Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 1 of 30 RESEARCH ARTICLE A multidisciplinary biostratigraphic framework for the Lower to Middle Miocene of the Norwegian North Sea – the siliceous succession of the Valhall–Hod area Emma Sheldon1* , Karen Dybkjær1 , Erik Skovbjerg Rasmussen2 , Mimmi Oksman3 1Department of Geo-energy and Storage, Geological Survey of Denmark and Greenland (GEUS), Copenhagen, Denmark; 2Department of Geophysics and Sedimentary Basins, Geological Survey of Denmark and Greenland (GEUS), Copenhagen, Denmark; 3Department of Glaciology and Climate, Geological Survey of Denmark and Greenland (GEUS), Copenhagen, Denmark Abstract A new multidisciplinary biostratigraphic framework, combining dinoflagellate cysts, microfossils, calcareous nannofossils, diatoms and silicoflagellates, is established for the Early to Middle Mio- cene deep marine clay and siliceous ooze in the southern Norwegian sector of the North Sea, based on core samples from the Valhall and Hod hydrocarbon fields. The framework was successfully tested on the equivalent chronostratigraphic level of several wells based on ditch cutting samples. New biostratigraphic events for the Danish and Norwegian North Sea resulting from this study are successfully used to correlate between the Valhall and Hod areas and supplement published zonation schemes. To our knowledge, this is the first time that diatoms and silicoflagellates from the fine fraction of microfossil samples have been used as correlation tools in the North Sea Basin. Dating of the siliceous/diatomite-rich interval results in a high-resolution (5–15 m intervals) bio- stratigraphic subdivision. The successful application of the new framework across the Valhall and Hod areas implies that it could also be useful in a more regional context. The new biostratigra- phy enables the correlation of the lithostratigraphic units recently defined for the Danish offshore Neogene succession to the study area and the correlation of the sequence stratigraphic surfaces defined for the Danish sector to the southern Norwegian sector. *Correspondence: es@geus.dk Received: 21 Aug 2024 Revised: 07 May 2025 Accepted: 11 Jun 2025 Published: 19 Dec 2025 Keywords: biostratigraphy, diatomite, Miocene, North Sea, Norway Abbreviations: FO: first occurence FSST: falling stage systems tract GEUS: Geological Survey of Denmark and Greenland HST: highstand systems tract LO: last occurence LST: lowstand systems tract TST: transgressive systems tract MMCT: Middle Miocene Climatic Transition MCO: Miocene Climatic Optimum PRZ: partial range zone GEUS Bulletin (eISSN: 2597-2154) is an open access, peer-reviewed journal published by the Geological Survey of Denmark and Greenland (GEUS). This article is distributed under a CC-BY 4.0 licence, permitting free redistribution, and reproduction for any purpose, even commercial, provided proper citation of the original work. Author(s) retain copyright. Edited by: Mette Olivarius (GEUS, Denmark) Reviewed by: Haydon Bailey (Independent Researcher, UK), Erik Anthonissen (Equinor ASA, Norway) Funding: See page 26 Competing interests: See page 26 Additional files: See page 26 1. Introduction A new, multidisciplinary biostratigraphic study of the Lower and Middle Mio- cene sections of six wells from the Valhall and Hod fields is presented. These Upper Cretaceous – Danian chalk hydrocarbon fields are located on salt structures in the southernmost part of the Norwegian North Sea (Fig. 1). The Miocene succession above the chalk comprises deep-marine clay with a vari- able content of siliceous ooze. The silica content reaches 50% in some inter- vals, which are often referred to as diatomite. The focus is on the siliceous ooze or diatomite in some areas, such as the Valhall–Hod area, because of its hydrocarbon reservoir potential and because its geomechanical properties are critical in connection with well abandonment. Two of the six studied well sections, 2/11–12S (Hod Field) and 2/8–G10A (Valhall Field), were cored through the Lower and Middle Miocene succes- sions, providing an exceptional and continuous record, unique in the North Sea area. The other four studied wells, 2/8–N4, 2/8–V6, 2/8–8 (all from Valhall Field) and 2/11–1 (in the saddle between the Valhall and Hod fields), were not cored. The locations of the wells are shown in Fig. 2. The purpose of this study is to establish a high-resolution multidisciplinary biostratigraphy for the siliceous or diatomite-rich succession represented by the unique Hod and Valhall cores. The resulting biostratigraphic framework is then applied to the four non-cored wells using fossil assemblages from ditch cutting samples. https://doi.org/10.34194/5k9dv133 https://orcid.org/0000-0003-4353-8241 https://orcid.org/0000-0002-8420-3379 https://orcid.org/0000-0001-8603-8429 https://orcid.org/0000-0002-8386-516X mailto:es@geus.dk https://creativecommons.org/licenses/by/4.0/deed.en Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 2 of 30 GEUSBULLETIN.ORG The study was performed combining five biostrati- graphic groups: dinoflagellate cysts (dinocysts), micro- fossils (primarily foraminifera but also including large diatoms and Bolboforma), small fraction diatoms, silico- flagellates and calcareous nannofossils, mostly on the same series of closely spaced sediment samples. This is the first time, to our knowledge, that a detailed North Sea Miocene biostratigraphic study includes small sili- ceous diatoms and silicoflagellates. 2. Geological setting and palaeoclimate The Oligocene–Miocene transition was character- ised by inversion tectonism resulting in shallower waters in the north-eastern part of the North Sea Basin (Ziegler 1990; Rasmussen 2009, 2013; Knox et  al. 2010). The initial uplift of the Southern Scandes (Fig. 1) re-exposed the present-day Norway and cen- tral Sweden, which formed a low relief landscape at the end of the Oligocene (Thyberg et al. 2000; Løseth & Henriksen 2005; Gabrielsen et al. 2009). The uplift of the hinterland in the Early Miocene and the shallow- ing of the north-eastern North Sea Basin resulted in progradation of large delta systems from Scandinavia (Rasmussen et al. 2010; Fig. 1). In the northern North Sea Basin, delta progradation occurred from the west, the Shetland Platform, coincident with the eastern system (Eidvin et al. 2014a). The southern North Sea Basin was dominated by a coastal plain, and swamp environments formed the margin of a low-relief cen- tral European landscape, which was separated from the Alps by a foreland basin (Fig. 1). During the Middle Miocene, a major transgression occurred, and the deltas established during the Early Miocene were flooded. The flooding commenced coin- cident with a global climatic deterioration (Zachos et al. 2001) and the initiation of a new tectonic regime in the North Atlantic. Huge inversion structures were formed off west Norway, and the main phase of uplift of the Sole Pit structure in the western part of the North Sea Basin took place (Knox et  al. 2010; Løseth et  al. 2017 and references therein). Iceland also formed at this time (Rasmussen et  al. 2008), so branches of the Ice- landic Plume (Schoonman et  al. 2017) may also have reshaped the landscape around the northern North Sea Basin. The late Early Miocene to Middle Miocene was also an important phase in the uplift of the Carpathian Fig. 2 Depth map to the top Miocene of the Valhall and Hod structures and locations of the six studied wells (yellow dots). Wells circled in red are cored. Credit: Aker BP. Valhall Field 2/8-N4 2/8-8 2/8-G10A –1325 –1350 –1375 –1400 –1425 –1450 –1475 –1500 –1525 –1550 –1575 –1600 m 2/8-V6 Hod Field 2/11-1 2/11-12S 5000 m Fig. 1 Palaeogeography of the North Sea area in the Early Miocene. The red dot indicates the location of the study area. Arrows indicate sedi- ment influx. The Valhall/Hod area was located in the central part of the basin. North Sea sectors are as follows: D: Germany. DK: Denmark. N: Norway. NL: Netherlands. UK: United Kingdom. Modified from Rasmus- sen et al. (2008). 0°E 10°E 50°N 60°N 100 km Shetland Platform N UK DK D NL Valhall- Hod area Southern Scandes https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 3 of 30 GEUSBULLETIN.ORG Mountains in Central Europe (Oszczypko 2006). The establishment of the new tectonic regime resulted in accelerated subsidence of the North Sea Basin, and the deposition of marine mud dominated the Middle Mio- cene (Koch 1989; Rasmussen 2004a, 2004b; Rasmussen & Dybkjær 2014). Our study area was situated in a fully marine, outer shelf to upper bathyal, basin floor setting in a semi-closed basin with long distances to coastlines. Deposition was characterised by hemipelagic sedimen- tation in water depths of between 500 and 1000 m. During the Late Miocene, continued growth of the Carpathian Mountains, uplift of the Alpine Foreland Basin and formation of the Jura Mountains resulted in the formation of a massive new source area in Central Europe (Kuhlemann 2007; Fig. 1). In the Late Miocene, huge, braided river systems supplied the south-eastern North Sea Basin for the first time (Knox et al. 2010). The new central European river system evolved into the so-called Eridanos Delta system (Biljsma 1981; Overeem et al. 2001; Rasmussen & Dybkjær 2014), which began to fill the eastern North Sea Basin. Delta systems sourced from Scandinavia also began to prograde into the north-eastern part of the basin during the latest Late Miocene and reached the Central Graben area during the Messinian Stage when they coalesced with the Eridanos Delta. These delta sys- tems correlate with the Nordland Group of the Norwe- gian part of the North Sea (Fig. 3). The climate in the study area was warm-temperate to sub-tropical and humid during the Early and Middle Miocene (Utescher et  al. 2009; Larsson et  al. 2011; Sli- winska et al. 2024). Studies on the Sdr. Vium borehole, Jylland, Denmark, by Larsson et al. (2011), Herbert et al. (2020) and Sliwinska et al. (2024) indicate mean annual temperatures of around 17–18.5°C on land, mean annual precipitation of c. 750–1750 mm/yr and sea sur- face temperatures of 23–28°C, although with some fluc- tuations during the Miocene Climatic Optimum (MCO) (c.17–13 Ma; e.g. Larsson et al. 2011; Herbert et al. 2020; Sliwinska et al. 2024). At the end of the Middle Miocene, the global climate deteriorated, a period known as the Middle Miocene Climatic Transition (MMCT). In the Danish and German areas, a decrease in annual temperatures during the Serravalian Stage has been recognised (Utescher et al. 2009; Herbert et al. 2020, Sliwinska et al. 2024). Marked climatic deterioration in the Messinian Stage at the close of the Miocene Epoch resulted in the expansion of ice caps on Antarctica and probably also  in parts of the northern hemisphere (Utescher et al. 2009). 3. Lithostratigraphy The Neogene deposits in the North Sea area include marginal, fluvio-deltaic deposits, shoreface and offshore shelf deposits and basinal deep water hemipe- lagic and gravity-flow deposits. Silica- or diatomite-rich deposits are found locally in basinal areas (including the study area) in the Lower and Middle Miocene parts of the succession. A new lithostratigraphic subdivision of the Neogene succession in the Danish North Sea sector is presented in Rasmussen et al. (in press; mod- ified version presented here, Fig. 3). In this study, we correlate the new offshore Danish lithostratigraphy to the Norwegian sector of the North Sea. This litho- stratigraphy includes the new Lower Miocene Dany Formation, which comprises muddy and silty deposits, and the new Middle Miocene Nora Formation, which is defined based on its high content of silica or diatomite. The well sections presented in this study include the Dany, Nora and Hodde formations following the new subdivision. According to the Norwegian lithostratig- raphy for the Late Paleocene to Neogene, the studied Lower and Middle Miocene succession is subdivided into the Hordaland Group (Lark Formation) and the Nordland Group (Eidvin et al. 2022; Fig. 3). In this study, lithostratigraphic information including diatomite con- tent is available for the two cored wells 2/8–G10A and 2/11–12S, and thus, it has been possible to subdivide the Miocene succession into the lithostratigraphic units defined in the Danish sector. An attempt at a similar subdivision in the non-cored wells is made, based on gamma log responses. 4. Sequence stratigraphic framework In the Danish and southern Norwegian sectors, eight depositional sequence boundaries are found within the Miocene succession (Rasmussen et al. 1996, Rasmussen 2004b, 2017). The boundaries are defined on a combi- nation of studies of outcrop sections onshore Denmark, borehole logs and cores and seismic data, including high-resolution shallow seismic, multichannel, and for the Central Graben area, 3D seismic data (Rasmussen 1996; Rasmussen 2004b, 2017, Dybkjær et al. 2021). The eight sequence boundaries confine seven fully devel- oped sequences, named B, C, D1, D2, E, F1 and F2 (Ras- mussen 2004b, 2017). The two lowermost sequences, B and C, and the F2 sequence include all four systems tracts (LST: lowstand systems tract, TST: transgressive systems tract, HST: highstand systems tract and FSST: falling stage systems tract), whereas the upper Lower Miocene to lower Upper Miocene sequences only have a two-fold subdivision (TST and HST). The latter was due to increased subsidence of the North Sea Basin during the middle part of the Miocene outpacing eustatic sea-level fall (Rasmussen 2004b, 2017). The seismic surfaces can be seen for the 2/8–G10A and 2/11–125 wells in Figs 5 and 6 and for all wells in the Supplementary Files S1–S12. https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 4 of 30 GEUSBULLETIN.ORG Fi g. 3  T he n ew li th os tr at ig ra ph ic s ub di vi si on fo r th e N eo ge ne s uc ce ss io n in th e D an is h se ct or o f t he N or th S ea (m od ifi ed fr om R as m us se n et  a l., in p re ss ) w ith a d as he d re d bo x sh ow in g th e st ud ie d in te rv al . N ot e th e pr es en ce o f d ia to m ite in th e m id dl e M io ce ne , r ef er re d to a s th e N or a Fo rm at io n (s ol id r ed b ox ), an d so m e m in or o cc ur re nc es in th e Lo w er M io ce ne s uc ce ss io n w ith in th e D an y Fo rm at io n (r ed a rr ow s) . T he in te rv al s co ve re d by th e tw o co re d w el l s ec tio ns (2 /1 1– 12 S an d 2/ 8– G 10 A) a re s ho w n. T he ti m es ca le o f R affi e t a l. (2 02 0) is u se d in th is fi gu re . A qu it an ia n Bu rd ig al ia n La ng hi an Se rr av al lia n To rt on ia n M es si ni an Za nc le an Pi ac en zi an G el as ia n C al ab ri an NeogeneQuaternary Miocene MiddleUpper Lower Plio- cene Plei- sto- cene H ol oc en e 0 5 10 15 20 Måde Group Ribe Group Lark Formation Nordland Group Hordaland Group G ra m F m Ø rn hø j F m H od de F m A rn um F m Ba st ru p Fm V ej le F jo rd F m Bi llu nd F m Kl in ti ng ho ve d Fm Fl oo dp la in s ed im en t D el ta s an d O �s ho re d ia to m it e Sw am p Fl uv ia l s an d/ gr av el F2 F1 E D 2 D 1 C B A qu i10 0 A qu i2 0 0 Bu rd 10 0 Bu rd 20 0 La ng 10 0 To rt 10 0 To rt 20 0 M es s1 0 0 Sl op e sa nd O �s ho re g ra vi ty -� ow s an d O �s ho re m ud O �s ho re s an d D ee p m ar in e m ud Sh or ef ac e sa nd G ra m F m Em m a Fm El in F m Emma Fm Ø rn hø j F m H od de F m G ra m F m Ø rn hø j F m G 10 A 12 S H od de F m A rn um F m O dd er up F m N or a Fm O dd er up F m Kl in ti ng ho ve d Fm D an y Fm D an y Fm M ar bæ k Fm Li lle Jo hn M b M ar bæ k Fm V ag n Fm H . o bs cu ra (H .o .) S. ar m ag ed do ne ns is (S .a .) M . c ho an op ho ru m (M .c .) B. p lio ce ni cu m (B .p .) I. m ul tip le xu m (I. m .) G . v er ri cu la (G .v .) A . a nd al ou sie ns e (A .a .) A . u m br ac ul a (A .u .) U . a qu ae du ct um (U .a .) L. tr un ca tu m (L .t. ) C . c an th ar el lu s ( C .c .) E. in sig ne (E .i. ) C . a ub ry ae (C .a u. ) C . g al ea (C .g .) S. ha m ul at um (S .h .) T. pe la gi ca (T .p .) C . a m ic ul um (C .a m .) H om ot ry bl iu m sp p. (H .) O dd er up F m O dd er up F m V ag n Fm Lu na F m A ge W E (M a) Period Ep oc h A ge /S ta ge N or w eg ia n Li th o- st ra ti gr ap hy C en tr al G ra be n D in oc ys t zo na ti on (D yb kj æ r & Pi as ec ki 2 0 10 ) Ri ng kø bi ng – F yn H ig h N or w eg ia n - D an is h Ba si n/ on sh or e D en m ar k Se qu en ce s (R as m us se n 20 17 ) Se qu en ce s (In fo rm al ) https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 5 of 30 GEUSBULLETIN.ORG 5. Absolute dating Absolute dating by palaeomagnetic stratigraphy, radiometric dating or Sr isotope stratigraphy of the Miocene succession in the North Sea Basin is rare. Deeper levels were usually targeted for hydrocarbon exploration, while the younger ‘overburden’ was tra- ditionally considered uninteresting and therefore only rarely cored. Therefore, the absolute ages of dinocyst and micro- fossil events within the Miocene succession in the North Sea Basin as shown by Powell (1992), Munster- man & Brinkhuis (2004), Louwye et  al. (2007), Dybk- jær & Piasecki (2010), Köthe (2012), King (1989, 2016), Munsterman et al. (2019) and Dybkjær et al. (2019) are usually based on data from areas outside the North Sea Basin, where these events are found in wells where absolute dating has been carried out (e.g. Haq et al. 1987; De Verteuil & Norris 1996; De Verteuil 1997; Williams et al. 2004 and references therein). Also, cor- relation with other microfossil groups (e.g. nannofos- sils) and the global sea-level changes have been used to date the dinocyst and microfossil events recorded in the North Sea Basin (e.g. Dybkjær & Piasecki 2010; King 2016; Munsterman et al. 2019). However, the absolute dating of specific events varies from one reference to another, due to the diachronicity of first and last appearance datums and uncertainties of the datings. The most comprehensive Sr-isotope study of the Miocene North Sea Basin is that of Eidvin et al. (2014b) from onshore Denmark. The results of that study gener- ally supported the ages of the dinocyst zones of Dybkjær & Piasecki (2010) in the Early Miocene but also docu- mented the uncertainty of using Sr-isotopes for dating this stratigraphic level, especially the late Middle to Late Miocene part of the succession. Comparing the dinocyst event and zonation scheme of Dybkjær & Piasecki (2010, their fig. 6) with that of King (2016, their fig. 18) clearly reflects this uncertainty in chronostratigraphic correlation. Similarly, correlation between dinocyst and microfossil events and zones also differs in the two publications (e.g. Dybkjær & Piasecki 2010, their fig. 6; King 2016, their fig. 21). Correlation between Miocene nannofossil and microfossil zones is also problematic, as seen in and explained by King (2016), compare their figs. 21 and 27. Absolute dating has not been carried out on the studied wells. Due to this absence of an absolute age model, the events and biozonations for the five studied microfossil groups are presented against established zonations (Fig. 4), sample depth, sequence boundaries, lithostratigraphy and the gamma log, instead of against a timescale (Ma), see Figs 5, 6 and Supplementary Files S1–S12. 6. Previous studies The majority of biostratigraphic work that has been car- ried out on the Neogene succession of the North Sea area over the past 50 years is a direct result of extensive hydrocarbon exploration at deeper levels. Palynological studies on the Miocene succes- sion in the North Sea include Piasecki (1980), Strauss & Lund (1992), Powell (1992), Head (1996), Louwye et  al. (1999), Louwye (2002), Dybkjær & Rasmussen (2000, 2007), Strauss et  al. (2001), De Schepper et  al. (2004, 2009), Dybkjær (2004a, 2004b), Munsterman & Brinkhuis (2004), Schiøler (2005), Köthe & Piesker (2007), Louwye et al. (2007), Louwye & Laga (2008), Louwye & De Schepper (2010), Dybkjær & Piasecki (2010), Dybkjær et al. (2012), Köthe (2012), Eidvin et al. (2014a, 2014b), Śliwinska et al. (2014), King (2016), De Schepper & Man- gerud (2017), Grøsfjeld et al. (2019), Dybkjær et al. (2021) & Sliwinska et al. (2024). Microfossil studies of the Miocene of the North Sea area are also numerous (Von Daniels & Spiegler 1977; Doppert et al. 1979; Doppert 1980; Spiegler & Von Dan- iels 1991; Spiegler 1999; King 1983, 1989, 2016; Grad- stein et al. 1988; Gradstein & Backstrom 1996; Laursen & Kristoffersen 1999; Eidvin et al. 1999; Kaminski & Grad- stein 2005; Rundberg & Eidvin 2005; Eidvin & Rundberg 2007; Anthonissen 2012; Fox et al. 2018). Published Neogene calcareous nannofossil stud- ies for the North Sea area are not common due to the siliciclastic nature of much of the Neogene section and the successful application of high-resolution studies of other biostratigraphic disciplines. Neogene nannofos- sil studies of the broader North Atlantic region include Müller (1976), Steinmetz (1979), Gartner (1992), De Kaenel et  al. (2017), Boesiger et  al. (2017) and Bergen et  al. (2017). The global nannofossil ‘NN’ zonation of Martini (1971) is still widely used and is correlated with other nannofossil zonations in Young et al. (1994) and Young (1998). For the North Sea area, the use of diatoms for Palae- ogene and Neogene biostratigraphy was studied by Mit- lehner (2019). Several biostratigraphic studies of Upper Oligocene and Lower Miocene North Sea successions using large, pyritized diatoms in conjunction with other microfossils have also been published. King (1983, 2016) focused on the Cainozoic micropalaeontological biostra- tigraphy of the North Sea and adjacent areas. Laursen and Kristoffersen (1999) studied the Miocene succes- sions of onshore Denmark using foraminifera, and Dybk- jær et al. (2012) concentrated on the Oligocene–Miocene boundary in the eastern North Sea Basin using dino- cyst stratigraphy, micropalaeontology and δ13C-isotope data. Eidvin and Rundberg (2001) focused on the Late Cainozoic stratigraphy of the northern North Sea, and https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 6 of 30 GEUSBULLETIN.ORG Anthonissen (2012) compiled an integrated Miocene biostratigraphy for the northeastern North Atlantic. Small siliceous diatoms are not generally used in the North Sea area for routine biostratigraphy as their minute size results in them being washed through standard sieves, and their delicate structures are easily destroyed by the harsh preparation techniques used in industry. A diatom zonation scheme for the North Sea does not exist. However, studies with potential use in biostratigraphy in the Neogene of the North Sea area include Eidvin et al. (1998), Thyberg et al. (1999) and Sheldon et al. (2018). Siliceous diatom stud- ies from the wider high northern latitudes that are useful for correlation with this study include those of Schrader & Fenner (1976), Koç & Scherer (1996) and Suto (2006) from the Norwegian and Iceland Seas, Dzinoridze et al. (1979) from the Norwegian Basin and Baldauf (1985) from the Rockall Plateau. Silicoflagellates only comprise a small percentage of the siliceous component of marine sediments and therefore have limited biostratigraphic use. A silicoflagel- late zonation scheme for the North Sea does not exist. Martini & Muller (1976), Locker & Martini (1989), Ciesiel- ski et al. (1989) and Amigo (1999) investigated Miocene successions in the Norwegian-Greenland Sea and the Ice- land-Rockall Plateau areas to the north of the study area. 7. Zonation schemes in this study The biostratigraphic zonations used in this study are the North Sea dinocyst zonation of Dybkjær & Piasecki (2010), the North Sea microfossil zonations of King (1989, 2016), the global calcareous nannoplankton zonation of Martini (1971), the Norwegian Sea diatom zonation of Schrader & Fenner (1976) and the Norwegian Sea sili- coflagellate zonation of Locker & Martini (1989), Fig. 4. In their study of the Neogene of the Iceland Sea, Koç & Scherer (1996) revised the biostratigraphy of Schrader & Fenner (1976). In this study, however, we revert to the zonation of Schrader & Fenner (1976) due to the similar- ity of diatom assemblages therein with those from the Valhall–Hod area. The zonation schemes follow a chro- nostratigraphy that was current when the zonation was published. The North Sea dinocyst zonation of Dybkjær & Piasecki (2010) follows the chronostratigraphy of Lou- rens et al. (2004). The North Sea microfossil zonations of King (1989, 2016) follow the chronostratigraphy of Hilgen et al. (2012) in King (2016). The global calcareous nannoplankton zonation of Martini (1971) is correlated with the chronostratigraphy of Raffi et  al. (2020). The Norwegian Sea diatom zonation of Schrader & Fenner (1976) follows the chronostratigraphy of Berggren (1972), and the Norwegian Sea silicoflagellate zonation of Locker & Martini (1989) follows the chronostratigra- phy of Berggren et al. (1985). It is beyond the scope of this paper to attempt an up-to-date chronostratigraphic correlation of the five zonation schemes. Dinocyst tax- onomy follows the ‘Lentin & Williams Index’ (Williams et al. 2017). Microfossil taxonomy follows that used in King (1989, 2016) and Young et  al. (2024b). Nannofos- sil taxonomy is based on Young et al. (2024a). Diatom taxonomy follows that of Schrader & Fenner (1976) and Barron (1985), and silicoflagellate taxonomy follows that of Perch-Nielsen (1985) and Locker & Martini (1989). 8. Materials and methods Cores from the 2/11–12S (Hod Field) and 2/8–G10A (Val- hall Field) wells were sampled with a spacing of approx- imately 4 to 5 m. Ditch cutting samples were taken from the core gap in 2/8–G10A. For the non-cored wells 2/8– N4, 2/8–V6, 2/8–8 (all Valhall Field) and 2/11–1 (in the saddle between the Valhall and Hod fields), ditch cutting samples were taken every 10 m (Table 1). The position of the analysed samples in each well is shown in Figs 5, 6 and Supplementary Files S1–S12. Most of the samples were analysed for palynology (dinocysts), microfossils (large fraction) and siliceous microfossils (small fraction: diatoms and silicoflagel- lates). Only around ten samples per well were selected from minor calcareous-rich intervals and analysed for nannofossils as they are not routinely used for bio- stratigraphy in the Miocene of the North Sea. 8.1. Preparation methods The sample preparation methods for each biostrati- graphic discipline are described below. All sediment sam- ples were processed in the Stratigraphic Laboratory at the Geological Survey of Denmark and Greenland (GEUS). 8.1.1. Palynology Approximately 20 g of sample was dried and crushed until all particles were <2 mm in size. Dissolution of Table 1 Number of samples for each discipline for each well. Well 2/11-12S Well 2/8-G10A Well 2/8-V6 Well 2/8-N4 Well 2/8-8 Well 2/11-1 Palynology (dinocysts): 40 40 CO, 15 DCS 27 25 49 32 Microfossils: 47 39 CO, 13 DCS 27 25 51 34 Diatoms & Silicoflagellates: 36 39 CO, 13 DCS 27 25 51 34 Nannofossils: 10 10 CO 10 11 11 11 CO: core samples. DCS: ditch cutting samples. The samples from 2/11-12S are all from core material. Those from 2/8-G10A are mostly from core material and the remaining well samples are ditch cutting samples. https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 7 of 30 GEUSBULLETIN.ORG Fi g. 4  B io st ra tig ra ph ic z on at io ns u se d in th is s tu dy . D in oc ys t z on at io n of D yb kj æ r & P ia se ck i ( 20 10 ), na nn of os si l z on at io n of M ar tin i ( 19 71 ), m ic ro fo ss il zo na tio ns o f K in g (1 98 9, 2 01 6) , d ia to m z on at io n of S ch ra de r & Fe nn er (1 97 6) & th e si lic ofl ag el la te z on at io n of L oc ke r & M ar tin i ( 19 89 ). Re fe r to in di vi du al r ef er en ce s fo r tim es ca le s ap pl ie d. Ep oc h St ag e D in o� ag el la te s Zo ne A ge (M a) A ge (M a) A ge (M a) A ge (M a) A ge (M a) 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 11 12 13 14 15 16 17 18 19 20 21 22 23 10 11 12 13 14 15 16 17 18 19 20 21 22 23 13 14 15 16 17 18 19 20 21 22 9 10 11 12 13 14 15 16 17 18 19 20 21 Ep oc h St ag e N an no fo ss ils Zo ne Ep oc h St ag e M ic ro fo ss ils N SB Z on e N S Zo ne Ep oc h D ia to m s Pa rt ia l R an ge Z on e Ep oc hSi lic o� ag el la te s Zo ne Miocene Serravalian Langhian Burdigalian AquitanianTortonian Miocene Miocene Serravalian Langhian Burdigalian AquitanianTortonian N S4 0 N S3 9 N S3 8 N S3 7 N S3 6 N S3 5 bb cc aaa N S3 4 Th al as sio sir a fr ag a N itz ec hi a m al ei nt er pr et ar ia C os oi no di so us v ig ila ns Rh iz os en ia n or w eg ic a Sy ne dr a jo us ea na Ps eu do di m er og ra m m a el eg an s C os ci no di sc us p lic at us D en tic ul a hy al in a U pp er C . t ria ca nt ha H ia tu s P. c irc ul us a pi cu la ta Lo w er C . t ria ca nt ha N . n av ic ul a Rh iz os ol en ia b ul bo sa 9101112 b 12 c 12 a N S3 3 N N 8 N N 7 N N 6 N N 5 N N 4 N N 3 N N 2 N N 1 Serravalian Langhian Burdigalian AquitanianTortonian Early MioceneMiddle Miocene Early MioceneMiddle MioceneLate Miocene A . u m br ac ul a G . v er ric ul a A . a nd al ou sie ns e H . o bs cu ra U . a qu ae du ct um L. tr un ca tu m C . a ub ry ae E. in sig ne C . c an th ar el lu s S. h am ul at um T. p el ag ic a C . a m ic ul um H om ot ry bl iu m s pp . C . g al ea https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 8 of 30 GEUSBULLETIN.ORG Fi g. 5  C ap tio n on n ex t p ag e. •• A bu nd an t • C om m on ° C on si st en t W el l n am e: 2 /8 - G 10 A - Fi g 5A 14 60 14 50 14 70 14 80 14 90 15 0 0 15 10 15 20 15 30 15 40 15 50 15 60 15 70 15 80 15 90 Gamma log Cores Li th o- st ra ti gr ap hy Chrono- stratigraphy D in o� ag el la te c ys t e ve nt s Samples North Sea Cenozoic Microfossils M ic ro fo ss il ev en ts SB E m fs D 2 m rs E Nordland Gp. Måde Group Lower LanghianUpper Langhian Middle Miocene Dany Fm. Nora Fm. Ribe Group Ørnhøj Fm. Hodde Fm. O�shore Denmark O�shore Norway Hordaland Group Lark Fm. * ** ** * ? M .M . Se r. A . a nd . ? ? ? ? ? Labyrinthodinium truncatumUnipontidinium aquaeductum Nannofossil Zone Nannofossil samples N an no fo ss il ev en ts Diatom P.R.Z. D ia to m e ve nt s Silico�agellate Zone Si lic o� ag el la te e ve nt s S. a �. k itt on ia nu s D . h us te dt ii, O . g em m at a, Sy ne dr a sp p. , S . g ru no w ii, D en tic ul op sis s pp . • h ig h ab un d. & d iv er si ty d ia to m a ss em bl ag es A . i ng en s, D . s m ith ii P. e lo ng at a, D . h ya lin a C . b ih ar en sis T. n itz sc hi od es •• R. w ic om ic oe ns is D en tic ul op sis s pp . • , D . h us te dt ii • A . i ng en s ° S. h or rid us •• S. g ru no w ii °, Sy ne dr a sp p. (i n� ux ) D . c ru x T. fragaRhizosolenia bulbosaDenticulopsis hyalina lower Corbisema triacanthaupper Corbisema triacantha R. m io ce ni ca B. d io do n B. d io do n, O . s pe cu lu m C . t ria ca nt ha C . b ih ar en sis C . p ep lu m North Sea Cenozoic Microfossils C . a ng ul io � ci na lis T. q ua dr ilo ba tu s B. p la ty re tic ul at a A . g . s ta es ch ei , L. si nu os um , E. in �a tu m , C . s ub co ni cu s, L. p er eg rin a E. in �a tu m T. tr ilo bu s, C . d . p ee le ns is, G . e ol ab ia cr as sa ta , P. n an a, C . c ip er oe ns is Undi� Undi�erentiated NSB12a NSB11 Undi�erentiated NS36c NS36a-b H . s ci ss ur a (o cc ), D . e xi lis (o cc ), H . w al tr an s ( oc c) , D . d isc iss us (o cc ) D . c au li� or is, H . b ip un ct a, C . � or id an us , C . p el ag ic us , H . c ar te ri D . d ru gi i ( oc c) , D . d e� an dr ei (o cc ) H . b ip un ct a S. n eo ab ie s U . a qu a. , I m p. s p. A W & K 19 96 •, P. m io ca en ic um A . a nd al ou sie ns is, O . t eg ill at um Im pa gi di ni um s p. A W re nn & Ko ki no s 19 96 •, I. la cr ym os a, O . e iri ki an um H . t ec ta ta • H . t ec ta ta U . a qu ae du ct um C . a ub ry ae I. ta bu la ta D . h ya lin a T. fr ag a N N 4 N N 4- 5 Sequence stratigraphy Dino�agellate cyst Zone Ki ng 19 89 Ki ng 20 16 U nd i� C . p as ., C . p ou l., C . p la c. (o cc ) A . a nd al ou sie ns is https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 9 of 30 GEUSBULLETIN.ORG Fi g. 5  ( Co nt in ue d) S um m ar y di ag ra m fo r th e 2/ 8– G 10 A w el l w ith b io st ra tig ra ph ic z on es , p re vi ou sl y pu bl is he d ev en ts (b la ck t ex t) a nd n ew e ve nt s (r ed t ex t) , f or fu ll fo ss il na m es s ee S up pl em en ta ry F ile S 1. L ith os tr a- tig ra ph y fo r th e N or w eg ia n Se ct or fr om E id vi n et  a l. (2 02 2) , o ns ho re D en m ar k fr om R as m us se n et  a l. (2 01 0) a nd o ff sh or e D en m ar k fr om R as m us se n et  a l. (in p re ss ). Se qu en ce s tr at ig ra ph y fr om R as m us se n (2 00 4a ; 20 17 ) a nd D yb kj æ r et  a l. (2 02 1) . G am m a lo g c/ o Ak er B P. C or ed in te rv al s ar e in di ca te d. T he m ai n sa m pl e co lu m n in di ca te s sa m pl es fo r di no cy st s, m ic ro fo ss ils , d ia to m s an d si lic ofl ag el la te s. N an no fo ss il sa m pl es a re in di ca te d se pa ra te ly . I n th e ‘C or es ’ c ol um n, c or e sa m pl es a re m ar ke d w ith a d as h- do t sy m bo l, an d di tc h cu tt in g sa m pl es a re m ar ke d w ith a d as h. C hr on os tr at ig ra ph y of R affi e t a l. (2 02 0) , d in oc ys t z on at io n of D yb kj æ r an d Pi as ec ki (2 01 0) , m ic ro fo ss il zo na tio ns o f K in g (1 98 9; 2 01 6) , d ia to m z on at io n of S ch ra de r an d Fe nn er (1 97 6) , s ili co fla ge lla te z on at io n of L oc ke r an d M ar tin i ( 19 89 ) a nd n an no fo ss il zo na tio n of M ar tin i ( 19 71 ). *: L at e M io ce ne , * *: la te T or to ni an , * ** : H ys tr ic ho sp ha er op si s ob sc ur a zo ne . F M : F or m at io n. M .M . a nd M . M ic oe ne : M id dl e M io ce ne . S er .: Se rr av al lia n. L an g. : L an gh ia n. P .R .Z .: Pa rt ia l R an ge Z on e. U .: U pp er . L w r. : L ow er . A qu i.: Aq ui ta ni an . B ur d. : B ur di ga lia n. U nd iff .: U nd iff er en tia te d. Lower Lang. M. Miocene L. truncatum L. tr un ca tu m ° 17 0 0 17 10 17 30 17 40 17 20 SB D 1 Lwr. Aqui.U. Aqui. – Lwr. Burd Lw r. S. h am . – C . a m ic . St ic to di sc us a �. k itt on ia nu s, O . g em m at a H ig h ab un da nc e & di ve rs ity d ia to m as se m bl ag es Undi� P. se m in ud a, A . a llo rg ei , A . a em ul an s S. h am ul at um T. p el ag ic a (o cc ) E. b ur di ga le ns is E. b ur di ga le ns is C hi ro pt er id iu m s pp . ?C . g al ea N SB 9 N S3 4 ?H om ot r. sp p. 16 80 16 90 Upper S. hamulatum P. e lo ng at a Undi� NN3-4 NS35a A . g . s ta es ch ei E. in sig ne T. ro ta D . c la do id es W el l n am e: 2 /8 - G 10 A - Fi g 5B 15 90 16 0 0 16 10 16 20 16 30 16 40 16 50 16 60 16 70 Gamma log Cores Li th o- st ra ti gr ap hy Chrono- stratigraphy D in o� ag el la te c ys t e ve nt s Samples North Sea Cenozoic Microfossils M ic ro fo ss il ev en ts SB D 2 m fs D 1 m rs D 1 Lower Miocene Burdigalian Dany Fm. Ribe Group O�shore Denmark O�shore Norway Hordaland Group Lark Fm. C. canth.Cousteaudinium aubryae Nannofossil Zone N an no fo ss il ev en ts Diatom P.R.Z. D ia to m e ve nt s Silico�agellate Zone Si lic o� ag el la te e ve nt s R. m ar ga rit al im ba ta , R. ro bu st at a, R. m ar ga rit al im ba ta T. n itz sc hi od es •- •• R. g al id a, S . o ss ifo rm is R. norwegica - C. vigilans - N. maleinterpretariaThalassiosira fraga lower Corbisema triacantha O . s pe cu lu m , D . c ru x C . t ria ca nt ha S. h or rid us •- •• North Sea Cenozoic Microfossils U . t en ui pu st ul a, G . z ea la nd ic a, G . p ra es ci tu la A . w ol te rs to r� G . z ea la nd ic a T. b ra dy i, N . c f. gr an os um C . c on tr ar ia , G . o bl iq uu s U . t en ui pu st ul a, G . e ol ab ia cr as sa ta (o cc ) G . p ra es ci tu la NN4 NSB10 NS35b D . c au li� or is, S. h et er om or ph us (o cc ) H . a m pl ia pe rt a H . w al be rs do rf en sis (o cc ) H . a m pl ia pe rt a P. m io ca en ic um C . p ou lse ni i C . a ub ry ae C . c an th ar el lu s E. in sig ne Sequence stratigraphy Dino�agellate cyst Zone E. in sig ne Ki ng 19 89 Ki ng 20 16 R. m ar yl an ic us (o cc ) H . m ed ite rr an ea (o cc ) D . e m bl em at ic us , C . p el ag ic us , H . c ar te ri, S . n eo ab ie s C . � or id an us D . e m bl em at ic us Nannofossil samples •• A bu nd an t • C om m on ° C on si st en t https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 10 of 30 GEUSBULLETIN.ORG Fi g. 6  S um m ar y di ag ra m fo r t he 2 /1 1– 12 S w el l w ith b io st ra tig ra ph ic z on es , p re vi ou sl y pu bl is he d ev en ts (b la ck te xt ) a nd n ew e ve nt s (r ed te xt ), fo r f ul l f os si l n am es s ee S up pl em en ta ry F ile S 2. L ith os tr at ig ra ph y fo r t he N or w eg ia n Se ct or fr om E id vi n et  a l. (2 02 2) , o ns ho re D en m ar k fr om R as m us se n et  a l. (2 01 0) a nd o ff sh or e D en m ar k fr om R as m us se n et  a l. (in p re ss ). Se qu en ce s tr at ig ra ph y fr om R as m us se n (2 00 4a ; 2 01 7) a nd D yb kj æ r et  a l. (2 02 1) . G am m a lo g c/ o Ak er B P. C or ed in te rv al s ar e in di ca te d. T he m ai n sa m pl e co lu m n in di ca te s sa m pl es fo r di no cy st s, m ic ro fo ss ils , d ia to m s an d si lic ofl ag el la te s. N an no fo ss il sa m pl es a re in di ca te d se pa ra te ly . Co re s am pl es a re m ar ke d w ith a d as h w it h a do t, an d di tc h cu tt in g sa m pl es a re m ar ke d w ith a d as h. C hr on os tr at ig ra ph y of R affi e t a l. (2 02 0) , d in oc ys t z on at io n of D yb kj æ r a nd P ia se ck i ( 20 10 ), m ic ro fo ss il zo na tio ns o f Ki ng (1 98 9, 2 01 6) , d ia to m z on at io n of S ch ra de r an d Fe nn er (1 97 6) , s ili co fla ge lla te z on at io n of L oc ke r an d M ar tin i ( 19 89 ) a nd n an no fo ss il zo na tio n of M ar tin i ( 19 71 ). FM : F or m at io n. U nd iff .: U nd iff er en tia te d. Im pa gi di ni um s p. A W re nn & Ko ki no s 19 86 • S. h am ul at um , U . a qu ae du ct um Gamma log Cores Li th o- st ra ti gr ap hy Chrono- stratigraphy D in o� ag el la te c ys t e ve nt s Samples North Sea Cenozoic Microfossils M ic ro fo ss il ev en ts O�shore Denmark O�shore Norway Nannofossil Zone N an no fo ss il ev en ts Diatom P.R.Z. D ia to m e ve nt s Silico�agellate Zone Si lic o� ag el la te e ve nt s North Sea Cenozoic Microfossils Sequence stratigraphy Dino�agellate cyst Zone Ki ng 19 89 Ki ng 20 16 W el l n am e: 2 /1 1 - 12 S - F ig 6 14 80 14 90 15 0 0 15 10 15 20 15 30 15 40 15 50 15 60 15 70 15 80 15 90 16 0 0 16 10 16 20 16 30 16 40 16 50 16 60 16 70 16 80 SB F m fs E m rs E SB E m fs D 2 SB D 2 m fs D 1 m rs D 1 Nordland Gp. Lower Miocene BurdigalianLower LanghianUpper LanghianSerravallian Middle Miocene Dany Fm. Nora Fm.Hodde Fm.Ørnhøj Fm. Måde Group Ribe Group Hordaland Group Lark Fm. S. h am . G . ve rr ic ul a A . a nd . C . ca nt h.Cousteaudinium aubryaeLabyrinthodinium truncatumUnipontidinium aquaeductum Undifferentiated Undifferentiated NN3-4NN4Undi� Undi� NS36c-38NS38-39 NS36a-b NS35aNS35b-c NSB12c NSB12a-b NSB10NSB11 Sy ne dr a sp p. D . c ru x D . c ru x O . s pe cu lu m H ig h ab un . & d iv . d ia to m a ss em ., D . h us te dt ii, S . g ru no w ii, D en tic ul op sis s pp . • O . g em m at a D . s m ith ii T. n itz sc hi od es •• A . i ng en s D . h ya lin a, P. e lo ng at a D en tic ul op sis s pp . • •, D . h us te dt ii R. m ar yl an di cu s S. g ru no w ii °, A . i ng en s, S. h or rid us •• S. h or rid us •• T. n itz sc hi od es •• O . g em m at a H ig h ab un da nc e an d di ve rs ity d ia to m as se m bl ag es C . � or id an us H . b ip un ct a, H . a m pl ia pe rt a, D . l en tic ul at a, R. p se ud ou m bi lic us H . s ci ss ur a, D . e m bl em at ic us , D . d ru gg ii, D . c au li� or is, D . e xi lis (o cc ), D . m oo re i ( oc c) , H . w al be rs do rf en sis (o cc ) S. p ro ce ru s ( oc c) , S. h et er om or ph us , S . c on ic us H . m ed ite rr an ea (o cc ), D . p et al lif . ( oc c) , S . a po xi s ( oc c) U . k in gi , B . p la ty re tic ul at a U . k in gi C . a ng ul io � ci na lis , T. q ua dr ilo ba tu s S. d isj un ct a, A . g . s ta es ch ei , L . s in uo su m , C . s ub co ni cu s L. p er eg rin a, C . d . p ee le ns is G . e ol ab ia cr as sa ta , T. tr ilo bu s, E. in �a tu m S. d isj un ct a U . t en ui pu st ul at a, N on io n cf . g ra no su m E. in �a tu m C . c on tr ar ia , G . p ra es ci tu la , G . z ea la nd ic a T. b ra dy i, A . w ol te rs do r� G . o bl iq uu s G . z ea la nd ic a U . t en ui pu st ul a G . p ra es ci tu la G . v er ric ul a, C . p ou lse ni i ( oc c) C . p as sio A . a nd al ou sie ns is C . p as sio , I . l ac ry m os a P. m io ca en ic um , C . p la ca nt hu m O . e iri ki an um , H . t ec ta ta • H . t ec ta ta U . a qu ae du ct um C . a ub ry ae I. ta bu la ta L. tr un ca tu m P. m io ca en ic um C . p ou lse ni i E. in sig ne C . a ub ry ae C . c an th ar el lu s E. in sig ne S. h am ul at um , ( oc c) T. ro ta P. n an a, C . c ip er oe ns is H . b ip un ct a D . d ru gg ii, D . c au li� or is, S. h et er om or ph us Denticulopsis hyalina D . h ya lin a, R. m ar ga rit al im ba ta (o cc ) R. a ng ul at a (o cc ) R. w ic om ic oe ns is Sy ne dr a sp p. (i n� ux ) E. in sig ne T. fragaRhizosolenia bulbosa C . p el ag ic us , H . c ar te ri D . e m bl em at ic us , H . s ci ss ur a S. p un ic eu s, D . d e� an dr ei S. p un ic eu s H . w al tr an s C . � or id an us , H . a m pl ia pe rt a, R. p se ud ou m bi lic us , H . c ar te ri, C . p el ag ic us A . g . s ta es ch ei Nannofossil samples •• A bu nd an t • C om m on ° C on si st en t https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 11 of 30 GEUSBULLETIN.ORG carbonates was carried out using 1M HCl until bubbling ceased, followed by 5M HCl for 24 h. This was followed by treatment with a mild solution of citric acid heated to 70°C. The silicate fraction was dissolved using cold HF (40%) for a minimum of 6 days, followed by treat- ment with a mild solution of citric acid heated to 70°C. The acid treatment was followed by brief oxidation with concentrated HNO3 and KOH 5% and by heavy liquid separation with ZnBr (2.3 g/ml). Each step was followed by filtration through an 11 µm nylon net. A final filtra- tion through a 20 µm nylon net was carried out before mounting the acid-resistant organic particles in glycerin jelly on glass slides. The dinocyst slides were examined using a Leica DM2000 normal light microscope at 400x and 1000x magnification. A minimum of 200 dinocysts were identified to species level. Finally, the slide used for counting and an additional slide were scanned to include rare dinocyst taxa. 8.1.2. Microfossils (large fraction) Approximately 50 g of sample was boiled to disaggre- gate it, and a little washing-up liquid was added to help remove drilling mud. The sediment was washed through a 63 µm sieve and dried at 60°C. The residues were sieved into >250 µm, >100 µm and >63 µm fractions. The larger fractions were picked for microfossils (primarily foraminifera, but also large diatoms and Bolboforma), and the >63 µm fraction was scanned. The microfossils were examined using a Leica M205C microscope and semi-quantitative counting. 8.1.3. Calcareous Nannofossils Nannofossil smear slides were prepared using the sim- ple smear slide technique described in Bown & Young (1998). The prepared slides were examined using a Leica DM2500P light microscope under x1000 magnification with cross-polarised light. Several slide traverses were analysed, and simple presence–absence recording was undertaken. 8.1.4. Siliceous microfossils (small fraction) The siliceous microfossil (small fraction) preparation method used in this study was developed at GEUS. Hydrogen peroxide was used to remove the organic material. The sample was then boiled for several hours and cooled, and then, a small amount of 10% HCl was added. The sample was then cleaned with distilled water and allowed to rest, several times over, for a few days. Microspheres were added and the mixture pipetted onto a glass coverslip and dried overnight. The coverslip was mounted onto a glass slide with Naphrax and heated on a hot plate to set. The prepared slides were examined using a Leica DM2500P light microscope under x1000 magnification. For the two cored wells, the whole of each slide was analysed, and all diatoms and silicoflagellates were counted. For the non-cored wells, the five ‘longest traverses’ of the circular slide were counted. 9. Results High-quality biostratigraphic data were produced from the two cored wells, 2/11–12S and 2/8–G10A; in that cored material is not subjected to harsh mechanical and chem- ical processes that can destroy fossil assemblages. The data from the ditch cutting samples from the 2/8–N4, 2/8– V6, 2/8–8 and 2/11–1 wells and from the core gap in 2/8– G10A are also generally of good quality, although caved dinocysts and microfossils occur. Natural processes such as reworking and fluctuations in abundance and diversity of the dinocysts and microfossils due to climatic or palaeo- environmental changes also influence the data. The multidisciplinary biostratigraphic study of the Lower and Middle Miocene succession of the 2/8–G10A and 2/11–12S cored sections resulted in the successful application of established regional dinocyst and microfos- sil biozonations (Dybkjær & Piasecki 2010; King 1989, 2016) and the global nannofossil zonation of Martini (1971), in addition to the recognition of several new key bioevents that appear to have stratigraphic potential (Figs. 4–8, Sup- plementary Files S1, S2, S13). The biozones and new events identified in the cored wells were tested on the non-cored sections with a high degree of success and are discussed here (see also Fig. 8 and Supplementary Files S3–S13). To our knowledge, diatoms (small fraction) and silicoflagel- lates have not previously been applied to routine biostra- tigraphy in the North Sea and are therefore discussed in some detail here. FO denotes the first (oldest) stratigraphic occurrence in this study, and LO denotes the last (young- est) stratigraphic occurrence in this study. Established bio- stratigraphic events are indicated by black text in figures, and new events are indicated in red text in figures. 9.1. Palynology In total, the studied succession from the six wells covers 14 of the dinocyst zones defined by Dybkjær & Piasecki (2010), comprising the Chiropteridium galea Zone (early Aquitanian) to the Hystrichosphaeropsis obscura Zone (late Tortonian; Figs 5, 6 and Supplementary Files S1–S6). In the 2/11–12S core, eight dinocyst zones were recorded, spanning the early Burdigalian Sumatradinium hamulatum Zone to the Serravallian Gramocysta verricula Zone (Fig. 6). The studied succession in the partly cored 2/8–G10A well starts somewhat lower, possibly in the early Aqui- tanian Chiropteridium galea Zone (Fig. 5). The palynos- tratigraphy of this lower part is not well defined, and the presence of the lower two zones, the C. galea Zone and the Homotryblium spp. Zone, are questionable as the index taxa occur in very low numbers and may be reworked. However, the occurrence of microfossil Zone NSB9 (King 1989) around this level supports an early https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 12 of 30 GEUSBULLETIN.ORG Aquitanian age for this interval. The succession up to the Unipontidinium aquaeductum Zone seems to be com- plete. Above the U. aquaeductum Zone, three dinocyst zones, the Achomosphaera andalousiense Zone, Gramo- cysta verricula Zone and Amiculosphaera umbracula Zone, are missing corresponding to the upper part of the Ser- ravallian and the lower part of the Tortonian succession. The U. aquaeductum Zone is overlain by an interval referred to as the late Tortonian Hystrichosphaeropsis obscura Zone. Above the H. obscura Zone is an interval referred to as the early Serravallian A. andalousiense Zone, indicating a repeated section possibly due to faulting. 9.2. Dinocyst events In addition to the FOs and LOs defining zonal boundar- ies, Dybkjær & Piasecki (2010, their figs. 6, 7) also pre- sented selected additional stratigraphically useful events in their zonation. These events were also found in this study, including the base of Ectosphaeropsis burdigalensis, the top of Thalassiphora rota, the base of Sumatradinium hamulatum, the top of Exochosphaeridium insigne, the base of Cerebrocysta poulsenii, the top of Cousteaudinium aubryae, the base of Palaeocystodinium miocaenicum, the top of Unipontidinium aquaeductum, the occurrence of Cannosphaeropsis passio and the top of Palaeocystodinium miocaenicum. This study has revealed some differences when com- paring the positions of these events with those of Dybk- jær & Piasecki (2010). Firstly, the top of Exochosphaeridium insigne is found in both the 2/11–12S core and the 2/8G10A core above the base of Cousteaudinium aubryae and thus within the C. aubryae Zone. In Dybkjær & Piasecki (2010), this event was located below the base of C. aubryae, within the E. insigne Zone. Next, the top of Cousteaudinium aubryae is located in both the 2/11–12S core and the 2/8–G10A core, some- what above the FO of Labyrinthodinium truncatum, and is thus within the L. truncatum Zone. These two events were erroneously shown to occur at the same level (Dybkjær & Piasecki 2010, their figs. 6, 7), while in the description of the L. truncatum Zone, it was stated that the LO of C. aubryae occurs within that zone. The data from this study thus support the latter. Acccording to Dybkjær & Piasecki (2010), the top of Cerebrocysta poulsenii is found at the top of the Achomos- phaera andalousiense Zone. In this study, in the 2/11–12S core, two specimens of C. poulsenii were found in the same sample as the base of Gramocysta verricula, the latter defining the base of the G. verricula Zone. Unfortu- nately, that sample was the highest sample analysed, so it is uncertain if the top of C. poulsenii continues further up in the G. verricula Zone in the study area. Lastly, the LO of Cleistosphaeridium placacanthum was erroneously located at two different levels in the zonation of Dybkjær & Piasecki (2010): at the top of the Achomosphaera andalousiense Zone (p. 18) and coincid- ing with the upper boundary of the A. umbraculum Zone (p. 19) and the lower boundary of the H. obscura Zone (p. 21). The location of the top of Cleistosphaeridium pla- cacanthum is clearly not a good stratigraphic marker. This study indicates a gradual decrease in abundance of this species, resulting in an indistinct and poorly defined top. In the 2/11–12S core, the top of C. placa- canthum was found in the upper part of the U. aquae- ductum Zone. In the 2/8–G10A core, it is present in the interval referred to as the A. andalousiense Zone. In the 2/8–8 well, this species was found consistently in the upper part of U. aquaeductum Zone and sporadically at least up to and within the uppermost analysed sam- ple at the base of the G. verricula Zone. In the 2/8–N4 well, the top occurrence of this species was found in an interval which either belongs to the upper U. aquaeduc- tum Zone or the lower A. andalousiense Zone, and in the 2/11–1 well, the top of C. placacanthum was found in the upper part of the U. aquaeductum Zone. However, the succession above the base of the G. verricula Zone was not included in the cored sections in this study, and so, our data set does not provide a well-documented location for the LO of this species. New events for the Danish and southern Norwegian North Sea area with potential stratigraphic use found in this study are given as follows (see Figs 5, 6, 7, 8 and Supplementary Files S1–S13): • The LO of Ectosphaeropsis burdigalensis in either the  Homotryblium spp. Zone or in the Caligodinium amiculum Zone in the 2/8-G10A well. • The LO of Membranilarnacea cf. picena group in the upper Caligodinium amiculum Zone in the 2/11-1 well. • The LO of Leptodinium italicum in the upper Sumatra- dinium hamulatum Zone in the 2/11-1 well. • The LO of Dinopterygidium cladoides in the upper S. hamulatum Zone in the 2/8G10A core (and also in the 2/8–8 and 2/11–1 wells). • The LO of Hystrichokolpoma cinctum in the upper Sumatradinium hamulatum Zone in the 2/11-1 well. • The FO of Invertocysta tabulata in the lower part of the Labyrinthodinium truncatum Zone in all wells. • The FO of Habibacysta tectata in the lower part of the Unipontidinium aquaeductum Zone in the 2/11–12S and 2/8–G10A cored wells and also in the 2/8–8 well. • The FO of Palaeocystodinium powellense in the Uniponti- dinium aquaeductum Zone in the 2/11-1 well. • The FO of Operculodinium eirikianum in the middle part of the Unipontidinium aquaeductum Zone in the 2/11– 12S, 2/8–8 and 2/11–1 wells and in the middle to ?upper https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 13 of 30 GEUSBULLETIN.ORG part of the Unipontidinium aquaeductum Zone in the 2/8–G10A well (the upper part of the zone has probably been removed by erosion or faulting, see Fig. 5). • The FO of common Habibacysta tectata in the middle part of the Unipontidinium aquaeductum Zone in the 2/11–12S, 2/8–8 and 2/11–1 wells and in the middle to ?upper part of the Unipontidinium aqueaductum Zone in the 2/8–G10A well. • The LO of Sumatradinium hamulatum in the upper part of the Unipontidinium aquaeductum Zone in the 2/11–12S well, and in the Achomosphaera andalou- siense Zone in the 2/8–N4, 2/8–8 and 2/11–1 wells. • The FO of common Impagidinium sp. A Wrenn & Kokinos 1996 in the uppermost part of the Unipon- tidinium aquaeductum Zone in the 2/11–12S and 2/8– G10A wells. • The FO of Invertocysta lacrymosa in the basal part of the Achomosphaera andalousiense Zone in the 2/11–12S well. • The FO of Operculodinium tegillatum in the Hystrichos- phaeropsis obscura Zone in the 2/8-G10A well. • The LO of Impagidinium sp. A Wrenn & Kokinos 1996 (common) in the Achomosphaera andalousiense Zone in the 2/8-G10A well. Established and new dinocyst events noted in this study are illustrated in Figs 5, 6 and Supplementary Files S1– S6. The most useful dinocyst events for correlation on a local level are marked using purple correlation lines in Supplementary Files S7–S12 and combined in a correla- tion figure (Fig. 8 and Supplementary File S13). 9.3. Microfossils Application of the North Sea microfossil zonations, NSB (King 1989) and NS (King 2016), referred the combined stud- ied interval of the six wells to Aquitanian to Early Tortonian Zones NSB9 (NS34) to NSB12c (NS38–39; Figs 4–6, Supple- mentary Files S1–S12). The calcareous benthic foraminifera that mark the top of NSB9 (NS34; LO of Plectofrondicularia seminuda), top of NSB10 (NS35; LO of Uvigerina teniupus- tulata), top of NSB11 (NS36b; LO of Asterigerina guerichi staeschei) and top of NSB12 (NS39; LO of Uvigerina kingii), along with other marker planktonic microfossils from King (1989, 2016) are applied successfully in this study. The 2/11–12S core spans Burdigalian Zone NSB10 (NS35a) to Serravallian Subzone NSB12c (NS39). The 2/8–G10A core covers Aquitanian Zone NSB9 (NS34) Langhian to Subzone NSB12a (NS36c). While FOs and LOs of planktonic and calcareous ben- thic foraminifera are useful for biostratigraphic correla- tion in the Valhall–Hod area, abundance variations in other microfossil groups, such as agglutinating foramin- fera, diatoms, radiolaria and sponge spicules, are also potentially useful for correlation. In addition, variations in the siliceous versus calcareous microfossil compo- nents are of interest regarding our understanding of the diatomite reservoir architecture and palaeoenvironment and form the basis for ongoing detailed studies. 9.4. Microfossil events In addition to the FOs and LOs of microfossils defin- ing zone boundaries, King (1989, 2016) also presented selected additional stratigraphically useful events. This study reveals some local observations and differences in the relative positions of some of these events (see Figs. 5, 6, 8 and Supplementary Files S1–S13) as follows: • The FO of Globorotalia praescitula below the FO of Uvi- gerina tenuipustulata in the 2/11–12S and 2/8–G10A wells. In King (1983), these two events are in reverse order, although their ranges are marked as uncertain. • The FO of Globorotalia zealandica above the FO of Uvigerina tenuipustulata in the 2/11–12S and 2/8– G10A wells. In King (1983), these two events coincide, although their ranges are noted as uncertain. • The coinciding LOs of Globorotalia zealandica and Glo- borotalia praescitula just below the top of Zone NSB10 (NS35b-c) in the 2/11–12S and 2/8–G10A wells. In King (1983, 2016), the LO of Globorotalia zealandica occurs slightly earlier in NSB10 than the LO of Globorotalia praescitula. • The LO of Trilobatus trilobus is noted within Zone NSB11 (NS36a-b) in this study. King (1983) places its LO in the lower part of Subzone NSB12a. • The LO of Loxostomum sinuosum is seen at the top of Zone NSB11 (NS36a-b) in the 2/8–G10A, 2/11–12S, 2/8–N4 and 2/11–1 wells. The LO of Loxostomum sin- uosum is found in the lower part of subzone NSB10 (Laursen & Kristoffersen 1999) onshore Denmark, at the top of NSB11 in King (1989) and at the top of NSB9 (NS34) in King (2016). • The FO of Uvigerina kingi marks the base of NSB12c (NS38–39) in this study in the absence of Elphidium antoninum (King 1989). In King (1989), the FO of Uvigerina kingi is within subzone NSB12c with uncertainty. We also identified new microfossil events not noted in the zonations of King (1983, 1989, 2016). These events have potential stratigraphic use and are found in the cored intervals in this study and also in some of the non-cored wells (see Figs. 5, 6, 8 and Supplementary Files S1–S13, where they are noted in red). These events are as follows: • The LO of Globigerinelloides obliquus in the upper part of NSB10 (NS35b-c) below the LOs of Alabamina wolter- storffi and Trifarina bradyi and above the FO of Uviger- https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 14 of 30 GEUSBULLETIN.ORG ina tenuipustulata in the 2/8–G10A and 2/11–12S wells. • The LO of Ceratobulimina contraria just below (2/8–G10A) or close to (2/11–12S, 2/8–V6, 2/11–1) the LO of Uvigerina tenuipustulata in upper NSB10 (NS35b-c). Onshore Denmark, in a shallower setting, the LO of Ceratobulimina contraria is a younger event marking the top of NSB12a (Laursen & Kristoffersen 1999). • The LOs of Alabamina wolterstorffi and Trifarina bradyi in the upper part of NSB10 (NS35b-c) below the LOs of Globorotalia zealandica and Globorotalia praescit- ula in the 2/8–G10A and 2/11–12S wells. The LO of Trifirina bradyi is found in the Upper Pliocene succes- sion in the North Sea (King 1989). Its consistent LO towards the top of NSB10 in the study area (also in the 2/8–N4, 2/8–8 and 2/11–1 wells) may be useful for local correlation. • The LOs of Ciperoella ciperoensis and Paragloborotalia nana close to the base of NSB11 (NS36a-b) in the 2/8– G10A, 2/11–12S, 2/8–N4 and 2/8–V6 wells. The LO of Ciperoella ciperoensis was noted in the Lower Miocene succession of the Ekofisk Field (Eidvin et al. 1999). • The LO of Globoturborotalita eolabiacrassata in the lower part of NSB11 (NS36a-b) in the 2/8–G10A and 2/11–12S wells. • The LO of Lenticulina peregrina at or just below the top of NSB11 (NS36a-b) in the 2/8–G10A, 2/11–12S and 2/8–N4 wells. • The LO of Cancris subconicus at the top of NSB11 (NS36a-b) in the 2/8–G10A and 2/11–12S wells. Two additional events are identified that may have correlation potential (also marked in red in Supple- mentary Files S1–S12). These include the LO of Non- ion cf. granosum towards the top of NSB10 (NS35b-c) in the 2/8–G10A, 2/11–12S, 2/8–N4, 2/8–8 and 2/8–V6 wells and the LOs of Ciperoella anguliofficionalis and Trilobatus quadrilobatus within subones NSB12a-b (NS36c-38) in the 2/8–G10A, 2/11–12S, 2/8–N4 and 2/8–8 wells. Established and new microfossil events noted in this study are illustrated in Figs. 5, 6 and Supplementary Files S1–S6. The most useful microfossil events for correla- tion on a local level are marked using blue correlation lines in Supplementary Files S7–S12 and combined into a correlation figure (Fig. 8 and Supplementary File S13). 9.5. Calcareous Nannofossils Approximately ten samples per well were analysed for calcareous nannofossils using basic presence–absence observations, which enabled a broad biostratigraphic breakdown into fairly long-ranging nannofossil zones, overall spanning Early to Middle Miocene zones NN3– NN6 (Martini 1971). Calcareous nannofossil ranges are based on observations of Young (1998), de Kaenel et al. (2017), Bergen et  al. (2017) and Boesiger et  al. (2017). Reworked nannofossils from the Upper Cretaceous and Palaeogene are present in all wells, and caved material was occasionally noted in the non-cored wells. Nanno- fossil events in Figs. 5, 6 and Supplementary Files S1–S12 are in black text and are not included in the correlation in Fig. 8 and Supplementary File S13 due to large sam- ple spacing and the basic counting method applied. The relative position of Early to Middle Miocene calcareous nannofossil events from the Valhall–Hod area is shown in Fig. 7. While these events are not necessarily new, they may be useful for local and regional correlation and biostratigraphy. The oldest nannofossil zone identified is Zone NN3 in the 2/8–8 well, recognised due to the co-occurrence of Helicosphaera bipuncta, Sphenolithus apoxis, Sphenolithus puniceus and Helicosphaera scissura. Joint zones NN3–4 are noted in the 2/8–G10A, 2/11–12S and 2/8–V6 wells based on an assemblage containing elements restricted to either zone. For example, Sphenolithus apoxis and Sphenolithus conicus (LOs in NN3), Discoaster emblemati- cus (FO in NN3) and Discoaster caulifloris, Discoaster pet- aliformis and Sphenolithus heteromorphus (FOs in NN4). Zone NN4 is identified in all wells and is characterised by the co-occurrence of Helicosphaera ampliaperta, vari- ably with Helicosphaera waltrans, Sphenolithus puniceus, Sphenolithus heteromorphus, Helicosphaera scissura, Discoaster emblematicus, Discoaster caulifloris and Sphe- nolithus abies. Joint zone NN4–5 is recorded in the 2/8– G10A well, based on the co-occurrence of Helicosphaera scissura, Discoaster caulifloris, Reticulofenestra pseudoum- bilicus and Discoaster discissus. The upper parts of the 2/8–N4, 2/8–8, 2/8–V6 and 2/11–1 wells are assigned a wide NN4–6 range based on the presence of Cyclicar- golithus bukryi, Cyclicargolithus floridanus and Helicos- phaera vedderi whose upper range is no younger than Zone NN6. 9.6. Calcareous Nannofossil events Nannofossil events with potential correlative use and other possible useful nannofossil biostratigraphic occur- rences are seen in Figs. 5, 6, 7 and Supplementary Files S1–12. They are described here in stratigraphic order in relation to NSB/NS microfossil zones (King 1989, 2016), and their relative positioning is shown in Fig. 7 against the nannofossil zonation of Martini (1971). Low resolu- tion sample spacing and potential caving can result in depressed FOs in the non-cored wells. The events are indicated by the: • FO of Coccolithus pelagicus in all wells at the base of the studied sections. • FO of Cyclicargolithus floridanus in all wells at or near https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 15 of 30 GEUSBULLETIN.ORG the base of the studied sections. • FO of Helicosphaera carteri at the base of the studied sections in the 2/8–G10A, 2/11–12S, 2/8–N4, 2/8–8 and 2/11–1 wells. • FO of Helicosphaera ampliaperta at the base of the studied sections in the 2/11–12S, 2/8–N4, 2/8–8 and 2/11–1 wells and slightly higher, in the upper part of NSB10 (near the base of NS35b) in the 2/8–G10A and 2/8–V6 wells. • FO of Discoaster emblematicus at the base of the stud- ied sections in the 2/8–G10A, 2/8–N4 and 2/8–8 wells and higher, in the upper part of NSB10 (NS35b-c) in the 2/11–12S and 2/8–V6 wells. • FO of Reticulofenestra pseudoumbilicus in the 2/11– 12S, 2/8–V6 and 2/11–1 wells towards the middle of NSB10 (NS35). • FO of Discoaster caulifloris in the upper part of NSB10 (near the base of NS35b) in the 2/8–G10A, 2/11–12S and 2/11–1 wells and slightly higher in the 2/8–V6 well. • FO of Discoaster exilis in the 2/8–N4, 2/8–8 and 2/11–1 wells towards the middle of NSB10 (near the base of NS35b). • LO of Discoaster emblematicus close to the boundary of NSB10 (NS35b-c) and NSB11 (NS36a) in the 2/11– 12S, 2/8–8 and 2/8–V6 wells, and slightly lower in the 2/G10A and 2/8–N4 wells. • FO of Helicosphaera bipuncta in the 2/8–G10A, 2/11– 12S and 2/8–V6 wells towards the base of NSB11 (NS36a-b) and lower, perhaps due to caving in the 2/11–1 well. • LO of Helicosphaera ampliaperta in NSB10 (NS35) in the 2/8–G10A, 2/8–N4, 2/8–8 and 2/11–1 wells and higher, within NSB12a (NS36c) in the 2/11–12S and 2/8–V6 wells. The discrepancy in age may be due to wide nannofossil sample spacing and the thin suc- cession represented by NSB11. • LO of Discoaster exilis in NSB12a-b (NS36c-38) in the 2/8–N4, 2/8–8 and 2/11–1 wells. • LO of Discoaster caulifloris within NSB11 (NS36a-b) in the 2/8–G10A and 2/11–12S wells and slightly lower, in upper NSB10 (NS35), in 2/11–1. • LO of Helicosphaera bipuncta in NSB12 (NS36–38) in the 2/11–12S and 2/11–1 wells, and lower in NSB11 (NS36a-b) in the 2/8–G10A and 2/8–V6 wells. • LO of Cyclicargolithus floridanus at or near the top of the studied sections in all wells. • LO of Helicosphaera carteri at or near the top of the studied sections in all wells. • LO of Coccolithus pelagicus in all wells at the top of the studied sections. A more extensive study of nannofossils of the calcar- eous intervals was unfortunately beyond the scope of this study but would be an interesting exercise to carry out in the future. 9.7. Diatoms Sufficiently well-preserved diatoms recorded in the cored and non-cored wells in this study allowed the application of the Early and Middle Miocene biostratig- raphy of Schrader and Fenner (1976) from the Norwe- gian Sea. In this study, the Early Miocene Rhizosolenia norwegica–Coscinodiscus vigilans–Nitzschia maleinterpre- taria joint Partial Range Zone (PRZ) to the Middle Mio- cene Denticulopsis hyalina PRZ are recognised (Figs. 5, 6, 8, Supplementary Files S1–S13). Zone definitions of Schrader & Fenner (1976) and observed assemblages in this study are described here. Apart from in the 2/8–G10A core, several of the diatom species that define the tops and bases of the PRZs of Schrader & Fenner (1976) are not consistently recorded in this study. However, the diatom assem- blages described in Schrader & Fenner (1976) in a par- ticular PRZ are similar to the assemblages found here and can provide an alternative method of identifying that specific PRZ when the markers for PRZ tops and bases are not seen. New diatom events are recognised in this study within the identified PRZs in several wells. Therefore, these new events correlate across the Val- hall and Hod areas and may potentially be of correlative Fig. 7 Overview of the relative position of Early to Middle Miocene cal- careous nannofossil events from the Valhall–Hod area. Nannofossil zonation of Martini (1971). Age (Ma) 11 12 13 14 15 16 17 18 19 20 21 22 23 Epoch Stage Nannofossils Zone M io ce ne Se rr av al lia n La ng hi an Bu rd ig al ia n A qu it an ia n To rt on ia n NN8 NN7 NN6 NN5 NN4 NN3 NN2 NN1 C. pelagicus, H. carteri, C. �oridanus, H. ampliaperta, D. emblematicus D. exilis, D. cauli�oris, R. pseudoumbilicus C. pelagicus, H. carteri, C. �oridanus H. bipuncta D. cauli�oris D. exilis H. ampliaperta D. emblematicus H. bipuncta https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 16 of 30 GEUSBULLETIN.ORG value further afield. The chronostratigraphic age of the diatom zonations in this study (Fig. 4) is linked to the dinocyst stratigraphy of Dybkjær & Piasecki (2010) in the description that follows. 9.8. Diatom zonation Rhizosolenia norwegica–Coscinodiscus vigilans–Nitzs- chia maleinterpretaria joint PRZ (Schrader & Fenner 1976) Definition. The base of the Rhizosolenia norwegica PRZ is based on the FO of Stictodiscus aff. kittonianus and Mac- rora stella. The top of the Nitzschia maleinterpretaria PRZ is based on the LOs of Dimerogramma fossile, Sceptroneis ossiformis, Thalassionema harosakiensis and Thalassi- osira spinosa var. aspinosa and the FO of Raphoneis margaritalimbata. Age. Early Miocene (Burdigalian). From the lower part of the upper Sumatradinium hamulatum Zone to the lower part of the Cousteaudinium aubryae dinocyst Zone (Dyb- kjær & Piasecki 2010) in the 2/8–G10A cored well. Diatom Assemblage (this study). Includes Stictodiscus aff. kittonianus, Raphoneis margaritalimbata, Opephora gemmata, Raphidodiscus marylandicus, Pseudodimero- gramma elongata, Thalassiosira fraga, Rocella gelida, Rhi- zosolenia hebetata, Sceptroneis ossiformis, Actinocyclus ehrenbergii, Dimerogramma fossile, Actinocyclus tenellus, Stephanopyxis horridus, Diploneis smithii, Thalassionema spp. (abundant in the uppermost part), Synedra jouse- ana, Raphoneis amphiceros, Paralia sulcata (abundant), Chaetoceros spp. (abundant), Pseudopodosira spp. (abun- dant, including Pseudoporosira westii) and Actinoptychus senarius. Observations. It was not possible to subdivide these three PRZs in this study. In the 2/8–G10A core, the base of this combined interval was based on the FO of Stictodiscus aff. kittonianus and the top by the FO of Raphoneis margaritalimbata. The FO of Stictodiscus aff. kittonianus is found with the FO of Opephora gemmata in the upper Sumatradinium hamulatum dinocyst Zone in the 2/8–G10A core. The FO of Opephora gemmata is found at a similar stratigraphic level in the 2/8–8, 2/8– V6 and 2/11–1 wells and is here used as an additional diatom event for the identification of the base of the R. norwegica–C. vigilans–N. maleinterpretaria joint PRZ. The top of this joint PRZ is marked by the FO of Raphoneis margaritalimbata in the 2/8–G10A, 2/8–8 and 2/11–1 wells (Figs. 5, 8, Supplementary Files S1, S4, S6, S7, S10, S12, S13), close to the FO of the dinocyst Cousteaudinium aubryae. The top of this PRZ in the 2/8–V6 well (Supple- mentary Files S5, S11) is comparatively high, but due to a lack of other marker diatoms is based on the FO  of common to abundant Stephanopyxis horridus in the overlying sample, an event which is characteristic of the overlying Thalassiosira fraga PRZ. The R. norwegica–C. vigilans–N. maleinterpretaria joint PRZ is only identified in the 2/8–G10A, 2/8–8, 2/8–V6 and 2/11–1 wells. Thalassiosira fraga PRZ (Schrader & Fenner 1976) Definition. The base of the Thalassiosira fraga PRZ is defined on the LOs of Dimerogramma fossile, Sceptroneis ossiformis, Thalassionema harosakiensis and Thalassiosira spinosa var. aspinosa, and the FO of Raphoneis margari- talimbata. The top is based on the FOs of Coscinodiscus lewisianus, Cymatosira biharensis, Dimerogramma aff. dubium and Hemiaulus malleus and the LO of Thalassi- osira fraga. Age. Early to Middle Miocene (Burdigalian to early Langhian). From the lower part of the Cousteaudinium aubryae Zone to the lower part of the Labyrinthodinium truncatum dinocyst Zone (Dybkjær & Piasecki 2010) in the 2/8–G10A cored well. Diatom Assemblage (this study). Includes Chaetoceros spp. (abundant), Thalassiosira fraga, Cymatosira biharensis, Raphoneis margaritalimbata, Thalassionema nitzschiodes (abundant), Paralia sulcata (abundant), Pseudopodosira spp. (common, including Pseudopodosira westii), Actino- cyclus ingens (rare), Actinocyclus ehrenbergii, Raphoneis robustata, Raphoneis angulata, Raphidodiscus marylandi- cus, Pseudodimerogramma elongata, Stephanopyxis horri- dus, Diploneis smithii, Stictodiscus aff. kittonianus, Synedra jouseana, Stephanopyxis grunowii, Rhizosolenia hebetata, Rhizosolenia miocenica (rare), Raphoneis amphiceros, Pterotheca reticulata, Opephora gemmata, Cestodiscus peplum and Actinoptychus senarius. Observations. In the 2/8–G10A, 2/8–8 and 2/11–1 wells, the base of this PRZ is based on the FO of Raphoneis margaritalimbata (Fig. 5, Supplementary Files S1, S4, S6, S7, S10, S12) close to the FO of the dinocyst Cous- teaudinium aubryae. In the 2/8–G10A well, the top of this PRZ is marked by the FO of Cymatosira biharensis coinciding with the LO of Thalassiosira fraga (Fig. 5, Sup- plementary Files S1, S7) and in the 2/8–8 well (Supple- mentary Files S4, S10) by the FO of Cymatosira biharensis, in the Labyrinthodinium truncatum dinocyst Zone, below the LO of the dinocyst Cousteaudinium aubryae. In the absence of the aforementioned established markers, alternative diatom events recognised in this study in the Thalassiosira fraga PRZ are the FO of common to abun- dant Stephanopyxis horridus close to the base of the Labyrinthodinium truncatum dinocyst Zone in the 2/8– G10A, 2/11–12S, 2/8–8, 2/8–V6 and 2/11–1 wells. The FO of common to abundant Thalassionema nitzschiodes https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 17 of 30 GEUSBULLETIN.ORG is also mainly noted within the Thalassiosira fraga PRZ in this study, but variably within the Cousteaudinium aubryae and Labyrinthodinium truncatum dinocyst zones, suggesting that this event is diachronous. The LO of Raphoneis margaritalimbata is noted towards the top of the Cousteaudinium aubryae dinocyst Zone in the 2/8– G10A, 2/8–8 and 2/11–1 wells, and the FO of Raphoneis robustata is noted at the same level in the 2/8–G10A and 2/11–1 wells. Both events are potentially useful. The T. fraga PRZ is identified in the 2/8–G10A, 2/11–12S, 2/8–8, 2/8–V6 and 2/11–1 wells (Figs. 5, 6, Supplementary Files S1–S2, S4–S8, S10–S12). Rhizosolenia bulbosa PRZ (Schrader & Fenner 1976) Definition. The base of the Rhizosolenia bulbosa PRZ is defined on the FOs of Coscinodiscus lewisianus, Cymato- sira biharensis, Dimerogramma aff. dubium and Hemiau- lus malleus and the LO of Thalassiosira fraga, and the top on the FOs of Denticulopsis hustedtii, Denticulopsis nor- wegica, Coscinodiscus endoi and Rhizosolenia miocenica. Age. Middle Miocene (Langhian). From the upper part of the Labyrinthodinium truncatum Zone to the lower part of the Unipontidinium aquaeductum dinocyst Zone (Dyb- kjær & Piasecki 2010) in the 2/8–G10A cored well. Diatom Assemblage (this study). Includes Cymatosira biharensis, Cestodiscus peplum, Raphoneis robustata, Raphoneis angulata, Pseudodimerogramma elongata, Actinocyclus tenellus, Actinocyclus ingens (rare), Diploneis smithii, Actinoptychus splendens, Synedra jouseana, Steph- anopyxis horridus (abundant), Stictodiscus aff. kittonianus, Thalassionema nitzschiodes (abundant), Chaetoceros spp. (abundant), Mediaria splendida (rare), Paralia sul- cata (common), Pseudopodosira spp. (common, includ- ing Pseudopodosira westii), Raphidodiscus marylandicus, Stephanopyxis turris, Stephanopyxis grunowii, Rhizosole- nia miocenica (rare), Rhizosolenia hebetata, Raphoneis amphiceros, Synedra spp., Pterotheca reticulata, Opephora gemmata, Denticulopsis spp. (rare) and Actinoptychus senarius. Observations. In the 2/8–G10A core, the base of this PRZ was based on the co-occurrence of the FO of Cymato- sira biharensis and the LO of Thalassiosira fraga, and in the 2/8–8 well by the FO of Cymatosira biharensis. In the 2/8–G10A, 2/11–12S, 2/8–N4, 2/8–8 and 2/8–V6 wells, the top of this PRZ is based on the FO of Denticulopsis hustedtii. The diatoms that define the base of the Rhi- zosolenia bulbosa PRZ were only recorded in the 2/8– G10A and 2/8–8 wells. In the absence of these marker diatoms in the other wells, alternative diatom events are recognised in this study in the Rhizosolenia bulbosa PRZ, which are potentially useful for correlation. An influx of Synedra spp. is noted in all wells towards the base of the Rhizosolenia bulbosa PRZ, and the FO of consistent Stephanopyxis grunowii is also noted towards the base of this zone in the 2/8–G10A, 2/8–8 and 2/8–V6 wells. The LO of Stephanopyxis horridus (common to abundant) is noted in all wells towards the middle of this PRZ (Figs. 5, 6, 8, Supplementary Files S1–13). The FO of Denticulopsis hustedtii coincides in all wells with the FO of common to abundant Denticulopsis spp. (including Denticulopsis hya- lina). This may be an easier event to recognise to mark the top of the Rhizosolenia bulbosa PRZ. The Rhizosolenia bulbosa PRZ is identified in all wells (Figs. 5, 6, Supple- mentary Files S1–S13). Denticulopsis hyalina PRZ (Schrader & Fenner 1976) Definition. The base of the Denticulopsis hyalina PRZ is defined on the FOs of Denticulopsis hustedtii, Denticu- lopsis norwegica, Coscinodiscus endoi and Rhizosolenia miocenica. The top is based on the LOs of Hemiaulus malleus, Hemiaulus malleolus and Pseudodimerogramma elongata and the FOs of the Coscinodiscus plicatus group and Rouxia californica. Age. Middle Miocene (Late Langhian). Unipontidinium aquaeductum dinocyst Zone (Dybkjær & Piasecki 2010) in the 2/8–G10A well. Diatom Assemblage (this study). Includes Denticulopsis hyalina, Denticulopsis hustedtii, Pseudodimerogramma elongata, Mediaria splendida (rare), Raphoneis wicomico- ensis, Actinocyclus tenellus, Rhizosolenia miocenica (rare), Rhizosolenia hebetata, Cymatosira biharensis (rare), Opephora gemmata, Stephanopyxis horridus, Diploneis smithii, Actinoptychus splendens (rare), Actinoptychus heliopelta (rare), Actinocyclus ingens (common), Stepha- nopyxis turris, Stictodiscus aff. kittonianus, Synedra jouse- ana (abundant), Thalassionema nitzschiodes (abundant), Stephanopyxis grunowii, Raphoneis amphiceros, Pteroth- eca reticulata, Paralia sulcata (common), Chaetoceros spp. (super-abundant) and Actinoptychus senarius. Pseudopo- dosira spp. (common, including Pseudopodosira westii) is present throughout this PRZ and becomes abundant towards the top of the studied section. Observations. In the 2/8–G10A, 2/11–12S, 2/8–N4, 2/8–8 and 2/8–V6 wells, the base of this PRZ was based on the FO of Denticulopsis hustedtii. This event is coincident with the FO of common Denticulopsis spp. (including Denticu- lopsis hyalina, Denticulopsis nicobarica (rare), Denticulopsis punctata (rare), Denticulopsis kanayae and Denticulop- sis lauta) on which the base of this PRZ is based in the 2/11–1 well. A series of local LO events occur within the Denticulopsis hyalina PRZ: the LO of Thalassionema nitzs- chiodes (abundant), Cymatosira biharensis, Denticulopsis https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 18 of 30 GEUSBULLETIN.ORG hyalina, Actinocyclus ingens, Diploneis smithii, Stictodiscus aff. kittonianus, Synedra spp. and common Denticulopsis spp. The top of the Denticulopsis hyalina PRZ in this study is based on the LO of high abundance and diversity dia- tom assemblages without evidence of species from the overlying Coscinodiscus plicatus Group PRZ and approx- imates to the upper part of the Unipontidinium aquae- ductum dinocyst Zone. The Denticulopsis hyalina PRZ is identified in all wells (Figs. 5, 6, 8, Supplementary Files S1–S13). 9.9. Diatom events and observations from this study Early to Middle Miocene diatom events found in the Norwegian and Iceland Seas (Schrader & Fenner 1976; Koç & Scherer 1996) used in this study are presented and discussed here and noted in black in Figs. 5, 6 and Supplementary Files S1–S12. The events are correlated with the dinocyst zonation of Dybkjær and Piasecki (2010; see Figs. 5, 6, 8 and Supplementary Files S1–S13). Selected diatom occurrences or events that were found in the Norwegian Sea and are noted sporadically in this study are found on Figs. 5, 6, and Supplementary Files S1–S12 and are also noted in black on the figures. Boundary-defining events of Schrader and Fenner’s (1976) zonation are discussed here as follows: • The FO of Stictodiscus aff. kittonianus defines the base of the Rhizosolenia norwegica PRZ (Schrader & Fenner 1976) and is only noted in the 2/8–G10A well at the base of the Burdigalian (upper part of the Sumatra- dinium hamulatum dinocyst Zone). • The FO of Raphoneis margaritalimbata defines the base of the Thalassiosira fraga PRZ (Schrader & Fen- ner 1976) and is found in the 2/8–G10A, 2/8–8 and 2/11–1 wells close to the FO of the Cousteaudinium aubryae dinocyst in the middle Burdigalian Stage (Fig. 5, Supplementary Files S1, S4, S6, S7, S10 and S12). The FO of Raphoneis margaritalimbata is found in the Synedra pulchella Interval (dated as late Middle Miocene by Koç & Scherer 1996) in the Iceland Sea. • The LO of Thalassiosira fraga marks the top of the Thalassiosira fraga PRZ and the base of the Rhizosole- nia bulbosa PRZ. This Early Miocene event (Schrader & Fenner 1976) was only noted in the 2/8–G10A well in the middle of the early Langhian Labyrinthodinium truncatum dinocyst Zone. • The FO of Cymatosira biharensis is recognised in the 2/8–G10A and 2/8–8 wells in the middle of the early Langhian Labyrinthodinium truncatum dinocyst Zone. Its FO defines the base of the Rhizosolenia bulbosa PRZ in the Norwegian Sea (Schrader & Fenner 1976), which they interpreted as a late Early Miocene event. In the Iceland Sea, its FO is noted towards the base of the Proboscia praebarboi Interval, dated as early Late Miocene by Koç & Scherer (1996). • The FO of Actinocylus ingens (2/11–12S, 2/8–N4) or the FO of consistent Actinocylus ingens (2/8–G10A, 2/8–8, 2/8–V6, 2/11–1) is seen in all wells in the middle to upper part of the Labyrinthodinium truncatum dino- cyst Zone, close to the LO of Cousteaudinium aubryae. In the Norwegian Sea, the FO of Actinocylus ingens is found in the Rhizosolenia bulbosa PRZ of Schrader & Fenner (1976), which they interpreted to be late Ear- ly Miocene in age. In the Iceland Sea, its FO is found close to the base of the Actinocylus ingens Interval (dated as late Middle Miocene by Koç & Scherer 1996). • In the Norwegian Sea, the FO of Rhizosolenia mioce- nica defines the base of the Denticulopsis hyalina PRZ (and the top of the underlying Rhizosolenia bulbosa PRZ), Schrader & Fenner (1976). The FO of Rhizosole- nia miocenica is seen in the 2/8–G10A, 2/8–N4, 2/8–V6 and 2/11–1 wells in the lower part of the Labyrintho- dinium truncatum dinocyst Zone, in the upper part of the Thalassiosira fraga and lowermost Rhizosolenia bulbosa PRZ’s. While noting that this event defines a zonal boundary in Schrader & Fenner (1976), in this study, other events are deemed to be more useful. Additional diatom events (FOs, LOs) noted in Schrader & Fenner (1976), and observations from this study with potential stratigraphic use in the North Sea, are pre- sented here (see Figs. 5, 6, 8 and Supplementary Files S1–S13; marked on the figures in red) as follows: • The FO of high abundance and diversity diatom assemblages is found in the upper part of the Suma- tradinium hamulatum dinocyst Zone in the 2/8–G10A, 2/8–8, 2/8–V6 and 2/11–1 wells. In the 2/11–12S well, the event is found at the base of the Cousteaudinium aubryae dinocyst Zone, and in the 2/8–N4 well, it is found towards the base of the Labyrinthodinium trun- catum dinocyst Zone, clearly displaying that this event is diachronous across the Valhall–Hod area. • The FO of Opephora gemmata is found in the 2/8–G10A, 2/8–8, 2/8–V6 and 2/11–1 wells close to or at the FO of high abundance and diversity diatom assemblages, in the upper part of the Sumatradinium hamulatum dino- cyst Zone. The FO of Opephora gemmata is found at the base of the Synedra jouseana PRZ (dated as Early Miocene by Schrader & Fenner 1976) in the Norwegian Sea. • The FO of common to abundant Thalassionema nitzschiodes is noted in all wells, usually in the mid- dle Cousteaudinium aubryae dinocyst Zone (wells 2/8–G10A, 2/8–8, 2/8–V6, 2/11–1) and occasion- ally in the lower Labyrinthodinium truncatum Zone (2/11–12S, 2/8–N4). Thalassionema nitzschiodes is https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 19 of 30 GEUSBULLETIN.ORG consistently abundant in the Iceland Sea from the top of the Synedra pulchella Interval to the top of the Denticulopsis hustedtii Interval, dated as late Middle Miocene to early Late Miocene by Koç & Scherer (1996). • The LO of Raphoneis margaritalimbata is recognised in the 2/8–G10A, 2/8–8 and 2/11–1 wells and wells towards the top of the Cousteaudinium aubryae dino- cyst Zone. In the Iceland Sea, this event occurs in the Probiscia praeparboi Interval, dated early Late Mio- cene according to Koç & Scherer (1996), and in the Norwegian Sea at the base of the Cymatosira bihar- ensis PRZ, dated Late Miocene by Schrader & Fenner (1976). • The FO of common to abundant Stephanopyxis hor- ridus is recognised in all wells, usually in the upper Cousteaudinium aubryae dinocyst Zone, occasionally in the lower Labyrinthodinium truncatum dinocyst Zone. In the Norwegian Sea, the base of Stephano- pyxis horridus is not well defined but it is present at least from the Actinocyclus ingens PRZ, dated as Mid- dle Miocene by Schrader & Fenner (1976). • The FO of Raphoneis robustata is recognized in the 2/8–G10A, 2/8–N4 and 2/11–1 wells in the upper part of the Cousteaudinium aubryae or lower Labyrinthodin- ium truncatum dinocyst zones. • The FO of an influx of Synedra spp. is noted in all wells at the base of the Rhizosolenia bulbosa PRZ in this study, in the lower to mid Labyrinthodinium truncatum dinocyst Zone. • The LO of Stephanopyxis horridus (common to abun- dant) is recognised in all wells in the Labyrinthodinium truncatum dinocyst Zone. In the Norwegian Sea, the LO of Stephanopyxis horridus is found towards the base of the Rhizosolenia miocenica PRZ, dated as early Late Miocene by Schrader & Fenner (1976). • The FO of consistent Stephanopyxis grunowii is recorded in all wells in the Labyrinthodinium trunca- tum dinocyst Zone, close to the LO of Stephanopyxis horridus (common to abundant). • The FO of Denticulopsis hyalina is noted close to the boundary of the Labyrinthodinium truncatum and Unipontidinium aquaeductum dinocyst zones in the 2/8–G10A, 2/11–12S, 2/8–N4, 2/8–8 and 2/11–1 wells, just below or at the FO of common to abundant Den- ticulopsis spp. In the Norwegian Sea, the FO of Dentic- ulopsis hyalina is close to the base of the Denticulop- sis hyalina PRZ, dated as Early to Middle Miocene by Schrader & Fenner (1976). • The FO of common to abundant Denticulopsis spp. is noted in all wells and is found towards the base of the Unipontidinium aquaeductum dinocyst Zone in the 2/11–12S, 2/8–G10A, 2/8–8 and 2/8–V6 wells and towards the top of the upper Labyrinthodinium trun- catum dinocyst Zone in the 2/8–N4 and 2/11–1 wells. Denticulopsis spp. is consistently common to abun- dant from the Coscinodiscus norwegicus Interval, dated as Middle Miocene by Koç & Scherer (1996) in the Ice- land Sea and the Denticulopsis hyalina PRZ, dated as early Middle Miocene by Schrader & Fenner (1976) in the Norwegian Sea. Common Denticulopsis spp. is noted in the upper Langhian interval in the E-8X well, Danish sector of the North Sea. • The LO of common-abundant Thalassionema nitzs- chiodes is recorded in the Unipontidinium aquaeduc- tum dinocyst Zone in most wells, not far above the FO of common to abundant Denticulopsis spp. In the Iceland Sea, this event is noted in the Proboscia bar- boi Interval, dated as Late Miocene by Koç & Scherer (1996). • The LO of Diploneis smithii is recognised at or between the LO of common-abundant Thalas- sionema nitzschiodes and the LO of high abundance and diversity diatom assemblages in the Uniponti- dinium aquaeductum dinocyst Zone in the 2/8–G10A, 2/11–12S, 2/8–V6 and 2/8–8 wells and at the top of the Labyrinthodinium truncatum dinocyst Zone in the 2/11–1 well. The ‘Diploneis group’ is noted as rare occurrences in the high-latitude North Atlantic Ocean region from the Early Miocene to the Early Pliocene in Baldauf (1982). • The LO of Actinocyclus ingens is recognised in all wells in the upper part of the Unipontidinium aquaeductum dinocyst Zone. In this study, its LO is close to the LO of Diploneis smithii. In the 2/8–N4 and 2/8–8 wells, its LO coincides with the LO of high abundance and diversity diatom assemblages. In the Norwegian Sea, the LO of Actinocyclus ingens marks the top of the Goniothecium tenue PRZ (Schrader & Fenner 1976), which is Middle Miocene in age according to Barron (1985). In the Ice- land Sea, its LO marks the top of the Proboscia prae- barboi Interval, dated as early Late Miocene by Koç & Scherer (1996). • The LO of Denticulopsis hyalina is recognised in the 2/8–G10A, 2/11–12S, 2/8–8, 2/11–1 and 2/8–N4 wells just below the LO of Actinocyclus ingens and in most sections just below the LO of high abundance and diversity diatom assemblages. In the Norwegian Sea, the LO of Denticulopsis hyalina is found at the top of the Coscinodiscus plicatus PRZ, dated as Middle Mio- cene by Schrader & Fenner (1976). • The LO of common Denticulopsis spp. (including Den- ticulopsis lauta and Denticulopsis hustedtii) is recorded in most wells in the upper part of the Unipontidin- ium aquaeductum dinocyst zone. In the Norwegian Sea and Iceland Sea regions, the event is found in the Denticulopsis hustedtii PRZ and Interval, dated as Late Miocene by Schrader & Fenner (1976) and Koç & https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 20 of 30 GEUSBULLETIN.ORG Scherer (1996). In the E-8X well in the Danish sector of the North Sea, the LO of Denticulopsis spp. is noted in the Late Langhian Stage. This event coincides with the LO of high abundance and diversity diatom assem- blages in the 2/8–G10A, 2/11–12S, 2/8–8 and 2/11–1 wells and is slightly below this level in the 2/8–N4 and 2/8–V6 wells. • The LO of high abundance and diversity diatom assemblages (including Opephora gemmata, Steph- anopyxis grunowii and Synedra spp.) is recognised in most wells in this study in the upper part of the Unipontidinium aquaeductum dinocyst Zone. This event coincides with the top of the Hodde Formation in the 2/8–G10A, 2/11–12S, 2/8–8 wells and the top of the Nora Formation in the 2/8–N4, 2/8–V6 and 2/11–1 wells. Diatom events noted in this study are illustrated in Figs.  5, 6 and Supplementary Files S1–S6. The most useful diatom events for correlation on a local level are marked using green correlation lines in Supplementary Files S7–S12 and combined in the correlation figure (Fig. 8 and Supplementary File S13). 9.10. Silicoflagellates Silicoflagellates commonly only make up about 2–3% of the biogenic part of siliceous sediments (McCartney et al. 2011). As a silicoflagellate zonation for the North Sea does not exist, the Early and Middle Miocene Nor- wegian Sea silicoflagellate biostratigraphy of Locker & Martini (1989) was applied, with some success, to the 2/8–G10A well and with limited success to the 2/8–N4, 2/8–8 and 2/11–1 wells (Fig. 5, and Supplementary Files S1, S3, S4, S6, S7, S9, S10, S12). The 2/11–12S and 2/8–V6 wells did not yield assemblages adequate for a silicoflagellate biostratigraphic breakdown. Two sil- icoflagellate zones are identified in this study: (1) the lower Corbisema triacantha Zone and (2) the upper Cor- bisema triacantha Zone. As this is the first time that sil- icoflagellate biostratigraphy has been applied to North Sea Lower-Middle Miocene deposits, zonal definitions and observed assemblages in this study are described in Section 9.10.1. The chronostratigraphy of the silico- flagellate zones is established via correlation with the dinocyst stratigraphy of Dybkjær & Piasecki (2010) and this study (Fig. 4). 9.10.1. Silicoflagellate zonation Lower Corbisema triacantha Zone (Locker & Martini 1989) Definition. From the LO of Naviculopsis quadratum to the FO of Mesocena diodon (now known as Bachmannocena diodon). Age. Early to Middle Miocene (this study, Burdigalian to early Langhian, Cordosphaeridium cantharellus – Lab- yrinthodinium truncatum dinocyst zones, Dybkjær & Piasecki 2010). Observations. In the 2/8–G10A well, the co-occurrence of Corbisema triacantha, Mesocena apiculata apiculata, Octatis speculum ssp. and Distephanus crux ssp. (common) with- out Bachmannocena diodon or Naviculopsis quadratum indicates the presence of the lower Corbisema triacantha Zone. Assemblages contained Octatis speculum hemis- phaericum, Octatis speculum speculum and Distephanus pentagona. Siliceous microfossil assemblages were bar- ren with respect to silicoflagellates below 1666.72 m. The Lower Corbisema triacantha Zone is only identified in the 2/8–G10A well (Fig. 5, Supplementary Files S1, S7). A joint upper and lower Corbisema triacantha ‘zone’ is assigned to the 2/8–N4, 2/8–8 and 2/11–1 wells (Supplementary Files S3, S4, S6, S9, S10, S12) as the marker for the top of the Lower Corbisema triacantha Zone, Bachmannocena diodon, was not seen in these wells. Upper Corbisema triacantha Zone (Locker & Martini 1989) Definition. From the FO of Mesocena diodon (now known as Bachmannocena diodon) to the LO of Corbisema triacantha. Age. Middle Miocene (this study, late Langhian to early Serravallian, Unipontidinium aquaeductum dinocyst Zone, Dybkjær & Piasecki 2010). Observation. In the 2/8–G10A well, the co-occurrence of Corbisema triacantha, Distephanus crux ssp. (common) and Bachmannocena diodon indicates the presence of the Upper Corbisema triacantha Zone. Assemblages also contain Octatis speculum ssp., Mesocena elliptica, Mesocena dumitricae, Dictyocha aspera, Octatis specu- lum hemisphaericum and Octatis speculum quintus. The total range of Bachmannocena diodon occurs within the Upper Corbisema triacantha Zone in this study. Its LO in the Norwegian Sea and Iceland and Rockall plateaux is younger, ranging up into the Late Miocene to Early Pliocene (Ciesielski et al. 1989; Amigo 1999). The Upper Corbisema triacantha Zone is only identified in the 2/8– G10A well (Fig. 5, Supplementary Files S1, S7). A joint upper and lower Corbisema triacantha ‘zone’ is assigned to the 2/8–N4, 2/8–8 and 2/11–1 wells (Supplementary Files S3, S4, S6, S9, S10, S12) as the marker for the base of the Upper Corbisema triacantha Zone, Bachmanno- cena diodon, was not seen in these wells. The base of the overlying Paramesocena circulus apiculata Zone is based on the LO of Corbisema triacantha. This event is seen in several wells but the only silicoflagellate species https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 21 of 30 GEUSBULLETIN.ORG Fig. 8 Caption on page 23. N SB 11 N SB 10 N SB 9 1460 1450 2/11 - 1 1470 1480 1490 1500 1510 1520 1530 1540 1550 1560 1570 1580 1590 1600 1610 1620 1630 1640 1650 1660 1670 1680 1690 1710 1720 1730 1740 1750 1760 1770 1780 1790 1800 1700 G am m a ra y lo gLitho- stratigraphy Se qu en ce s C hr on o- st ra ti gr ap hy D in oc ys t Z on e M ic ro fo ss ils (K in g 20 16 ) SBF1 mfs E SBE mfs D2 SB D2 mfs D1 SBD1 mrs D1 N or dl an d G ro up Lo w er M io ce ne Bu rd ig al ia n U pp er A qu it an ia n – Lo w er m os t B ur di ga lia n Lo w er A qu it an ia n La ng hi an Se rr av al lia n To rt on ia n M id dl e M io ce ne U pp er M io ce ne Ri be G ro up D an y Fo rm at io n N or a Fo rm at io n M åd e G ro up H od de F or m at io n O �s ho re D en m ar k O �s ho re N or w ay H or da la nd G ro up La rk F or m at io n H om ot . s pp . G. v. A. u. A. a. Ca lig od in iu m am ic ul um T. p. – L. s.h. U pp er S um at ra di ni um ha m ul at um C. c./ E. i. C ou st ea ud in iu m a ub ry ae La by rin th od in iu m tr un ca tu m U ni po nt id in iu m aq ua ed uc tu m D ia to m Z on e M ic ro fo ss ils (K in g 19 89 ) U nd i� er en ti at ed U nd i� er en ti at ed D en tic ul op sis h ya lin a Rh iz os ol en ia b ul bo sa Th al as sio sir a fr ag a N SB 12 c N S3 6c -3 8 N S3 8- 39 N SB 12 a- b N S3 6a N S3 5b -c N S3 5a N S3 4 2/11 - 12S Fig. 8A 1480 1490 1500 1510 1520 1530 1540 1550 1560 1570 1580 1590 1600 1610 1620 1630 1640 1650 1660 1670 1680 G am m a ra y lo g Se qu en ce s D ep th (m ) D in oc ys t Z on e M ic ro fo ss ils (K in g 20 16 ) SBF mfs E mrs E SBE mfs D2 SBD2 mrs D1 mfs D1 S. h. G. v. A. a. C. c. C ou st ea ud in iu m a ub ry ae La by rin th od in iu m tr un ca tu m U ni po nt id in iu m a qu ae du ct um D ia to m Z on e M ic ro fo ss ils (K in g 19 89 ) U nd iff er en ti at ed U nd i� N S3 6c -3 8 N S3 8- 39 N S3 6a -b N S3 5a N S3 5b -c N SB 12 c N SB 12 a- b N SB 10 N SB 11 D en tic ul op sis h ya lin a E. i. T. fr ag a Rh iz os ol en ia b ul bo sa C or es 1750 1740 2/8 - V6 1760 1770 1780 1790 1800 1810 1820 1830 1840 1850 1860 1870 1880 1890 1900 1910 1920 1930 1940 1950 1960 1970 1980 2000 2010 2020 2030 2040 2050 2060 1990 G am m a ra y lo g Se qu en ce s D in oc ys t Z on e M ic ro fo ss ils (K in g 20 16 ) SB F1 mfs E SBE mfs D2 SBD2 mfs D1 mrs D1 S. h am ul at um C . ca nt ha re llu s E. i. C ou st ea ud in iu m a ub ry ae La by rin th od in iu m tr un ca tu m U ni po nt id in iu m aq ua ed uc tu m D ia to m Z on e M ic ro fo ss ils (K in g 19 89 ) Rh iz os ol en ia b ul bo sa Th al as sio sir a fr ag a U nd i� . D . h ya lin a N S3 5a N S3 6a -b N S3 5b -c N SB 12 a- b N S3 6c -3 8 N SB 10 N SB 11 U. R. n or w eg ic a - C . v ig ila ns - N . m al ei nt er pr et ar ia R. n or w eg ic a - C . v ig ila ns - N . m al ei nt er pr et ar ia D ep th (m ) D ep th (m ) GR--> GR--> GR--> a, f c a, b, a a b d g, k i, j, l q a, b c a d e f d g, k h, r j, i, l d, g i, m, t k, m, j, n, q l, k o v n, l m r, w p, q, p r, s t, o b d e, g h k l m o, p q, x r u, x s, y, z aa, w, y e, f d, f, g, h, j, r i, k l, m, t, u, v n, o, p r s t d e c f e m, n o, p, q r s, t, u, v g, h g, d h i l m n q, x r, s, t u, v w, y, w, y z b, j, i, j, k d, g, k, l k m l,n o p, q r, o, p s t, x https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 22 of 30 GEUSBULLETIN.ORG 2010 2000 2/8 - N4 2020 2030 2040 2050 2060 2070 2080 2090 2100 2110 2120 2130 2140 2150 2160 2170 2180 2190 2200 2210 2220 2230 2240 2260 2270 2280 2290 2250 G am m a ra y lo g Se qu en ce s D in oc ys t Z on e M ic ro fo ss ils (K in g 20 16 ) SBF GR--> mfs E SBE mfs D2 SBD 2 mfs D1 mrs D1 U. s.h. E. i. C . c an t. C ou st ea ud in iu m au br ya e La by rin th od in iu m tr un ca tu m D ia to m Z on e M ic ro fo ss ils (K in g 19 89 ) U nd i� . U nd i� er en ti at ed N S3 5a N S3 6c -3 8 N S3 5b -c N S3 6a N SB 11 N SB 12 a- b N SB 10 1460 1450 2/8 - G10A 1470 1480 1490 1500 1510 1520 1530 1540 1550 1560 1570 1580 1590 1600 1610 1620 1630 1640 1650 1660 1670 1680 1690 1700 1710 1730 1740 1720 G am m a ra y lo g C or es M ic ro fo ss ils (K in g 20 16 ) SBE mfs D2 mrs E SBD2 mfs D1 mrs D1 SBD1 *** A. a. ? ? L. S. h – C. a U pp er S . h am ul at um C . c an th . C ou st ea ud in iu m a ub ry ae La by rin th od in iu m tr un ca tu m U ni po nt id in iu m a qu ae du ct um D ia to m Z on e R. n or w eg ic a - C . v ig ila ns - N itz sc hi a m al ei nt er pr et ar ia Th al as sio sir a fr ag a U nd i� er en ti at ed T. fr ag a Rh iz os ol en ia b ul bo sa D en tic ul op sis h ya lin a M ic ro fo ss ils (K in g 19 89 ) U nd i� N SB 12 a N SB 11 N SB 10 U nd i� N S3 6c N S3 6a -b N S3 5b N S3 5a Fig. 8B SBD 1 Se qu en ce s D in oc ys t Z on e E. i. ?C. g. NSB9 NS34 ?H. s. 1380 1370 2/8 - 8 1390 1400 1410 1420 1430 1440 1450 1460 1470 1480 1490 1500 1510 1520 1530 1540 1550 1560 1570 1580 1590 1600 1610 1630 1640 1650 1660 1670 1680 1690 1700 1710 1720 1620 G am m a ra y lo g Se qu en ce s D in oc ys t Z on e M ic ro fo ss ils (K in g 20 16 ) SBF mfs E SBE mfs D2 SBD 2 mfs D1 mrs D1 Lo w er S . h am ul at um U pp er S . h am ul at um E. i./ C. c. C ou st ea ud in iu m a ub ry ae La by rin th od in iu m tr un ca tu m U ni po nt id in iu m a qu ae du ct um G. v. A. a. D ia to m Z on e M ic ro fo ss ils (K in g 19 89 ) Th al as sio sir a fr ag a Rh iz os ol en ia b ul bo sa D en tic ul op sis h ya lin a U nd i� U nd i� er en ti at ed N S3 5a N S3 6c -3 8 N S3 7- 38 U nd i� . U nd i� . N SB 12 b N S3 5b NS35c N S3 4 N SB 9 N SB 12 a- b N SB 10 SBD 1 U . aq ua Ac ho m os ph ae ra an da lo us ie ns e Rh iz os ol en ia b ul bo sa D en tic ul op sis h ya lin a NS37-38NSB12b NS36a-bNSB11 R. n or w eg ic a - C . v ig ila ns - N . m al ei nt er pr et ar ia D ep th (m ) D ep th (m ) D ep th (m ) GR--> GR--> a, b, c b b a d g l n, o, v a b c, d f a e, e d, g f, j g, k, j, k, r a, b, i, m, o, q, r c d e f g d k h, i, j, g k, j, l, n, u l d, g, h, j, k, p, t, v l, m, n o p, q s, m r, n t, p, r, u, x o, q, s s, w j, k, l, p, t m m n o p r, u, y s w, z n, l o p, q w r s t, q a a, b b c d e h, d i, e, i f, g h k j m, n, o, p q, r, s, t, v w m n o r, v, x, y d, f, c e f, g, h, j, k, l i j, k m, l n o p, q r s, p t, u p, q, p Fig. 8 (Continued) Caption on next page. https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 23 of 30 GEUSBULLETIN.ORG present above the LO of Corbisema triacantha is rare Distephanus crux, which cannot be used to confirm the presence of the Paramesocena circulus apiculata Zone. 9.10.2. Silicoflagellate events Locker & Martini (1989) presented stratigraphically useful Early to Middle Miocene silicoflagellate events from the Norwegian Sea. Some of those events were recognised in this study (Figs. 5, 6, Supplementary File S1–12) and may be useful for correlation within the Valhall–Hod area. These are as follows: • The FO of Distephanus crux is found in the Burdigalian Cordosphaeridium cantharellus/Sumatradinium hamu- latum dinocyst zones in the 2/8–G10A, 2/8–8, 2/8–V6 and 2/11–1 wells. In the 2/11–12S and 2/8–N4 wells, the event occurs in the Early Langhian Labyrinthod- inium truncatum Zone. The apparent diachroneity of the event in the Valhall–Hod area may be due to the scarcity of silicoflagellates. • The FO of Corbisema triacantha is noted in the lower part of the Cousteadinium aubryae dinocyst Zone in the 2/8–G10A well and towards the base of the Laby- rinthodinium truncatum dinocyst Zone in the 2/8–N4, 2/8–8 and 2/11–1 wells (Supplementary Files S3, S4, S6, S9, S10, S12). The apparent diachroneity of this event in the Valhall–Hod region may be due to the scarcity of silicoflagellates. • The FO of Bachmannocena diodon is found locally towards the base of the early Langhian Unipontidinium aquaeductum dinocyst Zone in the 2/8–G10A well (Fig. 5, Supplementary Files S1, S7). It is only noted in the 2/8–G10A cored section but is an important event as it marks the base of the Upper Corbisema triacantha silicoflagellate Zone in the Norwegian Sea (Locker & Martini 1989). • The LO of Corbisema triacantha is an important event as it marks the top of the Upper Corbisema triacantha silicoflagellate Zone in the Norwegian Sea (Locker & Martini 1989). The LO of Corbisema triacantha is found below the LO of Distephanus crux in the early Langhian Unipontidinium aquaeductum dinocyst Zone in the 2/8–G10A and 2/8–8 wells (Fig. 5, Supple- mentary Files S1, S4, S7, S10), and lower, in the late Langhian Labyrinthodinium truncatum dinocyst Zone in the 2/8–N4 and 2/11–1 wells (Supplementary Files S3, S6, S9, S12). The apparent diachroneity of this event in the Valhall–Hod region may be due to the scarcity of silicoflagellates. • The LO of Distephanus crux is diachronous in the Valhall–Hod area. It is found within the Unipontidin- ium aquaeductum dinocyst Zone in the 2/8–G10A, 2/11–12S, 2/8–8, 2/8–V6 and 2/11–1 wells and slightly lower in the late Langhian Labyrinthodinium trunca- tum dinocyst Zone in the 2/8–N4 well. 10. Correlation and stratigraphy The biostratigraphic framework presented here is the result of the analysis of two cored sections (wells 2/11-12S and 2/8-G10A) and four non-cored sections Fig. 8 (Continued) Multidisciplinary biostratigraphic correlation diagram of the six studied wells, from south to north, from the Hod Field to the Valhall Field using dinocyst, microfossil and diatom events. Events are correlated using coloured correlation lines and are denoted using a letter as follows: Dinocysts (purple lines): a: FO Sumatradinium hamulatum, b: LO Dinopterygium cladoides, c: LO Thalassiophora rota, d: FO Exochosphaeridium insigne, e: LO Cordosphaeridium cantharellus, f: FO Cousteaudinium aubryae, g: LO Exochosphaeridium insigne, h: FO Cerebrocysta poulsenii, i: FO Palaeocystodinium miocaenicum, j: FO Labyrinthodinium truncatum, k: FO Invertocysta tabulata, l: LO Cousteaudinium aubryae, m: FO Unipontidinium aquaeductum, n: FO Habibacysta tectata, o: FO Habibacysta tectata (common), p: FO Operculodinium erikanium, q: LO Cleistosphaeridium placacanthum, r: LO Unipontidinium aquaeductum, s: LO Sumatradinium hamulatum, t: FO Impagidinium sp A Wrenn & Kokinos 1996 (common), u: FO Invertocysta lacrymosa, v: FO Can- nosphaeropsis passio, w: LO Cannosphaeropsis passio, x: LO Palaeocystodinium miocaenicum, y: FO Achomosphaera andalousiensis, z: FO Grammocysta verricula, aa: FO Amiculosphaera umbracula. Microfossils (blue lines): a: LO Plectofrondicularia seminuda, b: LO Aulacodiscus allorgei, c: FO Globorotalia praescitula, d: FO Uvigerina tenuipustulata, e: LO Globigerinoides obliquus, f: FO Globorotalia zealandica, g: LO Trifarina bradyi, h: LO Alabamina wolter- storffi, i: LO Ceratobulimina contraria, j: LO Globorotalia zealandica, k: LO Globorotalia praescitula, l: LO Uvigerina tenuipustulata, m: LO Paragloborotalia nana, n: LO Ciperoella ciperoensis, o: LO Trilobatus trilobus, p: LO Globoturborotalita eolabiacrassata, q: LO Elphidium inflatum, r: LO Lenticulina peregrina, s: LO Cancris subconicus, t: LO Loxostomum sinuosum, u: LO Sphaeroidinellopsis disjuncta, v: LO Asterigerina guerichi staeschei, w: LO Bolboforma platyre- ticulata, x: FO Uvigerina kingi, y: LO Uvigerina kingi. Diatoms (green lines): a: FO High abundance and diversity diatom assemblages, b: FO Opephora gemmata, c: FO Stictodiscus aff. kittonianus, d: FO Thalassionema nitzschiodes (common to abundant), e: FO Raphoneis margaritalambata, f: LO Raphoneis margaritalambata, g: FO Stephanopyxis horridus (common to abundant), h: FO Raphoneis robustata, i: FO Rhizosolenia miocenca, j: FO Cymatosira biha- rensis, k: FO Synedra spp. (influx), l: FO Stephanopyxis grunowii, m: LO Stephanopyxis horridus (common to abundant), n: FO Actinocyclus ingens, o: FO Denticulopsis hyalina, p: FO Denticulopsis spp. (common to abundant), q: FO Denticulopsis hustedtii, r: LO Thalassionema nitzschiodes (common to abun- dant), s: LO Diploneis smithii, t: LO High abundance and diversity diatom assemblages (including A. ingens, Denticulopsis spp., O. gemmata, S. grunowii, S. aff. kittonianus, Synedra spp.). Sequence boundary correlations are marked in brown. The datum line for the correlation is sequence boundary SBE. Lithostratigraphy for the Norwegian Sector from Eidvin et al. (2022), onshore Denmark from Rasmussen et al. (2010) and offshore Denmark from Rasmussen et al. (in press). Sequences from Rasmussen (2004a; 2017) and Dybkjær et al. (2021). Gamma (GR) log c/o Aker BP. Cored intervals are indicated. Chronostratigraphy of Raffi et al. (2020), dinocyst zonation of Dybkjær & Piasecki (2010), microfossil zonations of King (1989, 2016) and diatom zonation of Schrader & Fenner (1976). Dinocyst zone abbreviations: A. a.: Achomosphaera andalousiensis, A. u.: Amiculosphaera umbracula, C. a.: Caligodinium amiculum, C. c.: Cordosphaeridium cantharellus, C. g.: Chiropteridium galea, E. i.: Exochosphaeridium insigne, G. v.: Gramocysta verricula, H. s.: Homotryblium spp., L. S. h.: Lower Sumatradinium hamulatum, S. h.: Sumatradinium hamulatum, T. p.: Thalassiphora pelagica, U. S. h.: Upper Suma- tradinium hamulatum, ***: Hystrichosphaeropsis obscura, U.: undifferentiated. Grey shading denotes no information. A full-sized version of this figure is available as Supplementary File S13. https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 24 of 30 GEUSBULLETIN.ORG (wells 2/8-N4, 2/8-V6, 2/8-8 and 2/11-1). The resulting framework with new and established dinocyst, micro- fossil, calcareous nannofossil, diatom (small frac- tion) and silicoflagellate events is presented in Figs. 5, 6, 8 and Supplementary Files S1-S6. The new and established fossil events are shown in Supplementary Files S7–S12, denoted by coloured correlation lines (dinocysts-purple, microfossils-blue, diatoms-green, sequence boundaries-brown). These correlations are the basis for the conceptual Valhall–Hod correlation diagram (Fig. 8 and Supplementary File S13) showing a south to north correlation using dinocyst, microfos- sil and diatom (small fraction) events and sequence boundaries. The successful application of the biostra- tigraphic correlation of new and established events in the Valhall–Hod area implies that they could also be valuable in a more regional context. The studied succession comprises the Aquitanian– Serravallian (Early Miocene–Late Middle Miocene) inter- val of the Valhall–Hod area. Our biostratigraphic study has enabled the dating of the diatomite-rich succession in the Valhall–Hod area and correlation of the succes- sion with the Norwegian lithostratigraphy and the more detailed, newly established lithostratigraphy for the Danish sector (Rasmussen et al., in press). The study also forms the framework for correlating sequence strati- graphic surfaces and units from the central North Sea Basin to onshore Denmark (Rasmussen 2004b, 2017; Dybkjær et al. 2021). The thickest diatomite-rich interval within the Mio- cene succession (c. 100 m thick in the 2/8–G10A core on the crest of Valhall structure, Fig. 5, Supplementary File S1; c. 80 m thick in the 2/11–12S core on the crest of Hod structure, Fig. 6, Supplementary File S2) is of Langhian age and correlates with the Nora Formation. The Nora Formation was defined in the Danish sector of the North Sea (Rasmussen et al., in press) and correlates with the upper part of the Odderup Formation and lower part of the Hodde Formation defined for onshore Denmark (Fig. 3). The underlying minor diatomite-rich intervals are of Burdigalian age and correlate with the Danish Dany Formation (Rasmussen et al., in press). The Bastrup For- mation and the lower and upper Odderup Formation, respectively, are defined onshore Denmark (see Fig. 3). The Nora and Odderup Formations correlate with the uppermost part of the Lark Formation defined in the Norwegian sector of the North Sea, while the Dany For- mation and the time-equivalent lithostratigraphic units defined for onshore Denmark correlate with the upper part of the Lark Formation (Rasmussen et al. in press; Fig. 3). The combination of biozones and additional events provides a stratigraphic subdivision of the diatomite- rich Lower to Middle Miocene succession in the order of 5–15 m in the fully cored 2/11–12S well, somewhat less in the 2/8–G10A well (due to the core gap) and in the non-cored wells. This extremely high-resolution subdi- vision forms a solid basis for improved reservoir evalu- ation. No major hiatuses are recognised, though there appears to be a potential repeated section in the upper- most part of the core in the 2/8–G10A well. 11. Discussion 11.1. Improvements to regional and local biostratigraphy The multidisciplinary biostratigraphy, with numerous microfossil events presented in this study, is useful for reservoir subdivision and reservoir characterisa- tion in the Valhall–Hod area and for local and regional correlation. Dinocysts were found in all samples in this study. The zonation of Dybkjær & Piasecki (2010) was used suc- cessfully. All the events found in their study and 15 new dinocyst events are recognised. Fewer events are found in the interval where the highest content of silica occurs in the Nora Formation equivalent (L. truncatum Zone). The dinocyst study of the Valhall–Hod area has resulted in the improvement of the zonation of Dybkjær & Pias- ecki (2010). The microfossil zonations of King (1989, 2016) are applied to the studied sections. Twenty-five microfos- sil events are recognised in this study (Fig. 8 and Sup- plementary File S13), and most are concentrated in the upper part of Zone NSB10 (NS35b-c) and in NSB11 (NS36a-b). Nannofossils are not commonly used for biostratig- raphy in the North Sea Miocene interval due to the lack of calcareous sediments at this stratigraphic level and the successful application of dinocysts and microfossils. However, the global zonation of Martini (1971) is success- fully applied to the low-resolution nannofossil study and supports the biostratigraphy based on other microfossil groups. Seventeen nannofossil events are recognised in this study that have correlation potential (Fig. 7). Most of these nannofossil events are documented from the North Sea area for the first time in this study. They have been previously described from the Mediterranean area and the low latitudes (Young 1998) and the Gulf of Mex- ico (De Kaenel et al. 2017; Browning et al. 2017; Boesiger et al. 2017) and are potentially important for correlation beyond the North Sea Basin. The detailed study of siliceous microfossils from the fine fraction of the cored wells 2/8–G10A (Valhall Field) and 2/11–12S (Hod Field) has resulted in the recognition of a series of diatom and silicoflagellate events that correlate between the two cored wells situated on two neighbour- ing structures. Many of these siliceous microfossil events https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 25 of 30 GEUSBULLETIN.ORG are also found in nearby non-cored wells: 2/8–N4, 2/8–V6 and 2/8–8 (Valhall) and 2/11–1 (in the saddle between the Valhall and Hod fields). This highlights the diatom correla- tion potential in a local context and that they could form the basis of a North Sea siliceous microfossil zonation at a later stage. While not all the diatom species that mark the tops and bases of the zones of Schrader & Fenner (1976) are found in this study, the diatom assemblages from the Valhall–Hod area compare well with those from the Norwegian Sea. The diatom biostratigraphy and 20 diatom events described in this study are useful for the subdivision of the diatomite or silica-rich Nora Formation equivalent and Dany Formation equivalent (uppermost part of the Lark Formation) and potentially a valuable tool for hydrocarbon reservoir characterisation. The dia- tom biostratigraphy can potentially be used to correlate regionally, from the North Sea farther north into the Nor- wegian Sea. This is the first time, potentially, that siliceous microfossils have been used to correlate within the Mio- cene succession of the North Sea Basin and regionally. Silicoflagellate identification allowed the recogni- tion of two zones of Locker & Martini (1989) and five stratigraphically significant events are recognised in this study. Delicate silicoflagellate skeletons in our samples were often fragmented, perhaps due to harsh conditions the samples were subjected to during preparation. We are presently testing a less destruc- tive method, which will probably benefit future silico- flagellate studies. Diatoms and silicoflagellate biostratigraphic events have been discussed in detail in this study. Other sili- ceous components (e.g. sponge spicules, radiolaria and ebridians) are also prevalent in parts of the Lower to Middle Miocene ‘diatomite’ successions. Further detailed studies concentrating on the individual sili- ceous elements, their relative abundances and palae- oecology are currently underway, which will hopefully shed light on the conditions and driving forces neces- sary for mass siliceous microfossil production, which, in turn, will potentially aid prediction of siliceous res- ervoir properties and reservoir correlation across the Valhall–Hod area. For the Early to Middle Miocene diatomite-rich inter- val of the Valhall–Hod area, a combination of dinocyst, microfossil and diatom biostratigraphy appears to be particularly promising for detailed subdivision of the siliceous reservoir interval (Fig. 8 and Supplementary File S13). 11.2. Sequence stratigraphic and lithostratigraphic correlation This robust biostratigraphic framework produced in this study allows correlation of the sequence stratigra- phy defined onshore Denmark to the central parts of the North Sea Basin as presented in Rasmussen (2004, 2017) and Dybkjær et al. (2021) and also to the new Dan- ish offshore lithostratigraphy for the Neogene (Rasmus- sen et al., in press). 11.3. Age model A solid age model for the Miocene does not exist for the North Sea area. However, steps have been taken to attain this goal. Eidvin et  al. (2014b) used Sr iso- topes to analyse the whole of the Miocene section from samples from Jylland, onshore Denmark. Results from that study showed that Sr dating of samples from the Lower Miocene and lower Middle Miocene succes- sion supports the datings of the dinocyst zonation of Dybkjær & Piasecki (2010). However, in the upper Mid- dle Miocene and Upper Miocene parts of the section, the Sr ages are too old compared with ages based on dinocysts and Bolboforma. The authors of these publi- cations agree that there are discrepancies with the Sr dating. The lack of a robust age model for the Miocene of the North Sea area is also a major reason for the use of a selection of vintage, but strongly reliable, biostra- tigraphic zonations in this study. We considered it important to correlate fossil zones and events, rather than try to tackle the chronostratigraphic problem. The five biostratigraphic zonation schemes in Fig. 4, one for each fossil discipline, are correlated with the chronos- tratigraphic time scale that was relevant at the time of publication of the zonation. Our study has provided a direct, reliable correlation for the Lower and Middle Miocene succession of the Valhall–Hod area, based on five fossil groups using a series of events recognised in a selection of closely spaced wells, mostly on the same samples. We acknowledge the value of the Neogene times- cale of Raffi et al. (2020), which contains dinocyst, for- aminifera, calcareous nannofossil and diatom data and events from a multitude of global localities. However, our study is centred in the North Sea area, which was a semi-enclosed basin during the Miocene period. Mio- cene North Sea microfossil assemblages and events share more affinities to northern mid- to high-latitude and boreal assemblages than those from the low lati- tude and tropical locations cited in Raffi et  al. (2020). We conclude that a reliable and robust palaeomagnetic time frame for the North Sea area would be valuable for future stratigraphic studies. 11.4. Palaeoclimate The studied interval in this project includes the MCO (c.17–13 Ma; e.g. Larsson et  al. 2011; Herbert et  al. 2020; Sliwinska et al. 2024) and the MMCT (c. 14.7–13–8 https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 26 of 30 GEUSBULLETIN.ORG Ma; Flower & Kennett 1994). The high-resolution bio- stratigraphic framework thus forms a good basis for future studies of the climatic and palaeoenvironmental changes during these important time intervals. 12. Conclusions A new multidisciplinary biostratigraphic framework is established using core samples from the Lower-Middle Miocene succession of the 2/8–G10A and 2/11–12S wells from the Valhall and Hod Fields, respectively. The core study provides a correlation using five bio- stratigraphic disciplines: dinocysts, foraminifera, cal- careous nannofossils, diatoms and silicoflagellates. The correlation is based mostly on the same samples, providing a unique and exceptionally detailed, robust framework. This framework was successfully tested on the equiv- alent chronostratigraphic level of the 2/8–N4, 2/8–V6, 2/8–8 (Valhall Field) and 2/11–1 (in the saddle between the Valhall and Hod fields) wells based on ditch cutting samples. The studied interval spans the Chiropteridium galea Zone to the Hystrichosphaeropsis obscura dinocyst Zone (Dybkjær & Piasecki 2010), the NSB9 (NS34) to NSB12c (NS38–39) microfossil zones (King 1989, 2016), nanno- fossil zones NN3–NN6 (Martini 1971), the Rhizosolenia norwegica–Coscinodiscus vigilans–Nitzschia maleinterpre- taria joint PRZ to the Denticulopsis hyalina PRZ (diatoms, Schrader & Fenner 1976) and the lower and upper Cor- bisema triacantha silicoflagellates Zones (Locker & Mar- tini 1989). Twenty-seven dinocyst events, 25 microfossil events, 17 nannofossil events, 20 diatom events and 5 silicofla- gellate events are recognised for the studied interval. The new dinocyst, microfossil, nannofossil and dia- tom events (bases, tops and occurrences) from this study are successfully used to correlate on and between the Valhall Field and Hod Field and can be used to sup- plement published zonation schemes and events. The successful application of the new framework in the Val- hall–Hod area implies that it could also be valuable in a more regional context. A suite of diatom events is recognised for the Lower and Middle Miocene succession of the Valhall–Hod area and potentially forms the basis for a diatom zonation for the North Sea Basin. To the best of our knowledge, this diatom study (fine fraction) is the first of its kind cover- ing the Early and Middle Miocene interval of the North Sea and is potentially useful for regional correlation and local reservoir characterisation. Our low-resolution calcareous nannofossil study of the Valhall–Hod part of the North Sea Basin recognises events described from low-latitude and tropical locations. The siliceous or diatomite-rich reservoir interval has been dated, and a detailed (5–15 m interval) biostrati- graphic subdivision is provided. The detailed biostra- tigraphy has enabled the correlation of the sequence stratigraphic surfaces and lithostratigraphic units defined in the Danish sector to the southern Norwegian sector. Acknowledgements Aker BP and Pandion Energy are acknowledged for their initiation and funding of the study, as well as for their valuable contributions through discussions and the exchange of ideas throughout the project’s dura- tion. Special thanks go to GEUS laboratory technicians Annette Ryge and Charlotte Olsen for processing a huge number of samples. Jacob Lind Bendtsen is thanked for his unwavering patience when preparing the figures. Reviewers Haydon Bailey and Erik Anthonissen are thanked for their comments, which helped to develop the manuscript for publication. Additional Information Funding statement Aker BP and Pandion Energy funded this study. Author contributions ES and KD: Conceptualisation, writing – original draft, writing – review and editing. ES: Biostratigraphy (microfossils, calcareous nannofos- sils, diatoms, silicoflagellates). KD: Biostratigraphy (dinocysts). ESR: Lithostratigraphy, sequence stratigraphy. MO: Initial diatom supervi- sion and biostratigraphy (diatoms). Competing interests The authors declare no competing interests. Additional files Thirteen supplementary files are available at https://doi.org/10.22008/ FK2/VLO4LN: Supplementary Files S1–S6.pdf contain Fig. S1 Biostratigraphic sum- mary diagram for the 2/8–G10A well, Fig. S2 Biostratigraphic sum- mary diagram for the 2/11–12S well, Fig. S3 Biostratigraphic summary diagram for the 2/8–N4 well, Fig. S4 Biostratigraphic summary dia- gram for the 2/8–8 well, Fig. S5. Biostratigraphic summary diagram for the 2/8–V6 well and Fig. S6 Biostratigraphic summary diagram for the 2/11–1 well. Supplementary Files S7–S12.pdf include correlation lines Fig. S7 Biostratigraphic summary diagram for the 2/8–G10A well, Fig. S8 Biostratigraphic summary diagram for the 2/11–12S well, Fig. S9 Bio- stratigraphic summary diagram for the 2/8–N4 well, Fig. S10 Biostrati- graphic summary diagram for the 2/8–8 well, Fig. S11 Biostratigraphic summary diagram for the 2/8–V6 well and Fig. S12 Biostratigraphic summary diagram for the 2/11–1 well. Supplementary File S13.pdf contains Fig. S13 Multidisciplinary bio- stratigraphic correlation diagram (full size) of the six studied wells using dinocyst, microfossil and diatom events. References Amigo, A.E. 1999: Miocene silicoflagellate stratigraphy: Iceland and Rock- all Plateaus. In: Raymo, M.E. et al. (eds): Proceedings of the Ocean Drilling Program. Scientific Results 162, 1–19. https://doi.org/10.2973/odp.proc. sr.162.1999 Anthonissen, E.D. 2012: A new Miocene biostratigraphy for the north- eastern North Atlantic: an integrated foraminiferal, bolboformid, dinoflagellate and diatom zonation. Newsletters on Stratigraphy 45(3), 281–307. https://doi.org/10.1127/0078-0421/2012/0025 Baldauf, J.G. 1985: Cenozoic diatom biostratigraphy and paleocean- ography of the Rockall Plateau region, North Atlantic, Deep Sea https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ https://doi.org/10.22008/FK2/VLO4LN https://doi.org/10.22008/FK2/VLO4LN https://doi.org/10.2973/odp.proc.sr.162.1999 https://doi.org/10.2973/odp.proc.sr.162.1999 https://doi.org/10.1127/0078-0421/2012/0025 Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 27 of 30 GEUSBULLETIN.ORG Drilling Project Leg 81. In: Roberts, D.G. et al. (eds): Initial reports of the deep sea drilling project 81, 439–479. https://doi.org/10.2973/dsdp. proc.81.107.1984 Barron, J.A. 1985: Miocene to Holocene planktic diatoms. In: Bolli, H.M. et  al. (eds): Plankton Stratigraphy, 763–809. Cambridge University Press. https://doi.org/10.1017/s0016756800035214 Bergen, J.A., De Kaenel, E., Blair, S.A., Boesiger, T.M. & Browning, E. 2017: Oligocene-Pliocene taxonomy and stratigraphy of the genus Spheno- lithus in the circum North Atlantic Basin: Gulf of Mexico and ODP Leg 154. Journal of Nannoplankton Research 37(2–3), 77–112. https://doi. org/10.58998/jnr2016 Berggren, W.A. 1972: A Cenozoic time-scale, some implications for regional geology and paleobiogeography. Lethaia 5, 195–215. https://doi.org/10.1111/j.1502-3931.1972.tb00852.x Berggren, W.A., Kent, D.V., Flynn, J.J. & Van Couvering, J.A. 1985: Cenozoic geochronology. Geological Society of America Bulletin 96, 1407–1418. https://doi.org/10.1130/0016-7606(1985)96%3C1407:cg%3E2.0.co;2 Biljsma, S. 1981: Fluvial sedimentation from the Fennoscandian area into the North-West European Basin during the Late Cenozoic. In: Van Loon, A.J. (ed.): Quaternary geology: a farewell to A.J. Wiggers. Geolo- gie en Mijnbouw 60, 337–345. Boesiger, T.M., De Kaenel, E., Bergen, J.A., Browning, E. & Blair, S.A. 2017: Oligocene to Pleistocene taxonomy and stratigraphy of the genus Helicosphaera and other placolith taxa in the circum North Atlantic Basin. Journal of Nannoplankton Research 37(2–3), 145–175. https:// doi.org/10.58998/jnr2021 Bown, P. & Young, J. 1998: Techniques. In: Bown, P.R. (ed.): Calcareous Nannofossil biostratigraphy. British Micropalaeontological Soci- ety Series, 16–28. Chapman & Hall/Kluwer Academic. https://doi. org/10.1007/978-94-011-4902-0_2 Browning, E., Bergen, J., Blair, S., Boesiger, T. & de Kaenel, E. 2017: Late Miocene to Late Pliocene taxonomy and stratigraphy in the circum North Atlantic Basin: gulf of Mexico and ODP Leg 154. Journal of Nanno- plankton Research 37(2–3), 189–214. https://doi.org/10.58998/jnr2037 Ciesielski, P.F., Hasson, P. & Turner, J.W. 1989: The stratigraphy of neogene silicoflagellates from the Norwegian Sea, ODP Leg 104. In: Eldholm, O. et  al. (eds): Proceedings of the ocean drilling program. Scientific Results 104, 497–525. https://doi.org/10.2973/odp.proc. sr.104.164.1989 De Kaenel, E., Bergen, J.A., Browning, E., Blair, S.A. & Boesiger, T.M. 2017: Uppermost oligocene to middle Miocene discoaster and catin- aster taxonomy and stratigraphy in the circum North Atlantic Basin. Journal of Nannoplankton Research 37(2–3), 215–244. https://doi. org/10.58998/jnr2077 De Schepper, S., Head, M. & Louwye, S. 2004: New dinoflagellate cyst and Incertae Sedis taxa from the Pliocene of Northern Belgium, southern North Sea Basin. Journal of Paleontology 78(4), 625–644. https://doi. org/10.1666/0022-3360(2004)078%3C0625:ndcais%3E2.0.co;2 De Schepper, S., Head, M.J. & Louwye, S. 2009: Pliocene dinoflagel- late cyst stratigraphy, palaeoecology and sequence stratigraphy of the Tunnel-Canal Dock, Belgium. Geological Magazine 146, 92–112. https://doi.org/10.1017/s0016756808005438 De Schepper, S. & Mangerud, G. 2017: Age and palaeoenvironment of the Utsira Formation in the northern North Sea based on marine palynology. Norwegian Journal of Geology 97, 255–276. https://doi. org/10.17850/njg97-4-04 De Verteuil, L. 1997: Palynological delineation and regional correla- tion of Lower through Upper Miocene sequences in the Cape May and Atlantic City Boreholes, New Jersey Coastal Plain. In: Miller, K.G. & Snyder, S.W. (eds): Proceedings of the ocean drilling program, Scientific Results 150X, 129–145. https://doi.org/10.2973/odp.proc. sr.150x.310.1997 De Verteuil, L. & Norris, G. 1996: Miocene dinoflagellate stratigraphy and systematics of Maryland and Virginia. Micropaleontology 42(Suppl), 172 pp. https://doi.org/10.2307/1485926 Doppert, J.W.C. 1980: Lithostratigraphy and biostratigraphy of Marine Neogene deposits in the Netherlands. Mededelingen Rijks Geolo- gische Dienst 32(16), 255–311. Doppert, J.W.C., Laga, P.G. & De Meuter, F.J. 1979: Correlation of the biostratigraphy of marine Neogene deposits, based on benthonic foraminifera, established in Belgium and The Netherlands. Med- edelingen Rijks Geologische Dienst 31(1), 1–8. Dybkjær, K. 2004a: Dinocyst stratigraphy and palynofacies studies used for refining a sequence stratigraphic model – uppermost Oligocene to Lower Miocene, Jylland, Denmark. Review of Palaeobotany and Pal- ynology 131, 201–249. https://doi.org/10.1016/j.revpalbo.2004.03.006 Dybkjær, K. 2004b: Morphological and abundance variations in Homo- tryblium-cyst assemblages related to depositional environments; uppermost Oligocene–Lower Miocene, Jylland, Denmark. Palaeoge- ography, Palaeoclimatology, Palaeoecology 206, 41–58. https://doi. org/10.1016/j.palaeo.2003.12.021 Dybkjær, K. & Rasmussen, E.S. 2000: Palynological dating of the Oli- gocene–Miocene successions in the Lille Bælt area, Denmark. Bul- letin of the Geological Society of Denmark 47, 87–103. https://doi. org/10.37570/bgsd-2000-47-07 Dybkjær, K. & Rasmussen, E.S. 2007: Dinocyst stratigraphy in an expanded Oligocene–Miocene boundary section in the eastern North Sea Basin (the Frida-1 well, Denmark) and correlation from basinal to marginal areas. Journal of Micropaleontology 26, 1–17. https://doi. org/10.1144/jm.26.1.1 Dybkjær, K. & Piasecki, S. 2010: Neogene dinocyst zonation in the east- ern North Sea Basin, Denmark. Review of Palaeobotany and Palynol- ogy 161, 1–29. https://doi.org/10.1016/j.revpalbo.2010.02.005 Dybkjær, K., King, C. & Sheldon, E. 2012: Identification and character- isation of the Oligocene–Miocene boundary (base Neogene) in the eastern North Sea Basin – based on dinocyst stratigraphy, micro- palaeontology and δ13C-isotope data. Palaeogeography, Palae- oclimatology, Palaeoecology 363, 11–22. https://doi.org/10.1016/j. palaeo.2012.08.007 Dybkjær, K., Rasmussen, E.S., Śliwińska, K.K., Esbensen, K.H. & Mathiesen, A. 2019: A palynofacies study of past fluvio-deltaic and shelf environments, the Oligocene-Miocene succession, North Sea Basin: a reference data set for similar Cenozoic systems. Marine and Petroleum Geology 100, 111–147. https://doi.org/10.1016/j. marpetgeo.2018.08.012 Dybkjær, K., Rasmussen, E.S., Eidvin, T., Grøsfjeld, K., Riis, F., Piasecki, S. & Śliwińska, K.K. 2021: A new stratigraphic framework for the Miocene – lower Pliocene deposits offshore Scandinavia: a mul- tiscale approach. Geological Journal 56, 1699–1725. https://doi. org/10.1002/gj.3982 Dzinoridze, R.N., Jousé, A.P., Koroleva-Golikova, G.S., Kozlova, G.E., Nagaeva, G.S., Petraschvskaya, M.G. & Strelinikova, N.I. 1979: Diatom and radiolarian Cenozoic stratigraphy, Norwegian basin, DSDP Leg 38. In: Supko, P.R. et al. (eds): Initial Reports of the Deep Sea Drilling Proj- ect, Suppl. to Vols. 38, 39, 40 & 41, 289–427. https://doi.org/10.2973/ dsdp.proc.38394041s.119.1978 Eidvin, T., Koc, N., Smelrør, M. & Jansen, E. 1998: Biostratigraphical inves- tigation of borehole 6704/12-GB1 from the Gjallar Ridge on the Vøring Plateau. Report for the Seabed project. OD-98-22. Oljedirektoratet. Eidvin, T., Riis, F. & Rundberg, Y. 1999: Upper Cainozoic stratigraphy in the central North Sea (Ekofisk and Sleipner fields). Norsk Geologisk Tidsskrift 79, 97–127. https://doi.org/10.1080/002919699433843 Eidvin, T., Riis, F. & Rasmussen, E.S. 2014a: Oligocene to Lower Plio- cene deposits of the Norwegian continental shelf, Norwegian Sea, Svalbard; Denmark and their relation to the uplift of Fennoscandia: a synthesis. Marine and Petroleum Geology 56, 184–221. https://doi. org/10.1016/j.marpetgeo.2014.04.006 Eidvin, T., Ullmann, C.V., Dybkjær, K., Rasmussen, E.S. & Piasecki, S. 2014b: Discrepancy between Sr isotope and biostratigraphic datings of the upper middle and upper Miocene successions (eastern North Sea Basin, Denmark). Palaeogeography, Palaeoclimatology, Palaeo- ecology 411, 267–280. https://doi.org/10.1016/j.palaeo.2014.07.005 Eidvin, T. & Rundberg, Y. 2001: Late Cainozoic stratigraphy of the Tampen area (Snorre and Visund fields) in the northern North Sea, with emphasis on the chronology of early Neogene sands. Norsk Geologisk Tidsskrift 81, 119–160. Eidvin, T. & Rundberg, Y. 2007: Post-Eocene strata of the southern Viking Graben, northern North Sea; integrated biostratigraphic, strontium isotopic and lithostratigraphic study. Norwegian Journal of Geology 87, 391–450. https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ https://doi.org/10.2973/dsdp.proc.81.107.1984 https://doi.org/10.2973/dsdp.proc.81.107.1984 https://doi.org/10.1017/s0016756800035214 https://doi.org/10.58998/jnr2016 https://doi.org/10.58998/jnr2016 https://doi.org/10.1111/j.1502-3931.1972.tb00852.x https://doi.org/10.1130/0016-7606(1985)96%3C1407:cg%3E2.0.co;2 https://doi.org/10.58998/jnr2021 https://doi.org/10.58998/jnr2021 https://doi.org/10.1007/978-94-011-4902-0_2 https://doi.org/10.1007/978-94-011-4902-0_2 https://doi.org/10.58998/jnr2037 https://doi.org/10.2973/odp.proc.sr.104.164.1989 https://doi.org/10.2973/odp.proc.sr.104.164.1989 https://doi.org/10.58998/jnr2077 https://doi.org/10.58998/jnr2077 https://doi.org/10.1666/0022-3360(2004)078%3C0625:ndcais%3E2.0.co;2 https://doi.org/10.1666/0022-3360(2004)078%3C0625:ndcais%3E2.0.co;2 https://doi.org/10.1017/s0016756808005438 https://doi.org/10.17850/njg97-4-04 https://doi.org/10.17850/njg97-4-04 https://doi.org/10.2973/odp.proc.sr.150x.310.1997 https://doi.org/10.2973/odp.proc.sr.150x.310.1997 https://doi.org/10.2307/1485926 https://doi.org/10.1016/j.revpalbo.2004.03.006 https://doi.org/10.1016/j.palaeo.2003.12.021 https://doi.org/10.1016/j.palaeo.2003.12.021 https://doi.org/10.37570/bgsd-2000-47-07 https://doi.org/10.37570/bgsd-2000-47-07 https://doi.org/10.1144/jm.26.1.1 https://doi.org/10.1144/jm.26.1.1 https://doi.org/10.1016/j.revpalbo.2010.02.005 https://doi.org/10.1016/j.palaeo.2012.08.007 https://doi.org/10.1016/j.palaeo.2012.08.007 https://doi.org/10.1016/j.marpetgeo.2018.08.012 https://doi.org/10.1016/j.marpetgeo.2018.08.012 https://doi.org/10.1002/gj.3982 https://doi.org/10.1002/gj.3982 https://doi.org/10.2973/dsdp.proc.38394041s.119.1978 https://doi.org/10.2973/dsdp.proc.38394041s.119.1978 https://doi.org/10.1080/002919699433843 https://doi.org/10.1016/j.marpetgeo.2014.04.006 https://doi.org/10.1016/j.marpetgeo.2014.04.006 https://doi.org/10.1016/j.palaeo.2014.07.005 Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 28 of 30 GEUSBULLETIN.ORG Eidvin, T., Riis, F., Brekke, H. & Smelror, M. 2022: A revised lithostrati- graphic scheme for the Eocene to Pleistocene succession on the Norwegian Continental shelf. Norwegian Journal of Geology, Special Publication 1, 1–132. https://doi.org/10.17850/njgsp1 Flower, B.P. & Kennett, J.P. 1994: The middle Miocene climatic transition: East Antarctic ice sheet development, deep ocean circulation and global carbon cycling. Palaeogeography, Palaeoclimatology, Palaeo- ecology 108, 537–555. https://doi.org/10.1016/0031-0182(94)90251-8 Fox, L.R., Stukins, S., Hill, T. & Bailey, H.W. 2018: New species of Cenozoic benthic foraminifera from the former British Petroleum micropalae- ontology collection. Journal of Micropalaeontology 37(1), 11–16. https:// doi.org/10.5194/jm-37-11-2018 Gabrielsen, R.-H., Faleide, J.I., Pascal, C., Braathen, A., Nystuen, J.P., Etzelmuller, B. & O’Dennel, S. 2009: Latest Caledonian to present tectonomorphological development of southern Norway. Marine and Petroleum Geology 27, 709–723. https://doi.org/10.1016/j. marpetgeo.2009.06.004 Gartner, S. 1992: Miocene nannofossil chronology in the North Atlantic, DSDP Site 608. Marine Micropaleontology 18, 307–331. https://doi. org/10.1016/0377-8398(92)90045-l Gradstein, F.M. & Backstrom, S. 1996: Cainozoic biostratigraphy and palaeobathymetry, northern North Sea and Haltenbanken. Norsk Geologisk Tidsskrift 76, 3–32. Gradstein, F.M., Kaminski, M.A. & Berggren, W.A. 1988: Cenozoic fora- miniferal biostratigraphy of the central North Sea. Abhandlungen der Geologischen Bundesanstalt 41, 97–108. Grøsfjeld, K., Dybkjær, K., Eidvin, T., Riis, F., Rasmussen, E.S. & Knies, J. 2019: A Miocene age for the Molo Formation, Norwegian shelf off Vestfjorden, based on marine palynology. Norwegian Journal of Geol- ogy 99(3), 1–20. https://doi.org/10.17850/njg99-3-6 Haq, B.U., Hardenbol, J. & Vail, P.R. 1987: Chronology of fluctuating sea-levels since the Triassic. Science 235, 1156–1167. https://doi. org/10.1126/science.235.4793.1156 Head, M.J. 1996: Late Cenozoic dinoflagellates from the Royal Society Borehole at Ludham, Norfolk, Eastern England. Journal of Paleontol- ogy 70, 543–570. https://doi.org/10.1017/s0022336000023532 Herbert, T.D., Rose, R., Dybkjær, K., Rasmussen, E.S. & Śliwińska, K.K. 2020: Bihemispheric warming in the Miocene Climatic Optimum as seen from the Danish North Sea. Paleoceanog- raphy and Paleoclimatology 35, e2020PA003935. https://doi. org/10.1029/2020pa003935 Hilgen, F.J. et al. 2012: Chapter 29. The Neogene period. In: Gradstein, F.M. et al. (eds): The geologic time scale, 923–978. Elsevier. https://doi. org/10.1016/b978-0-444-59425-9.00029-9 Kaminski, M.A. & Gradstein, F.M. 2005: Atlas of Paleogene cosmo- politan deep-water agglutinated foraminifera. Grzybowski Foun- dation Special Publication 10, 547 pp. https://doi.org/10.2113/ gsmicropal.52.6.555 King, C. 1983: Cainozoic Micropalaeontological biostratigraphy of the North Sea. Report of the Institute of Geological Sciences 82(7), 1–40. King, C. 1989: Cenozoic of the North Sea, In: Jenkins, D.G. & Murray, J.W. (eds): Stratigraphical atlas of Fossil Foraminifera, 2nd ed., 418–489. Ellis Horwood Ltd. https://doi.org/10.1017/s0016756800014655 King, C. 2016: A revised correlation of Tertiary rocks in the British Isles and adjacent areas of NW Europe. In: Gale, A.S. & Barry, T.L. (eds): Geological Society Special Report 27, 719 pp. https://doi.org/10.1144/ sr27 Knox, R. et  al. 2010: Cenozoic. In: Doornenbal, J.C. & Stevenson, A.G. (eds): Petroleum geological atlas of the Southern Permian Basin Area, 210–323. EAGE Publications. Koç, N. & Scherer, P.R. 1996: Neogene diatom biostratigraphy of the Ice- land Sea site 9071. In: Thiede, J. et al. (eds): Proceedings of the ocean drilling program. Scientific Results 151, 61–74. https://doi.org/10.2973/ odp.proc.sr.151.108.1996 Koch, B.E. 1989: Geology of the Søby-Fasterholt area. Danmarks Geolo- giske Undersøgelse Serie A 22, 171 pp. https://doi.org/10.34194/seriea. v22.7042 Kuhlemann, J. 2007: Paleogeographic and paleotopographic evolution of the Swiss and Eastern Alps since the Oligocene. Global and Planetary Change 58, 224–236. https://doi.org/10.1016/j.gloplacha.2007.03.007 Köthe, A. 2012: A revised Cenozoic dinoflagellate cyst and calcareous nannoplankton zonation for the German sector of the southeastern North Sea Basin. Newsletters on Stratigraphy 45(3), 189–220. https:// doi.org/10.1127/0078-0421/2012/0021 Köthe, A. & Piesker, B. 2007: Stratigraphic distribution of Paleogene and Miocene dinocysts in Germany. Revue de Paléobiologie 26, 1–39. Larsson, L.M., Dybkjær, K., Rasmussen, E.S., Piasecki, S., Utescher, T. & Vajda, V. 2011: Miocene climate evolution of northern Europe: a palynological investigation from Denmark. Palaegeography, Palae- oclimatology, Palaeoecology 309, 161–175. https://doi.org/10.1016/j. palaeo.2011.05.003 Laursen, G. & Kristoffersen, F.N. 1999: Detailed foraminiferal biostratig- raphy of Miocene formations in Denmark. Contributions to Tertiary and Quaternary Geology 36, 73–107. Locker, S. & Martini, E. 1989: Cenozoic Silicoflagellates, Ebridians and Actiniscidians from the Vøring Plateau (ODP Leg 104). In: Eld- holm, O. et  al. (eds): Proceedings of the ocean drilling program. Scientific Results 104, 543–585. https://doi.org/10.2973/odp.proc. sr.104.204.1989 Lourens, L., Hilgen, F., Shackleton, N.J., Laskar, J. & Wilson, D. 2004: The neogene period. In: Gradstein, F. et  al. (eds): A geologic time scale 2004, 409–440. Cambridge University Press. https://doi.org/10.1017/ cbo9780511536045.022 Louwye, S. 2002: Dinoflagellate cyst biostratigraphy of the upper Mio- cene Deurne Sands (Diest Formation) of northern Belgium, southern North Sea Basin. Geological Journal 37, 55–67. https://doi.org/10.1002/ gj.900 Louwye, S., De Coninck, J. & Verniers, J. 1999: Dinoflagellate cyst stratig- raphy and depositional history of Miocene and Lower Pliocene for- mations in northern Belgium (southern North Sea Basin). Geologie en Mijnbouw 78, 31–46. Louwye, S., De Schepper, S., Laga, P. & Vandenberghe, N. 2007: The upper Miocene of the southern North Sea Basin (northern Belgium): a palaeoenvironmental and stratigraphical reconstruction using dinoflagellate cysts. Geological Magazine 144, 33–52. https://doi. org/10.1017/s0016756806002627 Louwye, S. & Laga, P. 2008: Dinoflagellate cyst stratigraphy and palae- oenvironment of the marginal marine middle and upper Miocene of the eastern Campine area, northern Belgium (southern North Sea Basin). Geological Journal 43, 75–94. https://doi.org/10.1002/gj.1103 Louwye, S. & De Schepper, S. 2010: The Miocene–Pliocene hiatus in the southern North Sea Basin (northern Belgium) revealed by dino- flagellate cysts. Geological Magazine 147(5), 760–776. https://doi. org/10.1017/S0016756810000191 Løseth, H. & Henriksen, S. 2005: A Middle to Late Miocene compres- sion phase along the Norwegian passive margin. In: Doré, A.G. & Vinding, B.A. (eds): Petroleum geology: Northwest Europe and global perspectives – Proceedings of the 6th Petroleum Geology Conference 6, 845–859. Geological Society, London. https://doi. org/10.1144/0060845 Løseth, H., Kyrkjebø, R., Hilde, E., Wild, J.W. & Bunkholt, H. 2017: 500 m of rapid base level rise along an inner passive margin. Seismic observations from the Pliocene Molo Formation, mid Norway. Marine and Petroleum Geology 86, 268–287. https://doi.org/10.1016/j. marpetgeo.2017.05.039 Martini, E. 1971: Standard tertiary and quaternary calcareous nanno- plankton zonation. In: Farinacci, A. (ed.): Proceedings of the Second Planktonic Conference Roma 1970 2, 739–785. Tecnoscienza. Martini, E. & Muller, C. 1976: Eocene to Pleistocene silicoflagellates from the Norwegian-Greenland Sea (DSDP Leg 38). Initial Reports of the Deep Sea Drilling Project 38, 857–895. https://doi.org/10.2973/dsdp. proc.38.128.1976 McCartney, K., Witkowski, J. & Harwood, D. 2011: Late Cretaceous silico- flagellate taxonomy and biostratigraphy of the Arctic margin, North- west Territories, Canada. Micropaleontology 57(1), 61–86. https://doi. org/10.47894/mpal.57.1.03 Mitlehner, A.G. 2019: Species of the diatom taxa Aulacodiscus and Trin- acria with biostratigraphic utility in Palaeogene and Neogene North Sea sediments. Journal of Micropalaeontology 38(1), 67–81. https:// doi.org/10.5194/jm-38-67-2019 https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ https://doi.org/10.17850/njgsp1 https://doi.org/10.1016/0031-0182(94)90251-8 https://doi.org/10.5194/jm-37-11-2018 https://doi.org/10.5194/jm-37-11-2018 https://doi.org/10.1016/j.marpetgeo.2009.06.004 https://doi.org/10.1016/j.marpetgeo.2009.06.004 https://doi.org/10.1016/0377-8398(92)90045-l https://doi.org/10.1016/0377-8398(92)90045-l https://doi.org/10.17850/njg99-3-6 https://doi.org/10.1126/science.235.4793.1156 https://doi.org/10.1126/science.235.4793.1156 https://doi.org/10.1017/s0022336000023532 https://doi.org/10.1029/2020pa003935 https://doi.org/10.1029/2020pa003935 https://doi.org/10.1016/b978-0-444-59425-9.00029-9 https://doi.org/10.1016/b978-0-444-59425-9.00029-9 https://doi.org/10.2113/gsmicropal.52.6.555 https://doi.org/10.2113/gsmicropal.52.6.555 https://doi.org/10.1017/s0016756800014655 https://doi.org/10.1144/sr27 https://doi.org/10.1144/sr27 https://doi.org/10.2973/odp.proc.sr.151.108.1996 https://doi.org/10.2973/odp.proc.sr.151.108.1996 https://doi.org/10.34194/seriea.v22.7042 https://doi.org/10.34194/seriea.v22.7042 https://doi.org/10.1016/j.gloplacha.2007.03.007 https://doi.org/10.1127/0078-0421/2012/0021 https://doi.org/10.1127/0078-0421/2012/0021 https://doi.org/10.1016/j.palaeo.2011.05.003 https://doi.org/10.1016/j.palaeo.2011.05.003 https://doi.org/10.2973/odp.proc.sr.104.204.1989 https://doi.org/10.2973/odp.proc.sr.104.204.1989 https://doi.org/10.1017/cbo9780511536045.022 https://doi.org/10.1017/cbo9780511536045.022 https://doi.org/10.1002/gj.900 https://doi.org/10.1002/gj.900 https://doi.org/10.1017/s0016756806002627 https://doi.org/10.1017/s0016756806002627 https://doi.org/10.1002/gj.1103 https://doi.org/10.1017/S0016756810000191 https://doi.org/10.1017/S0016756810000191 https://doi.org/10.1144/0060845 https://doi.org/10.1144/0060845 https://doi.org/10.1016/j.marpetgeo.2017.05.039 https://doi.org/10.1016/j.marpetgeo.2017.05.039 https://doi.org/10.2973/dsdp.proc.38.128.1976 https://doi.org/10.2973/dsdp.proc.38.128.1976 https://doi.org/10.47894/mpal.57.1.03 https://doi.org/10.47894/mpal.57.1.03 https://doi.org/10.5194/jm-38-67-2019 https://doi.org/10.5194/jm-38-67-2019 Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 29 of 30 GEUSBULLETIN.ORG Müller, C. 1976: Tertiary and Quaternary Calcareous nannoplankton in the Norwegian-Greenland Sea, DSDP Leg 38. Initial Reports of the Deep Sea Drilling Project 38, 823–841. https://doi.org/10.2973/dsdp. proc.38.126.1976 Munsterman, D.K. & Brinkhuis, H. 2004: A southern North Sea Miocene dinoflagellate cyst zonation. Netherlands Journal of Geosciences 83(4), 267–285. https://doi.org/10.1017/s0016774600020369 Munsterman, D.K., ten Veen, J.H., Menkovic, A., Deckers, J., Witmans, N., Verhaegen, J., Kerstholt-Boegehold, S.J., van de Ven, T. & Busschers, F.S. (2019): An updated and revised stratigraphic framework for the Miocene and earliest Pliocene strata of the Roer Valley Graben and adjacent blocks. Netherlands Journal of Geosciences 98,1–23. https:// doi.org/10.1017/njg.2019.10 Oszczypko, N. 2006: Late Jurassic-Miocene evolution of the Outer Carpa- tian fold-and-thrust belt and its foredeep basin (Western Carpatians, Poland). Geological Quarterly 50, 169–194. Overeem, I., Weltje, G.J., Bishop-Kay, C. & Kroonenberg, S.B. 2001: The Late Cenozoic Eridanos delta system in the Southern North Sea Basin: a climate signal in sediment supply? Basin Research 13, 293–312. https://doi.org/10.1046/j.1365-2117.2001.00151.x Perch-Nielsen, K. 1985: Silicoflagellates. In: Bolli, H.M. et  al. (eds.): Plankton Stratigraphy, 811–846. Cambridge University Press. https:// doi.org/10.1017/s0016756800035214 Piasecki, S. 1980: Dinoflagellate cyst stratigraphy of the Miocene Hodde and Gram Formations, Denmark. Bulletin of the Geological Society of Denmark 29, 53–76. https://doi.org/10.37570/bgsd-1980-29-03 Powell, A.J. 1992: Dinoflagellate cysts of the Tertiary System. In: Powell, A.J. (ed.): A stratigraphic index of dinoflagellate cysts, 155–251. Chap- man & Hall. https://doi.org/10.1007/978-94-011-2386-0_4 Raffi, I., Wade, B.S. & Pälike, H. 2020: The neogene period. In: Gradstein, F. et  al. (eds): Geological Time Scale 2020 2, 1141–1215. https://doi. org/10.1016/b978-0-12-824360-2.00029-2 Rasmussen, E.S. 1996: Sequence stratigraphic subdivision of the Oligo- cene and Miocene succession in South Jutland. Bulletin of the Geo- logical Society of Denmark 43, 143–155. https://doi.org/10.37570/ bgsd-1996-43-14 Rasmussen, E.S. 2004a: The interplay between true eustatic sea- level changes, tectonics, and climatical changes: What is the dominating factor in sequence formation of the Upper Oligocene – Miocene succession in the eastern North Sea Basin, Denmark? Global and Planetary Change 41, 15–30. https://doi.org/10.1016/j. gloplacha.2003.08.004 Rasmussen, E.S. 2004b: Stratigraphy and depositional evolution of the uppermost Oligocene – Miocene succession in Denmark. Bul- letin of the Geological Society of Denmark 51, 89–109. https://doi. org/10.37570/bgsd-2004-51-07 Rasmussen, E.S. 2009: Neogene inversion of the north-eastern North Sea. Tectonophysics 465, 84–97. https://doi.org/10.1016/j. tecto.2008.10.025 Rasmussen, E.S. 2013: Cenozoic structures in the North Sea Basin – a case for salt tectonics: discussion. Tectonophysics 601, 226–233. https://doi.org/10.1016/j.tecto.2012.10.038 Rasmussen, E.S. 2017: Sedimentology and sequence stratigraphy of the uppermost upper Oligocene – Miocene fluvio-deltaic system in the eastern North Sea Basin: the influence of tectonism, eustacy and cli- mate. 67 pp + 15 Papers. Unpublished doctoral thesis, University of Copenhagen. Rasmussen, E.S. & Dybkjær, K. 2014: Patterns of Cenozoic sediment flux from western Scandinavia: discussion. Basin Research 26, 338–346. https://doi.org/10.1111/bre.12024 Rasmussen, E.S., Heilmann-Clausen, C., Waagstein, R. & Eidvin, T. 2008: Tertiary of Norden. Episodes 2008 31, 66–72. https://doi.org/10.18814/ epiiugs/2008/v31i1/010 Rasmussen, E.S., Dybkjær K. & Piasecki, S. 2010: Lithostratigraphy of the upper Oligocene – Miocene succession in Denmark. Geological Survey of Denmark and Greenland Bulletin 22, 92 pp. https://doi. org/10.34194/geusb.v22.4733 Rasmussen, E.S., Dybkjær, K., Toft, J.C., Nielsen, O.B., Sheldon, E. & Mørk, F. in press: Lithostratigraphy of the Neogene succession of the Danish North Sea. GEUS Bulletin 61. https://doi.org/10.34194/0nydbt40 Rundberg, Y. & Eidvin, T. 2005: Controls on depositional history and architecture of Oligocene-Miocene deposition, northern North Sea basin. In: Wandas, B. (ed.): Onshore-Offshore relationships on the North Atlantic Margin. Norwegian Petroleum Society Special Publica- tion 12, 207–239. https://doi.org/10.1016/s0928-8937(05)80050-5 Schiøler, P. 2005: Dinoflagellate cysts and acritarchs from the Oligocene – lower Miocene interval of the Alma-1X well, Danish North Sea. Jour- nal of Micropalaeontology 24, 1–37. https://doi.org/10.1144/jm.24.1.1 Schoonman, C.M., White, N.J. & Pritchard, D. 2017: Radial viscous fin- gering of hot asthenosphere within the Icelandic plume beneath the North Atlantic Ocean. Earth and Planetary Science Letters 468, 51–61. https://doi.org/10.1016/j.epsl.2017.03.036 Schrader, H.-J. & Fenner, J. 1976: Norwegian Sea Cenozoic diatom bio- stratigraphy and taxonomy. In: Talwani, M. et al. (eds): Initial Reports of the Deep Sea Drilling Project 38, 921–1099. https://doi.org/10.2973/ dsdp.proc.38.130.1976 Sheldon, E., Rasmussen, E.S., Dybkjær, K., Eidvin, T., Riis, F. & Weibel, R. 2018: Miocene oil-bearing diatom ooze from the North Sea. Geologi- cal Survey of Denmark and Greenland Bulletin 41, 29–32. https://doi. org/10.34194/geusb.v41.4335 Śliwinska, K.K., Dybkjær, K., Schoon, P.L., Beyer, C., King, C., Schouten, S. & Nielsen, O.B. 2014: Paleoclimatic and paleoenvironmen- tal records of the Oligocene–Miocene transition, central Jylland, Denmark. Marine Geology 350, 1–15. https://doi.org/10.1016/j. margeo.2013.12.014 Sliwinska, K.K., Denk, T., Dybkjær, K., Fredborg, J.M., Lindström, S., Pias- ecki, S. & Rasmussen, E.S. 2024: Miocene vegetation and climate in the eastern North Sea Basin, onshore Denmark, compared to the pres- ent. GEUS Bulletin 57, 8365. https://doi.org/10.34194/geusb.v57.8365 Spiegler, D. 1999: Bolboforma Biostratigraphy from the Hatton-Rock- all Basin (North Atlantic). In: Raymo, M.E. et al. (eds): Proceedings of the Ocean Drilling Program, Scientific Results 162, 35–49. https://doi. org/10.2973/odp.proc.sr.162.013.1999 Spiegler D. & Von Daniels, C.H. 1991: Stratigraphic and taxonomic atlas of Bolboforma (Protophytes, Incertae sedis, Tertiary). Journal of Fora- miniferal Research 21(2), 126–158. https://doi.org/10.2113/gsjfr.21.2.126 Steinmetz, J.C. 1979: Calcareous nannofossils from the North Atlantic Ocean, Leg 49, Deep Sea Drilling Project. In: Luyendyk, B.P. et al. (eds): Initial Reports of the Deep Sea Drilling Project 49, 519–532. https://doi. org/10.2973/dsdp.proc.49.116.1979 Strauss, C. & Lund, J.J. 1992: A middle Miocene dinoflagellate cyst micro- flora from Papendorf near Hamburg, Germany. Mitteilungen Aus dem Geologisch-Paläontologischen Institut der Universität Hamburg 73, 159–189. Strauss, C., Lund, J.J. & Lund-Christensen, J. 2001: Miocene dinoflagellate cyst stratigraphy of the Nieder Octenhausen research borehole (NW Germany). Geologishe Jahrbuch Reihe A 152, 395–448. Suto, I. 2006: The explosive diversification of the diatom genus Chaeto- ceros across the Eocene/Oligocene and Oligocene/Miocene boundar- ies in the Norwegian Sea. Marine Micropaleontology 58(4), 259–269. https://doi.org/10.1016/j.marmicro.2005.11.004 Thyberg, B.I., Stabell, B., Faleide, J.I. & Bjørlykke, K. 1999: Upper Oligo- cene diatomaceous deposits in the northern North Sea – silica dia- genesis and paleogeographic implications. Norsk Geologisk Tidsskrift 79(1), 3–18. https://doi.org/10.1080/002919699433870 Thyberg, B.I., Jordt, H., Bjørlykke, K. & Faleide, J.I. 2000: Relationships between sequence stratigraphy, mineralogy and geochemistry in Cenozoic sediments of the northern North Sea. In: Nøttvedt, A. et al. (eds): Dynamics of the Norwegian Margin. Geological Society, Lon- don, Special Publications 167, 245–272. https://doi.org/10.1144/gsl. sp.2000.167.01.10 Utescher, T., Mosbrugger, V., Ivanov, D. & Dilcher, D.L. 2009: Pres- ent-day climatic equivalents of European Cenozoic climates. Earth and Planetary Science Letters 284, 544–552. https://doi.org/10.1016/j. epsl.2009.05.021 Von Daniels, C.H. & Spiegler, D. 1977: Uvigerinen (Foram.) im Neogen Nordwestdeutschlands (Das Nordwestdeutsch Tertiärbecken, Beitrag Nr. 23). Geologisches Jahrbuch A 40, 3–59. Williams, G.L., Brinkhuis, H., Pearce, M.A., Fensome, R.A. & Weegink, J.W. 2004: Southern Ocean and global dinoflagellate cyst events https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ https://doi.org/10.2973/dsdp.proc.38.126.1976 https://doi.org/10.2973/dsdp.proc.38.126.1976 https://doi.org/10.1017/s0016774600020369 https:// doi.org/10.1017/njg.2019.10 https://doi.org/10.1046/j.1365-2117.2001.00151.x https://doi.org/10.1017/s0016756800035214 https://doi.org/10.1017/s0016756800035214 https://doi.org/10.37570/bgsd-1980-29-03 https://doi.org/10.1007/978-94-011-2386-0_4 https://doi.org/10.1016/b978-0-12-824360-2.00029-2 https://doi.org/10.1016/b978-0-12-824360-2.00029-2 https://doi.org/10.37570/bgsd-1996-43-14 https://doi.org/10.37570/bgsd-1996-43-14 https://doi.org/10.1016/j.gloplacha.2003.08.004 https://doi.org/10.1016/j.gloplacha.2003.08.004 https://doi.org/10.37570/bgsd-2004-51-07 https://doi.org/10.37570/bgsd-2004-51-07 https://doi.org/10.1016/j.tecto.2008.10.025 https://doi.org/10.1016/j.tecto.2008.10.025 https://doi.org/10.1016/j.tecto.2012.10.038 https://doi.org/10.1111/bre.12024 https://doi.org/10.18814/epiiugs/2008/v31i1/010 https://doi.org/10.18814/epiiugs/2008/v31i1/010 https://doi.org/10.34194/geusb.v22.4733 https://doi.org/10.34194/geusb.v22.4733 https://doi.org/10.34194/0nydbt40 https://doi.org/10.1016/s0928-8937(05)80050-5 https://doi.org/10.1144/jm.24.1.1 https://doi.org/10.1016/j.epsl.2017.03.036 https://doi.org/10.2973/dsdp.proc.38.130.1976 https://doi.org/10.2973/dsdp.proc.38.130.1976 https://doi.org/10.34194/geusb.v41.4335 https://doi.org/10.34194/geusb.v41.4335 https://doi.org/10.1016/j.margeo.2013.12.014 https://doi.org/10.1016/j.margeo.2013.12.014 https://doi.org/10.34194/geusb.v57.8365 https://doi.org/10.2973/odp.proc.sr.162.013.1999 https://doi.org/10.2973/odp.proc.sr.162.013.1999 https://doi.org/10.2113/gsjfr.21.2.126 https://doi.org/10.2973/dsdp.proc.49.116.1979 https://doi.org/10.2973/dsdp.proc.49.116.1979 https://doi.org/10.1016/j.marmicro.2005.11.004 https://doi.org/10.1080/002919699433870 https://doi.org/10.1144/gsl.sp.2000.167.01.10 https://doi.org/10.1144/gsl.sp.2000.167.01.10 https://doi.org/10.1016/j.epsl.2009.05.021 https://doi.org/10.1016/j.epsl.2009.05.021 Sheldon et al. 2025: GEUS Bulletin 59. 8381. https://doi.org/10.34194/5k9dv133 30 of 30 GEUSBULLETIN.ORG compared: Index events for the Late Cretaceous – Neogene. In: Exon, N.F. et al. (eds): Proceedings of the Ocean Drilling Program, Scientific Results 189, 1–98. https://doi.org/10.2973/odp.proc.sr. 189.107.2004 Williams, G.L., Fensome, R.A. & MacRae, R.A. 2017: The Lentin and Williams index of fossil dinoflagellates 2017 edition. AASP Contribu- tion Series 48. American Association of Stratigraphic Palynologists Foundation. https://doi.org/10.4095/103330 Wrenn, J.H., & Kokinos, J.P. (1986): Preliminary comments on Miocene through Pleistocene dinoflagellate cysts from De Soto Canyon, Gulf of Mexico. In: Wrenn, J.H., Duffield, S.L. & Stein, J.A. (eds): Papers from the first Symposium on Neogene dinoflagellate cyst biostratigraphy, American Association of Stratigraphic Palynologists, Contribution Series 17, 169–225. Young, J.R., Flores, J.A. & Wei, W. 1994: A summary chart of Neogene Nannofossil magnetostratigraphy. Journal of Nannoplankton Research 16(1), 21–27. https://doi.org/10.58998/jnr2294 Young, J.R. 1998: Neogene. In: Bown, P.R. (ed.): Calcareous Nan- nofossil biostratigraphy. British Micropalaeontological Society Series, 226–265. Chapman & Hall/Kluwer Academic. https://doi. org/10.1007/978-94-011-4902-0_8 Young, J.R., Bown, P.R. & Lees, J.A. 2024a: Nannotax3 website. Interna- tional Nannoplankton Association. https://www.mikrotax.org/Nanno- tax3 (accessed December 2024). Young, J.R., Wade, B.S. & Huber, B.T. 2024b: Mikrotax.org website. https://www.mikrotax.org/pforams/index.php?dir=pf_cenozoic (accessed December 2024). Zachos, J.C., Pagani, M., Sloan, L.C., Thomas, E. & Billups, K. 2001: Trends, rhythms, and aberrations in global climate 65 Ma to present. Science 292, 686–693. https://doi.org/10.1126/science.1059412 Ziegler, P.A. 1990: Geological Atlas of Western and Central Europe. In: Mij, B.V. (ed.): 2nd. Ed. Shell International Petroleum, 1–239, Geologi- cal Society Publication. London. https://doi.org/10.34194/5k9dv133 http://www.geusbulletin.org/ https://doi.org/10.2973/odp.proc.sr.189.107.2004 https://doi.org/10.4095/103330 https://doi.org/10.58998/jnr2294 https://doi.org/10.1007/978-94-011-4902-0_8 https://doi.org/10.1007/978-94-011-4902-0_8 https://www.mikrotax.org/Nannotax3 https://www.mikrotax.org/Nannotax3 http://Mikrotax.org https://www.mikrotax.org/pforams/index.php?dir=pf_cenozoic https://doi.org/10.1126/science.1059412 A multidisciplinary biostratigraphic framework for the Lower to Middle Miocene of the Norwegian Nort 1. Introduction 2. Geological setting and palaeoclimate 3. Lithostratigraphy 4. Sequence stratigraphic framework 5. Absolute dating 6. Previous studies 7. Zonation schemesin this study 8. Materials and methods 8.1. Preparation methods 8.1.1. Palynology 8.1.2. Microfossils (large fraction) 8.1.3. Calcareous Nannofossils 8.1.4. Siliceous microfossils (small fraction) 9. Results 9.1. Palynology 9.2. Dinocyst events 9.3. Microfossils 9.4. Microfossil events 9.5. Calcareous Nannofossils 9.6. Calcareous Nannofossil events 9.7. Diatoms 9.8. Diatom zonation 9.9. Diatom events and observations from this study 9.10. Silicoflagellates 9.10.1. Silicoflagellate zonation 9.10.2. Silicoflagellate events 10. Correlation and stratigraphy 11. Discussion 11.1. Improvements to regional and local biostratigraphy 11.2. Sequence stratigraphic and lithostratigraphic correlation 11.3. Age model 11.4. Palaeoclimate 12. Conclusions Acknowledgements Additional Information Funding statement Author contributions Competing interests Additional files References Figures Fig. 1 Palaeogeography of the North Sea area in the Early Miocene. The red dot indicates the locati Fig. 2 Depth map to the top Miocene of the Valhall and Hod structures and locations of the six studi Fig. 3 The new lithostratigraphic subdivision for the Neogene succession in the Danish sector of the Fig. 4 (Continued) Summary diagram for the 2/8-G10A well with biostratigraphic zones, previously pub Fig. 5 Summary diagram for the 2/11-12S well with biostratigraphic zones, previously published event Fig. 8 Multidisciplinary biostratigraphic correlation diagram of the six studied wells, from south