ANTICANCER COMPOUNDS FROM MEDICINAL PLANTS BIOLOGICA NYSSANA 2 (2)  December 2011: 00-00 Ljupković R.B. et al..  Removal Cu(II) ions from water... 21 Review Article Checklist of the genus Bracon (Hymenoptera: Braconidae) in Serbia Vladimir Žikić 1* , Saša S. Stanković 1 , Marijana Ilić 1 1 Faculty of Science and Mathematics, Department of Biology and Ecology, University of Niš, Višegradska 33, 18000 Niš, Serbia * E-mail: vzikic@yahoo.com Abstract: Žikić, V., Stanković, S., Ilić, M.: Checklist of the genus Bracon (Hymenoptera: Braconidae) in Serbia, Biologica Nyssana, 3 (1), September 2012: 21-29. This is the first checklist of the genus Bracon for the territory of Serbia. Here we present 80 species of this cosmopolitan genus of the subfamily Braconinae which is one of the cyclostome braconid groups. We connected the parasitoids with their hosts where the data about them were available. There are 290 host species in total belonging to 39 families of four insect orders: Diptera, Coleoptera, Hymenoptera and Lepidoptera. Key words: Cyclostome Braconidae, Bracon, Serbia, checklist Introduction Braconinae belongs to the cyclostome group of the family Braconidae. This subfamily assembles ectoparasitoid wasps which lay their eggs on the host body. According to their biology, ectoparasitic braconid wasps belong to the group of idiobionts; which means that they do not permit the host to continue its development after oviposition (M a t h e w s , 1974). Oviposition in this group is split in two acts: paralisation of the host, than laying eggs which is opposite in koinobions. As a cosmopolitan group of parasitoids they represent a large and powerful biological weapon (eg. B a i r d , 1958; L e w i s et al., 1990) against various pests from the following holometabolous insect orders: Diptera, Coleoptera, Hymenoptera and Lepidoptera. The species of the genus Bracon show different degree of the host specificity, ranging from a strict specialization to only one host species, whereas other can be polyphagous, when parasitism occurs in more than a hundred hosts in different types of habitats and geographical areas (S h a w & H u d d l e s t o n , 1991). Because of their important role in biological control, biology of some species is very well studied: for B. intercessor (van A c h t e r b e r g et al., 1990; G e o r g i e v et al., 2005), for B. brevicornis (T e m e r a k , 1982), or many publications for B. hebetor and its host Plodia interpunctella (Pyralidae) (R e i n e r t et al., 1971; T a y l o r , 1988a, b; Y u et al., 1999; M i l o n a s , 2005; D w e c k et al. 2010). According to the Fauna Europaea (v a n A c h t e r b e r g , 2012), about 250 species of the genus Bracon are recorded in Europe. The present publication stemmed from the need to make the list of all registered Bracon species on the territory of Serbia after many years of investigation and also to connect the parasitoids with their hosts. Investigation of the genus Bracon in Serbia in different aspects is presented in following publications by the authors B e l o k o b y s k i j & Ž i k i ć (2009); B r a j k o v i ć (1989); B r a j k o v i ć et al. (1991, 1994); S t a n k o v i ć et al. (2010) and Ž i k i ć et al. (2000, 2010, 2012). 3 (1) • September 2012: 21-29 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)… 22 Materials and methods The majority of analyzed specimens were taken from the collection of the Faculty of Biology, University of Belgrade and the Faculty of Sciences and Mathematics, University of Niš, Serbia. A part of specimens is deposited at the Zoological Institute RAS (St. Petersburg, Russia) and The Natural History Museum of Budapest, Hungary. The data about braconid hosts are compiled using the available literature; the list of references is bellow. The basic key for the identification of the species we used is one by Tobias et al. (1986), updated Fauna Europaea (http://faunaeur.org) and the database Taxapad (Y u et al. 2012). A lot of information about the hosts and the distribution of the investigated genus we collected from P a p p (1973, 1977, 2008) and B e y a r s l a n et al. (2005). Results We present 80 species of the genus Bracon registered on the territory of Serbia. The data regarding parasitoids are connected with the adequate data about their hosts. The species of Bracon are listed alphabetically according to their subgenera. For 25 braconid species there were no host records for Serbia nor for other parts of Europe. In addition, we give the geographical distribution of all registered parasitoids in continental part of Europe and Russia. Bracon (Bracon) alutaceus Szepligeti 1901 – no host data. Distribution: Austria, Great Britain, Bulgaria, Cyprus, Czech Republic, Denmark, Germany, Hungary, Lithuania, Poland, Russia (Northwestern), Slovakia, Spain, Sweden, Ukraine, Serbia. Bracon (Bracon) crassiceps Thomson 1892 – no host data. Distribution: Austria, Finland, Germany, Hungary, Russia (Northwestern), Sweden, Switzerland, Serbia. Bracon (Bracon) crassungula Thomson 1892 – no host data. Distribution: Austria, Hungary, Italy, Sweden, Switzerland, Serbia. Bracon (Bracon) filicornis Thomson 1892 – no host data. Distribution: Austria, Sweden, Serbia. Bracon (Bracon) flagellaris Thomson 1892 – no host data. Distribution: Croatia, Germany, Hungary, Romania, Russia (South), Slovenia, Sweden, Switzerland, Ukraine, Serbia. Bracon (Bracon) fulvipes Nees 1834 on Apion minimum, Cistapion cyanescens (Apionidae); Curculio villosus, Mononychus punctumalbum (Curculionidae); Tetramesa brevicornis T. hyalipennis (Eurytomidae); Coleophora coronillae (Coleophoridae); Paranthrene tabaniformis (Sesiidae). Distribution: Austria, Belgium, Great Britain, Bulgaria, Croatia, Turkey, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Poland, Russia (Central and South), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Bracon) grandiceps Thomson 1892 on Bembecia scopigera (Sesiidae). Distribution: Austria, Croatia, Turkey, Finland, France, Germany, Greece, Hungary, Italy, Macedonia, Poland, Russia (South), Slovenia, Sweden, Serbia. Bracon (Bracon) intercessor Nees 1834 on Apion opeticum (Apionidae); Agapanthia villosoviridescens (Cerambycidae); Anthonomus pedicularius, A. pomorum, A. sorbi, Lixus brevirostris, L. incanescens, L. junci, L. scabricollis, Microlarinus lareynii, M. lypriformis (Curculionidae); Rhynchites bacchus (Rhynchitidae); Tetramesa hyalipennis, T. rossica (Eurytomidae); Haplochrois theae (Agonoxenidae); Augasma atraphaxidellum (Coleophoridae); Synanthedon culiciformis, Paranthrene tabaniformis (Sesiidae); Sparganothis pilleriana (Tortricidae). Distribution: Albania, Austria, Great Britain, Bulgaria, Croatia, Cyprus, Turkey, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Macedonia, Norway, Poland, Romania, Russia (Central, East, Northwestern and South), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Ukraine, Serbia. Bracon (Bracon) laetus Wesmael 1838 on Anthonomus pomorum, Magdalis nitida (Curculionidae); Amblypalpis tamaricella (Gelechiidae); Argyresthia chrysidella, Prays citri; Prays oleae (Yponomeutidae). Distribution: Belgium, Great Britain, Bulgaria, Croatia, Cyprus, Czech Republic, France, Germany, Greece, Hungary, Spain, Serbia. Bracon (Bracon) leptus Marshall 1897 on Metzneria lapella (Gelechiidae). Distribution: Austria, Great Britain, Bulgaria, Croatia, Cyprus, Germany, Greece, Hungary, Lithuania, Macedonia, Romania, Russia (Central and South), Slovakia, Slovenia, Spain, Switzerland, Netherlands, Serbia. Bracon (Bracon) longicollis Wesmael 1838 on Chlorops pumilionis (Chloropidae). Distribution: Austria, Belgium, Great Britain, Bulgaria, Croatia, Cyprus, Turkey, Finland, France, Germany, Greece, Hungary, Italy, Lithuania, Macedonia, Norway, Poland, Russia (Central, Northwestern and South), BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)… 23 Slovenia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Bracon) longulus Thomson 1892 on Oscinella frit (Chloropidae). Distribution: Austria, Bulgaria, Croatia, Cyprus, Czech Republic, Denmark, Finland, France, Germany, Greece, Hungary, Italy, Macedonia, Norway, Slovenia, Spain, Sweden, Netherlands, Serbia. Bracon (Bracon) luteator Spinola 1808 on Urophora solstitialis (Tephritidae); Metzneria aestivella, M. lappella (Gelechiidae). Distribution: Austria, Great Britain, Bulgaria, Croatia, Cyprus, Turkey, France, Greece, Hungary, Italy, Latvia, Macedonia, Russia (Central, Northwestern and South), Slovakia, Spain, Switzerland, Ukraine, Serbia. Bracon (Bracon) mariae Dalla Torre 1898 on Chaetorellia jaceae (Tephritidae). Distribution: Bulgaria, Croatia, Germany, Hungary, Italy, Russia (South), Slovakia, Spain, Sweden, Switzerland, Ukraine, Serbia. Bracon (Bracon) mediostriatus Papp 1968 – no host data. Distribution: Hungary, Serbia. Bracon (Bracon) nigratus Wesmael 1838 on Chaetostomella cylindrica (Tephritidae); Coleophora millefolii (Coleophoridae); Grapholita compositella (Tortricidae); Zygaena loti (Zygaenidae). Distribution: Austria, Belgium, Great Britain, Croatia, Czech Republic, France, Germany, Hungary, Ireland, Italy, Lithuania, Romania, Russia (Central and North), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Bracon) pallicarpus Thomson 1892 – no host data. Distribution: Great Britain, Bulgaria, Finland, Germany, Hungary, Italy, Spain, Sweden, Serbia. Bracon (Bracon) pectoralis Wesmael 1838 on Chrysobothis affisinis, Sphenoptera gossypii, S. lobicollis, S. montana (Buprestidae); Plagionotus arcuatus, Plagionotus bobelayei (Cerambycidae). Distribution: Austria, Belgium, Great Britain, Bulgaria, Croatia, Cyprus, Turkey, France, Germany, Greece, Hungary, Italy, Macedonia, Romania, Russia (Central and South), Slovakia, Spain, Switzerland, Serbia. Bracon (Bracon) scabriusculus Dalla Torre 1898 – no host data. Distribution: Austria, Germany, Hungary, Italy, Romania, Russia (Central), Spain, Serbia. Bracon (Bracon) scutellaris Wesmael 1838 on Anobium abietis (Anobiidae); Euura mucronata, Pontania pedunculi, P. viminallis (Tenthredinidae). Distribution: Belgium, Great Britain, Finland, France, Hungary, Ireland, Italy, Poland, Sweden, Netherlands, Serbia. Bracon (Bracon) speerschneideri Schmiedeknecht 1897 on Argyresthia thuiella, Ocnerostoma sp. (Yponomeutidae). Distribution: Austria, Great Britain, France, Germany, Norway, Sweden, Switzerland, Serbia. Bracon (Bracon) subglaber Szepligeti 1901 on Noeeta pupillata, Tephritis leontodontis, Terellia serratulae, Urophora cardui, U. quadrifasciata, U solstitialis (Tephritidae); Metzneria lappella (Gelechidae); Epichnopterix sieboldi (Psychidae); Paranthrene tabaniformis, Pennisetia hylaeiformis (Sesiidae); Aethes williana (Tortricidae). Distribution: Croatia; Germany; Hungary; Italy, Kazakhstan, Moldova, Russia (Russia-Krasnodar, Yaroslavl Oblast), Slovenia, Switzerland, Ukraine, Serbia. Bracon (Bracon) trucidator Marshall 1888 on Metzneria lappella (Gelechiidae). Distribution: Albania, Austria, Belgium, Great Britain, Bulgaria, Croatia, Denmark, Turkey, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Poland, Romania, Russia (Central, Northwestern and South), Slovenia, Spain, Switzerland, Netherlands, Serbia. Bracon (Bracon) variegator Spinola 1808 on Ernobius abietis (Anobiidae); Coleophora iutipennella (Coleophoridae); Agonopterix propinquella, (Depressariidae); Anarsia lineatella, Recurvaria nanella, Teleiodes luculella (Gelechiidae); Phyllonorycter klemannella, Ph. mespillela (Gracillariidae); Endrosis sarcitrella (Oecophoridae); Acrobasis obsutella (Pyralidae); Archips rosana, Cydia strobilella, Tortrix viridana (Tortricidae); Yponomeuta padella (Yponomeutidae). Distribution: Albania, Austria, Croatia, Cyprus, France, Hungary, Italy, Macedonia, Russia (South), Sardinia, Spain, Switzerland, Ukraine, Serbia. Bracon (Cyanopterobracon) sabulosus Szepligeti 1896 on Tephritidae. Distribution: Bulgaria, Croatia, Cyprus, Greece, Hungary, Macedonia, Romania, Russia (South), Serbia. Bracon (Cyanopterobracon) urinator (Fabricius 1798) on Larinus saussureae, L. sturnus, Rhinocyllus conicus (Curculilonidae); Tephritis pulchra (Tephritidae). Distribution: Albania, Austria, Belgium, Bosnia and Herzegovina, Great Britain, Bulgaria, Croatia, Cyprus, Turkey, France, Germany, Greece, Hungary, Italy, Macedonia, Portugal, Romania, Russia (Central, East, BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)… 24 Northwestern and South), Slovakia, Spain, Sweden, Switzerland, Netherlands, Ukraine, Serbia. Bracon (Cyanopterobracon) uromelas Costa 1888 – no host data. Distribution: Albania, Austria, Croatia, Cyprus, France, Hungary, Italy, Macedonia, Russia (South), Sardinia, Spain, Switzerland, Ukraina, Serbia. Bracon (Glabrobracon) abbreviator Nees 1834 on Anthonomus pomorum (Curculionidae); Lipara lucens (Chloropidae); Cephus pygmaeus (Cephidae); Coleophora trogldytella (Coleophoridae); Mesapamea secalis, Oria musculosa, Sesamia nonagrioides (Noctuidae); Pennisetia hylaeiformis (Sesiidae); Cochylis posterna, Pandemis heparana, Rhyacionia buoliana (Tortricidae). Distribution: Austria, Great Britain, Croatia, Cyprus, Czech Republic, Finland, Germany, Hungary, Poland, Romania, Slovakia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) arcuatus Thomson 1892 – no host data. Distribution: Austria, Great Britain, Bulgaria, Croatia, Cyprus, Czech Republic, Germany, Greece, Hungary, Slovenia, Sweden, Netherlands, Serbia. Bracon (Glabrobracon) atrator Nees 1834 on Omphalapion buddebergi (Apionidae); Gymnetron villosulum (Curculionidae); Amauromyza flavifrons (Agromyzidae); Tephritis conura, T. separata (Tephritidae); Coleophora coronillae (Coleophoridae). Distribution: Austria, Belgium, Great Britain, Bulgaria, Croatia, Cyprus, Turkey, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Macedonia, Poland, Russia (Central, Northwestern and South), Slovakia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) caudatus Ratzeburg 1848 on Hylesinus fraxini (Curculionidae). Distribution: Austria, Great Britain, Bulgaria, Czech Republic, Germany, Greece, Hungary, Poland, Serbia. Bracon (Glabrobracon) caudiger Nees 1834 on Coleophora obscenella (Coleophoridae); Aphelia amplana, Cydia splendana, C. strobilella (Tortricidae). Distribution: Austria, Belgium, Czech Republic, Turkey, France, Germany, Hungary, Italy, Poland, Romania, Russia (Central, North, Northwestern and South), Sweden, The Netherland, Serbia. Bracon (Glabrobracon) colpophorus Wesmael 1838 on Coleophora sp. (Coleophoridae). Distribution: Austria, Belgium, Great Britain, Czech Republic, France, Germany, Hungary, Italy, Moldova, Poland, Romania, Russia (Northwestern), Netherlands, Ukraine, Serbia. Bracon (Glabrobracon) coniferarum Fahringer 1928 on Millieria dolosalis (Choreutidae); Coleophora gallipennella, C. serratella (Coleophoridae). Distribution: Great Britain, Bulgaria, Czech Republic, Germany, Italy, Poland, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) delibator Haliday 1833 on Gymnetron campanulae, Hylobius piceus (Curculionidae); Olibrus aeneus (Phalacridae); Urophora cuspidata (Tephritidae); Cydia strobilella (Tortricidae). Distribution: Austria, Belgium, Bosnia and Herzegovina, Great Britain, Bulgaria, Croatia, Cyprus, Czech Republic, Turkey, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Macedonia, Poland, Russia (Central and South), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Ukraine, Serbia. Bracon (Glabrobracon) delusor Spinola 1808 – no host data. Distribution: Austria, Croatia, France, Germany, Italy, Latvia, Poland, Russia (Northwestern), Serbia. Bracon (Glabrobracon) dichromus Wesmael 1838 on Gymnetron asellus (Curculionidae); Platyedra subcinerea (Gelechidae); Grapholita funebrana (Tortricidae). Distribution: Austria, Belgium, Bulgaria, Croatia, Cyprus, France, Germany, Hungary, Italy, Poland, Romania, Slovakia, Spain, Serbia. Bracon (Glabrobracon) discoideus Wesmael 1838 on Saperda populnea (Cerambycidae); Anthonomus pomorum, A. spilotus, Ceutorhynchus assimilis, C. quadridens, Curculio crux, C. nucum, C. pyrrhoceras, C. salicivorus, Pissodes notatus (Curculionidae); Byctiscus betulae, B. populi, Rhynchites pauxillus (Rhynchitidae); Chlorops pumilionis (Chloropidae); Biorhiza pallida (Cynipidae); Euura mucronata, Pontania bridgmanii, P. collactanea, P. foetidae, P. pedunculi, P. proxima, P. vesicator, P. viminalis (Tenthredinidae); Anticlea badiata (Geometridae); Adoxophyes orana, Rhyacionia buoliana (Tortricidae). Distribution: Austria, Belgium, Great Britain, Bulgaria, Czech Republic, France, Germany, Hungary, Ireland, Italy, Poland, Romania, Russia (Northwestern), Slovakia, Slovenia, Sweden, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) fumatus Szepligeti 1901 – no host data. Distribution: Austria, Croatia, Cyprus, Turkey, France, Greece, Hungary, Russia (South), Spain, Switzerland, Serbia. Bracon (Glabrobracon) fuscipennis Wesmael 1838 on Gastrophysa polygoni, G. viridula, Plagiodera versicolora (Chrysomelidae). Distribution: Austria, http://www.faunaeur.org/full_results.php?id=11049 http://www.faunaeur.org/full_results.php?id=11175 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)… 25 Belgium, Croatia, France, Germany, Hungary, Italy, Poland, Slovenia, Serbia. Bracon (Glabrobracon) hyalinipennis Szepligeti 1901 – no host data. Distribution: Hungary, Serbia. Bracon (Glabrobracon) immutator Nees 1834 on Anthonomus rubi, Ceutorhynchus pleurostigma, Cryptorhynchus lapathi, Curculio villosus, Pissodes notatus (Curculionidae); Urophora stylata (Tephritidae); Biorhiza pallida, (Cynipidae). Distribution: Austria, Belgium, Bosnia and Herzegovina, Great Britain, Croatia, Cyprus, Czech Republic, Finland, France, Germany, Hungary, Italy, Macedonia, Poland, Romania, Russia (Central, North, and Northwestern), Slovenia, Sweden, Switzerland, Netherlands, Ukraine, Serbia. Bracon (Glabrobracon) larvicida Wesmael 1838 – no host data. Distribution: Austria, Belgium, Great Britain, Bulgaria, Croatia, Turkey, France, Germany, Greece, Hungary, Italy, Poland, Romania, Russia (Central, Northwestern and South), Slovakia, Slovenia, Serbia. Bracon (Glabrobracon) marshalli Szepligeti 1901 on Trachys sp. (Buprestidae); Myopites inulaedyssentericae (Tephritidae); Cydia nigricana (Tortricidae). Distribution: Great Britain, Bulgaria, Croatia, Cyprus, Denmark, Finland, Germany, Greece, Hungary, Italy, Norway, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden, Switzerland, Serbia. Bracon (Glabrobracon) minutator (Fabricius 1798) on Anthonomus pomorum (Curculionidae); Lipara lucens (Chloropidae); Trypeta jaceae (Tephritidae); Cephus pygmaeus (Cephidae); Caloptilia syringella (Gracillariidae); Mesapamea secalis, Oria musculosa, Sesamia nonagrioides (Noctuidae); Pennisetia hylaeiformis (Sesiidae); Cochylis posterana, Leguminivora glycinivorella, Rhyacionia buoliana (Tortricidae). Distribution: Albania, Austria, Belgium, Great Britain, Bulgaria, Cyprus, Czech Republic, Denmark, Turkey, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Macedonia, Norway, Poland, Russia (Central and South), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) obscurator Nees 1811 on Trachys pumilus, T. troglodytes (Buprestidae); Hylesinus fraxini, Ips typographus, Orthotomicus suturalis, Trypodendron domesticum, T. signatum (Curculionidae); Coleophora caespititiella (Colephoridae); Epermenia fulviguttella (Epermeniidae); Homoeosoma sinuella (Pyralidae). Distribution: Austria, Belgium, Bosnia and Herzegovina, Great Britain, Bulgaria, Croatia, Cyprus, Czech Republic, Denmark, Turkey, Finland, France, Germany, Greece, Hungary, Italy, Macedonia, Poland, Romania, Russia (Central, Northwestern and South), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) osculator Nees 1811 on Millieria dolosalis (Choreutidae); Coleophora albicostella, C. alticolella, C. caespititiella, C. colutella, C. discordella, C. flaviella, C. frischella, C. gallipennella, C. lutipennella, C. milvipennis, C. obscenella, C. ochrea, C. saponariella, C. serenella, C. serratella, C. solitariella, C. spinella, Co. spiraeella, C. therinella, C. trifariella, C. vibicella, C. vicinella, C. vulpecula, Coleophora sp.1, Coleophora sp.2 (Coleophoridae); Cosmopterix eximia (Cosmopterigidae); Elachista contaminatella, Elachista sp. (Elachistidae); Phyllonorycter sp. (Gracillariidae); Mompha conturbatella (Momphidae); Ectoedemia agrimoniae (Nepticulidae); Retinia perangustana (Tortricidae). Distribution: Austria, Belgium, Bosnia and Herzegovina, Great Britain, Bulgaria, Croatia, Cyprus, Czech Republic, Denmark, Turkey, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Macedonia, Poland, Romania, Russia (Central, East, Northwestern and South), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) peroculatus Wesmael 1838 on Cnephasia chrysantheana (Tortricidae). Distribution: Belgium, Finland, France, Germany, Hungary, Italy, Russia (Central), Serbia. Bracon (Glabrobracon) picticornis Wesmael 1838 on Clytus sp., Plagionotus arcuatus (Cerambicidae); Nematus bipartitus, N. salicis, Pontania pedunculi, P. proxima, P. vesicator, P. viminallis (Tenthredinidae); Cochylis pallidana (Tortricidae). Distribution: Austria, Belgium, Great Britain, Bulgaria, Turkey, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Lithuania, Poland, Russia (Central, Northwestern and South), Slovakia, Spain, Sweden, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) pineti Thomson 1892 on Ernobius abietis, E. longicornis (Anobiidae); Cydia strobilella (Tortricidae). Distribution: Austria, Bulgaria, Croatia, Czech Republic, Finland, Germany, Hungary, Lithuania, Moldova, Poland, Russia (Central, North, Northwestern and South), Slovakia, Sweden, Serbia. Bracon (Glabrobracon) ratzeburgii Dalla Torre 1898 on Ceutorhynchus maculaalba, Hylesinus fraxini, Scolytus rugulosus (Curculionidae). Distribution: Great Britain, Germany, Italy, Serbia. BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)… 26 Bracon (Glabrobracon) terebella Wesmael 1838 on Miarus campanulae (Curculionidae); Cephus pygmaeus, Trachelus tabidus (Cephidae). Distribution: Austria, Belgium, Great Britain, Denmark, Finland, Germany, Hungary, Italy, Lithuania, Poland, Romania, Russia (Central,Northwestern and South), Slovakia, Sweden, Switzerland, Serbia. Bracon (Glabrobracon) titubans Wesmael 1838 – no host data. Distribution: Austria, Belgium, Bulgaria, Finland, France, Germany, Hungary, Italy, Poland, Russia (Northwestern), Slovakia, Sweden, Switzerland, Netherlands, Serbia. Bracon (Glabrobracon) tornator Marshall 1885 – no host data. Distribution: Austria, Great Britain, Bulgaria, Croatia, Finland, Germany, Greece, Hungary, Italy, Sweden, Switzerland, Serbia. Bracon (Glabrobracon) variator Nees 1811 on Bruchidius lividimanus, Bruchus atomarius, B. laticollis, B. lentis, B. viciae (Bruchidae); Anthonomus pomorum, Baris chlorizans, B. cuprirostris, B. laticollis, Ceutorhynchus punctiger, Gymnetron asellus, G. tetrum, Larinus jaceae, L. turbinatus, Magdalis rufa, Miarus campanulae, Pissodes validirostris, Sibinia viscariae, Sitona longulus (Curculionidae); Chaetostomella cylindrica, Noeeta pupillata, Sphenella marginata, Tephritis leontodontis (Tephritidae); Hoplocampa brevis, H. flava (Tenthrenidae); Colephora coronillae, C. medelichenis (Coleophoridae); Mesophleps corsicella, Pexicopia malvella, Platyedra subcinerea, (Gelechiidae); Phyllonorycter mespilella (Gracillariidae); Hadena bicruris (Noctuidae); Dioryctria abietella, Etiella zinckenella, Myelois circumvoluta (Pyralidae); Synathedon andrenaeformis (Sesiidae); Barbara herrichiana, Cydia strobilella, Eucosma cana, Grapholita funebrana, G. lunulana, Pandemis cerasana, Retinia resinella (Tortricidae). Distribution: Austria, Belgium, Bosnia and Herzegovina, Great Britain, Bulgaria, Canary Is., Croatia, Cyprus, Czech Republic, Turkey, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Macedonia, Moldova, Poland, Romania, Russia (Central, East, North, Northwestern and South), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Ukraine, Serbia. Bracon (Habrobracon) brevicornis Wesmael 1838 on Sesamia cretica (Noctuidae). Distribution: Belgium, Great Britain, Bulgaria, Czech Republic, France, Germany, Hungary, Italy, Poland, Portugal, Romania, Russia (Central and South), Slovakia, Spain, Switzerland, Netherlands, Serbia. Bracon (Habrobracon) hebetor Say 1836 on Ostrinia nubilalis (Crambidae); Pexicopia malvella (Gelechiidae); Helicoverpa armigera, Heliothis peltigera, Spodoptera littoralis (Noctuidae); Ephestia elutella, E. kuehniella, Etiella zinckenella, Galleria mellonella, Plodia interpunctella (Pyralidae); Adoxophyes orana, Lobesia botrana (Tortricidae). Distribution: Austria, Belgium, Great Britain, Bulgaria, Croatia, Cyprus, Czech Republic, Turkey, Finland, France, Germany, Greece, Hungary, Italy, Lithuania, Macedonia, Poland, Russia (Central and South), Slovakia, Slovenia, Spain, Netherlands, Serbia. Bracon (Lucobracon) apricus Schmiedeknecht 1897 – no host data. Distribution: Germany, Greece, Serbia. Bracon (Lucobracon) exarator Marshall 1885 – no host data. Distribution: Great Britain, Germany, Serbia. Bracon (Lucobracon) femoralis Brullé, – no host data. Distribution: Cyprus, Greece, Hungary, Spain, Switzerland, Serbia. Bracon (Lucobracon) fortipes Wesmael 1838 on Plagionotus floralis (Cerambycidae); Lixus anguinus (Curculionidae); Gortyna xanthenes (Noctuidae); Bembecia scopigera (Sesiidae). Distribution: Belgium, Bulgaria, Croatia, Cyprus, France, Germany, Greece, Hungary, Italy, Poland, Slovenia, Switzerland, Serbia. Bracon (Lucobracon) fumarius Szepligeti 1901 – no host data. Distribution: Austria, Croatia, Cyprus, Hungary, Romania, Serbia. Bracon (Lucobracon) hungaricus (Szepligeti 1896) on Clytus arietis (Cerambycidae). Distribution: Bulgaria, Croatia, Turkey, Hungary, Russia (Central and South), Slovakia, Ukraine, Serbia. Bracon (Lucobracon) hylobii Ratzeburg 1848 on Hylobius abietis, H. piceus, Pissodes harcyniae, P. notatus, P. piceae, P. pini, Polygraphus polygraphus (Curculionidae). Distribution: Austria, Great Britain, Bulgaria, Czech Republic, Denmark, Germany, Hungary, Italy, Lithuania, Poland, Russia (Central and Northwestern), Sweden, Switzerland, Netherlands, Serbia. Bracon (Lucobracon) megapterus Wesmael 1838 on Glaphyra umbellatarum (Cerambycidae); Chaetorellia jaceae (Tephritidae). Distribution: Belgium, Bulgaria, Czech Republic, Finland, France, Germany, Sweden, Serbia. Bracon (Lucobracon) nigriventris Wesmael 1838 on Plagionotus floralis (Cerambycidae); Hylobius piceus, Pissodes pini (Curculionidae). Distribution: BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)… 27 Albania, Austria, Belgium, Bulgaria, Croatia, Cyprus, Turkey, Germany, Hungary, Italy, Poland, Romania, Russia (South), Slovakia, Switzerland, Ukraine, Serbia. Bracon (Lucobracon) ochraceus Szepligeti 1896 – no host data. Distribution: Croatia, Hungary, Russia (South), Spain, Switzerland, Serbia. Bracon (Lucobracon) planiceps Telenga 1936 – no host data. Distribution: Russia (North), Serbia. Bracon (Lucobracon) romani Fahringer 1927 on Chamaesphecia bibioniformis, Paranthrene tabaniformis, Pennisetia hylaeiformis, Sesia apiformis, S. bembeciformis, Synanthedon culiciformis, S. spheciformis (Sesiidae). Distribution: Greece, Hungary, Slovakia, Slovenia, Sweden, Netherlands, Serbia. Bracon (Lucobracon) thuringiacus Schmiedeknecht 1896 – no host data. Distribution: Great Britain, Germany, Greece, Italy, Serbia. Bracon (Lucobracon) triangularis Nees 1834 on Synanthedon tipuliformis (Sesiidae). Distribution: Great Britain, Bulgaria, Czech Republic, Finland, Germany, Hungary, Ireland, Italy, Poland, Slovakia, Switzerland, Netherlands, Serbia. Bracon (Orthobracon) epitriptus Marshall 1885 on Cryptorhynchus lapathi, Hylobius transversovittatus, Pissodes spp. (Curculionidae); Agromyza flaviceps (Agromyzidae); Iteomyia capreae (Cecidomyiidae); Fenusa pumila (Tenthredinidae). Distribution: Austria, Great Britain, Bulgaria, Germany, Greece, Hungary, Italy, Lithuania, Poland, Russia (Central, Northwestern and South), Slovakia, Slovenia, Switzerland, Netherlands, Serbia. Bracon (Orthobracon) erraticus Wesmael 1838 on Gastrophysa viridula, Phaedon cochleariae, Phyllotreta nemorum (Chrysomelidae); Chaetostomella cylindrca, Urophora eriolepidis, Terellia serratulae (Tephritidae); Coleophora laricella, C. lutipennella (Coleophoridae); Metzneria lappella (Gelechiidae); Pennisetia hylaeiformis (Sesiidae); Acleris rhombana (Tortricidae). Distribution: Austria, Belgium, Bosnia and Herzegovina, Great Britain, Bulgaria, Croatia, Cyprus, Turkey, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Lithuania, Macedonia, Poland, Russia (Central, East, North, Northwestern, and South), Slovakia, Slovenia, Spain, Sweden, Switzerland, Netherlands, Ukraine, Serbia. Bracon (Orthobracon) exhilarator Nees 1834 on Omphalapion hookerorum (Apionidae); Nanna sp. (Scatophagidae); Acleris rhombana (Tortricidae). Distribution: Belgium, Great Britain, Bulgaria, Cyprus, Turkey, Finland, France, Germany, Hungary, Italy, Latvia, Lithuania, Norway, Poland, Russia (Central, North, Northwestern and South), Slovakia, Sweden, Switzerland, Netherlands, Serbia. Bracon (Orthobracon) laevigatissimus Dalla Torre 1898 on Plagionotus arcuatus (Cerambycidae); Pontania pedunculi, P. proxima, P. vesicator, P. viminalis (Tenthredinidae). Distribution: Austria, Great Britain, Finland, France, Germany, Italy, Poland, Romania, Switzerland, Netherlands, Serbia. Bracon (Orthobracon) ochropus Nees 1834 – no host data. Distribution: Austria, France, Germany, Italy, Serbia. Bracon (Orthobracon) roberti Wesmael 1838 on Synanthedon andrenaeformis (Sesiidae); Cydia strobilella (Tortricidae). Distribution: Austria, Belgium, Great Britain, Turkey, France, Germany, Ireland, Russia (Northwestern), Netherlands, Serbia. Bracon (Orthobracon) subcylindricus Wesmael 1838 – no host data. Distribution: Belgium, Great Britain, France, Poland, Russia (Northwestern), Serbia. Bracon (Orthobracon) tenuicornis Wesmael 1838 on Phloetribus scarabaeoides (Curculionidae). Distribution: Belgium, Great Britain, France, Hungary, Russia (South), Sweden, Netherlands, Serbia. Bracon (Striobracon) idrianus Fahringer 1934 on Hylesinus oleiperda (Curculionidae). Distribution: Serbia. Discussion The present account is the first summary of all available data about the genus Bracon and its hosts until now on the territory of Serbia. Out of total number 250 Bracon species in Europe, 80 species are present in Serbian fauna. We have registered 290 hosts for 55 parasitoid species connected in 409 trophic associations. The hosts come from four insect orders. The most frequent hosts belong to Lepidoptera and Coleoptera, whereas Diptera and Hymenoptera are less parasitized. Numbers in brackets refer to the number of registered host species. 1. Diptera (28) with five families: Agromyzidae (2); Chloropidae (3); Cecidomyiidae (1) Scatophagidae (1) and Tephritidae (21). 2. Coleoptera (109) with nine families: Anobiidae (3); Apionidae (5); Bruchidae (5); Buprestidae BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)… 28 (7); Cerambycidae (12); Chrysomelidae (6); Curculionidae (66); Phalacridae (1) and Rhynchitidae (4). 3. Hymenoptera (26) with four families: Cephidae (2); Cynipidae (1); Eurytomidae (3) and Tenthredinidae (20). 4. Lepidoptera (128) with 21 families: Agonoxenidae (1); Choreutidae (1); Coleophoridae (37); Cosmopterigidae (1); Crambidae (1); Depressariidae (1); Elachistidae (1); Epermeniidae (1); Gelechiidae (11); Geometridae (1); Gracillariidae (4); Momphidae (1); Nepticulidae (1); Noctuidae (5); Oecophoridae (1); Psychidae (1); Pyralidae (9); Sesiidae (14); Tortricidae (24); Yponomeutidae (6) and Zygaenidae (1). The most suitable host species in these trophic relations parasitoid-host, belongs to the coleopteran family Curculionidae and lepidopteran family Coleophoridae especially the genus Coleophora which has 11 parasitoids registered in this work. The torticid moth Cydia strobilella has remarkably wide range of parasitoids, six Bracon species were registered here as parasitoids of this moth (B. (Bracon) variegator, B. (Glabrobracon) caudiger, B. (Glabrobracon) delibator, B. (Glabrobracon) pineti, B. (Glabrobracon) variator and B. (Orthobracon) roberti). We noted that Bracon idrianus is present only on the territory of Serbia. This basic research shows better insight into the trophic interactions about each parasitoid species which can help in further biological control programs. Nonetheless, it can ease the identification of some parasitoid species according to their hosts. Acknowledgements. The authors wish to express their deepest gratitude to Dr. Sergey A. Belokobylskij from the Zoological Institute RAS (St. Petersburg, Russia) and Dr. Jenő Papp from The Natural History Museum of Budapest, Hungary for the identification of parasitoid specimens and to Dr. Aleksandar Ćetković from the Faculty of Biology, University of Belgrade for the loan of some specimens from his collection. For the present work the authors were supported by the Ministry of Science and Environment Protection of the Republic of Serbia (Grant No. III43001). References Achterberg, C. van 2012: Fauna Europaea: Braconidae. In: van Achterberg (ed.): Fauna Europaea: Hymenoptera: Symphyta + Ichneumonoidea. Fauna Europaea, Version 2.5. http://www.faunaeur.org Achterberg, C. van, Hemminga, M.A., Soelen, J. van 1990: New host record of Bracon intercessor Nees f. megasomides Strand (Hymenoptera: Braconidae), a parasite of Agapanthia villosoviridescens DeGeer (Coleoptera: Cerambycidae) in salt marshes. Zoologische Mededelingen, 64 (3): 25-29. Achterberg, C. van. 1985: Notes On Braconidae I- IV. Zoologische Mededelingen, 59 (15): 163- 167. Baird, A.B. 1958: Biological control of insects and plant pests in Canada. In: Baecker, E.C., (ed.), Proceedings of 10 th International Congress of Entomologist, Ottawa, 4: 483-485. Belokobylskij, S.A., Žikić, V. 2009: New data on cyclostome braconid subfamilies Doryctinae, Exothecinae, Rogadinae and Braconinae (Braconidae: Hymenoptera) of Serbia and neighbouring territories. Acta Entomologica Serbica, 14 (1): 65-71. Beyarslan, A., Çetin Erdoğan, Ö., Aydoğdu, M. 2005: A survey of Braconinae (Hymenoptera, Braconidae) of Turkish Western Black Sea Region. Linzer Biologische Beiträge, 37 (1): 195–213. Brajković, M., 1989: Knowledge of the Braconidae (Hymenoptera) fauna in Yugoslavia. Glasnik Prirodnjačkog muzeja, B (43/44): 127-138. Brajković, M., Krunić, M., Tomanović, Ž., 1991: A Contribution to the braconid Fauna (Braconidae: Hymenoptera) of Yugoslavia. In: Hawksworth, D.L. (ed.): Proceedings of the Fourth European Congress of Entomology and the XIII Internationale Symposium für Entomofaunistik Mitteleuropas, Hungarian Natural History Museum, Budapest, (2): 424-428. Brajković, M., Krunić, M., Tomanović, Ž., 1994: Research of braconids (Braconidae: Hymenoptera) of Deliblatska peščara. In: Marinković, P. (ed.): Proceedings of Deliblatski pesak, Srbijašume, 6: 197-502 Dweck, H.K., Svensson, G.P., Gündüz, E.A., Anderbrant, O. 2010: Kairomonal response of the parasitoid, Bracon hebetor Say, to the male- produced sex pheromone of its host, the greater waxmoth, Galleria mellonella (L.). Journal of Chemical Ecology, 36: 171-178. Fischer, M. 1959: Neue und wenig bekannte Braconiden aus Jugoslawien (Hymenoptera). Acta Musei Macedonici Scientiarum Naturalium, 6 (1): 53: 1-25. Georgiev, G. 2005: Bioecological characteristics of Bracon intercessor Nees (Hymenoptera: Braconidae) as a parasitoid of the poplar clearwing moth, Paranthrene tabaniformis http://www.faunaeur.org/full_results.php?id=11049 BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)… 29 (Rott.) (Lepidoptera: Sesiidae) in Bulgaria. Journal of Pest Science, 78: 161-165. Lewis, W.J., Vet, L.E.M., Tumlinson, J.H., van Lenteren, J.C., Papaj, D.R. 1990: Variations in parasitoid foraging behavior: essential element of a sound biological control theory. Environmental Entomology, 19: 1183-1193. Mathews, R.W. 1974: Biology of Braconidae. Annual Review of Entomology, 19: 15-32. Milonas, P.G. 2005: Influence of initial egg density and host size on the development of the gregarious parasitoid Bracon hebetor on three different host species. Biocontrol, 50: 415-428. Papp, J. 1973: Contributions to the braconid fauna of Yugoslavia (Hymenoptera, Braconidae) I. Acta Musei Macedonici Scientarium Naturalium, Skopje, 14 (1: 119): 1-23. Papp, J. 1977: Contribution to the Braconid fauna of Yugoslavia. (Hymenoptera: Braconidae) III. Folia Entomologica Hungarica (Series nova), 30: 105-117. Papp, J. 2008: A revision of the Bracon (subgenera Bracon s. str., Cyanopterobracon, Glabrobracon, Lucobracon, Osculobracon, subgen. n., Pigeria) species described by Szépligeti from the western Palaearctic region. Biologia, 65 (1): 110-112. Reinert, J.A., King, E.W. 1971. Action of Bracon hehetor Say as a parasite of Plodia interpunctella at controlled densities. Annals of the Entomological Society of America, 64: 1335- 1340. Shaw, M.R. and Huddleston, T. 1991: Classification and biology of braconid wasps (Hymenoptera: Braconidae). Royal Entomological Society of London, London. 126p. Stanković, S., Žikić, V., Ilić, M. 2010: Betula species as host plants for various insects parasitized by braconids (Hymenoptera: Braconidae) in Serbia. Biologica Nyssana, 1 (1- 2): 117-122. Taylor, A.D. 1988a: Host effects on larval competition in the gregarious parasitoid Bracon hehetor. Journal of Animal Ecology, 57: 163- 172. Taylor, A.D. 1988b: Host effects on functional and ovipositional responses of Bracon hebetor. Journal of Animal Ecology 57: 173-184. Temerak, A.S. 1982: Laboruntersuchungen zur Wirtshabitat-Wahl von Bracon brevicornis Wesm. (Hym., Braconidae), einem Parasitoiden der Raupen von Sesamia cretica Led. (Lep., Noctuidae) in Sorghum-Stengeln. Anzeiger für Schädlingskunde, Berlin und Hamburg, 55: 152- 154. Tobias, V.I., Belokobylskij, S.A., Kotenko, A.G. 1986: Family Braconidae. - In: Medvedev G.S. (ed.). A Key to Insects of the European Part of the USSR. Hymenoptera 3: 4. Leningrad: Nauka. 500 p. [In Russian]. Yu, D.S., Achterberg, C. van, Horstmann, K. 2012: World Ichneumonoidea 2011. Taxonomy, biology, Morphology and distribution (Braconidae). Taxapad (Scientific names for information management) Interactive catalogue on DVD/CDROM. Vancouver. Yu, S.-H., Ryoo, M.II., Na, J.H. 1999: Life History of Bracon hebetor (Hymenoptera: Braconidae) on Plodia interpunctella (Lepidoptera: Pyralidae) on a Dried Vegetable Commodity. Journal of Asia-Pacific Entomology, 2 (2): 149-152. Žikić, V., Achterberg, C. van, Stanković, S. 2010: A contribution to Braconidae, Hybrizontidae (Ichneumonoidea: Hymenoptera) and Stephanidae (Stephanoidea: Hymenoptera) from the south- West Balkans. Acta Entomologica Serbica, 15 (2): 227-235. Žikić, V., Brajković, M., Tomanović, Ž. 2000: Preliminary results of Braconid fauna research (Braconidae: Hymenoptera). Acta Entomologica Serbica, 5 (1/2): 95-111. Žikić, V., Stanković, S.S., Ilić, M., Kavallieratos, N.G. 2012: Braconid parasitoids (Hymenoptera: Braconidae) on poplars and aspen (Populus spp.) in Serbia and Montenegro. North-Western Journal of Zoology [in press]. BIOLOGICA NYSSANA 3 (1)  September 2012: 21-29 Žikić, V. et al.  Checklist of the genus Bracon (Hymenoptera: Braconidae)… 30