Xhulaj 2022, Biologica Nyssana 13(2) 13 (2) December 2022: 173-178 DOI: 10.5281/zenodo.7437326 Various records of lichens from southeastern Albania Original Article Skerdilaid Xhulaj Research Center of Flora and Fauna, Faculty of Natural Sciences, University of Tirana, Bulevardi Zog I, Nr. 25/1, Tirana, Albania skerdilaid.xhulaj@fshn.edu.al (corresponding author) Received: October 10, 2022 Revised: November 16, 2022 Accepted: November 23, 2022 Abstract: 117 taxa of lichens from two neighboring districts (Pogradec and Korçë) in the southeast of Albania are presented. 62 taxa are new for the study area and 27 taxa are new to Albania. Key words: biodiversity, lichens, flora of Albania, new records Apstrakt: Različiti zapisi o lišajima iz jugoistočne Albanije Predstavljeno je 117 taksona lišaja iz dva susedna okruga (Pogradec i Korçë) na jugoistoku Albanije. 62 taksona su nova za područje istraživanja i 27 taksona su nova za Albaniju. Ključne reči: biodiverzitet, lišaji, flora Albanije, novi zapisi Introduction Hitherto, no thorough investigation about the lichens of this area has been carried out. As a result, publications are quite sporadic or lacking. Markgraf (1927, 1931) has produced a few records from Korçë district as part of his expedition focused on the flora and vegetation of Albania. Hafellner (2007) provides a checklist of lichens based on historical records by several authors. This list does not include any species of lichen from the area specifically covered in this investigation, except those reported by Markgraf (1927, 1931). Svoboda et al. (2012) have reported a greater number of lichens from 5 localities visited in this area (1 in the district of Pogradec and 4 in the district of Korça) compared to the above-mentioned authors. It should be noted that their localities in the district of Korça were positioned within the Bredhi i Drenovës National Park, leaving other areas or habitats of this district uncovered. The results presented below are based on fieldwork undertaken by author encouraged by the need to increase and improve knowledge on the diversity of lichens in Albania. Materials and Methods Specimens were collected by the author during field trips carried out in both districts. The sampling localities were chosen according to diversity of the habitats paying attention to the substrates. Specimens were determined using lichenological routine light microscopy methods and based on relevant literature according to Poelt (1969), Poelt & Vězda (1977, 1981), Clauzade & Roux (1985), Wirth (1995), Nimis & Martellos (2004), Smith et al. (2009), Wirth et al. (2013). The nomenclature follows Nimis & Martellos (2008) and reflects a more modern concept of genus. The specimens are deposited in the author’s personal herbarium. List of sampling localities 1: Pogradec district, calcareous hills W of Ohrid lake c. 20 km N of Pogradec, 41°03’40”N/20°36’36.2”E, alt. c. 700 m, 10.06.2020. 2: Pogradec district, calcareous rocks on Carpinus scrub, near the village Rëmenj, c. 16 km S of Pogradec, an area characterized by drought but influenced by human activity, 40°51’05.6”N/20°36’49.4”E, alt. c. 830 m, 10.06.2020. 3: Pogradec district, near the village Peshkëpi, c. 20 km E of Pogradec, 40°53’45.9”N/20°46’45.7”E, alt. c. 940 m, 11.06.2020. 4: Pogradec district, near the village Guri i Bardhë, c. 22 km W of Pogradec, hedges between pastures, 40°52’53.8”N/20°33’03.6”E, alt. c. 1050 m, 11.06.2020. 5: Korçë district, on roadside trees (Populus), c. 5 km NW of Korça, 40°39’13.8”N/20°44’48”E, alt. c. 850 m, 12.06.2020. 6: Korçë district, on Robinia behind the Memorial of Martyrs in Korçë, 40°37’10.3”N/20°47’22.4”E, alt. 950 m, 12.06.2020. © 2022 Xhulaj. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and build upon your work non-commercially under the same license as the original. 173 14th Symposium on the Flora of Southeastern Serbia and Neighboring Regions 174 BIOLOGICA NYSSANA ● 13 (2) December 2022: 173-178 Xhulaj ● Various records of lichens from southeastern Albania 7: Korçë district, Bozdovec, Bredhi i Drenovës National Park, c. 7 km E of the village Drenovë, on hill wooded by middle-aged mixed forest with dominance of Abies borisii-regis in the northern part of the park, 40°34’36.8”N/20°50’34.5”E, alt. 1360 m, 13.06.2020. 8: Korçë district, Korçë, Bredhi i Drenovës National Park, tourist route in the southern part of the park, 40°32’33.3”N/20°49’18.2”E, alt. 1240-1300 m, 14.06.2020. 9: Korçë district, near the village Vithkuq, c. 32 km SW of Korçë, limestone outcrops, mixed deciduous forest, 40°31’05.3”N/20°35’09.4”E, alt. c. 1230 m, 15.06.2020. 10: Korçë district, near the village Dardhë, c. 18 km SE of Korçë, mixed forest, 40°30’54”N/20°49’25”E, alt. c. 1250 m, 16.06.2020. List of substrates and their abbreviations: cor: on bark of trees cal: on limestone ter-cal: on calcareous soil sil: on siliceous rocks bry: on bryophytes Abb: Abies borisii-regis Aps: Acer pseudoplatanus Car: Carpinus sp. Fag: Fagus sylvatica Pop: Populus sp. Qce: Quercus cerris Rps: Robinia pseudoacacia Sal: Salix sp. Results A total of 117 taxa of lichens were recorded, including 62 lichens new to the investigation area and 27 new to the country. Species are arranged alphabetically. Numbers from 1 to 10 correspond to the localities listed above; the abbreviations following this number correspond to the substrates listed above. Author’s initials and numbers in brackets show individual consecutive numbering systems used by the author to identify and locate individual specimens. New records for the investigation area are indicated with an asterisk (*), and new records for Albania with a ring (o). Acarospora cervina A. Massal. 1, 2, 5, 9 cal (SX 7108, 7211, 8521, 8964) *Acarospora glaucocarpa (Ach.) Körb. 1 cal (SX 7133) Anaptychia ciliaris (L.) Flot. 1 Car, 6 Rps (SX 7113, 8656) *oArthonia apatetica (A. Massal.) Th. Fr. 3, 4, 5, cor (SX 7314, 7404, 8593) Arthonia didyma Körb. 7, 8, 10 Abb (SX 8772, 8811, 8123) Athallia alnetorum (Giralt, Nimis & Poelt) Arup, Frödén & Søchting 4 Qce, 5 Pop (SX 7415, 8542) *oAthallia cerinella (Nyl.) Arup, Frödén & Søchting 6 cor (SX 8619) Athallia holocarpa (Hoffm.) Arup, Frödén & Søchting 5 cor (SX 8565) *oAthallia pyracea (Ach.) Arup, Frödén & Søchting 4, 7, 8, 10 Aps (SX 7423, 8735, 8809, 8176) *Bagliettoa marmorea (Scop.) Gueidan & Cl. Roux 1 cal (SX 7186) *oBagliettoa parmigera (J. Steiner) Vězda & Poelt 1 cal (SX 7153) *oBagliettoa parmigerella (Zahlbr.) Vězda & Poelt 1 cal (SX 7104) Blastenia crenularia (With.) Arup, Søchting & Frödén 1, 2, 4 cal (SX 7111, 7270, 7441) Blastenia ferruginea (Huds.) A. Massal. 3, 4, 6, 9 cor (SX 7315, 7476, 8655, 8911) Calicium salicinum Pers. 7, 8 Abb (SX 8768, 8832) Caloplaca cerina (Hedw.) Th. Fr. 1 cor, 7 Fag, 8 Fag, 9 Sal (SX 7174, 8761, 8844, 8924) *Caloplaca conversa (Kremp.) Jatta 1 cal (SX 7168) *Candelariella aurella (Hoffm.) Zahlbr. 1 cal (SX 7156) *Candelariella vitellina (Hoffm.) Müll. Arg 6, 10 cor (SX 8660, 8111) Candelariella xanthostigma (Ach.) Lettau 3, 4, 6 cor, 7, 8 Abb (SX 7310, 7437, 8692, 8750, 8812) *Circinaria calcarea (L.) A. Nordin, Savić & Tibell 1, 2, 3, 6, 9, 10 cal (SX 7137, 7261, 7303, 8605, 8922, 8108) *Circinaria contorta (Hoffm.) A. Nordin, Savić & Tibell 1, 2, 3, 6, 9 cal (SX 7157, 7224, 7302, 8613, 8957) *oCircinaria hoffmanniana (S. Ekman & Fröberg ex R. Sant.) A. Nordin 1, 2 cal (SX 7107, 7223) 175 BIOLOGICA NYSSANA ● 13 (2) December 2022: 173-178 Xhulaj ● Various records of lichens from southeastern Albania *Cladonia pyxidata (L.) Hoffm. f. pocillum (Ach.) Nyl. 1, 2, ter-cal (SX 7105, 7219) *Cladonia pyxidata (L.) Hoffm. f. pyxidata 1, 2, 3, 4 ter-cal, 7, 9, 10 bry (SX 7152, 7261, 7308, 7430, 8708, 8964, 8152) Collema flaccidum (Ach.) Ach. 1, 2, 3, 4 ter-cal (SX 7170, 7252, 7344, 7408) *oEnchylium polycarpon (Hoffm.) Otálora, P.M. Jørg. & Wedin subsp. polycarpon 1, 2, 4 cal (SX 7114, 7216, 7426) Enchylium tenax (Sw.) Gray 1, 2, 3, 4 ter-cal (SX 7175, 7246, 7322, 7467) *Fuscopannaria praetermissa (Nyl.) P.M. Jørg. 1 ter-cal, 9 bry (SX 7169, 8913) *oGyalolechia flavorubescens (Huds.) Søchting, Frödén & Arup var. flavorubescens 2 Car, 5 Pop (SX 7266, 8590) Hypogymnia farinacea Zopf 7, 8 Abb (SX 8718, 8875) *Lathagrium auriforme (With.) Otálora, P.M. Jørg. & Wedin 1 cal (SX 7103) *Lathagrium cristatum (L.) Otálora, P.M. Jørg. & Wedin 1, 2, 9 cal (SX 7161, 7230, 8959) Lecanora allophana (Ach.) Nyl. f. allophana 3, 5, 6 Pop, 7, 8, 10 Fag (SX 7317, 8546, 8643, 8715, 8808, 8151) Lecanora allophana f. sorediata Vain. 3, 5 Pop (SX 7347, 8513) *Lecanora argentata (Ach.) Malme 7, 8, 9 Fag (SX 8780, 8867, 8934) *Lecanora carpinea (L.) Vain. 3, 4, 10 cor (SX 7319, 7440, 8105) Lecanora chlarotera Nyl. subsp. chlarotera 3, 4 cor, 8 Fag (SX 7324, 7449, 8833) Lecanora intumescens (Rebent.) Rabenh. 7, 8 Fag (SX 8762, 8843) Lecanora leptyrodes (Nyl.) Degel. 1, 3, 4, 8, 9, 10 cor (SX 7102, 7366, 7465, 8812, 8918, 8116) Lecanora salicicola H. Magn. 4, 9 cor (SX 7409, 8912) Lecanora subcarpinea Szatala 7, 8, 10 Abb (SX 8709, 8822, 8160) Lecidea atrobrunnea (DC.) Schaer. subsp. atrobrunnea 3, 10 sil (SX 7313, 8149) *Lecidella elaeochroma (Ach.) M. Choisy var. elaeochroma 1, 3, 5, 6, 10 cor (SX 7127, 7329, 8597, 8617, 8121) *Lecidella euphorea (Flörke) Hertel 1, 2, 7, 9 cor (SX 7106, 7255, 8731, 8961) Lecidella flavosorediata (Vězda) Hertel & Leuckert 5, 6, 9 cor (SX 8536, 8671, 8953) Lepra albescens (Huds.) Hafellner 9, 10 cor (SX 8926, 8172) Leptogium saturninum (Dicks.) Nyl. 4 cor, 8 Fag (SX 7413, 8841) *Lobothallia radiosa (Hoffm.) Hafellner 1, 2, 9 cal (SX 7119, 7279, 8948) Melanohalea exasperata (De Not.) O. Blanco, A. Crespo, Divakar, Essl., D. Hawksw. & Lumbsch 9, 10 cor (SX 8931, 8124) Melanohalea laciniatula (H. Olivier) O. Blanco, A. Crespo, Divakar, Essl., D. Hawksw. & Lumbsch 8 Abb (SX 8847) *oMicarea globulosella (Nyl.) Coppins 10 Abb (SX 8131) *Myriolecis dispersa (Pers.) Sliwa, Zhao Xin & Lumbsch 2, 9 cal (SX 7253, 8938) Myriolecis hagenii (Ach.) Sliwa, Zhao Xin & Lumbsch 3, 5, 6 cor (SX 7316, 8594, 8612) Nephroma resupinatum (L.) Ach. 4, 9, 10 cor (SX 7432, 8925, 8137) Ochrolechia pallescens (L.) A. Massal. 7 Aps, 8 Abb (SX 8710, 8859) Ochrolechia szatalaensis Verseghy 10 Abb (SX 8128) Parmelia submontana Hale 7 Fag, 10 Abb (SX 8724, 8162) Phaeophyscia nigricans (Flörke) Moberg 2 cor, 5 Pop (SX 7247, 8567) Phaeophyscia orbicularis (Neck.) Moberg 1, 2, 5, 6 cor (SX 7158, 7271, 8525, 8644) *Physcia adscendens H. Olivier 2, 6, 9 cor (SX 7281, 8672, 8903) Physcia aipolia (Humb.) Fürnr. 7 Abb, 9 cor (SX 8704, 8962) *Physcia biziana (A. Massal.) Zahlbr. var. biziana 176 BIOLOGICA NYSSANA ● 13 (2) December 2022: 173-178 Xhulaj ● Various records of lichens from southeastern Albania 4, 6, 9, 10 cor (SX 7405, 8640, 8958, 8136) *Physcia dubia (Hoffm.) Lettau 2 cal, 6 cor (SX 7267, 8673) *Physcia leptalea (Ach.) DC. 9 cor (SX 8919) *Physcia stellaris (L.) Nyl. 5, 9 cor (SX 8553, 8914) *Physciella chloantha (Ach.) Essl. 9 cor (SX 8933) Physconia distorta (With.) J.R. Laundon 7 Fag, 8 Aps, 9 Abb (SX 8759, 8848, 8927) Physconia perisidiosa (Erichsen) Moberg 8 Abb (SX 8814) *oPlacidium lachneum (Ach.) B. de Lesd. 1, 2 ter-cal, 9 bry (SX 7139, 7222, 8924) *oPlacocarpus schaereri (Fr.) Breuss 1, 2 cal (SX 7199, 7286) *oPlacolecis opaca (Dufour) Hafellner 2 cal (SX 7280) *Placynthium nigrum (Huds.) Gray 2 cal (SX 7278) Pleurosticta acetabulum (Neck.) Elix & Lumbsch 7, 8 Abb (SX 8725, 8846) *oPolycauliona polycarpa (Hoffm.) Frödén, Arup & Søchting 5 cor (SX 8518) Protoblastenia incrustans (DC.) J. Steiner var. incrustans 1, 2, 9 cal (SX 7116, 7265, 8952) *oProtoparmeliopsis graeca (J.Steiner) Sipman & Cl. Roux 2 cal (SX 7251) *Protoparmeliopsis muralis (Schreb.) M. Choisy var. muralis 1, 9 cal (SX 7177, 8906) Psora rubiformis (Ach.) Hook. 7 sil, 10 bry (SX 8757, 8125) *oPyrenodesmia chalybaea (Fr.) A. Massal. 1, 2, 4 cal (SX 7163, 7249, 7463) *Pyrenodesmia variabilis (Pers.) A. Massal. 2, 4, 9 cal (SX 7245, 7433, 8923) Ramalina calicaris (L.) Fr. 3, 4 cor, 8, 10 Abb (SX 7345, 7434, 8823, 8142) Ramalina canariensis J. Steiner 3, 4, 9 cor (SX 7328, 7481, 8907) Ramalina fraxinea (L.) Ach. 7, 8, 9 cor (SX 8792, 8836, 8951) *Rinodina albana (A. Massal.) A. Massal. 3, 6 cor (SX 7320, 8606) *oRinodina anomala (Zahlbr.) H. Mayrhofer & Giralt 2 cor (SX 7204) *oRinodina bischoffii (Hepp) A. Massal. 1, 2 cal (SX 7162, 7222) *Rinodina exigua (Ach.) Gray 3, 9 cor (SX 7338, 8929) Rinodina immersa (Körb.) J. Steiner 1, 2, 4, 9 cal (SX 7190, 7283, 7444, 8902) *oRinodina pyrina (Ach.) Arnold 2 cor (SX 7286) *oRinodinella dubyanoides (Hepp) H. Mayrhofer & Poelt 1, 2 cal (SX 7193, 7232) Sanguineodiscus haematites (Chaub. ex St.-Amans) I.V. Frolov & Vondrák 1, 2, 4, 8 cor (SX 7130, 7243, 7461, 8834) *Sarcogyne regularis Körb. var. regularis 2 cal (SX 7273) Scoliciosporum umbrinum (Ach.) Arnold 7, 8, 10 Abb (SX 8736, 8882, 8153) *oScytinium fragile (Taylor) Otálora, P.M. Jørg. & Wedin 1, 2 cal (SX 7144, 7213) Scytinium lichenoides (L.) Otálora, P.M. Jørg. & Wedin 1, 2 ter-cal, 6, 10 bry (SX 7164, 7248, 8639, 8117) Scytinium pulvinatum (Hoffm.) Otálora, P.M. Jørg. & Wedin 1, 2, 7 ter-cal (SX 7165, 7250, 8732) Scytinium tenuissimum (Hoffm.) Otálora, P.M. Jørg. & Wedin 1, 2 cor (SX 7166, 7254) Squamarina cartilaginea (With.) P. James 1, 2 ter-cal (SX 7124, 7257) Staurothele areolata (Ach.) Lettau 7, 8, 10 sil (SX 8734, 8845, 8143) *oSynalissa ramulosa (Bernh.) Fr. 2 cal (SX 7264) *Thalloidima candidum (Weber) A.Massal. 1, 2 cal (SX 7112, 7238) *oThalloidima diffractum (A. Massal.) A. Massal. 1 cal (SX 7129) BIOLOGICA NYSSANA ● 13 (2) December 2022: 173-178 Xhulaj ● Various records of lichens from southeastern Albania Thalloidima opuntioides (Vill.) Kistenich, Timdal, Bendiksby & S.Ekman 2 cal, 9 bry (SX 7229, 8963) Usnea barbata (L.) F.H. Wigg. 8 Abb (SX 8821) Usnea perplexans Stirt. 10 Abb (SX 8104) *Variospora aurantia (Pers.) Arup, Frödén & Søchting 1, 9 cal (SX 7146, 8928) *oVariospora dalmatica (A. Massal.) 1, 2 cal (SX 7142, 7233) *oVariospora flavescens (Huds.) Arup, Frödén & Søchting 2 cal (SX 7236) *oVerrucaria caerulea DC. 1, 2 cal (SX 7191, 7260) *Verrucaria nigrescens Pers. 1, 2 cal (SX 7192, 7258) *Xanthocarpia ochracea (Schaer.) A. Massal. & De Not. 1, 2 cal (SX 7115, 7242) *oXanthomendoza fallax (Hepp) Søchting, Kärnefelt & S.Y. Kondr. 5 cor (SX 8508) Xanthomendoza fulva (Hoffm.) Søchting, Kärnefelt & S.Y. Kondr. 8, 10 cor (SX 8863, 8133) *oXanthoparmelia glabrans (Nyl.) O. Blanco, A. Crespo, Elix, D. Hawksw. & Lumbsch 7 sil (SX 8755) Xanthoparmelia pulla (Ach.) O. Blanco, A. Crespo, Elix, D. Hawksw. & Lumbsch 7, 10 sil (SX 8758, 8122) *Xanthoria parietina (L.) Th. Fr. 1, 2 cal, 4, 5, 6, 9 cor (SX 7109, 7263, 7435, 8541, 8654, 8921) Discussion Thus far, 55 taxa of lichens were known from both districts (Markgraf, 1927, 1931; Svoboda et al., 2012). This study is able to add a further 62 taxa. More than 60% of the lichen taxa added by this contribution are crustose and many of them are widely distributed in the Mediterranean region. Regarding the substrate ecology of the newly recorded lichen taxa, 58% of the added taxa are saxicolous, 35% are corticolous and only 6% are terricolous or muscicolous. New lichen species for the Albania are Arthonia apatetica, Athallia cerinella, A. pyracea, Bagliettoa parmigera, B. parmigerella, Circinaria hoffmanniana, Enchylium polycarpon subsp. polycarpon, Gyalolechia flavorubescens var. flavorubescens, Micarea globulosella, Placidium lachneum, Placocarpus schaereri, Placolecis opaca, Polycauliona polycarpa, Protoparmeliopsis graeca, Pyrenodesmia chalybaea, Rinodina anomala, R. bischoffii, R. pyrina, Rinodinella dubyanoides, Scytinium fragile, Synalissa ramulosa, Thalloidima diffractum, Variospora dalmatica, V. flavescens, Verrucaria caerulea, Xanthomendoza fallax and Xanthoparmelia glabrans. It should be noted: In Korça town itself were only found three species, miserably developed (Physcia biziana, Physciella chloantha and Phaeophyscia orbicularis). Compared to the coastal areas visited so far this is poor, presumably because of the dry climate far from the coast. Calcareous rocks on Carpinus scrub south of Pogradec, an area characterized by drought but influenced by human activity. To be noticed on Carpinus: Anaptychia ciliaris, Gyalolechia flavorubescens, Physcia leptalea, Rinodina pyrina. On limestone: Placocarpus schaereri, Placolecis opaca, Scytinium fragile, Synalissa ramulosa and Thalloidima diffractum. Conclusions Despite the considerable number of taxa found and localities visited, it is expected that the lichen flora should be richer due to the fact that some areas are difficult to reach and the large extent of both districts. Although many factors influence the species composition of the investigated localities, it may be concluded that the general climate and the potential natural vegetation/vegetation zones adjust the general pattern of the lichenic flora, while the stand parameters and the conditions of the instant surroundings, especially the topography-influenced local climatic conditions and the present vegetation surrounding the habitats, may affect the species composition to a considerable extent. References Clauzade, G., Roux, C. 1985: Likenoj de okcidenta Eŭropo. Ilustrita determinlibro. Bulletin de la Société Botanique du Centre-Ouest. Nouvelle Série. Numéro Spécial, 7: 1-893. Hafellner, J. 2007: Checklist and bibliography of lichenized and lichenicolous fungi so far reported from Albania (version 05-2007). 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