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Specimen records for North American Lepidoptera (Insecta) in the Oregon State 
Arthropod Collection. Pyraloidea: Crambidae Latreille 1810 and Pyralidae Linnaeus, 
1758

Jon H. Shepard
Paul C. Hammond
Christopher J. Marshall

Oregon State Arthropod Collection, Department of Integrative Biology, Oregon State University, 
Corvallis OR 97331

Cite this work, including the attached dataset, as: 

Shepard, J. S, P. C. Hammond, C. J. Marshall. 2019. Specimen records for North American Lepidoptera (Insecta) in the 
Oregon State Arthropod Collection. Pyraloidea: Crambidae Latreille 1810 and Pyralidae Linnaeus, 1758.  
Catalog: Oregon State Arthropod Collection 3(2)  (beta version).  
http://dx.doi.org/10.5399/osu/cat_osac.3.2.4596

Introduction

These records were generated using funds from the LepNet project (Seltmann et al. 2017) - a national 
effort to create digital records for North American Lepidoptera. The dataset published herein contains 
the label data for all North American specimens of Crambidae in the collection as of March 2019.  This 
dataset is not complete, as portions of our North American Pyralidae remain to be digitized.  This will 
be accomplished with a future dataset.  A beta version of these data records will be made available on 
the OSAC server (http://osac.oregonstate.edu/IPT) at the time of this publication.  The beta version 
will be replaced in the near future with an official release (version 1.0), which will be archived as a 
supplemental file to this paper. 

Methods

Basic digitization protocols and metadata standards can be found in (Shepard et al. 2018).  
Identifications for this family are tricky, especially in the Pacific Northwest where the pyraloid fauna 
remains relatively unstudied.   As a result, providing determinations to species, or even genus, was 
not always possible.  In the interest of making these specimens visible to taxonomists, we include 
even those specimens identified only to family. Generic and species determinations were reviewed 
prior to digitization by Jon Shepard and Paul Hammond using (Group 2019).  Nomenclature follows 
that of the Digital Guide to Moth Identification (Group 2019).  
 

Results

Crambidae: There are 3461 specimens identified at species-level (species or subspecies), with another 
123 specimens identifiable only to genus.  The following genera are present: Achroia Hübner,  1819, 
Achyra Guenée,  1849, Agathodes Guenée,  1854, Agriphila Hübner,  1825, Anageshna Munroe,  1956, 
Anania Hübner, 1923, Anatralata Munroe,  1961, Blepharomastix Lederer,  1863, Catoptria Hübner,  
1825, Chalcoela Zeller,  1872, Choristostigma Warren,  1892, Chrysoteuchia Hübner,  1825, Clydonopteron 
Riley,  1880, Cosipara Munroe,  1972, Crambus Fabricius,  1798, Crocidophora Lederer,  1863, Desmia 



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Westwood,  1832, Diaphania Hübner,  1818, Diastictis Hübner,  1818, Dicymolomia Zeller,  1872, 
Donacaula Meyrick,  1890, Euchromius Guenée,  1845, Eudonia Billberg,  1820, Eurrhypara Hübner,  1825, 
Evergestis Hübner,  1825, Fissicrambus Bleszynski,  1963, Frechinia Munroe,  1961, Fumibotys Munroe,  
1976, Galleria Fabricius,  1798, Gesneria Hübner, 1925, Gyros Edwards,  1881, Hahncappsia Munroe,  
1976, Herpetogramma Lederer,  1863, Loxostege Hübner,  1825, Macrotheca Ragonot,  1891, Mecyna 
Doubleday,  1849, Microcrambus Bleszynski,  1963, Microtheoris Meyrick,  1932, Mimoschinia Warren,  
1892, Neodactria Landry,  1995, Noctueliopsis Munroe,  1961, Nomophila Hübner,  1825, Occidentalia Dyar 
& Heinrich,  1927, Orenaia Duponchel,  1845, Ostrinia Hübner,  1825, Palpita Hübner,  1808, Parapeiacia 
Bleszynski, 1966, Parapoynx Hübner,  1825, Pediasia Hübner,  1825, Perispasta Zeller,  1875, Petrophila 
Guilding, 1830, Phlyctaenia Kützing,  1849, Pogonogenys Munroe,  1961, Procymbopteryx Munroe,  1961, 
Prorasea Grote,  1878, Pseudoschoenobius Fernald,  1891, Pyrausta Schrank,  1802, Saucrobotys Munroe,  
1976, Scoparia Haworth 1811, Sitochroa Hübner,  1825, Spoladea Guenée,  1854, Tehama Hulst,  1888, 
Thaumatopsis Morrison,  1874, Tosale Walker,  1863, Udea Guenée,  1845, Uresiphita Hübner,  1825, Urola 
Walker,  1863 (Table 1).

Pyralidae:  Of the pyralid specimens digitized at this time, only 24, belonging to the subfamily 
Phycitinae, could not be identified to genus; an additional 157 specimens could only be identified at 
the generic level. To date, 1383 specimens of North American Pyralidae were identified to species-
level (species or subspecies) and digitized, belonging to the following genera: Acrobasis Zeller, 
1839, Acrobasis Zeller, 1839, Acroncosa Barnes & McDunnough, 1917, Aglossa Latreille, 1796, Alberada 
Heinrich, 1939, Ambesa Grote, 1880, Amyelois Amsel, 1956, Ancylosis Zeller, 1839, Bandera Ragonot, 
1887, Dasypyga Ragonot, 1887, Dioryctria Zeller, 1846, Ephestia Guenée, 1845, Ephestiodes Ragonot, 
1887, Etiella Zeller, 1839, Euzophera Zeller, 1867, Homoeosoma Curtis, 1833, Honora Grote, 1878, 
Hypsopygia Hübner, 1825, Interjectio Heinrich, 1956, Laetilia Ragonot, 1889, Lipographis Ragonot, 1887, 
Macalla Walker, 1859, Melitara Walker, 1863, Myelopsis Heinrich, 1956, Olybria Heinrich, 1956, Oneida 
Hulst, 1889, Phobus Heinrich, 1956, Phycitodes Hampson, 1917, Pima Hulst, 1888, Plodia Guenée, 1845, 
Pococera Zeller, 1848, Promylea Ragonot, 1887, Pyla Grote, 1882, Pyralis Linnaeus, 1758, Rostrolaetilia 
Blanchard & Ferguson, 1975, Sarata Ragonot, 1887, Sciota Hulst, 1888, Toripalpus Grote, 1878, Vitula 
Ragonot, 1887, Zophodia Hübner, 1825 (Table 2).

Table 1. Crambidae

Taxon     #specimens

Achroia grisella (Fabricius, 1794) 17
Achyra occidentalis Packard, 1873 5
Achyra rantalis Guenée, 1854 7
Agathodes designalis Guenée, 1854 5
Agriphila anceps Grote, 1880 1
Agriphila attenuatus Grote, 1880 6
Agriphila biothanatalis Hulst, 1886 132
Agriphila plumbifimbriellus Dyar, 1904 79
Agriphila undatus Grote 2
Anageshna primordialis Dyar, 1906 1
Anania funebris (Strom, 1768) 7
Anania mysippusalis (Walker, 1859) 8



Catalog: Oregon State Arthropod Collection   2019 Vol 3(2)

Anatralata versicolor Warren, 1892 1
Blepharomastix ranalis (Guenée, 1854) 2
Catoptria latiradiellus Walker, 1863 2
Catoptria oregonicus Grote 17
Chalcoela iphitalis Walker, 1859 15
Choristostigma elegantalis Warren, 1892 4
Chrysoteuchia topiarius Zeller, 1866 173
Clydonopteron undetermined 1
Cosipara tricoloralis Dyar, 1904 27
Crambus agitatellus Clemens, 1861 2
Crambus dimidiatus Haworth, 1809 1
Crambus gausapalis Hulst, 1886 3
Crambus girardellus Clemens, 1860 1
Crambus leachellus Zincken, 1818 210
Crambus pascuella floridus Zeller, 1873 3
Crambus perlella innotatellus Walker, 1863 5
Crambus sperryellus Klots, 1940 20
Crambus tutillus McDunnough, 1921 129
Crambus unistriatellus Packard, 1868 44
Crambus undetermined 24
Crocidophora serratissimalis Zeller, 1872 1
Desmia funeralis Hübner 2
Desmia maculalis Westwood, 1831 13
Diaphania hyalinata (Linnaeus, 1767) 1
Diaphania nitidalis (Stoll, 1781) 2
Diastictis fracturalis (Zeller, 1872) 7
Diastictis robustior Munroe, 1956 5
Diastictis sperryorum Munroe, 1956 1
Dicymolomia metalliferalis Packard, 1873 51
Donacaula melinellus Clemens, 1861 1
Donacaula undetermined 1
Euchromius ocellea (Haworth, 1811) 148
Eudonia commortalis Dyar, 1921 3
Eudonia echo Dyar, 1928 1
Eudonia leucophthalma Dyar, 1928 2
Eudonia rectilineata Li, Li & Nuss, 2012 1

Table 1 (continued) 
                   
                   Taxon          #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Eudonia spenceri Munroe, 1972 18
Eudonia torniplagalis Dyar, 1904 1
Eurrhypara hortulata Linnaeus, 1758 5
Evergestis eurekalis Barnes & McDunnough, 1914 7
Evergestis funalis (Grote, 1878) 14
Evergestis pallidata (Hufnagel, 1767) 22
Evergestis rimosalis Guenée, 1854 1
Evergestis simulatilis Grote, 1880 7
Evergestis subterminalis Barnes & McDunnough, 1914 10
Fissicrambus mutabilis (Clemens, 1860) 2
Frechinia helianthiales (Murt., 1897) 1
Frechinia laetalis Barnes & McDunnough, 1914 1
Fumibotys fumalis Guenée, 1854 13
Galleria mellonella (Linnaeus, 1758) 72
Gesneria centuriella (Denis & Schiffermuller, 1775) 4
Gyros muiri Edwards, 1881 19
Hahncappsia coloradensis Grote & Robinson, 1867 3
Herpetogramma abdominalis Zeller 4
Herpetogramma pertextalis Lederer, 1863 174
Herpetogramma thestealis Walker, 1859 1
Loxostege anartalis Grote, 1877 9
Loxostege cereralis Zeller, 1872 30
Loxostege ephippialis (Zetterstedt, 1839) 1
Loxostege indentalis (Grote, 1883) 39
Loxostege lepidalis Hulst, 1886 79
Loxostege sierralis Munroe, 1976 12
Loxostege sticticalis (Linnaeus, 1761) 15
Loxostege terpnalis Barnes & McDunnough, 1918 2
Macrotheca ponda (Dyar, 1907) 2
Mecyna mustelinalis Packard, 1873 66
Microcrambus elegans Clemens, 1860 2
Microtheoris ophionalis Walker, 1859 26
Mimoschinia rufofascialis (Stephens, 1834) 21
Neodactria luteolellus Clemens, 1860 43
Noctueliopsis virula Barnes & McDunnough, 1918 2
Nomophila nearctica Munroe, 1973 76

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3(2)

Occidentalia comptulatalis Hulst, 1886 4
Orenaia arcticalis Munroe, 1974 1
Ostrinia nubilalis (Hubner, 1796) 6
Ostrinia penitalis Grote, 1876 5
Palpita magniferalis Walker, 1861 9
Parapediasia teterrellus (Zincken, 1821) 71
Parapoynx badiusalis Walker, 1859 9
Parapoynx obscuralis Grote, 1881 2
Pediasia browerellus Klots, 1942 1
Pediasia dorsipunctellus Kearfott, 1908 133
Pediasia trisecta Walker, 1856 145
Perispasta caeculalis Zeller, 1875 12
Petrophila avernalis Grote, 1878 3
Petrophila confusalis Walker, 1865 65
Phlyctaenia coronata Hufnagel, 1767 29
Pogonogenys proximalis Fernald 2
Procymbopteryx belialis Druce, 1899 1
Prorasea praeia Dyar, 1917 2
Prorasea simalis Grote, 1878 9
Pseudoschoenobius opalescalis Hulst, 1886 10
Pyrausta acrionalis Walker, 1859 1
Pyrausta californicalis Packard, 1873 81
Pyrausta dapalis Grote, 1881 2
Pyrausta fodinalis Lederer, 1863 41
Pyrausta grotei Munroe, 1876 9
Pyrausta nexalis(Hulst 1886) 6
Pyrausta laticlavia Grote & Robinson, 1867 4
Pyrausta linealis Fernald, 1894 1
Pyrausta nicalis Grote, 1878 29
Pyrausta perrubralis Packard, 1873 38
Pyrausta pilatealis Barnes & McDunnough, 1914 1
Pyrausta pseuderosnealis Munroe, 1976 1
Pyrausta semirubralis Packard, 1873 20
Pyrausta signatalis Walker, 1865 2
Pyrausta subsequalis Guenée, 1854 48
Pyrausta unifascialis Packard, 1873 57

Table 1 (continued) 
                   
                   Taxon          #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Pyrausta volupialis Grote, 1877 21
Saucrobotys fumoferalis Hulst, 1886 21
Saucrobotys futilalis Lederer, 1863 5
Scoparia biplagialis Walker, 1865 11
Sitochroa chortalis Grote, 1874 11
Sitochroa palealis (Denis & Schiffermuller, 1775) 3
Spoladea recurvalis (Fabricius, 1775) 2
Tehama bonifatellus Hulst, 1887 236
Thaumatopsis fernaldella Kearfott, 1905 4
Thaumatopsis repandus Grote, 1880 2
Tosale oviplagalis Walker, 1866 4
Udea undetermined 1
Udea itysalis Walker, 1859 7
Udea itysalis tillialis 15
Udea profundalis Packard, 1873 257
Udea rubigalis (Guenee, 1854) 2
Udea vacunalis (Grote, 1881) 1
Udea washingtonalis (Grote, 1881) 20
Uresiphita reversalis Guenée, 1854 3
Urola nivalis (Drury, 1773) 1

Table 1 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3(2)

Taxon                                                                        #specimens
Acrobasis caliginella (Hulst, 1878) 19
Acrobasis tricolorella Grote, 1878 98
Acroncosa albiflavella Barnes & McDunnough, 1917 1
Aglossa cacamica Dyar, 1914 7
Aglossa pinguinalis (Linnaeus, 1758) 6
Alberada californiensis Neunzig, 1997 2
Ambesa laetella Grote, 1880 25
Ambesa walsinghami Ragonot, 1887 22
Amyelois transitella (Walker, 1863) 5
Ancylosis albipenella (Hulst, 1887) 1
Ancylosis morrisonella Ragonot, 1887 55
Ancylosis undulatella Clemens, 1860 15
Bandera virginella Dyar, 1908 1
Dasypyga alternosquamella Ragonot, 1887 29
Dioryctria undetermined 54
Dioryctria abietivorella Grote, 1878 116
Dioryctria albovittella Hulst, 1890 21
Dioryctria auranticella Grote, 1883 36
Dioryctria cambiicola Dyar, 1914 48
Dioryctria durangoensis Mutuura & Neunzig, 1986 1
Dioryctria erythropasa Dyar, 1914 1
Dioryctria pentictonella Mutuura, Munroe & Ross, 1969 74
Dioryctria ponderosae Dyar, 1914 3
Dioryctria pseudotsugella Munroe, 1959 219
Dioryctria rossi Munroe, 1959 4
Dioryctria tumicolella Mutuura, Munroe & Ross, 1969 4
Ephestia elutella (Hubner, 1796) 5
Ephestia kuehniella Zeller, 1879 24
Ephestiodes gilvescentella Ragonot, 1887 8
Ephestiodes  undetermined 1
Etiella zinckenella (Treitschke, 1832) 9
Euzophera semifuneralis Walker, 1863 20
Homoeosoma electella Hulst, 1887 10
Honora dotella Dyar, 1910 3
Honora mellinella Grote, 1878 7
Hypsopygia costalis (Fabricius, 1775) 28
Hypsopygia undetermined 1

Table 2 



Catalog: Oregon State Arthropod Collection   2019 Vol 3 (2)

Hypsopygia phoezalis (Dyar, 1908) 19
Interjectio undetermined 17
Interjectio columbiella McDunnough, 1935 2
Interjectio denticulella Ragonot, 1887 5
Interjectio niviella Hulst, 1888 2
Laetilia dilatifasciella (Ragonot, 1887) 1
Laetilia zamacrella Dyar, 1925 23
Lipographis fenestrella Packard, 1874 14
Macalla zelleri Grote, 1876 3
Melitara subumbiella Dyar 11
Myelopsis alatella Hulst, 1887 1
Myelopsis minutularia Hulst, 1887 1
Olybria aliculella Hulst, 1887 3
Oneida undetermined 3
Phobus brucei Hulst, 1895 5
Phobus curvatella Ragonot, 1887 6
Phycitodes albatella (Ragonot, 1887) 2
Phycitodes mucidella (Ragonot, 1887) 4
Phycitodes reliquellum (Dyar, 1904) 17
Pima albiplagiatella Packard, 1874 2
Pima fergusoni Neunzig, 2003 3
Pima fosterella Hulst, 1888 10
Pima occidentalis Heinrich, 1956 2
Plodia undetermined 22
Plodia interpunctella (Hubner, 1813) 45
Pococera aplastella Hulst, 1888 1
Promylea lunigerella Ragonot, 1887 2
Pyla aeneoviridella Ragonot, 1887 28
Pyla aenigmatica Heinrich, 1956 19
Pyla fusca Haworth, 1811 45
Pyla undetermined 24
Pyla scintillans Grote, 1881 4
Pyla westerlandi Wilterding & Balogh, 2002 5
Pyralis farinalis (Linnaeus, 1758) 58
Rostrolaetilia minimella Blanchard & Ferguson, 1975 2
Sarata caudellella Dyar, 1904 1

Table 2 (continued). 

       Taxon             #specimens



Catalog: Oregon State Arthropod Collection   2019 Vol 3(2)

Sarata edwardsialis Hulst, 1886 1
Sarata incanella Hulst, 1895 2
Sarata perfuscalis Hulst, 1886 4
Sarata pullatella Ragonot, 1887 12
Sciota undetermined 15
Sciota levigatella (Hulst, 1892) 1
Sciota tarmitalis (Hulst, 1886) 4
Toripalpus breviornatalis Grote, 1878 2
Toripalpus trabalis Grote 23
Vitula edmandsii (Packard, 1865) 25
Vitula pinei Heinrich, 1956 6
Vitula undetermined 29
Zophodia grossulariella (Hubner, 1809) 5
Pyralidae: Phycitinae undetermined 24

Table 2 (continued) 
                   
                   Taxon          #specimens

Additional material from outside of North America is present, but was not digitized and catalogued 
at this time.

Label data for these specimens can be accessed via online data portals that serve osac IPT data (e.g., 
www.gbif.org) or by direct download via the archived datasets stored in the supplemental files of this 
paper.

Discussion: 

Minor corrections and other small modifications made in the future to this dataset will be 
accomplished by issuing a new version and providing a file of modified records to the supplemental 
files.  Major additions, such as new records, new data fields, new contributors or other significant 
modifications will be accomplished through the generation and release of new datasets.

References

Moth Photographers Group (2019) Digital Guide to Moth Identification. https://
mothphotographersgroup.msstate.edu/  [accessed April 1, 2019.2019]

Seltmann, K. C. et al. 2017.  LepNet: The Lepidoptera of North America Network. Zootaxa 4247 (1): 
073-077. https://doi.org/10.11646/zootaxa.4247.1.10.

Shepard J.H., C.J. Marshall, P.C. Hammond. 2018. North American Lepidoptera in the Oregon State 
Arthropod Collection: Hesperiidae, Pieridae, Papilionidae. Catalog: Oregon State 
Arthropod Collection 2(2): 1-2 and supplement file.  
doi: http://dx.doi.org/10.5399/osu/cat_osac.2.2.4480


