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Specimen records for North American Lepidoptera (Insecta) in the Oregon State 
Arthropod Collection. Lycaenidae Leach, 1815 and Riodinidae Grote, 1895

Jon H. Shepard
Paul C. Hammond
Christopher J. Marshall

Oregon State Arthropod Collection, Department of Integrative Biology, Oregon State University, 
Corvallis OR 97331

Cite this work, including the attached dataset, as: 

Shepard, J. S, P. C. Hammond, C. J. Marshall. 2019. Specimen records for North American Lepidoptera (Insecta) in the 
Oregon State Arthropod Collection. Lycaenidae Leach, 1815 and Riodinidae Grote, 1895.  Catalog: Oregon 
State Arthropod Collection 3(2). (beta version).  
http://dx.doi.org/10.5399/osu/cat_osac.3.2.4594

Introduction

These records were generated using funds from the LepNet project (Seltmann) - a national effort to 
create digital records for North American Lepidoptera. The dataset published herein contains the 
label data for all North American specimens of Lycaenidae and Riodinidae residing at the Oregon 
State Arthropod Collection as of March 2019. A beta version of these data records will be made 
available on the OSAC server (http://osac.oregonstate.edu/IPT) at the time of this publication.  The 
beta version will be replaced in the near future with an official release (version 1.0), which will be 
archived as a supplemental file to this paper. 

Methods

Basic digitization protocols and metadata standards can be found in (Shepard et al. 2018).  
Identifications were confirmed by Jon Shepard and Paul Hammond prior to digitization. 
Nomenclature follows that of (Pelham 2008).
 
Results

The holdings in these two families are extensive.  Combined, they make up 25,743 specimens (24,598 
Lycanidae and 1145 Riodinidae).  While the majority of specimens are from the Pacific Northwest, the 
collection possesses examples of species from throughout North America. 

Within the Lycaenidae, the collection holds 316 distinct species-level taxa (species and supspecies), 
classified into the following genera: Atlides Hübner, 1819, Brephidium Scudder, 1876, Callophrys 
Billberg, 1820, Calycopis Scudder, 1876, Celastrina Tutt, 1906, Chlorostrymon Clench, 1961, Cupido 
Schrank, 1801, Cyanophrys Clench, 1961, Cyclargus Nabokov, 1945, Echinargus Nabokov, 1945, 
Electrostrymon Clench, 1961, Erora Scudder, 1872, Euphilotes Mattoni, 1978, Glaucopsyche Scudder, 1872, 
Hemiargus Hübner, 1818, Lampides Hübner, 1819, Leptotes Scudder, 1857, Ministrymon Clench, 1961, 
Parrhasius Hübner, 1819, Philotes Scudder, 1876, Philotiella Mattoni, 1978, Plebejus Kluk, 1780, Rekoa 
Kaye, 1904, Strymon Hübner, 1818, Tmolus Hübner, 1819, Vaga Zimmerman, 1958, Zizula Chapman, 



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Taxon             #specimens
Atlides halesus corcorani Clench, 1942 115
Atlides halesus halesus (Cramer, 1777) 33
Brephidium exilis (Boisduval, 1852) 142
Brephidium pseudofea (Morrison, 1873) 22
Callophrys affinis undetermined sspp. 67
Callophrys affinis affinis(Edwards, 1862) 5
Callophrys affinis apama (Edwards, 1882) 30
Callophrys affinis homoperplexa Barnes & Benjamin, 1923 42
Callophrys affinis washingtonia Clench, 1944 23
Callophrys augustinus annetteae (Passos, 1943) 19
Callophrys augustinus augustinus (Westwood, 1852) 23
Callophrys augustinus croesioides (Scudder, 1876) 30
Callophrys augustinus iroides (Boisduval, 1852) 380
Callophrys dumetorum undetermined sspp 46
Callophrys dumetorum dumetorum 411
Callophrys dumetorum oregonensis Górelick, 1968 171
Callophrys eryphon eryphon (Boisduval, 1852) 343
Callophrys eryphon fusca (Austin, 1998) 5
Callophrys eryphon pallescens (Austin, 1998) 6
Callophrys eryphon sheltonensis (Chermock & Frechin, 1949) 64
Callophrys fotis undetermined sspp. 57
Callophrys gryneus castalis (Edwards, 1871) 36
Callophrys gryneus chalcosiva Clench, 1981 353
Callophrys gryneus gryneus (Hübner, 1819) 70
Callophrys gryneus juniperaria (Comstock, 1925) 24
Callophrys gryneus loki (Skinner, 1907) 46
Callophrys gryneus mansfieldi (Tilden, 1951) 9
Callophrys gryneus nelsoni (Boisduval, 1869) 193
Callophrys gryneus plicataria Johnson, 1976 77
Callophrys gryneus rosneri Johnson, 1976 3
Callophrys gryneus siva (Edwards, 1874) 79
Callophrys henrici henrici (Grote & Robinson, 1867) 35

1910 (Table 1).  

The Riodinidae are less extensive, with onlh 20 species-level taxa classified into the following genera: 
Apodemia C. Felder & R. Felder, 1865, Calephelis Grote & Robinson, 1869, Caria Hübner, 1823, Emesis 
Fabricius, 1807, Lasaia H. Bates, 1868, (Table 2).

Table 1.  



Catalog: Oregon State Arthropod Collection   2019 Vol 3(2)

Callophrys henrici margaretae (dos Passos, 1943) 4
Callophrys henrici solatus (Cook & Watson, 1909) 12
Callophrys henrici turneri (Clench, 1943) 4
Callophrys hesseli (Rawson & Ziegler, 1950) 7
Callophrys irus arsace (Boisduval & LeConte, 1835) 11
Callophrys irus hadros (Cook & Watson, 1907) 41
Callophrys johnsoni (Skinner, 1904) 88
Callophrys lanoraieensis (Sheppard, 1934) 35
Callophrys mcfarlandi Ehrlich & Clench, 1960 15
Callophrys mossii bayensis Brown, 1969 44
Callophrys mossii hidakupa (Emmel, Emmel & Mattoon, 1998) 1
Callophrys mossii mossii (Edwards, 1881) 118
Callophrys mossii schryveri (Cross, 1937) 200
Callophrys mossii windi (Clench, 1943) 77
Callophrys muiri (Edwards, 1881) 54
Callophrys niphon (Hübner, 1819) 13
Callophrys niphon clarki (Freeman, 1938) 58
Callophrys polios maritima (Emmel, Emmel & Mattoon, 1998) 36
Callophrys polios obscurus Ferris & Fisher, 1973 194
Callophrys polios polios (Cook & Watson, 1907) 148
Callophrys sheridanii interrupta Austin, 1998 48
Callophrys sheridanii lemberti Tilden, 1963 157
Callophrys sheridanii newcomeri Clench, 1963 245
Callophrys sheridanii paradoxa Scott, 1986 2
Callophrys sheridanii pseudodumetorum Emmel, Emmel & Mattoon, 1998 190
Callophrys sheridanii sheridanii (Edwards, 1877) 71
Callophrys spinetorum spinetorum (Hewitson, 1867) 231
Callophrys xami xami (Reakirt, 1867) 11
Calycopis cecrops (Fabricius, 1793) 61
Calycopis isobeon (Butler & Druce, 1872) 58
Celastrina echo cinerea (Edwards, 1883) 44
Celastrina echo echo (Edwards, 1864) 33
Celastrina echo nigrescens (Fletcher, 1903) 585
Celastrina echo sidara (Clench, 1944) 13
Celastrina idella Wright & Pavulaan, 1999 3
Celastrina ladon (Cramer, 1780) 25

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Celastrina lucia (Kirby, 1837) 65
Celastrina neglecta (Edwards, 1862) 75
Celastrina neglectamajor Opler & Krizek, 1984 1
Celastrina nigra (Forbes, 1960) 7
Chlorostrymon simaethis (Drury, 1773) 31
Cupido amyntula albrighti (Clench, 1944) 11
Cupido amyntula amyntula (Boisduval, 1852) 270
Cupido amyntula herri (Grinnell, 1901) 16
Cupido amyntula montanorum (Austin, 1998) 27
Cupido amyntula valeriae (Clench, 1944) 10
Cupido comyntas comyntas (Godart, 1824) 63
Cupido comyntas sissona (Wright, 1905) 82
Cupido comyntas texana (Chermock, 1945) 24
Cyanophrys goodsoni (Clench, 1946) 32
Cyanophrys miserabilis (Clench, 1946) 11
Cyclargus thomasi bethunebakeri (Comstock & Huntington, 1943) 36
Echinargus isola (Reakirt, 1866) 112
Electrostrymon angelia angelia (Hewitson, 1874) 15
Erora laeta (Edwards, 1862) 18
Erora quaderna sanfordi Passos, 1940 22
Eumaeus atala (Poey, 1832) 19
Euphilotes battoides undetermined sspp (Behr, 1867) 67
Euphilotes battoides battoides (Behr, 1867) 33
Euphilotes battoides fusimaculata Austin, 1998 3
Euphilotes battoides mazourka Pratt & Emmel, 1998 9
Euphilotes battoides vernalis Pratt & Emmel, 1998 8
Euphilotes bernardino (Barnes & McDunnough, 1916) 99
Euphilotes centralis centralis (Barnes & McDunnough, 1917) 11
Euphilotes centralis hadrochilus Pratt & Emmel, 1998 13
Euphilotes enoptes undetermined sspp. 366
Euphilotes enoptes ancilla  (Barnes & McDunnough, 1918)               (see discussion)
Euphilotes enoptes bayensis (Langston, 1964) 97
Euphilotes enoptes columbiae (Mattoni, 1954)                                      (see discussion)
Euphilotes enoptes dammersi (Comstock & Henne, 1933) 2
Euphilotes enoptes enoptes (Boisduval, 1852) 254
Euphilotes enoptes langstoni (Shields, 1975) 16

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3(2)

Euphilotes enoptes tildeni (Langston, 1964) 23
Euphilotes glaucon undetermined sspp. 63
Euphilotes glaucon comstocki (Shields, 1975) 3
Euphilotes glaucon glaucon (Edwards, 1871) 159
Euphilotes glaucon oregonensis (Barnes & McDunnough, 1917) 321
Euphilotes mojave mojave (Watson & Comstock, 1920) 11
Euphilotes pallescens arenamontana Austin, 1998 7
Euphilotes pallescens calneva Emmel & Emmel, 1998 4
Euphilotes pallescens confusa Pratt & Emmel, 1998 5
Euphilotes pallescens mattonii (Shields, 1975) 2
Euphilotes pallescens ricei Austin, 1998 65
Euphilotes rita undetermined sspp. 3
Euphilotes rita coloradensis (Mattoni, 1966) 15
Euphilotes spaldingi (Barnes & McDunnough, 1917) 14
Feniseca tarquinius tarquinius (Fabricius, 1793) 42
Glaucopsyche lygdamus arizonensis McDunnough, 1936 14
Glaucopsyche lygdamus australis Grinnell, 1917 59
Glaucopsyche lygdamus columbia (Skinner, 1917) 561
Glaucopsyche lygdamus couperi Grote, 1873 53
Glaucopsyche lygdamus deserticola Austin & Emmel, 1998 13
Glaucopsyche lygdamus incognitus Tilden, 1974 55
Glaucopsyche lygdamus jacki Stallings & Turner, 1947 52
Glaucopsyche lygdamus lygdamus (Doubleday, 1842) 4
Glaucopsyche lygdamus mildredae Chermock, 1944 5
Glaucopsyche lygdamus orcus (Edwards, 1869) 31
Glaucopsyche lygdamus xerces (Boisduval, 1852) 15
Glaucopsyche piasus (Boisduval, 1852) 279
Habrodais grunus undetermined sspp. 174
Habrodais grunus grunus (Boisduval, 1852) 11
Habrodais grunus herri Field, 1938 131
Habrodais grunus lorquini Field, 1938 103
Hemiargus ceraunus undetermined sspp. 25
Hemiargus ceraunus antibubastus Hübner, 1818 16
Hemiargus ceraunus astenidas (Lucas, 1857) 27
Hemiargus ceraunus gyas (Edwards, 1871) 14
Hypaurotis crysalus undetermined sspp. 140

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Lampides boeticus (Linnaeus, 1767) 48
Leptotes cassius cassidula (Boisduval, 1870) 10
Leptotes cassius theonus (Lucas, 1857) 49
Leptotes marina (Reakirt, 1868) 114
Lycaena arota arota 199
Lycaena arota nubila (Comstock, 1926) 43
Lycaena arota schellbachi (Tilden, 1955) 25
Lycaena arota virginiensis (Edwards, 1870) 13
Lycaena cupreus undetermined sspp 13
Lycaena cupreus cupreus (Edwards, 1870) 100
Lycaena cupreus lapidicola Emmel & Pratt, 1998 73
Lycaena cupreus snowi (Edwards, 1881) 22
Lycaena dione (Scudder, 1868) 51
Lycaena dorcas arcticus (Ferris, 1977) 1
Lycaena dorcas castro (Reakirt, 1866) 37
Lycaena dorcas claytoni Brower, 1940 17
Lycaena dorcas dorcas Kirby, 1837 37
Lycaena dorcas florus (Edwards, 1884) 24
Lycaena dospassosi McDunnough, 1940 17
Lycaena editha editha (Mead, 1878) 43
Lycaena editha obscuramaculata Austin, 1989 2
Lycaena editha pseudonexa Emmel & Pratt, 1998 65
Lycaena editha vurali Koçak, 1984 306
Lycaena epixanthe epixanthe (Boisduval & Le Conte, 1835) 46
Lycaena epixanthe michiganensis Rawson, 1948 13
Lycaena epixanthe phaedrus (Hall, 1924) 13
Lycaena gorgon dorothea Emmel & Pratt, 1998 75
Lycaena gorgon gorgon (Boisduval, 1852) 78
Lycaena gorgon jacquelineae Emmel & Pratt, 1998 65
Lycaena helloides (Boisduval, 1852) 450
Lycaena hermes (Edwards, 1870) 32
Lycaena heteronea undetermined sspp. 43
Lycaena heteronea clara Edwards, 1877 2
Lycaena heteronea gravenotata Klots, 1930 28
Lycaena heteronea heteronea Boisduval, 1852 4
Lycaena heteronea klotsi Field, 1936 283

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3(2)

Lampides boeticus (Linnaeus, 1767) 48
Leptotes cassius cassidula (Boisduval, 1870) 10
Leptotes cassius theonus (Lucas, 1857) 49
Leptotes marina (Reakirt, 1868) 114
Lycaena arota arota 199
Lycaena arota nubila (Comstock, 1926) 43
Lycaena arota schellbachi (Tilden, 1955) 25
Lycaena arota virginiensis (Edwards, 1870) 13
Lycaena cupreus undetermined sspp 13
Lycaena cupreus cupreus (Edwards, 1870) 100
Lycaena cupreus lapidicola Emmel & Pratt, 1998 73
Lycaena cupreus snowi (Edwards, 1881) 22
Lycaena dione (Scudder, 1868) 51
Lycaena dorcas arcticus (Ferris, 1977) 1
Lycaena dorcas castro (Reakirt, 1866) 37
Lycaena dorcas claytoni Brower, 1940 17
Lycaena dorcas dorcas Kirby, 1837 37
Lycaena dorcas florus (Edwards, 1884) 24
Lycaena dospassosi McDunnough, 1940 17
Lycaena editha editha (Mead, 1878) 43
Lycaena editha obscuramaculata Austin, 1989 2
Lycaena editha pseudonexa Emmel & Pratt, 1998 65
Lycaena editha vurali Koçak, 1984 306
Lycaena epixanthe epixanthe (Boisduval & Le Conte, 1835) 46
Lycaena epixanthe michiganensis Rawson, 1948 13
Lycaena epixanthe phaedrus (Hall, 1924) 13
Lycaena gorgon dorothea Emmel & Pratt, 1998 75
Lycaena gorgon gorgon (Boisduval, 1852) 78
Lycaena gorgon jacquelineae Emmel & Pratt, 1998 65
Lycaena helloides (Boisduval, 1852) 450
Lycaena hermes (Edwards, 1870) 32
Lycaena heteronea undetermined sspp. 43
Lycaena heteronea clara Edwards, 1877 2
Lycaena heteronea gravenotata Klots, 1930 28
Lycaena heteronea heteronea Boisduval, 1852 4
Lycaena heteronea klotsi Field, 1936 283

Lycaena heteronea northi Emmel & Pratt, 1998 60
Lycaena heteronea rava Austin, 1998 61
Lycaena heteronea submaculata Emmel & Pratt, 1998 35
Lycaena hyllus (Cramer, 1775) 58
Lycaena mariposa charlottensis (Holland, 1930) 48
Lycaena mariposa mariposa (Reakirt, 1866) 278
Lycaena mariposa penroseae Field, 1938 104
Lycaena nivalis bichroma Emmel & Pratt, 1998 287
Lycaena nivalis browni Passos, 1938 151
Lycaena nivalis nivalis (Boisduval, 1869) 37
Lycaena nivalis praetexta Austin, 1998 30
Lycaena nivalis warnermontana Emmel & Pratt, 1998 31
Lycaena phlaeas alpestris Emmel & Pratt, 1998 3
Lycaena phlaeas arethusa (Walley-Dod, 1907) 4
Lycaena phlaeas feildeni (M’Lachlan, 1878) 2
Lycaena phlaeas hypophlaeas (Boisduval, 1852) 97
Lycaena rubidus duofascies Johnson & Balogh, 1977 112
Lycaena rubidus ferrisi Johnson & Balogh, 1977 2
Lycaena rubidus longi Johnson & Balogh, 1977 18
Lycaena rubidus perkinsorum Johnson & Balogh, 1977 51
Lycaena rubidus rubidus (Behr, 1866) 58
Lycaena rubidus sirius (Edwards, 1871) 135
Lycaena xanthoides nigromaculata Emmel & Pratt, 1998 59
Lycaena xanthoides xanthoides (Boisduval, 1852) 58
Ministrymon clytie (Edwards, 1877) 23
Ministrymon leda (Edwards, 1882) 51
Parrhasius m-album (Boisduval and Le Conte, 1833) 31
Philotes sonorensis (Felder & Felder, 1865) 135
Philotiella leona Hammond & McCorkle, 2000 146
Philotiella speciosa purisima Priestaf & Emmel, 1998 4
Philotiella speciosa septentrionalis Austin, 1998 15
Philotiella speciosa speciosa (Edwards, 1877) 66
Plebejus acmon (Westwood, 1852) 604
Plebejus anna undetermined sspp. 37
Plebejus anna anna (Edwards, 1861) 118
Plebejus anna azureus (Emmel, Emmel & Mattoon, 1998) 19

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Plebejus anna ricei Cross, 1937 511
Plebejus anna vancouverensis (Guppy & Shepard, 2001) 8
Plebejus emigdionis (Grinnell, 1905) 89
Plebejus glandon undetermined sspp. 31
Plebejus glandon megalo McDunnough, 1927 83
Plebejus glandon rustica (Edwards, 1865) 153
Plebejus icarioides undetermined sspp. 37
Plebejus icarioides albihalos (Emmel, Emmel & Mattoon, 1998) 9
Plebejus icarioides atascadero (Emmel, Emmel & Mattoon, 1998) 1
Plebejus icarioides blackmorei Barnes & McDunnough, 1919 42
Plebejus icarioides buchholzi Passos, 1938 21
Plebejus icarioides fenderi Macy, 1931 340
Plebejus icarioides fulla (Edwards, 1870) 206
Plebejus icarioides helios (Edwards, 1871) 175
Plebejus icarioides icarioides (Boisduval, 1852) 54
Plebejus icarioides inyo (Emmel, Emmel & Mattoon, 1998) 8
Plebejus icarioides lycea (Edwards, 1864) 41
Plebejus icarioides missionensis Hovanitz, 1937 26
Plebejus icarioides montis Blackmore, 1923 264
Plebejus icarioides pardalis (Behr, 1867) 72
Plebejus icarioides pembina (Edwards, 1862) 583
Plebejus icarioides pheres (Boisduval, 1852) 12
Plebejus idas alaskensis Chermock, 1945 55
Plebejus idas atrapraetextus Field, 1939 165
Plebejus idas empetri Freeman, 1938 6
Plebejus idas longinus (Nabokov, 1949) 18
Plebejus idas scudderii (Edwards, 1861) 25
Plebejus idas sublivens (Nabokov, 1949) 12
Plebejus lupini undetermined sspp. 37
Plebejus lupini alpicola (Emmel, Emmel & Mattoon, 1998) 5
Plebejus lupini chlorina (Skinner, 1902) 70
Plebejus lupini dedeckera (Emmel, Emmel & Mattoon, 1998) 4
Plebejus lupini goodpasturei (Austin, 1998) 5
Plebejus lupini lutzi Passos, 1938 355
Plebejus lupini spangelatus (Burdick, 1942) 27
Plebejus lupini texanus (Goodpasture, 1973) 37

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3(2)

Plebejus anna ricei Cross, 1937 511
Plebejus anna vancouverensis (Guppy & Shepard, 2001) 8
Plebejus emigdionis (Grinnell, 1905) 89
Plebejus glandon undetermined sspp. 31
Plebejus glandon megalo McDunnough, 1927 83
Plebejus glandon rustica (Edwards, 1865) 153
Plebejus icarioides undetermined sspp. 37
Plebejus icarioides albihalos (Emmel, Emmel & Mattoon, 1998) 9
Plebejus icarioides atascadero (Emmel, Emmel & Mattoon, 1998) 1
Plebejus icarioides blackmorei Barnes & McDunnough, 1919 42
Plebejus icarioides buchholzi Passos, 1938 21
Plebejus icarioides fenderi Macy, 1931 340
Plebejus icarioides fulla (Edwards, 1870) 206
Plebejus icarioides helios (Edwards, 1871) 175
Plebejus icarioides icarioides (Boisduval, 1852) 54
Plebejus icarioides inyo (Emmel, Emmel & Mattoon, 1998) 8
Plebejus icarioides lycea (Edwards, 1864) 41
Plebejus icarioides missionensis Hovanitz, 1937 26
Plebejus icarioides montis Blackmore, 1923 264
Plebejus icarioides pardalis (Behr, 1867) 72
Plebejus icarioides pembina (Edwards, 1862) 583
Plebejus icarioides pheres (Boisduval, 1852) 12
Plebejus idas alaskensis Chermock, 1945 55
Plebejus idas atrapraetextus Field, 1939 165
Plebejus idas empetri Freeman, 1938 6
Plebejus idas longinus (Nabokov, 1949) 18
Plebejus idas scudderii (Edwards, 1861) 25
Plebejus idas sublivens (Nabokov, 1949) 12
Plebejus lupini undetermined sspp. 37
Plebejus lupini alpicola (Emmel, Emmel & Mattoon, 1998) 5
Plebejus lupini chlorina (Skinner, 1902) 70
Plebejus lupini dedeckera (Emmel, Emmel & Mattoon, 1998) 4
Plebejus lupini goodpasturei (Austin, 1998) 5
Plebejus lupini lutzi Passos, 1938 355
Plebejus lupini spangelatus (Burdick, 1942) 27
Plebejus lupini texanus (Goodpasture, 1973) 37

Plebejus melissa undetermined sspp. 44
Plebejus melissa annetta (Edwards, 1882) 3
Plebejus melissa inyoensis (Nabokov, 1949) 14
Plebejus melissa melissa (Edwards, 1873) 408
Plebejus melissa samuelis (Nabokov, 1944) 2
Plebejus neurona (Skinner, 1902) 88
Plebejus optilete (Knoch, 1781) 4
Plebejus podarce cilla (Behr, 1867) 44
Plebejus podarce klamathensis (Emmel & Emmel, 1998) 116
Plebejus podarce podarce (Felder & Felder, 1865) 15
Plebejus saepiolus undetermined sspp 864
Plebejus saepiolus amica (Edwards, 1863) 56
Plebejus saepiolus hilda (Grinnell, 1907) 24
Plebejus shasta undetermined sspp. 21
Plebejus shasta minnehaha (Scudder, 1874) 4
Plebejus shasta pallidissima (Austin, 1998) 4
Plebejus shasta pitkinensis Ferris, 1976 14
Plebejus shasta shasta (Edwards, 1862) 192
Rekoa marius (Lucas, 1857) 1
Satyrium acadica (Edwards, 1862) 64
Satyrium acadica montanensis (Watson & Comstock, 1920) 3
Satyrium alcestis alcestis 22
Satyrium alcestis oslari 2
Satyrium auretorum (Boisduval, 1852) 79
Satyrium behrii undetermined sspp. 40
Satyrium behrii behrii (Edwards, 1870) 26
Satyrium behrii columbia (McDunnough, 1944) 175
Satyrium behrii crossi (Field, 1938) 52
Satyrium calanus undetermined sspp. 127
Satyrium californica undetermined sspp 410
Satyrium caryaevorus (McDunnough, 1942) 28
Satyrium edwardsii undetermined sspp. 65
Satyrium favonius autolycus (Edwards, 1871) 90
Satyrium favonius favonius (Smith, 1797) 40
Satyrium favonius ontario (Edwards, 1868) 16
Satyrium fuliginosa (Edwards, 1861) 18

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Satyrium fuliginosa tildeni Mattoon & Austin, 1998 16
Satyrium ilavia (Beutenmüller, 1899) 1
Satyrium kingi (Klots & Clench, 1952) 15
Satyrium liparops (LeConte, 1833) 44
Satyrium liparops floridensis Gatrelle, 2001 1
Satyrium polingi organensis Ferris, 1980 2
Satyrium polingi polingi (Barnes & Benjamin, 1926) 17
Satyrium saepium (Boisduval, 1852) 389
Satyrium semiluna semiluna Klots, 1930 230
Satyrium sylvinus undetermined sspp. 257
Satyrium sylvinus desertorum (Grinnell, 1917) 6
Satyrium sylvinus dryope (Edwards, 1870) 35
Satyrium sylvinus itys (Edwards, 1882) 2
Satyrium sylvinus megapallidum Austin, 1998 89
Satyrium tetra (Edwards, 1870) 113
Satyrium titus campus (Gatrelle, 2004) 5
Satyrium titus immaculosus (Comstock, 1913) 88
Satyrium titus occidentalis (Austin & Emmel, 1998) 1
Satyrium titus titus (Fabricius, 1793) 15
Satyrium titus watsoni (Barnes & Benjamin, 1926) 4
Satyrium titus winteri (Gatrelle, 2004) 49
Strymon acis (Drury, 1773) 9
Strymon alea (Godman & Salvin, 1887) 9
Strymon avalona (Wright, 1905) 4
Strymon bazochii bazochii (Godart, 1824) 38
Strymon istapa (Reakirt, 1867) 55
Strymon martialis (Herrich-Schäffer, 1864) 15
Strymon melinus Hübner, 1818 470
Strymon melinus melinus Hübner, 1818 2
Strymon melinus setonia McDunnough, 1927 2
Strymon rufofusca (Hewitson, 1877) 6
Tmolus echion (Linnaeus, 1767) 6
Vaga blackburni (Tuely, 1878) 28
Zizula cyna (Edwards, 1881) 3

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3(2)

Table 2. 
                   

                   Taxon          
#specimens
Apodemia duryi (Edwards, 1882) 1
Apodemia mejicanus mejicanus (Behr, 1865) 47
Apodemia mejicanus pueblo Scott, 1998 6
Apodemia mormo autumnalis Austin, 1998 80
Apodemia mormo cythera Edwards, 1873 25
Apodemia mormo undetermined sspp. 1
Apodemia mormo langei Comstock, 1939 65
Apodemia mormo mormo Felder, 1859 184
Apodemia mormo parva Austin, 1998 4
Apodemia nais Edwards, 1871 54
Apodemia palmerii arizona Austin, 1989 81
Apodemia palmerii palmerii Edwards, 1870 9
Apodemia virgulti davenporti Emmel, Emmel & Pratt, 1998 13
Apodemia virgulti virgulti (Behr, 1865) 130
Calephelis arizonensis McAlpine, 1971 40
Calephelis borealis (Grote & Robinson, 1866) 31
Calephelis freemani McAlpine, 1971 28
Calephelis muticum McAlpine, 1937 13
Calephelis nemesis Edwards, 1871 65
Calephelis perditalis perditalis Barnes & McDunnough, 1918 89
Calephelis rawsoni McAlpine, 1939 2
Calephelis virginiensis Gray, 1832 19
Calephelis wrighti Holland, 1930 1
Caria ino melicerta Schaus, 1890 44
Emesis ares (Edwards, 1882) 27
Emesis emesia Hewitson, 1867 1
Emesis zela cleis (Edwards, 1882) 62
Lasaia sula peninsularis Clench, 1971 23

Additional material from outside of North America is present, but was not digitized and 
catalogued at this time.



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Label data for these specimens can be accessed via online data portals that serve osac IPT data (e.g., 
www.gbif.org) or by direct download via the archived datasets stored in the supplemental files of this 
paper.

Discussion: 

Today the Oregon State Arthropod Collection maintains a relatively comprehensive representation 
of North American Lycaenidae, built up over many decades through the contributions of at least 20 
private collectors.  

A total of 156 species of Lycaenidae have been recorded from North America and Hawaii north of 
Mexico as reported in Pelham (2008) catalogue.  Of these, 149 (96%) are represented in the collection 
at the time of this writing, although 16 of these are represented by specimens from outside of this 
geographic range (e.g. further to the south) and were not included in this dataset.

The majority of specimens is made up of taxa that occur in the Pacific Northwest and California, with 
generally good geographic coverage within these regions.  In numerous cases (Table 1) subspecies 
were not always formally recognized in the database given uncertainties as to their validity and/
or diagnosability.   For example, while Euphilotes columbiae and Euphilotes ancilla are treated as valid 
species in Pelham (2008) there is some disagreement among lepidopterists as to their proper rank 
(e.g,, the are often treated as subspecies of Euphilotes enoptes) and not all lepidopterists consider them 
valid.  The OSAC does possess determined material corresponding to all of these nominative taxa, 
but at the time of this publication we opted to treat them as Euphilotes enoptes undetermined sspp. 
until we have more time to carefully review the taxonomy in greater detail.  

Minor corrections and other small modifications made in the future to this dataset will be 
accomplished by issuing a new version and providing a file of modified records to the supplemental 
files.  Major additions, such as new records, new data fields, new contributors or other significant 
modifications will be accomplished through the generation and release of new datasets.

Acknowledgments

The records published herein were generated with funds from NSF: DBI#1601888: Digitization TCN: 
Collaborative Research: Lepidoptera of North America Network: Documenting Diversity in the 
Largest Clade of Herbivores. 

References

Pelham JP (2008) A catalogue of the butterflies of the United States and Canada. Journal of Research 
on the Lepidoptera 40: 1-658

Seltmann, K. C. et al. (2017) LepNet: The Lepidoptera of North America Network. Zootaxa 4247 (1): 
073-077. https://doi.org/10.11646/zootaxa.4247.1.10.

Shepard J.H., C.J. Marshall, P.C. Hammond. 2018. North American Lepidoptera in the Oregon State 
Arthropod Collection: Hesperiidae, Pieridae, Papilionidae. Catalog: Oregon State 
Arthropod Collection 2(2): 1-2 and supplement file.  
doi: http://dx.doi.org/10.5399/osu/cat_osac.2.2.4480


