4-1 - - - + --- I T 1 I1 - I. - i - * 1 -- C#il . '. 1- r-2.4 r - ,-11- 4 -I . - M 1 . , I r„ 2., L . + 1 4-4 of the FLORIDA MUSEUM OF NATURAL HISTORY LAND SNAILS OF THE GENUS COELOCENTRUM FROM NORTHEASTERN MEXICO Fred G. Thompson and Alfonso Correa Biological Sciences Volume 36, Number 5, pf 141-173 1994 UNIVERSrrY OF FLORIDA GAINESVILLE Numbers of the BULLETIN OF THE FLORIDA MUSEUM OF NATURAL HISTORY. BIOLOGICAL SCIENCES, are published at irregular intervals. Volumes contain about 300 pages and are not necessarily completed in any one calendar year. F . WAYNE KING , EDITOR RHODA J . BRYANT, MANAGING EDITOR Communications concerning purchase or exchange of the publications and all manuscripts should be addressed to: Managing Editor, Bulletin; Florida Museum of Natural History; University of Florida; Gainesville FL 32611-2035; U.S.A. This public document was promulgated at an annual cost of $1455.40 OR $1.212 per copy. It makes available to libraries, scholars, and all interested persons the results of researches in the natural sciences, emphasizing the circum-Caribbean region. ISSN: 0071-6154 CODEN: BF SBAS Publication date: February 17,1994 Price: $ 1.30 LAND SNAILS OF THE GENUS COELOCENTRUM FROM NORTHEASTERN MEXICO Fred G. Thompsoni and Alfonso Correa2 ABSTRACT Eleven species of Coelocentrum (GASTROPODA, PULMONATA Urocoptidae) are recorded from northeasternchodonta are known only from a relatively small area in northwestern Hidalgo and adjacent Quer6taro, a geologically isolated region of Cretaceous limestones, which is parted in the middle by the Rio Moctezuma. Coe/ocentrum astrophorea and C endo/ophus occur south of the Rio Moctezuma; C telescopium and C brachyacron occur north of the river. Considering that all other known species of Coelocentrum have very restricted distributions it is not plausible that C endolophus could be so widely distributed as to occur in Guerrero and Hidalgo. The type locality given in Pilsbry (1953: 134) is considered by us to be incorrect. Conversely, there is no apparent reason to doubt the reliability of the record from Km. 251, Hidalgo. THOMPSON & CORREO: COELOCEN7RUM FROM NORTHEASTERN MEXICO 153 Subgenus Crossostephanus Dall, 1908 TYPE SPECIES: Coelocentrumpatmeri Dall and Bartsch, 1908. This subgenus is characterized by the sculpture of the embryonic whorls and by having a turgid evagination on the columella which bears various solid ornate knobs and scallops. The embryonic whorls are moderate-sized and bear coarse riblets below the suture. The riblets are discontinuous across the whorls, but are weakly expressed along the base (Fig. 49). The evagination and its sculpture become enlarged in the lower whorls. In the less specialized species the evagination is low and bears a few coarse nodes. In the most elaborately evolved species, such as C. pa/meri, the evagination is greatly extended as a pendulant fold, and the knobs consist of large forward-directed overlapping leafs or plates. Intermediate stages occur in other species. Within a given sample there may be considerable individual variation in the degree of sculpturing on the columella. Some specimens are nearly devoid of any tuberculation. This is correlated with incomplete definitive growth. These are individuals that have definitive development of the outer shell, but have not yet completed development internally. Crossostephanus is known only from southeastern Tamaulipas and immediately adjacent San Luis Potosi. The species inhabit a variety of vegetational zones on limestone strata, including submesic scrub, mesic oak and pine forests and mountain rain forests. Species appear to be numerous. The area within the known range of the subgenus is poorly explored for mollusks, and large areas of seemingly suitable habitat to the west and north remain completely unexplored. Coelocentrum hinkleyi Pilsbry, 1909 Coelocentrum hinkleyi Pilsbly, 1909. Nautilus, 22: 138-139. This species has not been illustrated previously (Figs. 20,21,33,42-43). The sculpture on the columella relates it to the following three species. As with other Crossostephanus some individuals are devoid of nodular sculpture on the columella; others may be quite rugose. Measurements of ten shells selected to show variation are given in Table 5. TYPE LOCALITY.- SAN LUiS POTOSi: highest mountain on south side of river at Mecos Falls. Holotype in the ANSP. DISTRIBUTION.- Recorded only from the type locality and San Dieguito, San Luis Potosi, M6xico. 154 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL 36(3) TABLE 5.- Coelocentrum hinkleyi Pilsbry, 1909. Shell measurements of ten paratypes (UF 50153). Abbreviations as in Table 1 (page 146). Specimen L W ApH ApW Wh RP R5 R10 paratype 38.9 8.2 5.7 5.6 12.5 58 59 57 paratype 37.1 8.3 5.1 5.1 12.5 52 58 47 paralype 37.1 8.0 5.2 5.5 11.8 54 57 47 paratype 36.6 8.1 5.0 5.4 12.0 53 55 45 paratype 36.4 7.9 5.3 5.4 13.7 63 62 56 paratype 35.0 8.0 4.8 5.2 11.3 50 49 50 paratype 34.7 8.0 5.6 5.6 12.2 52 54 43 paratype 34.6 7.7 4.9 5.1 11.9 55 47 44 paratype 34.1 7.2 4.5 5.0 13.3 65 64 54 paratype 33.2 7.6 4.6 5.0 12.4 59 60 52 Coelocentrum affu:is, new species DESCRIPTION (Figs. 16-19, 40).- Medium-sized about 38-44 mm in length and about 9.240.4 mm wide; 0.22-0.26 times as wide as long. Shell tapered or cylindric-tapered in shape with the upper half tapering and the lower half nearly cylindrical; the last whorl is only slightly narrower than the penultimate whorl. Periostracum dull brown in color; the interior of the aperture tinged brown, becoming nearly white around the peristome. The umbilicus and the apical whorl are imperforate. Adult shells have 12.2-14.2 moderately arched whorls that are narrowly shouldered below the suture. The last whorl has a short neck, the length of which is about 1/6-1/4 the diameter of the last whorl (Figs. 17-18). The neck is slightly compressed dorso-ventrally and has a rounded base lacking any trace of a basal crest. The suture is moderately impressed and is accentuated by a thin, wavy spiral thread bounding the shoulder of the whorl. The whorls are sculptured with thin low riblets that are about 1/6-1/10 the width of their interspaces; 52-69 riblets on penultimate whorl; 55-81 on fifth whorl; 47-59 on tenth. The interspaces have irregularly spaced and coarser short segments of spiral threads. The aperture is wider than high and is about 0.62-0.71 times the width of the shell. It is variable in shape but usually is ovate or triangular-ovate in shape with a slightly concave parietal wall. A distinct angle occurs at the posterior corner. The peristome is THOMPSON & CORREO: COELOCEN7RUM FROM NORTHEASTERN MEXICO 155 TABLE 6.- Coelocentrum Ollims, new species. Shell measurements ofthe holotype (UF 159634) and ten paratypes (UF 193543). Abbreviations as in Table 1 (page). Specimen L W ApH ApW Wh RP R5 RIO holotype 40.6 9.6 6.5 6.7 12.0 52 56 47 paratype 43.7 10.1 7.1 6.8 12.3 71 62 54 paratype 43.0 9.8 5.7 6.5 14.7 64 70 47 paratype 42.5 9.6 6.2 6.4 14.2 65 62 51 paratype 42.1 9.2 6.4 6.4 13.7 69 62 52 paratype 41.5 9.4 5.9 6.4 13.6 69 66 53 paratype 41.5 9.2 6.4 6.5 13.3 69 81 59 paratype 41.4 9.5 6.7 6.5 13.3 80 77 39 paratype 39.7 10.4 7.2 6.8 12.2 64 - 45 paratype 38.0 9.5 6.5 5.9 12.6 62 74 54 paratype 37.6 9.2 6.5 6.6 12.2 56 55 49 slightly thickened and rounded and is nearly uniformly and narrowly expanded around the aperture. The columella is relatively narrow, being about 0.18-0.28 times the width of the shell. It gradually increases in diameter down to the penultimate whorl and then becoming narrower in the last whorl (Fig. 40). The middle of the columella between septa has a slight spiral bulge that is sculptured in the last 3-4 whorls with about 8-10 sparse, oblique white bars or elongate nodes. Measurements for adult shells are given in Table 6. The juvenile shell remains unknown. TYPE LOCALITY.- Sierra Tamalave, 7 km west-southwest of Adolfo Lop6z Mateos, Tamaulipas; 350 m altitude. HOLOTYPE: UF 159634; collected 17 January 1990 by Fred G. Thompson and Steven P. Christman. PARATYPES: UF 193543 (16 complete shells plus 5 fragments)), ITCVZ 1006 (10 adult shells); same data as the holotype. The Sierra Tamalave is a low limestone range with a north-south axis. The type locality is in a steep ravine on the east slope of the sierra along the highway from El Lim6n to Tula. The ravine was forested with a dense xeric scrub thicket. Snail shells were found in the leaf litter and under boulders. DISTRIBUTION.- Known from the Siem Tamalave at the type locality, and at 12 km WSW of Adolfo Lop6z Mateos, 450 m alt. (UF 159664). 156 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 36(5) REMARKS.- This species is most similar in appearance to Coe/ocentrum hinkleyi Pilsbry. The latter species occurs at about 30 km to the southwest in the Sierra Grande. The Sierra Tamalave and the Sierra Grande are separated by a dry alluvial basin. Coelocentrum qmnis differs from C hinkleyi primarily by its larger size (see Table 5), and by having a less constricted last whorl. Other aspects of the shell including external sculpture and the columella are alike. ETYMOLOGY.- The species name 00?nis is from the Latin meaning related to, in allusion to its close affinity to the following species. Coelocentrum priosculpta, new spedes DESCRIPTION (Figs. 22-27, 52).- Adult shell small; about 21-29 mm long and about 0.22-0.24 times as wide as long. Shell tapered, slightly convex in outline. Color light brown becoming whitish behind the peristome; interior of aperture tan. Apical whorl and umbilicus imperforate. Adult shell with 10.8-13.3 moderately arched whorls. Base of last whorl with a nearly obsolete spiral ridge along outer edge. Aperture extended forward on a short rounded neck that lacks a basal ridge (Fig. 23); length of neck about a fourth the width of the last whorl; flattened above and strongly angular in front of suture. Aperture about 0.63-0.71 times the width of the shell; obovate in shape; angular at posterior corner and less so at parietal-columellar corner. Peristome slightly thickened and narrowly reflected; slightly wider along base and columellar margin; narrowest along parietal margin and upper outer lip. Suture deep; whorls with a narrow but abrupt nearly horizontal shoulder that is accentuated by a wavy spiral thread connecting upper ends of ribs. Whorls sculptured with low, rounded, recurved riblets that are 1B-1/5 the width of their interspaces; 47-62 riblets on penultimate whorl; 4844 on fifth whorl; 38-54 on tenth whorl. Bases of riblets expanded and Oattened, tending to be forked. Interspaces between riblets with numerous irregular rather course segments of spiral striations. Columella rather slender, about 0.16-0.22 times the width of the shell (Fig. 24, 27); uniformly tai)ered to the penultimate whorl; columella with a low swelling in center of each whorl. The swelling may be nearly smooth throughout the length of the columella (Fig. 24), or it may bear a few scattered small nodes in the last two or three whorls (Fig. 27). Measurements of adult shells are given in Table 7. The rate of growth of juvenile shell indicates that about 21 whorls are lost above the point of decollation. Embryonic whorls 5, button-like in appearance; first slightly higher but narrower; next three nearly cylindrical, 1.4 mm wide; fifth slightly constricted. Sculpture and form of embryonic and juvenile shell similar to C. penion. THOMPSON & CORREO: COELOCEN7RUM FROM NORTHEASTERN M~)Cleo 157 TABLE 7.- Coelocentrumpnosculpta, new species. Shellmeasurements oftheholotype (UF 159672) and ten paratypes (UF 193538). Abbreviations as in Table 1 (page 146). Specimen L W ApH ApW Wh RP R5 R10 holotype 27.9 6.3 4.4 4.5 13.0 56 54 51 paratype 28.5 6.3 4.2 4.1 13.3 50 59 47 paratype 28.1 6.6 4.3 4.6 13.6 47 52 45 paratype 28.1 6.6 4.0 4.2 12.6 57 55 46 paratype 27.6 6.7 4.5 4.7 10.8 55 56 paratype 27.5 6.0 - 4.1 4.1 13.0 62 50 paratype 26.7 6.0 4.1 4.1 13.6 57 48 paratype 26.2 6.0 4.1 4.0 12.0 57 64 paratype 25.9 6.3 4.0 4.0 12.5 53 50 paratype 23.8 5.8 3.9 4.0 11.6 49 51 paralype 21.8 5.3 3.5 3.6 12.2 54 60 44 TYPE LOCALITY.- TAMAULIPAS: Sierra Grande, 2 km west of Santa Maria de Guadalupe, 21 km west of Ocampo; 1100 m alt. HOLOTYPE: UF 159672; collected 18 January 1990 by Fred G. Thompson. PARATYPES: UF 193538 (15), ITCVZ 1007 (9); same data as the holotype. The type locality is in a submesic limestone ravine that was forested with slender tall oaks (Quercus sp.). No understory was present. Snail shells were common in the leaf-litter. Live specimens were found under large boulders. DISTRIBUTION.- This species is known from Tamaulipas and immediately adjacent San Luis Potosi at an altitude of 1000-1100 m in submesic habitats. OTHER LOCALITIES.- SAN LUiS POTOSi: hwy. from Naranjos to Cd. El Maiz (22°29'06" N, 99°25'00"W), 1000 m alt. UF 193537). REMARKS.- Coelocentrum priosculpta is distinguished from other species of Crossostephanus by its small size, its costulate sculpture with the lower ends of the riblets flattened and expanded to form short inverted Y's and its simple, nearly smooth columella with only a few small nodes in the lower whorls. It is most similar to C hinkleyi Pilsbry, 1969 in the latter two characters. C hinkleyi is a 158 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL 36(5) much larger and relatively more attenuate species, and has a relatively longer neck on the last whorl. ETYMOLOGY.- The species name is from the Latin, prior meaning early, and sculpta, a carving, in reference to the primordial stage of sculpture on the columella. Coe/ocentrum paucinoda, new species DESCRIPTION (Figs. 28-32).- Moderately large and stocky; about 33-40 mm long and 8.8-10.6 mm wide; about 0.25-0.28 times as wide as high. Spire nearly cylindrical; last whorl narrower; upper half tapered. Color light brown and nearly lusterless. Adult shell with 11.3-12.7 whorls. Apical whorl imperforate; 4.9-6.1 mm wide. Umbilicus very narrowly perforate or rimate. Base of last whorl rounded; with a weak ridge along outer margin. Aperture extended forward on a short rounded neck (Fig. 29). Aperture broadly ovate in shape; parietal wall flattened and forming a weak angle internally with the outer lip. Width of aperture 0.57-0.66 times the width of the shell. Peristome narrowly and uniformly reflected. Whorls moderately arched and shouldered below the suture; shoulder accentuated by a thin wavy spiral thread connecting the upper ends of the axial riblets. Sculpture consisting of narrow, recurved, rounded riblets that are about 1/5-1/7 the width of their interspaces; penultimate whorl with 55-75 riblets; fifth whorl, 65-84; tenth whorl, 45-61. Located between the riblets are irregular, fine, spiral thread-segments. In most specimens these are quite numerous, although they may be reduced or absent on the lower whorls in some individuals. Columella tapered, 0.28-0.33 times the width of the shell at penultimate whorl (Fig. 30). Columella with a low turgid spiral ridge which in the penultimate and antipenultimate whorls bears about 7-9 white nodes per whorl; nodes round or slightly elongate; length of nodes about equal to or less than the width of the spiral swelling. External sculpture and other adult shell features are similar to C. torosum. The juvenile shell is unknown. Measurements and other meristic data are given in Table 8. TYPE LOCALITY.- TAMAULIPAS: limestone escarpment 3 km east of and ascending to the village of Gdmez Farias (23°01'21" N, 99°08'00"W); 300 m alt. HOLOTYPE: UF 193544; collected 31 August 1991 by Alfonso Correa PARATYPES: UF 193545 (3), UF 193547 (20), ITCVZ 1008 (10); same data as the holotype. DISTRIBUTION.- This species is known for certain only from the type locality. We have before us a dead shell (UF 193546) collected from along the Rio THOMPSON & CORREO: COELOC817RUM FROM NORTHEASTERN Mbaco 159 Frio, a few lung NE of the type locality (22°07'48"N, 99°04'24"W). Its point of origin is uncertain. REMARKS.- This species differs from the following species by its larger size, its cylindric-tapered shape, its narrowly perforate umbilicus, and its wider columella with sparser, more nearly rounded nodose sculpture. Although the two tan are quite similar, they differ sufficiently to warrant specific recognition. This is supported by their geographic and ecological isolation. The ranges of the two are separated by the Rio Guayalejo Basin, and by a distance of about 60 km. Both species occupy upland limestone habitats, whereas the lowlands of the Guayalejo Basin consists of alluvial and metamorphic deposits that are uninhabitable to Coelocentrum. ETYMOLOGY.- The name paucinoda is derived from the Latin paucus, few, and nodus, a knot or swelling, in reference to the columellar sculpture. TABLE 8.- Coetocentrum paucinoda, new species. Shell measurements ofthe holotype (UF 193544) and ten paratypes (UF 193545) Abbreviations as in Table l (page 146). Specimen L W ApH ApW Wh RP R3 R10 holotype 38.5 10.5 6.2 6.2 11.7 67 76 51 paratype 39.7 9.9 6.7 6.4 12.7 67 80 55 paratype 38.4 10.0 6.6 6.2 12.5 64 84 61 paratype 37.6 10.6 6.0 6.3 12.8 64 68 56 paratype 35.7 9.7 6.3 6.1 12.0 62 70 45 paratype 35.3 8.8 5.4 3.6 12.7 70 77 49 paratype 35.2 9.9 5.5 5.7 11.9 62 65 72 paratype 35.1 9.5 3.8 5.6 12.5 61 70 57 paratype 34.6 9.3 6.1 6.0 11.6 75 89 53 paratype 34.2 9.1 5.8 6.0 12.1 64 74 58 paratype 33.2 9.7 6.6 6.4 11.3 55 65 4 160 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL 36(5) Coelocentrum torosum, new species DESCRIPTION (Figs. 34-37, 41).- Adult shell medium sized, about 26.8- 32.7 mm long and 8.0-9.0 mm wide; about 0.26-0.31 times as wide as high. Club- shaped; upper lB - 1/2 of spire tapered; top whorl at apical plug 4.2-5.5 mm wide; lower portion of spire nearly cylindrical. Umbilicus and apical whorl imperforate. Last whorl with an obsolete angle along periphery and ending with a short neck about 1 mm long behind the aperture (Fig. 35). Color brown with a whitish peristome and aperture; umbilical wall gray. About 8.8-11.6 whorls below apical plug; whorls weakly convex; with conspicuous shoulders that are margined and accentuated by a light-colored spiral thread. Suture moderately impressed. Whorls sculptured with fine, recurved riblets that are about 1/4 - 1/5 the width of their interspaces; 61-86 riblets on penultimate whorl, 62-86 on fifth and 43-65 on tenth whorl above aperture; riblets connected at their upper ends by a narrower, lighter- colored, wavy, spiral thread. Aperture rounded with a weak posterior angle; usually wider than high and relatively large; about 0.6-0.7 times the width of the shell. Peristome slightly thickened and moderately reflected. Columella slender; about a tenth the width of the whorl near the apex, becoming enlarged in the lower third of shell so that the sculptured columella is about 0.23-0.28 the width of the whorl; axis with a slight spiral swelling that becomes enlarged in the lower whorls; swelling with 6-8 large, elongate, white knobs per whorl in lower part of shell; knobs about twice as long as wide; knobs usually confined to penultimate whorl; occasionally on previous two whorls (Figs. 40). Measurements of adult shells are given in Table 9. Embryonic shell consisting of six whorls. First whorl rounded but not conspicuously enlarged (in contrast to C penion). Next two whorls gradually increase in width to about 1.5-1.6 mm; following three whorls narrower. First embryonic whorl smooth; next five whorls smooth but with coarse denticles along the shoulder that crenulate the suture. Postembryonic whorls forming an attenuate, weakly concave spire; sculptured with fine regularly spaced riblets, which by about the twentieth whorl become connected along the upper suture by a thin spiral thread. Rate of growth ofjuvenile shell indicates that about 19-21 whorls are lost above apical plug. TYPE LOCALITY.- Sierra de Tamaulipas, 7 km northwest of Piruli, Tamaulipas, M6xico; 350 m alt. (23°34'27"N, 98°33'50"W). HOLOTYPE: UF 189695; collected 8 February 1991 by Alfonso Correa and Fred G. Thompson. PARATYPES: UF 193525 (75 adult shells and numerous fragments), ITCVZ 1009 (25); same data as the holotype. The type locality lies in a deep ravine near the top of a limestone ridge. This locality is on the highway from Cd. Victoria to Soto La Marina, and is about 54 km west of Soto La Marina. Snails were found under dead leaves, logs and stones near the bottom of the ravine. THOMPSON & CORREO: COELOCEN7RUM FROM NORTHEASTERN MbaCO 161 TABLE 9.- Coelocentrum torosum, new species. Shell measurements ofthe holotype (UF 189695) and ten paratypes (UF 193525) Abbreviations as in Table 1 (page 146). Specimen L W ApH ApW Wh RP R5 R10 holotype 32.1 9.0 6.2 6.3 10.2 68 71 55 paratype 32.7 8.9 5.5 5.9 11.3 61 66 52 paratype 32.7 8.4 5.2 5.6 12.3 73 86 64 paratype 32.3 8.7 6.1 6.1 11.6 76 76 54 paratype 30.2 8.0 5.1 5.2 11.2 64 64 49 paratype 28.7 8.1 5.3 5.5 10.2 86 75 65 paratype 28.6 8.0 4.9 5.3 11.6 67 62 43 paralype 28.0 8.1 4.9 5.6 10.1 79 76 43 pat·atype 27.5 8.0 5.1 5.3 10.3 66 63 48 paratype 27.4 8.6 5.2 5.1 10.4 74 75 50 paratype 26.8 8.3 5.5 5.5 8.8 77 77 - DISTRIBUTION.- This species appears to be widely distributed within the Sierra de Tamaulipas. It is known from 50-480 m altitude, and is common within subcaducifolias forests. OTHER SPECIMENS.- TAMAULIPAS. Sierra de Tamaulipas: Rancho El Carrizo, Soto La Maring 50 m alt. (UF 193526); El Moro, km 69, on road from Cd. Victoria to Soto La Marina; 480 m alt., 23°34'10"N, 98°32'56"W (UF 193528, UF193529); El Sabinito, km 97, on road from Cd. Victoria to Soto La Marina, 300 m alt. (UF 193527); 2 km W of El Sabinito, 27 km SW of Soto La Marina, 260 m alt. (UF 189715). REMARKS - Coelocentrum torosum is distinguished from other Crossostephanus by its relatively small club-shaped shell, its relatively slender columella bearing only a few elongate knobs or scallops in the lower 2-3 whorls, and its imperforate umbilicus and apical whort. As with other Coelocentrum some populations differ significantly in shell features. Specimens from 2 km west of El Sabinito attain a length of 30.4-35.8 mm, and the columella has scalloped nodes within the lower four whorls. Other population samples we have examined are very similar to the type series. 162 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL 36(5) ETYMOLOGY.- The species name torosum is from the Latin torosus, meaning bulging, and refers to the knobby protuberances on the axis. Coetocentrum palmeri Dall and Bartsch, 1908 Coelocentrum pahneri Dall and Bartsch, 1908. in Dall, 1908. Proceedings of the United States National Museum, 35: 177-178; pl. 29, figs. 2,5.- Solem, 1954; Nautilus, 68: 7. The original description was based on only a few specimens. Additional material collected during recent years adds significantly to our knowledge of the species and provides a basis for its redescription. DESCRIPT[ON (Figs. 38,39,44,45,49).- This species is characterized by its large stocky size and the sculpture on the columella. Color brown. Club- shaped and stocky, being about 0.25-0.32 times as wide as high. Shell about 42-63 mm long; normally about 12.2-14.7 mm wide; apical whorl about 5.0-6.9 mm wide. Aperture with a short neck the length of which is about a fifth the diameter of the last whorl (Fig. 39). Base rounded and without a basal keel. Umbilicus imperforate or narrowly rimate; apical whorl narrowly rimate. Aperture ovate- triangular in shape; about 0.53-0.66 times the width of the shell. Peristome slightly thickened and narrowly expanded. Whorls 11.3-13.5; separated by a moderately impressed suture that is accentuated by a conspicuous wavy, spiral thread bounding a narrow shoulder on the whorls. First whorl 4.9-6.9 mm wide; subsequent whorls gradually increasing in size to the lower quarter of the shell, and then the remaining 2-3 whorls becoming narrower. Whorls sculptured with low, narrow, protracted, concave riblets that are about 1/5 the width of their interspaces; 62-85 riblets on penultimate whorl, 58-81 on fifth whorl and 39-52 on tenth whorl above the aperture. Upper ends of riblets connected at the shoulder by a wavy, spiral thread. Occasional specimens have sparse spiral striations between the riblets (most conspicuous in weathered shells and hardly distinguishable in fresh specimens). Axis (Figs. 44,45) hollow, slightly less than half the width of the shell; with a wide evaginated ridge that begins at about 2-3 whorls below apical plug; ridge increasing in size and overhanging to become deeply concave along its lower wall in the last 34 whorls; ridge sculptures initially with indistinct ripples which in lower whorls gradually form into nodes and forward-directed overlapping leaves. Adult shell measurements are given in Table 10. The rate of growth ofjuvenile shells indicates that about 21-23 whorls are lost above the axial plug of the adult shell. Embryonic shell consisting of about 5.0 whorls. First whorl smooth, large and bulbous; next whorl slightly wider but lower, 1.6 mm wide; subsequent three whorls slightly shorter between THOMPSON & CORREO: COELOCEmRUM FROM NORTHEASTERN Mtaco 163 TABLE 10.- Coetocentrum palmen Dall and Bartsch. 1908. Shell measurements of fourteen specimens from the Sima de Cucharas, west of 66mez Farlas, Tamaulipas at 430 m alt (UF 1935311 and of the LECTOTYPE (USNM 198083) and a LECTOPARATYPE (FMNH 62392). Abbrevialions as in Table 1 (page 146). Specimen L W ApH ApW Wh RP RS R10 UF 193531 63.3 12.5 8.0 7.6 17.0 62 66 50 51.3 13.9 7.6 8.2 13.0 67 64 39 50.0 12.5 7.0 8.2 13.0 68 80 49 49.8 12.8 7.8 7.6 11.3 59 79 46 47.2 12.8 7.6 7.8 12.8 69 81 45 46.7 12.4 7.2 7.0 12.7 85 58 39 46.6 13.8 7.7 8.5 11.8 - - 43 46.5 12.5 7.0 7.3 13.0 81 71 43 46.5 12.3 7.0 7.1 12.8 63 64 51 44.6 13.0 7.0 7.4 12.8 72 64 50 42.8 12.2 6.7 7.3 11.5 65 66 39 42.0 13.5 8.7 8.4 11.0 74 89 52 - 17.0 9.5 9.0 - - - Lcctotype 51.1 14.7 9.0 8.2 13.0 82 70 50 Paratype 47.4 13.4 7.1 7.8 11.8 77 68 45 sutures and narrower in diameter, following whorls gradually increase in size to form a slightly concave spire. Second whorl with fine radial striations along upper suture. Sculpture on third and fourth whorl consisting of low but coarse grow- wrinkles along the upper suture. Subsequent juvenile whorls with fine, regularly spaced riblets that grade into the adult sculpture. TYPE LOCALITY.- Dall and Bartsch (in Dall, 1908) give Tamaulipas, M6xico as the type locality. The original label accompanying the lectotype states that it was collected at (}6mez Farias, Tamaulipas by Edward Palmer; 14-15 April 1907. The type series consisted of five specimens, but no holotype was designated. LECTOTYPE by present designation: USNM 198083. This is the specimen figured by Dall and Bartsch (1908: pl. 29, fig. 2). Two LECTOPARATYPES are in the Field Museum of Natural History (FMNH 62392). Other type specimens cited by Dall and Bartsch could not be located. 164 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL 36(5) DISTRIBUTION.- Known only from the Sierra de Cucharas, west of Gamez Farias, Tamaulipas, from 350-730 m altitude. This is a small mountain range that has an area of approximately 900 km sq. and lies due west of Gdmez Farias and north of Ocampo. The range rises to a rolling plateau at about 1800 m with numerous isolated hills and peaks as high as 2200 m. The range is inhabited by several species of Coelocentrum representing two subgeneg Coelocentrum s. s. and Crossostephanus. SPECIMENS EXAMINED.- TAMAULIPAS. Sierra de Cucharas: (36mez Farias, 350 m alt. (UF 189680, UF 193534); east slope of Sierra de Cucharas W of Gamez Farias, 470 m alt. (23°01'43"W, 99°09'24"W) (UF 193531); 7 km by road W of Gdmez Farias, 730 m alt. (23°03'20"N, 99°10'21"W) (UF 190972); road from Gdmez Farias to Rancho El Cielo (Alta Cimas), 430 m alt. (UF 193535); Gamez Farias (USNM 198083, LECTOTYPE; FMNH 62392, LECTOPARATYPES). REMARKS.- This is the largest known species of Crossostephanus, and it columella is the most ornately sculptured of the subgenus. The measurements given in Table 10 encompass most of the variation we have seen in other samples. LITERATURE CITED Bartsch, P. 1906. The urocoptid mollusks from the mainland of America in the collection of the United States National Museum. Proc. U. S. Natl. Mus. 31: 109-190; pls. 3-5. Dall, Wm H. 1897. New land mails from Mexico and New Mexico. Nautilus, 11: 61-62. Dall„ W. H. 1908. Descriptions and figures of sorne land and freshwater shells from Mexico, believed to be new. Proc. U. S. Natl. Mus. 35: 177-182; pls. 29-30. Frost, D. R. and D. M. Hillis. 1990. Species in concept and practice: herpetological application- Herpetologica, 46: 87-104. Madens. E. von 1890-1901. Biologia Centrali Americana: 1-706; pls. 1-44. Pilsbry, H. A 1903. Manual of Conchology, Ser. II, 15: 1-323; pls. 1-64. Pilsbry, H. A 1909. New mollusks collected by Mr. A A Hinkley in San Luis Potosi, M6xico. Nautilus, 22: 138-140. Pilsbry, H. A 1953. Inland MOLLUSCA of northern M6xico. II. Urocoptidae, Pupillidae, Strobilopsidae, Valloniidae and Cionellidae. Proc. Acad. NaL Sci. Philadelphia, 105: 133-167; pls. 3-10. Solem, A 1954. Notes on Mexican mollusks. I Durango, Coahuila and Tamaulipas with description of two newHumboldtiana. Nautilus, 68: 3-10. Thompson, F. G. 1968. Some Mexican land snails of the family Urocoptidae. Bull. Florida State Mus., Biol. Sci. 12(3): 125-183. THOMPSON & CORREO: COELOCE?V7RUM FROM NORTHEASTERN MbaCO 165 r \,· v 4 14·,0 493%..147 ->'\ Nort. VI i i l 0 4 i '0 ..41 + 0 C. (Coelocentrum) penion.- 2 1, 0 C. (C) tanyde#a 4 C. ptychodonta~ astrophoma 0 C. (P ) endolophus V C. (P ) brachyacron 9 C. (P.) telescopium -2 * C. (Crossostephanu~ palmeri '- a C. (Cr.) pnosculptaa f A C. lcO hinkleyi o C. (Cr.) amnis0 - 9 C. (Cr.) torosum* I C. (Cr.) paucinoda 4 0 i/0 f GULF OF ..0,- # r I U / . 4'fJ 2---J to 3 40 1 a / r.+ IEXICO r·3 4 j k"omoto„ 0 50 100 150 i / SOURCE /-Nalional 0•ogrophic Figure 1.- The distributions of Coelocentrum s. s. Jychodonta and Crosiodephamis in Northeastern M6xico. 166 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL 36(5) 1 06 08 el 0 0 Figures 2-6.- Coelocentnim penion, new species. Figures 2-3.- Holotype, front and side (UF 190974). Figures 4-5.- Paratypes, front and side (UF 193540). Figure 6.- Paratype, opened shell showing axis (UF 193541). Figure 7.- Coelocentrum mnydeira Thompson, 1968. Holotype, side view (UF 19040). THOMPSON & CORREO: COELOCENYRUM FROM NORTHEASTERN MEXICO 167 0 0100 Ar - . f,0 60 0 + 60 Figures 8-11.- Coelocentn,m telescopium, new species. Figures 8-9.- Holotype, front and side views (UF 34493). Figures 10-11.- Paratypes, (UF 193539). Figures 12-15.- Coelocentrum brachyacron, new species. Figures 12-13 - Holotype, front and side views (UF 159627). Figures 14-15 - Paratypes (UF 193542). 168 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL 36(5) 0e Ve Figures 16-19.- Coelocentrum a~nis, new species. Figures 16-17.- Front and side views of holotype (UF 159634). Figures 18-19.- Front view ofparatypes (UF 193543). Figures 20-21.- Coelocentrum hinkleyi Pilsbry, 1909. Paratypes (UF 50153). THOMPSON & CORREO: COELOCENTRUM FROM NORTHEASTERN MEXICO 169 ,/. 00 A- Figures 22-27.- Coelocentrum priosculptum, new species. Figures 22-23.- Front and side views of holotype (UF 159672) Figures 24-27.-- Paratypes (UF 193538) 170 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 36(5) 1 e iFF*.1 v a O"0 00 4 Figures 28-32.- Coelocentrum paucinoda, new species. Figures 28-29.- Front and side view of hototype (UF 193544). Figures 30-32.- Paratypes (UF 193545). Figure 33.- Coelocentrum hinkleyi Pilsbry, paratype (UF 50153). THOMPSON &CORREO: COELOCENTRUMFROM NORTHEASTERN MEXICO 171 4 er Figures 34-37.- Coelocentrum torosum, new species. Figures 34-35.- Front and side views of holotype (UF 189695). Figures 36.37.- Paratypes (UF 193525). Figures 38-39.- Coelocentrumpalmen Dall and Bartsch, 1908; (UF 193535). 172 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL, 36(5) Figure 40.- Coelocentrum €#ims, new species. Opened shell showing columella (UF 193543) Figure 41.- Coelocentrum torosum, new species. Paratype (UF 193525). Figures 42-43.- Coetocentrum hinkleyi Pilsbry, 1909. Paratypes (UF 50153). Figures 44-45.- Coelocentrumpalmeri Dall and Bartsch, 1908. Opened shells showing axis (193331). Figure 46 (next page).- Coelocentrum telescopium, new species, X14. Figure 47(next page).- Coelocentrum tanydeira Thompson, 1968 (UF 190410, X14. THOMPSON & CORREO: COELOCENTRUM FROM NORTHEASTERN MEXICO 173 4 . X l e f/ & m' W ele Figure 48.- Coetocentrum turns (Pfeiffer, 1856) (UF 190726), X14. Figure 49.- Coelocentrumpalmeri Dall and Bartsch, 1906. (UF 190972), X14. Figure 50.- Coelocentrum telescopium, new species. Paratype, juvenile shell (UF 193539). Figure 51.- Coelocentrum brachyacron, new species. Paratype, juvenile shell (UF 193542). Figure 52.- Coelocentrumpriosculpta, new species. Paratype juvenile shell (UF 193538). Contributions to the BULLETIN OF THE FLORIDA MUSEUM OF NATURAL HISTORY, BIOLOGICAL SCIENCES, may be in any field of biology. 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