Schlagintweit.indd 1. INTRODUCTION The Upper Jurassic to Lower Cretaceous shallow water platform carbonates of the Northern Calcareous Alps (Plassen Formation, “Lärchberg Formation”) and the resedimented deposits of the latter within basinal series (Barmstein Limestone, Tressenstein Limestone) are currently under re-investigation (SCHLAGINTWEIT et al., 2003; GAWLICK et al., 2005). This shallow water development is directly linked to tectonic activity in the Western Tethyan domain causing early nappe forma- tions (GAWLICK et al., 1999; FRISCH & GAWLICK, 2003). Already in the Middle Jurassic, the geodynamic and sedimentological framework was mainly governed by the closure of parts of the Tethyan ocean leading to the first thrust nappes and the formation of a rise-and- basin pattern (e.g. GAWLICK et al., 1999). Starting in the southern parts of the Northern Calcareous Alps, the compressional tectonic regime governed uplift proc- esses producing shallowing-upwards successions from basinal to external and finally internal carbonate plat- form deposits. In the same way as there were different Neogyroporella? gawlicki n.sp., a New Dasycladale from the Upper Jurassic–Lower Cretaceous “Lärchberg Formation” of the Northern Calcareous Alps, Austria Felix SCHLAGINTWEIT basins with cherty sediments (“radiolarite basins”), the new results show that there was not only one, but sev- eral independent areas with shallow water carbonates within the former deeper-marine area (e.g. KÜGLER et al., 2003 – fig. 3; FRISCH & GAWLICK, 2003; GAWLICK & FRISCH, 2003 – for further details, and unpubl. data). Current research should help to increase the preci- sion of existing palaeogeographic and geodynamic inter- pretations and models. However, there is still a lack of knowledge of the micropalaeontological composition of these platform carbonates, in contrast to the compara- bly good data base of the Alpine Triassic. In the course of recent biostratigraphic and microfacies analyses of the Alpine “Upper Jurassic” platform carbonates, a Dasycladacean alga, that has already been figured sev- eral times from the Northern Calcareous Alps since the early 1990’s as Macroporella praturloni DRA- GASTAN, has been recognized as a new species and is established as Neogyroporella? gawlicki n.sp. 2. GEOLOGICAL SETTING In the Salzburg Calcareous Alps, near the German– Austrian border, there are several isolated occurrences of shallow water limestones known as the “Lärchberg Formation” (FERNECK, 1962), currently a non-formal- ized unit. For a long time, these carbonates have been assigned to the Triassic, e.g. Dachstein Limestone, and it was FERNECK (1962) who first recognized their Upper Jurassic–Lower Cretaceous age. He proposed the subdivision of the “Lärchberg Formation” into two parts: a clastic basal complex (Lofer beds or L. mem- ber) and the Lerchkogel Limestone. These litholo- gies have been referred to in the works of FERNECK (1962), DARGA & SCHLAGINTWEIT (1991) and DYA (1992). Another Upper Jurassic carbonate plat- form evolutionary series in the Northern Calcareous Alps is known as the Plassen Formation, with the type- locality Mount Plassen near Hallstatt in the Austrian Salzkammergut (e.g. TOLLMANN, 1976). In contrast to the latter, the “Lärchberg Formation” partly shows a terrigenous influx unknown from the former. However, detailed discussions on their possible correlation are Geologia Croatica 58/2 103–117 3 Figs. 3 Tabs. 3 Pls. ZAGREB 2005 Key words: Calcareous algae (Dasycladales), Taxon- omy, Upper Jurassic, Lower Cretaceous, “Lärchberg Formation”, Northern Calcareous Alps, Austria. University Leoben, Institute for Geosciences, Prospektion and Applied Sedimentology, Peter-Tunner-Str. 5, A-8700 Leoben, Austria; e-mail: EF.Schlagintweit@t-online.de. Abstract A new Dasycladale (calcareous green alga) is described as Neogyro- porella? gawlicki n.sp. from the Upper Jurassic–Lower Cretaceous “Lärchberg Formation” of the Northern Calcareous Alps, Austria. It is distinguished from the type-species Neogyroporella elegans YABE & TOYAMA (Upper Jurassic of Japan; currently the only representa- tive of the genus), by its distinctly smaller dimensions and a higher number of laterals. The generic attribution of the new taxon is dis- cussed, including similarities/differences to allied genera, especially Humiella SOKAČ & VELIĆ. Neogyroporella? gawlicki n.sp. is so far only known from the Northern Calcareous Alps. In addition, some other Dasycladales accompanying Neogyroporella? gawlicki n.sp. are also illustrated. 104 Geologia Croatica 58/2 beyond the scope of this paper. The marly limestones of the “Lärchberg Formation” are characterized by a rich assemblage of Dasycladales together with “stromatopo- roids”, benthic foraminifera and cyanophycean algae. In some parts, representatives of the protohalimedacean alga Pinnatiporidium (sensu DRAGASTAN et al., 2002) are dominant and can be termed Pinnatiporidium wackestones to floatstones. These lithologies yielded the new Dasycladale Neogyroporella? gawlicki n.sp. detected at the Dietrichshorn and Litzelkogel (Fig. 1). The marly limestones containing the new alga out- crop near the summit of the Litzelkogel (1625 m a.s.l.), which forms a plateau-like mountain, together with the Gerhardstein (1541 m a.s.l. – Fig. 2). Due to sporadic outcrops and vegetation as well as steep flanks border- ing the plateau, a lithological profile cannot be provid- ed. According to DYA (1992), the “Lärchberg Forma- tion” of the Litzelkogel-Gerhardstein has a thickness of about 250 m. Neogyroporella? gawlicki n.sp. was figured by DYA (1992 – Profile 11) as Macroporella? praturloni DRAGASTAN (see synonymy below) from the northern part of the Gerhardstein–Litzelkogel pla- teau. Detailed sampling carried out in recent times, has shown that Neogyroporella? gawlicki n.sp. is missing at the Gerhardstein. Here, the youngest sediments exposed can be attributed to the uppermost Kimmeridgian or lowermost Tithonian due to subaerial exposure (unpub- lished data). At the Dietrichshorn (1542 m a.s.l.), located about 11 km northwest of the Litzelkogel, Neogyropore- lla? gawlicki n.sp. was also figured by DARGA & SCHLAGINTWEIT (1991) as Macroporella? pratur- loni DRAGASTAN (see synonymy below) from marly limestones that have been attributed to the Lofer Mem- ber, the clastic influenced part of the “Lärchberg For- mation”. For further details see DARGA & SCHLAG- INTWEIT (1991). 3. SYSTEMATICS Order Dasycladales Tribe Salpingoporelleae Genus Neogyroporella YABE & TOYAMA, 1949 Remarks: As will be discussed below, we tentative- ly assign our specimens to the genus Neogyroporella YABE & TOYAMA. The genus Neogyroporella with the type-species Neogyroporella elegans has been estab- lished by YABE & TOYAMA (1949), from the Upper Jurassic Torinosu Limestone of the Sakawa Basin, Japan. The age of the Torinosu Limestone, previously referred to the Oxfordian–Tithonian interval, has more recently been precisely dated as Tithonian–Berriasian (e.g. OHGA & IRYU, 2003; SHIRAISHI & KANO, 2004). Therefore the exact stratigraphic position of Neogyroporella elegans is not known in detail. Neo- Fig. 1 Generalized geographic sketch map of the localities where the new Dasycladale Neogyroporella? gawlicki n.sp. has been detected. 105Schlagintweit: Neogyroporella? gawlicki n.sp., A New Dasycladale... gyroporella elegans has not been recorded again since 1949 from its type-locality nor from any other occur- rence. The original figures, however, have been re- illustrated by JOHNSON (1964), BASSOULLET et al. (1978) and DELOFFRE (1988). BASSOULLET et al. (1978) regard the “weakly known alga” as well charac- terized by the shape and arrangement of the primaries. The latter are of the cladosporous type (BARATTOLO, 1991 – p. 523). In his classification of the Dasycladales the genus Neogyroporella has been included in the sub- tribus Salpingoporellinae BASSOULLET et al. 1978 by DELOFFRE (1988), characterized by phloiopho- rous first order laterals. The primaries can also be com- pared with elongated vesiculiferous laterals comparable for example with the Triassic Gyroporella ampleforata PIA (see Pl. II, Fig. 5). OTT & FLAVIANI (1983 – p. 25) state that on two original figures of Neogyroporella elegans, the existence of a corona at the proximal part of the laterals could be assumed. YABE & TOYAMA (1949 – p. 162) just indicate that the calcareous crust “more or less thickens at the very basal part” of the lat- erals. Anyway, the existence of a corona structure in Neogyroporella has not so far been observed. It should be noted that the presence/absence of such proximal modifications of the laterals are regarded as generic characteristics. For example Clypeina-type dasycladales with a short basal portion below the junction point were removed from Clypeina and refered to the genus Hamu- lusella ELLIOTT by BARATTOLO (1998). Similar structures were also illustrated by DE CASTRO (1997 – pl. 20, fig. 1) from Clypeina sulcata (ALTH) and called “small lower protuberance (lp)”. A second species of Neogyroporella has been described as Neogyroporella servaisi by MICHAUD (1988) from the Maastrichtian of Mexico. Due to the existence of pedunculated primaries it has been trans- ferred to the genus Morelletpora by GRANIER et al. (1991). BARATTOLO (2002 – p. 25), however, pre- fers to keep the species in the genus Neogyroporella although with some doubts. In revision of the Jurassic– Cretaceous Dasycladales by GRANIER & DELOFFRE (1993), Neogyroporella elegans has been treated as a valid taxon and the only species of the genus Neogyro- porella. Although never discussed previously, another genus very similar to Neogyroporella is Humiella SOKAČ & VELIĆ (1981), with the type-species Humiella catenae- formis (RADOIČIĆ) from the Neocomian of Southern Herzegovina. In fact, representatives of Humiella are characterized by alternating fertile laterals (ampullae), closed at the tips which are of variable shape (mostly bulbous). In both the original description, and others, “a thin corticular envelope enwrapping the soft parts of the plant” has been proposed by SOKAČ & VELIĆ (1981 – p. 104) as the main generic characteristic. This feature has also been remarked for Neogyroporella by YABE & TOYAMA (1949 – “surface is covered by a thin calcar- eous crust”). Later, the main characteristic of Humiella was said to be represented by tiny pores or indentations in the calcareous wall of the laterals with the main axis corresponding to sterile laterals (MASSE et al., 1984; SOKAČ, 1987). In fact, these “pores” or indentations (= “kleine Zähnchen” – OTT & FLAVIANI, 1983) obviously only seldom cut through the whole wall and can therefore be termed “blind pores” as described from the genus Drobnella BARATTOLO, 1998 (Chlo- rophyta or Charophyta). The existence of such blind pores or real pores, however, is not clearly observed in the specimens from the Northern Calcareous Alps. Due to strong recrystallization the boundary between adja- Fig. 2 General position of the sample localities with Neogyroporella? gawlicki n.sp. at the Litzelkogel type locality. The sample A-190 contains the holo- type. 106 Geologia Croatica 58/2 cent calcite crystals may suggest a finely perforated wall (diagenetic origin!). This uncertainty has been the reason to only tentatively refer these specimens to the genus Neogyroporella. However, if a re-investigation of the type-material of Neogyroporella elegans YABE & TOYAMA shows both types of pores in the calcare- ous walls, as could perhaps be assumed in, for example, fig. 5 from YABE & TOYAMA (1949) and any kind of corona structure is absent, the genus Humiella would become very close to Neogyroporella or perhaps even a synonym of the former. Finally, it should be mentioned that the genus Humiella has been included in the tribe Clypeineae with “verticilles espacés” by DELOFFRE (1988). The genus Humiella, however, includes both species with well spaced-out verticals, such as Humiella sardinien- sis (OTT & FLAVIANI), or forms with closely packed touching laterals, such as Humiella japodica recently described by SOKAČ (2001) from the Lower Jurassic of Croatia. For the Clypeina species with slightly touch- ing whorls (“overlapping whorls”), the genus Similicly- peina has been established by BUCUR (1993). Last but not least, the systematic framework beco- mes more complex as the genus Humiella SOKAČ & VELIĆ has been regarded as a synonym of Sarfatiella CONRAD & PEYBERNES by CONRAD (1982). Sar- fatiella, however, is treated as a synonym of Holospore- lla PIA by BASSOULLET (1987) and by GRANIER & DELOFFRE (1993). The whole brief discussion of current taxonomic incongruencies is not pursued further here; it is just mentioned for completeness, but it also expresses the uncertainties we have in the systematic attribution of these specimens (see also Table 1). Neogyroporella? gawlicki n.sp. Pl. I, Figs. 1–9; Pl. II, Figs. 1–2, 4; Pl. III, Figs. 2–5 1991 “Macroporella” praturloni DRAGASTAN – DARGA & SCHLAGINTWEIT: p. 209; pl. 3, figs. 2–3 1991 “Macroporella” praturloni DRAGASTAN – SCHLAGINTWEIT: pl. 1, fig. 7 1992 “Macroporella” praturloni DRAGASTAN – DYA: p. 83; pl. 7, fig. 15; pl. 8, figs. 4–5, 8 Origin of the name: Dedicated to Prof. H.-J. Gaw- lick, University of Leoben, in recognition of his stim- ulating new concepts and ideas concerning the geody- namic evolution of the Alpine Jurassic. Type-locality: Litzelkogel, Topographic Map of Aus- tria ÖK 92 sheet Lofer, marly limestones cropping out in the area of the summit (Fig. 2). The location of the summit of the Litzelkogel is 12º46’ geographic longi- tude and 47º38’ geographic latitude. Type strata: Light brownish, marly limestones of the “Lärchberg Formation”, partly displaying a siliciclastic influx. The Dasycladales include Salpingoporella gr. pygmaea (GÜMBEL) (common), Salpingoporella cf. istriana (GUŠIĆ) (rare; Pl. III, Fig. 1), Chinianella? sp. (rare; Pl. III, Fig. 10), Montenegrella florifera BERNI- ER (Pl. III, Fig. 6), Neogyroporella? gawlicki n.sp., with protohalimedacean algae (genus Pinnatiporidium) and benthic foraminifera with Comaliamma? cf. gediki (TASLI), Rheopax sp. and Protopeneroplis ultragran- ulata (GORBATCHIK) (Pl. III, Fig. 7). In addition, remains of gastropods, stromatoporoids, oncoids and nodules formed by Lithocodium aggregatum ELLIOTT are present. At the Dietrichshorn locality, Neogyroporella? gaw- licki n.sp. is associated with Clypeina aff. parasolkani FARINACCI & RADOIČIĆ (Pl. II, Fig. 6), Clypeina catinula CAROZZI (Pl. II, Fig. 13), Clypeina aff. isa- bellae MASSE, BUCUR, VIRGONE & DALMASSO (Pl. II, Fig. 11), Cylindroporella? sp. (Pl. II, Fig. 9), Zer- gabriella embergeri (BOUROULLEC & DELOFFRE) (Pl. II, Fig. 7), Salpingoporella annulata CAROZZI, Rajkaella bartheli (BERNIER) and Deloffrella querci- foliipora GRANIER & MICHAUD (Pl. II, Figs. 8, 10, 12) and the benthic foraminifera Anchispirocyclina lusi- tanica (EGGER) and Kastamonina abanica SIREL (Pl. III, Figs. 8–9). At both the Dietrichshorn and Litzelko- gel localities, the Dasycladalean remains typically show micritic envelopes of presumably cyanophycean origin. Material: One thin-section containing about 30 speci- mens from Mount Dietrichshorn and several thin-sec- tions each with 2–5 specimens from Mt. Litzelkogel. Genus Neogyroporella Humiella Clypeina Similclypeina Actinoporella Laterals in mutual contact in mutual contact spaced-out in mutual contact spaced-out or spaced-out Corona ? (1) missing missing (1) missing present structure Blind pores ? present maybe present (2) missing missing (sterile branches) Table 1 Main characteristics of Neogyroporella and allied Dasycladale genera. Legend: (1): see remarks in the text, (2): as shown in some Clypeina species (e.g. RADOIČIĆ, 1969). 107Schlagintweit: Neogyroporella? gawlicki n.sp., A New Dasycladale... Holotype: Oblique-tangential section figured on Pl. I, Fig. 5, thin section labelled A–190 (exact location see Fig. 2). The material from the Litzelkogel is stored at the “Institut für Geowissenschaften, Universität Leoben”. Isotypes are represented by different sections shown in Pls. I–III. Diagnosis: Thallus euspondyl, cylindrical with a com- parable small main axis and numerous elongated, gently broadening laterals with swollen endings. The laterals, closed at tips, are arranged more or less perpendicular or slightly inclined to the axis. The microstructure of the individual calcified sheaths enclosing the laterals has a fibrous appearance. The whorls are close-set with alternating laterals touching those from adjacent whorls. Reproductive organs presumably cladosporous. Description: Thallus simple and cylindrical. The axial cavity is comparatively small, most frequently comprising 27–30% of the total diameter. The simple, undivided and elongated laterals communicate with the axial cavity by means of narrow connecting pores that broaden gently, then swell into sphere-like distally closed ends. The arrangement of laterals in consecu- tive whorls is alternating. The main axis and the laterals are each covered with their own thin calcareous enve- lope. Due to secondary calcification, the laterals are in contact at their proximal parts. At the most distal parts, adjacent laterals are not in contact with each other. The outer part of the thallus is composed of the distal ends of individual laterals of alternating arrangement; the inner surface of the thallus is straight. The laterals are slightly inclined to the axis at an angle between 5 to 15º. In tangential sections, the laterals appear as a rounded pore meaning that they are neither longitudinally nor transversally compressed. Although cysts have not been observed in our specimens, the swollen laterals suggest a cladosporous type of reproduction. Dimensions: see Table 2 and Fig. 3. Remarks: The species in question has repeatedly been confused with Macroporella praturloni DRAGASTAN (see synonymy). In fact, the laterals with individual calcareous sheets are very similar in tangential sections and also the d/D ratio is the same. M. praturloni, how- ever, is on the whole different from Neogyroporella? gawlicki n.sp. by possessing phloiophorous laterals, open at the tips and distinctly inclined to the axis. Addi- tionally, M. praturloni is much larger than N.? gawlicki n.sp. with a thallus diameter reaching up to several mil- limetres (see also BUCUR, 1985 – table 1). The only species to which Neogyroporella? gawlicki n.sp. can be compared to is the type-species N. elegans YABE & TOYAMA. Generally, this new species can be regarded as a “mini version” of the latter. N. elegans is distinctly larger than N.? gawlicki n.sp. (see Table 3). The d and D values do not show any overlapping areas but are distinctly separated from each other. Thus, both species are clearly individual due to their dimensions. In addition, the type-species possess a lower number of laterals (w, see Table 3) which more or less thicken at their proximal parts. In addition, a second taxon has been briefly recorded by YABE & TOYAMA (1949) No. D d d/D pmax l h w L 1 0.4 0.1 0.26 2 0.52 0.09 0.28 22 3 0.56 0.18 0.31 4 0.56 0.16 0.28 5 0.59 0.2 0.34 0.09 0.22 24 6 0.62 0.17 0.27 0.055 7 0.64 0.19 0.3 23 8 0.64 0.29 0.185 9 0.67 0.19 0.29 10 0.68 0.2 0.29 21 11 0.68 0.19 0.28 22 12 0.7 0.14 0.2 0.22 13 0.74 0.24 0.32 0.25 14 0.8 0.22 0.28 2.65 15 0.8 0.22 0.28 24 16 0.8 0.2 0.25 24 17 0.8 0.22 0.28 0.08 0.28 0.07–0.08 2.75 18 0.98 0.37 0.38 0.1 0.08–0.09 5.68 19 0.08 20 1.2 0.2 0.17 Table 2 The main biometric parameters of Neogyroporella? gawlicki n.sp. (in mm). Legend: D – outer thallus diameter; d – inner diameter; pmax – maximum diameter of laterals; l – length of laterals; h – distance between whorls; w – number of laterals per whorl; L – maximum observed length. No. 20 – data given by DYA (1992). 108 Geologia Croatica 58/2 as Neogyroporella sp. from the same locality as N. ele- gans. While being distinctly larger than N.? gawlicki n.sp., the number of laterals per whorl (~34) approach the maximum value measured for N.? gawlicki (~26) (see Table 3). Stratigraphy: Based on the aforementioned micro- fossil assemblages and results of previous and recent investigations, the samples containing Neogyroporella? gawlicki n.sp. can be assigned to the Late Tithonian or Early Berriasian. The evaluation of the total stratigraph- ic range, however, cannot be established reliably and surely needs further data. In this connection it should be mentioned that Neogyroporella? gawlicki n.sp. has not been detected in the samples of the Gerhardstein where the youngest sediments match the Kimmeridgian–Titho- nian boundary or the earliest Tithonian. However, the marly limestones containing Neogyroporella? gawlicki n.sp. which crop out at the Litzelkogel summit repre- sent the final part of the exposed succession. Facies and palaeoenvironmental interpreta- tion: The marly limestones that crop out around the summit of the Litzelkogel (see Fig. 2) follow micrite dominated low energy platform margin deposits that are represented by Bacinella–Lithocodium bindstones associated with “stromatoporoid” wackestones (genus Milleporidium). As no high energy outer central reef deposits characterized by Actinostromariids were found, it is assumed that the barrier reef model of TURNŠEK et al. (1981) cannot be applied. It should be noted that the protohalimedacean alga Pinnatiporidium frequent- ly associated with Neogyroporella? gawlicki n.sp., is according to our own observations, also a typical back- reef inhabitant. In a Late Jurassic–Early Cretaceous platform model from the East Carpathians of Roma- nia, DRAGASTAN (1999 – fig. 6) also reported on a bindstone facies with Lithocodium and Pinnatiporidium assigned to a reef-flat facies that passes to a back-reef lagoon facies. Therefore, the wackestones, which are partly oncoidal, containing Neogyroporella? gawlicki Fig. 3 Neogyroporella? gawlicki n.sp.: outer diameter of the thallus against the inner diameter of the thallus (d). The param- eters d and D are well correlated except the value indicated by DYA (1992 – D = 1.2 mm). Species Neogyroporella elegans Neogyroporella sp. Neogyroporella? gawlicki n.sp. YABE & TOYAMA YABE & TOYAMA Dimensions D 3.5–5.5 3.5 0.4–1.2 d 1.1–1.5 2.1 0.1–0.37 d/D – – 0.185–0.38 p – – 0.055–0.1 l 1.5–2.0 – 0.22–0.28 h – – 0.07–0.09 w about 13 ~34 21–26 L – – 5.68 Table 3 Dimensions of Neogyroporella elegans YABE & TOYAMA, Neogy- roporella sp. and Neogyroporella? gawlicki n.sp. (in mm). 109Schlagintweit: Neogyroporella? gawlicki n.sp., A New Dasycladale... n.sp. may be assigned to a sheltered “back-reef” envi- ronment with terrigenous influence (siliciclastic detri- tus, terrestrial? plant remains). At the Dietrichshorn locality, the co-occurrence with Zergabriella embergeri (BOUROULLEC & DELOF- FRE) may point to freshwater influences (CONRAD, 1977). Indications of a very shallow water setting with signs of subaerial exposure are indicated by components showing circumgranular cracking sensu ESTEBAN & KLAPPA (1983). For Neogyroporella elegans, YABE & TOYAMA (1949) did not give comments on the facies or associ- ated microflora and microfauna, but the illustrated figu- rations show a micritic microfacies (wackestones) indi- cating a quiet water setting. Thus, the Torinosu Forma- tion does not only comprise reefal facies (SHIRAISHI & KANO, 2004) but also lagoonal facies, as indicated by the occurrence of wackestones with Cladocoropsis mirabilis FELIX (YABE & TOYAMA, 1927). 4. CONCLUSIONS Neogyroporella? gawlicki n.sp. represents another spe- cies of the so-far poorly known genus Neogyroporella YABE & TOYAMA. It occurs in marly limestones of the “Lärchberg Formation” of Late Tithonian or Early Berriasian age. This terrigeneously influenced litho- facies, rich in calcareous algae has up to now not been recorded from the Alpine Plassen Formation. Thus, many Dasycladales that occur together with Neogyro- porella? gawlicki n.sp., such as Zergabriella embergeri (BOUROULLEC & DELOFFRE), Deloffrella quercifo- liipora GRANIER & MICHAUD or Clypeina catinula CAROZZI, are therefore also missing in the Plassen Formation. The new species Neogyroporella? gawlicki n.sp. furthers the knowledge not only of the poorly known genus Neogyroporella YABE & TOYAMA but also of Upper Jurassic–Lower Cretaceous Dasycladales of the Northern Calcareous Alps. Acknowledgements Financial support from the Austrian Science Foun- dation, FWF-project P 16812–B06 is kindly acknow- ledged. The samples from the Dietrichshorn have been provided by R. DARGA (Naturkundemuseum of Siegs- dorf, Bavaria). The material from the Litzelkogel has been collected by H.-J. GAWLICK and S. MISSONI (University of Leoben, Austria). Both are also thanked for logistic support. The discussion concerning the close affinity of Neogyroporella and Humiella has been stimulated by M.A. CONRAD (Geneva, Switzerland). Helpful comments of reviewers M.A. CONRAD (Gene- va, Switzerland), B. GRANIER (Brest) and B. SOKAČ (Zagreb) are kindly acknowledged. 5. REFERENCES BARATTOLO, F. (1991): Mesozoic and Cenozoic marine benthic calcareous algae.– In: RIDING, R. (ed.): Calca- reous Algae and Stromatolites. Springer–Verlag, Berlin, 504–540. BARATTOLO, F. (1998): Dasycladacean green algae and microproblematica of the uppermost Cretaceous–Pale- ocene in the Karst Area (NE Italy and Slovenia).– Dela – Opera SAZU 4. razr., 34/2, 65–127. BARATTOLO, F. (2002): Late Cretaceous–Paleogene Dasy- cladaleans and the K/T boundary problem. – In: BUCUR, I.I. & FILIPESCU, S. 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Revised manuscript accepted November 11, 2005. 112 Geologia Croatica 58/2 PLATE I Neogyroporella? gawlicki n.sp. from the Late Tithonian–Early Berriasian of the Northern Calcareous Alps 1 Various oblique sections, Dietrichshorn locality. 2 Oblique section together with the benthic foraminifera Anchispirocyclina lusitanica (EGGER), Dietrich- shorn locality. 3 Transverse section (left) and various algal fragments, Litzelkogel locality, thin-section A–190. 4 Oblique section, Dietrichshorn locality. 5 Oblique–tangential section, holotype, Litzelkogel locality, thin-section A–190. 6 Detail from Fig. 5 showing the close and alternating laterals, Litzelkogel locality, thin-section A–190. 7 Oblique section, Dietrichshorn locality. 8 Fragmentary transverse section, Dietrichshorn locality. 9 Transverse section, Dietrichshorn locality. 113Schlagintweit PLATE I 1 2 3 4 6 5 7 8 9 114 Geologia Croatica 58/2 PLATE II Neogyroporella? gawlicki n.sp. and other Dasycladales from the Late Tithonian–Early Berriasian of the Northern Calcareous Alps 1 Neogyroporella? gawlicki n.sp., transverse section, Dietrichshorn locality. 2 Neogyroporella? gawlicki n.sp., tangential section, Litzelkogel locality, thin-section A–190. 3 Unknown dasycladale with a calcareous envelope consisting of radial-fibrous calcite, Dietrichshorn loca- lity. 4 Neogyroporella? gawlicki n.sp., Detail from Pl. I, Fig. 8, showing elongated vesiculiferous branch morpho- logy, Dietrichshorn locality. 5 Branch morphology of Gyroporella ampleforata PIA (from PIA, 1927). 6 Clypeina aff. parasolkani FARINACCI & RADOIČIĆ, transverse section, Dietrichshorn locality. 7 Zergabriella embergeri (BOUROULLEC & DELOFFRE), Dietrichshorn locality. 8, 10, 12 Different sections of Deloffrella quercifoliipora GRANIER & MICHAUD, Dietrichshorn locality. 9 Cylindroporella? sp., Dietrichshorn locality. 11 Clypeina aff. isabellae MASSE, BUCUR & VERON, Dietrichshorn locality. 13 Clypeina catinula CAROZZI, longitudinal section showing the swelling of the main axis at the vertical plane, Dietrichshorn locality. 115Schlagintweit PLATE II 1 2 3 6 7 4 5 8 9 10 11 12 13 116 Geologia Croatica 58/2 PLATE III Neogyroporella? gawlicki n.sp. and other Dasycladales and benthic foraminifera from the Late Tithonian–Early Berriasian of the Northern Calcareous Alps 1 Salpingoporella cf. istriana (GUŠIĆ), longitudinal section, Litzelkogel locality, thin-section Ber–17a. 2–5 Neogyroporella? gawlicki n.sp. 2–4: oblique sections, thin-sections A–149 and A–182, 5: longitudinal– tangential section, thin-section A–191 (all from Litzelkogel locality). 6 Montenegrella florifera BERNIER, Litzelkogel locality, thin-section A–189–2. 7 Protopeneroplis ultragranulata (GORBATCHIK), axial section, Litzelkogel locality, thin-section Ber– 100–2. 8–9 Kastamonina abanica SIREL, transverse section (8) and subaxial section (9), Dietrichshorn locality. 10 Chinianella? sp., oblique transverse section, Litzelkogel locality, thin-section A–191. 117Schlagintweit PLATE III 1 2 3 4 5 8 6 7 9 10 118 Geologia Croatica 58/2