id	sid	tid	token	lemma	pos
biotropia-309	1	1	5	5	X
biotropia-309	1	2	.	.	X
biotropia-309	1	3	shahabuddin	shahabuddin	NOUN
biotropia-309	1	4	(	(	PUNCT
biotropia-309	1	5	effectiveness	effectiveness	NOUN
biotropia-309	1	6	...	...	PUNCT
biotropia-309	1	7	biotropia	biotropia	ADJ
biotropia-309	1	8	vol	vol	NOUN
biotropia-309	1	9	.	.	PROPN
biotropia-309	1	10	21	21	NUM
biotropia-309	1	11	no	no	NOUN
biotropia-309	1	12	.	.	NOUN
biotropia-309	1	13	1	1	NUM
biotropia-309	1	14	,	,	PUNCT
biotropia-309	1	15	20114	20114	NUM
biotropia-309	1	16	:	:	PUNCT
biotropia-309	1	17	48	48	NUM
biotropia-309	1	18	58	58	NUM
biotropia-309	1	19	effectiveness	effectiveness	NOUN
biotropia-309	1	20	of	of	ADP
biotropia-309	1	21	dung	dung	NOUN
biotropia-309	1	22	beetles	beetle	NOUN
biotropia-309	1	23	as	as	ADP
biotropia-309	1	24	bioindicators	bioindicator	NOUN
biotropia-309	1	25	of	of	ADP
biotropia-309	1	26	environmental	environmental	ADJ
biotropia-309	1	27	changes	change	NOUN
biotropia-309	1	28	in	in	ADP
biotropia-309	1	29	land	land	NOUN
biotropia-309	1	30	-	-	PUNCT
biotropia-309	1	31	use	use	NOUN
biotropia-309	1	32	gradient	gradient	NOUN
biotropia-309	1	33	in	in	ADP
biotropia-309	1	34	sulawesi	sulawesi	PROPN
biotropia-309	1	35	,	,	PUNCT
biotropia-309	1	36	indonesia	indonesia	PROPN
biotropia-309	1	37	shahabuddin	shahabuddin	NOUN
biotropia-309	1	38	,	,	PUNCT
biotropia-309	1	39	uswah	uswah	VERB
biotropia-309	1	40	hasanah	hasanah	PROPN
biotropia-309	1	41	,	,	PUNCT
biotropia-309	1	42	and	and	CCONJ
biotropia-309	1	43	elijonnahdi	elijonnahdi	PROPN
biotropia-309	1	44	received	receive	VERB
biotropia-309	1	45	13	13	NUM
biotropia-309	1	46	june	june	PROPN
biotropia-309	1	47	2013	2013	NUM
biotropia-309	1	48	/	/	SYM
biotropia-309	1	49	accepted	accept	VERB
biotropia-309	1	50	09	09	NUM
biotropia-309	1	51	november	november	PROPN
biotropia-309	1	52	2013	2013	NUM
biotropia-309	1	53	bioindicators	bioindicator	NOUN
biotropia-309	1	54	have	have	AUX
biotropia-309	1	55	been	be	AUX
biotropia-309	1	56	widely	widely	ADV
biotropia-309	1	57	accepted	accept	VERB
biotropia-309	1	58	as	as	ADP
biotropia-309	1	59	useful	useful	ADJ
biotropia-309	1	60	tools	tool	NOUN
biotropia-309	1	61	for	for	ADP
biotropia-309	1	62	monitoring	monitor	VERB
biotropia-309	1	63	and	and	CCONJ
biotropia-309	1	64	detecting	detect	VERB
biotropia-309	1	65	changes	change	NOUN
biotropia-309	1	66	in	in	ADP
biotropia-309	1	67	the	the	DET
biotropia-309	1	68	environment	environment	NOUN
biotropia-309	1	69	or	or	CCONJ
biotropia-309	1	70	habitat	habitat	NOUN
biotropia-309	1	71	condition	condition	NOUN
biotropia-309	1	72	.	.	PUNCT
biotropia-309	2	1	by	by	ADP
biotropia-309	2	2	using	use	VERB
biotropia-309	2	3	bioindicators	bioindicator	NOUN
biotropia-309	2	4	,	,	PUNCT
biotropia-309	2	5	it	it	PRON
biotropia-309	2	6	is	be	AUX
biotropia-309	2	7	possible	possible	ADJ
biotropia-309	2	8	to	to	PART
biotropia-309	2	9	assess	assess	VERB
biotropia-309	2	10	the	the	DET
biotropia-309	2	11	impact	impact	NOUN
biotropia-309	2	12	of	of	ADP
biotropia-309	2	13	human	human	ADJ
biotropia-309	2	14	activities	activity	NOUN
biotropia-309	2	15	on	on	ADP
biotropia-309	2	16	the	the	DET
biotropia-309	2	17	biota	biota	NOUN
biotropia-309	2	18	,	,	PUNCT
biotropia-309	2	19	instead	instead	ADV
biotropia-309	2	20	of	of	ADP
biotropia-309	2	21	examining	examine	VERB
biotropia-309	2	22	the	the	DET
biotropia-309	2	23	entire	entire	ADJ
biotropia-309	2	24	biota	biota	NOUN
biotropia-309	2	25	.	.	PUNCT
biotropia-309	3	1	in	in	ADP
biotropia-309	3	2	this	this	DET
biotropia-309	3	3	paper	paper	NOUN
biotropia-309	3	4	we	we	PRON
biotropia-309	3	5	analyzed	analyze	VERB
biotropia-309	3	6	diversity	diversity	NOUN
biotropia-309	3	7	of	of	ADP
biotropia-309	3	8	dung	dung	NOUN
biotropia-309	3	9	beetles	beetle	NOUN
biotropia-309	3	10	(	(	PUNCT
biotropia-309	3	11	coleoptera	coleoptera	NOUN
biotropia-309	3	12	:	:	PUNCT
biotropia-309	3	13	scarabaeidae	scarabaeidae	NOUN
biotropia-309	3	14	)	)	PUNCT
biotropia-309	3	15	across	across	ADP
biotropia-309	3	16	land	land	NOUN
biotropia-309	3	17	use	use	NOUN
biotropia-309	3	18	gradient	gradient	NOUN
biotropia-309	3	19	in	in	ADP
biotropia-309	3	20	central	central	ADJ
biotropia-309	3	21	sulawesi	sulawesi	NOUN
biotropia-309	3	22	and	and	CCONJ
biotropia-309	3	23	tested	test	VERB
biotropia-309	3	24	the	the	DET
biotropia-309	3	25	suitability	suitability	NOUN
biotropia-309	3	26	of	of	ADP
biotropia-309	3	27	dung	dung	NOUN
biotropia-309	3	28	beetles	beetle	NOUN
biotropia-309	3	29	as	as	ADP
biotropia-309	3	30	bioindicators	bioindicator	NOUN
biotropia-309	3	31	for	for	ADP
biotropia-309	3	32	environmental	environmental	ADJ
biotropia-309	3	33	changes	change	NOUN
biotropia-309	3	34	.	.	PUNCT
biotropia-309	4	1	ninety	ninety	NUM
biotropia-309	4	2	baited	bait	VERB
biotropia-309	4	3	pitfall	pitfall	NOUN
biotropia-309	4	4	traps	trap	NOUN
biotropia-309	4	5	were	be	AUX
biotropia-309	4	6	placed	place	VERB
biotropia-309	4	7	and	and	CCONJ
biotropia-309	4	8	several	several	ADJ
biotropia-309	4	9	habitat	habitat	NOUN
biotropia-309	4	10	parameters	parameter	NOUN
biotropia-309	4	11	were	be	AUX
biotropia-309	4	12	measured	measure	VERB
biotropia-309	4	13	at	at	ADP
biotropia-309	4	14	five	five	NUM
biotropia-309	4	15	land	land	NOUN
biotropia-309	4	16	-	-	PUNCT
biotropia-309	4	17	use	use	NOUN
biotropia-309	4	18	types	type	NOUN
biotropia-309	4	19	ranging	range	VERB
biotropia-309	4	20	from	from	ADP
biotropia-309	4	21	natural	natural	ADJ
biotropia-309	4	22	forest	forest	NOUN
biotropia-309	4	23	to	to	PART
biotropia-309	4	24	cacao	cacao	VERB
biotropia-309	4	25	agroforestry	agroforestry	NOUN
biotropia-309	4	26	systems	system	NOUN
biotropia-309	4	27	to	to	PART
biotropia-309	4	28	open	open	VERB
biotropia-309	4	29	areas	area	NOUN
biotropia-309	4	30	in	in	ADP
biotropia-309	4	31	2009	2009	NUM
biotropia-309	4	32	and	and	CCONJ
biotropia-309	4	33	2012	2012	NUM
biotropia-309	4	34	.	.	PUNCT
biotropia-309	5	1	the	the	DET
biotropia-309	5	2	effectiveness	effectiveness	NOUN
biotropia-309	5	3	of	of	ADP
biotropia-309	5	4	dung	dung	NOUN
biotropia-309	5	5	beetles	beetle	NOUN
biotropia-309	5	6	as	as	ADP
biotropia-309	5	7	bioindicators	bioindicator	NOUN
biotropia-309	5	8	of	of	ADP
biotropia-309	5	9	environmental	environmental	ADJ
biotropia-309	5	10	changes	change	NOUN
biotropia-309	5	11	was	be	AUX
biotropia-309	5	12	evaluated	evaluate	VERB
biotropia-309	5	13	by	by	ADP
biotropia-309	5	14	the	the	DET
biotropia-309	5	15	method	method	NOUN
biotropia-309	5	16	,	,	PUNCT
biotropia-309	5	17	a	a	DET
biotropia-309	5	18	method	method	NOUN
biotropia-309	5	19	combining	combine	VERB
biotropia-309	5	20	the	the	DET
biotropia-309	5	21	specificity	specificity	NOUN
biotropia-309	5	22	and	and	CCONJ
biotropia-309	5	23	fidelity	fidelity	NOUN
biotropia-309	5	24	of	of	ADP
biotropia-309	5	25	certain	certain	ADJ
biotropia-309	5	26	species	specie	NOUN
biotropia-309	5	27	with	with	ADP
biotropia-309	5	28	particular	particular	ADJ
biotropia-309	5	29	types	type	NOUN
biotropia-309	5	30	of	of	ADP
biotropia-309	5	31	habitat	habitat	NOUN
biotropia-309	5	32	or	or	CCONJ
biotropia-309	5	33	environmental	environmental	ADJ
biotropia-309	5	34	conditions	condition	NOUN
biotropia-309	5	35	.	.	PUNCT
biotropia-309	6	1	surprisingly	surprisingly	ADV
biotropia-309	6	2	,	,	PUNCT
biotropia-309	6	3	the	the	DET
biotropia-309	6	4	results	result	NOUN
biotropia-309	6	5	showed	show	VERB
biotropia-309	6	6	that	that	SCONJ
biotropia-309	6	7	the	the	DET
biotropia-309	6	8	diversity	diversity	NOUN
biotropia-309	6	9	of	of	ADP
biotropia-309	6	10	dung	dung	NOUN
biotropia-309	6	11	beetles	beetle	NOUN
biotropia-309	6	12	in	in	ADP
biotropia-309	6	13	two	two	NUM
biotropia-309	6	14	types	type	NOUN
biotropia-309	6	15	of	of	ADP
biotropia-309	6	16	cacao	cacao	NOUN
biotropia-309	6	17	plantations	plantation	NOUN
biotropia-309	6	18	were	be	AUX
biotropia-309	6	19	similar	similar	ADJ
biotropia-309	6	20	to	to	ADP
biotropia-309	6	21	the	the	DET
biotropia-309	6	22	forest	forest	NOUN
biotropia-309	6	23	sites	site	NOUN
biotropia-309	6	24	and	and	CCONJ
biotropia-309	6	25	were	be	AUX
biotropia-309	6	26	significantly	significantly	ADV
biotropia-309	6	27	higher	high	ADJ
biotropia-309	6	28	than	than	ADP
biotropia-309	6	29	the	the	DET
biotropia-309	6	30	open	open	ADJ
biotropia-309	6	31	cultivated	cultivate	VERB
biotropia-309	6	32	area	area	NOUN
biotropia-309	6	33	.	.	PUNCT
biotropia-309	7	1	of	of	ADP
biotropia-309	7	2	the	the	DET
biotropia-309	7	3	16	16	NUM
biotropia-309	7	4	dung	dung	NOUN
biotropia-309	7	5	beetles	beetle	NOUN
biotropia-309	7	6	species	specie	NOUN
biotropia-309	7	7	analyzed	analyze	VERB
biotropia-309	7	8	only	only	ADV
biotropia-309	7	9	four	four	NUM
biotropia-309	7	10	species	specie	NOUN
biotropia-309	7	11	could	could	AUX
biotropia-309	7	12	be	be	AUX
biotropia-309	7	13	suggested	suggest	VERB
biotropia-309	7	14	as	as	ADP
biotropia-309	7	15	indicator	indicator	NOUN
biotropia-309	7	16	(	(	PUNCT
biotropia-309	7	17	characteristic	characteristic	ADJ
biotropia-309	7	18	)	)	PUNCT
biotropia-309	7	19	species	specie	NOUN
biotropia-309	7	20	while	while	SCONJ
biotropia-309	7	21	the	the	DET
biotropia-309	7	22	majority	majority	NOUN
biotropia-309	7	23	of	of	ADP
biotropia-309	7	24	collected	collect	VERB
biotropia-309	7	25	species	specie	NOUN
biotropia-309	7	26	were	be	AUX
biotropia-309	7	27	categorized	categorize	VERB
biotropia-309	7	28	as	as	ADP
biotropia-309	7	29	detector	detector	NOUN
biotropia-309	7	30	species	specie	NOUN
biotropia-309	7	31	.	.	PUNCT
biotropia-309	8	1	two	two	NUM
biotropia-309	8	2	of	of	ADP
biotropia-309	8	3	them	they	PRON
biotropia-309	8	4	(	(	PUNCT
biotropia-309	8	5	and	and	CCONJ
biotropia-309	8	6	)	)	PUNCT
biotropia-309	8	7	were	be	AUX
biotropia-309	8	8	associated	associate	VERB
biotropia-309	8	9	with	with	ADP
biotropia-309	8	10	natural	natural	ADJ
biotropia-309	8	11	forest	forest	NOUN
biotropia-309	8	12	and	and	CCONJ
biotropia-309	8	13	cacao	cacao	NOUN
biotropia-309	8	14	agroforestry	agroforestry	NOUN
biotropia-309	8	15	system	system	NOUN
biotropia-309	8	16	,	,	PUNCT
biotropia-309	8	17	thus	thus	ADV
biotropia-309	8	18	were	be	AUX
biotropia-309	8	19	suggested	suggest	VERB
biotropia-309	8	20	as	as	ADP
biotropia-309	8	21	the	the	DET
biotropia-309	8	22	indicator	indicator	NOUN
biotropia-309	8	23	of	of	ADP
biotropia-309	8	24	shaded	shaded	ADJ
biotropia-309	8	25	and	and	CCONJ
biotropia-309	8	26	cooler	cooler	ADJ
biotropia-309	8	27	habitats	habitat	NOUN
biotropia-309	8	28	whereas	whereas	SCONJ
biotropia-309	8	29	and	and	CCONJ
biotropia-309	8	30	can	can	AUX
biotropia-309	8	31	be	be	AUX
biotropia-309	8	32	suggested	suggest	VERB
biotropia-309	8	33	as	as	ADP
biotropia-309	8	34	indicator	indicator	NOUN
biotropia-309	8	35	of	of	ADP
biotropia-309	8	36	unshaded	unshaded	ADJ
biotropia-309	8	37	and	and	CCONJ
biotropia-309	8	38	warmer	warm	ADJ
biotropia-309	8	39	habitats	habitat	NOUN
biotropia-309	8	40	(	(	PUNCT
biotropia-309	8	41	bare	bare	ADJ
biotropia-309	8	42	land	land	NOUN
biotropia-309	8	43	area	area	NOUN
biotropia-309	8	44	)	)	PUNCT
biotropia-309	8	45	.	.	PUNCT
biotropia-309	9	1	bioindicators	bioindicator	NOUN
biotropia-309	9	2	,	,	PUNCT
biotropia-309	9	3	diversity	diversity	NOUN
biotropia-309	9	4	,	,	PUNCT
biotropia-309	9	5	scarabaeidae	scarabaeidae	NOUN
biotropia-309	9	6	,	,	PUNCT
biotropia-309	9	7	habitat	habitat	NOUN
biotropia-309	9	8	preferences	preference	NOUN
biotropia-309	9	9	,	,	PUNCT
biotropia-309	9	10	.	.	PUNCT
biotropia-309	10	1	1	1	NUM
biotropia-309	10	2	*	*	SYM
biotropia-309	10	3	1	1	NUM
biotropia-309	10	4	2	2	NUM
biotropia-309	10	5	1	1	NUM
biotropia-309	10	6	2	2	NUM
biotropia-309	10	7	department	department	NOUN
biotropia-309	10	8	of	of	ADP
biotropia-309	10	9	agrotechnology	agrotechnology	NOUN
biotropia-309	10	10	faculty	faculty	NOUN
biotropia-309	10	11	of	of	ADP
biotropia-309	10	12	agriculture	agriculture	PROPN
biotropia-309	10	13	university	university	PROPN
biotropia-309	10	14	of	of	ADP
biotropia-309	10	15	tadulako	tadulako	PROPN
biotropia-309	10	16	,	,	PUNCT
biotropia-309	10	17	palu	palu	PROPN
biotropia-309	10	18	94118	94118	NUM
biotropia-309	10	19	,	,	PUNCT
biotropia-309	10	20	central	central	ADJ
biotropia-309	10	21	sulawesi	sulawesi	NOUN
biotropia-309	10	22	,	,	PUNCT
biotropia-309	10	23	indonesia	indonesia	PROPN
biotropia-309	10	24	department	department	PROPN
biotropia-309	10	25	of	of	ADP
biotropia-309	10	26	biology	biology	NOUN
biotropia-309	10	27	faculty	faculty	NOUN
biotropia-309	10	28	of	of	ADP
biotropia-309	10	29	mathematics	mathematic	NOUN
biotropia-309	10	30	and	and	CCONJ
biotropia-309	10	31	science	science	PROPN
biotropia-309	10	32	university	university	PROPN
biotropia-309	10	33	of	of	ADP
biotropia-309	10	34	tadulako	tadulako	PROPN
biotropia-309	10	35	,	,	PUNCT
biotropia-309	10	36	palu	palu	PROPN
biotropia-309	10	37	94118	94118	NUM
biotropia-309	10	38	,	,	PUNCT
biotropia-309	10	39	central	central	ADJ
biotropia-309	10	40	sulawesi	sulawesi	NOUN
biotropia-309	10	41	,	,	PUNCT
biotropia-309	10	42	indonesia	indonesia	PROPN
biotropia-309	10	43	indval	indval	PROPN
biotropia-309	10	44	copris	copris	PROPN
biotropia-309	10	45	saundersi	saundersi	PROPN
biotropia-309	10	46	onthophagus	onthophagus	VERB
biotropia-309	10	47	forsteni	forsteni	PROPN
biotropia-309	10	48	o.	o.	PROPN
biotropia-309	10	49	limbatus	limbatus	PROPN
biotropia-309	11	1	o.	o.	PROPN
biotropia-309	11	2	trituber	trituber	PROPN
biotropia-309	11	3	indval	indval	ADJ
biotropia-309	11	4	abstract	abstract	ADJ
biotropia-309	11	5	keywords	keyword	NOUN
biotropia-309	11	6	:	:	PUNCT
biotropia-309	11	7	*	*	PUNCT
biotropia-309	11	8	corresponding	correspond	VERB
biotropia-309	11	9	author	author	NOUN
biotropia-309	11	10	:	:	PUNCT
biotropia-309	11	11	shahabuddin_slh@yahoo.com	shahabuddin_slh@yahoo.com	X
biotropia-309	12	1	doi	doi	NOUN
biotropia-309	12	2	:	:	PUNCT
biotropia-309	12	3	10.11598	10.11598	NUM
biotropia-309	12	4	/	/	SYM
biotropia-309	12	5	btb.2014.21.1.5	btb.2014.21.1.5	NOUN
biotropia-309	12	6	48	48	NUM
biotropia-309	12	7	introduction	introduction	NOUN
biotropia-309	12	8	the	the	DET
biotropia-309	12	9	conversion	conversion	NOUN
biotropia-309	12	10	of	of	ADP
biotropia-309	12	11	natural	natural	ADJ
biotropia-309	12	12	habitat	habitat	NOUN
biotropia-309	12	13	to	to	ADP
biotropia-309	12	14	other	other	ADJ
biotropia-309	12	15	land	land	NOUN
biotropia-309	12	16	uses	use	NOUN
biotropia-309	12	17	and	and	CCONJ
biotropia-309	12	18	its	its	PRON
biotropia-309	12	19	consequences	consequence	NOUN
biotropia-309	12	20	on	on	ADP
biotropia-309	12	21	environmental	environmental	ADJ
biotropia-309	12	22	changes	change	NOUN
biotropia-309	12	23	has	have	AUX
biotropia-309	12	24	been	be	AUX
biotropia-309	12	25	acknowledged	acknowledge	VERB
biotropia-309	12	26	as	as	ADP
biotropia-309	12	27	the	the	DET
biotropia-309	12	28	main	main	ADJ
biotropia-309	12	29	driver	driver	NOUN
biotropia-309	12	30	of	of	ADP
biotropia-309	12	31	biodiversity	biodiversity	NOUN
biotropia-309	12	32	loss	loss	NOUN
biotropia-309	12	33	at	at	ADP
biotropia-309	12	34	the	the	DET
biotropia-309	12	35	southeast	southeast	PROPN
biotropia-309	12	36	asia	asia	PROPN
biotropia-309	12	37	and	and	CCONJ
biotropia-309	12	38	global	global	ADJ
biotropia-309	12	39	scales	scale	NOUN
biotropia-309	12	40	(	(	PUNCT
biotropia-309	12	41	sodhi	sodhi	NOUN
biotropia-309	12	42	.	.	PUNCT
biotropia-309	13	1	2004	2004	NUM
biotropia-309	13	2	)	)	PUNCT
biotropia-309	14	1	and	and	CCONJ
biotropia-309	14	2	by	by	ADP
biotropia-309	14	3	2100	2100	NUM
biotropia-309	14	4	the	the	DET
biotropia-309	14	5	impact	impact	NOUN
biotropia-309	14	6	of	of	ADP
biotropia-309	14	7	land	land	NOUN
biotropia-309	14	8	use	use	NOUN
biotropia-309	14	9	changes	change	NOUN
biotropia-309	14	10	on	on	ADP
biotropia-309	14	11	biodiversity	biodiversity	NOUN
biotropia-309	14	12	is	be	AUX
biotropia-309	14	13	likely	likely	ADJ
biotropia-309	14	14	to	to	PART
biotropia-309	14	15	be	be	AUX
biotropia-309	14	16	more	more	ADV
biotropia-309	14	17	significant	significant	ADJ
biotropia-309	14	18	than	than	ADP
biotropia-309	14	19	that	that	PRON
biotropia-309	14	20	of	of	ADP
biotropia-309	14	21	climate	climate	NOUN
biotropia-309	14	22	change	change	NOUN
biotropia-309	14	23	,	,	PUNCT
biotropia-309	14	24	nitrogen	nitrogen	NOUN
biotropia-309	14	25	deposition	deposition	NOUN
biotropia-309	14	26	,	,	PUNCT
biotropia-309	14	27	species	species	NOUN
biotropia-309	14	28	introductions	introduction	NOUN
biotropia-309	14	29	and	and	CCONJ
biotropia-309	14	30	changing	change	VERB
biotropia-309	14	31	atmospheric	atmospheric	ADJ
biotropia-309	14	32	concentrations	concentration	NOUN
biotropia-309	14	33	(	(	PUNCT
biotropia-309	14	34	sala	sala	PROPN
biotropia-309	14	35	2000	2000	NUM
biotropia-309	14	36	;	;	PUNCT
biotropia-309	14	37	young	young	ADJ
biotropia-309	14	38	2009	2009	NUM
biotropia-309	14	39	)	)	PUNCT
biotropia-309	14	40	.	.	PUNCT
biotropia-309	15	1	therefore	therefore	ADV
biotropia-309	15	2	,	,	PUNCT
biotropia-309	15	3	detecting	detect	VERB
biotropia-309	15	4	environmental	environmental	ADJ
biotropia-309	15	5	changes	change	NOUN
biotropia-309	15	6	due	due	ADP
biotropia-309	15	7	to	to	ADP
biotropia-309	15	8	land	land	NOUN
biotropia-309	15	9	-	-	PUNCT
biotropia-309	15	10	use	use	NOUN
biotropia-309	15	11	change	change	NOUN
biotropia-309	15	12	is	be	AUX
biotropia-309	15	13	needed	need	VERB
biotropia-309	15	14	in	in	ADP
biotropia-309	15	15	order	order	NOUN
biotropia-309	15	16	to	to	PART
biotropia-309	15	17	avoid	avoid	VERB
biotropia-309	15	18	the	the	DET
biotropia-309	15	19	continuous	continuous	ADJ
biotropia-309	15	20	loss	loss	NOUN
biotropia-309	15	21	of	of	ADP
biotropia-309	15	22	biodiversity	biodiversity	NOUN
biotropia-309	15	23	.	.	PUNCT
biotropia-309	16	1	bioindicators	bioindicator	NOUN
biotropia-309	16	2	have	have	AUX
biotropia-309	16	3	been	be	AUX
biotropia-309	16	4	proven	prove	VERB
biotropia-309	16	5	to	to	PART
biotropia-309	16	6	be	be	AUX
biotropia-309	16	7	useful	useful	ADJ
biotropia-309	16	8	tools	tool	NOUN
biotropia-309	16	9	for	for	ADP
biotropia-309	16	10	monitoring	monitor	VERB
biotropia-309	16	11	and	and	CCONJ
biotropia-309	16	12	detecting	detect	VERB
biotropia-309	16	13	changes	change	NOUN
biotropia-309	16	14	in	in	ADP
biotropia-309	16	15	the	the	DET
biotropia-309	16	16	environment	environment	NOUN
biotropia-309	16	17	.	.	PUNCT
biotropia-309	17	1	species	specie	NOUN
biotropia-309	17	2	have	have	VERB
biotropia-309	17	3	different	different	ADJ
biotropia-309	17	4	ecological	ecological	ADJ
biotropia-309	17	5	requirements	requirement	NOUN
biotropia-309	17	6	and	and	CCONJ
biotropia-309	17	7	their	their	PRON
biotropia-309	17	8	reactions	reaction	NOUN
biotropia-309	17	9	to	to	ADP
biotropia-309	17	10	environmental	environmental	ADJ
biotropia-309	17	11	variation	variation	NOUN
biotropia-309	17	12	are	be	AUX
biotropia-309	17	13	different	different	ADJ
biotropia-309	17	14	from	from	ADP
biotropia-309	17	15	one	one	NUM
biotropia-309	17	16	another	another	DET
biotropia-309	17	17	.	.	PUNCT
biotropia-309	18	1	therefore	therefore	ADV
biotropia-309	18	2	,	,	PUNCT
biotropia-309	18	3	some	some	DET
biotropia-309	18	4	species	specie	NOUN
biotropia-309	18	5	are	be	AUX
biotropia-309	18	6	better	well	ADJ
biotropia-309	18	7	indicators	indicator	NOUN
biotropia-309	18	8	than	than	ADP
biotropia-309	18	9	others	other	NOUN
biotropia-309	18	10	(	(	PUNCT
biotropia-309	18	11	dufrêne	dufrêne	PROPN
biotropia-309	18	12	&	&	CCONJ
biotropia-309	18	13	legendre	legendre	PROPN
biotropia-309	18	14	1997	1997	NUM
biotropia-309	18	15	)	)	PUNCT
biotropia-309	18	16	.	.	PUNCT
biotropia-309	19	1	some	some	DET
biotropia-309	19	2	species	specie	NOUN
biotropia-309	19	3	are	be	AUX
biotropia-309	19	4	generalists	generalist	NOUN
biotropia-309	19	5	occurring	occur	VERB
biotropia-309	19	6	in	in	ADP
biotropia-309	19	7	a	a	DET
biotropia-309	19	8	wide	wide	ADJ
biotropia-309	19	9	range	range	NOUN
biotropia-309	19	10	of	of	ADP
biotropia-309	19	11	habitats	habitat	NOUN
biotropia-309	19	12	(	(	PUNCT
biotropia-309	19	13	ubiquitous	ubiquitous	ADJ
biotropia-309	19	14	)	)	PUNCT
biotropia-309	19	15	,	,	PUNCT
biotropia-309	19	16	while	while	SCONJ
biotropia-309	19	17	others	other	NOUN
biotropia-309	19	18	are	be	AUX
biotropia-309	19	19	more	more	ADV
biotropia-309	19	20	specialized	specialized	ADJ
biotropia-309	19	21	,	,	PUNCT
biotropia-309	19	22	requiring	require	VERB
biotropia-309	19	23	certain	certain	ADJ
biotropia-309	19	24	habitat	habitat	NOUN
biotropia-309	19	25	characteristics	characteristic	NOUN
biotropia-309	19	26	(	(	PUNCT
biotropia-309	19	27	)	)	PUNCT
biotropia-309	19	28	.	.	PUNCT
biotropia-309	20	1	a	a	DET
biotropia-309	20	2	bioindicator	bioindicator	NOUN
biotropia-309	20	3	can	can	AUX
biotropia-309	20	4	be	be	AUX
biotropia-309	20	5	defined	define	VERB
biotropia-309	20	6	as	as	ADP
biotropia-309	20	7	a	a	DET
biotropia-309	20	8	species	specie	NOUN
biotropia-309	20	9	or	or	CCONJ
biotropia-309	20	10	a	a	DET
biotropia-309	20	11	species	species	NOUN
biotropia-309	20	12	group	group	NOUN
biotropia-309	20	13	that	that	PRON
biotropia-309	20	14	reflects	reflect	VERB
biotropia-309	20	15	the	the	DET
biotropia-309	20	16	abiotic	abiotic	ADJ
biotropia-309	20	17	or	or	CCONJ
biotropia-309	20	18	biotic	biotic	ADJ
biotropia-309	20	19	state	state	NOUN
biotropia-309	20	20	of	of	ADP
biotropia-309	20	21	the	the	DET
biotropia-309	20	22	environment	environment	NOUN
biotropia-309	20	23	(	(	PUNCT
biotropia-309	20	24	)	)	PUNCT
biotropia-309	20	25	,	,	PUNCT
biotropia-309	20	26	represents	represent	VERB
biotropia-309	20	27	the	the	DET
biotropia-309	20	28	impact	impact	NOUN
biotropia-309	20	29	of	of	ADP
biotropia-309	20	30	environmental	environmental	ADJ
biotropia-309	20	31	change	change	NOUN
biotropia-309	20	32	on	on	ADP
biotropia-309	20	33	a	a	DET
biotropia-309	20	34	habitat	habitat	NOUN
biotropia-309	20	35	,	,	PUNCT
biotropia-309	20	36	community	community	NOUN
biotropia-309	20	37	or	or	CCONJ
biotropia-309	20	38	ecosystems	ecosystem	NOUN
biotropia-309	20	39	(	(	PUNCT
biotropia-309	20	40	)	)	PUNCT
biotropia-309	20	41	,	,	PUNCT
biotropia-309	20	42	or	or	CCONJ
biotropia-309	20	43	indicates	indicate	VERB
biotropia-309	20	44	the	the	DET
biotropia-309	20	45	diversity	diversity	NOUN
biotropia-309	20	46	of	of	ADP
biotropia-309	20	47	other	other	ADJ
biotropia-309	20	48	species	specie	NOUN
biotropia-309	20	49	(	(	PUNCT
biotropia-309	20	50	)	)	PUNCT
biotropia-309	20	51	(	(	PUNCT
biotropia-309	20	52	mcgeoch	mcgeoch	NOUN
biotropia-309	20	53	1998	1998	NUM
biotropia-309	20	54	)	)	PUNCT
biotropia-309	20	55	.	.	PUNCT
biotropia-309	21	1	one	one	NUM
biotropia-309	21	2	of	of	ADP
biotropia-309	21	3	the	the	DET
biotropia-309	21	4	reasons	reason	NOUN
biotropia-309	21	5	for	for	ADP
biotropia-309	21	6	using	use	VERB
biotropia-309	21	7	bioindicators	bioindicator	NOUN
biotropia-309	21	8	is	be	AUX
biotropia-309	21	9	their	their	PRON
biotropia-309	21	10	cost	cost	NOUN
biotropia-309	21	11	-	-	PUNCT
biotropia-309	21	12	effectiveness	effectiveness	NOUN
biotropia-309	21	13	.	.	PUNCT
biotropia-309	22	1	by	by	ADP
biotropia-309	22	2	using	use	VERB
biotropia-309	22	3	bioindicators	bioindicator	NOUN
biotropia-309	22	4	it	it	PRON
biotropia-309	22	5	is	be	AUX
biotropia-309	22	6	possible	possible	ADJ
biotropia-309	22	7	to	to	PART
biotropia-309	22	8	assess	assess	VERB
biotropia-309	22	9	the	the	DET
biotropia-309	22	10	impact	impact	NOUN
biotropia-309	22	11	of	of	ADP
biotropia-309	22	12	human	human	ADJ
biotropia-309	22	13	activities	activity	NOUN
biotropia-309	22	14	on	on	ADP
biotropia-309	22	15	the	the	DET
biotropia-309	22	16	biota	biota	NOUN
biotropia-309	22	17	,	,	PUNCT
biotropia-309	22	18	instead	instead	ADV
biotropia-309	22	19	of	of	ADP
biotropia-309	22	20	examining	examine	VERB
biotropia-309	22	21	the	the	DET
biotropia-309	22	22	entire	entire	ADJ
biotropia-309	22	23	biota	biota	NOUN
biotropia-309	22	24	.	.	PUNCT
biotropia-309	23	1	especially	especially	ADV
biotropia-309	23	2	useful	useful	ADJ
biotropia-309	23	3	are	be	AUX
biotropia-309	23	4	species	specie	NOUN
biotropia-309	23	5	that	that	PRON
biotropia-309	23	6	provide	provide	VERB
biotropia-309	23	7	early	early	ADJ
biotropia-309	23	8	warning	warning	NOUN
biotropia-309	23	9	of	of	ADP
biotropia-309	23	10	change	change	NOUN
biotropia-309	23	11	(	(	PUNCT
biotropia-309	23	12	spellerberg	spellerberg	NOUN
biotropia-309	23	13	1993	1993	NUM
biotropia-309	23	14	)	)	PUNCT
biotropia-309	23	15	environmental	environmental	ADJ
biotropia-309	23	16	changes	change	NOUN
biotropia-309	23	17	can	can	AUX
biotropia-309	23	18	cause	cause	VERB
biotropia-309	23	19	different	different	ADJ
biotropia-309	23	20	kinds	kind	NOUN
biotropia-309	23	21	of	of	ADP
biotropia-309	23	22	effects	effect	NOUN
biotropia-309	23	23	in	in	ADP
biotropia-309	23	24	the	the	DET
biotropia-309	23	25	indicator	indicator	NOUN
biotropia-309	23	26	,	,	PUNCT
biotropia-309	23	27	including	include	VERB
biotropia-309	23	28	physiological	physiological	ADJ
biotropia-309	23	29	changes	change	NOUN
biotropia-309	23	30	or	or	CCONJ
biotropia-309	23	31	changes	change	NOUN
biotropia-309	23	32	in	in	ADP
biotropia-309	23	33	species	species	NOUN
biotropia-309	23	34	number	number	NOUN
biotropia-309	23	35	or	or	CCONJ
biotropia-309	23	36	abundance	abundance	NOUN
biotropia-309	23	37	.	.	PUNCT
biotropia-309	24	1	the	the	DET
biotropia-309	24	2	response	response	NOUN
biotropia-309	24	3	of	of	ADP
biotropia-309	24	4	the	the	DET
biotropia-309	24	5	species	specie	NOUN
biotropia-309	24	6	can	can	AUX
biotropia-309	24	7	be	be	AUX
biotropia-309	24	8	seen	see	VERB
biotropia-309	24	9	within	within	ADP
biotropia-309	24	10	the	the	DET
biotropia-309	24	11	organism	organism	NOUN
biotropia-309	24	12	(	(	PUNCT
biotropia-309	24	13	e.g.	e.g.	ADV
biotropia-309	24	14	heavy	heavy	ADJ
biotropia-309	24	15	-	-	PUNCT
biotropia-309	24	16	metal	metal	NOUN
biotropia-309	24	17	concentrations	concentration	NOUN
biotropia-309	24	18	)	)	PUNCT
biotropia-309	24	19	,	,	PUNCT
biotropia-309	24	20	at	at	ADP
biotropia-309	24	21	the	the	DET
biotropia-309	24	22	species	species	NOUN
biotropia-309	24	23	level	level	NOUN
biotropia-309	24	24	(	(	PUNCT
biotropia-309	24	25	species	species	NOUN
biotropia-309	24	26	number	number	NOUN
biotropia-309	24	27	and	and	CCONJ
biotropia-309	24	28	abundance	abundance	NOUN
biotropia-309	24	29	)	)	PUNCT
biotropia-309	24	30	or	or	CCONJ
biotropia-309	24	31	at	at	ADP
biotropia-309	24	32	the	the	DET
biotropia-309	24	33	community	community	NOUN
biotropia-309	24	34	level	level	NOUN
biotropia-309	24	35	(	(	PUNCT
biotropia-309	24	36	relations	relation	NOUN
biotropia-309	24	37	between	between	ADP
biotropia-309	24	38	species	specie	NOUN
biotropia-309	24	39	,	,	PUNCT
biotropia-309	24	40	e.g.	e.g.	ADV
biotropia-309	24	41	pestpredator	pestpredator	NOUN
biotropia-309	24	42	)	)	PUNCT
biotropia-309	24	43	.	.	PUNCT
biotropia-309	25	1	increase	increase	VERB
biotropia-309	25	2	or	or	CCONJ
biotropia-309	25	3	decrease	decrease	NOUN
biotropia-309	25	4	or	or	CCONJ
biotropia-309	25	5	abundance	abundance	NOUN
biotropia-309	25	6	of	of	ADP
biotropia-309	25	7	species	specie	NOUN
biotropia-309	25	8	number	number	NOUN
biotropia-309	25	9	might	might	AUX
biotropia-309	25	10	be	be	AUX
biotropia-309	25	11	directly	directly	ADV
biotropia-309	25	12	caused	cause	VERB
biotropia-309	25	13	by	by	ADP
biotropia-309	25	14	change	change	NOUN
biotropia-309	25	15	in	in	ADP
biotropia-309	25	16	abiotic	abiotic	ADJ
biotropia-309	25	17	and/or	and/or	CCONJ
biotropia-309	25	18	biotic	biotic	ADJ
biotropia-309	25	19	factors	factor	NOUN
biotropia-309	25	20	or	or	CCONJ
biotropia-309	25	21	indirectly	indirectly	ADV
biotropia-309	25	22	by	by	ADP
biotropia-309	25	23	change	change	NOUN
biotropia-309	25	24	of	of	ADP
biotropia-309	25	25	species	specie	NOUN
biotropia-309	25	26	assemblage	assemblage	NOUN
biotropia-309	25	27	of	of	ADP
biotropia-309	25	28	other	other	ADJ
biotropia-309	25	29	species	specie	NOUN
biotropia-309	25	30	(	(	PUNCT
biotropia-309	25	31	davis	davis	PROPN
biotropia-309	25	32	.	.	PUNCT
biotropia-309	26	1	2001	2001	NUM
biotropia-309	26	2	;	;	PUNCT
biotropia-309	26	3	rainio	rainio	PROPN
biotropia-309	26	4	&	&	CCONJ
biotropia-309	26	5	niemela	niemela	PROPN
biotropia-309	26	6	2003	2003	NUM
biotropia-309	26	7	;	;	PUNCT
biotropia-309	26	8	hambler	hambler	NOUN
biotropia-309	26	9	.	.	PUNCT
biotropia-309	27	1	2011	2011	NUM
biotropia-309	27	2	;	;	PUNCT
biotropia-309	27	3	gerlach	gerlach	PROPN
biotropia-309	27	4	.	.	PUNCT
biotropia-309	28	1	2013	2013	NUM
biotropia-309	28	2	)	)	PUNCT
biotropia-309	28	3	.	.	PUNCT
biotropia-309	29	1	a	a	DET
biotropia-309	29	2	good	good	ADJ
biotropia-309	29	3	bioindicator	bioindicator	NOUN
biotropia-309	29	4	must	must	AUX
biotropia-309	29	5	fulfill	fulfill	VERB
biotropia-309	29	6	several	several	ADJ
biotropia-309	29	7	criteria	criterion	NOUN
biotropia-309	29	8	.	.	PUNCT
biotropia-309	30	1	it	it	PRON
biotropia-309	30	2	has	have	VERB
biotropia-309	30	3	to	to	PART
biotropia-309	30	4	be	be	AUX
biotropia-309	30	5	well	well	ADV
biotropia-309	30	6	-	-	PUNCT
biotropia-309	30	7	known	know	VERB
biotropia-309	30	8	taxonomically	taxonomically	ADV
biotropia-309	30	9	and	and	CCONJ
biotropia-309	30	10	ecologically	ecologically	ADV
biotropia-309	30	11	,	,	PUNCT
biotropia-309	30	12	be	be	AUX
biotropia-309	30	13	distributed	distribute	VERB
biotropia-309	30	14	over	over	ADP
biotropia-309	30	15	a	a	DET
biotropia-309	30	16	broad	broad	ADJ
biotropia-309	30	17	geographic	geographic	ADJ
biotropia-309	30	18	area	area	NOUN
biotropia-309	30	19	,	,	PUNCT
biotropia-309	30	20	have	have	VERB
biotropia-309	30	21	specialization	specialization	NOUN
biotropia-309	30	22	to	to	ADP
biotropia-309	30	23	certain	certain	ADJ
biotropia-309	30	24	habitat	habitat	NOUN
biotropia-309	30	25	requirements	requirement	NOUN
biotropia-309	30	26	,	,	PUNCT
biotropia-309	30	27	provide	provide	VERB
biotropia-309	30	28	early	early	ADJ
biotropia-309	30	29	warning	warning	NOUN
biotropia-309	30	30	of	of	ADP
biotropia-309	30	31	change	change	NOUN
biotropia-309	30	32	,	,	PUNCT
biotropia-309	30	33	be	be	AUX
biotropia-309	30	34	easy	easy	ADJ
biotropia-309	30	35	and	and	CCONJ
biotropia-309	30	36	cost	cost	NOUN
biotropia-309	30	37	-	-	PUNCT
biotropia-309	30	38	effective	effective	ADJ
biotropia-309	30	39	to	to	PART
biotropia-309	30	40	survey	survey	VERB
biotropia-309	30	41	,	,	PUNCT
biotropia-309	30	42	be	be	AUX
biotropia-309	30	43	relatively	relatively	ADV
biotropia-309	30	44	independent	independent	ADJ
biotropia-309	30	45	of	of	ADP
biotropia-309	30	46	sample	sample	NOUN
biotropia-309	30	47	size	size	NOUN
biotropia-309	30	48	.	.	PUNCT
biotropia-309	31	1	its	its	PRON
biotropia-309	31	2	response	response	NOUN
biotropia-309	31	3	should	should	AUX
biotropia-309	31	4	reflect	reflect	VERB
biotropia-309	31	5	the	the	DET
biotropia-309	31	6	response	response	NOUN
biotropia-309	31	7	of	of	ADP
biotropia-309	31	8	other	other	ADJ
biotropia-309	31	9	species	specie	NOUN
biotropia-309	31	10	,	,	PUNCT
biotropia-309	31	11	one	one	PRON
biotropia-309	31	12	should	should	AUX
biotropia-309	31	13	be	be	AUX
biotropia-309	31	14	able	able	ADJ
biotropia-309	31	15	to	to	PART
biotropia-309	31	16	differentiate	differentiate	VERB
biotropia-309	31	17	between	between	ADP
biotropia-309	31	18	natural	natural	ADJ
biotropia-309	31	19	cycles	cycle	NOUN
biotropia-309	31	20	or	or	CCONJ
biotropia-309	31	21	trends	trend	NOUN
biotropia-309	31	22	and	and	CCONJ
biotropia-309	31	23	those	those	PRON
biotropia-309	31	24	induced	induce	VERB
biotropia-309	31	25	by	by	ADP
biotropia-309	31	26	anthropogenic	anthropogenic	ADJ
biotropia-309	31	27	stress	stress	NOUN
biotropia-309	31	28	,	,	PUNCT
biotropia-309	31	29	and	and	CCONJ
biotropia-309	31	30	it	it	PRON
biotropia-309	31	31	should	should	AUX
biotropia-309	31	32	be	be	AUX
biotropia-309	31	33	of	of	ADP
biotropia-309	31	34	potential	potential	ADJ
biotropia-309	31	35	economic	economic	ADJ
biotropia-309	31	36	importance	importance	NOUN
biotropia-309	31	37	(	(	PUNCT
biotropia-309	31	38	e.g.	e.g.	ADV
biotropia-309	31	39	noss	noss	PROPN
biotropia-309	31	40	1990	1990	NUM
biotropia-309	31	41	;	;	PUNCT
biotropia-309	31	42	pearson	pearson	PROPN
biotropia-309	31	43	&	&	CCONJ
biotropia-309	31	44	cassola	cassola	PROPN
biotropia-309	31	45	1992	1992	NUM
biotropia-309	31	46	)	)	PUNCT
biotropia-309	31	47	.	.	PUNCT
biotropia-309	32	1	however	however	ADV
biotropia-309	32	2	,	,	PUNCT
biotropia-309	32	3	it	it	PRON
biotropia-309	32	4	is	be	AUX
biotropia-309	32	5	difficult	difficult	ADJ
biotropia-309	32	6	to	to	PART
biotropia-309	32	7	find	find	VERB
biotropia-309	32	8	species	specie	NOUN
biotropia-309	32	9	or	or	CCONJ
biotropia-309	32	10	species	species	NOUN
biotropia-309	32	11	groups	group	NOUN
biotropia-309	32	12	which	which	PRON
biotropia-309	32	13	would	would	AUX
biotropia-309	32	14	have	have	VERB
biotropia-309	32	15	all	all	PRON
biotropia-309	32	16	of	of	ADP
biotropia-309	32	17	these	these	DET
biotropia-309	32	18	criteria	criterion	NOUN
biotropia-309	32	19	(	(	PUNCT
biotropia-309	32	20	noss	noss	PROPN
biotropia-309	32	21	1990	1990	NUM
biotropia-309	32	22	;	;	PUNCT
biotropia-309	32	23	pearson	pearson	PROPN
biotropia-309	32	24	&	&	CCONJ
biotropia-309	32	25	cassola	cassola	PROPN
biotropia-309	32	26	1992	1992	NUM
biotropia-309	32	27	)	)	PUNCT
biotropia-309	32	28	.	.	PUNCT
biotropia-309	33	1	requirements	requirement	NOUN
biotropia-309	33	2	needed	need	VERB
biotropia-309	33	3	depend	depend	VERB
biotropia-309	33	4	on	on	ADP
biotropia-309	33	5	the	the	DET
biotropia-309	33	6	goal	goal	NOUN
biotropia-309	33	7	of	of	ADP
biotropia-309	33	8	the	the	DET
biotropia-309	33	9	survey	survey	NOUN
biotropia-309	33	10	and	and	CCONJ
biotropia-309	33	11	the	the	DET
biotropia-309	33	12	sensitivity	sensitivity	NOUN
biotropia-309	33	13	to	to	ADP
biotropia-309	33	14	the	the	DET
biotropia-309	33	15	anthropogenic	anthropogenic	ADJ
biotropia-309	33	16	disturbance	disturbance	NOUN
biotropia-309	33	17	is	be	AUX
biotropia-309	33	18	the	the	DET
biotropia-309	33	19	most	most	ADV
biotropia-309	33	20	important	important	ADJ
biotropia-309	33	21	criteria	criterion	NOUN
biotropia-309	33	22	for	for	ADP
biotropia-309	33	23	monitoring	monitor	VERB
biotropia-309	33	24	environmental	environmental	ADJ
biotropia-309	33	25	changes	change	NOUN
biotropia-309	33	26	(	(	PUNCT
biotropia-309	33	27	kremen	kreman	NOUN
biotropia-309	33	28	.	.	PUNCT
biotropia-309	34	1	1993	1993	NUM
biotropia-309	34	2	)	)	PUNCT
biotropia-309	34	3	.	.	PUNCT
biotropia-309	35	1	et	et	PROPN
biotropia-309	35	2	al	al	PROPN
biotropia-309	35	3	et	et	PROPN
biotropia-309	35	4	al	al	PROPN
biotropia-309	35	5	stenotopic	stenotopic	PROPN
biotropia-309	35	6	environmental	environmental	PROPN
biotropia-309	35	7	indicator	indicator	PROPN
biotropia-309	35	8	ecological	ecological	ADJ
biotropia-309	35	9	indicator	indicator	NOUN
biotropia-309	35	10	biodiversity	biodiversity	NOUN
biotropia-309	35	11	indicator	indicator	NOUN
biotropia-309	35	12	.	.	PUNCT
biotropia-309	36	1	et	et	PROPN
biotropia-309	36	2	al	al	PROPN
biotropia-309	36	3	et	et	PROPN
biotropia-309	36	4	al	al	PROPN
biotropia-309	36	5	et	et	PROPN
biotropia-309	36	6	al	al	PROPN
biotropia-309	36	7	et	et	PROPN
biotropia-309	36	8	al	al	PROPN
biotropia-309	36	9	.	.	PUNCT
biotropia-309	37	1	49	49	NUM
biotropia-309	37	2	effectiveness	effectiveness	NOUN
biotropia-309	37	3	of	of	ADP
biotropia-309	37	4	dung	dung	NOUN
biotropia-309	37	5	beetles	beetle	NOUN
biotropia-309	37	6	as	as	ADP
biotropia-309	37	7	bioindicators	bioindicator	NOUN
biotropia-309	37	8	of	of	ADP
biotropia-309	37	9	environmental	environmental	ADJ
biotropia-309	37	10	changes	change	NOUN
biotropia-309	37	11	in	in	ADP
biotropia-309	37	12	land	land	NOUN
biotropia-309	37	13	shahabuddin	shahabuddin	NOUN
biotropia-309	37	14	–	–	PUNCT
biotropia-309	37	15	et	et	PROPN
biotropia-309	37	16	al	al	PROPN
biotropia-309	37	17	.	.	PUNCT
biotropia-309	38	1	in	in	ADP
biotropia-309	38	2	this	this	DET
biotropia-309	38	3	study	study	NOUN
biotropia-309	38	4	dung	dung	NOUN
biotropia-309	38	5	beetles	beetle	NOUN
biotropia-309	38	6	(	(	PUNCT
biotropia-309	38	7	coleoptera	coleoptera	NOUN
biotropia-309	38	8	:	:	PUNCT
biotropia-309	38	9	scarabaeidae	scarabaeidae	NOUN
biotropia-309	38	10	)	)	PUNCT
biotropia-309	38	11	was	be	AUX
biotropia-309	38	12	selected	select	VERB
biotropia-309	38	13	as	as	ADP
biotropia-309	38	14	a	a	DET
biotropia-309	38	15	bioindicator	bioindicator	NOUN
biotropia-309	38	16	group	group	NOUN
biotropia-309	38	17	because	because	SCONJ
biotropia-309	38	18	it	it	PRON
biotropia-309	38	19	has	have	AUX
biotropia-309	38	20	been	be	AUX
biotropia-309	38	21	widely	widely	ADV
biotropia-309	38	22	known	know	VERB
biotropia-309	38	23	as	as	ADP
biotropia-309	38	24	one	one	NUM
biotropia-309	38	25	of	of	ADP
biotropia-309	38	26	the	the	DET
biotropia-309	38	27	best	good	ADJ
biotropia-309	38	28	bioindicator	bioindicator	NOUN
biotropia-309	38	29	groups	group	NOUN
biotropia-309	38	30	(	(	PUNCT
biotropia-309	38	31	e.g.	e.g.	ADV
biotropia-309	38	32	halffter	halffter	NOUN
biotropia-309	38	33	&	&	CCONJ
biotropia-309	38	34	favila	favila	PROPN
biotropia-309	38	35	1993	1993	NUM
biotropia-309	38	36	;	;	PUNCT
biotropia-309	38	37	mcgeoch	mcgeoch	PROPN
biotropia-309	38	38	2002	2002	NUM
biotropia-309	38	39	)	)	PUNCT
biotropia-309	38	40	.	.	PUNCT
biotropia-309	39	1	dung	dung	NOUN
biotropia-309	39	2	beetles	beetle	NOUN
biotropia-309	39	3	have	have	AUX
biotropia-309	39	4	been	be	AUX
biotropia-309	39	5	proven	prove	VERB
biotropia-309	39	6	to	to	PART
biotropia-309	39	7	be	be	AUX
biotropia-309	39	8	very	very	ADV
biotropia-309	39	9	suitable	suitable	ADJ
biotropia-309	39	10	to	to	PART
biotropia-309	39	11	assess	assess	VERB
biotropia-309	39	12	effects	effect	NOUN
biotropia-309	39	13	of	of	ADP
biotropia-309	39	14	disturbances	disturbance	NOUN
biotropia-309	39	15	on	on	ADP
biotropia-309	39	16	tropical	tropical	ADJ
biotropia-309	39	17	ecosystems	ecosystem	NOUN
biotropia-309	39	18	(	(	PUNCT
biotropia-309	39	19	nichols	nichols	PROPN
biotropia-309	39	20	.	.	PUNCT
biotropia-309	39	21	2007	2007	NUM
biotropia-309	39	22	;	;	PUNCT
biotropia-309	39	23	shahabuddin	shahabuddin	VERB
biotropia-309	39	24	2010	2010	NUM
biotropia-309	39	25	)	)	PUNCT
biotropia-309	39	26	and	and	CCONJ
biotropia-309	39	27	human	human	ADJ
biotropia-309	39	28	habitat	habitat	PROPN
biotropia-309	39	29	modification	modification	NOUN
biotropia-309	39	30	(	(	PUNCT
biotropia-309	39	31	shahabuddin	shahabuddin	VERB
biotropia-309	39	32	2010	2010	NUM
biotropia-309	39	33	;	;	PUNCT
biotropia-309	39	34	harvey	harvey	NOUN
biotropia-309	39	35	2006	2006	NUM
biotropia-309	39	36	)	)	PUNCT
biotropia-309	39	37	due	due	ADP
biotropia-309	39	38	to	to	ADP
biotropia-309	39	39	their	their	PRON
biotropia-309	39	40	abundance	abundance	NOUN
biotropia-309	39	41	,	,	PUNCT
biotropia-309	39	42	highly	highly	ADV
biotropia-309	39	43	varied	varied	ADJ
biotropia-309	39	44	with	with	ADP
biotropia-309	39	45	respect	respect	NOUN
biotropia-309	39	46	to	to	ADP
biotropia-309	39	47	species	species	NOUN
biotropia-309	39	48	traits	trait	NOUN
biotropia-309	39	49	,	,	PUNCT
biotropia-309	39	50	and	and	CCONJ
biotropia-309	39	51	rapid	rapid	ADJ
biotropia-309	39	52	responses	response	NOUN
biotropia-309	39	53	to	to	ADP
biotropia-309	39	54	environmental	environmental	ADJ
biotropia-309	39	55	change	change	NOUN
biotropia-309	39	56	(	(	PUNCT
biotropia-309	39	57	slade	slade	PROPN
biotropia-309	39	58	.	.	PUNCT
biotropia-309	40	1	2011	2011	NUM
biotropia-309	40	2	;	;	PUNCT
biotropia-309	41	1	shahabuddin	shahabuddin	VERB
biotropia-309	41	2	.	.	PUNCT
biotropia-309	42	1	2005	2005	NUM
biotropia-309	42	2	&	&	CCONJ
biotropia-309	42	3	2010	2010	NUM
biotropia-309	42	4	;	;	PUNCT
biotropia-309	42	5	spector	spector	PROPN
biotropia-309	42	6	2006	2006	NUM
biotropia-309	42	7	;	;	PUNCT
biotropia-309	42	8	slade	slade	PROPN
biotropia-309	42	9	.	.	PUNCT
biotropia-309	43	1	2011	2011	NUM
biotropia-309	43	2	)	)	PUNCT
biotropia-309	43	3	.	.	PUNCT
biotropia-309	44	1	another	another	DET
biotropia-309	44	2	advantage	advantage	NOUN
biotropia-309	44	3	is	be	AUX
biotropia-309	44	4	their	their	PRON
biotropia-309	44	5	relatively	relatively	ADV
biotropia-309	44	6	complete	complete	ADJ
biotropia-309	44	7	species	specie	NOUN
biotropia-309	44	8	inventories	inventory	NOUN
biotropia-309	44	9	and	and	CCONJ
biotropia-309	44	10	their	their	PRON
biotropia-309	44	11	data	datum	NOUN
biotropia-309	44	12	on	on	ADP
biotropia-309	44	13	the	the	DET
biotropia-309	44	14	abundance	abundance	NOUN
biotropia-309	44	15	of	of	ADP
biotropia-309	44	16	individual	individual	ADJ
biotropia-309	44	17	species	specie	NOUN
biotropia-309	44	18	can	can	AUX
biotropia-309	44	19	be	be	AUX
biotropia-309	44	20	achieved	achieve	VERB
biotropia-309	44	21	rapidly	rapidly	ADV
biotropia-309	44	22	with	with	ADP
biotropia-309	44	23	standardized	standardized	ADJ
biotropia-309	44	24	methods	method	NOUN
biotropia-309	44	25	(	(	PUNCT
biotropia-309	44	26	larsen	larsen	PROPN
biotropia-309	44	27	&	&	CCONJ
biotropia-309	44	28	forsyth	forsyth	PROPN
biotropia-309	44	29	,	,	PUNCT
biotropia-309	44	30	2005	2005	NUM
biotropia-309	44	31	)	)	PUNCT
biotropia-309	44	32	.	.	PUNCT
biotropia-309	45	1	recently	recently	ADV
biotropia-309	45	2	,	,	PUNCT
biotropia-309	45	3	dung	dung	NOUN
biotropia-309	45	4	beetles	beetle	NOUN
biotropia-309	45	5	have	have	AUX
biotropia-309	45	6	been	be	AUX
biotropia-309	45	7	identified	identify	VERB
biotropia-309	45	8	as	as	ADP
biotropia-309	45	9	one	one	NUM
biotropia-309	45	10	of	of	ADP
biotropia-309	45	11	the	the	DET
biotropia-309	45	12	most	most	ADV
biotropia-309	45	13	cost	cost	NOUN
biotropia-309	45	14	-	-	PUNCT
biotropia-309	45	15	effective	effective	ADJ
biotropia-309	45	16	group	group	NOUN
biotropia-309	45	17	for	for	ADP
biotropia-309	45	18	biodiversity	biodiversity	NOUN
biotropia-309	45	19	survey	survey	NOUN
biotropia-309	45	20	in	in	ADP
biotropia-309	45	21	tropical	tropical	ADJ
biotropia-309	45	22	forests	forest	NOUN
biotropia-309	45	23	(	(	PUNCT
biotropia-309	45	24	kessler	kessler	PROPN
biotropia-309	45	25	.	.	PUNCT
biotropia-309	46	1	2011	2011	NUM
biotropia-309	46	2	)	)	PUNCT
biotropia-309	47	1	and	and	CCONJ
biotropia-309	47	2	contributors	contributor	NOUN
biotropia-309	47	3	in	in	ADP
biotropia-309	47	4	improving	improve	VERB
biotropia-309	47	5	the	the	DET
biotropia-309	47	6	level	level	NOUN
biotropia-309	47	7	of	of	ADP
biotropia-309	47	8	soil	soil	NOUN
biotropia-309	47	9	carbon	carbon	NOUN
biotropia-309	47	10	stocks	stock	NOUN
biotropia-309	47	11	(	(	PUNCT
biotropia-309	47	12	kessler	kessler	PROPN
biotropia-309	47	13	.	.	PUNCT
biotropia-309	47	14	2012	2012	NUM
biotropia-309	47	15	)	)	PUNCT
biotropia-309	47	16	.	.	PUNCT
biotropia-309	48	1	one	one	NUM
biotropia-309	48	2	method	method	NOUN
biotropia-309	48	3	used	use	VERB
biotropia-309	48	4	to	to	PART
biotropia-309	48	5	quantify	quantify	VERB
biotropia-309	48	6	the	the	DET
biotropia-309	48	7	'	'	PUNCT
biotropia-309	48	8	bioindicator	bioindicator	NOUN
biotropia-309	48	9	value	value	NOUN
biotropia-309	48	10	'	'	PUNCT
biotropia-309	48	11	of	of	ADP
biotropia-309	48	12	a	a	DET
biotropia-309	48	13	range	range	NOUN
biotropia-309	48	14	of	of	ADP
biotropia-309	48	15	taxa	taxa	NOUN
biotropia-309	48	16	is	be	AUX
biotropia-309	48	17	the	the	DET
biotropia-309	48	18	indicator	indicator	NOUN
biotropia-309	48	19	value	value	NOUN
biotropia-309	48	20	(	(	PUNCT
biotropia-309	48	21	)	)	PUNCT
biotropia-309	48	22	method	method	NOUN
biotropia-309	48	23	developed	develop	VERB
biotropia-309	48	24	by	by	ADP
biotropia-309	48	25	dufrene	dufrene	NOUN
biotropia-309	48	26	and	and	CCONJ
biotropia-309	48	27	legendre	legendre	PROPN
biotropia-309	48	28	(	(	PUNCT
biotropia-309	48	29	1997	1997	NUM
biotropia-309	48	30	)	)	PUNCT
biotropia-309	48	31	.	.	PUNCT
biotropia-309	49	1	this	this	DET
biotropia-309	49	2	method	method	NOUN
biotropia-309	49	3	combines	combine	VERB
biotropia-309	49	4	measurements	measurement	NOUN
biotropia-309	49	5	of	of	ADP
biotropia-309	49	6	the	the	DET
biotropia-309	49	7	degree	degree	NOUN
biotropia-309	49	8	of	of	ADP
biotropia-309	49	9	specificity	specificity	NOUN
biotropia-309	49	10	of	of	ADP
biotropia-309	49	11	a	a	DET
biotropia-309	49	12	species	species	NOUN
biotropia-309	49	13	to	to	ADP
biotropia-309	49	14	an	an	DET
biotropia-309	49	15	ecological	ecological	ADJ
biotropia-309	49	16	state	state	NOUN
biotropia-309	49	17	,	,	PUNCT
biotropia-309	49	18	for	for	ADP
biotropia-309	49	19	example	example	NOUN
biotropia-309	49	20	a	a	DET
biotropia-309	49	21	habitat	habitat	NOUN
biotropia-309	49	22	type	type	NOUN
biotropia-309	49	23	,	,	PUNCT
biotropia-309	49	24	and	and	CCONJ
biotropia-309	49	25	its	its	PRON
biotropia-309	49	26	fidelity	fidelity	NOUN
biotropia-309	49	27	within	within	ADP
biotropia-309	49	28	that	that	DET
biotropia-309	49	29	state	state	NOUN
biotropia-309	49	30	(	(	PUNCT
biotropia-309	49	31	dufrene	dufrene	PROPN
biotropia-309	49	32	&	&	CCONJ
biotropia-309	49	33	legendre	legendre	PROPN
biotropia-309	49	34	1997	1997	NUM
biotropia-309	49	35	)	)	PUNCT
biotropia-309	49	36	.	.	PUNCT
biotropia-309	50	1	species	specie	NOUN
biotropia-309	50	2	with	with	ADP
biotropia-309	50	3	a	a	DET
biotropia-309	50	4	high	high	ADJ
biotropia-309	50	5	specificity	specificity	NOUN
biotropia-309	50	6	and	and	CCONJ
biotropia-309	50	7	high	high	ADJ
biotropia-309	50	8	fidelity	fidelity	NOUN
biotropia-309	50	9	within	within	ADP
biotropia-309	50	10	a	a	DET
biotropia-309	50	11	habitat	habitat	NOUN
biotropia-309	50	12	will	will	AUX
biotropia-309	50	13	have	have	VERB
biotropia-309	50	14	a	a	DET
biotropia-309	50	15	high	high	ADJ
biotropia-309	50	16	indicator	indicator	NOUN
biotropia-309	50	17	value	value	NOUN
biotropia-309	50	18	.	.	PUNCT
biotropia-309	51	1	high	high	ADJ
biotropia-309	51	2	fidelity	fidelity	PROPN
biotropia-309	51	3	(	(	PUNCT
biotropia-309	51	4	frequency	frequency	NOUN
biotropia-309	51	5	of	of	ADP
biotropia-309	51	6	occurrence	occurrence	NOUN
biotropia-309	51	7	)	)	PUNCT
biotropia-309	51	8	of	of	ADP
biotropia-309	51	9	a	a	DET
biotropia-309	51	10	species	specie	NOUN
biotropia-309	51	11	across	across	ADP
biotropia-309	51	12	sample	sample	NOUN
biotropia-309	51	13	sites	site	NOUN
biotropia-309	51	14	is	be	AUX
biotropia-309	51	15	generally	generally	ADV
biotropia-309	51	16	associated	associate	VERB
biotropia-309	51	17	with	with	ADP
biotropia-309	51	18	large	large	ADJ
biotropia-309	51	19	abundance	abundance	NOUN
biotropia-309	51	20	of	of	ADP
biotropia-309	51	21	individuals	individual	NOUN
biotropia-309	51	22	.	.	PUNCT
biotropia-309	52	1	both	both	DET
biotropia-309	52	2	characteristics	characteristic	NOUN
biotropia-309	52	3	facilitate	facilitate	NOUN
biotropia-309	52	4	sampling	sample	VERB
biotropia-309	52	5	and	and	CCONJ
biotropia-309	52	6	monitoring	monitoring	NOUN
biotropia-309	52	7	,	,	PUNCT
biotropia-309	52	8	which	which	PRON
biotropia-309	52	9	are	be	AUX
biotropia-309	52	10	important	important	ADJ
biotropia-309	52	11	requirements	requirement	NOUN
biotropia-309	52	12	for	for	ADP
biotropia-309	52	13	a	a	DET
biotropia-309	52	14	useful	useful	ADJ
biotropia-309	52	15	bioindicator	bioindicator	NOUN
biotropia-309	52	16	(	(	PUNCT
biotropia-309	52	17	kremen	kreman	NOUN
biotropia-309	52	18	.	.	PUNCT
biotropia-309	53	1	1994	1994	NUM
biotropia-309	53	2	;	;	PUNCT
biotropia-309	54	1	mcgeoch	mcgeoch	PROPN
biotropia-309	54	2	.	.	PUNCT
biotropia-309	55	1	2002	2002	NUM
biotropia-309	55	2	)	)	PUNCT
biotropia-309	55	3	.	.	PUNCT
biotropia-309	56	1	the	the	DET
biotropia-309	56	2	indicator	indicator	NOUN
biotropia-309	56	3	value	value	NOUN
biotropia-309	56	4	method	method	NOUN
biotropia-309	56	5	is	be	AUX
biotropia-309	56	6	important	important	ADJ
biotropia-309	56	7	to	to	ADP
biotropia-309	56	8	conservation	conservation	NOUN
biotropia-309	56	9	biology	biology	NOUN
biotropia-309	56	10	because	because	SCONJ
biotropia-309	56	11	it	it	PRON
biotropia-309	56	12	is	be	AUX
biotropia-309	56	13	conceptually	conceptually	ADV
biotropia-309	56	14	straightforward	straightforward	ADJ
biotropia-309	56	15	and	and	CCONJ
biotropia-309	56	16	allows	allow	VERB
biotropia-309	56	17	researchers	researcher	NOUN
biotropia-309	56	18	to	to	PART
biotropia-309	56	19	identify	identify	VERB
biotropia-309	56	20	bioindicators	bioindicator	NOUN
biotropia-309	56	21	for	for	ADP
biotropia-309	56	22	any	any	DET
biotropia-309	56	23	combination	combination	NOUN
biotropia-309	56	24	of	of	ADP
biotropia-309	56	25	habitat	habitat	ADJ
biotropia-309	56	26	types	type	NOUN
biotropia-309	56	27	or	or	CCONJ
biotropia-309	56	28	areas	area	NOUN
biotropia-309	56	29	of	of	ADP
biotropia-309	56	30	interest	interest	NOUN
biotropia-309	56	31	,	,	PUNCT
biotropia-309	56	32	e.g.	e.g.	ADV
biotropia-309	56	33	existing	exist	VERB
biotropia-309	56	34	conservation	conservation	NOUN
biotropia-309	56	35	areas	area	NOUN
biotropia-309	56	36	,	,	PUNCT
biotropia-309	56	37	or	or	CCONJ
biotropia-309	56	38	groups	group	NOUN
biotropia-309	56	39	of	of	ADP
biotropia-309	56	40	sites	site	NOUN
biotropia-309	56	41	based	base	VERB
biotropia-309	56	42	on	on	ADP
biotropia-309	56	43	the	the	DET
biotropia-309	56	44	outcome	outcome	NOUN
biotropia-309	56	45	of	of	ADP
biotropia-309	56	46	a	a	DET
biotropia-309	56	47	classification	classification	NOUN
biotropia-309	56	48	procedure	procedure	NOUN
biotropia-309	56	49	(	(	PUNCT
biotropia-309	56	50	mcgeoch	mcgeoch	PROPN
biotropia-309	56	51	&	&	CCONJ
biotropia-309	56	52	chown	chown	ADJ
biotropia-309	56	53	1998	1998	NUM
biotropia-309	56	54	)	)	PUNCT
biotropia-309	56	55	.	.	PUNCT
biotropia-309	57	1	accordingly	accordingly	ADV
biotropia-309	57	2	,	,	PUNCT
biotropia-309	57	3	method	method	NOUN
biotropia-309	57	4	has	have	AUX
biotropia-309	57	5	become	become	VERB
biotropia-309	57	6	the	the	DET
biotropia-309	57	7	most	most	ADV
biotropia-309	57	8	robust	robust	ADJ
biotropia-309	57	9	and	and	CCONJ
biotropia-309	57	10	popular	popular	ADJ
biotropia-309	57	11	method	method	NOUN
biotropia-309	57	12	used	use	VERB
biotropia-309	57	13	to	to	PART
biotropia-309	57	14	measure	measure	VERB
biotropia-309	57	15	indicator	indicator	NOUN
biotropia-309	57	16	species	species	NOUN
biotropia-309	57	17	analysis	analysis	NOUN
biotropia-309	57	18	(	(	PUNCT
biotropia-309	57	19	mc.geoch	mc.geoch	NOUN
biotropia-309	57	20	.	.	PUNCT
biotropia-309	58	1	2002	2002	NUM
biotropia-309	58	2	;	;	PUNCT
biotropia-309	58	3	aydin	aydin	NOUN
biotropia-309	58	4	&	&	CCONJ
biotropia-309	58	5	kazak	kazak	PROPN
biotropia-309	58	6	2010	2010	NUM
biotropia-309	58	7	;	;	PUNCT
biotropia-309	58	8	negro	negro	PROPN
biotropia-309	58	9	.	.	PUNCT
biotropia-309	59	1	2011	2011	NUM
biotropia-309	59	2	)	)	PUNCT
biotropia-309	59	3	.	.	PUNCT
biotropia-309	60	1	in	in	ADP
biotropia-309	60	2	central	central	ADJ
biotropia-309	60	3	sulawesi	sulawesi	NOUN
biotropia-309	60	4	,	,	PUNCT
biotropia-309	60	5	forest	forest	NOUN
biotropia-309	60	6	habitats	habitat	NOUN
biotropia-309	60	7	especially	especially	ADV
biotropia-309	60	8	in	in	ADP
biotropia-309	60	9	the	the	DET
biotropia-309	60	10	interior	interior	NOUN
biotropia-309	60	11	of	of	ADP
biotropia-309	60	12	lore	lore	PROPN
biotropia-309	60	13	lindu	lindu	PROPN
biotropia-309	60	14	national	national	PROPN
biotropia-309	60	15	park	park	NOUN
biotropia-309	60	16	(	(	PUNCT
biotropia-309	60	17	llnp	llnp	PROPN
biotropia-309	60	18	)	)	PUNCT
biotropia-309	60	19	are	be	AUX
biotropia-309	60	20	still	still	ADV
biotropia-309	60	21	relatively	relatively	ADV
biotropia-309	60	22	undisturbed	undisturbed	ADJ
biotropia-309	60	23	while	while	SCONJ
biotropia-309	60	24	the	the	DET
biotropia-309	60	25	margins	margin	NOUN
biotropia-309	60	26	of	of	ADP
biotropia-309	60	27	the	the	DET
biotropia-309	60	28	park	park	NOUN
biotropia-309	60	29	are	be	AUX
biotropia-309	60	30	characterized	characterize	VERB
biotropia-309	60	31	by	by	ADP
biotropia-309	60	32	a	a	DET
biotropia-309	60	33	mosaic	mosaic	ADJ
biotropia-309	60	34	land	land	NOUN
biotropia-309	60	35	-	-	PUNCT
biotropia-309	60	36	use	use	NOUN
biotropia-309	60	37	type	type	NOUN
biotropia-309	60	38	such	such	ADJ
biotropia-309	60	39	as	as	ADP
biotropia-309	60	40	near	near	ADP
biotropia-309	60	41	-	-	PUNCT
biotropia-309	60	42	primary	primary	ADJ
biotropia-309	60	43	forests	forest	NOUN
biotropia-309	60	44	,	,	PUNCT
biotropia-309	60	45	secondary	secondary	ADJ
biotropia-309	60	46	forests	forest	NOUN
biotropia-309	60	47	,	,	PUNCT
biotropia-309	60	48	forest	forest	NOUN
biotropia-309	60	49	gardens	garden	NOUN
biotropia-309	60	50	and	and	CCONJ
biotropia-309	60	51	plantations	plantation	NOUN
biotropia-309	60	52	of	of	ADP
biotropia-309	60	53	cacao	cacao	NOUN
biotropia-309	60	54	,	,	PUNCT
biotropia-309	60	55	maize	maize	NOUN
biotropia-309	60	56	and	and	CCONJ
biotropia-309	60	57	paddy	paddy	NOUN
biotropia-309	60	58	rice	rice	NOUN
biotropia-309	60	59	fields	field	NOUN
biotropia-309	60	60	(	(	PUNCT
biotropia-309	60	61	gerold	gerold	VERB
biotropia-309	60	62	.	.	PUNCT
biotropia-309	61	1	2004	2004	NUM
biotropia-309	61	2	)	)	PUNCT
biotropia-309	61	3	.	.	PUNCT
biotropia-309	62	1	this	this	DET
biotropia-309	62	2	study	study	NOUN
biotropia-309	62	3	aimed	aim	VERB
biotropia-309	62	4	to	to	PART
biotropia-309	62	5	evaluate	evaluate	VERB
biotropia-309	62	6	the	the	DET
biotropia-309	62	7	effectiveness	effectiveness	NOUN
biotropia-309	62	8	of	of	ADP
biotropia-309	62	9	dung	dung	NOUN
biotropia-309	62	10	beetles	beetle	NOUN
biotropia-309	62	11	as	as	ADP
biotropia-309	62	12	bioindicator	bioindicator	NOUN
biotropia-309	62	13	of	of	ADP
biotropia-309	62	14	environmental	environmental	ADJ
biotropia-309	62	15	changes	change	NOUN
biotropia-309	62	16	across	across	ADP
biotropia-309	62	17	land	land	NOUN
biotropia-309	62	18	-	-	PUNCT
biotropia-309	62	19	use	use	NOUN
biotropia-309	62	20	gradient	gradient	NOUN
biotropia-309	62	21	in	in	ADP
biotropia-309	62	22	the	the	DET
biotropia-309	62	23	margins	margin	NOUN
biotropia-309	62	24	of	of	ADP
biotropia-309	62	25	llnp	llnp	PROPN
biotropia-309	62	26	.	.	PUNCT
biotropia-309	63	1	this	this	DET
biotropia-309	63	2	study	study	NOUN
biotropia-309	63	3	was	be	AUX
biotropia-309	63	4	carried	carry	VERB
biotropia-309	63	5	out	out	ADP
biotropia-309	63	6	at	at	ADP
biotropia-309	63	7	the	the	DET
biotropia-309	63	8	northern	northern	ADJ
biotropia-309	63	9	margin	margin	NOUN
biotropia-309	63	10	of	of	ADP
biotropia-309	63	11	the	the	DET
biotropia-309	63	12	llnp	llnp	NOUN
biotropia-309	63	13	in	in	ADP
biotropia-309	63	14	central	central	ADJ
biotropia-309	63	15	sulawesi	sulawesi	NOUN
biotropia-309	63	16	,	,	PUNCT
biotropia-309	63	17	indonesia	indonesia	PROPN
biotropia-309	63	18	.	.	PUNCT
biotropia-309	64	1	the	the	DET
biotropia-309	64	2	park	park	NOUN
biotropia-309	64	3	,	,	PUNCT
biotropia-309	64	4	a	a	DET
biotropia-309	64	5	local	local	ADJ
biotropia-309	64	6	biodiversity	biodiversity	NOUN
biotropia-309	64	7	hot	hot	ADJ
biotropia-309	64	8	spot	spot	NOUN
biotropia-309	64	9	covers	cover	VERB
biotropia-309	64	10	an	an	DET
biotropia-309	64	11	area	area	NOUN
biotropia-309	64	12	of	of	ADP
biotropia-309	64	13	229,000	229,000	NUM
biotropia-309	64	14	ha	ha	INTJ
biotropia-309	64	15	and	and	CCONJ
biotropia-309	64	16	is	be	AUX
biotropia-309	64	17	located	locate	VERB
biotropia-309	64	18	southeast	southeast	ADV
biotropia-309	64	19	of	of	ADP
biotropia-309	64	20	palu	palu	PROPN
biotropia-309	64	21	,	,	PUNCT
biotropia-309	64	22	the	the	DET
biotropia-309	64	23	capital	capital	NOUN
biotropia-309	64	24	of	of	ADP
biotropia-309	64	25	central	central	ADJ
biotropia-309	64	26	sulawesi	sulawesi	PROPN
biotropia-309	64	27	province	province	NOUN
biotropia-309	64	28	.	.	PUNCT
biotropia-309	65	1	all	all	DET
biotropia-309	65	2	study	study	NOUN
biotropia-309	65	3	sites	site	NOUN
biotropia-309	65	4	were	be	AUX
biotropia-309	65	5	selected	select	VERB
biotropia-309	65	6	at	at	ADP
biotropia-309	65	7	the	the	DET
biotropia-309	65	8	surrounding	surrounding	NOUN
biotropia-309	65	9	of	of	ADP
biotropia-309	65	10	the	the	DET
biotropia-309	65	11	palolo	palolo	ADJ
biotropia-309	65	12	valley	valley	NOUN
biotropia-309	65	13	in	in	ADP
biotropia-309	65	14	the	the	DET
biotropia-309	65	15	vicinity	vicinity	NOUN
biotropia-309	65	16	of	of	ADP
biotropia-309	65	17	the	the	DET
biotropia-309	65	18	villages	village	NOUN
biotropia-309	65	19	of	of	ADP
biotropia-309	65	20	bobo	bobo	PROPN
biotropia-309	65	21	et	et	PROPN
biotropia-309	65	22	al	al	PROPN
biotropia-309	65	23	.	.	PROPN
biotropia-309	65	24	et	et	PROPN
biotropia-309	66	1	al	al	PROPN
biotropia-309	66	2	et	et	PROPN
biotropia-309	66	3	al	al	PROPN
biotropia-309	66	4	.	.	PROPN
biotropia-309	66	5	et	et	PROPN
biotropia-309	67	1	al	al	PROPN
biotropia-309	67	2	.	.	PROPN
biotropia-309	67	3	et	et	PROPN
biotropia-309	67	4	al	al	PROPN
biotropia-309	67	5	et	et	PROPN
biotropia-309	67	6	al	al	PROPN
biotropia-309	67	7	et	et	PROPN
biotropia-309	67	8	al	al	PROPN
biotropia-309	67	9	et	et	PROPN
biotropia-309	67	10	al	al	PROPN
biotropia-309	67	11	et	et	PROPN
biotropia-309	67	12	al	al	PROPN
biotropia-309	67	13	indval	indval	PROPN
biotropia-309	67	14	et	et	PROPN
biotropia-309	67	15	al	al	PROPN
biotropia-309	67	16	et	et	PROPN
biotropia-309	67	17	al	al	PROPN
biotropia-309	67	18	indval	indval	PROPN
biotropia-309	67	19	et	et	PROPN
biotropia-309	67	20	al	al	PROPN
biotropia-309	67	21	et	et	PROPN
biotropia-309	67	22	al	al	PROPN
biotropia-309	67	23	et	et	PROPN
biotropia-309	67	24	al	al	PROPN
biotropia-309	67	25	materials	materials	PROPN
biotropia-309	67	26	and	and	CCONJ
biotropia-309	67	27	methods	method	NOUN
biotropia-309	67	28	study	study	NOUN
biotropia-309	67	29	area	area	NOUN
biotropia-309	67	30	50	50	NUM
biotropia-309	67	31	biotropia	biotropia	ADJ
biotropia-309	67	32	vol	vol	NOUN
biotropia-309	67	33	.	.	PROPN
biotropia-309	67	34	21	21	NUM
biotropia-309	67	35	no	no	NOUN
biotropia-309	67	36	.	.	NOUN
biotropia-309	67	37	1	1	NUM
biotropia-309	67	38	,	,	PUNCT
biotropia-309	67	39	2014	2014	NUM
biotropia-309	67	40	(	(	PUNCT
biotropia-309	67	41	01	01	NUM
biotropia-309	67	42	07'0.46	07'0.46	NUM
biotropia-309	67	43	"	"	PUNCT
biotropia-309	67	44	s	s	PART
biotropia-309	67	45	119	119	NUM
biotropia-309	67	46	59'702	59'702	NUM
biotropia-309	67	47	"	"	PUNCT
biotropia-309	67	48	e	e	NOUN
biotropia-309	67	49	)	)	PUNCT
biotropia-309	67	50	and	and	CCONJ
biotropia-309	67	51	were	be	AUX
biotropia-309	67	52	situated	situate	VERB
biotropia-309	67	53	at	at	ADP
biotropia-309	67	54	an	an	DET
biotropia-309	67	55	altitude	altitude	NOUN
biotropia-309	67	56	between	between	ADP
biotropia-309	67	57	790	790	NUM
biotropia-309	67	58	and	and	CCONJ
biotropia-309	67	59	985	985	NUM
biotropia-309	67	60	m	m	NOUN
biotropia-309	67	61	asl	asl	PROPN
biotropia-309	67	62	.	.	PUNCT
biotropia-309	68	1	dung	dung	NOUN
biotropia-309	68	2	beetle	beetle	NOUN
biotropia-309	68	3	communities	community	NOUN
biotropia-309	68	4	were	be	AUX
biotropia-309	68	5	studied	study	VERB
biotropia-309	68	6	in	in	ADP
biotropia-309	68	7	five	five	NUM
biotropia-309	68	8	land	land	NOUN
biotropia-309	68	9	-	-	PUNCT
biotropia-309	68	10	use	use	NOUN
biotropia-309	68	11	types	type	NOUN
biotropia-309	68	12	:	:	PUNCT
biotropia-309	68	13	natural	natural	ADJ
biotropia-309	68	14	forest	forest	NOUN
biotropia-309	68	15	(	(	PUNCT
biotropia-309	68	16	nf	nf	INTJ
biotropia-309	68	17	)	)	PUNCT
biotropia-309	68	18	;	;	PUNCT
biotropia-309	68	19	secondary	secondary	ADJ
biotropia-309	68	20	forest	forest	NOUN
biotropia-309	68	21	(	(	PUNCT
biotropia-309	68	22	sf	sf	PROPN
biotropia-309	68	23	)	)	PUNCT
biotropia-309	68	24	;	;	PUNCT
biotropia-309	68	25	cacao	cacao	NOUN
biotropia-309	68	26	agroforestry	agroforestry	NOUN
biotropia-309	68	27	systems	system	NOUN
biotropia-309	68	28	(	(	PUNCT
biotropia-309	68	29	cacao	cacao	NOUN
biotropia-309	68	30	cultivated	cultivate	VERB
biotropia-309	68	31	under	under	ADP
biotropia-309	68	32	natural	natural	ADJ
biotropia-309	68	33	shade	shade	NOUN
biotropia-309	68	34	tree	tree	NOUN
biotropia-309	68	35	at	at	ADP
biotropia-309	68	36	the	the	DET
biotropia-309	68	37	forest	forest	NOUN
biotropia-309	68	38	margin	margin	NOUN
biotropia-309	68	39	(	(	PUNCT
biotropia-309	68	40	ac	ac	PROPN
biotropia-309	68	41	)	)	PUNCT
biotropia-309	68	42	;	;	PUNCT
biotropia-309	68	43	cacao	cacao	NOUN
biotropia-309	68	44	plantation	plantation	NOUN
biotropia-309	68	45	under	under	ADP
biotropia-309	68	46	monospecific	monospecific	ADJ
biotropia-309	68	47	shade	shade	NOUN
biotropia-309	68	48	tree	tree	NOUN
biotropia-309	68	49	dominated	dominate	VERB
biotropia-309	68	50	by	by	ADP
biotropia-309	68	51	(	(	PUNCT
biotropia-309	68	52	cp	cp	NOUN
biotropia-309	68	53	)	)	PUNCT
biotropia-309	68	54	;	;	PUNCT
biotropia-309	68	55	and	and	CCONJ
biotropia-309	68	56	open	open	ADJ
biotropia-309	68	57	area	area	NOUN
biotropia-309	68	58	(	(	PUNCT
biotropia-309	68	59	oa	oa	NOUN
biotropia-309	68	60	)	)	PUNCT
biotropia-309	68	61	,	,	PUNCT
biotropia-309	68	62	cultivated	cultivate	VERB
biotropia-309	68	63	either	either	ADV
biotropia-309	68	64	by	by	ADP
biotropia-309	68	65	or	or	CCONJ
biotropia-309	68	66	.	.	PUNCT
biotropia-309	69	1	three	three	NUM
biotropia-309	69	2	replications	replication	NOUN
biotropia-309	69	3	sites	site	NOUN
biotropia-309	69	4	for	for	ADP
biotropia-309	69	5	each	each	DET
biotropia-309	69	6	land	land	NOUN
biotropia-309	69	7	-	-	PUNCT
biotropia-309	69	8	use	use	NOUN
biotropia-309	69	9	type	type	NOUN
biotropia-309	69	10	were	be	AUX
biotropia-309	69	11	selected	select	VERB
biotropia-309	69	12	with	with	ADP
biotropia-309	69	13	distance	distance	NOUN
biotropia-309	69	14	at	at	ADP
biotropia-309	69	15	least	least	ADJ
biotropia-309	69	16	50	50	NUM
biotropia-309	69	17	m	m	NOUN
biotropia-309	69	18	from	from	ADP
biotropia-309	69	19	each	each	DET
biotropia-309	69	20	other	other	ADJ
biotropia-309	69	21	dung	dung	NOUN
biotropia-309	69	22	beetles	beetle	NOUN
biotropia-309	69	23	were	be	AUX
biotropia-309	69	24	sampled	sample	VERB
biotropia-309	69	25	in	in	ADP
biotropia-309	69	26	2500	2500	NUM
biotropia-309	69	27	m	m	NOUN
biotropia-309	69	28	plots	plot	NOUN
biotropia-309	69	29	at	at	ADP
biotropia-309	69	30	15	15	NUM
biotropia-309	69	31	sites	site	NOUN
biotropia-309	69	32	all	all	PRON
biotropia-309	69	33	using	use	VERB
biotropia-309	69	34	baited	bait	VERB
biotropia-309	69	35	pitfall	pitfall	NOUN
biotropia-309	69	36	traps	trap	NOUN
biotropia-309	69	37	as	as	SCONJ
biotropia-309	69	38	described	describe	VERB
biotropia-309	69	39	in	in	ADP
biotropia-309	69	40	shahabuddin	shahabuddin	VERB
biotropia-309	69	41	(	(	PUNCT
biotropia-309	69	42	2010	2010	NUM
biotropia-309	69	43	)	)	PUNCT
biotropia-309	69	44	.	.	PUNCT
biotropia-309	70	1	six	six	NUM
biotropia-309	70	2	traps	trap	NOUN
biotropia-309	70	3	were	be	AUX
biotropia-309	70	4	set	set	VERB
biotropia-309	70	5	up	up	ADP
biotropia-309	70	6	at	at	ADP
biotropia-309	70	7	the	the	DET
biotropia-309	70	8	centre	centre	NOUN
biotropia-309	70	9	of	of	ADP
biotropia-309	70	10	each	each	DET
biotropia-309	70	11	plot	plot	NOUN
biotropia-309	70	12	and	and	CCONJ
biotropia-309	70	13	placed	place	VERB
biotropia-309	70	14	with	with	ADP
biotropia-309	70	15	an	an	DET
biotropia-309	70	16	interval	interval	NOUN
biotropia-309	70	17	of	of	ADP
biotropia-309	70	18	10	10	NUM
biotropia-309	70	19	m.	m.	NOUN
biotropia-309	70	20	the	the	DET
biotropia-309	70	21	traps	trap	NOUN
biotropia-309	70	22	were	be	AUX
biotropia-309	70	23	baited	bait	VERB
biotropia-309	70	24	with	with	ADP
biotropia-309	70	25	ca	ca	PROPN
biotropia-309	70	26	.	.	PUNCT
biotropia-309	71	1	30	30	NUM
biotropia-309	71	2	g	g	NOUN
biotropia-309	71	3	of	of	ADP
biotropia-309	71	4	fresh	fresh	ADJ
biotropia-309	71	5	cattle	cattle	NOUN
biotropia-309	71	6	(	(	PUNCT
biotropia-309	71	7	)	)	PUNCT
biotropia-309	71	8	dung	dung	NOUN
biotropia-309	71	9	and	and	CCONJ
biotropia-309	71	10	exposed	expose	VERB
biotropia-309	71	11	six	six	NUM
biotropia-309	71	12	times	time	NOUN
biotropia-309	71	13	from	from	ADP
biotropia-309	71	14	april	april	PROPN
biotropia-309	71	15	to	to	ADP
biotropia-309	71	16	july	july	PROPN
biotropia-309	71	17	in	in	ADP
biotropia-309	71	18	2009	2009	NUM
biotropia-309	71	19	and	and	CCONJ
biotropia-309	71	20	march	march	PROPN
biotropia-309	71	21	to	to	ADP
biotropia-309	71	22	june	june	PROPN
biotropia-309	71	23	in	in	ADP
biotropia-309	71	24	2012	2012	NUM
biotropia-309	71	25	.	.	PUNCT
biotropia-309	72	1	cattle	cattle	NOUN
biotropia-309	72	2	dung	dung	NOUN
biotropia-309	72	3	has	have	AUX
biotropia-309	72	4	been	be	AUX
biotropia-309	72	5	widely	widely	ADV
biotropia-309	72	6	used	use	VERB
biotropia-309	72	7	as	as	ADP
biotropia-309	72	8	bait	bait	NOUN
biotropia-309	72	9	for	for	ADP
biotropia-309	72	10	dung	dung	NOUN
biotropia-309	72	11	beetles	beetle	NOUN
biotropia-309	72	12	,	,	PUNCT
biotropia-309	72	13	aside	aside	ADV
biotropia-309	72	14	from	from	ADP
biotropia-309	72	15	human	human	ADJ
biotropia-309	72	16	faeces	faece	NOUN
biotropia-309	72	17	(	(	PUNCT
biotropia-309	72	18	e.g.	e.g.	ADV
biotropia-309	72	19	erroizi	erroizi	NOUN
biotropia-309	72	20	2004	2004	NUM
biotropia-309	72	21	;	;	PUNCT
biotropia-309	72	22	andresen	andresen	PROPN
biotropia-309	72	23	2005	2005	NUM
biotropia-309	72	24	;	;	PUNCT
biotropia-309	72	25	mendoza	mendoza	PROPN
biotropia-309	72	26	2005	2005	NUM
biotropia-309	72	27	)	)	PUNCT
biotropia-309	72	28	.	.	PUNCT
biotropia-309	73	1	our	our	PRON
biotropia-309	73	2	previous	previous	ADJ
biotropia-309	73	3	study	study	NOUN
biotropia-309	73	4	also	also	ADV
biotropia-309	73	5	showed	show	VERB
biotropia-309	73	6	that	that	SCONJ
biotropia-309	73	7	at	at	ADP
biotropia-309	73	8	the	the	DET
biotropia-309	73	9	same	same	ADJ
biotropia-309	73	10	weight	weight	NOUN
biotropia-309	73	11	of	of	ADP
biotropia-309	73	12	bait	bait	NOUN
biotropia-309	73	13	(	(	PUNCT
biotropia-309	73	14	ca	ca	NOUN
biotropia-309	73	15	.	.	NOUN
biotropia-309	73	16	30	30	NUM
biotropia-309	73	17	g	g	NOUN
biotropia-309	73	18	)	)	PUNCT
biotropia-309	73	19	cattle	cattle	NOUN
biotropia-309	73	20	dung	dung	NOUN
biotropia-309	73	21	attracts	attract	VERB
biotropia-309	73	22	the	the	DET
biotropia-309	73	23	dung	dung	NOUN
biotropia-309	73	24	beetles	beetle	NOUN
biotropia-309	73	25	with	with	ADP
biotropia-309	73	26	similar	similar	ADJ
biotropia-309	73	27	species	species	NOUN
biotropia-309	73	28	composition	composition	NOUN
biotropia-309	73	29	found	find	VERB
biotropia-309	73	30	in	in	ADP
biotropia-309	73	31	the	the	DET
biotropia-309	73	32	dung	dung	NOUN
biotropia-309	73	33	of	of	ADP
biotropia-309	73	34	anoa	anoa	PROPN
biotropia-309	73	35	(	(	PUNCT
biotropia-309	73	36	)	)	PUNCT
biotropia-309	73	37	,	,	PUNCT
biotropia-309	73	38	an	an	DET
biotropia-309	73	39	endemic	endemic	ADJ
biotropia-309	73	40	herbivore	herbivore	NOUN
biotropia-309	73	41	of	of	ADP
biotropia-309	73	42	sulawesi	sulawesi	NOUN
biotropia-309	73	43	(	(	PUNCT
biotropia-309	73	44	shahabuddin	shahabuddin	NOUN
biotropia-309	73	45	.	.	PUNCT
biotropia-309	74	1	2010	2010	NUM
biotropia-309	74	2	)	)	PUNCT
biotropia-309	74	3	.	.	PUNCT
biotropia-309	75	1	the	the	DET
biotropia-309	75	2	trapped	trap	VERB
biotropia-309	75	3	specimens	specimen	NOUN
biotropia-309	75	4	were	be	AUX
biotropia-309	75	5	removed	remove	VERB
biotropia-309	75	6	after	after	ADP
biotropia-309	75	7	two	two	NUM
biotropia-309	75	8	days	day	NOUN
biotropia-309	75	9	and	and	CCONJ
biotropia-309	75	10	preserved	preserve	VERB
biotropia-309	75	11	in	in	ADP
biotropia-309	75	12	scheerpelz	scheerpelz	NOUN
biotropia-309	75	13	solution	solution	NOUN
biotropia-309	75	14	(	(	PUNCT
biotropia-309	75	15	krell	krell	PROPN
biotropia-309	75	16	2007	2007	NUM
biotropia-309	75	17	)	)	PUNCT
biotropia-309	75	18	.	.	PUNCT
biotropia-309	76	1	later	later	ADV
biotropia-309	76	2	on	on	ADV
biotropia-309	76	3	,	,	PUNCT
biotropia-309	76	4	the	the	DET
biotropia-309	76	5	samples	sample	NOUN
biotropia-309	76	6	were	be	AUX
biotropia-309	76	7	identified	identify	VERB
biotropia-309	76	8	in	in	ADP
biotropia-309	76	9	the	the	DET
biotropia-309	76	10	laboratory	laboratory	NOUN
biotropia-309	76	11	using	use	VERB
biotropia-309	76	12	available	available	ADJ
biotropia-309	76	13	identification	identification	NOUN
biotropia-309	76	14	keys	key	NOUN
biotropia-309	76	15	(	(	PUNCT
biotropia-309	76	16	e.g.	e.g.	ADV
biotropia-309	76	17	balthasar	balthasar	VERB
biotropia-309	76	18	1963	1963	NUM
biotropia-309	76	19	)	)	PUNCT
biotropia-309	76	20	and	and	CCONJ
biotropia-309	76	21	by	by	ADP
biotropia-309	76	22	comparing	compare	VERB
biotropia-309	76	23	to	to	ADP
biotropia-309	76	24	the	the	DET
biotropia-309	76	25	reference	reference	NOUN
biotropia-309	76	26	collection	collection	NOUN
biotropia-309	76	27	of	of	ADP
biotropia-309	76	28	the	the	DET
biotropia-309	76	29	center	center	NOUN
biotropia-309	76	30	for	for	ADP
biotropia-309	76	31	biodiversity	biodiversity	NOUN
biotropia-309	76	32	research	research	NOUN
biotropia-309	76	33	tadulako	tadulako	PROPN
biotropia-309	76	34	university	university	PROPN
biotropia-309	76	35	.	.	PUNCT
biotropia-309	77	1	species	specie	NOUN
biotropia-309	77	2	which	which	PRON
biotropia-309	77	3	could	could	AUX
biotropia-309	77	4	not	not	PART
biotropia-309	77	5	be	be	AUX
biotropia-309	77	6	identified	identify	VERB
biotropia-309	77	7	,	,	PUNCT
biotropia-309	77	8	were	be	AUX
biotropia-309	77	9	sorted	sort	VERB
biotropia-309	77	10	to	to	ADP
biotropia-309	77	11	morphospecies	morphospecie	NOUN
biotropia-309	77	12	.	.	PUNCT
biotropia-309	78	1	several	several	ADJ
biotropia-309	78	2	habitat	habitat	ADJ
biotropia-309	78	3	parameters	parameter	NOUN
biotropia-309	78	4	(	(	PUNCT
biotropia-309	78	5	i.e.	i.e.	X
biotropia-309	78	6	vegetation	vegetation	NOUN
biotropia-309	78	7	structure	structure	NOUN
biotropia-309	78	8	and	and	CCONJ
biotropia-309	78	9	microclimate	microclimate	NOUN
biotropia-309	78	10	)	)	PUNCT
biotropia-309	78	11	affecting	affect	VERB
biotropia-309	78	12	the	the	DET
biotropia-309	78	13	dung	dung	NOUN
biotropia-309	78	14	beetles	beetles	NOUN
biotropia-309	78	15	diversity	diversity	NOUN
biotropia-309	78	16	(	(	PUNCT
biotropia-309	78	17	see	see	VERB
biotropia-309	78	18	davis	davis	PROPN
biotropia-309	78	19	.	.	PUNCT
biotropia-309	79	1	2001	2001	NUM
biotropia-309	79	2	;	;	PUNCT
biotropia-309	79	3	shahabuddin	shahabuddin	VERB
biotropia-309	79	4	2010	2010	NUM
biotropia-309	79	5	)	)	PUNCT
biotropia-309	79	6	were	be	AUX
biotropia-309	79	7	measured	measure	VERB
biotropia-309	79	8	to	to	PART
biotropia-309	79	9	characterize	characterize	VERB
biotropia-309	79	10	the	the	DET
biotropia-309	79	11	land	land	NOUN
biotropia-309	79	12	-	-	PUNCT
biotropia-309	79	13	use	use	NOUN
biotropia-309	79	14	types	type	NOUN
biotropia-309	79	15	including	include	VERB
biotropia-309	79	16	air	air	NOUN
biotropia-309	79	17	temperature	temperature	NOUN
biotropia-309	79	18	,	,	PUNCT
biotropia-309	79	19	relative	relative	ADJ
biotropia-309	79	20	humidity	humidity	NOUN
biotropia-309	79	21	,	,	PUNCT
biotropia-309	79	22	canopy	canopy	NOUN
biotropia-309	79	23	cover	cover	NOUN
biotropia-309	79	24	,	,	PUNCT
biotropia-309	79	25	and	and	CCONJ
biotropia-309	79	26	herb	herb	NOUN
biotropia-309	79	27	layer	layer	NOUN
biotropia-309	79	28	coverage	coverage	NOUN
biotropia-309	79	29	.	.	PUNCT
biotropia-309	80	1	the	the	DET
biotropia-309	80	2	relative	relative	ADJ
biotropia-309	80	3	humidity	humidity	NOUN
biotropia-309	80	4	at	at	ADP
biotropia-309	80	5	the	the	DET
biotropia-309	80	6	start	start	NOUN
biotropia-309	80	7	and	and	CCONJ
biotropia-309	80	8	end	end	NOUN
biotropia-309	80	9	of	of	ADP
biotropia-309	80	10	the	the	DET
biotropia-309	80	11	exposure	exposure	NOUN
biotropia-309	80	12	period	period	NOUN
biotropia-309	80	13	were	be	AUX
biotropia-309	80	14	measured	measure	VERB
biotropia-309	80	15	using	use	VERB
biotropia-309	80	16	a	a	DET
biotropia-309	80	17	digital	digital	ADJ
biotropia-309	80	18	thermo	thermo	NOUN
biotropia-309	80	19	-	-	PUNCT
biotropia-309	80	20	hygrometer	hygrometer	NOUN
biotropia-309	80	21	(	(	PUNCT
biotropia-309	80	22	corona	corona	NOUN
biotropia-309	80	23	model	model	NOUN
biotropia-309	80	24	:	:	PUNCT
biotropia-309	80	25	gl	gl	PROPN
biotropia-309	80	26	99	99	NUM
biotropia-309	80	27	)	)	PUNCT
biotropia-309	80	28	1	1	NUM
biotropia-309	80	29	m	m	NOUN
biotropia-309	80	30	above	above	ADP
biotropia-309	80	31	ground	ground	NOUN
biotropia-309	80	32	while	while	SCONJ
biotropia-309	80	33	the	the	DET
biotropia-309	80	34	canopy	canopy	NOUN
biotropia-309	80	35	cover	cover	NOUN
biotropia-309	80	36	was	be	AUX
biotropia-309	80	37	visually	visually	ADV
biotropia-309	80	38	estimated	estimate	VERB
biotropia-309	80	39	at	at	ADP
biotropia-309	80	40	four	four	NUM
biotropia-309	80	41	locations	location	NOUN
biotropia-309	80	42	per	per	ADP
biotropia-309	80	43	site	site	NOUN
biotropia-309	80	44	for	for	ADP
biotropia-309	80	45	a	a	DET
biotropia-309	80	46	corridor	corridor	NOUN
biotropia-309	80	47	of	of	ADP
biotropia-309	80	48	ca	ca	PROPN
biotropia-309	80	49	.	.	PUNCT
biotropia-309	81	1	10	10	NUM
biotropia-309	81	2	m	m	NOUN
biotropia-309	81	3	inside	inside	ADP
biotropia-309	81	4	the	the	DET
biotropia-309	81	5	plot	plot	NOUN
biotropia-309	81	6	.	.	PUNCT
biotropia-309	82	1	the	the	DET
biotropia-309	82	2	herb	herb	NOUN
biotropia-309	82	3	coverage	coverage	NOUN
biotropia-309	82	4	was	be	AUX
biotropia-309	82	5	estimated	estimate	VERB
biotropia-309	82	6	at	at	ADP
biotropia-309	82	7	four	four	NUM
biotropia-309	82	8	plots	plot	NOUN
biotropia-309	82	9	of	of	ADP
biotropia-309	82	10	2x2	2x2	NUM
biotropia-309	82	11	m	m	AUX
biotropia-309	82	12	randomly	randomly	ADV
biotropia-309	82	13	placed	place	VERB
biotropia-309	82	14	at	at	ADP
biotropia-309	82	15	ca	ca	NOUN
biotropia-309	82	16	.	.	PROPN
biotropia-309	83	1	5	5	NUM
biotropia-309	83	2	m	m	NOUN
biotropia-309	83	3	inside	inside	ADP
biotropia-309	83	4	the	the	DET
biotropia-309	83	5	plot	plot	NOUN
biotropia-309	83	6	.	.	PUNCT
biotropia-309	84	1	based	base	VERB
biotropia-309	84	2	on	on	ADP
biotropia-309	84	3	the	the	DET
biotropia-309	84	4	environmental	environmental	ADJ
biotropia-309	84	5	variables	variable	NOUN
biotropia-309	84	6	measured	measure	VERB
biotropia-309	84	7	all	all	DET
biotropia-309	84	8	land	land	NOUN
biotropia-309	84	9	-	-	PUNCT
biotropia-309	84	10	use	use	NOUN
biotropia-309	84	11	types	type	NOUN
biotropia-309	84	12	were	be	AUX
biotropia-309	84	13	then	then	ADV
biotropia-309	84	14	grouped	group	VERB
biotropia-309	84	15	using	use	VERB
biotropia-309	84	16	a	a	DET
biotropia-309	84	17	two	two	NUM
biotropia-309	84	18	-	-	PUNCT
biotropia-309	84	19	dimensional	dimensional	ADJ
biotropia-309	84	20	scaling	scaling	NOUN
biotropia-309	84	21	(	(	PUNCT
biotropia-309	84	22	clarke	clarke	PROPN
biotropia-309	84	23	1993	1993	NUM
biotropia-309	84	24	;	;	PUNCT
biotropia-309	84	25	statsoft	statsoft	PROPN
biotropia-309	84	26	2001	2001	NUM
biotropia-309	84	27	)	)	PUNCT
biotropia-309	84	28	.	.	PUNCT
biotropia-309	85	1	the	the	DET
biotropia-309	85	2	three	three	NUM
biotropia-309	85	3	most	most	ADV
biotropia-309	85	4	widely	widely	ADV
biotropia-309	85	5	used	use	VERB
biotropia-309	85	6	measures	measure	NOUN
biotropia-309	85	7	of	of	ADP
biotropia-309	85	8	species	specie	NOUN
biotropia-309	85	9	diversity	diversity	NOUN
biotropia-309	85	10	were	be	AUX
biotropia-309	85	11	species	specie	NOUN
biotropia-309	85	12	richness	richness	ADJ
biotropia-309	85	13	,	,	PUNCT
biotropia-309	85	14	shannon	shannon	ADJ
biotropia-309	85	15	-	-	PUNCT
biotropia-309	85	16	wiener	wiener	NOUN
biotropia-309	85	17	index	index	NOUN
biotropia-309	85	18	and	and	CCONJ
biotropia-309	85	19	simpson	simpson	PROPN
biotropia-309	85	20	index	index	PROPN
biotropia-309	85	21	(	(	PUNCT
biotropia-309	85	22	si	si	X
biotropia-309	85	23	=	=	SYM
biotropia-309	85	24	∑	∑	PROPN
biotropia-309	85	25	pi	pi	PROPN
biotropia-309	85	26	)	)	PUNCT
biotropia-309	85	27	(	(	PUNCT
biotropia-309	85	28	lande	lande	NOUN
biotropia-309	85	29	1996	1996	NUM
biotropia-309	85	30	)	)	PUNCT
biotropia-309	85	31	.	.	PUNCT
biotropia-309	86	1	species	specie	NOUN
biotropia-309	86	2	richness	richness	NOUN
biotropia-309	86	3	of	of	ADP
biotropia-309	86	4	dung	dung	NOUN
biotropia-309	86	5	beetles	beetle	NOUN
biotropia-309	86	6	was	be	AUX
biotropia-309	86	7	estimated	estimate	VERB
biotropia-309	86	8	using	use	VERB
biotropia-309	86	9	the	the	DET
biotropia-309	86	10	second	second	ADJ
biotropia-309	86	11	-	-	PUNCT
biotropia-309	86	12	order	order	NOUN
biotropia-309	86	13	jackknife	jackknife	NOUN
biotropia-309	86	14	extrapolation	extrapolation	NOUN
biotropia-309	86	15	method	method	NOUN
biotropia-309	86	16	(	(	PUNCT
biotropia-309	86	17	colwell	colwell	PROPN
biotropia-309	86	18	,	,	PUNCT
biotropia-309	86	19	2004	2004	NUM
biotropia-309	86	20	)	)	PUNCT
biotropia-309	86	21	,	,	PUNCT
biotropia-309	86	22	one	one	NUM
biotropia-309	86	23	of	of	ADP
biotropia-309	86	24	the	the	DET
biotropia-309	86	25	best	good	ADJ
biotropia-309	86	26	species	specie	NOUN
biotropia-309	86	27	richness	richness	NOUN
biotropia-309	86	28	predictor	predictor	NOUN
biotropia-309	86	29	with	with	ADP
biotropia-309	86	30	respect	respect	NOUN
biotropia-309	86	31	to	to	ADP
biotropia-309	86	32	accuracy	accuracy	NOUN
biotropia-309	86	33	(	(	PUNCT
biotropia-309	86	34	e.g.	e.g.	ADV
biotropia-309	86	35	brose	brose	NOUN
biotropia-309	86	36	.	.	PUNCT
biotropia-309	87	1	2003	2003	NUM
biotropia-309	87	2	)	)	PUNCT
biotropia-309	87	3	.	.	PUNCT
biotropia-309	88	1	as	as	ADP
biotropia-309	88	2	units	unit	NOUN
biotropia-309	88	3	for	for	ADP
biotropia-309	88	4	estimating	estimate	VERB
biotropia-309	88	5	the	the	DET
biotropia-309	88	6	total	total	ADJ
biotropia-309	88	7	species	specie	NOUN
biotropia-309	88	8	richness	richness	NOUN
biotropia-309	88	9	of	of	ADP
biotropia-309	88	10	land	land	NOUN
biotropia-309	88	11	-	-	PUNCT
biotropia-309	88	12	use	use	NOUN
biotropia-309	88	13	type	type	NOUN
biotropia-309	88	14	,	,	PUNCT
biotropia-309	88	15	samples	sample	NOUN
biotropia-309	88	16	from	from	ADP
biotropia-309	88	17	all	all	DET
biotropia-309	88	18	traps	trap	NOUN
biotropia-309	88	19	and	and	CCONJ
biotropia-309	88	20	replicates	replicate	NOUN
biotropia-309	88	21	were	be	AUX
biotropia-309	88	22	pooled	pool	VERB
biotropia-309	88	23	for	for	ADP
biotropia-309	88	24	individual	individual	ADJ
biotropia-309	88	25	sample	sample	NOUN
biotropia-309	88	26	o	o	NOUN
biotropia-309	88	27	o	o	NOUN
biotropia-309	89	1	2	2	NUM
biotropia-309	89	2	r	r	NOUN
biotropia-309	89	3	2	2	NUM
biotropia-309	89	4	2	2	NUM
biotropia-309	89	5	gliricidia	gliricidia	NOUN
biotropia-309	89	6	sepium	sepium	PROPN
biotropia-309	89	7	zea	zea	PROPN
biotropia-309	89	8	mays	mays	PROPN
biotropia-309	89	9	morus	morus	PROPN
biotropia-309	89	10	alba	alba	PROPN
biotropia-309	89	11	et	et	PROPN
biotropia-309	89	12	al	al	PROPN
biotropia-309	89	13	.	.	PROPN
biotropia-309	89	14	bos	bos	PROPN
biotropia-309	89	15	taurus	taurus	PROPN
biotropia-309	89	16	et	et	PROPN
biotropia-309	89	17	al	al	PROPN
biotropia-309	89	18	.	.	PROPN
biotropia-309	89	19	et	et	PROPN
biotropia-309	90	1	al	al	PROPN
biotropia-309	90	2	.	.	PUNCT
biotropia-309	91	1	bubalus	bubalus	PROPN
biotropia-309	91	2	deppresicornis	deppresicornis	PROPN
biotropia-309	91	3	et	et	PROPN
biotropia-309	91	4	al	al	PROPN
biotropia-309	91	5	et	et	PROPN
biotropia-309	91	6	al	al	PROPN
biotropia-309	91	7	et	et	PROPN
biotropia-309	91	8	al	al	PROPN
biotropia-309	91	9	specimens	specimens	PROPN
biotropia-309	91	10	collection	collection	PROPN
biotropia-309	91	11	environmental	environmental	ADJ
biotropia-309	91	12	variables	variable	NOUN
biotropia-309	91	13	measured	measure	VERB
biotropia-309	91	14	data	datum	NOUN
biotropia-309	91	15	analysis	analysis	NOUN
biotropia-309	91	16	.	.	PUNCT
biotropia-309	92	1	51	51	NUM
biotropia-309	92	2	effectiveness	effectiveness	NOUN
biotropia-309	92	3	of	of	ADP
biotropia-309	92	4	dung	dung	NOUN
biotropia-309	92	5	beetles	beetle	NOUN
biotropia-309	92	6	as	as	ADP
biotropia-309	92	7	bioindicators	bioindicator	NOUN
biotropia-309	92	8	of	of	ADP
biotropia-309	92	9	environmental	environmental	ADJ
biotropia-309	92	10	changes	change	NOUN
biotropia-309	92	11	in	in	ADP
biotropia-309	92	12	land	land	NOUN
biotropia-309	92	13	shahabuddin	shahabuddin	NOUN
biotropia-309	92	14	–	–	PUNCT
biotropia-309	92	15	et	et	PROPN
biotropia-309	92	16	al	al	PROPN
biotropia-309	92	17	.	.	PROPN
biotropia-309	92	18	times	times	PROPN
biotropia-309	92	19	(	(	PUNCT
biotropia-309	92	20	=	=	NOUN
biotropia-309	92	21	6	6	NUM
biotropia-309	92	22	)	)	PUNCT
biotropia-309	92	23	due	due	ADP
biotropia-309	92	24	to	to	ADP
biotropia-309	92	25	the	the	DET
biotropia-309	92	26	close	close	ADJ
biotropia-309	92	27	proximity	proximity	NOUN
biotropia-309	92	28	between	between	ADP
biotropia-309	92	29	each	each	DET
biotropia-309	92	30	trap	trap	NOUN
biotropia-309	92	31	and	and	CCONJ
biotropia-309	92	32	site	site	NOUN
biotropia-309	92	33	.	.	PUNCT
biotropia-309	93	1	effects	effect	NOUN
biotropia-309	93	2	of	of	ADP
biotropia-309	93	3	habitat	habitat	ADJ
biotropia-309	93	4	type	type	NOUN
biotropia-309	93	5	on	on	ADP
biotropia-309	93	6	diversity	diversity	NOUN
biotropia-309	93	7	were	be	AUX
biotropia-309	93	8	tested	test	VERB
biotropia-309	93	9	using	use	VERB
biotropia-309	93	10	one	one	NUM
biotropia-309	93	11	-	-	PUNCT
biotropia-309	93	12	way	way	NOUN
biotropia-309	93	13	anova	anova	PROPN
biotropia-309	93	14	.	.	PUNCT
biotropia-309	94	1	abundance	abundance	NOUN
biotropia-309	94	2	data	datum	NOUN
biotropia-309	94	3	were	be	AUX
biotropia-309	94	4	transformed	transform	VERB
biotropia-309	94	5	by	by	ADP
biotropia-309	94	6	log	log	NOUN
biotropia-309	94	7	(	(	PUNCT
biotropia-309	94	8	n+1	n+1	NOUN
biotropia-309	94	9	)	)	PUNCT
biotropia-309	94	10	before	before	ADP
biotropia-309	94	11	analysis	analysis	NOUN
biotropia-309	94	12	(	(	PUNCT
biotropia-309	94	13	zar	zar	NOUN
biotropia-309	94	14	1999	1999	NUM
biotropia-309	94	15	)	)	PUNCT
biotropia-309	94	16	.	.	PUNCT
biotropia-309	95	1	statsoft	statsoft	ADJ
biotropia-309	95	2	6.0	6.0	NUM
biotropia-309	95	3	software	software	NOUN
biotropia-309	95	4	(	(	PUNCT
biotropia-309	95	5	2001	2001	NUM
biotropia-309	95	6	)	)	PUNCT
biotropia-309	95	7	was	be	AUX
biotropia-309	95	8	used	use	VERB
biotropia-309	95	9	to	to	PART
biotropia-309	95	10	perform	perform	VERB
biotropia-309	95	11	all	all	DET
biotropia-309	95	12	statistical	statistical	ADJ
biotropia-309	95	13	analyses	analysis	NOUN
biotropia-309	95	14	.	.	PUNCT
biotropia-309	96	1	all	all	DET
biotropia-309	96	2	diversities	diversity	NOUN
biotropia-309	96	3	measured	measure	VERB
biotropia-309	96	4	were	be	AUX
biotropia-309	96	5	computed	compute	VERB
biotropia-309	96	6	with	with	ADP
biotropia-309	96	7	estimates	estimate	NOUN
biotropia-309	96	8	version	version	NOUN
biotropia-309	96	9	7.00	7.00	NUM
biotropia-309	96	10	program	program	NOUN
biotropia-309	96	11	(	(	PUNCT
biotropia-309	96	12	colwell	colwell	PROPN
biotropia-309	96	13	2004	2004	NUM
biotropia-309	96	14	)	)	PUNCT
biotropia-309	96	15	by	by	ADP
biotropia-309	96	16	randomizing	randomize	VERB
biotropia-309	96	17	the	the	DET
biotropia-309	96	18	ranking	ranking	NOUN
biotropia-309	96	19	of	of	ADP
biotropia-309	96	20	samples	sample	NOUN
biotropia-309	96	21	50	50	NUM
biotropia-309	96	22	times	time	NOUN
biotropia-309	96	23	.	.	PUNCT
biotropia-309	97	1	only	only	ADV
biotropia-309	97	2	species	specie	NOUN
biotropia-309	97	3	sampled	sample	VERB
biotropia-309	97	4	in	in	ADP
biotropia-309	97	5	both	both	DET
biotropia-309	97	6	sample	sample	NOUN
biotropia-309	97	7	periods	period	NOUN
biotropia-309	97	8	(	(	PUNCT
biotropia-309	97	9	2009	2009	NUM
biotropia-309	97	10	and	and	CCONJ
biotropia-309	97	11	2012	2012	NUM
biotropia-309	97	12	)	)	PUNCT
biotropia-309	97	13	were	be	AUX
biotropia-309	97	14	analyzed	analyze	VERB
biotropia-309	97	15	.	.	PUNCT
biotropia-309	98	1	the	the	DET
biotropia-309	98	2	effectiveness	effectiveness	NOUN
biotropia-309	98	3	of	of	ADP
biotropia-309	98	4	each	each	DET
biotropia-309	98	5	dung	dung	NOUN
biotropia-309	98	6	beetles	beetle	NOUN
biotropia-309	98	7	species	specie	NOUN
biotropia-309	98	8	as	as	ADP
biotropia-309	98	9	bioindicators	bioindicator	NOUN
biotropia-309	98	10	were	be	AUX
biotropia-309	98	11	identified	identify	VERB
biotropia-309	98	12	for	for	ADP
biotropia-309	98	13	each	each	DET
biotropia-309	98	14	habitat	habitat	NOUN
biotropia-309	98	15	type	type	NOUN
biotropia-309	98	16	using	use	VERB
biotropia-309	98	17	the	the	DET
biotropia-309	98	18	indicator	indicator	NOUN
biotropia-309	98	19	value	value	NOUN
biotropia-309	98	20	(	(	PUNCT
biotropia-309	98	21	)	)	PUNCT
biotropia-309	98	22	method	method	NOUN
biotropia-309	98	23	(	(	PUNCT
biotropia-309	98	24	dufrene	dufrene	PROPN
biotropia-309	98	25	&	&	CCONJ
biotropia-309	98	26	legendre	legendre	PROPN
biotropia-309	98	27	1997).this	1997).this	PROPN
biotropia-309	98	28	method	method	NOUN
biotropia-309	98	29	combines	combine	VERB
biotropia-309	98	30	measures	measure	NOUN
biotropia-309	98	31	of	of	ADP
biotropia-309	98	32	specificity	specificity	NOUN
biotropia-309	98	33	and	and	CCONJ
biotropia-309	98	34	fidelity	fidelity	NOUN
biotropia-309	98	35	and	and	CCONJ
biotropia-309	98	36	provides	provide	VERB
biotropia-309	98	37	an	an	PRON
biotropia-309	98	38	for	for	ADP
biotropia-309	98	39	each	each	DET
biotropia-309	98	40	species	specie	NOUN
biotropia-309	98	41	,	,	PUNCT
biotropia-309	98	42	as	as	ADP
biotropia-309	98	43	a	a	DET
biotropia-309	98	44	percentage	percentage	NOUN
biotropia-309	98	45	(	(	PUNCT
biotropia-309	98	46	dufrene	dufrene	NOUN
biotropia-309	98	47	&	&	CCONJ
biotropia-309	98	48	legendre	legendre	PROPN
biotropia-309	98	49	1997	1997	NUM
biotropia-309	98	50	)	)	PUNCT
biotropia-309	98	51	.	.	PUNCT
biotropia-309	99	1	specificity	specificity	NOUN
biotropia-309	99	2	measure	measure	NOUN
biotropia-309	99	3	:	:	PUNCT
biotropia-309	100	1	=	=	PUNCT
biotropia-309	100	2	/	/	SYM
biotropia-309	100	3	where	where	SCONJ
biotropia-309	100	4	is	be	AUX
biotropia-309	100	5	the	the	DET
biotropia-309	100	6	mean	mean	ADJ
biotropia-309	100	7	number	number	NOUN
biotropia-309	100	8	of	of	ADP
biotropia-309	100	9	species	specie	NOUN
biotropia-309	100	10	across	across	ADP
biotropia-309	100	11	sites	site	NOUN
biotropia-309	100	12	of	of	ADP
biotropia-309	100	13	group	group	NOUN
biotropia-309	100	14	,	,	PUNCT
biotropia-309	100	15	and	and	CCONJ
biotropia-309	100	16	is	be	AUX
biotropia-309	100	17	the	the	DET
biotropia-309	100	18	sum	sum	NOUN
biotropia-309	100	19	of	of	ADP
biotropia-309	100	20	the	the	DET
biotropia-309	100	21	mean	mean	ADJ
biotropia-309	100	22	numbers	number	NOUN
biotropia-309	100	23	of	of	ADP
biotropia-309	100	24	individuals	individual	NOUN
biotropia-309	100	25	of	of	ADP
biotropia-309	100	26	species	specie	NOUN
biotropia-309	100	27	over	over	ADP
biotropia-309	100	28	all	all	DET
biotropia-309	100	29	groups	group	NOUN
biotropia-309	100	30	.	.	PUNCT
biotropia-309	101	1	fidelity	fidelity	NOUN
biotropia-309	101	2	measure	measure	NOUN
biotropia-309	101	3	:	:	PUNCT
biotropia-309	102	1	=	=	PUNCT
biotropia-309	102	2	/	/	SYM
biotropia-309	102	3	where	where	SCONJ
biotropia-309	102	4	is	be	AUX
biotropia-309	102	5	the	the	DET
biotropia-309	102	6	number	number	NOUN
biotropia-309	102	7	of	of	ADP
biotropia-309	102	8	sites	site	NOUN
biotropia-309	102	9	in	in	ADP
biotropia-309	102	10	cluster	cluster	NOUN
biotropia-309	102	11	(	(	PUNCT
biotropia-309	102	12	habitat	habitat	PROPN
biotropia-309	102	13	)	)	PUNCT
biotropia-309	102	14	j	j	NOUN
biotropia-309	102	15	where	where	SCONJ
biotropia-309	102	16	species	specie	NOUN
biotropia-309	102	17	is	be	AUX
biotropia-309	102	18	present	present	ADJ
biotropia-309	102	19	,	,	PUNCT
biotropia-309	102	20	and	and	CCONJ
biotropia-309	102	21	is	be	AUX
biotropia-309	102	22	the	the	DET
biotropia-309	102	23	total	total	ADJ
biotropia-309	102	24	number	number	NOUN
biotropia-309	102	25	of	of	ADP
biotropia-309	102	26	sites	site	NOUN
biotropia-309	102	27	in	in	ADP
biotropia-309	102	28	that	that	DET
biotropia-309	102	29	cluster	cluster	NOUN
biotropia-309	102	30	.	.	PUNCT
biotropia-309	103	1	the	the	DET
biotropia-309	103	2	percentage	percentage	NOUN
biotropia-309	103	3	indicator	indicator	NOUN
biotropia-309	103	4	value	value	NOUN
biotropia-309	103	5	for	for	ADP
biotropia-309	103	6	species	specie	NOUN
biotropia-309	103	7	in	in	ADP
biotropia-309	103	8	cluster	cluster	NOUN
biotropia-309	103	9	(	(	PUNCT
biotropia-309	103	10	habitat	habitat	PROPN
biotropia-309	103	11	)	)	PUNCT
biotropia-309	103	12	is	be	AUX
biotropia-309	103	13	then	then	ADV
biotropia-309	103	14	:	:	PUNCT
biotropia-309	103	15	=	=	PUNCT
biotropia-309	103	16	×	×	PROPN
biotropia-309	103	17	×	×	NOUN
biotropia-309	103	18	100	100	NUM
biotropia-309	103	19	.	.	PUNCT
biotropia-309	104	1	the	the	DET
biotropia-309	104	2	indicator	indicator	NOUN
biotropia-309	104	3	values	value	NOUN
biotropia-309	104	4	are	be	AUX
biotropia-309	104	5	the	the	DET
biotropia-309	104	6	highest	high	ADJ
biotropia-309	104	7	(	(	PUNCT
biotropia-309	104	8	100	100	NUM
biotropia-309	104	9	)	)	PUNCT
biotropia-309	104	10	when	when	SCONJ
biotropia-309	104	11	all	all	DET
biotropia-309	104	12	individuals	individual	NOUN
biotropia-309	104	13	of	of	ADP
biotropia-309	104	14	a	a	DET
biotropia-309	104	15	species	specie	NOUN
biotropia-309	104	16	are	be	AUX
biotropia-309	104	17	found	find	VERB
biotropia-309	104	18	in	in	ADP
biotropia-309	104	19	a	a	DET
biotropia-309	104	20	single	single	ADJ
biotropia-309	104	21	habitat	habitat	NOUN
biotropia-309	104	22	(	(	PUNCT
biotropia-309	104	23	high	high	ADJ
biotropia-309	104	24	specificity	specificity	NOUN
biotropia-309	104	25	)	)	PUNCT
biotropia-309	104	26	and	and	CCONJ
biotropia-309	104	27	when	when	SCONJ
biotropia-309	104	28	the	the	DET
biotropia-309	104	29	species	specie	NOUN
biotropia-309	104	30	occurs	occur	VERB
biotropia-309	104	31	in	in	ADP
biotropia-309	104	32	all	all	DET
biotropia-309	104	33	samples	sample	NOUN
biotropia-309	104	34	of	of	ADP
biotropia-309	104	35	that	that	DET
biotropia-309	104	36	habitat	habitat	NOUN
biotropia-309	104	37	(	(	PUNCT
biotropia-309	104	38	high	high	ADJ
biotropia-309	104	39	fidelity	fidelity	NOUN
biotropia-309	104	40	)	)	PUNCT
biotropia-309	104	41	.	.	PUNCT
biotropia-309	105	1	species	specie	NOUN
biotropia-309	105	2	with	with	ADP
biotropia-309	105	3	between	between	ADP
biotropia-309	105	4	50	50	NUM
biotropia-309	105	5	%	%	NOUN
biotropia-309	105	6	to	to	ADP
biotropia-309	105	7	less	less	ADJ
biotropia-309	105	8	than	than	ADP
biotropia-309	105	9	70	70	NUM
biotropia-309	105	10	%	%	NOUN
biotropia-309	105	11	is	be	AUX
biotropia-309	105	12	categorized	categorize	VERB
biotropia-309	105	13	as	as	ADP
biotropia-309	105	14	the	the	DET
biotropia-309	105	15	detector	detector	NOUN
biotropia-309	105	16	or	or	CCONJ
biotropia-309	105	17	generalist	generalist	ADJ
biotropia-309	105	18	species	specie	NOUN
biotropia-309	105	19	while	while	SCONJ
biotropia-309	105	20	those	those	DET
biotropia-309	105	21	species	specie	NOUN
biotropia-309	105	22	with	with	ADP
biotropia-309	105	23	significant	significant	ADJ
biotropia-309	105	24	of	of	ADP
biotropia-309	105	25	greater	great	ADJ
biotropia-309	105	26	than	than	ADP
biotropia-309	105	27	70	70	NUM
biotropia-309	105	28	%	%	NOUN
biotropia-309	105	29	were	be	AUX
biotropia-309	105	30	regarded	regard	VERB
biotropia-309	105	31	as	as	ADP
biotropia-309	105	32	characteristic	characteristic	ADJ
biotropia-309	105	33	indicator	indicator	NOUN
biotropia-309	105	34	species	specie	NOUN
biotropia-309	105	35	for	for	ADP
biotropia-309	105	36	the	the	DET
biotropia-309	105	37	particular	particular	ADJ
biotropia-309	105	38	habitat	habitat	NOUN
biotropia-309	105	39	type	type	NOUN
biotropia-309	105	40	(	(	PUNCT
biotropia-309	105	41	mc.geoch	mc.geoch	NOUN
biotropia-309	105	42	.	.	PUNCT
biotropia-309	105	43	2002	2002	NUM
biotropia-309	105	44	)	)	PUNCT
biotropia-309	105	45	.	.	PUNCT
biotropia-309	106	1	a	a	DET
biotropia-309	106	2	total	total	NOUN
biotropia-309	106	3	of	of	ADP
biotropia-309	106	4	1996	1996	NUM
biotropia-309	106	5	dung	dung	NOUN
biotropia-309	106	6	beetles	beetle	NOUN
biotropia-309	106	7	specimens	specimen	NOUN
biotropia-309	106	8	were	be	AUX
biotropia-309	106	9	collected	collect	VERB
biotropia-309	106	10	during	during	ADP
biotropia-309	106	11	the	the	DET
biotropia-309	106	12	study	study	NOUN
biotropia-309	106	13	period	period	NOUN
biotropia-309	106	14	.	.	PUNCT
biotropia-309	107	1	they	they	PRON
biotropia-309	107	2	belongs	belong	VERB
biotropia-309	107	3	to	to	ADP
biotropia-309	107	4	four	four	NUM
biotropia-309	107	5	genera	genera	NOUN
biotropia-309	107	6	(	(	PUNCT
biotropia-309	107	7	dominated	dominate	VERB
biotropia-309	107	8	by	by	ADP
biotropia-309	107	9	)	)	PUNCT
biotropia-309	107	10	and	and	CCONJ
biotropia-309	107	11	28	28	NUM
biotropia-309	107	12	species	specie	NOUN
biotropia-309	107	13	(	(	PUNCT
biotropia-309	107	14	for	for	ADP
biotropia-309	107	15	complete	complete	ADJ
biotropia-309	107	16	species	species	NOUN
biotropia-309	107	17	list	list	NOUN
biotropia-309	107	18	see	see	VERB
biotropia-309	107	19	shahabuddin	shahabuddin	VERB
biotropia-309	107	20	2013	2013	NUM
biotropia-309	107	21	)	)	PUNCT
biotropia-309	107	22	.	.	PUNCT
biotropia-309	108	1	however	however	ADV
biotropia-309	108	2	,	,	PUNCT
biotropia-309	108	3	only	only	ADV
biotropia-309	108	4	16	16	NUM
biotropia-309	108	5	species	specie	NOUN
biotropia-309	108	6	were	be	AUX
biotropia-309	108	7	recorded	record	VERB
biotropia-309	108	8	in	in	ADP
biotropia-309	108	9	both	both	DET
biotropia-309	108	10	sample	sample	NOUN
biotropia-309	108	11	periods	period	NOUN
biotropia-309	108	12	(	(	PUNCT
biotropia-309	108	13	2009	2009	NUM
biotropia-309	108	14	and	and	CCONJ
biotropia-309	108	15	2012	2012	NUM
biotropia-309	108	16	)	)	PUNCT
biotropia-309	108	17	.	.	PUNCT
biotropia-309	109	1	the	the	DET
biotropia-309	109	2	diversity	diversity	NOUN
biotropia-309	109	3	of	of	ADP
biotropia-309	109	4	dung	dung	NOUN
biotropia-309	109	5	beetles	beetle	NOUN
biotropia-309	109	6	changed	change	VERB
biotropia-309	109	7	from	from	ADP
biotropia-309	109	8	natural	natural	ADJ
biotropia-309	109	9	forest	forest	NOUN
biotropia-309	109	10	,	,	PUNCT
biotropia-309	109	11	to	to	ADP
biotropia-309	109	12	agroforestry	agroforestry	NOUN
biotropia-309	109	13	cacao	cacao	NOUN
biotropia-309	109	14	and	and	CCONJ
biotropia-309	109	15	to	to	PART
biotropia-309	109	16	open	open	VERB
biotropia-309	109	17	area	area	NOUN
biotropia-309	109	18	.	.	PUNCT
biotropia-309	110	1	interestingly	interestingly	ADV
biotropia-309	110	2	,	,	PUNCT
biotropia-309	110	3	the	the	DET
biotropia-309	110	4	diversity	diversity	NOUN
biotropia-309	110	5	of	of	ADP
biotropia-309	110	6	dung	dung	NOUN
biotropia-309	110	7	beetles	beetle	NOUN
biotropia-309	110	8	in	in	ADP
biotropia-309	110	9	both	both	DET
biotropia-309	110	10	forest	forest	NOUN
biotropia-309	110	11	types	type	NOUN
biotropia-309	110	12	and	and	CCONJ
biotropia-309	110	13	the	the	DET
biotropia-309	110	14	two	two	NUM
biotropia-309	110	15	types	type	NOUN
biotropia-309	110	16	of	of	ADP
biotropia-309	110	17	cacao	cacao	NOUN
biotropia-309	110	18	plantations	plantation	NOUN
biotropia-309	110	19	tend	tend	VERB
biotropia-309	110	20	to	to	PART
biotropia-309	110	21	be	be	AUX
biotropia-309	110	22	similar	similar	ADJ
biotropia-309	110	23	but	but	CCONJ
biotropia-309	110	24	significantly	significantly	ADV
biotropia-309	110	25	higher	high	ADJ
biotropia-309	110	26	than	than	ADP
biotropia-309	110	27	that	that	PRON
biotropia-309	110	28	in	in	ADP
biotropia-309	110	29	open	open	ADJ
biotropia-309	110	30	cultivated	cultivate	VERB
biotropia-309	110	31	area	area	NOUN
biotropia-309	110	32	(	(	PUNCT
biotropia-309	110	33	table	table	NOUN
biotropia-309	110	34	1	1	NUM
biotropia-309	110	35	)	)	PUNCT
biotropia-309	110	36	.	.	PUNCT
biotropia-309	111	1	hence	hence	ADV
biotropia-309	111	2	,	,	PUNCT
biotropia-309	111	3	this	this	DET
biotropia-309	111	4	study	study	NOUN
biotropia-309	111	5	showed	show	VERB
biotropia-309	111	6	that	that	SCONJ
biotropia-309	111	7	secondary	secondary	ADJ
biotropia-309	111	8	forest	forest	NOUN
biotropia-309	111	9	and	and	CCONJ
biotropia-309	111	10	agroforestry	agroforestry	NOUN
biotropia-309	111	11	system	system	NOUN
biotropia-309	111	12	may	may	AUX
biotropia-309	111	13	support	support	VERB
biotropia-309	111	14	a	a	DET
biotropia-309	111	15	high	high	ADJ
biotropia-309	111	16	portion	portion	NOUN
biotropia-309	111	17	of	of	ADP
biotropia-309	111	18	tropical	tropical	ADJ
biotropia-309	111	19	dung	dung	NOUN
biotropia-309	111	20	beetles	beetle	NOUN
biotropia-309	111	21	species	specie	NOUN
biotropia-309	111	22	than	than	ADP
biotropia-309	111	23	in	in	ADP
biotropia-309	111	24	the	the	DET
biotropia-309	111	25	bare	bare	ADJ
biotropia-309	111	26	land	land	NOUN
biotropia-309	111	27	and	and	CCONJ
biotropia-309	111	28	thereby	thereby	ADV
biotropia-309	111	29	in	in	ADP
biotropia-309	111	30	line	line	NOUN
biotropia-309	111	31	with	with	ADP
biotropia-309	111	32	the	the	DET
biotropia-309	111	33	findings	finding	NOUN
biotropia-309	111	34	by	by	ADP
biotropia-309	111	35	nichols	nichols	PROPN
biotropia-309	111	36	.	.	PUNCT
biotropia-309	112	1	(	(	PUNCT
biotropia-309	112	2	2007	2007	NUM
biotropia-309	112	3	)	)	PUNCT
biotropia-309	112	4	and	and	CCONJ
biotropia-309	112	5	our	our	PRON
biotropia-309	112	6	previous	previous	ADJ
biotropia-309	112	7	study	study	NOUN
biotropia-309	112	8	(	(	PUNCT
biotropia-309	112	9	shahabuddin	shahabuddin	NOUN
biotropia-309	112	10	.	.	PUNCT
biotropia-309	112	11	2010	2010	NUM
biotropia-309	112	12	)	)	PUNCT
biotropia-309	112	13	.	.	PUNCT
biotropia-309	113	1	the	the	DET
biotropia-309	113	2	fact	fact	NOUN
biotropia-309	113	3	that	that	SCONJ
biotropia-309	113	4	agroforestry	agroforestry	NOUN
biotropia-309	113	5	system	system	NOUN
biotropia-309	113	6	has	have	AUX
biotropia-309	113	7	high	high	ADJ
biotropia-309	113	8	potency	potency	NOUN
biotropia-309	113	9	for	for	ADP
biotropia-309	113	10	conserving	conserve	VERB
biotropia-309	113	11	high	high	ADJ
biotropia-309	113	12	biodiversity	biodiversity	NOUN
biotropia-309	113	13	supported	support	VERB
biotropia-309	113	14	by	by	ADP
biotropia-309	113	15	previous	previous	ADJ
biotropia-309	113	16	study	study	NOUN
biotropia-309	113	17	(	(	PUNCT
biotropia-309	113	18	e.g.	e.g.	ADV
biotropia-309	113	19	mcnely	mcnely	PROPN
biotropia-309	113	20	&	&	CCONJ
biotropia-309	113	21	scroth	scroth	PROPN
biotropia-309	113	22	2006	2006	NUM
biotropia-309	113	23	;	;	PUNCT
biotropia-309	113	24	schulze	schulze	NOUN
biotropia-309	113	25	.	.	PUNCT
biotropia-309	113	26	2010	2010	NUM
biotropia-309	113	27	)	)	PUNCT
biotropia-309	113	28	.	.	PUNCT
biotropia-309	114	1	however	however	ADV
biotropia-309	114	2	,	,	PUNCT
biotropia-309	114	3	the	the	DET
biotropia-309	114	4	results	result	NOUN
biotropia-309	114	5	may	may	AUX
biotropia-309	114	6	also	also	ADV
biotropia-309	114	7	be	be	AUX
biotropia-309	114	8	related	relate	VERB
biotropia-309	114	9	with	with	ADP
biotropia-309	114	10	the	the	DET
biotropia-309	114	11	spatial	spatial	ADJ
biotropia-309	114	12	distribution	distribution	NOUN
biotropia-309	114	13	of	of	ADP
biotropia-309	114	14	our	our	PRON
biotropia-309	114	15	study	study	NOUN
biotropia-309	114	16	sites	site	NOUN
biotropia-309	114	17	.	.	PUNCT
biotropia-309	115	1	the	the	DET
biotropia-309	115	2	agroforestry	agroforestry	NOUN
biotropia-309	115	3	cacao	cacao	NOUN
biotropia-309	115	4	sites	site	NOUN
biotropia-309	115	5	were	be	AUX
biotropia-309	115	6	closer	close	ADJ
biotropia-309	115	7	to	to	ADP
biotropia-309	115	8	the	the	DET
biotropia-309	115	9	natural	natural	ADJ
biotropia-309	115	10	and	and	CCONJ
biotropia-309	115	11	secondary	secondary	ADJ
biotropia-309	115	12	forest	forest	NOUN
biotropia-309	115	13	than	than	ADP
biotropia-309	115	14	to	to	ADP
biotropia-309	115	15	the	the	DET
biotropia-309	115	16	open	open	ADJ
biotropia-309	115	17	area	area	NOUN
biotropia-309	115	18	and	and	CCONJ
biotropia-309	115	19	has	have	VERB
biotropia-309	115	20	a	a	DET
biotropia-309	115	21	high	high	ADJ
biotropia-309	115	22	opportunity	opportunity	NOUN
biotropia-309	115	23	to	to	PART
biotropia-309	115	24	be	be	AUX
biotropia-309	115	25	colonized	colonize	VERB
biotropia-309	115	26	by	by	ADP
biotropia-309	115	27	dung	dung	NOUN
biotropia-309	115	28	beetles	beetle	NOUN
biotropia-309	115	29	coming	come	VERB
biotropia-309	115	30	from	from	ADP
biotropia-309	115	31	the	the	DET
biotropia-309	115	32	forest	forest	NOUN
biotropia-309	115	33	sites	site	NOUN
biotropia-309	115	34	.	.	PUNCT
biotropia-309	116	1	therefore	therefore	ADV
biotropia-309	116	2	,	,	PUNCT
biotropia-309	116	3	the	the	DET
biotropia-309	116	4	high	high	ADJ
biotropia-309	116	5	diversity	diversity	NOUN
biotropia-309	116	6	of	of	ADP
biotropia-309	116	7	dung	dung	NOUN
biotropia-309	116	8	beetles	beetle	NOUN
biotropia-309	116	9	at	at	ADP
biotropia-309	116	10	the	the	DET
biotropia-309	116	11	cacao	cacao	NOUN
biotropia-309	116	12	agroforestry	agroforestry	NOUN
biotropia-309	116	13	system	system	NOUN
biotropia-309	116	14	is	be	AUX
biotropia-309	116	15	also	also	ADV
biotropia-309	116	16	related	relate	VERB
biotropia-309	116	17	to	to	ADP
biotropia-309	116	18	their	their	PRON
biotropia-309	116	19	close	close	ADJ
biotropia-309	116	20	proximity	proximity	NOUN
biotropia-309	116	21	to	to	ADP
biotropia-309	116	22	the	the	DET
biotropia-309	116	23	forest	forest	NOUN
biotropia-309	116	24	sites	site	NOUN
biotropia-309	116	25	.	.	PUNCT
biotropia-309	117	1	it	it	PRON
biotropia-309	117	2	has	have	VERB
biotropia-309	117	3	n	n	NUM
biotropia-309	117	4	indval	indval	ADJ
biotropia-309	117	5	indicator	indicator	NOUN
biotropia-309	117	6	value	value	VERB
biotropia-309	117	7	a	a	DET
biotropia-309	117	8	nindividuals	nindividual	NOUN
biotropia-309	117	9	nindividuals	nindividual	NOUN
biotropia-309	117	10	nindividuals	nindividual	NOUN
biotropia-309	118	1	i	i	PRON
biotropia-309	118	2	j	j	PROPN
biotropia-309	118	3	nindividuals	nindividual	NOUN
biotropia-309	118	4	i	i	PRON
biotropia-309	118	5	bij	bij	VERB
biotropia-309	118	6	nsites	nsites	PROPN
biotropia-309	118	7	nsites	nsites	PROPN
biotropia-309	118	8	nsites	nsite	NOUN
biotropia-309	118	9	i	i	PRON
biotropia-309	118	10	nsites	nsite	VERB
biotropia-309	118	11	i	i	PRON
biotropia-309	118	12	j	j	PROPN
biotropia-309	118	13	indval	indval	VERB
biotropia-309	118	14	a	a	DET
biotropia-309	118	15	b	b	PROPN
biotropia-309	118	16	indval	indval	ADJ
biotropia-309	118	17	indval	indval	NOUN
biotropia-309	118	18	et	et	PROPN
biotropia-309	118	19	al	al	PROPN
biotropia-309	118	20	onthophagus	onthophagus	VERB
biotropia-309	119	1	et	et	PROPN
biotropia-309	119	2	al	al	PROPN
biotropia-309	119	3	et	et	PROPN
biotropia-309	119	4	al	al	PROPN
biotropia-309	119	5	et	et	PROPN
biotropia-309	119	6	al	al	PROPN
biotropia-309	120	1	ij	ij	INTJ
biotropia-309	120	2	ij	ij	INTJ
biotropia-309	121	1	i	i	PRON
biotropia-309	121	2	ij	ij	INTJ
biotropia-309	122	1	i	i	PRON
biotropia-309	122	2	ij	ij	INTJ
biotropia-309	123	1	j	j	PROPN
biotropia-309	123	2	ij	ij	INTJ
biotropia-309	123	3	j	j	PROPN
biotropia-309	123	4	ij	ij	PROPN
biotropia-309	123	5	ijij	ijij	ADJ
biotropia-309	123	6	results	result	NOUN
biotropia-309	123	7	and	and	CCONJ
biotropia-309	123	8	discussions	discussion	NOUN
biotropia-309	123	9	diversity	diversity	NOUN
biotropia-309	123	10	of	of	ADP
biotropia-309	123	11	dung	dung	NOUN
biotropia-309	123	12	beetles	beetle	NOUN
biotropia-309	123	13	52	52	NUM
biotropia-309	123	14	biotropia	biotropia	ADJ
biotropia-309	123	15	vol	vol	NOUN
biotropia-309	123	16	.	.	PROPN
biotropia-309	123	17	21	21	NUM
biotropia-309	124	1	no	no	NOUN
biotropia-309	124	2	.	.	NOUN
biotropia-309	124	3	1	1	NUM
biotropia-309	124	4	,	,	PUNCT
biotropia-309	124	5	2014	2014	NUM
biotropia-309	124	6	been	be	AUX
biotropia-309	124	7	reported	report	VERB
biotropia-309	124	8	that	that	SCONJ
biotropia-309	124	9	neighboring	neighboring	NOUN
biotropia-309	124	10	forest	forest	NOUN
biotropia-309	124	11	or	or	CCONJ
biotropia-309	124	12	isolation	isolation	NOUN
biotropia-309	124	13	from	from	ADP
biotropia-309	124	14	forest	forest	NOUN
biotropia-309	124	15	may	may	AUX
biotropia-309	124	16	determine	determine	VERB
biotropia-309	124	17	insect	insect	NOUN
biotropia-309	124	18	communities	community	NOUN
biotropia-309	124	19	in	in	ADP
biotropia-309	124	20	tropical	tropical	ADJ
biotropia-309	124	21	land	land	NOUN
biotropia-309	124	22	-	-	PUNCT
biotropia-309	124	23	use	use	NOUN
biotropia-309	124	24	systems	system	NOUN
biotropia-309	124	25	(	(	PUNCT
biotropia-309	124	26	tscharntke	tscharntke	NOUN
biotropia-309	124	27	.	.	PUNCT
biotropia-309	125	1	2005	2005	NUM
biotropia-309	125	2	;	;	PUNCT
biotropia-309	125	3	klein	klein	PROPN
biotropia-309	125	4	.	.	PUNCT
biotropia-309	126	1	2006	2006	NUM
biotropia-309	126	2	)	)	PUNCT
biotropia-309	126	3	.	.	PUNCT
biotropia-309	127	1	agroforestry	agroforestry	NOUN
biotropia-309	127	2	systems	system	NOUN
biotropia-309	127	3	can	can	AUX
biotropia-309	127	4	be	be	AUX
biotropia-309	127	5	part	part	NOUN
biotropia-309	127	6	of	of	ADP
biotropia-309	127	7	the	the	DET
biotropia-309	127	8	habitat	habitat	NOUN
biotropia-309	127	9	for	for	ADP
biotropia-309	127	10	many	many	ADJ
biotropia-309	127	11	forest	forest	NOUN
biotropia-309	127	12	species	specie	NOUN
biotropia-309	127	13	using	use	VERB
biotropia-309	127	14	it	it	PRON
biotropia-309	127	15	for	for	ADP
biotropia-309	127	16	foraging	forage	VERB
biotropia-309	127	17	,	,	PUNCT
biotropia-309	127	18	but	but	CCONJ
biotropia-309	127	19	they	they	PRON
biotropia-309	127	20	may	may	AUX
biotropia-309	127	21	also	also	ADV
biotropia-309	127	22	harbour	harbour	VERB
biotropia-309	127	23	largely	largely	ADV
biotropia-309	127	24	independent	independent	ADJ
biotropia-309	127	25	populations	population	NOUN
biotropia-309	127	26	.	.	PUNCT
biotropia-309	128	1	nonetheless	nonetheless	ADV
biotropia-309	128	2	,	,	PUNCT
biotropia-309	128	3	this	this	DET
biotropia-309	128	4	study	study	NOUN
biotropia-309	128	5	suggests	suggest	VERB
biotropia-309	128	6	that	that	SCONJ
biotropia-309	128	7	the	the	DET
biotropia-309	128	8	preservation	preservation	NOUN
biotropia-309	128	9	of	of	ADP
biotropia-309	128	10	environmental	environmental	ADJ
biotropia-309	128	11	heterogeneity	heterogeneity	NOUN
biotropia-309	128	12	should	should	AUX
biotropia-309	128	13	be	be	AUX
biotropia-309	128	14	encouraged	encourage	VERB
biotropia-309	128	15	for	for	ADP
biotropia-309	128	16	conserving	conserve	VERB
biotropia-309	128	17	dung	dung	NOUN
biotropia-309	128	18	beetles	beetle	NOUN
biotropia-309	128	19	in	in	ADP
biotropia-309	128	20	the	the	DET
biotropia-309	128	21	llnp	llnp	NOUN
biotropia-309	128	22	,	,	PUNCT
biotropia-309	128	23	central	central	ADJ
biotropia-309	128	24	sulawesi	sulawesi	NOUN
biotropia-309	128	25	.	.	PUNCT
biotropia-309	129	1	this	this	PRON
biotropia-309	129	2	is	be	AUX
biotropia-309	129	3	important	important	ADJ
biotropia-309	129	4	because	because	SCONJ
biotropia-309	129	5	high	high	ADJ
biotropia-309	129	6	diversity	diversity	NOUN
biotropia-309	129	7	of	of	ADP
biotropia-309	129	8	dung	dung	NOUN
biotropia-309	129	9	beetles	beetle	NOUN
biotropia-309	129	10	in	in	ADP
biotropia-309	129	11	tropical	tropical	ADJ
biotropia-309	129	12	land	land	NOUN
biotropia-309	129	13	-	-	PUNCT
biotropia-309	129	14	use	use	NOUN
biotropia-309	129	15	will	will	AUX
biotropia-309	129	16	enhance	enhance	VERB
biotropia-309	129	17	its	its	PRON
biotropia-309	129	18	ecosystem	ecosystem	NOUN
biotropia-309	129	19	function	function	NOUN
biotropia-309	129	20	and	and	CCONJ
biotropia-309	129	21	this	this	DET
biotropia-309	129	22	ecological	ecological	ADJ
biotropia-309	129	23	services	service	NOUN
biotropia-309	129	24	will	will	AUX
biotropia-309	129	25	be	be	AUX
biotropia-309	129	26	diminished	diminish	VERB
biotropia-309	129	27	by	by	ADP
biotropia-309	129	28	increasing	increase	VERB
biotropia-309	129	29	human	human	ADJ
biotropia-309	129	30	dominated	dominate	VERB
biotropia-309	129	31	land	land	NOUN
biotropia-309	129	32	-	-	PUNCT
biotropia-309	129	33	use	use	NOUN
biotropia-309	129	34	(	(	PUNCT
biotropia-309	129	35	shahabuddin	shahabuddin	VERB
biotropia-309	129	36	2011	2011	NUM
biotropia-309	129	37	;	;	PUNCT
biotropia-309	129	38	slade	slade	PROPN
biotropia-309	129	39	.	.	PUNCT
biotropia-309	130	1	2011	2011	NUM
biotropia-309	130	2	;	;	PUNCT
biotropia-309	130	3	kudavidanage	kudavidanage	NOUN
biotropia-309	130	4	2012	2012	NUM
biotropia-309	130	5	)	)	PUNCT
biotropia-309	130	6	.	.	PUNCT
biotropia-309	131	1	in	in	ADP
biotropia-309	131	2	more	more	ADV
biotropia-309	131	3	natural	natural	ADJ
biotropia-309	131	4	and	and	CCONJ
biotropia-309	131	5	heterogeneous	heterogeneous	ADJ
biotropia-309	131	6	habitats	habitat	NOUN
biotropia-309	131	7	,	,	PUNCT
biotropia-309	131	8	such	such	ADJ
biotropia-309	131	9	as	as	ADP
biotropia-309	131	10	natural	natural	ADJ
biotropia-309	131	11	forest	forest	NOUN
biotropia-309	131	12	and	and	CCONJ
biotropia-309	131	13	agroforestry	agroforestry	NOUN
biotropia-309	131	14	system	system	NOUN
biotropia-309	131	15	,	,	PUNCT
biotropia-309	131	16	dung	dung	NOUN
biotropia-309	131	17	removal	removal	NOUN
biotropia-309	131	18	,	,	PUNCT
biotropia-309	131	19	biological	biological	ADJ
biotropia-309	131	20	control	control	NOUN
biotropia-309	131	21	and	and	CCONJ
biotropia-309	131	22	seed	seed	NOUN
biotropia-309	131	23	dispersal	dispersal	NOUN
biotropia-309	131	24	activities	activity	NOUN
biotropia-309	131	25	of	of	ADP
biotropia-309	131	26	dung	dung	NOUN
biotropia-309	131	27	beetles	beetle	NOUN
biotropia-309	131	28	were	be	AUX
biotropia-309	131	29	higher	high	ADJ
biotropia-309	131	30	than	than	ADP
biotropia-309	131	31	in	in	ADP
biotropia-309	131	32	homogenous	homogenous	ADJ
biotropia-309	131	33	or	or	CCONJ
biotropia-309	131	34	disturbed	disturbed	ADJ
biotropia-309	131	35	habitats	habitat	NOUN
biotropia-309	131	36	(	(	PUNCT
biotropia-309	131	37	slade	slade	PROPN
biotropia-309	131	38	.	.	PUNCT
biotropia-309	132	1	2007	2007	NUM
biotropia-309	132	2	;	;	PUNCT
biotropia-309	132	3	nichols	nichols	PROPN
biotropia-309	132	4	.	.	PUNCT
biotropia-309	132	5	2008	2008	NUM
biotropia-309	132	6	;	;	PUNCT
biotropia-309	132	7	shahabuddin	shahabuddin	VERB
biotropia-309	132	8	2011	2011	NUM
biotropia-309	132	9	;	;	PUNCT
biotropia-309	132	10	slade	slade	PROPN
biotropia-309	132	11	.	.	PUNCT
biotropia-309	133	1	2011	2011	NUM
biotropia-309	133	2	)	)	PUNCT
biotropia-309	133	3	.	.	PUNCT
biotropia-309	134	1	because	because	SCONJ
biotropia-309	134	2	the	the	DET
biotropia-309	134	3	dung	dung	NOUN
biotropia-309	134	4	beetles	beetle	VERB
biotropia-309	134	5	diversity	diversity	NOUN
biotropia-309	134	6	at	at	ADP
biotropia-309	134	7	the	the	DET
biotropia-309	134	8	forest	forest	NOUN
biotropia-309	134	9	sites	site	NOUN
biotropia-309	134	10	(	(	PUNCT
biotropia-309	134	11	natural	natural	ADJ
biotropia-309	134	12	forest	forest	NOUN
biotropia-309	134	13	and	and	CCONJ
biotropia-309	134	14	secondary	secondary	ADJ
biotropia-309	134	15	forest	forest	NOUN
biotropia-309	134	16	)	)	PUNCT
biotropia-309	134	17	and	and	CCONJ
biotropia-309	134	18	the	the	DET
biotropia-309	134	19	cacao	cacao	NOUN
biotropia-309	134	20	plantation	plantation	NOUN
biotropia-309	134	21	sites	site	NOUN
biotropia-309	134	22	(	(	PUNCT
biotropia-309	134	23	agroforestry	agroforestry	NOUN
biotropia-309	134	24	cacao	cacao	NOUN
biotropia-309	134	25	and	and	CCONJ
biotropia-309	134	26	cacao	cacao	NOUN
biotropia-309	134	27	plantation	plantation	NOUN
biotropia-309	134	28	)	)	PUNCT
biotropia-309	134	29	were	be	AUX
biotropia-309	134	30	similar	similar	ADJ
biotropia-309	134	31	to	to	ADP
biotropia-309	134	32	each	each	DET
biotropia-309	134	33	other	other	ADJ
biotropia-309	134	34	but	but	CCONJ
biotropia-309	134	35	significantly	significantly	ADV
biotropia-309	134	36	higher	high	ADJ
biotropia-309	134	37	than	than	ADP
biotropia-309	134	38	in	in	ADP
biotropia-309	134	39	the	the	DET
biotropia-309	134	40	open	open	ADJ
biotropia-309	134	41	area	area	NOUN
biotropia-309	134	42	(	(	PUNCT
biotropia-309	134	43	table	table	NOUN
biotropia-309	134	44	1	1	NUM
biotropia-309	134	45	)	)	PUNCT
biotropia-309	134	46	,	,	PUNCT
biotropia-309	134	47	the	the	DET
biotropia-309	134	48	indicator	indicator	NOUN
biotropia-309	134	49	species	specie	VERB
biotropia-309	134	50	analysis	analysis	NOUN
biotropia-309	134	51	land	land	NOUN
biotropia-309	134	52	-	-	PUNCT
biotropia-309	134	53	use	use	NOUN
biotropia-309	134	54	type	type	NOUN
biotropia-309	134	55	is	be	AUX
biotropia-309	134	56	only	only	ADV
biotropia-309	134	57	classified	classify	VERB
biotropia-309	134	58	into	into	ADP
biotropia-309	134	59	three	three	NUM
biotropia-309	134	60	groups	group	NOUN
biotropia-309	134	61	,	,	PUNCT
biotropia-309	134	62	:	:	PUNCT
biotropia-309	134	63	forest	forest	NOUN
biotropia-309	134	64	,	,	PUNCT
biotropia-309	134	65	agroforestry	agroforestry	NOUN
biotropia-309	134	66	cacao	cacao	NOUN
biotropia-309	134	67	and	and	CCONJ
biotropia-309	134	68	open	open	ADJ
biotropia-309	134	69	area	area	NOUN
biotropia-309	134	70	.	.	PUNCT
biotropia-309	135	1	the	the	DET
biotropia-309	135	2	high	high	ADJ
biotropia-309	135	3	similarity	similarity	NOUN
biotropia-309	135	4	of	of	ADP
biotropia-309	135	5	environmental	environmental	ADJ
biotropia-309	135	6	parameter	parameter	NOUN
biotropia-309	135	7	measured	measure	VERB
biotropia-309	135	8	among	among	ADP
biotropia-309	135	9	these	these	DET
biotropia-309	135	10	three	three	NUM
biotropia-309	135	11	groups	group	NOUN
biotropia-309	135	12	of	of	ADP
biotropia-309	135	13	land	land	NOUN
biotropia-309	135	14	-	-	PUNCT
biotropia-309	135	15	use	use	NOUN
biotropia-309	135	16	types	type	NOUN
biotropia-309	135	17	was	be	AUX
biotropia-309	135	18	supported	support	VERB
biotropia-309	135	19	by	by	ADP
biotropia-309	135	20	anova	anova	PROPN
biotropia-309	135	21	showing	show	VERB
biotropia-309	135	22	that	that	PRON
biotropia-309	135	23	temperature	temperature	NOUN
biotropia-309	135	24	and	and	CCONJ
biotropia-309	135	25	herb	herb	NOUN
biotropia-309	135	26	coverage	coverage	NOUN
biotropia-309	135	27	decreased	decrease	VERB
biotropia-309	135	28	from	from	ADP
biotropia-309	135	29	the	the	DET
biotropia-309	135	30	natural	natural	ADJ
biotropia-309	135	31	and	and	CCONJ
biotropia-309	135	32	secondary	secondary	ADJ
biotropia-309	135	33	forest	forest	NOUN
biotropia-309	135	34	to	to	ADP
biotropia-309	135	35	the	the	DET
biotropia-309	135	36	open	open	ADJ
biotropia-309	135	37	area	area	NOUN
biotropia-309	135	38	while	while	SCONJ
biotropia-309	135	39	canopy	canopy	NOUN
biotropia-309	135	40	and	and	CCONJ
biotropia-309	135	41	humidity	humidity	NOUN
biotropia-309	135	42	showed	show	VERB
biotropia-309	135	43	a	a	DET
biotropia-309	135	44	reverse	reverse	ADJ
biotropia-309	135	45	pattern	pattern	NOUN
biotropia-309	135	46	.	.	PUNCT
biotropia-309	136	1	however	however	ADV
biotropia-309	136	2	,	,	PUNCT
biotropia-309	136	3	a	a	DET
biotropia-309	136	4	highly	highly	ADV
biotropia-309	136	5	significant	significant	ADJ
biotropia-309	136	6	difference	difference	NOUN
biotropia-309	136	7	of	of	ADP
biotropia-309	136	8	environmental	environmental	ADJ
biotropia-309	136	9	parameter	parameter	NOUN
biotropia-309	136	10	measured	measure	VERB
biotropia-309	136	11	was	be	AUX
biotropia-309	136	12	only	only	ADV
biotropia-309	136	13	recorded	record	VERB
biotropia-309	136	14	in	in	ADP
biotropia-309	136	15	the	the	DET
biotropia-309	136	16	open	open	ADJ
biotropia-309	136	17	area	area	NOUN
biotropia-309	136	18	(	(	PUNCT
biotropia-309	136	19	fig	fig	NOUN
biotropia-309	136	20	.	.	PUNCT
biotropia-309	137	1	1	1	NUM
biotropia-309	137	2	)	)	PUNCT
biotropia-309	137	3	.	.	PUNCT
biotropia-309	138	1	this	this	DET
biotropia-309	138	2	land	land	NOUN
biotropia-309	138	3	-	-	PUNCT
biotropia-309	138	4	use	use	NOUN
biotropia-309	138	5	type	type	NOUN
biotropia-309	138	6	grouping	grouping	NOUN
biotropia-309	138	7	was	be	AUX
biotropia-309	138	8	also	also	ADV
biotropia-309	138	9	supported	support	VERB
biotropia-309	138	10	by	by	ADP
biotropia-309	138	11	the	the	DET
biotropia-309	138	12	ordination	ordination	NOUN
biotropia-309	138	13	technique	technique	NOUN
biotropia-309	138	14	using	use	VERB
biotropia-309	138	15	multidimensional	multidimensional	ADJ
biotropia-309	138	16	scaling	scaling	NOUN
biotropia-309	138	17	(	(	PUNCT
biotropia-309	138	18	fig	fig	NOUN
biotropia-309	138	19	.	.	PUNCT
biotropia-309	139	1	2	2	NUM
biotropia-309	139	2	)	)	PUNCT
biotropia-309	139	3	.	.	PUNCT
biotropia-309	140	1	based	base	VERB
biotropia-309	140	2	on	on	ADP
biotropia-309	140	3	analysis	analysis	NOUN
biotropia-309	140	4	of	of	ADP
biotropia-309	140	5	16	16	NUM
biotropia-309	140	6	dung	dung	NOUN
biotropia-309	140	7	beetles	beetle	NOUN
biotropia-309	140	8	species	specie	NOUN
biotropia-309	140	9	collected	collect	VERB
biotropia-309	140	10	in	in	ADP
biotropia-309	140	11	two	two	NUM
biotropia-309	140	12	sampling	sample	VERB
biotropia-309	140	13	years	year	NOUN
biotropia-309	140	14	(	(	PUNCT
biotropia-309	140	15	2009	2009	NUM
biotropia-309	140	16	and	and	CCONJ
biotropia-309	140	17	2012	2012	NUM
biotropia-309	140	18	)	)	PUNCT
biotropia-309	140	19	,	,	PUNCT
biotropia-309	140	20	this	this	DET
biotropia-309	140	21	study	study	NOUN
biotropia-309	140	22	recorded	record	VERB
biotropia-309	140	23	four	four	NUM
biotropia-309	140	24	species	specie	NOUN
biotropia-309	140	25	having	have	VERB
biotropia-309	140	26	less	less	ADJ
biotropia-309	140	27	than	than	ADP
biotropia-309	140	28	70	70	NUM
biotropia-309	140	29	%	%	NOUN
biotropia-309	140	30	and	and	CCONJ
biotropia-309	140	31	therefore	therefore	ADV
biotropia-309	140	32	,	,	PUNCT
biotropia-309	140	33	can	can	AUX
biotropia-309	140	34	be	be	AUX
biotropia-309	140	35	used	use	VERB
biotropia-309	140	36	as	as	ADP
biotropia-309	140	37	indicator	indicator	NOUN
biotropia-309	140	38	(	(	PUNCT
biotropia-309	140	39	characteristic	characteristic	ADJ
biotropia-309	140	40	)	)	PUNCT
biotropia-309	140	41	species	specie	NOUN
biotropia-309	140	42	that	that	PRON
biotropia-309	140	43	are	be	AUX
biotropia-309	140	44	:	:	PUNCT
biotropia-309	140	45	at	at	ADP
biotropia-309	140	46	forest	forest	NOUN
biotropia-309	140	47	sites	site	NOUN
biotropia-309	140	48	,	,	PUNCT
biotropia-309	140	49	at	at	ADP
biotropia-309	140	50	cacao	cacao	NOUN
biotropia-309	140	51	plantation	plantation	NOUN
biotropia-309	140	52	,	,	PUNCT
biotropia-309	140	53	as	as	ADV
biotropia-309	140	54	well	well	ADV
biotropia-309	140	55	as	as	ADP
biotropia-309	140	56	and	and	CCONJ
biotropia-309	140	57	at	at	ADP
biotropia-309	140	58	open	open	ADJ
biotropia-309	140	59	area	area	NOUN
biotropia-309	140	60	(	(	PUNCT
biotropia-309	140	61	table	table	NOUN
biotropia-309	140	62	2	2	NUM
biotropia-309	140	63	)	)	PUNCT
biotropia-309	140	64	.	.	PUNCT
biotropia-309	141	1	and	and	CCONJ
biotropia-309	141	2	are	be	AUX
biotropia-309	141	3	suggested	suggest	VERB
biotropia-309	141	4	to	to	PART
biotropia-309	141	5	be	be	AUX
biotropia-309	141	6	used	use	VERB
biotropia-309	141	7	as	as	ADP
biotropia-309	141	8	indicators	indicator	NOUN
biotropia-309	141	9	of	of	ADP
biotropia-309	141	10	cooler	cool	ADJ
biotropia-309	141	11	and	and	CCONJ
biotropia-309	141	12	shaded	shaded	ADJ
biotropia-309	141	13	habitats	habitat	NOUN
biotropia-309	141	14	such	such	ADJ
biotropia-309	141	15	as	as	ADP
biotropia-309	141	16	forest	forest	NOUN
biotropia-309	141	17	sites	site	NOUN
biotropia-309	141	18	and	and	CCONJ
biotropia-309	141	19	cacao	cacao	NOUN
biotropia-309	141	20	agroforestry	agroforestry	NOUN
biotropia-309	141	21	et	et	NOUN
biotropia-309	141	22	al	al	PROPN
biotropia-309	141	23	et	et	PROPN
biotropia-309	141	24	al	al	PROPN
biotropia-309	141	25	et	et	PROPN
biotropia-309	141	26	al	al	PROPN
biotropia-309	141	27	et	et	PROPN
biotropia-309	141	28	al	al	PROPN
biotropia-309	141	29	.	.	PROPN
biotropia-309	142	1	et	et	PROPN
biotropia-309	142	2	al	al	PROPN
biotropia-309	142	3	et	et	PROPN
biotropia-309	142	4	al	al	PROPN
biotropia-309	142	5	et	et	PROPN
biotropia-309	142	6	al	al	PROPN
biotropia-309	142	7	i.e	i.e	PROPN
biotropia-309	142	8	indval	indval	ADJ
biotropia-309	142	9	indval	indval	ADJ
biotropia-309	142	10	copris	copris	PROPN
biotropia-309	142	11	saundersi	saundersi	PROPN
biotropia-309	142	12	onthophagus	onthophagus	VERB
biotropia-309	142	13	forsteni	forsteni	PROPN
biotropia-309	142	14	o.	o.	PROPN
biotropia-309	142	15	trituber	trituber	PROPN
biotropia-309	142	16	o.	o.	PROPN
biotropia-309	142	17	limbatus	limbatus	PROPN
biotropia-309	142	18	copris	copris	PROPN
biotropia-309	142	19	saundersi	saundersi	PROPN
biotropia-309	142	20	onthophagus	onthophagus	VERB
biotropia-309	142	21	forsteni	forsteni	PROPN
biotropia-309	142	22	indicator	indicator	NOUN
biotropia-309	142	23	species	species	NOUN
biotropia-309	142	24	table	table	VERB
biotropia-309	142	25	1	1	NUM
biotropia-309	142	26	.	.	PUNCT
biotropia-309	142	27	diversity	diversity	NOUN
biotropia-309	142	28	of	of	ADP
biotropia-309	142	29	dung	dung	NOUN
biotropia-309	142	30	beetles	beetle	NOUN
biotropia-309	142	31	at	at	ADP
biotropia-309	142	32	five	five	NUM
biotropia-309	142	33	habitat	habitat	ADJ
biotropia-309	142	34	type	type	NOUN
biotropia-309	142	35	.	.	PUNCT
biotropia-309	143	1	nf	nf	NOUN
biotropia-309	143	2	=	=	SYM
biotropia-309	143	3	natural	natural	ADJ
biotropia-309	143	4	forest	forest	NOUN
biotropia-309	143	5	;	;	PUNCT
biotropia-309	143	6	sf	sf	NOUN
biotropia-309	143	7	=	=	PUNCT
biotropia-309	143	8	secondary	secondary	ADJ
biotropia-309	143	9	forest	forest	NOUN
biotropia-309	143	10	;	;	PUNCT
biotropia-309	143	11	ac	ac	PROPN
biotropia-309	143	12	=	=	PUNCT
biotropia-309	143	13	cacao	cacao	NOUN
biotropia-309	143	14	agroforestry	agroforestry	NOUN
biotropia-309	143	15	system	system	NOUN
biotropia-309	143	16	;	;	PUNCT
biotropia-309	143	17	cp	cp	NOUN
biotropia-309	143	18	=	=	NOUN
biotropia-309	143	19	cacao	cacao	NOUN
biotropia-309	143	20	plantation	plantation	NOUN
biotropia-309	143	21	;	;	PUNCT
biotropia-309	144	1	oa	oa	PROPN
biotropia-309	144	2	=	=	PUNCT
biotropia-309	144	3	open	open	ADJ
biotropia-309	144	4	area	area	NOUN
biotropia-309	144	5	land	land	NOUN
biotropia-309	144	6	-	-	PUNCT
biotropia-309	144	7	use	use	NOUN
biotropia-309	144	8	type	type	NOUN
biotropia-309	144	9	estimated	estimate	VERB
biotropia-309	144	10	species	specie	NOUN
biotropia-309	144	11	richness	richness	NOUN
biotropia-309	144	12	(	(	PUNCT
biotropia-309	144	13	jack-2	jack-2	NOUN
biotropia-309	144	14	)	)	PUNCT
biotropia-309	144	15	(	(	PUNCT
biotropia-309	144	16	f4,10=6.27	f4,10=6.27	PROPN
biotropia-309	144	17	,	,	PUNCT
biotropia-309	144	18	p	p	X
biotropia-309	144	19	<	<	NOUN
biotropia-309	144	20	0.05	0.05	NUM
biotropia-309	144	21	)	)	PUNCT
biotropia-309	144	22	number	number	NOUN
biotropia-309	144	23	of	of	ADP
biotropia-309	144	24	species	specie	NOUN
biotropia-309	144	25	recorded	record	VERB
biotropia-309	144	26	f4,10	f4,10	NOUN
biotropia-309	144	27	=	=	SYM
biotropia-309	144	28	3·60	3·60	NUM
biotropia-309	144	29	,	,	PUNCT
biotropia-309	144	30	p	p	X
biotropia-309	144	31	<	<	X
biotropia-309	144	32	0·005	0·005	NUM
biotropia-309	145	1	h	h	NOUN
biotropia-309	145	2	'	'	PUNCT
biotropia-309	145	3	simpsom	simpsom	NOUN
biotropia-309	145	4	nf	nf	NUM
biotropia-309	145	5	14.3a	14.3a	NUM
biotropia-309	145	6	9ab	9ab	NOUN
biotropia-309	145	7	1.6	1.6	NUM
biotropia-309	145	8	4.2	4.2	NUM
biotropia-309	145	9	sf	sf	NOUN
biotropia-309	145	10	14.4a	14.4a	NUM
biotropia-309	145	11	10.7ab	10.7ab	NUM
biotropia-309	145	12	1.7	1.7	NUM
biotropia-309	145	13	4.2	4.2	NUM
biotropia-309	145	14	ac	ac	PROPN
biotropia-309	145	15	15a	15a	NOUN
biotropia-309	145	16	11a	11a	NOUN
biotropia-309	145	17	1.7	1.7	NUM
biotropia-309	145	18	4.3	4.3	NUM
biotropia-309	145	19	cp	cp	INTJ
biotropia-309	145	20	13.6a	13.6a	NUM
biotropia-309	145	21	10ab	10ab	NOUN
biotropia-309	145	22	1.7	1.7	NUM
biotropia-309	145	23	4.5	4.5	NUM
biotropia-309	145	24	oa	oa	NOUN
biotropia-309	145	25	8.7b	8.7b	NUM
biotropia-309	145	26	6.3b	6.3b	NUM
biotropia-309	145	27	1.3	1.3	NUM
biotropia-309	145	28	3.1	3.1	NUM
biotropia-309	145	29	53	53	NUM
biotropia-309	145	30	effectiveness	effectiveness	NOUN
biotropia-309	145	31	of	of	ADP
biotropia-309	145	32	dung	dung	NOUN
biotropia-309	145	33	beetles	beetle	NOUN
biotropia-309	145	34	as	as	ADP
biotropia-309	145	35	bioindicators	bioindicator	NOUN
biotropia-309	145	36	of	of	ADP
biotropia-309	145	37	environmental	environmental	ADJ
biotropia-309	145	38	changes	change	NOUN
biotropia-309	145	39	in	in	ADP
biotropia-309	145	40	land	land	NOUN
biotropia-309	145	41	shahabuddin	shahabuddin	NOUN
biotropia-309	145	42	–	–	PUNCT
biotropia-309	145	43	et	et	PROPN
biotropia-309	145	44	al	al	PROPN
biotropia-309	145	45	.	.	PROPN
biotropia-309	145	46	system	system	NOUN
biotropia-309	145	47	,	,	PUNCT
biotropia-309	145	48	while	while	SCONJ
biotropia-309	145	49	and	and	CCONJ
biotropia-309	145	50	most	most	ADV
biotropia-309	145	51	likely	likely	ADV
biotropia-309	145	52	indicate	indicate	VERB
biotropia-309	145	53	warmer	warm	ADJ
biotropia-309	145	54	and	and	CCONJ
biotropia-309	145	55	unshaded	unshaded	ADJ
biotropia-309	145	56	habitats	habitat	NOUN
biotropia-309	145	57	(	(	PUNCT
biotropia-309	145	58	e.g.	e.g.	ADV
biotropia-309	145	59	open	open	ADJ
biotropia-309	145	60	cultivated	cultivate	VERB
biotropia-309	145	61	area	area	NOUN
biotropia-309	145	62	)	)	PUNCT
biotropia-309	145	63	.	.	PUNCT
biotropia-309	146	1	these	these	DET
biotropia-309	146	2	characteristic	characteristic	ADJ
biotropia-309	146	3	species	specie	NOUN
biotropia-309	146	4	(	(	PUNCT
biotropia-309	146	5	of	of	ADP
biotropia-309	146	6	>	>	X
biotropia-309	146	7	70	70	NUM
biotropia-309	146	8	%	%	NOUN
biotropia-309	146	9	)	)	PUNCT
biotropia-309	146	10	are	be	AUX
biotropia-309	146	11	unlikely	unlikely	ADJ
biotropia-309	146	12	to	to	PART
biotropia-309	146	13	move	move	VERB
biotropia-309	146	14	from	from	ADP
biotropia-309	146	15	their	their	PRON
biotropia-309	146	16	requisite	requisite	NOUN
biotropia-309	146	17	to	to	ADP
biotropia-309	146	18	other	other	ADJ
biotropia-309	146	19	habitat	habitat	NOUN
biotropia-309	146	20	types	type	NOUN
biotropia-309	146	21	,	,	PUNCT
biotropia-309	146	22	even	even	ADV
biotropia-309	146	23	under	under	ADP
biotropia-309	146	24	changing	change	VERB
biotropia-309	146	25	conditions	condition	NOUN
biotropia-309	146	26	within	within	ADP
biotropia-309	146	27	this	this	DET
biotropia-309	146	28	habitat	habitat	NOUN
biotropia-309	146	29	.	.	PUNCT
biotropia-309	147	1	accordingly	accordingly	ADV
biotropia-309	147	2	,	,	PUNCT
biotropia-309	147	3	populations	population	NOUN
biotropia-309	147	4	of	of	ADP
biotropia-309	147	5	these	these	DET
biotropia-309	147	6	species	specie	NOUN
biotropia-309	147	7	need	need	VERB
biotropia-309	147	8	only	only	ADV
biotropia-309	147	9	to	to	PART
biotropia-309	147	10	be	be	AUX
biotropia-309	147	11	monitored	monitor	VERB
biotropia-309	147	12	within	within	ADP
biotropia-309	147	13	the	the	DET
biotropia-309	147	14	specific	specific	ADJ
biotropia-309	147	15	habitat	habitat	NOUN
biotropia-309	147	16	.	.	PUNCT
biotropia-309	148	1	beside	beside	ADP
biotropia-309	148	2	indicator	indicator	PROPN
biotropia-309	148	3	species	specie	NOUN
biotropia-309	148	4	,	,	PUNCT
biotropia-309	148	5	this	this	DET
biotropia-309	148	6	study	study	NOUN
biotropia-309	148	7	has	have	AUX
biotropia-309	148	8	recorded	record	VERB
biotropia-309	148	9	several	several	ADJ
biotropia-309	148	10	generalist	generalist	ADJ
biotropia-309	148	11	or	or	CCONJ
biotropia-309	148	12	moderate	moderate	ADJ
biotropia-309	148	13	species	specie	NOUN
biotropia-309	148	14	with	with	ADP
biotropia-309	148	15	between	between	ADP
biotropia-309	148	16	50	50	NUM
biotropia-309	148	17	%	%	NOUN
biotropia-309	148	18	to	to	ADP
biotropia-309	148	19	less	less	ADJ
biotropia-309	148	20	than	than	ADP
biotropia-309	148	21	70	70	NUM
biotropia-309	148	22	%	%	NOUN
biotropia-309	148	23	.	.	PUNCT
biotropia-309	149	1	these	these	DET
biotropia-309	149	2	species	specie	NOUN
biotropia-309	149	3	were	be	AUX
biotropia-309	149	4	therefore	therefore	ADV
biotropia-309	149	5	,	,	PUNCT
biotropia-309	149	6	not	not	PART
biotropia-309	149	7	characteristic	characteristic	ADJ
biotropia-309	149	8	species	specie	NOUN
biotropia-309	149	9	,	,	PUNCT
biotropia-309	149	10	as	as	SCONJ
biotropia-309	149	11	they	they	PRON
biotropia-309	149	12	do	do	AUX
biotropia-309	149	13	not	not	PART
biotropia-309	149	14	have	have	VERB
biotropia-309	149	15	high	high	ADJ
biotropia-309	149	16	of	of	ADP
biotropia-309	149	17	more	more	ADJ
biotropia-309	149	18	than	than	ADP
biotropia-309	149	19	70	70	NUM
biotropia-309	149	20	%	%	NOUN
biotropia-309	149	21	for	for	ADP
biotropia-309	149	22	any	any	DET
biotropia-309	149	23	particular	particular	ADJ
biotropia-309	149	24	habitat	habitat	NOUN
biotropia-309	149	25	.	.	PUNCT
biotropia-309	150	1	species	specie	NOUN
biotropia-309	150	2	meeting	meet	VERB
biotropia-309	150	3	these	these	DET
biotropia-309	150	4	criteria	criterion	NOUN
biotropia-309	150	5	are	be	AUX
biotropia-309	150	6	unlikely	unlikely	ADJ
biotropia-309	150	7	to	to	PART
biotropia-309	150	8	respond	respond	VERB
biotropia-309	150	9	very	very	ADV
biotropia-309	150	10	rapidly	rapidly	ADV
biotropia-309	150	11	to	to	ADP
biotropia-309	150	12	changing	change	VERB
biotropia-309	150	13	habitat	habitat	ADJ
biotropia-309	150	14	conditions	condition	NOUN
biotropia-309	150	15	.	.	PUNCT
biotropia-309	151	1	they	they	PRON
biotropia-309	151	2	can	can	AUX
biotropia-309	151	3	invade	invade	VERB
biotropia-309	151	4	either	either	CCONJ
biotropia-309	151	5	close	close	ADJ
biotropia-309	151	6	canopy	canopy	NOUN
biotropia-309	151	7	or	or	CCONJ
biotropia-309	151	8	moist	moist	NOUN
biotropia-309	151	9	habitat	habitat	NOUN
biotropia-309	151	10	e.g.	e.g.	ADJ
biotropia-309	151	11	natural	natural	ADJ
biotropia-309	151	12	forest	forest	NOUN
biotropia-309	151	13	,	,	PUNCT
biotropia-309	151	14	cacao	cacao	NOUN
biotropia-309	151	15	agroforestry	agroforestry	NOUN
biotropia-309	151	16	or	or	CCONJ
biotropia-309	151	17	cacao	cacao	NOUN
biotropia-309	151	18	plantation	plantation	NOUN
biotropia-309	151	19	but	but	CCONJ
biotropia-309	151	20	also	also	ADV
biotropia-309	151	21	open	open	ADJ
biotropia-309	151	22	or	or	CCONJ
biotropia-309	151	23	warmer	warm	ADJ
biotropia-309	151	24	environment	environment	NOUN
biotropia-309	151	25	such	such	ADJ
biotropia-309	151	26	us	us	PROPN
biotropia-309	151	27	open	open	ADJ
biotropia-309	151	28	area	area	NOUN
biotropia-309	151	29	.	.	PUNCT
biotropia-309	152	1	furthermore	furthermore	ADV
biotropia-309	152	2	,	,	PUNCT
biotropia-309	152	3	these	these	DET
biotropia-309	152	4	species	specie	NOUN
biotropia-309	152	5	are	be	AUX
biotropia-309	152	6	less	less	ADV
biotropia-309	152	7	likely	likely	ADJ
biotropia-309	152	8	to	to	PART
biotropia-309	152	9	become	become	VERB
biotropia-309	152	10	more	more	ADV
biotropia-309	152	11	vulnerable	vulnerable	ADJ
biotropia-309	152	12	than	than	ADP
biotropia-309	152	13	indicator	indicator	NOUN
biotropia-309	152	14	species	specie	NOUN
biotropia-309	152	15	,	,	PUNCT
biotropia-309	152	16	because	because	SCONJ
biotropia-309	152	17	a	a	DET
biotropia-309	152	18	variety	variety	NOUN
biotropia-309	152	19	of	of	ADP
biotropia-309	152	20	habitats	habitat	NOUN
biotropia-309	152	21	or	or	CCONJ
biotropia-309	152	22	ecological	ecological	ADJ
biotropia-309	152	23	states	state	NOUN
biotropia-309	152	24	,	,	PUNCT
biotropia-309	152	25	rather	rather	ADV
biotropia-309	152	26	than	than	ADP
biotropia-309	152	27	only	only	ADV
biotropia-309	152	28	a	a	DET
biotropia-309	152	29	single	single	ADJ
biotropia-309	152	30	one	one	NOUN
biotropia-309	152	31	,	,	PUNCT
biotropia-309	152	32	provide	provide	VERB
biotropia-309	152	33	suitable	suitable	ADJ
biotropia-309	152	34	resources	resource	NOUN
biotropia-309	152	35	for	for	ADP
biotropia-309	152	36	them	they	PRON
biotropia-309	152	37	and	and	CCONJ
biotropia-309	152	38	accordingly	accordingly	ADV
biotropia-309	152	39	this	this	DET
biotropia-309	152	40	group	group	NOUN
biotropia-309	152	41	of	of	ADP
biotropia-309	152	42	species	specie	NOUN
biotropia-309	152	43	will	will	AUX
biotropia-309	152	44	be	be	AUX
biotropia-309	152	45	useful	useful	ADJ
biotropia-309	152	46	for	for	ADP
biotropia-309	152	47	longer	long	ADJ
biotropia-309	152	48	-	-	PUNCT
biotropia-309	152	49	term	term	NOUN
biotropia-309	152	50	monitoring	monitoring	NOUN
biotropia-309	152	51	.	.	PUNCT
biotropia-309	153	1	o.	o.	PROPN
biotropia-309	153	2	trituber	trituber	PROPN
biotropia-309	153	3	o.	o.	PROPN
biotropia-309	153	4	limbatus	limbatus	PROPN
biotropia-309	153	5	indval	indval	PROPN
biotropia-309	153	6	indval	indval	PROPN
biotropia-309	153	7	indval	indval	ADJ
biotropia-309	153	8	figure	figure	NOUN
biotropia-309	153	9	1	1	NUM
biotropia-309	153	10	.	.	PUNCT
biotropia-309	154	1	the	the	DET
biotropia-309	154	2	effects	effect	NOUN
biotropia-309	154	3	of	of	ADP
biotropia-309	154	4	land	land	NOUN
biotropia-309	154	5	use	use	NOUN
biotropia-309	154	6	change	change	NOUN
biotropia-309	154	7	on	on	ADP
biotropia-309	154	8	a	a	DET
biotropia-309	154	9	)	)	PUNCT
biotropia-309	154	10	temperature	temperature	NOUN
biotropia-309	154	11	(	(	PUNCT
biotropia-309	154	12	anova	anova	PROPN
biotropia-309	154	13	:	:	PUNCT
biotropia-309	154	14	f(4	f(4	PROPN
biotropia-309	154	15	,	,	PUNCT
biotropia-309	154	16	10)=25.15	10)=25.15	NOUN
biotropia-309	154	17	,	,	PUNCT
biotropia-309	154	18	p<0.01	p<0.01	PROPN
biotropia-309	154	19	)	)	PUNCT
biotropia-309	154	20	,	,	PUNCT
biotropia-309	154	21	b	b	X
biotropia-309	154	22	)	)	PUNCT
biotropia-309	154	23	relative	relative	ADJ
biotropia-309	154	24	humidity	humidity	NOUN
biotropia-309	154	25	(	(	PUNCT
biotropia-309	154	26	anova	anova	PROPN
biotropia-309	154	27	:	:	PUNCT
biotropia-309	154	28	f(4,10)=62.88	f(4,10)=62.88	PROPN
biotropia-309	154	29	,	,	PUNCT
biotropia-309	154	30	p<0.01	p<0.01	PROPN
biotropia-309	154	31	)	)	PUNCT
biotropia-309	154	32	,	,	PUNCT
biotropia-309	154	33	c	c	X
biotropia-309	154	34	)	)	PUNCT
biotropia-309	154	35	canopy	canopy	NOUN
biotropia-309	154	36	cover	cover	NOUN
biotropia-309	154	37	(	(	PUNCT
biotropia-309	154	38	anova	anova	PROPN
biotropia-309	154	39	:	:	PUNCT
biotropia-309	154	40	f(4,10)=139.81	f(4,10)=139.81	PROPN
biotropia-309	154	41	,	,	PUNCT
biotropia-309	154	42	p<0.01	p<0.01	PROPN
biotropia-309	154	43	)	)	PUNCT
biotropia-309	154	44	,	,	PUNCT
biotropia-309	154	45	and	and	CCONJ
biotropia-309	154	46	d	d	X
biotropia-309	154	47	)	)	PUNCT
biotropia-309	154	48	herb	herb	NOUN
biotropia-309	154	49	layer	layer	NOUN
biotropia-309	154	50	coverage	coverage	NOUN
biotropia-309	154	51	(	(	PUNCT
biotropia-309	154	52	anova	anova	PROPN
biotropia-309	154	53	:	:	PUNCT
biotropia-309	154	54	f(4	f(4	NOUN
biotropia-309	154	55	,	,	PUNCT
biotropia-309	154	56	14)=52.15,p<0.01	14)=52.15,p<0.01	NUM
biotropia-309	154	57	)	)	PUNCT
biotropia-309	154	58	.	.	PUNCT
biotropia-309	155	1	all	all	DET
biotropia-309	155	2	variables	variable	NOUN
biotropia-309	155	3	were	be	AUX
biotropia-309	155	4	averaged	average	VERB
biotropia-309	155	5	per	per	ADP
biotropia-309	155	6	land	land	NOUN
biotropia-309	155	7	-	-	PUNCT
biotropia-309	155	8	use	use	NOUN
biotropia-309	155	9	type	type	NOUN
biotropia-309	155	10	.	.	PUNCT
biotropia-309	156	1	nf=	nf=	PROPN
biotropia-309	156	2	natural	natural	ADJ
biotropia-309	156	3	forest	forest	NOUN
biotropia-309	156	4	;	;	PUNCT
biotropia-309	156	5	sf=	sf=	PROPN
biotropia-309	156	6	secondary	secondary	ADJ
biotropia-309	156	7	forest	forest	NOUN
biotropia-309	156	8	;	;	PUNCT
biotropia-309	156	9	ac=	ac=	PROPN
biotropia-309	156	10	agroforestry	agroforestry	NOUN
biotropia-309	156	11	cacao	cacao	NOUN
biotropia-309	156	12	;	;	PUNCT
biotropia-309	156	13	cp=	cp=	NOUN
biotropia-309	156	14	cacao	cacao	NOUN
biotropia-309	156	15	plantation	plantation	NOUN
biotropia-309	156	16	;	;	PUNCT
biotropia-309	156	17	oa=	oa=	NOUN
biotropia-309	156	18	open	open	ADJ
biotropia-309	156	19	area	area	NOUN
biotropia-309	156	20	.	.	PUNCT
biotropia-309	157	1	significant	significant	ADJ
biotropia-309	157	2	differences	difference	NOUN
biotropia-309	157	3	between	between	ADP
biotropia-309	157	4	habitat	habitat	ADJ
biotropia-309	157	5	types	type	NOUN
biotropia-309	157	6	were	be	AUX
biotropia-309	157	7	indicated	indicate	VERB
biotropia-309	157	8	by	by	ADP
biotropia-309	157	9	different	different	ADJ
biotropia-309	157	10	letters	letter	NOUN
biotropia-309	157	11	over	over	ADP
biotropia-309	157	12	the	the	DET
biotropia-309	157	13	standard	standard	ADJ
biotropia-309	157	14	error	error	NOUN
biotropia-309	157	15	(	(	PUNCT
biotropia-309	157	16	based	base	VERB
biotropia-309	157	17	on	on	ADP
biotropia-309	157	18	tukey	tukey	PROPN
biotropia-309	157	19	's	's	PART
biotropia-309	157	20	hsd	hsd	NOUN
biotropia-309	157	21	post	post	ADJ
biotropia-309	157	22	-	-	ADJ
biotropia-309	157	23	hoc	hoc	ADJ
biotropia-309	157	24	test	test	NOUN
biotropia-309	157	25	)	)	PUNCT
biotropia-309	157	26	54	54	NUM
biotropia-309	157	27	biotropia	biotropia	NOUN
biotropia-309	157	28	vol	vol	NOUN
biotropia-309	157	29	.	.	PROPN
biotropia-309	158	1	21	21	NUM
biotropia-309	159	1	no	no	NOUN
biotropia-309	159	2	.	.	NOUN
biotropia-309	159	3	1	1	NUM
biotropia-309	159	4	,	,	PUNCT
biotropia-309	159	5	2014	2014	NUM
biotropia-309	159	6	nf1	nf1	NOUN
biotropia-309	159	7	nf2	nf2	VERB
biotropia-309	159	8	nf3	nf3	PROPN
biotropia-309	159	9	sf1sf2sf3	sf1sf2sf3	NOUN
biotropia-309	159	10	ac1	ac1	PROPN
biotropia-309	159	11	ac2	ac2	PROPN
biotropia-309	159	12	ac3	ac3	PROPN
biotropia-309	159	13	cp1cp2cp3	cp1cp2cp3	PROPN
biotropia-309	159	14	oa1oa2oa3	oa1oa2oa3	PROPN
biotropia-309	159	15	-1.0	-1.0	PROPN
biotropia-309	159	16	-0.8	-0.8	PROPN
biotropia-309	159	17	-0.6	-0.6	PROPN
biotropia-309	159	18	-0.4	-0.4	X
biotropia-309	160	1	-0.2	-0.2	PROPN
biotropia-309	160	2	0.0	0.0	NUM
biotropia-309	160	3	0.2	0.2	NUM
biotropia-309	160	4	0.4	0.4	NUM
biotropia-309	160	5	0.6	0.6	NUM
biotropia-309	160	6	0.8	0.8	NUM
biotropia-309	160	7	dimension	dimension	NOUN
biotropia-309	160	8	2	2	NUM
biotropia-309	160	9	-1.2	-1.2	PROPN
biotropia-309	160	10	-1.0	-1.0	PROPN
biotropia-309	160	11	-0.8	-0.8	PROPN
biotropia-309	160	12	-0.6	-0.6	PROPN
biotropia-309	160	13	-0.4	-0.4	X
biotropia-309	160	14	-0.2	-0.2	PROPN
biotropia-309	160	15	0.0	0.0	NUM
biotropia-309	160	16	0.2	0.2	NUM
biotropia-309	160	17	0.4	0.4	NUM
biotropia-309	160	18	0.6	0.6	NUM
biotropia-309	160	19	0.8	0.8	NUM
biotropia-309	160	20	1.0	1.0	NUM
biotropia-309	160	21	1.2	1.2	NUM
biotropia-309	160	22	1.4	1.4	NUM
biotropia-309	160	23	1.6	1.6	NUM
biotropia-309	160	24	1.8	1.8	NUM
biotropia-309	160	25	d	d	NOUN
biotropia-309	160	26	i	i	PRON
biotropia-309	160	27	m	m	VERB
biotropia-309	160	28	e	e	NOUN
biotropia-309	160	29	n	n	PROPN
biotropia-309	160	30	s	s	PART
biotropia-309	160	31	io	io	X
biotropia-309	160	32	n	n	CCONJ
biotropia-309	160	33	1	1	NUM
biotropia-309	160	34	stress=0.007	stress=0.007	NOUN
biotropia-309	160	35	open	open	ADJ
biotropia-309	160	36	area	area	NOUN
biotropia-309	160	37	forest	forest	NOUN
biotropia-309	160	38	sites	site	NOUN
biotropia-309	160	39	agroforestry	agroforestry	NOUN
biotropia-309	160	40	cacao	cacao	NOUN
biotropia-309	160	41	figure	figure	NOUN
biotropia-309	160	42	2	2	NUM
biotropia-309	160	43	.	.	PUNCT
biotropia-309	160	44	multidimensional	multidimensional	ADJ
biotropia-309	160	45	scaling	scaling	NOUN
biotropia-309	160	46	(	(	PUNCT
biotropia-309	160	47	mds	mds	NOUN
biotropia-309	160	48	)	)	PUNCT
biotropia-309	160	49	placing	place	VERB
biotropia-309	160	50	the	the	DET
biotropia-309	160	51	five	five	NUM
biotropia-309	160	52	land	land	NOUN
biotropia-309	160	53	-	-	PUNCT
biotropia-309	160	54	use	use	NOUN
biotropia-309	160	55	type	type	NOUN
biotropia-309	160	56	studied	study	VERB
biotropia-309	160	57	in	in	ADP
biotropia-309	160	58	three	three	NUM
biotropia-309	160	59	groups	group	NOUN
biotropia-309	160	60	based	base	VERB
biotropia-309	160	61	on	on	ADP
biotropia-309	160	62	several	several	ADJ
biotropia-309	160	63	habitat	habitat	NOUN
biotropia-309	160	64	parameters	parameter	NOUN
biotropia-309	160	65	measured	measure	VERB
biotropia-309	160	66	.	.	PUNCT
biotropia-309	161	1	nf=	nf=	PROPN
biotropia-309	161	2	natural	natural	ADJ
biotropia-309	161	3	forest	forest	NOUN
biotropia-309	161	4	;	;	PUNCT
biotropia-309	161	5	sf=	sf=	PROPN
biotropia-309	161	6	secondary	secondary	ADJ
biotropia-309	161	7	forest	forest	NOUN
biotropia-309	161	8	;	;	PUNCT
biotropia-309	161	9	,	,	PUNCT
biotropia-309	161	10	ac=	ac=	PROPN
biotropia-309	161	11	agroforestry	agroforestry	NOUN
biotropia-309	161	12	cacao	cacao	NOUN
biotropia-309	161	13	;	;	PUNCT
biotropia-309	161	14	oa=	oa=	NOUN
biotropia-309	161	15	open	open	ADJ
biotropia-309	161	16	area	area	NOUN
biotropia-309	161	17	table	table	NOUN
biotropia-309	161	18	2	2	NUM
biotropia-309	161	19	.	.	PUNCT
biotropia-309	161	20	indicator	indicator	NOUN
biotropia-309	161	21	values	value	NOUN
biotropia-309	161	22	(	(	PUNCT
biotropia-309	161	23	percentage	percentage	NOUN
biotropia-309	161	24	)	)	PUNCT
biotropia-309	161	25	for	for	ADP
biotropia-309	161	26	dung	dung	NOUN
biotropia-309	161	27	beetle	beetle	NOUN
biotropia-309	161	28	species	specie	NOUN
biotropia-309	161	29	recorded	record	VERB
biotropia-309	161	30	in	in	ADP
biotropia-309	161	31	three	three	NUM
biotropia-309	161	32	group	group	NOUN
biotropia-309	161	33	of	of	ADP
biotropia-309	161	34	habitats	habitat	NOUN
biotropia-309	161	35	.	.	PUNCT
biotropia-309	162	1	species	specie	NOUN
biotropia-309	162	2	with	with	ADP
biotropia-309	162	3	>	>	SYM
biotropia-309	162	4	70	70	NUM
biotropia-309	162	5	%	%	NOUN
biotropia-309	162	6	is	be	AUX
biotropia-309	162	7	categorized	categorize	VERB
biotropia-309	162	8	as	as	ADP
biotropia-309	162	9	a	a	DET
biotropia-309	162	10	characteristic	characteristic	ADJ
biotropia-309	162	11	species	specie	NOUN
biotropia-309	162	12	(	(	PUNCT
biotropia-309	162	13	c	c	NOUN
biotropia-309	162	14	)	)	PUNCT
biotropia-309	162	15	while	while	SCONJ
biotropia-309	162	16	species	specie	NOUN
biotropia-309	162	17	with	with	ADP
biotropia-309	162	18	50	50	NUM
biotropia-309	162	19	≤	≤	NUM
biotropia-309	162	20	70	70	NUM
biotropia-309	162	21	%	%	NOUN
biotropia-309	162	22	is	be	AUX
biotropia-309	162	23	a	a	DET
biotropia-309	162	24	detector	detector	NOUN
biotropia-309	162	25	species	specie	NOUN
biotropia-309	162	26	(	(	PUNCT
biotropia-309	162	27	d	d	NOUN
biotropia-309	162	28	)	)	PUNCT
biotropia-309	162	29	of	of	ADP
biotropia-309	162	30	certain	certain	ADJ
biotropia-309	162	31	habitat	habitat	NOUN
biotropia-309	162	32	type	type	NOUN
biotropia-309	162	33	indval	indval	ADJ
biotropia-309	162	34	indval	indval	ADJ
biotropia-309	162	35	indval	indval	PROPN
biotropia-309	162	36	species	specie	NOUN
biotropia-309	162	37	*	*	PUNCT
biotropia-309	162	38	forest	forest	NOUN
biotropia-309	162	39	sites	site	NOUN
biotropia-309	162	40	agroforestry	agroforestry	NOUN
biotropia-309	162	41	cacao	cacao	PROPN
biotropia-309	162	42	open	open	ADJ
biotropia-309	162	43	area	area	NOUN
biotropia-309	162	44	species	specie	NOUN
biotropia-309	162	45	category	category	NOUN
biotropia-309	162	46	2009	2009	NUM
biotropia-309	162	47	2012	2012	NUM
biotropia-309	162	48	2009	2009	NUM
biotropia-309	162	49	2012	2012	NUM
biotropia-309	162	50	2009	2009	NUM
biotropia-309	162	51	2012	2012	NUM
biotropia-309	162	52	copris	copris	PROPN
biotropia-309	162	53	macacus	macacus	VERB
biotropia-309	162	54	67.4	67.4	NUM
biotropia-309	162	55	13.3	13.3	NUM
biotropia-309	162	56	22.6	22.6	NUM
biotropia-309	162	57	20.0	20.0	NUM
biotropia-309	162	58	0.0	0.0	NUM
biotropia-309	162	59	0.0	0.0	NUM
biotropia-309	162	60	copris	copris	PROPN
biotropia-309	162	61	punctulatus	punctulatus	NOUN
biotropia-309	162	62	12.1	12.1	NUM
biotropia-309	162	63	22.2	22.2	NUM
biotropia-309	162	64	30.3	30.3	NUM
biotropia-309	162	65	44.4	44.4	NUM
biotropia-309	162	66	24.2	24.2	NUM
biotropia-309	162	67	0.0	0.0	NUM
biotropia-309	162	68	copris	copris	NOUN
biotropia-309	162	69	saundersi	saundersi	NOUN
biotropia-309	162	70	95.8	95.8	NUM
biotropia-309	162	71	91.7	91.7	NUM
biotropia-309	162	72	0.0	0.0	NUM
biotropia-309	162	73	0.0	0.0	NUM
biotropia-309	162	74	0.0	0.0	NUM
biotropia-309	162	75	0.0	0.0	NUM
biotropia-309	162	76	c	c	NOUN
biotropia-309	162	77	onthophagus	onthophagus	ADJ
biotropia-309	162	78	cf.wallacei	cf.wallacei	NOUN
biotropia-309	162	79	22.4	22.4	NUM
biotropia-309	162	80	41.5	41.5	NUM
biotropia-309	162	81	10.5	10.5	NUM
biotropia-309	162	82	41.5	41.5	NUM
biotropia-309	162	83	67.0	67.0	NUM
biotropia-309	162	84	1.5	1.5	NUM
biotropia-309	162	85	onthophagus	onthophagus	ADJ
biotropia-309	162	86	forsteni	forsteni	NOUN
biotropia-309	162	87	0.0	0.0	NUM
biotropia-309	162	88	3.7	3.7	NUM
biotropia-309	162	89	33.3	33.3	NUM
biotropia-309	162	90	74.1	74.1	NUM
biotropia-309	162	91	0.0	0.0	NUM
biotropia-309	162	92	0.0	0.0	NUM
biotropia-309	162	93	c	c	PROPN
biotropia-309	162	94	onthophagus	onthophagus	NOUN
biotropia-309	162	95	fulvus	fulvus	VERB
biotropia-309	162	96	0.0	0.0	NUM
biotropia-309	162	97	16.7	16.7	NUM
biotropia-309	162	98	53.6	53.6	NUM
biotropia-309	162	99	61.8	61.8	NUM
biotropia-309	162	100	7.1	7.1	NUM
biotropia-309	162	101	21.5	21.5	NUM
biotropia-309	162	102	d	d	NOUN
biotropia-309	162	103	onthophagus	onthophagus	ADJ
biotropia-309	162	104	rectecornutus	rectecornutus	NOUN
biotropia-309	162	105	0.0	0.0	NUM
biotropia-309	162	106	0.0	0.0	NUM
biotropia-309	162	107	0.0	0.0	NUM
biotropia-309	162	108	33.3	33.3	NUM
biotropia-309	162	109	66.7	66.7	NUM
biotropia-309	162	110	0.0	0.0	NUM
biotropia-309	162	111	onthophagus	onthophagus	ADJ
biotropia-309	162	112	ribbei	ribbei	VERB
biotropia-309	162	113	60.5	60.5	NUM
biotropia-309	162	114	32.4	32.4	NUM
biotropia-309	162	115	8.4	8.4	NUM
biotropia-309	162	116	37.1	37.1	NUM
biotropia-309	162	117	0.4	0.4	NUM
biotropia-309	162	118	1.0	1.0	NUM
biotropia-309	162	119	d	d	PROPN
biotropia-309	162	120	onthophagus	onthophagus	ADJ
biotropia-309	162	121	rudis	rudi	NOUN
biotropia-309	163	1	5.3	5.3	NUM
biotropia-309	163	2	39.5	39.5	NUM
biotropia-309	163	3	29.3	29.3	NUM
biotropia-309	163	4	51.2	51.2	NUM
biotropia-309	163	5	13.3	13.3	NUM
biotropia-309	163	6	9.3	9.3	NUM
biotropia-309	163	7	d	d	NOUN
biotropia-309	163	8	onthophagus	onthophagus	ADJ
biotropia-309	163	9	scrutator	scrutator	NOUN
biotropia-309	163	10	1.3	1.3	NUM
biotropia-309	163	11	37.5	37.5	NUM
biotropia-309	163	12	67.8	67.8	NUM
biotropia-309	163	13	57.1	57.1	NUM
biotropia-309	163	14	5.8	5.8	NUM
biotropia-309	163	15	0.0	0.0	NUM
biotropia-309	163	16	d	d	NOUN
biotropia-309	163	17	onthophagus	onthophagus	ADJ
biotropia-309	163	18	sp.1	sp.1	PROPN
biotropia-309	163	19	63.6	63.6	NUM
biotropia-309	163	20	62.8	62.8	NUM
biotropia-309	163	21	19.4	19.4	NUM
biotropia-309	163	22	30.2	30.2	NUM
biotropia-309	163	23	6.9	6.9	NUM
biotropia-309	163	24	4.7	4.7	NUM
biotropia-309	163	25	d	d	NOUN
biotropia-309	163	26	onthophagus	onthophagus	ADJ
biotropia-309	163	27	sp.2	sp.2	PROPN
biotropia-309	163	28	33.3	33.3	NUM
biotropia-309	163	29	0.0	0.0	NUM
biotropia-309	163	30	0.0	0.0	NUM
biotropia-309	163	31	33.3	33.3	NUM
biotropia-309	163	32	0.0	0.0	NUM
biotropia-309	163	33	0.0	0.0	NUM
biotropia-309	163	34	onthophagus	onthophagus	ADJ
biotropia-309	163	35	sp.3	sp.3	NOUN
biotropia-309	163	36	0.0	0.0	NUM
biotropia-309	163	37	46.9	46.9	NUM
biotropia-309	163	38	66.7	66.7	NUM
biotropia-309	163	39	38.6	38.6	NUM
biotropia-309	163	40	0.0	0.0	NUM
biotropia-309	163	41	14.5	14.5	NUM
biotropia-309	163	42	d	d	NOUN
biotropia-309	163	43	onthophagus	onthophagus	ADJ
biotropia-309	163	44	trituber	trituber	VERB
biotropia-309	163	45	0.0	0.0	NUM
biotropia-309	163	46	4.2	4.2	NUM
biotropia-309	163	47	7.1	7.1	NUM
biotropia-309	163	48	4.2	4.2	NUM
biotropia-309	163	49	92.9	92.9	NUM
biotropia-309	163	50	75.0	75.0	NUM
biotropia-309	163	51	c	c	NOUN
biotropia-309	163	52	phaechrous	phaechrous	ADJ
biotropia-309	163	53	emarginatus	emarginatus	NOUN
biotropia-309	163	54	58.3	58.3	NUM
biotropia-309	163	55	35.7	35.7	NUM
biotropia-309	163	56	0.0	0.0	NUM
biotropia-309	163	57	41.9	41.9	NUM
biotropia-309	163	58	0.0	0.0	NUM
biotropia-309	163	59	22.4	22.4	NUM
biotropia-309	163	60	onthophagus	onthophagus	ADJ
biotropia-309	163	61	limbatus	limbatus	PROPN
biotropia-309	163	62	0.7	0.7	NUM
biotropia-309	163	63	0.3	0.3	NUM
biotropia-309	163	64	0.2	0.2	NUM
biotropia-309	163	65	19.6	19.6	NUM
biotropia-309	163	66	97.3	97.3	NUM
biotropia-309	163	67	77.3	77.3	NUM
biotropia-309	163	68	c	c	NOUN
biotropia-309	163	69	*	*	PUNCT
biotropia-309	163	70	only	only	ADV
biotropia-309	163	71	species	specie	NOUN
biotropia-309	163	72	recorded	record	VERB
biotropia-309	163	73	in	in	ADP
biotropia-309	163	74	both	both	DET
biotropia-309	163	75	sample	sample	NOUN
biotropia-309	163	76	years	year	NOUN
biotropia-309	163	77	(	(	PUNCT
biotropia-309	163	78	2009	2009	NUM
biotropia-309	163	79	and	and	CCONJ
biotropia-309	163	80	2012	2012	NUM
biotropia-309	163	81	)	)	PUNCT
biotropia-309	163	82	were	be	AUX
biotropia-309	163	83	included	include	VERB
biotropia-309	163	84	in	in	ADP
biotropia-309	163	85	the	the	DET
biotropia-309	163	86	analysesindval	analysesindval	NOUN
biotropia-309	163	87	55	55	NUM
biotropia-309	163	88	effectiveness	effectiveness	NOUN
biotropia-309	163	89	of	of	ADP
biotropia-309	163	90	dung	dung	NOUN
biotropia-309	163	91	beetles	beetle	NOUN
biotropia-309	163	92	as	as	ADP
biotropia-309	163	93	bioindicators	bioindicator	NOUN
biotropia-309	163	94	of	of	ADP
biotropia-309	163	95	environmental	environmental	ADJ
biotropia-309	163	96	changes	change	NOUN
biotropia-309	163	97	in	in	ADP
biotropia-309	163	98	land	land	NOUN
biotropia-309	163	99	shahabuddin	shahabuddin	NOUN
biotropia-309	163	100	–	–	PUNCT
biotropia-309	163	101	et	et	PROPN
biotropia-309	163	102	al	al	PROPN
biotropia-309	163	103	.	.	PUNCT
biotropia-309	164	1	conclusions	conclusion	NOUN
biotropia-309	164	2	acknowledgements	acknowledgement	NOUN
biotropia-309	164	3	references	reference	VERB
biotropia-309	164	4	the	the	DET
biotropia-309	164	5	fact	fact	NOUN
biotropia-309	164	6	that	that	SCONJ
biotropia-309	164	7	diversity	diversity	NOUN
biotropia-309	164	8	of	of	ADP
biotropia-309	164	9	dung	dung	NOUN
biotropia-309	164	10	beetles	beetle	NOUN
biotropia-309	164	11	at	at	ADP
biotropia-309	164	12	secondary	secondary	ADJ
biotropia-309	164	13	forest	forest	NOUN
biotropia-309	164	14	and	and	CCONJ
biotropia-309	164	15	agroforestry	agroforestry	NOUN
biotropia-309	164	16	system	system	NOUN
biotropia-309	164	17	has	have	AUX
biotropia-309	164	18	no	no	DET
biotropia-309	164	19	significant	significant	ADJ
biotropia-309	164	20	differences	difference	NOUN
biotropia-309	164	21	with	with	ADP
biotropia-309	164	22	natural	natural	ADJ
biotropia-309	164	23	forest	forest	NOUN
biotropia-309	164	24	has	have	VERB
biotropia-309	164	25	important	important	ADJ
biotropia-309	164	26	implications	implication	NOUN
biotropia-309	164	27	on	on	ADP
biotropia-309	164	28	landscape	landscape	NOUN
biotropia-309	164	29	management	management	NOUN
biotropia-309	164	30	aiming	aim	VERB
biotropia-309	164	31	at	at	ADP
biotropia-309	164	32	maintaining	maintain	VERB
biotropia-309	164	33	a	a	DET
biotropia-309	164	34	high	high	ADJ
biotropia-309	164	35	biodiversity	biodiversity	NOUN
biotropia-309	164	36	.	.	PUNCT
biotropia-309	165	1	besides	besides	SCONJ
biotropia-309	165	2	natural	natural	ADJ
biotropia-309	165	3	forest	forest	NOUN
biotropia-309	165	4	sites	site	NOUN
biotropia-309	165	5	,	,	PUNCT
biotropia-309	165	6	certain	certain	ADJ
biotropia-309	165	7	agroecosystems	agroecosystem	NOUN
biotropia-309	165	8	like	like	ADP
biotropia-309	165	9	cacao	cacao	NOUN
biotropia-309	165	10	agroforestry	agroforestry	NOUN
biotropia-309	165	11	also	also	ADV
biotropia-309	165	12	have	have	VERB
biotropia-309	165	13	potency	potency	NOUN
biotropia-309	165	14	to	to	PART
biotropia-309	165	15	maintain	maintain	VERB
biotropia-309	165	16	a	a	DET
biotropia-309	165	17	high	high	ADJ
biotropia-309	165	18	local	local	ADJ
biotropia-309	165	19	diversity	diversity	NOUN
biotropia-309	165	20	.	.	PUNCT
biotropia-309	166	1	dung	dung	NOUN
biotropia-309	166	2	beetles	beetle	NOUN
biotropia-309	166	3	have	have	VERB
biotropia-309	166	4	high	high	ADJ
biotropia-309	166	5	potency	potency	NOUN
biotropia-309	166	6	to	to	PART
biotropia-309	166	7	be	be	AUX
biotropia-309	166	8	used	use	VERB
biotropia-309	166	9	as	as	ADP
biotropia-309	166	10	bioindicator	bioindicator	NOUN
biotropia-309	166	11	(	(	PUNCT
biotropia-309	166	12	i.e.	i.e.	X
biotropia-309	166	13	ecological	ecological	ADJ
biotropia-309	166	14	indicator	indicator	NOUN
biotropia-309	166	15	)	)	PUNCT
biotropia-309	166	16	in	in	ADP
biotropia-309	166	17	relation	relation	NOUN
biotropia-309	166	18	with	with	ADP
biotropia-309	166	19	their	their	PRON
biotropia-309	166	20	fidelity	fidelity	NOUN
biotropia-309	166	21	and	and	CCONJ
biotropia-309	166	22	specificity	specificity	NOUN
biotropia-309	166	23	to	to	ADP
biotropia-309	166	24	a	a	DET
biotropia-309	166	25	particular	particular	ADJ
biotropia-309	166	26	type	type	NOUN
biotropia-309	166	27	of	of	ADP
biotropia-309	166	28	habitat	habitat	NOUN
biotropia-309	166	29	.	.	PUNCT
biotropia-309	167	1	and	and	CCONJ
biotropia-309	167	2	can	can	AUX
biotropia-309	167	3	be	be	AUX
biotropia-309	167	4	suggested	suggest	VERB
biotropia-309	167	5	as	as	ADP
biotropia-309	167	6	indicator	indicator	NOUN
biotropia-309	167	7	(	(	PUNCT
biotropia-309	167	8	characteristic	characteristic	ADJ
biotropia-309	167	9	)	)	PUNCT
biotropia-309	167	10	species	specie	NOUN
biotropia-309	167	11	of	of	ADP
biotropia-309	167	12	cooler	cool	ADJ
biotropia-309	167	13	and	and	CCONJ
biotropia-309	167	14	shaded	shaded	ADJ
biotropia-309	167	15	habitats	habitat	NOUN
biotropia-309	167	16	(	(	PUNCT
biotropia-309	167	17	e.g.	e.g.	ADV
biotropia-309	167	18	forest	forest	NOUN
biotropia-309	167	19	sites	site	NOUN
biotropia-309	167	20	and	and	CCONJ
biotropia-309	167	21	cacao	cacao	NOUN
biotropia-309	167	22	agroforestry	agroforestry	NOUN
biotropia-309	167	23	system	system	NOUN
biotropia-309	167	24	)	)	PUNCT
biotropia-309	167	25	,	,	PUNCT
biotropia-309	167	26	while	while	SCONJ
biotropia-309	167	27	and	and	CCONJ
biotropia-309	167	28	can	can	AUX
biotropia-309	167	29	be	be	AUX
biotropia-309	167	30	suggested	suggest	VERB
biotropia-309	167	31	as	as	ADP
biotropia-309	167	32	indicator	indicator	NOUN
biotropia-309	167	33	of	of	ADP
biotropia-309	167	34	warmer	warm	ADJ
biotropia-309	167	35	and	and	CCONJ
biotropia-309	167	36	unshaded	unshaded	ADJ
biotropia-309	167	37	habitats	habitat	NOUN
biotropia-309	167	38	(	(	PUNCT
biotropia-309	167	39	e.g.	e.g.	ADV
biotropia-309	167	40	open	open	ADJ
biotropia-309	167	41	cultivated	cultivate	VERB
biotropia-309	167	42	area	area	NOUN
biotropia-309	167	43	)	)	PUNCT
biotropia-309	167	44	.	.	PUNCT
biotropia-309	168	1	the	the	DET
biotropia-309	168	2	remaining	remain	VERB
biotropia-309	168	3	dung	dung	NOUN
biotropia-309	168	4	beetles	beetle	NOUN
biotropia-309	168	5	species	specie	NOUN
biotropia-309	168	6	found	find	VERB
biotropia-309	168	7	across	across	ADP
biotropia-309	168	8	the	the	DET
biotropia-309	168	9	land	land	NOUN
biotropia-309	168	10	-	-	PUNCT
biotropia-309	168	11	use	use	NOUN
biotropia-309	168	12	types	type	NOUN
biotropia-309	168	13	in	in	ADP
biotropia-309	168	14	the	the	DET
biotropia-309	168	15	margin	margin	NOUN
biotropia-309	168	16	of	of	ADP
biotropia-309	168	17	llnp	llnp	PROPN
biotropia-309	168	18	,	,	PUNCT
biotropia-309	168	19	central	central	ADJ
biotropia-309	168	20	sulawesi	sulawesi	NOUN
biotropia-309	168	21	are	be	AUX
biotropia-309	168	22	better	well	ADJ
biotropia-309	168	23	to	to	PART
biotropia-309	168	24	be	be	AUX
biotropia-309	168	25	categorized	categorize	VERB
biotropia-309	168	26	as	as	ADP
biotropia-309	168	27	detector	detector	NOUN
biotropia-309	168	28	or	or	CCONJ
biotropia-309	168	29	moderate	moderate	ADJ
biotropia-309	168	30	species	specie	NOUN
biotropia-309	168	31	.	.	PUNCT
biotropia-309	169	1	the	the	DET
biotropia-309	169	2	authors	author	NOUN
biotropia-309	169	3	gratefully	gratefully	ADV
biotropia-309	169	4	acknowledge	acknowledge	VERB
biotropia-309	169	5	the	the	DET
biotropia-309	169	6	directorate	directorate	NOUN
biotropia-309	169	7	of	of	ADP
biotropia-309	169	8	higher	high	ADJ
biotropia-309	169	9	education	education	PROPN
biotropia-309	169	10	ministry	ministry	PROPN
biotropia-309	169	11	of	of	ADP
biotropia-309	169	12	education	education	PROPN
biotropia-309	169	13	and	and	CCONJ
biotropia-309	169	14	culture	culture	NOUN
biotropia-309	169	15	indonesia	indonesia	PROPN
biotropia-309	169	16	for	for	ADP
biotropia-309	169	17	providing	provide	VERB
biotropia-309	169	18	the	the	DET
biotropia-309	169	19	national	national	ADJ
biotropia-309	169	20	strategies	strategy	NOUN
biotropia-309	169	21	research	research	NOUN
biotropia-309	169	22	grant	grant	NOUN
biotropia-309	169	23	(	(	PUNCT
biotropia-309	169	24	contract	contract	NOUN
biotropia-309	169	25	no	no	INTJ
biotropia-309	169	26	.	.	PUNCT
biotropia-309	170	1	027	027	NUM
biotropia-309	170	2	/	/	SYM
biotropia-309	170	3	sp2h	sp2h	NOUN
biotropia-309	170	4	/	/	SYM
biotropia-309	170	5	dit.litabmas	dit.litabmas	PROPN
biotropia-309	170	6	/	/	SYM
biotropia-309	170	7	iii/2012	iii/2012	NOUN
biotropia-309	170	8	)	)	PUNCT
biotropia-309	170	9	,	,	PUNCT
biotropia-309	170	10	and	and	CCONJ
biotropia-309	170	11	to	to	ADP
biotropia-309	170	12	the	the	DET
biotropia-309	170	13	head	head	NOUN
biotropia-309	170	14	of	of	ADP
biotropia-309	170	15	lore	lore	PROPN
biotropia-309	170	16	lindu	lindu	PROPN
biotropia-309	170	17	national	national	ADJ
biotropia-309	170	18	park	park	NOUN
biotropia-309	170	19	for	for	ADP
biotropia-309	170	20	his	his	PRON
biotropia-309	170	21	permission	permission	NOUN
biotropia-309	170	22	to	to	PART
biotropia-309	170	23	conduct	conduct	VERB
biotropia-309	170	24	this	this	DET
biotropia-309	170	25	study	study	NOUN
biotropia-309	170	26	.	.	PUNCT
biotropia-309	171	1	copris	copris	PROPN
biotropia-309	171	2	saundersi	saundersi	PROPN
biotropia-309	171	3	onthophagus	onthophagus	VERB
biotropia-309	171	4	forsteni	forsteni	PROPN
biotropia-309	171	5	o.	o.	PROPN
biotropia-309	171	6	trituber	trituber	PROPN
biotropia-309	171	7	o.	o.	PROPN
biotropia-309	171	8	limbatus	limbatus	PROPN
biotropia-309	171	9	andresen	andresen	PROPN
biotropia-309	171	10	e.	e.	PROPN
biotropia-309	171	11	2005	2005	NUM
biotropia-309	171	12	.	.	PUNCT
biotropia-309	172	1	effects	effect	NOUN
biotropia-309	172	2	of	of	ADP
biotropia-309	172	3	season	season	NOUN
biotropia-309	172	4	and	and	CCONJ
biotropia-309	172	5	vegetation	vegetation	NOUN
biotropia-309	172	6	type	type	NOUN
biotropia-309	172	7	on	on	ADP
biotropia-309	172	8	community	community	NOUN
biotropia-309	172	9	organization	organization	NOUN
biotropia-309	172	10	of	of	ADP
biotropia-309	172	11	dung	dung	NOUN
biotropia-309	172	12	beetles	beetle	NOUN
biotropia-309	172	13	in	in	ADP
biotropia-309	172	14	a	a	DET
biotropia-309	172	15	tropical	tropical	ADJ
biotropia-309	172	16	dry	dry	ADJ
biotropia-309	172	17	forest	forest	NOUN
biotropia-309	172	18	.	.	PUNCT
biotropia-309	173	1	biotropica	biotropica	PROPN
biotropia-309	173	2	37(2	37(2	PRON
biotropia-309	173	3	):	):	PUNCT
biotropia-309	173	4	291	291	NUM
biotropia-309	173	5	-	-	SYM
biotropia-309	173	6	300	300	NUM
biotropia-309	173	7	.	.	PUNCT
biotropia-309	174	1	aydin	aydin	VERB
biotropia-309	174	2	g	g	PROPN
biotropia-309	174	3	,	,	PUNCT
biotropia-309	174	4	kazak	kazak	PROPN
biotropia-309	174	5	c.	c.	PROPN
biotropia-309	174	6	2010	2010	NUM
biotropia-309	174	7	.	.	PUNCT
biotropia-309	175	1	selecting	select	VERB
biotropia-309	175	2	indicator	indicator	NOUN
biotropia-309	175	3	species	specie	NOUN
biotropia-309	175	4	habitat	habitat	ADJ
biotropia-309	175	5	description	description	NOUN
biotropia-309	175	6	and	and	CCONJ
biotropia-309	175	7	sustainable	sustainable	ADJ
biotropia-309	175	8	land	land	NOUN
biotropia-309	175	9	utilization	utilization	NOUN
biotropia-309	175	10	:	:	PUNCT
biotropia-309	175	11	a	a	DET
biotropia-309	175	12	case	case	NOUN
biotropia-309	175	13	study	study	NOUN
biotropia-309	175	14	in	in	ADP
biotropia-309	175	15	amediterranean	amediterranean	PROPN
biotropia-309	175	16	delta.international	delta.international	PROPN
biotropia-309	175	17	journal	journal	NOUN
biotropia-309	175	18	of	of	ADP
biotropia-309	175	19	agriculture	agriculture	PROPN
biotropia-309	175	20	&	&	CCONJ
biotropia-309	175	21	biology	biology	NOUN
biotropia-309	175	22	12	12	NUM
biotropia-309	175	23	:	:	PUNCT
biotropia-309	175	24	931	931	NUM
biotropia-309	175	25	-	-	SYM
biotropia-309	175	26	934	934	NUM
biotropia-309	175	27	.	.	PUNCT
biotropia-309	176	1	balthasar	balthasar	VERB
biotropia-309	176	2	v.1963	v.1963	PROPN
biotropia-309	176	3	.	.	PUNCT
biotropia-309	177	1	monographie	monographie	PROPN
biotropia-309	177	2	der	der	PROPN
biotropia-309	177	3	scarabaeidae	scarabaeidae	VERB
biotropia-309	177	4	und	und	PROPN
biotropia-309	177	5	aphodiidae	aphodiidae	PROPN
biotropia-309	177	6	der	der	NOUN
biotropia-309	177	7	palaearktischen	palaearktischen	PROPN
biotropia-309	177	8	und	und	VERB
biotropia-309	177	9	orientalischen	orientalischen	ADV
biotropia-309	177	10	region.band	region.band	PRON
biotropia-309	177	11	1.verlag	1.verlag	NUM
biotropia-309	177	12	der	der	NOUN
biotropia-309	177	13	tschechoslawakischen	tschechoslawakischen	PROPN
biotropia-309	177	14	akademie	akademie	PROPN
biotropia-309	177	15	der	der	PROPN
biotropia-309	177	16	wissenschaften	wissenschaften	NOUN
biotropia-309	177	17	.	.	PUNCT
biotropia-309	178	1	prague	prague	PROPN
biotropia-309	178	2	.	.	PUNCT
biotropia-309	179	1	brose	brose	PROPN
biotropia-309	179	2	u	u	PROPN
biotropia-309	179	3	,	,	PUNCT
biotropia-309	179	4	martinez	martinez	PROPN
biotropia-309	179	5	nd	nd	PROPN
biotropia-309	179	6	,	,	PUNCT
biotropia-309	179	7	williams	williams	PROPN
biotropia-309	179	8	rj	rj	PROPN
biotropia-309	179	9	.	.	PROPN
biotropia-309	179	10	2003	2003	NUM
biotropia-309	179	11	.	.	PUNCT
biotropia-309	180	1	estimating	estimate	VERB
biotropia-309	180	2	species	specie	NOUN
biotropia-309	180	3	richness	richness	NOUN
biotropia-309	180	4	:	:	PUNCT
biotropia-309	180	5	sensitivity	sensitivity	NOUN
biotropia-309	180	6	to	to	PART
biotropia-309	180	7	sample	sample	VERB
biotropia-309	180	8	coverage	coverage	NOUN
biotropia-309	180	9	and	and	CCONJ
biotropia-309	180	10	insensitivity	insensitivity	NOUN
biotropia-309	180	11	to	to	ADP
biotropia-309	180	12	spatial	spatial	ADJ
biotropia-309	180	13	patterns	pattern	NOUN
biotropia-309	180	14	.	.	PUNCT
biotropia-309	181	1	ecology	ecology	NOUN
biotropia-309	181	2	84	84	NUM
biotropia-309	181	3	(	(	PUNCT
biotropia-309	181	4	9	9	NUM
biotropia-309	181	5	):	):	PUNCT
biotropia-309	181	6	2364	2364	NUM
biotropia-309	181	7	-	-	SYM
biotropia-309	181	8	2377	2377	NUM
biotropia-309	181	9	.	.	PUNCT
biotropia-309	182	1	clarke	clarke	PROPN
biotropia-309	182	2	kr	kr	PROPN
biotropia-309	182	3	.	.	PROPN
biotropia-309	182	4	1993	1993	NUM
biotropia-309	182	5	.	.	PUNCT
biotropia-309	183	1	non	non	ADJ
biotropia-309	183	2	-	-	ADJ
biotropia-309	183	3	parametric	parametric	ADJ
biotropia-309	183	4	multivariate	multivariate	NOUN
biotropia-309	183	5	analyses	analysis	NOUN
biotropia-309	183	6	of	of	ADP
biotropia-309	183	7	changes	change	NOUN
biotropia-309	183	8	in	in	ADP
biotropia-309	183	9	community	community	NOUN
biotropia-309	183	10	structure	structure	NOUN
biotropia-309	183	11	.	.	PUNCT
biotropia-309	184	1	australian	australian	ADJ
biotropia-309	184	2	journal	journal	PROPN
biotropia-309	184	3	ecology	ecology	NOUN
biotropia-309	184	4	18	18	NUM
biotropia-309	184	5	:	:	SYM
biotropia-309	184	6	117	117	NUM
biotropia-309	184	7	-	-	SYM
biotropia-309	184	8	143	143	NUM
biotropia-309	184	9	.	.	PUNCT
biotropia-309	185	1	colwell	colwell	PROPN
biotropia-309	185	2	rk	rk	PROPN
biotropia-309	185	3	.	.	PROPN
biotropia-309	185	4	2004	2004	NUM
biotropia-309	185	5	.	.	PUNCT
biotropia-309	186	1	estimates	estimate	VERB
biotropia-309	186	2	:	:	PUNCT
biotropia-309	186	3	statistical	statistical	ADJ
biotropia-309	186	4	estimation	estimation	NOUN
biotropia-309	186	5	of	of	ADP
biotropia-309	186	6	species	specie	NOUN
biotropia-309	186	7	richnes	richne	NOUN
biotropia-309	186	8	and	and	CCONJ
biotropia-309	186	9	shared	share	VERB
biotropia-309	186	10	species	specie	NOUN
biotropia-309	186	11	from	from	ADP
biotropia-309	186	12	samples	sample	NOUN
biotropia-309	186	13	.	.	PUNCT
biotropia-309	187	1	version	version	NOUN
biotropia-309	187	2	7.user	7.user	NUM
biotropia-309	187	3	's	's	PART
biotropia-309	187	4	guide	guide	NOUN
biotropia-309	187	5	and	and	CCONJ
biotropia-309	187	6	application	application	NOUN
biotropia-309	187	7	.	.	PUNCT
biotropia-309	187	8	.	.	PUNCT
biotropia-309	188	1	davis	davis	PROPN
biotropia-309	188	2	aj	aj	PROPN
biotropia-309	188	3	,	,	PUNCT
biotropia-309	188	4	holloway	holloway	PROPN
biotropia-309	188	5	jd	jd	PROPN
biotropia-309	188	6	,	,	PUNCT
biotropia-309	188	7	huijbregts	huijbregts	PROPN
biotropia-309	188	8	h	h	PROPN
biotropia-309	188	9	,	,	PUNCT
biotropia-309	188	10	krikken	krikken	PROPN
biotropia-309	188	11	j	j	PROPN
biotropia-309	188	12	,	,	PUNCT
biotropia-309	188	13	kirk	kirk	PROPN
biotropia-309	188	14	-	-	PUNCT
biotropia-309	188	15	spriggs	spriggs	PROPN
biotropia-309	188	16	ah	ah	INTJ
biotropia-309	188	17	,	,	PUNCT
biotropia-309	188	18	sutton	sutton	PROPN
biotropia-309	188	19	sl	sl	PROPN
biotropia-309	188	20	.	.	PROPN
biotropia-309	188	21	2001	2001	NUM
biotropia-309	188	22	.	.	PUNCT
biotropia-309	189	1	dung	dung	NOUN
biotropia-309	189	2	beetles	beetle	NOUN
biotropia-309	189	3	as	as	ADP
biotropia-309	189	4	indicators	indicator	NOUN
biotropia-309	189	5	of	of	ADP
biotropia-309	189	6	change	change	NOUN
biotropia-309	189	7	in	in	ADP
biotropia-309	189	8	the	the	DET
biotropia-309	189	9	forests	forest	NOUN
biotropia-309	189	10	of	of	ADP
biotropia-309	189	11	northern	northern	ADJ
biotropia-309	189	12	borneo	borneo	NOUN
biotropia-309	189	13	.	.	PUNCT
biotropia-309	190	1	journal	journal	PROPN
biotropia-309	190	2	of	of	ADP
biotropia-309	190	3	applied	applied	ADJ
biotropia-309	190	4	ecology	ecology	NOUN
biotropia-309	190	5	38	38	NUM
biotropia-309	190	6	:	:	SYM
biotropia-309	190	7	593	593	NUM
biotropia-309	190	8	-	-	SYM
biotropia-309	190	9	616	616	NUM
biotropia-309	190	10	.	.	PUNCT
biotropia-309	191	1	dufrêne	dufrêne	PROPN
biotropia-309	191	2	m	m	PROPN
biotropia-309	191	3	,	,	PUNCT
biotropia-309	191	4	legendre	legendre	PROPN
biotropia-309	191	5	p.	p.	PROPN
biotropia-309	191	6	1997	1997	NUM
biotropia-309	191	7	.	.	PUNCT
biotropia-309	192	1	species	specie	NOUN
biotropia-309	192	2	assemblages	assemblage	NOUN
biotropia-309	192	3	and	and	CCONJ
biotropia-309	192	4	indicator	indicator	NOUN
biotropia-309	192	5	species	specie	NOUN
biotropia-309	192	6	:	:	PUNCT
biotropia-309	192	7	the	the	DET
biotropia-309	192	8	need	need	NOUN
biotropia-309	192	9	for	for	ADP
biotropia-309	192	10	a	a	DET
biotropia-309	192	11	flexible	flexible	ADJ
biotropia-309	192	12	asymmetrical	asymmetrical	ADJ
biotropia-309	192	13	approach	approach	NOUN
biotropia-309	192	14	.	.	PUNCT
biotropia-309	193	1	ecological	ecological	ADJ
biotropia-309	193	2	monographs	monograph	NOUN
biotropia-309	193	3	67	67	NUM
biotropia-309	193	4	:	:	SYM
biotropia-309	193	5	345	345	NUM
biotropia-309	193	6	-	-	SYM
biotropia-309	193	7	366	366	NUM
biotropia-309	193	8	.	.	PUNCT
biotropia-309	194	1	errouissi	errouissi	PROPN
biotropia-309	194	2	f	f	PROPN
biotropia-309	194	3	,	,	PUNCT
biotropia-309	194	4	haloti	haloti	PROPN
biotropia-309	194	5	s	s	PROPN
biotropia-309	194	6	,	,	PUNCT
biotropia-309	194	7	robert	robert	PROPN
biotropia-309	194	8	pj	pj	PROPN
biotropia-309	194	9	,	,	PUNCT
biotropia-309	194	10	idrissi	idrissi	PROPN
biotropia-309	194	11	aj	aj	PROPN
biotropia-309	194	12	,	,	PUNCT
biotropia-309	194	13	lumaret	lumaret	PROPN
biotropia-309	194	14	jp	jp	PROPN
biotropia-309	194	15	.	.	PUNCT
biotropia-309	194	16	2004	2004	NUM
biotropia-309	194	17	.	.	PUNCT
biotropia-309	195	1	effect	effect	NOUN
biotropia-309	195	2	of	of	ADP
biotropia-309	195	3	the	the	DET
biotropia-309	195	4	attractiveness	attractiveness	NOUN
biotropia-309	195	5	for	for	ADP
biotropia-309	195	6	dung	dung	NOUN
biotropia-309	195	7	beetles	beetle	NOUN
biotropia-309	195	8	of	of	ADP
biotropia-309	195	9	dung	dung	NOUN
biotropia-309	195	10	pat	pat	NOUN
biotropia-309	195	11	origin	origin	NOUN
biotropia-309	195	12	and	and	CCONJ
biotropia-309	195	13	size	size	NOUN
biotropia-309	195	14	along	along	ADP
biotropia-309	195	15	climatic	climatic	ADJ
biotropia-309	195	16	gradient	gradient	NOUN
biotropia-309	195	17	.	.	PUNCT
biotropia-309	196	1	environ	environ	PROPN
biotropia-309	196	2	.	.	PUNCT
biotropia-309	196	3	entomol	entomol	PROPN
biotropia-309	196	4	.	.	PUNCT
biotropia-309	197	1	33(1	33(1	NUM
biotropia-309	197	2	):	):	PUNCT
biotropia-309	197	3	45	45	NUM
biotropia-309	197	4	-	-	SYM
biotropia-309	197	5	53	53	NUM
biotropia-309	197	6	.	.	PUNCT
biotropia-309	197	7	gerold	gerold	VERB
biotropia-309	197	8	g	g	NOUN
biotropia-309	197	9	,	,	PUNCT
biotropia-309	197	10	fremery	fremery	NOUN
biotropia-309	197	11	m	m	NOUN
biotropia-309	197	12	,	,	PUNCT
biotropia-309	197	13	guhardja	guhardja	PROPN
biotropia-309	197	14	,	,	PUNCT
biotropia-309	197	15	e.	e.	PROPN
biotropia-309	197	16	2004	2004	PROPN
biotropia-309	197	17	.	.	PUNCT
biotropia-309	198	1	land	land	NOUN
biotropia-309	198	2	-	-	PUNCT
biotropia-309	198	3	use	use	NOUN
biotropia-309	198	4	,	,	PUNCT
biotropia-309	198	5	nature	nature	NOUN
biotropia-309	198	6	conservation	conservation	NOUN
biotropia-309	198	7	and	and	CCONJ
biotropia-309	198	8	the	the	DET
biotropia-309	198	9	stability	stability	NOUN
biotropia-309	198	10	of	of	ADP
biotropia-309	198	11	rainforest	rainforest	NOUN
biotropia-309	198	12	margins	margin	NOUN
biotropia-309	198	13	in	in	ADP
biotropia-309	198	14	southeast	southeast	PROPN
biotropia-309	198	15	asia.springer	asia.springer	PROPN
biotropia-309	198	16	,	,	PUNCT
biotropia-309	198	17	berlin	berlin	PROPN
biotropia-309	198	18	.	.	PUNCT
biotropia-309	199	1	534	534	NUM
biotropia-309	199	2	pp	pp	NOUN
biotropia-309	199	3	.	.	PUNCT
biotropia-309	200	1	http://purl.oclc.org/estimates	http://purl.oclc.org/estimate	NOUN
biotropia-309	200	2	56	56	NUM
biotropia-309	200	3	biotropia	biotropia	ADJ
biotropia-309	200	4	vol	vol	NOUN
biotropia-309	200	5	.	.	PROPN
biotropia-309	201	1	21	21	NUM
biotropia-309	201	2	no	no	NOUN
biotropia-309	201	3	.	.	NOUN
biotropia-309	201	4	1	1	NUM
biotropia-309	201	5	,	,	PUNCT
biotropia-309	201	6	2014	2014	NUM
biotropia-309	201	7	gerlach	gerlach	PROPN
biotropia-309	201	8	j	j	PROPN
biotropia-309	201	9	,	,	PUNCT
biotropia-309	201	10	samways	samways	PROPN
biotropia-309	201	11	m	m	PROPN
biotropia-309	201	12	,	,	PUNCT
biotropia-309	201	13	pryke	pryke	NOUN
biotropia-309	201	14	j.	j.	PROPN
biotropia-309	201	15	2013	2013	NUM
biotropia-309	201	16	.	.	PUNCT
biotropia-309	202	1	terrestrial	terrestrial	ADJ
biotropia-309	202	2	invertebrates	invertebrate	NOUN
biotropia-309	202	3	as	as	ADP
biotropia-309	202	4	bioindicators	bioindicator	NOUN
biotropia-309	202	5	:	:	PUNCT
biotropia-309	202	6	an	an	DET
biotropia-309	202	7	overview	overview	NOUN
biotropia-309	202	8	of	of	ADP
biotropia-309	202	9	available	available	ADJ
biotropia-309	202	10	taxonomic	taxonomic	ADJ
biotropia-309	202	11	groups	group	NOUN
biotropia-309	202	12	.	.	PUNCT
biotropia-309	203	1	journal	journal	NOUN
biotropia-309	203	2	of	of	ADP
biotropia-309	203	3	insect	insect	NOUN
biotropia-309	203	4	conservation,17	conservation,17	NOUN
biotropia-309	203	5	,	,	PUNCT
biotropia-309	203	6	831	831	NUM
biotropia-309	203	7	-	-	SYM
biotropia-309	203	8	850	850	NUM
biotropia-309	203	9	.	.	PUNCT
biotropia-309	204	1	halffter	halffter	NOUN
biotropia-309	204	2	g	g	PROPN
biotropia-309	204	3	,	,	PUNCT
biotropia-309	204	4	favila	favila	VERB
biotropia-309	204	5	me	i	PRON
biotropia-309	204	6	.	.	PUNCT
biotropia-309	205	1	1993	1993	NUM
biotropia-309	205	2	.	.	PUNCT
biotropia-309	206	1	the	the	DET
biotropia-309	206	2	scarabaeinae	scarabaeinae	NOUN
biotropia-309	206	3	(	(	PUNCT
biotropia-309	206	4	coleoptera	coleoptera	NOUN
biotropia-309	206	5	):	):	PUNCT
biotropia-309	206	6	an	an	DET
biotropia-309	206	7	animal	animal	NOUN
biotropia-309	206	8	group	group	NOUN
biotropia-309	206	9	for	for	ADP
biotropia-309	206	10	analysing	analyse	VERB
biotropia-309	206	11	,	,	PUNCT
biotropia-309	206	12	inventorying	inventory	VERB
biotropia-309	206	13	and	and	CCONJ
biotropia-309	206	14	monitoring	monitor	VERB
biotropia-309	206	15	biodiversity	biodiversity	NOUN
biotropia-309	206	16	in	in	ADP
biotropia-309	206	17	tropical	tropical	ADJ
biotropia-309	206	18	rainforest	rainforest	NOUN
biotropia-309	206	19	and	and	CCONJ
biotropia-309	206	20	modified	modify	VERB
biotropia-309	206	21	landscapes	landscape	NOUN
biotropia-309	206	22	.	.	PUNCT
biotropia-309	207	1	biology	biology	NOUN
biotropia-309	207	2	international	international	ADJ
biotropia-309	207	3	27:15	27:15	NUM
biotropia-309	207	4	-	-	SYM
biotropia-309	207	5	21	21	NUM
biotropia-309	207	6	hambler	hambler	NOUN
biotropia-309	207	7	c	c	PROPN
biotropia-309	207	8	,	,	PUNCT
biotropia-309	207	9	henderson	henderson	PROPN
biotropia-309	207	10	pa	pa	PROPN
biotropia-309	207	11	,	,	PUNCT
biotropia-309	207	12	speight	speight	PROPN
biotropia-309	207	13	mr	mr	PROPN
biotropia-309	207	14	.	.	PROPN
biotropia-309	207	15	2011	2011	NUM
biotropia-309	207	16	.	.	PUNCT
biotropia-309	208	1	extinction	extinction	NOUN
biotropia-309	208	2	rates	rate	NOUN
biotropia-309	208	3	,	,	PUNCT
biotropia-309	208	4	extinction	extinction	NOUN
biotropia-309	208	5	-	-	PUNCT
biotropia-309	208	6	prone	prone	ADJ
biotropia-309	208	7	habitats	habitat	NOUN
biotropia-309	208	8	,	,	PUNCT
biotropia-309	208	9	and	and	CCONJ
biotropia-309	208	10	indicator	indicator	NOUN
biotropia-309	208	11	groups	group	NOUN
biotropia-309	208	12	in	in	ADP
biotropia-309	208	13	britain	britain	PROPN
biotropia-309	208	14	and	and	CCONJ
biotropia-309	208	15	at	at	ADP
biotropia-309	208	16	larger	large	ADJ
biotropia-309	208	17	scales	scale	NOUN
biotropia-309	208	18	.	.	PUNCT
biotropia-309	209	1	biological	biological	ADJ
biotropia-309	209	2	conservation,144	conservation,144	PROPN
biotropia-309	209	3	,	,	PUNCT
biotropia-309	209	4	713	713	NUM
biotropia-309	209	5	-	-	SYM
biotropia-309	209	6	721	721	NUM
biotropia-309	209	7	.	.	PUNCT
biotropia-309	209	8	harvey	harvey	PROPN
biotropia-309	209	9	ca	ca	PROPN
biotropia-309	209	10	,	,	PUNCT
biotropia-309	209	11	gonzalez	gonzalez	PROPN
biotropia-309	209	12	j	j	PROPN
biotropia-309	209	13	,	,	PUNCT
biotropia-309	209	14	somarriba	somarriba	PROPN
biotropia-309	209	15	e.	e.	PROPN
biotropia-309	209	16	2006	2006	NUM
biotropia-309	209	17	.	.	PUNCT
biotropia-309	210	1	dung	dung	NOUN
biotropia-309	210	2	beetle	beetle	NOUN
biotropia-309	210	3	and	and	CCONJ
biotropia-309	210	4	terrestrial	terrestrial	ADJ
biotropia-309	210	5	mammal	mammal	ADJ
biotropia-309	210	6	diversity	diversity	NOUN
biotropia-309	210	7	in	in	ADP
biotropia-309	210	8	forests	forest	NOUN
biotropia-309	210	9	,	,	PUNCT
biotropia-309	210	10	indigenous	indigenous	ADJ
biotropia-309	210	11	agroforestry	agroforestry	NOUN
biotropia-309	210	12	systems	system	NOUN
biotropia-309	210	13	and	and	CCONJ
biotropia-309	210	14	plantain	plantain	NOUN
biotropia-309	210	15	monocultures	monoculture	NOUN
biotropia-309	210	16	in	in	ADP
biotropia-309	210	17	talamanca	talamanca	NOUN
biotropia-309	210	18	,	,	PUNCT
biotropia-309	210	19	costa	costa	PROPN
biotropia-309	210	20	rica	rica	PROPN
biotropia-309	210	21	.	.	PUNCT
biotropia-309	211	1	biodiversity	biodiversity	NOUN
biotropia-309	211	2	and	and	CCONJ
biotropia-309	211	3	conservatio	conservatio	NOUN
biotropia-309	211	4	15	15	NUM
biotropia-309	211	5	:	:	PUNCT
biotropia-309	211	6	555	555	NUM
biotropia-309	211	7	-	-	SYM
biotropia-309	211	8	585	585	NUM
biotropia-309	211	9	.	.	PUNCT
biotropia-309	212	1	kessler	kessler	PROPN
biotropia-309	212	2	m	m	PROPN
biotropia-309	212	3	,	,	PUNCT
biotropia-309	212	4	abrahamczyk	abrahamczyk	PROPN
biotropia-309	212	5	s	s	PROPN
biotropia-309	212	6	,	,	PUNCT
biotropia-309	212	7	bos	bos	NOUN
biotropia-309	212	8	m	m	PROPN
biotropia-309	212	9	,	,	PUNCT
biotropia-309	212	10	buchori	buchori	ADJ
biotropia-309	212	11	d	d	PROPN
biotropia-309	212	12	,	,	PUNCT
biotropia-309	212	13	putra	putra	PROPN
biotropia-309	212	14	dd	dd	PROPN
biotropia-309	212	15	,	,	PUNCT
biotropia-309	212	16	gradstein	gradstein	PROPN
biotropia-309	212	17	sr	sr	PROPN
biotropia-309	212	18	,	,	PUNCT
biotropia-309	212	19	höhn	höhn	PROPN
biotropia-309	212	20	p	p	X
biotropia-309	212	21	,	,	PUNCT
biotropia-309	212	22	kluge	kluge	ADJ
biotropia-309	212	23	j	j	PROPN
biotropia-309	212	24	,	,	PUNCT
biotropia-309	212	25	orend	orend	ADJ
biotropia-309	212	26	f	f	NOUN
biotropia-309	212	27	,	,	PUNCT
biotropia-309	212	28	pitopang	pitopang	PROPN
biotropia-309	212	29	r	r	NOUN
biotropia-309	212	30	,	,	PUNCT
biotropia-309	212	31	shahabuddin	shahabuddin	VERB
biotropia-309	212	32	s	s	PRON
biotropia-309	212	33	,	,	PUNCT
biotropia-309	212	34	schulze	schulze	NOUN
biotropia-309	212	35	ch	ch	PROPN
biotropia-309	212	36	,	,	PUNCT
biotropia-309	212	37	sporn	sporn	PROPN
biotropia-309	212	38	sg	sg	PROPN
biotropia-309	212	39	,	,	PUNCT
biotropia-309	212	40	steffan	steffan	NOUN
biotropia-309	212	41	-	-	PUNCT
biotropia-309	212	42	dewenteri	dewenteri	NOUN
biotropia-309	212	43	,	,	PUNCT
biotropia-309	212	44	tjitrosoedirdjo	tjitrosoedirdjo	PROPN
biotropia-309	212	45	ss	ss	NOUN
biotropia-309	212	46	,	,	PUNCT
biotropia-309	212	47	tscharntke	tscharntke	NOUN
biotropia-309	212	48	t.	t.	PROPN
biotropia-309	212	49	2011	2011	NUM
biotropia-309	212	50	.	.	PUNCT
biotropia-309	213	1	cost	cost	NOUN
biotropia-309	213	2	-	-	PUNCT
biotropia-309	213	3	effectiveness	effectiveness	NOUN
biotropia-309	213	4	of	of	ADP
biotropia-309	213	5	plant	plant	NOUN
biotropia-309	213	6	and	and	CCONJ
biotropia-309	213	7	animal	animal	NOUN
biotropia-309	213	8	biodiversity	biodiversity	NOUN
biotropia-309	213	9	indicators	indicator	NOUN
biotropia-309	213	10	in	in	ADP
biotropia-309	213	11	tropical	tropical	ADJ
biotropia-309	213	12	forest	forest	NOUN
biotropia-309	213	13	and	and	CCONJ
biotropia-309	213	14	agroforest	agroforest	NOUN
biotropia-309	213	15	habitats	habitat	NOUN
biotropia-309	213	16	.	.	PUNCT
biotropia-309	214	1	journal	journal	NOUN
biotropia-309	214	2	of	of	ADP
biotropia-309	214	3	applied	applied	ADJ
biotropia-309	214	4	ecology	ecology	NOUN
biotropia-309	214	5	48	48	NUM
biotropia-309	214	6	:	:	SYM
biotropia-309	214	7	330	330	NUM
biotropia-309	214	8	339	339	NUM
biotropia-309	214	9	.	.	PUNCT
biotropia-309	215	1	kessler	kessler	PROPN
biotropia-309	215	2	m	m	PROPN
biotropia-309	215	3	,	,	PUNCT
biotropia-309	215	4	herte	herte	PROPN
biotropia-309	215	5	d	d	NOUN
biotropia-309	215	6	,	,	PUNCT
biotropia-309	215	7	jungkunst	jungkunst	ADJ
biotropia-309	215	8	hf	hf	PROPN
biotropia-309	215	9	,	,	PUNCT
biotropia-309	215	10	kluge	kluge	ADJ
biotropia-309	215	11	j	j	PROPN
biotropia-309	215	12	,	,	PUNCT
biotropia-309	215	13	abrahamczyk	abrahamczyk	NOUN
biotropia-309	215	14	s	s	PROPN
biotropia-309	215	15	,	,	PUNCT
biotropia-309	215	16	bos	bos	NOUN
biotropia-309	215	17	m	m	PROPN
biotropia-309	215	18	,	,	PUNCT
biotropia-309	215	19	buchori	buchori	ADJ
biotropia-309	215	20	d	d	NOUN
biotropia-309	215	21	,	,	PUNCT
biotropia-309	215	22	gerold	gerold	VERB
biotropia-309	215	23	g	g	NOUN
biotropia-309	215	24	,	,	PUNCT
biotropia-309	215	25	gradstein	gradstein	PROPN
biotropia-309	215	26	sr	sr	PROPN
biotropia-309	215	27	,	,	PUNCT
biotropia-309	215	28	ko¨	ko¨	PROPN
biotropia-309	215	29	hler	hler	NOUN
biotropia-309	215	30	s	s	PART
biotropia-309	215	31	,	,	PUNCT
biotropia-309	215	32	leuschner	leuschner	NOUN
biotropia-309	215	33	c	c	PROPN
biotropia-309	215	34	,	,	PUNCT
biotropia-309	215	35	moser	moser	PROPN
biotropia-309	215	36	g	g	PROPN
biotropia-309	215	37	,	,	PUNCT
biotropia-309	215	38	pitopang	pitopang	PROPN
biotropia-309	215	39	r	r	NOUN
biotropia-309	215	40	,	,	PUNCT
biotropia-309	215	41	shahabuddin	shahabuddin	VERB
biotropia-309	215	42	s	s	PRON
biotropia-309	215	43	,	,	PUNCT
biotropia-309	215	44	schulze	schulze	NOUN
biotropia-309	215	45	ch	ch	PROPN
biotropia-309	215	46	,	,	PUNCT
biotropia-309	215	47	sporn	sporn	PROPN
biotropia-309	215	48	sg	sg	PROPN
biotropia-309	215	49	,	,	PUNCT
biotropia-309	215	50	steffan	steffan	NOUN
biotropia-309	215	51	-	-	ADJ
biotropia-309	215	52	dewenter	dewenter	ADJ
biotropia-309	215	53	i	i	NOUN
biotropia-309	215	54	,	,	PUNCT
biotropia-309	215	55	tjitrosoedirdjo	tjitrosoedirdjo	PROPN
biotropia-309	215	56	ss	ss	PROPN
biotropia-309	215	57	,	,	PUNCT
biotropia-309	215	58	tscharntke	tscharntke	NOUN
biotropia-309	215	59	t.2012	t.2012	NOUN
biotropia-309	215	60	.	.	PUNCT
biotropia-309	216	1	can	can	AUX
biotropia-309	216	2	joint	joint	VERB
biotropia-309	216	3	carbon	carbon	NOUN
biotropia-309	216	4	and	and	CCONJ
biotropia-309	216	5	biodiversity	biodiversity	NOUN
biotropia-309	216	6	management	management	NOUN
biotropia-309	216	7	in	in	ADP
biotropia-309	216	8	tropical	tropical	ADJ
biotropia-309	216	9	agroforestry	agroforestry	NOUN
biotropia-309	216	10	landscapes	landscape	NOUN
biotropia-309	216	11	be	be	AUX
biotropia-309	216	12	optimized	optimize	VERB
biotropia-309	216	13	?	?	PUNCT
biotropia-309	217	1	plosone7(10	plosone7(10	NOUN
biotropia-309	217	2	):	):	PUNCT
biotropia-309	217	3	e47192	e47192	NOUN
biotropia-309	217	4	.	.	PUNCT
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biotropia-309	217	6	/	/	SYM
biotropia-309	218	1	journal.pone.0047192	journal.pone.0047192	PROPN
biotropia-309	218	2	.	.	PROPN
biotropia-309	218	3	klein	klein	PROPN
biotropia-309	218	4	am	be	AUX
biotropia-309	218	5	,	,	PUNCT
biotropia-309	218	6	steffan	steffan	NOUN
biotropia-309	218	7	-	-	ADJ
biotropia-309	218	8	dewenter	dewenter	ADJ
biotropia-309	218	9	i	i	NOUN
biotropia-309	218	10	,	,	PUNCT
biotropia-309	218	11	buchori	buchori	ADJ
biotropia-309	218	12	d	d	NOUN
biotropia-309	218	13	,	,	PUNCT
biotropia-309	218	14	tscharntke	tscharntke	NOUN
biotropia-309	218	15	t.	t.	PROPN
biotropia-309	218	16	2006	2006	NUM
biotropia-309	218	17	.	.	PUNCT
biotropia-309	219	1	rain	rain	NOUN
biotropia-309	219	2	forest	forest	NOUN
biotropia-309	219	3	promotes	promote	VERB
biotropia-309	219	4	trophic	trophic	ADJ
biotropia-309	219	5	interactions	interaction	NOUN
biotropia-309	219	6	and	and	CCONJ
biotropia-309	219	7	diversity	diversity	NOUN
biotropia-309	219	8	of	of	ADP
biotropia-309	219	9	trap	trap	NOUN
biotropia-309	219	10	-	-	PUNCT
biotropia-309	219	11	nesting	nest	VERB
biotropia-309	219	12	hymenoptera	hymenoptera	NOUN
biotropia-309	219	13	in	in	ADP
biotropia-309	219	14	adjacent	adjacent	ADJ
biotropia-309	219	15	agroforestry	agroforestry	PROPN
biotropia-309	219	16	.	.	PUNCT
biotropia-309	220	1	journal	journal	PROPN
biotropia-309	220	2	of	of	ADP
biotropia-309	220	3	animal	animal	NOUN
biotropia-309	220	4	ecology	ecology	NOUN
biotropia-309	220	5	75	75	NUM
biotropia-309	220	6	:	:	SYM
biotropia-309	220	7	315323	315323	NUM
biotropia-309	220	8	.	.	PUNCT
biotropia-309	221	1	kudavidanage	kudavidanage	PROPN
biotropia-309	221	2	ep	ep	PROPN
biotropia-309	221	3	,	,	PUNCT
biotropia-309	221	4	qie	qie	PROPN
biotropia-309	221	5	l	l	PROPN
biotropia-309	221	6	,	,	PUNCT
biotropia-309	221	7	lie	lie	NOUN
biotropia-309	221	8	jsh	jsh	PROPN
biotropia-309	221	9	.	.	PROPN
biotropia-309	221	10	2012	2012	NUM
biotropia-309	221	11	.	.	PUNCT
biotropia-309	222	1	linking	link	VERB
biotropia-309	222	2	biodiversity	biodiversity	NOUN
biotropia-309	222	3	and	and	CCONJ
biotropia-309	222	4	ecosystem	ecosystem	NOUN
biotropia-309	222	5	functioning	functioning	NOUN
biotropia-309	222	6	of	of	ADP
biotropia-309	222	7	dungbeetles	dungbeetle	NOUN
biotropia-309	222	8	in	in	ADP
biotropia-309	222	9	south	south	PROPN
biotropia-309	222	10	and	and	CCONJ
biotropia-309	222	11	southeast	southeast	ADJ
biotropia-309	222	12	asian	asian	ADJ
biotropia-309	222	13	tropical	tropical	ADJ
biotropia-309	222	14	rainforests	rainforest	NOUN
biotropia-309	222	15	.	.	PUNCT
biotropia-309	223	1	the	the	DET
biotropia-309	223	2	raffles	raffle	NOUN
biotropia-309	223	3	bulletin	bulletin	NOUN
biotropia-309	223	4	of	of	ADP
biotropia-309	223	5	zoology	zoology	NOUN
biotropia-309	223	6	.	.	PUNCT
biotropia-309	224	1	supplement	supplement	VERB
biotropia-309	224	2	no	no	INTJ
biotropia-309	224	3	.	.	PROPN
biotropia-309	225	1	25	25	NUM
biotropia-309	225	2	:	:	SYM
biotropia-309	225	3	141	141	NUM
biotropia-309	225	4	-	-	SYM
biotropia-309	225	5	154	154	NUM
biotropia-309	225	6	krell	krell	PROPN
biotropia-309	225	7	ft	ft	PROPN
biotropia-309	225	8	.	.	PROPN
biotropia-309	225	9	2007	2007	NUM
biotropia-309	225	10	.	.	PUNCT
biotropia-309	226	1	dung	dung	NOUN
biotropia-309	226	2	beetle	beetle	NOUN
biotropia-309	226	3	sampling	sample	VERB
biotropia-309	226	4	protocols	protocol	NOUN
biotropia-309	226	5	.	.	PUNCT
biotropia-309	227	1	denvermuseum	denvermuseum	NOUN
biotropia-309	227	2	of	of	ADP
biotropia-309	227	3	nature	nature	NOUN
biotropia-309	227	4	&	&	CCONJ
biotropia-309	227	5	science	science	PROPN
biotropia-309	227	6	(	(	PUNCT
biotropia-309	227	7	dmns	dmn	NOUN
biotropia-309	227	8	)	)	PUNCT
biotropia-309	227	9	technical	technical	ADJ
biotropia-309	227	10	report200	report200	PROPN
biotropia-309	227	11	-	-	PUNCT
biotropia-309	227	12	6:1	6:1	NUM
biotropia-309	227	13	-	-	NUM
biotropia-309	227	14	11	11	NUM
biotropia-309	227	15	.	.	PUNCT
biotropia-309	228	1	http://www.dmns.org/media/5782/130-dmnstechrep2007-6.pdf	http://www.dmns.org/media/5782/130-dmnstechrep2007-6.pdf	NOUN
biotropia-309	228	2	kremen	kremen	PROPN
biotropia-309	228	3	c	c	PROPN
biotropia-309	228	4	,	,	PUNCT
biotropia-309	228	5	merenlander	merenlander	NOUN
biotropia-309	228	6	am	be	AUX
biotropia-309	228	7	,	,	PUNCT
biotropia-309	228	8	murphy	murphy	PROPN
biotropia-309	228	9	dd	dd	PROPN
biotropia-309	228	10	.	.	PROPN
biotropia-309	228	11	1994	1994	NUM
biotropia-309	228	12	.	.	PUNCT
biotropia-309	229	1	ecological	ecological	ADJ
biotropia-309	229	2	monitoring	monitoring	NOUN
biotropia-309	229	3	:	:	PUNCT
biotropia-309	229	4	a	a	DET
biotropia-309	229	5	vital	vital	ADJ
biotropia-309	229	6	need	need	NOUN
biotropia-309	229	7	forintegrated	forintegrate	VERB
biotropia-309	229	8	conservation	conservation	NOUN
biotropia-309	229	9	and	and	CCONJ
biotropia-309	229	10	development	development	NOUN
biotropia-309	229	11	programs	program	NOUN
biotropia-309	229	12	in	in	ADP
biotropia-309	229	13	the	the	DET
biotropia-309	229	14	tropics	tropic	NOUN
biotropia-309	229	15	.	.	PUNCT
biotropia-309	230	1	conservation	conservation	NOUN
biotropia-309	230	2	biology	biology	NOUN
biotropia-309	230	3	8	8	NUM
biotropia-309	230	4	:	:	PUNCT
biotropia-309	230	5	388	388	NUM
biotropia-309	230	6	-	-	SYM
biotropia-309	230	7	397	397	NUM
biotropia-309	230	8	.	.	PUNCT
biotropia-309	231	1	lande	lande	PROPN
biotropia-309	231	2	r.	r.	PROPN
biotropia-309	231	3	1996	1996	NUM
biotropia-309	231	4	.	.	PUNCT
biotropia-309	232	1	statistics	statistic	NOUN
biotropia-309	232	2	and	and	CCONJ
biotropia-309	232	3	partitioning	partitioning	NOUN
biotropia-309	232	4	of	of	ADP
biotropia-309	232	5	species	specie	NOUN
biotropia-309	232	6	diversity	diversity	NOUN
biotropia-309	232	7	,	,	PUNCT
biotropia-309	232	8	and	and	CCONJ
biotropia-309	232	9	similarity	similarity	NOUN
biotropia-309	232	10	among	among	ADP
biotropia-309	232	11	multiple	multiple	ADJ
biotropia-309	232	12	communities.oikos	communities.oikos	PROPN
biotropia-309	232	13	76	76	NUM
biotropia-309	232	14	:	:	PUNCT
biotropia-309	232	15	5	5	NUM
biotropia-309	232	16	-	-	SYM
biotropia-309	232	17	13	13	NUM
biotropia-309	232	18	.	.	PUNCT
biotropia-309	233	1	larsen	larsen	PROPN
biotropia-309	233	2	th	th	PROPN
biotropia-309	233	3	,	,	PUNCT
biotropia-309	233	4	forsyth	forsyth	PROPN
biotropia-309	233	5	a.	a.	NOUN
biotropia-309	233	6	2005	2005	NUM
biotropia-309	233	7	.	.	PUNCT
biotropia-309	234	1	trap	trap	NOUN
biotropia-309	234	2	spacing	spacing	NOUN
biotropia-309	234	3	and	and	CCONJ
biotropia-309	234	4	transect	transect	NOUN
biotropia-309	234	5	design	design	NOUN
biotropia-309	234	6	for	for	ADP
biotropia-309	234	7	dung	dung	NOUN
biotropia-309	234	8	beetle	beetle	NOUN
biotropia-309	234	9	biodiversity	biodiversity	NOUN
biotropia-309	234	10	studies	study	NOUN
biotropia-309	234	11	.	.	PUNCT
biotropia-309	235	1	biotropica	biotropica	PROPN
biotropia-309	235	2	37(2	37(2	PRON
biotropia-309	235	3	)	)	PUNCT
biotropia-309	235	4	:	:	PUNCT
biotropia-309	235	5	322	322	NUM
biotropia-309	235	6	-	-	SYM
biotropia-309	235	7	325	325	NUM
biotropia-309	235	8	mcgeoch	mcgeoch	PROPN
biotropia-309	235	9	ma	ma	PROPN
biotropia-309	235	10	.	.	PROPN
biotropia-309	235	11	1998	1998	NUM
biotropia-309	235	12	.	.	PUNCT
biotropia-309	236	1	the	the	DET
biotropia-309	236	2	selection	selection	NOUN
biotropia-309	236	3	,	,	PUNCT
biotropia-309	236	4	testing	testing	NOUN
biotropia-309	236	5	and	and	CCONJ
biotropia-309	236	6	application	application	NOUN
biotropia-309	236	7	of	of	ADP
biotropia-309	236	8	terrestrial	terrestrial	ADJ
biotropia-309	236	9	insects	insect	NOUN
biotropia-309	236	10	as	as	ADP
biotropia-309	236	11	bioindicators	bioindicator	NOUN
biotropia-309	236	12	.	.	PUNCT
biotropia-309	237	1	biological	biological	ADJ
biotropia-309	237	2	review	review	NOUN
biotropia-309	237	3	73	73	NUM
biotropia-309	237	4	:	:	PUNCT
biotropia-309	237	5	181	181	NUM
biotropia-309	237	6	-	-	SYM
biotropia-309	237	7	201	201	NUM
biotropia-309	237	8	.	.	PUNCT
biotropia-309	238	1	mcgeoch	mcgeoch	PROPN
biotropia-309	238	2	ma	ma	PROPN
biotropia-309	238	3	,	,	PUNCT
biotropia-309	238	4	van	van	PROPN
biotropia-309	238	5	rensburg	rensburg	PROPN
biotropia-309	238	6	bj	bj	PROPN
biotropia-309	238	7	,	,	PUNCT
biotropia-309	238	8	botes	botes	PROPN
biotropia-309	238	9	a.	a.	NOUN
biotropia-309	238	10	2002	2002	NUM
biotropia-309	238	11	.	.	PUNCT
biotropia-309	239	1	the	the	DET
biotropia-309	239	2	verification	verification	NOUN
biotropia-309	239	3	and	and	CCONJ
biotropia-309	239	4	application	application	NOUN
biotropia-309	239	5	of	of	ADP
biotropia-309	239	6	bioindicators	bioindicator	NOUN
biotropia-309	239	7	:	:	PUNCT
biotropia-309	239	8	a	a	DET
biotropia-309	239	9	case	case	NOUN
biotropia-309	239	10	study	study	NOUN
biotropia-309	239	11	of	of	ADP
biotropia-309	239	12	dung	dung	NOUN
biotropia-309	239	13	beetles	beetle	NOUN
biotropia-309	239	14	in	in	ADP
biotropia-309	239	15	a	a	DET
biotropia-309	239	16	savanna	savanna	NOUN
biotropia-309	239	17	ecosystem	ecosystem	NOUN
biotropia-309	239	18	.	.	PUNCT
biotropia-309	239	19	journal	journal	PROPN
biotropia-309	239	20	of	of	ADP
biotropia-309	239	21	applied	apply	VERB
biotropia-309	239	22	ecology39	ecology39	NOUN
biotropia-309	239	23	:	:	PUNCT
biotropia-309	239	24	661	661	NUM
biotropia-309	239	25	-	-	SYM
biotropia-309	239	26	672	672	NUM
biotropia-309	239	27	.	.	PUNCT
biotropia-309	240	1	mcgeoch	mcgeoch	PROPN
biotropia-309	240	2	ma	ma	PROPN
biotropia-309	240	3	,	,	PUNCT
biotropia-309	240	4	chown	chown	PROPN
biotropia-309	240	5	sl	sl	NOUN
biotropia-309	240	6	.	.	PROPN
biotropia-309	240	7	1998	1998	NUM
biotropia-309	240	8	.	.	PUNCT
biotropia-309	241	1	scaling	scale	VERB
biotropia-309	241	2	up	up	ADP
biotropia-309	241	3	the	the	DET
biotropia-309	241	4	value	value	NOUN
biotropia-309	241	5	obioindicators	obioindicator	NOUN
biotropia-309	241	6	.	.	PUNCT
biotropia-309	242	1	46	46	NUM
biotropia-309	242	2	-	-	SYM
biotropia-309	242	3	47	47	NUM
biotropia-309	242	4	.	.	PUNCT
biotropia-309	243	1	mcneely	mcneely	PROPN
biotropia-309	243	2	,	,	PUNCT
biotropia-309	243	3	ja	ja	PROPN
biotropia-309	243	4	and	and	CCONJ
biotropia-309	243	5	g	g	NOUN
biotropia-309	243	6	,	,	PUNCT
biotropia-309	243	7	schroth	schroth	ADJ
biotropia-309	243	8	.	.	PUNCT
biotropia-309	244	1	2006	2006	NUM
biotropia-309	244	2	.	.	PUNCT
biotropia-309	245	1	agroforestry	agroforestry	NOUN
biotropia-309	245	2	and	and	CCONJ
biotropia-309	245	3	biodiversity	biodiversity	NOUN
biotropia-309	245	4	conservation	conservation	NOUN
biotropia-309	245	5	traditional	traditional	ADJ
biotropia-309	245	6	practices	practice	NOUN
biotropia-309	245	7	,	,	PUNCT
biotropia-309	245	8	present	present	ADJ
biotropia-309	245	9	dynamics	dynamic	NOUN
biotropia-309	245	10	,	,	PUNCT
biotropia-309	245	11	and	and	CCONJ
biotropia-309	245	12	lessons	lesson	NOUN
biotropia-309	245	13	for	for	ADP
biotropia-309	245	14	the	the	DET
biotropia-309	245	15	future	future	NOUN
biotropia-309	245	16	.	.	PUNCT
biotropia-309	246	1	biodiversity	biodiversity	NOUN
biotropia-309	246	2	and	and	CCONJ
biotropia-309	246	3	conservation	conservation	NOUN
biotropia-309	246	4	15:549	15:549	NUM
biotropia-309	246	5	-	-	SYM
biotropia-309	246	6	554	554	NUM
biotropia-309	246	7	.	.	PUNCT
biotropia-309	247	1	mendoza	mendoza	PROPN
biotropia-309	247	2	cao	cao	PROPN
biotropia-309	247	3	,	,	PUNCT
biotropia-309	247	4	rios	rios	PROPN
biotropia-309	247	5	am	be	AUX
biotropia-309	247	6	,	,	PUNCT
biotropia-309	247	7	cano	cano	PROPN
biotropia-309	247	8	eb	eb	PROPN
biotropia-309	247	9	,	,	PUNCT
biotropia-309	247	10	corte	corte	PROPN
biotropia-309	247	11	's	's	PART
biotropia-309	247	12	jln	jln	NOUN
biotropia-309	247	13	.	.	PROPN
biotropia-309	247	14	2005	2005	NUM
biotropia-309	247	15	.	.	PUNCT
biotropia-309	248	1	dung	dung	NOUN
biotropia-309	248	2	beetle	beetle	NOUN
biotropia-309	248	3	community	community	NOUN
biotropia-309	248	4	(	(	PUNCT
biotropia-309	248	5	coleoptera	coleoptera	NOUN
biotropia-309	248	6	:	:	PUNCT
biotropia-309	248	7	scarabaeidae	scarabaeidae	NOUN
biotropia-309	248	8	:	:	PUNCT
biotropia-309	248	9	scarabaeinae	scarabaeinae	NOUN
biotropia-309	248	10	)	)	PUNCT
biotropia-309	248	11	in	in	ADP
biotropia-309	248	12	a	a	DET
biotropia-309	248	13	tropical	tropical	ADJ
biotropia-309	248	14	landscape	landscape	NOUN
biotropia-309	248	15	at	at	ADP
biotropia-309	248	16	the	the	DET
biotropia-309	248	17	lachua	lachua	PROPN
biotropia-309	248	18	region	region	PROPN
biotropia-309	248	19	,	,	PUNCT
biotropia-309	248	20	guatemala	guatemala	PROPN
biotropia-309	248	21	.	.	PUNCT
biotropia-309	249	1	biodiversity	biodiversity	NOUN
biotropia-309	249	2	and	and	CCONJ
biotropia-309	249	3	conservation	conservation	NOUN
biotropia-309	249	4	14	14	NUM
biotropia-309	249	5	:	:	PUNCT
biotropia-309	249	6	801	801	NUM
biotropia-309	249	7	-	-	SYM
biotropia-309	249	8	82	82	NUM
biotropia-309	249	9	.	.	PUNCT
biotropia-309	250	1	negro	negro	PROPN
biotropia-309	250	2	m	m	PROPN
biotropia-309	250	3	,	,	PUNCT
biotropia-309	250	4	palestrini	palestrini	PROPN
biotropia-309	250	5	c	c	PROPN
biotropia-309	250	6	,	,	PUNCT
biotropia-309	250	7	giraudo	giraudo	PROPN
biotropia-309	250	8	mt	mt	PROPN
biotropia-309	250	9	,	,	PUNCT
biotropia-309	250	10	rolando	rolando	PROPN
biotropia-309	250	11	a.	a.	PROPN
biotropia-309	250	12	2011	2011	NUM
biotropia-309	250	13	.	.	PUNCT
biotropia-309	251	1	the	the	DET
biotropia-309	251	2	effect	effect	NOUN
biotropia-309	251	3	of	of	ADP
biotropia-309	251	4	local	local	ADJ
biotropia-309	251	5	environmental	environmental	ADJ
biotropia-309	251	6	heterogeneity	heterogeneity	NOUN
biotropia-309	251	7	on	on	ADP
biotropia-309	251	8	species	specie	NOUN
biotropia-309	251	9	diversity	diversity	NOUN
biotropia-309	251	10	of	of	ADP
biotropia-309	251	11	alpine	alpine	NOUN
biotropia-309	251	12	dung	dung	NOUN
biotropia-309	251	13	beetles	beetle	NOUN
biotropia-309	251	14	(	(	PUNCT
biotropia-309	251	15	coleoptera	coleoptera	NOUN
biotropia-309	251	16	:	:	PUNCT
biotropia-309	251	17	scarabaeidae	scarabaeidae	NOUN
biotropia-309	251	18	)	)	PUNCT
biotropia-309	251	19	.	.	PUNCT
biotropia-309	252	1	eur	eur	PROPN
biotropia-309	252	2	.	.	PUNCT
biotropia-309	253	1	j.	j.	PROPN
biotropia-309	253	2	entomol	entomol	PROPN
biotropia-309	253	3	.	.	PUNCT
biotropia-309	254	1	108	108	NUM
biotropia-309	254	2	:	:	PUNCT
biotropia-309	254	3	91	91	NUM
biotropia-309	254	4	-	-	SYM
biotropia-309	254	5	98	98	NUM
biotropia-309	254	6	.	.	PUNCT
biotropia-309	255	1	nichols	nichols	PROPN
biotropia-309	255	2	e	e	PROPN
biotropia-309	255	3	,	,	PUNCT
biotropia-309	255	4	larsen	larsen	PROPN
biotropia-309	255	5	t	t	PROPN
biotropia-309	255	6	,	,	PUNCT
biotropia-309	255	7	spector	spector	PROPN
biotropia-309	255	8	s	s	PROPN
biotropia-309	255	9	,	,	PUNCT
biotropia-309	255	10	davis	davis	PROPN
biotropia-309	255	11	al	al	PROPN
biotropia-309	255	12	,	,	PUNCT
biotropia-309	255	13	escobar	escobar	PROPN
biotropia-309	255	14	f	f	PROPN
biotropia-309	255	15	,	,	PUNCT
biotropia-309	255	16	favila	favila	PROPN
biotropia-309	255	17	m	m	PROPN
biotropia-309	255	18	,	,	PUNCT
biotropia-309	255	19	vulinec	vulinec	PROPN
biotropia-309	255	20	k.	k.	PROPN
biotropia-309	255	21	2007	2007	NUM
biotropia-309	255	22	.	.	PUNCT
biotropia-309	256	1	global	global	ADJ
biotropia-309	256	2	dung	dung	NOUN
biotropia-309	256	3	beetle	beetle	NOUN
biotropia-309	256	4	response	response	NOUN
biotropia-309	256	5	to	to	ADP
biotropia-309	256	6	tropical	tropical	ADJ
biotropia-309	256	7	forest	forest	NOUN
biotropia-309	256	8	modification	modification	NOUN
biotropia-309	256	9	and	and	CCONJ
biotropia-309	256	10	fragmentation	fragmentation	NOUN
biotropia-309	256	11	:	:	PUNCT
biotropia-309	256	12	a	a	DET
biotropia-309	256	13	quantitative	quantitative	ADJ
biotropia-309	256	14	literature	literature	NOUN
biotropia-309	256	15	review	review	NOUN
biotropia-309	256	16	and	and	CCONJ
biotropia-309	256	17	meta	meta	ADJ
biotropia-309	256	18	-	-	PUNCT
biotropia-309	256	19	analysis	analysis	NOUN
biotropia-309	256	20	.	.	PUNCT
biotropia-309	257	1	biological	biological	ADJ
biotropia-309	257	2	conservation	conservation	NOUN
biotropia-309	257	3	137	137	NUM
biotropia-309	257	4	:	:	PUNCT
biotropia-309	257	5	1	1	NUM
biotropia-309	257	6	-	-	SYM
biotropia-309	257	7	19	19	NUM
biotropia-309	257	8	.	.	PUNCT
biotropia-309	258	1	nichols	nichols	PROPN
biotropia-309	258	2	e	e	PROPN
biotropia-309	258	3	,	,	PUNCT
biotropia-309	258	4	spector	spector	PROPN
biotropia-309	258	5	s	s	PROPN
biotropia-309	258	6	,	,	PUNCT
biotropia-309	258	7	louzada	louzada	PROPN
biotropia-309	258	8	j	j	PROPN
biotropia-309	258	9	,	,	PUNCT
biotropia-309	258	10	larsen	larsen	PROPN
biotropia-309	258	11	t	t	PROPN
biotropia-309	258	12	,	,	PUNCT
biotropia-309	258	13	amesquita	amesquita	NOUN
biotropia-309	258	14	s	s	PART
biotropia-309	258	15	,	,	PUNCT
biotropia-309	258	16	favila	favila	VERB
biotropia-309	258	17	me	i	PRON
biotropia-309	258	18	.	.	PUNCT
biotropia-309	259	1	2008	2008	NUM
biotropia-309	259	2	.	.	PUNCT
biotropia-309	260	1	ecological	ecological	ADJ
biotropia-309	260	2	functions	function	NOUN
biotropia-309	260	3	and	and	CCONJ
biotropia-309	260	4	ecosystem	ecosystem	NOUN
biotropia-309	260	5	services	service	NOUN
biotropia-309	260	6	provided	provide	VERB
biotropia-309	260	7	by	by	ADP
biotropia-309	260	8	scarabaeinae	scarabaeinae	ADJ
biotropia-309	260	9	dung	dung	NOUN
biotropia-309	260	10	beetles	beetle	NOUN
biotropia-309	260	11	.	.	PUNCT
biotropia-309	261	1	biological	biological	ADJ
biotropia-309	261	2	conservation	conservation	NOUN
biotropia-309	261	3	141	141	NUM
biotropia-309	261	4	(	(	PUNCT
biotropia-309	261	5	6	6	NUM
biotropia-309	261	6	):	):	PUNCT
biotropia-309	261	7	1461	1461	NUM
biotropia-309	261	8	-	-	SYM
biotropia-309	261	9	1474	1474	NUM
biotropia-309	261	10	.	.	PUNCT
biotropia-309	262	1	noss	noss	PROPN
biotropia-309	262	2	rf	rf	PROPN
biotropia-309	262	3	.	.	PROPN
biotropia-309	262	4	1990	1990	NUM
biotropia-309	262	5	.	.	PUNCT
biotropia-309	263	1	indicators	indicator	NOUN
biotropia-309	263	2	for	for	ADP
biotropia-309	263	3	monitoring	monitor	VERB
biotropia-309	263	4	biodiversity	biodiversity	NOUN
biotropia-309	263	5	:	:	PUNCT
biotropia-309	263	6	a	a	DET
biotropia-309	263	7	hierarchical	hierarchical	ADJ
biotropia-309	263	8	approach	approach	NOUN
biotropia-309	263	9	.	.	PUNCT
biotropia-309	264	1	conservation	conservation	NOUN
biotropia-309	264	2	biology4	biology4	PROPN
biotropia-309	264	3	:	:	PUNCT
biotropia-309	264	4	355	355	NUM
biotropia-309	264	5	-	-	SYM
biotropia-309	264	6	364	364	NUM
biotropia-309	264	7	.	.	PUNCT
biotropia-309	264	8	.	.	PUNCT
biotropia-309	264	9	.	.	PUNCT
biotropia-309	265	1	trends	trend	NOUN
biotropia-309	265	2	in	in	ADP
biotropia-309	265	3	ecology	ecology	NOUN
biotropia-309	265	4	and	and	CCONJ
biotropia-309	265	5	evolution13	evolution13	NOUN
biotropia-309	265	6	:	:	PUNCT
biotropia-309	265	7	57	57	NUM
biotropia-309	265	8	effectiveness	effectiveness	NOUN
biotropia-309	265	9	of	of	ADP
biotropia-309	265	10	dung	dung	NOUN
biotropia-309	265	11	beetles	beetle	NOUN
biotropia-309	265	12	as	as	ADP
biotropia-309	265	13	bioindicators	bioindicator	NOUN
biotropia-309	265	14	of	of	ADP
biotropia-309	265	15	environmental	environmental	ADJ
biotropia-309	265	16	changes	change	NOUN
biotropia-309	265	17	in	in	ADP
biotropia-309	265	18	land	land	NOUN
biotropia-309	265	19	shahabuddin	shahabuddin	NOUN
biotropia-309	265	20	–	–	PUNCT
biotropia-309	265	21	et	et	PROPN
biotropia-309	265	22	al	al	PROPN
biotropia-309	265	23	.	.	PROPN
biotropia-309	265	24	pearson	pearson	PROPN
biotropia-309	265	25	dl	dl	PROPN
biotropia-309	265	26	,	,	PUNCT
biotropia-309	265	27	cassola	cassola	PROPN
biotropia-309	265	28	f.	f.	PROPN
biotropia-309	265	29	1992	1992	NUM
biotropia-309	265	30	.	.	PUNCT
biotropia-309	266	1	world	world	NOUN
biotropia-309	266	2	-	-	PUNCT
biotropia-309	266	3	wide	wide	ADJ
biotropia-309	266	4	species	specie	NOUN
biotropia-309	266	5	richness	richness	NOUN
biotropia-309	266	6	patterns	pattern	NOUN
biotropia-309	266	7	of	of	ADP
biotropia-309	266	8	tiger	tiger	NOUN
biotropia-309	266	9	beetles	beetle	NOUN
biotropia-309	266	10	(	(	PUNCT
biotropia-309	266	11	coleoptera	coleoptera	NOUN
biotropia-309	266	12	:	:	PUNCT
biotropia-309	266	13	cicindelidae	cicindelidae	PROPN
biotropia-309	266	14	):	):	PUNCT
biotropia-309	266	15	indicator	indicator	NOUN
biotropia-309	266	16	taxon	taxon	ADJ
biotropia-309	266	17	for	for	ADP
biotropia-309	266	18	biodiversity	biodiversity	NOUN
biotropia-309	266	19	and	and	CCONJ
biotropia-309	266	20	conservation	conservation	NOUN
biotropia-309	266	21	studies	study	NOUN
biotropia-309	266	22	.	.	PUNCT
biotropia-309	267	1	conservation	conservation	NOUN
biotropia-309	267	2	biology	biology	NOUN
biotropia-309	267	3	6:376	6:376	NOUN
biotropia-309	267	4	-	-	SYM
biotropia-309	267	5	391	391	NUM
biotropia-309	267	6	.	.	PUNCT
biotropia-309	268	1	rainio	rainio	PROPN
biotropia-309	268	2	j	j	PROPN
biotropia-309	268	3	,	,	PUNCT
biotropia-309	268	4	niemelä	niemelä	NOUN
biotropia-309	268	5	j.	j.	PROPN
biotropia-309	268	6	2003	2003	NUM
biotropia-309	268	7	.	.	PUNCT
biotropia-309	269	1	ground	ground	NOUN
biotropia-309	269	2	beetles	beetle	NOUN
biotropia-309	269	3	(	(	PUNCT
biotropia-309	269	4	coleoptera	coleoptera	NOUN
biotropia-309	269	5	:	:	PUNCT
biotropia-309	269	6	carabidae	carabidae	NOUN
biotropia-309	269	7	)	)	PUNCT
biotropia-309	269	8	as	as	ADP
biotropia-309	269	9	bioindicators.biodiversityand	bioindicators.biodiversityand	PROPN
biotropia-309	269	10	conservation12	conservation12	VERB
biotropia-309	269	11	:	:	PUNCT
biotropia-309	269	12	487	487	NUM
biotropia-309	269	13	-	-	SYM
biotropia-309	269	14	506	506	NUM
biotropia-309	269	15	.	.	PUNCT
biotropia-309	270	1	sala	sala	PROPN
biotropia-309	270	2	oe	oe	PROPN
biotropia-309	270	3	,	,	PUNCT
biotropia-309	270	4	chapin	chapin	PROPN
biotropia-309	270	5	fs	fs	PROPN
biotropia-309	270	6	,	,	PUNCT
biotropia-309	270	7	armesto	armesto	PROPN
biotropia-309	270	8	jj	jj	PROPN
biotropia-309	270	9	,	,	PUNCT
biotropia-309	270	10	berlow	berlow	NOUN
biotropia-309	270	11	e	e	NOUN
biotropia-309	270	12	,	,	PUNCT
biotropia-309	270	13	bloomfield	bloomfield	PROPN
biotropia-309	270	14	j	j	PROPN
biotropia-309	270	15	,	,	PUNCT
biotropia-309	270	16	dirzo	dirzo	PROPN
biotropia-309	270	17	r	r	NOUN
biotropia-309	270	18	,	,	PUNCT
biotropia-309	270	19	huber	huber	PROPN
biotropia-309	270	20	-	-	PUNCT
biotropia-309	270	21	sanwald	sanwald	PROPN
biotropia-309	270	22	e	e	NOUN
biotropia-309	270	23	,	,	PUNCT
biotropia-309	270	24	huenneke	huenneke	PROPN
biotropia-309	270	25	lf	lf	PROPN
biotropia-309	270	26	,	,	PUNCT
biotropia-309	270	27	jackson	jackson	PROPN
biotropia-309	270	28	rb	rb	PROPN
biotropia-309	270	29	,	,	PUNCT
biotropia-309	270	30	kinzig	kinzig	PROPN
biotropia-309	270	31	a	a	PROPN
biotropia-309	270	32	,	,	PUNCT
biotropia-309	270	33	leemans	leemans	PROPN
biotropia-309	270	34	r	r	NOUN
biotropia-309	270	35	,	,	PUNCT
biotropia-309	270	36	lodge	lodge	PROPN
biotropia-309	270	37	dm	dm	PROPN
biotropia-309	270	38	,	,	PUNCT
biotropia-309	270	39	mooney	mooney	PROPN
biotropia-309	270	40	ha	ha	PROPN
biotropia-309	270	41	,	,	PUNCT
biotropia-309	270	42	oesterheld	oesterheld	PROPN
biotropia-309	270	43	m	m	PROPN
biotropia-309	270	44	,	,	PUNCT
biotropia-309	270	45	poff	poff	PROPN
biotropia-309	270	46	nl	nl	PROPN
biotropia-309	270	47	,	,	PUNCT
biotropia-309	270	48	sykes	sykes	PROPN
biotropia-309	270	49	mt	mt	PROPN
biotropia-309	270	50	,	,	PUNCT
biotropia-309	270	51	walker	walker	PROPN
biotropia-309	270	52	bh	bh	PROPN
biotropia-309	270	53	,	,	PUNCT
biotropia-309	270	54	walker	walker	PROPN
biotropia-309	270	55	m	m	PROPN
biotropia-309	270	56	,	,	PUNCT
biotropia-309	270	57	wall	wall	PROPN
biotropia-309	270	58	dh	dh	PROPN
biotropia-309	270	59	.	.	PROPN
biotropia-309	270	60	2000	2000	NUM
biotropia-309	270	61	.	.	PUNCT
biotropia-309	271	1	review	review	NOUN
biotropia-309	271	2	:	:	PUNCT
biotropia-309	271	3	global	global	ADJ
biotropia-309	271	4	biodiversity	biodiversity	NOUN
biotropia-309	271	5	scenarios	scenario	NOUN
biotropia-309	271	6	for	for	ADP
biotropia-309	271	7	the	the	DET
biotropia-309	271	8	year	year	NOUN
biotropia-309	271	9	2100	2100	NUM
biotropia-309	271	10	.	.	PUNCT
biotropia-309	272	1	science	science	NOUN
biotropia-309	272	2	287	287	NUM
biotropia-309	272	3	:	:	PUNCT
biotropia-309	272	4	1770	1770	NUM
biotropia-309	272	5	-	-	SYM
biotropia-309	272	6	1774	1774	NUM
biotropia-309	272	7	.	.	PUNCT
biotropia-309	273	1	schulze	schulze	PROPN
biotropia-309	273	2	ch	ch	PROPN
biotropia-309	273	3	,	,	PUNCT
biotropia-309	273	4	schneeweihs	schneeweihs	VERB
biotropia-309	273	5	s	s	PART
biotropia-309	273	6	,	,	PUNCT
biotropia-309	273	7	fiedler	fiedler	PROPN
biotropia-309	273	8	k.	k.	PROPN
biotropia-309	274	1	the	the	DET
biotropia-309	274	2	potential	potential	NOUN
biotropia-309	274	3	of	of	ADP
biotropia-309	274	4	land	land	NOUN
biotropia-309	274	5	-	-	PUNCT
biotropia-309	274	6	use	use	NOUN
biotropia-309	274	7	systems	system	NOUN
biotropia-309	274	8	for	for	ADP
biotropia-309	274	9	maintaining	maintain	VERB
biotropia-309	274	10	tropical	tropical	ADJ
biotropia-309	274	11	forest	forest	NOUN
biotropia-309	274	12	butterfly	butterfly	NOUN
biotropia-309	274	13	diversity	diversity	NOUN
biotropia-309	274	14	.	.	PUNCT
biotropia-309	275	1	in	in	ADP
biotropia-309	275	2	tropical	tropical	ADJ
biotropia-309	275	3	rainforests	rainforest	NOUN
biotropia-309	275	4	and	and	CCONJ
biotropia-309	275	5	agroforests	agroforest	NOUN
biotropia-309	275	6	under	under	ADP
biotropia-309	275	7	global	global	ADJ
biotropia-309	275	8	change	change	NOUN
biotropia-309	275	9	73	73	NUM
biotropia-309	275	10	-	-	SYM
biotropia-309	275	11	96	96	NUM
biotropia-309	275	12	.	.	PUNCT
biotropia-309	275	13	edited	edit	VERB
biotropia-309	275	14	by	by	ADP
biotropia-309	275	15	tschartnke	tschartnke	NOUN
biotropia-309	275	16	.	.	PUNCT
biotropia-309	276	1	new	new	PROPN
biotropia-309	276	2	york	york	PROPN
biotropia-309	276	3	:	:	PUNCT
biotropia-309	276	4	springer	springer	NOUN
biotropia-309	276	5	2010:73	2010:73	PROPN
biotropia-309	276	6	-	-	SYM
biotropia-309	276	7	96	96	NUM
biotropia-309	276	8	.	.	PUNCT
biotropia-309	277	1	shahabuddin	shahabuddin	NOUN
biotropia-309	277	2	,	,	PUNCT
biotropia-309	277	3	schulze	schulze	NOUN
biotropia-309	277	4	ch	ch	NOUN
biotropia-309	277	5	,	,	PUNCT
biotropia-309	277	6	tscharntke	tscharntke	NOUN
biotropia-309	277	7	t.	t.	PROPN
biotropia-309	277	8	2005	2005	NUM
biotropia-309	277	9	.	.	PUNCT
biotropia-309	278	1	changes	change	NOUN
biotropia-309	278	2	of	of	ADP
biotropia-309	278	3	dung	dung	NOUN
biotropia-309	278	4	beetle	beetle	NOUN
biotropia-309	278	5	communities	community	NOUN
biotropia-309	278	6	from	from	ADP
biotropia-309	278	7	rainforests	rainforest	NOUN
biotropia-309	278	8	towards	towards	ADP
biotropia-309	278	9	agroforestry	agroforestry	NOUN
biotropia-309	278	10	systems	system	NOUN
biotropia-309	278	11	and	and	CCONJ
biotropia-309	278	12	annual	annual	ADJ
biotropia-309	278	13	cultures	culture	NOUN
biotropia-309	278	14	.	.	PUNCT
biotropia-309	279	1	biodiversity	biodiversity	NOUN
biotropia-309	279	2	and	and	CCONJ
biotropia-309	279	3	conservation	conservation	NOUN
biotropia-309	279	4	14	14	NUM
biotropia-309	279	5	:	:	PUNCT
biotropia-309	279	6	863	863	NUM
biotropia-309	279	7	-	-	SYM
biotropia-309	279	8	877	877	NUM
biotropia-309	279	9	.	.	PUNCT
biotropia-309	280	1	shahabuddin	shahabuddin	VERB
biotropia-309	280	2	.	.	PUNCT
biotropia-309	281	1	2010	2010	NUM
biotropia-309	281	2	.	.	PUNCT
biotropia-309	282	1	diversity	diversity	NOUN
biotropia-309	282	2	and	and	CCONJ
biotropia-309	282	3	community	community	NOUN
biotropia-309	282	4	structure	structure	NOUN
biotropia-309	282	5	of	of	ADP
biotropia-309	282	6	dung	dung	NOUN
biotropia-309	282	7	beetles	beetle	NOUN
biotropia-309	282	8	(	(	PUNCT
biotropia-309	282	9	coleoptera	coleoptera	NOUN
biotropia-309	282	10	:	:	PUNCT
biotropia-309	282	11	scarabaeidae	scarabaeidae	NOUN
biotropia-309	282	12	)	)	PUNCT
biotropia-309	282	13	across	across	ADP
biotropia-309	282	14	a	a	DET
biotropia-309	282	15	habitat	habitat	NOUN
biotropia-309	282	16	disturbance	disturbance	NOUN
biotropia-309	282	17	gradient	gradient	NOUN
biotropia-309	282	18	in	in	ADP
biotropia-309	282	19	lore	lore	PROPN
biotropia-309	282	20	lindu	lindu	PROPN
biotropia-309	282	21	national	national	PROPN
biotropia-309	282	22	park	park	PROPN
biotropia-309	282	23	,	,	PUNCT
biotropia-309	282	24	central	central	ADJ
biotropia-309	282	25	sulawesi	sulawesi	NOUN
biotropia-309	282	26	.	.	PUNCT
biotropia-309	283	1	11	11	NUM
biotropia-309	283	2	(	(	PUNCT
biotropia-309	283	3	1	1	NUM
biotropia-309	283	4	):	):	PUNCT
biotropia-309	283	5	29	29	NUM
biotropia-309	283	6	-	-	SYM
biotropia-309	283	7	33	33	NUM
biotropia-309	283	8	.	.	PUNCT
biotropia-309	284	1	shahabuddin	shahabuddin	NOUN
biotropia-309	284	2	,	,	PUNCT
biotropia-309	284	3	hidayat	hidayat	PROPN
biotropia-309	284	4	p	p	PROPN
biotropia-309	284	5	,	,	PUNCT
biotropia-309	284	6	noerdjito	noerdjito	X
biotropia-309	284	7	wa	wa	PROPN
biotropia-309	284	8	,	,	PUNCT
biotropia-309	284	9	manuwoto	manuwoto	PROPN
biotropia-309	284	10	s	s	PART
biotropia-309	284	11	,	,	PUNCT
biotropia-309	284	12	schulze	schulze	NOUN
biotropia-309	284	13	ch	ch	PROPN
biotropia-309	284	14	.	.	PROPN
biotropia-309	284	15	2010	2010	NUM
biotropia-309	284	16	.	.	PUNCT
biotropia-309	285	1	diversity	diversity	NOUN
biotropia-309	285	2	and	and	CCONJ
biotropia-309	285	3	body	body	NOUN
biotropia-309	285	4	size	size	NOUN
biotropia-309	285	5	of	of	ADP
biotropia-309	285	6	dung	dung	NOUN
biotropia-309	285	7	beetles	beetle	NOUN
biotropia-309	285	8	attracted	attract	VERB
biotropia-309	285	9	to	to	ADP
biotropia-309	285	10	different	different	ADJ
biotropia-309	285	11	dung	dung	NOUN
biotropia-309	285	12	types	type	NOUN
biotropia-309	285	13	along	along	ADP
biotropia-309	285	14	a	a	DET
biotropia-309	285	15	tropical	tropical	ADJ
biotropia-309	285	16	land	land	NOUN
biotropia-309	285	17	-	-	PUNCT
biotropia-309	285	18	use	use	NOUN
biotropia-309	285	19	gradient	gradient	NOUN
biotropia-309	285	20	in	in	ADP
biotropia-309	285	21	sulawesi	sulawesi	PROPN
biotropia-309	285	22	,	,	PUNCT
biotropia-309	285	23	indonesia	indonesia	PROPN
biotropia-309	285	24	.	.	PROPN
biotropia-309	286	1	journal	journal	PROPN
biotropia-309	286	2	of	of	ADP
biotropia-309	286	3	tropical	tropical	ADJ
biotropia-309	286	4	ecology	ecology	NOUN
biotropia-309	286	5	26	26	NUM
biotropia-309	286	6	(	(	PUNCT
biotropia-309	286	7	1	1	NUM
biotropia-309	286	8	)	)	PUNCT
biotropia-309	286	9	:	:	PUNCT
biotropia-309	286	10	53	53	NUM
biotropia-309	286	11	-	-	SYM
biotropia-309	286	12	65	65	NUM
biotropia-309	286	13	.	.	PUNCT
biotropia-309	287	1	shahabuddin	shahabuddin	VERB
biotropia-309	287	2	.	.	PUNCT
biotropia-309	288	1	2011	2011	NUM
biotropia-309	288	2	.	.	PUNCT
biotropia-309	289	1	effect	effect	NOUN
biotropia-309	289	2	of	of	ADP
biotropia-309	289	3	land	land	NOUN
biotropia-309	289	4	use	use	NOUN
biotropia-309	289	5	change	change	NOUN
biotropia-309	289	6	on	on	ADP
biotropia-309	289	7	ecosystem	ecosystem	NOUN
biotropia-309	289	8	function	function	NOUN
biotropia-309	289	9	of	of	ADP
biotropia-309	289	10	dung	dung	NOUN
biotropia-309	289	11	beetles	beetle	NOUN
biotropia-309	289	12	:	:	PUNCT
biotropia-309	289	13	experimental	experimental	ADJ
biotropia-309	289	14	evidence	evidence	NOUN
biotropia-309	289	15	from	from	ADP
biotropia-309	289	16	wallacea	wallacea	PROPN
biotropia-309	289	17	region	region	PROPN
biotropia-309	289	18	in	in	ADP
biotropia-309	289	19	sulawesi	sulawesi	PROPN
biotropia-309	289	20	,	,	PUNCT
biotropia-309	289	21	indonesia.jurnalbiodiversitas	indonesia.jurnalbiodiversitas	NUM
biotropia-309	289	22	12	12	NUM
biotropia-309	289	23	(	(	PUNCT
biotropia-309	289	24	3	3	NUM
biotropia-309	289	25	):	):	PUNCT
biotropia-309	289	26	177	177	NUM
biotropia-309	289	27	-	-	SYM
biotropia-309	289	28	181	181	NUM
biotropia-309	289	29	shahabuddin	shahabuddin	NOUN
biotropia-309	289	30	.	.	PUNCT
biotropia-309	290	1	2013	2013	NUM
biotropia-309	290	2	.	.	PUNCT
biotropia-309	291	1	contribution	contribution	NOUN
biotropia-309	291	2	of	of	ADP
biotropia-309	291	3	alpha	alpha	NOUN
biotropia-309	291	4	and	and	CCONJ
biotropia-309	291	5	beta	beta	NOUN
biotropia-309	291	6	diversity	diversity	NOUN
biotropia-309	291	7	across	across	ADP
biotropia-309	291	8	land	land	NOUN
biotropia-309	291	9	-	-	PUNCT
biotropia-309	291	10	use	use	NOUN
biotropia-309	291	11	type	type	NOUN
biotropia-309	291	12	to	to	ADP
biotropia-309	291	13	the	the	DET
biotropia-309	291	14	regional	regional	ADJ
biotropia-309	291	15	diversity	diversity	NOUN
biotropia-309	291	16	of	of	ADP
biotropia-309	291	17	dung	dung	NOUN
biotropia-309	291	18	beetles	beetle	NOUN
biotropia-309	291	19	in	in	ADP
biotropia-309	291	20	central	central	ADJ
biotropia-309	291	21	sulawesi	sulawesi	NOUN
biotropia-309	291	22	.	.	PUNCT
biotropia-309	292	1	hayati	hayati	PROPN
biotropia-309	292	2	journal	journal	PROPN
biotropia-309	292	3	of	of	ADP
biotropia-309	292	4	biosciences	bioscience	NOUN
biotropia-309	292	5	20	20	NUM
biotropia-309	292	6	(	(	PUNCT
biotropia-309	292	7	2):72	2):72	PROPN
biotropia-309	292	8	-	-	SYM
biotropia-309	292	9	79	79	NUM
biotropia-309	292	10	.	.	PUNCT
biotropia-309	293	1	slade	slade	PROPN
biotropia-309	293	2	em	em	PRON
biotropia-309	293	3	,	,	PUNCT
biotropia-309	293	4	mann	mann	PROPN
biotropia-309	293	5	dj	dj	PROPN
biotropia-309	293	6	,	,	PUNCT
biotropia-309	293	7	villanueva	villanueva	PROPN
biotropia-309	293	8	jf	jf	PROPN
biotropia-309	293	9	,	,	PUNCT
biotropia-309	293	10	lewis	lewis	PROPN
biotropia-309	293	11	ot	ot	PROPN
biotropia-309	293	12	.	.	PROPN
biotropia-309	293	13	2007	2007	NUM
biotropia-309	293	14	.	.	PUNCT
biotropia-309	294	1	experimental	experimental	ADJ
biotropia-309	294	2	evidence	evidence	NOUN
biotropia-309	294	3	for	for	ADP
biotropia-309	294	4	the	the	DET
biotropia-309	294	5	effects	effect	NOUN
biotropia-309	294	6	of	of	ADP
biotropia-309	294	7	dung	dung	NOUN
biotropia-309	294	8	beetle	beetle	VERB
biotropia-309	294	9	functional	functional	ADJ
biotropia-309	294	10	group	group	NOUN
biotropia-309	294	11	richness	richness	NOUN
biotropia-309	294	12	and	and	CCONJ
biotropia-309	294	13	composition	composition	NOUN
biotropia-309	294	14	on	on	ADP
biotropia-309	294	15	ecosystem	ecosystem	NOUN
biotropia-309	294	16	function	function	NOUN
biotropia-309	294	17	in	in	ADP
biotropia-309	294	18	a	a	DET
biotropia-309	294	19	tropical	tropical	ADJ
biotropia-309	294	20	forest	forest	NOUN
biotropia-309	294	21	.	.	PUNCT
biotropia-309	295	1	journal	journal	NOUN
biotropia-309	295	2	of	of	ADP
biotropia-309	295	3	animal	animal	NOUN
biotropia-309	295	4	ecology76	ecology76	NOUN
biotropia-309	295	5	:	:	PUNCT
biotropia-309	295	6	1094	1094	NUM
biotropia-309	295	7	-	-	SYM
biotropia-309	295	8	1104	1104	NUM
biotropia-309	295	9	.	.	PUNCT
biotropia-309	296	1	slade	slade	PROPN
biotropia-309	296	2	em	em	PRON
biotropia-309	296	3	,	,	PUNCT
biotropia-309	296	4	mann	mann	PROPN
biotropia-309	296	5	dj	dj	PROPN
biotropia-309	296	6	,	,	PUNCT
biotropia-309	296	7	lewis	lewis	PROPN
biotropia-309	296	8	ot	ot	PROPN
biotropia-309	296	9	.	.	PROPN
biotropia-309	296	10	2011	2011	NUM
biotropia-309	296	11	.	.	PUNCT
biotropia-309	297	1	biodiversity	biodiversity	NOUN
biotropia-309	297	2	and	and	CCONJ
biotropia-309	297	3	ecosystem	ecosystem	NOUN
biotropia-309	297	4	function	function	NOUN
biotropia-309	297	5	of	of	ADP
biotropia-309	297	6	tropical	tropical	ADJ
biotropia-309	297	7	forest	forest	NOUN
biotropia-309	297	8	dung	dung	NOUN
biotropia-309	297	9	beetles	beetle	NOUN
biotropia-309	297	10	under	under	ADP
biotropia-309	297	11	contrasting	contrast	VERB
biotropia-309	297	12	logging	log	VERB
biotropia-309	297	13	regimes	regime	NOUN
biotropia-309	297	14	.	.	PUNCT
biotropia-309	298	1	144	144	NUM
biotropia-309	298	2	:	:	SYM
biotropia-309	298	3	166	166	NUM
biotropia-309	298	4	-	-	SYM
biotropia-309	298	5	174	174	NUM
biotropia-309	298	6	.	.	PUNCT
biotropia-309	299	1	sodhi	sodhi	NOUN
biotropia-309	299	2	ns	ns	NUM
biotropia-309	299	3	,	,	PUNCT
biotropia-309	299	4	koh	koh	PROPN
biotropia-309	299	5	lp	lp	PROPN
biotropia-309	299	6	,	,	PUNCT
biotropia-309	299	7	brook	brook	PROPN
biotropia-309	299	8	bw	bw	PROPN
biotropia-309	299	9	,	,	PUNCT
biotropia-309	299	10	peter	peter	PROPN
biotropia-309	299	11	klng	klng	PROPN
biotropia-309	299	12	.	.	PUNCT
biotropia-309	300	1	2004	2004	NUM
biotropia-309	300	2	.	.	PUNCT
biotropia-309	301	1	southeast	southeast	ADJ
biotropia-309	301	2	asian	asian	ADJ
biotropia-309	301	3	biodiversity	biodiversity	NOUN
biotropia-309	301	4	:	:	PUNCT
biotropia-309	301	5	the	the	DET
biotropia-309	301	6	impending	impending	ADJ
biotropia-309	301	7	disaster	disaster	NOUN
biotropia-309	301	8	.	.	PUNCT
biotropia-309	302	1	trends	trend	NOUN
biotropia-309	302	2	in	in	ADP
biotropia-309	302	3	ecology	ecology	NOUN
biotropia-309	302	4	and	and	CCONJ
biotropia-309	302	5	evolution	evolution	NOUN
biotropia-309	302	6	.	.	PUNCT
biotropia-309	303	1	19	19	NUM
biotropia-309	303	2	(	(	PUNCT
biotropia-309	303	3	12	12	NUM
biotropia-309	303	4	)	)	PUNCT
biotropia-309	303	5	:	:	PUNCT
biotropia-309	303	6	654	654	NUM
biotropia-309	303	7	-	-	SYM
biotropia-309	303	8	660	660	NUM
biotropia-309	303	9	.	.	PUNCT
biotropia-309	304	1	spector	spector	PROPN
biotropia-309	304	2	s.	s.	PROPN
biotropia-309	304	3	2006	2006	NUM
biotropia-309	304	4	.	.	PUNCT
biotropia-309	305	1	scarabaeine	scarabaeine	NOUN
biotropia-309	305	2	dung	dung	NOUN
biotropia-309	305	3	beetles	beetle	NOUN
biotropia-309	305	4	(	(	PUNCT
biotropia-309	305	5	coleoptera	coleoptera	NOUN
biotropia-309	305	6	:	:	PUNCT
biotropia-309	305	7	scarabaeidae	scarabaeidae	NOUN
biotropia-309	305	8	:	:	PUNCT
biotropia-309	305	9	scarabaeinae	scarabaeinae	ADV
biotropia-309	305	10	):	):	PUNCT
biotropia-309	305	11	an	an	DET
biotropia-309	305	12	invertebrate	invertebrate	ADJ
biotropia-309	305	13	focal	focal	ADJ
biotropia-309	305	14	taxon	taxon	NOUN
biotropia-309	305	15	for	for	ADP
biotropia-309	305	16	biodiversity	biodiversity	NOUN
biotropia-309	305	17	research	research	NOUN
biotropia-309	305	18	and	and	CCONJ
biotropia-309	305	19	conservation	conservation	NOUN
biotropia-309	305	20	.	.	PUNCT
biotropia-309	306	1	the	the	DET
biotropia-309	306	2	coleopterists	coleopterist	NOUN
biotropia-309	306	3	bulletin	bulletin	VERB
biotropia-309	306	4	5	5	NUM
biotropia-309	306	5	:	:	SYM
biotropia-309	306	6	71	71	NUM
biotropia-309	306	7	-	-	SYM
biotropia-309	306	8	78	78	NUM
biotropia-309	306	9	.	.	PUNCT
biotropia-309	307	1	spellerberg	spellerberg	PROPN
biotropia-309	307	2	if	if	SCONJ
biotropia-309	307	3	.	.	PROPN
biotropia-309	307	4	1993	1993	NUM
biotropia-309	307	5	.	.	PUNCT
biotropia-309	308	1	monitoring	monitor	VERB
biotropia-309	308	2	ecological	ecological	ADJ
biotropia-309	308	3	change	change	NOUN
biotropia-309	308	4	.	.	PUNCT
biotropia-309	309	1	cambridge	cambridge	PROPN
biotropia-309	309	2	university	university	PROPN
biotropia-309	309	3	press	press	PROPN
biotropia-309	309	4	,	,	PUNCT
biotropia-309	309	5	cambridge	cambridge	PROPN
biotropia-309	309	6	,	,	PUNCT
biotropia-309	309	7	uk	uk	PROPN
biotropia-309	309	8	.	.	PROPN
biotropia-309	309	9	statsoft	statsoft	PROPN
biotropia-309	309	10	inc	inc	PROPN
biotropia-309	309	11	.	.	PROPN
biotropia-309	309	12	2001	2001	NUM
biotropia-309	309	13	.	.	PUNCT
biotropia-309	310	1	statistica	statistica	PROPN
biotropia-309	310	2	(	(	PUNCT
biotropia-309	310	3	data	datum	NOUN
biotropia-309	310	4	analysis	analysis	NOUN
biotropia-309	310	5	software	software	NOUN
biotropia-309	310	6	system	system	NOUN
biotropia-309	310	7	)	)	PUNCT
biotropia-309	310	8	,	,	PUNCT
biotropia-309	310	9	version	version	NOUN
biotropia-309	310	10	6.www.statsoft.com	6.www.statsoft.com	NOUN
biotropia-309	310	11	.	.	PUNCT
biotropia-309	310	12	tscharntke	tscharntke	PROPN
biotropia-309	310	13	t	t	PROPN
biotropia-309	310	14	,	,	PUNCT
biotropia-309	310	15	klein	klein	PROPN
biotropia-309	310	16	am	am	PROPN
biotropia-309	310	17	,	,	PUNCT
biotropia-309	310	18	kruess	kruess	VERB
biotropia-309	310	19	a	a	PRON
biotropia-309	310	20	,	,	PUNCT
biotropia-309	310	21	steffan	steffan	NOUN
biotropia-309	310	22	-	-	PUNCT
biotropia-309	310	23	dewenter	dewenter	ADJ
biotropia-309	310	24	i	i	NOUN
biotropia-309	310	25	,	,	PUNCT
biotropia-309	310	26	thies	thies	PROPN
biotropia-309	310	27	c.	c.	PROPN
biotropia-309	310	28	2005	2005	NUM
biotropia-309	310	29	.	.	PUNCT
biotropia-309	311	1	landscape	landscape	NOUN
biotropia-309	311	2	perspectives	perspective	NOUN
biotropia-309	311	3	on	on	ADP
biotropia-309	311	4	agricultural	agricultural	ADJ
biotropia-309	311	5	intensification	intensification	NOUN
biotropia-309	311	6	and	and	CCONJ
biotropia-309	311	7	biodiversity	biodiversity	NOUN
biotropia-309	311	8	-	-	PUNCT
biotropia-309	311	9	ecosystem	ecosystem	NOUN
biotropia-309	311	10	service	service	NOUN
biotropia-309	311	11	management	management	NOUN
biotropia-309	311	12	.	.	PUNCT
biotropia-309	312	1	ecology	ecology	NOUN
biotropia-309	312	2	letter	letter	NOUN
biotropia-309	312	3	8	8	NUM
biotropia-309	312	4	:	:	PUNCT
biotropia-309	312	5	857	857	NUM
biotropia-309	312	6	-	-	SYM
biotropia-309	312	7	874	874	NUM
biotropia-309	312	8	.	.	PUNCT
biotropia-309	313	1	young	young	PROPN
biotropia-309	313	2	rh	rh	PROPN
biotropia-309	313	3	.	.	PROPN
biotropia-309	313	4	2009	2009	NUM
biotropia-309	313	5	.	.	PUNCT
biotropia-309	314	1	land	land	NOUN
biotropia-309	314	2	use	use	NOUN
biotropia-309	314	3	and	and	CCONJ
biotropia-309	314	4	biodiversity	biodiversity	NOUN
biotropia-309	314	5	relationships	relationship	NOUN
biotropia-309	314	6	.	.	PUNCT
biotropia-309	315	1	journal	journal	NOUN
biotropia-309	315	2	of	of	ADP
biotropia-309	315	3	land	land	NOUN
biotropia-309	315	4	use	use	NOUN
biotropia-309	315	5	policy	policy	NOUN
biotropia-309	315	6	26s	26	NOUN
biotropia-309	315	7	:	:	PUNCT
biotropia-309	315	8	s178	s178	PROPN
biotropia-309	315	9	-	-	PUNCT
biotropia-309	315	10	s186	s186	PROPN
biotropia-309	315	11	.	.	PUNCT
biotropia-309	316	1	zar	zar	PROPN
biotropia-309	316	2	jh	jh	PROPN
biotropia-309	316	3	.	.	PROPN
biotropia-309	316	4	1999	1999	NUM
biotropia-309	316	5	.	.	PUNCT
biotropia-309	317	1	biostatistical	biostatistical	ADJ
biotropia-309	317	2	analysis	analysis	NOUN
biotropia-309	317	3	.	.	PUNCT
biotropia-309	318	1	4th	4th	ADJ
biotropia-309	318	2	edition	edition	PROPN
biotropia-309	318	3	.	.	PUNCT
biotropia-309	319	1	prentice	prentice	PROPN
biotropia-309	319	2	-	-	PUNCT
biotropia-309	319	3	hall	hall	PROPN
biotropia-309	319	4	inc	inc	PROPN
biotropia-309	319	5	.	.	PUNCT
biotropia-309	320	1	new	new	ADJ
biotropia-309	320	2	jersey.usa	jersey.usa	X
biotropia-309	320	3	et	et	NOUN
biotropia-309	320	4	al	al	PROPN
biotropia-309	320	5	biodiversitas	biodiversita	NOUN
biotropia-309	320	6	biological	biological	ADJ
biotropia-309	320	7	conservation	conservation	NOUN
biotropia-309	320	8	.	.	PUNCT
biotropia-309	321	1	58	58	NUM
biotropia-309	321	2	biotropia	biotropia	ADJ
biotropia-309	321	3	vol	vol	NOUN
biotropia-309	321	4	.	.	PROPN
biotropia-309	321	5	21	21	NUM
biotropia-309	321	6	no	no	NOUN
biotropia-309	321	7	.	.	NOUN
biotropia-309	321	8	1	1	NUM
biotropia-309	321	9	,	,	PUNCT
biotropia-309	321	10	2014	2014	NUM