id	sid	tid	token	lemma	pos
elements-9012	1	1	do	do	AUX
elements-9012	1	2	all	all	DET
elements-9012	1	3	mammals	mammal	NOUN
elements-9012	1	4	exhibit	exhibit	VERB
elements-9012	1	5	heterogeneity	heterogeneity	NOUN
elements-9012	1	6	in	in	ADP
elements-9012	1	7	their	their	PRON
elements-9012	1	8	mutation	mutation	NOUN
elements-9012	1	9	rates	rate	NOUN
elements-9012	1	10	?	?	PUNCT
elements-9012	2	1	do	do	AUX
elements-9012	2	2	all	all	DET
elements-9012	2	3	yeasts	yeast	NOUN
elements-9012	2	4	exhibit	exhibit	VERB
elements-9012	2	5	uniformity	uniformity	NOUN
elements-9012	2	6	?	?	PUNCT
elements-9012	2	7	"	"	PUNCT
elements-9012	3	1	v	v	ADP
elements-9012	3	2	a	a	DET
elements-9012	3	3	r	r	NOUN
elements-9012	3	4	y	y	NOUN
elements-9012	3	5	i	i	PRON
elements-9012	3	6	n	n	CCONJ
elements-9012	3	7	g	g	NOUN
elements-9012	3	8	patterns	pattern	NOUN
elements-9012	3	9	of	of	ADP
elements-9012	3	10	m	m	VERB
elements-9012	3	11	u	u	NOUN
elements-9012	3	12	t	t	PROPN
elements-9012	3	13	a	a	DET
elements-9012	3	14	t	t	NOUN
elements-9012	4	1	i	i	PRON
elements-9012	4	2	o	o	VERB
elements-9012	4	3	n	n	VERB
elements-9012	4	4	measuring	measure	VERB
elements-9012	4	5	the	the	DET
elements-9012	4	6	universality	universality	NOUN
elements-9012	4	7	of	of	ADP
elements-9012	4	8	regional	regional	ADJ
elements-9012	4	9	mutation	mutation	NOUN
elements-9012	4	10	rates	rate	NOUN
elements-9012	4	11	a	a	DET
elements-9012	4	12	l	l	NOUN
elements-9012	4	13	e	e	NOUN
elements-9012	4	14	a	a	DET
elements-9012	4	15	h	h	NOUN
elements-9012	5	1	f	f	NOUN
elements-9012	5	2	o	o	NOUN
elements-9012	5	3	x	x	X
elements-9012	5	4	m	m	VERB
elements-9012	5	5	u	u	NOUN
elements-9012	5	6	t	t	PROPN
elements-9012	6	1	a	a	DET
elements-9012	6	2	t	t	NOUN
elements-9012	7	1	i	i	PRON
elements-9012	7	2	o	o	VERB
elements-9012	8	1	n	n	X
elements-9012	8	2	s	s	VERB
elements-9012	8	3	a	a	DET
elements-9012	8	4	r	r	NOUN
elements-9012	8	5	e	e	NOUN
elements-9012	8	6	t	t	NOUN
elements-9012	8	7	h	h	NOUN
elements-9012	8	8	e	e	X
elements-9012	8	9	u	u	NOUN
elements-9012	8	10	l	l	NOUN
elements-9012	8	11	t	t	NOUN
elements-9012	9	1	i	i	PRON
elements-9012	9	2	m	m	VERB
elements-9012	9	3	a	a	DET
elements-9012	9	4	t	t	NOUN
elements-9012	9	5	e	e	NOUN
elements-9012	9	6	s	s	PROPN
elements-9012	10	1	o	o	X
elements-9012	10	2	u	u	NOUN
elements-9012	10	3	r	r	NOUN
elements-9012	10	4	c	c	NOUN
elements-9012	10	5	e	e	NOUN
elements-9012	10	6	o	o	NOUN
elements-9012	10	7	f	f	X
elements-9012	10	8	g	g	PROPN
elements-9012	10	9	e	e	PROPN
elements-9012	10	10	n	n	PROPN
elements-9012	10	11	e	e	NOUN
elements-9012	10	12	t	t	NOUN
elements-9012	11	1	i	i	PRON
elements-9012	11	2	c	c	VERB
elements-9012	11	3	v	v	ADP
elements-9012	11	4	a	a	DET
elements-9012	11	5	r	r	NOUN
elements-9012	11	6	i	i	PRON
elements-9012	12	1	a	a	DET
elements-9012	12	2	t	t	NOUN
elements-9012	13	1	i	i	PRON
elements-9012	13	2	o	o	VERB
elements-9012	13	3	n	n	VERB
elements-9012	13	4	i	i	PRON
elements-9012	13	5	n	n	VERB
elements-9012	13	6	d	d	PROPN
elements-9012	13	7	n	n	ADV
elements-9012	13	8	a	a	PRON
elements-9012	13	9	.	.	PUNCT
elements-9012	14	1	t	t	NOUN
elements-9012	14	2	h	h	NOUN
elements-9012	15	1	e	e	PROPN
elements-9012	15	2	p	p	X
elements-9012	15	3	a	a	DET
elements-9012	15	4	t	t	NOUN
elements-9012	15	5	t	t	NOUN
elements-9012	15	6	e	e	NOUN
elements-9012	15	7	r	r	NOUN
elements-9012	15	8	n	n	PROPN
elements-9012	15	9	s	s	PART
elements-9012	15	10	o	o	NOUN
elements-9012	15	11	f	f	X
elements-9012	15	12	m	m	VERB
elements-9012	15	13	u	u	NOUN
elements-9012	15	14	t	t	PROPN
elements-9012	15	15	a	a	DET
elements-9012	15	16	t	t	NOUN
elements-9012	16	1	i	i	PRON
elements-9012	16	2	o	o	VERB
elements-9012	16	3	n	n	INTJ
elements-9012	16	4	,	,	PUNCT
elements-9012	16	5	h	h	NOUN
elements-9012	17	1	o	o	NOUN
elements-9012	17	2	w	w	PROPN
elements-9012	17	3	e	e	X
elements-9012	17	4	v	v	X
elements-9012	17	5	e	e	NOUN
elements-9012	17	6	r	r	NOUN
elements-9012	17	7	,	,	PUNCT
elements-9012	17	8	c	c	PROPN
elements-9012	17	9	a	a	DET
elements-9012	17	10	n	n	NOUN
elements-9012	17	11	v	v	NOUN
elements-9012	18	1	a	a	DET
elements-9012	18	2	r	r	NOUN
elements-9012	18	3	y	y	PROPN
elements-9012	18	4	b	b	PROPN
elements-9012	18	5	o	o	NOUN
elements-9012	18	6	t	t	NOUN
elements-9012	19	1	h	h	NOUN
elements-9012	19	2	w	w	PROPN
elements-9012	20	1	i	i	PRON
elements-9012	20	2	t	t	NOUN
elements-9012	21	1	h	h	NOUN
elements-9012	21	2	i	i	PRON
elements-9012	21	3	n	n	PROPN
elements-9012	21	4	a	a	DET
elements-9012	21	5	n	n	NOUN
elements-9012	21	6	d	d	NOUN
elements-9012	21	7	a	a	DET
elements-9012	21	8	c	c	NOUN
elements-9012	21	9	r	r	NOUN
elements-9012	21	10	o	o	NOUN
elements-9012	21	11	s	s	X
elements-9012	21	12	s	s	X
elements-9012	21	13	g	g	NOUN
elements-9012	21	14	e	e	NOUN
elements-9012	21	15	n	n	NOUN
elements-9012	21	16	o	o	NOUN
elements-9012	21	17	m	m	NOUN
elements-9012	21	18	e	e	NOUN
elements-9012	21	19	s	s	PROPN
elements-9012	21	20	.	.	PUNCT
elements-9012	22	1	i	i	PRON
elements-9012	22	2	t	t	VERB
elements-9012	22	3	h	h	NOUN
elements-9012	22	4	a	a	DET
elements-9012	22	5	s	s	X
elements-9012	22	6	p	p	NOUN
elements-9012	22	7	r	r	NOUN
elements-9012	22	8	e	e	NOUN
elements-9012	22	9	v	v	NOUN
elements-9012	23	1	i	i	PRON
elements-9012	23	2	o	o	VERB
elements-9012	23	3	u	u	NOUN
elements-9012	23	4	s	s	PROPN
elements-9012	23	5	l	l	NOUN
elements-9012	23	6	y	y	PROPN
elements-9012	23	7	b	b	PROPN
elements-9012	23	8	e	e	PROPN
elements-9012	23	9	e	e	ADP
elements-9012	23	10	n	n	PROPN
elements-9012	23	11	s	s	NOUN
elements-9012	23	12	h	h	NOUN
elements-9012	24	1	o	o	NOUN
elements-9012	24	2	w	w	NOUN
elements-9012	24	3	n	n	PROPN
elements-9012	24	4	t	t	PROPN
elements-9012	24	5	h	h	NOUN
elements-9012	24	6	a	a	DET
elements-9012	24	7	t	t	NOUN
elements-9012	24	8	s	s	X
elements-9012	24	9	e	e	X
elements-9012	24	10	v	v	ADP
elements-9012	24	11	e	e	NOUN
elements-9012	24	12	r	r	NOUN
elements-9012	24	13	a	a	PROPN
elements-9012	24	14	l	l	NOUN
elements-9012	24	15	m	m	VERB
elements-9012	24	16	a	a	DET
elements-9012	24	17	m	m	NOUN
elements-9012	24	18	m	m	VERB
elements-9012	24	19	a	a	DET
elements-9012	24	20	l	l	NOUN
elements-9012	24	21	s	s	NOUN
elements-9012	24	22	h	h	NOUN
elements-9012	24	23	a	a	DET
elements-9012	24	24	v	v	NOUN
elements-9012	24	25	e	e	NOUN
elements-9012	24	26	h	h	NOUN
elements-9012	24	27	e	e	NOUN
elements-9012	24	28	t	t	NOUN
elements-9012	24	29	e	e	X
elements-9012	24	30	r	r	NOUN
elements-9012	24	31	o	o	X
elements-9012	24	32	g	g	NOUN
elements-9012	24	33	e	e	PROPN
elements-9012	24	34	n	n	NOUN
elements-9012	24	35	e	e	X
elements-9012	24	36	o	o	X
elements-9012	24	37	u	u	NOUN
elements-9012	24	38	s	s	NOUN
elements-9012	24	39	m	m	VERB
elements-9012	24	40	u	u	NOUN
elements-9012	24	41	t	t	NOUN
elements-9012	24	42	a	a	DET
elements-9012	24	43	t	t	NOUN
elements-9012	25	1	i	i	PRON
elements-9012	25	2	o	o	VERB
elements-9012	26	1	n	n	CCONJ
elements-9012	26	2	r	r	NOUN
elements-9012	26	3	a	a	DET
elements-9012	26	4	t	t	NOUN
elements-9012	26	5	e	e	X
elements-9012	26	6	s	s	X
elements-9012	26	7	,	,	PUNCT
elements-9012	26	8	w	w	NOUN
elements-9012	26	9	h	h	NOUN
elements-9012	27	1	i	i	NOUN
elements-9012	27	2	l	l	NOUN
elements-9012	28	1	e	e	NOUN
elements-9012	28	2	f	f	X
elements-9012	28	3	o	o	X
elements-9012	28	4	u	u	NOUN
elements-9012	28	5	r	r	NOUN
elements-9012	28	6	y	y	PROPN
elements-9012	28	7	e	e	PROPN
elements-9012	28	8	a	a	PRON
elements-9012	28	9	s	s	ADP
elements-9012	28	10	t	t	NOUN
elements-9012	28	11	s	s	NOUN
elements-9012	28	12	h	h	NOUN
elements-9012	28	13	a	a	PRON
elements-9012	28	14	v	v	NOUN
elements-9012	28	15	e	e	X
elements-9012	28	16	b	b	PROPN
elements-9012	28	17	e	e	X
elements-9012	28	18	e	e	X
elements-9012	28	19	n	n	NOUN
elements-9012	28	20	o	o	X
elements-9012	28	21	b	b	X
elements-9012	28	22	s	s	X
elements-9012	28	23	e	e	NOUN
elements-9012	28	24	r	r	NOUN
elements-9012	28	25	v	v	NOUN
elements-9012	28	26	e	e	X
elements-9012	28	27	d	d	NOUN
elements-9012	28	28	t	t	X
elements-9012	28	29	o	o	X
elements-9012	28	30	h	h	NOUN
elements-9012	29	1	a	a	DET
elements-9012	29	2	v	v	NOUN
elements-9012	29	3	e	e	X
elements-9012	29	4	u	u	NOUN
elements-9012	29	5	n	n	ADV
elements-9012	30	1	i	i	PRON
elements-9012	30	2	f	f	NOUN
elements-9012	31	1	o	o	NOUN
elements-9012	31	2	r	r	NOUN
elements-9012	31	3	m	m	VERB
elements-9012	31	4	r	r	NOUN
elements-9012	31	5	a	a	DET
elements-9012	31	6	t	t	NOUN
elements-9012	31	7	e	e	X
elements-9012	31	8	s	s	PROPN
elements-9012	31	9	.	.	PUNCT
elements-9012	32	1	t	t	PROPN
elements-9012	32	2	h	h	NOUN
elements-9012	32	3	e	e	PROPN
elements-9012	32	4	g	g	PROPN
elements-9012	32	5	e	e	PROPN
elements-9012	32	6	n	n	X
elements-9012	32	7	e	e	NOUN
elements-9012	32	8	r	r	NOUN
elements-9012	32	9	a	a	PROPN
elements-9012	32	10	l	l	NOUN
elements-9012	33	1	i	i	PRON
elements-9012	33	2	t	t	VERB
elements-9012	34	1	y	y	INTJ
elements-9012	35	1	o	o	X
elements-9012	35	2	f	f	NOUN
elements-9012	35	3	t	t	NOUN
elements-9012	35	4	h	h	NOUN
elements-9012	36	1	e	e	X
elements-9012	36	2	s	s	PROPN
elements-9012	36	3	e	e	X
elements-9012	36	4	o	o	X
elements-9012	36	5	b	b	X
elements-9012	36	6	s	s	X
elements-9012	36	7	e	e	NOUN
elements-9012	36	8	r	r	NOUN
elements-9012	36	9	v	v	PROPN
elements-9012	36	10	a	a	DET
elements-9012	36	11	t	t	NOUN
elements-9012	37	1	i	i	PRON
elements-9012	37	2	o	o	VERB
elements-9012	37	3	n	n	NOUN
elements-9012	37	4	s	s	NOUN
elements-9012	37	5	h	h	NOUN
elements-9012	37	6	a	a	DET
elements-9012	37	7	s	s	NOUN
elements-9012	37	8	n	n	PRON
elements-9012	37	9	o	o	NOUN
elements-9012	37	10	t	t	NOUN
elements-9012	37	11	b	b	PROPN
elements-9012	37	12	e	e	PROPN
elements-9012	37	13	e	e	PROPN
elements-9012	37	14	n	n	CCONJ
elements-9012	37	15	k	k	PROPN
elements-9012	37	16	n	n	CCONJ
elements-9012	37	17	o	o	NOUN
elements-9012	37	18	w	w	NOUN
elements-9012	37	19	n	n	PROPN
elements-9012	37	20	.	.	PUNCT
elements-9012	38	1	h	h	NOUN
elements-9012	39	1	e	e	NOUN
elements-9012	39	2	r	r	NOUN
elements-9012	39	3	e	e	PROPN
elements-9012	39	4	w	w	NOUN
elements-9012	39	5	e	e	NOUN
elements-9012	39	6	e	e	X
elements-9012	39	7	x	x	X
elements-9012	39	8	a	a	PRON
elements-9012	39	9	m	m	VERB
elements-9012	39	10	i	i	NOUN
elements-9012	39	11	n	n	NOUN
elements-9012	39	12	e	e	NOUN
elements-9012	39	13	s	s	VERB
elements-9012	39	14	i	i	NOUN
elements-9012	39	15	l	l	NOUN
elements-9012	39	16	e	e	NOUN
elements-9012	39	17	n	n	ADP
elements-9012	39	18	t	t	NOUN
elements-9012	39	19	s	s	X
elements-9012	40	1	i	i	INTJ
elements-9012	40	2	t	t	NOUN
elements-9012	40	3	e	e	NOUN
elements-9012	40	4	s	s	PROPN
elements-9012	40	5	u	u	PROPN
elements-9012	40	6	b	b	PROPN
elements-9012	40	7	s	s	ADP
elements-9012	40	8	t	t	X
elements-9012	41	1	i	i	PRON
elements-9012	41	2	t	t	VERB
elements-9012	41	3	u	u	NOUN
elements-9012	42	1	t	t	NOUN
elements-9012	43	1	i	i	PRON
elements-9012	43	2	o	o	VERB
elements-9012	44	1	n	n	VERB
elements-9012	44	2	s	s	X
elements-9012	44	3	i	i	PRON
elements-9012	44	4	n	n	NOUN
elements-9012	44	5	c	c	NOUN
elements-9012	44	6	o	o	X
elements-9012	45	1	d	d	NOUN
elements-9012	45	2	i	i	PRON
elements-9012	45	3	n	n	CCONJ
elements-9012	45	4	g	g	NOUN
elements-9012	45	5	r	r	NOUN
elements-9012	45	6	e	e	NOUN
elements-9012	45	7	g	g	NOUN
elements-9012	46	1	i	i	PRON
elements-9012	46	2	o	o	VERB
elements-9012	47	1	n	n	NOUN
elements-9012	47	2	s	s	X
elements-9012	47	3	o	o	X
elements-9012	47	4	f	f	NUM
elements-9012	47	5	20	20	NUM
elements-9012	47	6	m	m	NOUN
elements-9012	47	7	a	a	DET
elements-9012	47	8	m	m	NOUN
elements-9012	47	9	m	m	VERB
elements-9012	47	10	a	a	DET
elements-9012	47	11	l	l	NOUN
elements-9012	47	12	s	s	NOUN
elements-9012	47	13	,	,	PUNCT
elements-9012	47	14	27	27	NUM
elements-9012	47	15	y	y	NOUN
elements-9012	47	16	e	e	PROPN
elements-9012	47	17	a	a	PRON
elements-9012	47	18	s	s	PROPN
elements-9012	47	19	t	t	NOUN
elements-9012	47	20	,	,	PUNCT
elements-9012	47	21	a	a	DET
elements-9012	47	22	n	n	NOUN
elements-9012	47	23	d	d	ADP
elements-9012	47	24	4	4	NUM
elements-9012	47	25	i	i	NOUN
elements-9012	47	26	n	n	VERB
elements-9012	47	27	s	s	NOUN
elements-9012	47	28	e	e	X
elements-9012	47	29	c	c	PROPN
elements-9012	47	30	t	t	PROPN
elements-9012	47	31	s	s	PART
elements-9012	47	32	,	,	PUNCT
elements-9012	47	33	t	t	NOUN
elements-9012	47	34	o	o	X
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elements-9012	48	3	t	t	NOUN
elements-9012	48	4	e	e	PROPN
elements-9012	48	5	r	r	NOUN
elements-9012	48	6	m	m	VERB
elements-9012	48	7	i	i	PRON
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elements-9012	48	9	e	e	NOUN
elements-9012	48	10	w	w	NOUN
elements-9012	48	11	h	h	NOUN
elements-9012	49	1	i	i	NOUN
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elements-9012	49	3	h	h	NOUN
elements-9012	49	4	g	g	PROPN
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elements-9012	49	6	n	n	ADV
elements-9012	49	7	o	o	NOUN
elements-9012	49	8	m	m	NOUN
elements-9012	50	1	e	e	NOUN
elements-9012	50	2	s	s	PROPN
elements-9012	50	3	d	d	X
elements-9012	50	4	e	e	X
elements-9012	50	5	m	m	NOUN
elements-9012	50	6	o	o	NOUN
elements-9012	51	1	n	n	X
elements-9012	51	2	s	s	PROPN
elements-9012	51	3	t	t	NOUN
elements-9012	51	4	r	r	NOUN
elements-9012	51	5	a	a	DET
elements-9012	51	6	t	t	NOUN
elements-9012	51	7	e	e	NOUN
elements-9012	51	8	t	t	NOUN
elements-9012	52	1	h	h	NOUN
elements-9012	53	1	i	i	PRON
elements-9012	53	2	s	s	VERB
elements-9012	53	3	m	m	VERB
elements-9012	53	4	o	o	NOUN
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elements-9012	53	6	a	a	PRON
elements-9012	54	1	i	i	PRON
elements-9012	54	2	c	c	NOUN
elements-9012	54	3	r	r	NOUN
elements-9012	54	4	a	a	DET
elements-9012	54	5	t	t	NOUN
elements-9012	54	6	e	e	X
elements-9012	55	1	d	d	X
elements-9012	55	2	i	i	PRON
elements-9012	55	3	s	s	VERB
elements-9012	55	4	t	t	NOUN
elements-9012	56	1	r	r	NOUN
elements-9012	57	1	i	i	NOUN
elements-9012	58	1	b	b	PROPN
elements-9012	59	1	u	u	NOUN
elements-9012	60	1	t	t	X
elements-9012	61	1	i	i	PRON
elements-9012	61	2	o	o	VERB
elements-9012	61	3	n	n	CCONJ
elements-9012	61	4	a	a	DET
elements-9012	61	5	n	n	NOUN
elements-9012	61	6	d	d	NOUN
elements-9012	61	7	w	w	NOUN
elements-9012	62	1	h	h	NOUN
elements-9012	63	1	i	i	PRON
elements-9012	63	2	c	c	NOUN
elements-9012	63	3	h	h	NOUN
elements-9012	64	1	a	a	DET
elements-9012	64	2	r	r	NOUN
elements-9012	64	3	e	e	X
elements-9012	64	4	u	u	NOUN
elements-9012	64	5	n	n	ADV
elements-9012	65	1	i	i	PRON
elements-9012	65	2	f	f	NOUN
elements-9012	66	1	o	o	NOUN
elements-9012	66	2	r	r	NOUN
elements-9012	66	3	m	m	NOUN
elements-9012	66	4	.	.	PUNCT
elements-9012	67	1	o	o	X
elements-9012	68	1	u	u	NOUN
elements-9012	68	2	r	r	NOUN
elements-9012	68	3	f	f	NOUN
elements-9012	68	4	i	i	PRON
elements-9012	68	5	n	n	VERB
elements-9012	69	1	d	d	NOUN
elements-9012	69	2	i	i	PRON
elements-9012	69	3	n	n	CCONJ
elements-9012	69	4	g	g	PROPN
elements-9012	69	5	s	s	NOUN
elements-9012	69	6	s	s	NOUN
elements-9012	69	7	h	h	NOUN
elements-9012	69	8	o	o	PROPN
elements-9012	69	9	w	w	PROPN
elements-9012	69	10	t	t	PROPN
elements-9012	69	11	h	h	NOUN
elements-9012	70	1	a	a	DET
elements-9012	70	2	t	t	NOUN
elements-9012	70	3	m	m	VERB
elements-9012	70	4	u	u	NOUN
elements-9012	70	5	t	t	PROPN
elements-9012	70	6	a	a	DET
elements-9012	70	7	t	t	NOUN
elements-9012	71	1	i	i	PRON
elements-9012	71	2	o	o	INTJ
elements-9012	71	3	n	n	CCONJ
elements-9012	71	4	a	a	DET
elements-9012	71	5	l	l	NOUN
elements-9012	71	6	h	h	NOUN
elements-9012	71	7	e	e	NOUN
elements-9012	71	8	t	t	NOUN
elements-9012	71	9	e	e	X
elements-9012	71	10	r	r	NOUN
elements-9012	71	11	o	o	X
elements-9012	71	12	g	g	NOUN
elements-9012	71	13	e	e	PROPN
elements-9012	71	14	n	n	NOUN
elements-9012	71	15	e	e	NOUN
elements-9012	72	1	i	i	PRON
elements-9012	72	2	t	t	VERB
elements-9012	72	3	y	y	INTJ
elements-9012	72	4	o	o	NOUN
elements-9012	72	5	c	c	NOUN
elements-9012	72	6	c	c	NOUN
elements-9012	72	7	u	u	NOUN
elements-9012	72	8	r	r	NOUN
elements-9012	72	9	s	s	VERB
elements-9012	72	10	i	i	PRON
elements-9012	72	11	n	n	CCONJ
elements-9012	72	12	a	a	DET
elements-9012	72	13	l	l	NOUN
elements-9012	72	14	l	l	NOUN
elements-9012	73	1	b	b	NOUN
elements-9012	73	2	r	r	NOUN
elements-9012	73	3	a	a	DET
elements-9012	73	4	n	n	NOUN
elements-9012	73	5	c	c	NOUN
elements-9012	73	6	h	h	NOUN
elements-9012	73	7	e	e	NOUN
elements-9012	73	8	s	s	PROPN
elements-9012	73	9	o	o	X
elements-9012	73	10	f	f	X
elements-9012	73	11	t	t	NOUN
elements-9012	73	12	h	h	NOUN
elements-9012	74	1	e	e	NOUN
elements-9012	74	2	m	m	VERB
elements-9012	74	3	a	a	PRON
elements-9012	74	4	m	m	NOUN
elements-9012	74	5	m	m	VERB
elements-9012	74	6	a	a	DET
elements-9012	74	7	l	l	NOUN
elements-9012	74	8	i	i	PRON
elements-9012	74	9	a	a	PRON
elements-9012	74	10	n	n	NOUN
elements-9012	74	11	p	p	NOUN
elements-9012	74	12	h	h	NOUN
elements-9012	74	13	y	y	PROPN
elements-9012	74	14	l	l	NOUN
elements-9012	74	15	o	o	NOUN
elements-9012	74	16	g	g	X
elements-9012	74	17	e	e	PROPN
elements-9012	74	18	n	n	CCONJ
elements-9012	74	19	y	y	PROPN
elements-9012	74	20	,	,	PUNCT
elements-9012	74	21	a	a	DET
elements-9012	74	22	s	s	NOUN
elements-9012	74	23	w	w	NOUN
elements-9012	74	24	e	e	NOUN
elements-9012	74	25	l	l	NOUN
elements-9012	74	26	l	l	NOUN
elements-9012	75	1	a	a	PRON
elements-9012	75	2	s	s	X
elements-9012	75	3	i	i	PRON
elements-9012	75	4	n	n	NOUN
elements-9012	75	5	f	f	NOUN
elements-9012	75	6	l	l	NOUN
elements-9012	76	1	i	i	PRON
elements-9012	76	2	e	e	X
elements-9012	76	3	s	s	VERB
elements-9012	76	4	a	a	DET
elements-9012	76	5	n	n	NOUN
elements-9012	77	1	d	d	NOUN
elements-9012	77	2	m	m	NOUN
elements-9012	78	1	o	o	NOUN
elements-9012	78	2	s	s	X
elements-9012	78	3	q	q	X
elements-9012	78	4	u	u	NOUN
elements-9012	78	5	i	i	PRON
elements-9012	78	6	t	t	VERB
elements-9012	78	7	o	o	X
elements-9012	78	8	e	e	X
elements-9012	78	9	s	s	PROPN
elements-9012	78	10	.	.	PUNCT
elements-9012	79	1	a	a	DET
elements-9012	79	2	l	l	NOUN
elements-9012	79	3	l	l	X
elements-9012	79	4	y	y	PROPN
elements-9012	79	5	e	e	PROPN
elements-9012	79	6	a	a	PRON
elements-9012	79	7	s	s	ADP
elements-9012	79	8	t	t	NOUN
elements-9012	79	9	s	s	NOUN
elements-9012	79	10	h	h	NOUN
elements-9012	79	11	a	a	DET
elements-9012	79	12	v	v	X
elements-9012	79	13	e	e	NOUN
elements-9012	79	14	a	a	DET
elements-9012	79	15	u	u	NOUN
elements-9012	80	1	n	n	NOUN
elements-9012	81	1	i	i	PRON
elements-9012	81	2	f	f	NOUN
elements-9012	82	1	o	o	NOUN
elements-9012	82	2	r	r	NOUN
elements-9012	82	3	m	m	VERB
elements-9012	82	4	r	r	NOUN
elements-9012	82	5	a	a	DET
elements-9012	82	6	t	t	NOUN
elements-9012	82	7	e	e	NOUN
elements-9012	82	8	a	a	DET
elements-9012	82	9	c	c	NOUN
elements-9012	82	10	r	r	NOUN
elements-9012	82	11	o	o	NOUN
elements-9012	82	12	s	s	X
elements-9012	82	13	s	s	X
elements-9012	82	14	t	t	NOUN
elements-9012	82	15	h	h	NOUN
elements-9012	83	1	e	e	NOUN
elements-9012	83	2	i	i	PRON
elements-9012	83	3	r	r	VERB
elements-9012	83	4	g	g	NOUN
elements-9012	83	5	e	e	PROPN
elements-9012	83	6	n	n	NOUN
elements-9012	83	7	o	o	NOUN
elements-9012	83	8	m	m	NOUN
elements-9012	83	9	e	e	NOUN
elements-9012	83	10	s	s	X
elements-9012	83	11	w	w	VERB
elements-9012	83	12	i	i	PRON
elements-9012	84	1	t	t	PROPN
elements-9012	84	2	h	h	NOUN
elements-9012	84	3	t	t	NOUN
elements-9012	84	4	h	h	NOUN
elements-9012	84	5	e	e	NOUN
elements-9012	84	6	e	e	X
elements-9012	84	7	x	x	X
elements-9012	84	8	c	c	NOUN
elements-9012	84	9	e	e	X
elements-9012	84	10	p	p	NOUN
elements-9012	84	11	t	t	X
elements-9012	85	1	i	i	PRON
elements-9012	85	2	o	o	VERB
elements-9012	86	1	n	n	X
elements-9012	87	1	o	o	X
elements-9012	87	2	f	f	NOUN
elements-9012	88	1	t	t	NOUN
elements-9012	88	2	h	h	NOUN
elements-9012	88	3	r	r	NOUN
elements-9012	88	4	e	e	PROPN
elements-9012	88	5	e	e	X
elements-9012	88	6	candida	candida	PROPN
elements-9012	88	7	s	s	PART
elements-9012	88	8	p	p	NOUN
elements-9012	88	9	e	e	X
elements-9012	88	10	c	c	NOUN
elements-9012	88	11	i	i	NOUN
elements-9012	88	12	e	e	X
elements-9012	88	13	s	s	X
elements-9012	88	14	!	!	PUNCT
elements-9012	89	1	c	c	NOUN
elements-9012	89	2	.	.	PUNCT
elements-9012	90	1	albicans	albican	NOUN
elements-9012	90	2	,	,	PUNCT
elements-9012	90	3	c.	c.	PROPN
elements-9012	90	4	d	d	PROPN
elements-9012	90	5	u	u	PROPN
elements-9012	90	6	b	b	PROPN
elements-9012	90	7	l	l	NOUN
elements-9012	91	1	i	i	PRON
elements-9012	92	1	n	n	VERB
elements-9012	93	1	i	i	NOUN
elements-9012	93	2	e	e	PROPN
elements-9012	93	3	n	n	NUM
elements-9012	93	4	sis	sis	NOUN
elements-9012	93	5	,	,	PUNCT
elements-9012	93	6	a	a	DET
elements-9012	93	7	n	n	NOUN
elements-9012	93	8	d	d	NOUN
elements-9012	93	9	c	c	PROPN
elements-9012	93	10	.	.	PUNCT
elements-9012	94	1	tropicalis	tropicalis	INTJ
elements-9012	94	2	.	.	PUNCT
elements-9012	95	1	w	w	PROPN
elements-9012	95	2	e	e	PROPN
elements-9012	95	3	h	h	NOUN
elements-9012	95	4	y	y	PROPN
elements-9012	95	5	p	p	X
elements-9012	96	1	o	o	X
elements-9012	96	2	t	t	NOUN
elements-9012	96	3	h	h	NOUN
elements-9012	97	1	e	e	X
elements-9012	97	2	s	s	VERB
elements-9012	97	3	i	i	X
elements-9012	97	4	z	z	NOUN
elements-9012	97	5	e	e	NOUN
elements-9012	97	6	t	t	PROPN
elements-9012	97	7	h	h	PROPN
elements-9012	97	8	a	a	DET
elements-9012	97	9	t	t	NOUN
elements-9012	97	10	t	t	NOUN
elements-9012	98	1	h	h	NOUN
elements-9012	99	1	i	i	PRON
elements-9012	99	2	s	s	VERB
elements-9012	100	1	i	i	PRON
elements-9012	100	2	s	s	VERB
elements-9012	100	3	d	d	X
elements-9012	100	4	u	u	X
elements-9012	100	5	e	e	NOUN
elements-9012	100	6	t	t	NOUN
elements-9012	100	7	o	o	X
elements-9012	100	8	t	t	NOUN
elements-9012	100	9	h	h	NOUN
elements-9012	100	10	e	e	NOUN
elements-9012	100	11	l	l	NOUN
elements-9012	101	1	a	a	X
elements-9012	101	2	c	c	NOUN
elements-9012	101	3	k	k	NOUN
elements-9012	101	4	o	o	X
elements-9012	101	5	f	f	X
elements-9012	101	6	s	s	X
elements-9012	101	7	e	e	X
elements-9012	101	8	x	x	X
elements-9012	101	9	u	u	PROPN
elements-9012	101	10	a	a	DET
elements-9012	101	11	l	l	NOUN
elements-9012	101	12	r	r	NOUN
elements-9012	101	13	e	e	NOUN
elements-9012	101	14	c	c	NOUN
elements-9012	101	15	o	o	NOUN
elements-9012	101	16	m	m	NOUN
elements-9012	101	17	b	b	NOUN
elements-9012	102	1	i	i	PRON
elements-9012	102	2	n	n	PROPN
elements-9012	102	3	a	a	DET
elements-9012	102	4	t	t	NOUN
elements-9012	103	1	i	i	PRON
elements-9012	103	2	o	o	INTJ
elements-9012	104	1	n	n	VERB
elements-9012	105	1	i	i	PRON
elements-9012	105	2	n	n	VERB
elements-9012	105	3	t	t	NOUN
elements-9012	105	4	h	h	NOUN
elements-9012	105	5	e	e	NOUN
elements-9012	105	6	s	s	PROPN
elements-9012	105	7	e	e	X
elements-9012	105	8	s	s	X
elements-9012	105	9	p	p	X
elements-9012	105	10	e	e	NOUN
elements-9012	105	11	c	c	NOUN
elements-9012	106	1	i	i	NOUN
elements-9012	106	2	e	e	NOUN
elements-9012	106	3	s	s	X
elements-9012	106	4	,	,	PUNCT
elements-9012	106	5	l	l	PROPN
elements-9012	106	6	e	e	NOUN
elements-9012	106	7	a	a	X
elements-9012	106	8	d	d	X
elements-9012	106	9	i	i	PRON
elements-9012	106	10	n	n	CCONJ
elements-9012	106	11	g	g	NOUN
elements-9012	106	12	t	t	X
elements-9012	106	13	o	o	X
elements-9012	106	14	t	t	NOUN
elements-9012	106	15	h	h	NOUN
elements-9012	106	16	e	e	NOUN
elements-9012	106	17	r	r	NOUN
elements-9012	106	18	e	e	NOUN
elements-9012	106	19	g	g	NOUN
elements-9012	107	1	i	i	PRON
elements-9012	107	2	o	o	VERB
elements-9012	108	1	n	n	CCONJ
elements-9012	108	2	a	a	DET
elements-9012	108	3	l	l	NOUN
elements-9012	109	1	a	a	DET
elements-9012	109	2	c	c	NOUN
elements-9012	109	3	c	c	NOUN
elements-9012	109	4	u	u	NOUN
elements-9012	109	5	m	m	NOUN
elements-9012	109	6	u	u	NOUN
elements-9012	109	7	l	l	NOUN
elements-9012	109	8	a	a	NOUN
elements-9012	109	9	t	t	NOUN
elements-9012	110	1	i	i	PRON
elements-9012	110	2	o	o	VERB
elements-9012	111	1	n	n	X
elements-9012	111	2	o	o	X
elements-9012	112	1	f	f	VERB
elements-9012	112	2	m	m	VERB
elements-9012	112	3	u	u	NOUN
elements-9012	112	4	t	t	PROPN
elements-9012	112	5	a	a	DET
elements-9012	112	6	t	t	NOUN
elements-9012	113	1	i	i	PRON
elements-9012	113	2	o	o	X
elements-9012	113	3	n	n	X
elements-9012	113	4	s	s	NOUN
elements-9012	113	5	.	.	PUNCT
elements-9012	114	1	b	b	X
elements-9012	115	1	a	a	PRON
elements-9012	115	2	c	c	NOUN
elements-9012	116	1	k	k	NOUN
elements-9012	116	2	g	g	PROPN
elements-9012	116	3	r	r	NOUN
elements-9012	116	4	o	o	X
elements-9012	116	5	u	u	NOUN
elements-9012	116	6	n	n	NOUN
elements-9012	116	7	d	d	PROPN
elements-9012	116	8	popular	popular	ADJ
elements-9012	116	9	understanding	understanding	NOUN
elements-9012	116	10	of	of	ADP
elements-9012	116	11	gene	gene	NOUN
elements-9012	116	12	mutations	mutation	NOUN
elements-9012	116	13	often	often	ADV
elements-9012	116	14	focuses	focus	VERB
elements-9012	116	15	on	on	ADP
elements-9012	116	16	those	those	PRON
elements-9012	116	17	that	that	PRON
elements-9012	116	18	have	have	VERB
elements-9012	116	19	obvious	obvious	ADJ
elements-9012	116	20	effects	effect	NOUN
elements-9012	116	21	,	,	PUNCT
elements-9012	116	22	such	such	ADJ
elements-9012	116	23	as	as	ADP
elements-9012	116	24	insects	insect	NOUN
elements-9012	116	25	that	that	PRON
elements-9012	116	26	develop	develop	VERB
elements-9012	116	27	resistance	resistance	NOUN
elements-9012	116	28	to	to	ADP
elements-9012	116	29	pesticides	pesticide	NOUN
elements-9012	116	30	.	.	PUNCT
elements-9012	117	1	however	however	ADV
elements-9012	117	2	,	,	PUNCT
elements-9012	117	3	the	the	DET
elements-9012	117	4	majority	majority	NOUN
elements-9012	117	5	of	of	ADP
elements-9012	117	6	mutations	mutation	NOUN
elements-9012	117	7	have	have	VERB
elements-9012	117	8	neither	neither	CCONJ
elements-9012	117	9	a	a	DET
elements-9012	117	10	positive	positive	ADJ
elements-9012	117	11	nor	nor	CCONJ
elements-9012	117	12	a	a	DET
elements-9012	117	13	negative	negative	ADJ
elements-9012	117	14	impact	impact	NOUN
elements-9012	117	15	because	because	SCONJ
elements-9012	117	16	they	they	PRON
elements-9012	117	17	do	do	AUX
elements-9012	117	18	not	not	PART
elements-9012	117	19	affect	affect	VERB
elements-9012	117	20	functional	functional	ADJ
elements-9012	117	21	regulatory	regulatory	ADJ
elements-9012	117	22	elements	element	NOUN
elements-9012	117	23	,	,	PUNCT
elements-9012	117	24	which	which	PRON
elements-9012	117	25	generally	generally	ADV
elements-9012	117	26	make	make	VERB
elements-9012	117	27	up	up	ADP
elements-9012	117	28	a	a	DET
elements-9012	117	29	small	small	ADJ
elements-9012	117	30	percentage	percentage	NOUN
elements-9012	117	31	of	of	ADP
elements-9012	117	32	the	the	DET
elements-9012	117	33	genome	genome	NOUN
elements-9012	117	34	.	.	PUNCT
elements-9012	118	1	these	these	DET
elements-9012	118	2	mutations	mutation	NOUN
elements-9012	118	3	are	be	AUX
elements-9012	118	4	known	know	VERB
elements-9012	118	5	as	as	ADP
elements-9012	118	6	"	"	PUNCT
elements-9012	118	7	neutral	neutral	ADJ
elements-9012	118	8	mutations	mutation	NOUN
elements-9012	118	9	.	.	PUNCT
elements-9012	118	10	"	"	PUNCT
elements-9012	119	1	the	the	DET
elements-9012	119	2	presence	presence	NOUN
elements-9012	119	3	and	and	CCONJ
elements-9012	119	4	rate	rate	NOUN
elements-9012	119	5	at	at	ADP
elements-9012	119	6	which	which	PRON
elements-9012	119	7	these	these	DET
elements-9012	119	8	mutations	mutation	NOUN
elements-9012	119	9	occur	occur	VERB
elements-9012	119	10	can	can	AUX
elements-9012	119	11	be	be	AUX
elements-9012	119	12	used	use	VERB
elements-9012	119	13	to	to	PART
elements-9012	119	14	illuminate	illuminate	VERB
elements-9012	119	15	evolutionary	evolutionary	ADJ
elements-9012	119	16	relationships	relationship	NOUN
elements-9012	119	17	between	between	ADP
elements-9012	119	18	different	different	ADJ
elements-9012	119	19	species	specie	NOUN
elements-9012	119	20	,	,	PUNCT
elements-9012	119	21	as	as	ADV
elements-9012	119	22	well	well	ADV
elements-9012	119	23	as	as	ADP
elements-9012	119	24	to	to	PART
elements-9012	119	25	distinguish	distinguish	VERB
elements-9012	119	26	areas	area	NOUN
elements-9012	119	27	of	of	ADP
elements-9012	119	28	functional	functional	ADJ
elements-9012	119	29	regulatory	regulatory	ADJ
elements-9012	119	30	elements	element	NOUN
elements-9012	119	31	from	from	ADP
elements-9012	119	32	non	non	ADJ
elements-9012	119	33	-	-	ADJ
elements-9012	119	34	functional	functional	ADJ
elements-9012	119	35	dna	dna	NOUN
elements-9012	119	36	,	,	PUNCT
elements-9012	119	37	a	a	DET
elements-9012	119	38	process	process	NOUN
elements-9012	119	39	called	call	VERB
elements-9012	119	40	"	"	PUNCT
elements-9012	119	41	phylogenetic	phylogenetic	ADJ
elements-9012	119	42	footprinting	footprinting	NOUN
elements-9012	119	43	.	.	PUNCT
elements-9012	119	44	"	"	PUNCT
elements-9012	120	1	though	though	SCONJ
elements-9012	120	2	neutral	neutral	ADJ
elements-9012	120	3	mutation	mutation	NOUN
elements-9012	120	4	rates	rate	NOUN
elements-9012	120	5	were	be	AUX
elements-9012	120	6	once	once	ADV
elements-9012	120	7	considered	consider	VERB
elements-9012	120	8	to	to	PART
elements-9012	120	9	be	be	AUX
elements-9012	120	10	uniform	uniform	ADJ
elements-9012	120	11	,	,	PUNCT
elements-9012	120	12	it	it	PRON
elements-9012	120	13	has	have	AUX
elements-9012	120	14	been	be	AUX
elements-9012	120	15	discovered	discover	VERB
elements-9012	120	16	that	that	SCONJ
elements-9012	120	17	they	they	PRON
elements-9012	120	18	can	can	AUX
elements-9012	120	19	vary	vary	VERB
elements-9012	120	20	dramatically	dramatically	ADV
elements-9012	120	21	,	,	PUNCT
elements-9012	120	22	not	not	PART
elements-9012	120	23	only	only	ADV
elements-9012	120	24	from	from	ADP
elements-9012	120	25	one	one	NUM
elements-9012	120	26	species	specie	NOUN
elements-9012	120	27	to	to	ADP
elements-9012	120	28	the	the	DET
elements-9012	120	29	next	next	ADJ
elements-9012	120	30	but	but	CCONJ
elements-9012	120	31	also	also	ADV
elements-9012	120	32	within	within	ADP
elements-9012	120	33	a	a	DET
elements-9012	120	34	single	single	ADJ
elements-9012	120	35	genome.1	genome.1	PROPN
elements-9012	120	36	within	within	ADP
elements-9012	120	37	-	-	PUNCT
elements-9012	120	38	genome	genome	NOUN
elements-9012	120	39	heterogeneity	heterogeneity	NOUN
elements-9012	120	40	has	have	AUX
elements-9012	120	41	been	be	AUX
elements-9012	120	42	demonstrated	demonstrate	VERB
elements-9012	120	43	in	in	ADP
elements-9012	120	44	several	several	ADJ
elements-9012	120	45	investigations	investigation	NOUN
elements-9012	120	46	of	of	ADP
elements-9012	120	47	mammalian	mammalian	ADJ
elements-9012	120	48	species.u"vn	species.u"vn	PROPN
elements-9012	120	49	i	i	PRON
elements-9012	120	50	n	n	VERB
elements-9012	120	51	these	these	DET
elements-9012	120	52	species	specie	NOUN
elements-9012	120	53	,	,	PUNCT
elements-9012	120	54	the	the	DET
elements-9012	120	55	rate	rate	NOUN
elements-9012	120	56	of	of	ADP
elements-9012	120	57	neutral	neutral	ADJ
elements-9012	120	58	mutations	mutation	NOUN
elements-9012	120	59	is	be	AUX
elements-9012	120	60	different	different	ADJ
elements-9012	120	61	in	in	ADP
elements-9012	120	62	different	different	ADJ
elements-9012	120	63	regions	region	NOUN
elements-9012	120	64	of	of	ADP
elements-9012	120	65	the	the	DET
elements-9012	120	66	genome	genome	NOUN
elements-9012	120	67	,	,	PUNCT
elements-9012	120	68	such	such	ADJ
elements-9012	120	69	as	as	ADP
elements-9012	120	70	that	that	PRON
elements-9012	120	71	of	of	ADP
elements-9012	120	72	chromosome	chromosome	NOUN
elements-9012	120	73	i	i	PRON
elements-9012	120	74	compared	compare	VERB
elements-9012	120	75	to	to	ADP
elements-9012	120	76	that	that	PRON
elements-9012	120	77	of	of	ADP
elements-9012	120	78	chromosome	chromosome	NOUN
elements-9012	120	79	4	4	NUM
elements-9012	120	80	.	.	PUNCT
elements-9012	121	1	in	in	ADP
elements-9012	121	2	contrast	contrast	NOUN
elements-9012	121	3	,	,	PUNCT
elements-9012	121	4	uniform	uniform	ADJ
elements-9012	121	5	mutation	mutation	NOUN
elements-9012	121	6	rates	rate	NOUN
elements-9012	121	7	have	have	AUX
elements-9012	121	8	been	be	AUX
elements-9012	121	9	observed	observe	VERB
elements-9012	121	10	only	only	ADV
elements-9012	121	11	in	in	ADP
elements-9012	121	12	the	the	DET
elements-9012	121	13	phylogeny	phylogeny	NOUN
elements-9012	121	14	of	of	ADP
elements-9012	121	15	the	the	DET
elements-9012	121	16	sensu	sensu	NOUN
elements-9012	121	17	stricto	stricto	X
elements-9012	121	18	yeasts	yeast	NOUN
elements-9012	121	19	s.	s.	PROPN
elements-9012	121	20	cerevisiae	cerevisiae	PROPN
elements-9012	121	21	,	,	PUNCT
elements-9012	121	22	s.	s.	PROPN
elements-9012	121	23	paradoxus	paradoxus	PROPN
elements-9012	121	24	,	,	PUNCT
elements-9012	121	25	s.	s.	PROPN
elements-9012	121	26	bayanus	bayanus	PROPN
elements-9012	121	27	,	,	PUNCT
elements-9012	121	28	and	and	CCONJ
elements-9012	121	29	s.	s.	PROPN
elements-9012	121	30	mikatae.vm	mikatae.vm	PROPN
elements-9012	122	1	i	i	PRON
elements-9012	122	2	n	n	VERB
elements-9012	122	3	these	these	DET
elements-9012	122	4	species	specie	NOUN
elements-9012	122	5	,	,	PUNCT
elements-9012	122	6	neutral	neutral	ADJ
elements-9012	122	7	mutations	mutation	NOUN
elements-9012	122	8	occur	occur	VERB
elements-9012	122	9	at	at	ADP
elements-9012	122	10	the	the	DET
elements-9012	122	11	same	same	ADJ
elements-9012	122	12	rate	rate	NOUN
elements-9012	122	13	in	in	ADP
elements-9012	122	14	the	the	DET
elements-9012	122	15	entire	entire	ADJ
elements-9012	122	16	genome	genome	NOUN
elements-9012	122	17	.	.	PUNCT
elements-9012	123	1	the	the	DET
elements-9012	123	2	reason	reason	NOUN
elements-9012	123	3	for	for	ADP
elements-9012	123	4	this	this	DET
elements-9012	123	5	differing	differ	VERB
elements-9012	123	6	mutational	mutational	ADJ
elements-9012	123	7	behavior	behavior	NOUN
elements-9012	123	8	is	be	AUX
elements-9012	123	9	not	not	PART
elements-9012	123	10	known	know	VERB
elements-9012	123	11	,	,	PUNCT
elements-9012	123	12	and	and	CCONJ
elements-9012	123	13	little	little	ADJ
elements-9012	123	14	is	be	AUX
elements-9012	123	15	known	know	VERB
elements-9012	123	16	about	about	ADP
elements-9012	123	17	regional	regional	ADJ
elements-9012	123	18	biases	bias	NOUN
elements-9012	123	19	in	in	ADP
elements-9012	123	20	other	other	ADJ
elements-9012	123	21	species	specie	NOUN
elements-9012	123	22	.	.	PUNCT
elements-9012	124	1	a	a	DET
elements-9012	124	2	better	well	ADJ
elements-9012	124	3	characterization	characterization	NOUN
elements-9012	124	4	of	of	ADP
elements-9012	124	5	regional	regional	ADJ
elements-9012	124	6	biases	bias	NOUN
elements-9012	124	7	would	would	AUX
elements-9012	124	8	improve	improve	VERB
elements-9012	124	9	fundamental	fundamental	ADJ
elements-9012	124	10	understanding	understanding	NOUN
elements-9012	124	11	of	of	ADP
elements-9012	124	12	dna	dna	PROPN
elements-9012	124	13	mutation	mutation	NOUN
elements-9012	124	14	.	.	PUNCT
elements-9012	125	1	it	it	PRON
elements-9012	125	2	would	would	AUX
elements-9012	125	3	also	also	ADV
elements-9012	125	4	aid	aid	VERB
elements-9012	125	5	i	i	PRON
elements-9012	125	6	n	n	CCONJ
elements-9012	125	7	the	the	DET
elements-9012	125	8	calibration	calibration	NOUN
elements-9012	125	9	of	of	ADP
elements-9012	125	10	phylogenetic	phylogenetic	ADJ
elements-9012	125	11	footprinting	footprinting	NOUN
elements-9012	125	12	methods	method	NOUN
elements-9012	125	13	,	,	PUNCT
elements-9012	125	14	which	which	PRON
elements-9012	125	15	are	be	AUX
elements-9012	125	16	used	use	VERB
elements-9012	125	17	to	to	PART
elements-9012	125	18	detect	detect	VERB
elements-9012	125	19	sequences	sequence	NOUN
elements-9012	125	20	under	under	ADP
elements-9012	125	21	purifying	purify	VERB
elements-9012	125	22	selection	selection	NOUN
elements-9012	125	23	.	.	PUNCT
elements-9012	126	1	for	for	ADP
elements-9012	126	2	example	example	NOUN
elements-9012	126	3	,	,	PUNCT
elements-9012	126	4	uncertainties	uncertainty	NOUN
elements-9012	126	5	in	in	ADP
elements-9012	126	6	regional	regional	ADJ
elements-9012	126	7	mutation	mutation	NOUN
elements-9012	126	8	biases	bias	NOUN
elements-9012	126	9	have	have	AUX
elements-9012	126	10	hindered	hinder	VERB
elements-9012	126	11	estimates	estimate	NOUN
elements-9012	126	12	of	of	ADP
elements-9012	126	13	the	the	DET
elements-9012	126	14	amount	amount	NOUN
elements-9012	126	15	of	of	ADP
elements-9012	126	16	functional	functional	ADJ
elements-9012	126	17	human	human	ADJ
elements-9012	126	18	dna	dna	NOUN
elements-9012	126	19	.	.	PUNCT
elements-9012	127	1	i	i	PRON
elements-9012	127	2	x	x	PUNCT
elements-9012	127	3	most	most	ADV
elements-9012	127	4	previous	previous	ADJ
elements-9012	127	5	studies	study	NOUN
elements-9012	127	6	of	of	ADP
elements-9012	127	7	mutation	mutation	NOUN
elements-9012	127	8	rate	rate	NOUN
elements-9012	127	9	variation	variation	NOUN
elements-9012	127	10	have	have	AUX
elements-9012	127	11	focused	focus	VERB
elements-9012	127	12	on	on	ADP
elements-9012	127	13	mammals	mammal	NOUN
elements-9012	127	14	,	,	PUNCT
elements-9012	127	15	including	include	VERB
elements-9012	127	16	human	human	NOUN
elements-9012	127	17	,	,	PUNCT
elements-9012	127	18	chimp	chimp	PROPN
elements-9012	127	19	,	,	PUNCT
elements-9012	127	20	mouse	mouse	PROPN
elements-9012	127	21	,	,	PUNCT
elements-9012	127	22	rat	rat	NOUN
elements-9012	127	23	,	,	PUNCT
elements-9012	127	24	dog	dog	NOUN
elements-9012	127	25	,	,	PUNCT
elements-9012	127	26	and	and	CCONJ
elements-9012	127	27	cow	cow	NOUN
elements-9012	127	28	.	.	PUNCT
elements-9012	128	1	such	such	ADJ
elements-9012	128	2	studies	study	NOUN
elements-9012	128	3	have	have	AUX
elements-9012	128	4	identified	identify	VERB
elements-9012	128	5	regional	regional	ADJ
elements-9012	128	6	effects	effect	NOUN
elements-9012	128	7	by	by	ADP
elements-9012	128	8	studying	study	VERB
elements-9012	128	9	the	the	DET
elements-9012	128	10	correlation	correlation	NOUN
elements-9012	128	11	of	of	ADP
elements-9012	128	12	independent	independent	ADJ
elements-9012	128	13	neutral	neutral	ADJ
elements-9012	128	14	measures	measure	NOUN
elements-9012	128	15	,	,	PUNCT
elements-9012	128	16	such	such	ADJ
elements-9012	128	17	as	as	ADP
elements-9012	128	18	single	single	ADJ
elements-9012	128	19	nucleotide	nucleotide	ADJ
elements-9012	128	20	polymorphism	polymorphism	NOUN
elements-9012	128	21	(	(	PUNCT
elements-9012	128	22	snp	snp	ADJ
elements-9012	128	23	)	)	PUNCT
elements-9012	128	24	density	density	NOUN
elements-9012	128	25	,	,	PUNCT
elements-9012	128	26	insertion	insertion	NOUN
elements-9012	128	27	-	-	PUNCT
elements-9012	128	28	deletion	deletion	NOUN
elements-9012	128	29	(	(	PUNCT
elements-9012	128	30	indel	indel	NOUN
elements-9012	128	31	)	)	PUNCT
elements-9012	128	32	density	density	NOUN
elements-9012	128	33	,	,	PUNCT
elements-9012	128	34	substitution	substitution	NOUN
elements-9012	128	35	rate	rate	NOUN
elements-9012	128	36	in	in	ADP
elements-9012	128	37	ancestral	ancestral	ADJ
elements-9012	128	38	repeats	repeat	NOUN
elements-9012	128	39	,	,	PUNCT
elements-9012	128	40	or	or	CCONJ
elements-9012	128	41	substitution	substitution	NOUN
elements-9012	128	42	rates	rate	NOUN
elements-9012	128	43	in	in	ADP
elements-9012	128	44	silent	silent	ADJ
elements-9012	128	45	sites.x	sites.x	PROPN
elements-9012	128	46	regional	regional	ADJ
elements-9012	128	47	effects	effect	NOUN
elements-9012	128	48	have	have	AUX
elements-9012	128	49	also	also	ADV
elements-9012	128	50	been	be	AUX
elements-9012	128	51	characterized	characterize	VERB
elements-9012	128	52	via	via	ADP
elements-9012	128	53	the	the	DET
elements-9012	128	54	length	length	NOUN
elements-9012	128	55	scales	scale	NOUN
elements-9012	128	56	at	at	ADP
elements-9012	128	57	which	which	PRON
elements-9012	128	58	nearby	nearby	ADJ
elements-9012	128	59	neutral	neutral	ADJ
elements-9012	128	60	sequences	sequence	NOUN
elements-9012	128	61	are	be	AUX
elements-9012	128	62	correlated	correlate	VERB
elements-9012	128	63	,	,	PUNCT
elements-9012	128	64	as	as	ADV
elements-9012	128	65	well	well	ADV
elements-9012	128	66	as	as	ADP
elements-9012	128	67	via	via	ADP
elements-9012	128	68	the	the	DET
elements-9012	128	69	strength	strength	NOUN
elements-9012	128	70	of	of	ADP
elements-9012	128	71	correlations	correlation	NOUN
elements-9012	128	72	within	within	ADP
elements-9012	128	73	single	single	ADJ
elements-9012	128	74	genes.x1	genes.x1	PROPN
elements-9012	128	75	proposed	propose	VERB
elements-9012	128	76	causes	cause	NOUN
elements-9012	128	77	of	of	ADP
elements-9012	128	78	mutation	mutation	NOUN
elements-9012	128	79	rate	rate	NOUN
elements-9012	128	80	variation	variation	NOUN
elements-9012	128	81	have	have	AUX
elements-9012	128	82	included	include	VERB
elements-9012	128	83	regional	regional	ADJ
elements-9012	128	84	variations	variation	NOUN
elements-9012	128	85	in	in	ADP
elements-9012	128	86	base	base	NOUN
elements-9012	128	87	composition	composition	NOUN
elements-9012	128	88	,	,	PUNCT
elements-9012	128	89	recombination	recombination	NOUN
elements-9012	128	90	,	,	PUNCT
elements-9012	128	91	gene	gene	NOUN
elements-9012	128	92	density	density	NOUN
elements-9012	128	93	,	,	PUNCT
elements-9012	128	94	or	or	CCONJ
elements-9012	128	95	pattern	pattern	NOUN
elements-9012	128	96	of	of	ADP
elements-9012	128	97	gene	gene	NOUN
elements-9012	128	98	expression/1	expression/1	NOUN
elements-9012	128	99	1	1	NUM
elements-9012	128	100	unfortunately	unfortunately	ADV
elements-9012	128	101	,	,	PUNCT
elements-9012	128	102	the	the	DET
elements-9012	128	103	causes	cause	NOUN
elements-9012	128	104	of	of	ADP
elements-9012	128	105	variation	variation	NOUN
elements-9012	128	106	in	in	ADP
elements-9012	128	107	these	these	DET
elements-9012	128	108	other	other	ADJ
elements-9012	128	109	features	feature	NOUN
elements-9012	128	110	are	be	AUX
elements-9012	128	111	not	not	PART
elements-9012	128	112	clearly	clearly	ADV
elements-9012	128	113	understood	understand	VERB
elements-9012	128	114	either	either	ADV
elements-9012	128	115	.	.	PUNCT
elements-9012	129	1	regional	regional	ADJ
elements-9012	129	2	effects	effect	NOUN
elements-9012	129	3	must	must	AUX
elements-9012	129	4	be	be	AUX
elements-9012	129	5	different	different	ADJ
elements-9012	129	6	in	in	ADP
elements-9012	129	7	yeast	yeast	NOUN
elements-9012	129	8	since	since	SCONJ
elements-9012	129	9	yeast	yeast	NOUN
elements-9012	129	10	chromosomes	chromosome	NOUN
elements-9012	129	11	are	be	AUX
elements-9012	129	12	typically	typically	ADV
elements-9012	129	13	only	only	ADV
elements-9012	129	14	one	one	NUM
elements-9012	129	15	-	-	PUNCT
elements-9012	129	16	hundredth	hundredth	ADJ
elements-9012	129	17	as	as	ADV
elements-9012	129	18	long	long	ADV
elements-9012	129	19	as	as	ADP
elements-9012	129	20	human	human	ADJ
elements-9012	129	21	chromosomes	chromosome	NOUN
elements-9012	129	22	.	.	PUNCT
elements-9012	130	1	the	the	DET
elements-9012	130	2	length	length	NOUN
elements-9012	130	3	scales	scale	NOUN
elements-9012	130	4	of	of	ADP
elements-9012	130	5	these	these	DET
elements-9012	130	6	various	various	ADJ
elements-9012	130	7	types	type	NOUN
elements-9012	130	8	of	of	ADP
elements-9012	130	9	mammalian	mammalian	ADJ
elements-9012	130	10	regional	regional	ADJ
elements-9012	130	11	variation	variation	NOUN
elements-9012	130	12	are	be	AUX
elements-9012	130	13	often	often	ADV
elements-9012	130	14	as	as	ADV
elements-9012	130	15	large	large	ADJ
elements-9012	130	16	as	as	ADP
elements-9012	130	17	a	a	DET
elements-9012	130	18	yeast	yeast	NOUN
elements-9012	130	19	chromosome	chromosome	NOUN
elements-9012	130	20	.	.	PUNCT
elements-9012	131	1	a	a	DET
elements-9012	131	2	n	n	CCONJ
elements-9012	131	3	important	important	ADJ
elements-9012	131	4	step	step	NOUN
elements-9012	131	5	toward	toward	ADP
elements-9012	131	6	understanding	understand	VERB
elements-9012	131	7	the	the	DET
elements-9012	131	8	causes	cause	NOUN
elements-9012	131	9	of	of	ADP
elements-9012	131	10	mutational	mutational	ADJ
elements-9012	131	11	heterogeneity	heterogeneity	NOUN
elements-9012	131	12	is	be	AUX
elements-9012	131	13	to	to	PART
elements-9012	131	14	measure	measure	VERB
elements-9012	131	15	which	which	DET
elements-9012	131	16	species	specie	NOUN
elements-9012	131	17	have	have	VERB
elements-9012	131	18	heterogeneous	heterogeneous	ADJ
elements-9012	131	19	mutation	mutation	NOUN
elements-9012	131	20	rates	rate	NOUN
elements-9012	131	21	and	and	CCONJ
elements-9012	131	22	which	which	DET
elements-9012	131	23	species	specie	NOUN
elements-9012	131	24	have	have	VERB
elements-9012	131	25	homogeneous	homogeneous	ADJ
elements-9012	131	26	mutation	mutation	NOUN
elements-9012	131	27	rates	rate	NOUN
elements-9012	131	28	.	.	PUNCT
elements-9012	132	1	in	in	ADP
elements-9012	132	2	this	this	DET
elements-9012	132	3	work	work	NOUN
elements-9012	132	4	,	,	PUNCT
elements-9012	132	5	we	we	PRON
elements-9012	132	6	analyze	analyze	VERB
elements-9012	132	7	regional	regional	ADJ
elements-9012	132	8	mutational	mutational	ADJ
elements-9012	132	9	biases	bias	NOUN
elements-9012	132	10	in	in	ADP
elements-9012	132	11	20	20	NUM
elements-9012	132	12	mammalian	mammalian	NOUN
elements-9012	132	13	,	,	PUNCT
elements-9012	132	14	27	27	NUM
elements-9012	132	15	yeast	yeast	NOUN
elements-9012	132	16	,	,	PUNCT
elements-9012	132	17	and	and	CCONJ
elements-9012	132	18	4	4	NUM
elements-9012	132	19	insect	insect	NOUN
elements-9012	132	20	genomes	genome	NOUN
elements-9012	132	21	,	,	PUNCT
elements-9012	132	22	determining	determine	VERB
elements-9012	132	23	which	which	DET
elements-9012	132	24	clades	clade	NOUN
elements-9012	132	25	(	(	PUNCT
elements-9012	132	26	taxonomic	taxonomic	ADJ
elements-9012	132	27	groups	group	NOUN
elements-9012	132	28	)	)	PUNCT
elements-9012	132	29	have	have	VERB
elements-9012	132	30	uniform	uniform	NOUN
elements-9012	132	31	and	and	CCONJ
elements-9012	132	32	which	which	PRON
elements-9012	132	33	have	have	VERB
elements-9012	132	34	heterogeneous	heterogeneous	ADJ
elements-9012	132	35	mutation	mutation	NOUN
elements-9012	132	36	rates	rate	NOUN
elements-9012	132	37	.	.	PUNCT
elements-9012	133	1	we	we	PRON
elements-9012	133	2	answer	answer	VERB
elements-9012	133	3	the	the	DET
elements-9012	133	4	questions	question	NOUN
elements-9012	133	5	:	:	PUNCT
elements-9012	133	6	do	do	AUX
elements-9012	133	7	all	all	DET
elements-9012	133	8	mammals	mammal	NOUN
elements-9012	133	9	exhibit	exhibit	VERB
elements-9012	133	10	heterogeneity	heterogeneity	NOUN
elements-9012	133	11	in	in	ADP
elements-9012	133	12	their	their	PRON
elements-9012	133	13	mutation	mutation	NOUN
elements-9012	133	14	rates	rate	NOUN
elements-9012	133	15	?	?	PUNCT
elements-9012	134	1	do	do	AUX
elements-9012	134	2	all	all	DET
elements-9012	134	3	yeasts	yeast	NOUN
elements-9012	134	4	exhibit	exhibit	VERB
elements-9012	134	5	uniformity	uniformity	NOUN
elements-9012	134	6	?	?	PUNCT
elements-9012	135	1	studying	study	VERB
elements-9012	135	2	these	these	DET
elements-9012	135	3	phylogenies	phylogeny	NOUN
elements-9012	135	4	together	together	ADV
elements-9012	135	5	provides	provide	VERB
elements-9012	135	6	a	a	DET
elements-9012	135	7	valuable	valuable	ADJ
elements-9012	135	8	contrast	contrast	NOUN
elements-9012	135	9	.	.	PUNCT
elements-9012	136	1	results	result	NOUN
elements-9012	136	2	there	there	PRON
elements-9012	136	3	are	be	VERB
elements-9012	136	4	many	many	ADJ
elements-9012	136	5	sequences	sequence	NOUN
elements-9012	136	6	in	in	ADP
elements-9012	136	7	the	the	DET
elements-9012	136	8	genome	genome	NOUN
elements-9012	136	9	that	that	PRON
elements-9012	136	10	can	can	AUX
elements-9012	136	11	be	be	AUX
elements-9012	136	12	used	use	VERB
elements-9012	136	13	to	to	PART
elements-9012	136	14	determine	determine	VERB
elements-9012	136	15	the	the	DET
elements-9012	136	16	neutral	neutral	ADJ
elements-9012	136	17	mutation	mutation	NOUN
elements-9012	136	18	rate	rate	NOUN
elements-9012	136	19	,	,	PUNCT
elements-9012	136	20	such	such	ADJ
elements-9012	136	21	as	as	ADP
elements-9012	136	22	pseudogenes	pseudogene	NOUN
elements-9012	136	23	,	,	PUNCT
elements-9012	136	24	ancestral	ancestral	ADJ
elements-9012	136	25	repeats	repeat	NOUN
elements-9012	136	26	,	,	PUNCT
elements-9012	136	27	intergenic	intergenic	ADJ
elements-9012	136	28	regions	region	NOUN
elements-9012	136	29	and	and	CCONJ
elements-9012	136	30	synonymous	synonymous	ADJ
elements-9012	136	31	sites	site	NOUN
elements-9012	136	32	.	.	PUNCT
elements-9012	137	1	in	in	ADP
elements-9012	137	2	this	this	DET
elements-9012	137	3	work	work	NOUN
elements-9012	137	4	,	,	PUNCT
elements-9012	137	5	we	we	PRON
elements-9012	137	6	focus	focus	VERB
elements-9012	137	7	on	on	ADP
elements-9012	137	8	synonymous	synonymous	ADJ
elements-9012	137	9	sites	site	NOUN
elements-9012	137	10	,	,	PUNCT
elements-9012	137	11	and	and	CCONJ
elements-9012	137	12	in	in	ADP
elements-9012	137	13	particular	particular	ADJ
elements-9012	137	14	,	,	PUNCT
elements-9012	137	15	four	four	NUM
elements-9012	137	16	-	-	ADJ
elements-9012	137	17	fold	fold	ADJ
elements-9012	137	18	degenerate	degenerate	ADJ
elements-9012	137	19	sites	site	NOUN
elements-9012	137	20	,	,	PUNCT
elements-9012	137	21	from	from	ADP
elements-9012	137	22	coding	code	VERB
elements-9012	137	23	regions	region	NOUN
elements-9012	137	24	in	in	ADP
elements-9012	137	25	51	51	NUM
elements-9012	137	26	species	specie	NOUN
elements-9012	137	27	.	.	PUNCT
elements-9012	138	1	these	these	DET
elements-9012	138	2	sites	site	NOUN
elements-9012	138	3	can	can	AUX
elements-9012	138	4	be	be	AUX
elements-9012	138	5	altered	alter	VERB
elements-9012	138	6	without	without	ADP
elements-9012	138	7	affecting	affect	VERB
elements-9012	138	8	the	the	DET
elements-9012	138	9	encoded	encode	VERB
elements-9012	138	10	amino	amino	NOUN
elements-9012	138	11	acid	acid	NOUN
elements-9012	138	12	sequence	sequence	NOUN
elements-9012	138	13	.	.	PUNCT
elements-9012	139	1	while	while	SCONJ
elements-9012	139	2	recent	recent	ADJ
elements-9012	139	3	studies	study	NOUN
elements-9012	139	4	indicate	indicate	VERB
elements-9012	139	5	that	that	SCONJ
elements-9012	139	6	some	some	DET
elements-9012	139	7	synonymous	synonymous	ADJ
elements-9012	139	8	sites	site	NOUN
elements-9012	139	9	are	be	AUX
elements-9012	139	10	under	under	ADP
elements-9012	139	11	selection	selection	NOUN
elements-9012	139	12	,	,	PUNCT
elements-9012	139	13	the	the	DET
elements-9012	139	14	majority	majority	NOUN
elements-9012	139	15	are	be	AUX
elements-9012	139	16	still	still	ADV
elements-9012	139	17	likely	likely	ADJ
elements-9012	139	18	to	to	PART
elements-9012	139	19	be	be	AUX
elements-9012	139	20	neutral	neutral	ADJ
elements-9012	139	21	.	.	PUNCT
elements-9012	140	1	x	x	PUNCT
elements-9012	141	1	i	i	PRON
elements-9012	141	2	l	l	VERB
elements-9012	142	1	i	i	PRON
elements-9012	142	2	to	to	PART
elements-9012	142	3	isolate	isolate	VERB
elements-9012	142	4	lineage	lineage	NOUN
elements-9012	142	5	-	-	PUNCT
elements-9012	142	6	specific	specific	ADJ
elements-9012	142	7	effects	effect	NOUN
elements-9012	142	8	,	,	PUNCT
elements-9012	142	9	species	specie	NOUN
elements-9012	142	10	were	be	AUX
elements-9012	142	11	analyzed	analyze	VERB
elements-9012	142	12	in	in	ADP
elements-9012	142	13	pairs	pair	NOUN
elements-9012	142	14	with	with	ADP
elements-9012	142	15	species	specie	NOUN
elements-9012	142	16	close	close	ADJ
elements-9012	142	17	to	to	ADP
elements-9012	142	18	them	they	PRON
elements-9012	142	19	in	in	ADP
elements-9012	142	20	the	the	DET
elements-9012	142	21	phylogenetic	phylogenetic	ADJ
elements-9012	142	22	tree	tree	NOUN
elements-9012	142	23	.	.	PUNCT
elements-9012	143	1	yeast	yeast	NOUN
elements-9012	143	2	species	specie	NOUN
elements-9012	143	3	pairs	pair	NOUN
elements-9012	143	4	were	be	AUX
elements-9012	143	5	determined	determine	VERB
elements-9012	143	6	from	from	ADP
elements-9012	143	7	the	the	DET
elements-9012	143	8	tree	tree	NOUN
elements-9012	143	9	of	of	ADP
elements-9012	143	10	42	42	NUM
elements-9012	143	11	fungal	fungal	ADJ
elements-9012	143	12	species	specie	NOUN
elements-9012	143	13	of	of	ADP
elements-9012	143	14	fitzpatrick	fitzpatrick	PROPN
elements-9012	143	15	et	et	PROPN
elements-9012	143	16	a	a	DET
elements-9012	143	17	l	l	NOUN
elements-9012	143	18	.	.	PUNCT
elements-9012	144	1	x	x	PUNCT
elements-9012	144	2	i	i	PRON
elements-9012	144	3	v	v	VERB
elements-9012	144	4	for	for	ADP
elements-9012	144	5	the	the	DET
elements-9012	144	6	mammals	mammal	NOUN
elements-9012	144	7	,	,	PUNCT
elements-9012	144	8	species	specie	NOUN
elements-9012	144	9	pairs	pair	NOUN
elements-9012	144	10	were	be	AUX
elements-9012	144	11	chosen	choose	VERB
elements-9012	144	12	based	base	VERB
elements-9012	144	13	on	on	ADP
elements-9012	144	14	recent	recent	ADJ
elements-9012	144	15	encode	encode	ADJ
elements-9012	144	16	(	(	PUNCT
elements-9012	144	17	the	the	DET
elements-9012	144	18	e	e	X
elements-9012	144	19	l	l	NOUN
elements-9012	144	20	e	e	X
elements-9012	144	21	m	m	PROPN
elements-9012	144	22	e	e	NOUN
elements-9012	144	23	n	n	NOUN
elements-9012	144	24	t	t	PROPN
elements-9012	144	25	s	s	NOUN
elements-9012	145	1	s	s	X
elements-9012	145	2	p	p	NOUN
elements-9012	145	3	r	r	NOUN
elements-9012	146	1	i	i	PRON
elements-9012	147	1	n	n	CCONJ
elements-9012	147	2	g	g	NOUN
elements-9012	147	3	08	08	NUM
elements-9012	147	4	encyclopedia	encyclopedia	NOUN
elements-9012	147	5	of	of	ADP
elements-9012	147	6	dna	dna	PROPN
elements-9012	147	7	elements	element	NOUN
elements-9012	147	8	project	project	NOUN
elements-9012	147	9	)	)	PUNCT
elements-9012	147	10	and	and	CCONJ
elements-9012	147	11	wholegenome	wholegenome	NOUN
elements-9012	147	12	-	-	PUNCT
elements-9012	147	13	based	base	VERB
elements-9012	147	14	phytogenies	phytogenie	NOUN
elements-9012	147	15	x	x	SYM
elements-9012	147	16	v	v	NUM
elements-9012	147	17	x	x	SYM
elements-9012	147	18	v	v	ADP
elements-9012	147	19	1	1	NUM
elements-9012	147	20	raw	raw	ADJ
elements-9012	147	21	substitution	substitution	NOUN
elements-9012	147	22	rates	rate	NOUN
elements-9012	147	23	were	be	AUX
elements-9012	147	24	calculated	calculate	VERB
elements-9012	147	25	for	for	ADP
elements-9012	147	26	each	each	DET
elements-9012	147	27	orthologous	orthologous	ADJ
elements-9012	147	28	gene	gene	NOUN
elements-9012	147	29	pair	pair	NOUN
elements-9012	147	30	(	(	PUNCT
elements-9012	147	31	pairs	pair	NOUN
elements-9012	147	32	of	of	ADP
elements-9012	147	33	similar	similar	ADJ
elements-9012	147	34	genes	gene	NOUN
elements-9012	147	35	in	in	ADP
elements-9012	147	36	different	different	ADJ
elements-9012	147	37	species	specie	NOUN
elements-9012	147	38	)	)	PUNCT
elements-9012	147	39	by	by	ADP
elements-9012	147	40	counting	count	VERB
elements-9012	147	41	the	the	DET
elements-9012	147	42	fraction	fraction	NOUN
elements-9012	147	43	of	of	ADP
elements-9012	147	44	observed	observed	ADJ
elements-9012	147	45	substitutions	substitution	NOUN
elements-9012	147	46	at	at	ADP
elements-9012	147	47	four	four	NUM
elements-9012	147	48	-	-	ADJ
elements-9012	147	49	fold	fold	ADJ
elements-9012	147	50	sites	site	NOUN
elements-9012	147	51	and	and	CCONJ
elements-9012	147	52	then	then	ADV
elements-9012	147	53	normalizing	normalize	VERB
elements-9012	147	54	to	to	ADP
elements-9012	147	55	z	z	NOUN
elements-9012	147	56	-	-	PUNCT
elements-9012	147	57	score	score	NOUN
elements-9012	147	58	values	value	NOUN
elements-9012	147	59	(	(	PUNCT
elements-9012	147	60	see	see	VERB
elements-9012	147	61	methods	method	NOUN
elements-9012	147	62	)	)	PUNCT
elements-9012	147	63	.	.	PUNCT
elements-9012	148	1	x	x	PUNCT
elements-9012	148	2	v	v	ADP
elements-9012	148	3	n	n	DET
elements-9012	148	4	this	this	DET
elements-9012	148	5	z	z	NOUN
elements-9012	148	6	-	-	PUNCT
elements-9012	148	7	score	score	NOUN
elements-9012	148	8	normalization	normalization	NOUN
elements-9012	148	9	corrects	correct	NOUN
elements-9012	148	10	for	for	ADP
elements-9012	148	11	the	the	DET
elements-9012	148	12	stochastic	stochastic	ADJ
elements-9012	148	13	finite	finite	ADJ
elements-9012	148	14	-	-	PUNCT
elements-9012	148	15	size	size	NOUN
elements-9012	148	16	effects	effect	NOUN
elements-9012	148	17	that	that	PRON
elements-9012	148	18	result	result	VERB
elements-9012	148	19	from	from	ADP
elements-9012	148	20	genes	gene	NOUN
elements-9012	148	21	having	have	VERB
elements-9012	148	22	different	different	ADJ
elements-9012	148	23	numbers	number	NOUN
elements-9012	148	24	of	of	ADP
elements-9012	148	25	fourfold	fourfold	ADJ
elements-9012	148	26	sites	site	NOUN
elements-9012	148	27	.	.	PUNCT
elements-9012	149	1	y	y	PROPN
elements-9012	149	2	e	e	PROPN
elements-9012	149	3	a	a	DET
elements-9012	149	4	s	s	X
elements-9012	149	5	t	t	NOUN
elements-9012	149	6	the	the	DET
elements-9012	149	7	distribution	distribution	NOUN
elements-9012	149	8	of	of	ADP
elements-9012	149	9	normalized	normalize	VERB
elements-9012	149	10	substitution	substitution	NOUN
elements-9012	149	11	rates	rate	NOUN
elements-9012	149	12	provides	provide	VERB
elements-9012	149	13	a	a	DET
elements-9012	149	14	test	test	NOUN
elements-9012	149	15	of	of	ADP
elements-9012	149	16	whether	whether	SCONJ
elements-9012	149	17	substitution	substitution	NOUN
elements-9012	149	18	rates	rate	NOUN
elements-9012	149	19	are	be	AUX
elements-9012	149	20	uniform	uniform	ADJ
elements-9012	149	21	throughout	throughout	ADP
elements-9012	149	22	a	a	DET
elements-9012	149	23	genome	genome	NOUN
elements-9012	149	24	.	.	PUNCT
elements-9012	150	1	our	our	PRON
elements-9012	150	2	yeast	yeast	NOUN
elements-9012	150	3	phylogeny	phylogeny	NOUN
elements-9012	150	4	is	be	AUX
elements-9012	150	5	comprised	comprise	VERB
elements-9012	150	6	of	of	ADP
elements-9012	150	7	27	27	NUM
elements-9012	150	8	species	specie	NOUN
elements-9012	150	9	,	,	PUNCT
elements-9012	150	10	and	and	CCONJ
elements-9012	150	11	we	we	PRON
elements-9012	150	12	have	have	AUX
elements-9012	150	13	calculated	calculate	VERB
elements-9012	150	14	substitution	substitution	NOUN
elements-9012	150	15	rates	rate	NOUN
elements-9012	150	16	between	between	ADP
elements-9012	150	17	26	26	NUM
elements-9012	150	18	species	specie	NOUN
elements-9012	150	19	pairs	pair	NOUN
elements-9012	150	20	,	,	PUNCT
elements-9012	150	21	based	base	VERB
elements-9012	150	22	on	on	ADP
elements-9012	150	23	choosing	choose	VERB
elements-9012	150	24	those	those	PRON
elements-9012	150	25	which	which	PRON
elements-9012	150	26	are	be	AUX
elements-9012	150	27	closely	closely	ADV
elements-9012	150	28	related	relate	VERB
elements-9012	150	29	in	in	ADP
elements-9012	150	30	the	the	DET
elements-9012	150	31	phylogeny	phylogeny	NOUN
elements-9012	150	32	.	.	PUNCT
elements-9012	151	1	in	in	ADP
elements-9012	151	2	genomes	genome	NOUN
elements-9012	151	3	with	with	ADP
elements-9012	151	4	uniform	uniform	ADJ
elements-9012	151	5	rates	rate	NOUN
elements-9012	151	6	,	,	PUNCT
elements-9012	151	7	such	such	ADJ
elements-9012	151	8	as	as	ADP
elements-9012	151	9	s.	s.	PROPN
elements-9012	151	10	cerevisiae	cerevisiae	PROPN
elements-9012	151	11	,	,	PUNCT
elements-9012	151	12	the	the	DET
elements-9012	151	13	distribution	distribution	NOUN
elements-9012	151	14	of	of	ADP
elements-9012	151	15	z	z	NOUN
elements-9012	151	16	-	-	PUNCT
elements-9012	151	17	scores	score	NOUN
elements-9012	151	18	is	be	AUX
elements-9012	151	19	very	very	ADV
elements-9012	151	20	close	close	ADJ
elements-9012	151	21	to	to	ADP
elements-9012	151	22	a	a	DET
elements-9012	151	23	normal	normal	ADJ
elements-9012	151	24	distribution	distribution	NOUN
elements-9012	151	25	with	with	ADP
elements-9012	151	26	a	a	DET
elements-9012	151	27	standard	standard	ADJ
elements-9012	151	28	deviation	deviation	NOUN
elements-9012	151	29	of	of	ADP
elements-9012	151	30	one	one	NUM
elements-9012	151	31	.	.	PUNCT
elements-9012	152	1	x	x	PUNCT
elements-9012	152	2	v	v	X
elements-9012	152	3	m	m	VERB
elements-9012	152	4	however	however	ADV
elements-9012	152	5	,	,	PUNCT
elements-9012	152	6	in	in	ADP
elements-9012	152	7	heterogeneous	heterogeneous	ADJ
elements-9012	152	8	genomes	genome	NOUN
elements-9012	152	9	such	such	ADJ
elements-9012	152	10	as	as	ADP
elements-9012	152	11	mouse	mouse	NOUN
elements-9012	152	12	and	and	CCONJ
elements-9012	152	13	human	human	NOUN
elements-9012	152	14	,	,	PUNCT
elements-9012	152	15	the	the	DET
elements-9012	152	16	width	width	NOUN
elements-9012	152	17	of	of	ADP
elements-9012	152	18	the	the	DET
elements-9012	152	19	z	z	NOUN
elements-9012	152	20	-	-	PUNCT
elements-9012	152	21	score	score	NOUN
elements-9012	152	22	distribution	distribution	NOUN
elements-9012	152	23	is	be	AUX
elements-9012	152	24	considerably	considerably	ADV
elements-9012	152	25	larger	large	ADJ
elements-9012	152	26	,	,	PUNCT
elements-9012	152	27	due	due	ADP
elements-9012	152	28	to	to	ADP
elements-9012	152	29	the	the	DET
elements-9012	152	30	tendency	tendency	NOUN
elements-9012	152	31	of	of	ADP
elements-9012	152	32	sites	site	NOUN
elements-9012	152	33	in	in	ADP
elements-9012	152	34	the	the	DET
elements-9012	152	35	same	same	ADJ
elements-9012	152	36	gene	gene	NOUN
elements-9012	152	37	to	to	PART
elements-9012	152	38	be	be	AUX
elements-9012	152	39	subject	subject	ADJ
elements-9012	152	40	to	to	ADP
elements-9012	152	41	similar	similar	ADJ
elements-9012	152	42	mutational	mutational	ADJ
elements-9012	152	43	pressures	pressure	NOUN
elements-9012	152	44	(	(	PUNCT
elements-9012	152	45	figure	figure	VERB
elements-9012	152	46	ia	ia	PROPN
elements-9012	152	47	)	)	PUNCT
elements-9012	152	48	.	.	PUNCT
elements-9012	153	1	we	we	PRON
elements-9012	153	2	find	find	VERB
elements-9012	153	3	that	that	SCONJ
elements-9012	153	4	the	the	DET
elements-9012	153	5	distributions	distribution	NOUN
elements-9012	153	6	of	of	ADP
elements-9012	153	7	normalized	normalize	VERB
elements-9012	153	8	rates	rate	NOUN
elements-9012	153	9	for	for	ADP
elements-9012	153	10	22	22	NUM
elements-9012	153	11	of	of	ADP
elements-9012	153	12	the	the	DET
elements-9012	153	13	27	27	NUM
elements-9012	153	14	yeast	yeast	NOUN
elements-9012	153	15	species	specie	NOUN
elements-9012	153	16	(	(	PUNCT
elements-9012	153	17	23	23	NUM
elements-9012	153	18	species	species	NOUN
elements-9012	153	19	pairs	pair	NOUN
elements-9012	153	20	)	)	PUNCT
elements-9012	153	21	fit	fit	VERB
elements-9012	153	22	the	the	DET
elements-9012	153	23	normal	normal	ADJ
elements-9012	153	24	distribution	distribution	NOUN
elements-9012	153	25	with	with	ADP
elements-9012	153	26	unit	unit	NOUN
elements-9012	153	27	standard	standard	ADJ
elements-9012	153	28	deviation	deviation	NOUN
elements-9012	153	29	(	(	PUNCT
elements-9012	153	30	average	average	NOUN
elements-9012	153	31	s	s	PART
elements-9012	153	32	=	=	SYM
elements-9012	153	33	1.32	1.32	NUM
elements-9012	153	34	,	,	PUNCT
elements-9012	153	35	range	range	NOUN
elements-9012	153	36	=	=	PUNCT
elements-9012	154	1	[	[	X
elements-9012	154	2	1.02	1.02	NUM
elements-9012	154	3	1.40	1.40	NUM
elements-9012	154	4	]	]	PUNCT
elements-9012	154	5	)	)	PUNCT
elements-9012	154	6	.	.	PUNCT
elements-9012	155	1	this	this	PRON
elements-9012	155	2	indicates	indicate	VERB
elements-9012	155	3	that	that	SCONJ
elements-9012	155	4	there	there	PRON
elements-9012	155	5	are	be	VERB
elements-9012	155	6	generally	generally	ADV
elements-9012	155	7	not	not	PART
elements-9012	155	8	regions	region	NOUN
elements-9012	155	9	of	of	ADP
elements-9012	155	10	high	high	ADJ
elements-9012	155	11	or	or	CCONJ
elements-9012	155	12	low	low	ADJ
elements-9012	155	13	neutral	neutral	ADJ
elements-9012	155	14	mutation	mutation	NOUN
elements-9012	155	15	rates	rate	NOUN
elements-9012	155	16	within	within	ADP
elements-9012	155	17	these	these	DET
elements-9012	155	18	genomes	genome	NOUN
elements-9012	155	19	(	(	PUNCT
elements-9012	155	20	figure	figure	NOUN
elements-9012	155	21	ib	ib	NOUN
elements-9012	155	22	)	)	PUNCT
elements-9012	155	23	.	.	PUNCT
elements-9012	156	1	as	as	ADP
elements-9012	156	2	in	in	ADP
elements-9012	156	3	the	the	DET
elements-9012	156	4	sensu	sensu	NOUN
elements-9012	156	5	stricto	stricto	X
elements-9012	156	6	yeasts	yeast	NOUN
elements-9012	156	7	,	,	PUNCT
elements-9012	156	8	these	these	DET
elements-9012	156	9	22	22	NUM
elements-9012	156	10	yeast	yeast	NOUN
elements-9012	156	11	species	specie	NOUN
elements-9012	156	12	tend	tend	VERB
elements-9012	156	13	to	to	PART
elements-9012	156	14	have	have	VERB
elements-9012	156	15	a	a	DET
elements-9012	156	16	slight	slight	ADJ
elements-9012	156	17	excess	excess	NOUN
elements-9012	156	18	of	of	ADP
elements-9012	156	19	genes	gene	NOUN
elements-9012	156	20	with	with	ADP
elements-9012	156	21	low	low	ADJ
elements-9012	156	22	silent	silent	ADJ
elements-9012	156	23	substitution	substitution	NOUN
elements-9012	156	24	rates	rate	NOUN
elements-9012	156	25	.	.	PUNCT
elements-9012	157	1	this	this	PRON
elements-9012	157	2	can	can	AUX
elements-9012	157	3	be	be	AUX
elements-9012	157	4	explained	explain	VERB
elements-9012	157	5	by	by	ADP
elements-9012	157	6	codon	codon	ADJ
elements-9012	157	7	usage	usage	NOUN
elements-9012	157	8	selection	selection	NOUN
elements-9012	157	9	.	.	PUNCT
elements-9012	158	1	the	the	DET
elements-9012	158	2	tail	tail	NOUN
elements-9012	158	3	at	at	ADP
elements-9012	158	4	negative	negative	ADJ
elements-9012	158	5	z	z	NOUN
elements-9012	158	6	-	-	PUNCT
elements-9012	158	7	scores	score	NOUN
elements-9012	158	8	is	be	AUX
elements-9012	158	9	mainly	mainly	ADV
elements-9012	158	10	comprised	comprise	VERB
elements-9012	158	11	of	of	ADP
elements-9012	158	12	ribosomal	ribosomal	NOUN
elements-9012	158	13	and	and	CCONJ
elements-9012	158	14	carbohydrate	carbohydrate	NOUN
elements-9012	158	15	catabolism	catabolism	NOUN
elements-9012	158	16	genes	gene	NOUN
elements-9012	158	17	that	that	PRON
elements-9012	158	18	are	be	AUX
elements-9012	158	19	under	under	ADP
elements-9012	158	20	selection	selection	NOUN
elements-9012	158	21	for	for	ADP
elements-9012	158	22	codon	codon	ADJ
elements-9012	158	23	usage	usage	NOUN
elements-9012	158	24	bias	bias	NOUN
elements-9012	158	25	.	.	PUNCT
elements-9012	159	1	x	x	X
elements-9012	160	1	l	l	NOUN
elements-9012	160	2	x	x	PUNCT
elements-9012	160	3	of	of	ADP
elements-9012	160	4	the	the	DET
elements-9012	160	5	432	432	NUM
elements-9012	160	6	genes	gene	NOUN
elements-9012	160	7	with	with	ADP
elements-9012	160	8	z	z	NOUN
elements-9012	160	9	-	-	PUNCT
elements-9012	160	10	scores	score	NOUN
elements-9012	160	11	<	<	X
elements-9012	160	12	-2	-2	INTJ
elements-9012	160	13	in	in	ADP
elements-9012	160	14	the	the	DET
elements-9012	160	15	s.	s.	PROPN
elements-9012	160	16	cerevisiae	cerevisiae	PROPN
elements-9012	160	17	s.	s.	PROPN
elements-9012	160	18	bayanus	bayanus	PROPN
elements-9012	160	19	comparison	comparison	NOUN
elements-9012	160	20	,	,	PUNCT
elements-9012	160	21	201	201	NUM
elements-9012	160	22	(	(	PUNCT
elements-9012	160	23	46.5	46.5	NUM
elements-9012	160	24	%	%	NOUN
elements-9012	160	25	)	)	PUNCT
elements-9012	160	26	have	have	VERB
elements-9012	160	27	a	a	DET
elements-9012	160	28	ribosomal	ribosomal	NOUN
elements-9012	160	29	or	or	CCONJ
elements-9012	160	30	metabolism	metabolism	NOUN
elements-9012	160	31	go	go	VERB
elements-9012	160	32	annotation	annotation	NOUN
elements-9012	160	33	.	.	PUNCT
elements-9012	161	1	similarly	similarly	ADV
elements-9012	161	2	,	,	PUNCT
elements-9012	161	3	in	in	ADP
elements-9012	161	4	the	the	DET
elements-9012	161	5	distinct	distinct	ADJ
elements-9012	161	6	lineage	lineage	NOUN
elements-9012	161	7	d.	d.	PROPN
elements-9012	161	8	hansenii	hansenii	PROPN
elements-9012	161	9	c.	c.	PROPN
elements-9012	161	10	guilliermondii	guilliermondii	PROPN
elements-9012	161	11	,	,	PUNCT
elements-9012	161	12	138	138	NUM
elements-9012	161	13	of	of	ADP
elements-9012	161	14	253	253	NUM
elements-9012	161	15	genes	gene	NOUN
elements-9012	161	16	(	(	PUNCT
elements-9012	161	17	54.5	54.5	NUM
elements-9012	161	18	%	%	NOUN
elements-9012	161	19	)	)	PUNCT
elements-9012	161	20	with	with	ADP
elements-9012	161	21	z	z	NOUN
elements-9012	161	22	-	-	PUNCT
elements-9012	161	23	score	score	VERB
elements-9012	161	24	<	<	X
elements-9012	161	25	-2	-2	NOUN
elements-9012	161	26	map	map	NOUN
elements-9012	161	27	to	to	ADP
elements-9012	161	28	r	r	NOUN
elements-9012	161	29	i	i	PRON
elements-9012	161	30	bosomal	bosomal	ADJ
elements-9012	161	31	or	or	CCONJ
elements-9012	161	32	metabolism	metabolism	NOUN
elements-9012	161	33	go	go	VERB
elements-9012	161	34	categories	category	NOUN
elements-9012	161	35	.	.	PUNCT
elements-9012	162	1	one	one	NUM
elements-9012	162	2	caveat	caveat	NOUN
elements-9012	162	3	is	be	AUX
elements-9012	162	4	that	that	SCONJ
elements-9012	162	5	this	this	DET
elements-9012	162	6	z	z	NOUN
elements-9012	162	7	-	-	PUNCT
elements-9012	162	8	score	score	NOUN
elements-9012	162	9	approach	approach	NOUN
elements-9012	162	10	is	be	AUX
elements-9012	162	11	less	less	ADV
elements-9012	162	12	applicable	applicable	ADJ
elements-9012	162	13	for	for	ADP
elements-9012	162	14	comparisons	comparison	NOUN
elements-9012	162	15	of	of	ADP
elements-9012	162	16	species	specie	NOUN
elements-9012	162	17	with	with	ADP
elements-9012	162	18	saturated	saturated	ADJ
elements-9012	162	19	divergence	divergence	NOUN
elements-9012	162	20	.	.	PUNCT
elements-9012	163	1	of	of	ADP
elements-9012	163	2	the	the	DET
elements-9012	163	3	full	full	ADJ
elements-9012	163	4	set	set	NOUN
elements-9012	163	5	of	of	ADP
elements-9012	163	6	26	26	NUM
elements-9012	163	7	species	specie	NOUN
elements-9012	163	8	pairs	pair	NOUN
elements-9012	163	9	,	,	PUNCT
elements-9012	163	10	we	we	PRON
elements-9012	163	11	observe	observe	VERB
elements-9012	163	12	8	8	NUM
elements-9012	163	13	with	with	ADP
elements-9012	163	14	less	less	ADJ
elements-9012	163	15	than	than	ADP
elements-9012	163	16	90	90	NUM
elements-9012	163	17	%	%	NOUN
elements-9012	163	18	of	of	ADP
elements-9012	163	19	the	the	DET
elements-9012	163	20	divergence	divergence	NOUN
elements-9012	163	21	that	that	PRON
elements-9012	163	22	would	would	AUX
elements-9012	163	23	be	be	AUX
elements-9012	163	24	expected	expect	VERB
elements-9012	163	25	at	at	ADP
elements-9012	163	26	saturation	saturation	NOUN
elements-9012	163	27	,	,	PUNCT
elements-9012	163	28	given	give	VERB
elements-9012	163	29	the	the	DET
elements-9012	163	30	base	base	NOUN
elements-9012	163	31	this	this	PRON
elements-9012	163	32	is	be	AUX
elements-9012	163	33	the	the	DET
elements-9012	163	34	distribution	distribution	NOUN
elements-9012	163	35	of	of	ADP
elements-9012	163	36	normalized	normalize	VERB
elements-9012	163	37	4	4	NUM
elements-9012	163	38	-	-	ADJ
elements-9012	163	39	fold	fold	ADJ
elements-9012	163	40	substitution	substitution	NOUN
elements-9012	163	41	rates	rate	NOUN
elements-9012	163	42	for	for	ADP
elements-9012	163	43	mammals	mammal	NOUN
elements-9012	163	44	and	and	CCONJ
elements-9012	163	45	yeasts	yeast	NOUN
elements-9012	163	46	.	.	PUNCT
elements-9012	164	1	the	the	DET
elements-9012	164	2	normal	normal	ADJ
elements-9012	164	3	gaussian	gaussian	ADJ
elements-9012	164	4	distribution	distribution	NOUN
elements-9012	164	5	(	(	PUNCT
elements-9012	164	6	smooth	smooth	ADJ
elements-9012	164	7	cray	cray	NOUN
elements-9012	164	8	line	line	NOUN
elements-9012	164	9	)	)	PUNCT
elements-9012	164	10	is	be	AUX
elements-9012	164	11	what	what	PRON
elements-9012	164	12	would	would	AUX
elements-9012	164	13	be	be	AUX
elements-9012	164	14	expected	expect	VERB
elements-9012	164	15	if	if	SCONJ
elements-9012	164	16	all	all	DET
elements-9012	164	17	4	4	ADJ
elements-9012	164	18	-	-	ADJ
elements-9012	164	19	fold	fold	ADJ
elements-9012	164	20	sites	site	NOUN
elements-9012	164	21	in	in	ADP
elements-9012	164	22	each	each	DET
elements-9012	164	23	cene	cene	NOUN
elements-9012	164	24	have	have	VERB
elements-9012	164	25	an	an	DET
elements-9012	164	26	equal	equal	ADJ
elements-9012	164	27	and	and	CCONJ
elements-9012	164	28	independent	independent	ADJ
elements-9012	164	29	probability	probability	NOUN
elements-9012	164	30	of	of	ADP
elements-9012	164	31	being	be	AUX
elements-9012	164	32	substituted	substitute	VERB
elements-9012	164	33	.	.	PUNCT
elements-9012	165	1	top	top	ADJ
elements-9012	165	2	:	:	PUNCT
elements-9012	165	3	all	all	DET
elements-9012	165	4	mammalian	mammalian	ADJ
elements-9012	165	5	distributions	distribution	NOUN
elements-9012	165	6	have	have	VERB
elements-9012	165	7	a	a	DET
elements-9012	165	8	bias	bias	NOUN
elements-9012	165	9	toward	toward	ADP
elements-9012	165	10	high	high	ADJ
elements-9012	165	11	and	and	CCONJ
elements-9012	165	12	low	low	ADJ
elements-9012	165	13	substitution	substitution	NOUN
elements-9012	165	14	rates	rate	NOUN
elements-9012	165	15	,	,	PUNCT
elements-9012	165	16	consistent	consistent	ADJ
elements-9012	165	17	with	with	ADP
elements-9012	165	18	regional	regional	ADJ
elements-9012	165	19	mutation	mutation	NOUN
elements-9012	165	20	biases	bias	NOUN
elements-9012	165	21	in	in	ADP
elements-9012	165	22	mammalian	mammalian	ADJ
elements-9012	165	23	genomes	genome	NOUN
elements-9012	165	24	.	.	PUNCT
elements-9012	166	1	middle	middle	ADJ
elements-9012	166	2	:	:	PUNCT
elements-9012	166	3	most	most	ADJ
elements-9012	166	4	yeast	yeast	NOUN
elements-9012	166	5	distributions	distribution	NOUN
elements-9012	166	6	fit	fit	ADJ
elements-9012	166	7	more	more	ADV
elements-9012	166	8	closely	closely	ADV
elements-9012	166	9	to	to	ADP
elements-9012	166	10	the	the	DET
elements-9012	166	11	gaussian	gaussian	NOUN
elements-9012	166	12	,	,	PUNCT
elements-9012	166	13	except	except	SCONJ
elements-9012	166	14	for	for	ADP
elements-9012	166	15	a	a	DET
elements-9012	166	16	tail	tail	NOUN
elements-9012	166	17	of	of	ADP
elements-9012	166	18	genes	gene	NOUN
elements-9012	166	19	with	with	ADP
elements-9012	166	20	low	low	ADJ
elements-9012	166	21	substitution	substitution	NOUN
elements-9012	166	22	rates	rate	NOUN
elements-9012	166	23	stemming	stem	VERB
elements-9012	166	24	for	for	ADP
elements-9012	166	25	codon	codon	ADJ
elements-9012	166	26	usage	usage	NOUN
elements-9012	166	27	selection	selection	NOUN
elements-9012	166	28	.	.	PUNCT
elements-9012	167	1	bottom	bottom	NOUN
elements-9012	167	2	:	:	PUNCT
elements-9012	168	1	this	this	PRON
elements-9012	168	2	i	i	PRON
elements-9012	168	3	s	s	VERB
elements-9012	168	4	the	the	DET
elements-9012	168	5	distribution	distribution	NOUN
elements-9012	168	6	of	of	ADP
elements-9012	168	7	c.	c.	NOUN
elements-9012	168	8	albicans	albican	NOUN
elements-9012	168	9	,	,	PUNCT
elements-9012	168	10	c.	c.	NOUN
elements-9012	168	11	dubliensis	dubliensis	PROPN
elements-9012	168	12	,	,	PUNCT
elements-9012	168	13	c.	c.	PROPN
elements-9012	168	14	tropicalis	tropicalis	PROPN
elements-9012	168	15	,	,	PUNCT
elements-9012	168	16	n.	n.	NOUN
elements-9012	168	17	crassa	crassa	PROPN
elements-9012	168	18	,	,	PUNCT
elements-9012	168	19	a	a	DET
elements-9012	168	20	n	n	NOUN
elements-9012	168	21	d	d	PROPN
elements-9012	168	22	c.	c.	PROPN
elements-9012	168	23	clobosum	clobosum	PROPN
elements-9012	168	24	.	.	PUNCT
elements-9012	169	1	these	these	DET
elements-9012	169	2	species	specie	NOUN
elements-9012	169	3	have	have	VERB
elements-9012	169	4	a	a	DET
elements-9012	169	5	wider	wide	ADJ
elements-9012	169	6	distribution	distribution	NOUN
elements-9012	169	7	similar	similar	ADJ
elements-9012	169	8	to	to	ADP
elements-9012	169	9	the	the	DET
elements-9012	169	10	mammalian	mammalian	ADJ
elements-9012	169	11	distribution	distribution	NOUN
elements-9012	169	12	.	.	PUNCT
elements-9012	170	1	m	m	PROPN
elements-9012	170	2	e	e	X
elements-9012	170	3	a	a	PRON
elements-9012	170	4	s	s	X
elements-9012	170	5	u	u	NOUN
elements-9012	170	6	r	r	NOUN
elements-9012	170	7	i	i	PRON
elements-9012	170	8	n	n	CCONJ
elements-9012	170	9	g	g	PROPN
elements-9012	170	10	t	t	NOUN
elements-9012	170	11	h	h	NOUN
elements-9012	170	12	e	e	X
elements-9012	170	13	u	u	NOUN
elements-9012	170	14	n	n	PROPN
elements-9012	170	15	i	i	PRON
elements-9012	170	16	v	v	VERB
elements-9012	170	17	e	e	NOUN
elements-9012	170	18	r	r	NOUN
elements-9012	170	19	s	s	PROPN
elements-9012	170	20	a	a	DET
elements-9012	170	21	l	l	NOUN
elements-9012	171	1	i	i	PRON
elements-9012	171	2	t	t	VERB
elements-9012	172	1	y	y	NOUN
elements-9012	173	1	o	o	X
elements-9012	173	2	f	f	NOUN
elements-9012	174	1	r	r	NOUN
elements-9012	174	2	e	e	NOUN
elements-9012	174	3	g	g	NOUN
elements-9012	175	1	i	i	PRON
elements-9012	175	2	o	o	VERB
elements-9012	176	1	n	n	CCONJ
elements-9012	176	2	a	a	DET
elements-9012	176	3	l	l	NOUN
elements-9012	176	4	m	m	VERB
elements-9012	177	1	u	u	NOUN
elements-9012	177	2	t	t	PROPN
elements-9012	177	3	a	a	DET
elements-9012	177	4	t	t	NOUN
elements-9012	178	1	i	i	PRON
elements-9012	178	2	o	o	VERB
elements-9012	179	1	n	n	CCONJ
elements-9012	179	2	r	r	NOUN
elements-9012	179	3	a	a	DET
elements-9012	179	4	t	t	NOUN
elements-9012	180	1	e	e	NOUN
elements-9012	180	2	s	s	PROPN
elements-9012	180	3	t	t	PROPN
elements-9012	180	4	a	a	DET
elements-9012	180	5	b	b	NOUN
elements-9012	180	6	l	l	NOUN
elements-9012	180	7	e	e	NOUN
elements-9012	180	8	1	1	NUM
elements-9012	180	9	:	:	PUNCT
elements-9012	180	10	p	p	X
elements-9012	180	11	e	e	X
elements-9012	180	12	a	a	DET
elements-9012	180	13	r	r	NOUN
elements-9012	180	14	s	s	NOUN
elements-9012	180	15	o	o	NOUN
elements-9012	180	16	n	n	ADP
elements-9012	180	17	c	c	NOUN
elements-9012	181	1	o	o	NOUN
elements-9012	181	2	r	r	NOUN
elements-9012	181	3	r	r	NOUN
elements-9012	181	4	e	e	NOUN
elements-9012	181	5	l	l	NOUN
elements-9012	181	6	a	a	NOUN
elements-9012	181	7	t	t	NOUN
elements-9012	182	1	i	i	PRON
elements-9012	182	2	o	o	VERB
elements-9012	183	1	n	n	NOUN
elements-9012	184	1	f	f	NOUN
elements-9012	184	2	o	o	NOUN
elements-9012	184	3	r	r	NOUN
elements-9012	184	4	t	t	NOUN
elements-9012	184	5	h	h	NOUN
elements-9012	184	6	e	e	X
elements-9012	184	7	s	s	PROPN
elements-9012	184	8	u	u	PROPN
elements-9012	184	9	b	b	PROPN
elements-9012	184	10	s	s	ADP
elements-9012	184	11	t	t	X
elements-9012	185	1	i	i	PRON
elements-9012	185	2	t	t	VERB
elements-9012	185	3	u	u	NOUN
elements-9012	186	1	t	t	NOUN
elements-9012	187	1	i	i	PRON
elements-9012	187	2	o	o	VERB
elements-9012	188	1	n	n	CCONJ
elements-9012	188	2	r	r	NOUN
elements-9012	188	3	a	a	DET
elements-9012	188	4	t	t	NOUN
elements-9012	188	5	e	e	X
elements-9012	188	6	o	o	NOUN
elements-9012	188	7	f	f	VERB
elements-9012	188	8	n	n	CCONJ
elements-9012	188	9	e	e	NOUN
elements-9012	188	10	i	i	PRON
elements-9012	188	11	g	g	PROPN
elements-9012	188	12	h	h	NOUN
elements-9012	189	1	b	b	NOUN
elements-9012	189	2	o	o	NOUN
elements-9012	190	1	r	r	VERB
elements-9012	191	1	i	i	PRON
elements-9012	192	1	n	n	CCONJ
elements-9012	193	1	g	g	NOUN
elements-9012	193	2	g	g	PROPN
elements-9012	193	3	e	e	PROPN
elements-9012	193	4	n	n	ADP
elements-9012	193	5	e	e	NOUN
elements-9012	193	6	s	s	PROPN
elements-9012	193	7	a	a	PRON
elements-9012	193	8	v	v	NOUN
elements-9012	193	9	e	e	NOUN
elements-9012	193	10	r	r	NOUN
elements-9012	193	11	a	a	DET
elements-9012	193	12	g	g	NOUN
elements-9012	193	13	e	e	PROPN
elements-9012	193	14	n	n	ADP
elements-9012	193	15	u	u	NOUN
elements-9012	193	16	m	m	NOUN
elements-9012	193	17	b	b	PROPN
elements-9012	193	18	e	e	NOUN
elements-9012	193	19	r	r	NOUN
elements-9012	193	20	o	o	X
elements-9012	193	21	f	f	X
elements-9012	193	22	p	p	X
elements-9012	193	23	e	e	PROPN
elements-9012	193	24	a	a	PRON
elements-9012	193	25	r	r	NOUN
elements-9012	193	26	s	s	NOUN
elements-9012	193	27	o	o	NOUN
elements-9012	193	28	n	n	X
elements-9012	193	29	s	s	PROPN
elements-9012	193	30	p	p	X
elements-9012	193	31	e	e	NOUN
elements-9012	193	32	c	c	NOUN
elements-9012	193	33	i	i	NOUN
elements-9012	193	34	e	e	NOUN
elements-9012	193	35	s	s	VERB
elements-9012	193	36	1	1	NUM
elements-9012	193	37	s	s	PART
elements-9012	193	38	p	p	NOUN
elements-9012	193	39	e	e	NOUN
elements-9012	193	40	c	c	NOUN
elements-9012	193	41	i	i	NOUN
elements-9012	193	42	e	e	NOUN
elements-9012	193	43	s	s	PROPN
elements-9012	193	44	2	2	NUM
elements-9012	193	45	d	d	NOUN
elements-9012	193	46	i	i	PRON
elements-9012	193	47	v	v	VERB
elements-9012	193	48	e	e	NOUN
elements-9012	193	49	r	r	NOUN
elements-9012	193	50	g	g	NOUN
elements-9012	193	51	e	e	NOUN
elements-9012	193	52	n	n	NOUN
elements-9012	193	53	c	c	NOUN
elements-9012	193	54	e	e	NOUN
elements-9012	193	55	o	o	NOUN
elements-9012	193	56	r	r	NOUN
elements-9012	193	57	t	t	NOUN
elements-9012	194	1	h	h	NOUN
elements-9012	195	1	o	o	NOUN
elements-9012	195	2	l	l	NOUN
elements-9012	196	1	o	o	X
elements-9012	196	2	g	g	NOUN
elements-9012	196	3	s	s	X
elements-9012	196	4	c	c	NOUN
elements-9012	196	5	o	o	NOUN
elements-9012	196	6	r	r	NOUN
elements-9012	196	7	r	r	NOUN
elements-9012	196	8	e	e	NOUN
elements-9012	196	9	l	l	NOUN
elements-9012	196	10	a	a	NOUN
elements-9012	196	11	t	t	NOUN
elements-9012	197	1	i	i	PRON
elements-9012	197	2	o	o	VERB
elements-9012	198	1	n	n	CCONJ
elements-9012	198	2	p	p	X
elements-9012	198	3	v	v	ADP
elements-9012	198	4	a	a	DET
elements-9012	198	5	l	l	NOUN
elements-9012	198	6	s_cerevisiae	s_cerevisiae	X
elements-9012	198	7	s_paradoxus	s_paradoxus	X
elements-9012	198	8	0	0	NUM
elements-9012	198	9	.	.	PUNCT
elements-9012	199	1	2	2	NUM
elements-9012	199	2	5	5	NUM
elements-9012	199	3	9	9	NUM
elements-9012	199	4	2	2	NUM
elements-9012	199	5	4	4	NUM
elements-9012	199	6	4	4	NUM
elements-9012	199	7	3	3	NUM
elements-9012	199	8	4	4	NUM
elements-9012	199	9	0	0	NUM
elements-9012	199	10	.	.	PUNCT
elements-9012	199	11	0	0	NUM
elements-9012	200	1	2	2	NUM
elements-9012	200	2	3	3	NUM
elements-9012	200	3	4	4	NUM
elements-9012	200	4	0	0	NUM
elements-9012	200	5	.	.	PUNCT
elements-9012	201	1	1	1	NUM
elements-9012	201	2	1	1	NUM
elements-9012	201	3	8	8	NUM
elements-9012	201	4	8	8	NUM
elements-9012	201	5	the	the	DET
elements-9012	201	6	data	datum	NOUN
elements-9012	201	7	collected	collect	VERB
elements-9012	201	8	was	be	AUX
elements-9012	201	9	from	from	ADP
elements-9012	201	10	s_cerevisiae	s_cerevisiae	PROPN
elements-9012	201	11	s_bayan	s_bayan	NOUN
elements-9012	201	12	yus	yus	NOUN
elements-9012	201	13	0	0	NUM
elements-9012	201	14	.	.	PUNCT
elements-9012	202	1	4	4	NUM
elements-9012	202	2	7	7	NUM
elements-9012	202	3	0	0	NUM
elements-9012	202	4	9	9	NUM
elements-9012	202	5	3781	3781	NUM
elements-9012	202	6	0	0	NUM
elements-9012	203	1	.0305	.0305	NOUN
elements-9012	203	2	0	0	NUM
elements-9012	203	3	.	.	PUNCT
elements-9012	203	4	0	0	NUM
elements-9012	204	1	6	6	NUM
elements-9012	204	2	0	0	NUM
elements-9012	204	3	6	6	NUM
elements-9012	204	4	2	2	NUM
elements-9012	204	5	6	6	NUM
elements-9012	204	6	species	specie	NOUN
elements-9012	204	7	pairs	pair	NOUN
elements-9012	204	8	a	a	DET
elements-9012	204	9	m	m	NOUN
elements-9012	204	10	o	o	NOUN
elements-9012	204	11	n	n	CCONJ
elements-9012	204	12	g	g	NOUN
elements-9012	204	13	2	2	NUM
elements-9012	204	14	7	7	NUM
elements-9012	204	15	s_castellii	s_castellii	NOUN
elements-9012	204	16	s_m	s_m	PROPN
elements-9012	204	17	1	1	NUM
elements-9012	204	18	katae	katae	NOUN
elements-9012	204	19	0	0	NUM
elements-9012	204	20	.	.	PUNCT
elements-9012	205	1	6	6	NUM
elements-9012	205	2	5	5	NUM
elements-9012	205	3	8	8	NUM
elements-9012	205	4	5	5	NUM
elements-9012	205	5	3	3	NUM
elements-9012	205	6	2	2	NUM
elements-9012	205	7	4	4	NUM
elements-9012	205	8	7	7	NUM
elements-9012	205	9	0	0	NUM
elements-9012	205	10	.	.	PUNCT
elements-9012	206	1	0	0	NUM
elements-9012	206	2	0	0	NUM
elements-9012	206	3	2	2	NUM
elements-9012	206	4	9	9	NUM
elements-9012	206	5	0	0	NUM
elements-9012	206	6	.	.	PUNCT
elements-9012	206	7	8	8	NUM
elements-9012	206	8	6	6	NUM
elements-9012	206	9	4	4	NUM
elements-9012	206	10	9	9	NUM
elements-9012	206	11	yeasts	yeast	NOUN
elements-9012	206	12	,	,	PUNCT
elements-9012	206	13	average	average	ADJ
elements-9012	206	14	divergence	divergence	NOUN
elements-9012	206	15	k_lactis	k_lactis	X
elements-9012	206	16	c_glabrata	c_glabrata	NOUN
elements-9012	206	17	0	0	NUM
elements-9012	206	18	.	.	PUNCT
elements-9012	207	1	6	6	NUM
elements-9012	207	2	5	5	NUM
elements-9012	207	3	4	4	NUM
elements-9012	207	4	5	5	NUM
elements-9012	207	5	4	4	NUM
elements-9012	207	6	0	0	NUM
elements-9012	207	7	7	7	NUM
elements-9012	207	8	7	7	NUM
elements-9012	207	9	-0	-0	PUNCT
elements-9012	207	10	.0135	.0135	INTJ
elements-9012	207	11	0	0	NUM
elements-9012	207	12	.	.	PUNCT
elements-9012	208	1	3	3	NUM
elements-9012	208	2	8	8	NUM
elements-9012	208	3	8	8	NUM
elements-9012	208	4	7	7	NUM
elements-9012	208	5	is	be	AUX
elements-9012	208	6	the	the	DET
elements-9012	208	7	fraction	fraction	NOUN
elements-9012	208	8	of	of	ADP
elements-9012	208	9	all	all	PRON
elements-9012	208	10	d_han	d_han	PRON
elements-9012	208	11	s	s	X
elements-9012	208	12	en	en	PROPN
elements-9012	208	13	1	1	NUM
elements-9012	208	14	t	t	NOUN
elements-9012	208	15	c_lusitaniae	c_lusitaniae	NOUN
elements-9012	208	16	0	0	NUM
elements-9012	208	17	.	.	PUNCT
elements-9012	209	1	6	6	NUM
elements-9012	209	2	9	9	NUM
elements-9012	209	3	2	2	NUM
elements-9012	209	4	6	6	NUM
elements-9012	209	5	3	3	NUM
elements-9012	209	6	4	4	NUM
elements-9012	209	7	2	2	NUM
elements-9012	209	8	5	5	NUM
elements-9012	209	9	0	0	NUM
elements-9012	209	10	.041	.041	NUM
elements-9012	209	11	7	7	NUM
elements-9012	209	12	0	0	NUM
elements-9012	209	13	.	.	PUNCT
elements-9012	209	14	0	0	NUM
elements-9012	209	15	1	1	NUM
elements-9012	209	16	4	4	NUM
elements-9012	209	17	4	4	NUM
elements-9012	209	18	al	al	PROPN
elements-9012	209	19	igned	igned	ADJ
elements-9012	209	20	4	4	NUM
elements-9012	209	21	-	-	ADJ
elements-9012	209	22	fold	fold	ADJ
elements-9012	209	23	sites	site	NOUN
elements-9012	209	24	w	w	NOUN
elements-9012	209	25	h	h	NOUN
elements-9012	210	1	i	i	NOUN
elements-9012	210	2	c	c	NOUN
elements-9012	210	3	h	h	PRON
elements-9012	210	4	d_hansenii	d_hansenii	NOUN
elements-9012	210	5	c_guilliermondii	c_guilliermondii	VERB
elements-9012	210	6	0	0	PUNCT
elements-9012	210	7	.	.	PUNCT
elements-9012	211	1	6	6	NUM
elements-9012	211	2	8	8	NUM
elements-9012	211	3	4	4	NUM
elements-9012	211	4	2	2	NUM
elements-9012	211	5	3	3	NUM
elements-9012	211	6	5	5	NUM
elements-9012	211	7	3	3	NUM
elements-9012	211	8	8	8	NUM
elements-9012	211	9	0	0	NUM
elements-9012	211	10	.0221	.0221	PROPN
elements-9012	211	11	0	0	PUNCT
elements-9012	211	12	.	.	PUNCT
elements-9012	212	1	1	1	NUM
elements-9012	212	2	8	8	NUM
elements-9012	212	3	8	8	NUM
elements-9012	212	4	3	3	NUM
elements-9012	212	5	differ	differ	VERB
elements-9012	212	6	between	between	ADP
elements-9012	212	7	the	the	DET
elements-9012	212	8	two	two	NUM
elements-9012	212	9	d_han	d_han	PROPN
elements-9012	212	10	s	s	PROPN
elements-9012	212	11	e	e	NOUN
elements-9012	212	12	n	n	CCONJ
elements-9012	212	13	11	11	NUM
elements-9012	212	14	l_elongisporus	l_elongisporus	NOUN
elements-9012	212	15	0	0	NUM
elements-9012	212	16	.	.	PUNCT
elements-9012	213	1	6	6	NUM
elements-9012	213	2	6	6	NUM
elements-9012	213	3	2	2	NUM
elements-9012	213	4	8	8	NUM
elements-9012	213	5	3	3	NUM
elements-9012	213	6	3	3	NUM
elements-9012	213	7	4	4	NUM
elements-9012	213	8	5	5	NUM
elements-9012	213	9	0	0	NUM
elements-9012	213	10	.0083	.0083	NOUN
elements-9012	213	11	0	0	PUNCT
elements-9012	213	12	.	.	PUNCT
elements-9012	214	1	6	6	NUM
elements-9012	214	2	2	2	NUM
elements-9012	214	3	7	7	NUM
elements-9012	214	4	5	5	NUM
elements-9012	214	5	species	specie	NOUN
elements-9012	214	6	.	.	PUNCT
elements-9012	215	1	pearson	pearson	NOUN
elements-9012	215	2	d_hansenii	d_hansenii	NOUN
elements-9012	215	3	c_parapsilosis	c_parapsilosis	NOUN
elements-9012	215	4	0	0	NUM
elements-9012	215	5	.6561	.6561	NOUN
elements-9012	215	6	3	3	NUM
elements-9012	215	7	3	3	NUM
elements-9012	215	8	2	2	NUM
elements-9012	215	9	4	4	NUM
elements-9012	215	10	0	0	NUM
elements-9012	215	11	.	.	PUNCT
elements-9012	216	1	0	0	NUM
elements-9012	217	1	2	2	NUM
elements-9012	217	2	5	5	NUM
elements-9012	217	3	7	7	NUM
elements-9012	217	4	0	0	NUM
elements-9012	217	5	.	.	PUNCT
elements-9012	218	1	1	1	NUM
elements-9012	218	2	3	3	NUM
elements-9012	218	3	7	7	NUM
elements-9012	218	4	8	8	NUM
elements-9012	218	5	correlations	correlation	NOUN
elements-9012	218	6	forthe	forthe	VERB
elements-9012	218	7	nord_han	nord_han	PUNCT
elements-9012	218	8	s	s	X
elements-9012	218	9	en	en	X
elements-9012	218	10	11	11	NUM
elements-9012	218	11	c_tropicalis	c_tropicali	NOUN
elements-9012	218	12	0	0	NUM
elements-9012	218	13	.	.	PUNCT
elements-9012	219	1	6	6	NUM
elements-9012	219	2	1	1	NUM
elements-9012	219	3	0	0	NUM
elements-9012	219	4	4	4	NUM
elements-9012	219	5	3	3	NUM
elements-9012	219	6	3	3	NUM
elements-9012	219	7	5	5	NUM
elements-9012	219	8	8	8	NUM
elements-9012	219	9	0	0	NUM
elements-9012	219	10	.0073	.0073	NOUN
elements-9012	219	11	0	0	PUNCT
elements-9012	219	12	.6685	.6685	NOUN
elements-9012	219	13	malized	malize	VERB
elements-9012	219	14	substitution	substitution	NOUN
elements-9012	219	15	rates	rate	NOUN
elements-9012	219	16	e_gossypii	e_gossypii	VERB
elements-9012	219	17	s_pombe	s_pombe	ADJ
elements-9012	219	18	0	0	NUM
elements-9012	219	19	.	.	PUNCT
elements-9012	220	1	7	7	NUM
elements-9012	220	2	6	6	NUM
elements-9012	220	3	1	1	NUM
elements-9012	220	4	6	6	NUM
elements-9012	220	5	1	1	NUM
elements-9012	220	6	5	5	NUM
elements-9012	220	7	3	3	NUM
elements-9012	220	8	6	6	NUM
elements-9012	220	9	0	0	NUM
elements-9012	220	10	.	.	PUNCT
elements-9012	220	11	0	0	NUM
elements-9012	220	12	8	8	NUM
elements-9012	220	13	3	3	NUM
elements-9012	220	14	7	7	NUM
elements-9012	220	15	0	0	NUM
elements-9012	220	16	.	.	PUNCT
elements-9012	220	17	0	0	NUM
elements-9012	220	18	0	0	NUM
elements-9012	220	19	1	1	NUM
elements-9012	220	20	0	0	NUM
elements-9012	220	21	of	of	ADP
elements-9012	220	22	neighboring	neighboring	NOUN
elements-9012	220	23	genes	gene	NOUN
elements-9012	220	24	are	be	AUX
elements-9012	220	25	e_cossypii	e_cossypii	NOUN
elements-9012	220	26	y_lipolytica	y_lipolytica	NOUN
elements-9012	220	27	0	0	NUM
elements-9012	220	28	.	.	PUNCT
elements-9012	221	1	7	7	NUM
elements-9012	221	2	1	1	NUM
elements-9012	221	3	1	1	NUM
elements-9012	221	4	4	4	NUM
elements-9012	221	5	2	2	NUM
elements-9012	221	6	4	4	NUM
elements-9012	221	7	4	4	NUM
elements-9012	221	8	1	1	NUM
elements-9012	221	9	0	0	NUM
elements-9012	221	10	.	.	PUNCT
elements-9012	221	11	0	0	NUM
elements-9012	221	12	2	2	NUM
elements-9012	221	13	9	9	NUM
elements-9012	221	14	7	7	NUM
elements-9012	221	15	0	0	NUM
elements-9012	221	16	.	.	PUNCT
elements-9012	221	17	1	1	NUM
elements-9012	221	18	4	4	NUM
elements-9012	221	19	1	1	NUM
elements-9012	221	20	4	4	NUM
elements-9012	221	21	shown	show	VERB
elements-9012	221	22	in	in	ADP
elements-9012	221	23	column	column	NOUN
elements-9012	221	24	5	5	NUM
elements-9012	221	25	.	.	PUNCT
elements-9012	222	1	the	the	DET
elements-9012	222	2	y_lipolytica	y_lipolytica	NOUN
elements-9012	222	3	m_grisea	m_grisea	PROPN
elements-9012	222	4	0	0	PUNCT
elements-9012	222	5	.6751	.6751	PROPN
elements-9012	222	6	2	2	NUM
elements-9012	222	7	4	4	NUM
elements-9012	222	8	6	6	NUM
elements-9012	222	9	4	4	NUM
elements-9012	222	10	0	0	NUM
elements-9012	222	11	.	.	PUNCT
elements-9012	223	1	0	0	NUM
elements-9012	224	1	4	4	NUM
elements-9012	224	2	1	1	NUM
elements-9012	224	3	0	0	NUM
elements-9012	224	4	0	0	NUM
elements-9012	224	5	.	.	PUNCT
elements-9012	225	1	0	0	NUM
elements-9012	225	2	4	4	NUM
elements-9012	225	3	1	1	NUM
elements-9012	225	4	8	8	NUM
elements-9012	225	5	only	only	ADJ
elements-9012	225	6	species	specie	NOUN
elements-9012	225	7	with	with	ADP
elements-9012	225	8	pearson	pearson	PROPN
elements-9012	225	9	y_lipolytica	y_lipolytica	PROPN
elements-9012	225	10	a_n1du	a_n1du	AUX
elements-9012	225	11	lans	lans	PROPN
elements-9012	225	12	0	0	NUM
elements-9012	225	13	.	.	PUNCT
elements-9012	226	1	7	7	NUM
elements-9012	226	2	0	0	NUM
elements-9012	226	3	7	7	NUM
elements-9012	226	4	2	2	NUM
elements-9012	226	5	2	2	NUM
elements-9012	226	6	5	5	NUM
elements-9012	226	7	1	1	NUM
elements-9012	226	8	5	5	NUM
elements-9012	226	9	0	0	NUM
elements-9012	226	10	.	.	PUNCT
elements-9012	227	1	0	0	NUM
elements-9012	228	1	1	1	NUM
elements-9012	228	2	6	6	NUM
elements-9012	228	3	2	2	NUM
elements-9012	228	4	0	0	NUM
elements-9012	228	5	.	.	PUNCT
elements-9012	229	1	4	4	NUM
elements-9012	229	2	1	1	NUM
elements-9012	229	3	5	5	NUM
elements-9012	229	4	6	6	NUM
elements-9012	229	5	correlation	correlation	NOUN
elements-9012	229	6	with	with	ADP
elements-9012	229	7	y_lipolytica	y_lipolytica	PROPN
elements-9012	229	8	c_immitis	c_immitis	PROPN
elements-9012	229	9	0	0	NUM
elements-9012	229	10	.	.	PUNCT
elements-9012	230	1	7	7	NUM
elements-9012	230	2	1	1	NUM
elements-9012	230	3	0	0	NUM
elements-9012	230	4	9	9	NUM
elements-9012	230	5	2	2	NUM
elements-9012	230	6	5	5	NUM
elements-9012	230	7	8	8	NUM
elements-9012	230	8	4	4	NUM
elements-9012	230	9	0	0	NUM
elements-9012	230	10	.0021	.0021	NOUN
elements-9012	230	11	0	0	NUM
elements-9012	230	12	.	.	PUNCT
elements-9012	230	13	9	9	NUM
elements-9012	230	14	1	1	NUM
elements-9012	230	15	1	1	NUM
elements-9012	230	16	7	7	NUM
elements-9012	230	17	significance	significance	NOUN
elements-9012	230	18	<	<	X
elements-9012	230	19	0	0	NUM
elements-9012	230	20	.001	.001	NUM
elements-9012	230	21	are	be	AUX
elements-9012	230	22	y_lipolytica	y_lipolytica	NOUN
elements-9012	230	23	s_pombe	s_pombe	ADJ
elements-9012	230	24	0	0	PUNCT
elements-9012	231	1	.7271	.7271	SYM
elements-9012	231	2	1	1	NUM
elements-9012	231	3	2	2	NUM
elements-9012	231	4	7	7	NUM
elements-9012	231	5	0	0	NUM
elements-9012	231	6	0	0	NUM
elements-9012	231	7	.	.	PUNCT
elements-9012	232	1	0	0	NUM
elements-9012	232	2	4	4	NUM
elements-9012	232	3	0	0	NUM
elements-9012	232	4	4	4	NUM
elements-9012	232	5	0	0	NUM
elements-9012	232	6	.1495	.1495	NOUN
elements-9012	232	7	those	those	PRON
elements-9012	232	8	among	among	ADP
elements-9012	232	9	c.	c.	PROPN
elements-9012	232	10	0	0	PROPN
elements-9012	232	11	.6551	.6551	PUNCT
elements-9012	232	12	1461	1461	NUM
elements-9012	232	13	0	0	NUM
elements-9012	232	14	.	.	PUNCT
elements-9012	232	15	0	0	NUM
elements-9012	232	16	5	5	NUM
elements-9012	232	17	4	4	NUM
elements-9012	232	18	9	9	NUM
elements-9012	232	19	0	0	NUM
elements-9012	232	20	.	.	PUNCT
elements-9012	232	21	0	0	NUM
elements-9012	232	22	3	3	NUM
elements-9012	232	23	5	5	NUM
elements-9012	232	24	8	8	NUM
elements-9012	232	25	dubliniensis	dubliniensis	NOUN
elements-9012	232	26	,	,	PUNCT
elements-9012	232	27	c.	c.	PROPN
elements-9012	232	28	albican	albican	PROPN
elements-9012	232	29	,	,	PUNCT
elements-9012	232	30	and	and	CCONJ
elements-9012	232	31	s_pombe	s_pombe	ADJ
elements-9012	232	32	sjaponicus	sjaponicus	NOUN
elements-9012	232	33	0	0	NUM
elements-9012	232	34	.6551	.6551	PUNCT
elements-9012	232	35	1461	1461	NUM
elements-9012	232	36	0	0	NUM
elements-9012	232	37	.	.	PUNCT
elements-9012	232	38	0	0	NUM
elements-9012	232	39	5	5	NUM
elements-9012	232	40	4	4	NUM
elements-9012	232	41	9	9	NUM
elements-9012	232	42	0	0	NUM
elements-9012	232	43	.	.	PUNCT
elements-9012	232	44	0	0	NUM
elements-9012	232	45	3	3	NUM
elements-9012	232	46	5	5	NUM
elements-9012	232	47	8	8	NUM
elements-9012	232	48	dubliniensis	dubliniensis	NOUN
elements-9012	232	49	,	,	PUNCT
elements-9012	232	50	c.	c.	PROPN
elements-9012	232	51	albican	albican	PROPN
elements-9012	232	52	,	,	PUNCT
elements-9012	232	53	and	and	CCONJ
elements-9012	232	54	sjaponicus	sjaponicus	VERB
elements-9012	232	55	0	0	NUM
elements-9012	232	56	.2893	.2893	PROPN
elements-9012	232	57	3	3	NUM
elements-9012	232	58	7	7	NUM
elements-9012	232	59	9	9	NUM
elements-9012	232	60	6	6	NUM
elements-9012	232	61	0	0	NUM
elements-9012	232	62	.	.	PUNCT
elements-9012	232	63	2	2	NUM
elements-9012	232	64	1	1	NUM
elements-9012	232	65	7	7	NUM
elements-9012	232	66	6	6	NUM
elements-9012	232	67	6.3	6.3	NUM
elements-9012	232	68	e	e	NOUN
elements-9012	232	69	-42	-42	PROPN
elements-9012	232	70	c.	c.	PROPN
elements-9012	232	71	tropicalis	tropicalis	PROPN
elements-9012	232	72	.	.	PUNCT
elements-9012	233	1	c_dubliniensis	c_dubliniensis	NOUN
elements-9012	233	2	c_albicans	c_albicans	PROPN
elements-9012	233	3	0	0	NUM
elements-9012	234	1	.2893	.2893	PROPN
elements-9012	234	2	3	3	NUM
elements-9012	234	3	7	7	NUM
elements-9012	234	4	9	9	NUM
elements-9012	234	5	6	6	NUM
elements-9012	234	6	0	0	NUM
elements-9012	234	7	.	.	PUNCT
elements-9012	235	1	2	2	NUM
elements-9012	235	2	1	1	NUM
elements-9012	235	3	7	7	NUM
elements-9012	235	4	6	6	NUM
elements-9012	235	5	6.3	6.3	NUM
elements-9012	235	6	e	e	NOUN
elements-9012	235	7	-42	-42	PROPN
elements-9012	235	8	c.	c.	PROPN
elements-9012	235	9	tropicalis	tropicalis	PROPN
elements-9012	235	10	.	.	PUNCT
elements-9012	236	1	c_dubliniensis	c_dubliniensis	PROPN
elements-9012	236	2	c_tropicalis	c_tropicali	VERB
elements-9012	236	3	0	0	PUNCT
elements-9012	237	1	.5522	.5522	NOUN
elements-9012	237	2	3	3	NUM
elements-9012	237	3	5	5	NUM
elements-9012	237	4	1	1	NUM
elements-9012	237	5	4	4	NUM
elements-9012	237	6	0	0	NUM
elements-9012	237	7	.	.	PUNCT
elements-9012	238	1	1	1	NUM
elements-9012	238	2	0	0	NUM
elements-9012	238	3	2	2	NUM
elements-9012	238	4	9	9	NUM
elements-9012	238	5	9.8	9.8	NUM
elements-9012	238	6	e	e	NOUN
elements-9012	238	7	1	1	NUM
elements-9012	238	8	0	0	NUM
elements-9012	238	9	c_tropicalis	c_tropicali	NOUN
elements-9012	238	10	c_parapsilosis	c_parapsilosis	NOUN
elements-9012	238	11	0	0	PUNCT
elements-9012	239	1	.	.	PUNCT
elements-9012	240	1	6	6	NUM
elements-9012	240	2	0	0	NUM
elements-9012	240	3	5	5	NUM
elements-9012	240	4	9	9	NUM
elements-9012	240	5	3	3	NUM
elements-9012	240	6	3	3	NUM
elements-9012	240	7	9	9	NUM
elements-9012	240	8	0	0	NUM
elements-9012	240	9	0	0	NUM
elements-9012	240	10	.	.	PUNCT
elements-9012	240	11	0	0	NUM
elements-9012	241	1	4	4	NUM
elements-9012	241	2	2	2	NUM
elements-9012	241	3	6	6	NUM
elements-9012	241	4	0	0	NUM
elements-9012	241	5	.	.	PUNCT
elements-9012	241	6	0	0	NUM
elements-9012	241	7	1	1	NUM
elements-9012	241	8	3	3	NUM
elements-9012	241	9	0	0	NUM
elements-9012	241	10	ctropicalis	ctropicalis	X
elements-9012	241	11	l_elongisporus	l_elongisporus	NOUN
elements-9012	241	12	0	0	NUM
elements-9012	241	13	.	.	PUNCT
elements-9012	242	1	6	6	NUM
elements-9012	242	2	2	2	NUM
elements-9012	242	3	5	5	NUM
elements-9012	242	4	7	7	NUM
elements-9012	242	5	3	3	NUM
elements-9012	242	6	4	4	NUM
elements-9012	242	7	4	4	NUM
elements-9012	242	8	2	2	NUM
elements-9012	242	9	0	0	NUM
elements-9012	242	10	.	.	PUNCT
elements-9012	243	1	0	0	NUM
elements-9012	244	1	2	2	NUM
elements-9012	244	2	2	2	NUM
elements-9012	244	3	7	7	NUM
elements-9012	244	4	0	0	NUM
elements-9012	244	5	.	.	PUNCT
elements-9012	245	1	1	1	NUM
elements-9012	245	2	8	8	NUM
elements-9012	245	3	1	1	NUM
elements-9012	245	4	8	8	NUM
elements-9012	245	5	n_crassa	n_crassa	NOUN
elements-9012	245	6	c_globosum	c_globosum	NOUN
elements-9012	245	7	0	0	NUM
elements-9012	245	8	.	.	PUNCT
elements-9012	246	1	5	5	NUM
elements-9012	246	2	8	8	NUM
elements-9012	246	3	4	4	NUM
elements-9012	246	4	2	2	NUM
elements-9012	246	5	2	2	NUM
elements-9012	246	6	3	3	NUM
elements-9012	246	7	6	6	NUM
elements-9012	246	8	9	9	NUM
elements-9012	246	9	-0	-0	SYM
elements-9012	246	10	.0085	.0085	NOUN
elements-9012	246	11	0	0	NUM
elements-9012	246	12	.	.	PUNCT
elements-9012	246	13	6	6	NUM
elements-9012	246	14	7	7	NUM
elements-9012	246	15	5	5	NUM
elements-9012	246	16	7	7	NUM
elements-9012	246	17	u_reesii	u_reesii	ADV
elements-9012	246	18	c_immitis	c_immitis	NOUN
elements-9012	246	19	0	0	PUNCT
elements-9012	247	1	.5293	.5293	NOUN
elements-9012	247	2	9	9	NUM
elements-9012	247	3	1	1	NUM
elements-9012	247	4	7	7	NUM
elements-9012	247	5	0	0	NUM
elements-9012	247	6	.	.	PUNCT
elements-9012	248	1	0	0	NUM
elements-9012	248	2	5	5	NUM
elements-9012	248	3	1	1	NUM
elements-9012	248	4	3	3	NUM
elements-9012	248	5	0	0	NUM
elements-9012	248	6	.1201	.1201	NOUN
elements-9012	248	7	a_n1du	a_n1du	AUX
elements-9012	248	8	lans	lan	VERB
elements-9012	248	9	a_terreus	a_terreus	PROPN
elements-9012	248	10	0	0	NUM
elements-9012	248	11	.	.	PUNCT
elements-9012	249	1	6	6	NUM
elements-9012	249	2	3	3	NUM
elements-9012	249	3	6	6	NUM
elements-9012	249	4	4	4	NUM
elements-9012	249	5	1121	1121	NUM
elements-9012	249	6	0	0	NUM
elements-9012	249	7	.	.	PUNCT
elements-9012	249	8	0	0	NUM
elements-9012	250	1	3	3	NUM
elements-9012	250	2	7	7	NUM
elements-9012	250	3	0	0	NUM
elements-9012	250	4	0	0	NUM
elements-9012	250	5	.	.	PUNCT
elements-9012	250	6	2	2	NUM
elements-9012	250	7	1	1	NUM
elements-9012	250	8	4	4	NUM
elements-9012	250	9	7	7	NUM
elements-9012	250	10	h_capsu	h_capsu	NOUN
elements-9012	250	11	latum	latum	NOUN
elements-9012	250	12	c_immitis	c_immitis	NOUN
elements-9012	250	13	0	0	NUM
elements-9012	250	14	.	.	PUNCT
elements-9012	251	1	6	6	NUM
elements-9012	251	2	8	8	NUM
elements-9012	251	3	3	3	NUM
elements-9012	251	4	9	9	NUM
elements-9012	251	5	8	8	NUM
elements-9012	251	6	4	4	NUM
elements-9012	251	7	7	7	NUM
elements-9012	251	8	0	0	NUM
elements-9012	251	9	.	.	PUNCT
elements-9012	251	10	0	0	NUM
elements-9012	252	1	3	3	NUM
elements-9012	252	2	5	5	NUM
elements-9012	252	3	6	6	NUM
elements-9012	252	4	0	0	NUM
elements-9012	252	5	.	.	PUNCT
elements-9012	253	1	2	2	NUM
elements-9012	253	2	9	9	NUM
elements-9012	253	3	9	9	NUM
elements-9012	253	4	9	9	NUM
elements-9012	253	5	h_capsu	h_capsu	NOUN
elements-9012	253	6	latum	latum	NOUN
elements-9012	253	7	u_reesii	u_reesii	X
elements-9012	253	8	0	0	NUM
elements-9012	253	9	.	.	PUNCT
elements-9012	254	1	6	6	NUM
elements-9012	254	2	6	6	NUM
elements-9012	254	3	5	5	NUM
elements-9012	254	4	5	5	NUM
elements-9012	254	5	393	393	NUM
elements-9012	254	6	0	0	NUM
elements-9012	254	7	.	.	PUNCT
elements-9012	255	1	0	0	NUM
elements-9012	255	2	0	0	NUM
elements-9012	255	3	3	3	NUM
elements-9012	255	4	4	4	NUM
elements-9012	255	5	0	0	NUM
elements-9012	255	6	.	.	PUNCT
elements-9012	256	1	9	9	NUM
elements-9012	256	2	4	4	NUM
elements-9012	256	3	5	5	NUM
elements-9012	256	4	9	9	NUM
elements-9012	256	5	l_elongisporus	l_elongisporus	NOUN
elements-9012	256	6	c_parapsilosis	c_parapsilosis	NOUN
elements-9012	256	7	0	0	NUM
elements-9012	256	8	.6363	.6363	NOUN
elements-9012	256	9	3	3	NUM
elements-9012	256	10	1	1	NUM
elements-9012	256	11	5	5	NUM
elements-9012	256	12	2	2	NUM
elements-9012	256	13	0	0	NUM
elements-9012	256	14	.0265	.0265	NOUN
elements-9012	256	15	0	0	NUM
elements-9012	256	16	.	.	PUNCT
elements-9012	257	1	1	1	NUM
elements-9012	257	2	3	3	NUM
elements-9012	257	3	6	6	NUM
elements-9012	257	4	6	6	NUM
elements-9012	257	5	t	t	NOUN
elements-9012	257	6	a	a	DET
elements-9012	257	7	b	b	NOUN
elements-9012	257	8	l	l	NOUN
elements-9012	257	9	e	e	NOUN
elements-9012	257	10	2	2	NUM
elements-9012	257	11	:	:	PUNCT
elements-9012	257	12	m	m	VERB
elements-9012	257	13	a	a	DET
elements-9012	257	14	m	m	NOUN
elements-9012	257	15	m	m	VERB
elements-9012	257	16	a	a	DET
elements-9012	257	17	l	l	NOUN
elements-9012	257	18	i	i	PRON
elements-9012	257	19	a	a	PRON
elements-9012	257	20	n	n	NOUN
elements-9012	257	21	p	p	NOUN
elements-9012	257	22	a	a	PRON
elements-9012	258	1	i	i	NOUN
elements-9012	258	2	r	r	VERB
elements-9012	258	3	w	w	NOUN
elements-9012	259	1	i	i	NOUN
elements-9012	259	2	s	s	X
elements-9012	259	3	e	e	NOUN
elements-9012	259	4	c	c	NOUN
elements-9012	259	5	o	o	NOUN
elements-9012	259	6	m	m	VERB
elements-9012	259	7	p	p	NOUN
elements-9012	260	1	a	a	DET
elements-9012	260	2	r	r	NOUN
elements-9012	261	1	i	i	PRON
elements-9012	261	2	s	s	VERB
elements-9012	261	3	o	o	NOUN
elements-9012	262	1	n	n	X
elements-9012	262	2	s	s	PROPN
elements-9012	262	3	p	p	X
elements-9012	262	4	a	a	PRON
elements-9012	263	1	i	i	NOUN
elements-9012	263	2	r	r	VERB
elements-9012	263	3	a	a	DET
elements-9012	263	4	v	v	NOUN
elements-9012	263	5	e	e	NOUN
elements-9012	263	6	r	r	NOUN
elements-9012	263	7	a	a	DET
elements-9012	263	8	g	g	NOUN
elements-9012	263	9	e	e	PROPN
elements-9012	263	10	n	n	ADP
elements-9012	263	11	u	u	NOUN
elements-9012	263	12	m	m	NOUN
elements-9012	263	13	b	b	NOUN
elements-9012	263	14	e	e	NOUN
elements-9012	263	15	r	r	NOUN
elements-9012	263	16	p	p	PROPN
elements-9012	263	17	e	e	PROPN
elements-9012	263	18	a	a	PRON
elements-9012	263	19	r	r	NOUN
elements-9012	263	20	s	s	PROPN
elements-9012	263	21	o	o	NOUN
elements-9012	263	22	n	n	CCONJ
elements-9012	263	23	n	n	CCONJ
elements-9012	263	24	u	u	NOUN
elements-9012	263	25	m	m	PROPN
elements-9012	263	26	b	b	NOUN
elements-9012	263	27	e	e	NOUN
elements-9012	263	28	r	r	NOUN
elements-9012	263	29	s	s	X
elements-9012	263	30	p	p	X
elements-9012	263	31	e	e	NOUN
elements-9012	263	32	c	c	NOUN
elements-9012	264	1	i	i	NOUN
elements-9012	264	2	e	e	NOUN
elements-9012	264	3	s	s	VERB
elements-9012	264	4	1	1	NUM
elements-9012	264	5	s	s	PART
elements-9012	264	6	p	p	NOUN
elements-9012	264	7	e	e	NOUN
elements-9012	264	8	c	c	NOUN
elements-9012	265	1	i	i	NOUN
elements-9012	265	2	e	e	NOUN
elements-9012	265	3	s	s	PROPN
elements-9012	265	4	2	2	NUM
elements-9012	265	5	d	d	NOUN
elements-9012	266	1	i	i	PRON
elements-9012	267	1	v	v	VERB
elements-9012	267	2	e	e	NOUN
elements-9012	267	3	r	r	NOUN
elements-9012	267	4	g	g	NOUN
elements-9012	267	5	e	e	NOUN
elements-9012	267	6	n	n	NOUN
elements-9012	267	7	c	c	NOUN
elements-9012	267	8	e	e	NOUN
elements-9012	267	9	o	o	NOUN
elements-9012	267	10	r	r	NOUN
elements-9012	267	11	t	t	NOUN
elements-9012	268	1	h	h	NOUN
elements-9012	269	1	o	o	NOUN
elements-9012	269	2	l	l	NOUN
elements-9012	270	1	o	o	X
elements-9012	270	2	g	g	NOUN
elements-9012	270	3	s	s	X
elements-9012	270	4	c	c	NOUN
elements-9012	270	5	o	o	NOUN
elements-9012	270	6	r	r	NOUN
elements-9012	270	7	r	r	NOUN
elements-9012	270	8	e	e	NOUN
elements-9012	270	9	l	l	NOUN
elements-9012	270	10	a	a	NOUN
elements-9012	270	11	t	t	NOUN
elements-9012	271	1	i	i	PRON
elements-9012	271	2	o	o	VERB
elements-9012	272	1	n	n	CCONJ
elements-9012	272	2	p	p	X
elements-9012	272	3	v	v	X
elements-9012	272	4	a	a	DET
elements-9012	272	5	l	l	NOUN
elements-9012	272	6	1	1	NUM
elements-9012	272	7	h	h	NOUN
elements-9012	272	8	u	u	NOUN
elements-9012	272	9	m	m	PROPN
elements-9012	272	10	a	a	PRON
elements-9012	272	11	n	n	NUM
elements-9012	272	12	mouse	mouse	NOUN
elements-9012	272	13	0	0	NUM
elements-9012	272	14	.	.	PUNCT
elements-9012	273	1	3	3	NUM
elements-9012	273	2	3	3	NUM
elements-9012	273	3	1	1	NUM
elements-9012	273	4	7	7	NUM
elements-9012	273	5	1	1	NUM
elements-9012	273	6	7	7	NUM
elements-9012	273	7	1	1	NUM
elements-9012	273	8	1	1	NUM
elements-9012	273	9	2	2	NUM
elements-9012	273	10	0	0	NUM
elements-9012	273	11	.	.	PUNCT
elements-9012	274	1	2	2	NUM
elements-9012	274	2	3	3	NUM
elements-9012	274	3	0	0	NUM
elements-9012	274	4	6	6	NUM
elements-9012	274	5	2.0e	2.0e	NUM
elements-9012	274	6	-205	-205	PROPN
elements-9012	274	7	mammalian	mammalian	ADJ
elements-9012	274	8	pairwise	pairwise	PROPN
elements-9012	274	9	compar2	compar2	PROPN
elements-9012	275	1	h	h	NOUN
elements-9012	275	2	u	u	NOUN
elements-9012	275	3	m	m	VERB
elements-9012	275	4	a	a	DET
elements-9012	275	5	n	n	NUM
elements-9012	275	6	dog	dog	NOUN
elements-9012	275	7	0	0	NUM
elements-9012	275	8	.2451	.2451	NOUN
elements-9012	275	9	1	1	NUM
elements-9012	275	10	6	6	NUM
elements-9012	275	11	5	5	NUM
elements-9012	275	12	3	3	NUM
elements-9012	275	13	2	2	NUM
elements-9012	275	14	0	0	NUM
elements-9012	275	15	.	.	PUNCT
elements-9012	276	1	3	3	NUM
elements-9012	276	2	0	0	NUM
elements-9012	276	3	1	1	NUM
elements-9012	276	4	7	7	NUM
elements-9012	276	5	9.8e	9.8e	NUM
elements-9012	276	6	-247	-247	NUM
elements-9012	276	7	isons	ison	NOUN
elements-9012	276	8	.	.	PUNCT
elements-9012	277	1	4	4	X
elements-9012	277	2	-	-	ADJ
elements-9012	277	3	fold	fold	ADJ
elements-9012	277	4	site	site	NOUN
elements-9012	277	5	3	3	NUM
elements-9012	277	6	h	h	NOUN
elements-9012	277	7	u	u	NOUN
elements-9012	277	8	m	m	PROPN
elements-9012	277	9	a	a	DET
elements-9012	277	10	n	n	X
elements-9012	277	11	cat	cat	NOUN
elements-9012	277	12	0	0	NUM
elements-9012	277	13	.	.	PUNCT
elements-9012	278	1	2	2	NUM
elements-9012	278	2	4	4	NUM
elements-9012	278	3	1	1	NUM
elements-9012	278	4	7	7	NUM
elements-9012	278	5	7	7	NUM
elements-9012	278	6	5	5	NUM
elements-9012	278	7	2	2	NUM
elements-9012	278	8	9	9	NUM
elements-9012	278	9	0	0	NUM
elements-9012	278	10	.3263	.3263	PROPN
elements-9012	278	11	2	2	NUM
elements-9012	278	12	.	.	X
elements-9012	278	13	5	5	NUM
elements-9012	278	14	e	e	NOUN
elements-9012	278	15	1	1	NUM
elements-9012	278	16	8	8	NUM
elements-9012	278	17	6	6	NUM
elements-9012	278	18	divergence	divergence	NOUN
elements-9012	278	19	between	between	ADP
elements-9012	278	20	the	the	DET
elements-9012	278	21	two	two	NUM
elements-9012	278	22	4	4	NUM
elements-9012	278	23	h	h	NOUN
elements-9012	278	24	u	u	NOUN
elements-9012	278	25	m	m	PROPN
elements-9012	278	26	a	a	PRON
elements-9012	278	27	n	n	X
elements-9012	278	28	cow	cow	NOUN
elements-9012	278	29	0	0	NUM
elements-9012	278	30	.5761	.5761	NOUN
elements-9012	278	31	6	6	NUM
elements-9012	278	32	9	9	NUM
elements-9012	278	33	7	7	NUM
elements-9012	278	34	2	2	NUM
elements-9012	278	35	0	0	NUM
elements-9012	278	36	.	.	PUNCT
elements-9012	278	37	0	0	NUM
elements-9012	278	38	5	5	NUM
elements-9012	278	39	4	4	NUM
elements-9012	278	40	0	0	NUM
elements-9012	278	41	6	6	NUM
elements-9012	278	42	.	.	SYM
elements-9012	278	43	2	2	NUM
elements-9012	278	44	e	e	NOUN
elements-9012	278	45	0	0	NUM
elements-9012	278	46	6	6	NUM
elements-9012	278	47	species	specie	NOUN
elements-9012	278	48	is	be	AUX
elements-9012	278	49	shown	show	VERB
elements-9012	278	50	in	in	ADP
elements-9012	278	51	column	column	NOUN
elements-9012	278	52	5	5	NUM
elements-9012	278	53	dog	dog	NOUN
elements-9012	278	54	cat	cat	NOUN
elements-9012	278	55	0	0	NUM
elements-9012	279	1	.1831	.1831	PROPN
elements-9012	279	2	12931	12931	NUM
elements-9012	279	3	0	0	NUM
elements-9012	279	4	.	.	PUNCT
elements-9012	280	1	2	2	NUM
elements-9012	280	2	4	4	NUM
elements-9012	280	3	3	3	NUM
elements-9012	280	4	8	8	NUM
elements-9012	280	5	1	1	NUM
elements-9012	280	6	.	.	PUNCT
elements-9012	280	7	9	9	NUM
elements-9012	280	8	e	e	NOUN
elements-9012	280	9	1	1	NUM
elements-9012	280	10	7	7	NUM
elements-9012	280	11	4	4	NUM
elements-9012	280	12	3	3	NUM
elements-9012	280	13	.	.	PUNCT
elements-9012	281	1	pearson	pearson	PROPN
elements-9012	281	2	correlations	correlation	NOUN
elements-9012	281	3	for	for	ADP
elements-9012	281	4	6	6	NUM
elements-9012	281	5	cat	cat	NOUN
elements-9012	281	6	mouse	mouse	PROPN
elements-9012	281	7	0	0	NUM
elements-9012	281	8	.3581	.3581	PROPN
elements-9012	281	9	6	6	NUM
elements-9012	281	10	7	7	NUM
elements-9012	281	11	2	2	NUM
elements-9012	281	12	6	6	NUM
elements-9012	281	13	0	0	NUM
elements-9012	281	14	.	.	PUNCT
elements-9012	281	15	2	2	NUM
elements-9012	281	16	2	2	NUM
elements-9012	281	17	9	9	NUM
elements-9012	281	18	2	2	NUM
elements-9012	281	19	6	6	NUM
elements-9012	281	20	.	.	PUNCT
elements-9012	281	21	0	0	NUM
elements-9012	282	1	e	e	NOUN
elements-9012	282	2	8	8	NUM
elements-9012	282	3	1	1	NUM
elements-9012	282	4	the	the	DET
elements-9012	282	5	normalized	normalize	VERB
elements-9012	282	6	0	0	NUM
elements-9012	282	7	.	.	PUNCT
elements-9012	283	1	1	1	NUM
elements-9012	283	2	5	5	NUM
elements-9012	283	3	8	8	NUM
elements-9012	283	4	6	6	NUM
elements-9012	283	5	substitution	substitution	NOUN
elements-9012	283	6	rates	rate	NOUN
elements-9012	283	7	of	of	ADP
elements-9012	283	8	neigh7	neigh7	PROPN
elements-9012	283	9	rat	rat	PROPN
elements-9012	283	10	mouse	mouse	PROPN
elements-9012	283	11	0	0	NUM
elements-9012	283	12	.	.	PUNCT
elements-9012	284	1	1	1	NUM
elements-9012	284	2	5	5	NUM
elements-9012	284	3	8	8	NUM
elements-9012	284	4	6	6	NUM
elements-9012	284	5	2	2	NUM
elements-9012	284	6	0	0	NUM
elements-9012	284	7	0	0	NUM
elements-9012	284	8	5	5	NUM
elements-9012	284	9	4	4	NUM
elements-9012	284	10	0	0	NUM
elements-9012	284	11	.	.	PUNCT
elements-9012	284	12	1	1	NUM
elements-9012	284	13	5	5	NUM
elements-9012	284	14	7	7	NUM
elements-9012	284	15	6	6	NUM
elements-9012	284	16	i	i	PRON
elements-9012	284	17	.	.	PUNCT
elements-9012	285	1	o	o	X
elements-9012	285	2	e	e	NOUN
elements-9012	285	3	1	1	NUM
elements-9012	285	4	1	1	NUM
elements-9012	285	5	1	1	NUM
elements-9012	285	6	boring	boring	ADJ
elements-9012	285	7	genes	gene	NOUN
elements-9012	285	8	are	be	AUX
elements-9012	285	9	shown	show	VERB
elements-9012	285	10	in	in	ADP
elements-9012	285	11	8	8	NUM
elements-9012	285	12	mouse	mouse	NOUN
elements-9012	285	13	rabbit	rabbit	NOUN
elements-9012	285	14	0	0	NUM
elements-9012	285	15	.	.	PUNCT
elements-9012	286	1	3	3	NUM
elements-9012	286	2	6	6	NUM
elements-9012	286	3	0	0	NUM
elements-9012	286	4	6	6	NUM
elements-9012	286	5	1	1	NUM
elements-9012	286	6	2	2	NUM
elements-9012	286	7	4	4	NUM
elements-9012	286	8	4	4	NUM
elements-9012	286	9	2	2	NUM
elements-9012	286	10	0	0	NUM
elements-9012	286	11	.1851	.1851	NOUN
elements-9012	286	12	2	2	NUM
elements-9012	286	13	.	.	SYM
elements-9012	286	14	2	2	NUM
elements-9012	286	15	e	e	NOUN
elements-9012	286	16	9	9	NUM
elements-9012	286	17	6	6	NUM
elements-9012	286	18	column	column	NOUN
elements-9012	286	19	5	5	NUM
elements-9012	286	20	.	.	PUNCT
elements-9012	287	1	every	every	DET
elements-9012	287	2	mammalian	mammalian	NOUN
elements-9012	287	3	0	0	NUM
elements-9012	287	4	.1851	.1851	NOUN
elements-9012	287	5	2	2	NUM
elements-9012	287	6	.	.	SYM
elements-9012	287	7	2	2	NUM
elements-9012	287	8	e	e	NOUN
elements-9012	287	9	9	9	NUM
elements-9012	287	10	6	6	NUM
elements-9012	287	11	column	column	NOUN
elements-9012	287	12	5	5	NUM
elements-9012	287	13	.	.	PUNCT
elements-9012	288	1	every	every	DET
elements-9012	288	2	mammalian	mammalian	NOUN
elements-9012	288	3	9	9	NUM
elements-9012	288	4	euro	euro	NOUN
elements-9012	288	5	.	.	PUNCT
elements-9012	289	1	hedgehog	hedgehog	NOUN
elements-9012	289	2	tenrec	tenrec	NOUN
elements-9012	289	3	0	0	NUM
elements-9012	289	4	.	.	PUNCT
elements-9012	290	1	3	3	NUM
elements-9012	290	2	6	6	NUM
elements-9012	290	3	7	7	NUM
elements-9012	290	4	9	9	NUM
elements-9012	290	5	2	2	NUM
elements-9012	290	6	3	3	NUM
elements-9012	290	7	6	6	NUM
elements-9012	290	8	0	0	NUM
elements-9012	290	9	0	0	NUM
elements-9012	290	10	.	.	PUNCT
elements-9012	291	1	1	1	NUM
elements-9012	291	2	9	9	NUM
elements-9012	291	3	6	6	NUM
elements-9012	291	4	8	8	NUM
elements-9012	291	5	4	4	NUM
elements-9012	291	6	.	.	PUNCT
elements-9012	291	7	6	6	NUM
elements-9012	291	8	e	e	NOUN
elements-9012	291	9	2	2	NUM
elements-9012	291	10	2	2	NUM
elements-9012	291	11	comparison	comparison	NOUN
elements-9012	291	12	shows	show	VERB
elements-9012	291	13	a	a	DET
elements-9012	291	14	signifi1	signifi1	NOUN
elements-9012	291	15	0	0	NUM
elements-9012	291	16	euro	euro	NOUN
elements-9012	291	17	.	.	PUNCT
elements-9012	292	1	hedgehog	hedgehog	NOUN
elements-9012	292	2	tree	tree	PROPN
elements-9012	292	3	shrew	shrew	PROPN
elements-9012	292	4	0	0	NUM
elements-9012	292	5	.	.	PUNCT
elements-9012	293	1	3	3	NUM
elements-9012	293	2	3	3	NUM
elements-9012	293	3	2	2	NUM
elements-9012	293	4	2	2	NUM
elements-9012	293	5	1	1	NUM
elements-9012	293	6	6	6	NUM
elements-9012	293	7	5	5	NUM
elements-9012	293	8	0	0	NUM
elements-9012	293	9	0	0	NUM
elements-9012	293	10	.	.	PUNCT
elements-9012	293	11	3	3	NUM
elements-9012	293	12	1	1	NUM
elements-9012	293	13	3	3	NUM
elements-9012	293	14	2	2	NUM
elements-9012	293	15	6	6	NUM
elements-9012	293	16	.	.	PUNCT
elements-9012	293	17	7	7	NUM
elements-9012	293	18	e	e	NOUN
elements-9012	293	19	3	3	NUM
elements-9012	293	20	9	9	NUM
elements-9012	293	21	ca	can	AUX
elements-9012	293	22	nt	not	PART
elements-9012	293	23	correlation	correlation	VERB
elements-9012	293	24	between	between	ADP
elements-9012	293	25	1	1	NUM
elements-9012	293	26	1	1	NUM
elements-9012	293	27	bushbaby	bushbaby	PROPN
elements-9012	293	28	opossum	opossum	NOUN
elements-9012	293	29	0	0	NUM
elements-9012	293	30	.	.	PUNCT
elements-9012	294	1	4	4	NUM
elements-9012	294	2	5	5	NUM
elements-9012	294	3	9	9	NUM
elements-9012	294	4	9	9	NUM
elements-9012	294	5	1	1	NUM
elements-9012	294	6	2	2	NUM
elements-9012	294	7	9	9	NUM
elements-9012	294	8	2	2	NUM
elements-9012	294	9	2	2	NUM
elements-9012	294	10	0	0	NUM
elements-9012	294	11	.	.	PUNCT
elements-9012	295	1	2	2	NUM
elements-9012	295	2	0	0	NUM
elements-9012	295	3	9	9	NUM
elements-9012	295	4	7	7	NUM
elements-9012	295	5	2	2	NUM
elements-9012	295	6	.	.	PUNCT
elements-9012	295	7	0	0	NUM
elements-9012	296	1	e	e	NOUN
elements-9012	296	2	1	1	NUM
elements-9012	296	3	2	2	NUM
elements-9012	296	4	8	8	NUM
elements-9012	296	5	rates	rate	NOUN
elements-9012	296	6	of	of	ADP
elements-9012	296	7	neighboring	neighboring	NOUN
elements-9012	296	8	genes	gene	NOUN
elements-9012	296	9	1	1	NUM
elements-9012	296	10	2	2	NUM
elements-9012	296	11	cow	cow	NOUN
elements-9012	296	12	dog	dog	NOUN
elements-9012	296	13	0	0	NUM
elements-9012	296	14	.	.	PUNCT
elements-9012	297	1	2	2	NUM
elements-9012	297	2	5	5	NUM
elements-9012	297	3	3	3	NUM
elements-9012	297	4	2	2	NUM
elements-9012	297	5	16941	16941	NUM
elements-9012	297	6	0	0	NUM
elements-9012	297	7	.	.	PUNCT
elements-9012	298	1	2	2	NUM
elements-9012	298	2	6	6	NUM
elements-9012	298	3	0	0	NUM
elements-9012	298	4	6	6	NUM
elements-9012	298	5	4.4e	4.4e	PROPN
elements-9012	298	6	-261	-261	PROPN
elements-9012	298	7	(	(	PUNCT
elements-9012	298	8	column	column	NOUN
elements-9012	298	9	6	6	NUM
elements-9012	298	10	)	)	PUNCT
elements-9012	298	11	.	.	PUNCT
elements-9012	299	1	1	1	NUM
elements-9012	299	2	3	3	NUM
elements-9012	299	3	cow	cow	NOUN
elements-9012	299	4	cat	cat	NOUN
elements-9012	299	5	0	0	NUM
elements-9012	299	6	.	.	PUNCT
elements-9012	299	7	2	2	NUM
elements-9012	299	8	5	5	NUM
elements-9012	299	9	3	3	NUM
elements-9012	299	10	0	0	NUM
elements-9012	299	11	13355	13355	NUM
elements-9012	299	12	0	0	NUM
elements-9012	299	13	.	.	PUNCT
elements-9012	299	14	2	2	NUM
elements-9012	299	15	6	6	NUM
elements-9012	299	16	2	2	NUM
elements-9012	299	17	6	6	NUM
elements-9012	299	18	1	1	NUM
elements-9012	299	19	.5e	.5e	X
elements-9012	299	20	-209	-209	NOUN
elements-9012	299	21	1	1	NUM
elements-9012	299	22	4	4	NUM
elements-9012	299	23	dog	dog	NOUN
elements-9012	299	24	mouse	mouse	NOUN
elements-9012	299	25	0	0	NUM
elements-9012	299	26	.	.	PUNCT
elements-9012	300	1	3	3	NUM
elements-9012	300	2	5	5	NUM
elements-9012	300	3	5	5	NUM
elements-9012	300	4	0	0	NUM
elements-9012	300	5	1	1	NUM
elements-9012	300	6	6	6	NUM
elements-9012	300	7	5	5	NUM
elements-9012	300	8	2	2	NUM
elements-9012	300	9	2	2	NUM
elements-9012	300	10	0	0	NUM
elements-9012	300	11	.	.	PUNCT
elements-9012	300	12	2	2	NUM
elements-9012	300	13	1	1	NUM
elements-9012	300	14	5	5	NUM
elements-9012	300	15	2	2	NUM
elements-9012	300	16	2	2	NUM
elements-9012	300	17	.	.	PUNCT
elements-9012	300	18	4	4	NUM
elements-9012	300	19	e	e	NOUN
elements-9012	300	20	1	1	NUM
elements-9012	300	21	7	7	NUM
elements-9012	300	22	2	2	NUM
elements-9012	300	23	1	1	NUM
elements-9012	300	24	5	5	NUM
elements-9012	300	25	cow	cow	NOUN
elements-9012	300	26	mouse	mouse	NOUN
elements-9012	300	27	0	0	NUM
elements-9012	300	28	.	.	PUNCT
elements-9012	301	1	3	3	NUM
elements-9012	301	2	6	6	NUM
elements-9012	301	3	4	4	NUM
elements-9012	301	4	6	6	NUM
elements-9012	301	5	1	1	NUM
elements-9012	301	6	7	7	NUM
elements-9012	301	7	3	3	NUM
elements-9012	301	8	4	4	NUM
elements-9012	301	9	0	0	NUM
elements-9012	301	10	0	0	NUM
elements-9012	301	11	.	.	PUNCT
elements-9012	302	1	2	2	NUM
elements-9012	302	2	1	1	NUM
elements-9012	302	3	5	5	NUM
elements-9012	302	4	2	2	NUM
elements-9012	302	5	6	6	NUM
elements-9012	302	6	.	.	PUNCT
elements-9012	302	7	5	5	NUM
elements-9012	302	8	e	e	NOUN
elements-9012	302	9	1	1	NUM
elements-9012	302	10	8	8	NUM
elements-9012	302	11	1	1	NUM
elements-9012	302	12	1	1	NUM
elements-9012	302	13	6	6	NUM
elements-9012	302	14	h	h	NOUN
elements-9012	302	15	u	u	NOUN
elements-9012	302	16	m	m	NOUN
elements-9012	302	17	a	a	PRON
elements-9012	302	18	n	n	ADV
elements-9012	302	19	macaque	macaque	ADJ
elements-9012	302	20	0	0	NUM
elements-9012	303	1	.0673	.0673	PROPN
elements-9012	303	2	1	1	NUM
elements-9012	303	3	6	6	NUM
elements-9012	303	4	0	0	NUM
elements-9012	303	5	0	0	NUM
elements-9012	303	6	0	0	NUM
elements-9012	303	7	0	0	NUM
elements-9012	303	8	.	.	PUNCT
elements-9012	304	1	2	2	NUM
elements-9012	304	2	7	7	NUM
elements-9012	304	3	5	5	NUM
elements-9012	304	4	4	4	NUM
elements-9012	304	5	2	2	NUM
elements-9012	304	6	.	.	PUNCT
elements-9012	304	7	0	0	NUM
elements-9012	305	1	e	e	NOUN
elements-9012	305	2	2	2	NUM
elements-9012	305	3	7	7	NUM
elements-9012	305	4	6	6	NUM
elements-9012	305	5	1	1	NUM
elements-9012	305	6	7	7	NUM
elements-9012	305	7	bat	bat	NOUN
elements-9012	305	8	rat	rat	NOUN
elements-9012	305	9	0	0	NUM
elements-9012	305	10	.	.	PUNCT
elements-9012	306	1	3	3	NUM
elements-9012	306	2	6	6	NUM
elements-9012	306	3	2	2	NUM
elements-9012	306	4	7	7	NUM
elements-9012	306	5	5	5	NUM
elements-9012	306	6	9	9	NUM
elements-9012	306	7	3	3	NUM
elements-9012	306	8	8	8	NUM
elements-9012	306	9	0	0	NUM
elements-9012	306	10	.	.	PUNCT
elements-9012	307	1	1	1	NUM
elements-9012	307	2	9	9	NUM
elements-9012	307	3	9	9	NUM
elements-9012	307	4	5	5	NUM
elements-9012	307	5	2	2	NUM
elements-9012	307	6	.	.	PUNCT
elements-9012	307	7	2	2	NUM
elements-9012	307	8	e	e	NOUN
elements-9012	307	9	5	5	NUM
elements-9012	307	10	4	4	NUM
elements-9012	307	11	1	1	NUM
elements-9012	307	12	8	8	NUM
elements-9012	307	13	macaque	macaque	ADJ
elements-9012	308	1	c	c	NOUN
elements-9012	308	2	h	h	NOUN
elements-9012	309	1	i	i	PRON
elements-9012	309	2	m	m	VERB
elements-9012	309	3	p	p	ADJ
elements-9012	309	4	0	0	NUM
elements-9012	309	5	.	.	PUNCT
elements-9012	310	1	0	0	NUM
elements-9012	311	1	7	7	NUM
elements-9012	311	2	1	1	NUM
elements-9012	311	3	0	0	NUM
elements-9012	311	4	1	1	NUM
elements-9012	311	5	7	7	NUM
elements-9012	311	6	6	6	NUM
elements-9012	311	7	2	2	NUM
elements-9012	311	8	1	1	NUM
elements-9012	311	9	0	0	NUM
elements-9012	311	10	.	.	PUNCT
elements-9012	311	11	2	2	NUM
elements-9012	311	12	2	2	NUM
elements-9012	311	13	3	3	NUM
elements-9012	311	14	1	1	NUM
elements-9012	311	15	1.1	1.1	NUM
elements-9012	311	16	e	e	NOUN
elements-9012	311	17	1	1	NUM
elements-9012	311	18	9	9	NUM
elements-9012	311	19	7	7	NUM
elements-9012	311	20	1	1	NUM
elements-9012	311	21	9	9	NUM
elements-9012	311	22	armadillo	armadillo	NOUN
elements-9012	311	23	elephant	elephant	NOUN
elements-9012	311	24	0	0	NUM
elements-9012	311	25	.	.	PUNCT
elements-9012	312	1	2	2	NUM
elements-9012	312	2	5	5	NUM
elements-9012	312	3	2	2	NUM
elements-9012	312	4	7	7	NUM
elements-9012	312	5	2	2	NUM
elements-9012	312	6	0	0	NUM
elements-9012	312	7	4	4	NUM
elements-9012	312	8	5	5	NUM
elements-9012	312	9	0	0	NUM
elements-9012	312	10	.	.	PUNCT
elements-9012	313	1	1	1	NUM
elements-9012	313	2	9	9	NUM
elements-9012	313	3	4	4	NUM
elements-9012	313	4	8	8	NUM
elements-9012	313	5	5	5	NUM
elements-9012	313	6	.	.	PUNCT
elements-9012	314	1	9	9	NUM
elements-9012	314	2	e	e	NOUN
elements-9012	314	3	1	1	NUM
elements-9012	314	4	9	9	NUM
elements-9012	314	5	2	2	NUM
elements-9012	314	6	0	0	NUM
elements-9012	314	7	mouse	mouse	NOUN
elements-9012	314	8	squirrel	squirrel	NOUN
elements-9012	314	9	0	0	NUM
elements-9012	314	10	.	.	PUNCT
elements-9012	315	1	3	3	NUM
elements-9012	315	2	3	3	NUM
elements-9012	315	3	0	0	NUM
elements-9012	315	4	4	4	NUM
elements-9012	315	5	1	1	NUM
elements-9012	315	6	2	2	NUM
elements-9012	315	7	7	7	NUM
elements-9012	315	8	9	9	NUM
elements-9012	315	9	4	4	NUM
elements-9012	315	10	0	0	NUM
elements-9012	315	11	.	.	PUNCT
elements-9012	316	1	1	1	NUM
elements-9012	316	2	7	7	NUM
elements-9012	316	3	8	8	NUM
elements-9012	316	4	9	9	NUM
elements-9012	316	5	1	1	NUM
elements-9012	316	6	.4e-92	.4e-92	NOUN
elements-9012	316	7	21	21	NUM
elements-9012	316	8	tenrec	tenrec	NOUN
elements-9012	316	9	elephant	elephant	NOUN
elements-9012	316	10	0	0	NUM
elements-9012	316	11	.2741	.2741	NOUN
elements-9012	316	12	2	2	NUM
elements-9012	316	13	2	2	NUM
elements-9012	316	14	4	4	NUM
elements-9012	316	15	4	4	NUM
elements-9012	316	16	0	0	NUM
elements-9012	316	17	.	.	PUNCT
elements-9012	317	1	1	1	NUM
elements-9012	317	2	6	6	NUM
elements-9012	317	3	6	6	NUM
elements-9012	317	4	7	7	NUM
elements-9012	317	5	1	1	NUM
elements-9012	317	6	.	.	PUNCT
elements-9012	317	7	8	8	NUM
elements-9012	317	8	e	e	NOUN
elements-9012	317	9	1	1	NUM
elements-9012	317	10	5	5	NUM
elements-9012	317	11	22	22	NUM
elements-9012	318	1	c	c	NOUN
elements-9012	318	2	o	o	NOUN
elements-9012	318	3	m	m	VERB
elements-9012	318	4	m	m	VERB
elements-9012	318	5	o	o	NOUN
elements-9012	318	6	n	n	PRON
elements-9012	318	7	shrew	shrew	NOUN
elements-9012	318	8	tree	tree	NOUN
elements-9012	318	9	shrew	shrew	NOUN
elements-9012	318	10	0	0	NUM
elements-9012	318	11	.	.	PUNCT
elements-9012	319	1	3	3	NUM
elements-9012	319	2	4	4	NUM
elements-9012	319	3	4	4	NUM
elements-9012	319	4	2	2	NUM
elements-9012	319	5	2	2	NUM
elements-9012	319	6	1	1	NUM
elements-9012	319	7	2	2	NUM
elements-9012	319	8	4	4	NUM
elements-9012	319	9	0	0	NUM
elements-9012	319	10	.2503	.2503	NOUN
elements-9012	319	11	i	i	PRON
elements-9012	319	12	.oe-31	.oe-31	PUNCT
elements-9012	319	13	23	23	NUM
elements-9012	319	14	doc	doc	PROPN
elements-9012	319	15	bat	bat	NOUN
elements-9012	319	16	0	0	NUM
elements-9012	319	17	.	.	PUNCT
elements-9012	319	18	2	2	NUM
elements-9012	319	19	4	4	NUM
elements-9012	319	20	5	5	NUM
elements-9012	319	21	9	9	NUM
elements-9012	319	22	1	1	NUM
elements-9012	319	23	3	3	NUM
elements-9012	319	24	6	6	NUM
elements-9012	319	25	6	6	NUM
elements-9012	319	26	9	9	NUM
elements-9012	319	27	0	0	NUM
elements-9012	319	28	.	.	PUNCT
elements-9012	320	1	3	3	NUM
elements-9012	320	2	5	5	NUM
elements-9012	320	3	2	2	NUM
elements-9012	320	4	7	7	NUM
elements-9012	320	5	<	<	X
elements-9012	320	6	e	e	X
elements-9012	320	7	2	2	NUM
elements-9012	320	8	7	7	NUM
elements-9012	320	9	6	6	NUM
elements-9012	320	10	24	24	NUM
elements-9012	320	11	cow	cow	NOUN
elements-9012	320	12	bat	bat	NOUN
elements-9012	320	13	0	0	NUM
elements-9012	320	14	.	.	PUNCT
elements-9012	320	15	2	2	NUM
elements-9012	320	16	5	5	NUM
elements-9012	320	17	7	7	NUM
elements-9012	320	18	9	9	NUM
elements-9012	320	19	1	1	NUM
elements-9012	320	20	4	4	NUM
elements-9012	320	21	1	1	NUM
elements-9012	320	22	7	7	NUM
elements-9012	320	23	4	4	NUM
elements-9012	320	24	0	0	NUM
elements-9012	320	25	.	.	PUNCT
elements-9012	321	1	3	3	NUM
elements-9012	321	2	3	3	NUM
elements-9012	321	3	9	9	NUM
elements-9012	321	4	8	8	NUM
elements-9012	321	5	<	<	X
elements-9012	321	6	e	e	X
elements-9012	321	7	2	2	NUM
elements-9012	321	8	7	7	NUM
elements-9012	321	9	6	6	NUM
elements-9012	321	10	25	25	NUM
elements-9012	321	11	opossum	opossum	NOUN
elements-9012	321	12	platypus	platypus	NOUN
elements-9012	321	13	0	0	NUM
elements-9012	321	14	.	.	PUNCT
elements-9012	322	1	4	4	NUM
elements-9012	322	2	9	9	NUM
elements-9012	322	3	9	9	NUM
elements-9012	322	4	2	2	NUM
elements-9012	322	5	9	9	NUM
elements-9012	322	6	7	7	NUM
elements-9012	322	7	8	8	NUM
elements-9012	322	8	6	6	NUM
elements-9012	322	9	0	0	NUM
elements-9012	322	10	.2701	.2701	PROPN
elements-9012	322	11	3	3	NUM
elements-9012	322	12	.	.	PUNCT
elements-9012	322	13	3	3	NUM
elements-9012	322	14	e	e	NOUN
elements-9012	322	15	1	1	NUM
elements-9012	322	16	6	6	NUM
elements-9012	322	17	3	3	NUM
elements-9012	322	18	f	f	NOUN
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elements-9012	323	5	e	e	NOUN
elements-9012	323	6	2	2	NUM
elements-9012	323	7	:	:	PUNCT
elements-9012	323	8	a	a	DET
elements-9012	323	9	u	u	NOUN
elements-9012	323	10	t	t	X
elements-9012	323	11	o	o	NOUN
elements-9012	324	1	c	c	NOUN
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elements-9012	325	5	a	a	NOUN
elements-9012	325	6	t	t	NOUN
elements-9012	326	1	i	i	PRON
elements-9012	326	2	o	o	VERB
elements-9012	327	1	n	n	X
elements-9012	328	1	o	o	X
elements-9012	328	2	f	f	NOUN
elements-9012	328	3	n	n	CCONJ
elements-9012	328	4	o	o	NOUN
elements-9012	328	5	r	r	NOUN
elements-9012	328	6	m	m	VERB
elements-9012	328	7	a	a	DET
elements-9012	328	8	l	l	NOUN
elements-9012	329	1	i	i	NOUN
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elements-9012	329	3	e	e	PROPN
elements-9012	330	1	d	d	X
elements-9012	330	2	s	s	VERB
elements-9012	330	3	u	u	PROPN
elements-9012	330	4	b	b	PROPN
elements-9012	330	5	s	s	ADP
elements-9012	330	6	t	t	X
elements-9012	330	7	i	i	PRON
elements-9012	330	8	t	t	VERB
elements-9012	330	9	u	u	NOUN
elements-9012	331	1	t	t	NOUN
elements-9012	332	1	i	i	PRON
elements-9012	332	2	o	o	VERB
elements-9012	333	1	n	n	CCONJ
elements-9012	333	2	r	r	NOUN
elements-9012	333	3	a	a	DET
elements-9012	333	4	t	t	NOUN
elements-9012	333	5	e	e	X
elements-9012	333	6	s	s	NOUN
elements-9012	333	7	f	f	X
elements-9012	333	8	o	o	NOUN
elements-9012	333	9	r	r	NOUN
elements-9012	334	1	p	p	NOUN
elements-9012	334	2	a	a	PRON
elements-9012	335	1	i	i	NOUN
elements-9012	335	2	r	r	VERB
elements-9012	335	3	w	w	NOUN
elements-9012	336	1	i	i	NOUN
elements-9012	336	2	s	s	PROPN
elements-9012	336	3	e	e	NOUN
elements-9012	336	4	a	a	PRON
elements-9012	336	5	n	n	NOUN
elements-9012	336	6	a	a	DET
elements-9012	336	7	l	l	NOUN
elements-9012	337	1	y	y	NOUN
elements-9012	337	2	s	s	PROPN
elements-9012	337	3	e	e	NOUN
elements-9012	337	4	s	s	X
elements-9012	337	5	f	f	X
elements-9012	337	6	i	i	PRON
elements-9012	337	7	g	g	NOUN
elements-9012	337	8	u	u	NOUN
elements-9012	337	9	r	r	NOUN
elements-9012	337	10	e	e	NOUN
elements-9012	337	11	3	3	NUM
elements-9012	337	12	:	:	PUNCT
elements-9012	337	13	c	c	NOUN
elements-9012	337	14	a	a	PRON
elements-9012	337	15	i	i	NOUN
elements-9012	337	16	v	v	NOUN
elements-9012	337	17	s	s	NOUN
elements-9012	337	18	.	.	PUNCT
elements-9012	338	1	n	n	CCONJ
elements-9012	339	1	o	o	NOUN
elements-9012	339	2	r	r	NOUN
elements-9012	339	3	m	m	VERB
elements-9012	339	4	a	a	DET
elements-9012	339	5	l	l	NOUN
elements-9012	340	1	i	i	NOUN
elements-9012	340	2	z	z	NOUN
elements-9012	340	3	e	e	PROPN
elements-9012	341	1	d	d	X
elements-9012	341	2	s	s	VERB
elements-9012	341	3	u	u	PROPN
elements-9012	341	4	b	b	PROPN
elements-9012	341	5	s	s	ADP
elements-9012	341	6	t	t	X
elements-9012	341	7	i	i	PRON
elements-9012	341	8	t	t	VERB
elements-9012	341	9	u	u	NOUN
elements-9012	342	1	t	t	NOUN
elements-9012	343	1	i	i	PRON
elements-9012	343	2	o	o	VERB
elements-9012	344	1	n	n	CCONJ
elements-9012	344	2	r	r	NOUN
elements-9012	344	3	a	a	DET
elements-9012	344	4	t	t	NOUN
elements-9012	344	5	e	e	X
elements-9012	344	6	s	s	PROPN
elements-9012	344	7	f	f	X
elements-9012	345	1	o	o	NOUN
elements-9012	345	2	r	r	NOUN
elements-9012	345	3	sensu	sensu	NOUN
elements-9012	345	4	stricto	stricto	X
elements-9012	345	5	y	y	PROPN
elements-9012	345	6	e	e	PROPN
elements-9012	345	7	a	a	PRON
elements-9012	345	8	s	s	PROPN
elements-9012	345	9	t	t	NOUN
elements-9012	345	10	s	s	PROPN
elements-9012	345	11	a	a	DET
elements-9012	345	12	n	n	NOUN
elements-9012	345	13	d	d	PROPN
elements-9012	345	14	n.	n.	NOUN
elements-9012	345	15	crassa	crassa	NOUN
elements-9012	345	16	albicans	albican	VERB
elements-9012	345	17	a	a	DET
elements-9012	345	18	n	n	NOUN
elements-9012	345	19	d	d	PROPN
elements-9012	345	20	c.	c.	PROPN
elements-9012	345	21	dubliensis	dubliensis	PROPN
elements-9012	345	22	5	5	NUM
elements-9012	345	23	0	0	SYM
elements-9012	345	24	0	0	NUM
elements-9012	345	25	0	0	NUM
elements-9012	345	26	0	0	NUM
elements-9012	345	27	1	1	NUM
elements-9012	345	28	e	e	NOUN
elements-9012	345	29	+	+	NOUN
elements-9012	345	30	0	0	NUM
elements-9012	345	31	5	5	NUM
elements-9012	345	32	0	0	NUM
elements-9012	345	33	5	5	NUM
elements-9012	345	34	0	0	NUM
elements-9012	345	35	0	0	NUM
elements-9012	345	36	0	0	NUM
elements-9012	345	37	0	0	NUM
elements-9012	345	38	1	1	NUM
elements-9012	345	39	e	e	NOUN
elements-9012	345	40	+	+	NOUN
elements-9012	345	41	0	0	NUM
elements-9012	345	42	5	5	NUM
elements-9012	346	1	d	d	NOUN
elements-9012	346	2	i	i	PRON
elements-9012	346	3	s	s	PROPN
elements-9012	346	4	t	t	PROPN
elements-9012	346	5	a	a	PRON
elements-9012	346	6	n	n	NOUN
elements-9012	346	7	c	c	NOUN
elements-9012	346	8	e	e	X
elements-9012	346	9	b	b	PROPN
elements-9012	346	10	e	e	PROPN
elements-9012	346	11	t	t	PROPN
elements-9012	346	12	w	w	PROPN
elements-9012	346	13	e	e	PROPN
elements-9012	346	14	e	e	X
elements-9012	346	15	n	n	CCONJ
elements-9012	346	16	g	g	PROPN
elements-9012	346	17	e	e	PROPN
elements-9012	346	18	n	n	ADP
elements-9012	346	19	e	e	X
elements-9012	346	20	s	s	X
elements-9012	346	21	(	(	PUNCT
elements-9012	346	22	b	b	PROPN
elements-9012	346	23	p	p	NOUN
elements-9012	346	24	)	)	PUNCT
elements-9012	346	25	rates	rate	NOUN
elements-9012	346	26	are	be	AUX
elements-9012	346	27	not	not	PART
elements-9012	346	28	correlated	correlate	VERB
elements-9012	346	29	along	along	ADP
elements-9012	346	30	the	the	DET
elements-9012	346	31	genomes	genome	NOUN
elements-9012	346	32	in	in	ADP
elements-9012	346	33	figure	figure	NOUN
elements-9012	346	34	2a	2a	NUM
elements-9012	346	35	.	.	PUNCT
elements-9012	347	1	analagous	analagous	ADJ
elements-9012	347	2	graphs	graph	NOUN
elements-9012	347	3	were	be	AUX
elements-9012	347	4	produced	produce	VERB
elements-9012	347	5	for	for	ADP
elements-9012	347	6	all	all	DET
elements-9012	347	7	yeast	yeast	NOUN
elements-9012	347	8	species	specie	NOUN
elements-9012	347	9	pairs	pair	NOUN
elements-9012	347	10	,	,	PUNCT
elements-9012	347	11	each	each	PRON
elements-9012	347	12	indicating	indicate	VERB
elements-9012	347	13	no	no	DET
elements-9012	347	14	autocorrelation	autocorrelation	NOUN
elements-9012	347	15	along	along	ADP
elements-9012	347	16	the	the	DET
elements-9012	347	17	genome	genome	NOUN
elements-9012	347	18	,	,	PUNCT
elements-9012	347	19	except	except	SCONJ
elements-9012	347	20	for	for	ADP
elements-9012	347	21	the	the	DET
elements-9012	347	22	c.	c.	PROPN
elements-9012	347	23	dubli	dubli	PROPN
elements-9012	347	24	ensis	ensis	PROPN
elements-9012	347	25	/	/	SYM
elements-9012	347	26	c.	c.	PROPN
elements-9012	347	27	albicans	albican	VERB
elements-9012	347	28	a	a	DET
elements-9012	347	29	n	n	NOUN
elements-9012	347	30	d	d	PROPN
elements-9012	347	31	c.	c.	PROPN
elements-9012	347	32	tropicalis	tropicalis	PROPN
elements-9012	347	33	/	/	SYM
elements-9012	347	34	c.	c.	PROPN
elements-9012	347	35	dubliensis	dubliensis	PROPN
elements-9012	347	36	comparisons	comparison	NOUN
elements-9012	347	37	.	.	PUNCT
elements-9012	348	1	those	those	DET
elements-9012	348	2	pairs	pair	NOUN
elements-9012	348	3	show	show	VERB
elements-9012	348	4	that	that	SCONJ
elements-9012	348	5	rates	rate	NOUN
elements-9012	348	6	of	of	ADP
elements-9012	348	7	neighboring	neighboring	NOUN
elements-9012	348	8	genes	gene	NOUN
elements-9012	348	9	are	be	AUX
elements-9012	348	10	correlated	correlate	VERB
elements-9012	348	11	up	up	ADP
elements-9012	348	12	to	to	ADP
elements-9012	348	13	50	50	NUM
elements-9012	348	14	kb	kb	NOUN
elements-9012	348	15	apart	apart	ADV
elements-9012	348	16	.	.	PUNCT
elements-9012	349	1	f	f	X
elements-9012	350	1	i	i	PRON
elements-9012	350	2	g	g	VERB
elements-9012	350	3	u	u	VERB
elements-9012	350	4	r	r	NOUN
elements-9012	350	5	e	e	NOUN
elements-9012	350	6	4	4	NUM
elements-9012	350	7	:	:	PUNCT
elements-9012	350	8	a	a	DET
elements-9012	350	9	u	u	NOUN
elements-9012	350	10	t	t	X
elements-9012	350	11	o	o	NOUN
elements-9012	351	1	c	c	NOUN
elements-9012	351	2	o	o	NOUN
elements-9012	352	1	r	r	NOUN
elements-9012	352	2	r	r	NOUN
elements-9012	352	3	e	e	NOUN
elements-9012	352	4	l	l	NOUN
elements-9012	352	5	a	a	NOUN
elements-9012	352	6	t	t	NOUN
elements-9012	353	1	i	i	PRON
elements-9012	353	2	o	o	VERB
elements-9012	354	1	n	n	X
elements-9012	355	1	o	o	X
elements-9012	355	2	f	f	NOUN
elements-9012	355	3	n	n	CCONJ
elements-9012	355	4	o	o	NOUN
elements-9012	355	5	r	r	NOUN
elements-9012	355	6	m	m	VERB
elements-9012	355	7	a	a	DET
elements-9012	355	8	l	l	NOUN
elements-9012	356	1	i	i	NOUN
elements-9012	356	2	z	z	NOUN
elements-9012	356	3	e	e	PROPN
elements-9012	357	1	d	d	X
elements-9012	357	2	s	s	VERB
elements-9012	357	3	u	u	PROPN
elements-9012	357	4	b	b	PROPN
elements-9012	357	5	s	s	ADP
elements-9012	357	6	t	t	X
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elements-9012	357	8	t	t	VERB
elements-9012	357	9	u	u	NOUN
elements-9012	358	1	t	t	NOUN
elements-9012	359	1	i	i	PRON
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elements-9012	360	1	n	n	CCONJ
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elements-9012	360	3	a	a	DET
elements-9012	360	4	t	t	NOUN
elements-9012	361	1	e	e	NOUN
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elements-9012	361	3	a	a	DET
elements-9012	361	4	t	t	NOUN
elements-9012	361	5	4	4	NUM
elements-9012	361	6	f	f	NOUN
elements-9012	361	7	o	o	NOUN
elements-9012	361	8	l	l	NOUN
elements-9012	362	1	d	d	X
elements-9012	362	2	d	d	X
elements-9012	362	3	e	e	X
elements-9012	362	4	g	g	PROPN
elements-9012	362	5	e	e	PROPN
elements-9012	362	6	n	n	X
elements-9012	362	7	e	e	NOUN
elements-9012	362	8	r	r	NOUN
elements-9012	362	9	a	a	DET
elements-9012	362	10	t	t	NOUN
elements-9012	362	11	e	e	X
elements-9012	362	12	s	s	X
elements-9012	362	13	i	i	INTJ
elements-9012	362	14	t	t	NOUN
elements-9012	362	15	e	e	X
elements-9012	362	16	s	s	X
elements-9012	362	17	cai	cai	X
elements-9012	362	18	vs.	vs.	ADP
elements-9012	362	19	z	z	NOUN
elements-9012	362	20	-	-	PUNCT
elements-9012	362	21	score	score	NOUN
elements-9012	362	22	for	for	ADP
elements-9012	362	23	n.	n.	NOUN
elements-9012	362	24	crassa	crassa	PROPN
elements-9012	363	1	c	c	NOUN
elements-9012	364	1	o	o	X
elements-9012	364	2	w	w	NOUN
elements-9012	364	3	a	a	DET
elements-9012	364	4	n	n	NOUN
elements-9012	365	1	d	d	NOUN
elements-9012	365	2	d	d	NOUN
elements-9012	365	3	o	o	X
elements-9012	365	4	g	g	PROPN
elements-9012	365	5	a	a	DET
elements-9012	365	6	u	u	NOUN
elements-9012	365	7	t	t	NOUN
elements-9012	365	8	o	o	NOUN
elements-9012	366	1	c	c	NOUN
elements-9012	366	2	o	o	NOUN
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elements-9012	367	4	l	l	NOUN
elements-9012	367	5	a	a	NOUN
elements-9012	367	6	t	t	NOUN
elements-9012	368	1	i	i	PRON
elements-9012	368	2	o	o	VERB
elements-9012	369	1	n	n	CCONJ
elements-9012	369	2	h	h	NOUN
elements-9012	370	1	u	u	NOUN
elements-9012	370	2	m	m	VERB
elements-9012	370	3	a	a	PRON
elements-9012	370	4	n	n	ADV
elements-9012	370	5	a	a	DET
elements-9012	370	6	n	n	NOUN
elements-9012	370	7	d	d	NOUN
elements-9012	370	8	m	m	PROPN
elements-9012	370	9	a	a	DET
elements-9012	370	10	c	c	NOUN
elements-9012	370	11	a	a	DET
elements-9012	370	12	q	q	X
elements-9012	370	13	u	u	NOUN
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elements-9012	370	15	a	a	DET
elements-9012	370	16	u	u	NOUN
elements-9012	370	17	t	t	X
elements-9012	371	1	o	o	NOUN
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elements-9012	372	1	o	o	NOUN
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elements-9012	372	7	t	t	NOUN
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elements-9012	373	4	1.2	1.2	NUM
elements-9012	373	5	5e+06	5e+06	NUM
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elements-9012	373	7	e	e	NOUN
elements-9012	373	8	+	+	CCONJ
elements-9012	373	9	0	0	NUM
elements-9012	373	10	7	7	NUM
elements-9012	373	11	1.5e+07	1.5e+07	NUM
elements-9012	373	12	0	0	NUM
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elements-9012	374	3	d	d	X
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elements-9012	375	7	e	e	X
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elements-9012	375	13	e	e	X
elements-9012	375	14	n	n	CCONJ
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elements-9012	375	17	n	n	ADP
elements-9012	375	18	e	e	X
elements-9012	375	19	s	s	X
elements-9012	375	20	(	(	PUNCT
elements-9012	375	21	b	b	PROPN
elements-9012	375	22	p	p	NOUN
elements-9012	375	23	)	)	PUNCT
elements-9012	375	24	autocorrelation	autocorrelation	NOUN
elements-9012	375	25	for	for	ADP
elements-9012	375	26	normalized	normalize	VERB
elements-9012	375	27	substitution	substitution	NOUN
elements-9012	375	28	rates	rate	NOUN
elements-9012	375	29	at	at	ADP
elements-9012	375	30	4	4	NUM
elements-9012	375	31	-	-	ADJ
elements-9012	375	32	fold	fold	ADJ
elements-9012	375	33	degenerate	degenerate	ADJ
elements-9012	375	34	sites	site	NOUN
elements-9012	375	35	between	between	ADP
elements-9012	375	36	cow	cow	NOUN
elements-9012	375	37	and	and	CCONJ
elements-9012	375	38	doc	doc	PROPN
elements-9012	375	39	(	(	PUNCT
elements-9012	375	40	left	left	ADJ
elements-9012	375	41	)	)	PUNCT
elements-9012	375	42	and	and	CCONJ
elements-9012	375	43	human	human	NOUN
elements-9012	375	44	and	and	CCONJ
elements-9012	375	45	macaque	macaque	ADJ
elements-9012	375	46	(	(	PUNCT
elements-9012	375	47	right	right	NOUN
elements-9012	375	48	)	)	PUNCT
elements-9012	375	49	.	.	PUNCT
elements-9012	376	1	both	both	DET
elements-9012	376	2	graphs	graph	NOUN
elements-9012	376	3	show	show	VERB
elements-9012	376	4	correlation	correlation	NOUN
elements-9012	376	5	of	of	ADP
elements-9012	376	6	rates	rate	NOUN
elements-9012	376	7	for	for	ADP
elements-9012	376	8	genes	gene	NOUN
elements-9012	376	9	within	within	ADP
elements-9012	376	10	wmb	wmb	PROPN
elements-9012	376	11	of	of	ADP
elements-9012	376	12	each	each	DET
elements-9012	376	13	other	other	ADJ
elements-9012	376	14	.	.	PUNCT
elements-9012	377	1	z	z	X
elements-9012	377	2	-	-	PUNCT
elements-9012	377	3	score	score	NOUN
elements-9012	377	4	these	these	PRON
elements-9012	377	5	are	be	AUX
elements-9012	377	6	graphs	graph	NOUN
elements-9012	377	7	of	of	ADP
elements-9012	377	8	cai	cai	PROPN
elements-9012	377	9	vs.	vs.	ADP
elements-9012	377	10	normalized	normalize	VERB
elements-9012	377	11	substitution	substitution	NOUN
elements-9012	377	12	rates	rate	NOUN
elements-9012	377	13	for	for	ADP
elements-9012	377	14	sensu	sensu	NOUN
elements-9012	377	15	s	s	PART
elements-9012	378	1	t	t	NOUN
elements-9012	378	2	r	r	NOUN
elements-9012	379	1	i	i	NOUN
elements-9012	379	2	c	c	NOUN
elements-9012	380	1	t	t	X
elements-9012	380	2	o	o	X
elements-9012	380	3	y	y	PROPN
elements-9012	380	4	e	e	PROPN
elements-9012	380	5	a	a	PRON
elements-9012	380	6	s	s	X
elements-9012	380	7	t	t	NOUN
elements-9012	380	8	s	s	PART
elements-9012	380	9	(	(	PUNCT
elements-9012	380	10	s.	s.	PROPN
elements-9012	380	11	cerevisiae	cerevisiae	PROPN
elements-9012	380	12	,	,	PUNCT
elements-9012	380	13	s.	s.	PROPN
elements-9012	380	14	bayanus	bayanus	PROPN
elements-9012	380	15	,	,	PUNCT
elements-9012	380	16	s.	s.	PROPN
elements-9012	380	17	mikatae	mikatae	PROPN
elements-9012	380	18	,	,	PUNCT
elements-9012	380	19	s.	s.	PROPN
elements-9012	380	20	paradoxus	paradoxus	PROPN
elements-9012	380	21	)	)	PUNCT
elements-9012	380	22	(	(	PUNCT
elements-9012	380	23	bottom	bottom	NOUN
elements-9012	380	24	)	)	PUNCT
elements-9012	380	25	and	and	CCONJ
elements-9012	380	26	n.	n.	PROPN
elements-9012	380	27	crassa	crassa	PROPN
elements-9012	380	28	(	(	PUNCT
elements-9012	380	29	top	top	NOUN
elements-9012	380	30	)	)	PUNCT
elements-9012	380	31	.	.	PUNCT
elements-9012	381	1	in	in	ADP
elements-9012	381	2	each	each	DET
elements-9012	381	3	graph	graph	NOUN
elements-9012	381	4	,	,	PUNCT
elements-9012	381	5	there	there	PRON
elements-9012	381	6	is	be	VERB
elements-9012	381	7	a	a	DET
elements-9012	381	8	large	large	ADJ
elements-9012	381	9	cluster	cluster	NOUN
elements-9012	381	10	of	of	ADP
elements-9012	381	11	genes	gene	NOUN
elements-9012	381	12	with	with	ADP
elements-9012	381	13	lower	low	ADJ
elements-9012	381	14	cai	cai	PROPN
elements-9012	381	15	values	value	NOUN
elements-9012	381	16	and	and	CCONJ
elements-9012	381	17	substitution	substitution	NOUN
elements-9012	381	18	rates	rate	NOUN
elements-9012	381	19	distributed	distribute	VERB
elements-9012	381	20	around	around	ADP
elements-9012	381	21	zero	zero	NUM
elements-9012	381	22	.	.	PUNCT
elements-9012	382	1	each	each	DET
elements-9012	382	2	graph	graph	NOUN
elements-9012	382	3	also	also	ADV
elements-9012	382	4	contains	contain	VERB
elements-9012	382	5	a	a	DET
elements-9012	382	6	second	second	ADJ
elements-9012	382	7	group	group	NOUN
elements-9012	382	8	of	of	ADP
elements-9012	382	9	genes	gene	NOUN
elements-9012	382	10	with	with	ADP
elements-9012	382	11	higher	high	ADJ
elements-9012	382	12	cai	cai	NOUN
elements-9012	382	13	values	value	NOUN
elements-9012	382	14	and	and	CCONJ
elements-9012	382	15	lower	low	ADJ
elements-9012	382	16	substitution	substitution	NOUN
elements-9012	382	17	rates	rate	NOUN
elements-9012	382	18	,	,	PUNCT
elements-9012	382	19	and	and	CCONJ
elements-9012	382	20	such	such	ADJ
elements-9012	382	21	genes	gene	NOUN
elements-9012	382	22	are	be	AUX
elements-9012	382	23	l	l	NOUN
elements-9012	382	24	i	i	NOUN
elements-9012	383	1	k	k	PROPN
elements-9012	383	2	e	e	X
elements-9012	383	3	l	l	X
elements-9012	383	4	y	y	X
elements-9012	383	5	to	to	ADP
elements-9012	383	6	b	b	PROPN
elements-9012	383	7	e	e	X
elements-9012	383	8	under	under	ADP
elements-9012	383	9	codon	codon	NOUN
elements-9012	383	10	usage	usage	NOUN
elements-9012	383	11	selection	selection	NOUN
elements-9012	383	12	.	.	PUNCT
elements-9012	384	1	the	the	DET
elements-9012	384	2	sensu	sensu	NOUN
elements-9012	384	3	stricto	stricto	X
elements-9012	384	4	y	y	PROPN
elements-9012	384	5	e	e	PROPN
elements-9012	384	6	a	a	PRON
elements-9012	384	7	s	s	ADP
elements-9012	384	8	t	t	NOUN
elements-9012	384	9	s	s	X
elements-9012	384	10	have	have	VERB
elements-9012	384	11	727	727	NUM
elements-9012	384	12	genes	gene	NOUN
elements-9012	384	13	that	that	PRON
elements-9012	384	14	have	have	VERB
elements-9012	384	15	cai	cai	PROPN
elements-9012	384	16	values	value	NOUN
elements-9012	384	17	greater	great	ADJ
elements-9012	384	18	than	than	ADP
elements-9012	384	19	0.40	0.40	NUM
elements-9012	384	20	while	while	SCONJ
elements-9012	384	21	n.	n.	NOUN
elements-9012	384	22	crassa	crassa	PROPN
elements-9012	384	23	has	have	VERB
elements-9012	384	24	949	949	NUM
elements-9012	384	25	with	with	ADP
elements-9012	384	26	cai	cai	PROPN
elements-9012	384	27	values	value	NOUN
elements-9012	384	28	greater	great	ADJ
elements-9012	384	29	than	than	ADP
elements-9012	384	30	0.70	0.70	NUM
elements-9012	384	31	,	,	PUNCT
elements-9012	384	32	suggesting	suggest	VERB
elements-9012	384	33	that	that	SCONJ
elements-9012	384	34	more	more	ADJ
elements-9012	384	35	genes	gene	NOUN
elements-9012	384	36	a	a	DET
elements-9012	384	37	r	r	NOUN
elements-9012	384	38	e	e	NOUN
elements-9012	384	39	under	under	ADP
elements-9012	384	40	codon	codon	NOUN
elements-9012	384	41	usage	usage	NOUN
elements-9012	384	42	selection	selection	NOUN
elements-9012	384	43	in	in	ADP
elements-9012	384	44	n.	n.	PROPN
elements-9012	384	45	crassa	crassa	NOUN
elements-9012	384	46	.	.	PUNCT
elements-9012	385	1	the	the	DET
elements-9012	385	2	n.	n.	PROPN
elements-9012	385	3	crassa	crassa	NOUN
elements-9012	385	4	codon	codon	ADJ
elements-9012	385	5	usage	usage	NOUN
elements-9012	385	6	table	table	NOUN
elements-9012	385	7	was	be	AUX
elements-9012	385	8	used	use	VERB
elements-9012	385	9	to	to	PART
elements-9012	385	10	calculate	calculate	VERB
elements-9012	385	11	cai	cai	PROPN
elements-9012	385	12	values	value	NOUN
elements-9012	385	13	in	in	ADP
elements-9012	385	14	the	the	DET
elements-9012	385	15	top	top	ADJ
elements-9012	385	16	graph	graph	NOUN
elements-9012	385	17	and	and	CCONJ
elements-9012	385	18	the	the	DET
elements-9012	385	19	s.	s.	PROPN
elements-9012	385	20	cerevisiae	cerevisiae	PROPN
elements-9012	385	21	codon	codon	ADJ
elements-9012	385	22	usage	usage	NOUN
elements-9012	385	23	table	table	NOUN
elements-9012	385	24	was	be	AUX
elements-9012	385	25	used	use	VERB
elements-9012	385	26	to	to	PART
elements-9012	385	27	compute	compute	VERB
elements-9012	385	28	the	the	DET
elements-9012	385	29	cai	cai	PROPN
elements-9012	385	30	values	value	NOUN
elements-9012	385	31	in	in	ADP
elements-9012	385	32	the	the	DET
elements-9012	385	33	bottom	bottom	ADJ
elements-9012	385	34	graph	graph	NOUN
elements-9012	385	35	.	.	PUNCT
elements-9012	386	1	cai	cai	PROPN
elements-9012	386	2	vs.	vs.	ADP
elements-9012	386	3	z	z	NOUN
elements-9012	386	4	-	-	PUNCT
elements-9012	386	5	score	score	NOUN
elements-9012	386	6	for	for	ADP
elements-9012	386	7	sensu	sensu	NOUN
elements-9012	386	8	stricto	stricto	X
elements-9012	386	9	yeast	yeast	NOUN
elements-9012	386	10	z	z	NOUN
elements-9012	386	11	-	-	PUNCT
elements-9012	386	12	score	score	NOUN
elements-9012	387	1	f	f	NOUN
elements-9012	388	1	i	i	PRON
elements-9012	388	2	g	g	NOUN
elements-9012	388	3	u	u	NOUN
elements-9012	388	4	r	r	NOUN
elements-9012	388	5	e	e	NOUN
elements-9012	388	6	5	5	NUM
elements-9012	388	7	:	:	PUNCT
elements-9012	388	8	c	c	NOUN
elements-9012	388	9	o	o	NOUN
elements-9012	389	1	m	m	VERB
elements-9012	389	2	p	p	NOUN
elements-9012	390	1	a	a	DET
elements-9012	390	2	r	r	NOUN
elements-9012	391	1	i	i	PRON
elements-9012	391	2	s	s	VERB
elements-9012	391	3	o	o	NOUN
elements-9012	392	1	n	n	ADP
elements-9012	392	2	o	o	X
elements-9012	393	1	f	f	NOUN
elements-9012	393	2	t	t	NOUN
elements-9012	393	3	h	h	NOUN
elements-9012	394	1	e	e	PROPN
elements-9012	394	2	w	w	NOUN
elements-9012	394	3	i	i	NOUN
elements-9012	394	4	d	d	NOUN
elements-9012	394	5	t	t	PROPN
elements-9012	394	6	h	h	NOUN
elements-9012	394	7	s	s	PART
elements-9012	394	8	o	o	X
elements-9012	394	9	f	f	X
elements-9012	394	10	t	t	NOUN
elements-9012	394	11	h	h	NOUN
elements-9012	394	12	e	e	NOUN
elements-9012	394	13	r	r	NOUN
elements-9012	394	14	a	a	DET
elements-9012	394	15	t	t	NOUN
elements-9012	394	16	e	e	X
elements-9012	395	1	d	d	X
elements-9012	395	2	i	i	PRON
elements-9012	395	3	s	s	VERB
elements-9012	395	4	t	t	NOUN
elements-9012	396	1	r	r	NOUN
elements-9012	397	1	i	i	NOUN
elements-9012	398	1	b	b	PROPN
elements-9012	399	1	u	u	NOUN
elements-9012	400	1	t	t	X
elements-9012	401	1	i	i	PRON
elements-9012	401	2	o	o	VERB
elements-9012	401	3	n	n	X
elements-9012	401	4	s	s	NOUN
elements-9012	401	5	m	m	VERB
elements-9012	401	6	a	a	DET
elements-9012	401	7	m	m	NOUN
elements-9012	401	8	m	m	VERB
elements-9012	401	9	a	a	DET
elements-9012	401	10	l	l	NOUN
elements-9012	401	11	s	s	PART
elements-9012	401	12	u	u	NOUN
elements-9012	402	1	n	n	NOUN
elements-9012	403	1	i	i	PRON
elements-9012	403	2	f	f	NOUN
elements-9012	404	1	o	o	NOUN
elements-9012	404	2	r	r	NOUN
elements-9012	404	3	m	m	VERB
elements-9012	404	4	n	n	ADV
elements-9012	404	5	o	o	NOUN
elements-9012	404	6	n	n	CCONJ
elements-9012	404	7	u	u	NOUN
elements-9012	405	1	n	n	CCONJ
elements-9012	406	1	i	i	PRON
elements-9012	406	2	f	f	NOUN
elements-9012	407	1	o	o	NOUN
elements-9012	407	2	r	r	NOUN
elements-9012	407	3	m	m	VERB
elements-9012	407	4	y	y	NOUN
elements-9012	407	5	e	e	PROPN
elements-9012	407	6	a	a	DET
elements-9012	407	7	s	s	X
elements-9012	407	8	t	t	X
elements-9012	407	9	y	y	PROPN
elements-9012	407	10	e	e	PROPN
elements-9012	407	11	a	a	DET
elements-9012	407	12	s	s	ADP
elements-9012	407	13	t	t	NOUN
elements-9012	407	14	the	the	DET
elements-9012	407	15	blue	blue	ADJ
elements-9012	407	16	line	line	NOUN
elements-9012	407	17	at	at	ADP
elements-9012	407	18	s	s	NOUN
elements-9012	407	19	=	=	SYM
elements-9012	407	20	1	1	NUM
elements-9012	407	21	shows	show	VERB
elements-9012	407	22	the	the	DET
elements-9012	407	23	expected	expect	VERB
elements-9012	407	24	standard	standard	ADJ
elements-9012	407	25	deviation	deviation	NOUN
elements-9012	407	26	for	for	ADP
elements-9012	407	27	a	a	DET
elements-9012	407	28	normal	normal	ADJ
elements-9012	407	29	distribution	distribution	NOUN
elements-9012	407	30	.	.	PUNCT
elements-9012	408	1	the	the	DET
elements-9012	408	2	mammals	mammal	NOUN
elements-9012	408	3	have	have	VERB
elements-9012	408	4	an	an	DET
elements-9012	408	5	average	average	ADJ
elements-9012	408	6	standard	standard	ADJ
elements-9012	408	7	deviation	deviation	NOUN
elements-9012	408	8	of	of	ADP
elements-9012	408	9	2.056	2.056	NUM
elements-9012	408	10	,	,	PUNCT
elements-9012	408	11	indicating	indicate	VERB
elements-9012	408	12	their	their	PRON
elements-9012	408	13	bias	bias	NOUN
elements-9012	408	14	toward	toward	ADP
elements-9012	408	15	both	both	CCONJ
elements-9012	408	16	high	high	ADJ
elements-9012	408	17	and	and	CCONJ
elements-9012	408	18	low	low	ADJ
elements-9012	408	19	rates	rate	NOUN
elements-9012	408	20	.	.	PUNCT
elements-9012	409	1	there	there	PRON
elements-9012	409	2	are	be	VERB
elements-9012	409	3	three	three	NUM
elements-9012	409	4	yeast	yeast	NOUN
elements-9012	409	5	pairwise	pairwise	NOUN
elements-9012	409	6	comparisons	comparison	NOUN
elements-9012	409	7	(	(	PUNCT
elements-9012	409	8	c.	c.	NOUN
elements-9012	409	9	dubliniensis	dubliniensis	NOUN
elements-9012	409	10	/	/	SYM
elements-9012	409	11	c.	c.	NOUN
elements-9012	409	12	albicans	albican	NOUN
elements-9012	409	13	,	,	PUNCT
elements-9012	409	14	c.	c.	PROPN
elements-9012	409	15	tropicalis	tropicalis	PROPN
elements-9012	409	16	/	/	SYM
elements-9012	409	17	c.	c.	PROPN
elements-9012	409	18	dubli	dubli	VERB
elements-9012	409	19	n	n	ADV
elements-9012	409	20	i	i	NOUN
elements-9012	409	21	e	e	PROPN
elements-9012	409	22	n	n	PRON
elements-9012	409	23	sis	sis	NOUN
elements-9012	409	24	,	,	PUNCT
elements-9012	409	25	and	and	CCONJ
elements-9012	409	26	n.	n.	NOUN
elements-9012	409	27	crassa	crassa	PROPN
elements-9012	409	28	/	/	SYM
elements-9012	409	29	c	c	NOUN
elements-9012	409	30	globosum	globosum	NOUN
elements-9012	409	31	)	)	PUNCT
elements-9012	409	32	which	which	PRON
elements-9012	409	33	have	have	VERB
elements-9012	409	34	unusually	unusually	ADV
elements-9012	409	35	wide	wide	ADJ
elements-9012	409	36	rate	rate	NOUN
elements-9012	409	37	distributions	distribution	NOUN
elements-9012	409	38	(	(	PUNCT
elements-9012	409	39	see	see	VERB
elements-9012	409	40	figure	figure	NOUN
elements-9012	409	41	lc	lc	PROPN
elements-9012	409	42	)	)	PUNCT
elements-9012	409	43	.	.	PUNCT
elements-9012	410	1	for	for	ADP
elements-9012	410	2	these	these	DET
elements-9012	410	3	comparisons	comparison	NOUN
elements-9012	410	4	,	,	PUNCT
elements-9012	410	5	the	the	DET
elements-9012	410	6	average	average	NOUN
elements-9012	410	7	s	s	VERB
elements-9012	410	8	is	be	AUX
elements-9012	410	9	1.825	1.825	NUM
elements-9012	410	10	.	.	PUNCT
elements-9012	411	1	the	the	DET
elements-9012	411	2	remaining	remain	VERB
elements-9012	411	3	22	22	NUM
elements-9012	411	4	yeasts	yeast	NOUN
elements-9012	411	5	have	have	VERB
elements-9012	411	6	a	a	DET
elements-9012	411	7	noticeably	noticeably	ADV
elements-9012	411	8	lower	low	ADJ
elements-9012	411	9	value	value	NOUN
elements-9012	411	10	of	of	ADP
elements-9012	411	11	s	s	PRON
elements-9012	411	12	(	(	PUNCT
elements-9012	411	13	average	average	ADJ
elements-9012	411	14	s=7.32	s=7.32	NOUN
elements-9012	411	15	,	,	PUNCT
elements-9012	411	16	rance	rance	NOUN
elements-9012	411	17	=	=	PUNCT
elements-9012	412	1	[	[	X
elements-9012	412	2	1.021.40	1.021.40	NUM
elements-9012	412	3	]	]	X
elements-9012	412	4	,	,	PUNCT
elements-9012	412	5	consistent	consistent	ADJ
elements-9012	412	6	with	with	ADP
elements-9012	412	7	their	their	PRON
elements-9012	412	8	having	have	VERB
elements-9012	412	9	m	m	NOUN
elements-9012	412	10	compositions	composition	NOUN
elements-9012	412	11	of	of	ADP
elements-9012	412	12	these	these	DET
elements-9012	412	13	species	specie	NOUN
elements-9012	412	14	(	(	PUNCT
elements-9012	412	15	see	see	VERB
elements-9012	412	16	methods	method	NOUN
elements-9012	412	17	)	)	PUNCT
elements-9012	412	18	.	.	PUNCT
elements-9012	413	1	the	the	DET
elements-9012	413	2	inference	inference	NOUN
elements-9012	413	3	that	that	SCONJ
elements-9012	413	4	there	there	PRON
elements-9012	413	5	is	be	VERB
elements-9012	413	6	no	no	DET
elements-9012	413	7	neutral	neutral	ADJ
elements-9012	413	8	mutational	mutational	ADJ
elements-9012	413	9	variation	variation	NOUN
elements-9012	413	10	within	within	ADP
elements-9012	413	11	these	these	DET
elements-9012	413	12	22	22	NUM
elements-9012	413	13	yeast	yeast	NOUN
elements-9012	413	14	genomes	genome	NOUN
elements-9012	413	15	is	be	AUX
elements-9012	413	16	further	far	ADV
elements-9012	413	17	supported	support	VERB
elements-9012	413	18	by	by	ADP
elements-9012	413	19	a	a	DET
elements-9012	413	20	neighboring	neighboring	NOUN
elements-9012	413	21	gene	gene	NOUN
elements-9012	413	22	analysis	analysis	NOUN
elements-9012	413	23	.	.	PUNCT
elements-9012	414	1	for	for	ADP
elements-9012	414	2	each	each	DET
elements-9012	414	3	pair	pair	NOUN
elements-9012	414	4	of	of	ADP
elements-9012	414	5	species	specie	NOUN
elements-9012	414	6	,	,	PUNCT
elements-9012	414	7	the	the	DET
elements-9012	414	8	pearson	pearson	NOUN
elements-9012	414	9	correlation	correlation	NOUN
elements-9012	414	10	was	be	AUX
elements-9012	414	11	calculated	calculate	VERB
elements-9012	414	12	for	for	ADP
elements-9012	414	13	the	the	DET
elements-9012	414	14	rates	rate	NOUN
elements-9012	414	15	of	of	ADP
elements-9012	414	16	substitution	substitution	NOUN
elements-9012	414	17	of	of	ADP
elements-9012	414	18	neighboring	neighboring	NOUN
elements-9012	414	19	genes	gene	NOUN
elements-9012	414	20	(	(	PUNCT
elements-9012	414	21	see	see	VERB
elements-9012	414	22	methods	method	NOUN
elements-9012	414	23	)	)	PUNCT
elements-9012	414	24	.	.	PUNCT
elements-9012	415	1	these	these	DET
elements-9012	415	2	results	result	NOUN
elements-9012	415	3	are	be	AUX
elements-9012	415	4	shown	show	VERB
elements-9012	415	5	in	in	ADP
elements-9012	415	6	table	table	NOUN
elements-9012	415	7	1	1	NUM
elements-9012	415	8	,	,	PUNCT
elements-9012	415	9	with	with	ADP
elements-9012	415	10	the	the	DET
elements-9012	415	11	22	22	NUM
elements-9012	415	12	homogeneous	homogeneous	ADJ
elements-9012	415	13	genomes	genome	NOUN
elements-9012	415	14	shown	show	VERB
elements-9012	415	15	in	in	ADP
elements-9012	415	16	black	black	ADJ
elements-9012	415	17	.	.	PUNCT
elements-9012	416	1	there	there	PRON
elements-9012	416	2	was	be	VERB
elements-9012	416	3	no	no	DET
elements-9012	416	4	significant	significant	ADJ
elements-9012	416	5	correlation	correlation	NOUN
elements-9012	416	6	for	for	ADP
elements-9012	416	7	any	any	PRON
elements-9012	416	8	of	of	ADP
elements-9012	416	9	the	the	DET
elements-9012	416	10	species	species	NOUN
elements-9012	416	11	pairs	pair	NOUN
elements-9012	416	12	made	make	VERB
elements-9012	416	13	up	up	ADP
elements-9012	416	14	of	of	ADP
elements-9012	416	15	two	two	NUM
elements-9012	416	16	members	member	NOUN
elements-9012	416	17	from	from	ADP
elements-9012	416	18	these	these	DET
elements-9012	416	19	22	22	NUM
elements-9012	416	20	genomes	genome	NOUN
elements-9012	416	21	.	.	PUNCT
elements-9012	417	1	the	the	DET
elements-9012	417	2	same	same	ADJ
elements-9012	417	3	results	result	NOUN
elements-9012	417	4	were	be	AUX
elements-9012	417	5	obtained	obtain	VERB
elements-9012	417	6	irrespective	irrespective	ADV
elements-9012	417	7	of	of	ADP
elements-9012	417	8	which	which	DET
elements-9012	417	9	species	specie	NOUN
elements-9012	417	10	in	in	ADP
elements-9012	417	11	the	the	DET
elements-9012	417	12	pair	pair	NOUN
elements-9012	417	13	was	be	AUX
elements-9012	417	14	used	use	VERB
elements-9012	417	15	to	to	PART
elements-9012	417	16	specify	specify	VERB
elements-9012	417	17	gene	gene	NOUN
elements-9012	417	18	locations	location	NOUN
elements-9012	417	19	.	.	PUNCT
elements-9012	418	1	more	more	ADV
elements-9012	418	2	generally	generally	ADV
elements-9012	418	3	,	,	PUNCT
elements-9012	418	4	an	an	DET
elements-9012	418	5	autocorrelation	autocorrelation	NOUN
elements-9012	418	6	function	function	NOUN
elements-9012	418	7	<	<	X
elements-9012	418	8	r	r	NOUN
elements-9012	418	9	x	x	PUNCT
elements-9012	418	10	^r(o	^r(o	NUM
elements-9012	418	11	)	)	PUNCT
elements-9012	418	12	>	>	X
elements-9012	418	13	was	be	AUX
elements-9012	418	14	computed	compute	VERB
elements-9012	418	15	,	,	PUNCT
elements-9012	418	16	where	where	SCONJ
elements-9012	418	17	r(o	r(o	NOUN
elements-9012	418	18	)	)	PUNCT
elements-9012	418	19	is	be	AUX
elements-9012	418	20	the	the	DET
elements-9012	418	21	normalized	normalize	VERB
elements-9012	418	22	substitution	substitution	NOUN
elements-9012	418	23	rate	rate	NOUN
elements-9012	418	24	of	of	ADP
elements-9012	418	25	a	a	DET
elements-9012	418	26	gene	gene	NOUN
elements-9012	418	27	and	and	CCONJ
elements-9012	418	28	r	r	NOUN
elements-9012	418	29	1	1	NUM
elements-9012	418	30	^	^	PUNCT
elements-9012	418	31	is	be	AUX
elements-9012	418	32	the	the	DET
elements-9012	418	33	normalized	normalize	VERB
elements-9012	418	34	substitution	substitution	NOUN
elements-9012	418	35	rate	rate	NOUN
elements-9012	418	36	of	of	ADP
elements-9012	418	37	a	a	DET
elements-9012	418	38	gene	gene	NOUN
elements-9012	418	39	that	that	PRON
elements-9012	418	40	is	be	AUX
elements-9012	418	41	x	x	PART
elements-9012	418	42	base	base	NOUN
elements-9012	418	43	pairs	pair	NOUN
elements-9012	418	44	downstream	downstream	ADV
elements-9012	418	45	of	of	ADP
elements-9012	418	46	the	the	DET
elements-9012	418	47	first	first	ADJ
elements-9012	418	48	gene.x	gene.x	PROPN
elements-9012	418	49	x	x	SYM
elements-9012	418	50	the	the	DET
elements-9012	418	51	rates	rate	NOUN
elements-9012	418	52	are	be	AUX
elements-9012	418	53	normalized	normalize	VERB
elements-9012	418	54	around	around	ADP
elements-9012	418	55	r	r	NOUN
elements-9012	418	56	=	=	SYM
elements-9012	418	57	0	0	NUM
elements-9012	419	1	so	so	ADV
elements-9012	419	2	we	we	PRON
elements-9012	419	3	would	would	AUX
elements-9012	419	4	expect	expect	VERB
elements-9012	419	5	<	<	X
elements-9012	419	6	r	r	X
elements-9012	419	7	(	(	PUNCT
elements-9012	419	8	o	o	NOUN
elements-9012	419	9	)	)	PUNCT
elements-9012	419	10	r	r	NOUN
elements-9012	419	11	l	l	NOUN
elements-9012	419	12	6	6	NUM
elements-9012	419	13	>	>	PUNCT
elements-9012	419	14	~	~	PUNCT
elements-9012	419	15	o	o	X
elements-9012	420	1	i	i	PRON
elements-9012	420	2	f	f	VERB
elements-9012	420	3	there	there	PRON
elements-9012	420	4	are	be	VERB
elements-9012	420	5	no	no	DET
elements-9012	420	6	regional	regional	ADJ
elements-9012	420	7	rate	rate	NOUN
elements-9012	420	8	biases	bias	NOUN
elements-9012	420	9	(	(	PUNCT
elements-9012	420	10	see	see	VERB
elements-9012	420	11	methods	method	NOUN
elements-9012	420	12	)	)	PUNCT
elements-9012	420	13	.	.	PUNCT
elements-9012	421	1	the	the	DET
elements-9012	421	2	autocorrelation	autocorrelation	NOUN
elements-9012	421	3	reveals	reveal	VERB
elements-9012	421	4	no	no	DET
elements-9012	421	5	significant	significant	ADJ
elements-9012	421	6	correlation	correlation	NOUN
elements-9012	421	7	between	between	ADP
elements-9012	421	8	neighboring	neighboring	NOUN
elements-9012	421	9	genes	gene	NOUN
elements-9012	421	10	at	at	ADP
elements-9012	421	11	any	any	DET
elements-9012	421	12	finite	finite	ADJ
elements-9012	421	13	separation	separation	NOUN
elements-9012	421	14	(	(	PUNCT
elements-9012	421	15	figure	figure	NOUN
elements-9012	421	16	2a	2a	NUM
elements-9012	421	17	)	)	PUNCT
elements-9012	421	18	.	.	PUNCT
elements-9012	422	1	this	this	PRON
elements-9012	422	2	shows	show	VERB
elements-9012	422	3	that	that	SCONJ
elements-9012	422	4	these	these	DET
elements-9012	422	5	yeast	yeast	NOUN
elements-9012	422	6	species	specie	NOUN
elements-9012	422	7	do	do	AUX
elements-9012	422	8	not	not	PART
elements-9012	422	9	demonstrate	demonstrate	VERB
elements-9012	422	10	the	the	DET
elements-9012	422	11	heterogeneous	heterogeneous	ADJ
elements-9012	422	12	mutation	mutation	NOUN
elements-9012	422	13	rates	rate	NOUN
elements-9012	422	14	that	that	PRON
elements-9012	422	15	have	have	AUX
elements-9012	422	16	been	be	AUX
elements-9012	422	17	observed	observe	VERB
elements-9012	422	18	in	in	ADP
elements-9012	422	19	mammals	mammal	NOUN
elements-9012	422	20	.	.	PUNCT
elements-9012	423	1	species	specie	NOUN
elements-9012	423	2	pairs	pair	NOUN
elements-9012	423	3	from	from	ADP
elements-9012	423	4	two	two	NUM
elements-9012	423	5	subclades	subclade	NOUN
elements-9012	423	6	[	[	X
elements-9012	423	7	(	(	PUNCT
elements-9012	423	8	c.	c.	NOUN
elements-9012	423	9	albicans	albican	NOUN
elements-9012	423	10	,	,	PUNCT
elements-9012	423	11	c.	c.	NOUN
elements-9012	423	12	dubliniensis	dubliniensis	NOUN
elements-9012	423	13	,	,	PUNCT
elements-9012	423	14	c.	c.	PROPN
elements-9012	423	15	tropicalis	tropicalis	PROPN
elements-9012	423	16	)	)	PUNCT
elements-9012	423	17	and	and	CCONJ
elements-9012	423	18	(	(	PUNCT
elements-9012	423	19	n.	n.	NOUN
elements-9012	423	20	crassa	crassa	PROPN
elements-9012	423	21	,	,	PUNCT
elements-9012	423	22	c.	c.	PROPN
elements-9012	423	23	globosum	globosum	PROPN
elements-9012	423	24	)	)	PUNCT
elements-9012	423	25	]	]	PUNCT
elements-9012	423	26	were	be	AUX
elements-9012	423	27	found	find	VERB
elements-9012	423	28	to	to	PART
elements-9012	423	29	have	have	VERB
elements-9012	423	30	distributions	distribution	NOUN
elements-9012	423	31	that	that	PRON
elements-9012	423	32	did	do	AUX
elements-9012	423	33	not	not	PART
elements-9012	423	34	fit	fit	VERB
elements-9012	423	35	the	the	DET
elements-9012	423	36	normal	normal	ADJ
elements-9012	423	37	curve	curve	NOUN
elements-9012	423	38	.	.	PUNCT
elements-9012	424	1	these	these	DET
elements-9012	424	2	species	specie	NOUN
elements-9012	424	3	have	have	VERB
elements-9012	424	4	substitution	substitution	NOUN
elements-9012	424	5	rate	rate	NOUN
elements-9012	424	6	distributions	distribution	NOUN
elements-9012	424	7	biased	bias	VERB
elements-9012	424	8	toward	toward	ADP
elements-9012	424	9	high	high	ADJ
elements-9012	424	10	and	and	CCONJ
elements-9012	424	11	low	low	ADJ
elements-9012	424	12	rates	rate	NOUN
elements-9012	424	13	(	(	PUNCT
elements-9012	424	14	2.07	2.07	NUM
elements-9012	424	15	<	<	X
elements-9012	424	16	s	s	X
elements-9012	424	17	<	<	X
elements-9012	424	18	2.20	2.20	NUM
elements-9012	424	19	)	)	PUNCT
elements-9012	424	20	(	(	PUNCT
elements-9012	424	21	figure	figure	NOUN
elements-9012	424	22	ic	ic	PROPN
elements-9012	424	23	)	)	PUNCT
elements-9012	424	24	.	.	PUNCT
elements-9012	425	1	this	this	DET
elements-9012	425	2	phenomenon	phenomenon	NOUN
elements-9012	425	3	mirrors	mirror	VERB
elements-9012	425	4	that	that	SCONJ
elements-9012	425	5	which	which	PRON
elements-9012	425	6	has	have	AUX
elements-9012	425	7	been	be	AUX
elements-9012	425	8	observed	observe	VERB
elements-9012	425	9	for	for	ADP
elements-9012	425	10	human	human	NOUN
elements-9012	425	11	-	-	PUNCT
elements-9012	425	12	mouse	mouse	NOUN
elements-9012	425	13	substitution	substitution	NOUN
elements-9012	425	14	rates	rate	NOUN
elements-9012	425	15	(	(	PUNCT
elements-9012	425	16	see	see	AUX
elements-9012	425	17	also	also	ADV
elements-9012	425	18	figure	figure	VERB
elements-9012	425	19	i	i	PRON
elements-9012	425	20	a	a	PRON
elements-9012	425	21	)	)	PUNCT
elements-9012	425	22	.	.	PUNCT
elements-9012	426	1	x	x	PUNCT
elements-9012	426	2	x	x	PUNCT
elements-9012	426	3	l	l	NOUN
elements-9012	426	4	for	for	ADP
elements-9012	426	5	the	the	DET
elements-9012	426	6	three	three	NUM
elements-9012	426	7	candida	candida	PROPN
elements-9012	426	8	species	specie	NOUN
elements-9012	426	9	,	,	PUNCT
elements-9012	426	10	the	the	DET
elements-9012	426	11	autocorrelation	autocorrelation	NOUN
elements-9012	426	12	analysis	analysis	NOUN
elements-9012	426	13	shows	show	VERB
elements-9012	426	14	that	that	SCONJ
elements-9012	426	15	substitution	substitution	NOUN
elements-9012	426	16	rates	rate	NOUN
elements-9012	426	17	are	be	AUX
elements-9012	426	18	significantly	significantly	ADV
elements-9012	426	19	correlated	correlate	VERB
elements-9012	426	20	for	for	ADP
elements-9012	426	21	genes	gene	NOUN
elements-9012	426	22	within	within	ADP
elements-9012	426	23	50,000	50,000	NUM
elements-9012	426	24	base	base	NOUN
elements-9012	426	25	pairs	pair	NOUN
elements-9012	426	26	of	of	ADP
elements-9012	426	27	each	each	DET
elements-9012	426	28	other	other	ADJ
elements-9012	426	29	(	(	PUNCT
elements-9012	426	30	figure	figure	NOUN
elements-9012	426	31	2b	2b	NOUN
elements-9012	426	32	)	)	PUNCT
elements-9012	426	33	.	.	PUNCT
elements-9012	427	1	this	this	DET
elements-9012	427	2	result	result	NOUN
elements-9012	427	3	implies	imply	VERB
elements-9012	427	4	that	that	SCONJ
elements-9012	427	5	regional	regional	ADJ
elements-9012	427	6	biases	bias	NOUN
elements-9012	427	7	extend	extend	VERB
elements-9012	427	8	over	over	ADP
elements-9012	427	9	scales	scale	NOUN
elements-9012	427	10	encompassing	encompass	VERB
elements-9012	427	11	over	over	ADP
elements-9012	427	12	20	20	NUM
elements-9012	427	13	genes	gene	NOUN
elements-9012	427	14	,	,	PUNCT
elements-9012	427	15	since	since	SCONJ
elements-9012	427	16	in	in	ADP
elements-9012	427	17	c.	c.	NOUN
elements-9012	427	18	albicans	albican	VERB
elements-9012	427	19	the	the	DET
elements-9012	427	20	spacing	spacing	NOUN
elements-9012	427	21	between	between	ADP
elements-9012	427	22	genes	gene	NOUN
elements-9012	427	23	is	be	AUX
elements-9012	427	24	2300	2300	NUM
elements-9012	427	25	b	b	NOUN
elements-9012	427	26	p	p	NOUN
elements-9012	427	27	.	.	PUNCT
elements-9012	428	1	™	™	VERB
elements-9012	428	2	1	1	NUM
elements-9012	428	3	the	the	DET
elements-9012	428	4	neighboring	neighbor	VERB
elements-9012	428	5	gene	gene	NOUN
elements-9012	428	6	pearson	pearson	NOUN
elements-9012	428	7	correlations	correlation	NOUN
elements-9012	428	8	were	be	AUX
elements-9012	428	9	also	also	ADV
elements-9012	428	10	significant	significant	ADJ
elements-9012	428	11	(	(	PUNCT
elements-9012	428	12	c.	c.	NOUN
elements-9012	428	13	dubliniensis	dubliniensis	NOUN
elements-9012	428	14	/	/	SYM
elements-9012	428	15	c.	c.	NOUN
elements-9012	428	16	albicans	albican	VERB
elements-9012	428	17	pearson	pearson	NOUN
elements-9012	428	18	correlation	correlation	NOUN
elements-9012	428	19	=	=	PROPN
elements-9012	428	20	0.218	0.218	NUM
elements-9012	428	21	,	,	PUNCT
elements-9012	428	22	p	p	X
elements-9012	428	23	=	=	NUM
elements-9012	428	24	6.4	6.4	NUM
elements-9012	428	25	x	x	SYM
elements-9012	428	26	io"4	io"4	PROPN
elements-9012	428	27	2	2	NUM
elements-9012	428	28	;	;	PUNCT
elements-9012	428	29	c.	c.	PROPN
elements-9012	428	30	tropicalis	tropicalis	PROPN
elements-9012	428	31	/	/	SYM
elements-9012	428	32	c.	c.	PROPN
elements-9012	428	33	dubliniensis	dubliniensis	NOUN
elements-9012	428	34	pearson	pearson	NOUN
elements-9012	428	35	correlation	correlation	NOUN
elements-9012	428	36	=	=	PUNCT
elements-9012	428	37	0.103	0.103	NUM
elements-9012	428	38	,	,	PUNCT
elements-9012	428	39	p	p	NOUN
elements-9012	428	40	=	=	NOUN
elements-9012	428	41	9	9	NUM
elements-9012	428	42	-	-	SYM
elements-9012	428	43	8	8	NUM
elements-9012	428	44	x	x	SYM
elements-9012	428	45	10	10	NUM
elements-9012	428	46	-	-	SYM
elements-9012	428	47	10	10	NUM
elements-9012	428	48	)	)	PUNCT
elements-9012	428	49	.	.	PUNCT
elements-9012	429	1	snps	snps	PROPN
elements-9012	429	2	within	within	ADP
elements-9012	429	3	the	the	DET
elements-9012	429	4	c.	c.	NOUN
elements-9012	429	5	albicans	albican	NOUN
elements-9012	429	6	genome	genome	NOUN
elements-9012	429	7	have	have	AUX
elements-9012	429	8	been	be	AUX
elements-9012	429	9	shown	show	VERB
elements-9012	429	10	to	to	PART
elements-9012	429	11	be	be	AUX
elements-9012	429	12	unevenly	unevenly	ADV
elements-9012	429	13	distributed	distribute	VERB
elements-9012	429	14	along	along	ADP
elements-9012	429	15	contigs	contig	NOUN
elements-9012	429	16	,	,	PUNCT
elements-9012	429	17	x	x	PUNCT
elements-9012	429	18	x	x	VERB
elements-9012	429	19	m	m	VERB
elements-9012	429	20	in	in	ADP
elements-9012	429	21	agreement	agreement	NOUN
elements-9012	429	22	with	with	ADP
elements-9012	429	23	regional	regional	ADJ
elements-9012	429	24	mutational	mutational	ADJ
elements-9012	429	25	biases	bias	NOUN
elements-9012	429	26	.	.	PUNCT
elements-9012	430	1	these	these	DET
elements-9012	430	2	regional	regional	ADJ
elements-9012	430	3	effects	effect	NOUN
elements-9012	430	4	are	be	AUX
elements-9012	430	5	not	not	PART
elements-9012	430	6	due	due	ADJ
elements-9012	430	7	to	to	ADP
elements-9012	430	8	a	a	DET
elements-9012	430	9	cpg	cpg	NOUN
elements-9012	430	10	dinucleotide	dinucleotide	NOUN
elements-9012	430	11	effect	effect	NOUN
elements-9012	430	12	.	.	PUNCT
elements-9012	431	1	when	when	SCONJ
elements-9012	431	2	we	we	PRON
elements-9012	431	3	excluded	exclude	VERB
elements-9012	431	4	cpg	cpg	NOUN
elements-9012	431	5	sites	site	NOUN
elements-9012	431	6	from	from	ADP
elements-9012	431	7	the	the	DET
elements-9012	431	8	analysis	analysis	NOUN
elements-9012	431	9	,	,	PUNCT
elements-9012	431	10	neighboring	neighboring	NOUN
elements-9012	431	11	genes	gene	NOUN
elements-9012	431	12	were	be	AUX
elements-9012	431	13	still	still	ADV
elements-9012	431	14	observed	observe	VERB
elements-9012	431	15	to	to	PART
elements-9012	431	16	have	have	AUX
elements-9012	431	17	correlated	correlate	VERB
elements-9012	431	18	substitution	substitution	NOUN
elements-9012	431	19	rates	rate	NOUN
elements-9012	431	20	(	(	PUNCT
elements-9012	431	21	pearson	pearson	NOUN
elements-9012	431	22	-	-	PUNCT
elements-9012	431	23	correlation	correlation	NOUN
elements-9012	431	24	=	=	NOUN
elements-9012	431	25	0.1096	0.1096	NUM
elements-9012	431	26	,	,	PUNCT
elements-9012	431	27	p	p	X
elements-9012	431	28	<	<	X
elements-9012	431	29	10"^	10"^	NUM
elements-9012	431	30	)	)	PUNCT
elements-9012	431	31	.	.	PUNCT
elements-9012	432	1	these	these	DET
elements-9012	432	2	candida	candida	PROPN
elements-9012	432	3	species	specie	NOUN
elements-9012	432	4	translate	translate	ADJ
elements-9012	432	5	cug	cug	NOUN
elements-9012	432	6	as	as	ADP
elements-9012	432	7	serine	serine	NOUN
elements-9012	432	8	instead	instead	ADV
elements-9012	432	9	of	of	ADP
elements-9012	432	10	the	the	DET
elements-9012	432	11	usual	usual	ADJ
elements-9012	432	12	leucine	leucine	NOUN
elements-9012	432	13	,	,	PUNCT
elements-9012	432	14	and	and	CCONJ
elements-9012	432	15	it	it	PRON
elements-9012	432	16	might	might	AUX
elements-9012	432	17	be	be	AUX
elements-9012	432	18	hypothesized	hypothesize	VERB
elements-9012	432	19	that	that	SCONJ
elements-9012	432	20	this	this	PRON
elements-9012	432	21	is	be	AUX
elements-9012	432	22	relevant	relevant	ADJ
elements-9012	432	23	to	to	ADP
elements-9012	432	24	the	the	DET
elements-9012	432	25	rate	rate	NOUN
elements-9012	432	26	inferences.xx1v	inferences.xx1v	VERB
elements-9012	432	27	however	however	ADV
elements-9012	432	28	,	,	PUNCT
elements-9012	432	29	ignoring	ignore	VERB
elements-9012	432	30	these	these	DET
elements-9012	432	31	codons	codon	NOUN
elements-9012	432	32	does	do	AUX
elements-9012	432	33	not	not	PART
elements-9012	432	34	significantly	significantly	ADV
elements-9012	432	35	diminish	diminish	VERB
elements-9012	432	36	the	the	DET
elements-9012	432	37	correlation	correlation	NOUN
elements-9012	432	38	(	(	PUNCT
elements-9012	432	39	c.	c.	NOUN
elements-9012	432	40	e	e	NOUN
elements-9012	432	41	l	l	PROPN
elements-9012	432	42	e	e	X
elements-9012	432	43	m	m	PROPN
elements-9012	432	44	e	e	NOUN
elements-9012	432	45	n	n	PROPN
elements-9012	432	46	t	t	PROPN
elements-9012	432	47	s	s	PART
elements-9012	432	48	:	:	PUNCT
elements-9012	432	49	:	:	PUNCT
elements-9012	432	50	s	s	AUX
elements-9012	432	51	p	p	NOUN
elements-9012	432	52	r	r	NOUN
elements-9012	432	53	i	i	PRON
elements-9012	432	54	n	n	CCONJ
elements-9012	432	55	g	g	NOUN
elements-9012	432	56	08	08	NUM
elements-9012	432	57	dubliniensis	dubliniensis	NOUN
elements-9012	432	58	/	/	SYM
elements-9012	432	59	c.	c.	NOUN
elements-9012	432	60	albicans	albican	VERB
elements-9012	432	61	pearson	pearson	NOUN
elements-9012	432	62	correlation	correlation	NOUN
elements-9012	432	63	=	=	NOUN
elements-9012	432	64	0.208	0.208	NUM
elements-9012	432	65	,	,	PUNCT
elements-9012	432	66	p	p	NOUN
elements-9012	432	67	=	=	NOUN
elements-9012	432	68	10	10	NUM
elements-9012	432	69	"	"	PUNCT
elements-9012	432	70	*	*	NOUN
elements-9012	432	71	7	7	NUM
elements-9012	432	72	)	)	PUNCT
elements-9012	432	73	.	.	PUNCT
elements-9012	433	1	like	like	ADP
elements-9012	433	2	the	the	DET
elements-9012	433	3	candida	candida	PROPN
elements-9012	433	4	species	specie	NOUN
elements-9012	433	5	,	,	PUNCT
elements-9012	433	6	the	the	DET
elements-9012	433	7	species	specie	NOUN
elements-9012	433	8	n.	n.	NOUN
elements-9012	433	9	crassa	crassa	NOUN
elements-9012	433	10	and	and	CCONJ
elements-9012	433	11	c.	c.	PROPN
elements-9012	433	12	globosum	globosum	PROPN
elements-9012	433	13	have	have	VERB
elements-9012	433	14	a	a	DET
elements-9012	433	15	rate	rate	NOUN
elements-9012	433	16	distribution	distribution	NOUN
elements-9012	433	17	that	that	PRON
elements-9012	433	18	is	be	AUX
elements-9012	433	19	wider	wide	ADJ
elements-9012	433	20	than	than	ADP
elements-9012	433	21	the	the	DET
elements-9012	433	22	normal	normal	ADJ
elements-9012	433	23	gaussian	gaussian	NOUN
elements-9012	433	24	.	.	PUNCT
elements-9012	434	1	however	however	ADV
elements-9012	434	2	,	,	PUNCT
elements-9012	434	3	there	there	PRON
elements-9012	434	4	is	be	VERB
elements-9012	434	5	no	no	DET
elements-9012	434	6	significant	significant	ADJ
elements-9012	434	7	rate	rate	NOUN
elements-9012	434	8	correlation	correlation	NOUN
elements-9012	434	9	(	(	PUNCT
elements-9012	434	10	pearson	pearson	NOUN
elements-9012	434	11	correlation	correlation	NOUN
elements-9012	434	12	=	=	PUNCT
elements-9012	434	13	-0.0085	-0.0085	NOUN
elements-9012	434	14	,	,	PUNCT
elements-9012	434	15	p	p	X
elements-9012	434	16	=	=	NOUN
elements-9012	434	17	0	0	NUM
elements-9012	434	18	-	-	SYM
elements-9012	434	19	6757	6757	NUM
elements-9012	434	20	)	)	PUNCT
elements-9012	434	21	between	between	ADP
elements-9012	434	22	neighboring	neighboring	NOUN
elements-9012	434	23	genes	gene	NOUN
elements-9012	434	24	,	,	PUNCT
elements-9012	434	25	and	and	CCONJ
elements-9012	434	26	the	the	DET
elements-9012	434	27	autocorrelation	autocorrelation	NOUN
elements-9012	434	28	plot	plot	NOUN
elements-9012	434	29	appears	appear	VERB
elements-9012	434	30	nearly	nearly	ADV
elements-9012	434	31	identical	identical	ADJ
elements-9012	434	32	to	to	ADP
elements-9012	434	33	those	those	PRON
elements-9012	434	34	of	of	ADP
elements-9012	434	35	the	the	DET
elements-9012	434	36	yeast	yeast	NOUN
elements-9012	434	37	with	with	ADP
elements-9012	434	38	uniform	uniform	ADJ
elements-9012	434	39	rates	rate	NOUN
elements-9012	434	40	.	.	PUNCT
elements-9012	435	1	these	these	DET
elements-9012	435	2	seemingly	seemingly	ADV
elements-9012	435	3	contradictory	contradictory	ADJ
elements-9012	435	4	results	result	NOUN
elements-9012	435	5	suggest	suggest	VERB
elements-9012	435	6	that	that	SCONJ
elements-9012	435	7	the	the	DET
elements-9012	435	8	wide	wide	ADJ
elements-9012	435	9	distribution	distribution	NOUN
elements-9012	435	10	of	of	ADP
elements-9012	435	11	rates	rate	NOUN
elements-9012	435	12	(	(	PUNCT
elements-9012	435	13	s	s	NOUN
elements-9012	435	14	=	=	NOUN
elements-9012	435	15	2.07	2.07	NUM
elements-9012	435	16	)	)	PUNCT
elements-9012	435	17	is	be	AUX
elements-9012	435	18	due	due	ADJ
elements-9012	435	19	to	to	ADP
elements-9012	435	20	pressures	pressure	NOUN
elements-9012	435	21	on	on	ADP
elements-9012	435	22	individual	individual	ADJ
elements-9012	435	23	genes	gene	NOUN
elements-9012	435	24	,	,	PUNCT
elements-9012	435	25	rather	rather	ADV
elements-9012	435	26	than	than	ADP
elements-9012	435	27	regional	regional	ADJ
elements-9012	435	28	effects	effect	NOUN
elements-9012	435	29	.	.	PUNCT
elements-9012	436	1	we	we	PRON
elements-9012	436	2	hypothesized	hypothesize	VERB
elements-9012	436	3	that	that	SCONJ
elements-9012	436	4	the	the	DET
elements-9012	436	5	large	large	ADJ
elements-9012	436	6	s	s	NOUN
elements-9012	436	7	without	without	ADP
elements-9012	436	8	apparent	apparent	ADJ
elements-9012	436	9	regional	regional	ADJ
elements-9012	436	10	correlation	correlation	NOUN
elements-9012	436	11	could	could	AUX
elements-9012	436	12	be	be	AUX
elements-9012	436	13	related	relate	VERB
elements-9012	436	14	to	to	ADP
elements-9012	436	15	increased	increase	VERB
elements-9012	436	16	selection	selection	NOUN
elements-9012	436	17	on	on	ADP
elements-9012	436	18	the	the	DET
elements-9012	436	19	silent	silent	ADJ
elements-9012	436	20	sites	site	NOUN
elements-9012	436	21	of	of	ADP
elements-9012	436	22	n.	n.	NOUN
elements-9012	436	23	crassa	crassa	NOUN
elements-9012	436	24	and	and	CCONJ
elements-9012	436	25	c.	c.	PROPN
elements-9012	436	26	globosum	globosum	PROPN
elements-9012	436	27	genes	gene	NOUN
elements-9012	436	28	.	.	PUNCT
elements-9012	437	1	such	such	ADJ
elements-9012	437	2	selection	selection	NOUN
elements-9012	437	3	could	could	AUX
elements-9012	437	4	cause	cause	VERB
elements-9012	437	5	some	some	DET
elements-9012	437	6	genes	gene	NOUN
elements-9012	437	7	to	to	PART
elements-9012	437	8	have	have	VERB
elements-9012	437	9	more	more	ADJ
elements-9012	437	10	extreme	extreme	ADJ
elements-9012	437	11	conservation	conservation	NOUN
elements-9012	437	12	,	,	PUNCT
elements-9012	437	13	broadening	broaden	VERB
elements-9012	437	14	the	the	DET
elements-9012	437	15	rate	rate	NOUN
elements-9012	437	16	distribution	distribution	NOUN
elements-9012	437	17	.	.	PUNCT
elements-9012	438	1	to	to	PART
elements-9012	438	2	test	test	VERB
elements-9012	438	3	this	this	PRON
elements-9012	438	4	,	,	PUNCT
elements-9012	438	5	we	we	PRON
elements-9012	438	6	considered	consider	VERB
elements-9012	438	7	the	the	DET
elements-9012	438	8	codon	codon	ADJ
elements-9012	438	9	usage	usage	NOUN
elements-9012	438	10	bias	bias	NOUN
elements-9012	438	11	in	in	ADP
elements-9012	438	12	these	these	DET
elements-9012	438	13	species	specie	NOUN
elements-9012	438	14	,	,	PUNCT
elements-9012	438	15	which	which	PRON
elements-9012	438	16	is	be	AUX
elements-9012	438	17	the	the	DET
elements-9012	438	18	best	well	ADV
elements-9012	438	19	understood	understand	VERB
elements-9012	438	20	type	type	NOUN
elements-9012	438	21	of	of	ADP
elements-9012	438	22	silent	silent	ADJ
elements-9012	438	23	site	site	NOUN
elements-9012	438	24	selection	selection	NOUN
elements-9012	438	25	in	in	ADP
elements-9012	438	26	yeasts	yeast	NOUN
elements-9012	438	27	.	.	PUNCT
elements-9012	439	1	our	our	PRON
elements-9012	439	2	results	result	NOUN
elements-9012	439	3	indicate	indicate	VERB
elements-9012	439	4	that	that	SCONJ
elements-9012	439	5	codon	codon	ADJ
elements-9012	439	6	usage	usage	NOUN
elements-9012	439	7	selection	selection	NOUN
elements-9012	439	8	is	be	AUX
elements-9012	439	9	a	a	DET
elements-9012	439	10	stronger	strong	ADJ
elements-9012	439	11	effect	effect	NOUN
elements-9012	439	12	in	in	ADP
elements-9012	439	13	n.	n.	NOUN
elements-9012	439	14	crassa	crassa	PROPN
elements-9012	439	15	and	and	CCONJ
elements-9012	439	16	c.	c.	PROPN
elements-9012	439	17	globosum	globosum	PROPN
elements-9012	439	18	genomes	genome	NOUN
elements-9012	439	19	than	than	ADP
elements-9012	439	20	in	in	ADP
elements-9012	439	21	sensu	sensu	NOUN
elements-9012	439	22	stricto	stricto	X
elements-9012	439	23	yeasts	yeast	NOUN
elements-9012	439	24	.	.	PUNCT
elements-9012	440	1	cai	cai	PROPN
elements-9012	440	2	values	value	NOUN
elements-9012	440	3	were	be	AUX
elements-9012	440	4	calculated	calculate	VERB
elements-9012	440	5	for	for	ADP
elements-9012	440	6	each	each	PRON
elements-9012	440	7	of	of	ADP
elements-9012	440	8	n.	n.	NOUN
elements-9012	440	9	crassa	crassa	NOUN
elements-9012	440	10	and	and	CCONJ
elements-9012	440	11	s.	s.	PROPN
elements-9012	440	12	cerevisiae	cerevisiae	PROPN
elements-9012	440	13	based	base	VERB
elements-9012	440	14	on	on	ADP
elements-9012	440	15	their	their	PRON
elements-9012	440	16	respective	respective	ADJ
elements-9012	440	17	codon	codon	ADJ
elements-9012	440	18	biases	bias	NOUN
elements-9012	440	19	(	(	PUNCT
elements-9012	440	20	figure	figure	NOUN
elements-9012	440	21	3	3	NUM
elements-9012	440	22	)	)	PUNCT
elements-9012	440	23	.	.	PUNCT
elements-9012	441	1	n.	n.	PROPN
elements-9012	441	2	crassa	crassa	PROPN
elements-9012	441	3	genes	gene	NOUN
elements-9012	441	4	generally	generally	ADV
elements-9012	441	5	have	have	VERB
elements-9012	441	6	higher	high	ADJ
elements-9012	441	7	cai	cai	PROPN
elements-9012	441	8	values	value	NOUN
elements-9012	441	9	(	(	PUNCT
elements-9012	441	10	median	median	NOUN
elements-9012	441	11	0.63	0.63	NUM
elements-9012	441	12	)	)	PUNCT
elements-9012	441	13	than	than	ADP
elements-9012	441	14	s.	s.	PROPN
elements-9012	441	15	cerevisiae	cerevisiae	NOUN
elements-9012	441	16	genes	gene	NOUN
elements-9012	441	17	(	(	PUNCT
elements-9012	441	18	median	median	NOUN
elements-9012	441	19	0.14	0.14	NUM
elements-9012	441	20	)	)	PUNCT
elements-9012	441	21	.	.	PUNCT
elements-9012	442	1	in	in	ADP
elements-9012	442	2	each	each	PRON
elements-9012	442	3	of	of	ADP
elements-9012	442	4	the	the	DET
elements-9012	442	5	genomes	genome	NOUN
elements-9012	442	6	,	,	PUNCT
elements-9012	442	7	there	there	PRON
elements-9012	442	8	is	be	VERB
elements-9012	442	9	one	one	NUM
elements-9012	442	10	main	main	ADJ
elements-9012	442	11	cluster	cluster	NOUN
elements-9012	442	12	of	of	ADP
elements-9012	442	13	genes	gene	NOUN
elements-9012	442	14	having	have	VERB
elements-9012	442	15	z	z	NOUN
elements-9012	442	16	-	-	PUNCT
elements-9012	442	17	scores	score	NOUN
elements-9012	442	18	distributed	distribute	VERB
elements-9012	442	19	around	around	ADP
elements-9012	442	20	zero	zero	NUM
elements-9012	442	21	and	and	CCONJ
elements-9012	442	22	low	low	ADJ
elements-9012	442	23	cai	cai	PROPN
elements-9012	442	24	values	value	NOUN
elements-9012	442	25	.	.	PUNCT
elements-9012	443	1	then	then	ADV
elements-9012	443	2	there	there	PRON
elements-9012	443	3	is	be	VERB
elements-9012	443	4	another	another	DET
elements-9012	443	5	group	group	NOUN
elements-9012	443	6	having	have	VERB
elements-9012	443	7	negative	negative	ADJ
elements-9012	443	8	z	z	NOUN
elements-9012	443	9	-	-	PUNCT
elements-9012	443	10	scores	score	NOUN
elements-9012	443	11	and	and	CCONJ
elements-9012	443	12	high	high	ADJ
elements-9012	443	13	cai	cai	X
elements-9012	443	14	,	,	PUNCT
elements-9012	443	15	and	and	CCONJ
elements-9012	443	16	this	this	DET
elements-9012	443	17	group	group	NOUN
elements-9012	443	18	is	be	AUX
elements-9012	443	19	presumably	presumably	ADV
elements-9012	443	20	under	under	ADP
elements-9012	443	21	codon	codon	ADJ
elements-9012	443	22	usage	usage	NOUN
elements-9012	443	23	selection	selection	NOUN
elements-9012	443	24	.	.	PUNCT
elements-9012	444	1	n.	n.	PROPN
elements-9012	444	2	crassa	crassa	PROPN
elements-9012	444	3	appears	appear	VERB
elements-9012	444	4	to	to	PART
elements-9012	444	5	have	have	VERB
elements-9012	444	6	more	more	ADJ
elements-9012	444	7	genes	gene	NOUN
elements-9012	444	8	i	i	PRON
elements-9012	444	9	n	n	VERB
elements-9012	444	10	the	the	DET
elements-9012	444	11	group	group	NOUN
elements-9012	444	12	under	under	ADP
elements-9012	444	13	codon	codon	NOUN
elements-9012	444	14	usage	usage	NOUN
elements-9012	444	15	selection	selection	NOUN
elements-9012	444	16	(	(	PUNCT
elements-9012	444	17	949	949	NUM
elements-9012	444	18	genes	gene	NOUN
elements-9012	444	19	above	above	ADP
elements-9012	444	20	cai	cai	X
elements-9012	444	21	=	=	NOUN
elements-9012	444	22	0.7	0.7	NUM
elements-9012	444	23	)	)	PUNCT
elements-9012	444	24	than	than	ADP
elements-9012	444	25	s.	s.	PROPN
elements-9012	444	26	cerevisiae	cerevisiae	PROPN
elements-9012	444	27	(	(	PUNCT
elements-9012	444	28	121	121	NUM
elements-9012	444	29	genes	gene	NOUN
elements-9012	444	30	above	above	ADP
elements-9012	444	31	cai	cai	X
elements-9012	444	32	=	=	PROPN
elements-9012	444	33	0.4	0.4	NUM
elements-9012	444	34	)	)	PUNCT
elements-9012	444	35	.	.	PUNCT
elements-9012	445	1	this	this	PRON
elements-9012	445	2	suggests	suggest	VERB
elements-9012	445	3	that	that	SCONJ
elements-9012	445	4	codon	codon	ADJ
elements-9012	445	5	usage	usage	NOUN
elements-9012	445	6	selection	selection	NOUN
elements-9012	445	7	affects	affect	VERB
elements-9012	445	8	more	more	ADJ
elements-9012	445	9	genes	gene	NOUN
elements-9012	445	10	in	in	ADP
elements-9012	445	11	n.	n.	PROPN
elements-9012	445	12	crassa/	crassa/	NUM
elements-9012	445	13	c.	c.	PROPN
elements-9012	445	14	globosum	globosum	PROPN
elements-9012	445	15	,	,	PUNCT
elements-9012	445	16	and	and	CCONJ
elements-9012	445	17	could	could	AUX
elements-9012	445	18	be	be	AUX
elements-9012	445	19	responsible	responsible	ADJ
elements-9012	445	20	for	for	ADP
elements-9012	445	21	the	the	DET
elements-9012	445	22	large	large	ADJ
elements-9012	445	23	s	s	NOUN
elements-9012	445	24	in	in	ADP
elements-9012	445	25	these	these	DET
elements-9012	445	26	genomes	genome	NOUN
elements-9012	445	27	.	.	PUNCT
elements-9012	446	1	m	m	AUX
elements-9012	446	2	a	a	DET
elements-9012	446	3	m	m	NOUN
elements-9012	446	4	m	m	VERB
elements-9012	446	5	a	a	DET
elements-9012	446	6	l	l	NOUN
elements-9012	446	7	s	s	AUX
elements-9012	446	8	al	al	PROPN
elements-9012	446	9	l	l	PROPN
elements-9012	446	10	20	20	NUM
elements-9012	446	11	mammalian	mammalian	ADJ
elements-9012	446	12	species	specie	NOUN
elements-9012	446	13	appear	appear	VERB
elements-9012	446	14	to	to	PART
elements-9012	446	15	have	have	VERB
elements-9012	446	16	heterogeneous	heterogeneous	ADJ
elements-9012	446	17	mutation	mutation	NOUN
elements-9012	446	18	patterns	pattern	NOUN
elements-9012	446	19	,	,	PUNCT
elements-9012	446	20	as	as	SCONJ
elements-9012	446	21	evidenced	evidence	VERB
elements-9012	446	22	by	by	ADP
elements-9012	446	23	their	their	PRON
elements-9012	446	24	wide	wide	ADJ
elements-9012	446	25	rate	rate	NOUN
elements-9012	446	26	distributions	distribution	NOUN
elements-9012	446	27	(	(	PUNCT
elements-9012	446	28	1.80	1.80	NUM
elements-9012	446	29	<	<	X
elements-9012	446	30	s	s	X
elements-9012	446	31	<	<	X
elements-9012	446	32	2.23	2.23	NUM
elements-9012	446	33	)	)	PUNCT
elements-9012	446	34	(	(	PUNCT
elements-9012	446	35	figures	figure	NOUN
elements-9012	446	36	ia	ia	PROPN
elements-9012	446	37	and	and	CCONJ
elements-9012	446	38	5	5	NUM
elements-9012	446	39	)	)	PUNCT
elements-9012	446	40	.	.	PUNCT
elements-9012	447	1	the	the	DET
elements-9012	447	2	pearson	pearson	PROPN
elements-9012	447	3	correlations	correlation	NOUN
elements-9012	447	4	(	(	PUNCT
elements-9012	447	5	table	table	NOUN
elements-9012	447	6	2	2	NUM
elements-9012	447	7	)	)	PUNCT
elements-9012	447	8	and	and	CCONJ
elements-9012	447	9	autocorrelations	autocorrelation	NOUN
elements-9012	447	10	of	of	ADP
elements-9012	447	11	nearby	nearby	ADJ
elements-9012	447	12	genes	gene	NOUN
elements-9012	447	13	(	(	PUNCT
elements-9012	447	14	figure	figure	NOUN
elements-9012	447	15	4	4	NUM
elements-9012	447	16	)	)	PUNCT
elements-9012	447	17	for	for	ADP
elements-9012	447	18	each	each	DET
elements-9012	447	19	pair	pair	NOUN
elements-9012	447	20	-	-	PUNCT
elements-9012	447	21	wise	wise	ADJ
elements-9012	447	22	mammalian	mammalian	ADJ
elements-9012	447	23	comparison	comparison	NOUN
elements-9012	447	24	are	be	AUX
elements-9012	447	25	all	all	ADV
elements-9012	447	26	significant	significant	ADJ
elements-9012	447	27	.	.	PUNCT
elements-9012	448	1	for	for	ADP
elements-9012	448	2	example	example	NOUN
elements-9012	448	3	,	,	PUNCT
elements-9012	448	4	in	in	ADP
elements-9012	448	5	the	the	DET
elements-9012	448	6	cow	cow	NOUN
elements-9012	448	7	/	/	SYM
elements-9012	448	8	human	human	ADJ
elements-9012	448	9	comparison	comparison	NOUN
elements-9012	448	10	,	,	PUNCT
elements-9012	448	11	the	the	DET
elements-9012	448	12	autocorrelation	autocorrelation	NOUN
elements-9012	448	13	graph	graph	NOUN
elements-9012	448	14	suggests	suggest	VERB
elements-9012	448	15	mutational	mutational	ADJ
elements-9012	448	16	blocks	block	NOUN
elements-9012	448	17	along	along	ADP
elements-9012	448	18	each	each	DET
elements-9012	448	19	chromosome	chromosome	NOUN
elements-9012	448	20	as	as	ADV
elements-9012	448	21	large	large	ADJ
elements-9012	448	22	as	as	ADP
elements-9012	448	23	10	10	NUM
elements-9012	448	24	mb	mb	ADP
elements-9012	448	25	,	,	PUNCT
elements-9012	448	26	similar	similar	ADJ
elements-9012	448	27	to	to	ADP
elements-9012	448	28	the	the	DET
elements-9012	448	29	length	length	NOUN
elements-9012	448	30	scale	scale	NOUN
elements-9012	448	31	that	that	PRON
elements-9012	448	32	has	have	AUX
elements-9012	448	33	previously	previously	ADV
elements-9012	448	34	been	be	AUX
elements-9012	448	35	observed	observe	VERB
elements-9012	448	36	in	in	ADP
elements-9012	448	37	mouse	mouse	NOUN
elements-9012	448	38	and	and	CCONJ
elements-9012	448	39	human	human	NOUN
elements-9012	448	40	.	.	PUNCT
elements-9012	449	1	the	the	DET
elements-9012	449	2	regional	regional	ADJ
elements-9012	449	3	variations	variation	NOUN
elements-9012	449	4	are	be	AUX
elements-9012	449	5	also	also	ADV
elements-9012	449	6	apparent	apparent	ADJ
elements-9012	449	7	from	from	ADP
elements-9012	449	8	correlation	correlation	NOUN
elements-9012	449	9	analysis	analysis	NOUN
elements-9012	449	10	of	of	ADP
elements-9012	449	11	ancestral	ancestral	ADJ
elements-9012	449	12	repeats	repeat	NOUN
elements-9012	449	13	,	,	PUNCT
elements-9012	449	14	another	another	DET
elements-9012	449	15	largely	largely	ADV
elements-9012	449	16	neutral	neutral	ADJ
elements-9012	449	17	scattered	scatter	VERB
elements-9012	449	18	throughout	throughout	ADP
elements-9012	449	19	the	the	DET
elements-9012	449	20	genome	genome	NOUN
elements-9012	449	21	.	.	PUNCT
elements-9012	450	1	x	x	PUNCT
elements-9012	450	2	x	x	X
elements-9012	450	3	v	v	NOUN
elements-9012	450	4	we	we	PRON
elements-9012	450	5	analyzed	analyze	VERB
elements-9012	450	6	the	the	DET
elements-9012	450	7	18	18	NUM
elements-9012	450	8	-	-	PUNCT
elements-9012	450	9	way	way	NOUN
elements-9012	450	10	vertebrate	vertebrate	ADJ
elements-9012	450	11	multi	multi	ADJ
elements-9012	450	12	-	-	ADJ
elements-9012	450	13	species	species	ADJ
elements-9012	450	14	alignments	alignment	NOUN
elements-9012	450	15	from	from	ADP
elements-9012	450	16	ucsc	ucsc	PROPN
elements-9012	450	17	to	to	PART
elements-9012	450	18	obtain	obtain	VERB
elements-9012	450	19	table	table	NOUN
elements-9012	450	20	3	3	NUM
elements-9012	450	21	:	:	PUNCT
elements-9012	451	1	c	c	NOUN
elements-9012	451	2	o	o	NOUN
elements-9012	451	3	r	r	NOUN
elements-9012	451	4	r	r	NOUN
elements-9012	451	5	e	e	NOUN
elements-9012	451	6	l	l	NOUN
elements-9012	451	7	a	a	NOUN
elements-9012	451	8	t	t	NOUN
elements-9012	452	1	i	i	PRON
elements-9012	452	2	o	o	VERB
elements-9012	453	1	n	n	X
elements-9012	454	1	o	o	X
elements-9012	454	2	f	f	X
elements-9012	454	3	s	s	PROPN
elements-9012	454	4	u	u	PROPN
elements-9012	454	5	b	b	PROPN
elements-9012	454	6	s	s	ADP
elements-9012	454	7	t	t	X
elements-9012	455	1	i	i	PRON
elements-9012	455	2	t	t	VERB
elements-9012	455	3	u	u	NOUN
elements-9012	456	1	t	t	NOUN
elements-9012	457	1	i	i	PRON
elements-9012	457	2	o	o	VERB
elements-9012	457	3	n	n	CCONJ
elements-9012	457	4	a	a	DET
elements-9012	457	5	l	l	NOUN
elements-9012	457	6	r	r	NOUN
elements-9012	457	7	a	a	DET
elements-9012	457	8	t	t	NOUN
elements-9012	457	9	e	e	X
elements-9012	457	10	s	s	X
elements-9012	457	11	in	in	ADP
elements-9012	457	12	n	n	PRON
elements-9012	457	13	e	e	NOUN
elements-9012	457	14	i	i	PRON
elements-9012	457	15	g	g	PROPN
elements-9012	457	16	h	h	NOUN
elements-9012	458	1	b	b	NOUN
elements-9012	458	2	o	o	NOUN
elements-9012	459	1	r	r	VERB
elements-9012	460	1	i	i	PRON
elements-9012	461	1	n	n	CCONJ
elements-9012	462	1	g	g	PROPN
elements-9012	462	2	a	a	PRON
elements-9012	462	3	n	n	NOUN
elements-9012	463	1	c	c	VERB
elements-9012	463	2	i	i	NOUN
elements-9012	463	3	e	e	PROPN
elements-9012	463	4	n	n	PROPN
elements-9012	463	5	t	t	NOUN
elements-9012	463	6	r	r	NOUN
elements-9012	463	7	e	e	NOUN
elements-9012	463	8	p	p	PROPN
elements-9012	463	9	e	e	PROPN
elements-9012	463	10	a	a	DET
elements-9012	463	11	t	t	PROPN
elements-9012	463	12	s	s	PART
elements-9012	463	13	n	n	ADP
elements-9012	463	14	u	u	NOUN
elements-9012	463	15	m	m	NOUN
elements-9012	463	16	b	b	NOUN
elements-9012	463	17	e	e	NOUN
elements-9012	463	18	r	r	NOUN
elements-9012	463	19	p	p	PROPN
elements-9012	463	20	e	e	PROPN
elements-9012	463	21	a	a	DET
elements-9012	463	22	r	r	NOUN
elements-9012	463	23	s	s	NOUN
elements-9012	463	24	o	o	NOUN
elements-9012	463	25	n	n	X
elements-9012	463	26	s	s	PROPN
elements-9012	463	27	p	p	X
elements-9012	463	28	e	e	NOUN
elements-9012	463	29	c	c	NOUN
elements-9012	464	1	i	i	NOUN
elements-9012	464	2	e	e	NOUN
elements-9012	464	3	s	s	VERB
elements-9012	464	4	1	1	NUM
elements-9012	464	5	s	s	PART
elements-9012	464	6	p	p	NOUN
elements-9012	464	7	e	e	NOUN
elements-9012	464	8	c	c	NOUN
elements-9012	465	1	i	i	NOUN
elements-9012	465	2	e	e	NOUN
elements-9012	465	3	s	s	X
elements-9012	465	4	2	2	NUM
elements-9012	465	5	o	o	NOUN
elements-9012	465	6	f	f	NOUN
elements-9012	465	7	r	r	NOUN
elements-9012	465	8	e	e	PROPN
elements-9012	465	9	p	p	PROPN
elements-9012	465	10	e	e	PROPN
elements-9012	465	11	a	a	DET
elements-9012	465	12	t	t	NOUN
elements-9012	465	13	s	s	X
elements-9012	465	14	c	c	NOUN
elements-9012	465	15	o	o	NOUN
elements-9012	465	16	r	r	NOUN
elements-9012	465	17	r	r	NOUN
elements-9012	465	18	e	e	NOUN
elements-9012	465	19	l	l	NOUN
elements-9012	465	20	a	a	NOUN
elements-9012	465	21	t	t	NOUN
elements-9012	466	1	i	i	PRON
elements-9012	466	2	o	o	VERB
elements-9012	467	1	n	n	CCONJ
elements-9012	467	2	p	p	X
elements-9012	467	3	v	v	X
elements-9012	467	4	a	a	DET
elements-9012	467	5	l	l	NOUN
elements-9012	467	6	5	5	NUM
elements-9012	467	7	0	0	NUM
elements-9012	467	8	8	8	NUM
elements-9012	467	9	1	1	NUM
elements-9012	467	10	4	4	NUM
elements-9012	467	11	2	2	NUM
elements-9012	467	12	6	6	NUM
elements-9012	467	13	0	0	NUM
elements-9012	467	14	.	.	PUNCT
elements-9012	468	1	3	3	NUM
elements-9012	468	2	2	2	NUM
elements-9012	468	3	1	1	NUM
elements-9012	468	4	9	9	NUM
elements-9012	468	5	2	2	NUM
elements-9012	468	6	<	<	X
elements-9012	468	7	e	e	X
elements-9012	468	8	2	2	NUM
elements-9012	468	9	7	7	NUM
elements-9012	468	10	6	6	NUM
elements-9012	468	11	1	1	NUM
elements-9012	468	12	3	3	NUM
elements-9012	468	13	7	7	NUM
elements-9012	468	14	8	8	NUM
elements-9012	468	15	6	6	NUM
elements-9012	468	16	9	9	NUM
elements-9012	468	17	9	9	NUM
elements-9012	468	18	0	0	NUM
elements-9012	468	19	.	.	PUNCT
elements-9012	469	1	1	1	NUM
elements-9012	469	2	8	8	NUM
elements-9012	469	3	8	8	NUM
elements-9012	469	4	2	2	NUM
elements-9012	469	5	9	9	NUM
elements-9012	469	6	<	<	X
elements-9012	469	7	e	e	X
elements-9012	469	8	2	2	NUM
elements-9012	469	9	7	7	NUM
elements-9012	469	10	6	6	NUM
elements-9012	469	11	elephant	elephant	NOUN
elements-9012	469	12	2	2	NUM
elements-9012	469	13	4	4	NUM
elements-9012	469	14	8	8	NUM
elements-9012	469	15	5	5	NUM
elements-9012	469	16	7	7	NUM
elements-9012	469	17	7	7	NUM
elements-9012	469	18	0	0	NUM
elements-9012	469	19	.	.	PUNCT
elements-9012	470	1	1	1	NUM
elements-9012	470	2	2	2	NUM
elements-9012	470	3	1	1	NUM
elements-9012	470	4	5	5	NUM
elements-9012	470	5	8	8	NUM
elements-9012	470	6	<	<	X
elements-9012	470	7	e	e	X
elements-9012	470	8	2	2	NUM
elements-9012	470	9	7	7	NUM
elements-9012	470	10	6	6	NUM
elements-9012	470	11	2	2	NUM
elements-9012	470	12	3	3	NUM
elements-9012	470	13	1	1	NUM
elements-9012	470	14	2	2	NUM
elements-9012	470	15	8	8	NUM
elements-9012	470	16	3	3	NUM
elements-9012	470	17	0	0	NUM
elements-9012	470	18	.	.	PUNCT
elements-9012	471	1	1	1	NUM
elements-9012	471	2	2	2	NUM
elements-9012	471	3	9	9	NUM
elements-9012	471	4	2	2	NUM
elements-9012	471	5	<	<	X
elements-9012	471	6	e	e	X
elements-9012	471	7	2	2	NUM
elements-9012	471	8	7	7	NUM
elements-9012	471	9	6	6	NUM
elements-9012	471	10	m	m	NOUN
elements-9012	471	11	e	e	NOUN
elements-9012	471	12	a	a	PRON
elements-9012	471	13	s	s	X
elements-9012	471	14	u	u	NOUN
elements-9012	471	15	r	r	NOUN
elements-9012	471	16	i	i	PRON
elements-9012	471	17	n	n	CCONJ
elements-9012	471	18	g	g	PROPN
elements-9012	471	19	t	t	NOUN
elements-9012	471	20	h	h	NOUN
elements-9012	471	21	e	e	X
elements-9012	471	22	u	u	NOUN
elements-9012	471	23	n	n	PROPN
elements-9012	471	24	i	i	PRON
elements-9012	471	25	v	v	VERB
elements-9012	471	26	e	e	NOUN
elements-9012	471	27	r	r	NOUN
elements-9012	471	28	s	s	PROPN
elements-9012	471	29	a	a	DET
elements-9012	471	30	l	l	NOUN
elements-9012	472	1	i	i	PRON
elements-9012	472	2	t	t	VERB
elements-9012	473	1	y	y	NOUN
elements-9012	474	1	o	o	X
elements-9012	474	2	f	f	NOUN
elements-9012	475	1	r	r	NOUN
elements-9012	475	2	e	e	NOUN
elements-9012	475	3	g	g	NOUN
elements-9012	476	1	i	i	PRON
elements-9012	476	2	o	o	VERB
elements-9012	477	1	n	n	CCONJ
elements-9012	477	2	a	a	DET
elements-9012	477	3	l	l	NOUN
elements-9012	477	4	m	m	VERB
elements-9012	478	1	u	u	NOUN
elements-9012	478	2	t	t	PROPN
elements-9012	478	3	a	a	DET
elements-9012	478	4	t	t	NOUN
elements-9012	479	1	i	i	PRON
elements-9012	479	2	o	o	VERB
elements-9012	480	1	n	n	CCONJ
elements-9012	480	2	r	r	NOUN
elements-9012	480	3	a	a	DET
elements-9012	480	4	t	t	NOUN
elements-9012	480	5	e	e	X
elements-9012	480	6	s	s	X
elements-9012	480	7	in	in	ADP
elements-9012	480	8	all	all	DET
elements-9012	480	9	four	four	NUM
elements-9012	480	10	mammalian	mammalian	ADJ
elements-9012	480	11	pairs	pair	NOUN
elements-9012	480	12	considered	consider	VERB
elements-9012	480	13	(	(	PUNCT
elements-9012	480	14	human	human	NOUN
elements-9012	480	15	-	-	PUNCT
elements-9012	480	16	macaque	macaque	NOUN
elements-9012	480	17	)	)	PUNCT
elements-9012	480	18	,	,	PUNCT
elements-9012	480	19	(	(	PUNCT
elements-9012	480	20	doc	doc	PROPN
elements-9012	480	21	-	-	NOUN
elements-9012	480	22	cow	cow	NOUN
elements-9012	480	23	)	)	PUNCT
elements-9012	480	24	,	,	PUNCT
elements-9012	480	25	(	(	PUNCT
elements-9012	480	26	elephant	elephant	NOUN
elements-9012	480	27	-	-	PUNCT
elements-9012	480	28	tenrec	tenrec	NOUN
elements-9012	480	29	)	)	PUNCT
elements-9012	480	30	,	,	PUNCT
elements-9012	480	31	and	and	CCONJ
elements-9012	480	32	(	(	PUNCT
elements-9012	480	33	rat	rat	NOUN
elements-9012	480	34	-	-	PUNCT
elements-9012	480	35	mouse	mouse	NOUN
elements-9012	480	36	)	)	PUNCT
elements-9012	480	37	,	,	PUNCT
elements-9012	480	38	the	the	DET
elements-9012	480	39	correlations	correlation	NOUN
elements-9012	480	40	were	be	AUX
elements-9012	480	41	e	e	NOUN
elements-9012	480	42	x	x	X
elements-9012	480	43	t	t	NOUN
elements-9012	480	44	r	r	NOUN
elements-9012	480	45	e	e	NOUN
elements-9012	480	46	m	m	NOUN
elements-9012	480	47	e	e	NOUN
elements-9012	480	48	l	l	X
elements-9012	480	49	y	y	X
elements-9012	480	50	significant	significant	ADJ
elements-9012	480	51	(	(	PUNCT
elements-9012	480	52	<	<	X
elements-9012	480	53	1(t^^	1(t^^	NUM
elements-9012	480	54	)	)	PUNCT
elements-9012	480	55	.	.	PUNCT
elements-9012	481	1	this	this	PRON
elements-9012	481	2	supports	support	VERB
elements-9012	481	3	the	the	DET
elements-9012	481	4	widespread	widespread	ADJ
elements-9012	481	5	heterogeneity	heterogeneity	NOUN
elements-9012	481	6	of	of	ADP
elements-9012	481	7	mutation	mutation	NOUN
elements-9012	481	8	rates	rate	NOUN
elements-9012	481	9	in	in	ADP
elements-9012	481	10	mammalian	mammalian	ADJ
elements-9012	481	11	genomes	genome	NOUN
elements-9012	481	12	.	.	PUNCT
elements-9012	482	1	48	48	NUM
elements-9012	482	2	a	a	DET
elements-9012	482	3	stringent	stringent	ADJ
elements-9012	482	4	set	set	NOUN
elements-9012	482	5	of	of	ADP
elements-9012	482	6	ancestral	ancestral	ADJ
elements-9012	482	7	repeats	repeat	NOUN
elements-9012	482	8	aligned	align	VERB
elements-9012	482	9	orthologously	orthologously	ADV
elements-9012	482	10	across	across	ADP
elements-9012	482	11	the	the	DET
elements-9012	482	12	species	species	NOUN
elements-9012	482	13	pairs	pair	NOUN
elements-9012	482	14	(	(	PUNCT
elements-9012	482	15	human	human	ADJ
elements-9012	482	16	,	,	PUNCT
elements-9012	482	17	macaque	macaque	ADJ
elements-9012	482	18	)	)	PUNCT
elements-9012	482	19	,	,	PUNCT
elements-9012	482	20	(	(	PUNCT
elements-9012	482	21	mouse	mouse	NOUN
elements-9012	482	22	,	,	PUNCT
elements-9012	482	23	rat	rat	NOUN
elements-9012	482	24	)	)	PUNCT
elements-9012	482	25	,	,	PUNCT
elements-9012	482	26	(	(	PUNCT
elements-9012	482	27	elephant	elephant	NOUN
elements-9012	482	28	,	,	PUNCT
elements-9012	482	29	tenrec	tenrec	NOUN
elements-9012	482	30	)	)	PUNCT
elements-9012	482	31	,	,	PUNCT
elements-9012	482	32	and	and	CCONJ
elements-9012	482	33	(	(	PUNCT
elements-9012	482	34	dog	dog	NOUN
elements-9012	482	35	,	,	PUNCT
elements-9012	482	36	cow	cow	PROPN
elements-9012	482	37	)	)	PUNCT
elements-9012	482	38	.	.	PUNCT
elements-9012	483	1	we	we	PRON
elements-9012	483	2	observed	observe	VERB
elements-9012	483	3	that	that	SCONJ
elements-9012	483	4	an	an	DET
elements-9012	483	5	ancestral	ancestral	ADJ
elements-9012	483	6	repeat	repeat	NOUN
elements-9012	483	7	's	's	PART
elements-9012	483	8	substitution	substitution	NOUN
elements-9012	483	9	rate	rate	NOUN
elements-9012	483	10	is	be	AUX
elements-9012	483	11	significantly	significantly	ADV
elements-9012	483	12	correlated	correlate	VERB
elements-9012	483	13	with	with	ADP
elements-9012	483	14	that	that	PRON
elements-9012	483	15	of	of	ADP
elements-9012	483	16	the	the	DET
elements-9012	483	17	neighboring	neighbor	VERB
elements-9012	483	18	ancestral	ancestral	ADJ
elements-9012	483	19	repeat	repeat	NOUN
elements-9012	483	20	along	along	ADP
elements-9012	483	21	the	the	DET
elements-9012	483	22	genome	genome	NOUN
elements-9012	483	23	,	,	PUNCT
elements-9012	483	24	for	for	ADP
elements-9012	483	25	each	each	PRON
elements-9012	483	26	of	of	ADP
elements-9012	483	27	the	the	DET
elements-9012	483	28	species	species	NOUN
elements-9012	483	29	pairs	pair	NOUN
elements-9012	483	30	(	(	PUNCT
elements-9012	483	31	table	table	NOUN
elements-9012	483	32	3	3	NUM
elements-9012	483	33	)	)	PUNCT
elements-9012	483	34	.	.	PUNCT
elements-9012	484	1	in	in	ADP
elements-9012	484	2	each	each	DET
elements-9012	484	3	case	case	NOUN
elements-9012	484	4	the	the	DET
elements-9012	484	5	significance	significance	NOUN
elements-9012	484	6	of	of	ADP
elements-9012	484	7	the	the	DET
elements-9012	484	8	correlation	correlation	NOUN
elements-9012	484	9	was	be	AUX
elements-9012	484	10	at	at	ADP
elements-9012	484	11	the	the	DET
elements-9012	484	12	l	l	PROPN
elements-9012	484	13	imi	imi	PROPN
elements-9012	484	14	t	t	PROPN
elements-9012	484	15	of	of	ADP
elements-9012	484	16	computational	computational	ADJ
elements-9012	484	17	precision	precision	NOUN
elements-9012	484	18	(	(	PUNCT
elements-9012	484	19	p	p	X
elements-9012	484	20	<	<	X
elements-9012	484	21	10	10	NUM
elements-9012	484	22	•	•	NUM
elements-9012	484	23	2	2	NUM
elements-9012	484	24	76	76	NUM
elements-9012	484	25	)	)	PUNCT
elements-9012	484	26	.	.	PUNCT
elements-9012	485	1	the	the	DET
elements-9012	485	2	wide	wide	ADJ
elements-9012	485	3	distribution	distribution	NOUN
elements-9012	485	4	of	of	ADP
elements-9012	485	5	ancestral	ancestral	ADJ
elements-9012	485	6	repeat	repeat	NOUN
elements-9012	485	7	normalized	normalize	VERB
elements-9012	485	8	rates	rate	NOUN
elements-9012	485	9	mimics	mimic	VERB
elements-9012	485	10	those	those	PRON
elements-9012	485	11	in	in	ADP
elements-9012	485	12	figures	figure	NOUN
elements-9012	485	13	ia	ia	PROPN
elements-9012	485	14	and	and	CCONJ
elements-9012	485	15	ic	ic	PROPN
elements-9012	485	16	,	,	PUNCT
elements-9012	485	17	with	with	ADP
elements-9012	485	18	standard	standard	ADJ
elements-9012	485	19	deviations	deviation	NOUN
elements-9012	485	20	ranging	range	VERB
elements-9012	485	21	from	from	ADP
elements-9012	485	22	1.41	1.41	NUM
elements-9012	485	23	(	(	PUNCT
elements-9012	485	24	elephant	elephant	NOUN
elements-9012	485	25	-	-	PUNCT
elements-9012	485	26	tenrec	tenrec	NOUN
elements-9012	485	27	)	)	PUNCT
elements-9012	485	28	to	to	ADP
elements-9012	485	29	1.96	1.96	NUM
elements-9012	485	30	(	(	PUNCT
elements-9012	485	31	human	human	NOUN
elements-9012	485	32	-	-	PUNCT
elements-9012	485	33	macaque	macaque	ADJ
elements-9012	485	34	)	)	PUNCT
elements-9012	485	35	.	.	PUNCT
elements-9012	486	1	1n	1n	NUM
elements-9012	486	2	s	s	X
elements-9012	486	3	e	e	NOUN
elements-9012	486	4	c	c	PROPN
elements-9012	486	5	t	t	PROPN
elements-9012	486	6	s	s	VERB
elements-9012	486	7	the	the	DET
elements-9012	486	8	yeast	yeast	NOUN
elements-9012	486	9	species	specie	NOUN
elements-9012	486	10	we	we	PRON
elements-9012	486	11	have	have	AUX
elements-9012	486	12	studied	study	VERB
elements-9012	486	13	are	be	AUX
elements-9012	486	14	at	at	ADP
elements-9012	486	15	phylogenetic	phylogenetic	ADJ
elements-9012	486	16	distances	distance	NOUN
elements-9012	486	17	that	that	PRON
elements-9012	486	18	are	be	AUX
elements-9012	486	19	generally	generally	ADV
elements-9012	486	20	larger	large	ADJ
elements-9012	486	21	than	than	ADP
elements-9012	486	22	those	those	PRON
elements-9012	486	23	for	for	ADP
elements-9012	486	24	the	the	DET
elements-9012	486	25	mammalian	mammalian	ADJ
elements-9012	486	26	species	specie	NOUN
elements-9012	486	27	.	.	PUNCT
elements-9012	487	1	in	in	ADP
elements-9012	487	2	principle	principle	NOUN
elements-9012	487	3	,	,	PUNCT
elements-9012	487	4	it	it	PRON
elements-9012	487	5	is	be	AUX
elements-9012	487	6	more	more	ADV
elements-9012	487	7	difficult	difficult	ADJ
elements-9012	487	8	to	to	PART
elements-9012	487	9	measure	measure	VERB
elements-9012	487	10	regional	regional	ADJ
elements-9012	487	11	variations	variation	NOUN
elements-9012	487	12	when	when	SCONJ
elements-9012	487	13	inferring	infer	VERB
elements-9012	487	14	rates	rate	NOUN
elements-9012	487	15	from	from	ADP
elements-9012	487	16	more	more	ADV
elements-9012	487	17	distantly	distantly	ADV
elements-9012	487	18	related	relate	VERB
elements-9012	487	19	species	specie	NOUN
elements-9012	487	20	.	.	PUNCT
elements-9012	488	1	this	this	PRON
elements-9012	488	2	is	be	AUX
elements-9012	488	3	because	because	SCONJ
elements-9012	488	4	as	as	SCONJ
elements-9012	488	5	species	specie	NOUN
elements-9012	488	6	approach	approach	NOUN
elements-9012	488	7	saturated	saturate	VERB
elements-9012	488	8	divergence	divergence	NOUN
elements-9012	488	9	,	,	PUNCT
elements-9012	488	10	all	all	DET
elements-9012	488	11	mutation	mutation	NOUN
elements-9012	488	12	rate	rate	NOUN
elements-9012	488	13	inferences	inference	NOUN
elements-9012	488	14	become	become	VERB
elements-9012	488	15	increasingly	increasingly	ADV
elements-9012	488	16	uncertain	uncertain	ADJ
elements-9012	488	17	.	.	PUNCT
elements-9012	489	1	therefore	therefore	ADV
elements-9012	489	2	one	one	PRON
elements-9012	489	3	might	might	AUX
elements-9012	489	4	be	be	AUX
elements-9012	489	5	concerned	concern	VERB
elements-9012	489	6	that	that	SCONJ
elements-9012	489	7	the	the	DET
elements-9012	489	8	greater	great	ADJ
elements-9012	489	9	divergence	divergence	NOUN
elements-9012	489	10	among	among	ADP
elements-9012	489	11	the	the	DET
elements-9012	489	12	yeast	yeast	NOUN
elements-9012	489	13	species	specie	NOUN
elements-9012	489	14	obscures	obscure	VERB
elements-9012	489	15	regional	regional	ADJ
elements-9012	489	16	effects	effect	NOUN
elements-9012	489	17	.	.	PUNCT
elements-9012	490	1	this	this	DET
elements-9012	490	2	hypothesis	hypothesis	NOUN
elements-9012	490	3	can	can	AUX
elements-9012	490	4	be	be	AUX
elements-9012	490	5	tested	test	VERB
elements-9012	490	6	by	by	ADP
elements-9012	490	7	using	use	VERB
elements-9012	490	8	insect	insect	NOUN
elements-9012	490	9	genomes	genome	NOUN
elements-9012	490	10	,	,	PUNCT
elements-9012	490	11	several	several	ADJ
elements-9012	490	12	of	of	ADP
elements-9012	490	13	which	which	PRON
elements-9012	490	14	are	be	AUX
elements-9012	490	15	at	at	ADP
elements-9012	490	16	phylogenetic	phylogenetic	ADJ
elements-9012	490	17	divergences	divergence	NOUN
elements-9012	490	18	as	as	ADV
elements-9012	490	19	large	large	ADJ
elements-9012	490	20	as	as	ADP
elements-9012	490	21	that	that	PRON
elements-9012	490	22	of	of	ADP
elements-9012	490	23	the	the	DET
elements-9012	490	24	yeasts	yeast	NOUN
elements-9012	490	25	.	.	PUNCT
elements-9012	491	1	in	in	ADP
elements-9012	491	2	particular	particular	ADJ
elements-9012	491	3	,	,	PUNCT
elements-9012	491	4	d.	d.	NOUN
elements-9012	491	5	melanogaster	melanogaster	NOUN
elements-9012	491	6	and	and	CCONJ
elements-9012	491	7	d.	d.	PROPN
elements-9012	491	8	pseudoobscura	pseudoobscura	NOUN
elements-9012	491	9	are	be	AUX
elements-9012	491	10	at	at	ADP
elements-9012	491	11	a	a	DET
elements-9012	491	12	separation	separation	NOUN
elements-9012	491	13	generally	generally	ADV
elements-9012	491	14	larger	large	ADJ
elements-9012	491	15	than	than	ADP
elements-9012	491	16	that	that	PRON
elements-9012	491	17	of	of	ADP
elements-9012	491	18	the	the	DET
elements-9012	491	19	mammalian	mammalian	ADJ
elements-9012	491	20	species	specie	NOUN
elements-9012	491	21	(	(	PUNCT
elements-9012	491	22	four	four	NUM
elements-9012	491	23	-	-	ADJ
elements-9012	491	24	fold	fold	ADJ
elements-9012	491	25	divergence	divergence	NOUN
elements-9012	491	26	=	=	NOUN
elements-9012	491	27	0.514	0.514	NUM
elements-9012	491	28	)	)	PUNCT
elements-9012	491	29	and	and	CCONJ
elements-9012	491	30	comparable	comparable	ADJ
elements-9012	491	31	to	to	ADP
elements-9012	491	32	that	that	PRON
elements-9012	491	33	of	of	ADP
elements-9012	491	34	most	most	ADJ
elements-9012	491	35	of	of	ADP
elements-9012	491	36	the	the	DET
elements-9012	491	37	yeast	yeast	NOUN
elements-9012	491	38	pairs	pair	NOUN
elements-9012	491	39	,	,	PUNCT
elements-9012	491	40	as	as	SCONJ
elements-9012	491	41	are	be	AUX
elements-9012	491	42	the	the	DET
elements-9012	491	43	two	two	NUM
elements-9012	491	44	mosquitoes	mosquito	NOUN
elements-9012	491	45	a.	a.	NOUN
elements-9012	491	46	aegypti	aegypti	NOUN
elements-9012	491	47	and	and	CCONJ
elements-9012	491	48	a.	a.	NOUN
elements-9012	491	49	gambiae	gambiae	NOUN
elements-9012	491	50	(	(	PUNCT
elements-9012	491	51	4	4	NUM
elements-9012	491	52	-	-	ADJ
elements-9012	491	53	fold	fold	ADJ
elements-9012	491	54	divergence	divergence	NOUN
elements-9012	491	55	=	=	PROPN
elements-9012	491	56	0.632	0.632	NUM
elements-9012	491	57	)	)	PUNCT
elements-9012	491	58	.	.	PUNCT
elements-9012	492	1	in	in	ADP
elements-9012	492	2	contrast	contrast	NOUN
elements-9012	492	3	to	to	ADP
elements-9012	492	4	the	the	DET
elements-9012	492	5	yeasts	yeast	NOUN
elements-9012	492	6	,	,	PUNCT
elements-9012	492	7	flies	fly	NOUN
elements-9012	492	8	and	and	CCONJ
elements-9012	492	9	mosquitoes	mosquito	NOUN
elements-9012	492	10	both	both	PRON
elements-9012	492	11	show	show	VERB
elements-9012	492	12	clear	clear	ADJ
elements-9012	492	13	evidence	evidence	NOUN
elements-9012	492	14	of	of	ADP
elements-9012	492	15	regional	regional	ADJ
elements-9012	492	16	effects	effect	NOUN
elements-9012	492	17	.	.	PUNCT
elements-9012	493	1	d.	d.	NOUN
elements-9012	493	2	melanogaster	melanogaster	PROPN
elements-9012	493	3	and	and	CCONJ
elements-9012	493	4	d.	d.	PROPN
elements-9012	493	5	pseudoobscura	pseudoobscura	NOUN
elements-9012	493	6	have	have	VERB
elements-9012	493	7	neighboring	neighboring	NOUN
elements-9012	493	8	genes	gene	NOUN
elements-9012	493	9	with	with	ADP
elements-9012	493	10	significant	significant	ADJ
elements-9012	493	11	rate	rate	NOUN
elements-9012	493	12	correlations	correlation	NOUN
elements-9012	493	13	at	at	ADP
elements-9012	493	14	distances	distance	NOUN
elements-9012	493	15	up	up	ADP
elements-9012	493	16	to	to	PART
elements-9012	493	17	~imb	~imb	VERB
elements-9012	493	18	apart	apart	ADV
elements-9012	493	19	(	(	PUNCT
elements-9012	493	20	pearson	pearson	NOUN
elements-9012	493	21	correlation	correlation	NOUN
elements-9012	493	22	=	=	PUNCT
elements-9012	493	23	0.1642	0.1642	NUM
elements-9012	493	24	,	,	PUNCT
elements-9012	493	25	p	p	X
elements-9012	493	26	=	=	PROPN
elements-9012	493	27	9.06	9.06	NUM
elements-9012	493	28	-	-	SYM
elements-9012	493	29	59	59	NUM
elements-9012	493	30	)	)	PUNCT
elements-9012	493	31	and	and	CCONJ
elements-9012	493	32	a	a	DET
elements-9012	493	33	wide	wide	ADJ
elements-9012	493	34	rate	rate	NOUN
elements-9012	493	35	distribution	distribution	NOUN
elements-9012	493	36	(	(	PUNCT
elements-9012	493	37	s	s	NOUN
elements-9012	493	38	=	=	NOUN
elements-9012	493	39	2.152	2.152	NUM
elements-9012	493	40	)	)	PUNCT
elements-9012	493	41	.	.	PUNCT
elements-9012	494	1	this	this	DET
elements-9012	494	2	effect	effect	NOUN
elements-9012	494	3	is	be	AUX
elements-9012	494	4	also	also	ADV
elements-9012	494	5	observed	observe	VERB
elements-9012	494	6	in	in	ADP
elements-9012	494	7	a.	a.	NOUN
elements-9012	494	8	aegypti	aegypti	NOUN
elements-9012	494	9	and	and	CCONJ
elements-9012	494	10	a.	a.	NOUN
elements-9012	494	11	gambiae	gambiae	NOUN
elements-9012	494	12	(	(	PUNCT
elements-9012	494	13	pearson	pearson	NOUN
elements-9012	494	14	correlation	correlation	NOUN
elements-9012	494	15	=	=	SYM
elements-9012	494	16	0.2099	0.2099	NUM
elements-9012	494	17	,	,	PUNCT
elements-9012	494	18	p	p	NOUN
elements-9012	494	19	=	=	NOUN
elements-9012	494	20	2.16	2.16	NUM
elements-9012	494	21	-	-	SYM
elements-9012	494	22	89	89	NUM
elements-9012	494	23	)	)	PUNCT
elements-9012	494	24	.	.	PUNCT
elements-9012	495	1	the	the	DET
elements-9012	495	2	width	width	NOUN
elements-9012	495	3	of	of	ADP
elements-9012	495	4	the	the	DET
elements-9012	495	5	distribution	distribution	NOUN
elements-9012	495	6	of	of	ADP
elements-9012	495	7	normalized	normalize	VERB
elements-9012	495	8	rates	rate	NOUN
elements-9012	495	9	is	be	AUX
elements-9012	495	10	s	s	NOUN
elements-9012	495	11	=	=	NOUN
elements-9012	495	12	1.804	1.804	NUM
elements-9012	495	13	.	.	PUNCT
elements-9012	496	1	each	each	PRON
elements-9012	496	2	of	of	ADP
elements-9012	496	3	these	these	DET
elements-9012	496	4	insect	insect	NOUN
elements-9012	496	5	lineages	lineage	NOUN
elements-9012	496	6	demonstrate	demonstrate	VERB
elements-9012	496	7	correlations	correlation	NOUN
elements-9012	496	8	which	which	PRON
elements-9012	496	9	are	be	AUX
elements-9012	496	10	more	more	ADV
elements-9012	496	11	statistically	statistically	ADV
elements-9012	496	12	significant	significant	ADJ
elements-9012	496	13	than	than	ADP
elements-9012	496	14	the	the	DET
elements-9012	496	15	most	most	ADV
elements-9012	496	16	significant	significant	ADJ
elements-9012	496	17	yeast	yeast	NOUN
elements-9012	496	18	comparison	comparison	NOUN
elements-9012	496	19	,	,	PUNCT
elements-9012	496	20	c.	c.	NOUN
elements-9012	496	21	albicans	albican	NOUN
elements-9012	496	22	and	and	CCONJ
elements-9012	496	23	c.	c.	NOUN
elements-9012	496	24	dubliniensis	dubliniensis	NOUN
elements-9012	496	25	,	,	PUNCT
elements-9012	496	26	despite	despite	SCONJ
elements-9012	496	27	the	the	DET
elements-9012	496	28	fact	fact	NOUN
elements-9012	496	29	that	that	SCONJ
elements-9012	496	30	the	the	DET
elements-9012	496	31	two	two	NUM
elements-9012	496	32	candida	candida	PROPN
elements-9012	496	33	species	specie	NOUN
elements-9012	496	34	have	have	AUX
elements-9012	496	35	a	a	DET
elements-9012	496	36	lower	low	ADJ
elements-9012	496	37	divergence	divergence	NOUN
elements-9012	496	38	(	(	PUNCT
elements-9012	496	39	0.289	0.289	NUM
elements-9012	496	40	)	)	PUNCT
elements-9012	496	41	than	than	ADP
elements-9012	496	42	the	the	DET
elements-9012	496	43	insects	insect	NOUN
elements-9012	496	44	.	.	PUNCT
elements-9012	497	1	therefore	therefore	ADV
elements-9012	497	2	,	,	PUNCT
elements-9012	497	3	we	we	PRON
elements-9012	497	4	can	can	AUX
elements-9012	497	5	conclude	conclude	VERB
elements-9012	497	6	that	that	SCONJ
elements-9012	497	7	the	the	DET
elements-9012	497	8	predominant	predominant	ADJ
elements-9012	497	9	lack	lack	NOUN
elements-9012	497	10	of	of	ADP
elements-9012	497	11	correlations	correlation	NOUN
elements-9012	497	12	for	for	ADP
elements-9012	497	13	the	the	DET
elements-9012	497	14	yeasts	yeast	NOUN
elements-9012	497	15	is	be	AUX
elements-9012	497	16	not	not	PART
elements-9012	497	17	an	an	DET
elements-9012	497	18	artifact	artifact	NOUN
elements-9012	497	19	of	of	ADP
elements-9012	497	20	their	their	PRON
elements-9012	497	21	larger	large	ADJ
elements-9012	497	22	phylogenetic	phylogenetic	ADJ
elements-9012	497	23	separations	separation	NOUN
elements-9012	497	24	.	.	PUNCT
elements-9012	498	1	c	c	NOUN
elements-9012	499	1	o	o	NOUN
elements-9012	499	2	n	n	ADP
elements-9012	499	3	c	c	NOUN
elements-9012	499	4	l	l	NOUN
elements-9012	499	5	u	u	NOUN
elements-9012	499	6	s	s	VERB
elements-9012	499	7	i	i	PRON
elements-9012	499	8	o	o	NOUN
elements-9012	500	1	n	n	X
elements-9012	500	2	s	s	PROPN
elements-9012	500	3	a	a	DET
elements-9012	500	4	n	n	NOUN
elements-9012	501	1	d	d	NOUN
elements-9012	501	2	d	d	NOUN
elements-9012	501	3	i	i	PRON
elements-9012	501	4	s	s	VERB
elements-9012	501	5	c	c	NOUN
elements-9012	501	6	u	u	NOUN
elements-9012	501	7	s	s	X
elements-9012	501	8	s	s	X
elements-9012	501	9	i	i	PRON
elements-9012	501	10	o	o	VERB
elements-9012	501	11	n	n	VERB
elements-9012	501	12	based	base	VERB
elements-9012	501	13	on	on	ADP
elements-9012	501	14	our	our	PRON
elements-9012	501	15	examination	examination	NOUN
elements-9012	501	16	of	of	ADP
elements-9012	501	17	20	20	NUM
elements-9012	501	18	mammalian	mammalian	ADJ
elements-9012	501	19	species	specie	NOUN
elements-9012	501	20	,	,	PUNCT
elements-9012	501	21	we	we	PRON
elements-9012	501	22	conclude	conclude	VERB
elements-9012	501	23	that	that	SCONJ
elements-9012	501	24	all	all	DET
elements-9012	501	25	mammals	mammal	NOUN
elements-9012	501	26	have	have	VERB
elements-9012	501	27	regional	regional	ADJ
elements-9012	501	28	biases	bias	NOUN
elements-9012	501	29	of	of	ADP
elements-9012	501	30	neutral	neutral	ADJ
elements-9012	501	31	mutation	mutation	NOUN
elements-9012	501	32	rates	rate	NOUN
elements-9012	501	33	.	.	PUNCT
elements-9012	502	1	while	while	SCONJ
elements-9012	502	2	the	the	DET
elements-9012	502	3	factors	factor	NOUN
elements-9012	502	4	controlling	control	VERB
elements-9012	502	5	these	these	DET
elements-9012	502	6	regional	regional	ADJ
elements-9012	502	7	biases	bias	NOUN
elements-9012	502	8	are	be	AUX
elements-9012	502	9	still	still	ADV
elements-9012	502	10	not	not	PART
elements-9012	502	11	well	well	ADV
elements-9012	502	12	understood	understand	VERB
elements-9012	502	13	(	(	PUNCT
elements-9012	502	14	e.g.	e.g.	ADV
elements-9012	502	15	base	base	NOUN
elements-9012	502	16	composition	composition	NOUN
elements-9012	502	17	,	,	PUNCT
elements-9012	502	18	local	local	ADJ
elements-9012	502	19	recombination	recombination	NOUN
elements-9012	502	20	rate	rate	NOUN
elements-9012	502	21	,	,	PUNCT
elements-9012	502	22	pattern	pattern	NOUN
elements-9012	502	23	of	of	ADP
elements-9012	502	24	gene	gene	NOUN
elements-9012	502	25	expression	expression	NOUN
elements-9012	502	26	,	,	PUNCT
elements-9012	502	27	gene	gene	NOUN
elements-9012	502	28	density	density	NOUN
elements-9012	502	29	and	and	CCONJ
elements-9012	502	30	dna	dna	PROPN
elements-9012	502	31	repair	repair	NOUN
elements-9012	502	32	domain	domain	NOUN
elements-9012	502	33	)	)	PUNCT
elements-9012	502	34	,	,	PUNCT
elements-9012	502	35	x	x	PUNCT
elements-9012	502	36	x	x	X
elements-9012	502	37	v	v	X
elements-9012	502	38	1	1	NUM
elements-9012	502	39	our	our	PRON
elements-9012	502	40	findings	finding	NOUN
elements-9012	502	41	indicate	indicate	VERB
elements-9012	502	42	that	that	SCONJ
elements-9012	502	43	any	any	DET
elements-9012	502	44	valid	valid	ADJ
elements-9012	502	45	explanations	explanation	NOUN
elements-9012	502	46	must	must	AUX
elements-9012	502	47	occur	occur	VERB
elements-9012	502	48	throughout	throughout	ADP
elements-9012	502	49	the	the	DET
elements-9012	502	50	mammalian	mammalian	ADJ
elements-9012	502	51	phylogeny	phylogeny	NOUN
elements-9012	502	52	.	.	PUNCT
elements-9012	503	1	in	in	ADP
elements-9012	503	2	contrast	contrast	NOUN
elements-9012	503	3	,	,	PUNCT
elements-9012	503	4	we	we	PRON
elements-9012	503	5	find	find	VERB
elements-9012	503	6	that	that	SCONJ
elements-9012	503	7	almost	almost	ADV
elements-9012	503	8	all	all	DET
elements-9012	503	9	yeasts	yeast	NOUN
elements-9012	503	10	have	have	VERB
elements-9012	503	11	a	a	DET
elements-9012	503	12	uniform	uniform	ADJ
elements-9012	503	13	neutral	neutral	ADJ
elements-9012	503	14	mutation	mutation	NOUN
elements-9012	503	15	rate	rate	NOUN
elements-9012	503	16	.	.	PUNCT
elements-9012	504	1	this	this	DET
elements-9012	504	2	conclusion	conclusion	NOUN
elements-9012	504	3	is	be	AUX
elements-9012	504	4	supported	support	VERB
elements-9012	504	5	by	by	ADP
elements-9012	504	6	a	a	DET
elements-9012	504	7	recent	recent	ADJ
elements-9012	504	8	report	report	NOUN
elements-9012	504	9	that	that	SCONJ
elements-9012	504	10	s.	s.	PROPN
elements-9012	504	11	cerevisiae	cerevisiae	PROPN
elements-9012	504	12	polymorphism	polymorphism	NOUN
elements-9012	504	13	rates	rate	NOUN
elements-9012	504	14	are	be	AUX
elements-9012	504	15	not	not	PART
elements-9012	504	16	correlated	correlate	VERB
elements-9012	504	17	along	along	ADP
elements-9012	504	18	the	the	DET
elements-9012	504	19	genome	genome	NOUN
elements-9012	504	20	.	.	PUNCT
elements-9012	505	1	x	x	PUNCT
elements-9012	505	2	x	x	SYM
elements-9012	505	3	v	v	ADP
elements-9012	505	4	n	n	CCONJ
elements-9012	505	5	the	the	DET
elements-9012	505	6	monophyletic	monophyletic	ADJ
elements-9012	505	7	group	group	NOUN
elements-9012	505	8	of	of	ADP
elements-9012	505	9	the	the	DET
elements-9012	505	10	three	three	NUM
elements-9012	505	11	candida	candida	PROPN
elements-9012	505	12	species	specie	NOUN
elements-9012	505	13	(	(	PUNCT
elements-9012	505	14	c.	c.	NOUN
elements-9012	505	15	albicans	albican	NOUN
elements-9012	505	16	,	,	PUNCT
elements-9012	505	17	c.	c.	NOUN
elements-9012	505	18	dubliniensis	dubliniensis	NOUN
elements-9012	505	19	,	,	PUNCT
elements-9012	505	20	and	and	CCONJ
elements-9012	505	21	c.	c.	PROPN
elements-9012	505	22	tropicalis	tropicalis	PROPN
elements-9012	505	23	)	)	PUNCT
elements-9012	505	24	is	be	AUX
elements-9012	505	25	an	an	DET
elements-9012	505	26	exception	exception	NOUN
elements-9012	505	27	to	to	ADP
elements-9012	505	28	the	the	DET
elements-9012	505	29	other	other	ADJ
elements-9012	505	30	yeasts	yeast	NOUN
elements-9012	505	31	.	.	PUNCT
elements-9012	506	1	both	both	CCONJ
elements-9012	506	2	the	the	DET
elements-9012	506	3	distribution	distribution	NOUN
elements-9012	506	4	of	of	ADP
elements-9012	506	5	rates	rate	NOUN
elements-9012	506	6	and	and	CCONJ
elements-9012	506	7	gene	gene	NOUN
elements-9012	506	8	-	-	PUNCT
elements-9012	506	9	togene	togene	NOUN
elements-9012	506	10	correlations	correlation	NOUN
elements-9012	506	11	indicate	indicate	VERB
elements-9012	506	12	that	that	SCONJ
elements-9012	506	13	these	these	DET
elements-9012	506	14	species	specie	NOUN
elements-9012	506	15	have	have	VERB
elements-9012	506	16	heterogeneous	heterogeneous	ADJ
elements-9012	506	17	mutation	mutation	NOUN
elements-9012	506	18	rates	rate	NOUN
elements-9012	506	19	,	,	PUNCT
elements-9012	506	20	a	a	DET
elements-9012	506	21	trait	trait	NOUN
elements-9012	506	22	which	which	PRON
elements-9012	506	23	had	have	AUX
elements-9012	506	24	not	not	PART
elements-9012	506	25	previously	previously	ADV
elements-9012	506	26	been	be	AUX
elements-9012	506	27	observed	observe	VERB
elements-9012	506	28	outside	outside	ADP
elements-9012	506	29	the	the	DET
elements-9012	506	30	mammals	mammal	NOUN
elements-9012	506	31	.	.	PUNCT
elements-9012	507	1	previous	previous	ADJ
elements-9012	507	2	studies	study	NOUN
elements-9012	507	3	of	of	ADP
elements-9012	507	4	snp	snp	ADJ
elements-9012	507	5	data	datum	NOUN
elements-9012	507	6	also	also	ADV
elements-9012	507	7	indicate	indicate	VERB
elements-9012	507	8	hotspots	hotspot	NOUN
elements-9012	507	9	of	of	ADP
elements-9012	507	10	polymorphism	polymorphism	NOUN
elements-9012	507	11	,	,	PUNCT
elements-9012	507	12	supporting	support	VERB
elements-9012	507	13	the	the	DET
elements-9012	507	14	concept	concept	NOUN
elements-9012	507	15	of	of	ADP
elements-9012	507	16	regional	regional	ADJ
elements-9012	507	17	biases	bias	NOUN
elements-9012	507	18	.	.	PUNCT
elements-9012	508	1	x	x	PUNCT
elements-9012	509	1	x	x	X
elements-9012	509	2	v	v	ADP
elements-9012	509	3	m	m	NOUN
elements-9012	509	4	what	what	PRON
elements-9012	509	5	characteristic	characteristic	ADJ
elements-9012	509	6	sets	set	VERB
elements-9012	509	7	these	these	DET
elements-9012	509	8	three	three	NUM
elements-9012	509	9	yeasts	yeast	NOUN
elements-9012	509	10	apart	apart	ADV
elements-9012	509	11	from	from	ADP
elements-9012	509	12	the	the	DET
elements-9012	509	13	others	other	NOUN
elements-9012	509	14	?	?	PUNCT
elements-9012	510	1	one	one	NUM
elements-9012	510	2	intriguing	intriguing	ADJ
elements-9012	510	3	trait	trait	NOUN
elements-9012	510	4	is	be	AUX
elements-9012	510	5	meiosis	meiosis	ADJ
elements-9012	510	6	.	.	PUNCT
elements-9012	511	1	unlike	unlike	ADP
elements-9012	511	2	other	other	ADJ
elements-9012	511	3	yeasts	yeast	NOUN
elements-9012	511	4	,	,	PUNCT
elements-9012	511	5	sexual	sexual	ADJ
elements-9012	511	6	reproduction	reproduction	NOUN
elements-9012	511	7	appears	appear	VERB
elements-9012	511	8	to	to	PART
elements-9012	511	9	be	be	AUX
elements-9012	511	10	rare	rare	ADJ
elements-9012	511	11	in	in	ADP
elements-9012	511	12	c.	c.	NOUN
elements-9012	511	13	albicans	albican	NOUN
elements-9012	511	14	,	,	PUNCT
elements-9012	511	15	c.	c.	NOUN
elements-9012	511	16	dubliniensis	dubliniensis	NOUN
elements-9012	511	17	,	,	PUNCT
elements-9012	511	18	and	and	CCONJ
elements-9012	511	19	c.	c.	PROPN
elements-9012	511	20	tropicalis	tropicalis	PROPN
elements-9012	511	21	.	.	PUNCT
elements-9012	512	1	sexual	sexual	ADJ
elements-9012	512	2	reproduction	reproduction	NOUN
elements-9012	512	3	in	in	ADP
elements-9012	512	4	c.	c.	NOUN
elements-9012	512	5	albicans	albican	NOUN
elements-9012	512	6	and	and	CCONJ
elements-9012	512	7	c.	c.	PROPN
elements-9012	512	8	dubliniensis	dubliniensis	NOUN
elements-9012	512	9	was	be	AUX
elements-9012	512	10	recentiy	recentiy	VERB
elements-9012	512	11	discovered	discover	VERB
elements-9012	512	12	under	under	ADP
elements-9012	512	13	specialized	specialized	ADJ
elements-9012	512	14	laboratory	laboratory	NOUN
elements-9012	512	15	conditions	condition	NOUN
elements-9012	512	16	;	;	PUNCT
elements-9012	512	17	however	however	ADV
elements-9012	512	18	,	,	PUNCT
elements-9012	512	19	no	no	DET
elements-9012	512	20	evidence	evidence	NOUN
elements-9012	512	21	for	for	ADP
elements-9012	512	22	meiosis	meiosis	NOUN
elements-9012	512	23	has	have	AUX
elements-9012	512	24	been	be	AUX
elements-9012	512	25	found	find	VERB
elements-9012	512	26	.	.	PUNCT
elements-9012	513	1	x	x	PUNCT
elements-9012	513	2	x	x	PUNCT
elements-9012	513	3	l	l	NOUN
elements-9012	513	4	x	x	VERB
elements-9012	513	5	the	the	DET
elements-9012	513	6	sexual	sexual	ADJ
elements-9012	513	7	cycle	cycle	NOUN
elements-9012	513	8	of	of	ADP
elements-9012	513	9	c.	c.	PROPN
elements-9012	513	10	tropicalis	tropicalis	PROPN
elements-9012	513	11	has	have	AUX
elements-9012	513	12	not	not	PART
elements-9012	513	13	been	be	AUX
elements-9012	513	14	extensively	extensively	ADV
elements-9012	513	15	explored	explore	VERB
elements-9012	513	16	,	,	PUNCT
elements-9012	513	17	but	but	CCONJ
elements-9012	513	18	meiosis	meiosis	NOUN
elements-9012	513	19	has	have	AUX
elements-9012	513	20	not	not	PART
elements-9012	513	21	been	be	AUX
elements-9012	513	22	observed	observe	VERB
elements-9012	513	23	for	for	ADP
elements-9012	513	24	it	it	PRON
elements-9012	513	25	either	either	ADV
elements-9012	513	26	.	.	PUNCT
elements-9012	514	1	on	on	ADP
elements-9012	514	2	the	the	DET
elements-9012	514	3	other	other	ADJ
elements-9012	514	4	hand	hand	NOUN
elements-9012	514	5	,	,	PUNCT
elements-9012	514	6	comparative	comparative	ADJ
elements-9012	514	7	genomic	genomic	NOUN
elements-9012	514	8	studies	study	NOUN
elements-9012	514	9	of	of	ADP
elements-9012	514	10	the	the	DET
elements-9012	514	11	uniformly	uniformly	ADV
elements-9012	514	12	mutating	mutate	VERB
elements-9012	514	13	c.	c.	PROPN
elements-9012	514	14	glabrata	glabrata	PROPN
elements-9012	514	15	support	support	VERB
elements-9012	514	16	a	a	DET
elements-9012	514	17	complete	complete	ADJ
elements-9012	514	18	sexual	sexual	ADJ
elements-9012	514	19	cycle	cycle	NOUN
elements-9012	514	20	,	,	PUNCT
elements-9012	514	21	x	x	PUNCT
elements-9012	514	22	x	x	PUNCT
elements-9012	514	23	x	x	X
elements-9012	514	24	and	and	CCONJ
elements-9012	514	25	so	so	ADV
elements-9012	514	26	-	-	PUNCT
elements-9012	514	27	called	call	VERB
elements-9012	514	28	"	"	PUNCT
elements-9012	514	29	defects	defect	NOUN
elements-9012	514	30	"	"	PUNCT
elements-9012	514	31	in	in	ADP
elements-9012	514	32	e	e	X
elements-9012	514	33	l	l	NOUN
elements-9012	514	34	e	e	X
elements-9012	514	35	m	m	PROPN
elements-9012	514	36	e	e	NOUN
elements-9012	514	37	n	n	NOUN
elements-9012	514	38	t	t	PROPN
elements-9012	514	39	s	s	NOUN
elements-9012	514	40	s	s	X
elements-9012	514	41	p	p	NOUN
elements-9012	514	42	r	r	NOUN
elements-9012	514	43	i	i	PRON
elements-9012	514	44	n	n	CCONJ
elements-9012	514	45	g	g	PROPN
elements-9012	514	46	08	08	NUM
elements-9012	514	47	the	the	DET
elements-9012	514	48	mating	mating	NOUN
elements-9012	514	49	type	type	NOUN
elements-9012	514	50	locus	locus	NOUN
elements-9012	514	51	of	of	ADP
elements-9012	514	52	c.	c.	PROPN
elements-9012	514	53	parapsiolosis	parapsiolosis	PROPN
elements-9012	514	54	suggest	suggest	VERB
elements-9012	514	55	that	that	SCONJ
elements-9012	514	56	it	it	PRON
elements-9012	514	57	is	be	AUX
elements-9012	514	58	unlikely	unlikely	ADJ
elements-9012	514	59	to	to	PART
elements-9012	514	60	have	have	VERB
elements-9012	514	61	mating	mate	VERB
elements-9012	514	62	similar	similar	ADJ
elements-9012	514	63	to	to	ADP
elements-9012	514	64	c.	c.	NOUN
elements-9012	514	65	albicans	albican	NOUN
elements-9012	514	66	and	and	CCONJ
elements-9012	514	67	c.	c.	NOUN
elements-9012	514	68	dubliniens	dublinien	NOUN
elements-9012	514	69	is	be	AUX
elements-9012	514	70	.	.	PUNCT
elements-9012	514	71	5	5	NUM
elements-9012	514	72	™	™	NOUN
elements-9012	514	73	thus	thus	ADV
elements-9012	514	74	,	,	PUNCT
elements-9012	514	75	the	the	DET
elements-9012	514	76	candida	candida	PROPN
elements-9012	514	77	species	specie	NOUN
elements-9012	514	78	without	without	ADP
elements-9012	514	79	evidence	evidence	NOUN
elements-9012	514	80	for	for	ADP
elements-9012	514	81	meiosis	meiosis	NOUN
elements-9012	514	82	are	be	AUX
elements-9012	514	83	the	the	DET
elements-9012	514	84	ones	one	NOUN
elements-9012	514	85	with	with	ADP
elements-9012	514	86	heterogeneous	heterogeneous	ADJ
elements-9012	514	87	neutral	neutral	ADJ
elements-9012	514	88	mutation	mutation	NOUN
elements-9012	514	89	rates	rate	NOUN
elements-9012	514	90	.	.	PUNCT
elements-9012	515	1	how	how	SCONJ
elements-9012	515	2	could	could	AUX
elements-9012	515	3	a	a	DET
elements-9012	515	4	lack	lack	NOUN
elements-9012	515	5	of	of	ADP
elements-9012	515	6	meiosis	meiosis	NOUN
elements-9012	515	7	influence	influence	NOUN
elements-9012	515	8	regional	regional	ADJ
elements-9012	515	9	mutation	mutation	NOUN
elements-9012	515	10	rates	rate	NOUN
elements-9012	515	11	?	?	PUNCT
elements-9012	516	1	one	one	NUM
elements-9012	516	2	possible	possible	ADJ
elements-9012	516	3	explanation	explanation	NOUN
elements-9012	516	4	is	be	AUX
elements-9012	516	5	that	that	SCONJ
elements-9012	516	6	the	the	DET
elements-9012	516	7	lack	lack	NOUN
elements-9012	516	8	of	of	ADP
elements-9012	516	9	meiosis	meiosis	NOUN
elements-9012	516	10	prevents	prevent	VERB
elements-9012	516	11	recombination	recombination	NOUN
elements-9012	516	12	between	between	ADP
elements-9012	516	13	homologous	homologous	ADJ
elements-9012	516	14	chromosomes	chromosome	NOUN
elements-9012	516	15	.	.	PUNCT
elements-9012	517	1	i	i	PRON
elements-9012	517	2	f	f	NOUN
elements-9012	517	3	recombination	recombination	NOUN
elements-9012	517	4	were	be	AUX
elements-9012	517	5	to	to	PART
elements-9012	517	6	cause	cause	VERB
elements-9012	517	7	random	random	ADJ
elements-9012	517	8	fluctuations	fluctuation	NOUN
elements-9012	517	9	in	in	ADP
elements-9012	517	10	chromosome	chromosome	NOUN
elements-9012	517	11	length	length	NOUN
elements-9012	517	12	,	,	PUNCT
elements-9012	517	13	this	this	PRON
elements-9012	517	14	could	could	AUX
elements-9012	517	15	have	have	VERB
elements-9012	517	16	the	the	DET
elements-9012	517	17	effect	effect	NOUN
elements-9012	517	18	of	of	ADP
elements-9012	517	19	smoothing	smooth	VERB
elements-9012	517	20	out	out	ADP
elements-9012	517	21	regional	regional	ADJ
elements-9012	517	22	mutational	mutational	ADJ
elements-9012	517	23	biases	bias	NOUN
elements-9012	517	24	.	.	PUNCT
elements-9012	518	1	a	a	DET
elements-9012	518	2	comparison	comparison	NOUN
elements-9012	518	3	of	of	ADP
elements-9012	518	4	n.	n.	NOUN
elements-9012	518	5	crassa	crassa	NOUN
elements-9012	518	6	and	and	CCONJ
elements-9012	518	7	c.	c.	PROPN
elements-9012	518	8	globosum	globosum	PROPN
elements-9012	518	9	also	also	ADV
elements-9012	518	10	fails	fail	VERB
elements-9012	518	11	to	to	PART
elements-9012	518	12	give	give	VERB
elements-9012	518	13	a	a	DET
elements-9012	518	14	normal	normal	ADJ
elements-9012	518	15	distribution	distribution	NOUN
elements-9012	518	16	of	of	ADP
elements-9012	518	17	substitution	substitution	NOUN
elements-9012	518	18	rate	rate	NOUN
elements-9012	518	19	z	z	NOUN
elements-9012	518	20	-	-	PUNCT
elements-9012	518	21	scores	score	NOUN
elements-9012	518	22	,	,	PUNCT
elements-9012	518	23	but	but	CCONJ
elements-9012	518	24	this	this	PRON
elements-9012	518	25	is	be	AUX
elements-9012	518	26	due	due	ADJ
elements-9012	518	27	to	to	ADP
elements-9012	518	28	stronger	strong	ADJ
elements-9012	518	29	codon	codon	ADJ
elements-9012	518	30	usage	usage	NOUN
elements-9012	518	31	selection	selection	NOUN
elements-9012	518	32	,	,	PUNCT
elements-9012	518	33	rather	rather	ADV
elements-9012	518	34	than	than	ADP
elements-9012	518	35	regional	regional	ADJ
elements-9012	518	36	mutation	mutation	NOUN
elements-9012	518	37	effects	effect	NOUN
elements-9012	518	38	.	.	PUNCT
elements-9012	519	1	this	this	DET
elements-9012	519	2	stronger	strong	ADJ
elements-9012	519	3	codon	codon	NOUN
elements-9012	519	4	usage	usage	NOUN
elements-9012	519	5	selection	selection	NOUN
elements-9012	519	6	is	be	AUX
elements-9012	519	7	more	more	ADV
elements-9012	519	8	apparent	apparent	ADJ
elements-9012	519	9	in	in	ADP
elements-9012	519	10	the	the	DET
elements-9012	519	11	context	context	NOUN
elements-9012	519	12	of	of	ADP
elements-9012	519	13	a	a	DET
elements-9012	519	14	mutational	mutational	ADJ
elements-9012	519	15	defense	defense	NOUN
elements-9012	519	16	mechanism	mechanism	NOUN
elements-9012	519	17	that	that	PRON
elements-9012	519	18	n.	n.	PROPN
elements-9012	519	19	crassa	crassa	PROPN
elements-9012	519	20	employs	employ	VERB
elements-9012	519	21	against	against	ADP
elements-9012	519	22	selfish	selfish	ADJ
elements-9012	519	23	dna	dna	NOUN
elements-9012	519	24	.	.	PUNCT
elements-9012	520	1	the	the	DET
elements-9012	520	2	mechanism	mechanism	NOUN
elements-9012	520	3	,	,	PUNCT
elements-9012	520	4	known	know	VERB
elements-9012	520	5	as	as	ADP
elements-9012	520	6	repeat	repeat	NOUN
elements-9012	520	7	-	-	PUNCT
elements-9012	520	8	induced	induce	VERB
elements-9012	520	9	point	point	NOUN
elements-9012	520	10	mutation	mutation	NOUN
elements-9012	520	11	(	(	PUNCT
elements-9012	520	12	rip	rip	NOUN
elements-9012	520	13	)	)	PUNCT
elements-9012	520	14	,	,	PUNCT
elements-9012	520	15	protects	protect	VERB
elements-9012	520	16	against	against	ADP
elements-9012	520	17	selfish	selfish	ADJ
elements-9012	520	18	dna	dna	NOUN
elements-9012	520	19	by	by	ADP
elements-9012	520	20	inducing	induce	VERB
elements-9012	520	21	g	g	NOUN
elements-9012	520	22	:	:	PUNCT
elements-9012	520	23	c	c	NOUN
elements-9012	520	24	to	to	ADP
elements-9012	520	25	a	a	DET
elements-9012	520	26	:	:	PUNCT
elements-9012	520	27	t	t	NOUN
elements-9012	520	28	mutations	mutation	NOUN
elements-9012	520	29	during	during	ADP
elements-9012	520	30	sexual	sexual	ADJ
elements-9012	520	31	reproduction	reproduction	NOUN
elements-9012	520	32	and	and	CCONJ
elements-9012	520	33	by	by	ADP
elements-9012	520	34	methylating	methylate	VERB
elements-9012	520	35	cytosine	cytosine	NOUN
elements-9012	520	36	residues	residue	NOUN
elements-9012	520	37	in	in	ADP
elements-9012	520	38	duplicated	duplicate	VERB
elements-9012	520	39	sequences/	sequences/	NUM
elements-9012	520	40	™	™	ADJ
elements-9012	520	41	1	1	NUM
elements-9012	520	42	this	this	DET
elements-9012	520	43	process	process	NOUN
elements-9012	520	44	affects	affect	VERB
elements-9012	520	45	all	all	DET
elements-9012	520	46	duplicated	duplicate	VERB
elements-9012	520	47	regions	region	NOUN
elements-9012	520	48	except	except	SCONJ
elements-9012	520	49	for	for	ADP
elements-9012	520	50	ribosomal	ribosomal	NOUN
elements-9012	520	51	rna	rna	NOUN
elements-9012	520	52	genes	gene	NOUN
elements-9012	520	53	,	,	PUNCT
elements-9012	520	54	even	even	ADV
elements-9012	520	55	though	though	SCONJ
elements-9012	520	56	these	these	PRON
elements-9012	520	57	occur	occur	VERB
elements-9012	520	58	i	i	PRON
elements-9012	520	59	n	n	CCONJ
elements-9012	520	60	large	large	ADJ
elements-9012	520	61	copy	copy	NOUN
elements-9012	520	62	numb	numb	ADJ
elements-9012	520	63	e	e	NOUN
elements-9012	520	64	r	r	NOUN
elements-9012	520	65	.	.	PUNCT
elements-9012	521	1	™	™	NOUN
elements-9012	521	2	1	1	NUM
elements-9012	521	3	this	this	DET
elements-9012	521	4	observation	observation	NOUN
elements-9012	521	5	is	be	AUX
elements-9012	521	6	consistent	consistent	ADJ
elements-9012	521	7	with	with	ADP
elements-9012	521	8	ribosomal	ribosomal	NOUN
elements-9012	521	9	genes	gene	NOUN
elements-9012	521	10	being	be	AUX
elements-9012	521	11	under	under	ADP
elements-9012	521	12	strong	strong	ADJ
elements-9012	521	13	codon	codon	ADJ
elements-9012	521	14	usage	usage	NOUN
elements-9012	521	15	selection	selection	NOUN
elements-9012	521	16	and	and	CCONJ
elements-9012	521	17	hence	hence	ADV
elements-9012	521	18	being	be	AUX
elements-9012	521	19	more	more	ADV
elements-9012	521	20	conserved	conserved	ADJ
elements-9012	521	21	at	at	ADP
elements-9012	521	22	synonymous	synonymous	ADJ
elements-9012	521	23	sites	site	NOUN
elements-9012	521	24	.	.	PUNCT
elements-9012	522	1	rip	rip	NOUN
elements-9012	522	2	drives	drive	NOUN
elements-9012	522	3	other	other	ADJ
elements-9012	522	4	duplicated	duplicate	VERB
elements-9012	522	5	genes	gene	NOUN
elements-9012	522	6	in	in	ADP
elements-9012	522	7	n.	n.	NOUN
elements-9012	522	8	crassa	crassa	NOUN
elements-9012	522	9	to	to	ADP
elements-9012	522	10	unusually	unusually	ADV
elements-9012	522	11	high	high	ADJ
elements-9012	522	12	substitution	substitution	NOUN
elements-9012	522	13	rates	rate	NOUN
elements-9012	522	14	,	,	PUNCT
elements-9012	522	15	which	which	PRON
elements-9012	522	16	would	would	AUX
elements-9012	522	17	explain	explain	VERB
elements-9012	522	18	the	the	DET
elements-9012	522	19	genes	gene	NOUN
elements-9012	522	20	with	with	ADP
elements-9012	522	21	high	high	ADJ
elements-9012	522	22	scores	score	NOUN
elements-9012	522	23	i	i	PRON
elements-9012	522	24	n	n	VERB
elements-9012	522	25	the	the	DET
elements-9012	522	26	rate	rate	NOUN
elements-9012	522	27	distribution	distribution	NOUN
elements-9012	522	28	.	.	PUNCT
elements-9012	523	1	the	the	DET
elements-9012	523	2	wide	wide	ADJ
elements-9012	523	3	substitution	substitution	NOUN
elements-9012	523	4	rate	rate	NOUN
elements-9012	523	5	distribution	distribution	NOUN
elements-9012	523	6	is	be	AUX
elements-9012	523	7	likely	likely	ADJ
elements-9012	523	8	due	due	ADJ
elements-9012	523	9	to	to	PART
elements-9012	523	10	rip	rip	VERB
elements-9012	523	11	in	in	ADP
elements-9012	523	12	just	just	ADV
elements-9012	523	13	n.	n.	NOUN
elements-9012	523	14	crassa	crassa	NOUN
elements-9012	523	15	,	,	PUNCT
elements-9012	523	16	as	as	SCONJ
elements-9012	523	17	the	the	DET
elements-9012	523	18	phenomenon	phenomenon	NOUN
elements-9012	523	19	has	have	AUX
elements-9012	523	20	not	not	PART
elements-9012	523	21	been	be	AUX
elements-9012	523	22	observed	observe	VERB
elements-9012	523	23	in	in	ADP
elements-9012	523	24	c.	c.	PROPN
elements-9012	523	25	globosum	globosum	PROPN
elements-9012	523	26	.	.	PUNCT
elements-9012	524	1	m	m	PROPN
elements-9012	525	1	e	e	NOUN
elements-9012	525	2	t	t	NOUN
elements-9012	525	3	h	h	NOUN
elements-9012	526	1	o	o	X
elements-9012	526	2	d	d	X
elements-9012	526	3	s	s	X
elements-9012	526	4	o	o	X
elements-9012	526	5	r	r	NOUN
elements-9012	526	6	t	t	NOUN
elements-9012	526	7	h	h	NOUN
elements-9012	527	1	o	o	NOUN
elements-9012	527	2	l	l	NOUN
elements-9012	528	1	o	o	NOUN
elements-9012	528	2	c	c	NOUN
elements-9012	528	3	g	g	NOUN
elements-9012	528	4	e	e	PROPN
elements-9012	528	5	n	n	X
elements-9012	528	6	e	e	NOUN
elements-9012	528	7	r	r	NOUN
elements-9012	528	8	a	a	DET
elements-9012	528	9	t	t	NOUN
elements-9012	529	1	i	i	PRON
elements-9012	529	2	o	o	X
elements-9012	529	3	n	n	X
elements-9012	529	4	for	for	ADP
elements-9012	529	5	the	the	DET
elements-9012	529	6	yeast	yeast	NOUN
elements-9012	529	7	analysis	analysis	NOUN
elements-9012	529	8	,	,	PUNCT
elements-9012	529	9	fasta	fasta	PROPN
elements-9012	529	10	files	file	NOUN
elements-9012	529	11	of	of	ADP
elements-9012	529	12	coding	code	VERB
elements-9012	529	13	regions	region	NOUN
elements-9012	529	14	and	and	CCONJ
elements-9012	529	15	an	an	DET
elements-9012	529	16	ortholog	ortholog	NOUN
elements-9012	529	17	tree	tree	NOUN
elements-9012	529	18	listing	list	VERB
elements-9012	529	19	predicted	predict	VERB
elements-9012	529	20	gene	gene	NOUN
elements-9012	529	21	relationships	relationship	NOUN
elements-9012	529	22	between	between	ADP
elements-9012	529	23	all	all	DET
elements-9012	529	24	32	32	NUM
elements-9012	529	25	yeast	yeast	NOUN
elements-9012	529	26	species	specie	NOUN
elements-9012	529	27	were	be	AUX
elements-9012	529	28	obtained	obtain	VERB
elements-9012	529	29	as	as	SCONJ
elements-9012	529	30	described	describe	VERB
elements-9012	529	31	in	in	ADP
elements-9012	529	32	tsong	tsong	ADP
elements-9012	529	33	et	et	PROPN
elements-9012	529	34	a	a	DET
elements-9012	529	35	l	l	NOUN
elements-9012	529	36	.	.	PUNCT
elements-9012	530	1	x	x	PUNCT
elements-9012	530	2	x	x	PUNCT
elements-9012	530	3	x	x	PUNCT
elements-9012	530	4	l	l	NOUN
elements-9012	530	5	v	v	X
elements-9012	530	6	mammalian	mammalian	NOUN
elements-9012	530	7	and	and	CCONJ
elements-9012	530	8	mosquito	mosquito	ADJ
elements-9012	530	9	genes	gene	NOUN
elements-9012	530	10	were	be	AUX
elements-9012	530	11	obtained	obtain	VERB
elements-9012	530	12	using	use	VERB
elements-9012	530	13	the	the	DET
elements-9012	530	14	ensembl	ensembl	NOUN
elements-9012	530	15	biomart	biomart	NOUN
elements-9012	530	16	(	(	PUNCT
elements-9012	530	17	release	release	VERB
elements-9012	530	18	45	45	NUM
elements-9012	530	19	)	)	PUNCT
elements-9012	530	20	database	database	NOUN
elements-9012	530	21	.	.	PUNCT
elements-9012	531	1	fly	fly	VERB
elements-9012	531	2	cds	cd	NOUN
elements-9012	531	3	and	and	CCONJ
elements-9012	531	4	peptide	peptide	ADJ
elements-9012	531	5	files	file	NOUN
elements-9012	531	6	were	be	AUX
elements-9012	531	7	downloaded	download	VERB
elements-9012	531	8	from	from	ADP
elements-9012	531	9	flybase	flybase	NOUN
elements-9012	531	10	(	(	PUNCT
elements-9012	531	11	version	version	NOUN
elements-9012	531	12	fb2oo6_oi	fb2oo6_oi	PROPN
elements-9012	531	13	)	)	PUNCT
elements-9012	531	14	.	.	PUNCT
elements-9012	532	1	for	for	ADP
elements-9012	532	2	the	the	DET
elements-9012	532	3	fly	fly	NOUN
elements-9012	532	4	species	specie	NOUN
elements-9012	532	5	,	,	PUNCT
elements-9012	532	6	the	the	DET
elements-9012	532	7	amino	amino	NOUN
elements-9012	532	8	acid	acid	NOUN
elements-9012	532	9	sequences	sequence	NOUN
elements-9012	532	10	were	be	AUX
elements-9012	532	11	run	run	VERB
elements-9012	532	12	through	through	ADP
elements-9012	532	13	blast	blast	NOUN
elements-9012	532	14	and	and	CCONJ
elements-9012	532	15	tagged	tag	VERB
elements-9012	532	16	as	as	ADP
elements-9012	532	17	true	true	ADJ
elements-9012	532	18	orthologs	ortholog	NOUN
elements-9012	533	1	i	i	PRON
elements-9012	533	2	f	f	VERB
elements-9012	534	1	they	they	PRON
elements-9012	534	2	were	be	AUX
elements-9012	534	3	each	each	DET
elements-9012	534	4	other	other	ADJ
elements-9012	534	5	's	's	PART
elements-9012	534	6	mutual	mutual	ADJ
elements-9012	534	7	best	good	ADJ
elements-9012	534	8	hit	hit	VERB
elements-9012	534	9	when	when	SCONJ
elements-9012	534	10	applying	apply	VERB
elements-9012	534	11	blastall	blastall	NOUN
elements-9012	534	12	with	with	ADP
elements-9012	534	13	a	a	DET
elements-9012	534	14	worst	bad	ADJ
elements-9012	534	15	-	-	PUNCT
elements-9012	534	16	case	case	NOUN
elements-9012	534	17	e	e	NOUN
elements-9012	534	18	-	-	NOUN
elements-9012	534	19	value	value	ADJ
elements-9012	534	20	cutoff	cutoff	NOUN
elements-9012	534	21	of	of	ADP
elements-9012	534	22	10	10	NUM
elements-9012	534	23	"	"	SYM
elements-9012	534	24	1	1	NUM
elements-9012	534	25	0	0	NUM
elements-9012	534	26	.	.	PUNCT
elements-9012	535	1	c	c	NOUN
elements-9012	535	2	a	a	DET
elements-9012	535	3	l	l	NOUN
elements-9012	535	4	c	c	NOUN
elements-9012	535	5	u	u	NOUN
elements-9012	535	6	l	l	NOUN
elements-9012	535	7	a	a	NOUN
elements-9012	535	8	t	t	NOUN
elements-9012	536	1	i	i	PRON
elements-9012	536	2	o	o	VERB
elements-9012	537	1	n	n	X
elements-9012	538	1	o	o	X
elements-9012	538	2	f	f	X
elements-9012	538	3	s	s	PROPN
elements-9012	538	4	u	u	PROPN
elements-9012	538	5	b	b	PROPN
elements-9012	538	6	s	s	ADP
elements-9012	538	7	t	t	X
elements-9012	539	1	i	i	PRON
elements-9012	539	2	t	t	VERB
elements-9012	539	3	u	u	NOUN
elements-9012	540	1	t	t	NOUN
elements-9012	541	1	i	i	PRON
elements-9012	541	2	o	o	VERB
elements-9012	542	1	n	n	CCONJ
elements-9012	542	2	r	r	NOUN
elements-9012	543	1	a	a	DET
elements-9012	543	2	t	t	NOUN
elements-9012	543	3	e	e	NOUN
elements-9012	543	4	s	s	VERB
elements-9012	543	5	our	our	PRON
elements-9012	543	6	substitution	substitution	NOUN
elements-9012	543	7	rate	rate	NOUN
elements-9012	543	8	calculations	calculation	NOUN
elements-9012	543	9	parallel	parallel	VERB
elements-9012	543	10	those	those	PRON
elements-9012	543	11	in	in	ADP
elements-9012	543	12	previous	previous	ADJ
elements-9012	543	13	w	w	PROPN
elements-9012	543	14	o	o	NOUN
elements-9012	543	15	r	r	NOUN
elements-9012	543	16	k	k	PROPN
elements-9012	543	17	s	s	PROPN
elements-9012	543	18	.	.	PUNCT
elements-9012	544	1	x	x	PUNCT
elements-9012	544	2	x	x	PUNCT
elements-9012	544	3	x	x	SYM
elements-9012	544	4	v	v	X
elements-9012	544	5	x	x	SYM
elements-9012	544	6	x	x	SYM
elements-9012	544	7	x	x	SYM
elements-9012	544	8	v	v	ADP
elements-9012	544	9	l	l	NOUN
elements-9012	544	10	nucleotide	nucleotide	NOUN
elements-9012	544	11	sequences	sequence	NOUN
elements-9012	544	12	of	of	ADP
elements-9012	544	13	orthologous	orthologous	ADJ
elements-9012	544	14	coding	code	VERB
elements-9012	544	15	regions	region	NOUN
elements-9012	544	16	were	be	AUX
elements-9012	544	17	translated	translate	VERB
elements-9012	544	18	into	into	ADP
elements-9012	544	19	amino	amino	NOUN
elements-9012	544	20	acid	acid	NOUN
elements-9012	544	21	residues	residue	NOUN
elements-9012	544	22	,	,	PUNCT
elements-9012	544	23	aligned	align	VERB
elements-9012	544	24	using	use	VERB
elements-9012	544	25	clustalw	clustalw	NOUN
elements-9012	544	26	,	,	PUNCT
elements-9012	544	27	and	and	CCONJ
elements-9012	544	28	then	then	ADV
elements-9012	544	29	back	back	ADV
elements-9012	544	30	-	-	PUNCT
elements-9012	544	31	translated	translate	VERB
elements-9012	544	32	to	to	PART
elements-9012	544	33	determine	determine	VERB
elements-9012	544	34	the	the	DET
elements-9012	544	35	aligned	aligned	ADJ
elements-9012	544	36	dna	dna	NOUN
elements-9012	544	37	sequence	sequence	NOUN
elements-9012	544	38	.	.	PUNCT
elements-9012	545	1	the	the	DET
elements-9012	545	2	four	four	NUM
elements-9012	545	3	-	-	ADJ
elements-9012	545	4	fold	fold	ADJ
elements-9012	545	5	synonymous	synonymous	ADJ
elements-9012	545	6	sites	site	NOUN
elements-9012	545	7	—	—	PUNCT
elements-9012	545	8	the	the	DET
elements-9012	545	9	third	third	ADJ
elements-9012	545	10	base	base	NOUN
elements-9012	545	11	i	i	PRON
elements-9012	545	12	n	n	VERB
elements-9012	545	13	a	a	DET
elements-9012	545	14	codon	codon	NOUN
elements-9012	545	15	for	for	ADP
elements-9012	545	16	which	which	PRON
elements-9012	545	17	the	the	DET
elements-9012	545	18	amino	amino	NOUN
elements-9012	545	19	acid	acid	NOUN
elements-9012	545	20	is	be	AUX
elements-9012	545	21	determined	determine	VERB
elements-9012	545	22	by	by	ADP
elements-9012	545	23	the	the	DET
elements-9012	545	24	first	first	ADJ
elements-9012	545	25	two	two	NUM
elements-9012	545	26	positions	position	NOUN
elements-9012	545	27	—	—	PUNCT
elements-9012	545	28	were	be	AUX
elements-9012	545	29	analyzed	analyze	VERB
elements-9012	545	30	.	.	PUNCT
elements-9012	546	1	i	i	PRON
elements-9012	546	2	f	f	PROPN
elements-9012	547	1	a	a	DET
elements-9012	547	2	sequence	sequence	NOUN
elements-9012	547	3	contained	contain	VERB
elements-9012	547	4	fewer	few	ADJ
elements-9012	547	5	than	than	ADP
elements-9012	547	6	20	20	NUM
elements-9012	547	7	four	four	NUM
elements-9012	547	8	-	-	ADJ
elements-9012	547	9	fold	fold	ADJ
elements-9012	547	10	sites	site	NOUN
elements-9012	547	11	before	before	ADP
elements-9012	547	12	occurrence	occurrence	NOUN
elements-9012	547	13	of	of	ADP
elements-9012	547	14	a	a	DET
elements-9012	547	15	stop	stop	NOUN
elements-9012	547	16	codon	codon	NOUN
elements-9012	547	17	,	,	PUNCT
elements-9012	547	18	the	the	DET
elements-9012	547	19	entire	entire	ADJ
elements-9012	547	20	sequence	sequence	NOUN
elements-9012	547	21	was	be	AUX
elements-9012	547	22	discarded	discard	VERB
elements-9012	547	23	.	.	PUNCT
elements-9012	548	1	to	to	PART
elements-9012	548	2	ensure	ensure	VERB
elements-9012	548	3	equivalent	equivalent	ADJ
elements-9012	548	4	sequence	sequence	NOUN
elements-9012	548	5	context	context	NOUN
elements-9012	548	6	,	,	PUNCT
elements-9012	548	7	only	only	ADV
elements-9012	548	8	four	four	NUM
elements-9012	548	9	-	-	ADJ
elements-9012	548	10	fold	fold	ADJ
elements-9012	548	11	sites	site	NOUN
elements-9012	548	12	for	for	ADP
elements-9012	548	13	which	which	PRON
elements-9012	548	14	both	both	CCONJ
elements-9012	548	15	the	the	DET
elements-9012	548	16	preceding	precede	VERB
elements-9012	548	17	and	and	CCONJ
elements-9012	548	18	succeeding	succeed	VERB
elements-9012	548	19	base	base	NOUN
elements-9012	548	20	matched	match	VERB
elements-9012	548	21	for	for	ADP
elements-9012	548	22	the	the	DET
elements-9012	548	23	two	two	NUM
elements-9012	548	24	species	specie	NOUN
elements-9012	548	25	were	be	AUX
elements-9012	548	26	considered	consider	VERB
elements-9012	548	27	.	.	PUNCT
elements-9012	549	1	the	the	DET
elements-9012	549	2	raw	raw	ADJ
elements-9012	549	3	neutral	neutral	ADJ
elements-9012	549	4	substitution	substitution	NOUN
elements-9012	549	5	rate	rate	NOUN
elements-9012	549	6	was	be	AUX
elements-9012	549	7	calculated	calculate	VERB
elements-9012	549	8	based	base	VERB
elements-9012	549	9	on	on	ADP
elements-9012	549	10	the	the	DET
elements-9012	549	11	fraction	fraction	NOUN
elements-9012	549	12	of	of	ADP
elements-9012	549	13	observed	observe	VERB
elements-9012	549	14	differences	difference	NOUN
elements-9012	549	15	at	at	ADP
elements-9012	549	16	silent	silent	ADJ
elements-9012	549	17	sites	site	NOUN
elements-9012	549	18	within	within	ADP
elements-9012	549	19	a	a	DET
elements-9012	549	20	gene	gene	NOUN
elements-9012	549	21	.	.	PUNCT
elements-9012	550	1	individual	individual	ADJ
elements-9012	550	2	gene	gene	NOUN
elements-9012	550	3	rates	rate	NOUN
elements-9012	550	4	were	be	AUX
elements-9012	550	5	then	then	ADV
elements-9012	550	6	normalized	normalize	VERB
elements-9012	550	7	i	i	PRON
elements-9012	550	8	n	n	ADV
elements-9012	550	9	order	order	NOUN
elements-9012	550	10	to	to	PART
elements-9012	550	11	correct	correct	VERB
elements-9012	550	12	for	for	ADP
elements-9012	550	13	the	the	DET
elements-9012	550	14	finite	finite	NOUN
elements-9012	550	15	-	-	NOUN
elements-9012	550	16	size	size	NOUN
elements-9012	550	17	of	of	ADP
elements-9012	550	18	each	each	DET
elements-9012	550	19	gene	gene	NOUN
elements-9012	550	20	(	(	PUNCT
elements-9012	550	21	table	table	NOUN
elements-9012	550	22	1	1	NUM
elements-9012	550	23	&	&	CCONJ
elements-9012	550	24	2	2	NUM
elements-9012	550	25	)	)	PUNCT
elements-9012	550	26	,	,	PUNCT
elements-9012	550	27	and	and	CCONJ
elements-9012	550	28	this	this	DET
elements-9012	550	29	new	new	ADJ
elements-9012	550	30	rate	rate	NOUN
elements-9012	550	31	was	be	AUX
elements-9012	550	32	defined	define	VERB
elements-9012	550	33	to	to	PART
elements-9012	550	34	be	be	AUX
elements-9012	550	35	r=(p-<p>)/s(n	r=(p-<p>)/s(n	NOUN
elements-9012	550	36	)	)	PUNCT
elements-9012	550	37	,	,	PUNCT
elements-9012	550	38	where	where	SCONJ
elements-9012	550	39	p	p	NOUN
elements-9012	550	40	is	be	AUX
elements-9012	550	41	the	the	DET
elements-9012	550	42	observed	observe	VERB
elements-9012	550	43	four	four	NUM
elements-9012	550	44	-	-	ADJ
elements-9012	550	45	fold	fold	ADJ
elements-9012	550	46	substitution	substitution	NOUN
elements-9012	550	47	rate	rate	NOUN
elements-9012	550	48	for	for	ADP
elements-9012	550	49	the	the	DET
elements-9012	550	50	gene	gene	NOUN
elements-9012	550	51	i	i	PRON
elements-9012	550	52	n	n	PRON
elements-9012	550	53	question	question	NOUN
elements-9012	550	54	and	and	CCONJ
elements-9012	550	55	<	<	X
elements-9012	550	56	p	p	X
elements-9012	550	57	>	>	X
elements-9012	550	58	is	be	AUX
elements-9012	550	59	the	the	DET
elements-9012	550	60	average	average	ADJ
elements-9012	550	61	substitution	substitution	NOUN
elements-9012	550	62	rate	rate	NOUN
elements-9012	550	63	for	for	ADP
elements-9012	550	64	all	all	DET
elements-9012	550	65	ortholog	ortholog	NOUN
elements-9012	550	66	pairs	pair	NOUN
elements-9012	550	67	for	for	ADP
elements-9012	550	68	the	the	DET
elements-9012	550	69	two	two	NUM
elements-9012	550	70	species	specie	NOUN
elements-9012	550	71	in	in	ADP
elements-9012	550	72	question	question	NOUN
elements-9012	550	73	.	.	PUNCT
elements-9012	551	1	s(n	s(n	NOUN
elements-9012	551	2	)	)	PUNCT
elements-9012	551	3	was	be	AUX
elements-9012	551	4	defined	define	VERB
elements-9012	551	5	to	to	PART
elements-9012	551	6	be	be	AUX
elements-9012	551	7	the	the	DET
elements-9012	551	8	expected	expect	VERB
elements-9012	551	9	standard	standard	ADJ
elements-9012	551	10	deviation	deviation	NOUN
elements-9012	551	11	for	for	ADP
elements-9012	551	12	a	a	DET
elements-9012	551	13	gene	gene	NOUN
elements-9012	551	14	with	with	ADP
elements-9012	551	15	n	n	CCONJ
elements-9012	551	16	independent	independent	ADJ
elements-9012	551	17	four	four	NUM
elements-9012	551	18	-	-	ADJ
elements-9012	551	19	fold	fold	ADJ
elements-9012	551	20	sites	site	NOUN
elements-9012	551	21	,	,	PUNCT
elements-9012	551	22	i.e.	i.e.	X
elements-9012	551	23	s(n	s(n	NOUN
elements-9012	551	24	)	)	PUNCT
elements-9012	551	25	=	=	PUNCT
elements-9012	552	1	(	(	PUNCT
elements-9012	552	2	<	<	X
elements-9012	552	3	p>(i	p>(i	X
elements-9012	552	4	<	<	X
elements-9012	552	5	p	p	X
elements-9012	552	6	>	>	X
elements-9012	552	7	)	)	PUNCT
elements-9012	552	8	/n	/n	SYM
elements-9012	552	9	)	)	PUNCT
elements-9012	552	10	1	1	NUM
elements-9012	552	11	/	/	SYM
elements-9012	552	12	2	2	NUM
elements-9012	552	13	.	.	PUNCT
elements-9012	553	1	the	the	DET
elements-9012	553	2	distribution	distribution	NOUN
elements-9012	553	3	of	of	ADP
elements-9012	553	4	the	the	DET
elements-9012	553	5	normalized	normalize	VERB
elements-9012	553	6	substitution	substitution	NOUN
elements-9012	553	7	rates	rate	NOUN
elements-9012	553	8	would	would	AUX
elements-9012	553	9	be	be	AUX
elements-9012	553	10	expected	expect	VERB
elements-9012	553	11	to	to	PART
elements-9012	553	12	follow	follow	VERB
elements-9012	553	13	a	a	DET
elements-9012	553	14	normal	normal	ADJ
elements-9012	553	15	gaussian	gaussian	NOUN
elements-9012	553	16	(	(	PUNCT
elements-9012	553	17	f(x	f(x	PROPN
elements-9012	553	18	)	)	PUNCT
elements-9012	553	19	=	=	PUNCT
elements-9012	554	1	e	e	X
elements-9012	554	2	x	x	PUNCT
elements-9012	554	3	"	"	PUNCT
elements-9012	554	4	x	x	SYM
elements-9012	554	5	/	/	SYM
elements-9012	554	6	2	2	NUM
elements-9012	554	7	/	/	SYM
elements-9012	554	8	v(2tt	v(2tt	NOUN
elements-9012	554	9	)	)	PUNCT
elements-9012	554	10	)	)	PUNCT
elements-9012	555	1	i	i	PRON
elements-9012	555	2	f	f	PROPN
elements-9012	556	1	each	each	DET
elements-9012	556	2	four	four	NUM
elements-9012	556	3	-	-	ADJ
elements-9012	556	4	fold	fold	ADJ
elements-9012	556	5	site	site	NOUN
elements-9012	556	6	was	be	AUX
elements-9012	556	7	mutating	mutate	VERB
elements-9012	556	8	at	at	ADP
elements-9012	556	9	the	the	DET
elements-9012	556	10	same	same	ADJ
elements-9012	556	11	rate	rate	NOUN
elements-9012	556	12	and	and	CCONJ
elements-9012	556	13	independently	independently	ADV
elements-9012	556	14	of	of	ADP
elements-9012	556	15	each	each	DET
elements-9012	556	16	other	other	ADJ
elements-9012	556	17	.	.	PUNCT
elements-9012	557	1	c	c	NOUN
elements-9012	558	1	o	o	NOUN
elements-9012	558	2	r	r	NOUN
elements-9012	558	3	r	r	NOUN
elements-9012	558	4	e	e	NOUN
elements-9012	558	5	l	l	NOUN
elements-9012	558	6	a	a	NOUN
elements-9012	558	7	t	t	NOUN
elements-9012	559	1	i	i	PRON
elements-9012	559	2	o	o	VERB
elements-9012	560	1	n	n	VERB
elements-9012	560	2	c	c	NOUN
elements-9012	560	3	a	a	DET
elements-9012	560	4	l	l	NOUN
elements-9012	560	5	c	c	NOUN
elements-9012	561	1	u	u	NOUN
elements-9012	561	2	l	l	NOUN
elements-9012	561	3	a	a	NOUN
elements-9012	561	4	t	t	NOUN
elements-9012	562	1	i	i	PRON
elements-9012	562	2	o	o	VERB
elements-9012	563	1	n	n	CCONJ
elements-9012	563	2	s	s	VERB
elements-9012	563	3	we	we	PRON
elements-9012	563	4	tested	test	VERB
elements-9012	563	5	whether	whether	SCONJ
elements-9012	563	6	or	or	CCONJ
elements-9012	563	7	not	not	PART
elements-9012	563	8	neighboring	neighboring	NOUN
elements-9012	563	9	genes	gene	NOUN
elements-9012	563	10	had	have	VERB
elements-9012	563	11	similar	similar	ADJ
elements-9012	563	12	substitution	substitution	NOUN
elements-9012	563	13	rates	rate	NOUN
elements-9012	563	14	by	by	ADP
elements-9012	563	15	calculating	calculate	VERB
elements-9012	563	16	a	a	DET
elements-9012	563	17	pearson	pearson	NOUN
elements-9012	563	18	correlation	correlation	NOUN
elements-9012	563	19	between	between	ADP
elements-9012	563	20	the	the	DET
elements-9012	563	21	rate	rate	NOUN
elements-9012	563	22	of	of	ADP
elements-9012	563	23	gene	gene	NOUN
elements-9012	563	24	r(o	r(o	PROPN
elements-9012	563	25	)	)	PUNCT
elements-9012	563	26	and	and	CCONJ
elements-9012	563	27	the	the	DET
elements-9012	563	28	rate	rate	NOUN
elements-9012	563	29	of	of	ADP
elements-9012	563	30	gene	gene	NOUN
elements-9012	563	31	r	r	NOUN
elements-9012	563	32	1	1	NUM
elements-9012	563	33	®	®	NOUN
elements-9012	563	34	which	which	PRON
elements-9012	563	35	is	be	AUX
elements-9012	563	36	located	locate	VERB
elements-9012	563	37	x	x	PUNCT
elements-9012	563	38	base	base	NOUN
elements-9012	563	39	pairs	pair	NOUN
elements-9012	563	40	downstream	downstream	ADJ
elements-9012	563	41	.	.	PUNCT
elements-9012	564	1	gene	gene	NOUN
elements-9012	564	2	pairs	pair	NOUN
elements-9012	564	3	in	in	ADP
elements-9012	564	4	orthologous	orthologous	ADJ
elements-9012	564	5	blocks	block	NOUN
elements-9012	564	6	up	up	ADP
elements-9012	564	7	to	to	ADP
elements-9012	564	8	the	the	DET
elements-9012	564	9	35th	35th	ADJ
elements-9012	564	10	gene	gene	NOUN
elements-9012	564	11	downstream	downstream	ADJ
elements-9012	564	12	from	from	ADP
elements-9012	564	13	the	the	DET
elements-9012	564	14	starting	starting	NOUN
elements-9012	564	15	gene	gene	NOUN
elements-9012	564	16	were	be	AUX
elements-9012	564	17	considered	consider	VERB
elements-9012	564	18	.	.	PUNCT
elements-9012	565	1	blocks	block	NOUN
elements-9012	565	2	were	be	AUX
elements-9012	565	3	determined	determine	VERB
elements-9012	565	4	by	by	ADP
elements-9012	565	5	m	m	PROPN
elements-9012	565	6	e	e	NOUN
elements-9012	565	7	a	a	PRON
elements-9012	565	8	s	s	X
elements-9012	565	9	u	u	NOUN
elements-9012	565	10	r	r	NOUN
elements-9012	565	11	i	i	PRON
elements-9012	565	12	n	n	CCONJ
elements-9012	565	13	g	g	PROPN
elements-9012	565	14	t	t	NOUN
elements-9012	565	15	h	h	NOUN
elements-9012	566	1	e	e	X
elements-9012	566	2	u	u	NOUN
elements-9012	566	3	n	n	PROPN
elements-9012	566	4	i	i	PRON
elements-9012	566	5	v	v	VERB
elements-9012	566	6	e	e	NOUN
elements-9012	566	7	r	r	NOUN
elements-9012	566	8	s	s	PROPN
elements-9012	566	9	a	a	DET
elements-9012	566	10	l	l	NOUN
elements-9012	567	1	i	i	PRON
elements-9012	567	2	t	t	VERB
elements-9012	568	1	y	y	NOUN
elements-9012	569	1	o	o	X
elements-9012	569	2	f	f	NOUN
elements-9012	570	1	r	r	NOUN
elements-9012	570	2	e	e	NOUN
elements-9012	570	3	g	g	NOUN
elements-9012	571	1	i	i	PRON
elements-9012	571	2	o	o	VERB
elements-9012	572	1	n	n	CCONJ
elements-9012	572	2	a	a	DET
elements-9012	572	3	l	l	NOUN
elements-9012	572	4	m	m	VERB
elements-9012	573	1	u	u	NOUN
elements-9012	573	2	t	t	PROPN
elements-9012	573	3	a	a	DET
elements-9012	573	4	t	t	NOUN
elements-9012	574	1	i	i	PRON
elements-9012	574	2	o	o	VERB
elements-9012	575	1	n	n	CCONJ
elements-9012	575	2	r	r	NOUN
elements-9012	575	3	a	a	DET
elements-9012	575	4	t	t	NOUN
elements-9012	575	5	e	e	NOUN
elements-9012	575	6	s	s	NOUN
elements-9012	575	7	genes	gene	NOUN
elements-9012	575	8	located	locate	VERB
elements-9012	575	9	on	on	ADP
elements-9012	575	10	the	the	DET
elements-9012	575	11	same	same	ADJ
elements-9012	575	12	chromosome	chromosome	NOUN
elements-9012	575	13	(	(	PUNCT
elements-9012	575	14	scaffolds	scaffold	NOUN
elements-9012	575	15	were	be	AUX
elements-9012	575	16	used	use	VERB
elements-9012	575	17	when	when	SCONJ
elements-9012	575	18	chromosomal	chromosomal	ADJ
elements-9012	575	19	data	datum	NOUN
elements-9012	575	20	was	be	AUX
elements-9012	575	21	not	not	PART
elements-9012	575	22	available	available	ADJ
elements-9012	575	23	)	)	PUNCT
elements-9012	575	24	.	.	PUNCT
elements-9012	576	1	correlations	correlation	NOUN
elements-9012	576	2	were	be	AUX
elements-9012	576	3	measured	measure	VERB
elements-9012	576	4	twice	twice	ADV
elements-9012	576	5	,	,	PUNCT
elements-9012	576	6	in	in	ADP
elements-9012	576	7	each	each	DET
elements-9012	576	8	case	case	NOUN
elements-9012	576	9	using	use	VERB
elements-9012	576	10	location	location	NOUN
elements-9012	576	11	data	datum	NOUN
elements-9012	576	12	from	from	ADP
elements-9012	576	13	one	one	NUM
elements-9012	576	14	of	of	ADP
elements-9012	576	15	the	the	DET
elements-9012	576	16	species	specie	NOUN
elements-9012	576	17	,	,	PUNCT
elements-9012	576	18	except	except	SCONJ
elements-9012	576	19	in	in	ADP
elements-9012	576	20	cases	case	NOUN
elements-9012	576	21	where	where	SCONJ
elements-9012	576	22	location	location	NOUN
elements-9012	576	23	data	datum	NOUN
elements-9012	576	24	was	be	AUX
elements-9012	576	25	available	available	ADJ
elements-9012	576	26	in	in	ADP
elements-9012	576	27	only	only	ADV
elements-9012	576	28	one	one	NUM
elements-9012	576	29	species	specie	NOUN
elements-9012	576	30	.	.	PUNCT
elements-9012	577	1	for	for	ADP
elements-9012	577	2	yeast	yeast	NOUN
elements-9012	577	3	,	,	PUNCT
elements-9012	577	4	the	the	DET
elements-9012	577	5	data	datum	NOUN
elements-9012	577	6	for	for	ADP
elements-9012	577	7	each	each	DET
elements-9012	577	8	pairwise	pairwise	NOUN
elements-9012	577	9	calculation	calculation	NOUN
elements-9012	577	10	was	be	AUX
elements-9012	577	11	binned	bin	VERB
elements-9012	577	12	into	into	ADP
elements-9012	577	13	50	50	NUM
elements-9012	577	14	uniformly	uniformly	ADV
elements-9012	577	15	spaced	space	VERB
elements-9012	577	16	groups	group	NOUN
elements-9012	577	17	covering	cover	VERB
elements-9012	577	18	x	x	X
elements-9012	577	19	=	=	PRON
elements-9012	577	20	{	{	PUNCT
elements-9012	577	21	o	o	NOUN
elements-9012	577	22	,	,	PUNCT
elements-9012	577	23	300000	300000	NUM
elements-9012	577	24	}	}	PUNCT
elements-9012	577	25	and	and	CCONJ
elements-9012	577	26	averaged	average	VERB
elements-9012	577	27	over	over	ADP
elements-9012	577	28	each	each	DET
elements-9012	577	29	bin	bin	NOUN
elements-9012	577	30	to	to	PART
elements-9012	577	31	determine	determine	VERB
elements-9012	577	32	the	the	DET
elements-9012	577	33	autocorrelation	autocorrelation	NOUN
elements-9012	577	34	function	function	NOUN
elements-9012	577	35	<	<	X
elements-9012	577	36	r(o)r	r(o)r	PROPN
elements-9012	577	37	x	x	SYM
elements-9012	577	38	^	^	SYM
elements-9012	577	39	>	>	PROPN
elements-9012	577	40	.	.	PUNCT
elements-9012	578	1	error	error	NOUN
elements-9012	578	2	bars	bar	NOUN
elements-9012	578	3	were	be	AUX
elements-9012	578	4	assigned	assign	VERB
elements-9012	578	5	based	base	VERB
elements-9012	578	6	on	on	ADP
elements-9012	578	7	the	the	DET
elements-9012	578	8	standard	standard	ADJ
elements-9012	578	9	deviation	deviation	NOUN
elements-9012	578	10	of	of	ADP
elements-9012	578	11	the	the	DET
elements-9012	578	12	values	value	NOUN
elements-9012	578	13	in	in	ADP
elements-9012	578	14	each	each	DET
elements-9012	578	15	bin	bin	NOUN
elements-9012	578	16	.	.	PUNCT
elements-9012	579	1	for	for	ADP
elements-9012	579	2	the	the	DET
elements-9012	579	3	larger	large	ADJ
elements-9012	579	4	genomes	genome	NOUN
elements-9012	579	5	of	of	ADP
elements-9012	579	6	mammals	mammal	NOUN
elements-9012	579	7	and	and	CCONJ
elements-9012	579	8	insects	insect	NOUN
elements-9012	579	9	,	,	PUNCT
elements-9012	579	10	data	datum	NOUN
elements-9012	579	11	was	be	AUX
elements-9012	579	12	binned	bin	VERB
elements-9012	579	13	into	into	ADP
elements-9012	579	14	200	200	NUM
elements-9012	579	15	groups	group	NOUN
elements-9012	579	16	where	where	SCONJ
elements-9012	579	17	x	x	PRON
elements-9012	579	18	ranged	range	VERB
elements-9012	579	19	from	from	ADP
elements-9012	579	20	o	o	NOUN
elements-9012	579	21	to	to	ADP
elements-9012	579	22	15	15	NUM
elements-9012	579	23	mb	mb	NOUN
elements-9012	579	24	.	.	PUNCT
elements-9012	580	1	c	c	NOUN
elements-9012	581	1	a	a	PRON
elements-9012	581	2	i	i	PRON
elements-9012	581	3	codonw	codonw	NOUN
elements-9012	581	4	(	(	PUNCT
elements-9012	581	5	peden	peden	PROPN
elements-9012	581	6	1999	1999	NUM
elements-9012	581	7	)	)	PUNCT
elements-9012	581	8	was	be	AUX
elements-9012	581	9	downloaded	download	VERB
elements-9012	581	10	and	and	CCONJ
elements-9012	581	11	used	use	VERB
elements-9012	581	12	to	to	PART
elements-9012	581	13	calculate	calculate	VERB
elements-9012	581	14	the	the	DET
elements-9012	581	15	cai	cai	PROPN
elements-9012	581	16	values	value	NOUN
elements-9012	581	17	for	for	ADP
elements-9012	581	18	the	the	DET
elements-9012	581	19	yeast	yeast	NOUN
elements-9012	581	20	species	specie	NOUN
elements-9012	581	21	.	.	PUNCT
elements-9012	582	1	the	the	DET
elements-9012	582	2	input	input	NOUN
elements-9012	582	3	file	file	NOUN
elements-9012	582	4	for	for	ADP
elements-9012	582	5	each	each	DET
elements-9012	582	6	species	specie	NOUN
elements-9012	582	7	was	be	AUX
elements-9012	582	8	a	a	DET
elements-9012	582	9	cds	cd	NOUN
elements-9012	582	10	fasta	fasta	PROPN
elements-9012	582	11	file	file	NOUN
elements-9012	582	12	of	of	ADP
elements-9012	582	13	all	all	DET
elements-9012	582	14	genes	gene	NOUN
elements-9012	582	15	(	(	PUNCT
elements-9012	582	16	predicted	predict	VERB
elements-9012	582	17	and	and	CCONJ
elements-9012	582	18	known	know	VERB
elements-9012	582	19	)	)	PUNCT
elements-9012	582	20	and	and	CCONJ
elements-9012	582	21	the	the	DET
elements-9012	582	22	background	background	NOUN
elements-9012	582	23	cai	cai	PROPN
elements-9012	582	24	was	be	AUX
elements-9012	582	25	set	set	VERB
elements-9012	582	26	to	to	ADP
elements-9012	582	27	saccharomyces	saccharomyce	NOUN
elements-9012	582	28	cerevisiae	cerevisiae	NOUN
elements-9012	582	29	for	for	ADP
elements-9012	582	30	all	all	DET
elements-9012	582	31	sensu	sensu	NOUN
elements-9012	582	32	stricto	stricto	X
elements-9012	582	33	.	.	PUNCT
elements-9012	583	1	the	the	DET
elements-9012	583	2	emboss	emboss	NOUN
elements-9012	583	3	package	package	NOUN
elements-9012	583	4	was	be	AUX
elements-9012	583	5	also	also	ADV
elements-9012	583	6	downloaded	download	VERB
elements-9012	583	7	locally	locally	ADV
elements-9012	583	8	.	.	PUNCT
elements-9012	584	1	this	this	PRON
elements-9012	584	2	includes	include	VERB
elements-9012	584	3	codon	codon	ADJ
elements-9012	584	4	usage	usage	NOUN
elements-9012	584	5	tables	table	NOUN
elements-9012	584	6	for	for	ADP
elements-9012	584	7	a	a	DET
elements-9012	584	8	number	number	NOUN
elements-9012	584	9	of	of	ADP
elements-9012	584	10	species	specie	NOUN
elements-9012	584	11	including	include	VERB
elements-9012	584	12	n.	n.	NOUN
elements-9012	584	13	crassa.xxxvn	crassa.xxxvn	ADP
elements-9012	584	14	this	this	DET
elements-9012	584	15	table	table	NOUN
elements-9012	584	16	was	be	AUX
elements-9012	584	17	used	use	VERB
elements-9012	584	18	to	to	PART
elements-9012	584	19	calculate	calculate	VERB
elements-9012	584	20	the	the	DET
elements-9012	584	21	cai	cai	NOUN
elements-9012	584	22	for	for	ADP
elements-9012	584	23	the	the	DET
elements-9012	584	24	genes	gene	NOUN
elements-9012	584	25	in	in	ADP
elements-9012	584	26	n.	n.	NOUN
elements-9012	584	27	crassa	crassa	NOUN
elements-9012	584	28	.	.	PUNCT
elements-9012	585	1	e	e	NOUN
elements-9012	585	2	n	n	PROPN
elements-9012	585	3	d	d	PROPN
elements-9012	585	4	n	n	NUM
elements-9012	585	5	o	o	NOUN
elements-9012	585	6	t	t	X
elements-9012	585	7	e	e	X
elements-9012	585	8	s	s	VERB
elements-9012	585	9	i	i	PRON
elements-9012	585	10	.	.	PUNCT
elements-9012	586	1	baer	baer	PROPN
elements-9012	586	2	2007	2007	NUM
elements-9012	587	1	i	i	PRON
elements-9012	587	2	i	i	INTJ
elements-9012	587	3	.	.	PUNCT
elements-9012	588	1	chuang	chuang	PROPN
elements-9012	588	2	2004	2004	NUM
elements-9012	589	1	i	i	PRON
elements-9012	589	2	i	i	PRON
elements-9012	589	3	i	i	PRON
elements-9012	589	4	.	.	PUNCT
elements-9012	590	1	lercher	lercher	PROPN
elements-9012	590	2	2001	2001	NUM
elements-9012	590	3	vi	vi	NOUN
elements-9012	590	4	.	.	PUNCT
elements-9012	591	1	liu	liu	PROPN
elements-9012	591	2	2006	2006	NUM
elements-9012	591	3	v.	v.	CCONJ
elements-9012	591	4	malcom	malcom	PROPN
elements-9012	591	5	2003	2003	NUM
elements-9012	591	6	vi	vi	PROPN
elements-9012	591	7	.	.	PUNCT
elements-9012	592	1	matassi	matassi	PROPN
elements-9012	592	2	1999	1999	NUM
elements-9012	592	3	v	v	NOUN
elements-9012	592	4	i	i	PRON
elements-9012	592	5	i	i	PRON
elements-9012	592	6	.	.	PUNCT
elements-9012	593	1	wolfe	wolfe	PROPN
elements-9012	593	2	1989	1989	NUM
elements-9012	593	3	v	v	PROPN
elements-9012	594	1	i	i	PRON
elements-9012	594	2	i	i	PRON
elements-9012	595	1	i	i	PRON
elements-9012	595	2	.	.	PUNCT
elements-9012	596	1	chin	chin	NOUN
elements-9012	596	2	2005	2005	NUM
elements-9012	596	3	ix	ix	PROPN
elements-9012	596	4	.	.	PUNCT
elements-9012	596	5	pheasant	pheasant	NOUN
elements-9012	596	6	2007	2007	NUM
elements-9012	596	7	x.	x.	NOUN
elements-9012	596	8	hardison	hardison	PROPN
elements-9012	596	9	2003	2003	NUM
elements-9012	596	10	xi	xi	X
elements-9012	596	11	.	.	PUNCT
elements-9012	597	1	chuang	chuang	PROPN
elements-9012	597	2	2004	2004	NUM
elements-9012	597	3	xi	xi	INTJ
elements-9012	598	1	i	i	PROPN
elements-9012	598	2	.	.	PUNCT
elements-9012	599	1	baer	baer	PROPN
elements-9012	599	2	2007	2007	NUM
elements-9012	599	3	xi	xi	INTJ
elements-9012	600	1	i	i	PRON
elements-9012	601	1	i	i	INTJ
elements-9012	601	2	.	.	PUNCT
elements-9012	602	1	chamary	chamary	ADJ
elements-9012	602	2	2006	2006	NUM
elements-9012	602	3	xiv	xiv	PROPN
elements-9012	602	4	.	.	PUNCT
elements-9012	603	1	fitzpatrick	fitzpatrick	PROPN
elements-9012	603	2	2006	2006	NUM
elements-9012	603	3	xv	xv	PROPN
elements-9012	603	4	.	.	PUNCT
elements-9012	604	1	murphy	murphy	PROPN
elements-9012	604	2	2007	2007	NUM
elements-9012	604	3	xvi	xvi	NOUN
elements-9012	604	4	.	.	PUNCT
elements-9012	605	1	nikolaev	nikolaev	X
elements-9012	605	2	2007	2007	NUM
elements-9012	605	3	xvii	xvii	PROPN
elements-9012	605	4	.	.	PUNCT
elements-9012	606	1	chuang	chuang	PROPN
elements-9012	606	2	2004	2004	NUM
elements-9012	606	3	xvii	xvii	VERB
elements-9012	606	4	i	i	PRON
elements-9012	606	5	.	.	PUNCT
elements-9012	607	1	chin	chin	NOUN
elements-9012	607	2	2005	2005	NUM
elements-9012	607	3	xix	xix	PROPN
elements-9012	607	4	.	.	PUNCT
elements-9012	607	5	ibid	ibid	PROPN
elements-9012	607	6	.	.	PUNCT
elements-9012	608	1	xx	xx	X
elements-9012	608	2	.	.	PUNCT
elements-9012	608	3	ibid	ibid	PROPN
elements-9012	608	4	.	.	PUNCT
elements-9012	609	1	xxi	xxi	PROPN
elements-9012	609	2	.	.	PUNCT
elements-9012	610	1	chuang	chuang	PROPN
elements-9012	610	2	2004	2004	NUM
elements-9012	610	3	xxii	xxii	PROPN
elements-9012	610	4	.	.	PUNCT
elements-9012	611	1	m	m	VERB
elements-9012	611	2	i	i	PRON
elements-9012	611	3	2007	2007	NUM
elements-9012	611	4	xxii	xxii	NOUN
elements-9012	612	1	i	i	PRON
elements-9012	612	2	.	.	PUNCT
elements-9012	613	1	lott	lott	PROPN
elements-9012	613	2	2005	2005	NUM
elements-9012	614	1	xxiv	xxiv	PROPN
elements-9012	614	2	.	.	PUNCT
elements-9012	615	1	yokogawa	yokogawa	PROPN
elements-9012	615	2	1992	1992	NUM
elements-9012	615	3	xxv	xxv	PROPN
elements-9012	615	4	.	.	PUNCT
elements-9012	616	1	lunter	lunter	PROPN
elements-9012	616	2	2006	2006	NUM
elements-9012	616	3	xxvi	xxvi	NOUN
elements-9012	616	4	.	.	PUNCT
elements-9012	617	1	baer	baer	PROPN
elements-9012	617	2	2007	2007	NUM
elements-9012	617	3	xxvh	xxvh	PROPN
elements-9012	617	4	.	.	PUNCT
elements-9012	618	1	ruderfe	ruderfe	PROPN
elements-9012	618	2	2006	2006	NUM
elements-9012	618	3	xxviii	xxviii	PROPN
elements-9012	618	4	.	.	PUNCT
elements-9012	619	1	jones	jones	PROPN
elements-9012	619	2	2004	2004	NUM
elements-9012	619	3	xxix	xxix	PROPN
elements-9012	619	4	.	.	PUNCT
elements-9012	620	1	pujol	pujol	PROPN
elements-9012	620	2	2004	2004	NUM
elements-9012	620	3	xxx	xxx	NOUN
elements-9012	620	4	.	.	PUNCT
elements-9012	621	1	wong	wong	PROPN
elements-9012	621	2	2003	2003	NUM
elements-9012	621	3	xxxi	xxxi	PROPN
elements-9012	621	4	.	.	PUNCT
elements-9012	622	1	logue	logue	PROPN
elements-9012	622	2	2005	2005	NUM
elements-9012	622	3	xxxii	xxxii	PROPN
elements-9012	622	4	.	.	PUNCT
elements-9012	623	1	selke	selke	ADJ
elements-9012	623	2	1990	1990	NUM
elements-9012	623	3	xxxiii	xxxiii	NOUN
elements-9012	623	4	.	.	PUNCT
elements-9012	624	1	vyas	vyas	PROPN
elements-9012	624	2	2005	2005	NUM
elements-9012	624	3	xxxiv	xxxiv	PROPN
elements-9012	624	4	.	.	PUNCT
elements-9012	625	1	tsong	tsong	ADP
elements-9012	625	2	2006	2006	NUM
elements-9012	625	3	xxxv	xxxv	PROPN
elements-9012	625	4	.	.	PUNCT
elements-9012	626	1	chuang	chuang	PROPN
elements-9012	626	2	2004	2004	NUM
elements-9012	626	3	xxxvi	xxxvi	PROPN
elements-9012	626	4	.	.	PUNCT
elements-9012	627	1	chin	chin	PROPN
elements-9012	627	2	2005	2005	NUM
elements-9012	627	3	xxxvii	xxxvii	PROPN
elements-9012	627	4	.	.	PUNCT
elements-9012	628	1	ikemura	ikemura	PROPN
elements-9012	628	2	1985	1985	NUM
elements-9012	628	3	references	reference	NOUN
elements-9012	628	4	baer	baer	PROPN
elements-9012	628	5	,	,	PUNCT
elements-9012	628	6	c.	c.	PROPN
elements-9012	628	7	f.	f.	PROPN
elements-9012	628	8	,	,	PUNCT
elements-9012	628	9	miyamoto	miyamoto	NOUN
elements-9012	628	10	,	,	PUNCT
elements-9012	628	11	m	m	PROPN
elements-9012	628	12	.	.	PUNCT
elements-9012	629	1	m	m	PROPN
elements-9012	629	2	.	.	PUNCT
elements-9012	629	3	&	&	CCONJ
elements-9012	629	4	denver	denver	PROPN
elements-9012	629	5	,	,	PUNCT
elements-9012	629	6	d.	d.	PROPN
elements-9012	629	7	r.	r.	PROPN
elements-9012	629	8	mutation	mutation	PROPN
elements-9012	629	9	rate	rate	NOUN
elements-9012	629	10	variation	variation	NOUN
elements-9012	630	1	i	i	PRON
elements-9012	630	2	n	n	VERB
elements-9012	630	3	multicellular	multicellular	ADJ
elements-9012	630	4	eukaryotes	eukaryote	NOUN
elements-9012	630	5	:	:	PUNCT
elements-9012	630	6	causes	cause	NOUN
elements-9012	630	7	and	and	CCONJ
elements-9012	630	8	consequences	consequence	NOUN
elements-9012	630	9	.	.	PUNCT
elements-9012	631	1	nat	nat	PROPN
elements-9012	631	2	rev	rev	PROPN
elements-9012	631	3	genet	genet	PROPN
elements-9012	631	4	8	8	NUM
elements-9012	631	5	,	,	PUNCT
elements-9012	631	6	619	619	NUM
elements-9012	631	7	-	-	SYM
elements-9012	631	8	31	31	NUM
elements-9012	631	9	(	(	PUNCT
elements-9012	631	10	2007	2007	NUM
elements-9012	631	11	)	)	PUNCT
elements-9012	631	12	.	.	PUNCT
elements-9012	632	1	chamary	chamary	PROPN
elements-9012	632	2	,	,	PUNCT
elements-9012	632	3	j.	j.	PROPN
elements-9012	632	4	v.	v.	PROPN
elements-9012	632	5	,	,	PUNCT
elements-9012	632	6	parmley	parmley	PROPN
elements-9012	632	7	,	,	PUNCT
elements-9012	632	8	j.	j.	PROPN
elements-9012	632	9	l.	l.	PROPN
elements-9012	632	10	&	&	CCONJ
elements-9012	632	11	hurst	hurst	PROPN
elements-9012	632	12	,	,	PUNCT
elements-9012	632	13	l.	l.	PROPN
elements-9012	632	14	d.	d.	PROPN
elements-9012	632	15	hearing	hearing	PROPN
elements-9012	632	16	silence	silence	NOUN
elements-9012	632	17	:	:	PUNCT
elements-9012	632	18	non	non	ADJ
elements-9012	632	19	-	-	ADJ
elements-9012	632	20	neutral	neutral	ADJ
elements-9012	632	21	evolution	evolution	NOUN
elements-9012	632	22	at	at	ADP
elements-9012	632	23	synonymous	synonymous	ADJ
elements-9012	632	24	sites	site	NOUN
elements-9012	632	25	i	i	PRON
elements-9012	632	26	n	n	VERB
elements-9012	632	27	mammals	mammal	NOUN
elements-9012	632	28	.	.	PUNCT
elements-9012	633	1	nat	nat	PROPN
elements-9012	633	2	rev	rev	PROPN
elements-9012	633	3	genet	genet	PROPN
elements-9012	633	4	7	7	NUM
elements-9012	633	5	,	,	PUNCT
elements-9012	633	6	98	98	NUM
elements-9012	633	7	-	-	SYM
elements-9012	633	8	108	108	NUM
elements-9012	633	9	(	(	PUNCT
elements-9012	633	10	2006	2006	NUM
elements-9012	633	11	)	)	PUNCT
elements-9012	633	12	.	.	PUNCT
elements-9012	634	1	chin	chin	PROPN
elements-9012	634	2	,	,	PUNCT
elements-9012	634	3	c.	c.	PROPN
elements-9012	634	4	s.	s.	PROPN
elements-9012	634	5	,	,	PUNCT
elements-9012	634	6	chuang	chuang	PROPN
elements-9012	634	7	,	,	PUNCT
elements-9012	634	8	j.	j.	PROPN
elements-9012	634	9	h	h	PROPN
elements-9012	634	10	.	.	PUNCT
elements-9012	634	11	&	&	CCONJ
elements-9012	634	12	li	li	PROPN
elements-9012	634	13	,	,	PUNCT
elements-9012	634	14	h	h	NOUN
elements-9012	634	15	.	.	PUNCT
elements-9012	635	1	genome	genome	NOUN
elements-9012	635	2	-	-	PUNCT
elements-9012	635	3	wide	wide	ADJ
elements-9012	635	4	regulatory	regulatory	ADJ
elements-9012	635	5	complexity	complexity	NOUN
elements-9012	635	6	i	i	PRON
elements-9012	635	7	n	n	VERB
elements-9012	635	8	yeast	yeast	NOUN
elements-9012	635	9	promoters	promoter	NOUN
elements-9012	635	10	:	:	PUNCT
elements-9012	635	11	separation	separation	NOUN
elements-9012	635	12	of	of	ADP
elements-9012	635	13	functionally	functionally	ADV
elements-9012	635	14	conserved	conserved	ADJ
elements-9012	635	15	and	and	CCONJ
elements-9012	635	16	neutral	neutral	ADJ
elements-9012	635	17	sequence	sequence	NOUN
elements-9012	635	18	.	.	PUNCT
elements-9012	636	1	genome	genome	NOUN
elements-9012	636	2	res	re	NOUN
elements-9012	636	3	15	15	NUM
elements-9012	636	4	,	,	PUNCT
elements-9012	636	5	205	205	NUM
elements-9012	636	6	-	-	SYM
elements-9012	636	7	13	13	NUM
elements-9012	636	8	(	(	PUNCT
elements-9012	636	9	2005	2005	NUM
elements-9012	636	10	)	)	PUNCT
elements-9012	636	11	.	.	PUNCT
elements-9012	637	1	chuang	chuang	PROPN
elements-9012	637	2	,	,	PUNCT
elements-9012	637	3	j.	j.	PROPN
elements-9012	637	4	h	h	PROPN
elements-9012	637	5	.	.	PUNCT
elements-9012	637	6	&	&	CCONJ
elements-9012	637	7	li	li	PROPN
elements-9012	637	8	,	,	PUNCT
elements-9012	637	9	h	h	NOUN
elements-9012	637	10	.	.	PUNCT
elements-9012	638	1	functional	functional	ADJ
elements-9012	638	2	bias	bias	NOUN
elements-9012	638	3	and	and	CCONJ
elements-9012	638	4	spatial	spatial	ADJ
elements-9012	638	5	organization	organization	NOUN
elements-9012	638	6	of	of	ADP
elements-9012	638	7	genes	gene	NOUN
elements-9012	638	8	i	i	PRON
elements-9012	638	9	n	n	VERB
elements-9012	638	10	mutational	mutational	ADJ
elements-9012	638	11	hot	hot	ADJ
elements-9012	638	12	and	and	CCONJ
elements-9012	638	13	cold	cold	ADJ
elements-9012	638	14	regions	region	NOUN
elements-9012	638	15	i	i	PRON
elements-9012	638	16	n	n	VERB
elements-9012	638	17	the	the	DET
elements-9012	638	18	human	human	ADJ
elements-9012	638	19	genome	genome	NOUN
elements-9012	638	20	.	.	PUNCT
elements-9012	639	1	plos	plos	PROPN
elements-9012	639	2	biol	biol	PROPN
elements-9012	639	3	2	2	NUM
elements-9012	639	4	,	,	PUNCT
elements-9012	639	5	e29	e29	PROPN
elements-9012	639	6	(	(	PUNCT
elements-9012	639	7	2004	2004	NUM
elements-9012	639	8	)	)	PUNCT
elements-9012	639	9	.	.	PUNCT
elements-9012	640	1	fitzpatrick	fitzpatrick	PROPN
elements-9012	640	2	,	,	PUNCT
elements-9012	640	3	d.	d.	PROPN
elements-9012	640	4	,	,	PUNCT
elements-9012	640	5	logue	logue	PROPN
elements-9012	640	6	,	,	PUNCT
elements-9012	640	7	m	m	PROPN
elements-9012	640	8	.	.	PUNCT
elements-9012	640	9	,	,	PUNCT
elements-9012	640	10	stajich	stajich	PROPN
elements-9012	640	11	,	,	PUNCT
elements-9012	640	12	j.	j.	PROPN
elements-9012	640	13	&	&	CCONJ
elements-9012	640	14	butler	butler	PROPN
elements-9012	640	15	,	,	PUNCT
elements-9012	640	16	g.	g.	PROPN
elements-9012	641	1	a	a	DET
elements-9012	641	2	fungal	fungal	ADJ
elements-9012	641	3	phylogeny	phylogeny	NOUN
elements-9012	641	4	based	base	VERB
elements-9012	641	5	on	on	ADP
elements-9012	641	6	42	42	NUM
elements-9012	641	7	complete	complete	ADJ
elements-9012	641	8	genomes	genome	NOUN
elements-9012	641	9	derived	derive	VERB
elements-9012	641	10	from	from	ADP
elements-9012	641	11	supertree	supertree	ADJ
elements-9012	641	12	and	and	CCONJ
elements-9012	641	13	combined	combined	ADJ
elements-9012	641	14	gene	gene	NOUN
elements-9012	641	15	analysis	analysis	NOUN
elements-9012	641	16	.	.	PUNCT
elements-9012	642	1	bmc	bmc	ADJ
elements-9012	642	2	evolutionary	evolutionary	ADJ
elements-9012	642	3	biology	biology	NOUN
elements-9012	642	4	6	6	NUM
elements-9012	642	5	,	,	PUNCT
elements-9012	642	6	99	99	NUM
elements-9012	642	7	(	(	PUNCT
elements-9012	642	8	2006	2006	NUM
elements-9012	642	9	)	)	PUNCT
elements-9012	642	10	.	.	PUNCT
elements-9012	643	1	hardison	hardison	PROPN
elements-9012	643	2	,	,	PUNCT
elements-9012	643	3	r.	r.	PROPN
elements-9012	643	4	c.	c.	PROPN
elements-9012	643	5	et	et	PROPN
elements-9012	643	6	al	al	PROPN
elements-9012	643	7	.	.	PROPN
elements-9012	643	8	covariation	covariation	PROPN
elements-9012	643	9	in	in	ADP
elements-9012	643	10	frequencies	frequency	NOUN
elements-9012	643	11	of	of	ADP
elements-9012	643	12	substitution	substitution	NOUN
elements-9012	643	13	,	,	PUNCT
elements-9012	643	14	deletion	deletion	NOUN
elements-9012	643	15	,	,	PUNCT
elements-9012	643	16	transposition	transposition	NOUN
elements-9012	643	17	,	,	PUNCT
elements-9012	643	18	and	and	CCONJ
elements-9012	643	19	recombination	recombination	NOUN
elements-9012	643	20	during	during	ADP
elements-9012	643	21	eutherian	eutherian	ADJ
elements-9012	643	22	evolution	evolution	NOUN
elements-9012	643	23	.	.	PUNCT
elements-9012	644	1	genome	genome	NOUN
elements-9012	644	2	res	re	NOUN
elements-9012	644	3	13	13	NUM
elements-9012	644	4	,	,	PUNCT
elements-9012	644	5	13	13	NUM
elements-9012	644	6	-	-	SYM
elements-9012	644	7	26	26	NUM
elements-9012	644	8	(	(	PUNCT
elements-9012	644	9	2003	2003	NUM
elements-9012	644	10	)	)	PUNCT
elements-9012	644	11	.	.	PUNCT
elements-9012	645	1	ikemura	ikemura	NOUN
elements-9012	645	2	,	,	PUNCT
elements-9012	645	3	t.	t.	NOUN
elements-9012	645	4	codon	codon	ADJ
elements-9012	645	5	usage	usage	NOUN
elements-9012	645	6	and	and	CCONJ
elements-9012	645	7	trna	trna	NOUN
elements-9012	645	8	content	content	NOUN
elements-9012	645	9	in	in	ADP
elements-9012	645	10	unicellular	unicellular	ADJ
elements-9012	645	11	and	and	CCONJ
elements-9012	645	12	multicellular	multicellular	ADJ
elements-9012	645	13	organisms	organism	NOUN
elements-9012	645	14	.	.	PUNCT
elements-9012	646	1	mol	mol	ADP
elements-9012	646	2	biol	biol	PROPN
elements-9012	646	3	evol	evol	NOUN
elements-9012	646	4	2,13	2,13	PROPN
elements-9012	646	5	-	-	SYM
elements-9012	646	6	34	34	NUM
elements-9012	646	7	(	(	PUNCT
elements-9012	646	8	1985	1985	NUM
elements-9012	646	9	)	)	PUNCT
elements-9012	646	10	.	.	PUNCT
elements-9012	647	1	jones	jones	PROPN
elements-9012	647	2	,	,	PUNCT
elements-9012	647	3	t.	t.	PROPN
elements-9012	647	4	et	et	PROPN
elements-9012	647	5	al	al	PROPN
elements-9012	647	6	.	.	PUNCT
elements-9012	648	1	the	the	DET
elements-9012	648	2	diploid	diploid	ADJ
elements-9012	648	3	genome	genome	NOUN
elements-9012	648	4	sequence	sequence	NOUN
elements-9012	648	5	of	of	ADP
elements-9012	648	6	candida	candida	NOUN
elements-9012	648	7	albicans	albican	NOUN
elements-9012	648	8	.	.	PUNCT
elements-9012	649	1	proceedings	proceeding	NOUN
elements-9012	649	2	of	of	ADP
elements-9012	649	3	the	the	DET
elements-9012	649	4	national	national	PROPN
elements-9012	649	5	academy	academy	PROPN
elements-9012	649	6	of	of	ADP
elements-9012	649	7	sciences	science	NOUN
elements-9012	649	8	101	101	NUM
elements-9012	649	9	,	,	PUNCT
elements-9012	649	10	7329	7329	NUM
elements-9012	649	11	-	-	SYM
elements-9012	649	12	7334	7334	NUM
elements-9012	649	13	(	(	PUNCT
elements-9012	649	14	2004	2004	NUM
elements-9012	649	15	)	)	PUNCT
elements-9012	649	16	.	.	PUNCT
elements-9012	650	1	lercher	lercher	PROPN
elements-9012	650	2	,	,	PUNCT
elements-9012	650	3	m	m	PROPN
elements-9012	650	4	.	.	PUNCT
elements-9012	651	1	j.	j.	PROPN
elements-9012	651	2	,	,	PUNCT
elements-9012	651	3	williams	williams	PROPN
elements-9012	651	4	,	,	PUNCT
elements-9012	651	5	e.	e.	PROPN
elements-9012	651	6	j.	j.	PROPN
elements-9012	651	7	&	&	CCONJ
elements-9012	651	8	hurst	hurst	PROPN
elements-9012	651	9	,	,	PUNCT
elements-9012	651	10	l.	l.	PROPN
elements-9012	651	11	d.	d.	PROPN
elements-9012	651	12	local	local	PROPN
elements-9012	651	13	similarity	similarity	NOUN
elements-9012	651	14	in	in	ADP
elements-9012	651	15	evolutionary	evolutionary	ADJ
elements-9012	651	16	rates	rate	NOUN
elements-9012	651	17	extends	extend	VERB
elements-9012	651	18	over	over	ADP
elements-9012	651	19	whole	whole	ADJ
elements-9012	651	20	chromosomes	chromosome	NOUN
elements-9012	651	21	in	in	ADP
elements-9012	651	22	humanrodent	humanrodent	NOUN
elements-9012	651	23	and	and	CCONJ
elements-9012	651	24	mouse	mouse	NOUN
elements-9012	651	25	-	-	PUNCT
elements-9012	651	26	rat	rat	NOUN
elements-9012	651	27	comparisons	comparison	NOUN
elements-9012	651	28	:	:	PUNCT
elements-9012	651	29	implications	implication	NOUN
elements-9012	651	30	for	for	ADP
elements-9012	651	31	e	e	NOUN
elements-9012	651	32	l	l	NOUN
elements-9012	651	33	e	e	X
elements-9012	651	34	m	m	PROPN
elements-9012	651	35	e	e	NOUN
elements-9012	651	36	n	n	NOUN
elements-9012	651	37	t	t	PROPN
elements-9012	651	38	s	s	NOUN
elements-9012	651	39	s	s	X
elements-9012	652	1	p	p	NOUN
elements-9012	652	2	r	r	NOUN
elements-9012	653	1	i	i	PRON
elements-9012	654	1	n	n	CCONJ
elements-9012	654	2	g	g	NOUN
elements-9012	654	3	08	08	NUM
elements-9012	654	4	understanding	understand	VERB
elements-9012	654	5	the	the	DET
elements-9012	654	6	mechanistic	mechanistic	ADJ
elements-9012	654	7	basis	basis	NOUN
elements-9012	654	8	of	of	ADP
elements-9012	654	9	the	the	DET
elements-9012	654	10	male	male	ADJ
elements-9012	654	11	mutation	mutation	NOUN
elements-9012	654	12	bias	bias	NOUN
elements-9012	654	13	.	.	PUNCT
elements-9012	655	1	mol	mol	ADP
elements-9012	655	2	biol	biol	PROPN
elements-9012	655	3	evol	evol	NOUN
elements-9012	655	4	18	18	NUM
elements-9012	655	5	,	,	PUNCT
elements-9012	655	6	2032	2032	NUM
elements-9012	655	7	-	-	SYM
elements-9012	655	8	9	9	NUM
elements-9012	655	9	(	(	PUNCT
elements-9012	655	10	2001	2001	NUM
elements-9012	655	11	)	)	PUNCT
elements-9012	655	12	.	.	PUNCT
elements-9012	656	1	liu	liu	PROPN
elements-9012	656	2	,	,	PUNCT
elements-9012	656	3	g.	g.	PROPN
elements-9012	656	4	e.	e.	PROPN
elements-9012	656	5	,	,	PUNCT
elements-9012	656	6	matukumalli	matukumalli	PROPN
elements-9012	656	7	,	,	PUNCT
elements-9012	656	8	l.	l.	PROPN
elements-9012	656	9	k.	k.	PROPN
elements-9012	656	10	,	,	PUNCT
elements-9012	656	11	sonstegard	sonstegard	NOUN
elements-9012	656	12	,	,	PUNCT
elements-9012	656	13	t.	t.	PROPN
elements-9012	656	14	s.	s.	PROPN
elements-9012	656	15	,	,	PUNCT
elements-9012	656	16	shade	shade	PROPN
elements-9012	656	17	,	,	PUNCT
elements-9012	656	18	l.	l.	PROPN
elements-9012	656	19	l.	l.	PROPN
elements-9012	656	20	&	&	CCONJ
elements-9012	656	21	van	van	PROPN
elements-9012	656	22	tassell	tassell	PROPN
elements-9012	656	23	,	,	PUNCT
elements-9012	656	24	c.	c.	PROPN
elements-9012	656	25	p.	p.	NOUN
elements-9012	656	26	genomic	genomic	ADJ
elements-9012	656	27	divergences	divergence	NOUN
elements-9012	656	28	among	among	ADP
elements-9012	656	29	cattle	cattle	NOUN
elements-9012	656	30	,	,	PUNCT
elements-9012	656	31	dog	dog	NOUN
elements-9012	656	32	and	and	CCONJ
elements-9012	656	33	human	human	NOUN
elements-9012	656	34	estimated	estimate	VERB
elements-9012	656	35	from	from	ADP
elements-9012	656	36	large	large	ADJ
elements-9012	656	37	-	-	PUNCT
elements-9012	656	38	scale	scale	NOUN
elements-9012	656	39	alignments	alignment	NOUN
elements-9012	656	40	of	of	ADP
elements-9012	656	41	genomic	genomic	ADJ
elements-9012	656	42	sequences	sequence	NOUN
elements-9012	656	43	.	.	PUNCT
elements-9012	657	1	bmc	bmc	PROPN
elements-9012	657	2	genomics	genomic	NOUN
elements-9012	657	3	7,140	7,140	NUM
elements-9012	657	4	(	(	PUNCT
elements-9012	657	5	2006	2006	NUM
elements-9012	657	6	)	)	PUNCT
elements-9012	657	7	.	.	PUNCT
elements-9012	658	1	logue	logue	PROPN
elements-9012	658	2	,	,	PUNCT
elements-9012	658	3	m	m	PROPN
elements-9012	658	4	.	.	PUNCT
elements-9012	659	1	e.	e.	PROPN
elements-9012	659	2	,	,	PUNCT
elements-9012	659	3	wong	wong	PROPN
elements-9012	659	4	,	,	PUNCT
elements-9012	659	5	s.	s.	PROPN
elements-9012	659	6	,	,	PUNCT
elements-9012	659	7	wolfe	wolfe	PROPN
elements-9012	659	8	,	,	PUNCT
elements-9012	659	9	k.	k.	PROPN
elements-9012	659	10	h	h	PROPN
elements-9012	659	11	.	.	PUNCT
elements-9012	660	1	&	&	CCONJ
elements-9012	660	2	butler	butler	PROPN
elements-9012	660	3	,	,	PUNCT
elements-9012	660	4	g.	g.	PROPN
elements-9012	660	5	a	a	DET
elements-9012	660	6	genome	genome	NOUN
elements-9012	660	7	sequence	sequence	NOUN
elements-9012	660	8	survey	survey	NOUN
elements-9012	660	9	shows	show	VERB
elements-9012	660	10	that	that	SCONJ
elements-9012	660	11	the	the	DET
elements-9012	660	12	pathogenic	pathogenic	ADJ
elements-9012	660	13	yeast	yeast	NOUN
elements-9012	660	14	candida	candida	NOUN
elements-9012	660	15	parapsilosis	parapsilosis	NOUN
elements-9012	660	16	has	have	VERB
elements-9012	660	17	a	a	DET
elements-9012	660	18	defective	defective	ADJ
elements-9012	660	19	mtlai	mtlai	ADJ
elements-9012	660	20	allele	allele	NOUN
elements-9012	660	21	at	at	ADP
elements-9012	660	22	its	its	PRON
elements-9012	660	23	mating	mating	NOUN
elements-9012	660	24	type	type	NOUN
elements-9012	660	25	locus	locus	NOUN
elements-9012	660	26	.	.	PUNCT
elements-9012	661	1	eukaryot	eukaryot	NOUN
elements-9012	661	2	cell	cell	NOUN
elements-9012	661	3	4	4	NUM
elements-9012	661	4	,	,	PUNCT
elements-9012	661	5	1009	1009	NUM
elements-9012	661	6	-	-	SYM
elements-9012	661	7	17	17	NUM
elements-9012	661	8	(	(	PUNCT
elements-9012	661	9	2005	2005	NUM
elements-9012	661	10	)	)	PUNCT
elements-9012	661	11	.	.	PUNCT
elements-9012	662	1	lott	lott	PROPN
elements-9012	662	2	,	,	PUNCT
elements-9012	662	3	t.	t.	PROPN
elements-9012	662	4	j.	j.	PROPN
elements-9012	662	5	,	,	PUNCT
elements-9012	662	6	fundyga	fundyga	PROPN
elements-9012	662	7	,	,	PUNCT
elements-9012	662	8	r.	r.	PROPN
elements-9012	662	9	e.	e.	PROPN
elements-9012	662	10	,	,	PUNCT
elements-9012	662	11	kuykendall	kuykendall	PROPN
elements-9012	662	12	,	,	PUNCT
elements-9012	662	13	r.	r.	PROPN
elements-9012	662	14	j.	j.	PROPN
elements-9012	662	15	&	&	CCONJ
elements-9012	662	16	arnold	arnold	PROPN
elements-9012	662	17	,	,	PUNCT
elements-9012	662	18	j.	j.	PROPN
elements-9012	662	19	the	the	DET
elements-9012	662	20	human	human	ADJ
elements-9012	662	21	commensal	commensal	NOUN
elements-9012	662	22	yeast	yeast	NOUN
elements-9012	662	23	,	,	PUNCT
elements-9012	662	24	candida	candida	NOUN
elements-9012	662	25	albicans	albican	NOUN
elements-9012	662	26	,	,	PUNCT
elements-9012	662	27	has	have	VERB
elements-9012	662	28	an	an	DET
elements-9012	662	29	ancient	ancient	ADJ
elements-9012	662	30	origin	origin	NOUN
elements-9012	662	31	.	.	PUNCT
elements-9012	663	1	fungal	fungal	ADJ
elements-9012	663	2	genet	genet	PROPN
elements-9012	663	3	biol	biol	PROPN
elements-9012	663	4	42	42	NUM
elements-9012	663	5	,	,	PUNCT
elements-9012	663	6	444	444	NUM
elements-9012	663	7	-	-	SYM
elements-9012	663	8	51	51	NUM
elements-9012	663	9	(	(	PUNCT
elements-9012	663	10	2005	2005	NUM
elements-9012	663	11	)	)	PUNCT
elements-9012	663	12	.	.	PUNCT
elements-9012	664	1	lunter	lunter	PROPN
elements-9012	664	2	,	,	PUNCT
elements-9012	664	3	g.	g.	PROPN
elements-9012	664	4	,	,	PUNCT
elements-9012	664	5	ponting	ponting	NOUN
elements-9012	664	6	,	,	PUNCT
elements-9012	664	7	c.	c.	PROPN
elements-9012	664	8	p.	p.	PROPN
elements-9012	664	9	&	&	CCONJ
elements-9012	664	10	hein	hein	PROPN
elements-9012	664	11	,	,	PUNCT
elements-9012	664	12	j.	j.	PROPN
elements-9012	664	13	genome	genome	NOUN
elements-9012	664	14	-	-	PUNCT
elements-9012	664	15	wide	wide	ADJ
elements-9012	664	16	identification	identification	NOUN
elements-9012	664	17	of	of	ADP
elements-9012	664	18	human	human	ADJ
elements-9012	664	19	functional	functional	ADJ
elements-9012	664	20	dna	dna	NOUN
elements-9012	664	21	using	use	VERB
elements-9012	664	22	a	a	DET
elements-9012	664	23	neutral	neutral	ADJ
elements-9012	664	24	indel	indel	NOUN
elements-9012	664	25	model	model	NOUN
elements-9012	664	26	.	.	PUNCT
elements-9012	665	1	plos	plos	PROPN
elements-9012	665	2	comput	comput	PROPN
elements-9012	665	3	biol	biol	PROPN
elements-9012	665	4	2	2	NUM
elements-9012	665	5	,	,	PUNCT
elements-9012	665	6	e5	e5	PROPN
elements-9012	665	7	(	(	PUNCT
elements-9012	665	8	2006	2006	NUM
elements-9012	665	9	)	)	PUNCT
elements-9012	665	10	.	.	PUNCT
elements-9012	666	1	malcom	malcom	PROPN
elements-9012	666	2	,	,	PUNCT
elements-9012	666	3	c.	c.	PROPN
elements-9012	666	4	m	m	PROPN
elements-9012	666	5	.	.	PUNCT
elements-9012	666	6	,	,	PUNCT
elements-9012	666	7	wyckoff	wyckoff	PROPN
elements-9012	666	8	,	,	PUNCT
elements-9012	666	9	g.	g.	PROPN
elements-9012	666	10	j.	j.	PROPN
elements-9012	666	11	&	&	CCONJ
elements-9012	666	12	lahn	lahn	PROPN
elements-9012	666	13	,	,	PUNCT
elements-9012	666	14	b.	b.	PROPN
elements-9012	666	15	t.	t.	PROPN
elements-9012	666	16	genie	genie	PROPN
elements-9012	666	17	mutation	mutation	NOUN
elements-9012	666	18	rates	rate	NOUN
elements-9012	666	19	i	i	PRON
elements-9012	666	20	n	n	VERB
elements-9012	666	21	mammals	mammal	NOUN
elements-9012	666	22	:	:	PUNCT
elements-9012	666	23	local	local	ADJ
elements-9012	666	24	similarity	similarity	NOUN
elements-9012	666	25	,	,	PUNCT
elements-9012	666	26	chromosomal	chromosomal	ADJ
elements-9012	666	27	heterogeneity	heterogeneity	NOUN
elements-9012	666	28	,	,	PUNCT
elements-9012	666	29	and	and	CCONJ
elements-9012	666	30	x	x	X
elements-9012	666	31	-	-	ADJ
elements-9012	666	32	versus	versus	ADP
elements-9012	666	33	-	-	PUNCT
elements-9012	666	34	autosome	autosome	NOUN
elements-9012	666	35	disparity	disparity	NOUN
elements-9012	666	36	.	.	PUNCT
elements-9012	667	1	mol	mol	ADP
elements-9012	667	2	biol	biol	PROPN
elements-9012	667	3	evol	evol	NOUN
elements-9012	667	4	20	20	NUM
elements-9012	667	5	,	,	PUNCT
elements-9012	667	6	1633	1633	NUM
elements-9012	667	7	-	-	SYM
elements-9012	667	8	41	41	NUM
elements-9012	667	9	(	(	PUNCT
elements-9012	667	10	2003	2003	NUM
elements-9012	667	11	)	)	PUNCT
elements-9012	667	12	.	.	PUNCT
elements-9012	668	1	matassi	matassi	PROPN
elements-9012	668	2	,	,	PUNCT
elements-9012	668	3	g.	g.	PROPN
elements-9012	668	4	,	,	PUNCT
elements-9012	668	5	sharp	sharp	ADJ
elements-9012	668	6	,	,	PUNCT
elements-9012	668	7	p.	p.	NOUN
elements-9012	668	8	m	m	PROPN
elements-9012	668	9	.	.	PUNCT
elements-9012	668	10	&	&	CCONJ
elements-9012	668	11	gautier	gautier	PROPN
elements-9012	668	12	,	,	PUNCT
elements-9012	668	13	c.	c.	NOUN
elements-9012	668	14	chromosomal	chromosomal	ADJ
elements-9012	668	15	location	location	NOUN
elements-9012	668	16	effects	effect	NOUN
elements-9012	668	17	on	on	ADP
elements-9012	668	18	gene	gene	NOUN
elements-9012	668	19	sequence	sequence	NOUN
elements-9012	668	20	evolution	evolution	NOUN
elements-9012	668	21	in	in	ADP
elements-9012	668	22	mammals	mammal	NOUN
elements-9012	668	23	.	.	PUNCT
elements-9012	669	1	curr	curr	PROPN
elements-9012	669	2	biol	biol	PROPN
elements-9012	669	3	9	9	NUM
elements-9012	669	4	,	,	PUNCT
elements-9012	669	5	786	786	NUM
elements-9012	669	6	-	-	SYM
elements-9012	669	7	91	91	NUM
elements-9012	669	8	(	(	PUNCT
elements-9012	669	9	1999	1999	NUM
elements-9012	669	10	)	)	PUNCT
elements-9012	669	11	.	.	PUNCT
elements-9012	670	1	mit	mit	PROPN
elements-9012	670	2	,	,	PUNCT
elements-9012	670	3	b.	b.	PROPN
elements-9012	671	1	i	i	PRON
elements-9012	671	2	.	.	PUNCT
elements-9012	672	1	o.	o.	PROPN
elements-9012	672	2	h	h	PROPN
elements-9012	672	3	.	.	PUNCT
elements-9012	673	1	a.	a.	NOUN
elements-9012	673	2	(	(	PUNCT
elements-9012	673	3	2007	2007	NUM
elements-9012	673	4	)	)	PUNCT
elements-9012	673	5	.	.	PUNCT
elements-9012	674	1	murphy	murphy	PROPN
elements-9012	674	2	,	,	PUNCT
elements-9012	674	3	w.	w.	PROPN
elements-9012	674	4	j.	j.	PROPN
elements-9012	674	5	,	,	PUNCT
elements-9012	674	6	pringle	pringle	PROPN
elements-9012	674	7	,	,	PUNCT
elements-9012	674	8	t.	t.	NOUN
elements-9012	674	9	h	h	PROPN
elements-9012	674	10	.	.	PUNCT
elements-9012	674	11	,	,	PUNCT
elements-9012	674	12	crider	crider	NOUN
elements-9012	674	13	,	,	PUNCT
elements-9012	674	14	t.	t.	PROPN
elements-9012	674	15	a.	a.	PROPN
elements-9012	674	16	,	,	PUNCT
elements-9012	674	17	springer	springer	NOUN
elements-9012	674	18	,	,	PUNCT
elements-9012	674	19	m	m	PROPN
elements-9012	674	20	.	.	PUNCT
elements-9012	675	1	s.	s.	PROPN
elements-9012	675	2	&	&	CCONJ
elements-9012	675	3	miller	miller	PROPN
elements-9012	675	4	,	,	PUNCT
elements-9012	675	5	w	w	ADP
elements-9012	675	6	using	use	VERB
elements-9012	675	7	genomic	genomic	ADJ
elements-9012	675	8	data	datum	NOUN
elements-9012	675	9	to	to	PART
elements-9012	675	10	unravel	unravel	VERB
elements-9012	675	11	the	the	DET
elements-9012	675	12	root	root	NOUN
elements-9012	675	13	of	of	ADP
elements-9012	675	14	the	the	DET
elements-9012	675	15	placental	placental	ADJ
elements-9012	675	16	mammal	mammal	ADJ
elements-9012	675	17	phylogeny	phylogeny	NOUN
elements-9012	675	18	.	.	PUNCT
elements-9012	676	1	genome	genome	NOUN
elements-9012	676	2	res	re	NOUN
elements-9012	676	3	17	17	NUM
elements-9012	676	4	,	,	PUNCT
elements-9012	676	5	413	413	NUM
elements-9012	676	6	-	-	SYM
elements-9012	676	7	21	21	NUM
elements-9012	676	8	(	(	PUNCT
elements-9012	676	9	2007	2007	NUM
elements-9012	676	10	)	)	PUNCT
elements-9012	676	11	.	.	PUNCT
elements-9012	677	1	nikolaev	nikolaev	INTJ
elements-9012	677	2	,	,	PUNCT
elements-9012	677	3	s.	s.	PROPN
elements-9012	677	4	et	et	PROPN
elements-9012	677	5	al	al	PROPN
elements-9012	677	6	.	.	PROPN
elements-9012	678	1	early	early	ADJ
elements-9012	678	2	history	history	NOUN
elements-9012	678	3	of	of	ADP
elements-9012	678	4	mammals	mammal	NOUN
elements-9012	678	5	is	be	AUX
elements-9012	678	6	elucidated	elucidate	VERB
elements-9012	678	7	with	with	ADP
elements-9012	678	8	the	the	DET
elements-9012	678	9	encode	encode	ADJ
elements-9012	678	10	multiple	multiple	ADJ
elements-9012	678	11	species	specie	NOUN
elements-9012	678	12	sequencing	sequence	VERB
elements-9012	678	13	data	datum	NOUN
elements-9012	678	14	.	.	PUNCT
elements-9012	679	1	plos	plos	PROPN
elements-9012	679	2	genet	genet	PROPN
elements-9012	679	3	3	3	NUM
elements-9012	679	4	,	,	PUNCT
elements-9012	679	5	e2	e2	PROPN
elements-9012	679	6	(	(	PUNCT
elements-9012	679	7	2007	2007	NUM
elements-9012	679	8	)	)	PUNCT
elements-9012	679	9	.	.	PUNCT
elements-9012	680	1	pheasant	pheasant	NOUN
elements-9012	680	2	,	,	PUNCT
elements-9012	680	3	m	m	PROPN
elements-9012	680	4	.	.	PUNCT
elements-9012	680	5	&	&	CCONJ
elements-9012	680	6	mattick	mattick	PROPN
elements-9012	680	7	,	,	PUNCT
elements-9012	681	1	j.	j.	PROPN
elements-9012	681	2	s.	s.	PROPN
elements-9012	681	3	raising	raise	VERB
elements-9012	681	4	the	the	DET
elements-9012	681	5	estimate	estimate	NOUN
elements-9012	681	6	of	of	ADP
elements-9012	681	7	functional	functional	ADJ
elements-9012	681	8	human	human	ADJ
elements-9012	681	9	sequences	sequence	NOUN
elements-9012	681	10	.	.	PUNCT
elements-9012	682	1	genome	genome	NOUN
elements-9012	682	2	res	re	NOUN
elements-9012	682	3	.	.	PUNCT
elements-9012	683	1	17,1245	17,1245	NUM
elements-9012	683	2	-	-	SYM
elements-9012	683	3	1253	1253	NUM
elements-9012	683	4	(	(	PUNCT
elements-9012	683	5	2007	2007	NUM
elements-9012	683	6	)	)	PUNCT
elements-9012	683	7	.	.	PUNCT
elements-9012	684	1	pujol	pujol	NOUN
elements-9012	684	2	,	,	PUNCT
elements-9012	684	3	c.	c.	PROPN
elements-9012	684	4	et	et	PROPN
elements-9012	684	5	al	al	PROPN
elements-9012	684	6	.	.	PUNCT
elements-9012	685	1	the	the	DET
elements-9012	685	2	closely	closely	ADV
elements-9012	685	3	related	relate	VERB
elements-9012	685	4	species	specie	NOUN
elements-9012	685	5	candida	candida	NOUN
elements-9012	685	6	albicans	albican	NOUN
elements-9012	685	7	and	and	CCONJ
elements-9012	685	8	candida	candida	NOUN
elements-9012	685	9	dubliniensis	dubliniensis	NOUN
elements-9012	685	10	can	can	AUX
elements-9012	685	11	mate	mate	VERB
elements-9012	685	12	.	.	PUNCT
elements-9012	686	1	eukaryot	eukaryot	PROPN
elements-9012	686	2	cell	cell	NOUN
elements-9012	686	3	3	3	NUM
elements-9012	686	4	,	,	PUNCT
elements-9012	686	5	1015	1015	NUM
elements-9012	686	6	-	-	SYM
elements-9012	686	7	27	27	NUM
elements-9012	686	8	(	(	PUNCT
elements-9012	686	9	2004	2004	NUM
elements-9012	686	10	)	)	PUNCT
elements-9012	686	11	.	.	PUNCT
elements-9012	687	1	ruderfer	ruderfer	PROPN
elements-9012	687	2	,	,	PUNCT
elements-9012	687	3	d.	d.	PROPN
elements-9012	687	4	m	m	PROPN
elements-9012	687	5	.	.	PUNCT
elements-9012	687	6	,	,	PUNCT
elements-9012	687	7	pratt	pratt	PROPN
elements-9012	687	8	,	,	PUNCT
elements-9012	687	9	s.	s.	PROPN
elements-9012	687	10	c	c	PROPN
elements-9012	687	11	,	,	PUNCT
elements-9012	687	12	seidel	seidel	PROPN
elements-9012	687	13	,	,	PUNCT
elements-9012	687	14	h	h	NOUN
elements-9012	687	15	.	.	PUNCT
elements-9012	688	1	s.	s.	PROPN
elements-9012	688	2	&	&	CCONJ
elements-9012	688	3	kruglyak	kruglyak	PROPN
elements-9012	688	4	,	,	PUNCT
elements-9012	688	5	l.	l.	PROPN
elements-9012	688	6	population	population	PROPN
elements-9012	688	7	genomic	genomic	NOUN
elements-9012	688	8	analysis	analysis	NOUN
elements-9012	688	9	of	of	ADP
elements-9012	688	10	outcrossing	outcrossing	NOUN
elements-9012	688	11	and	and	CCONJ
elements-9012	688	12	recombination	recombination	NOUN
elements-9012	688	13	i	i	PRON
elements-9012	688	14	n	n	PRON
elements-9012	688	15	yeast	yeast	NOUN
elements-9012	688	16	.	.	PUNCT
elements-9012	689	1	nat	nat	PROPN
elements-9012	689	2	genet	genet	PROPN
elements-9012	689	3	38	38	NUM
elements-9012	689	4	,	,	PUNCT
elements-9012	689	5	1077	1077	NUM
elements-9012	689	6	-	-	SYM
elements-9012	689	7	81	81	NUM
elements-9012	689	8	(	(	PUNCT
elements-9012	689	9	2006	2006	NUM
elements-9012	689	10	)	)	PUNCT
elements-9012	689	11	.	.	PUNCT
elements-9012	690	1	selker	selker	PROPN
elements-9012	690	2	,	,	PUNCT
elements-9012	690	3	e.	e.	PROPN
elements-9012	690	4	u	u	PROPN
elements-9012	690	5	.	.	PUNCT
elements-9012	691	1	premeiotic	premeiotic	ADJ
elements-9012	691	2	instability	instability	NOUN
elements-9012	691	3	of	of	ADP
elements-9012	691	4	repeated	repeat	VERB
elements-9012	691	5	sequences	sequence	NOUN
elements-9012	691	6	in	in	ADP
elements-9012	691	7	neurospora	neurospora	NOUN
elements-9012	691	8	crassa	crassa	NOUN
elements-9012	691	9	.	.	PUNCT
elements-9012	692	1	a	a	DET
elements-9012	692	2	n	n	CCONJ
elements-9012	692	3	n	n	PRON
elements-9012	692	4	u	u	NOUN
elements-9012	692	5	rev	rev	VERB
elements-9012	692	6	genet	genet	PROPN
elements-9012	692	7	24	24	NUM
elements-9012	692	8	,	,	PUNCT
elements-9012	692	9	579	579	NUM
elements-9012	692	10	-	-	SYM
elements-9012	692	11	613	613	NUM
elements-9012	692	12	(	(	PUNCT
elements-9012	692	13	1990	1990	NUM
elements-9012	692	14	)	)	PUNCT
elements-9012	692	15	.	.	PUNCT
elements-9012	693	1	tsong	tsong	PROPN
elements-9012	693	2	,	,	PUNCT
elements-9012	693	3	a.	a.	PROPN
elements-9012	693	4	e.	e.	PROPN
elements-9012	693	5	,	,	PUNCT
elements-9012	693	6	tuch	tuch	PROPN
elements-9012	693	7	,	,	PUNCT
elements-9012	693	8	b.	b.	PROPN
elements-9012	693	9	b.	b.	PROPN
elements-9012	693	10	,	,	PUNCT
elements-9012	693	11	li	li	PROPN
elements-9012	693	12	,	,	PUNCT
elements-9012	693	13	h	h	PROPN
elements-9012	693	14	.	.	PUNCT
elements-9012	693	15	&	&	CCONJ
elements-9012	693	16	johnson	johnson	PROPN
elements-9012	693	17	,	,	PUNCT
elements-9012	693	18	a.	a.	PROPN
elements-9012	693	19	d.	d.	PROPN
elements-9012	693	20	evolution	evolution	PROPN
elements-9012	693	21	of	of	ADP
elements-9012	693	22	alternative	alternative	ADJ
elements-9012	693	23	transcriptional	transcriptional	ADJ
elements-9012	693	24	circuits	circuit	NOUN
elements-9012	693	25	wi	wi	PROPN
elements-9012	693	26	th	th	X
elements-9012	693	27	identical	identical	ADJ
elements-9012	693	28	logic	logic	NOUN
elements-9012	693	29	.	.	PUNCT
elements-9012	694	1	nature	nature	NOUN
elements-9012	694	2	443	443	NUM
elements-9012	694	3	,	,	PUNCT
elements-9012	694	4	415	415	NUM
elements-9012	694	5	(	(	PUNCT
elements-9012	694	6	2006	2006	NUM
elements-9012	694	7	)	)	PUNCT
elements-9012	694	8	.	.	PUNCT
elements-9012	695	1	vyas	vyas	PROPN
elements-9012	695	2	,	,	PUNCT
elements-9012	695	3	m	m	PROPN
elements-9012	695	4	.	.	PUNCT
elements-9012	695	5	&	&	CCONJ
elements-9012	695	6	kasbekar	kasbekar	PROPN
elements-9012	695	7	,	,	PUNCT
elements-9012	695	8	d.	d.	PROPN
elements-9012	695	9	p.	p.	PROPN
elements-9012	695	10	collateral	collateral	ADJ
elements-9012	695	11	damage	damage	NOUN
elements-9012	695	12	:	:	PUNCT
elements-9012	695	13	spread	spread	NOUN
elements-9012	695	14	of	of	ADP
elements-9012	695	15	repeatinduced	repeatinduce	VERB
elements-9012	695	16	point	point	NOUN
elements-9012	695	17	mutation	mutation	NOUN
elements-9012	695	18	from	from	ADP
elements-9012	695	19	a	a	DET
elements-9012	695	20	duplicated	duplicate	VERB
elements-9012	695	21	dna	dna	NOUN
elements-9012	695	22	sequence	sequence	NOUN
elements-9012	695	23	into	into	ADP
elements-9012	695	24	an	an	DET
elements-9012	695	25	adjoining	adjoining	ADJ
elements-9012	695	26	single	single	ADJ
elements-9012	695	27	-	-	PUNCT
elements-9012	695	28	copy	copy	NOUN
elements-9012	695	29	gene	gene	NOUN
elements-9012	695	30	i	i	PRON
elements-9012	695	31	n	n	VERB
elements-9012	695	32	neurospora	neurospora	VERB
elements-9012	695	33	crassa	crassa	NOUN
elements-9012	695	34	.	.	PUNCT
elements-9012	696	1	j	j	PROPN
elements-9012	696	2	biosci	biosci	PROPN
elements-9012	696	3	30	30	NUM
elements-9012	696	4	,	,	PUNCT
elements-9012	696	5	15	15	NUM
elements-9012	696	6	-	-	SYM
elements-9012	696	7	20	20	NUM
elements-9012	696	8	(	(	PUNCT
elements-9012	696	9	2005	2005	NUM
elements-9012	696	10	)	)	PUNCT
elements-9012	696	11	.	.	PUNCT
elements-9012	697	1	wolfe	wolfe	PROPN
elements-9012	697	2	,	,	PUNCT
elements-9012	697	3	k.	k.	PROPN
elements-9012	697	4	h	h	PROPN
elements-9012	697	5	.	.	PUNCT
elements-9012	698	1	,	,	PUNCT
elements-9012	698	2	sharp	sharp	ADJ
elements-9012	698	3	,	,	PUNCT
elements-9012	698	4	p.	p.	NOUN
elements-9012	698	5	m	m	PROPN
elements-9012	698	6	.	.	PUNCT
elements-9012	699	1	&	&	CCONJ
elements-9012	699	2	li	li	PROPN
elements-9012	699	3	,	,	PUNCT
elements-9012	699	4	w.	w.	PROPN
elements-9012	699	5	h	h	PROPN
elements-9012	699	6	.	.	PUNCT
elements-9012	700	1	mutation	mutation	NOUN
elements-9012	700	2	rates	rate	NOUN
elements-9012	700	3	differ	differ	VERB
elements-9012	700	4	among	among	ADP
elements-9012	700	5	regions	region	NOUN
elements-9012	700	6	of	of	ADP
elements-9012	700	7	the	the	DET
elements-9012	700	8	mammalian	mammalian	ADJ
elements-9012	700	9	genome	genome	NOUN
elements-9012	700	10	.	.	PUNCT
elements-9012	701	1	nature	nature	NOUN
elements-9012	701	2	337	337	NUM
elements-9012	701	3	,	,	PUNCT
elements-9012	701	4	283	283	NUM
elements-9012	701	5	-	-	SYM
elements-9012	701	6	5	5	NUM
elements-9012	701	7	(	(	PUNCT
elements-9012	701	8	1989	1989	NUM
elements-9012	701	9	)	)	PUNCT
elements-9012	701	10	.	.	PUNCT
elements-9012	702	1	wong	wong	PROPN
elements-9012	702	2	,	,	PUNCT
elements-9012	702	3	s.	s.	PROPN
elements-9012	702	4	,	,	PUNCT
elements-9012	702	5	fares	fare	NOUN
elements-9012	702	6	,	,	PUNCT
elements-9012	702	7	m	m	PROPN
elements-9012	702	8	.	.	PUNCT
elements-9012	703	1	a.	a.	PROPN
elements-9012	703	2	,	,	PUNCT
elements-9012	703	3	zimmerman	zimmerman	PROPN
elements-9012	703	4	,	,	PUNCT
elements-9012	703	5	w.	w.	PROPN
elements-9012	703	6	,	,	PUNCT
elements-9012	703	7	butler	butler	PROPN
elements-9012	703	8	,	,	PUNCT
elements-9012	703	9	g.	g.	PROPN
elements-9012	703	10	&	&	CCONJ
elements-9012	703	11	wolfe	wolfe	PROPN
elements-9012	703	12	,	,	PUNCT
elements-9012	703	13	k	k	PROPN
elements-9012	703	14	.h	.h	PROPN
elements-9012	703	15	.	.	PUNCT
elements-9012	704	1	evidence	evidence	NOUN
elements-9012	704	2	from	from	ADP
elements-9012	704	3	comparative	comparative	ADJ
elements-9012	704	4	genomics	genomic	NOUN
elements-9012	704	5	for	for	ADP
elements-9012	704	6	a	a	DET
elements-9012	704	7	complete	complete	ADJ
elements-9012	704	8	sexual	sexual	ADJ
elements-9012	704	9	cycle	cycle	NOUN
elements-9012	704	10	in	in	ADP
elements-9012	704	11	the	the	DET
elements-9012	704	12	'	'	PUNCT
elements-9012	704	13	asexual	asexual	ADJ
elements-9012	704	14	'	'	PUNCT
elements-9012	704	15	pathogenic	pathogenic	ADJ
elements-9012	704	16	yeast	yeast	NOUN
elements-9012	704	17	candida	candida	NOUN
elements-9012	704	18	glabrata	glabrata	NOUN
elements-9012	704	19	.	.	PUNCT
elements-9012	705	1	genome	genome	NOUN
elements-9012	705	2	biol	biol	NOUN
elements-9012	705	3	4	4	NUM
elements-9012	705	4	,	,	PUNCT
elements-9012	705	5	rio	rio	NOUN
elements-9012	705	6	(	(	PUNCT
elements-9012	705	7	2003	2003	NUM
elements-9012	705	8	)	)	PUNCT
elements-9012	705	9	.	.	PUNCT
elements-9012	706	1	yokogawa	yokogawa	PROPN
elements-9012	706	2	,	,	PUNCT
elements-9012	706	3	t.	t.	PROPN
elements-9012	706	4	et	et	PROPN
elements-9012	706	5	al	al	PROPN
elements-9012	706	6	.	.	PROPN
elements-9012	706	7	serine	serine	PROPN
elements-9012	706	8	trna	trna	PROPN
elements-9012	706	9	complementary	complementary	ADJ
elements-9012	706	10	to	to	ADP
elements-9012	706	11	the	the	DET
elements-9012	706	12	nonuniversal	nonuniversal	ADJ
elements-9012	706	13	serine	serine	NOUN
elements-9012	706	14	codon	codon	NOUN
elements-9012	706	15	cug	cug	NOUN
elements-9012	706	16	i	i	PRON
elements-9012	706	17	n	n	VERB
elements-9012	706	18	candida	candida	PROPN
elements-9012	706	19	cylindracea	cylindracea	NOUN
elements-9012	706	20	:	:	PUNCT
elements-9012	706	21	evolutionary	evolutionary	ADJ
elements-9012	706	22	implications	implication	NOUN
elements-9012	706	23	.	.	PUNCT
elements-9012	707	1	proc	proc	PROPN
elements-9012	707	2	natl	natl	PROPN
elements-9012	707	3	acad	acad	PROPN
elements-9012	707	4	sci	sci	PROPN
elements-9012	707	5	u	u	PROPN
elements-9012	707	6	s	s	PROPN
elements-9012	707	7	a	a	DET
elements-9012	707	8	89	89	NUM
elements-9012	707	9	,	,	PUNCT
elements-9012	707	10	7408	7408	NUM
elements-9012	707	11	-	-	SYM
elements-9012	707	12	11	11	NUM
elements-9012	707	13	(	(	PUNCT
elements-9012	707	14	1992	1992	NUM
elements-9012	707	15	)	)	PUNCT
elements-9012	707	16	.	.	PUNCT
elements-9012	708	1	m	m	PROPN
elements-9012	708	2	e	e	X
elements-9012	708	3	a	a	PRON
elements-9012	708	4	s	s	X
elements-9012	708	5	u	u	NOUN
elements-9012	708	6	r	r	NOUN
elements-9012	708	7	i	i	PRON
elements-9012	708	8	n	n	CCONJ
elements-9012	708	9	g	g	PROPN
elements-9012	708	10	t	t	NOUN
elements-9012	708	11	h	h	NOUN
elements-9012	708	12	e	e	X
elements-9012	708	13	u	u	NOUN
elements-9012	708	14	n	n	PROPN
elements-9012	708	15	i	i	PRON
elements-9012	708	16	v	v	VERB
elements-9012	708	17	e	e	NOUN
elements-9012	708	18	r	r	NOUN
elements-9012	708	19	s	s	PROPN
elements-9012	708	20	a	a	DET
elements-9012	708	21	l	l	NOUN
elements-9012	709	1	i	i	PRON
elements-9012	709	2	t	t	VERB
elements-9012	710	1	y	y	NOUN
elements-9012	711	1	o	o	X
elements-9012	711	2	f	f	NOUN
elements-9012	712	1	r	r	NOUN
elements-9012	712	2	e	e	NOUN
elements-9012	712	3	g	g	NOUN
elements-9012	713	1	i	i	PRON
elements-9012	713	2	o	o	VERB
elements-9012	714	1	n	n	CCONJ
elements-9012	714	2	a	a	DET
elements-9012	714	3	l	l	NOUN
elements-9012	714	4	m	m	VERB
elements-9012	715	1	u	u	NOUN
elements-9012	715	2	t	t	PROPN
elements-9012	715	3	a	a	DET
elements-9012	715	4	t	t	NOUN
elements-9012	716	1	i	i	PRON
elements-9012	716	2	o	o	VERB
elements-9012	717	1	n	n	CCONJ
elements-9012	717	2	r	r	NOUN
elements-9012	717	3	a	a	DET
elements-9012	717	4	t	t	NOUN
elements-9012	717	5	e	e	NOUN
elements-9012	717	6	s	s	NOUN