www.geologia-croatica.hr Journal of the Croatian Geological Survey and the Croatian Geological Society 1. INTRODUCTION During the Late Jurassic, the Dasycladales (green algae) flourish­­ed­during­a­period­that­followed­crisis­moments­at­the­ end­of­the­Late­Triassic­and­the­final­part­of­the­Early­Jurassic,­ as­ well­ as­ a­ very­ slow­ recovery­ in­ the­ Middle­ Jurassic­ (GRANIER & DELOFFRE, 1993; BUCUR, 1999; BUCUR & REOLID,­2024).­In­the­Oxfordian–Tithonian­interval,­86­ dasycladalean­species­are­known,­72­of­which­are­mentioned­ in­ the­Tithonian­ (BUCUR­&­REOLID,­2024).­Thirty­ four­ species­continued­their­evolution­into­the­Early­Cretaceous.­ Regarding­the­Upper­Jurassic­dasycladaleans,­Aloisalthella sulcata (ALTH)­[ex­Clypeina jurassica FAVRE,­ex­Clypeina sulcata (ALTH), see GRANIER­&­LETHIERS, 2019] and Salpingoporella pygmaea (GÜMBEL)­are­among­the­most­fre­ quently­encountered­and,­as­such,­the­best­known.­A­special­ place­is­occupied­by­Neoteutloporella socialis (PRATURLON), a­dasycladalean­alga­that­develops­in­the­form­of­"mini­reefs"/ bioherms.­These­are­actually­agglomerations­of­thalli­in­the­ form­of­overlapping­"bushes"­that­can­reach­from­a­few­centi­ metres­to­a­few­decimetres­in­height­and­breadth. The­aim­of­the­present­work­is­to­describe­a­new­dasycla­ dalean alga, Schlagintweitella inopinata nov.­gen.,­nov.­sp.,­ which,­like­Neoteutloporella,­develops­in­the­form­of­thallus­ agglomerations.­Although,­agglomeration­is­not­uncommon­ among­extant­Dasycladales,­as­evidenced­by­numerous­recent­ illustrations­of­loose­agglomerations­(e.g.,­TROPICAL­FISH­ PLUS­(2024),­for­Neomeris),­none­develops­Neoteutloporella­ type­“mini­reefs”. 2. GEOLOGICAL BACKGROUND The­Mesozoic­deposits­in­the­Cioclovina­Băniţa­area­(also­ known­as­the­Haţeg­Pui­area;­Fig.­1)­constitute­a­fragment­of­ the­sedimentary­cover­of­the­Getic­nappe­(STILLĂ,­1985;­ PLEŞ­et­al.,­2019).­The­sedimentary­succession­of­this­area­ includes­ deposits­ from­ the­ Lower­ Jurassic­ to­ the­ Upper­ Schlagintweitella inopinata, a new genus and species of Dasycladales (green algae) from the Upper Jurassic limestones of Romania Ioan I. Bucur1,2,*, Emanoil Săsăran1, George Pleş1 1 Babeş-Bolyai University, Department of Geology and Center for Integrated Geological Studies, M. Kogălniceanu 1, 400084 Cluj-Napoca, Romania; (*corresponding author: ioan.bucur@ubbcluj.ro; emanoil.sasaran@ubbcluj.ro; george.ples@ubbcluj.ro) 2 Romanian Academy, Cluj Branch, Republicii 9, 400015 Cluj-Napoca, Romania doi: 10.4154/gc.2025.10 Abstract An outcrop identified within the Upper Jurassic deposits of the Cioclovina-Băniţa area (Southern Carpathians) most likely corresponds to a cavern fill. In several lithoclasts of this fill, including black-pebbles, a dasycladalean alga with morphological features that differ- entiate it from all other Jurassic–Cretaceous dasycladales with two orders of laterals was observed. It is described in the present work as a new species of a new genus: Schlagin­ tweitella inopinata nov. gen., nov. sp. Cretaceous­ (pre­Maastrichtian).­ The­ terrigenous­ Lower­ Jurassic­ is­ developed­ in­ the­ Gresten­ facies.­ The­Middle­ Jurassic­is­mainly­represented­by­siliciclastic­rocks­followed­ by­a­carbonate­sequence­in­the­upper­part.­The­Upper­Jurassic­ and­the­lowermost­Cretaceous­are­represented­by­carbonate­ rocks­(Figs.­1,­2).­Although­initially­included­within­an­Upper­ Jurassic–Aptian­carbonate­succession­(STILLĂ,­1985),­the­ Barremian–Aptian­is­present­only­as­clasts­in­the­conglo­me­ ratic­breccia­deposits­in­the­lower­part­of­the­Cenomanian­ (PLEȘ­ et­ al.,­ 2019).­During­ the­Albian,­ lateritic­ deposits­ formed,­leading­to­the­accumulation­of­bauxites.­The­Upper­ Cretaceous­ includes­a­marine­succession­ that­begins­with­ Ceno­manian­breccias­and­conglomerates,­and­ends­with­Cam­ panian­marls.­These­are­covered­by­the­Maastrichtian­conti­ nental­deposits­of­the­Haţeg­Basin. The­limestones­containing­the­new­dasycladalean­alga­ belong­to­the­Upper­Jurassic.­The­outcrop­is­located­in­the­ Lunca­Ohabei­plateau,­near­Albian­bauxite­lenses­(Fig.­3).­Ac­ cording­to­SĂSĂRAN­et­al.­(2021),­the­outcrop­represents­a­ cavern­fill.­The­clasts­in­the­cavern­fill­originated­from­different­ depositional­environments­and­were­later­transported­into­the­ karst­void.­They­consist­of­bioclasts,­intraclasts,­calcrete­clasts,­ fragments­of­altered­carbonate­ rocks­pigmented­with­ iron­ oxides­hydroxides.­Some­of­ these­ represent­black­pebbles­ formed­in­subaerially­exposed­areas­through­impregnation­ with­organic­matter. One­of­ these­clasts­(Fig.­4)­ is­mainly­composed­of­an­ agglo­meration­of­algal­thalli.­Their­study­revealed­the­dis­co­ very­of­a­new­genus­and­a­new­species­of­dasycladalean­green­ algae. 3. MATERIAL AND REPOSITORY The­new­alga­was­found­in­two­samples,­from­which­four­thin­ sections­were­made.­Additionally,­three­other­samples­were­ collected­from­the­type­locality,­resulting­in­thirteen­thin­ 2025 | 78/2 | 103–113 | 4 Figs. | 6 Plts. | Article history: Manuscript received: December 4, 2024 Revised manuscript accepted: March 12, 2025 Available online: June 3, 2025 Keywords: Dasycladales, Upper Jurassic, new genus, new species, Romania G eo lo gi a C ro at ic a 104 Geologia Croatica 78/2 sections­used­for­studying­the­accompanying­micro­palaeonto­ logical­association.­All­thin­sections­are­stored­in­the­collection­ of­the­Paleontology­Museum­of­Babeş­Bolyai­University,­Cluj­ Napoca. 4. PALAEONTOLOGICAL DESCRIPTION Phylum­Chlorophyta­PASCHER,­1914 Class­Ulvophyceae­MATTOX­&­STEWART,­1984 Order­Dasycladales­PASCHER,­1931 Family­Triploporellaceae­(PIA,­1920)­BERGER­&­KAEVER,­ 1992 Tribe­Triploporelleae­(PIA,­1920)­BASSOULLET­et­al.,­1979 Genus Schlagintweitella nov. gen. Type­species:­Schlagintweitella inopinata­nov.­gen.,­nov.­sp. 1997­Dasycladacean­with­unsegmented­primary­ and­ seg­ ment­ed­secondary­laterals­–­DE­CASTRO,­p.­200,­pl.­4,­figs.­ 1­3. Derivation­of­name:­dedicated­to­Dr.­Felix­SCHLAGINTWEIT (München)­for­his­significant­contributions­to­the­study­of­ Mesozoic­and­Cenozoic­calcareous­algae. Generic diagnosis:­Dasycladalean­ algae­with­ cylindrical­ thallus­and­two­orders­of­laterals.­The­primary­laterals­are­ cylindrical­or­ovoid­elon­gated,­shorter­than­the­secondaries.­ The­secondary­laterals­are­also­cylindrical­and­consist­of­two,­ rarely­three­successive­segments­separated­by­constrictions.­ The­proximal­part­of­the­secondary­laterals­is­covered­by­a­ common­calcareous­sheath­(collective­sheath). Remarks and comparisons: In­the­Jurassic­and­Cretaceous­ fossil­record,­29­genera­of­Dasy­cladales­with­two­orders­of­ laterals­are­known­(e.g.,­BUCUR­et­al.,­2010;­Table­1;­to­the­ genera­in­this­table­we­must­add­the­genera­Steinmanniporella, Bakalovaella, Eodasy cladus, Chinianella, Granieria, Diste­ fano polia, and Cylindro porella? liasica).­ The­ differences­ between­all­these­taxa­are­given­by­the­shape­and­dimensions­ of­the­laterals,­the­diff­eren­ces­in­shape­between­the­first­and­ second­order­laterals,­as­well­as­the­dimen­sional­ratio­between­ the­two­types­of­laterals.­In­addition,­the­type­of­reproduction­ is­considered.­The­genus­Linoporella,­with­the­type­species­ Linoporella capriotica (OPPENHEIM),­presents­three­orders­ of­ laterals­ and­ thus­ differentiates­ itself­ from­ this­ group­ (BARATTOLO­&­ROMANO,­2005).­We­can­also­add­to­this­ list the genus Palaeo dasycladus,­ which­ has­ segmented­ secondary­ laterals,­but­ it­presents­ three­or­more­orders­of­ laterals,­and­thus­differs­from­the­new­genus.­Three­of­the­ genera­ mentioned­ in­ Table­ 1­ by­ BUCUR­ et­ al.­ (2010)­ (Montenegrella, Crinella, and Helio porella)­are­most­likely­ recent­synonyms­of­the­genera­Suppilu liu maella­(the­first­two)­ and Neomeris,­respectively.­Of­the­remaining­genera,­those­in­ the­tribe­Neomeree­(Genotella, Montiella/Bakalovaella, Neo­ meris, and Cymopolia)­ are­ distin­guished­ by­ the­ type­ of­ choristospore­reproduction­and/or­by­the­shape­of­the­thallus­ (e.g., Cymopolia). Conradella, Chinia nella, Eodasycladus, Gra nieria, Distefanopolia, and Cylindro porella? liasica belong­to­the­same­group­(cf.­BARATTOLO­&­PARENTE,­ 2000;­BARATTOLO­et­al.,­2008,­2012,­2021a).­Among­the­ Dasy­cladales­ in­ the­ Triploporelleae­ tribe,­ Acroporella, Rajkaella, and Triploporella­all­have­much­longer­first­order­ laterals­com­pared­to­second­order­laterals.­Dis so cladella (tribe­ Dissocla­delleae)­includes­species­with­primary­and­secondary­ laterals­that­are­comparable­in­dimension­but­differ­in­shape.­ Selliporella­has­short,­globose­first­order­laterals­and­long,­ trichoform­second­order­laterals­(see­BARATTOLO­et­al.,­ 2021b).­Tersella and Pseudocymopolia­are­distinguished­by­ the­different­general­morphology­of­the­thallus­(club­shaped,­ very­clearly­differen­tiated­into­head­and­stipe,­and­moniliform,­ respectively). The­new­genus­Schlagintweitella­shows­the­greatest­simi­ larity­to­the­genus­Neoteutloporella (BASSOULLET­et­al.,­1978)­ Figure 1. A geological map of the Cioclovina-Băniţa area with the location of the studied section (red star). Modified after Pleş et al., 2019. G eologia C roatica 105Bucur et al.: Schlagintweitella inopinata, a new genus and species of Dasycladales (green algae) from the Upper Jurassic limestones of Romania DE­CASTRO,­1993.­The­genus­Neoteutloporella­includes­two­ species,­both­from­the­Upper­Jurassic:­N. obsoleta­(CAROZZI,­ 1954)­and­N. socialis­(PRATURLON,­1963).­Both­species­have­ relatively­short,­globose­or­ovoid­first­order­laterals­and­rela­ tively­long,­acrophorous­or­slightly­phloiophorous­second­order­ laterals­ that­ are­ segmented­ by­ several­ constrictions.­Both­ species­ occur­ as­ dense­ agglomerations­ of­ thalli,­ often­ in­ overlapping­ generations­ in­ the­ case­ of­N. socialis,­with­ a­ “micro­reef”/bioherm­appearance. From­the­current­material­obtained,­Schlagintweitella­nov.­ gen.­also­develops­as­an­agglomeration­of­thalli­(Pl.­1,­figs.­ A­C).­However,­it­differs­from­Neutloporella by­the­more­elon­ gated­cylindrical­shape­of­the­first­order­laterals­and­acrophore­ type­second­order­laterals,­which­have­rarer­constrictions­and­ often­an­irregular­arrangement.­These­laterals­are­included,­at­ least­in­their­lower­part,­in­a­collective­sheath.­Moreover,­the­ skeleton­of­Neoteutloporella­is­formed­„by­a­succession­of­very­ wide­articles,­roughly­funnel­shaped”­(DE­CASTRO,­1993,­p.­ 174),­and­the­thallus­is­branched. Schlagintweitella inopinata nov. sp. Pls.­1­5 Derivation­of­name:­inopinatus­(latin)­=­unexpected. Material:­ four­ thin­sections­from­two­samples,­containing­ more­than­50­specimens. Holotype:­the­specimen­in­Pl.­2­fig.­A,­thin­section­15207(2),­ stored­in­the­collection­of­the­Paleontology­Museum,­Babeş­ Bolyai­University­Cluj­Napoca­under­the­inventory­number­ 24412. Paratypes:­Specimens­in­Pl.­2­fig.­B,­Pl.3,­figs.­E­G,­ thin­ sections­3751(2),­15207(1)­and­3751(1)­stored­in­the­same­col­ lection­under­the­inventory­numbers­24410,­24411­and­24409­ respectivelly. Type­locality:­Lunca­Ohabei­plateau,­Cioclovina­Băniţa­zone,­ Southern­Carpathians,­Romania­(Fig.­1).­GPS­coordinates:­ 45°31'35.61"N;­23°­9'55.65"E. Type­level:­Upper­Jurassic,­most­probably­upper­Tithonian. Diagnosis:­Dasycladalean­alga­with­an­elongated­cylindrical­ thallus,­a­relatively­wide­axial­cavity­and­two­orders­of­lateral­ branches­arranged­obliquely­to­the­axial­cavity.­The­first­order­ laterals­are­cylindrical­(acrophores)­or­ovoid­to­elongated­and­ shorter­ than­ those­of­ the­ second­order.­The­ second­order­ laterals­are­also­cylindrical,­acrophorous,­longer­than­those­of­ the­first­order­and­have­one­or­more­constrictions­that­separate­ them­into­successive­segments.­The­calcification­consists­of­a­ calca­reous­sheath­surrounding­the­central­cavity­and­each­ lateral­branch.­In­their­lower­part,­the­second­order­laterals­are­ wrapped­in­a­collective­sheath.­ Dimensions­(in­mm,­except­w1­and­w2) D­(external­diameter)­=­1.30–2.60­(median­=­1.97) d­(internal­diameter)­=­0.50–1.70­mm­(median­=­1.05­mm) p1­(diameter­of­the­primary­laterals)­=­0.06–0.12­mm­(median­ =­0.09­mm) l1­(length­of­the­primary­laterals)­=­0.13–0.30­mm­(median­=­ 0.19­mm) p2 (diameter­of­the­secondary­laterals)­=­0.05–0.10­mm­(median­ =­0.06­mm) l2­(length­of­the­secondary­laterals)­=­0.28–0.60­mm­(median­ =­0.41­mm) h­(distance­between­two­consecutive­verticils)­=­0.10–0.15­ (median­=­0.13) w1­(number­of­primary­laterals­in­a­verticil)­=­approx.­25­to­60 w2­(number­of­secondary­laterals­per­primary­lateral)­=­2–3 Description:­The­new­alga­occurs­as­agglomerations­of­thalli­ within­cal­careous­clasts­or­black­pebbles­(Pl.­1,­figs.­A–C).­ Longitudinal­and­longitudinal­oblique­sections­(Pl.­2,­figs.­ A–E)­show­a­cylindrical­thallus,­sometimes­slightly­curved­ (Pl.­2A),­with­a­relatively­smooth­axial­cavity­and­a­rather­ irregular­external­surface,­marked­by­the­different­lengths­(and­ some­times­different­orientations)­of­the­secondary­laterals.­ These­features­are­also­visible­in­oblique­or­oblique­longitudinal­ Figure 2. The general stratigraphic succession of the Jurassic and Creta- ceous deposits from the Cioclovina-Băniţa area (Modified from PLEŞ et al., 2019). The arrow points to the stratigraphic position of the outcrop with the new alga. G eo lo gi a C ro at ic a 106 Geologia Croatica 78/2 sections­(Pl.­3,­figs.­A–I;­Pl.­4,­figs.­C,­I,­J)­and­in­transverse­ sections­(Pl.­2,­fig.­F;­Pl.­4.,­figs.­A,­B,­D,­G,­H).­The­primary­ laterals­are­tubular­(most­often­cylindrical;­e.g.,­Pl.­3,­figs.­C­F;­ sometimes­they­are­ovoid­elongated,­e.g.,­Pl.­5,­fig.­C).­The­ primary­laterals­are­provided­at­the­lower­part­with­a­relatively­ wide­pore­that­connects­them­with­the­axial­cavity.­From­the­ primary­ laterals­ two­(sometimes­ three)­secondary­ laterals­ originate­that­are­also­cylindrical­(acrophorous)­(Pl.­5,­figs.­A,­ G,­I)­and­present­at­certain­intervals­constrictions­that­separate­ them­into­successive­segments­delimited­by­a­fine­micritic­ fringe­(Pl.­5,­figs.­A,­D,­E,­arrows).­The­large­number­of­late­ rals­in­a­whorl­makes­it­difficult­to­observe­their­arrangement­ around­and­along­the­axial­cavity.­However,­some­sections­ through­smaller­specimens­(e.g.,­Pl.­3,­figs.­E­G;­Pl.­4,­figs.­D,­ H,­J),­where­the­thallus­is­tangentially­obliquely­sectioned,­ indicate­a­euspondyl­arrangement.­The­secondary­laterals,­ however,­occasionally­seem­to­orient­themselves­in­various­ directions­(Pl.­3,­figs.­C,­D,­F),­giving­an­irregular­appearance­ to­the­external­part­of­the­thallus.­The­calcification­consists­of­ a­calcareous­sheath,­observable­in­some­places­around­the­ axial­cavity­(Pl.­2,­figs.­A,­F;­Pl.­3,­figs.­A,­F,­H,­I;­Pl.­4,­figs.­ B,­H),­but­especially­around­the­laterals,­being­clearly­visible­ where­the­laterals­are­sectioned­more­or­less­transversely­(Pl.­ 2,­fig.­B­–­lower­part;­Pl.­3,­figs.­A,­D­I;­Pl.­4,­fig.­G).­One­of­ the­important­characteristics­of­the­new­alga­is­the­presence­ of­ a­ common­ calcareous­ sheath­ (collective­ sheath­ cf.­DE­ CASTRO,­1997)­that­envelops­the­proximal­part­of­the­secon­ dary­laterals­(e.g.,­Pl.­2,­fig.­G;­Pl.­4,­figs.­E,­J;­Pl.­5,­figs.­H,­J).­ The­term­collective­sheath­introduced­by­DE­CASTRO­(1997)­ was­illustrated­by­this­author­in­pl.­4,­figs.­1–3­through­two­ specimens­ of­ an­ alga­ defined­ as­ "Dasycladacean­ with­ unsegmented­primary­and­segmented­secondary­laterals"­and­ which­actually­represents­the­first­illustration­in­the­literature­ of­Schlagintweitela inopinata­nov.­gen.,­nov.­sp.­It­is­worth­ mentioning­that­while­some­specimens­present­a­calcareous­ sheath­of­sparitic­calcite­(Pl.­3,­fig.­G;­Pl.­4,­figs.­D,­F,­G;­Pl.­5,­ fig.­H),­sometimes­with­a­yellowish­brown­tint­(Pl.­3,­fig.­B;­ Pl.­4,­figs.­E,­H,­J;­Pl.­5,­fig.­J),­most­specimens­have­a­strongly­ blackened­sheath­due,­most­ likely,­ to­ the­accumulation­of­ organic­matter­during­the­blackening­process­that­affected­the­ black­pebbles­ in­which­ they­ are­ contained.­Reproductive­ structures­were­not­encountered,­but­their­presence­in­the­first­ order­laterals­cannot­be­excluded­(cladospore). Comparison:­Within­the­large­group­of­Jurassic–Cretaceous­ Dasycladales­with­ two­orders­of­ laterals,­Schlaginweitella inopoinata­nov.­gen.,­nov.­sp.­is­most­­similar­to­species­of­the­ genus Neoteut loporella (N. obsoleta and N. socialis)­and­with­ some­species­of­the­genus­Steinmanniporella (e.g., S. kapelen sis, S. taurica).­It­differs­from­Neoteutloporella­by­the­shape­and­ dimensional­ratio­of­the­primary­and­secondary­laterals,­as­ well­as­the­lack­of­articulation.­It­differs­from­Steinmanniporella by­the­seg­men­tation­of­the­secondary­laterals. Associated microfossils: In­ the­ outcrop­ from­ the­ Lunca­ Ohabei­plateau,­Schlagintweitella inopinata­nov.­gen.,­nov.­sp.­ is­accompanied­by­an­assemblage­of­foraminifera­and­cal­­ careous­ algae­ consisting­ of:­Everti cyclammina virguliana (KOECHLIN), Everticyclammina­sp.,­Redmondoides lugeoni (SEPTFONTAINE), Charentia­sp.,­Coscinoconus alpinus LEUPOLD­ (Pl.­ 6,­ figs.­ G,­ I),­ Salpingo porella annulata (CAROZZI)­ (Pl.­ 6,­ figs.­A–D),­Salpingopo rella pygmaea (GÜMBEL)­(Pl.­6,­figs.­E–F),­S.­cf.­praturloni (DRAGASTAN), Aloisalthella sulcata (ALTH)­ (Pl.­6,­fig.­ J),­Campbelliella striata (CAROZZI),­and­ rivulariacean­like­cyanobacteria.­ Figure 3. The outcrop from the Lunca Ohabei plateau (Cioclovina-Băniţa zone) representing most probably a palaeo-cave filling where the sample with Schlagintweitella inopinata nov. gen., nov. sp. has been found. Figure 4. A pebble of the cave filling showing the new alga on a weath- ered surface. G eologia C roatica 107Bucur et al.: Schlagintweitella inopinata, a new genus and species of Dasycladales (green algae) from the Upper Jurassic limestones of Romania Some­of­the­microfossils­in­this­association­are­coloured­with­ a­light­brown­tint,­likely­due­to­the­partial­incor­poration­of­ organic­matter.­The­entire­association­indicates­a­Late­Jurassic­ age,­most­likely­late­Tithonian. 5. CONCLUSION A­new­dasycladalean­alga,­Schlaginweitella inopinata­nov.­ gen.,­nov.­sp.­was­identified­in­the­Upper­Jurassic­limestones­ of­the­Cioclovina­Băniţa­area­(Southern­Carpathians,­Roma­ nia).­By­its­morphological­characteristics,­the­new­alga­differs­ from­all­other­genera­ and­ species­of­ Jurassic–Cretaceous­ Dasy­cladales­with­two­orders­of­laterals. ACKNOWLEDGEMENT We­thank­Bruno­Granier­(Brest)­and­Marc­A.­Conrad­(Geneve)­ for­their­remarks­and­suggested­corrections. 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STILLĂ,­A.­(1985):­Géologie­de­la­région­de­Haţeg­Cioclovina­Pui­Băniţa­ (Carpathes­Méridionales)­[Geology of the Haţeg-Cioclovina-Pui-Băniţa –­in­Romanian].–­Annuaire­de­l’Institut­de­Geologie­er­Geophysique,­66,­ 91–179. Web sources: TROPICAL­FISH­PLUS­(2024):­Neomeris.­Available­at:­https://­tropicalfi­ shplus.com/products/neomeris.­Accessed­on:­December­1st,­2024. https://doi.org/10.1016/j.earscirev.2023.104666 https://doi.org/10.1007/s10347-019-0574-3 https://doi.org/10.1007/s10347-019-0574-3 G eo lo gi a C ro at ic a 108 Geologia Croatica 78/2 Plate 1. A–C general views of an agglomeration of thalli of Schlagintweitella inopinata nov. gen., nov. sp. Thin-sections: A, 3751(1); B, 3751(2); C, 15207(1). G eologia C roatica 109Bucur et al.: Schlagintweitella inopinata, a new genus and species of Dasycladales (green algae) from the Upper Jurassic limestones of Romania Plate 2. Schlagintweitella inopinata nov. gen., nov. sp. A–E longitudinal (A) and longitudinal oblique (B–F) sections. A = holotype; B = paratype; F = transverse section. Thin-sections: A, D-F, 15207(2); B, 3751(2); C, 15207(1). G eo lo gi a C ro at ic a 110 Geologia Croatica 78/2 Plate 3. A-I Schlagintweitella inopinata nov. gen., nov. sp., oblique sections. E and G, paratypes. Thin-sections: A, C-F, H-I, 15207(1); B, 3751(2); G, 3751(1). G eologia C roatica 111Bucur et al.: Schlagintweitella inopinata, a new genus and species of Dasycladales (green algae) from the Upper Jurassic limestones of Romania Plate 4. Schlagintweitella inopinata nov. gen., nov. sp. A, B, D, G transverse sections. C, F, H-J oblique sections. E is a close-up view of the lower part of J. Thin-sections: A-C, H-I, 15207(1); D-G, J, 3751(2). G eo lo gi a C ro at ic a 112 Geologia Croatica 78/2 Plate 5. Schlagintweitella inopinata nov. gen., nov. sp. A-J details of laterals, showing the primary (r1) and the secondary laterals (r2). The arrows in A, D and E point to the constrictions separating segments inside the secondary laterals. The arrows in H and J point to the collective sheath around the sec- ondary laterals. Thin-sections: A, D-G, I-J, 15207(1); B, 15207(2); C, 3751(2); H, 3751(1). G eologia C roatica 113Bucur et al.: Schlagintweitella inopinata, a new genus and species of Dasycladales (green algae) from the Upper Jurassic limestones of Romania Plate 6. Microfossils associated with Schlagintweitella inopinata nov. gen., nov. sp. A-D Salpingoporella annulata (CAROZZI); E-F Salpingoporella pygmaea (GÜMBEL); H Frentzenella involuta (MANTSUROVA); G-I Coscinoconus alpinus LEUPOLD. J Aloisalthella sulcata (ALTH). Thin-sections: A, 3747(2); B-D, G-I, 3767; E, 3751(9); F, 3751(6); J, 3747(5).