNCT
alces-483	144	20	where	where	SCONJ
alces-483	144	21	deep	deep	ADJ
alces-483	144	22	soils	soil	NOUN
alces-483	144	23	support	support	VERB
alces-483	144	24	old	old	ADJ
alces-483	144	25	and	and	CCONJ
alces-483	144	26	tall	tall	ADJ
alces-483	144	27	deciduous	deciduous	ADJ
alces-483	144	28	forests	forest	NOUN
alces-483	144	29	that	that	PRON
alces-483	144	30	heavily	heavily	ADV
alces-483	144	31	shade	shade	VERB
alces-483	144	32	the	the	DET
alces-483	144	33	forest	forest	NOUN
alces-483	144	34	floor	floor	NOUN
alces-483	144	35	(	(	PUNCT
alces-483	144	36	fig	fig	NOUN
alces-483	144	37	.	.	PUNCT
alces-483	145	1	3	3	NUM
alces-483	145	2	)	)	PUNCT
alces-483	145	3	.	.	PUNCT
alces-483	146	1	balsam	balsam	NOUN
alces-483	146	2	fir	fir	PROPN
alces-483	146	3	(	(	PUNCT
alces-483	146	4	abies	abie	NOUN
alces-483	146	5	balsamea	balsamea	VERB
alces-483	146	6	)	)	PUNCT
alces-483	146	7	,	,	PUNCT
alces-483	146	8	a	a	DET
alces-483	146	9	key	key	ADJ
alces-483	146	10	winter	winter	NOUN
alces-483	146	11	forage	forage	NOUN
alces-483	146	12	species	specie	NOUN
alces-483	146	13	for	for	ADP
alces-483	146	14	moose	moose	NOUN
alces-483	146	15	fig	fig	NOUN
alces-483	146	16	.	.	PUNCT
alces-483	147	1	2	2	X
alces-483	147	2	.	.	NOUN
alces-483	147	3	wolf	wolf	PROPN
alces-483	147	4	population	population	NOUN
alces-483	147	5	size	size	NOUN
alces-483	147	6	and	and	CCONJ
alces-483	147	7	moose	moose	NOUN
alces-483	147	8	estimated	estimate	VERB
alces-483	147	9	population	population	NOUN
alces-483	147	10	size	size	NOUN
alces-483	147	11	,	,	PUNCT
alces-483	147	12	isle	isle	PROPN
alces-483	147	13	royale	royale	PROPN
alces-483	147	14	national	national	PROPN
alces-483	147	15	park	park	PROPN
alces-483	147	16	,	,	PUNCT
alces-483	147	17	1959	1959	NUM
alces-483	147	18	-	-	SYM
alces-483	147	19	1998	1998	NUM
alces-483	147	20	(	(	PUNCT
alces-483	147	21	upper	upper	ADJ
alces-483	147	22	panel	panel	NOUN
alces-483	147	23	)	)	PUNCT
alces-483	147	24	,	,	PUNCT
alces-483	147	25	and	and	CCONJ
alces-483	147	26	the	the	DET
alces-483	147	27	proportion	proportion	NOUN
alces-483	147	28	of	of	ADP
alces-483	147	29	calves	calf	NOUN
alces-483	147	30	(	(	PUNCT
alces-483	147	31	~6	~6	ADJ
alces-483	147	32	months	month	NOUN
alces-483	147	33	of	of	ADP
alces-483	147	34	age	age	NOUN
alces-483	147	35	)	)	PUNCT
alces-483	147	36	in	in	ADP
alces-483	147	37	the	the	DET
alces-483	147	38	moose	moose	NOUN
alces-483	147	39	population	population	NOUN
alces-483	147	40	(	(	PUNCT
alces-483	147	41	lower	low	ADJ
alces-483	147	42	panel	panel	NOUN
alces-483	147	43	)	)	PUNCT
alces-483	147	44	.	.	PUNCT
alces-483	148	1	each	each	DET
alces-483	148	2	annual	annual	ADJ
alces-483	148	3	estimate	estimate	NOUN
alces-483	148	4	of	of	ADP
alces-483	148	5	%	%	NOUN
alces-483	148	6	calves	calf	NOUN
alces-483	148	7	is	be	AUX
alces-483	148	8	an	an	DET
alces-483	148	9	average	average	NOUN
alces-483	148	10	of	of	ADP
alces-483	148	11	a	a	DET
alces-483	148	12	field	field	NOUN
alces-483	148	13	-	-	PUNCT
alces-483	148	14	based	base	VERB
alces-483	148	15	estimate	estimate	NOUN
alces-483	148	16	(	(	PUNCT
alces-483	148	17	mean	mean	NOUN
alces-483	148	18	of	of	ADP
alces-483	148	19	all	all	PRON
alces-483	148	20	available	available	ADJ
alces-483	148	21	counts	count	NOUN
alces-483	148	22	for	for	ADP
alces-483	148	23	each	each	DET
alces-483	148	24	cohort	cohort	NOUN
alces-483	148	25	,	,	PUNCT
alces-483	148	26	ranging	range	VERB
alces-483	148	27	from	from	ADP
alces-483	148	28	summer	summer	NOUN
alces-483	148	29	ground	ground	NOUN
alces-483	148	30	counts	count	NOUN
alces-483	148	31	to	to	ADP
alces-483	148	32	aerial	aerial	ADJ
alces-483	148	33	counts	count	NOUN
alces-483	148	34	in	in	ADP
alces-483	148	35	autumn	autumn	NOUN
alces-483	148	36	or	or	CCONJ
alces-483	148	37	winter	winter	NOUN
alces-483	148	38	)	)	PUNCT
alces-483	148	39	and	and	CCONJ
alces-483	148	40	an	an	DET
alces-483	148	41	estimate	estimate	NOUN
alces-483	148	42	based	base	VERB
alces-483	148	43	on	on	ADP
alces-483	148	44	population	population	NOUN
alces-483	148	45	reconstruction	reconstruction	NOUN
alces-483	148	46	(	(	PUNCT
alces-483	148	47	details	detail	NOUN
alces-483	148	48	at	at	ADP
alces-483	148	49	www.isleroyalewolf.org	www.isleroyalewolf.org	NOUN
alces-483	148	50	)	)	PUNCT
alces-483	148	51	.	.	PUNCT
alces-483	149	1	space	space	NOUN
alces-483	149	2	and	and	CCONJ
alces-483	149	3	time	time	NOUN
alces-483	149	4	in	in	ADP
alces-483	149	5	predation	predation	NOUN
alces-483	149	6	dynamics	dynamic	NOUN
alces-483	149	7	–	–	PUNCT
alces-483	149	8	peterson	peterson	PROPN
alces-483	149	9	et	et	PROPN
alces-483	149	10	al	al	PROPN
alces-483	149	11	.	.	PROPN
alces-483	150	1	alces	alce	NOUN
alces-483	150	2	vol	vol	NOUN
alces-483	150	3	.	.	PROPN
alces-483	150	4	39	39	NUM
alces-483	150	5	,	,	PUNCT
alces-483	150	6	2003	2003	NUM
alces-483	150	7	222	222	NUM
alces-483	150	8	(	(	PUNCT
alces-483	150	9	risenhoover	risenhoover	NOUN
alces-483	150	10	1987	1987	NUM
alces-483	150	11	)	)	PUNCT
alces-483	150	12	is	be	AUX
alces-483	150	13	regenerating	regenerate	VERB
alces-483	150	14	at	at	ADP
alces-483	150	15	high	high	ADJ
alces-483	150	16	density	density	NOUN
alces-483	150	17	at	at	ADP
alces-483	150	18	the	the	DET
alces-483	150	19	east	east	ADJ
alces-483	150	20	end	end	NOUN
alces-483	150	21	but	but	CCONJ
alces-483	150	22	not	not	PART
alces-483	150	23	at	at	ADP
alces-483	150	24	the	the	DET
alces-483	150	25	west	west	PROPN
alces-483	150	26	end	end	NOUN
alces-483	150	27	.	.	PUNCT
alces-483	151	1	somewhat	somewhat	ADV
alces-483	151	2	paradoxically	paradoxically	ADV
alces-483	151	3	,	,	PUNCT
alces-483	151	4	this	this	PRON
alces-483	151	5	browse	browse	NOUN
alces-483	151	6	species	specie	NOUN
alces-483	151	7	receives	receive	VERB
alces-483	151	8	proportionately	proportionately	ADV
alces-483	151	9	less	less	ADJ
alces-483	151	10	damage	damage	NOUN
alces-483	151	11	by	by	ADP
alces-483	151	12	moose	moose	NOUN
alces-483	151	13	herbivory	herbivory	NOUN
alces-483	151	14	in	in	ADP
alces-483	151	15	the	the	DET
alces-483	151	16	thin	thin	ADJ
alces-483	151	17	soils	soil	NOUN
alces-483	151	18	at	at	ADP
alces-483	151	19	the	the	DET
alces-483	151	20	east	east	ADJ
alces-483	151	21	end	end	NOUN
alces-483	151	22	of	of	ADP
alces-483	151	23	the	the	DET
alces-483	151	24	island	island	NOUN
alces-483	151	25	(	(	PUNCT
alces-483	151	26	mclaren	mclaren	PROPN
alces-483	151	27	1996	1996	NUM
alces-483	151	28	)	)	PUNCT
alces-483	151	29	.	.	PUNCT
alces-483	152	1	we	we	PRON
alces-483	152	2	used	use	VERB
alces-483	152	3	dendrochronology	dendrochronology	NOUN
alces-483	152	4	to	to	PART
alces-483	152	5	determine	determine	VERB
alces-483	152	6	the	the	DET
alces-483	152	7	pattern	pattern	NOUN
alces-483	152	8	of	of	ADP
alces-483	152	9	fir	fir	NOUN
alces-483	152	10	growth	growth	NOUN
alces-483	152	11	as	as	ADP
alces-483	152	12	a	a	DET
alces-483	152	13	key	key	ADJ
alces-483	152	14	component	component	NOUN
alces-483	152	15	of	of	ADP
alces-483	152	16	the	the	DET
alces-483	152	17	isle	isle	PROPN
alces-483	152	18	royale	royale	PROPN
alces-483	152	19	trophic	trophic	PROPN
alces-483	152	20	system	system	NOUN
alces-483	152	21	.	.	PUNCT
alces-483	153	1	fir	fir	NOUN
alces-483	153	2	saplings	sapling	NOUN
alces-483	153	3	and	and	CCONJ
alces-483	153	4	small	small	ADJ
alces-483	153	5	trees	tree	NOUN
alces-483	153	6	growing	grow	VERB
alces-483	153	7	with	with	ADP
alces-483	153	8	minimum	minimum	ADJ
alces-483	153	9	competition	competition	NOUN
alces-483	153	10	for	for	ADP
alces-483	153	11	light	light	NOUN
alces-483	153	12	were	be	AUX
alces-483	153	13	sampled	sample	VERB
alces-483	153	14	from	from	ADP
alces-483	153	15	the	the	DET
alces-483	153	16	west	west	NOUN
alces-483	153	17	and	and	CCONJ
alces-483	153	18	east	east	NOUN
alces-483	153	19	ends	end	NOUN
alces-483	153	20	of	of	ADP
alces-483	153	21	the	the	DET
alces-483	153	22	island	island	NOUN
alces-483	153	23	.	.	PUNCT
alces-483	154	1	for	for	ADP
alces-483	154	2	each	each	DET
alces-483	154	3	individual	individual	ADJ
alces-483	154	4	stem	stem	NOUN
alces-483	154	5	,	,	PUNCT
alces-483	154	6	ring	ring	NOUN
alces-483	154	7	widths	width	NOUN
alces-483	154	8	were	be	AUX
alces-483	154	9	measured	measure	VERB
alces-483	154	10	in	in	ADP
alces-483	154	11	cross	cross	NOUN
alces-483	154	12	-	-	NOUN
alces-483	154	13	sections	section	NOUN
alces-483	154	14	from	from	ADP
alces-483	154	15	the	the	DET
alces-483	154	16	base	base	NOUN
alces-483	154	17	of	of	ADP
alces-483	154	18	the	the	DET
alces-483	154	19	stem	stem	NOUN
alces-483	154	20	.	.	PUNCT
alces-483	155	1	each	each	DET
alces-483	155	2	ring	ring	NOUN
alces-483	155	3	-	-	PUNCT
alces-483	155	4	width	width	NOUN
alces-483	155	5	series	series	NOUN
alces-483	155	6	was	be	AUX
alces-483	155	7	indexed	index	VERB
alces-483	155	8	following	follow	VERB
alces-483	155	9	the	the	DET
alces-483	155	10	method	method	NOUN
alces-483	155	11	described	describe	VERB
alces-483	155	12	in	in	ADP
alces-483	155	13	chouinard	chouinard	NOUN
alces-483	155	14	and	and	CCONJ
alces-483	155	15	filion	filion	NOUN
alces-483	155	16	(	(	PUNCT
alces-483	155	17	2001	2001	NUM
alces-483	155	18	)	)	PUNCT
alces-483	155	19	.	.	PUNCT
alces-483	156	1	heights	height	NOUN
alces-483	156	2	of	of	ADP
alces-483	156	3	fir	fir	NOUN
alces-483	156	4	stems	stem	VERB
alces-483	156	5	on	on	ADP
alces-483	156	6	the	the	DET
alces-483	156	7	west	west	PROPN
alces-483	156	8	end	end	NOUN
alces-483	156	9	ranged	range	VERB
alces-483	156	10	from	from	ADP
alces-483	156	11	90200	90200	NUM
alces-483	156	12	cm	cm	NOUN
alces-483	156	13	,	,	PUNCT
alces-483	156	14	while	while	SCONJ
alces-483	156	15	on	on	ADP
alces-483	156	16	the	the	DET
alces-483	156	17	east	east	PROPN
alces-483	156	18	end	end	NOUN
alces-483	156	19	heights	height	NOUN
alces-483	156	20	were	be	AUX
alces-483	156	21	200	200	NUM
alces-483	156	22	-	-	SYM
alces-483	156	23	600	600	NUM
alces-483	156	24	cm	cm	NOUN
alces-483	156	25	.	.	PUNCT
alces-483	157	1	all	all	DET
alces-483	157	2	trees	tree	NOUN
alces-483	157	3	we	we	PRON
alces-483	157	4	sampled	sample	VERB
alces-483	157	5	were	be	AUX
alces-483	157	6	from	from	ADP
alces-483	157	7	sites	site	NOUN
alces-483	157	8	previously	previously	ADV
alces-483	157	9	studied	study	VERB
alces-483	157	10	by	by	ADP
alces-483	157	11	mclaren	mclaren	PROPN
alces-483	157	12	and	and	CCONJ
alces-483	157	13	peterson	peterson	PROPN
alces-483	157	14	(	(	PUNCT
alces-483	157	15	1994	1994	NUM
alces-483	157	16	)	)	PUNCT
alces-483	157	17	.	.	PUNCT
alces-483	158	1	tree	tree	NOUN
alces-483	158	2	-	-	PUNCT
alces-483	158	3	ring	ring	NOUN
alces-483	158	4	growth	growth	NOUN
alces-483	158	5	,	,	PUNCT
alces-483	158	6	which	which	PRON
alces-483	158	7	may	may	AUX
alces-483	158	8	index	index	NOUN
alces-483	158	9	abundance	abundance	NOUN
alces-483	158	10	of	of	ADP
alces-483	158	11	fir	fir	PROPN
alces-483	158	12	forage	forage	NOUN
alces-483	158	13	,	,	PUNCT
alces-483	158	14	differs	differ	VERB
alces-483	158	15	substantially	substantially	ADV
alces-483	158	16	across	across	ADP
alces-483	158	17	different	different	ADJ
alces-483	158	18	portions	portion	NOUN
alces-483	158	19	of	of	ADP
alces-483	158	20	isle	isle	PROPN
alces-483	158	21	royale	royale	PROPN
alces-483	158	22	in	in	ADP
alces-483	158	23	some	some	DET
alces-483	158	24	years	year	NOUN
alces-483	158	25	(	(	PUNCT
alces-483	158	26	fig	fig	NOUN
alces-483	158	27	.	.	PUNCT
alces-483	158	28	3a	3a	NUM
alces-483	158	29	)	)	PUNCT
alces-483	158	30	.	.	PUNCT
alces-483	159	1	spatial	spatial	ADJ
alces-483	159	2	variation	variation	NOUN
alces-483	159	3	in	in	ADP
alces-483	159	4	fir	fir	NOUN
alces-483	159	5	growth	growth	NOUN
alces-483	159	6	may	may	AUX
alces-483	159	7	be	be	AUX
alces-483	159	8	the	the	DET
alces-483	159	9	most	most	ADV
alces-483	159	10	significant	significant	ADJ
alces-483	159	11	aspect	aspect	NOUN
alces-483	159	12	of	of	ADP
alces-483	159	13	spatial	spatial	ADJ
alces-483	159	14	heterogeneity	heterogeneity	NOUN
alces-483	159	15	in	in	ADP
alces-483	159	16	the	the	DET
alces-483	159	17	vegetation	vegetation	NOUN
alces-483	159	18	-	-	PUNCT
alces-483	159	19	moose	moose	NOUN
alces-483	159	20	-	-	PUNCT
alces-483	159	21	wolf	wolf	NOUN
alces-483	159	22	system	system	NOUN
alces-483	159	23	on	on	ADP
alces-483	159	24	isle	isle	PROPN
alces-483	159	25	royale	royale	PROPN
alces-483	159	26	.	.	PUNCT
alces-483	160	1	is	be	AUX
alces-483	160	2	spatial	spatial	ADJ
alces-483	160	3	heterogeneity	heterogeneity	NOUN
alces-483	160	4	in	in	ADP
alces-483	160	5	balsam	balsam	NOUN
alces-483	160	6	fir	fir	NOUN
alces-483	160	7	manifested	manifest	VERB
alces-483	160	8	in	in	ADP
alces-483	160	9	the	the	DET
alces-483	160	10	dynamics	dynamic	NOUN
alces-483	160	11	of	of	ADP
alces-483	160	12	higher	high	ADJ
alces-483	160	13	trophic	trophic	ADJ
alces-483	160	14	levels	level	NOUN
alces-483	160	15	?	?	PUNCT
alces-483	161	1	to	to	PART
alces-483	161	2	gain	gain	VERB
alces-483	161	3	a	a	DET
alces-483	161	4	preliminary	preliminary	ADJ
alces-483	161	5	understanding	understanding	NOUN
alces-483	161	6	for	for	ADP
alces-483	161	7	how	how	SCONJ
alces-483	161	8	spatial	spatial	ADJ
alces-483	161	9	variation	variation	NOUN
alces-483	161	10	in	in	ADP
alces-483	161	11	fir	fir	NOUN
alces-483	161	12	could	could	AUX
alces-483	161	13	be	be	AUX
alces-483	161	14	manifest	manifest	ADJ
alces-483	161	15	in	in	ADP
alces-483	161	16	moose	moose	NOUN
alces-483	161	17	population	population	NOUN
alces-483	161	18	dynamics	dynamic	NOUN
alces-483	161	19	,	,	PUNCT
alces-483	161	20	we	we	PRON
alces-483	161	21	consider	consider	VERB
alces-483	161	22	how	how	SCONJ
alces-483	161	23	calf	calf	NOUN
alces-483	161	24	production	production	NOUN
alces-483	161	25	is	be	AUX
alces-483	161	26	influenced	influence	VERB
alces-483	161	27	by	by	ADP
alces-483	161	28	the	the	DET
alces-483	161	29	abundance	abundance	NOUN
alces-483	161	30	of	of	ADP
alces-483	161	31	fir	fir	NOUN
alces-483	161	32	forage	forage	NOUN
alces-483	161	33	.	.	PUNCT
alces-483	162	1	using	use	VERB
alces-483	162	2	our	our	PRON
alces-483	162	3	data	datum	NOUN
alces-483	162	4	(	(	PUNCT
alces-483	162	5	figs	fig	NOUN
alces-483	162	6	.	.	X
alces-483	162	7	2	2	NUM
alces-483	162	8	and	and	CCONJ
alces-483	162	9	3a	3a	NUM
alces-483	162	10	)	)	PUNCT
alces-483	162	11	and	and	CCONJ
alces-483	162	12	multiple	multiple	ADJ
alces-483	162	13	linear	linear	ADJ
alces-483	162	14	regression	regression	NOUN
alces-483	162	15	we	we	PRON
alces-483	162	16	obtained	obtain	VERB
alces-483	162	17	the	the	DET
alces-483	162	18	following	following	ADJ
alces-483	162	19	model	model	NOUN
alces-483	162	20	:	:	PUNCT
alces-483	162	21	ct	ct	PROPN
alces-483	162	22	=	=	SYM
alces-483	162	23	0.15	0.15	NUM
alces-483	162	24	–	–	PUNCT
alces-483	162	25	3.3×10	3.3×10	NUM
alces-483	162	26	-	-	SYM
alces-483	162	27	5mt-1	5mt-1	NUM
alces-483	162	28	+	+	NUM
alces-483	162	29	4.6×10	4.6×10	PROPN
alces-483	162	30	-	-	PUNCT
alces-483	162	31	2ft-1	2ft-1	NUM
alces-483	162	32	,	,	PUNCT
alces-483	162	33	(	(	PUNCT
alces-483	162	34	1	1	X
alces-483	162	35	)	)	PUNCT
alces-483	162	36	where	where	SCONJ
alces-483	162	37	percent	percent	NOUN
alces-483	162	38	calf	calf	NOUN
alces-483	162	39	production	production	NOUN
alces-483	162	40	in	in	ADP
alces-483	162	41	the	the	DET
alces-483	162	42	current	current	ADJ
alces-483	162	43	year	year	NOUN
alces-483	162	44	(	(	PUNCT
alces-483	162	45	ct	ct	NOUN
alces-483	162	46	)	)	PUNCT
alces-483	162	47	is	be	AUX
alces-483	162	48	dependent	dependent	ADJ
alces-483	162	49	on	on	ADP
alces-483	162	50	moose	moose	NOUN
alces-483	162	51	abundance	abundance	NOUN
alces-483	162	52	(	(	PUNCT
alces-483	162	53	m	m	NOUN
alces-483	162	54	)	)	PUNCT
alces-483	162	55	and	and	CCONJ
alces-483	162	56	fir	fir	NOUN
alces-483	162	57	tree	tree	NOUN
alces-483	162	58	-	-	PUNCT
alces-483	162	59	ring	ring	NOUN
alces-483	162	60	index	index	NOUN
alces-483	162	61	(	(	PUNCT
alces-483	162	62	f	f	X
alces-483	162	63	)	)	PUNCT
alces-483	162	64	in	in	ADP
alces-483	162	65	the	the	DET
alces-483	162	66	previous	previous	ADJ
alces-483	162	67	year	year	NOUN
alces-483	162	68	.	.	PUNCT
alces-483	163	1	for	for	ADP
alces-483	163	2	this	this	DET
alces-483	163	3	model	model	NOUN
alces-483	163	4	ft-1	ft-1	PROPN
alces-483	163	5	,	,	PUNCT
alces-483	163	6	represents	represent	VERB
alces-483	163	7	tree	tree	NOUN
alces-483	163	8	ring	ring	NOUN
alces-483	163	9	growth	growth	NOUN
alces-483	163	10	,	,	PUNCT
alces-483	163	11	averaged	average	VERB
alces-483	163	12	across	across	ADP
alces-483	163	13	all	all	PRON
alces-483	163	14	of	of	ADP
alces-483	163	15	isle	isle	PROPN
alces-483	163	16	royale	royale	PROPN
alces-483	163	17	.	.	PUNCT
alces-483	164	1	the	the	DET
alces-483	164	2	coefficients	coefficient	NOUN
alces-483	164	3	for	for	ADP
alces-483	164	4	moose	moose	NOUN
alces-483	164	5	(	(	PUNCT
alces-483	164	6	p	p	X
alces-483	164	7	=	=	NOUN
alces-483	164	8	0.03	0.03	NUM
alces-483	164	9	)	)	PUNCT
alces-483	164	10	and	and	CCONJ
alces-483	164	11	fir	fir	NOUN
alces-483	164	12	(	(	PUNCT
alces-483	164	13	p	p	X
alces-483	164	14	=	=	NOUN
alces-483	164	15	0.03	0.03	NUM
alces-483	164	16	)	)	PUNCT
alces-483	164	17	are	be	AUX
alces-483	164	18	statistically	statistically	ADV
alces-483	164	19	fig	fig	VERB
alces-483	164	20	.	.	PUNCT
alces-483	165	1	3	3	X
alces-483	165	2	.	.	X
alces-483	165	3	upper	upper	ADJ
alces-483	165	4	panel	panel	NOUN
alces-483	165	5	:	:	PUNCT
alces-483	165	6	index	index	NOUN
alces-483	165	7	of	of	ADP
alces-483	165	8	fir	fir	NOUN
alces-483	165	9	growth	growth	NOUN
alces-483	165	10	(	(	PUNCT
alces-483	165	11	see	see	VERB
alces-483	165	12	text	text	NOUN
alces-483	165	13	)	)	PUNCT
alces-483	165	14	for	for	ADP
alces-483	165	15	eight	eight	NUM
alces-483	165	16	trees	tree	NOUN
alces-483	165	17	located	locate	VERB
alces-483	165	18	on	on	ADP
alces-483	165	19	the	the	DET
alces-483	165	20	east	east	ADJ
alces-483	165	21	half	half	NOUN
alces-483	165	22	of	of	ADP
alces-483	165	23	isle	isle	PROPN
alces-483	165	24	royale	royale	PROPN
alces-483	165	25	and	and	CCONJ
alces-483	165	26	for	for	ADP
alces-483	165	27	eight	eight	NUM
alces-483	165	28	trees	tree	NOUN
alces-483	165	29	located	locate	VERB
alces-483	165	30	on	on	ADP
alces-483	165	31	the	the	DET
alces-483	165	32	west	west	NOUN
alces-483	165	33	half	half	NOUN
alces-483	165	34	of	of	ADP
alces-483	165	35	isle	isle	PROPN
alces-483	165	36	royale	royale	PROPN
alces-483	165	37	.	.	PUNCT
alces-483	166	1	middle	middle	ADJ
alces-483	166	2	panel	panel	NOUN
alces-483	166	3	:	:	PUNCT
alces-483	166	4	two	two	NUM
alces-483	166	5	predictions	prediction	NOUN
alces-483	166	6	of	of	ADP
alces-483	166	7	percent	percent	NOUN
alces-483	166	8	calves	calf	NOUN
alces-483	166	9	based	base	VERB
alces-483	166	10	on	on	ADP
alces-483	166	11	equation	equation	NOUN
alces-483	166	12	(	(	PUNCT
alces-483	166	13	1	1	NUM
alces-483	166	14	)	)	PUNCT
alces-483	166	15	and	and	CCONJ
alces-483	166	16	the	the	DET
alces-483	166	17	hypothetical	hypothetical	ADJ
alces-483	166	18	assumptions	assumption	NOUN
alces-483	166	19	that	that	PRON
alces-483	166	20	island	island	NOUN
alces-483	166	21	-	-	PUNCT
alces-483	166	22	wide	wide	ADJ
alces-483	166	23	fir	fir	NOUN
alces-483	166	24	growth	growth	NOUN
alces-483	166	25	is	be	AUX
alces-483	166	26	characterized	characterize	VERB
alces-483	166	27	by	by	ADP
alces-483	166	28	fir	fir	NOUN
alces-483	166	29	growth	growth	NOUN
alces-483	166	30	that	that	PRON
alces-483	166	31	actually	actually	ADV
alces-483	166	32	characterizes	characterize	VERB
alces-483	166	33	only	only	ADV
alces-483	166	34	the	the	DET
alces-483	166	35	east	east	ADJ
alces-483	166	36	half	half	NOUN
alces-483	166	37	of	of	ADP
alces-483	166	38	the	the	DET
alces-483	166	39	island	island	NOUN
alces-483	166	40	(	(	PUNCT
alces-483	166	41	open	open	ADJ
alces-483	166	42	circles	circle	NOUN
alces-483	166	43	)	)	PUNCT
alces-483	166	44	and	and	CCONJ
alces-483	166	45	only	only	ADV
alces-483	166	46	the	the	DET
alces-483	166	47	west	west	ADJ
alces-483	166	48	half	half	NOUN
alces-483	166	49	of	of	ADP
alces-483	166	50	the	the	DET
alces-483	166	51	island	island	NOUN
alces-483	166	52	(	(	PUNCT
alces-483	166	53	closed	close	VERB
alces-483	166	54	circles	circle	NOUN
alces-483	166	55	)	)	PUNCT
alces-483	166	56	.	.	PUNCT
alces-483	167	1	lower	low	ADJ
alces-483	167	2	panel	panel	NOUN
alces-483	167	3	:	:	PUNCT
alces-483	167	4	the	the	DET
alces-483	167	5	difference	difference	NOUN
alces-483	167	6	between	between	ADP
alces-483	167	7	the	the	DET
alces-483	167	8	two	two	NUM
alces-483	167	9	predicted	predict	VERB
alces-483	167	10	percent	percent	NOUN
alces-483	167	11	calf	calf	NOUN
alces-483	167	12	alces	alce	NOUN
alces-483	167	13	vol	vol	NOUN
alces-483	167	14	.	.	PROPN
alces-483	167	15	39	39	NUM
alces-483	167	16	,	,	PUNCT
alces-483	167	17	2003	2003	NUM
alces-483	167	18	peterson	peterson	PROPN
alces-483	167	19	et	et	PROPN
alces-483	167	20	al	al	PROPN
alces-483	167	21	.	.	PROPN
alces-483	167	22	–	–	PUNCT
alces-483	167	23	space	space	NOUN
alces-483	167	24	and	and	CCONJ
alces-483	167	25	time	time	NOUN
alces-483	167	26	in	in	ADP
alces-483	167	27	predation	predation	NOUN
alces-483	167	28	dynamics	dynamic	NOUN
alces-483	167	29	223	223	NUM
alces-483	167	30	significant	significant	ADJ
alces-483	167	31	,	,	PUNCT
alces-483	167	32	and	and	CCONJ
alces-483	167	33	this	this	DET
alces-483	167	34	model	model	NOUN
alces-483	167	35	explains	explain	VERB
alces-483	167	36	approximately	approximately	ADV
alces-483	167	37	20	20	NUM
alces-483	167	38	%	%	NOUN
alces-483	167	39	of	of	ADP
alces-483	167	40	the	the	DET
alces-483	167	41	variation	variation	NOUN
alces-483	167	42	in	in	ADP
alces-483	167	43	calf	calf	NOUN
alces-483	167	44	production	production	NOUN
alces-483	167	45	.	.	PUNCT
alces-483	168	1	the	the	DET
alces-483	168	2	residuals	residual	NOUN
alces-483	168	3	of	of	ADP
alces-483	168	4	this	this	DET
alces-483	168	5	model	model	NOUN
alces-483	168	6	do	do	AUX
alces-483	168	7	not	not	PART
alces-483	168	8	appear	appear	VERB
alces-483	168	9	to	to	PART
alces-483	168	10	deviate	deviate	VERB
alces-483	168	11	substantially	substantially	ADV
alces-483	168	12	from	from	ADP
alces-483	168	13	normal	normal	ADJ
alces-483	168	14	(	(	PUNCT
alces-483	168	15	p	p	X
alces-483	168	16	=	=	NOUN
alces-483	168	17	0.83	0.83	NUM
alces-483	168	18	)	)	PUNCT
alces-483	168	19	,	,	PUNCT
alces-483	168	20	nor	nor	CCONJ
alces-483	168	21	do	do	AUX
alces-483	168	22	they	they	PRON
alces-483	168	23	appear	appear	VERB
alces-483	168	24	to	to	PART
alces-483	168	25	be	be	AUX
alces-483	168	26	autocorrelated	autocorrelate	VERB
alces-483	168	27	(	(	PUNCT
alces-483	168	28	durbin	durbin	ADJ
alces-483	168	29	-	-	PUNCT
alces-483	168	30	watson	watson	NOUN
alces-483	168	31	statistic	statistic	NOUN
alces-483	168	32	=	=	NOUN
alces-483	168	33	0.93	0.93	NUM
alces-483	168	34	)	)	PUNCT
alces-483	168	35	.	.	PUNCT
alces-483	169	1	for	for	ADP
alces-483	169	2	this	this	DET
alces-483	169	3	model	model	NOUN
alces-483	169	4	,	,	PUNCT
alces-483	169	5	interannual	interannual	ADJ
alces-483	169	6	variation	variation	NOUN
alces-483	169	7	in	in	ADP
alces-483	169	8	tree	tree	NOUN
alces-483	169	9	ring	ring	NOUN
alces-483	169	10	growth	growth	NOUN
alces-483	169	11	accounts	account	VERB
alces-483	169	12	for	for	ADP
alces-483	169	13	approximately	approximately	ADV
alces-483	169	14	10	10	NUM
alces-483	169	15	%	%	NOUN
alces-483	169	16	of	of	ADP
alces-483	169	17	the	the	DET
alces-483	169	18	variation	variation	NOUN
alces-483	169	19	in	in	ADP
alces-483	169	20	calf	calf	NOUN
alces-483	169	21	production	production	NOUN
alces-483	169	22	.	.	PUNCT
alces-483	170	1	consider	consider	VERB
alces-483	170	2	,	,	PUNCT
alces-483	170	3	hypothetically	hypothetically	ADV
alces-483	170	4	,	,	PUNCT
alces-483	170	5	what	what	PRON
alces-483	170	6	the	the	DET
alces-483	170	7	temporal	temporal	ADJ
alces-483	170	8	dynamics	dynamic	NOUN
alces-483	170	9	of	of	ADP
alces-483	170	10	calf	calf	NOUN
alces-483	170	11	production	production	NOUN
alces-483	170	12	would	would	AUX
alces-483	170	13	be	be	AUX
alces-483	170	14	like	like	ADP
alces-483	170	15	if	if	SCONJ
alces-483	170	16	fir	fir	NOUN
alces-483	170	17	growth	growth	NOUN
alces-483	170	18	across	across	ADP
alces-483	170	19	the	the	DET
alces-483	170	20	entire	entire	ADJ
alces-483	170	21	island	island	NOUN
alces-483	170	22	were	be	AUX
alces-483	170	23	like	like	SCONJ
alces-483	170	24	it	it	PRON
alces-483	170	25	has	have	AUX
alces-483	170	26	been	be	AUX
alces-483	170	27	on	on	ADP
alces-483	170	28	just	just	ADV
alces-483	170	29	the	the	DET
alces-483	170	30	west	west	ADJ
alces-483	170	31	end	end	NOUN
alces-483	170	32	of	of	ADP
alces-483	170	33	isle	isle	PROPN
alces-483	170	34	royale	royale	PROPN
alces-483	170	35	,	,	PUNCT
alces-483	170	36	or	or	CCONJ
alces-483	170	37	just	just	ADV
alces-483	170	38	the	the	DET
alces-483	170	39	east	east	ADJ
alces-483	170	40	end	end	NOUN
alces-483	170	41	of	of	ADP
alces-483	170	42	isle	isle	PROPN
alces-483	170	43	royale	royale	PROPN
alces-483	170	44	.	.	PUNCT
alces-483	171	1	to	to	PART
alces-483	171	2	explore	explore	VERB
alces-483	171	3	this	this	DET
alces-483	171	4	hypothetical	hypothetical	ADJ
alces-483	171	5	scenario	scenario	NOUN
alces-483	171	6	,	,	PUNCT
alces-483	171	7	we	we	PRON
alces-483	171	8	predicted	predict	VERB
alces-483	171	9	2	2	NUM
alces-483	171	10	time	time	NOUN
alces-483	171	11	series	series	NOUN
alces-483	171	12	of	of	ADP
alces-483	171	13	percent	percent	NOUN
alces-483	171	14	calf	calf	NOUN
alces-483	171	15	production	production	NOUN
alces-483	171	16	using	use	VERB
alces-483	171	17	equation	equation	NOUN
alces-483	171	18	(	(	PUNCT
alces-483	171	19	1	1	NUM
alces-483	171	20	)	)	PUNCT
alces-483	171	21	,	,	PUNCT
alces-483	171	22	except	except	SCONJ
alces-483	171	23	that	that	PRON
alces-483	171	24	for	for	ADP
alces-483	171	25	one	one	NUM
alces-483	171	26	time	time	NOUN
alces-483	171	27	series	series	NOUN
alces-483	171	28	we	we	PRON
alces-483	171	29	replaced	replace	VERB
alces-483	171	30	ft-1	ft-1	PUNCT
alces-483	171	31	with	with	ADP
alces-483	171	32	a	a	DET
alces-483	171	33	series	series	NOUN
alces-483	171	34	of	of	ADP
alces-483	171	35	values	value	NOUN
alces-483	171	36	representing	represent	VERB
alces-483	171	37	growth	growth	NOUN
alces-483	171	38	at	at	ADP
alces-483	171	39	the	the	DET
alces-483	171	40	west	west	PROPN
alces-483	171	41	end	end	NOUN
alces-483	171	42	of	of	ADP
alces-483	171	43	isle	isle	PROPN
alces-483	171	44	royale	royale	PROPN
alces-483	171	45	,	,	PUNCT
alces-483	171	46	and	and	CCONJ
alces-483	171	47	for	for	ADP
alces-483	171	48	the	the	DET
alces-483	171	49	other	other	ADJ
alces-483	171	50	series	series	NOUN
alces-483	171	51	,	,	PUNCT
alces-483	171	52	we	we	PRON
alces-483	171	53	replaced	replace	VERB
alces-483	171	54	ft-1	ft-1	PUNCT
alces-483	171	55	with	with	ADP
alces-483	171	56	a	a	DET
alces-483	171	57	series	series	NOUN
alces-483	171	58	of	of	ADP
alces-483	171	59	values	value	NOUN
alces-483	171	60	representing	represent	VERB
alces-483	171	61	growth	growth	NOUN
alces-483	171	62	at	at	ADP
alces-483	171	63	the	the	DET
alces-483	171	64	east	east	ADJ
alces-483	171	65	end	end	NOUN
alces-483	171	66	of	of	ADP
alces-483	171	67	isle	isle	PROPN
alces-483	171	68	royale	royale	PROPN
alces-483	171	69	.	.	PUNCT
alces-483	172	1	these	these	PRON
alces-483	172	2	predicted	predict	VERB
alces-483	172	3	percent	percent	NOUN
alces-483	172	4	calf	calf	NOUN
alces-483	172	5	time	time	NOUN
alces-483	172	6	series	series	NOUN
alces-483	172	7	are	be	AUX
alces-483	172	8	depicted	depict	VERB
alces-483	172	9	in	in	ADP
alces-483	172	10	figure	figure	NOUN
alces-483	172	11	3b	3b	NUM
alces-483	172	12	,	,	PUNCT
alces-483	172	13	and	and	CCONJ
alces-483	172	14	the	the	DET
alces-483	172	15	absolute	absolute	ADJ
alces-483	172	16	difference	difference	NOUN
alces-483	172	17	between	between	ADP
alces-483	172	18	these	these	DET
alces-483	172	19	time	time	NOUN
alces-483	172	20	series	series	NOUN
alces-483	172	21	is	be	AUX
alces-483	172	22	depicted	depict	VERB
alces-483	172	23	in	in	ADP
alces-483	172	24	figure	figure	NOUN
alces-483	172	25	3c	3c	NUM
alces-483	172	26	.	.	PUNCT
alces-483	173	1	in	in	ADP
alces-483	173	2	absolute	absolute	ADJ
alces-483	173	3	terms	term	NOUN
alces-483	173	4	the	the	DET
alces-483	173	5	difference	difference	NOUN
alces-483	173	6	between	between	ADP
alces-483	173	7	the	the	DET
alces-483	173	8	two	two	NUM
alces-483	173	9	time	time	NOUN
alces-483	173	10	series	series	NOUN
alces-483	173	11	appears	appear	VERB
alces-483	173	12	minor	minor	ADJ
alces-483	173	13	.	.	PUNCT
alces-483	174	1	however	however	ADV
alces-483	174	2	,	,	PUNCT
alces-483	174	3	the	the	DET
alces-483	174	4	differences	difference	NOUN
alces-483	174	5	are	be	AUX
alces-483	174	6	highly	highly	ADV
alces-483	174	7	autocorrelated	autocorrelated	ADJ
alces-483	174	8	.	.	PUNCT
alces-483	175	1	for	for	ADP
alces-483	175	2	example	example	NOUN
alces-483	175	3	,	,	PUNCT
alces-483	175	4	for	for	ADP
alces-483	175	5	11	11	NUM
alces-483	175	6	consecutive	consecutive	ADJ
alces-483	175	7	years	year	NOUN
alces-483	175	8	(	(	PUNCT
alces-483	175	9	1968	1968	NUM
alces-483	175	10	-	-	SYM
alces-483	175	11	78	78	NUM
alces-483	175	12	)	)	PUNCT
alces-483	175	13	and	and	CCONJ
alces-483	175	14	for	for	ADP
alces-483	175	15	9	9	NUM
alces-483	175	16	consecutive	consecutive	ADJ
alces-483	175	17	years	year	NOUN
alces-483	175	18	(	(	PUNCT
alces-483	175	19	199098	199098	NUM
alces-483	175	20	)	)	PUNCT
alces-483	175	21	the	the	DET
alces-483	175	22	difference	difference	NOUN
alces-483	175	23	is	be	AUX
alces-483	175	24	positive	positive	ADJ
alces-483	175	25	.	.	PUNCT
alces-483	176	1	such	such	DET
alces-483	176	2	a	a	DET
alces-483	176	3	pattern	pattern	NOUN
alces-483	176	4	could	could	AUX
alces-483	176	5	lead	lead	VERB
alces-483	176	6	to	to	ADP
alces-483	176	7	substantial	substantial	ADJ
alces-483	176	8	differences	difference	NOUN
alces-483	176	9	in	in	ADP
alces-483	176	10	moose	moose	NOUN
alces-483	176	11	population	population	NOUN
alces-483	176	12	abundance	abundance	PROPN
alces-483	176	13	.	.	PUNCT
alces-483	177	1	investigation	investigation	NOUN
alces-483	177	2	beyond	beyond	ADP
alces-483	177	3	the	the	DET
alces-483	177	4	scope	scope	NOUN
alces-483	177	5	of	of	ADP
alces-483	177	6	this	this	DET
alces-483	177	7	manuscript	manuscript	NOUN
alces-483	177	8	is	be	AUX
alces-483	177	9	required	require	VERB
alces-483	177	10	to	to	PART
alces-483	177	11	accurately	accurately	ADV
alces-483	177	12	understand	understand	VERB
alces-483	177	13	the	the	DET
alces-483	177	14	extent	extent	NOUN
alces-483	177	15	to	to	PART
alces-483	177	16	which	which	PRON
alces-483	177	17	spatial	spatial	ADJ
alces-483	177	18	heterogeneity	heterogeneity	NOUN
alces-483	177	19	in	in	ADP
alces-483	177	20	fir	fir	NOUN
alces-483	177	21	could	could	AUX
alces-483	177	22	give	give	VERB
alces-483	177	23	rise	rise	NOUN
alces-483	177	24	to	to	ADP
alces-483	177	25	biologically	biologically	ADV
alces-483	177	26	significant	significant	ADJ
alces-483	177	27	spatial	spatial	ADJ
alces-483	177	28	heterogeneity	heterogeneity	NOUN
alces-483	177	29	in	in	ADP
alces-483	177	30	moose	moose	NOUN
alces-483	177	31	population	population	NOUN
alces-483	177	32	dynamics	dynamic	NOUN
alces-483	177	33	.	.	PUNCT
alces-483	178	1	nevertheless	nevertheless	ADV
alces-483	178	2	,	,	PUNCT
alces-483	178	3	our	our	PRON
alces-483	178	4	hypothetical	hypothetical	ADJ
alces-483	178	5	example	example	NOUN
alces-483	178	6	suggests	suggest	VERB
alces-483	178	7	that	that	SCONJ
alces-483	178	8	further	further	ADJ
alces-483	178	9	investigation	investigation	NOUN
alces-483	178	10	could	could	AUX
alces-483	178	11	reveal	reveal	VERB
alces-483	178	12	important	important	ADJ
alces-483	178	13	insights	insight	NOUN
alces-483	178	14	.	.	PUNCT
alces-483	179	1	a	a	DET
alces-483	179	2	convincing	convincing	ADJ
alces-483	179	3	example	example	NOUN
alces-483	179	4	of	of	ADP
alces-483	179	5	the	the	DET
alces-483	179	6	importance	importance	NOUN
alces-483	179	7	of	of	ADP
alces-483	179	8	spatial	spatial	ADJ
alces-483	179	9	differences	difference	NOUN
alces-483	179	10	in	in	ADP
alces-483	179	11	habitat	habitat	NOUN
alces-483	179	12	emerged	emerge	VERB
alces-483	179	13	as	as	ADP
alces-483	179	14	moose	moose	NOUN
alces-483	179	15	population	population	NOUN
alces-483	179	16	levels	level	NOUN
alces-483	179	17	at	at	ADP
alces-483	179	18	opposite	opposite	ADJ
alces-483	179	19	ends	end	NOUN
alces-483	179	20	of	of	ADP
alces-483	179	21	isle	isle	PROPN
alces-483	179	22	royale	royale	PROPN
alces-483	179	23	diverged	diverge	VERB
alces-483	179	24	as	as	ADP
alces-483	179	25	a	a	DET
alces-483	179	26	result	result	NOUN
alces-483	179	27	of	of	ADP
alces-483	179	28	the	the	DET
alces-483	179	29	moose	moose	NOUN
alces-483	179	30	die	die	NOUN
alces-483	179	31	-	-	PUNCT
alces-483	179	32	off	off	NOUN
alces-483	179	33	in	in	ADP
alces-483	179	34	1996	1996	NUM
alces-483	179	35	when	when	SCONJ
alces-483	179	36	many	many	ADJ
alces-483	179	37	moose	moose	NOUN
alces-483	179	38	survived	survive	VERB
alces-483	179	39	in	in	ADP
alces-483	179	40	the	the	DET
alces-483	179	41	thick	thick	ADJ
alces-483	179	42	fir	fir	NOUN
alces-483	179	43	stands	stand	VERB
alces-483	179	44	at	at	ADP
alces-483	179	45	the	the	DET
alces-483	179	46	east	east	ADJ
alces-483	179	47	end	end	NOUN
alces-483	179	48	but	but	CCONJ
alces-483	179	49	perished	perish	VERB
alces-483	179	50	at	at	ADP
alces-483	179	51	the	the	DET
alces-483	179	52	west	west	PROPN
alces-483	179	53	end	end	NOUN
alces-483	179	54	(	(	PUNCT
alces-483	179	55	r.	r.	PROPN
alces-483	179	56	o.	o.	PROPN
alces-483	179	57	peterson	peterson	PROPN
alces-483	179	58	and	and	CCONJ
alces-483	179	59	j.	j.	PROPN
alces-483	179	60	a.	a.	PROPN
alces-483	179	61	vucetich	vucetich	PROPN
alces-483	179	62	,	,	PUNCT
alces-483	179	63	unpublished	unpublished	ADJ
alces-483	179	64	data	datum	NOUN
alces-483	179	65	)	)	PUNCT
alces-483	179	66	.	.	PUNCT
alces-483	180	1	by	by	ADP
alces-483	180	2	1999	1999	NUM
alces-483	180	3	,	,	PUNCT
alces-483	180	4	20	20	NUM
alces-483	180	5	of	of	ADP
alces-483	180	6	the	the	DET
alces-483	180	7	25	25	NUM
alces-483	180	8	wolves	wolf	NOUN
alces-483	180	9	present	present	ADJ
alces-483	180	10	were	be	AUX
alces-483	180	11	also	also	ADV
alces-483	180	12	supported	support	VERB
alces-483	180	13	by	by	ADP
alces-483	180	14	moose	moose	NOUN
alces-483	180	15	at	at	ADP
alces-483	180	16	the	the	DET
alces-483	180	17	east	east	NOUN
alces-483	180	18	end	end	NOUN
alces-483	180	19	,	,	PUNCT
alces-483	180	20	so	so	CCONJ
alces-483	180	21	the	the	DET
alces-483	180	22	bottom	bottom	ADJ
alces-483	180	23	-	-	PUNCT
alces-483	180	24	up	up	ADP
alces-483	180	25	pulse	pulse	NOUN
alces-483	180	26	of	of	ADP
alces-483	180	27	productivity	productivity	NOUN
alces-483	180	28	was	be	AUX
alces-483	180	29	manifest	manifest	ADJ
alces-483	180	30	at	at	ADP
alces-483	180	31	all	all	DET
alces-483	180	32	3	3	NUM
alces-483	180	33	trophic	trophic	ADJ
alces-483	180	34	levels	level	NOUN
alces-483	180	35	.	.	PUNCT
alces-483	181	1	thus	thus	ADV
alces-483	181	2	,	,	PUNCT
alces-483	181	3	a	a	DET
alces-483	181	4	long	long	ADJ
alces-483	181	5	-	-	PUNCT
alces-483	181	6	term	term	NOUN
alces-483	181	7	and	and	CCONJ
alces-483	181	8	large	large	ADJ
alces-483	181	9	-	-	PUNCT
alces-483	181	10	scale	scale	NOUN
alces-483	181	11	pattern	pattern	NOUN
alces-483	181	12	of	of	ADP
alces-483	181	13	soil	soil	NOUN
alces-483	181	14	development	development	NOUN
alces-483	181	15	established	establish	VERB
alces-483	181	16	as	as	SCONJ
alces-483	181	17	glaciers	glacier	NOUN
alces-483	181	18	retreated	retreat	VERB
alces-483	181	19	was	be	AUX
alces-483	181	20	manifested	manifest	VERB
alces-483	181	21	in	in	ADP
alces-483	181	22	a	a	DET
alces-483	181	23	rather	rather	ADV
alces-483	181	24	peculiar	peculiar	ADJ
alces-483	181	25	historic	historic	ADJ
alces-483	181	26	fashion	fashion	NOUN
alces-483	181	27	as	as	ADP
alces-483	181	28	a	a	DET
alces-483	181	29	once	once	ADV
alces-483	181	30	-	-	PUNCT
alces-483	181	31	per	per	ADP
alces-483	181	32	-	-	PUNCT
alces-483	181	33	century	century	NOUN
alces-483	181	34	severe	severe	ADJ
alces-483	181	35	winter	winter	NOUN
alces-483	181	36	impacted	impact	VERB
alces-483	181	37	a	a	DET
alces-483	181	38	moose	moose	NOUN
alces-483	181	39	population	population	NOUN
alces-483	181	40	poised	poise	VERB
alces-483	181	41	at	at	ADP
alces-483	181	42	a	a	DET
alces-483	181	43	historic	historic	ADJ
alces-483	181	44	high	high	ADJ
alces-483	181	45	population	population	NOUN
alces-483	181	46	density	density	NOUN
alces-483	181	47	.	.	PUNCT
alces-483	182	1	this	this	DET
alces-483	182	2	example	example	NOUN
alces-483	182	3	of	of	ADP
alces-483	182	4	the	the	DET
alces-483	182	5	influence	influence	NOUN
alces-483	182	6	of	of	ADP
alces-483	182	7	spatial	spatial	ADJ
alces-483	182	8	heterogeneity	heterogeneity	NOUN
alces-483	182	9	in	in	ADP
alces-483	182	10	fir	fir	NOUN
alces-483	182	11	dynamics	dynamic	NOUN
alces-483	182	12	highlights	highlight	NOUN
alces-483	182	13	the	the	DET
alces-483	182	14	need	need	NOUN
alces-483	182	15	for	for	ADP
alces-483	182	16	an	an	DET
alces-483	182	17	improved	improved	ADJ
alces-483	182	18	understanding	understanding	NOUN
alces-483	182	19	of	of	ADP
alces-483	182	20	meso	meso	NOUN
alces-483	182	21	-	-	PUNCT
alces-483	182	22	scale	scale	NOUN
alces-483	182	23	spatial	spatial	ADJ
alces-483	182	24	heterogeneity	heterogeneity	NOUN
alces-483	182	25	in	in	ADP
alces-483	182	26	wolf	wolf	NOUN
alces-483	182	27	-	-	PUNCT
alces-483	182	28	ungulate	ungulate	ADJ
alces-483	182	29	dynamics	dynamic	NOUN
alces-483	182	30	across	across	ADP
alces-483	182	31	north	north	PROPN
alces-483	182	32	america	america	PROPN
alces-483	182	33	(	(	PUNCT
alces-483	182	34	cf	cf	NOUN
alces-483	182	35	.	.	PUNCT
alces-483	183	1	orians	orian	NOUN
alces-483	183	2	et	et	PROPN
alces-483	183	3	al	al	PROPN
alces-483	183	4	.	.	PROPN
alces-483	183	5	1997	1997	NUM
alces-483	183	6	)	)	PUNCT
alces-483	183	7	.	.	PUNCT
alces-483	184	1	what	what	PRON
alces-483	184	2	’s	’	VERB
alces-483	184	3	scale	scale	NOUN
alces-483	184	4	got	get	VERB
alces-483	184	5	to	to	PART
alces-483	184	6	do	do	VERB
alces-483	184	7	with	with	ADP
alces-483	184	8	it	it	PRON
alces-483	184	9	?	?	PUNCT
alces-483	185	1	with	with	ADP
alces-483	185	2	data	datum	NOUN
alces-483	185	3	on	on	ADP
alces-483	185	4	isle	isle	PROPN
alces-483	185	5	royale	royale	PROPN
alces-483	185	6	wolves	wolf	NOUN
alces-483	185	7	and	and	CCONJ
alces-483	185	8	moose	moose	NOUN
alces-483	185	9	from	from	ADP
alces-483	185	10	any	any	DET
alces-483	185	11	single	single	ADJ
alces-483	185	12	5	5	NUM
alces-483	185	13	-	-	PUNCT
alces-483	185	14	year	year	NOUN
alces-483	185	15	period	period	NOUN
alces-483	185	16	of	of	ADP
alces-483	185	17	the	the	DET
alces-483	185	18	last	last	ADJ
alces-483	185	19	40	40	NUM
alces-483	185	20	years	year	NOUN
alces-483	185	21	,	,	PUNCT
alces-483	185	22	it	it	PRON
alces-483	185	23	would	would	AUX
alces-483	185	24	be	be	AUX
alces-483	185	25	possible	possible	ADJ
alces-483	185	26	to	to	PART
alces-483	185	27	support	support	VERB
alces-483	185	28	almost	almost	ADV
alces-483	185	29	any	any	PRON
alces-483	185	30	interpretation	interpretation	NOUN
alces-483	185	31	of	of	ADP
alces-483	185	32	their	their	PRON
alces-483	185	33	interaction	interaction	NOUN
alces-483	185	34	,	,	PUNCT
alces-483	185	35	not	not	PART
alces-483	185	36	unlike	unlike	ADP
alces-483	185	37	the	the	DET
alces-483	185	38	fabled	fabled	ADJ
alces-483	185	39	10	10	NUM
alces-483	185	40	blind	blind	ADJ
alces-483	185	41	men	man	NOUN
alces-483	185	42	describing	describe	VERB
alces-483	185	43	an	an	DET
alces-483	185	44	elephant	elephant	NOUN
alces-483	185	45	.	.	PUNCT
alces-483	186	1	thus	thus	ADV
alces-483	186	2	varying	vary	VERB
alces-483	186	3	interpretations	interpretation	NOUN
alces-483	186	4	of	of	ADP
alces-483	186	5	predator	predator	NOUN
alces-483	186	6	-	-	PUNCT
alces-483	186	7	prey	prey	NOUN
alces-483	186	8	dynamics	dynamic	NOUN
alces-483	186	9	on	on	ADP
alces-483	186	10	isle	isle	PROPN
alces-483	186	11	royale	royale	PROPN
alces-483	186	12	(	(	PUNCT
alces-483	186	13	vanballenberghe	vanballenberghe	NOUN
alces-483	186	14	1987	1987	NUM
alces-483	186	15	)	)	PUNCT
alces-483	186	16	arose	arise	VERB
alces-483	186	17	in	in	ADP
alces-483	186	18	part	part	NOUN
alces-483	186	19	from	from	ADP
alces-483	186	20	the	the	DET
alces-483	186	21	slow	slow	ADJ
alces-483	186	22	rate	rate	NOUN
alces-483	186	23	of	of	ADP
alces-483	186	24	change	change	NOUN
alces-483	186	25	as	as	ADP
alces-483	186	26	a	a	DET
alces-483	186	27	system	system	NOUN
alces-483	186	28	initially	initially	ADV
alces-483	186	29	interpreted	interpret	VERB
alces-483	186	30	as	as	ADP
alces-483	186	31	being	be	AUX
alces-483	186	32	in	in	ADP
alces-483	186	33	equilibrium	equilibrium	NOUN
alces-483	186	34	(	(	PUNCT
alces-483	186	35	mech	mech	NOUN
alces-483	186	36	1966	1966	NUM
alces-483	186	37	,	,	PUNCT
alces-483	186	38	peterson	peterson	NOUN
alces-483	186	39	1977	1977	NUM
alces-483	186	40	)	)	PUNCT
alces-483	186	41	has	have	AUX
alces-483	186	42	since	since	SCONJ
alces-483	186	43	exhibited	exhibit	VERB
alces-483	186	44	long	long	ADJ
alces-483	186	45	-	-	PUNCT
alces-483	186	46	term	term	NOUN
alces-483	186	47	oscillatory	oscillatory	ADJ
alces-483	186	48	tendencies	tendency	NOUN
alces-483	186	49	(	(	PUNCT
alces-483	186	50	peterson	peterson	PROPN
alces-483	186	51	et	et	PROPN
alces-483	186	52	al	al	PROPN
alces-483	186	53	.	.	PROPN
alces-483	186	54	1984	1984	NUM
alces-483	186	55	,	,	PUNCT
alces-483	186	56	mclaren	mclaren	PROPN
alces-483	186	57	and	and	CCONJ
alces-483	186	58	peterson	peterson	PROPN
alces-483	186	59	1994	1994	NUM
alces-483	186	60	)	)	PUNCT
alces-483	186	61	.	.	PUNCT
alces-483	187	1	different	different	ADJ
alces-483	187	2	ecological	ecological	ADJ
alces-483	187	3	factors	factor	NOUN
alces-483	187	4	have	have	AUX
alces-483	187	5	prevailed	prevail	VERB
alces-483	187	6	at	at	ADP
alces-483	187	7	different	different	ADJ
alces-483	187	8	times	time	NOUN
alces-483	187	9	.	.	PUNCT
alces-483	188	1	temporal	temporal	ADJ
alces-483	188	2	fluctuations	fluctuation	NOUN
alces-483	188	3	at	at	ADP
alces-483	188	4	decadal	decadal	NOUN
alces-483	188	5	intervals	interval	NOUN
alces-483	188	6	may	may	AUX
alces-483	188	7	be	be	AUX
alces-483	188	8	the	the	DET
alces-483	188	9	norm	norm	NOUN
alces-483	188	10	for	for	ADP
alces-483	188	11	large	large	ADJ
alces-483	188	12	-	-	PUNCT
alces-483	188	13	bodied	bodied	ADJ
alces-483	188	14	ungulates	ungulate	NOUN
alces-483	188	15	and	and	CCONJ
alces-483	188	16	their	their	PRON
alces-483	188	17	prey	prey	NOUN
alces-483	188	18	,	,	PUNCT
alces-483	188	19	but	but	CCONJ
alces-483	188	20	this	this	PRON
alces-483	188	21	is	be	AUX
alces-483	188	22	much	much	ADV
alces-483	188	23	longer	long	ADJ
alces-483	188	24	than	than	ADP
alces-483	188	25	our	our	PRON
alces-483	188	26	usual	usual	ADJ
alces-483	188	27	framework	framework	NOUN
alces-483	188	28	for	for	ADP
alces-483	188	29	research	research	NOUN
alces-483	188	30	and	and	CCONJ
alces-483	188	31	management	management	NOUN
alces-483	188	32	.	.	PUNCT
alces-483	189	1	the	the	DET
alces-483	189	2	dilemma	dilemma	NOUN
alces-483	189	3	of	of	ADP
alces-483	189	4	scale	scale	NOUN
alces-483	189	5	-	-	PUNCT
alces-483	189	6	dependent	dependent	ADJ
alces-483	189	7	understanding	understanding	NOUN
alces-483	189	8	appears	appear	VERB
alces-483	189	9	to	to	PART
alces-483	189	10	be	be	AUX
alces-483	189	11	a	a	DET
alces-483	189	12	general	general	ADJ
alces-483	189	13	one	one	NUM
alces-483	189	14	for	for	ADP
alces-483	189	15	space	space	NOUN
alces-483	189	16	and	and	CCONJ
alces-483	189	17	time	time	NOUN
alces-483	189	18	in	in	ADP
alces-483	189	19	predation	predation	NOUN
alces-483	189	20	dynamics	dynamic	NOUN
alces-483	189	21	–	–	PUNCT
alces-483	189	22	peterson	peterson	PROPN
alces-483	189	23	et	et	PROPN
alces-483	189	24	al	al	PROPN
alces-483	189	25	.	.	PROPN
alces-483	189	26	alces	alce	NOUN
alces-483	189	27	vol	vol	NOUN
alces-483	189	28	.	.	PROPN
alces-483	189	29	39	39	NUM
alces-483	189	30	,	,	PUNCT
alces-483	189	31	2003	2003	NUM
alces-483	189	32	224	224	NUM
alces-483	189	33	animal	animal	NOUN
alces-483	189	34	ecology	ecology	NOUN
alces-483	189	35	.	.	PUNCT
alces-483	190	1	temporal	temporal	ADJ
alces-483	190	2	and	and	CCONJ
alces-483	190	3	spatial	spatial	ADJ
alces-483	190	4	variation	variation	NOUN
alces-483	190	5	in	in	ADP
alces-483	190	6	population	population	NOUN
alces-483	190	7	density	density	NOUN
alces-483	190	8	are	be	AUX
alces-483	190	9	not	not	PART
alces-483	190	10	wellunderstood	wellunderstood	NOUN
alces-483	190	11	(	(	PUNCT
alces-483	190	12	in	in	ADP
alces-483	190	13	relation	relation	NOUN
alces-483	190	14	to	to	ADP
alces-483	190	15	environment	environment	NOUN
alces-483	190	16	)	)	PUNCT
alces-483	190	17	,	,	PUNCT
alces-483	190	18	both	both	PRON
alces-483	190	19	for	for	ADP
alces-483	190	20	ungulates	ungulate	NOUN
alces-483	190	21	and	and	CCONJ
alces-483	190	22	other	other	ADJ
alces-483	190	23	animals	animal	NOUN
alces-483	190	24	in	in	ADP
alces-483	190	25	general	general	ADJ
alces-483	190	26	,	,	PUNCT
alces-483	190	27	but	but	CCONJ
alces-483	190	28	these	these	PRON
alces-483	190	29	lie	lie	VERB
alces-483	190	30	at	at	ADP
alces-483	190	31	the	the	DET
alces-483	190	32	core	core	NOUN
alces-483	190	33	of	of	ADP
alces-483	190	34	our	our	PRON
alces-483	190	35	understanding	understanding	NOUN
alces-483	190	36	of	of	ADP
alces-483	190	37	population	population	NOUN
alces-483	190	38	regulation	regulation	NOUN
alces-483	190	39	(	(	PUNCT
alces-483	190	40	lundberg	lundberg	PROPN
alces-483	190	41	et	et	PROPN
alces-483	190	42	al	al	PROPN
alces-483	190	43	.	.	PROPN
alces-483	190	44	2000	2000	NUM
alces-483	190	45	)	)	PUNCT
alces-483	190	46	.	.	PUNCT
alces-483	191	1	mechanisms	mechanism	NOUN
alces-483	191	2	underlying	underlie	VERB
alces-483	191	3	population	population	NOUN
alces-483	191	4	variability	variability	NOUN
alces-483	191	5	will	will	AUX
alces-483	191	6	be	be	AUX
alces-483	191	7	elucidated	elucidate	VERB
alces-483	191	8	only	only	ADV
alces-483	191	9	if	if	SCONJ
alces-483	191	10	appropriate	appropriate	ADJ
alces-483	191	11	response	response	NOUN
alces-483	191	12	variables	variable	NOUN
alces-483	191	13	are	be	AUX
alces-483	191	14	studied	study	VERB
alces-483	191	15	at	at	ADP
alces-483	191	16	appropriate	appropriate	ADJ
alces-483	191	17	scales	scale	NOUN
alces-483	191	18	(	(	PUNCT
alces-483	191	19	schmitz	schmitz	PROPN
alces-483	191	20	et	et	PROPN
alces-483	191	21	al	al	PROPN
alces-483	191	22	.	.	PROPN
alces-483	191	23	2000	2000	NUM
alces-483	191	24	)	)	PUNCT
alces-483	191	25	.	.	PUNCT
alces-483	192	1	long	long	ADJ
alces-483	192	2	response	response	NOUN
alces-483	192	3	times	time	NOUN
alces-483	192	4	may	may	AUX
alces-483	192	5	induce	induce	VERB
alces-483	192	6	long	long	ADJ
alces-483	192	7	lags	lag	NOUN
alces-483	192	8	;	;	PUNCT
alces-483	192	9	ungulate	ungulate	ADJ
alces-483	192	10	dynamics	dynamic	NOUN
alces-483	192	11	are	be	AUX
alces-483	192	12	drawn	draw	VERB
alces-483	192	13	-	-	PUNCT
alces-483	192	14	out	out	NOUN
alces-483	192	15	over	over	ADP
alces-483	192	16	decades	decade	NOUN
alces-483	192	17	,	,	PUNCT
alces-483	192	18	and	and	CCONJ
alces-483	192	19	response	response	NOUN
alces-483	192	20	times	time	NOUN
alces-483	192	21	for	for	ADP
alces-483	192	22	woody	woody	NOUN
alces-483	192	23	vegetation	vegetation	NOUN
alces-483	192	24	are	be	AUX
alces-483	192	25	even	even	ADV
alces-483	192	26	longer	long	ADV
alces-483	192	27	(	(	PUNCT
alces-483	192	28	holt	holt	PROPN
alces-483	192	29	2000	2000	NUM
alces-483	192	30	)	)	PUNCT
alces-483	192	31	.	.	PUNCT
alces-483	193	1	the	the	DET
alces-483	193	2	scale	scale	NOUN
alces-483	193	3	of	of	ADP
alces-483	193	4	study	study	NOUN
alces-483	193	5	impacts	impact	VERB
alces-483	193	6	interpretation	interpretation	NOUN
alces-483	193	7	of	of	ADP
alces-483	193	8	trophic	trophic	ADJ
alces-483	193	9	interactions	interaction	NOUN
alces-483	193	10	as	as	ADV
alces-483	193	11	well	well	ADV
alces-483	193	12	as	as	ADP
alces-483	193	13	dynamics	dynamic	NOUN
alces-483	193	14	of	of	ADP
alces-483	193	15	single	single	ADJ
alces-483	193	16	-	-	PUNCT
alces-483	193	17	species	specie	NOUN
alces-483	193	18	populations	population	NOUN
alces-483	193	19	(	(	PUNCT
alces-483	193	20	wiens	wien	NOUN
alces-483	193	21	1989	1989	NUM
alces-483	193	22	)	)	PUNCT
alces-483	193	23	.	.	PUNCT
alces-483	194	1	the	the	DET
alces-483	194	2	isle	isle	PROPN
alces-483	194	3	royale	royale	PROPN
alces-483	194	4	case	case	NOUN
alces-483	194	5	history	history	NOUN
alces-483	194	6	provides	provide	VERB
alces-483	194	7	a	a	DET
alces-483	194	8	particularly	particularly	ADV
alces-483	194	9	compelling	compelling	ADJ
alces-483	194	10	case	case	NOUN
alces-483	194	11	for	for	ADP
alces-483	194	12	temporal	temporal	ADJ
alces-483	194	13	variation	variation	NOUN
alces-483	194	14	in	in	ADP
alces-483	194	15	predator	predator	NOUN
alces-483	194	16	-	-	PUNCT
alces-483	194	17	prey	prey	NOUN
alces-483	194	18	interaction	interaction	NOUN
alces-483	194	19	,	,	PUNCT
alces-483	194	20	which	which	PRON
alces-483	194	21	can	can	AUX
alces-483	194	22	now	now	ADV
alces-483	194	23	be	be	AUX
alces-483	194	24	anticipated	anticipate	VERB
alces-483	194	25	for	for	ADP
alces-483	194	26	any	any	DET
alces-483	194	27	regularly	regularly	ADV
alces-483	194	28	fluctuating	fluctuate	VERB
alces-483	194	29	system	system	NOUN
alces-483	194	30	(	(	PUNCT
alces-483	194	31	sinclair	sinclair	NOUN
alces-483	194	32	et	et	PROPN
alces-483	194	33	al	al	PROPN
alces-483	194	34	.	.	PROPN
alces-483	194	35	2000	2000	NUM
alces-483	194	36	)	)	PUNCT
alces-483	194	37	.	.	PUNCT
alces-483	195	1	comparisons	comparison	NOUN
alces-483	195	2	of	of	ADP
alces-483	195	3	wolf	wolf	PROPN
alces-483	195	4	and	and	CCONJ
alces-483	195	5	moose	moose	NOUN
alces-483	195	6	status	status	NOUN
alces-483	195	7	over	over	ADP
alces-483	195	8	geographical	geographical	ADJ
alces-483	195	9	scales	scale	NOUN
alces-483	195	10	may	may	AUX
alces-483	195	11	not	not	PART
alces-483	195	12	always	always	ADV
alces-483	195	13	provide	provide	VERB
alces-483	195	14	mechanistic	mechanistic	ADJ
alces-483	195	15	insight	insight	NOUN
alces-483	195	16	into	into	ADP
alces-483	195	17	predatorprey	predatorprey	PROPN
alces-483	195	18	dynamics	dynamic	NOUN
alces-483	195	19	.	.	PUNCT
alces-483	196	1	for	for	ADP
alces-483	196	2	example	example	NOUN
alces-483	196	3	,	,	PUNCT
alces-483	196	4	a	a	DET
alces-483	196	5	“	"	PUNCT
alces-483	196	6	global	global	ADJ
alces-483	196	7	”	"	PUNCT
alces-483	196	8	correlation	correlation	NOUN
alces-483	196	9	between	between	ADP
alces-483	196	10	average	average	ADJ
alces-483	196	11	wolf	wolf	NOUN
alces-483	196	12	and	and	CCONJ
alces-483	196	13	prey	prey	PROPN
alces-483	196	14	density	density	NOUN
alces-483	196	15	exists	exist	VERB
alces-483	196	16	over	over	ADP
alces-483	196	17	a	a	DET
alces-483	196	18	wide	wide	ADJ
alces-483	196	19	range	range	NOUN
alces-483	196	20	of	of	ADP
alces-483	196	21	prey	prey	NOUN
alces-483	196	22	densities	density	NOUN
alces-483	196	23	(	(	PUNCT
alces-483	196	24	fuller	full	ADJ
alces-483	196	25	1989	1989	NUM
alces-483	196	26	)	)	PUNCT
alces-483	196	27	.	.	PUNCT
alces-483	197	1	this	this	PRON
alces-483	197	2	is	be	AUX
alces-483	197	3	widely	widely	ADV
alces-483	197	4	interpreted	interpret	VERB
alces-483	197	5	as	as	ADP
alces-483	197	6	the	the	DET
alces-483	197	7	“	"	PUNCT
alces-483	197	8	numerical	numerical	ADJ
alces-483	197	9	response	response	NOUN
alces-483	197	10	”	"	PUNCT
alces-483	197	11	of	of	ADP
alces-483	197	12	the	the	DET
alces-483	197	13	wolf	wolf	NOUN
alces-483	197	14	to	to	ADP
alces-483	197	15	fluctuations	fluctuation	NOUN
alces-483	197	16	in	in	ADP
alces-483	197	17	prey	prey	PROPN
alces-483	197	18	density	density	NOUN
alces-483	197	19	(	(	PUNCT
alces-483	197	20	messier	messy	ADJ
alces-483	197	21	1994	1994	NUM
alces-483	197	22	)	)	PUNCT
alces-483	197	23	,	,	PUNCT
alces-483	197	24	even	even	ADV
alces-483	197	25	as	as	ADP
alces-483	197	26	an	an	DET
alces-483	197	27	indication	indication	NOUN
alces-483	197	28	of	of	ADP
alces-483	197	29	ratio	ratio	NOUN
alces-483	197	30	-	-	PUNCT
alces-483	197	31	dependent	dependent	ADJ
alces-483	197	32	predation	predation	NOUN
alces-483	197	33	(	(	PUNCT
alces-483	197	34	arditi	arditi	INTJ
alces-483	197	35	et	et	PROPN
alces-483	197	36	al	al	PROPN
alces-483	197	37	.	.	PROPN
alces-483	197	38	1991	1991	NUM
alces-483	197	39	)	)	PUNCT
alces-483	197	40	,	,	PUNCT
alces-483	197	41	but	but	CCONJ
alces-483	197	42	the	the	DET
alces-483	197	43	temporal	temporal	ADJ
alces-483	197	44	trajectory	trajectory	NOUN
alces-483	197	45	followed	follow	VERB
alces-483	197	46	by	by	ADP
alces-483	197	47	local	local	ADJ
alces-483	197	48	populations	population	NOUN
alces-483	197	49	conforms	conform	VERB
alces-483	197	50	poorly	poorly	ADV
alces-483	197	51	to	to	ADP
alces-483	197	52	that	that	PRON
alces-483	197	53	which	which	PRON
alces-483	197	54	is	be	AUX
alces-483	197	55	extrapolated	extrapolate	VERB
alces-483	197	56	from	from	ADP
alces-483	197	57	the	the	DET
alces-483	197	58	large	large	ADJ
alces-483	197	59	spatial	spatial	NOUN
alces-483	197	60	(	(	PUNCT
alces-483	197	61	but	but	CCONJ
alces-483	197	62	temporally	temporally	ADV
alces-483	197	63	static	static	ADJ
alces-483	197	64	)	)	PUNCT
alces-483	197	65	global	global	ADJ
alces-483	197	66	pattern	pattern	NOUN
alces-483	197	67	.	.	PUNCT
alces-483	198	1	the	the	DET
alces-483	198	2	long	long	ADJ
alces-483	198	3	lives	life	NOUN
alces-483	198	4	and	and	CCONJ
alces-483	198	5	long	long	ADJ
alces-483	198	6	lags	lag	NOUN
alces-483	198	7	of	of	ADP
alces-483	198	8	large	large	ADJ
alces-483	198	9	mammals	mammal	NOUN
alces-483	198	10	may	may	AUX
alces-483	198	11	contribute	contribute	VERB
alces-483	198	12	to	to	ADP
alces-483	198	13	the	the	DET
alces-483	198	14	poor	poor	ADJ
alces-483	198	15	match	match	NOUN
alces-483	198	16	.	.	PUNCT
alces-483	199	1	what	what	PRON
alces-483	199	2	happens	happen	VERB
alces-483	199	3	to	to	ADP
alces-483	199	4	the	the	DET
alces-483	199	5	validity	validity	NOUN
alces-483	199	6	of	of	ADP
alces-483	199	7	models	model	NOUN
alces-483	199	8	(	(	PUNCT
alces-483	199	9	messier	messy	ADJ
alces-483	199	10	1994	1994	NUM
alces-483	199	11	)	)	PUNCT
alces-483	199	12	and	and	CCONJ
alces-483	199	13	management	management	NOUN
alces-483	199	14	programs	program	NOUN
alces-483	199	15	(	(	PUNCT
alces-483	199	16	gasaway	gasaway	NOUN
alces-483	199	17	et	et	PROPN
alces-483	199	18	al	al	PROPN
alces-483	199	19	.	.	PROPN
alces-483	199	20	1983	1983	NUM
alces-483	199	21	)	)	PUNCT
alces-483	199	22	when	when	SCONJ
alces-483	199	23	we	we	PRON
alces-483	199	24	ignore	ignore	VERB
alces-483	199	25	(	(	PUNCT
alces-483	199	26	lag	lag	NOUN
alces-483	199	27	-	-	PUNCT
alces-483	199	28	induced	induced	ADJ
alces-483	199	29	)	)	PUNCT
alces-483	199	30	temporal	temporal	ADJ
alces-483	199	31	dynamics	dynamic	NOUN
alces-483	199	32	?	?	PUNCT
alces-483	200	1	large	large	ADJ
alces-483	200	2	-	-	PUNCT
alces-483	200	3	scale	scale	NOUN
alces-483	200	4	geographic	geographic	ADJ
alces-483	200	5	comparisons	comparison	NOUN
alces-483	200	6	that	that	PRON
alces-483	200	7	ignore	ignore	VERB
alces-483	200	8	all	all	DET
alces-483	200	9	temporal	temporal	ADJ
alces-483	200	10	dynamics	dynamic	NOUN
alces-483	200	11	have	have	AUX
alces-483	200	12	also	also	ADV
alces-483	200	13	been	be	AUX
alces-483	200	14	used	use	VERB
alces-483	200	15	to	to	PART
alces-483	200	16	assess	assess	VERB
alces-483	200	17	predator	predator	NOUN
alces-483	200	18	-	-	PUNCT
alces-483	200	19	prey	prey	NOUN
alces-483	200	20	models	model	NOUN
alces-483	200	21	of	of	ADP
alces-483	200	22	temporal	temporal	ADJ
alces-483	200	23	dynamics	dynamic	NOUN
alces-483	200	24	.	.	PUNCT
alces-483	201	1	in	in	ADP
alces-483	201	2	a	a	DET
alces-483	201	3	study	study	NOUN
alces-483	201	4	from	from	ADP
alces-483	201	5	québec	québec	PROPN
alces-483	201	6	(	(	PUNCT
alces-483	201	7	crête	crête	ADJ
alces-483	201	8	and	and	CCONJ
alces-483	201	9	manseau	manseau	NOUN
alces-483	201	10	1996	1996	NUM
alces-483	201	11	)	)	PUNCT
alces-483	201	12	,	,	PUNCT
alces-483	201	13	spatial	spatial	ADJ
alces-483	201	14	patterns	pattern	NOUN
alces-483	201	15	were	be	AUX
alces-483	201	16	used	use	VERB
alces-483	201	17	to	to	PART
alces-483	201	18	support	support	VERB
alces-483	201	19	prey	prey	NOUN
alces-483	201	20	-	-	PUNCT
alces-483	201	21	dependent	dependent	ADJ
alces-483	201	22	models	model	NOUN
alces-483	201	23	of	of	ADP
alces-483	201	24	predation	predation	NOUN
alces-483	201	25	(	(	PUNCT
alces-483	201	26	sensu	sensu	NOUN
alces-483	201	27	,	,	PUNCT
alces-483	201	28	hairston	hairston	PROPN
alces-483	201	29	et	et	PROPN
alces-483	201	30	al	al	PROPN
alces-483	201	31	.	.	PROPN
alces-483	201	32	1960	1960	NUM
alces-483	201	33	)	)	PUNCT
alces-483	201	34	and	and	CCONJ
alces-483	201	35	to	to	PART
alces-483	201	36	reject	reject	VERB
alces-483	201	37	ratio	ratio	NOUN
alces-483	201	38	-	-	PUNCT
alces-483	201	39	dependent	dependent	ADJ
alces-483	201	40	models	model	NOUN
alces-483	201	41	of	of	ADP
alces-483	201	42	predation	predation	NOUN
alces-483	201	43	(	(	PUNCT
alces-483	201	44	sensu	sensu	NOUN
alces-483	201	45	,	,	PUNCT
alces-483	201	46	arditi	arditi	NOUN
alces-483	201	47	and	and	CCONJ
alces-483	201	48	ginzburg	ginzburg	NOUN
alces-483	201	49	1989	1989	NUM
alces-483	201	50	)	)	PUNCT
alces-483	201	51	.	.	PUNCT
alces-483	202	1	however	however	ADV
alces-483	202	2	,	,	PUNCT
alces-483	202	3	it	it	PRON
alces-483	202	4	seems	seem	VERB
alces-483	202	5	dubious	dubious	ADJ
alces-483	202	6	to	to	PART
alces-483	202	7	infer	infer	VERB
alces-483	202	8	processes	process	NOUN
alces-483	202	9	that	that	PRON
alces-483	202	10	occur	occur	VERB
alces-483	202	11	in	in	ADP
alces-483	202	12	one	one	NUM
alces-483	202	13	dimension	dimension	NOUN
alces-483	202	14	(	(	PUNCT
alces-483	202	15	time	time	NOUN
alces-483	202	16	)	)	PUNCT
alces-483	202	17	from	from	ADP
alces-483	202	18	observations	observation	NOUN
alces-483	202	19	made	make	VERB
alces-483	202	20	in	in	ADP
alces-483	202	21	another	another	DET
alces-483	202	22	dimension	dimension	NOUN
alces-483	202	23	(	(	PUNCT
alces-483	202	24	space	space	NOUN
alces-483	202	25	)	)	PUNCT
alces-483	202	26	(	(	PUNCT
alces-483	202	27	e.g.	e.g.	ADV
alces-483	202	28	,	,	PUNCT
alces-483	202	29	see	see	VERB
alces-483	202	30	also	also	ADV
alces-483	202	31	abrams	abrams	PROPN
alces-483	202	32	1994	1994	NUM
alces-483	202	33	,	,	PUNCT
alces-483	202	34	lundberg	lundberg	PROPN
alces-483	202	35	and	and	CCONJ
alces-483	202	36	fryxell	fryxell	NOUN
alces-483	202	37	1995	1995	NUM
alces-483	202	38	,	,	PUNCT
alces-483	202	39	abrams	abrams	NOUN
alces-483	202	40	and	and	CCONJ
alces-483	202	41	ginzburg	ginzburg	NOUN
alces-483	202	42	2000	2000	NUM
alces-483	202	43	)	)	PUNCT
alces-483	202	44	.	.	PUNCT
alces-483	203	1	in	in	ADP
alces-483	203	2	fact	fact	NOUN
alces-483	203	3	,	,	PUNCT
alces-483	203	4	our	our	PRON
alces-483	203	5	analysis	analysis	NOUN
alces-483	203	6	of	of	ADP
alces-483	203	7	the	the	DET
alces-483	203	8	temporal	temporal	ADJ
alces-483	203	9	dynamics	dynamic	NOUN
alces-483	203	10	of	of	ADP
alces-483	203	11	isle	isle	PROPN
alces-483	203	12	royale	royale	PROPN
alces-483	203	13	wolves	wolf	NOUN
alces-483	203	14	and	and	CCONJ
alces-483	203	15	moose	moose	NOUN
alces-483	203	16	appear	appear	VERB
alces-483	203	17	to	to	PART
alces-483	203	18	support	support	VERB
alces-483	203	19	ratio	ratio	NOUN
alces-483	203	20	-	-	PUNCT
alces-483	203	21	dependent	dependent	ADJ
alces-483	203	22	predation	predation	NOUN
alces-483	203	23	(	(	PUNCT
alces-483	203	24	vucetich	vucetich	NOUN
alces-483	203	25	et	et	PROPN
alces-483	203	26	al	al	PROPN
alces-483	203	27	.	.	PROPN
alces-483	203	28	2002	2002	NUM
alces-483	203	29	)	)	PUNCT
alces-483	203	30	.	.	PUNCT
alces-483	204	1	the	the	DET
alces-483	204	2	ecology	ecology	NOUN
alces-483	204	3	of	of	ADP
alces-483	204	4	moose	moose	NOUN
alces-483	204	5	at	at	ADP
alces-483	204	6	isle	isle	PROPN
alces-483	204	7	royale	royale	PROPN
alces-483	204	8	includes	include	VERB
alces-483	204	9	both	both	CCONJ
alces-483	204	10	slow	slow	ADJ
alces-483	204	11	and	and	CCONJ
alces-483	204	12	fast	fast	ADJ
alces-483	204	13	ecological	ecological	ADJ
alces-483	204	14	processes	process	NOUN
alces-483	204	15	,	,	PUNCT
alces-483	204	16	operating	operate	VERB
alces-483	204	17	at	at	ADP
alces-483	204	18	large	large	ADJ
alces-483	204	19	and	and	CCONJ
alces-483	204	20	small	small	ADJ
alces-483	204	21	spatial	spatial	ADJ
alces-483	204	22	scales	scale	NOUN
alces-483	204	23	.	.	PUNCT
alces-483	205	1	moose	moose	PROPN
alces-483	205	2	have	have	AUX
alces-483	205	3	virtually	virtually	ADV
alces-483	205	4	eliminated	eliminate	VERB
alces-483	205	5	some	some	DET
alces-483	205	6	plant	plant	NOUN
alces-483	205	7	species	specie	NOUN
alces-483	205	8	from	from	ADP
alces-483	205	9	the	the	DET
alces-483	205	10	forest	forest	NOUN
alces-483	205	11	(	(	PUNCT
alces-483	205	12	e.g.	e.g.	ADV
alces-483	205	13	,	,	PUNCT
alces-483	205	14	taxus	taxus	ADV
alces-483	205	15	)	)	PUNCT
alces-483	205	16	,	,	PUNCT
alces-483	205	17	and	and	CCONJ
alces-483	205	18	intensive	intensive	ADJ
alces-483	205	19	foraging	foraging	NOUN
alces-483	205	20	may	may	AUX
alces-483	205	21	eliminate	eliminate	VERB
alces-483	205	22	regeneration	regeneration	NOUN
alces-483	205	23	of	of	ADP
alces-483	205	24	many	many	ADJ
alces-483	205	25	species	specie	NOUN
alces-483	205	26	(	(	PUNCT
alces-483	205	27	e.g.	e.g.	ADV
alces-483	205	28	,	,	PUNCT
alces-483	205	29	abies	abie	NOUN
alces-483	205	30	)	)	PUNCT
alces-483	205	31	in	in	ADP
alces-483	205	32	the	the	DET
alces-483	205	33	tree	tree	NOUN
alces-483	205	34	layer	layer	NOUN
alces-483	205	35	.	.	PUNCT
alces-483	206	1	mesoscale	mesoscale	ADJ
alces-483	206	2	dynamics	dynamic	NOUN
alces-483	206	3	are	be	AUX
alces-483	206	4	dominated	dominate	VERB
alces-483	206	5	by	by	ADP
alces-483	206	6	cyclical	cyclical	ADJ
alces-483	206	7	fluctuations	fluctuation	NOUN
alces-483	206	8	in	in	ADP
alces-483	206	9	moose	moose	NOUN
alces-483	206	10	and	and	CCONJ
alces-483	206	11	wolves	wolf	NOUN
alces-483	206	12	with	with	ADP
alces-483	206	13	a	a	DET
alces-483	206	14	duration	duration	NOUN
alces-483	206	15	of	of	ADP
alces-483	206	16	about	about	ADV
alces-483	206	17	2	2	NUM
alces-483	206	18	decades	decade	NOUN
alces-483	206	19	,	,	PUNCT
alces-483	206	20	a	a	DET
alces-483	206	21	product	product	NOUN
alces-483	206	22	ultimately	ultimately	ADV
alces-483	206	23	of	of	ADP
alces-483	206	24	generation	generation	NOUN
alces-483	206	25	time	time	NOUN
alces-483	206	26	for	for	ADP
alces-483	206	27	predator	predator	NOUN
alces-483	206	28	and	and	CCONJ
alces-483	206	29	prey	prey	NOUN
alces-483	206	30	.	.	PUNCT
alces-483	207	1	yet	yet	CCONJ
alces-483	207	2	the	the	DET
alces-483	207	3	forest	forest	NOUN
alces-483	207	4	itself	itself	PRON
alces-483	207	5	has	have	AUX
alces-483	207	6	not	not	PART
alces-483	207	7	equilibrated	equilibrate	VERB
alces-483	207	8	following	follow	VERB
alces-483	207	9	the	the	DET
alces-483	207	10	arrival	arrival	NOUN
alces-483	207	11	of	of	ADP
alces-483	207	12	moose	moose	NOUN
alces-483	207	13	a	a	DET
alces-483	207	14	century	century	NOUN
alces-483	207	15	ago	ago	ADV
alces-483	207	16	–	–	PUNCT
alces-483	207	17	at	at	ADP
alces-483	207	18	the	the	DET
alces-483	207	19	island	island	NOUN
alces-483	207	20	’s	’s	PART
alces-483	207	21	west	west	PROPN
alces-483	207	22	end	end	VERB
alces-483	207	23	old	old	ADJ
alces-483	207	24	fir	fir	NOUN
alces-483	207	25	trees	tree	NOUN
alces-483	207	26	that	that	PRON
alces-483	207	27	established	establish	VERB
alces-483	207	28	in	in	ADP
alces-483	207	29	the	the	DET
alces-483	207	30	canopy	canopy	NOUN
alces-483	207	31	before	before	SCONJ
alces-483	207	32	moose	moose	PROPN
alces-483	207	33	arrived	arrive	VERB
alces-483	207	34	are	be	AUX
alces-483	207	35	now	now	ADV
alces-483	207	36	dying	die	VERB
alces-483	207	37	of	of	ADP
alces-483	207	38	old	old	ADJ
alces-483	207	39	age	age	NOUN
alces-483	207	40	without	without	ADP
alces-483	207	41	replacement	replacement	NOUN
alces-483	207	42	,	,	PUNCT
alces-483	207	43	and	and	CCONJ
alces-483	207	44	at	at	ADP
alces-483	207	45	the	the	DET
alces-483	207	46	east	east	PROPN
alces-483	207	47	end	end	NOUN
alces-483	207	48	spruce	spruce	NOUN
alces-483	207	49	-	-	PUNCT
alces-483	207	50	fir	fir	NOUN
alces-483	207	51	stands	stand	NOUN
alces-483	207	52	have	have	AUX
alces-483	207	53	only	only	ADV
alces-483	207	54	recently	recently	ADV
alces-483	207	55	emerged	emerge	VERB
alces-483	207	56	in	in	ADP
alces-483	207	57	extensive	extensive	ADJ
alces-483	207	58	19th	19th	ADJ
alces-483	207	59	century	century	NOUN
alces-483	207	60	burns	burn	NOUN
alces-483	207	61	initiated	initiate	VERB
alces-483	207	62	by	by	ADP
alces-483	207	63	mineral	mineral	NOUN
alces-483	207	64	prospectors	prospector	NOUN
alces-483	207	65	–	–	PUNCT
alces-483	207	66	while	while	SCONJ
alces-483	207	67	both	both	DET
alces-483	207	68	trends	trend	NOUN
alces-483	207	69	are	be	AUX
alces-483	207	70	superficially	superficially	ADV
alces-483	207	71	associated	associate	VERB
alces-483	207	72	with	with	ADP
alces-483	207	73	less	less	ADJ
alces-483	207	74	forage	forage	NOUN
alces-483	207	75	for	for	ADP
alces-483	207	76	moose	moose	NOUN
alces-483	207	77	,	,	PUNCT
alces-483	207	78	future	future	ADJ
alces-483	207	79	dynamics	dynamic	NOUN
alces-483	207	80	of	of	ADP
alces-483	207	81	wolves	wolf	NOUN
alces-483	207	82	and	and	CCONJ
alces-483	207	83	moose	moose	NOUN
alces-483	207	84	in	in	ADP
alces-483	207	85	response	response	NOUN
alces-483	207	86	to	to	ADP
alces-483	207	87	these	these	DET
alces-483	207	88	very	very	ADV
alces-483	207	89	slow	slow	ADJ
alces-483	207	90	changes	change	NOUN
alces-483	207	91	in	in	ADP
alces-483	207	92	vegetation	vegetation	NOUN
alces-483	207	93	are	be	AUX
alces-483	207	94	not	not	PART
alces-483	207	95	readily	readily	ADV
alces-483	207	96	predicted	predict	VERB
alces-483	207	97	.	.	PUNCT
alces-483	208	1	acknowledgements	acknowledgement	NOUN
alces-483	208	2	long	long	ADJ
alces-483	208	3	-	-	PUNCT
alces-483	208	4	term	term	NOUN
alces-483	208	5	studies	study	NOUN
alces-483	208	6	of	of	ADP
alces-483	208	7	population	population	NOUN
alces-483	208	8	dynamics	dynamic	NOUN
alces-483	208	9	of	of	ADP
alces-483	208	10	wolves	wolf	NOUN
alces-483	208	11	,	,	PUNCT
alces-483	208	12	moose	moose	NOUN
alces-483	208	13	and	and	CCONJ
alces-483	208	14	balsam	balsam	NOUN
alces-483	208	15	fir	fir	NOUN
alces-483	208	16	on	on	ADP
alces-483	208	17	isle	isle	PROPN
alces-483	208	18	alces	alces	PROPN
alces-483	208	19	vol	vol	NOUN
alces-483	208	20	.	.	PROPN
alces-483	208	21	39	39	NUM
alces-483	208	22	,	,	PUNCT
alces-483	208	23	2003	2003	NUM
alces-483	208	24	peterson	peterson	PROPN
alces-483	208	25	et	et	PROPN
alces-483	208	26	al	al	PROPN
alces-483	208	27	.	.	PROPN
alces-483	208	28	–	–	PUNCT
alces-483	208	29	space	space	NOUN
alces-483	208	30	and	and	CCONJ
alces-483	208	31	time	time	NOUN
alces-483	208	32	in	in	ADP
alces-483	208	33	predation	predation	NOUN
alces-483	208	34	dynamics	dynamic	NOUN
alces-483	208	35	225	225	NUM
alces-483	208	36	royale	royale	NOUN
alces-483	208	37	have	have	AUX
alces-483	208	38	been	be	AUX
alces-483	208	39	supported	support	VERB
alces-483	208	40	primarily	primarily	ADV
alces-483	208	41	by	by	ADP
alces-483	208	42	isle	isle	PROPN
alces-483	208	43	royale	royale	PROPN
alces-483	208	44	national	national	PROPN
alces-483	208	45	park	park	PROPN
alces-483	208	46	,	,	PUNCT
alces-483	208	47	the	the	DET
alces-483	208	48	national	national	PROPN
alces-483	208	49	science	science	PROPN
alces-483	208	50	foundation	foundation	PROPN
alces-483	208	51	(	(	PUNCT
alces-483	208	52	deb-9903671	deb-9903671	PROPN
alces-483	208	53	)	)	PUNCT
alces-483	208	54	,	,	PUNCT
alces-483	208	55	earthwatch	earthwatch	PROPN
alces-483	208	56	,	,	PUNCT
alces-483	208	57	inc	inc	PROPN
alces-483	208	58	.	.	PROPN
alces-483	208	59	,	,	PUNCT
alces-483	208	60	and	and	CCONJ
alces-483	208	61	numerous	numerous	ADJ
alces-483	208	62	private	private	ADJ
alces-483	208	63	donors	donor	NOUN
alces-483	208	64	.	.	PUNCT
alces-483	209	1	references	reference	NOUN
alces-483	209	2	abrams	abram	NOUN
alces-483	209	3	,	,	PUNCT
alces-483	209	4	p.	p.	NOUN
alces-483	209	5	a.	a.	NOUN
alces-483	209	6	1994	1994	NUM
alces-483	209	7	.	.	PUNCT
alces-483	210	1	the	the	DET
alces-483	210	2	fallacies	fallacy	NOUN
alces-483	210	3	of	of	ADP
alces-483	210	4	“	"	PUNCT
alces-483	210	5	ratio	ratio	NOUN
alces-483	210	6	-	-	PUNCT
alces-483	210	7	dependent	dependent	ADJ
alces-483	210	8	”	"	PUNCT
alces-483	210	9	predation	predation	NOUN
alces-483	210	10	.	.	PUNCT
alces-483	211	1	ecology	ecology	NOUN
alces-483	211	2	75:1842	75:1842	NOUN
alces-483	211	3	-	-	SYM
alces-483	211	4	1850	1850	NUM
alces-483	211	5	.	.	PUNCT
alces-483	212	1	,	,	PUNCT
alces-483	212	2	and	and	CCONJ
alces-483	212	3	l.	l.	PROPN
alces-483	212	4	r.	r.	PROPN
alces-483	212	5	ginzburg	ginzburg	PROPN
alces-483	212	6	.	.	PUNCT
alces-483	213	1	2000	2000	NUM
alces-483	213	2	.	.	PUNCT
alces-483	214	1	the	the	DET
alces-483	214	2	nature	nature	NOUN
alces-483	214	3	of	of	ADP
alces-483	214	4	predation	predation	NOUN
alces-483	214	5	:	:	PUNCT
alces-483	214	6	prey	prey	PROPN
alces-483	214	7	dependent	dependent	ADV
alces-483	214	8	,	,	PUNCT
alces-483	214	9	ratio	ratio	NOUN
alces-483	214	10	dependent	dependent	ADJ
alces-483	214	11	or	or	CCONJ
alces-483	214	12	neither	neither	PRON
alces-483	214	13	?	?	PUNCT
alces-483	215	1	trends	trend	NOUN
alces-483	215	2	in	in	ADP
alces-483	215	3	ecology	ecology	NOUN
alces-483	215	4	and	and	CCONJ
alces-483	215	5	evolution	evolution	NOUN
alces-483	215	6	15:337	15:337	PROPN
alces-483	215	7	-	-	SYM
alces-483	215	8	341	341	NUM
alces-483	215	9	.	.	PUNCT
alces-483	216	1	arditi	arditi	PROPN
alces-483	216	2	,	,	PUNCT
alces-483	216	3	r.	r.	PROPN
alces-483	216	4	,	,	PUNCT
alces-483	216	5	and	and	CCONJ
alces-483	216	6	l.	l.	PROPN
alces-483	216	7	r.	r.	PROPN
alces-483	216	8	ginzburg	ginzburg	PROPN
alces-483	216	9	.	.	PUNCT
alces-483	217	1	1989	1989	NUM
alces-483	217	2	.	.	PUNCT
alces-483	218	1	coupling	couple	VERB
alces-483	218	2	in	in	ADP
alces-483	218	3	predator	predator	NOUN
alces-483	218	4	-	-	PUNCT
alces-483	218	5	prey	prey	NOUN
alces-483	218	6	dynamics	dynamic	NOUN
alces-483	218	7	:	:	PUNCT
alces-483	218	8	ratio	ratio	NOUN
alces-483	218	9	dependence	dependence	NOUN
alces-483	218	10	.	.	PUNCT
alces-483	219	1	journal	journal	NOUN
alces-483	219	2	of	of	ADP
alces-483	219	3	theoretical	theoretical	ADJ
alces-483	219	4	biology	biology	NOUN
alces-483	219	5	139:311	139:311	NOUN
alces-483	219	6	-	-	PUNCT
alces-483	219	7	326	326	NUM
alces-483	219	8	.	.	PUNCT
alces-483	219	9	,	,	PUNCT
alces-483	219	10	,	,	PUNCT
alces-483	219	11	and	and	CCONJ
alces-483	219	12	h.	h.	PROPN
alces-483	219	13	r.	r.	PROPN
alces-483	219	14	akçakaya	akçakaya	PROPN
alces-483	219	15	.	.	PROPN
alces-483	219	16	1991	1991	NUM
alces-483	219	17	.	.	PUNCT
alces-483	220	1	variation	variation	NOUN
alces-483	220	2	in	in	ADP
alces-483	220	3	plankton	plankton	ADJ
alces-483	220	4	densities	density	NOUN
alces-483	220	5	among	among	ADP
alces-483	220	6	lakes	lake	NOUN
alces-483	220	7	:	:	PUNCT
alces-483	220	8	a	a	DET
alces-483	220	9	case	case	NOUN
alces-483	220	10	for	for	ADP
alces-483	220	11	ratio	ratio	NOUN
alces-483	220	12	-	-	PUNCT
alces-483	220	13	dependent	dependent	ADJ
alces-483	220	14	predation	predation	NOUN
alces-483	220	15	models	model	NOUN
alces-483	220	16	.	.	PUNCT
alces-483	221	1	american	american	ADJ
alces-483	221	2	naturalist	naturalist	PROPN
alces-483	221	3	138:1287	138:1287	PROPN
alces-483	221	4	-	-	SYM
alces-483	221	5	1296	1296	NUM
alces-483	221	6	.	.	PUNCT
alces-483	222	1	bailey	bailey	NOUN
alces-483	222	2	,	,	PUNCT
alces-483	222	3	t.	t.	PROPN
alces-483	222	4	n.	n.	PROPN
alces-483	222	5	1978	1978	NUM
alces-483	222	6	.	.	PUNCT
alces-483	223	1	moose	moose	NOUN
alces-483	223	2	populations	population	NOUN
alces-483	223	3	on	on	ADP
alces-483	223	4	the	the	DET
alces-483	223	5	kenai	kenai	PROPN
alces-483	223	6	national	national	PROPN
alces-483	223	7	moose	moose	PROPN
alces-483	223	8	range	range	NOUN
alces-483	223	9	.	.	PUNCT
alces-483	224	1	proceedings	proceeding	NOUN
alces-483	224	2	of	of	ADP
alces-483	224	3	the	the	DET
alces-483	224	4	north	north	ADJ
alces-483	224	5	american	american	PROPN
alces-483	224	6	moose	moose	PROPN
alces-483	224	7	conference	conference	NOUN
alces-483	224	8	and	and	CCONJ
alces-483	224	9	workshop	workshop	NOUN
alces-483	224	10	14:1	14:1	NUM
alces-483	224	11	-	-	SYM
alces-483	224	12	20	20	NUM
alces-483	224	13	.	.	PUNCT
alces-483	225	1	ballard	ballard	PROPN
alces-483	225	2	,	,	PUNCT
alces-483	225	3	w.	w.	PROPN
alces-483	225	4	b.	b.	PROPN
alces-483	225	5	,	,	PUNCT
alces-483	225	6	and	and	CCONJ
alces-483	225	7	d.	d.	PROPN
alces-483	225	8	g.	g.	PROPN
alces-483	225	9	larsen	larsen	PROPN
alces-483	225	10	.	.	PROPN
alces-483	225	11	1987	1987	NUM
alces-483	225	12	.	.	PUNCT
alces-483	226	1	implications	implication	NOUN
alces-483	226	2	of	of	ADP
alces-483	226	3	predator	predator	NOUN
alces-483	226	4	-	-	PUNCT
alces-483	226	5	prey	prey	NOUN
alces-483	226	6	relationships	relationship	NOUN
alces-483	226	7	to	to	ADP
alces-483	226	8	moose	moose	NOUN
alces-483	226	9	management	management	NOUN
alces-483	226	10	.	.	PUNCT
alces-483	227	1	swedish	swedish	ADJ
alces-483	227	2	wildlife	wildlife	NOUN
alces-483	227	3	research	research	NOUN
alces-483	227	4	supplement	supplement	NOUN
alces-483	227	5	1:581602	1:581602	NUM
alces-483	227	6	.	.	PUNCT
alces-483	227	7	,	,	PUNCT
alces-483	227	8	and	and	CCONJ
alces-483	227	9	v.	v.	ADP
alces-483	227	10	van	van	PROPN
alces-483	227	11	ballenberghe	ballenberghe	NOUN
alces-483	227	12	.	.	PUNCT
alces-483	228	1	1998	1998	NUM
alces-483	228	2	.	.	PUNCT
alces-483	229	1	predator	predator	NOUN
alces-483	229	2	/	/	SYM
alces-483	229	3	prey	prey	NOUN
alces-483	229	4	relationships	relationship	NOUN
alces-483	229	5	.	.	PUNCT
alces-483	230	1	pages	page	NOUN
alces-483	230	2	247273	247273	NUM
alces-483	230	3	in	in	ADP
alces-483	230	4	a.	a.	PROPN
alces-483	230	5	w.	w.	PROPN
alces-483	230	6	franzmann	franzmann	PROPN
alces-483	230	7	and	and	CCONJ
alces-483	230	8	c.	c.	PROPN
alces-483	230	9	c.	c.	PROPN
alces-483	230	10	schwartz	schwartz	PROPN
alces-483	230	11	,	,	PUNCT
alces-483	230	12	editors	editor	NOUN
alces-483	230	13	.	.	PUNCT
alces-483	231	1	ecology	ecology	NOUN
alces-483	231	2	and	and	CCONJ
alces-483	231	3	management	management	NOUN
alces-483	231	4	of	of	ADP
alces-483	231	5	the	the	DET
alces-483	231	6	north	north	ADJ
alces-483	231	7	american	american	ADJ
alces-483	231	8	moose	moose	NOUN
alces-483	231	9	.	.	PUNCT
alces-483	232	1	smithsonian	smithsonian	PROPN
alces-483	232	2	institution	institution	PROPN
alces-483	232	3	press	press	PROPN
alces-483	232	4	,	,	PUNCT
alces-483	232	5	washington	washington	PROPN
alces-483	232	6	,	,	PUNCT
alces-483	232	7	d.c	d.c	PROPN
alces-483	232	8	.	.	PROPN
alces-483	232	9	,	,	PUNCT
alces-483	232	10	usa	usa	PROPN
alces-483	232	11	.	.	PROPN
alces-483	232	12	,	,	PUNCT
alces-483	232	13	j.	j.	PROPN
alces-483	232	14	s.	s.	PROPN
alces-483	232	15	whitman	whitman	PROPN
alces-483	232	16	,	,	PUNCT
alces-483	232	17	and	and	CCONJ
alces-483	232	18	c.	c.	PROPN
alces-483	232	19	l.	l.	PROPN
alces-483	232	20	gardner	gardner	PROPN
alces-483	232	21	.	.	PUNCT
alces-483	233	1	1987	1987	NUM
alces-483	233	2	.	.	PUNCT
alces-483	234	1	ecology	ecology	NOUN
alces-483	234	2	of	of	ADP
alces-483	234	3	an	an	DET
alces-483	234	4	exploited	exploit	VERB
alces-483	234	5	wolf	wolf	NOUN
alces-483	234	6	population	population	NOUN
alces-483	234	7	in	in	ADP
alces-483	234	8	south	south	ADJ
alces-483	234	9	-	-	PUNCT
alces-483	234	10	central	central	ADJ
alces-483	234	11	alaska	alaska	PROPN
alces-483	234	12	.	.	PUNCT
alces-483	235	1	wildlife	wildlife	NOUN
alces-483	235	2	monographs	monograph	NOUN
alces-483	235	3	98	98	NUM
alces-483	235	4	.	.	PUNCT
alces-483	236	1	bangs	bang	NOUN
alces-483	236	2	,	,	PUNCT
alces-483	236	3	e.	e.	PROPN
alces-483	236	4	,	,	PUNCT
alces-483	236	5	and	and	CCONJ
alces-483	236	6	t.	t.	PROPN
alces-483	236	7	n.	n.	PROPN
alces-483	236	8	bailey	bailey	PROPN
alces-483	236	9	.	.	PUNCT
alces-483	237	1	1980	1980	NUM
alces-483	237	2	.	.	PUNCT
alces-483	238	1	interrelationships	interrelationship	NOUN
alces-483	238	2	of	of	ADP
alces-483	238	3	weather	weather	NOUN
alces-483	238	4	,	,	PUNCT
alces-483	238	5	fire	fire	NOUN
alces-483	238	6	,	,	PUNCT
alces-483	238	7	and	and	CCONJ
alces-483	238	8	moose	moose	VERB
alces-483	238	9	on	on	ADP
alces-483	238	10	the	the	DET
alces-483	238	11	kenai	kenai	PROPN
alces-483	238	12	national	national	PROPN
alces-483	238	13	moose	moose	PROPN
alces-483	238	14	range	range	PROPN
alces-483	238	15	,	,	PUNCT
alces-483	238	16	alaska	alaska	PROPN
alces-483	238	17	.	.	PUNCT
alces-483	239	1	proceedings	proceeding	NOUN
alces-483	239	2	of	of	ADP
alces-483	239	3	the	the	DET
alces-483	239	4	north	north	ADJ
alces-483	239	5	american	american	PROPN
alces-483	239	6	moose	moose	PROPN
alces-483	239	7	conference	conference	NOUN
alces-483	239	8	and	and	CCONJ
alces-483	239	9	workshop	workshop	NOUN
alces-483	239	10	16:255	16:255	NUM
alces-483	239	11	-	-	SYM
alces-483	239	12	274	274	NUM
alces-483	239	13	.	.	PUNCT
alces-483	240	1	berg	berg	PROPN
alces-483	240	2	,	,	PUNCT
alces-483	240	3	w.	w.	PROPN
alces-483	240	4	e.	e.	PROPN
alces-483	240	5	,	,	PUNCT
alces-483	240	6	and	and	CCONJ
alces-483	240	7	d.	d.	PROPN
alces-483	240	8	w.	w.	PROPN
alces-483	240	9	kuehn	kuehn	PROPN
alces-483	240	10	.	.	PUNCT
alces-483	241	1	1982	1982	NUM
alces-483	241	2	.	.	PUNCT
alces-483	242	1	ecology	ecology	NOUN
alces-483	242	2	of	of	ADP
alces-483	242	3	wolves	wolf	NOUN
alces-483	242	4	in	in	ADP
alces-483	242	5	north	north	NOUN
alces-483	242	6	-	-	PUNCT
alces-483	242	7	central	central	ADJ
alces-483	242	8	minnesota	minnesota	NOUN
alces-483	242	9	.	.	PUNCT
alces-483	243	1	pages	page	NOUN
alces-483	243	2	229	229	NUM
alces-483	243	3	-	-	SYM
alces-483	243	4	247	247	NUM
alces-483	243	5	in	in	ADP
alces-483	243	6	f.	f.	PROPN
alces-483	243	7	h.	h.	PROPN
alces-483	243	8	harrington	harrington	PROPN
alces-483	243	9	and	and	CCONJ
alces-483	243	10	p.	p.	PROPN
alces-483	243	11	c.	c.	PROPN
alces-483	243	12	paquet	paquet	PROPN
alces-483	243	13	,	,	PUNCT
alces-483	243	14	editors	editor	NOUN
alces-483	243	15	.	.	PUNCT
alces-483	244	1	wolves	wolf	NOUN
alces-483	244	2	of	of	ADP
alces-483	244	3	the	the	DET
alces-483	244	4	world	world	NOUN
alces-483	244	5	:	:	PUNCT
alces-483	244	6	perspectives	perspective	NOUN
alces-483	244	7	of	of	ADP
alces-483	244	8	behavior	behavior	NOUN
alces-483	244	9	,	,	PUNCT
alces-483	244	10	ecology	ecology	NOUN
alces-483	244	11	,	,	PUNCT
alces-483	244	12	and	and	CCONJ
alces-483	244	13	conservation	conservation	NOUN
alces-483	244	14	.	.	PUNCT
alces-483	245	1	noyes	noyes	PROPN
alces-483	245	2	,	,	PUNCT
alces-483	245	3	park	park	PROPN
alces-483	245	4	ridge	ridge	PROPN
alces-483	245	5	,	,	PUNCT
alces-483	245	6	new	new	PROPN
alces-483	245	7	jersey	jersey	PROPN
alces-483	245	8	,	,	PUNCT
alces-483	245	9	usa	usa	PROPN
alces-483	245	10	.	.	PROPN
alces-483	245	11	bergerud	bergerud	PROPN
alces-483	245	12	,	,	PUNCT
alces-483	245	13	a.	a.	NOUN
alces-483	245	14	t.	t.	PROPN
alces-483	245	15	1980	1980	NUM
alces-483	245	16	.	.	PUNCT
alces-483	246	1	a	a	DET
alces-483	246	2	review	review	NOUN
alces-483	246	3	of	of	ADP
alces-483	246	4	the	the	DET
alces-483	246	5	population	population	NOUN
alces-483	246	6	dynamics	dynamic	NOUN
alces-483	246	7	of	of	ADP
alces-483	246	8	caribou	caribou	NOUN
alces-483	246	9	and	and	CCONJ
alces-483	246	10	wild	wild	ADJ
alces-483	246	11	reindeer	reindeer	NOUN
alces-483	246	12	in	in	ADP
alces-483	246	13	north	north	PROPN
alces-483	246	14	america	america	PROPN
alces-483	246	15	.	.	PUNCT
alces-483	247	1	proceedings	proceeding	NOUN
alces-483	247	2	of	of	ADP
alces-483	247	3	the	the	DET
alces-483	247	4	international	international	ADJ
alces-483	247	5	reindeer	reindeer	NOUN
alces-483	247	6	/	/	SYM
alces-483	247	7	caribou	caribou	PROPN
alces-483	247	8	symposium	symposium	NOUN
alces-483	247	9	2:556	2:556	PROPN
alces-483	247	10	-	-	SYM
alces-483	247	11	581	581	NUM
alces-483	247	12	.	.	PUNCT
alces-483	247	13	.	.	PUNCT
alces-483	248	1	1983a	1983a	PROPN
alces-483	248	2	.	.	PUNCT
alces-483	249	1	the	the	DET
alces-483	249	2	natural	natural	ADJ
alces-483	249	3	population	population	NOUN
alces-483	249	4	control	control	NOUN
alces-483	249	5	of	of	ADP
alces-483	249	6	caribou	caribou	PROPN
alces-483	249	7	.	.	PUNCT
alces-483	250	1	pages	page	NOUN
alces-483	250	2	14	14	NUM
alces-483	250	3	-	-	SYM
alces-483	250	4	61	61	NUM
alces-483	250	5	in	in	ADP
alces-483	250	6	f.	f.	PROPN
alces-483	250	7	l.	l.	PROPN
alces-483	250	8	bunnell	bunnell	PROPN
alces-483	250	9	,	,	PUNCT
alces-483	250	10	d.	d.	PROPN
alces-483	250	11	s.	s.	PROPN
alces-483	250	12	eastman	eastman	PROPN
alces-483	250	13	,	,	PUNCT
alces-483	250	14	and	and	CCONJ
alces-483	250	15	j.	j.	PROPN
alces-483	250	16	m.	m.	PROPN
alces-483	250	17	peek	peek	PROPN
alces-483	250	18	,	,	PUNCT
alces-483	250	19	editors	editor	NOUN
alces-483	250	20	.	.	PUNCT
alces-483	251	1	symposium	symposium	NOUN
alces-483	251	2	on	on	ADP
alces-483	251	3	natural	natural	ADJ
alces-483	251	4	regulation	regulation	NOUN
alces-483	251	5	of	of	ADP
alces-483	251	6	wildlife	wildlife	NOUN
alces-483	251	7	populations	population	NOUN
alces-483	251	8	.	.	PUNCT
alces-483	252	1	proceedings	proceeding	NOUN
alces-483	252	2	number	number	NOUN
alces-483	252	3	14	14	NUM
alces-483	252	4	.	.	PUNCT
alces-483	253	1	forest	forest	NOUN
alces-483	253	2	,	,	PUNCT
alces-483	253	3	wildlife	wildlife	NOUN
alces-483	253	4	and	and	CCONJ
alces-483	253	5	range	range	NOUN
alces-483	253	6	experiment	experiment	NOUN
alces-483	253	7	station	station	NOUN
alces-483	253	8	,	,	PUNCT
alces-483	253	9	university	university	PROPN
alces-483	253	10	of	of	ADP
alces-483	253	11	idaho	idaho	PROPN
alces-483	253	12	,	,	PUNCT
alces-483	253	13	moscow	moscow	PROPN
alces-483	253	14	.	.	PUNCT
alces-483	253	15	.	.	PUNCT
alces-483	254	1	1983b	1983b	NOUN
alces-483	254	2	.	.	PUNCT
alces-483	255	1	caribou	caribou	NOUN
alces-483	255	2	,	,	PUNCT
alces-483	255	3	wolves	wolf	NOUN
alces-483	255	4	and	and	CCONJ
alces-483	255	5	man	man	NOUN
alces-483	255	6	.	.	PUNCT
alces-483	256	1	trends	trend	NOUN
alces-483	256	2	in	in	ADP
alces-483	256	3	ecology	ecology	NOUN
alces-483	256	4	and	and	CCONJ
alces-483	256	5	evolution	evolution	NOUN
alces-483	256	6	3:6872	3:6872	NUM
alces-483	256	7	.	.	NOUN
alces-483	256	8	,	,	PUNCT
alces-483	256	9	and	and	CCONJ
alces-483	256	10	f.	f.	PROPN
alces-483	256	11	manuel	manuel	PROPN
alces-483	256	12	.	.	PUNCT
alces-483	257	1	1968	1968	NUM
alces-483	257	2	.	.	PUNCT
alces-483	258	1	moose	moose	PROPN
alces-483	258	2	damage	damage	NOUN
alces-483	258	3	to	to	ADP
alces-483	258	4	balsam	balsam	NOUN
alces-483	258	5	fir	fir	PROPN
alces-483	258	6	-	-	PUNCT
alces-483	258	7	white	white	ADJ
alces-483	258	8	birch	birch	NOUN
alces-483	258	9	forests	forest	NOUN
alces-483	258	10	in	in	ADP
alces-483	258	11	central	central	ADJ
alces-483	258	12	newfoundland	newfoundland	PROPN
alces-483	258	13	.	.	PUNCT
alces-483	259	1	journal	journal	PROPN
alces-483	259	2	of	of	ADP
alces-483	259	3	wildlife	wildlife	NOUN
alces-483	259	4	management	management	NOUN
alces-483	259	5	32:729	32:729	NUM
alces-483	259	6	-	-	SYM
alces-483	259	7	746	746	NUM
alces-483	259	8	.	.	PUNCT
alces-483	260	1	,	,	PUNCT
alces-483	260	2	w.	w.	PROPN
alces-483	260	3	wyett	wyett	PROPN
alces-483	260	4	,	,	PUNCT
alces-483	260	5	and	and	CCONJ
alces-483	260	6	b.	b.	PROPN
alces-483	260	7	snider	snider	PROPN
alces-483	260	8	.	.	PUNCT
alces-483	261	1	1983	1983	NUM
alces-483	261	2	.	.	PUNCT
alces-483	262	1	the	the	DET
alces-483	262	2	role	role	NOUN
alces-483	262	3	of	of	ADP
alces-483	262	4	wolf	wolf	PROPN
alces-483	262	5	predation	predation	NOUN
alces-483	262	6	in	in	ADP
alces-483	262	7	limiting	limit	VERB
alces-483	262	8	a	a	DET
alces-483	262	9	moose	moose	NOUN
alces-483	262	10	population	population	NOUN
alces-483	262	11	.	.	PUNCT
alces-483	263	1	journal	journal	PROPN
alces-483	263	2	of	of	ADP
alces-483	263	3	wildlife	wildlife	NOUN
alces-483	263	4	management	management	NOUN
alces-483	263	5	47:977	47:977	NUM
alces-483	263	6	-	-	SYM
alces-483	263	7	988	988	NUM
alces-483	263	8	.	.	PUNCT
alces-483	264	1	boyce	boyce	PROPN
alces-483	264	2	,	,	PUNCT
alces-483	264	3	m.	m.	NOUN
alces-483	264	4	s.	s.	PROPN
alces-483	264	5	,	,	PUNCT
alces-483	264	6	and	and	CCONJ
alces-483	264	7	e.	e.	PROPN
alces-483	264	8	m.	m.	PROPN
alces-483	264	9	anderson	anderson	PROPN
alces-483	264	10	.	.	PUNCT
alces-483	264	11	1999	1999	NUM
alces-483	264	12	.	.	PUNCT
alces-483	265	1	evaluating	evaluate	VERB
alces-483	265	2	the	the	DET
alces-483	265	3	role	role	NOUN
alces-483	265	4	of	of	ADP
alces-483	265	5	carnivores	carnivore	NOUN
alces-483	265	6	in	in	ADP
alces-483	265	7	the	the	DET
alces-483	265	8	greater	great	ADJ
alces-483	265	9	yellowstone	yellowstone	PROPN
alces-483	265	10	ecosystem	ecosystem	NOUN
alces-483	265	11	.	.	PUNCT
alces-483	266	1	pages	page	NOUN
alces-483	266	2	265	265	NUM
alces-483	266	3	-	-	SYM
alces-483	266	4	283	283	NUM
alces-483	266	5	in	in	ADP
alces-483	266	6	t.	t.	PROPN
alces-483	266	7	w.	w.	PROPN
alces-483	266	8	clark	clark	PROPN
alces-483	266	9	,	,	PUNCT
alces-483	266	10	a.	a.	PROPN
alces-483	266	11	p.	p.	PROPN
alces-483	266	12	curlee	curlee	PROPN
alces-483	266	13	,	,	PUNCT
alces-483	266	14	s.	s.	PROPN
alces-483	266	15	c.	c.	PROPN
alces-483	266	16	minta	minta	PROPN
alces-483	266	17	,	,	PUNCT
alces-483	266	18	and	and	CCONJ
alces-483	266	19	p.	p.	PROPN
alces-483	266	20	m.	m.	PROPN
alces-483	266	21	kareiva	kareiva	PROPN
alces-483	266	22	,	,	PUNCT
alces-483	266	23	editors	editor	NOUN
alces-483	266	24	.	.	PUNCT
alces-483	267	1	carnivores	carnivore	NOUN
alces-483	267	2	in	in	ADP
alces-483	267	3	ecosystems	ecosystem	NOUN
alces-483	267	4	:	:	PUNCT
alces-483	267	5	the	the	DET
alces-483	267	6	yellowstone	yellowstone	PROPN
alces-483	267	7	experience	experience	NOUN
alces-483	267	8	.	.	PUNCT
alces-483	268	1	yale	yale	PROPN
alces-483	268	2	university	university	PROPN
alces-483	268	3	press	press	NOUN
alces-483	268	4	,	,	PUNCT
alces-483	268	5	new	new	ADJ
alces-483	268	6	haven	haven	NOUN
alces-483	268	7	,	,	PUNCT
alces-483	268	8	connecticut	connecticut	PROPN
alces-483	268	9	,	,	PUNCT
alces-483	268	10	usa	usa	PROPN
alces-483	268	11	.	.	PROPN
alces-483	268	12	cairns	cairns	PROPN
alces-483	268	13	,	,	PUNCT
alces-483	268	14	a.	a.	PROPN
alces-483	268	15	l.	l.	PROPN
alces-483	268	16	,	,	PUNCT
alces-483	268	17	and	and	CCONJ
alces-483	268	18	e.	e.	PROPN
alces-483	268	19	s.	s.	PROPN
alces-483	268	20	telfer	telfer	PROPN
alces-483	268	21	.	.	PUNCT
alces-483	269	1	1980	1980	NUM
alces-483	269	2	.	.	PUNCT
alces-483	270	1	habitat	habitat	NOUN
alces-483	270	2	use	use	NOUN
alces-483	270	3	by	by	ADP
alces-483	270	4	four	four	NUM
alces-483	270	5	sympatric	sympatric	ADJ
alces-483	270	6	ungulates	ungulate	NOUN
alces-483	270	7	in	in	ADP
alces-483	270	8	boreal	boreal	ADJ
alces-483	270	9	mixedwood	mixedwood	NOUN
alces-483	270	10	forest	forest	NOUN
alces-483	270	11	.	.	PUNCT
alces-483	271	1	journal	journal	PROPN
alces-483	271	2	of	of	ADP
alces-483	271	3	space	space	NOUN
alces-483	271	4	and	and	CCONJ
alces-483	271	5	time	time	NOUN
alces-483	271	6	in	in	ADP
alces-483	271	7	predation	predation	NOUN
alces-483	271	8	dynamics	dynamic	NOUN
alces-483	271	9	–	–	PUNCT
alces-483	271	10	peterson	peterson	PROPN
alces-483	271	11	et	et	PROPN
alces-483	271	12	al	al	PROPN
alces-483	271	13	.	.	PROPN
alces-483	272	1	alces	alce	NOUN
alces-483	272	2	vol	vol	NOUN
alces-483	272	3	.	.	PROPN
alces-483	272	4	39	39	NUM
alces-483	272	5	,	,	PUNCT
alces-483	272	6	2003	2003	NUM
alces-483	272	7	226	226	NUM
alces-483	272	8	wildlife	wildlife	NOUN
alces-483	272	9	management	management	NOUN
alces-483	272	10	47:977	47:977	NUM
alces-483	272	11	-	-	SYM
alces-483	272	12	988	988	NUM
alces-483	272	13	.	.	PUNCT
alces-483	273	1	carbyn	carbyn	PROPN
alces-483	273	2	,	,	PUNCT
alces-483	273	3	l.	l.	PROPN
alces-483	273	4	n.	n.	PROPN
alces-483	273	5	1983	1983	NUM
alces-483	273	6	.	.	PUNCT
alces-483	274	1	wolf	wolf	PROPN
alces-483	274	2	predation	predation	NOUN
alces-483	274	3	on	on	ADP
alces-483	274	4	elk	elk	PROPN
alces-483	274	5	in	in	ADP
alces-483	274	6	riding	ride	VERB
alces-483	274	7	mountain	mountain	NOUN
alces-483	274	8	national	national	ADJ
alces-483	274	9	park	park	PROPN
alces-483	274	10	,	,	PUNCT
alces-483	274	11	manitoba	manitoba	PROPN
alces-483	274	12	.	.	PROPN
alces-483	274	13	journal	journal	PROPN
alces-483	274	14	of	of	ADP
alces-483	274	15	wildlife	wildlife	NOUN
alces-483	274	16	management	management	NOUN
alces-483	274	17	47:963	47:963	NUM
alces-483	274	18	-	-	SYM
alces-483	274	19	976	976	NUM
alces-483	274	20	.	.	PUNCT
alces-483	275	1	caughley	caughley	PROPN
alces-483	275	2	,	,	PUNCT
alces-483	275	3	g.	g.	PROPN
alces-483	275	4	,	,	PUNCT
alces-483	275	5	and	and	CCONJ
alces-483	275	6	c.	c.	PROPN
alces-483	275	7	j.	j.	PROPN
alces-483	275	8	krebs	krebs	PROPN
alces-483	275	9	.	.	PUNCT
alces-483	276	1	1983	1983	NUM
alces-483	276	2	.	.	PUNCT
alces-483	276	3	are	be	AUX
alces-483	276	4	big	big	ADJ
alces-483	276	5	mammals	mammal	NOUN
alces-483	276	6	simply	simply	ADV
alces-483	276	7	little	little	ADJ
alces-483	276	8	mammals	mammal	NOUN
alces-483	276	9	writ	writ	VERB
alces-483	276	10	large	large	ADJ
alces-483	276	11	?	?	PUNCT
alces-483	277	1	oecologia	oecologia	ADJ
alces-483	277	2	59:7	59:7	NUM
alces-483	277	3	-	-	SYM
alces-483	277	4	17	17	NUM
alces-483	277	5	.	.	PUNCT
alces-483	278	1	cederlund	cederlund	PROPN
alces-483	278	2	,	,	PUNCT
alces-483	278	3	g.	g.	PROPN
alces-483	278	4	,	,	PUNCT
alces-483	278	5	and	and	CCONJ
alces-483	278	6	g.	g.	PROPN
alces-483	278	7	markgren	markgren	PROPN
alces-483	278	8	.	.	PUNCT
alces-483	279	1	1987	1987	NUM
alces-483	279	2	.	.	PUNCT
alces-483	280	1	the	the	DET
alces-483	280	2	development	development	NOUN
alces-483	280	3	of	of	ADP
alces-483	280	4	the	the	DET
alces-483	280	5	swedish	swedish	PROPN
alces-483	280	6	moose	moose	PROPN
alces-483	280	7	population	population	NOUN
alces-483	280	8	,	,	PUNCT
alces-483	280	9	1970	1970	NUM
alces-483	280	10	-	-	SYM
alces-483	280	11	1983	1983	NUM
alces-483	280	12	.	.	PUNCT
alces-483	281	1	swedish	swedish	ADJ
alces-483	281	2	wildlife	wildlife	NOUN
alces-483	281	3	research	research	NOUN
alces-483	281	4	supplement	supplement	NOUN
alces-483	281	5	1:55	1:55	NUM
alces-483	281	6	-	-	SYM
alces-483	281	7	61	61	NUM
alces-483	281	8	.	.	PUNCT
alces-483	282	1	chouinard	chouinard	PROPN
alces-483	282	2	,	,	PUNCT
alces-483	282	3	a.	a.	NOUN
alces-483	282	4	,	,	PUNCT
alces-483	282	5	and	and	CCONJ
alces-483	282	6	l.	l.	PROPN
alces-483	282	7	filion	filion	NOUN
alces-483	282	8	.	.	PUNCT
alces-483	283	1	2001	2001	NUM
alces-483	283	2	.	.	PUNCT
alces-483	284	1	detrimental	detrimental	ADJ
alces-483	284	2	effects	effect	NOUN
alces-483	284	3	of	of	ADP
alces-483	284	4	white	white	ADJ
alces-483	284	5	-	-	PUNCT
alces-483	284	6	tailed	tail	VERB
alces-483	284	7	deer	deer	NOUN
alces-483	284	8	browsing	browse	VERB
alces-483	284	9	on	on	ADP
alces-483	284	10	balsam	balsam	NOUN
alces-483	284	11	fir	fir	NOUN
alces-483	284	12	growth	growth	NOUN
alces-483	284	13	and	and	CCONJ
alces-483	284	14	recruitment	recruitment	NOUN
alces-483	284	15	in	in	ADP
alces-483	284	16	a	a	DET
alces-483	284	17	second	second	ADJ
alces-483	284	18	-	-	PUNCT
alces-483	284	19	growth	growth	NOUN
alces-483	284	20	stand	stand	NOUN
alces-483	284	21	on	on	ADP
alces-483	284	22	anticosti	anticosti	PROPN
alces-483	284	23	island	island	PROPN
alces-483	284	24	,	,	PUNCT
alces-483	284	25	quebec	quebec	PROPN
alces-483	284	26	.	.	PUNCT
alces-483	285	1	ecoscience	ecoscience	PROPN
alces-483	285	2	8:199	8:199	PROPN
alces-483	285	3	-	-	SYM
alces-483	285	4	210	210	NUM
alces-483	285	5	.	.	PUNCT
alces-483	286	1	crête	crête	PROPN
alces-483	286	2	,	,	PUNCT
alces-483	286	3	m.	m.	NOUN
alces-483	286	4	1987	1987	NUM
alces-483	286	5	.	.	PUNCT
alces-483	287	1	the	the	DET
alces-483	287	2	impact	impact	NOUN
alces-483	287	3	of	of	ADP
alces-483	287	4	sport	sport	NOUN
alces-483	287	5	hunting	hunting	NOUN
alces-483	287	6	on	on	ADP
alces-483	287	7	north	north	PROPN
alces-483	287	8	american	american	ADJ
alces-483	287	9	moose	moose	NOUN
alces-483	287	10	.	.	PUNCT
alces-483	288	1	swedish	swedish	ADJ
alces-483	288	2	wildlife	wildlife	NOUN
alces-483	288	3	research	research	NOUN
alces-483	288	4	supplement	supplement	NOUN
alces-483	288	5	1:553	1:553	NUM
alces-483	288	6	-	-	SYM
alces-483	288	7	563	563	NUM
alces-483	288	8	.	.	PUNCT
alces-483	288	9	.	.	PUNCT
alces-483	289	1	1989	1989	NUM
alces-483	289	2	.	.	PUNCT
alces-483	290	1	approximation	approximation	NOUN
alces-483	290	2	of	of	ADP
alces-483	290	3	k	k	PROPN
alces-483	290	4	carrying	carry	VERB
alces-483	290	5	capacity	capacity	NOUN
alces-483	290	6	for	for	ADP
alces-483	290	7	moose	moose	NOUN
alces-483	290	8	in	in	ADP
alces-483	290	9	eastern	eastern	PROPN
alces-483	290	10	quebec	quebec	PROPN
alces-483	290	11	.	.	PUNCT
alces-483	291	1	canadian	canadian	ADJ
alces-483	291	2	journal	journal	PROPN
alces-483	291	3	of	of	ADP
alces-483	291	4	zoology	zoology	NOUN
alces-483	291	5	67:373	67:373	PROPN
alces-483	291	6	-	-	SYM
alces-483	291	7	380	380	NUM
alces-483	291	8	.	.	PUNCT
alces-483	291	9	,	,	PUNCT
alces-483	291	10	and	and	CCONJ
alces-483	291	11	m.	m.	NOUN
alces-483	291	12	manseau	manseau	PROPN
alces-483	291	13	.	.	PUNCT
alces-483	292	1	1996	1996	NUM
alces-483	292	2	.	.	PUNCT
alces-483	293	1	natural	natural	ADJ
alces-483	293	2	regulation	regulation	NOUN
alces-483	293	3	of	of	ADP
alces-483	293	4	cervidae	cervidae	NOUN
alces-483	293	5	along	along	ADP
alces-483	293	6	a	a	DET
alces-483	293	7	1000	1000	NUM
alces-483	293	8	km	km	NOUN
alces-483	293	9	latitudinal	latitudinal	ADJ
alces-483	293	10	gradient	gradient	NOUN
alces-483	293	11	:	:	PUNCT
alces-483	293	12	change	change	NOUN
alces-483	293	13	in	in	ADP
alces-483	293	14	trophic	trophic	ADJ
alces-483	293	15	dominance	dominance	NOUN
alces-483	293	16	.	.	PUNCT
alces-483	294	1	evolutionary	evolutionary	ADJ
alces-483	294	2	ecology	ecology	NOUN
alces-483	294	3	10:51	10:51	NUM
alces-483	294	4	-	-	SYM
alces-483	294	5	62	62	NUM
alces-483	294	6	.	.	PUNCT
alces-483	295	1	crichton	crichton	PROPN
alces-483	295	2	,	,	PUNCT
alces-483	295	3	v.	v.	PROPN
alces-483	295	4	f.	f.	PROPN
alces-483	295	5	1977	1977	NUM
alces-483	295	6	.	.	PUNCT
alces-483	296	1	hecla	hecla	PROPN
alces-483	296	2	island	island	PROPN
alces-483	296	3	—	—	PUNCT
alces-483	296	4	manitoba	manitoba	PROPN
alces-483	296	5	’s	’s	PART
alces-483	296	6	answer	answer	NOUN
alces-483	296	7	to	to	ADP
alces-483	296	8	isle	isle	PROPN
alces-483	296	9	royale	royale	PROPN
alces-483	296	10	.	.	PUNCT
alces-483	297	1	proceedings	proceeding	NOUN
alces-483	297	2	of	of	ADP
alces-483	297	3	the	the	DET
alces-483	297	4	north	north	ADJ
alces-483	297	5	american	american	PROPN
alces-483	297	6	moose	moose	PROPN
alces-483	297	7	conference	conference	NOUN
alces-483	297	8	and	and	CCONJ
alces-483	297	9	workshop	workshop	NOUN
alces-483	297	10	13:191	13:191	NUM
alces-483	297	11	-	-	SYM
alces-483	297	12	199	199	NUM
alces-483	297	13	.	.	PUNCT
alces-483	297	14	delgiudice	delgiudice	NOUN
alces-483	297	15	,	,	PUNCT
alces-483	297	16	g.	g.	PROPN
alces-483	297	17	d.	d.	PROPN
alces-483	297	18	1998	1998	NUM
alces-483	297	19	.	.	PUNCT
alces-483	298	1	surplus	surplus	NOUN
alces-483	298	2	killing	killing	NOUN
alces-483	298	3	of	of	ADP
alces-483	298	4	white	white	ADJ
alces-483	298	5	-	-	PUNCT
alces-483	298	6	tailed	tail	VERB
alces-483	298	7	deer	deer	NOUN
alces-483	298	8	by	by	ADP
alces-483	298	9	wolves	wolf	NOUN
alces-483	298	10	in	in	ADP
alces-483	298	11	northcentral	northcentral	ADJ
alces-483	298	12	minnesota	minnesota	PROPN
alces-483	298	13	.	.	PUNCT
alces-483	299	1	journal	journal	PROPN
alces-483	299	2	of	of	ADP
alces-483	299	3	mammalogy	mammalogy	NOUN
alces-483	299	4	79:227	79:227	PROPN
alces-483	299	5	-	-	SYM
alces-483	299	6	235	235	NUM
alces-483	299	7	.	.	PUNCT
alces-483	299	8	,	,	PUNCT
alces-483	299	9	r.	r.	PROPN
alces-483	299	10	o.	o.	PROPN
alces-483	299	11	peterson	peterson	PROPN
alces-483	299	12	,	,	PUNCT
alces-483	299	13	and	and	CCONJ
alces-483	299	14	w.	w.	PROPN
alces-483	299	15	m.	m.	PROPN
alces-483	299	16	samuels	samuels	PROPN
alces-483	299	17	.	.	PROPN
alces-483	299	18	1997	1997	NUM
alces-483	299	19	.	.	PUNCT
alces-483	300	1	trends	trend	NOUN
alces-483	300	2	of	of	ADP
alces-483	300	3	winter	winter	NOUN
alces-483	300	4	nutritional	nutritional	ADJ
alces-483	300	5	restriction	restriction	NOUN
alces-483	300	6	,	,	PUNCT
alces-483	300	7	ticks	tick	NOUN
alces-483	300	8	and	and	CCONJ
alces-483	300	9	numbers	number	NOUN
alces-483	300	10	of	of	ADP
alces-483	300	11	moose	moose	NOUN
alces-483	300	12	on	on	ADP
alces-483	300	13	isle	isle	PROPN
alces-483	300	14	royale	royale	PROPN
alces-483	300	15	.	.	PUNCT
alces-483	301	1	journal	journal	PROPN
alces-483	301	2	of	of	ADP
alces-483	301	3	wildlife	wildlife	NOUN
alces-483	301	4	management	management	NOUN
alces-483	301	5	61:895	61:895	NUM
alces-483	301	6	-	-	SYM
alces-483	301	7	903	903	NUM
alces-483	301	8	.	.	PUNCT
alces-483	302	1	detling	detle	VERB
alces-483	302	2	,	,	PUNCT
alces-483	302	3	j.	j.	PROPN
alces-483	302	4	k.	k.	PROPN
alces-483	302	5	1998	1998	NUM
alces-483	302	6	.	.	PUNCT
alces-483	303	1	mammalian	mammalian	ADJ
alces-483	303	2	herbivores	herbivore	NOUN
alces-483	303	3	:	:	PUNCT
alces-483	303	4	ecosystem	ecosystem	NOUN
alces-483	303	5	-	-	PUNCT
alces-483	303	6	level	level	NOUN
alces-483	303	7	effects	effect	NOUN
alces-483	303	8	in	in	ADP
alces-483	303	9	two	two	NUM
alces-483	303	10	grassland	grassland	NOUN
alces-483	303	11	national	national	ADJ
alces-483	303	12	parks	park	NOUN
alces-483	303	13	.	.	PUNCT
alces-483	304	1	wildlife	wildlife	NOUN
alces-483	304	2	society	society	NOUN
alces-483	304	3	bulletin	bulletin	NOUN
alces-483	304	4	26:438	26:438	PROPN
alces-483	304	5	-	-	SYM
alces-483	304	6	448	448	NUM
alces-483	304	7	.	.	PUNCT
alces-483	305	1	eberhardt	eberhardt	PROPN
alces-483	305	2	,	,	PUNCT
alces-483	305	3	l.	l.	PROPN
alces-483	305	4	l.	l.	PROPN
alces-483	305	5	1997	1997	NUM
alces-483	305	6	.	.	PUNCT
alces-483	305	7	is	be	AUX
alces-483	305	8	wolf	wolf	PROPN
alces-483	305	9	predation	predation	NOUN
alces-483	305	10	ratio	ratio	NOUN
alces-483	305	11	-	-	PUNCT
alces-483	305	12	dependent	dependent	ADJ
alces-483	305	13	?	?	PUNCT
alces-483	306	1	canadian	canadian	ADJ
alces-483	306	2	journal	journal	NOUN
alces-483	306	3	of	of	ADP
alces-483	306	4	zoology	zoology	NOUN
alces-483	306	5	75:1940	75:1940	NOUN
alces-483	306	6	-	-	SYM
alces-483	306	7	1944	1944	NUM
alces-483	306	8	.	.	PUNCT
alces-483	307	1	ehrlich	ehrlich	NOUN
alces-483	307	2	,	,	PUNCT
alces-483	307	3	p.	p.	PROPN
alces-483	307	4	r.	r.	PROPN
alces-483	307	5	,	,	PUNCT
alces-483	307	6	and	and	CCONJ
alces-483	307	7	l.	l.	PROPN
alces-483	307	8	c.	c.	PROPN
alces-483	307	9	birch	birch	PROPN
alces-483	307	10	.	.	PUNCT
alces-483	308	1	1967	1967	NUM
alces-483	308	2	.	.	PUNCT
alces-483	309	1	the	the	DET
alces-483	309	2	balance	balance	NOUN
alces-483	309	3	of	of	ADP
alces-483	309	4	nature	nature	NOUN
alces-483	309	5	and	and	CCONJ
alces-483	309	6	population	population	NOUN
alces-483	309	7	control	control	NOUN
alces-483	309	8	.	.	PUNCT
alces-483	310	1	american	american	PROPN
alces-483	310	2	naturalist	naturalist	PROPN
alces-483	310	3	101:97	101:97	PROPN
alces-483	310	4	-	-	SYM
alces-483	310	5	107	107	NUM
alces-483	310	6	.	.	PUNCT
alces-483	311	1	elton	elton	PROPN
alces-483	311	2	,	,	PUNCT
alces-483	311	3	c.	c.	PROPN
alces-483	311	4	1927	1927	NUM
alces-483	311	5	.	.	PUNCT
alces-483	312	1	animal	animal	NOUN
alces-483	312	2	ecology	ecology	NOUN
alces-483	312	3	.	.	PUNCT
alces-483	313	1	macmillan	macmillan	PROPN
alces-483	313	2	,	,	PUNCT
alces-483	313	3	new	new	PROPN
alces-483	313	4	york	york	PROPN
alces-483	313	5	,	,	PUNCT
alces-483	313	6	new	new	PROPN
alces-483	313	7	york	york	PROPN
alces-483	313	8	,	,	PUNCT
alces-483	313	9	usa	usa	PROPN
alces-483	313	10	.	.	PROPN
alces-483	313	11	etzenhouser	etzenhouser	PROPN
alces-483	313	12	,	,	PUNCT
alces-483	313	13	m.	m.	PROPN
alces-483	313	14	j.	j.	PROPN
alces-483	313	15	,	,	PUNCT
alces-483	313	16	m.	m.	PROPN
alces-483	313	17	k.	k.	PROPN
alces-483	313	18	owens	owens	PROPN
alces-483	313	19	,	,	PUNCT
alces-483	313	20	d.e	d.e	PROPN
alces-483	313	21	.	.	PROPN
alces-483	313	22	spalinger	spalinger	NOUN
alces-483	313	23	,	,	PUNCT
alces-483	313	24	and	and	CCONJ
alces-483	313	25	d.	d.	PROPN
alces-483	313	26	e.	e.	PROPN
alces-483	313	27	murden	murden	PROPN
alces-483	313	28	.	.	PROPN
alces-483	313	29	1998	1998	NUM
alces-483	313	30	.	.	PUNCT
alces-483	314	1	foraging	forage	VERB
alces-483	314	2	behavior	behavior	NOUN
alces-483	314	3	of	of	ADP
alces-483	314	4	browsing	browse	VERB
alces-483	314	5	ruminants	ruminant	NOUN
alces-483	314	6	in	in	ADP
alces-483	314	7	a	a	DET
alces-483	314	8	heterogeneous	heterogeneous	ADJ
alces-483	314	9	landscape	landscape	NOUN
alces-483	314	10	.	.	PUNCT
alces-483	315	1	landscape	landscape	NOUN
alces-483	315	2	ecology	ecology	NOUN
alces-483	315	3	13:55	13:55	NUM
alces-483	315	4	-	-	SYM
alces-483	315	5	64	64	NUM
alces-483	315	6	.	.	PUNCT
alces-483	316	1	farnell	farnell	PROPN
alces-483	316	2	,	,	PUNCT
alces-483	316	3	r.	r.	PROPN
alces-483	316	4	,	,	PUNCT
alces-483	316	5	and	and	CCONJ
alces-483	316	6	j.	j.	PROPN
alces-483	316	7	mcdonald	mcdonald	PROPN
alces-483	316	8	.	.	PUNCT
alces-483	316	9	1988	1988	NUM
alces-483	316	10	.	.	PUNCT
alces-483	317	1	the	the	DET
alces-483	317	2	influence	influence	NOUN
alces-483	317	3	of	of	ADP
alces-483	317	4	wolf	wolf	PROPN
alces-483	317	5	predation	predation	NOUN
alces-483	317	6	on	on	ADP
alces-483	317	7	caribou	caribou	PROPN
alces-483	317	8	mortality	mortality	NOUN
alces-483	317	9	in	in	ADP
alces-483	317	10	yukon	yukon	PROPN
alces-483	317	11	’s	’s	PART
alces-483	317	12	finlayson	finlayson	PROPN
alces-483	317	13	caribou	caribou	PROPN
alces-483	317	14	herd	herd	PROPN
alces-483	317	15	.	.	PUNCT
alces-483	318	1	proceedings	proceeding	NOUN
alces-483	318	2	of	of	ADP
alces-483	318	3	the	the	DET
alces-483	318	4	north	north	PROPN
alces-483	318	5	american	american	PROPN
alces-483	318	6	caribou	caribou	PROPN
alces-483	318	7	workshop	workshop	NOUN
alces-483	318	8	3:52	3:52	NUM
alces-483	318	9	-	-	SYM
alces-483	318	10	70	70	NUM
alces-483	318	11	.	.	PUNCT
alces-483	319	1	fretwell	fretwell	PROPN
alces-483	319	2	,	,	PUNCT
alces-483	319	3	s.	s.	PROPN
alces-483	319	4	d.	d.	PROPN
alces-483	319	5	1987	1987	NUM
alces-483	319	6	.	.	PUNCT
alces-483	320	1	food	food	NOUN
alces-483	320	2	chain	chain	NOUN
alces-483	320	3	dynamics	dynamic	NOUN
alces-483	320	4	:	:	PUNCT
alces-483	320	5	the	the	DET
alces-483	320	6	central	central	ADJ
alces-483	320	7	theory	theory	NOUN
alces-483	320	8	of	of	ADP
alces-483	320	9	ecology	ecology	NOUN
alces-483	320	10	?	?	PUNCT
alces-483	321	1	oikos	oikos	PROPN
alces-483	321	2	50:291	50:291	PROPN
alces-483	321	3	-	-	PUNCT
alces-483	321	4	301	301	NUM
alces-483	321	5	.	.	PUNCT
alces-483	322	1	fryxell	fryxell	PROPN
alces-483	322	2	,	,	PUNCT
alces-483	322	3	j.	j.	PROPN
alces-483	322	4	m.	m.	PROPN
alces-483	322	5	,	,	PUNCT
alces-483	322	6	w.	w.	PROPN
alces-483	322	7	e.	e.	PROPN
alces-483	322	8	mercer	mercer	PROPN
alces-483	322	9	,	,	PUNCT
alces-483	322	10	and	and	CCONJ
alces-483	322	11	r.	r.	PROPN
alces-483	322	12	b.	b.	PROPN
alces-483	322	13	gellately	gellately	PROPN
alces-483	322	14	.	.	PUNCT
alces-483	323	1	1988	1988	NUM
alces-483	323	2	.	.	PUNCT
alces-483	324	1	population	population	NOUN
alces-483	324	2	dynamics	dynamic	NOUN
alces-483	324	3	of	of	ADP
alces-483	324	4	newfoundland	newfoundland	PROPN
alces-483	324	5	moose	moose	PROPN
alces-483	324	6	using	use	VERB
alces-483	324	7	cohort	cohort	NOUN
alces-483	324	8	analysis	analysis	NOUN
alces-483	324	9	.	.	PUNCT
alces-483	325	1	journal	journal	NOUN
alces-483	325	2	of	of	ADP
alces-483	325	3	wildlife	wildlife	NOUN
alces-483	325	4	management	management	NOUN
alces-483	325	5	52:14	52:14	NUM
alces-483	325	6	-	-	SYM
alces-483	325	7	21	21	NUM
alces-483	325	8	.	.	PUNCT
alces-483	325	9	,	,	PUNCT
alces-483	325	10	and	and	CCONJ
alces-483	325	11	a.	a.	PROPN
alces-483	325	12	r.	r.	PROPN
alces-483	325	13	e.	e.	PROPN
alces-483	325	14	sinclair	sinclair	PROPN
alces-483	325	15	.	.	PUNCT
alces-483	326	1	1988	1988	NUM
alces-483	326	2	.	.	PUNCT
alces-483	327	1	causes	cause	NOUN
alces-483	327	2	and	and	CCONJ
alces-483	327	3	consequences	consequence	NOUN
alces-483	327	4	of	of	ADP
alces-483	327	5	migration	migration	NOUN
alces-483	327	6	by	by	ADP
alces-483	327	7	large	large	ADJ
alces-483	327	8	herbivores	herbivore	NOUN
alces-483	327	9	.	.	PUNCT
alces-483	328	1	trends	trend	NOUN
alces-483	328	2	in	in	ADP
alces-483	328	3	ecology	ecology	NOUN
alces-483	328	4	and	and	CCONJ
alces-483	328	5	evolution	evolution	NOUN
alces-483	328	6	9:237	9:237	NUM
alces-483	328	7	-	-	SYM
alces-483	328	8	241	241	NUM
alces-483	328	9	.	.	PUNCT
alces-483	328	10	fuller	full	ADJ
alces-483	328	11	,	,	PUNCT
alces-483	328	12	t.	t.	PROPN
alces-483	328	13	k.	k.	PROPN
alces-483	328	14	1989	1989	NUM
alces-483	328	15	.	.	PUNCT
alces-483	329	1	population	population	NOUN
alces-483	329	2	dynamics	dynamic	NOUN
alces-483	329	3	of	of	ADP
alces-483	329	4	wolves	wolf	NOUN
alces-483	329	5	in	in	ADP
alces-483	329	6	northcentral	northcentral	ADJ
alces-483	329	7	minnesota	minnesota	PROPN
alces-483	329	8	.	.	PUNCT
alces-483	330	1	wildlife	wildlife	NOUN
alces-483	330	2	monographs	monograph	NOUN
alces-483	330	3	105	105	NUM
alces-483	330	4	.	.	PUNCT
alces-483	331	1	_	_	PUNCT
alces-483	332	1	_	_	PUNCT
alces-483	333	1	_	_	PUNCT
alces-483	334	1	_	_	PUNCT
alces-483	335	1	_	_	PUNCT
alces-483	335	2	,	,	PUNCT
alces-483	335	3	and	and	CCONJ
alces-483	335	4	l.	l.	PROPN
alces-483	335	5	b.	b.	PROPN
alces-483	335	6	keith	keith	PROPN
alces-483	335	7	.	.	PUNCT
alces-483	336	1	1980	1980	NUM
alces-483	336	2	.	.	PUNCT
alces-483	337	1	wolf	wolf	PROPN
alces-483	337	2	population	population	NOUN
alces-483	337	3	dynamics	dynamic	NOUN
alces-483	337	4	and	and	CCONJ
alces-483	337	5	prey	prey	VERB
alces-483	337	6	relationships	relationship	NOUN
alces-483	337	7	in	in	ADP
alces-483	337	8	northeastern	northeastern	ADJ
alces-483	337	9	alberta	alberta	PROPN
alces-483	337	10	.	.	PUNCT
alces-483	338	1	journal	journal	PROPN
alces-483	338	2	of	of	ADP
alces-483	338	3	wildlife	wildlife	NOUN
alces-483	338	4	management	management	NOUN
alces-483	338	5	44:583	44:583	PROPN
alces-483	338	6	-	-	PUNCT
alces-483	338	7	602	602	NUM
alces-483	338	8	.	.	PUNCT
alces-483	339	1	gasaway	gasaway	ADV
alces-483	339	2	,	,	PUNCT
alces-483	339	3	w.	w.	PROPN
alces-483	339	4	c.	c.	PROPN
alces-483	339	5	,	,	PUNCT
alces-483	339	6	r.	r.	PROPN
alces-483	339	7	d.	d.	PROPN
alces-483	339	8	boertje	boertje	PROPN
alces-483	339	9	,	,	PUNCT
alces-483	339	10	d.	d.	PROPN
alces-483	339	11	v.	v.	PROPN
alces-483	339	12	grangaard	grangaard	PROPN
alces-483	339	13	,	,	PUNCT
alces-483	339	14	d.	d.	PROPN
alces-483	339	15	g.	g.	PROPN
alces-483	339	16	kelleyhouse	kelleyhouse	PROPN
alces-483	339	17	,	,	PUNCT
alces-483	339	18	r.	r.	PROPN
alces-483	339	19	o.	o.	PROPN
alces-483	339	20	stephenson	stephenson	PROPN
alces-483	339	21	,	,	PUNCT
alces-483	339	22	and	and	CCONJ
alces-483	339	23	d.	d.	PROPN
alces-483	339	24	g.	g.	PROPN
alces-483	339	25	larsen	larsen	PROPN
alces-483	339	26	.	.	PROPN
alces-483	339	27	1992	1992	NUM
alces-483	339	28	.	.	PUNCT
alces-483	340	1	the	the	DET
alces-483	340	2	role	role	NOUN
alces-483	340	3	of	of	ADP
alces-483	340	4	predation	predation	NOUN
alces-483	340	5	in	in	ADP
alces-483	340	6	limiting	limit	VERB
alces-483	340	7	moose	moose	NOUN
alces-483	340	8	at	at	ADP
alces-483	340	9	low	low	ADJ
alces-483	340	10	densities	density	NOUN
alces-483	340	11	in	in	ADP
alces-483	340	12	alaska	alaska	PROPN
alces-483	340	13	and	and	CCONJ
alces-483	340	14	yukon	yukon	PROPN
alces-483	340	15	and	and	CCONJ
alces-483	340	16	implications	implication	NOUN
alces-483	340	17	for	for	ADP
alces-483	340	18	conservation	conservation	NOUN
alces-483	340	19	.	.	PUNCT
alces-483	341	1	wildalces	wildalce	NOUN
alces-483	341	2	vol	vol	NOUN
alces-483	341	3	.	.	PROPN
alces-483	341	4	39	39	NUM
alces-483	341	5	,	,	PUNCT
alces-483	341	6	2003	2003	NUM
alces-483	341	7	peterson	peterson	PROPN
alces-483	341	8	et	et	PROPN
alces-483	341	9	al	al	PROPN
alces-483	341	10	.	.	PROPN
alces-483	341	11	–	–	PUNCT
alces-483	341	12	space	space	NOUN
alces-483	341	13	and	and	CCONJ
alces-483	341	14	time	time	NOUN
alces-483	341	15	in	in	ADP
alces-483	341	16	predation	predation	NOUN
alces-483	341	17	dynamics	dynamic	NOUN
alces-483	341	18	227	227	NUM
alces-483	341	19	life	life	NOUN
alces-483	341	20	monographs	monograph	NOUN
alces-483	341	21	120	120	NUM
alces-483	341	22	.	.	PUNCT
alces-483	341	23	,	,	PUNCT
alces-483	341	24	r.	r.	PROPN
alces-483	341	25	o.	o.	PROPN
alces-483	341	26	stephenson	stephenson	PROPN
alces-483	341	27	,	,	PUNCT
alces-483	341	28	j.	j.	PROPN
alces-483	341	29	l.	l.	PROPN
alces-483	341	30	davis	davis	PROPN
alces-483	341	31	,	,	PUNCT
alces-483	341	32	p.	p.	PROPN
alces-483	341	33	e.	e.	PROPN
alces-483	341	34	k.	k.	PROPN
alces-483	341	35	shepherd	shepherd	PROPN
alces-483	341	36	,	,	PUNCT
alces-483	341	37	and	and	CCONJ
alces-483	341	38	o.	o.	PROPN
alces-483	341	39	e.	e.	PROPN
alces-483	341	40	burris	burris	PROPN
alces-483	341	41	.	.	PUNCT
alces-483	342	1	1983	1983	NUM
alces-483	342	2	.	.	PUNCT
alces-483	343	1	interrelationships	interrelationship	NOUN
alces-483	343	2	of	of	ADP
alces-483	343	3	wolves	wolf	NOUN
alces-483	343	4	,	,	PUNCT
alces-483	343	5	prey	prey	NOUN
alces-483	343	6	,	,	PUNCT
alces-483	343	7	and	and	CCONJ
alces-483	343	8	man	man	NOUN
alces-483	343	9	in	in	ADP
alces-483	343	10	interior	interior	ADJ
alces-483	343	11	alaska	alaska	PROPN
alces-483	343	12	.	.	PUNCT
alces-483	344	1	wildlife	wildlife	NOUN
alces-483	344	2	monographs	monograph	NOUN
alces-483	344	3	84	84	NUM
alces-483	344	4	.	.	PUNCT
alces-483	345	1	haber	haber	PROPN
alces-483	345	2	,	,	PUNCT
alces-483	345	3	g.	g.	PROPN
alces-483	345	4	c.	c.	PROPN
alces-483	345	5	1977	1977	NUM
alces-483	345	6	.	.	PUNCT
alces-483	346	1	socio	socio	NOUN
alces-483	346	2	-	-	PUNCT
alces-483	346	3	ecological	ecological	ADJ
alces-483	346	4	dynamics	dynamic	NOUN
alces-483	346	5	of	of	ADP
alces-483	346	6	wolves	wolf	NOUN
alces-483	346	7	and	and	CCONJ
alces-483	346	8	prey	prey	NOUN
alces-483	346	9	in	in	ADP
alces-483	346	10	a	a	DET
alces-483	346	11	subarctic	subarctic	ADJ
alces-483	346	12	ecosystem	ecosystem	NOUN
alces-483	346	13	.	.	PUNCT
alces-483	347	1	ph.d	ph.d	PROPN
alces-483	347	2	.	.	PUNCT
alces-483	348	1	thesis	thesis	NOUN
alces-483	348	2	,	,	PUNCT
alces-483	348	3	university	university	PROPN
alces-483	348	4	of	of	ADP
alces-483	348	5	british	british	PROPN
alces-483	348	6	columbia	columbia	PROPN
alces-483	348	7	,	,	PUNCT
alces-483	348	8	vancouver	vancouver	PROPN
alces-483	348	9	,	,	PUNCT
alces-483	348	10	british	british	PROPN
alces-483	348	11	columbia	columbia	PROPN
alces-483	348	12	,	,	PUNCT
alces-483	348	13	canada	canada	PROPN
alces-483	348	14	.	.	PUNCT
alces-483	349	1	hairston	hairston	PROPN
alces-483	349	2	,	,	PUNCT
alces-483	349	3	n.	n.	PROPN
alces-483	349	4	g.	g.	PROPN
alces-483	349	5	,	,	PUNCT
alces-483	349	6	sr	sr	PROPN
alces-483	349	7	.	.	PROPN
alces-483	349	8	,	,	PUNCT
alces-483	349	9	f.	f.	PROPN
alces-483	349	10	e.	e.	PROPN
alces-483	349	11	smith	smith	PROPN
alces-483	349	12	,	,	PUNCT
alces-483	349	13	and	and	CCONJ
alces-483	349	14	l.	l.	PROPN
alces-483	349	15	b.	b.	PROPN
alces-483	349	16	slobodkin	slobodkin	PROPN
alces-483	349	17	.	.	PUNCT
alces-483	350	1	1960	1960	NUM
alces-483	350	2	.	.	PUNCT
alces-483	351	1	community	community	NOUN
alces-483	351	2	structure	structure	NOUN
alces-483	351	3	,	,	PUNCT
alces-483	351	4	population	population	NOUN
alces-483	351	5	control	control	NOUN
alces-483	351	6	,	,	PUNCT
alces-483	351	7	and	and	CCONJ
alces-483	351	8	competition	competition	NOUN
alces-483	351	9	.	.	PUNCT
alces-483	352	1	american	american	PROPN
alces-483	352	2	naturalist	naturalist	PROPN
alces-483	352	3	94:421	94:421	PROPN
alces-483	352	4	-	-	SYM
alces-483	352	5	425	425	NUM
alces-483	352	6	.	.	PUNCT
alces-483	352	7	hampton	hampton	PROPN
alces-483	352	8	,	,	PUNCT
alces-483	352	9	b.	b.	PROPN
alces-483	352	10	1997	1997	NUM
alces-483	352	11	.	.	PUNCT
alces-483	353	1	the	the	DET
alces-483	353	2	great	great	ADJ
alces-483	353	3	american	american	ADJ
alces-483	353	4	wolf	wolf	PROPN
alces-483	353	5	.	.	PUNCT
alces-483	354	1	henry	henry	PROPN
alces-483	354	2	holt	holt	PROPN
alces-483	354	3	and	and	CCONJ
alces-483	354	4	company	company	NOUN
alces-483	354	5	,	,	PUNCT
alces-483	354	6	new	new	PROPN
alces-483	354	7	york	york	PROPN
alces-483	354	8	,	,	PUNCT
alces-483	354	9	new	new	PROPN
alces-483	354	10	york	york	PROPN
alces-483	354	11	,	,	PUNCT
alces-483	354	12	usa	usa	PROPN
alces-483	354	13	.	.	PROPN
alces-483	354	14	holt	holt	PROPN
alces-483	354	15	,	,	PUNCT
alces-483	354	16	r.	r.	PROPN
alces-483	354	17	d.	d.	PROPN
alces-483	354	18	2000	2000	NUM
alces-483	354	19	.	.	PUNCT
alces-483	355	1	trophic	trophic	ADJ
alces-483	355	2	cascades	cascade	NOUN
alces-483	355	3	in	in	ADP
alces-483	355	4	terrestrial	terrestrial	ADJ
alces-483	355	5	ecosystems	ecosystem	NOUN
alces-483	355	6	:	:	PUNCT
alces-483	355	7	reflections	reflection	NOUN
alces-483	355	8	on	on	ADP
alces-483	355	9	polis	polis	PROPN
alces-483	355	10	et	et	PROPN
alces-483	355	11	al	al	PROPN
alces-483	355	12	.	.	PUNCT
alces-483	355	13	trends	trend	NOUN
alces-483	355	14	in	in	ADP
alces-483	355	15	ecology	ecology	NOUN
alces-483	355	16	and	and	CCONJ
alces-483	355	17	evolution	evolution	NOUN
alces-483	355	18	15:444	15:444	NUM
alces-483	355	19	-	-	SYM
alces-483	355	20	445	445	NUM
alces-483	355	21	.	.	PUNCT
alces-483	356	1	kelsall	kelsall	PROPN
alces-483	356	2	,	,	PUNCT
alces-483	356	3	j.	j.	PROPN
alces-483	356	4	p.	p.	PROPN
alces-483	356	5	1968	1968	NUM
alces-483	356	6	.	.	PUNCT
alces-483	357	1	the	the	DET
alces-483	357	2	migratory	migratory	PROPN
alces-483	357	3	barrenground	barrenground	PROPN
alces-483	357	4	caribou	caribou	PROPN
alces-483	357	5	of	of	ADP
alces-483	357	6	canada	canada	PROPN
alces-483	357	7	.	.	PUNCT
alces-483	358	1	canadian	canadian	ADJ
alces-483	358	2	wildlife	wildlife	PROPN
alces-483	358	3	service	service	NOUN
alces-483	358	4	monograph	monograph	NOUN
alces-483	358	5	series	series	PROPN
alces-483	358	6	number	number	NOUN
alces-483	358	7	3	3	NUM
alces-483	358	8	.	.	PUNCT
alces-483	359	1	ottawa	ottawa	PROPN
alces-483	359	2	,	,	PUNCT
alces-483	359	3	ontario	ontario	PROPN
alces-483	359	4	,	,	PUNCT
alces-483	359	5	canada	canada	PROPN
alces-483	359	6	.	.	PUNCT
alces-483	360	1	krebs	krebs	PROPN
alces-483	360	2	,	,	PUNCT
alces-483	360	3	c.	c.	PROPN
alces-483	360	4	j.	j.	PROPN
alces-483	360	5	1995	1995	NUM
alces-483	360	6	.	.	PUNCT
alces-483	361	1	two	two	NUM
alces-483	361	2	paradigms	paradigm	NOUN
alces-483	361	3	of	of	ADP
alces-483	361	4	population	population	NOUN
alces-483	361	5	regulation	regulation	NOUN
alces-483	361	6	.	.	PUNCT
alces-483	362	1	wildlife	wildlife	NOUN
alces-483	362	2	research	research	NOUN
alces-483	362	3	22:1	22:1	NUM
alces-483	362	4	-	-	SYM
alces-483	362	5	10	10	NUM
alces-483	362	6	.	.	PUNCT
alces-483	363	1	krefting	krefte	VERB
alces-483	363	2	,	,	PUNCT
alces-483	363	3	l.	l.	PROPN
alces-483	363	4	1974	1974	NUM
alces-483	363	5	.	.	PUNCT
alces-483	364	1	the	the	DET
alces-483	364	2	ecology	ecology	NOUN
alces-483	364	3	of	of	ADP
alces-483	364	4	the	the	DET
alces-483	364	5	isle	isle	PROPN
alces-483	364	6	royale	royale	PROPN
alces-483	364	7	moose	moose	PROPN
alces-483	364	8	with	with	ADP
alces-483	364	9	special	special	ADJ
alces-483	364	10	reference	reference	NOUN
alces-483	364	11	to	to	ADP
alces-483	364	12	the	the	DET
alces-483	364	13	habitat	habitat	NOUN
alces-483	364	14	.	.	PUNCT
alces-483	365	1	agricultural	agricultural	ADJ
alces-483	365	2	experiment	experiment	NOUN
alces-483	365	3	station	station	NOUN
alces-483	365	4	technical	technical	ADJ
alces-483	365	5	bulletin	bulletin	PROPN
alces-483	365	6	297	297	NUM
alces-483	365	7	.	.	PUNCT
alces-483	366	1	university	university	NOUN
alces-483	366	2	of	of	ADP
alces-483	366	3	minnesota	minnesota	PROPN
alces-483	366	4	,	,	PUNCT
alces-483	366	5	minneapolis	minneapolis	PROPN
alces-483	366	6	,	,	PUNCT
alces-483	366	7	minnesota	minnesota	PROPN
alces-483	366	8	,	,	PUNCT
alces-483	366	9	usa	usa	PROPN
alces-483	366	10	.	.	PROPN
alces-483	366	11	larsen	larsen	PROPN
alces-483	366	12	,	,	PUNCT
alces-483	366	13	d.	d.	PROPN
alces-483	366	14	g.	g.	PROPN
alces-483	366	15	1982	1982	NUM
alces-483	366	16	.	.	PUNCT
alces-483	367	1	moose	moose	NOUN
alces-483	367	2	inventory	inventory	NOUN
alces-483	367	3	in	in	ADP
alces-483	367	4	the	the	DET
alces-483	367	5	southwest	southwest	PROPN
alces-483	367	6	yukon	yukon	PROPN
alces-483	367	7	.	.	PUNCT
alces-483	368	1	alces	alce	VERB
alces-483	368	2	18:142167	18:142167	NUM
alces-483	368	3	.	.	PUNCT
alces-483	369	1	lindemann	lindemann	PROPN
alces-483	369	2	,	,	PUNCT
alces-483	369	3	r.	r.	PROPN
alces-483	369	4	1942	1942	NUM
alces-483	369	5	.	.	PUNCT
alces-483	370	1	the	the	DET
alces-483	370	2	trophic	trophic	ADJ
alces-483	370	3	-	-	PUNCT
alces-483	370	4	dynamic	dynamic	ADJ
alces-483	370	5	aspect	aspect	NOUN
alces-483	370	6	of	of	ADP
alces-483	370	7	ecology	ecology	NOUN
alces-483	370	8	.	.	PUNCT
alces-483	371	1	ecology	ecology	NOUN
alces-483	371	2	23:399418	23:399418	PROPN
alces-483	371	3	.	.	PUNCT
alces-483	372	1	lundberg	lundberg	PROPN
alces-483	372	2	,	,	PUNCT
alces-483	372	3	p.	p.	PROPN
alces-483	372	4	,	,	PUNCT
alces-483	372	5	and	and	CCONJ
alces-483	372	6	j.	j.	PROPN
alces-483	372	7	m.	m.	PROPN
alces-483	372	8	fryxell	fryxell	PROPN
alces-483	372	9	.	.	PUNCT
alces-483	373	1	1995	1995	NUM
alces-483	373	2	.	.	PUNCT
alces-483	374	1	expected	expect	VERB
alces-483	374	2	population	population	NOUN
alces-483	374	3	density	density	NOUN
alces-483	374	4	versus	versus	ADP
alces-483	374	5	productivity	productivity	NOUN
alces-483	374	6	in	in	ADP
alces-483	374	7	ratio	ratio	NOUN
alces-483	374	8	-	-	PUNCT
alces-483	374	9	dependent	dependent	ADJ
alces-483	374	10	and	and	CCONJ
alces-483	374	11	preydependent	preydependent	NOUN
alces-483	374	12	models	model	NOUN
alces-483	374	13	.	.	PUNCT
alces-483	375	1	american	american	PROPN
alces-483	375	2	naturalist	naturalist	PROPN
alces-483	375	3	146:154	146:154	NOUN
alces-483	375	4	-	-	PUNCT
alces-483	375	5	161	161	NUM
alces-483	375	6	.	.	PUNCT
alces-483	376	1	,	,	PUNCT
alces-483	376	2	e.	e.	PROPN
alces-483	376	3	ranta	ranta	PROPN
alces-483	376	4	,	,	PUNCT
alces-483	376	5	j.	j.	PROPN
alces-483	376	6	ripa	ripa	PROPN
alces-483	376	7	,	,	PUNCT
alces-483	376	8	and	and	CCONJ
alces-483	377	1	v.	v.	ADP
alces-483	377	2	kaitala	kaitala	PROPN
alces-483	377	3	.	.	PUNCT
alces-483	377	4	2000	2000	NUM
alces-483	377	5	.	.	PUNCT
alces-483	378	1	population	population	NOUN
alces-483	378	2	variability	variability	NOUN
alces-483	378	3	in	in	ADP
alces-483	378	4	space	space	NOUN
alces-483	378	5	and	and	CCONJ
alces-483	378	6	time	time	NOUN
alces-483	378	7	.	.	PUNCT
alces-483	379	1	trends	trend	NOUN
alces-483	379	2	in	in	ADP
alces-483	379	3	ecology	ecology	NOUN
alces-483	379	4	and	and	CCONJ
alces-483	379	5	evolution	evolution	NOUN
alces-483	379	6	15:460	15:460	NUM
alces-483	379	7	-	-	SYM
alces-483	379	8	464	464	NUM
alces-483	379	9	.	.	PUNCT
alces-483	380	1	mclaren	mclaren	PROPN
alces-483	380	2	,	,	PUNCT
alces-483	380	3	b.	b.	PROPN
alces-483	380	4	e.	e.	PROPN
alces-483	380	5	1996	1996	PROPN
alces-483	380	6	.	.	PUNCT
alces-483	381	1	plant	plant	NOUN
alces-483	381	2	-	-	PUNCT
alces-483	381	3	specific	specific	ADJ
alces-483	381	4	response	response	NOUN
alces-483	381	5	to	to	ADP
alces-483	381	6	herbivory	herbivory	NOUN
alces-483	381	7	:	:	PUNCT
alces-483	381	8	simulated	simulate	VERB
alces-483	381	9	browsing	browsing	NOUN
alces-483	381	10	of	of	ADP
alces-483	381	11	suppressed	suppress	VERB
alces-483	381	12	balsam	balsam	NOUN
alces-483	381	13	fir	fir	NOUN
alces-483	381	14	on	on	ADP
alces-483	381	15	isle	isle	PROPN
alces-483	381	16	royale	royale	PROPN
alces-483	381	17	.	.	PUNCT
alces-483	382	1	ecology	ecology	NOUN
alces-483	382	2	77:228	77:228	NUM
alces-483	382	3	-	-	SYM
alces-483	382	4	235	235	NUM
alces-483	382	5	.	.	PUNCT
alces-483	382	6	,	,	PUNCT
alces-483	382	7	and	and	CCONJ
alces-483	382	8	r.	r.	PROPN
alces-483	382	9	a.	a.	PROPN
alces-483	382	10	janke	janke	PROPN
alces-483	382	11	.	.	PUNCT
alces-483	383	1	1996	1996	NUM
alces-483	383	2	.	.	PUNCT
alces-483	384	1	seedbed	seedbe	VERB
alces-483	384	2	and	and	CCONJ
alces-483	384	3	canopy	canopy	NOUN
alces-483	384	4	cover	cover	VERB
alces-483	384	5	effects	effect	NOUN
alces-483	384	6	on	on	ADP
alces-483	384	7	balsam	balsam	NOUN
alces-483	384	8	fir	fir	NOUN
alces-483	384	9	seedling	seedling	NOUN
alces-483	384	10	establishment	establishment	NOUN
alces-483	384	11	in	in	ADP
alces-483	384	12	isle	isle	PROPN
alces-483	384	13	royale	royale	PROPN
alces-483	384	14	national	national	PROPN
alces-483	384	15	park	park	PROPN
alces-483	384	16	.	.	PUNCT
alces-483	385	1	canadian	canadian	ADJ
alces-483	385	2	journal	journal	PROPN
alces-483	385	3	of	of	ADP
alces-483	385	4	forest	forest	PROPN
alces-483	385	5	research	research	NOUN
alces-483	385	6	26:782	26:782	PROPN
alces-483	385	7	-	-	SYM
alces-483	385	8	793	793	NUM
alces-483	385	9	.	.	PUNCT
alces-483	385	10	,	,	PUNCT
alces-483	385	11	and	and	CCONJ
alces-483	385	12	r.	r.	PROPN
alces-483	385	13	o.	o.	PROPN
alces-483	385	14	peterson	peterson	PROPN
alces-483	385	15	.	.	PROPN
alces-483	385	16	1994	1994	NUM
alces-483	385	17	.	.	PUNCT
alces-483	386	1	wolves	wolf	NOUN
alces-483	386	2	,	,	PUNCT
alces-483	386	3	moose	moose	NOUN
alces-483	386	4	and	and	CCONJ
alces-483	386	5	tree	tree	NOUN
alces-483	386	6	rings	ring	NOUN
alces-483	386	7	on	on	ADP
alces-483	386	8	isle	isle	PROPN
alces-483	386	9	royale	royale	PROPN
alces-483	386	10	.	.	PUNCT
alces-483	387	1	science	science	PROPN
alces-483	387	2	266:1555	266:1555	PROPN
alces-483	387	3	-	-	SYM
alces-483	387	4	1558	1558	NUM
alces-483	387	5	.	.	PUNCT
alces-483	388	1	mech	mech	PROPN
alces-483	388	2	,	,	PUNCT
alces-483	388	3	l.	l.	PROPN
alces-483	388	4	d.	d.	PROPN
alces-483	388	5	1966	1966	NUM
alces-483	388	6	.	.	PUNCT
alces-483	389	1	the	the	DET
alces-483	389	2	wolves	wolf	NOUN
alces-483	389	3	of	of	ADP
alces-483	389	4	isle	isle	PROPN
alces-483	389	5	royale	royale	PROPN
alces-483	389	6	.	.	PUNCT
alces-483	390	1	u.s	u.s	PROPN
alces-483	390	2	.	.	PROPN
alces-483	390	3	national	national	PROPN
alces-483	390	4	park	park	PROPN
alces-483	390	5	service	service	PROPN
alces-483	390	6	fauna	fauna	PROPN
alces-483	390	7	series	series	PROPN
alces-483	390	8	7:1	7:1	NUM
alces-483	390	9	-	-	SYM
alces-483	390	10	210	210	NUM
alces-483	390	11	.	.	PUNCT
alces-483	390	12	.	.	PUNCT
alces-483	391	1	1986	1986	NUM
alces-483	391	2	.	.	PUNCT
alces-483	392	1	wolf	wolf	NOUN
alces-483	392	2	numbers	number	NOUN
alces-483	392	3	and	and	CCONJ
alces-483	392	4	population	population	NOUN
alces-483	392	5	trend	trend	NOUN
alces-483	392	6	in	in	ADP
alces-483	392	7	the	the	DET
alces-483	392	8	central	central	ADJ
alces-483	392	9	superior	superior	ADJ
alces-483	392	10	national	national	ADJ
alces-483	392	11	forest	forest	NOUN
alces-483	392	12	,	,	PUNCT
alces-483	392	13	1967	1967	NUM
alces-483	392	14	-	-	SYM
alces-483	392	15	1985	1985	NUM
alces-483	392	16	.	.	PUNCT
alces-483	393	1	u.s	u.s	PROPN
alces-483	393	2	.	.	PROPN
alces-483	393	3	department	department	PROPN
alces-483	393	4	of	of	ADP
alces-483	393	5	agriculture	agriculture	PROPN
alces-483	393	6	forest	forest	PROPN
alces-483	393	7	service	service	PROPN
alces-483	393	8	research	research	NOUN
alces-483	393	9	paper	paper	NOUN
alces-483	393	10	no	no	INTJ
alces-483	393	11	.	.	PUNCT
alces-483	394	1	nc-270	nc-270	NOUN
alces-483	394	2	.	.	PUNCT
alces-483	395	1	north	north	NOUN
alces-483	395	2	central	central	PROPN
alces-483	395	3	forest	forest	NOUN
alces-483	395	4	experiment	experiment	NOUN
alces-483	395	5	station	station	NOUN
alces-483	395	6	,	,	PUNCT
alces-483	395	7	st	st	PROPN
alces-483	395	8	.	.	PROPN
alces-483	395	9	paul	paul	PROPN
alces-483	395	10	,	,	PUNCT
alces-483	395	11	minnesota	minnesota	PROPN
alces-483	395	12	,	,	PUNCT
alces-483	395	13	usa	usa	PROPN
alces-483	395	14	.	.	PROPN
alces-483	395	15	,	,	PUNCT
alces-483	395	16	and	and	CCONJ
alces-483	395	17	l.	l.	PROPN
alces-483	395	18	d.	d.	PROPN
alces-483	395	19	frenzel	frenzel	PROPN
alces-483	395	20	jr	jr	PROPN
alces-483	395	21	.	.	PROPN
alces-483	395	22	1971	1971	NUM
alces-483	395	23	.	.	PUNCT
alces-483	396	1	ecological	ecological	ADJ
alces-483	396	2	studies	study	NOUN
alces-483	396	3	of	of	ADP
alces-483	396	4	the	the	DET
alces-483	396	5	timber	timber	NOUN
alces-483	396	6	wolf	wolf	NOUN
alces-483	396	7	in	in	ADP
alces-483	396	8	northeastern	northeastern	ADJ
alces-483	396	9	minnesota	minnesota	PROPN
alces-483	396	10	.	.	PUNCT
alces-483	397	1	u.s	u.s	PROPN
alces-483	397	2	.	.	PROPN
alces-483	397	3	department	department	PROPN
alces-483	397	4	of	of	ADP
alces-483	397	5	agriculture	agriculture	PROPN
alces-483	397	6	forest	forest	PROPN
alces-483	397	7	service	service	PROPN
alces-483	397	8	research	research	NOUN
alces-483	397	9	paper	paper	NOUN
alces-483	397	10	nc-52	nc-52	ADV
alces-483	397	11	.	.	PUNCT
alces-483	398	1	north	north	PROPN
alces-483	398	2	central	central	PROPN
alces-483	398	3	forest	forest	NOUN
alces-483	398	4	experiment	experiment	NOUN
alces-483	398	5	station	station	NOUN
alces-483	398	6	,	,	PUNCT
alces-483	398	7	st	st	PROPN
alces-483	398	8	.	.	PROPN
alces-483	398	9	paul	paul	PROPN
alces-483	398	10	,	,	PUNCT
alces-483	398	11	minnesota	minnesota	PROPN
alces-483	398	12	,	,	PUNCT
alces-483	398	13	usa	usa	PROPN
alces-483	398	14	.	.	PROPN
alces-483	398	15	,	,	PUNCT
alces-483	398	16	and	and	CCONJ
alces-483	398	17	p.	p.	PROPN
alces-483	398	18	d.	d.	PROPN
alces-483	398	19	karns	karns	PROPN
alces-483	398	20	.	.	PUNCT
alces-483	399	1	1977	1977	NUM
alces-483	399	2	.	.	PUNCT
alces-483	400	1	role	role	NOUN
alces-483	400	2	of	of	ADP
alces-483	400	3	the	the	DET
alces-483	400	4	wolf	wolf	NOUN
alces-483	400	5	in	in	ADP
alces-483	400	6	a	a	DET
alces-483	400	7	deer	deer	NOUN
alces-483	400	8	decline	decline	NOUN
alces-483	400	9	in	in	ADP
alces-483	400	10	the	the	DET
alces-483	400	11	superior	superior	ADJ
alces-483	400	12	national	national	ADJ
alces-483	400	13	forest	forest	NOUN
alces-483	400	14	.	.	PUNCT
alces-483	401	1	u.s	u.s	PROPN
alces-483	401	2	.	.	PROPN
alces-483	401	3	department	department	PROPN
alces-483	401	4	of	of	ADP
alces-483	401	5	agriculture	agriculture	PROPN
alces-483	401	6	forest	forest	PROPN
alces-483	401	7	service	service	PROPN
alces-483	401	8	research	research	NOUN
alces-483	401	9	paper	paper	NOUN
alces-483	401	10	nc-148	nc-148	PROPN
alces-483	401	11	.	.	PUNCT
alces-483	402	1	north	north	PROPN
alces-483	402	2	central	central	PROPN
alces-483	402	3	forest	forest	NOUN
alces-483	402	4	experiment	experiment	NOUN
alces-483	402	5	station	station	NOUN
alces-483	402	6	,	,	PUNCT
alces-483	402	7	st	st	PROPN
alces-483	402	8	.	.	PROPN
alces-483	402	9	paul	paul	PROPN
alces-483	402	10	,	,	PUNCT
alces-483	402	11	minnesota	minnesota	PROPN
alces-483	402	12	,	,	PUNCT
alces-483	402	13	usa	usa	PROPN
alces-483	402	14	.	.	PROPN
alces-483	402	15	,	,	PUNCT
alces-483	402	16	r.	r.	PROPN
alces-483	402	17	e.	e.	PROPN
alces-483	402	18	mcroberts	mcroberts	PROPN
alces-483	402	19	,	,	PUNCT
alces-483	402	20	r.o	r.o	PROPN
alces-483	402	21	.	.	PROPN
alces-483	402	22	peterson	peterson	PROPN
alces-483	402	23	,	,	PUNCT
alces-483	402	24	and	and	CCONJ
alces-483	402	25	r.	r.	PROPN
alces-483	402	26	e.	e.	PROPN
alces-483	402	27	page	page	PROPN
alces-483	402	28	.	.	PUNCT
alces-483	403	1	1987	1987	NUM
alces-483	403	2	.	.	PUNCT
alces-483	404	1	relationship	relationship	NOUN
alces-483	404	2	of	of	ADP
alces-483	404	3	deer	deer	NOUN
alces-483	404	4	and	and	CCONJ
alces-483	404	5	moose	moose	NOUN
alces-483	404	6	populations	population	NOUN
alces-483	404	7	to	to	ADP
alces-483	404	8	previous	previous	ADJ
alces-483	404	9	winters	winter	NOUN
alces-483	404	10	’	'	PUNCT
alces-483	404	11	snow	snow	NOUN
alces-483	404	12	.	.	PUNCT
alces-483	405	1	journal	journal	NOUN
alces-483	405	2	of	of	ADP
alces-483	405	3	animal	animal	NOUN
alces-483	405	4	ecology	ecology	NOUN
alces-483	405	5	56:615	56:615	NUM
alces-483	405	6	-	-	SYM
alces-483	405	7	627	627	NUM
alces-483	405	8	.	.	NUM
alces-483	405	9	,	,	PUNCT
alces-483	405	10	t.	t.	PROPN
alces-483	405	11	j.	j.	PROPN
alces-483	405	12	meier	meier	PROPN
alces-483	405	13	,	,	PUNCT
alces-483	405	14	j.	j.	PROPN
alces-483	405	15	w.	w.	PROPN
alces-483	405	16	burch	burch	PROPN
alces-483	405	17	,	,	PUNCT
alces-483	405	18	and	and	CCONJ
alces-483	405	19	l.	l.	PROPN
alces-483	405	20	g.	g.	PROPN
alces-483	405	21	space	space	PROPN
alces-483	405	22	and	and	CCONJ
alces-483	405	23	time	time	NOUN
alces-483	405	24	in	in	ADP
alces-483	405	25	predation	predation	NOUN
alces-483	405	26	dynamics	dynamic	NOUN
alces-483	405	27	–	–	PUNCT
alces-483	405	28	peterson	peterson	PROPN
alces-483	405	29	et	et	PROPN
alces-483	405	30	al	al	PROPN
alces-483	405	31	.	.	PROPN
alces-483	406	1	alces	alce	NOUN
alces-483	406	2	vol	vol	NOUN
alces-483	406	3	.	.	PROPN
alces-483	406	4	39	39	NUM
alces-483	406	5	,	,	PUNCT
alces-483	406	6	2003	2003	NUM
alces-483	406	7	228	228	NUM
alces-483	406	8	adams	adam	NOUN
alces-483	406	9	.	.	PUNCT
alces-483	407	1	1995	1995	NUM
alces-483	407	2	.	.	PUNCT
alces-483	408	1	patterns	pattern	NOUN
alces-483	408	2	of	of	ADP
alces-483	408	3	prey	prey	PROPN
alces-483	408	4	selection	selection	NOUN
alces-483	408	5	by	by	ADP
alces-483	408	6	wolves	wolf	NOUN
alces-483	408	7	in	in	ADP
alces-483	408	8	denali	denali	PROPN
alces-483	408	9	national	national	PROPN
alces-483	408	10	park	park	PROPN
alces-483	408	11	,	,	PUNCT
alces-483	408	12	alaska	alaska	PROPN
alces-483	408	13	.	.	PUNCT
alces-483	409	1	pages	page	NOUN
alces-483	409	2	231	231	NUM
alces-483	409	3	-	-	SYM
alces-483	409	4	244	244	NUM
alces-483	409	5	in	in	ADP
alces-483	409	6	l.	l.	PROPN
alces-483	409	7	n.	n.	PROPN
alces-483	409	8	carbyn	carbyn	PROPN
alces-483	409	9	,	,	PUNCT
alces-483	409	10	s.	s.	PROPN
alces-483	409	11	h.	h.	PROPN
alces-483	409	12	fritts	fritts	PROPN
alces-483	409	13	,	,	PUNCT
alces-483	409	14	and	and	CCONJ
alces-483	409	15	d.	d.	PROPN
alces-483	409	16	r.	r.	PROPN
alces-483	409	17	seip	seip	PROPN
alces-483	409	18	,	,	PUNCT
alces-483	409	19	editors	editor	NOUN
alces-483	409	20	.	.	PUNCT
alces-483	410	1	ecology	ecology	NOUN
alces-483	410	2	and	and	CCONJ
alces-483	410	3	conservation	conservation	NOUN
alces-483	410	4	of	of	ADP
alces-483	410	5	wolves	wolf	NOUN
alces-483	410	6	in	in	ADP
alces-483	410	7	a	a	DET
alces-483	410	8	changing	change	VERB
alces-483	410	9	world	world	NOUN
alces-483	410	10	.	.	PUNCT
alces-483	411	1	canadian	canadian	PROPN
alces-483	411	2	circumpolar	circumpolar	PROPN
alces-483	411	3	institute	institute	NOUN
alces-483	411	4	occasional	occasional	ADJ
alces-483	411	5	publication	publication	NOUN
alces-483	411	6	no	no	INTJ
alces-483	411	7	.	.	PROPN
alces-483	411	8	35	35	NUM
alces-483	411	9	,	,	PUNCT
alces-483	411	10	edmonton	edmonton	PROPN
alces-483	411	11	,	,	PUNCT
alces-483	411	12	alberta	alberta	PROPN
alces-483	411	13	,	,	PUNCT
alces-483	411	14	canada	canada	PROPN
alces-483	411	15	.	.	PUNCT
alces-483	411	16	,	,	PUNCT
alces-483	411	17	m.	m.	PROPN
alces-483	411	18	e.	e.	PROPN
alces-483	411	19	nelson	nelson	PROPN
alces-483	411	20	,	,	PUNCT
alces-483	411	21	and	and	CCONJ
alces-483	411	22	r.	r.	PROPN
alces-483	411	23	e.	e.	PROPN
alces-483	411	24	mcroberts	mcroberts	PROPN
alces-483	411	25	.	.	PUNCT
alces-483	411	26	1991	1991	NUM
alces-483	411	27	.	.	PUNCT
alces-483	412	1	effects	effect	NOUN
alces-483	412	2	of	of	ADP
alces-483	412	3	maternal	maternal	ADJ
alces-483	412	4	and	and	CCONJ
alces-483	412	5	grandmaternal	grandmaternal	ADJ
alces-483	412	6	nutrition	nutrition	NOUN
alces-483	412	7	on	on	ADP
alces-483	412	8	deer	deer	NOUN
alces-483	412	9	mass	mass	NOUN
alces-483	412	10	and	and	CCONJ
alces-483	412	11	vulnerability	vulnerability	NOUN
alces-483	412	12	to	to	ADP
alces-483	412	13	wolf	wolf	PROPN
alces-483	412	14	predation	predation	NOUN
alces-483	412	15	.	.	PUNCT
alces-483	413	1	journal	journal	PROPN
alces-483	413	2	of	of	ADP
alces-483	413	3	mammalogy	mammalogy	NOUN
alces-483	413	4	72:146151	72:146151	PROPN
alces-483	413	5	.	.	PUNCT
alces-483	414	1	mercer	mercer	PROPN
alces-483	414	2	,	,	PUNCT
alces-483	414	3	w.	w.	PROPN
alces-483	414	4	e.	e.	PROPN
alces-483	414	5	,	,	PUNCT
alces-483	414	6	and	and	CCONJ
alces-483	414	7	f.	f.	PROPN
alces-483	414	8	manuel	manuel	PROPN
alces-483	414	9	.	.	PUNCT
alces-483	415	1	1974	1974	NUM
alces-483	415	2	.	.	PUNCT
alces-483	416	1	some	some	DET
alces-483	416	2	aspects	aspect	NOUN
alces-483	416	3	of	of	ADP
alces-483	416	4	moose	moose	NOUN
alces-483	416	5	management	management	NOUN
alces-483	416	6	in	in	ADP
alces-483	416	7	newfoundland	newfoundland	PROPN
alces-483	416	8	.	.	PUNCT
alces-483	417	1	naturaliste	naturaliste	PROPN
alces-483	417	2	canadien	canadien	PROPN
alces-483	417	3	101:657	101:657	PROPN
alces-483	417	4	-	-	PUNCT
alces-483	417	5	671	671	NUM
alces-483	417	6	.	.	PUNCT
alces-483	417	7	messier	messy	ADJ
alces-483	417	8	,	,	PUNCT
alces-483	417	9	f.	f.	PROPN
alces-483	417	10	1991	1991	NUM
alces-483	417	11	.	.	PUNCT
alces-483	418	1	the	the	DET
alces-483	418	2	significance	significance	NOUN
alces-483	418	3	of	of	ADP
alces-483	418	4	limiting	limit	VERB
alces-483	418	5	and	and	CCONJ
alces-483	418	6	regulating	regulate	VERB
alces-483	418	7	factors	factor	NOUN
alces-483	418	8	on	on	ADP
alces-483	418	9	the	the	DET
alces-483	418	10	demography	demography	NOUN
alces-483	418	11	of	of	ADP
alces-483	418	12	moose	moose	NOUN
alces-483	418	13	and	and	CCONJ
alces-483	418	14	white	white	ADJ
alces-483	418	15	-	-	PUNCT
alces-483	418	16	tailed	tail	VERB
alces-483	418	17	deer	deer	NOUN
alces-483	418	18	.	.	PUNCT
alces-483	419	1	journal	journal	NOUN
alces-483	419	2	of	of	ADP
alces-483	419	3	animal	animal	NOUN
alces-483	419	4	ecology	ecology	NOUN
alces-483	419	5	60:377393	60:377393	NOUN
alces-483	419	6	.	.	PUNCT
alces-483	419	7	.	.	PUNCT
alces-483	420	1	1994	1994	NUM
alces-483	420	2	.	.	PUNCT
alces-483	421	1	ungulate	ungulate	ADJ
alces-483	421	2	population	population	NOUN
alces-483	421	3	models	model	NOUN
alces-483	421	4	with	with	ADP
alces-483	421	5	predation	predation	NOUN
alces-483	421	6	:	:	PUNCT
alces-483	421	7	a	a	DET
alces-483	421	8	case	case	NOUN
alces-483	421	9	study	study	NOUN
alces-483	421	10	with	with	ADP
alces-483	421	11	the	the	DET
alces-483	421	12	north	north	ADJ
alces-483	421	13	american	american	ADJ
alces-483	421	14	moose	moose	NOUN
alces-483	421	15	.	.	PUNCT
alces-483	422	1	ecology	ecology	NOUN
alces-483	422	2	75:478	75:478	PROPN
alces-483	422	3	-	-	SYM
alces-483	422	4	488	488	NUM
alces-483	422	5	.	.	PUNCT
alces-483	422	6	,	,	PUNCT
alces-483	422	7	and	and	CCONJ
alces-483	422	8	m.	m.	NOUN
alces-483	422	9	crête	crête	PROPN
alces-483	422	10	.	.	PUNCT
alces-483	423	1	1985	1985	NUM
alces-483	423	2	.	.	PUNCT
alces-483	424	1	moose	moose	NOUN
alces-483	424	2	-	-	PUNCT
alces-483	424	3	wolf	wolf	NOUN
alces-483	424	4	dynamics	dynamic	NOUN
alces-483	424	5	and	and	CCONJ
alces-483	424	6	the	the	DET
alces-483	424	7	natural	natural	ADJ
alces-483	424	8	regulation	regulation	NOUN
alces-483	424	9	of	of	ADP
alces-483	424	10	moose	moose	NOUN
alces-483	424	11	populations	population	NOUN
alces-483	424	12	.	.	PUNCT
alces-483	425	1	oecologia	oecologia	PROPN
alces-483	425	2	65:503512	65:503512	NUM
alces-483	425	3	.	.	PUNCT
alces-483	426	1	murdoch	murdoch	PROPN
alces-483	426	2	,	,	PUNCT
alces-483	426	3	w.	w.	PROPN
alces-483	426	4	w.	w.	PROPN
alces-483	426	5	1966	1966	NUM
alces-483	426	6	.	.	PUNCT
alces-483	427	1	community	community	NOUN
alces-483	427	2	structure	structure	NOUN
alces-483	427	3	,	,	PUNCT
alces-483	427	4	population	population	NOUN
alces-483	427	5	control	control	NOUN
alces-483	427	6	,	,	PUNCT
alces-483	427	7	and	and	CCONJ
alces-483	427	8	competition	competition	NOUN
alces-483	427	9	.	.	PUNCT
alces-483	428	1	american	american	ADJ
alces-483	428	2	naturalist	naturalist	PROPN
alces-483	428	3	100:219	100:219	PROPN
alces-483	428	4	-	-	SYM
alces-483	428	5	226	226	NUM
alces-483	428	6	.	.	PUNCT
alces-483	429	1	nelson	nelson	PROPN
alces-483	429	2	,	,	PUNCT
alces-483	429	3	m.	m.	PROPN
alces-483	429	4	e.	e.	PROPN
alces-483	429	5	,	,	PUNCT
alces-483	429	6	and	and	CCONJ
alces-483	429	7	l.	l.	PROPN
alces-483	429	8	d.	d.	PROPN
alces-483	429	9	mech	mech	PROPN
alces-483	429	10	.	.	PUNCT
alces-483	430	1	1986	1986	NUM
alces-483	430	2	.	.	PUNCT
alces-483	431	1	white	white	ADJ
alces-483	431	2	-	-	PUNCT
alces-483	431	3	tailed	tail	VERB
alces-483	431	4	deer	deer	NOUN
alces-483	431	5	numbers	number	NOUN
alces-483	431	6	and	and	CCONJ
alces-483	431	7	population	population	NOUN
alces-483	431	8	trend	trend	NOUN
alces-483	431	9	in	in	ADP
alces-483	431	10	the	the	DET
alces-483	431	11	central	central	ADJ
alces-483	431	12	superior	superior	ADJ
alces-483	431	13	national	national	ADJ
alces-483	431	14	forest	forest	NOUN
alces-483	431	15	,	,	PUNCT
alces-483	431	16	1967	1967	NUM
alces-483	431	17	-	-	SYM
alces-483	431	18	1985	1985	NUM
alces-483	431	19	.	.	PUNCT
alces-483	432	1	u.s	u.s	PROPN
alces-483	432	2	.	.	PROPN
alces-483	432	3	department	department	PROPN
alces-483	432	4	of	of	ADP
alces-483	432	5	agriculture	agriculture	PROPN
alces-483	432	6	forest	forest	PROPN
alces-483	432	7	service	service	PROPN
alces-483	432	8	research	research	NOUN
alces-483	432	9	paper	paper	NOUN
alces-483	432	10	nc-271	nc-271	PROPN
alces-483	432	11	.	.	PUNCT
alces-483	433	1	north	north	PROPN
alces-483	433	2	central	central	PROPN
alces-483	433	3	forest	forest	NOUN
alces-483	433	4	experiment	experiment	NOUN
alces-483	433	5	station	station	NOUN
alces-483	433	6	,	,	PUNCT
alces-483	433	7	st	st	PROPN
alces-483	433	8	.	.	PROPN
alces-483	433	9	paul	paul	PROPN
alces-483	433	10	,	,	PUNCT
alces-483	433	11	minnesota	minnesota	PROPN
alces-483	433	12	,	,	PUNCT
alces-483	433	13	usa	usa	PROPN
alces-483	433	14	.	.	PROPN
alces-483	433	15	nygrén	nygrén	PROPN
alces-483	433	16	,	,	PUNCT
alces-483	433	17	t.	t.	PROPN
alces-483	433	18	1987	1987	NUM
alces-483	433	19	.	.	PUNCT
alces-483	434	1	the	the	DET
alces-483	434	2	history	history	NOUN
alces-483	434	3	of	of	ADP
alces-483	434	4	moose	moose	NOUN
alces-483	434	5	in	in	ADP
alces-483	434	6	finland	finland	PROPN
alces-483	434	7	.	.	PUNCT
alces-483	435	1	swedish	swedish	ADJ
alces-483	435	2	wildlife	wildlife	PROPN
alces-483	435	3	research	research	NOUN
alces-483	435	4	supplement	supplement	NOUN
alces-483	435	5	1:49	1:49	NUM
alces-483	435	6	-	-	SYM
alces-483	435	7	54	54	NUM
alces-483	435	8	.	.	PUNCT
alces-483	436	1	oksanen	oksanen	PROPN
alces-483	436	2	,	,	PUNCT
alces-483	436	3	l.	l.	PROPN
alces-483	436	4	1983	1983	NUM
alces-483	436	5	.	.	PUNCT
alces-483	437	1	trophic	trophic	ADJ
alces-483	437	2	exploitation	exploitation	NOUN
alces-483	437	3	and	and	CCONJ
alces-483	437	4	arctic	arctic	ADJ
alces-483	437	5	phytomass	phytomass	NOUN
alces-483	437	6	patterns	pattern	NOUN
alces-483	437	7	.	.	PUNCT
alces-483	438	1	american	american	PROPN
alces-483	438	2	naturalist	naturalist	PROPN
alces-483	438	3	122:360	122:360	PROPN
alces-483	438	4	-	-	PUNCT
alces-483	438	5	367	367	NUM
alces-483	438	6	.	.	PUNCT
alces-483	439	1	,	,	PUNCT
alces-483	439	2	s.	s.	PROPN
alces-483	439	3	fretwell	fretwell	PROPN
alces-483	439	4	,	,	PUNCT
alces-483	439	5	j.	j.	PROPN
alces-483	439	6	arruda	arruda	PROPN
alces-483	439	7	,	,	PUNCT
alces-483	439	8	and	and	CCONJ
alces-483	439	9	p.	p.	NOUN
alces-483	439	10	niemela	niemela	NOUN
alces-483	439	11	.	.	PUNCT
alces-483	440	1	1981	1981	NUM
alces-483	440	2	.	.	PUNCT
alces-483	441	1	exploitation	exploitation	NOUN
alces-483	441	2	ecosystems	ecosystem	NOUN
alces-483	441	3	in	in	ADP
alces-483	441	4	gradients	gradient	NOUN
alces-483	441	5	of	of	ADP
alces-483	441	6	primary	primary	ADJ
alces-483	441	7	productivity	productivity	NOUN
alces-483	441	8	.	.	PUNCT
alces-483	442	1	american	american	PROPN
alces-483	442	2	naturalist	naturalist	PROPN
alces-483	442	3	118:240	118:240	PROPN
alces-483	442	4	-	-	PUNCT
alces-483	442	5	261	261	NUM
alces-483	442	6	.	.	PUNCT
alces-483	443	1	oosenbrug	oosenbrug	PROPN
alces-483	443	2	,	,	PUNCT
alces-483	443	3	s.	s.	PROPN
alces-483	443	4	m.	m.	PROPN
alces-483	443	5	,	,	PUNCT
alces-483	443	6	and	and	CCONJ
alces-483	443	7	l.	l.	PROPN
alces-483	443	8	n.	n.	PROPN
alces-483	443	9	carbyn	carbyn	PROPN
alces-483	443	10	.	.	PUNCT
alces-483	444	1	1985	1985	NUM
alces-483	444	2	.	.	PUNCT
alces-483	445	1	wolf	wolf	PROPN
alces-483	445	2	predation	predation	NOUN
alces-483	445	3	on	on	ADP
alces-483	445	4	bison	bison	NOUN
alces-483	445	5	in	in	ADP
alces-483	445	6	wood	wood	NOUN
alces-483	445	7	buffalo	buffalo	PROPN
alces-483	445	8	national	national	PROPN
alces-483	445	9	park	park	PROPN
alces-483	445	10	.	.	PUNCT
alces-483	446	1	unpublished	unpublished	ADJ
alces-483	446	2	report	report	NOUN
alces-483	446	3	.	.	PUNCT
alces-483	447	1	canadian	canadian	ADJ
alces-483	447	2	wildlife	wildlife	NOUN
alces-483	447	3	service	service	PROPN
alces-483	447	4	,	,	PUNCT
alces-483	447	5	edmonton	edmonton	PROPN
alces-483	447	6	,	,	PUNCT
alces-483	447	7	alberta	alberta	PROPN
alces-483	447	8	,	,	PUNCT
alces-483	447	9	canada	canada	PROPN
alces-483	447	10	.	.	PUNCT
alces-483	447	11	orians	orians	PROPN
alces-483	447	12	,	,	PUNCT
alces-483	447	13	g.h	g.h	PROPN
alces-483	447	14	.	.	PROPN
alces-483	447	15	,	,	PUNCT
alces-483	447	16	et	et	PROPN
alces-483	447	17	al	al	PROPN
alces-483	447	18	.	.	PUNCT
alces-483	448	1	(	(	PUNCT
alces-483	448	2	13	13	NUM
alces-483	448	3	authors	author	NOUN
alces-483	448	4	)	)	PUNCT
alces-483	448	5	.	.	PUNCT
alces-483	449	1	1997	1997	NUM
alces-483	449	2	.	.	PUNCT
alces-483	450	1	wolves	wolf	NOUN
alces-483	450	2	,	,	PUNCT
alces-483	450	3	bears	bear	NOUN
alces-483	450	4	,	,	PUNCT
alces-483	450	5	and	and	CCONJ
alces-483	450	6	their	their	PRON
alces-483	450	7	prey	prey	NOUN
alces-483	450	8	in	in	ADP
alces-483	450	9	alaska	alaska	PROPN
alces-483	450	10	:	:	PUNCT
alces-483	450	11	biological	biological	ADJ
alces-483	450	12	and	and	CCONJ
alces-483	450	13	social	social	ADJ
alces-483	450	14	challenges	challenge	NOUN
alces-483	450	15	in	in	ADP
alces-483	450	16	wildlife	wildlife	NOUN
alces-483	450	17	management	management	NOUN
alces-483	450	18	.	.	PUNCT
alces-483	451	1	national	national	PROPN
alces-483	451	2	academy	academy	PROPN
alces-483	451	3	press	press	PROPN
alces-483	451	4	,	,	PUNCT
alces-483	451	5	washington	washington	PROPN
alces-483	451	6	,	,	PUNCT
alces-483	451	7	d.c	d.c	PROPN
alces-483	451	8	.	.	PROPN
alces-483	451	9	,	,	PUNCT
alces-483	451	10	usa	usa	PROPN
alces-483	451	11	.	.	PROPN
alces-483	451	12	page	page	PROPN
alces-483	451	13	,	,	PUNCT
alces-483	451	14	r.	r.	PROPN
alces-483	451	15	e.	e.	PROPN
alces-483	451	16	1989	1989	NUM
alces-483	451	17	.	.	PUNCT
alces-483	452	1	the	the	DET
alces-483	452	2	inverted	inverted	ADJ
alces-483	452	3	pyramid	pyramid	NOUN
alces-483	452	4	:	:	PUNCT
alces-483	452	5	ecosystem	ecosystem	NOUN
alces-483	452	6	dynamics	dynamic	NOUN
alces-483	452	7	of	of	ADP
alces-483	452	8	wolves	wolf	NOUN
alces-483	452	9	and	and	CCONJ
alces-483	452	10	moose	moose	NOUN
alces-483	452	11	on	on	ADP
alces-483	452	12	isle	isle	PROPN
alces-483	452	13	royale	royale	PROPN
alces-483	452	14	.	.	PUNCT
alces-483	453	1	ph.d	ph.d	PROPN
alces-483	453	2	.	.	PUNCT
alces-483	454	1	thesis	thesis	PROPN
alces-483	454	2	,	,	PUNCT
alces-483	454	3	michigan	michigan	PROPN
alces-483	454	4	technological	technological	ADJ
alces-483	454	5	university	university	PROPN
alces-483	454	6	,	,	PUNCT
alces-483	454	7	houghton	houghton	PROPN
alces-483	454	8	,	,	PUNCT
alces-483	454	9	michigan	michigan	PROPN
alces-483	454	10	,	,	PUNCT
alces-483	454	11	usa	usa	PROPN
alces-483	454	12	.	.	PROPN
alces-483	454	13	paine	paine	PROPN
alces-483	454	14	,	,	PUNCT
alces-483	454	15	r.	r.	PROPN
alces-483	454	16	t.	t.	PROPN
alces-483	454	17	1980	1980	NUM
alces-483	454	18	.	.	PUNCT
alces-483	455	1	food	food	NOUN
alces-483	455	2	webs	webs	NOUN
alces-483	455	3	:	:	PUNCT
alces-483	455	4	linkage	linkage	NOUN
alces-483	455	5	,	,	PUNCT
alces-483	455	6	interaction	interaction	NOUN
alces-483	455	7	strength	strength	NOUN
alces-483	455	8	,	,	PUNCT
alces-483	455	9	and	and	CCONJ
alces-483	455	10	community	community	NOUN
alces-483	455	11	structure	structure	NOUN
alces-483	455	12	.	.	PUNCT
alces-483	456	1	journal	journal	NOUN
alces-483	456	2	of	of	ADP
alces-483	456	3	animal	animal	NOUN
alces-483	456	4	ecology	ecology	NOUN
alces-483	456	5	49:667	49:667	NUM
alces-483	456	6	-	-	PUNCT
alces-483	456	7	685	685	NUM
alces-483	456	8	.	.	PUNCT
alces-483	456	9	.	.	PUNCT
alces-483	457	1	2000	2000	NUM
alces-483	457	2	.	.	PUNCT
alces-483	458	1	phycology	phycology	NOUN
alces-483	458	2	for	for	ADP
alces-483	458	3	the	the	DET
alces-483	458	4	mammalogist	mammalogist	NOUN
alces-483	458	5	:	:	PUNCT
alces-483	458	6	marine	marine	PROPN
alces-483	458	7	rocky	rocky	ADJ
alces-483	458	8	shores	shore	NOUN
alces-483	458	9	and	and	CCONJ
alces-483	458	10	mammal	mammal	ADJ
alces-483	458	11	-	-	PUNCT
alces-483	458	12	dominated	dominate	VERB
alces-483	458	13	communities	community	NOUN
alces-483	458	14	—	—	PUNCT
alces-483	458	15	how	how	SCONJ
alces-483	458	16	different	different	ADJ
alces-483	458	17	are	be	AUX
alces-483	458	18	the	the	DET
alces-483	458	19	structuring	structuring	NOUN
alces-483	458	20	processes	process	NOUN
alces-483	458	21	?	?	PUNCT
alces-483	459	1	journal	journal	PROPN
alces-483	459	2	of	of	ADP
alces-483	459	3	mammalogy	mammalogy	NOUN
alces-483	459	4	81:637	81:637	NUM
alces-483	459	5	-	-	SYM
alces-483	459	6	648	648	NUM
alces-483	459	7	.	.	PUNCT
alces-483	460	1	pastor	pastor	PROPN
alces-483	460	2	,	,	PUNCT
alces-483	460	3	j.	j.	PROPN
alces-483	460	4	,	,	PUNCT
alces-483	460	5	r.	r.	PROPN
alces-483	460	6	moen	moen	PROPN
alces-483	460	7	,	,	PUNCT
alces-483	460	8	and	and	CCONJ
alces-483	460	9	y.	y.	PROPN
alces-483	460	10	cohen	cohen	PROPN
alces-483	460	11	.	.	PUNCT
alces-483	461	1	1997	1997	NUM
alces-483	461	2	.	.	PUNCT
alces-483	462	1	spatial	spatial	ADJ
alces-483	462	2	heterogeneities	heterogeneity	NOUN
alces-483	462	3	,	,	PUNCT
alces-483	462	4	carrying	carry	VERB
alces-483	462	5	capacity	capacity	NOUN
alces-483	462	6	,	,	PUNCT
alces-483	462	7	and	and	CCONJ
alces-483	462	8	feedbacks	feedback	NOUN
alces-483	462	9	in	in	ADP
alces-483	462	10	animal	animal	NOUN
alces-483	462	11	-	-	PUNCT
alces-483	462	12	landscape	landscape	NOUN
alces-483	462	13	interactions	interaction	NOUN
alces-483	462	14	.	.	PUNCT
alces-483	463	1	journal	journal	NOUN
alces-483	463	2	of	of	ADP
alces-483	463	3	mammalogy	mammalogy	NOUN
alces-483	463	4	78:1040	78:1040	PROPN
alces-483	463	5	-	-	SYM
alces-483	463	6	1052	1052	NUM
alces-483	463	7	.	.	PUNCT
alces-483	464	1	peek	peek	NOUN
alces-483	464	2	,	,	PUNCT
alces-483	464	3	j.	j.	PROPN
alces-483	464	4	m.	m.	PROPN
alces-483	464	5	,	,	PUNCT
alces-483	464	6	d.	d.	PROPN
alces-483	464	7	l.	l.	PROPN
alces-483	464	8	ulrich	ulrich	PROPN
alces-483	464	9	,	,	PUNCT
alces-483	464	10	and	and	CCONJ
alces-483	464	11	r.	r.	PROPN
alces-483	464	12	j.	j.	PROPN
alces-483	464	13	mackie	mackie	PROPN
alces-483	464	14	.	.	PUNCT
alces-483	465	1	1976	1976	NUM
alces-483	465	2	.	.	PUNCT
alces-483	466	1	moose	moose	NOUN
alces-483	466	2	habitat	habitat	NOUN
alces-483	466	3	selection	selection	NOUN
alces-483	466	4	and	and	CCONJ
alces-483	466	5	relationships	relationship	NOUN
alces-483	466	6	to	to	ADP
alces-483	466	7	forest	forest	NOUN
alces-483	466	8	management	management	NOUN
alces-483	466	9	in	in	ADP
alces-483	466	10	northeastern	northeastern	ADJ
alces-483	466	11	minnesota	minnesota	PROPN
alces-483	466	12	.	.	PUNCT
alces-483	467	1	wildlife	wildlife	NOUN
alces-483	467	2	monographs	monograph	NOUN
alces-483	467	3	48	48	NUM
alces-483	467	4	.	.	PUNCT
alces-483	468	1	peterson	peterson	PROPN
alces-483	468	2	,	,	PUNCT
alces-483	468	3	r.	r.	PROPN
alces-483	468	4	o.	o.	PROPN
alces-483	468	5	1975	1975	NUM
alces-483	468	6	.	.	PUNCT
alces-483	469	1	wolf	wolf	PROPN
alces-483	469	2	response	response	NOUN
alces-483	469	3	to	to	ADP
alces-483	469	4	increased	increase	VERB
alces-483	469	5	moose	moose	NOUN
alces-483	469	6	vulnerability	vulnerability	NOUN
alces-483	469	7	in	in	ADP
alces-483	469	8	isle	isle	PROPN
alces-483	469	9	royale	royale	PROPN
alces-483	469	10	.	.	PUNCT
alces-483	470	1	proceedings	proceeding	NOUN
alces-483	470	2	of	of	ADP
alces-483	470	3	the	the	DET
alces-483	470	4	north	north	ADJ
alces-483	470	5	american	american	PROPN
alces-483	470	6	moose	moose	PROPN
alces-483	470	7	conference	conference	PROPN
alces-483	470	8	11:344368	11:344368	PROPN
alces-483	470	9	.	.	PUNCT
alces-483	470	10	alces	alce	NOUN
alces-483	470	11	vol	vol	NOUN
alces-483	470	12	.	.	PROPN
alces-483	470	13	39	39	NUM
alces-483	470	14	,	,	PUNCT
alces-483	470	15	2003	2003	NUM
alces-483	470	16	peterson	peterson	PROPN
alces-483	470	17	et	et	PROPN
alces-483	470	18	al	al	PROPN
alces-483	470	19	.	.	PROPN
alces-483	470	20	–	–	PUNCT
alces-483	470	21	space	space	NOUN
alces-483	470	22	and	and	CCONJ
alces-483	470	23	time	time	NOUN
alces-483	470	24	in	in	ADP
alces-483	470	25	predation	predation	NOUN
alces-483	470	26	dynamics	dynamic	NOUN
alces-483	470	27	229	229	NUM
alces-483	470	28	.	.	PUNCT
alces-483	471	1	1977	1977	NUM
alces-483	471	2	.	.	PUNCT
alces-483	472	1	wolf	wolf	PROPN
alces-483	472	2	ecology	ecology	NOUN
alces-483	472	3	and	and	CCONJ
alces-483	472	4	prey	prey	VERB
alces-483	472	5	relationships	relationship	NOUN
alces-483	472	6	on	on	ADP
alces-483	472	7	isle	isle	PROPN
alces-483	472	8	royale	royale	PROPN
alces-483	472	9	.	.	PUNCT
alces-483	473	1	u.s	u.s	PROPN
alces-483	473	2	.	.	PROPN
alces-483	473	3	national	national	PROPN
alces-483	473	4	park	park	PROPN
alces-483	473	5	service	service	PROPN
alces-483	473	6	scientific	scientific	PROPN
alces-483	473	7	monograph	monograph	NOUN
alces-483	473	8	series	series	PROPN
alces-483	473	9	11	11	NUM
alces-483	473	10	.	.	PUNCT
alces-483	474	1	washington	washington	PROPN
alces-483	474	2	,	,	PUNCT
alces-483	474	3	d.c	d.c	PROPN
alces-483	474	4	.	.	PROPN
alces-483	474	5	,	,	PUNCT
alces-483	474	6	usa	usa	PROPN
alces-483	474	7	.	.	PROPN
alces-483	474	8	.	.	PUNCT
alces-483	475	1	1995	1995	NUM
alces-483	475	2	.	.	PUNCT
alces-483	476	1	the	the	DET
alces-483	476	2	wolves	wolf	NOUN
alces-483	476	3	of	of	ADP
alces-483	476	4	isle	isle	PROPN
alces-483	476	5	royale	royale	PROPN
alces-483	476	6	—	—	PUNCT
alces-483	476	7	a	a	DET
alces-483	476	8	broken	broken	ADJ
alces-483	476	9	balance	balance	NOUN
alces-483	476	10	.	.	PUNCT
alces-483	477	1	willow	willow	PROPN
alces-483	477	2	creek	creek	PROPN
alces-483	477	3	press	press	PROPN
alces-483	477	4	,	,	PUNCT
alces-483	477	5	minocqua	minocqua	PROPN
alces-483	477	6	,	,	PUNCT
alces-483	477	7	wisconsin	wisconsin	PROPN
alces-483	477	8	,	,	PUNCT
alces-483	477	9	usa	usa	PROPN
alces-483	477	10	.	.	PROPN
alces-483	477	11	.	.	PUNCT
alces-483	478	1	1999	1999	NUM
alces-483	478	2	.	.	PUNCT
alces-483	479	1	wolf	wolf	NOUN
alces-483	479	2	-	-	PUNCT
alces-483	479	3	moose	moose	NOUN
alces-483	479	4	interaction	interaction	NOUN
alces-483	479	5	on	on	ADP
alces-483	479	6	isle	isle	PROPN
alces-483	479	7	royale	royale	PROPN
alces-483	479	8	:	:	PUNCT
alces-483	479	9	the	the	DET
alces-483	479	10	end	end	NOUN
alces-483	479	11	of	of	ADP
alces-483	479	12	natural	natural	ADJ
alces-483	479	13	regulation	regulation	NOUN
alces-483	479	14	?	?	PUNCT
alces-483	480	1	ecological	ecological	ADJ
alces-483	480	2	applications	application	NOUN
alces-483	480	3	9:10	9:10	NUM
alces-483	480	4	-	-	SYM
alces-483	480	5	16	16	NUM
alces-483	480	6	.	.	PUNCT
alces-483	480	7	,	,	PUNCT
alces-483	480	8	and	and	CCONJ
alces-483	480	9	r.	r.	PROPN
alces-483	480	10	e.	e.	PROPN
alces-483	480	11	page	page	PROPN
alces-483	480	12	.	.	PUNCT
alces-483	481	1	1983	1983	NUM
alces-483	481	2	.	.	PUNCT
alces-483	482	1	wolfmoose	wolfmoose	ADJ
alces-483	482	2	fluctuations	fluctuation	NOUN
alces-483	482	3	at	at	ADP
alces-483	482	4	isle	isle	PROPN
alces-483	482	5	royale	royale	PROPN
alces-483	482	6	national	national	PROPN
alces-483	482	7	park	park	PROPN
alces-483	482	8	,	,	PUNCT
alces-483	482	9	michigan	michigan	PROPN
alces-483	482	10	,	,	PUNCT
alces-483	482	11	u.s.a	u.s.a	PROPN
alces-483	482	12	.	.	PUNCT
alces-483	482	13	acta	acta	PROPN
alces-483	482	14	zoologica	zoologica	PROPN
alces-483	482	15	fennica	fennica	PROPN
alces-483	483	1	174:251	174:251	NUM
alces-483	483	2	-	-	PUNCT
alces-483	483	3	253	253	NUM
alces-483	483	4	.	.	PUNCT
alces-483	484	1	,	,	PUNCT
alces-483	484	2	n.	n.	PROPN
alces-483	484	3	j.	j.	PROPN
alces-483	484	4	thomas	thomas	PROPN
alces-483	484	5	,	,	PUNCT
alces-483	484	6	j.	j.	PROPN
alces-483	484	7	m.	m.	PROPN
alces-483	484	8	thurber	thurber	PROPN
alces-483	484	9	,	,	PUNCT
alces-483	484	10	j.	j.	PROPN
alces-483	484	11	a.	a.	PROPN
alces-483	484	12	vucetich	vucetich	PROPN
alces-483	484	13	,	,	PUNCT
alces-483	484	14	and	and	CCONJ
alces-483	484	15	t.	t.	PROPN
alces-483	484	16	a.	a.	PROPN
alces-483	484	17	waite	waite	PROPN
alces-483	484	18	.	.	PUNCT
alces-483	485	1	1998	1998	NUM
alces-483	485	2	.	.	PUNCT
alces-483	486	1	population	population	NOUN
alces-483	486	2	limitation	limitation	NOUN
alces-483	486	3	and	and	CCONJ
alces-483	486	4	the	the	DET
alces-483	486	5	wolves	wolf	NOUN
alces-483	486	6	of	of	ADP
alces-483	486	7	isle	isle	PROPN
alces-483	486	8	royale	royale	PROPN
alces-483	486	9	.	.	PUNCT
alces-483	487	1	journal	journal	PROPN
alces-483	487	2	of	of	ADP
alces-483	487	3	mammalogy	mammalogy	NOUN
alces-483	487	4	79:828	79:828	PROPN
alces-483	487	5	-	-	SYM
alces-483	487	6	841	841	NUM
alces-483	487	7	.	.	PUNCT
alces-483	487	8	,	,	PUNCT
alces-483	487	9	j.	j.	PROPN
alces-483	487	10	d.	d.	PROPN
alces-483	487	11	woolington	woolington	PROPN
alces-483	487	12	,	,	PUNCT
alces-483	487	13	and	and	CCONJ
alces-483	487	14	t.	t.	PROPN
alces-483	487	15	n.	n.	PROPN
alces-483	487	16	bailey	bailey	PROPN
alces-483	487	17	.	.	PUNCT
alces-483	487	18	1984	1984	NUM
alces-483	487	19	.	.	PUNCT
alces-483	488	1	wolves	wolf	NOUN
alces-483	488	2	of	of	ADP
alces-483	488	3	the	the	DET
alces-483	488	4	kenai	kenai	PROPN
alces-483	488	5	peninsula	peninsula	PROPN
alces-483	488	6	,	,	PUNCT
alces-483	488	7	alaska	alaska	PROPN
alces-483	488	8	.	.	PUNCT
alces-483	489	1	wildlife	wildlife	NOUN
alces-483	489	2	monographs	monograph	NOUN
alces-483	489	3	88	88	NUM
alces-483	489	4	.	.	PUNCT
alces-483	490	1	pimlott	pimlott	PROPN
alces-483	490	2	,	,	PUNCT
alces-483	490	3	d.	d.	PROPN
alces-483	490	4	h.	h.	PROPN
alces-483	490	5	1967	1967	NUM
alces-483	490	6	.	.	PUNCT
alces-483	491	1	wolf	wolf	PROPN
alces-483	491	2	predation	predation	NOUN
alces-483	491	3	and	and	CCONJ
alces-483	491	4	ungulate	ungulate	ADJ
alces-483	491	5	populations	population	NOUN
alces-483	491	6	.	.	PUNCT
alces-483	492	1	american	american	ADJ
alces-483	492	2	zoologist	zoologist	NOUN
alces-483	492	3	7:267	7:267	NOUN
alces-483	492	4	-	-	SYM
alces-483	492	5	278	278	NUM
alces-483	492	6	.	.	PUNCT
alces-483	492	7	,	,	PUNCT
alces-483	492	8	j.	j.	PROPN
alces-483	492	9	a.	a.	PROPN
alces-483	492	10	shannon	shannon	PROPN
alces-483	492	11	,	,	PUNCT
alces-483	492	12	and	and	CCONJ
alces-483	492	13	g.	g.	PROPN
alces-483	492	14	b.	b.	PROPN
alces-483	492	15	kolenosky	kolenosky	PROPN
alces-483	492	16	.	.	PUNCT
alces-483	493	1	1969	1969	NUM
alces-483	493	2	.	.	PUNCT
alces-483	494	1	the	the	DET
alces-483	494	2	ecology	ecology	NOUN
alces-483	494	3	of	of	ADP
alces-483	494	4	the	the	DET
alces-483	494	5	timber	timber	NOUN
alces-483	494	6	wolf	wolf	NOUN
alces-483	494	7	in	in	ADP
alces-483	494	8	algonquin	algonquin	PROPN
alces-483	494	9	provincial	provincial	ADJ
alces-483	494	10	park	park	NOUN
alces-483	494	11	.	.	PUNCT
alces-483	495	1	ontario	ontario	PROPN
alces-483	495	2	department	department	PROPN
alces-483	495	3	of	of	ADP
alces-483	495	4	lands	land	NOUN
alces-483	495	5	and	and	CCONJ
alces-483	495	6	forests	forest	VERB
alces-483	495	7	research	research	NOUN
alces-483	495	8	paper	paper	NOUN
alces-483	495	9	(	(	PUNCT
alces-483	495	10	wildlife	wildlife	NOUN
alces-483	495	11	)	)	PUNCT
alces-483	496	1	no	no	NOUN
alces-483	496	2	.	.	NOUN
alces-483	497	1	87	87	NUM
alces-483	497	2	.	.	PUNCT
alces-483	498	1	toronto	toronto	PROPN
alces-483	498	2	,	,	PUNCT
alces-483	498	3	ontario	ontario	PROPN
alces-483	498	4	,	,	PUNCT
alces-483	498	5	canada	canada	PROPN
alces-483	498	6	.	.	PUNCT
alces-483	499	1	polis	polis	PROPN
alces-483	499	2	,	,	PUNCT
alces-483	499	3	g.	g.	PROPN
alces-483	499	4	a.	a.	PROPN
alces-483	499	5	,	,	PUNCT
alces-483	499	6	a.	a.	PROPN
alces-483	499	7	l.	l.	PROPN
alces-483	499	8	w.	w.	PROPN
alces-483	499	9	sears	sears	PROPN
alces-483	499	10	,	,	PUNCT
alces-483	499	11	g.	g.	PROPN
alces-483	499	12	r.	r.	PROPN
alces-483	499	13	huxel	huxel	PROPN
alces-483	499	14	,	,	PUNCT
alces-483	499	15	d.	d.	PROPN
alces-483	499	16	r.	r.	PROPN
alces-483	499	17	strong	strong	PROPN
alces-483	499	18	,	,	PUNCT
alces-483	499	19	and	and	CCONJ
alces-483	499	20	j.	j.	PROPN
alces-483	499	21	maron	maron	PROPN
alces-483	499	22	.	.	PUNCT
alces-483	499	23	2000	2000	NUM
alces-483	499	24	.	.	PUNCT
alces-483	500	1	when	when	SCONJ
alces-483	500	2	is	be	AUX
alces-483	500	3	a	a	DET
alces-483	500	4	trophic	trophic	ADJ
alces-483	500	5	cascade	cascade	NOUN
alces-483	500	6	a	a	DET
alces-483	500	7	trophic	trophic	ADJ
alces-483	500	8	cascade	cascade	NOUN
alces-483	500	9	?	?	PUNCT
alces-483	501	1	trends	trend	NOUN
alces-483	501	2	in	in	ADP
alces-483	501	3	ecology	ecology	NOUN
alces-483	501	4	and	and	CCONJ
alces-483	501	5	evolution	evolution	NOUN
alces-483	501	6	15:473	15:473	NUM
alces-483	501	7	-	-	SYM
alces-483	501	8	475	475	NUM
alces-483	501	9	.	.	PUNCT
alces-483	502	1	post	post	PROPN
alces-483	502	2	,	,	PUNCT
alces-483	502	3	e.	e.	PROPN
alces-483	502	4	,	,	PUNCT
alces-483	502	5	r.	r.	PROPN
alces-483	502	6	o.	o.	PROPN
alces-483	502	7	peterson	peterson	PROPN
alces-483	502	8	,	,	PUNCT
alces-483	502	9	n.	n.	PROPN
alces-483	502	10	c.	c.	PROPN
alces-483	502	11	stenseth	stenseth	PROPN
alces-483	502	12	,	,	PUNCT
alces-483	502	13	and	and	CCONJ
alces-483	502	14	b.	b.	PROPN
alces-483	502	15	e.	e.	PROPN
alces-483	502	16	mclaren	mclaren	PROPN
alces-483	502	17	.	.	PUNCT
alces-483	503	1	1999	1999	NUM
alces-483	503	2	.	.	PUNCT
alces-483	504	1	ecosystem	ecosystem	NOUN
alces-483	504	2	consequences	consequence	NOUN
alces-483	504	3	of	of	ADP
alces-483	504	4	wolf	wolf	PROPN
alces-483	504	5	behavioural	behavioural	ADJ
alces-483	504	6	response	response	NOUN
alces-483	504	7	to	to	ADP
alces-483	504	8	climate	climate	NOUN
alces-483	504	9	.	.	PUNCT
alces-483	505	1	nature	nature	NOUN
alces-483	505	2	401:905	401:905	PROPN
alces-483	505	3	-	-	PUNCT
alces-483	505	4	907	907	NUM
alces-483	505	5	.	.	PUNCT
alces-483	506	1	potvin	potvin	PROPN
alces-483	506	2	,	,	PUNCT
alces-483	506	3	f.	f.	PROPN
alces-483	506	4	1988	1988	NUM
alces-483	506	5	.	.	PUNCT
alces-483	507	1	wolf	wolf	NOUN
alces-483	507	2	movements	movement	NOUN
alces-483	507	3	and	and	CCONJ
alces-483	507	4	population	population	NOUN
alces-483	507	5	dynamics	dynamic	NOUN
alces-483	507	6	in	in	ADP
alces-483	507	7	papineaulabelle	papineaulabelle	PROPN
alces-483	507	8	reserve	reserve	PROPN
alces-483	507	9	,	,	PUNCT
alces-483	507	10	quebec	quebec	PROPN
alces-483	507	11	.	.	PUNCT
alces-483	508	1	canadian	canadian	PROPN
alces-483	508	2	journal	journal	PROPN
alces-483	508	3	of	of	ADP
alces-483	508	4	zoology	zoology	NOUN
alces-483	508	5	66:1266	66:1266	NOUN
alces-483	508	6	-	-	PUNCT
alces-483	508	7	1273	1273	NUM
alces-483	508	8	.	.	PUNCT
alces-483	509	1	power	power	NOUN
alces-483	509	2	,	,	PUNCT
alces-483	509	3	m.	m.	NOUN
alces-483	509	4	e.	e.	PROPN
alces-483	509	5	2000	2000	NUM
alces-483	509	6	.	.	PUNCT
alces-483	510	1	what	what	PRON
alces-483	510	2	enables	enable	VERB
alces-483	510	3	trophic	trophic	ADJ
alces-483	510	4	cascades	cascade	NOUN
alces-483	510	5	?	?	PUNCT
alces-483	511	1	commentary	commentary	NOUN
alces-483	511	2	on	on	ADP
alces-483	511	3	polis	polis	PROPN
alces-483	511	4	et	et	PROPN
alces-483	511	5	al	al	PROPN
alces-483	511	6	.	.	PUNCT
alces-483	511	7	trends	trend	NOUN
alces-483	511	8	in	in	ADP
alces-483	511	9	ecology	ecology	NOUN
alces-483	511	10	and	and	CCONJ
alces-483	511	11	evolution	evolution	NOUN
alces-483	511	12	15:443444	15:443444	NUM
alces-483	511	13	.	.	PUNCT
alces-483	512	1	risenhoover	risenhoover	PROPN
alces-483	512	2	,	,	PUNCT
alces-483	512	3	k.	k.	PROPN
alces-483	512	4	l.	l.	PROPN
alces-483	512	5	1987	1987	NUM
alces-483	512	6	.	.	PUNCT
alces-483	513	1	winter	winter	NOUN
alces-483	513	2	foraging	foraging	NOUN
alces-483	513	3	strategies	strategy	NOUN
alces-483	513	4	of	of	ADP
alces-483	513	5	moose	moose	NOUN
alces-483	513	6	in	in	ADP
alces-483	513	7	subarctic	subarctic	ADJ
alces-483	513	8	and	and	CCONJ
alces-483	513	9	boreal	boreal	ADJ
alces-483	513	10	forest	forest	NOUN
alces-483	513	11	habitats	habitat	NOUN
alces-483	513	12	.	.	PUNCT
alces-483	514	1	ph.d	ph.d	PROPN
alces-483	514	2	.	.	PUNCT
alces-483	515	1	thesis	thesis	PROPN
alces-483	515	2	,	,	PUNCT
alces-483	515	3	michigan	michigan	PROPN
alces-483	515	4	technological	technological	ADJ
alces-483	515	5	university	university	PROPN
alces-483	515	6	,	,	PUNCT
alces-483	515	7	houghton	houghton	PROPN
alces-483	515	8	,	,	PUNCT
alces-483	515	9	michigan	michigan	PROPN
alces-483	515	10	,	,	PUNCT
alces-483	515	11	usa	usa	PROPN
alces-483	515	12	.	.	PROPN
alces-483	515	13	rolley	rolley	PROPN
alces-483	515	14	,	,	PUNCT
alces-483	515	15	r.	r.	PROPN
alces-483	515	16	e.	e.	PROPN
alces-483	515	17	,	,	PUNCT
alces-483	515	18	and	and	CCONJ
alces-483	515	19	l.	l.	PROPN
alces-483	515	20	b.	b.	PROPN
alces-483	515	21	keith	keith	PROPN
alces-483	515	22	.	.	PUNCT
alces-483	516	1	1980	1980	NUM
alces-483	516	2	.	.	PUNCT
alces-483	517	1	moose	moose	NOUN
alces-483	517	2	population	population	NOUN
alces-483	517	3	dynamics	dynamic	NOUN
alces-483	517	4	and	and	CCONJ
alces-483	517	5	winter	winter	NOUN
alces-483	517	6	habitat	habitat	NOUN
alces-483	517	7	use	use	NOUN
alces-483	517	8	at	at	ADP
alces-483	517	9	rochester	rochester	PROPN
alces-483	517	10	,	,	PUNCT
alces-483	517	11	alberta	alberta	PROPN
alces-483	517	12	,	,	PUNCT
alces-483	517	13	19651979	19651979	NUM
alces-483	517	14	.	.	PUNCT
alces-483	518	1	canadian	canadian	ADJ
alces-483	518	2	field	field	NOUN
alces-483	518	3	-	-	PUNCT
alces-483	518	4	naturalist	naturalist	NOUN
alces-483	518	5	89:4752	89:4752	NUM
alces-483	518	6	.	.	PUNCT
alces-483	519	1	rosenzweig	rosenzweig	PROPN
alces-483	519	2	,	,	PUNCT
alces-483	519	3	m.	m.	NOUN
alces-483	519	4	1968	1968	NUM
alces-483	519	5	.	.	PUNCT
alces-483	520	1	net	net	ADJ
alces-483	520	2	primary	primary	ADJ
alces-483	520	3	productivity	productivity	NOUN
alces-483	520	4	of	of	ADP
alces-483	520	5	terrestrial	terrestrial	ADJ
alces-483	520	6	communities	community	NOUN
alces-483	520	7	:	:	PUNCT
alces-483	520	8	prediction	prediction	NOUN
alces-483	520	9	from	from	ADP
alces-483	520	10	climatological	climatological	ADJ
alces-483	520	11	data	datum	NOUN
alces-483	520	12	.	.	PUNCT
alces-483	521	1	american	american	PROPN
alces-483	521	2	naturalist	naturalist	NOUN
alces-483	521	3	102:67	102:67	NUM
alces-483	521	4	-	-	SYM
alces-483	521	5	74	74	NUM
alces-483	521	6	.	.	PUNCT
alces-483	521	7	.	.	PUNCT
alces-483	522	1	1969	1969	NUM
alces-483	522	2	.	.	PUNCT
alces-483	523	1	why	why	SCONJ
alces-483	523	2	does	do	AUX
alces-483	523	3	the	the	DET
alces-483	523	4	prey	prey	NOUN
alces-483	523	5	isocline	isocline	NOUN
alces-483	523	6	have	have	VERB
alces-483	523	7	a	a	DET
alces-483	523	8	hump	hump	NOUN
alces-483	523	9	?	?	PUNCT
alces-483	524	1	american	american	ADJ
alces-483	524	2	naturalist	naturalist	PROPN
alces-483	524	3	103:81	103:81	NUM
alces-483	524	4	-	-	SYM
alces-483	524	5	87	87	NUM
alces-483	524	6	.	.	PUNCT
alces-483	524	7	.	.	PUNCT
alces-483	525	1	1971	1971	NUM
alces-483	525	2	.	.	PUNCT
alces-483	526	1	the	the	DET
alces-483	526	2	paradox	paradox	NOUN
alces-483	526	3	of	of	ADP
alces-483	526	4	enrichment	enrichment	NOUN
alces-483	526	5	:	:	PUNCT
alces-483	526	6	destabilization	destabilization	NOUN
alces-483	526	7	of	of	ADP
alces-483	526	8	exploitation	exploitation	NOUN
alces-483	526	9	ecosystems	ecosystem	NOUN
alces-483	526	10	in	in	ADP
alces-483	526	11	ecological	ecological	ADJ
alces-483	526	12	time	time	NOUN
alces-483	526	13	.	.	PUNCT
alces-483	527	1	science	science	PROPN
alces-483	527	2	171:385	171:385	PROPN
alces-483	527	3	-	-	SYM
alces-483	527	4	387	387	NUM
alces-483	527	5	.	.	PUNCT
alces-483	527	6	.	.	PUNCT
alces-483	528	1	1973	1973	NUM
alces-483	528	2	.	.	PUNCT
alces-483	529	1	exploitation	exploitation	NOUN
alces-483	529	2	in	in	ADP
alces-483	529	3	three	three	NUM
alces-483	529	4	trophic	trophic	ADJ
alces-483	529	5	levels	level	NOUN
alces-483	529	6	.	.	PUNCT
alces-483	530	1	american	american	ADJ
alces-483	530	2	naturalist	naturalist	PROPN
alces-483	530	3	107:275294	107:275294	NUM
alces-483	530	4	.	.	PUNCT
alces-483	531	1	schmitz	schmitz	PROPN
alces-483	531	2	,	,	PUNCT
alces-483	531	3	o.	o.	PROPN
alces-483	531	4	j.	j.	PROPN
alces-483	531	5	,	,	PUNCT
alces-483	531	6	p.	p.	NOUN
alces-483	531	7	a.	a.	NOUN
alces-483	531	8	hambaeck	hambaeck	PROPN
alces-483	531	9	,	,	PUNCT
alces-483	531	10	and	and	CCONJ
alces-483	531	11	a.	a.	PROPN
alces-483	531	12	p.	p.	PROPN
alces-483	531	13	beckerman	beckerman	NOUN
alces-483	531	14	.	.	PUNCT
alces-483	532	1	2000	2000	NUM
alces-483	532	2	.	.	PUNCT
alces-483	533	1	trophic	trophic	ADJ
alces-483	533	2	cascades	cascade	NOUN
alces-483	533	3	in	in	ADP
alces-483	533	4	terrestrial	terrestrial	ADJ
alces-483	533	5	systems	system	NOUN
alces-483	533	6	:	:	PUNCT
alces-483	533	7	a	a	DET
alces-483	533	8	review	review	NOUN
alces-483	533	9	of	of	ADP
alces-483	533	10	the	the	DET
alces-483	533	11	effects	effect	NOUN
alces-483	533	12	of	of	ADP
alces-483	533	13	carnivore	carnivore	ADJ
alces-483	533	14	removals	removal	NOUN
alces-483	533	15	on	on	ADP
alces-483	533	16	plants	plant	NOUN
alces-483	533	17	.	.	PUNCT
alces-483	534	1	american	american	PROPN
alces-483	534	2	naturalist	naturalist	PROPN
alces-483	534	3	155:141	155:141	PROPN
alces-483	534	4	-	-	SYM
alces-483	534	5	153	153	NUM
alces-483	534	6	.	.	PUNCT
alces-483	534	7	schwartz	schwartz	PROPN
alces-483	534	8	,	,	PUNCT
alces-483	534	9	c.	c.	PROPN
alces-483	534	10	c.	c.	PROPN
alces-483	534	11	,	,	PUNCT
alces-483	534	12	and	and	CCONJ
alces-483	534	13	a.	a.	NOUN
alces-483	534	14	w.	w.	PROPN
alces-483	534	15	franzmann	franzmann	PROPN
alces-483	534	16	.	.	PUNCT
alces-483	534	17	1991	1991	NUM
alces-483	534	18	.	.	PUNCT
alces-483	535	1	interrelationships	interrelationship	NOUN
alces-483	535	2	of	of	ADP
alces-483	535	3	black	black	ADJ
alces-483	535	4	bears	bear	NOUN
alces-483	535	5	to	to	AUX
alces-483	535	6	moose	moose	NOUN
alces-483	535	7	and	and	CCONJ
alces-483	535	8	forest	forest	NOUN
alces-483	535	9	succession	succession	NOUN
alces-483	535	10	in	in	ADP
alces-483	535	11	the	the	DET
alces-483	535	12	northern	northern	ADJ
alces-483	535	13	coniferous	coniferous	ADJ
alces-483	535	14	forest	forest	NOUN
alces-483	535	15	.	.	PUNCT
alces-483	536	1	wildlife	wildlife	NOUN
alces-483	536	2	monographs	monograph	NOUN
alces-483	536	3	113	113	NUM
alces-483	536	4	.	.	PUNCT
alces-483	537	1	seip	seip	PROPN
alces-483	537	2	,	,	PUNCT
alces-483	537	3	d.	d.	PROPN
alces-483	537	4	r.	r.	PROPN
alces-483	537	5	1991	1991	NUM
alces-483	537	6	.	.	PUNCT
alces-483	538	1	predation	predation	NOUN
alces-483	538	2	and	and	CCONJ
alces-483	538	3	caribou	caribou	NOUN
alces-483	538	4	populations	population	NOUN
alces-483	538	5	.	.	PUNCT
alces-483	539	1	proceedings	proceeding	NOUN
alces-483	539	2	the	the	DET
alces-483	539	3	north	north	PROPN
alces-483	539	4	american	american	PROPN
alces-483	539	5	caribou	caribou	PROPN
alces-483	539	6	workshop	workshop	NOUN
alces-483	539	7	5:46	5:46	NUM
alces-483	539	8	-	-	SYM
alces-483	539	9	52	52	NUM
alces-483	539	10	.	.	PUNCT
alces-483	539	11	.	.	PUNCT
alces-483	540	1	1992	1992	NUM
alces-483	540	2	.	.	PUNCT
alces-483	541	1	factors	factor	NOUN
alces-483	541	2	limiting	limit	VERB
alces-483	541	3	woodland	woodland	NOUN
alces-483	541	4	caribou	caribou	NOUN
alces-483	541	5	populations	population	NOUN
alces-483	541	6	and	and	CCONJ
alces-483	541	7	their	their	PRON
alces-483	541	8	interrelationships	interrelationship	NOUN
alces-483	541	9	with	with	ADP
alces-483	541	10	wolves	wolf	NOUN
alces-483	541	11	and	and	CCONJ
alces-483	541	12	moose	moose	NOUN
alces-483	541	13	in	in	ADP
alces-483	541	14	southeastern	southeastern	ADJ
alces-483	541	15	british	british	PROPN
alces-483	541	16	columbia	columbia	PROPN
alces-483	541	17	.	.	PUNCT
alces-483	542	1	canadian	canadian	ADJ
alces-483	542	2	journal	journal	PROPN
alces-483	542	3	of	of	ADP
alces-483	542	4	zoology	zoology	NOUN
alces-483	542	5	70:1494	70:1494	PROPN
alces-483	542	6	-	-	SYM
alces-483	542	7	1503	1503	NUM
alces-483	542	8	.	.	PUNCT
alces-483	543	1	space	space	NOUN
alces-483	543	2	and	and	CCONJ
alces-483	543	3	time	time	NOUN
alces-483	543	4	in	in	ADP
alces-483	543	5	predation	predation	NOUN
alces-483	543	6	dynamics	dynamic	NOUN
alces-483	543	7	–	–	PUNCT
alces-483	543	8	peterson	peterson	PROPN
alces-483	543	9	et	et	PROPN
alces-483	543	10	al	al	PROPN
alces-483	543	11	.	.	PROPN
alces-483	544	1	alces	alce	NOUN
alces-483	544	2	vol	vol	NOUN
alces-483	544	3	.	.	PROPN
alces-483	544	4	39	39	NUM
alces-483	544	5	,	,	PUNCT
alces-483	544	6	2003	2003	NUM
alces-483	544	7	230	230	NUM
alces-483	544	8	.	.	PUNCT
alces-483	545	1	1995	1995	NUM
alces-483	545	2	.	.	PUNCT
alces-483	546	1	introduction	introduction	NOUN
alces-483	546	2	to	to	ADP
alces-483	546	3	wolf	wolf	NOUN
alces-483	546	4	-	-	PUNCT
alces-483	546	5	prey	prey	NOUN
alces-483	546	6	interactions	interaction	NOUN
alces-483	546	7	.	.	PUNCT
alces-483	547	1	pages	page	NOUN
alces-483	547	2	179	179	NUM
alces-483	547	3	-	-	SYM
alces-483	547	4	186	186	NUM
alces-483	547	5	in	in	ADP
alces-483	547	6	l.	l.	PROPN
alces-483	547	7	n.	n.	PROPN
alces-483	547	8	carbyn	carbyn	PROPN
alces-483	547	9	,	,	PUNCT
alces-483	547	10	s.	s.	PROPN
alces-483	547	11	h.	h.	PROPN
alces-483	547	12	fritts	fritts	PROPN
alces-483	547	13	,	,	PUNCT
alces-483	547	14	and	and	CCONJ
alces-483	547	15	d.	d.	PROPN
alces-483	547	16	r.	r.	PROPN
alces-483	547	17	seip	seip	PROPN
alces-483	547	18	,	,	PUNCT
alces-483	547	19	editors	editor	NOUN
alces-483	547	20	.	.	PUNCT
alces-483	548	1	ecology	ecology	NOUN
alces-483	548	2	and	and	CCONJ
alces-483	548	3	conservation	conservation	NOUN
alces-483	548	4	of	of	ADP
alces-483	548	5	wolves	wolf	NOUN
alces-483	548	6	in	in	ADP
alces-483	548	7	a	a	DET
alces-483	548	8	changing	change	VERB
alces-483	548	9	world	world	NOUN
alces-483	548	10	.	.	PUNCT
alces-483	549	1	canadian	canadian	ADJ
alces-483	549	2	circumpolar	circumpolar	PROPN
alces-483	549	3	institute	institute	PROPN
alces-483	549	4	,	,	PUNCT
alces-483	549	5	occasional	occasional	ADJ
alces-483	549	6	publication	publication	NOUN
alces-483	549	7	no	no	INTJ
alces-483	549	8	.	.	PROPN
alces-483	549	9	35	35	NUM
alces-483	549	10	,	,	PUNCT
alces-483	549	11	edmonton	edmonton	PROPN
alces-483	549	12	,	,	PUNCT
alces-483	549	13	alberta	alberta	PROPN
alces-483	549	14	,	,	PUNCT
alces-483	549	15	canada	canada	PROPN
alces-483	549	16	.	.	PUNCT
alces-483	549	17	shipley	shipley	PROPN
alces-483	549	18	,	,	PUNCT
alces-483	549	19	l.	l.	PROPN
alces-483	549	20	a.	a.	PROPN
alces-483	549	21	,	,	PUNCT
alces-483	549	22	a.	a.	PROPN
alces-483	549	23	w.	w.	PROPN
alces-483	549	24	illius	illius	PROPN
alces-483	549	25	,	,	PUNCT
alces-483	549	26	k.	k.	PROPN
alces-483	549	27	danell	danell	PROPN
alces-483	549	28	,	,	PUNCT
alces-483	549	29	n.	n.	PROPN
alces-483	549	30	t.	t.	PROPN
alces-483	549	31	hobbs	hobbs	PROPN
alces-483	549	32	,	,	PUNCT
alces-483	549	33	and	and	CCONJ
alces-483	549	34	d.	d.	PROPN
alces-483	549	35	e.	e.	PROPN
alces-483	549	36	spalinger	spalinger	PROPN
alces-483	549	37	.	.	PUNCT
alces-483	550	1	1999	1999	NUM
alces-483	550	2	.	.	PUNCT
alces-483	551	1	predicting	predict	VERB
alces-483	551	2	bite	bite	NOUN
alces-483	551	3	size	size	NOUN
alces-483	551	4	selection	selection	NOUN
alces-483	551	5	of	of	ADP
alces-483	551	6	mammalian	mammalian	ADJ
alces-483	551	7	herbivores	herbivore	NOUN
alces-483	551	8	:	:	PUNCT
alces-483	551	9	a	a	DET
alces-483	551	10	test	test	NOUN
alces-483	551	11	of	of	ADP
alces-483	551	12	a	a	DET
alces-483	551	13	general	general	ADJ
alces-483	551	14	model	model	NOUN
alces-483	551	15	of	of	ADP
alces-483	551	16	diet	diet	PROPN
alces-483	551	17	optimization	optimization	NOUN
alces-483	551	18	.	.	PUNCT
alces-483	552	1	oikos	oikos	PROPN
alces-483	552	2	84:5568	84:5568	PROPN
alces-483	552	3	.	.	PUNCT
alces-483	553	1	sinclair	sinclair	PROPN
alces-483	553	2	,	,	PUNCT
alces-483	553	3	a.	a.	PROPN
alces-483	553	4	r.	r.	PROPN
alces-483	553	5	e.	e.	PROPN
alces-483	553	6	1985	1985	PROPN
alces-483	553	7	.	.	PUNCT
alces-483	554	1	does	do	VERB
alces-483	554	2	interspecific	interspecific	NOUN
alces-483	554	3	competition	competition	NOUN
alces-483	554	4	or	or	CCONJ
alces-483	554	5	predation	predation	NOUN
alces-483	554	6	shape	shape	NOUN
alces-483	554	7	the	the	DET
alces-483	554	8	african	african	ADJ
alces-483	554	9	ungulate	ungulate	ADJ
alces-483	554	10	community	community	NOUN
alces-483	554	11	?	?	PUNCT
alces-483	555	1	journal	journal	NOUN
alces-483	555	2	of	of	ADP
alces-483	555	3	animal	animal	NOUN
alces-483	555	4	ecology	ecology	NOUN
alces-483	555	5	54:899	54:899	NUM
alces-483	555	6	-	-	SYM
alces-483	555	7	918	918	NUM
alces-483	555	8	.	.	PUNCT
alces-483	555	9	,	,	PUNCT
alces-483	555	10	c.	c.	PROPN
alces-483	555	11	j.	j.	PROPN
alces-483	555	12	krebs	krebs	PROPN
alces-483	555	13	,	,	PUNCT
alces-483	555	14	j.	j.	PROPN
alces-483	555	15	m.	m.	PROPN
alces-483	555	16	fryxell	fryxell	PROPN
alces-483	555	17	,	,	PUNCT
alces-483	555	18	r.	r.	PROPN
alces-483	555	19	turkington	turkington	PROPN
alces-483	555	20	,	,	PUNCT
alces-483	555	21	s.	s.	PROPN
alces-483	555	22	boutin	boutin	PROPN
alces-483	555	23	,	,	PUNCT
alces-483	555	24	r.	r.	PROPN
alces-483	555	25	boonstra	boonstra	PROPN
alces-483	555	26	,	,	PUNCT
alces-483	555	27	p.	p.	NOUN
alces-483	555	28	seccombe	seccombe	NOUN
alces-483	555	29	-	-	PUNCT
alces-483	555	30	hett	hett	PROPN
alces-483	555	31	,	,	PUNCT
alces-483	555	32	p.	p.	PROPN
alces-483	555	33	lundberg	lundberg	PROPN
alces-483	555	34	,	,	PUNCT
alces-483	555	35	and	and	CCONJ
alces-483	555	36	l.	l.	PROPN
alces-483	555	37	oksanen	oksanen	PROPN
alces-483	555	38	.	.	PROPN
alces-483	555	39	2000	2000	NUM
alces-483	555	40	.	.	PUNCT
alces-483	556	1	testing	test	VERB
alces-483	556	2	hypotheses	hypothesis	NOUN
alces-483	556	3	of	of	ADP
alces-483	556	4	trophic	trophic	ADJ
alces-483	556	5	level	level	NOUN
alces-483	556	6	interactions	interaction	NOUN
alces-483	556	7	:	:	PUNCT
alces-483	556	8	a	a	DET
alces-483	556	9	boreal	boreal	ADJ
alces-483	556	10	forest	forest	NOUN
alces-483	556	11	ecosystem	ecosystem	NOUN
alces-483	556	12	.	.	PUNCT
alces-483	557	1	oikos	oikos	PROPN
alces-483	557	2	89:313	89:313	PROPN
alces-483	557	3	-	-	PUNCT
alces-483	557	4	328	328	NUM
alces-483	557	5	.	.	PUNCT
alces-483	558	1	singer	singer	NOUN
alces-483	558	2	,	,	PUNCT
alces-483	558	3	f.	f.	PROPN
alces-483	558	4	j.	j.	PROPN
alces-483	558	5	,	,	PUNCT
alces-483	558	6	and	and	CCONJ
alces-483	558	7	j.	j.	PROPN
alces-483	558	8	dalle	dalle	PROPN
alces-483	558	9	-	-	PROPN
alces-483	558	10	molle	molle	NOUN
alces-483	558	11	.	.	PUNCT
alces-483	559	1	1985	1985	NUM
alces-483	559	2	.	.	PUNCT
alces-483	560	1	the	the	DET
alces-483	560	2	denali	denali	NOUN
alces-483	560	3	ungulate	ungulate	ADJ
alces-483	560	4	-	-	PUNCT
alces-483	560	5	predator	predator	NOUN
alces-483	560	6	system	system	NOUN
alces-483	560	7	.	.	PUNCT
alces-483	561	1	alces	alce	NOUN
alces-483	561	2	21:339	21:339	NUM
alces-483	561	3	-	-	SYM
alces-483	561	4	358	358	NUM
alces-483	561	5	.	.	PUNCT
alces-483	562	1	,	,	PUNCT
alces-483	562	2	d.	d.	PROPN
alces-483	562	3	m.	m.	PROPN
alces-483	562	4	swift	swift	PROPN
alces-483	562	5	,	,	PUNCT
alces-483	562	6	m.	m.	PROPN
alces-483	562	7	b.	b.	PROPN
alces-483	562	8	coughenour	coughenour	PROPN
alces-483	562	9	,	,	PUNCT
alces-483	562	10	and	and	CCONJ
alces-483	562	11	j.	j.	PROPN
alces-483	562	12	d.	d.	PROPN
alces-483	562	13	varley	varley	PROPN
alces-483	562	14	.	.	PUNCT
alces-483	563	1	1998a	1998a	NUM
alces-483	563	2	.	.	PUNCT
alces-483	564	1	thunder	thunder	NOUN
alces-483	564	2	on	on	ADP
alces-483	564	3	the	the	DET
alces-483	564	4	yellowstone	yellowstone	NOUN
alces-483	564	5	revisited	revisit	VERB
alces-483	564	6	:	:	PUNCT
alces-483	564	7	an	an	DET
alces-483	564	8	assessment	assessment	NOUN
alces-483	564	9	of	of	ADP
alces-483	564	10	management	management	NOUN
alces-483	564	11	of	of	ADP
alces-483	564	12	native	native	ADJ
alces-483	564	13	ungulates	ungulate	NOUN
alces-483	564	14	by	by	ADP
alces-483	564	15	natural	natural	ADJ
alces-483	564	16	regulation	regulation	NOUN
alces-483	564	17	,	,	PUNCT
alces-483	564	18	1968	1968	NUM
alces-483	564	19	-	-	SYM
alces-483	564	20	1993	1993	NUM
alces-483	564	21	.	.	PUNCT
alces-483	565	1	wildlife	wildlife	NOUN
alces-483	565	2	society	society	NOUN
alces-483	565	3	bulletin	bulletin	NOUN
alces-483	565	4	26:375	26:375	NUM
alces-483	565	5	-	-	SYM
alces-483	565	6	390	390	NUM
alces-483	565	7	.	.	PUNCT
alces-483	565	8	,	,	PUNCT
alces-483	565	9	l.	l.	PROPN
alces-483	565	10	c.	c.	PROPN
alces-483	565	11	zeigenfuss	zeigenfuss	PROPN
alces-483	565	12	,	,	PUNCT
alces-483	565	13	r.	r.	PROPN
alces-483	565	14	g.	g.	PROPN
alces-483	565	15	cates	cat	VERB
alces-483	565	16	,	,	PUNCT
alces-483	565	17	and	and	CCONJ
alces-483	565	18	d.	d.	PROPN
alces-483	565	19	t.	t.	PROPN
alces-483	565	20	barnett	barnett	PROPN
alces-483	565	21	.	.	PUNCT
alces-483	566	1	1998b	1998b	PROPN
alces-483	566	2	.	.	PUNCT
alces-483	567	1	elk	elk	NOUN
alces-483	567	2	,	,	PUNCT
alces-483	567	3	multiple	multiple	ADJ
alces-483	567	4	factors	factor	NOUN
alces-483	567	5	,	,	PUNCT
alces-483	567	6	and	and	CCONJ
alces-483	567	7	persistence	persistence	NOUN
alces-483	567	8	of	of	ADP
alces-483	567	9	willows	willow	NOUN
alces-483	567	10	in	in	ADP
alces-483	567	11	national	national	ADJ
alces-483	567	12	parks	park	NOUN
alces-483	567	13	.	.	PUNCT
alces-483	568	1	wildlife	wildlife	NOUN
alces-483	568	2	society	society	NOUN
alces-483	568	3	bulletin	bulletin	NOUN
alces-483	568	4	26:419	26:419	NUM
alces-483	568	5	-	-	SYM
alces-483	568	6	428	428	NUM
alces-483	568	7	.	.	PUNCT
alces-483	569	1	spalinger	spalinger	NOUN
alces-483	569	2	,	,	PUNCT
alces-483	569	3	d.	d.	PROPN
alces-483	569	4	e.	e.	PROPN
alces-483	569	5	,	,	PUNCT
alces-483	569	6	and	and	CCONJ
alces-483	569	7	n.	n.	PROPN
alces-483	569	8	t.	t.	PROPN
alces-483	569	9	hobbs	hobbs	PROPN
alces-483	569	10	.	.	PUNCT
alces-483	570	1	1992	1992	NUM
alces-483	570	2	.	.	PUNCT
alces-483	570	3	mechanisms	mechanism	NOUN
alces-483	570	4	of	of	ADP
alces-483	570	5	foraging	forage	VERB
alces-483	570	6	in	in	ADP
alces-483	570	7	mammalian	mammalian	ADJ
alces-483	570	8	herbivores	herbivore	NOUN
alces-483	570	9	:	:	PUNCT
alces-483	570	10	new	new	ADJ
alces-483	570	11	models	model	NOUN
alces-483	570	12	of	of	ADP
alces-483	570	13	functional	functional	ADJ
alces-483	570	14	response	response	NOUN
alces-483	570	15	.	.	PUNCT
alces-483	571	1	american	american	PROPN
alces-483	571	2	naturalist	naturalist	PROPN
alces-483	571	3	140:325348	140:325348	NUM
alces-483	571	4	.	.	PUNCT
alces-483	572	1	stenlund	stenlund	PROPN
alces-483	572	2	,	,	PUNCT
alces-483	572	3	m.	m.	NOUN
alces-483	572	4	h.	h.	PROPN
alces-483	572	5	1955	1955	NUM
alces-483	572	6	.	.	PUNCT
alces-483	573	1	a	a	DET
alces-483	573	2	field	field	NOUN
alces-483	573	3	study	study	NOUN
alces-483	573	4	of	of	ADP
alces-483	573	5	the	the	DET
alces-483	573	6	timber	timber	NOUN
alces-483	573	7	wolf	wolf	NOUN
alces-483	573	8	(	(	PUNCT
alces-483	573	9	canis	canis	NOUN
alces-483	573	10	lupus	lupus	NOUN
alces-483	573	11	)	)	PUNCT
alces-483	573	12	on	on	ADP
alces-483	573	13	the	the	DET
alces-483	573	14	superior	superior	ADJ
alces-483	573	15	national	national	PROPN
alces-483	573	16	forest	forest	NOUN
alces-483	573	17	,	,	PUNCT
alces-483	573	18	minnesota	minnesota	PROPN
alces-483	573	19	.	.	PUNCT
alces-483	574	1	minnesota	minnesota	PROPN
alces-483	574	2	department	department	PROPN
alces-483	574	3	of	of	ADP
alces-483	574	4	conservation	conservation	PROPN
alces-483	574	5	technical	technical	ADJ
alces-483	574	6	bulletin	bulletin	NOUN
alces-483	574	7	no	no	INTJ
alces-483	574	8	.	.	NOUN
alces-483	575	1	4	4	NUM
alces-483	575	2	.	.	X
alces-483	575	3	st	st	PROPN
alces-483	575	4	.	.	PROPN
alces-483	575	5	paul	paul	PROPN
alces-483	575	6	,	,	PUNCT
alces-483	575	7	minnesota	minnesota	PROPN
alces-483	575	8	,	,	PUNCT
alces-483	575	9	usa	usa	PROPN
alces-483	575	10	.	.	PROPN
alces-483	575	11	thomas	thomas	PROPN
alces-483	575	12	,	,	PUNCT
alces-483	575	13	d.	d.	PROPN
alces-483	575	14	c.	c.	PROPN
alces-483	575	15	1995	1995	NUM
alces-483	575	16	.	.	PUNCT
alces-483	576	1	a	a	DET
alces-483	576	2	review	review	NOUN
alces-483	576	3	of	of	ADP
alces-483	576	4	wolfcaribou	wolfcaribou	PROPN
alces-483	576	5	relationships	relationship	NOUN
alces-483	576	6	and	and	CCONJ
alces-483	576	7	conservation	conservation	NOUN
alces-483	576	8	implications	implication	NOUN
alces-483	576	9	in	in	ADP
alces-483	576	10	canada	canada	PROPN
alces-483	576	11	.	.	PUNCT
alces-483	577	1	pages	page	NOUN
alces-483	577	2	261	261	NUM
alces-483	577	3	-	-	SYM
alces-483	577	4	273	273	NUM
alces-483	577	5	in	in	ADP
alces-483	577	6	l.	l.	PROPN
alces-483	577	7	n.	n.	PROPN
alces-483	577	8	carbyn	carbyn	PROPN
alces-483	577	9	,	,	PUNCT
alces-483	577	10	s.	s.	PROPN
alces-483	577	11	h.	h.	PROPN
alces-483	577	12	fritts	fritts	PROPN
alces-483	577	13	,	,	PUNCT
alces-483	577	14	and	and	CCONJ
alces-483	577	15	d.	d.	PROPN
alces-483	577	16	r.	r.	PROPN
alces-483	577	17	seip	seip	PROPN
alces-483	577	18	,	,	PUNCT
alces-483	577	19	editors	editor	NOUN
alces-483	577	20	.	.	PUNCT
alces-483	578	1	ecology	ecology	NOUN
alces-483	578	2	and	and	CCONJ
alces-483	578	3	conservation	conservation	NOUN
alces-483	578	4	of	of	ADP
alces-483	578	5	wolves	wolf	NOUN
alces-483	578	6	in	in	ADP
alces-483	578	7	a	a	DET
alces-483	578	8	changing	change	VERB
alces-483	578	9	world	world	NOUN
alces-483	578	10	.	.	PUNCT
alces-483	579	1	canadian	canadian	ADJ
alces-483	579	2	circumpolar	circumpolar	PROPN
alces-483	579	3	institute	institute	PROPN
alces-483	579	4	,	,	PUNCT
alces-483	579	5	occasional	occasional	ADJ
alces-483	579	6	publication	publication	NOUN
alces-483	579	7	no	no	INTJ
alces-483	579	8	.	.	PROPN
alces-483	579	9	35	35	NUM
alces-483	579	10	,	,	PUNCT
alces-483	579	11	edmonton	edmonton	PROPN
alces-483	579	12	,	,	PUNCT
alces-483	579	13	alberta	alberta	PROPN
alces-483	579	14	,	,	PUNCT
alces-483	579	15	canada	canada	PROPN
alces-483	579	16	.	.	PUNCT
alces-483	580	1	van	van	PROPN
alces-483	580	2	ballenberghe	ballenberghe	NOUN
alces-483	580	3	,	,	PUNCT
alces-483	580	4	v.	v.	ADP
alces-483	580	5	1987	1987	NUM
alces-483	580	6	.	.	PUNCT
alces-483	581	1	effects	effect	NOUN
alces-483	581	2	of	of	ADP
alces-483	581	3	predation	predation	NOUN
alces-483	581	4	on	on	ADP
alces-483	581	5	moose	moose	NOUN
alces-483	581	6	numbers	number	NOUN
alces-483	581	7	:	:	PUNCT
alces-483	581	8	a	a	DET
alces-483	581	9	review	review	NOUN
alces-483	581	10	of	of	ADP
alces-483	581	11	recent	recent	ADJ
alces-483	581	12	north	north	ADJ
alces-483	581	13	american	american	ADJ
alces-483	581	14	studies	study	NOUN
alces-483	581	15	.	.	PUNCT
alces-483	582	1	swedish	swedish	ADJ
alces-483	582	2	wildlife	wildlife	NOUN
alces-483	582	3	research	research	NOUN
alces-483	582	4	supplement	supplement	NOUN
alces-483	582	5	1:431	1:431	NUM
alces-483	582	6	-	-	SYM
alces-483	582	7	460	460	NUM
alces-483	582	8	.	.	PUNCT
alces-483	582	9	,	,	PUNCT
alces-483	582	10	and	and	CCONJ
alces-483	582	11	w.	w.	PROPN
alces-483	582	12	ballard	ballard	PROPN
alces-483	582	13	.	.	PUNCT
alces-483	582	14	1994	1994	NUM
alces-483	582	15	.	.	PUNCT
alces-483	583	1	limitation	limitation	NOUN
alces-483	583	2	and	and	CCONJ
alces-483	583	3	regulation	regulation	NOUN
alces-483	583	4	of	of	ADP
alces-483	583	5	moose	moose	NOUN
alces-483	583	6	populations	population	NOUN
alces-483	583	7	:	:	PUNCT
alces-483	583	8	the	the	DET
alces-483	583	9	role	role	NOUN
alces-483	583	10	of	of	ADP
alces-483	583	11	predation	predation	NOUN
alces-483	583	12	.	.	PUNCT
alces-483	584	1	canadian	canadian	ADJ
alces-483	584	2	journal	journal	PROPN
alces-483	584	3	of	of	ADP
alces-483	584	4	zoology	zoology	NOUN
alces-483	584	5	72:2071	72:2071	PROPN
alces-483	584	6	-	-	SYM
alces-483	584	7	2077	2077	NUM
alces-483	584	8	.	.	PUNCT
alces-483	584	9	,	,	PUNCT
alces-483	584	10	and	and	CCONJ
alces-483	584	11	.	.	PUNCT
alces-483	585	1	1998	1998	NUM
alces-483	585	2	.	.	PUNCT
alces-483	586	1	population	population	NOUN
alces-483	586	2	dynamics	dynamic	NOUN
alces-483	586	3	.	.	PUNCT
alces-483	587	1	pages	page	NOUN
alces-483	587	2	223	223	NUM
alces-483	587	3	-	-	SYM
alces-483	587	4	245	245	NUM
alces-483	587	5	in	in	ADP
alces-483	587	6	ecology	ecology	NOUN
alces-483	587	7	and	and	CCONJ
alces-483	587	8	management	management	NOUN
alces-483	587	9	of	of	ADP
alces-483	587	10	the	the	DET
alces-483	587	11	north	north	ADJ
alces-483	587	12	american	american	ADJ
alces-483	587	13	moose	moose	NOUN
alces-483	587	14	.	.	PUNCT
alces-483	588	1	smithsonian	smithsonian	PROPN
alces-483	588	2	institution	institution	PROPN
alces-483	588	3	press	press	PROPN
alces-483	588	4	,	,	PUNCT
alces-483	588	5	washington	washington	PROPN
alces-483	588	6	,	,	PUNCT
alces-483	588	7	d.c	d.c	PROPN
alces-483	588	8	.	.	PROPN
alces-483	588	9	,	,	PUNCT
alces-483	588	10	usa	usa	PROPN
alces-483	588	11	.	.	PROPN
alces-483	588	12	,	,	PUNCT
alces-483	588	13	a.	a.	PROPN
alces-483	588	14	w.	w.	PROPN
alces-483	588	15	erickson	erickson	PROPN
alces-483	588	16	,	,	PUNCT
alces-483	588	17	and	and	CCONJ
alces-483	588	18	d.	d.	PROPN
alces-483	588	19	byman	byman	NOUN
alces-483	588	20	.	.	PUNCT
alces-483	589	1	1975	1975	NUM
alces-483	589	2	.	.	PUNCT
alces-483	590	1	ecology	ecology	NOUN
alces-483	590	2	of	of	ADP
alces-483	590	3	the	the	DET
alces-483	590	4	timber	timber	NOUN
alces-483	590	5	wolf	wolf	NOUN
alces-483	590	6	in	in	ADP
alces-483	590	7	northeastern	northeastern	ADJ
alces-483	590	8	minnesota	minnesota	PROPN
alces-483	590	9	.	.	PUNCT
alces-483	591	1	wildlife	wildlife	NOUN
alces-483	591	2	monographs	monograph	NOUN
alces-483	591	3	43	43	NUM
alces-483	591	4	.	.	PUNCT
alces-483	591	5	vucetich	vucetich	PROPN
alces-483	591	6	,	,	PUNCT
alces-483	591	7	j.	j.	PROPN
alces-483	591	8	a.	a.	PROPN
alces-483	591	9	,	,	PUNCT
alces-483	591	10	r.	r.	PROPN
alces-483	591	11	o.	o.	PROPN
alces-483	591	12	peterson	peterson	PROPN
alces-483	591	13	,	,	PUNCT
alces-483	591	14	and	and	CCONJ
alces-483	591	15	c.	c.	PROPN
alces-483	591	16	l.	l.	PROPN
alces-483	591	17	schaefer	schaefer	PROPN
alces-483	591	18	.	.	PUNCT
alces-483	592	1	2002	2002	NUM
alces-483	592	2	.	.	PUNCT
alces-483	593	1	the	the	DET
alces-483	593	2	effect	effect	NOUN
alces-483	593	3	of	of	ADP
alces-483	593	4	prey	prey	NOUN
alces-483	593	5	and	and	CCONJ
alces-483	593	6	predator	predator	NOUN
alces-483	593	7	densities	density	NOUN
alces-483	593	8	on	on	ADP
alces-483	593	9	wolf	wolf	PROPN
alces-483	593	10	predation	predation	NOUN
alces-483	593	11	.	.	PUNCT
alces-483	594	1	ecology	ecology	NOUN
alces-483	594	2	83:3003	83:3003	NUM
alces-483	594	3	-	-	SYM
alces-483	594	4	3013	3013	NUM
alces-483	594	5	.	.	PUNCT
alces-483	595	1	white	white	ADJ
alces-483	595	2	,	,	PUNCT
alces-483	595	3	c.	c.	PROPN
alces-483	595	4	a.	a.	PROPN
alces-483	595	5	,	,	PUNCT
alces-483	595	6	c.	c.	PROPN
alces-483	595	7	e.	e.	PROPN
alces-483	595	8	olmsted	olmsted	PROPN
alces-483	595	9	,	,	PUNCT
alces-483	595	10	and	and	CCONJ
alces-483	595	11	c.	c.	PROPN
alces-483	595	12	e.	e.	PROPN
alces-483	595	13	kay	kay	PROPN
alces-483	595	14	.	.	PUNCT
alces-483	596	1	1998	1998	NUM
alces-483	596	2	.	.	PUNCT
alces-483	597	1	aspen	aspen	PROPN
alces-483	597	2	,	,	PUNCT
alces-483	597	3	elk	elk	NOUN
alces-483	597	4	,	,	PUNCT
alces-483	597	5	and	and	CCONJ
alces-483	597	6	fire	fire	NOUN
alces-483	597	7	in	in	ADP
alces-483	597	8	the	the	DET
alces-483	597	9	rocky	rocky	ADJ
alces-483	597	10	mountain	mountain	NOUN
alces-483	597	11	national	national	ADJ
alces-483	597	12	parks	park	NOUN
alces-483	597	13	of	of	ADP
alces-483	597	14	north	north	PROPN
alces-483	597	15	america	america	PROPN
alces-483	597	16	.	.	PUNCT
alces-483	598	1	wildlife	wildlife	NOUN
alces-483	598	2	society	society	PROPN
alces-483	598	3	bulletin	bulletin	PROPN
alces-483	598	4	26:449	26:449	PROPN
alces-483	598	5	-	-	PROPN
alces-483	598	6	462	462	NUM
alces-483	598	7	.	.	PUNCT
alces-483	599	1	white	white	PROPN
alces-483	599	2	,	,	PUNCT
alces-483	599	3	r.	r.	PROPN
alces-483	599	4	g.	g.	PROPN
alces-483	599	5	1983	1983	NUM
alces-483	599	6	.	.	PUNCT
alces-483	600	1	foraging	forage	VERB
alces-483	600	2	patterns	pattern	NOUN
alces-483	600	3	and	and	CCONJ
alces-483	600	4	their	their	PRON
alces-483	600	5	multiplier	multipli	ADJ
alces-483	600	6	effects	effect	NOUN
alces-483	600	7	on	on	ADP
alces-483	600	8	productivity	productivity	NOUN
alces-483	600	9	of	of	ADP
alces-483	600	10	northern	northern	ADJ
alces-483	600	11	ungulates	ungulate	NOUN
alces-483	600	12	.	.	PUNCT
alces-483	601	1	oikos	oikos	PROPN
alces-483	601	2	40:377384	40:377384	PUNCT
alces-483	601	3	.	.	PUNCT
alces-483	602	1	wiens	wiens	PROPN
alces-483	602	2	,	,	PUNCT
alces-483	602	3	j.	j.	PROPN
alces-483	602	4	a.	a.	PROPN
alces-483	602	5	1989	1989	NUM
alces-483	602	6	.	.	PUNCT
alces-483	603	1	spatial	spatial	ADJ
alces-483	603	2	scaling	scaling	NOUN
alces-483	603	3	in	in	ADP
alces-483	603	4	ecology	ecology	NOUN
alces-483	603	5	.	.	PUNCT
alces-483	604	1	functional	functional	ADJ
alces-483	604	2	ecology	ecology	NOUN
alces-483	604	3	3:385	3:385	NUM
alces-483	604	4	-	-	SYM
alces-483	604	5	397	397	NUM
alces-483	604	6	.	.	PUNCT
alces-483	605	1	alces	alce	NOUN
alces-483	605	2	vol	vol	NOUN
alces-483	605	3	.	.	PROPN
alces-483	605	4	39	39	NUM
alces-483	605	5	,	,	PUNCT
alces-483	605	6	2003	2003	NUM
alces-483	605	7	peterson	peterson	PROPN
alces-483	605	8	et	et	PROPN
alces-483	605	9	al	al	PROPN
alces-483	605	10	.	.	PROPN
alces-483	605	11	–	–	PUNCT
alces-483	605	12	space	space	NOUN
alces-483	605	13	and	and	CCONJ
alces-483	605	14	time	time	NOUN
alces-483	605	15	in	in	ADP
alces-483	605	16	predation	predation	NOUN
alces-483	605	17	dynamics	dynamic	NOUN
alces-483	605	18	231	231	NUM
alces-483	605	19	sl	sl	NOUN
alces-483	605	20	at	at	ADP
alces-483	605	21	e	e	PROPN
alces-483	605	22	is	be	AUX
alces-483	605	23	la	la	INTJ
alces-483	605	24	nd	nd	PRON
alces-483	605	25	s	s	PART
alces-483	605	26	ca	can	AUX
alces-483	605	27	rib	rib	NOUN
alces-483	605	28	ou	ou	ADP
alces-483	605	29	6	6	NUM
alces-483	605	30	0	0	NUM
alces-483	605	31	0	0	NUM
alces-483	605	32	0	0	NUM
alces-483	605	33	0	0	NUM
alces-483	605	34	0	0	NUM
alces-483	605	35	12	12	NUM
alces-483	605	36	se	se	ADV
alces-483	605	37	ip	ip	PROPN
alces-483	605	38	(	(	PUNCT
alces-483	605	39	1	1	NUM
alces-483	605	40	99	99	NUM
alces-483	605	41	1	1	NUM
alces-483	605	42	)	)	PUNCT
alces-483	605	43	n	n	NOUN
alces-483	605	44	or	or	CCONJ
alces-483	605	45	w	w	NOUN
alces-483	605	46	ay	ay	NOUN
alces-483	605	47	ca	can	AUX
alces-483	605	48	rib	rib	NOUN
alces-483	605	49	ou	ou	ADP
alces-483	605	50	3.5	3.5	NUM
alces-483	605	51	0	0	NUM
alces-483	605	52	1	1	NUM
alces-483	605	53	0	0	NUM
alces-483	605	54	0	0	NUM
alces-483	605	55	1	1	NUM
alces-483	605	56	7	7	NUM
alces-483	605	57	se	se	X
alces-483	605	58	ip	ip	PROPN
alces-483	605	59	(	(	PUNCT
alces-483	605	60	1	1	NUM
alces-483	605	61	99	99	NUM
alces-483	605	62	1	1	NUM
alces-483	605	63	)	)	PUNCT
alces-483	606	1	n	n	PRON
alces-483	607	1	ew	ew	INTJ
alces-483	607	2	fo	fo	INTJ
alces-483	607	3	un	un	PROPN
alces-483	607	4	dl	dl	PROPN
alces-483	608	1	an	an	PROPN
alces-483	608	2	d	d	PROPN
alces-483	608	3	ca	can	AUX
alces-483	608	4	rib	rib	NOUN
alces-483	608	5	ou	ou	ADP
alces-483	608	6	8.5	8.5	NUM
alces-483	608	7	0	0	NUM
alces-483	608	8	1	1	NUM
alces-483	608	9	0	0	NUM
alces-483	608	10	0	0	NUM
alces-483	608	11	1	1	NUM
alces-483	608	12	17	17	NUM
alces-483	608	13	se	se	X
alces-483	608	14	ip	ip	PROPN
alces-483	608	15	(	(	PUNCT
alces-483	608	16	1	1	NUM
alces-483	608	17	99	99	NUM
alces-483	608	18	1	1	NUM
alces-483	608	19	)	)	PUNCT
alces-483	609	1	so	so	CCONJ
alces-483	609	2	ut	ut	PROPN
alces-483	609	3	h	h	PROPN
alces-483	609	4	g	g	PROPN
alces-483	610	1	eo	eo	INTJ
alces-483	610	2	rg	rg	PROPN
alces-483	610	3	ia	ia	PROPN
alces-483	610	4	ca	can	AUX
alces-483	610	5	rib	rib	NOUN
alces-483	610	6	ou	ou	ADP
alces-483	610	7	2	2	NUM
alces-483	610	8	0	0	NUM
alces-483	610	9	1	1	NUM
alces-483	610	10	0	0	NUM
alces-483	610	11	0	0	NUM
alces-483	610	12	1	1	NUM
alces-483	610	13	4	4	NUM
alces-483	610	14	se	se	ADV
alces-483	610	15	ip	ip	NOUN
alces-483	610	16	(	(	PUNCT
alces-483	610	17	1	1	NUM
alces-483	610	18	99	99	NUM
alces-483	610	19	1	1	NUM
alces-483	610	20	)	)	PUNCT
alces-483	610	21	q	q	PROPN
alces-483	611	1	ue	ue	INTJ
alces-483	611	2	sn	sn	PROPN
alces-483	611	3	el	el	PROPN
alces-483	611	4	l	l	PROPN
alces-483	611	5	ak	ak	PROPN
alces-483	611	6	e	e	PROPN
alces-483	611	7	ca	can	AUX
alces-483	611	8	rib	rib	VERB
alces-483	611	9	ou	ou	ADP
alces-483	611	10	0.0	0.0	NUM
alces-483	611	11	3	3	NUM
alces-483	611	12	1	1	NUM
alces-483	611	13	0	0	NUM
alces-483	611	14	1	1	NUM
alces-483	611	15	0	0	NUM
alces-483	611	16	2	2	NUM
alces-483	611	17	0.0	0.0	NUM
alces-483	611	18	6	6	NUM
alces-483	611	19	se	se	ADV
alces-483	611	20	ip	ip	PROPN
alces-483	611	21	(	(	PUNCT
alces-483	611	22	1	1	NUM
alces-483	611	23	99	99	NUM
alces-483	611	24	1	1	NUM
alces-483	611	25	)	)	PUNCT
alces-483	611	26	o	o	NOUN
alces-483	612	1	nt	nt	INTJ
alces-483	612	2	ar	ar	PROPN
alces-483	612	3	io	io	PROPN
alces-483	612	4	ca	can	AUX
alces-483	612	5	rib	rib	NOUN
alces-483	612	6	ou	ou	ADP
alces-483	612	7	0.0	0.0	NUM
alces-483	612	8	3	3	NUM
alces-483	612	9	1	1	NUM
alces-483	612	10	1	1	NUM
alces-483	612	11	1	1	NUM
alces-483	612	12	0	0	NUM
alces-483	612	13	3	3	NUM
alces-483	612	14	0.0	0.0	NUM
alces-483	612	15	6	6	NUM
alces-483	612	16	se	se	ADV
alces-483	612	17	ip	ip	PROPN
alces-483	612	18	(	(	PUNCT
alces-483	612	19	1	1	NUM
alces-483	612	20	99	99	NUM
alces-483	612	21	1	1	NUM
alces-483	612	22	)	)	PUNCT
alces-483	612	23	sa	sa	NOUN
alces-483	612	24	sk	sk	NOUN
alces-483	612	25	at	at	ADP
alces-483	612	26	ch	ch	PROPN
alces-483	612	27	ew	ew	PROPN
alces-483	612	28	an	an	DET
alces-483	612	29	ca	ca	NOUN
alces-483	612	30	rib	rib	NOUN
alces-483	612	31	ou	ou	ADP
alces-483	612	32	0.0	0.0	NUM
alces-483	612	33	3	3	NUM
alces-483	612	34	1	1	NUM
alces-483	612	35	1	1	NUM
alces-483	612	36	1	1	NUM
alces-483	612	37	0	0	NUM
alces-483	612	38	3	3	NUM
alces-483	612	39	0.0	0.0	NUM
alces-483	612	40	6	6	NUM
alces-483	612	41	se	se	ADV
alces-483	612	42	ip	ip	PROPN
alces-483	612	43	(	(	PUNCT
alces-483	612	44	1	1	NUM
alces-483	612	45	99	99	NUM
alces-483	612	46	1	1	NUM
alces-483	612	47	)	)	PUNCT
alces-483	612	48	s.	s.	PROPN
alces-483	612	49	f	f	PROPN
alces-483	613	1	in	in	ADP
alces-483	613	2	la	la	PROPN
alces-483	613	3	nd	nd	ADV
alces-483	613	4	m	m	VERB
alces-483	613	5	oo	oo	INTJ
alces-483	613	6	se	se	ADV
alces-483	613	7	0.4	0.4	NUM
alces-483	613	8	0	0	NUM
alces-483	613	9	1	1	NUM
alces-483	613	10	0	0	NUM
alces-483	613	11	0	0	NUM
alces-483	613	12	1	1	NUM
alces-483	613	13	2.4	2.4	NUM
alces-483	613	14	n	n	NUM
alces-483	613	15	yg	yg	PROPN
alces-483	613	16	re	re	PROPN
alces-483	613	17	n	n	PROPN
alces-483	613	18	(	(	PUNCT
alces-483	613	19	19	19	NUM
alces-483	613	20	87	87	NUM
alces-483	613	21	)	)	PUNCT
alces-483	613	22	a	a	PRON
alces-483	613	23	lb	lb	INTJ
alces-483	613	24	er	er	INTJ
alces-483	613	25	ta	ta	PART
alces-483	613	26	m	m	ADV
alces-483	613	27	oo	oo	INTJ
alces-483	613	28	se	se	ADP
alces-483	613	29	0.8	0.8	NUM
alces-483	613	30	0	0	NUM
alces-483	613	31	1	1	NUM
alces-483	613	32	1	1	NUM
alces-483	613	33	0	0	NUM
alces-483	613	34	2	2	NUM
alces-483	613	35	4.8	4.8	NUM
alces-483	613	36	r	r	NOUN
alces-483	613	37	ol	ol	NOUN
alces-483	613	38	le	le	X
alces-483	613	39	y	y	PROPN
alces-483	613	40	a	a	DET
alces-483	613	41	nd	nd	INTJ
alces-483	613	42	k	k	PROPN
alces-483	613	43	ei	ei	X
alces-483	613	44	th	th	X
alces-483	613	45	(	(	PUNCT
alces-483	613	46	1	1	NUM
alces-483	613	47	98	98	NUM
alces-483	613	48	0	0	NUM
alces-483	613	49	)	)	PUNCT
alces-483	613	50	sw	sw	PROPN
alces-483	614	1	ed	ed	PROPN
alces-483	614	2	en	en	PROPN
alces-483	614	3	m	m	VERB
alces-483	614	4	oo	oo	INTJ
alces-483	614	5	se	se	PROPN
alces-483	614	6	1.5	1.5	NUM
alces-483	614	7	0	0	NUM
alces-483	614	8	1	1	NUM
alces-483	614	9	0	0	NUM
alces-483	614	10	0	0	NUM
alces-483	614	11	1	1	NUM
alces-483	614	12	9	9	NUM
alces-483	614	13	c	c	NOUN
alces-483	614	14	ed	ed	NOUN
alces-483	615	1	er	er	INTJ
alces-483	615	2	lu	lu	NOUN
alces-483	615	3	nd	nd	INTJ
alces-483	615	4	an	an	PROPN
alces-483	615	5	d	d	PROPN
alces-483	615	6	m	m	PROPN
alces-483	615	7	ar	ar	PROPN
alces-483	615	8	kg	kg	X
alces-483	615	9	re	re	ADP
alces-483	615	10	n	n	PROPN
alces-483	615	11	(	(	PUNCT
alces-483	615	12	19	19	NUM
alces-483	615	13	87	87	NUM
alces-483	615	14	)	)	PUNCT
alces-483	616	1	el	el	PROPN
alces-483	616	2	k	k	PROPN
alces-483	617	1	i	i	PRON
alces-483	617	2	sl	sl	VERB
alces-483	617	3	an	an	DET
alces-483	617	4	d	d	PROPN
alces-483	617	5	,	,	PUNCT
alces-483	617	6	a	a	DET
alces-483	617	7	lb	lb	INTJ
alces-483	617	8	er	er	INTJ
alces-483	617	9	ta	ta	PART
alces-483	617	10	m	m	ADV
alces-483	617	11	oo	oo	INTJ
alces-483	617	12	se	se	ADP
alces-483	617	13	1.5	1.5	NUM
alces-483	617	14	0	0	NUM
alces-483	617	15	1	1	NUM
alces-483	617	16	0	0	NUM
alces-483	617	17	0	0	NUM
alces-483	617	18	1	1	NUM
alces-483	617	19	9	9	NUM
alces-483	617	20	c	c	NOUN
alces-483	617	21	ai	ai	VERB
alces-483	617	22	rn	rn	PROPN
alces-483	617	23	s	s	PROPN
alces-483	617	24	a	a	PRON
alces-483	617	25	nd	nd	NOUN
alces-483	617	26	t	t	PROPN
alces-483	617	27	el	el	PROPN
alces-483	617	28	fe	fe	PROPN
alces-483	618	1	r	r	NOUN
alces-483	618	2	(	(	PUNCT
alces-483	618	3	19	19	NUM
alces-483	618	4	80	80	NUM
alces-483	618	5	)	)	PUNCT
alces-483	618	6	n	n	CCONJ
alces-483	619	1	ew	ew	INTJ
alces-483	619	2	fo	fo	INTJ
alces-483	619	3	un	un	PROPN
alces-483	619	4	dl	dl	PROPN
alces-483	620	1	an	an	PROPN
alces-483	620	2	d	d	NOUN
alces-483	620	3	m	m	VERB
alces-483	620	4	oo	oo	INTJ
alces-483	620	5	se	se	ADP
alces-483	620	6	1.8	1.8	NUM
alces-483	620	7	0	0	NUM
alces-483	620	8	1	1	NUM
alces-483	620	9	0	0	NUM
alces-483	620	10	0	0	NUM
alces-483	620	11	1	1	NUM
alces-483	620	12	10	10	NUM
alces-483	620	13	.8	.8	NUM
alces-483	620	14	b	b	PROPN
alces-483	620	15	er	er	INTJ
alces-483	620	16	ge	ge	PROPN
alces-483	620	17	ru	ru	PROPN
alces-483	621	1	d	d	PROPN
alces-483	621	2	a	a	DET
alces-483	621	3	nd	nd	NOUN
alces-483	621	4	m	m	NOUN
alces-483	621	5	an	an	DET
alces-483	621	6	ue	ue	ADJ
alces-483	621	7	l	l	NOUN
alces-483	621	8	(	(	PUNCT
alces-483	621	9	19	19	NUM
alces-483	621	10	68	68	NUM
alces-483	621	11	)	)	PUNCT
alces-483	621	12	,	,	PUNCT
alces-483	621	13	m	m	VERB
alces-483	621	14	er	er	INTJ
alces-483	621	15	ce	ce	PROPN
alces-483	621	16	r	r	NOUN
alces-483	621	17	a	a	DET
alces-483	621	18	nd	nd	NOUN
alces-483	621	19	m	m	NOUN
alces-483	621	20	an	an	DET
alces-483	621	21	ue	ue	ADJ
alces-483	621	22	l	l	NOUN
alces-483	622	1	(	(	PUNCT
alces-483	622	2	19	19	NUM
alces-483	622	3	74	74	NUM
alces-483	622	4	)	)	PUNCT
alces-483	622	5	,	,	PUNCT
alces-483	622	6	fr	fr	PROPN
alces-483	622	7	yx	yx	PROPN
alces-483	622	8	el	el	PROPN
alces-483	622	9	l	l	PROPN
alces-483	622	10	e	e	PROPN
alces-483	622	11	t	t	PROPN
alces-483	622	12	a	a	DET
alces-483	622	13	l.	l.	PROPN
alces-483	622	14	(	(	PUNCT
alces-483	622	15	19	19	NUM
alces-483	622	16	88	88	NUM
alces-483	622	17	)	)	PUNCT
alces-483	622	18	se	se	PROPN
alces-483	622	19	w	w	PROPN
alces-483	622	20	ar	ar	PROPN
alces-483	622	21	d	d	PROPN
alces-483	622	22	pe	pe	PROPN
alces-483	622	23	n	n	CCONJ
alces-483	622	24	,	,	PUNCT
alces-483	622	25	a	a	DET
alces-483	622	26	k	k	NOUN
alces-483	622	27	m	m	VERB
alces-483	622	28	oo	oo	INTJ
alces-483	622	29	se	se	ADV
alces-483	622	30	0.4	0.4	NUM
alces-483	622	31	0	0	NUM
alces-483	622	32	1	1	NUM
alces-483	622	33	0	0	NUM
alces-483	622	34	1	1	NUM
alces-483	622	35	2	2	NUM
alces-483	622	36	2.4	2.4	NUM
alces-483	622	37	g	g	NOUN
alces-483	622	38	as	as	ADP
alces-483	622	39	aw	aw	INTJ
alces-483	622	40	ay	ay	INTJ
alces-483	622	41	et	et	PROPN
alces-483	622	42	al	al	PROPN
alces-483	622	43	.	.	PUNCT
alces-483	623	1	(	(	PUNCT
alces-483	623	2	19	19	NUM
alces-483	623	3	92	92	NUM
alces-483	623	4	)	)	PUNCT
alces-483	623	5	r	r	NOUN
alces-483	623	6	i	i	PROPN
alces-483	623	7	d	d	PROPN
alces-483	623	8	in	in	ADP
alces-483	623	9	g	g	PROPN
alces-483	623	10	m	m	PROPN
alces-483	623	11	t	t	PROPN
alces-483	623	12	,	,	PUNCT
alces-483	623	13	m	m	VERB
alces-483	623	14	an	an	DET
alces-483	623	15	ito	ito	PROPN
alces-483	623	16	ba	ba	PROPN
alces-483	623	17	m	m	NOUN
alces-483	623	18	oo	oo	INTJ
alces-483	623	19	se	se	ADV
alces-483	623	20	0.8	0.8	NUM
alces-483	623	21	1	1	NUM
alces-483	623	22	1	1	NUM
alces-483	623	23	0	0	NUM
alces-483	623	24	0	0	NUM
alces-483	623	25	2	2	NUM
alces-483	623	26	4.8	4.8	NUM
alces-483	623	27	ca	ca	NOUN
alces-483	623	28	rb	rb	ADV
alces-483	624	1	yn	yn	PROPN
alces-483	625	1	(	(	PUNCT
alces-483	625	2	1	1	NUM
alces-483	625	3	98	98	NUM
alces-483	625	4	3	3	NUM
alces-483	625	5	)	)	PUNCT
alces-483	625	6	h	h	NOUN
alces-483	625	7	ec	ec	PROPN
alces-483	625	8	la	la	PROPN
alces-483	625	9	is	be	AUX
alces-483	625	10	la	la	PROPN
alces-483	625	11	nd	nd	PROPN
alces-483	625	12	,	,	PUNCT
alces-483	625	13	m	m	VERB
alces-483	625	14	an	an	DET
alces-483	625	15	ito	ito	PROPN
alces-483	625	16	ba	ba	PROPN
alces-483	625	17	m	m	NOUN
alces-483	625	18	oo	oo	INTJ
alces-483	625	19	se	se	ADJ
alces-483	625	20	1	1	NUM
alces-483	625	21	1	1	NUM
alces-483	625	22	1	1	NUM
alces-483	625	23	0	0	NUM
alces-483	625	24	0	0	NUM
alces-483	625	25	2	2	NUM
alces-483	625	26	6	6	NUM
alces-483	625	27	cr	cr	INTJ
alces-483	625	28	ic	ic	PROPN
alces-483	625	29	ht	ht	PROPN
alces-483	625	30	on	on	ADP
alces-483	625	31	(	(	PUNCT
alces-483	625	32	1	1	NUM
alces-483	625	33	97	97	NUM
alces-483	625	34	7	7	NUM
alces-483	625	35	)	)	PUNCT
alces-483	625	36	g	g	NOUN
alces-483	625	37	as	as	ADP
alces-483	625	38	pe	pe	X
alces-483	625	39	si	si	PROPN
alces-483	625	40	e	e	PROPN
alces-483	625	41	,	,	PUNCT
alces-483	625	42	q	q	PROPN
alces-483	625	43	ue	ue	INTJ
alces-483	625	44	be	be	AUX
alces-483	625	45	c	c	NOUN
alces-483	625	46	m	m	VERB
alces-483	625	47	oo	oo	INTJ
alces-483	625	48	se	se	ADP
alces-483	625	49	2	2	NUM
alces-483	625	50	0	0	SYM
alces-483	625	51	0	0	NUM
alces-483	625	52	1	1	NUM
alces-483	625	53	0	0	NUM
alces-483	625	54	1	1	NUM
alces-483	625	55	12	12	NUM
alces-483	625	56	cr	cr	NOUN
alces-483	625	57	êt	êt	ADP
alces-483	625	58	e	e	X
alces-483	625	59	(	(	PUNCT
alces-483	625	60	19	19	NUM
alces-483	625	61	89	89	NUM
alces-483	625	62	)	)	PUNCT
alces-483	625	63	is	be	AUX
alces-483	625	64	le	le	ADP
alces-483	625	65	r	r	NOUN
alces-483	625	66	oy	oy	PROPN
alces-483	625	67	al	al	PROPN
alces-483	625	68	e	e	PROPN
alces-483	625	69	,	,	PUNCT
alces-483	625	70	m	m	VERB
alces-483	625	71	i	i	PRON
alces-483	625	72	m	m	VERB
alces-483	625	73	oo	oo	INTJ
alces-483	625	74	se	se	ADJ
alces-483	625	75	2	2	NUM
alces-483	625	76	1	1	NUM
alces-483	625	77	0	0	NUM
alces-483	625	78	0	0	NUM
alces-483	625	79	0	0	NUM
alces-483	625	80	1	1	NUM
alces-483	625	81	12	12	NUM
alces-483	625	82	pe	pe	INTJ
alces-483	625	83	te	te	INTJ
alces-483	625	84	rs	rs	X
alces-483	625	85	on	on	ADP
alces-483	625	86	(	(	PUNCT
alces-483	625	87	1	1	NUM
alces-483	625	88	99	99	NUM
alces-483	625	89	9	9	NUM
alces-483	625	90	)	)	PUNCT
alces-483	625	91	a	a	DET
alces-483	625	92	ish	ish	NOUN
alces-483	625	93	ih	ih	PROPN
alces-483	625	94	ik	ik	PROPN
alces-483	625	95	,	,	PUNCT
alces-483	625	96	y	y	PROPN
alces-483	625	97	uk	uk	PROPN
alces-483	625	98	on	on	ADP
alces-483	625	99	m	m	PROPN
alces-483	625	100	oo	oo	INTJ
alces-483	625	101	se	se	PROPN
alces-483	625	102	0.1	0.1	NUM
alces-483	625	103	1	1	NUM
alces-483	625	104	1	1	NUM
alces-483	625	105	0	0	NUM
alces-483	625	106	1	1	NUM
alces-483	625	107	3	3	NUM
alces-483	625	108	0.6	0.6	NUM
alces-483	625	109	la	la	INTJ
alces-483	625	110	rs	rs	X
alces-483	625	111	en	en	X
alces-483	625	112	(	(	PUNCT
alces-483	625	113	1	1	NUM
alces-483	625	114	98	98	NUM
alces-483	625	115	2	2	NUM
alces-483	625	116	)	)	PUNCT
alces-483	626	1	k	k	NOUN
alces-483	626	2	lu	lu	VERB
alces-483	626	3	an	an	DET
alces-483	626	4	e	e	NOUN
alces-483	626	5	l	l	NOUN
alces-483	626	6	.	.	PUNCT
alces-483	627	1	,	,	PUNCT
alces-483	627	2	y	y	PROPN
alces-483	627	3	uk	uk	PROPN
alces-483	627	4	on	on	ADP
alces-483	627	5	m	m	PROPN
alces-483	627	6	oo	oo	INTJ
alces-483	627	7	se	se	PROPN
alces-483	627	8	0.1	0.1	NUM
alces-483	627	9	1	1	NUM
alces-483	627	10	1	1	NUM
alces-483	627	11	1	1	NUM
alces-483	627	12	1	1	NUM
alces-483	627	13	4	4	NUM
alces-483	627	14	0.6	0.6	NUM
alces-483	627	15	la	la	INTJ
alces-483	627	16	rs	rs	X
alces-483	627	17	en	en	X
alces-483	627	18	(	(	PUNCT
alces-483	627	19	1	1	NUM
alces-483	627	20	98	98	NUM
alces-483	627	21	2	2	NUM
alces-483	627	22	)	)	PUNCT
alces-483	627	23	d	d	NOUN
alces-483	627	24	en	en	PROPN
alces-483	627	25	al	al	PROPN
alces-483	627	26	i	i	PROPN
alces-483	627	27	,	,	PUNCT
alces-483	627	28	a	a	DET
alces-483	627	29	la	la	X
alces-483	627	30	sk	sk	VERB
alces-483	627	31	a	a	DET
alces-483	627	32	m	m	NOUN
alces-483	627	33	oo	oo	INTJ
alces-483	627	34	se	se	PROPN
alces-483	627	35	0.2	0.2	NUM
alces-483	627	36	1	1	NUM
alces-483	627	37	0	0	NUM
alces-483	627	38	0	0	NUM
alces-483	627	39	1	1	NUM
alces-483	627	40	2	2	NUM
alces-483	627	41	1.2	1.2	NUM
alces-483	627	42	si	si	NOUN
alces-483	627	43	ng	ng	PROPN
alces-483	628	1	er	er	INTJ
alces-483	629	1	an	an	PROPN
alces-483	629	2	d	d	X
alces-483	629	3	d	d	X
alces-483	629	4	al	al	PROPN
alces-483	629	5	le	le	PROPN
alces-483	629	6	-m	-m	PROPN
alces-483	629	7	ol	ol	PROPN
alces-483	629	8	le	le	X
alces-483	629	9	(	(	PUNCT
alces-483	629	10	1	1	NUM
alces-483	629	11	98	98	NUM
alces-483	629	12	5	5	NUM
alces-483	629	13	)	)	PUNCT
alces-483	629	14	n	n	PROPN
alces-483	629	15	el	el	PROPN
alces-483	629	16	ch	ch	PROPN
alces-483	629	17	in	in	ADP
alces-483	629	18	a	a	DET
alces-483	629	19	b	b	NOUN
alces-483	629	20	as	as	ADP
alces-483	629	21	in	in	ADV
alces-483	629	22	,	,	PUNCT
alces-483	629	23	a	a	DET
alces-483	629	24	k	k	NOUN
alces-483	629	25	m	m	VERB
alces-483	629	26	oo	oo	INTJ
alces-483	629	27	se	se	ADJ
alces-483	629	28	1	1	NUM
alces-483	629	29	1	1	NUM
alces-483	629	30	1	1	NUM
alces-483	629	31	0	0	NUM
alces-483	629	32	1	1	NUM
alces-483	629	33	3	3	NUM
alces-483	629	34	6	6	NUM
alces-483	629	35	g	g	NOUN
alces-483	629	36	as	as	ADP
alces-483	629	37	aw	aw	INTJ
alces-483	629	38	ay	ay	INTJ
alces-483	629	39	et	et	PROPN
alces-483	629	40	al	al	PROPN
alces-483	629	41	.	.	PUNCT
alces-483	630	1	(	(	PUNCT
alces-483	630	2	19	19	NUM
alces-483	630	3	92	92	NUM
alces-483	630	4	)	)	PUNCT
alces-483	631	1	k	k	PROPN
alces-483	631	2	en	en	INTJ
alces-483	631	3	ai	ai	VERB
alces-483	631	4	p	p	PROPN
alces-483	631	5	en	en	X
alces-483	631	6	,	,	PUNCT
alces-483	631	7	a	a	DET
alces-483	631	8	k	k	NOUN
alces-483	631	9	m	m	VERB
alces-483	631	10	oo	oo	INTJ
alces-483	631	11	se	se	PROPN
alces-483	631	12	1.1	1.1	NUM
alces-483	631	13	1	1	NUM
alces-483	631	14	1	1	NUM
alces-483	631	15	1	1	NUM
alces-483	631	16	1	1	NUM
alces-483	631	17	4	4	NUM
alces-483	631	18	6.6	6.6	NUM
alces-483	631	19	b	b	NOUN
alces-483	631	20	ai	ai	VERB
alces-483	631	21	le	le	X
alces-483	631	22	y	y	PROPN
alces-483	631	23	(	(	PUNCT
alces-483	631	24	19	19	NUM
alces-483	631	25	78	78	NUM
alces-483	631	26	)	)	PUNCT
alces-483	631	27	,	,	PUNCT
alces-483	631	28	pe	pe	AUX
alces-483	631	29	te	te	INTJ
alces-483	631	30	rs	rs	VERB
alces-483	631	31	on	on	ADP
alces-483	631	32	et	et	PROPN
alces-483	631	33	al	al	PROPN
alces-483	631	34	.	.	PUNCT
alces-483	632	1	(	(	PUNCT
alces-483	632	2	1	1	NUM
alces-483	632	3	98	98	NUM
alces-483	632	4	4	4	NUM
alces-483	632	5	)	)	PUNCT
alces-483	632	6	,	,	PUNCT
alces-483	632	7	s	s	PROPN
alces-483	632	8	ch	ch	PROPN
alces-483	632	9	w	w	PROPN
alces-483	632	10	ar	ar	PROPN
alces-483	632	11	tz	tz	PROPN
alces-483	632	12	an	an	PROPN
alces-483	632	13	d	d	X
alces-483	632	14	f	f	PROPN
alces-483	632	15	ra	ra	PROPN
alces-483	632	16	nz	nz	PROPN
alces-483	632	17	m	m	PROPN
alces-483	632	18	an	an	DET
alces-483	632	19	n	n	PROPN
alces-483	632	20	(	(	PUNCT
alces-483	632	21	1	1	NUM
alces-483	632	22	99	99	NUM
alces-483	632	23	1	1	NUM
alces-483	632	24	)	)	PUNCT
alces-483	633	1	s.	s.	PROPN
alces-483	634	1	q	q	PROPN
alces-483	634	2	ue	ue	PROPN
alces-483	634	3	be	be	AUX
alces-483	634	4	c	c	NOUN
alces-483	634	5	m	m	VERB
alces-483	634	6	oo	oo	INTJ
alces-483	634	7	se	se	X
alces-483	634	8	0.6	0.6	NUM
alces-483	634	9	1	1	NUM
alces-483	634	10	0	0	NUM
alces-483	634	11	1	1	NUM
alces-483	634	12	0	0	NUM
alces-483	634	13	2	2	NUM
alces-483	634	14	3.6	3.6	NUM
alces-483	634	15	po	po	NOUN
alces-483	634	16	tv	tv	NOUN
alces-483	634	17	in	in	ADP
alces-483	634	18	(	(	PUNCT
alces-483	634	19	1	1	NUM
alces-483	634	20	98	98	NUM
alces-483	634	21	8)	8)	NUM
alces-483	634	22	r	r	NOUN
alces-483	634	23	i	i	NOUN
alces-483	634	24	d	d	PROPN
alces-483	634	25	in	in	ADP
alces-483	634	26	g	g	PROPN
alces-483	634	27	m	m	PROPN
alces-483	634	28	t	t	PROPN
alces-483	634	29	,	,	PUNCT
alces-483	634	30	m	m	VERB
alces-483	634	31	an	an	DET
alces-483	634	32	ito	ito	PROPN
alces-483	634	33	ba	ba	PROPN
alces-483	634	34	m	m	NOUN
alces-483	634	35	oo	oo	INTJ
alces-483	634	36	se	se	ADV
alces-483	634	37	0.8	0.8	NUM
alces-483	634	38	1	1	NUM
alces-483	634	39	0	0	NUM
alces-483	634	40	1	1	NUM
alces-483	634	41	0	0	NUM
alces-483	634	42	2	2	NUM
alces-483	634	43	4.8	4.8	NUM
alces-483	634	44	ca	ca	NOUN
alces-483	634	45	rb	rb	ADV
alces-483	635	1	yn	yn	PROPN
alces-483	636	1	(	(	PUNCT
alces-483	636	2	1	1	NUM
alces-483	636	3	98	98	NUM
alces-483	636	4	3	3	NUM
alces-483	636	5	)	)	PUNCT
alces-483	637	1	s.	s.	PROPN
alces-483	637	2	c	c	PROPN
alces-483	637	3	en	en	PROPN
alces-483	637	4	tra	tra	PROPN
alces-483	637	5	l	l	NOUN
alces-483	637	6	o	o	X
alces-483	637	7	nt	not	PART
alces-483	637	8	ar	ar	PROPN
alces-483	637	9	io	io	PROPN
alces-483	637	10	m	m	PROPN
alces-483	637	11	oo	oo	INTJ
alces-483	637	12	se	se	PROPN
alces-483	637	13	0.3	0.3	NUM
alces-483	637	14	1	1	NUM
alces-483	637	15	0	0	NUM
alces-483	637	16	1	1	NUM
alces-483	637	17	0	0	NUM
alces-483	637	18	2	2	NUM
alces-483	637	19	1.8	1.8	NUM
alces-483	637	20	b	b	NUM
alces-483	637	21	er	er	INTJ
alces-483	637	22	ge	ge	PROPN
alces-483	637	23	ru	ru	PROPN
alces-483	637	24	d	d	PROPN
alces-483	637	25	e	e	PROPN
alces-483	637	26	t	t	PROPN
alces-483	637	27	a	a	DET
alces-483	637	28	l.	l.	PROPN
alces-483	637	29	(	(	PUNCT
alces-483	637	30	19	19	NUM
alces-483	637	31	83	83	NUM
alces-483	637	32	)	)	PUNCT
alces-483	637	33	sw	sw	PROPN
alces-483	637	34	q	q	PROPN
alces-483	637	35	ue	ue	PROPN
alces-483	637	36	be	be	AUX
alces-483	637	37	c	c	NOUN
alces-483	637	38	m	m	VERB
alces-483	637	39	oo	oo	INTJ
alces-483	637	40	se	se	PROPN
alces-483	637	41	0.3	0.3	NUM
alces-483	637	42	1	1	NUM
alces-483	637	43	0	0	NUM
alces-483	637	44	1	1	NUM
alces-483	637	45	0	0	NUM
alces-483	637	46	2	2	NUM
alces-483	637	47	1.8	1.8	NUM
alces-483	637	48	m	m	NOUN
alces-483	637	49	es	es	NOUN
alces-483	637	50	si	si	PROPN
alces-483	638	1	er	er	INTJ
alces-483	638	2	an	an	PRON
alces-483	638	3	d	d	X
alces-483	638	4	c	c	PROPN
alces-483	638	5	rê	rê	PROPN
alces-483	638	6	te	te	PROPN
alces-483	638	7	(	(	PUNCT
alces-483	638	8	1	1	NUM
alces-483	638	9	98	98	NUM
alces-483	638	10	5	5	NUM
alces-483	638	11	)	)	PUNCT
alces-483	638	12	a	a	DET
alces-483	638	13	pp	pp	X
alces-483	638	14	en	en	X
alces-483	638	15	di	di	X
alces-483	638	16	x	x	PROPN
alces-483	638	17	1	1	X
alces-483	638	18	.	.	PUNCT
alces-483	639	1	d	d	NOUN
alces-483	639	2	at	at	ADP
alces-483	639	3	a	a	DET
alces-483	639	4	us	us	PROPN
alces-483	639	5	ed	ed	NOUN
alces-483	639	6	in	in	ADP
alces-483	639	7	f	f	PROPN
alces-483	639	8	ig	ig	INTJ
alces-483	639	9	ur	ur	INTJ
alces-483	639	10	e	e	PROPN
alces-483	639	11	1	1	NUM
alces-483	639	12	.	.	PUNCT
alces-483	640	1	d	d	X
alces-483	640	2	en	en	ADP
alces-483	640	3	si	si	PROPN
alces-483	640	4	ty	ty	INTJ
alces-483	640	5	/	/	SYM
alces-483	640	6	bl	bl	PROPN
alces-483	640	7	ac	ac	PROPN
alces-483	641	1	k	k	PROPN
alces-483	641	2	br	br	INTJ
alces-483	641	3	ow	ow	INTJ
alces-483	641	4	n	n	PROPN
alces-483	641	5	n	n	PROPN
alces-483	642	1	o.	o.	NOUN
alces-483	643	1	d	d	X
alces-483	643	2	ee	ee	ADP
alces-483	643	3	r	r	NOUN
alces-483	643	4	-	-	PUNCT
alces-483	643	5	e	e	NOUN
alces-483	643	6	q.	q.	PROPN
alces-483	643	7	lo	lo	PROPN
alces-483	643	8	ca	can	AUX
alces-483	643	9	tio	tio	VERB
alces-483	643	10	n	n	INTJ
alces-483	643	11	sp	sp	ADP
alces-483	643	12	ec	ec	PROPN
alces-483	643	13	ie	ie	X
alces-483	643	14	s	s	PART
alces-483	643	15	km	km	NOUN
alces-483	643	16	2	2	NUM
alces-483	643	17	w	w	NOUN
alces-483	643	18	ol	ol	PROPN
alces-483	643	19	f	f	PROPN
alces-483	643	20	h	h	INTJ
alces-483	644	1	um	um	INTJ
alces-483	644	2	an	an	DET
alces-483	644	3	be	be	NOUN
alces-483	644	4	ar	ar	NOUN
alces-483	644	5	be	be	NOUN
alces-483	644	6	ar	ar	NOUN
alces-483	644	7	pr	pr	NOUN
alces-483	644	8	ed	ed	NOUN
alces-483	644	9	.	.	PUNCT
alces-483	645	1	de	de	PROPN
alces-483	645	2	ns	ns	PROPN
alces-483	645	3	.	.	PUNCT
alces-483	646	1	r	r	NOUN
alces-483	646	2	ef	ef	PROPN
alces-483	646	3	er	er	INTJ
alces-483	646	4	en	en	PROPN
alces-483	646	5	ce	ce	PROPN
alces-483	646	6	space	space	NOUN
alces-483	646	7	and	and	CCONJ
alces-483	646	8	time	time	NOUN
alces-483	646	9	in	in	ADP
alces-483	646	10	predation	predation	NOUN
alces-483	646	11	dynamics	dynamic	NOUN
alces-483	646	12	–	–	PUNCT
alces-483	646	13	peterson	peterson	PROPN
alces-483	646	14	et	et	PROPN
alces-483	646	15	al	al	PROPN
alces-483	646	16	.	.	PROPN
alces-483	647	1	alces	alce	NOUN
alces-483	647	2	vol	vol	NOUN
alces-483	647	3	.	.	PROPN
alces-483	647	4	39	39	NUM
alces-483	647	5	,	,	PUNCT
alces-483	647	6	2003	2003	NUM
alces-483	647	7	232	232	NUM
alces-483	648	1	n	n	NOUN
alces-483	648	2	.	.	PUNCT
alces-483	649	1	a	a	DET
alces-483	649	2	lb	lb	INTJ
alces-483	649	3	er	er	INTJ
alces-483	649	4	ta	ta	PART
alces-483	649	5	m	m	ADV
alces-483	649	6	oo	oo	INTJ
alces-483	649	7	se	se	ADV
alces-483	649	8	0.2	0.2	NUM
alces-483	649	9	1	1	NUM
alces-483	649	10	1	1	NUM
alces-483	649	11	1	1	NUM
alces-483	649	12	0	0	NUM
alces-483	649	13	3	3	NUM
alces-483	649	14	1.2	1.2	NUM
alces-483	649	15	o	o	NOUN
alces-483	649	16	os	os	NOUN
alces-483	649	17	en	en	INTJ
alces-483	649	18	br	br	X
alces-483	649	19	ug	ug	ADP
alces-483	649	20	an	an	DET
alces-483	649	21	d	d	X
alces-483	649	22	c	c	PROPN
alces-483	649	23	ar	ar	PROPN
alces-483	649	24	by	by	ADP
alces-483	649	25	n	n	PROPN
alces-483	649	26	(	(	PUNCT
alces-483	649	27	1	1	NUM
alces-483	649	28	98	98	NUM
alces-483	649	29	5	5	NUM
alces-483	649	30	)	)	PUNCT
alces-483	650	1	n	n	PRON
alces-483	650	2	e	e	NOUN
alces-483	650	3	a	a	PRON
alces-483	650	4	lb	lb	INTJ
alces-483	650	5	er	er	INTJ
alces-483	650	6	ta	ta	PART
alces-483	650	7	m	m	ADV
alces-483	650	8	oo	oo	INTJ
alces-483	650	9	se	se	ADV
alces-483	650	10	0.2	0.2	NUM
alces-483	650	11	1	1	NUM
alces-483	650	12	1	1	NUM
alces-483	650	13	1	1	NUM
alces-483	650	14	0	0	NUM
alces-483	650	15	3	3	NUM
alces-483	650	16	1.2	1.2	NUM
alces-483	650	17	fu	fu	NOUN
alces-483	650	18	lle	lle	PROPN
alces-483	650	19	r	r	PROPN
alces-483	650	20	a	a	DET
alces-483	650	21	nd	nd	NOUN
alces-483	650	22	k	k	PROPN
alces-483	650	23	ei	ei	X
alces-483	650	24	th	th	X
alces-483	650	25	(	(	PUNCT
alces-483	650	26	1	1	NUM
alces-483	650	27	98	98	NUM
alces-483	650	28	0	0	NUM
alces-483	650	29	)	)	PUNCT
alces-483	650	30	g	g	PROPN
alces-483	650	31	m	m	VERB
alces-483	650	32	u	u	NOUN
alces-483	650	33	20	20	NUM
alces-483	650	34	a	a	DET
alces-483	650	35	-a	-a	X
alces-483	650	36	k	k	NOUN
alces-483	650	37	m	m	VERB
alces-483	650	38	oo	oo	INTJ
alces-483	650	39	se	se	ADV
alces-483	650	40	0.2	0.2	NUM
alces-483	650	41	1	1	NUM
alces-483	650	42	1	1	NUM
alces-483	650	43	0	0	NUM
alces-483	650	44	1	1	NUM
alces-483	650	45	3	3	NUM
alces-483	650	46	1.2	1.2	NUM
alces-483	650	47	g	g	NOUN
alces-483	650	48	as	as	ADP
alces-483	650	49	aw	aw	INTJ
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alces-483	650	51	et	et	PROPN
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alces-483	651	1	(	(	PUNCT
alces-483	651	2	19	19	NUM
alces-483	651	3	83	83	NUM
alces-483	651	4	)	)	PUNCT
alces-483	652	1	d	d	NOUN
alces-483	652	2	en	en	PROPN
alces-483	652	3	al	al	PROPN
alces-483	652	4	i	i	PROPN
alces-483	652	5	,	,	PUNCT
alces-483	652	6	a	a	DET
alces-483	652	7	la	la	X
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alces-483	652	9	a	a	DET
alces-483	652	10	m	m	NOUN
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alces-483	652	13	0.2	0.2	NUM
alces-483	652	14	1	1	NUM
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alces-483	652	17	0	0	NUM
alces-483	652	18	1	1	NUM
alces-483	652	19	1.2	1.2	NUM
alces-483	652	20	h	h	NOUN
alces-483	652	21	ab	ab	PROPN
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alces-483	653	2	(	(	PUNCT
alces-483	653	3	1	1	NUM
alces-483	653	4	97	97	NUM
alces-483	653	5	7	7	NUM
alces-483	653	6	)	)	PUNCT
alces-483	653	7	n	n	PRON
alces-483	653	8	e	e	NOUN
alces-483	653	9	m	m	VERB
alces-483	653	10	n	n	PRON
alces-483	653	11	m	m	VERB
alces-483	653	12	oo	oo	INTJ
alces-483	653	13	se	se	X
alces-483	653	14	0.6	0.6	NUM
alces-483	653	15	1	1	NUM
alces-483	653	16	1	1	NUM
alces-483	653	17	1	1	NUM
alces-483	653	18	0	0	NUM
alces-483	653	19	3	3	NUM
alces-483	653	20	3.6	3.6	NUM
alces-483	653	21	m	m	NOUN
alces-483	653	22	ec	ec	PROPN
alces-483	653	23	h	h	NOUN
alces-483	653	24	a	a	DET
alces-483	653	25	nd	nd	NOUN
alces-483	653	26	f	f	X
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alces-483	653	30	(	(	PUNCT
alces-483	653	31	1	1	NUM
alces-483	653	32	97	97	NUM
alces-483	653	33	1	1	NUM
alces-483	653	34	)	)	PUNCT
alces-483	653	35	,	,	PUNCT
alces-483	653	36	p	p	X
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alces-483	653	38	k	k	PROPN
alces-483	653	39	e	e	PROPN
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alces-483	653	41	al	al	PROPN
alces-483	653	42	.	.	PUNCT
alces-483	654	1	(	(	PUNCT
alces-483	654	2	19	19	NUM
alces-483	654	3	76	76	NUM
alces-483	654	4	)	)	PUNCT
alces-483	654	5	,	,	PUNCT
alces-483	655	1	n	n	PROPN
alces-483	655	2	el	el	PROPN
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alces-483	656	2	n	n	CCONJ
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alces-483	656	4	d	d	NOUN
alces-483	656	5	m	m	NOUN
alces-483	656	6	ec	ec	PROPN
alces-483	656	7	h	h	NOUN
alces-483	656	8	(	(	PUNCT
alces-483	656	9	1	1	NUM
alces-483	656	10	98	98	NUM
alces-483	656	11	6	6	NUM
alces-483	656	12	)	)	PUNCT
alces-483	656	13	n	n	PRON
alces-483	656	14	e	e	NOUN
alces-483	656	15	m	m	VERB
alces-483	656	16	n	n	PRON
alces-483	656	17	m	m	VERB
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alces-483	656	20	0.7	0.7	NUM
alces-483	656	21	1	1	NUM
alces-483	656	22	1	1	NUM
alces-483	656	23	1	1	NUM
alces-483	656	24	0	0	NUM
alces-483	656	25	3	3	NUM
alces-483	656	26	4.2	4.2	NUM
alces-483	656	27	v	v	NOUN
alces-483	656	28	an	an	PRON
alces-483	656	29	b	b	PROPN
alces-483	656	30	al	al	PROPN
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alces-483	656	37	t	t	PROPN
alces-483	656	38	a	a	DET
alces-483	656	39	l.	l.	PROPN
alces-483	656	40	(	(	PUNCT
alces-483	656	41	19	19	NUM
alces-483	656	42	75	75	NUM
alces-483	656	43	)	)	PUNCT
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alces-483	656	49	o	o	X
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alces-483	656	56	0.2	0.2	NUM
alces-483	656	57	1	1	NUM
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alces-483	656	59	1	1	NUM
alces-483	656	60	0	0	NUM
alces-483	656	61	2	2	NUM
alces-483	656	62	1.2	1.2	NUM
alces-483	656	63	pi	pi	NOUN
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alces-483	656	67	e	e	PROPN
alces-483	656	68	t	t	PROPN
alces-483	656	69	a	a	DET
alces-483	656	70	l.	l.	PROPN
alces-483	656	71	(	(	PUNCT
alces-483	656	72	19	19	NUM
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alces-483	656	74	)	)	PUNCT
alces-483	657	1	e.	e.	PROPN
alces-483	657	2	c	c	PROPN
alces-483	657	3	en	en	PROPN
alces-483	657	4	tra	tra	PROPN
alces-483	657	5	l	l	NOUN
alces-483	657	6	o	o	X
alces-483	657	7	nt	not	PART
alces-483	657	8	ar	ar	PROPN
alces-483	657	9	io	io	PROPN
alces-483	657	10	w	w	PROPN
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alces-483	657	12	ee	ee	ADP
alces-483	657	13	r	r	NOUN
alces-483	657	14	3.1	3.1	NUM
alces-483	657	15	1	1	NUM
alces-483	657	16	0	0	NUM
alces-483	657	17	1	1	NUM
alces-483	657	18	0	0	NUM
alces-483	657	19	2	2	NUM
alces-483	657	20	3.1	3.1	NUM
alces-483	657	21	pi	pi	NOUN
alces-483	657	22	m	m	PROPN
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alces-483	657	24	tt	tt	PROPN
alces-483	657	25	e	e	PROPN
alces-483	657	26	t	t	PROPN
alces-483	657	27	a	a	DET
alces-483	657	28	l.	l.	PROPN
alces-483	657	29	(	(	PUNCT
alces-483	657	30	19	19	NUM
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alces-483	657	32	)	)	PUNCT
alces-483	658	1	s.	s.	PROPN
alces-483	659	1	q	q	PROPN
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alces-483	659	4	c	c	PROPN
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alces-483	659	6	td	td	NOUN
alces-483	659	7	ee	ee	PROPN
alces-483	659	8	r	r	NOUN
alces-483	659	9	3	3	NUM
alces-483	659	10	1	1	NUM
alces-483	659	11	0	0	NUM
alces-483	659	12	1	1	NUM
alces-483	659	13	0	0	NUM
alces-483	659	14	2	2	NUM
alces-483	659	15	3	3	NUM
alces-483	659	16	po	po	NOUN
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alces-483	659	18	in	in	ADP
alces-483	659	19	(	(	PUNCT
alces-483	659	20	1	1	NUM
alces-483	659	21	98	98	NUM
alces-483	659	22	8)	8)	NUM
alces-483	659	23	e.	e.	PROPN
alces-483	659	24	c	c	PROPN
alces-483	659	25	en	en	PROPN
alces-483	659	26	tra	tra	PROPN
alces-483	659	27	l	l	NOUN
alces-483	659	28	o	o	X
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alces-483	659	34	ee	ee	ADP
alces-483	659	35	r	r	NOUN
alces-483	659	36	5.8	5.8	NUM
alces-483	659	37	1	1	NUM
alces-483	659	38	0	0	NUM
alces-483	659	39	1	1	NUM
alces-483	659	40	0	0	NUM
alces-483	659	41	2	2	NUM
alces-483	659	42	5.8	5.8	NUM
alces-483	659	43	pi	pi	NOUN
alces-483	659	44	m	m	PROPN
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alces-483	660	1	tt	tt	PROPN
alces-483	660	2	e	e	PROPN
alces-483	660	3	t	t	PROPN
alces-483	660	4	a	a	DET
alces-483	660	5	l.	l.	PROPN
alces-483	660	6	(	(	PUNCT
alces-483	660	7	19	19	NUM
alces-483	660	8	69	69	NUM
alces-483	660	9	)	)	PUNCT
alces-483	661	1	n	n	CCONJ
alces-483	661	2	e	e	NOUN
alces-483	661	3	m	m	VERB
alces-483	661	4	n	n	PRON
alces-483	661	5	w	w	PROPN
alces-483	661	6	td	td	NOUN
alces-483	661	7	ee	ee	ADP
alces-483	661	8	r	r	NOUN
alces-483	661	9	5.1	5.1	NUM
alces-483	661	10	1	1	NUM
alces-483	661	11	1	1	NUM
alces-483	661	12	1	1	NUM
alces-483	661	13	0	0	NUM
alces-483	661	14	3	3	NUM
alces-483	661	15	5.1	5.1	NUM
alces-483	661	16	v	v	NOUN
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alces-483	661	18	b	b	PROPN
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alces-483	661	22	er	er	INTJ
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alces-483	661	25	e	e	PROPN
alces-483	661	26	t	t	PROPN
alces-483	661	27	a	a	DET
alces-483	661	28	l.	l.	PROPN
alces-483	661	29	(	(	PUNCT
alces-483	661	30	19	19	NUM
alces-483	661	31	75	75	NUM
alces-483	661	32	)	)	PUNCT
alces-483	662	1	n	n	PRON
alces-483	662	2	c	c	PROPN
alces-483	662	3	en	en	X
alces-483	662	4	tra	tra	PROPN
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alces-483	662	6	m	m	VERB
alces-483	662	7	n	n	PROPN
alces-483	662	8	w	w	PROPN
alces-483	662	9	td	td	NOUN
alces-483	662	10	ee	ee	ADP
alces-483	662	11	r	r	NOUN
alces-483	662	12	3.5	3.5	NUM
alces-483	662	13	1	1	NUM
alces-483	662	14	1	1	NUM
alces-483	662	15	1	1	NUM
alces-483	662	16	0	0	NUM
alces-483	662	17	3	3	NUM
alces-483	662	18	3.5	3.5	NUM
alces-483	662	19	st	st	PROPN
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alces-483	662	23	(	(	PUNCT
alces-483	662	24	1	1	NUM
alces-483	662	25	95	95	NUM
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alces-483	662	27	)	)	PUNCT
alces-483	663	1	n	n	DET
alces-483	663	2	c	c	PROPN
alces-483	663	3	en	en	X
alces-483	663	4	tra	tra	PROPN
alces-483	663	5	l	l	PROPN
alces-483	663	6	m	m	VERB
alces-483	663	7	n	n	PROPN
alces-483	663	8	w	w	PROPN
alces-483	663	9	td	td	NOUN
alces-483	663	10	ee	ee	ADP
alces-483	663	11	r	r	NOUN
alces-483	663	12	6	6	NUM
alces-483	663	13	1	1	NUM
alces-483	663	14	1	1	NUM
alces-483	663	15	1	1	NUM
alces-483	663	16	0	0	NUM
alces-483	663	17	3	3	NUM
alces-483	663	18	6	6	NUM
alces-483	663	19	b	b	NOUN
alces-483	663	20	er	er	INTJ
alces-483	663	21	g	g	PROPN
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alces-483	663	24	k	k	PROPN
alces-483	663	25	ue	ue	INTJ
alces-483	664	1	hn	hn	PROPN
alces-483	664	2	(	(	PUNCT
alces-483	664	3	1	1	NUM
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alces-483	664	6	)	)	PUNCT
alces-483	664	7	n	n	PRON
alces-483	664	8	c	c	PROPN
alces-483	664	9	en	en	X
alces-483	664	10	tra	tra	PROPN
alces-483	664	11	l	l	PROPN
alces-483	664	12	m	m	VERB
alces-483	664	13	n	n	PROPN
alces-483	664	14	w	w	PROPN
alces-483	664	15	td	td	NOUN
alces-483	664	16	ee	ee	ADP
alces-483	664	17	r	r	NOUN
alces-483	664	18	6.2	6.2	NUM
alces-483	664	19	1	1	NUM
alces-483	664	20	1	1	NUM
alces-483	664	21	1	1	NUM
alces-483	664	22	0	0	NUM
alces-483	664	23	3	3	NUM
alces-483	664	24	6.2	6.2	NUM
alces-483	664	25	fu	fu	NOUN
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alces-483	664	27	r	r	NOUN
alces-483	664	28	(	(	PUNCT
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alces-483	664	31	)	)	PUNCT
alces-483	664	32	a	a	PRON
alces-483	664	33	nt	not	PART
alces-483	665	1	ic	ic	PROPN
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alces-483	665	4	is	be	AUX
alces-483	665	5	la	la	ADP
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alces-483	665	7	,	,	PUNCT
alces-483	665	8	p	p	X
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alces-483	665	12	ee	ee	ADP
alces-483	665	13	r	r	NOUN
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alces-483	665	15	0	0	NUM
alces-483	665	16	1	1	NUM
alces-483	665	17	0	0	NUM
alces-483	665	18	0	0	NUM
alces-483	665	19	1	1	NUM
alces-483	665	20	15	15	NUM
alces-483	665	21	a	a	PRON
alces-483	665	22	.	.	PUNCT
alces-483	666	1	c	c	PROPN
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alces-483	667	1	(	(	PUNCT
alces-483	667	2	p	p	X
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alces-483	667	9	.	.	PUNCT
alces-483	667	10	)	)	PUNCT
alces-483	668	1	r	r	NOUN
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alces-483	668	8	,	,	PUNCT
alces-483	668	9	m	m	VERB
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alces-483	668	17	0.3	0.3	NUM
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alces-483	668	21	0	0	NUM
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alces-483	668	23	0.3	0.3	NUM
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alces-483	670	7	e	e	NOUN
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alces-483	670	9	n	n	PRON
alces-483	670	10	w	w	PROPN
alces-483	670	11	td	td	NOUN
alces-483	670	12	ee	ee	PROPN
alces-483	670	13	r	r	NOUN
alces-483	670	14	0.6	0.6	NUM
alces-483	670	15	1	1	NUM
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alces-483	670	17	1	1	NUM
alces-483	670	18	0	0	NUM
alces-483	670	19	3	3	NUM
alces-483	670	20	0.6	0.6	NUM
alces-483	670	21	m	m	NOUN
alces-483	670	22	ec	ec	PROPN
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alces-483	670	24	(	(	PUNCT
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alces-483	670	28	)	)	PUNCT
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alces-483	671	1	so	so	ADV
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alces-483	671	6	ec	ec	PROPN
alces-483	671	7	h	h	NOUN
alces-483	671	8	(	(	PUNCT
alces-483	671	9	1	1	NUM
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alces-483	671	12	)	)	PUNCT
alces-483	671	13	n	n	PRON
alces-483	671	14	e	e	NOUN
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alces-483	671	17	w	w	PROPN
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alces-483	671	19	ee	ee	ADP
alces-483	671	20	r	r	NOUN
alces-483	671	21	3.5	3.5	NUM
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alces-483	671	24	1	1	NUM
alces-483	671	25	0	0	NUM
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alces-483	675	8	)	)	PUNCT
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alces-483	678	20	0	0	NUM
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alces-483	679	4	1	1	NUM
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alces-483	680	25	2	2	NUM
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alces-483	683	1	w	w	INTJ
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alces-483	683	6	.	.	PUNCT
alces-483	684	1	(	(	PUNCT
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alces-483	684	4	)	)	PUNCT
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alces-483	685	20	.	.	PUNCT
alces-483	686	1	(	(	PUNCT
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alces-483	686	4	)	)	PUNCT
alces-483	686	5	...	...	PUNCT
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alces-483	689	12	.	.	PUNCT
alces-483	690	1	d	d	X
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alces-483	694	9	.	.	PUNCT
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