Assemblage-level structure in Morrison Formation dinosaurs, Western Interior, USA GEOLOGY OF THE INTERMOUNTAIN WEST an open-access journal of the Utah Geological Association ISSN 2380-7601 Volume 5 2018 © 2018 Utah Geological Association. All rights reserved. For permission to copy and distribute, see the following page or visit the UGA website at www.utahgeology.org for information. Email inquiries to GIW@utahgeology.org. ASSEMBLAGE-LEVEL STRUCTURE IN MORRISON FORMATION DINOSAURS, WESTERN INTERIOR, USA John A. Whitlock, Kelli C. Trujillo, and Gina M. Hanik �eme Issue An Ecosystem We �ought We Knew— �e Emerging Complexities of the Morrison Formation SOCIETY OF VERTEBRATE PALEONTOLOGY Annual Meeting, October 26 – 29, 2016 Grand America Hotel Salt Lake City, Utah, USA GEOLOGY OF THE INTERMOUNTAIN WEST an open-access journal of the Utah Geological Association ISSN 2380-7601 Production Cover Design and Desktop Publishing Douglas A. Sprinkel Cover Outcrop of Brushy Basin Member of the Morrison Formation in southeast Utah. Photograph by ReBecca Hunt-Foster. i 2018 President Paul Inkenbrandt paulinkenbrandt@utah.gov 801.537.3361 2018 President-Elect Peter Nielsen peternielsen@utah.gov 801.537.3359 2018 Program Chair Emily McDermott ekeller@utah.gov 801.537.3389 2018 Treasurer Zach Anderson zanderson@utah.gov 801.538.4779 2018 Secretary Christopher Kravits ckravitsgeo@gmail.com 2018 Past President Bill Loughlin bill@loughlinwater.com 435.649.4005 UGA Board UGA Committees Education/Scholarship Loren Morton lmorton@utah.gov 801.536.4262 Environmental A�airs Craig Eaton eaton@ihi-env.com 801.633.9396 Geologic Road Sign Terry Massoth twmassoth@hotmail.com 801.541.6258 Historian Paul Anderson paul@pbageo.com 801.364.6613 Membership Rick Ford rford@weber.edu 801.626.6942 Public Education Paul Jewell pwjewell@mines.utah.edu 801.581.6636 Matt A�olter g�247@yahoo.com Publications Roger Bon rogerbon@xmission.com 801.942.0533 Publicity Paul Inkenbrandt paulinkenbrandt@utah.gov 801.537.3361 Social/Recreation Roger Bon rogerbon@xmission.com 801.942.0533 AAPG House of Delegates 2017–2020 Term Tom Chidsey tomchidsey@utah.gov 801.537.3364 State Mapping Advisory Committe UGA Representative Jason Blake blake-j@comcast.net 435.658.3423 UGA Newsletter Newsletter Editor Bob Biek bobbiek@utah.gov 801.537.3356 UGA Website www.utahgeology.org Webmasters Paul Inkenbrandt paulinkenbrandt@utah.gov 801.537.3361 Lance Weaver lanceweaver@utah.gov 801.403.1636 Become a member of the UGA to help support the work of the Association and receive notices for monthly meetings, annual �eld conferences, and new publi- cations. 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Earthquake Safety Committe Chair Grant Willis gwillis@utah.gov 801.537.3355 Douglas A. Sprinkel Utah Geological Survey 801.391.1977 GIW@utahgeology.org Bart J. Kowallis Brigham Young University 801.422.2467 bkowallis@gmail.com �omas C. Chidsey, Jr. Utah Geological Survey 801.537.3364 tomchidsey@utah.gov Steven Schamel GeoX Consulting, Inc. 801.583-1146 geox-slc@comcast.net Society of Vertebrate Paleontology Editors Kelli C. Trujillo — University of Wyoming John Foster — Museum of Moab Cary Woodru� — University of Toronto Octavio Mateus — Universidade Nova de Lisboa Editors GEOLOGY OF THE INTERMOUNTAIN WEST an open-access journal of the Utah Geological Association Volume 5 2018 9 ABSTRACT �e Upper Jurassic Morrison Formation is both geographically extensive and well-sampled, making it an ideal candidate for biogeographic analysis at both coarse and �ner scales. Historically, however, this has not translated into a consensus on patterns of ecological structure and connectivity, particularly with regard to the characteristic dinosaur faunas. Here, we use both traditional (genus richness, alpha and beta diversity) and bipartite network-based (biogeographic connectivity, local endemism, and average occur- rence) measures to examine patterns of structure on a per-locality basis. Given the broad geographic range of the formation, we subdivide the Morrison Formation into four discrete regions based roughly on lat- itude and lithology—north (Montana, South Dakota, and northern Wyoming), west (Utah and western Colorado), east (central and eastern Colorado and southern Wyoming), and south (Arizona, New Mexico and Oklahoma). Further investigation revealed many coeval sites (ca. 152 Ma) in the east and west regions. Presence-absence data were also compared using network analysis to determine the presence and con- tent of discrete subassemblages within the larger region-level assemblages. Based on our results, we favor reconstructions of the Morrison Formation as a ‘mosaic’ type environment over most of its depositional history, with patches of open environments interspersed with more closed, forested regions. �is is sug- gested by relatively low rates of local endemism (patches are consistent in plant and animal structure) and connectivity across the majority of the formation, as well as the recovery of three non-overlapping assem- blages dominated by di�erent guilds of herbivorous dinosaurs. Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA John A. Whitlock1, Kelli C. Trujillo2, and Gina M. Hanik1 1Department of Science and Mathematics, Mount Aloysius College, Cresson, PA 16630; jwhitlock@mtaloy.edu; gmhst4@student.mtaloy.edu 2Haub School of Environmental and Natural Resources, �e University of Wyoming, Laramie, WY 82071; kellitrujillo@icloud.com Citation for this article. Whitlock, J.A., Trujillo, K.C., and Hanik, G.M., 2018, Assemblage-level structure in Morrison Formation dinosaurs, Western Interior, USA: Geology of the Intermountain West, v. 5, p. 9–22. © 2018 Utah Geological Association. All rights reserved. For permission to use, copy, or distribute see the preceeding page or the UGA website, www.utahgeology.org, for information. Email inquiries to GIW@utahgeology.org. INTRODUCTION �e Late Jurassic Morrison Formation (~157-149 Ma) of North America represents one of the most ex- tensive Mesozoic terrestrial depositional basins yet discovered, covering approximately 1.5 million square miles of territory, although much of the formation re- mains unexposed (Foster, 2003). As a result, the Mor- rison Formation has proven to be one of the most pro- ductive sources for dinosaur material, with dozens of named taxa inclusive of a wide range of bauplans, body sizes, and ecological habits. Despite the paleontological riches, however, little agreement on the basic ecosystem has been reached, with authors variously claiming a dominant conifer forest habitat, xeric plains-type hab- itats, or a “Jurassic savanna” habitat dominated by lush, low-growing vegetation and at least seasonally wet cli- mates, or a mosaic of at least two distinct habitat types. www.utahgeology.org 10 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 Here, we show contrasting patterns in distribution be- tween the largest bodied herbivores and all other dino- saurian taxa, and interpret the low degrees of endemism and connectivity as evidence for a ‘mosaic’ type land- scape throughout most of the depositional basin repre- sented by the Morrison Formation. Prior studies of broad-scale ecological reconstruc- tion in the Morrison Formation have primarily been focused on the abundant macro- and microfossil paleo- botanical evidence. Plant fossils representing a diverse �ora of cheirolepidiaceans, conifers, ferns, ginkophytes, horsetails, and podocarpaceans are known throughout the formation (Tidwell, 1990; Ash and Tidwell, 1998; Parrish and others, 2004). Between the abundance of drought intolerant plants (e.g., ferns, horsetails) and the known abundance of large herbivores (with their attendant forage requirements), some of the original interpretations of the Morrison Formation paleoenvi- ronment focused on it as a lush, wet �oodplain (Tid- well, 1990; Ash and Tidwell, 1998). Additionally, the ap- parent absence of arrested growth structures in woody remains and the presence of wood-rotting fungi have also been cited as evidence for a generally humid, mesic environment (Tidwell and others, 1998). More recently, however, two separate interpretations have taken root. One camp suggests that the paleoenvironment record- ed by the Morrison Formation was xeric or only sea- sonally wet, with a �ora comprised primarily of herba- ceous and low- to mid-height woody podocarpaceaean shrubs; taller woody plants would have been restricted to riparian environments (Engelmann and others, 2004; Parrish and others, 2004; Rees and others, 2004). �is view interprets the paleoenvironment as roughly equiv- alent to a modern tropical savanna grassland, with ferns and bryophytes occupying the ground-level browse lay- er currently dominated by grasses. More recently, some authors have presented evidence for the paleoenviron- ment of the Morrison Formation as a ‘mosaic,’ either with a matrix of conifer forest dominating interspersed open habitats (Gee, 2010; Hotton and Baghai-Riding, 2010), or as a primarily open environment with inter- spersed woodland environments (Whitlock, 2011). In all cases, it seems unlikely that dry-adapted plants such as cheirolepidiaceans would be restricted to riparian lands with wet soils, and indeed the opposite interpreta- tion is now typically accepted (Hotton and Baghai-Rid- ing, 2010). �ere has been some attempt to segregate the Mor- rison Formation into ecological regions, such that the northern exposures of the Morrison (particularly in Montana and South Dakota) represent a substantial- ly wetter paleoenvironment than the exposures to the south (Parrish and others, 2004; Noto and Grossman, 2010). Montana, in particular, houses some localities reconstructed as a unique depositional environment, dominated by mires, coal swamps, and peat bogs (Par- rish and others, 2004). Palynological evidence also in- dicates the presence of a more humid environment in the northern extremes with a potentially higher water table as well (Hotton and Baghai-Riding, 2010). �is is corroborated by the more frequent occurrence of semi-aquatic reptiles (e.g., crocodilians, turtles; Foster and McMullen, 2017) in the Morrison Formation of Montana, South Dakota, Wyoming, and eastern Colo- rado. Furthermore, there has been a suggestion of a lon- gitudinal gradient of drainage and precipitation as well (Demko and others, 2004; Turner and Peterson, 2004), with drier margins and a ‘center’ with the majority of standing water in the forms of lakes and wetlands. From the earliest stages of ecological assessment in the Morrison Formation, a ‘core’ fauna of large diplodo- cids, Stegosaurus, and Allosaurus has been recognized, with other taxa appearing with less regularity (Dodson and others, 1980a, 1980b), although there has been little consensus on how those faunas have been constructed. More recently, Noto and Grossman (2010) analyzed the paleoecology of dinosaurs in the Morrison Formation (and other formations), dividing the localities into six “assemblages” based on geographic proximity and us- ing Ecological Structure Analysis (ESA) to identify any potential relationships between the behavior or ecolo- gy of the dinosaurs (e.g., body size, locomotor habit) and the regional environment. Although they did not make explicit statements about generic constituencies of any inferred communities, they did identify what they termed a “semi-arid” set of assemblages from the latitudinal center of the formation, bounded by a more arid set of assemblages from the northern and southern extremes. �is distinction was largely recognized in the faunas as a function of body size (smaller overall body 11 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 size in the semi-arid environments) and locomotor habit (semi-arid environments were more dominated by bipedal organisms; Noto and Grossman, 2010). Here, we attempt to build on this prior work by ex- amining the Morrison Formation on a quarry level to try and pull out any signal that might indicate the pres- ence of distinct communities at the generic level. Using classical ecological inferences (alpha and beta diversity and shared taxonomic scope) in concert with newer bi- partite statistical methods (following Sidor and others, 2013), we characterize the dinosaur assemblages at both regional and individual locality scales, and interpret the results in the context of describing the overarching eco- logical conditions present at the time of Morrison For- mation deposition. METHODS Because the Morrison Formation incorporates both a large geographic and temporal range, the 291 locali- ties sampled were divided into four regions: north (65 localities), inclusive of localities in Montana, South Da- kota, and northern Wyoming (above 43°N); west (72 localities), inclusive of localities in Utah and Colora- do just east of the city of Gunnison, Colorado (Cabin Creek [see Turner and Petersen, 1999] and westward); east (128 localities), inclusive of Colorado east of Cabin Creek and Wyoming below 43°N and the lithological- ly similar Snowmass locality (Foster and Wedel, 2014); and south (26 localities), inclusive of localities in Arizo- na, New Mexico and Oklahoma (�gure 1). Di�erences in recognized members across the formation as well as limitations on the quality of intraformational correla- tions hamper further subdivision into more de�nitive age units at this time (Trujillo, 2006). New 40Ar/39Ar and U-Pb isotope ages, however, identify 28 localities across the east and west regions that are roughly coeval (ca. 152 Ma; Trujillo and others, 2014, 2015; Trujillo and Kowallis, 2015). �ese localities are examined in- dependent of the rest of the data set to provide further insight into potential regional di�erences independent of time dilation e�ects. Because we are predominantly interested in examining regional patterns of taxonomic structure, we refrain from attempts to group localities into bins of equal number or attempts to �lter localities by alpha diversity and other indicators. Taxonomic data was pooled from the Paleobiology Database (http://fossilworks.org; see supplemental in- formation) and supplemented with additional informa- tion from Foster (2003). One caveat with using data of this sort is that we are, at best, two steps removed from the original material (collector->compiler->present au- thors), each step having the potential to introduce er- rors ranging from incorrect taxonomic assignment to typographic issues. As it is reasonable to assume the general care of our colleagues with regard to data colla- tion, the biggest concern is misidenti�cation of the fos- sils themselves. Dinosaur workers in general and those familiar with the Morrison Formation speci�cally will no doubt already be aware of the di�culty in correctly assigning many fossil specimens, as well as the poten- tial for “hidden” taxonomic diversity (e.g., Galeamopus, “Elaphrosaurus;” Chure, 2001; Carrano and Sampson, 2008; Tschopp and others, 2015; Tschopp and Mateus, 2017). While recognizing the potential confounding e�ects of these issues, there is fundamentally little that can be done to resolve them at this level, and thus we note their existence here and calibrate our interpreta- tions accordingly. Similarly, we note the potential for taphonomic bias towards larger bodied taxa, which we attempt to mitigate again by limiting our scope to pres- ence-absence rather than abundance. Taxonomically unresolved taxa—e.g., “Elaphrosau- rus”—or records without identi�cation to the genus lev- el were omitted. �e recently proposed generic separa- tion between Apatosaurus and Brontosaurus (Tschopp and others, 2015) is not incorporated into this analysis for reasons of practicality; the records in question pre- date this proposal, and in lieu of attempting to examine all specimens of Apatosaurus in question (some 75 to- tal records composed of untold numbers of individu- al specimens), we elect to simply continue use of the genus Apatosaurus. To examine large-scale trends in connectivity and diversity, alpha diversity (using genus richness), beta diversity (as measured by Sørenson’s In- dex, or SI), and shared taxonomic scope were calculated using the pooled assemblage data from each major re- gion. Sørenson’s Index was chosen as a measure of beta diversity vis-à-vis continuity (sensu Kole� and others, 2003). Importantly, SI satis�es the requirement of be- 12 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 ing sensitive only to proportions of species, not to total number (Janson and Vegelius, 1981), the latter being a potential issue given the unequal areas and taxonom- ic compositions of the regions under consideration. To characterize locality-level connectivity, bipartite resam- pling methods (following Sidor and others, 2013) were used to calculate three measures: average occurrences per taxon (interpreted as range size, following Sidor and others, 2013), average endemic taxa per locality, and bio- geographic connectivity (the number of taxon linkages as a proportion of the maximum possible). Each anal- ysis presents a distribution of scores obtained via 1000 bootstrap replicates (see supplemental information for discussion of replicate choice). A Welch two-sample un- equal variances t-test was used to determine statistical signi�cance. To look for potentially distinct assemblag- es within the taxonomic samples, co-occurrence and edge matrices were computed in R using the cooccur package (Gri�th and others, 2016); resultant networks were analyzed for patterns of generic clustering using Gephi (Bastian and others, 2009). For the temporal groupings in the east and west regions, localities with good stratigraphic data were correlated with the nearest precise radiometric age by K.C. Trujillo. RESULTS Alpha diversity is consistently high in the north, west, and east regions (21-22 genera) and substantially lower in the south region (10 genera; table 1). Sauro- pod alpha diversity appears to decrease with decreasing latitude, but this trend is not seen in either theropods or ornithischians. In theropods, there appears to be a ‘bulge’ in diversity in a central region (west and east re- gions combined). Beta diversity is highest between the Figure 1. Map of localities by county, color coded by region. Heavy black line indicates approximate geographic extent of the Morrison Formation (modi�ed from Turner and Peterson, 2004). See supplemental information for full details. 13 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 north and east regions (SI = 83% similarity; table 2), but is also notable between the west and east regions (71%). Similarity is lower between the north and west regions (65%), and is lowest between the south region and all others (56-58%), with the north and south being most di�erent. �is trend is generally conserved when the samples are restricted by taxonomic group, with the exception of sauropods, where similarity is highest be- tween the south region and the west and east regions (83%), although again the north and south are least similar (63%). Shared taxonomic scope is reasonably high in the Morrison Formation (�gure 2). �is is exempli�ed by the large-bodied sauropods, where 5 of 12 taxa occur in all four regions and 8 of 12 occur in at least two. �e regionally endemic taxa, Dyslocosaurus, Dystrophaeus, Kaatedocus, and Suuwassea, are also known from only a single locality each, which suggests that their rarity is either a function of their rarity in the fauna as a whole or the rate at which they are misidenti�ed by collectors, or both. �eropod shared taxonomic scope is less robust, with only two of the largest bodied taxa (Allosaurus and Ceratosaurus) occurring in all four regions, and two others (the very large Torvosaurus and the small Coe- lurus) occurring in three. Nearly half (4/10) of theropod taxa are known from a single region only, and there is the possibility of a great many more singleton taxa that would increase this total (e.g., any potential taxa subdi- vided from the catch-all genus “Elaphrosaurus,” Kopar- ion). �ese regionally endemic theropods are invariably small-bodied. Ornithischian-shared taxonomic scope is somewhat intermediate between sauropods and thero- pods, in that regional endemism is high (4/10 taxa), as in theropods, but as many taxa occur in at least three regions (4/10), a distribution somewhat similar to that seen in sauropods. Two taxa (Camptosaurus and Stego- saurus) are known from all four regions. For the 152 Ma time bin, absolute taxonomic diversity is highest in the east region (18 taxa vs. 12 taxa in the west). Beta diver- sity is generally high (0.66), but this is largely driven by the sauropods (1.0). �eropod (0.5) and ornithischian (0.5) assemblages are both relatively less similar. �e di�erence in taxonomic scope is largely due to the ab- sence of many relatively small bodied taxa, particularly theropods (e.g., Coelurus) in the west. Bipartite analyses result in signi�cantly di�erent (p <0.0001; t and df reported in supplemental informa- tion) distributions of all three characteristics (average occurrence, average endemics, biogeographic connec- tivity) for all regions, excluding only biogeographic connectivity between north, east, and west regions (1 ≤ p ≤ 0.9994). Despite this statistical signi�cance, cer- tain characteristics of certain regions appear more sim- ilar to each other than others. As noted, biogeographic connectivity is statistically identical in the north, east, and west regions (�gure 3), and is signi�cantly lower Total Generic Diversity Sauropods only Theropods only Ornithischians only North 22 11 4 7 West 21 7 9 5 East 21 7 7 7 South 10 5 3 2 152Ma East 18 6 5 7 152Ma West 12 5 3 4 Table 1. Alpha diversity (generic richness) for each of the four study regions. North West East South AL L T AX A North West 65% East 83% 71% South 56% 58% 58% North West East South SA UR OP OD S North West 67% East 78% 71% South 63% 83% 83% North West East South TH ER OP OD S North West 67% East 73% 80% South 57% 36% 40% North West East South OR NI TH IS CH IA NS North West 62% East 86% 62% South 44% 50% 44% Table 2. Beta diversity (Sørenson’s Index) for each of the four study regions. 14 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 than in the south region, where connectivity is highest. Despite the lower connectivity scores, locality-level en- demism is essentially zero in the central and north re- gions, with an order of magnitude higher rates recorded in the south region. Average occurrence (AO) scores do increase as the number of localities sampled increases, but none are directly limited by the number of localities (i.e., all AO scores are less than 1/4 of the number of sampled localities). Time-restricted samples show the same general pattern as the broader samples, with the exception of an unusually high connectivity in the east 152 Ma time bin (�gure 3). Assemblage-level structure in the Morrison Forma- tion is generally built around a consistent “core” of taxa (Allosaurus, Apatosaurus, Camarasaurus, Diplodocus, and Stegosaurus) that are highly connected with the ma- jority of other taxa, and might be considered the ‘typi- cal’ Morrison fauna (Dodson and others, 1980a, 1980b). Camptosaurus and Ceratosaurus are also commonly found co-occurring with most other taxa. Larger her- bivores (i.e., sauropods, Camptosaurus) and some large predators (i.e., Allosaurus, Ceratosaurus) display little preference with regard to co-occurring taxa, although smaller carnivores (Coelurus, Tanycolagreus) and many other large predators (Fosterovenator, Stokesosaurus, Torvosaurus) do appear to show some segregation. �ese patterns are most distinct in the central region (�gure 4). �e most locality level segregation is found among the medium- to small-sized ornithischians in the central region, which appear to sort themselves into three general groups based upon non-overlapping ex- emplar taxa: (1) a ‘Mymoorapelta’ group, which co-oc- curs with no other smaller herbivores, only large sau- ropods; (2) an ’Othnielosaurus’ group, which co-occurs with Camarasaurus, Camptosaurus, Drinker, Nano- saurus, and Stegosaurus; and (3) a ‘Dryosaurus’ group, consisting of Apatosaurus, Camarasaurus, Camptosau- rus, Diplodocus, Gargoyleosaurus, and Stegosaurus. In the north region, where connectivity is relatively high, Dryosaurus and Othnielosaurus do co-occur in a single locality (Howe-Stevens, see supplemental information); the remaining combined 10 occurrences (as measured Figure 2. Shared taxonomic scope between the four regions of interest with distribution broken down by individual genera. 15 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 by presence/absence) in the north region and all 15 oc- currences in the central region are mutually exclusive. Othnielosaurus is a relatively rare taxon, but it is sub- stantially more common in the northern Morrison (�ve localities vs. two each in the west and east). Coelurus is found exclusively with the Othnielosaurus assemblage, and Tanycolagreus is found exclusively with the Dryo- saurus assemblage, although the two taxa do co-occur (in the absence of both Dryosaurus and Othnielosaurus) in the west region. Within the 152 Ma time bin, we observe this segre- gation in the east (�gures 4E and 4F), but not in the west Figure 3. Box and whisker plots representing the results of the bipartite statistical analyses for each sampled region as well as the 152 Ma sample of east and west. Average occurrence represents the average number of localities a randomly selected species would occur in. Average endemics reports the average number of per-locality endemic taxa recovered. Biogeographic connectivity measures the proportion of recovered connections (shared taxa) as a function of the total potential connections. Numbers reported represent distributions over 1000 replicates. 16 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 Figure 4. Network diagrams illustrating co-occurrence between genera in the central Morrison Formation. (A) Network diagram of the north region, with conjoined Dryosaurus and Othneilosaurus network highlighted. (B) Network diagram of the west region, with Mymoorapelta assemblage highlighted. (C) Network diagram of the west region, with Dryosaurus assemblage highlighted. (D) Network diagram of the west region, with Othnielosaurus assemblage highlighted. (E) Network diagram of the east region, with Dryosaurus assemblage highlighted. (F) Network diagram of the east region, with Othnielosaurus assemblage highlighted. Abbreviations: Al = Allosaurus; Ap = Apatosaurus; Ba = Barosaurus; Br = Brachiosaurus; Ca = Camarsaurus; Ce = Ceratosaurus; Co = Coelurus; Cp = Camptosaurus; Di = Diplodocus; Dr = Drinker; Ds = Dyslocosaurus; Dt = Dystrophaeus; Dy = Dryosaurus; Fo = Fosterovenator; Fr = Fruitadens; Ga = Gargoyleosaurus; Gp = Galeamopus; Ha = Haplocanthosaurus; He = Hesperosaurus; Ka = Kaatedocus; Ma = Marshosaurus; My = Mymoorapelta; Na = Nanosaurus; Or = Ornitholestes; Ot = Othneilosaurus; Sg = Stegosaurus; St = Stokesosaurus; Su = Suuwassea; Ta = Tanycolagreus; To = Torvosaurus. 17 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 (�gure 4D). Furthermore, Dryosaurus does not co-oc- cur with any other taxa in this sample and so provides no information relevant to its participation in discrete assemblages. Aside from a slightly lower taxonomic di- versity, the 152 Ma time bin is representative of the re- gions as a whole with regard to taxonomic assemblages. DISCUSSION Diversity metrics (genus richness, SI) generally sug- gest similar diversity levels in localities in the northern and central parts of the Morrison Formation, with the greatest diversity occurring in the central region. Re- gional interconnectivity appears to be tied to some com- bination of body size and ecological habit. Large-bodied herbivores (such as sauropods, and certain ornithiscians like Camptosaurus) tend to occur in more regions over- all (lower rates of endemism), and have a wider individ- ual distribution (weighted towards occurrence in four regions over occurrence in two). Camptosaurus is itself potentially cosmopolitan not simply within the Morri- son Formation, but also may appear in coeval basins in Europe as well (Foster, 2013). �is would be a substan- tial geographic range for a single taxon, although repre- sentatives of other Morrison Formation genera such as Allosaurus, Ceratosaurus, Stegosaurus, and Torvosaurus have been reported from Iberia (Mateus, 2006; Mateus and others, 2006; Escaso and others, 2007; Hendrickx and Mateus, 2014). Whether this is a true distribution or simply a matter of taxonomic ‘lumping’ (e.g., ‘Ela- phrosaurus;’ Carrano and Sampson 2008) is unclear, however. �ere does not appear to be a clear size-distribu- tion relationship for theropods, as in general large- and small-bodied individuals are recovered as both widely distributed (e.g., Allosaurus, Coelurus) and regionally endemic (e.g., Saurophaganax, Stokesosaurus). �is may be due to increased competitive pressures facing car- nivores, access to appropriate prey, and/or reinforcing behaviors such as territorialism. We agree with Foster (2000) in that we �nd little evidence to suggest drastic di�erences in environment with latitude, particularly with regard to the northern Morrison Formation as an environmental outlier, as has been previously suggested (Parrish and others, 2004; Hotton and Baghai-Riding, 2010; Whitlock, 2011). Sauropod diversity does decline with declining latitude, but the di�erences between the north and central regions are not substantial and are primarily driven by the presence of two north region endemics, Kaatedocus and Suuwassea. �ese two taxa are both very recently described (Harris and Dodson, 2004; Tschopp and Mateus, 2013) and are represented by a single specimen each; consequently, it is not im- plausible that additional specimens from other regions are either undiscovered or misidenti�ed in existing col- lections. Bipartite statistics predict greater provincialism when biogeographic connectivity and range sizes are small and average endemism is high (Sidor and others, 2013). In the south region, we �nd agreement between range size and endemism—both of which predict a high degree of provincialism—but disagreement with biogeographic connectivity, which would predict a very cosmopolitan distribution of taxa. Similarly, range sizes and endemism predict a more cosmopolitan distribu- tion in the north, east, and west regions, but the com- paratively lower biogeographic connectivity in those re- gions predicts provincialism. Some of these e�ects may be attributable to time dilation e�ects; i.e., substantial turnover in environmental structure may result in these statistics capturing discordant e�ects due to overlaying di�erent environmental structures on top of each oth- er, both in physical space (as a function of deposition) and in ‘statistical space’ (as a function of data pooling). However, we postulate that these results may be still interpretable due to a highly fractured environments. Extremely patchy ecosystems would result in smaller ranges and increased regional endemism. If substan- tial corridors connecting similar patches did not exist (or were outside the depositional area recorded), these patches would be isolated from each other in the fossil record, which is consistent with a low per-locality con- nectivity measurement, as observed. We do interpret some degree of connectivity, largely driven by the cos- mopolitan sauropod fauna, as complete balkanization of depositional environments has been shown to cor- respond to even lower degrees of connectivity than ob- served here (Sidor and others, 2013). In the case of the widely distributed taxa (sauropods, large theropods like Allosaurus), it seems likely that their large size would 18 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 have made them largely immune to barriers, inclusive of larger patches of less suitable habitat. In modern eco- systems, large rivers present barriers to dispersal for many organisms, including such varied groups as Afri- can Bu�alo (Naidoo and others, 2012), elephants (Rob- ertson, 2013), lions and other African carnivores (Cozzi and others, 2013), wolves (Blanco and others, 2005), passerine birds (Ayres and Clutton-Brock, 1992; Hayes and Sewlal, 2004; but see Gascon and others, 2000), and primates (Goodman and Ganzhorn, 2004; Harcourt and Wood, 2012), although there is some evidence that larger bodied taxa are less a�ected, particularly when the river in question is small or seasonally low (Cozzi and others, 2013; Robertson, 2013). �e river itself does not need to be impassable to be a barrier; in many cases the ecosystem on either side of the river is sig- ni�cantly di�erent as to be a barrier unto itself (Oates, 1998; Meijard and Groves, 2006). A more continuous environment would result in a signal with low regional endemism and high connectivity, as there would be no barrier regions where a variety of ecomorphs would be less likely to occur or be preserved. When restricted to localities deposited at or near 152 Ma, however, our results paint a more consistent picture. �e west region scores lower in range size and connectivity and higher in endemism than the east, all of which suggest that the west was more provincial at this time. �is fragmentation supports the idea that there was no ‘rain shadow’ e�ect caused by coeval and preceding upli� and erosion of ri� shoulders associated with a magmatic arc west of the Morrison Formation’s depositional area (Turner and Fishman, 1998; Dickin- son, 2006). Such upli� could have potentially created a ‘rain shadow’ on the leeward basin, resulting in an arid region without signi�cant permanent riparian struc- ture. Due to the high rate of sediment deposition at- tributable to erosion from those highlands (Turner and Fishman, 1998), however, it seems unlikely that there was a signi�cant rain shadow e�ect on the west region at this time (Turner and Peterson, 2004). In addition, a rain shadow would likely have resulted in a relatively homogenous, ‘savanna’ type environment without ma- jor river systems to serve as dispersal barriers for small- er dinosaurs (Ash and Tidwell, 1998; Parrish and others, 2004). Instead, the apparently robust riparian structure in that region may well have provided signi�cant (if ephemeral on geologic time scales) barriers to dispersal in the form of major river systems and alkaline-saline wetlands (e.g., “Lake” Too’di’chi’ in southwest Colorado and southeast Utah; Fishman and others, 1995; Turner and Peterson, 2004), which could potentially have lim- ited the range sizes and regional distribution of smaller dinosaurs (e.g., smaller theropods, ornithischians). �e riparian structure would have been less restrictive to the extremely large sauropods and larger theropods like Al- losaurus, which do indeed have a more cosmopolitan distribution. Range size is also strongly correlated with body size in extant mammals (McNab, 1966; Swihart and others, 1988), largely as a function of energetics, which would further explain the consistently cosmo- politan distribution of these larger organisms. �e larger range sizes and degree of connectivity overall in the east, however, suggest greater environ- mental homogeneity, consistent with reconstructions of this region dominated by a broad alluvial plain (Turner and Peterson, 2004). �erefore, we favor the nuanced interpretation many modern studies have put forward, such that the geographically large region recorded by the Morrison Formation likely included a diverse array of habitats and habitat structures inclusive of both drier, ‘savanna’-type environments and wetter, riparian-dom- inated environments, consistent with regional tectonic and eustatic changes at the time. However, the patchi- ness (or ‘mosaic’ distribution) of these environments as well as the e�ects of time dilation make direct compar- isons to modern ecosystems (e.g., ‘savanna’) somewhat problematic, particularly as the geographic scale of the study increases. �e interpretation of our bipartite analysis results is consistent with the network analysis which recovered three distinct ornithischian assemblages in the central region—a ‘Mymoorapelta’ assemblage, a ‘Dryosaurus’ assemblage, and an ‘Othnielosaurus’ assemblage—as well as with prior work (Noto and Grossman, 2010). �e Mymoorapelta group appears to be more ‘open’ adapt- ed, based upon the feeding preferences of the locally dominant diplodocid sauropods (Stevens and Parrish, 1999; Upchurch and Barrett, 2000; Stevens and Parrish, 2005; Whitlock, 2011), although the Mymoorapelta type locality does feature abundant evidence for conifers 19 Assemblage-level Structure in Morrison Formation Dinosaurs, Western Interior, USA Whitlock, J.A., Trujillo, K.C., and Hanik, G.M. Geology of the Intermountain West 2018 Volume 5 (Tidwell and others, 1999; Hotton and Baghai-Riding, 2010). �e Dryosaurus and Othnielosaurus assemblag- es both contain more closed-adapted genera, including a wide variety of other medium-sized ornithischians, but they di�er regarding sauropod content. Dryosaurus assemblages include both diplodocids and the mid- to high-browser Camarasaurus (Fiorillo, 1998; Upchurch and Barrett, 2000; Whitlock, 2011); Othnielosaurus as- semblages are strongly associated with Camarasaurus. �is suggests the preference of Dryosaurus for edge en- vironments (where open and closed adapted taxa would be expected to intermingle) and closed environments for Othnielosaurus. As Othnielosaurus is reconstructed as substantially smaller than Dryosaurus (e.g., Farlow and others, 2010), this is a reasonable assumption to make. �ere is some evidence for habitat preference in theropods as well, with many large-bodied taxa appear- ing to prefer open environments and medium- to small theropods choosing closed or edge environments. �is association is not as strong, however, suggesting that habitat choice is more labile. �is is consistent with ob- servations of modern animals, which o�en display vari- ance in habitat preference largely dependent on non- biotic environmental factors such as local temperature (May and others, 2010; van Beest and others, 2012). CONCLUSIONS A picture of the Morrison Formation paleoeco- system emerges as a mosaic of diverse communities and environmental structures in a massive, long-lived depositional basin, regionally dominated by distinct populations of relatively small-bodied ornithischian herbivores and theropod carnivores, with widespread and potentially migratory populations of large-bodied sauropods and certain other exemplar taxa such as Allo- saurus and Camptosaurus. Where temporal restriction is possible, distinct regional di�erences emerge between the western and eastern regions of the basin, consistent with coeval tectonic events, such that upli� and exten- sive riparian development to the west created greater habitat fractionation and provincialization in dinosau- rian assemblages. Direct comparisons to modern eco- systems (e.g., use of analogs such as ‘savanna’) are likely of limited application, at least at this geographic scale. �e apparent patchiness of much of the Morrison basin suggests a ‘mosaic’ type environment, and as such we must look to di�erent approaches, such as this one, to gain a better understanding of the community-level dy- namics in the megafauna. ACKNOWLEDGMENTS We thank C.V. Beightol (University of Washington), C.A. Sidor (University of Washington), and D. A. Vil- hena (Cedar AI) for discussion and technical assistance with bipartite methods. J.R. Foster (Museum of Moab), C.R. Noto (University of Wisconsin-Parkside), and M. J. Wedel (Western University of Health Sciences) pro- vided helpful commentary that greatly improved a pre- vious dra�. J.R. Foster, O. Mateus (Universidade Nova de Lisboa), and D.C. 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Whitlock, J.A., 2011, Inferences of diplodocoid (Sauropoda: Di- nosauria) feeding behavior from snout shape and microwear analyses: PLoS ONE, v. 6, no. 4, p. e18304. Descriptive Statistics Mean, Variance and Standard Deviation. Average Occurrence Average Endemics Biogeographic Connectivity M VAR SD M VAR SD M VAR SD North 10.43 0.12 0.35 0.0046 7.12E-05 0.00844 0.15 3.12E-05 0.00559 West 11.48 0.144 0.38 0.0014 1.78E-05 0.00422 0.15 2.78E-05 0.00527 East 18.44 0.25 0.50 1.60E-05 1.28E-07 0.00036 0.14 1.59E-05 0.00399 South 5.43 0.11 0.33 0.047 0.0013 0.03606 0.18 0.00019 0.01378 152Ma East 3.85 0.033 0.18166 0.26 0.0092 0.09592 0.19 0.00015 0.01225 152Ma West 2.327 0.0311 0.18 0.377 0.0136 0.11682 0.147 0.000383 0.01958 p, t-statistic and DF for for Welch's unequal variances t-test. Average Occurrence p t Df North-West <0.0001 64.27 1984 North-East <0.0001 415.02 1788 North-South <0.0001 346.44 1991 West-East <0.0001 350.46 1864 West-South <0.0001 380.13 1959 East-South <0.0001 686.74 1730 152Ma East-West <0.0001 205.161 1996 Average Endemics p t Df North-West <0.0001 10.72 1469 North-East <0.0001 17.16 1002 North-South <0.0001 32.75 1123 West-East <0.0001 10.33 1013 West-South <0.0001 37.78 1026 East-South <0.0001 41.27 999 152Ma East-West <0.0001 31.149 1850 Biogeographic Connectivity p T Df North-West 1 0 999 North-East 0.9995 0.0006 999 North-South <0.0001 67.08 1998 West-East 0.9994 0 999 West-South <0.0001 64.3 1285 East-South <0.0001 88.17 1165 152Ma East-West <0.0001 68.9153 1658 Diversity Alpha Diversity & STS SORENSON'S SIMILARITY INDEX N W E S 152E 152W Apatosaurus 1 1 1 1 1 1 SHARED Barosaurus 1 1 1 1 1 1 ALL SPODS ORNITHS TPODS Diplodocus 1 1 1 1 1 1 NW 14 6 4 4 Galeamopus 1 1 1 NE 17 7 6 4 Camarasaurus 1 1 1 1 1 1 NS 9 5 2 2 Haplocanthosaurus 1 1 EW 15 5 4 6 Supersaurus 1 1 ES 9 5 2 2 Brachiosaurus 1 1 1 1 1 1 WS 9 5 2 2 Dyslocosaurus 1 Suuwassea 1 SI Kaatedocus 1 N W E S Dystrophaeus 1 N Stegosaurus 1 1 1 1 1 1 ALL W 65.1162790698 Camptosaurus 1 1 1 1 1 1 E 82.9268292683 71.4285714286 Drinker 1 1 1 S 56.25 58.064516129 58.064516129 Dryosaurus 1 1 1 1 1 Nanosaurus 1 1 1 Othnielosaurus 1 1 1 1 W 66.6666666667 Gargoyleosaurus 1 1 SPODS E 77.7777777778 71.4285714286 Hesperosaurus 1 S 62.5 83.3333333333 83.3333333333 Fruitadens 1 1 Mymoorapelta 1 1 W 61.5384615385 Allosaurus 1 1 1 1 1 1 ORNITH E 85.7142857143 61.5384615385 Ceratosaurus 1 1 1 1 1 1 S 44.4444444444 50 44.4444444444 Coelurus 1 1 1 1 Saurophaganax 1 W 66.6666666667 Fosterovenator 1 TPODS E 72.7272727273 80 Ornitholestes 1 1 S 57.1428571429 36.3636363636 40 Stokesosaurus 1 Tanycolagreus 1 1 1 Marshosaurus 1 Torvosaurus 1 1 1 1 ALL 22 21 21 10 18 12 SAUROPODS 11 7 7 5 6 5 ORNITHS 7 6 7 2 7 5 THEROPODS 4 8 7 3 5 2 all NORTH 65 localities SOUTH 26 localities EAST 126 localities WEST 74 localities NORTH 65 localities PDBD # T&P 99 # Locality Name Age State County Member Apatosaurus Barosaurus Diplodocus Galeamopus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus PaleoDB 50714 Horse Coulee, AMHN 03-30 MT Big Horn -- 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38866 Rattlesnake RIdge MT Carbon -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 PaleoDB 49624 MT-2 Upper Strickland Creek MT Park -- 0 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 69890 MT-1 Mother's Day MT Carbon -- 0 0 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 T&J Quarry MT Gallatin -- 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Wittecombe's Ranch MT Wheatland -- 0 0 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49625 SD-1 Bear Butte SD Meade -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49630 SD-2 Fuller's 351, St. Onge SD Lawrence -- 1 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49633 Parker's Mystery SD Fall River -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57905 SD-3 Piedmont Butte, Hatch Ranch, Barosaurus type, SDSM V9140 SD Meade -- 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 132242 AHMN quarries, Sturgis area SD Meade -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 132243 USNM Quarry, Sturgis area SD Meade -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49631 SD-4 Wonderland, Piedmont SD Meade -- 0 1 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57904 SD-5 Wonderland North SD Meade -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 46468 WY-61 Big Al SIte WY Big Horn -- 0 0 1 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Big Butte WY Big Horn 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Smithsonian Quarry 1 WY Big Horn 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Dyer Ranch 1 and 2 WY Big Horn 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Freezeout Hills General WY Big Horn 0 0 1 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 57893 Freezeout Hills Quarry 4 WY Big Horn 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57894 Freezeout Hills Quarry 6 WY Big Horn 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57895 Freezeout Hills Quarry L WY Big Horn 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Freezeout Hills Quarry N WY Big Horn 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 PaleoDB 57896 Freezeout Hills Quarry O WY Big Horn 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 46460 WY-62 Howe Quarry WY Big Horn -- 1 1 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 46462 Howe Quarry 2, Sauriermuseum WY Big Horn -- 0 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 PaleoDB 46465 Howe-Stephens Quarry WY Big Horn -- 1 1 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 0 1 0 0 0 0 Lake's Quarry WY Big Horn 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 46469 Spring Hill, Howe Ranch WY Big Horn -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55333 Fox Mesa WY Big Horn -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Matt Quarry WY Big Horn 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Ninemile Hill WY Big Horn 0 0 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55323 ?reopened by C. Weege 1992? Ninemile Quarry (AMNH), Nine Mile Crossing, Nine Mile Quarry WY Big Horn -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Reed's Quarry 1.5 WY Big Horn 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Reed's Quarry 5 WY Big Horn 0 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Reed's Quarry 7 WY Big Horn 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Reed's Quarry 12 WY Big Horn 0 0 1 0 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 62561 Red Canyon Ranch, Hope College REU WY Big Horn -- 0 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 13214 Freeze Out Mtns Ks Expedition WY Converse -- 0 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 74664 Jimbo Quarry 2A WY Converse -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 PaleoDB 74665 Jimbo Quarry 2B WY Converse -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58511 Blacktail Creek WY Crook -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Lightning Rod Butte WY Crook 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49634 Inyan Kara Creek SDSM V9137 WY Crook -- 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56167 Little Houston, Mammal Quarry SDSM V941 WY Crook -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49636 WY-11 Little Houston, Main Quarry SDSM V9138 WY Crook -- 1 0 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Warm Springs Ranch I WY Crook 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56165 Mile 175, SDSM V931 WY Crook -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49637 SDSM V9139 WY Crook -- 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22650 BS Quarry + S Quarry, Warm Springs Ranch WY Hot Springs -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22678 RB Quarry, Warm Springs Ranch WY Hot Springs -- 0 1 1 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 68100 FS Quarry, Foot Site, Warm Springs Ranch WY Hot Springs -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55320 SI Unit 1, Warm Springs Ranch WY Hot Springs -- 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 142334 Brown's Ranch (Elk Mountain) WY Johnson -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55349 Buffalo Quarry, Smith Ranch WY Johnson -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 176983 WY-72 Poison Creek Quarry 1 WY Johnson -- 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 176984 WY-73 Poison Creek Quarry 2 WY Johnson -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 176985 WY-74 Poison Creek Quarry 3 WY Johnson -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58510 WY-71 Poison Creek WY Johnson -- 1 0 1 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 PaleoDB 48565 Red Fork Powder River Quarry A WY Johnson -- 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58509 Red Fork Powder River Quarry B WY Johnson -- 1 0 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Sheridan College Allosaurus WY Johnson 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Sheridan College 1 WY Johnson 1 0 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Sheridan College 2 WY Johnson 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 106532 Lance Creek Area, Dyslocosaurus type WY Niobrara -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 PaleoDB 58508 Paint Creek WY Park -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 126540 Dana WY Washakie -- 1 1 1 0 1 0 1 1 1 1 0 0 0 0 0 0 0 0 1 0 0 0 1 0 0 1 0 0 0 1 0 0 0 0 PaleoDB 58507 KU Camarasaurus WY Weston -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 SOUTH 26 localities PDBD # T&P 99 # Locality Name Age State County Member Apatosaurus Barosaurus Diplodocus Galeamopus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus PaleoDB 70024 Pinon AZ Not Reported 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49338 OK-1 Stovall's Pit 1, OMNH Quarry 1, Kenton (OMNH V92) OK Cimarron Brushy Basin 1 1 1 1 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 49598 OK-4 Stovall's Pit 8, Kenton OK Cimarron Brushy Basin 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49596 OK-2 Stovall's Pit 5, Kenton (OMNH) OK Cimarron Not Reported 1 0 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49597 OK-3 Stovall's Pit 6, Kenton OK Cimarron Not Reported 1 0 1 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56421 Stovall's Pit 3A, Kenton OK Cimarron Not Reported 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56422 Stovall's Pit 9, Kenton OK Cimarron Not Reported 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 51990 Hagan Basin, NMMNH locality L-210 NM Sandoval Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38872 Lee Acres, NMMNH L-369 NM Sandoval Brushy Basin 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 119021 McConnell I NM Sandoval Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22717 Peterson Quarry, NMMNH L-3282, ETE Locality 1593 NM Bernalillo Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 68099 NM-3 San Ysidro Camarasaurus, NMMNH L-555, Ridby's Camarasaurus NM Sandoval Brushy Basin 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 San Ysidro 2 NM Sandoval 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38871 NMMNH L-3285 NM Sandoval Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38867 NMMNH L-3281 NM Union Brushy Basin 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 51992 NM-4 Seismosaurus Quarry, NMMNH locality L-344 NM Sandoval Jackpile Sandstone 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57910 NM-1 Acoma Site NM Cibola Not Reported 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 36265 Suwanee Peak NM Cibola Not Reported 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Boney Canyon NM Cibola 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Concho Springs NM Cibola 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Bull Canyon NM Guadalupe 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Exeter NM Union Not Reported 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 51991 Hagan Basin, NMMNH locality L-211 NM Sandoval Recapture 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Quay County NM Quay 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Blue Peak NM McKinley 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 84308 Section 19 Mine NM McKinley Salt Wash 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 EAST 126 localities Age PDBD # T&P # Locality name State County Member Apatosaurus Barosaurus Diplodocus Galeamopus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus Garden of the Gods CO El Paso -- 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Cleveland Quarry CO Fremont -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 PaleoDB 48127 Cope Quarry 1, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48128 Cope Quarry II, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48129 Cope Quarry III, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48130 Cope Quarry IV, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48146 Cope Quarry V, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48147 Cope Quarry VI, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48148 Cope Quarry VII, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48149 CO-1 Cope Quarry VIII, The Fort, Cam Quarry No. 2 CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48153 Cope Quarry XIV CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48158 CO-76 Cope Quarry XV CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48154 CO-71 CopeQuarry XII CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48155 CO-5 Delfs CO Fremont -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 PaleoDB 48163 CO-6 Deweese slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48156 CO-72 Egg Gulch CO Fremont -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Eric's Tooth CO Fremont 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39377 CO-4 Felch Quarry 2 CO Fremont -- 0 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48164 Greg's Bone - Garden Park slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48170 CO-75 Kenny's Stegosaurus, Garden Park CO Fremont -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48160 CO-7 Kessler's slightly older than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48162 CO-8 Lindsey Quarry, Garden Park slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48168 Lucas's Site, Garden Park CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 28366 CO-3 Marsh- Felch Quarry, Garden Park CO Fremont -- 1 0 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0 1 0 0 0 0 PaleoDB 48172 CO-73 Meyer Site 1, Garden Park CO Fremont -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48171 CO-73 Meyer Site 2, Garden Park CO Fremont -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48169 CO-74 Meyer Site 3, Garden Park CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Meyer's Pump House CO Fremont 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Sauropod Quarry CO Fremont 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 PaleoDB 48161 CO-11 Small Quarry, Garden Park slightly older than 152.29 ± 0.27 Ma CO Fremont -- 1 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48166 CO-09 Valley of Death, Garden Park CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 70026 Wilson Creek CO Fremont -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 66909 Alameda Parkway Morrison dinosaur site CO Jefferson -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39376 Dinosaur Ridge, Lake's Quarry 8 CO Jefferson -- 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Lakes Quarry 1 CO Jefferson 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 13341 Lakes Quarry 10 CO Jefferson -- 1 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 28382 Lakes Quary 5, Beckwith Quarry CO Jefferson -- 0 0 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Horsetooth CO Larimer 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 70029 Masonville CO Larimer -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Purgatoire River CO Las Animas 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 70030 Villegreen CO Las Animas -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Webster Park CO Park -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 164787 Gordon-Bramson-Brothers - Snowmass CO Pitkin -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 Los Ochos Mine CO Saguache 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Heeney Site CO Summit 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 AMNH 550 WY Albany 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55486 WY-42 Bernice WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55487 WY-44 Bertha between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Albany -- 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22648 WY-78 Bone Cabin slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 1 1 1 1 1 1 0 0 0 0 0 0 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 49577 WY-79 Bone Cabin Quarry West slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 1 1 0 1 1 1 1 1 0 0 0 0 0 0 0 1 0 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0 Boris Quarry WY Albany 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 177492 Breakfast Bench WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Brown Quarry A WY Albany 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39372 Brown Quarry B WY Albany 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39373 Brown Quarry C WY Albany -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39374 Brown Quarry D WY Albany -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Brown Quarry G WY Albany 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Cam Bench WY Albany 0 0 0 0 1 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57902 Carnegie Quarry 4, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 142337 Carnegie Quarry 5, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 142335 Carnegie Quarry A, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 142336 Carnegie Quarry B, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 52815 Carnegie Quarry C, Sheep Creek WY Albany -- 1 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 13352 WY-91 Carnegie Quarry D WY Albany -- 1 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57897 Carnegie Quarry D(3), Sheep Creek WY Albany -- 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 52816 Carnegie Quarry E, Sheep Creek WY Albany -- 1 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57898 Carnegie Quarry F, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57899 Carnegie Quarry G, Sheep Creek WY Albany -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57900 Carnegie Quarry J, Sheep Creek WY Albany -- 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57901 Carnegie Quarry K, Sheep Creek WY Albany -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48837 Carnegie Quarry N, Sheep Creek WY Albany -- 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 Cassiopeia Quarry WY Albany 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 74000 Como Bluff (AMNH 222) Bone Cabin? slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 168056 Como Bluff (AMNH 223) Bone Cabin? slightly older than 152.51 ± 0.47 Ma WY Albany -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Cope's Como 1 WY Albany 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Cope's Como 3 WY Albany 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Cope's Como 4 WY Albany 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Cope's Como 5 WY Albany 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Convention site WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Drinker Quarry WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 47065 WY-86 East Camarasaurus 1993 (BCQ) slightly older than 152.51 ± 0.47 Ma WY Albany -- 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 EP Thompson WY Albany 0 0 1 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Haystack Quarry WY Albany 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Jeff P WY Albany 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49628 WY-22 Louise WY Albany -- 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Mummy WY Albany 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49627 WY-39 Nail Quarry older than 152.51 ± 0.47 Ma WY Albany -- 1 0 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56654 WY-82 Ninemile Hill very close to 152.51 ± 0.47 Ma WY Albany -- 0 0 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Okie WY Albany 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49629 op-KAT WY Albany -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Pat M WY Albany 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39368 WY-17 Reed's Quarry 10 slightly younger than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39369 WY-18 Reed's Quarry 11 WY Albany -- 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39253 WY-46 Reed's Quarry 13 between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Albany -- 0 0 1 0 1 1 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 39371 WY-16 Reed's Quarry 14 WY Albany -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39339 WY-23 Reed's Quarry 2 slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39340 WY-20 Reed's Quarry 3 WY Albany -- 0 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39347 WY-19 Reed's Quarry 8 WY Albany -- 0 0 1 0 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39431 Quarry 9 1968-1970 152.51 ± 0.47 Ma WY Albany -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57903 Reed's Quarry R WY Albany -- 1 0 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55330 Red Mountain WY Albany -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 13212 WY-21 Reed's Quarry 1 WY Albany -- 1 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39342 WY-40 Reed's Quarry 4 WY Albany -- 0 1 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39346 WY-28 Reed's Quarry 7, Three Trees WY Albany -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 12816 WY-15 Reed's Quarry 9 152.51 ± 0.47 Ma WY Albany -- 0 0 0 0 1 1 1 1 0 1 0 0 0 0 1 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 Regina Quarry WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55483 Stego 99 very close to 152.51 ± 0.47 Ma WY Albany -- 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Two Thigh WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Weege Boys WY Albany 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 75453 WY-92 Zane Quarry, Sheep Creek WY Albany -- 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 73998 Aurora Quarry 3 Como Bluff WY Carbon -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58512 Dyer Ranch Quarry 1 WY Carbon -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58513 Dyer Ranch Quarry 2 WY Carbon -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57893 Freezeout Hills Quarry 4 WY Carbon -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57894 Freezeout Hills Quarry 6 WY Carbon -- 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57895 Freezeout Hills Quarry L WY Carbon -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57896 Freezeout Hills Quarry O WY Carbon -- 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55484 Graff Ranch (Matt Quarry) between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Carbon -- 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39337 WY-27 Lakes' 1A, Big Canyon WY Carbon -- 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38884 WY-83 Meilyn Quarry between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Carbon -- 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39338 WY-30 Quarry 1.5 WY Carbon -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39344 WY-26 Quarry 5 WY Carbon -- 0 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39370 WY-29 Robber's Roost (Reed's Q 12) WY Carbon -- 0 0 1 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 74663 Alcova Lake - Dave's othnielia (Cottonwood Creek 1) WY Natrona -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 Allen's Alto WY Natrona 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Cottonwood Creek Quarry 2 WY Natrona 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Cottonwood Creek Quarry 3 WY Natrona 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48557 Stegosaurus longispinus type locality, Alcova WY Natrona -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 WEST 74 localities PDBD # T&P # Locality Name Age State County Member Apatosaurus Barosaurus Diplodocus Galeamopus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus PaleoDB 52872 Middle Park CO Grand -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 70027 Radium CO Grand -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Blue Mesa CO Gunnison Brushy Basin 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38883 CO-49 Dino Cove 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 1 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 154925 Dinosaur Beach 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 154924 Northern Dinosaur Beach 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 1 0 1 0 1 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49621 CO-48 Cabin Creek CO Gunnison Salt Wash 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56586 CO-27 Bollan Stegosaur site - Rabbit Valley CO Mesa -- 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 70032 CO-55 Cactus Park (BYU 681) CO Mesa -- 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 35293 CO-56 Dominguez/Jones CO Mesa Brushy Basin 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49652 CO-33 Eriksen Ceratosaur FPA 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22708 FPA Quarry 4 CO Mesa Brushy Basin 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 71476 FPA General SIte CO Mesa Brushy Basin 1 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 92360 FPA George's "Coelurosaur" Site, LACM Loc. 5576 CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 FPA Stego Knoll CO Mesa Brushy Basin 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56684 Riggs Hill (Holt Quarry) CO Mesa Brushy Basin 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 48124 CO-57 Hups - Cactus Park CO Mesa Brushy Basin 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 PaleoDB 55329 Kings View FruitaMWC L-2002-11 CO Mesa Brushy Basin 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 53040 CO-33 Main Callison Quarry, FPA 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 11611 CO-21 Mygatt-Moore 152.18 ± 0.29 Ma CO Mesa Brushy Basin 1 1 1 0 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 PaleoDB 56599 CO-26 Rabbit Valley East slightly older than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56596 CO-22 Rabbit Valley Camarasaurus slightly younger than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56590 CO-23 Rabbit Valley Iguanodon older than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 1 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 11924 Riggs Quarry 12, Redlands CO Mesa Brushy Basin 0 0 0 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 36576 CO-36 Riggs Quarry 15 CO Mesa Brushy Basin 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56683 CO-35 Riggs Quarry 13 CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 Split Rock 1 CO Mesa 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 Split Rock 2 CO Mesa 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Wolny CO Mesa 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 70031 Cactus Park 3A CO Mesa Not Reported 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 68097 Sinbad Valley CO Mesa Not Reported 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 53022 CO-34 Kirkland site slightly older than 152.0 ± 0.3 Ma CO Mesa Salt Wash 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 11925 Riggs Quarry 14 CO Mesa Salt Wash 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Homestead CO Moffat 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Lily Park CO Moffat 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 MF Amphitheater CO Moffat 0 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Witherell CO Moffat 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 Wolf Creek CO Moffat 0 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 25141 Calico Gulch - Lily Park CO Moffat Not Reported 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 164830 McElmo Canyon CO Montezuma Brushy Basin 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Yellowjacket Canyon CO Montezuma 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 45975 CO-58 Dry Mesa CO Montrose Brushy Basin 1 1 1 0 1 1 1 1 1 1 0 0 0 0 1 1 0 1 1 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 PaleoDB 49622 CO-59 Jones Hole Quarry, Dry Mesa 2 CO Montrose Brushy Basin 0 1 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49623 CO-60 Potter Creek CO Montrose Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 53041 CO-66 Sheetz Quarry 1, Uravan CO Montrose Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 Agate Basin UT Emery 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 46439 UT-7 Cleveland-Lloyd Dinosaur UT Emery Brushy Basin 1 0 1 0 1 1 1 1 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Duke Site UT Emery 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58551 Emery County sauropod site UT Emery Brushy Basin 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Ferron UT Emery 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49626 UT-9 Green River Dinosaur UT Emery Brushy Basin 0 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Mussentuchit Quarry UT Emery 0 0 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 San Rafael UT Emery 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 64053 UT-8 Sand Bench UT Emery Brushy Basin 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 66 Quarry UT Emery 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 47196 Willow Springs, BYU ESM-163R UT Emery Brushy Basin 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 Peterson Site UT Emery 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49614 Hinkle Site UT Grand Brushy Basin 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58516 Mill Canyon UT Grand Brushy Basin 1 0 0 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58518 Cisco sauropod site UT Grand Not Reported 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58517 Floy Junction UT Grand Not Reported 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 104715 La Sal Mountains UT San Juan Brushy Basin 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 28360 UT-4 East Canyon ~157 Ma UT San Juan Tidwell 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49595 Brush Creek (Marsh Allo) UT Uintah Brushy Basin 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Chevron's Morrison North UT Uintah 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Chevron's Morrison South UT Uintah 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22711 UT-25 Rainbow Park (DNM 94) younger than 151 Ma UT Uintah Brushy Basin 0 0 1 0 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 52110 UT-19 Dinosaur National Monument (DNM 315) 10 m above Quarry Ss) younger than 151 Ma UT Uintah Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 96826 UT-12 Jensen/Jensen UT Uintah Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 55350 Nielson Gulch, DMN, Marshosaurus UT Uintah Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57913 UT-26 Rainbow Park microsite younger than 151 Ma UT Uintah Brushy Basin 0 0 0 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 21852 UT-18 Dinosaur National Monument Carnegie Quarry 150.91 ± 0.43 Ma UT Uintah Brushy Basin 1 1 1 0 1 1 1 1 1 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58515 UT-13 Utah Field House UT Uintah Not Reported 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58514 Hanksville-Burpee UT Wayne Brushy Basin 1 1 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 NORTH NORTH 65 localities PDBD # T&P 99 # Locality Name State County Member Apatosaurus Barosaurus Diplodocus Galeamopus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus PaleoDB 50714 Horse Coulee, AMHN 03-30 MT Big Horn -- 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 38866 Rattlesnake RIdge MT Carbon -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 PaleoDB 69890 MT-1 Mother's Day MT Carbon -- 0 0 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 T&J Quarry MT Gallatin -- 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 49624 MT-2 Upper Strickland Creek MT Park -- 0 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 Wittecombe's Ranch MT Wheatland -- 0 0 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 49633 Parker's Mystery SD Fall River -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49630 SD-2 Fuller's 351, St. Onge SD Lawrence -- 1 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 49625 SD-1 Bear Butte SD Meade -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 57905 SD-3 Piedmont Butte, Hatch Ranch, Barosaurus type, SDSM V9140 SD Meade -- 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 132242 AHMN quarries, Sturgis area SD Meade -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 132243 USNM Quarry, Sturgis area SD Meade -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49631 SD-4 Wonderland, Piedmont SD Meade -- 0 1 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 57904 SD-5 Wonderland North SD Meade -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 46468 WY-61 Big Al SIte WY Big Horn -- 0 0 1 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 Big Butte WY Big Horn 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 Smithsonian Quarry 1 WY Big Horn 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Dyer Ranch 1 and 2 WY Big Horn 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 Freezeout Hills General WY Big Horn 0 0 1 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 5 PaleoDB 57893 Freezeout Hills Quarry 4 WY Big Horn 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 57894 Freezeout Hills Quarry 6 WY Big Horn 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 57895 Freezeout Hills Quarry L WY Big Horn 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Freezeout Hills Quarry N WY Big Horn 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 5 PaleoDB 57896 Freezeout Hills Quarry O WY Big Horn 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 46460 WY-62 Howe Quarry WY Big Horn -- 1 1 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 5 PaleoDB 46462 Howe Quarry 2, Sauriermuseum WY Big Horn -- 0 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 3 PaleoDB 46465 Howe-Stephens Quarry WY Big Horn -- 1 1 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0 0 1 0 0 0 1 0 0 0 10 Lake's Quarry WY Big Horn 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 46469 Spring Hill, Howe Ranch WY Big Horn -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 55333 Fox Mesa WY Big Horn -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Matt Quarry WY Big Horn 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 Ninemile Hill WY Big Horn 0 0 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 55323 ?reopened by C. Weege 1992? Ninemile Quarry (AMNH), Nine Mile Crossing, Nine Mile Quarry WY Big Horn -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Reed's Quarry 1.5 WY Big Horn 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Reed's Quarry 5 WY Big Horn 0 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 4 Reed's Quarry 7 WY Big Horn 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 Reed's Quarry 12 WY Big Horn 0 0 1 0 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 5 PaleoDB 62561 Red Canyon Ranch, Hope College REU WY Big Horn -- 0 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 13214 Freeze Out Mtns Ks Expedition WY Converse -- 0 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 74664 Jimbo Quarry 2A WY Converse -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 PaleoDB 74665 Jimbo Quarry 2B WY Converse -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 58511 Blacktail Creek WY Crook -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Lightning Rod Butte WY Crook 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49634 Inyan Kara Creek SDSM V9137 WY Crook -- 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 56167 Little Houston, Mammal Quarry SDSM V941 WY Crook -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 49636 WY-11 Little Houston, Main Quarry SDSM V9138 WY Crook -- 1 0 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 6 Warm Springs Ranch I WY Crook 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 56165 Mile 175, SDSM V931 WY Crook -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49637 SDSM V9139 WY Crook -- 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 22650 BS Quarry + S Quarry, Warm Springs Ranch WY Hot Springs -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 22678 RB Quarry, Warm Springs Ranch WY Hot Springs -- 0 1 1 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 6 PaleoDB 68100 FS Quarry, Foot Site, Warm Springs Ranch WY Hot Springs -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 55320 SI Unit 1, Warm Springs Ranch WY Hot Springs -- 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 142334 Brown's Ranch (Elk Mountain) WY Johnson -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 55349 Buffalo Quarry, Smith Ranch WY Johnson -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 176983 WY-72 Poison Creek Quarry 1 WY Johnson -- 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 176984 WY-73 Poison Creek Quarry 2 WY Johnson -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 176985 WY-74 Poison Creek Quarry 3 WY Johnson -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 58510 WY-71 Poison Creek WY Johnson -- 1 0 1 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 7 PaleoDB 48565 Red Fork Powder River Quarry A WY Johnson -- 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 58509 Red Fork Powder River Quarry B WY Johnson -- 1 0 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 Sheridan College Allosaurus WY Johnson 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Sheridan College 1 WY Johnson 1 0 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 5 Sheridan College 2 WY Johnson 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 106532 Lance Creek Area, Dyslocosaurus type WY Niobrara -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 PaleoDB 58508 Paint Creek WY Park -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 126540 Dana WY Washakie -- 1 1 1 0 1 0 1 1 1 1 0 0 0 0 0 0 0 0 1 0 0 0 1 0 0 1 0 0 0 1 0 0 0 12 PaleoDB 58507 KU Camarasaurus WY Weston -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 3 WEST WEST 72 localities PDBD # T&P # Locality Name Age State County Member Apatosaurus Barosaurus Diplodocus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus Blue Mesa CO Gunnison Brushy Basin 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 38883 CO-49 Dino Cove 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 1 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 154925 Dinosaur Beach 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 154924 Northern Dinosaur Beach 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 1 0 1 1 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 6 PaleoDB 49621 CO-48 Cabin Creek CO Gunnison Salt Wash 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 56586 CO-27 Bollan Stegosaur site - Rabbit Valley CO Mesa -- 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 70032 CO-55 Cactus Park (BYU 681) CO Mesa -- 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 35293 CO-56 Dominguez/Jones CO Mesa Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49652 CO-33 Eriksen Ceratosaur FPA 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 22708 FPA Quarry 4 CO Mesa Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 71476 FPA General SIte CO Mesa Brushy Basin 1 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 5 PaleoDB 92360 FPA George's "Coelurosaur" Site, LACM Loc. 5576 CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 FPA Stego Knoll CO Mesa Brushy Basin 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 56684 Riggs Hill (Holt Quarry) CO Mesa Brushy Basin 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 3 PaleoDB 48124 CO-57 Hups - Cactus Park CO Mesa Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 2 PaleoDB 55329 Kings View FruitaMWC L-2002-11 CO Mesa Brushy Basin 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 53040 CO-33 Main Callison Quarry, FPA 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 11611 CO-21 Mygatt-Moore 152.18 ± 0.29 Ma CO Mesa Brushy Basin 1 1 1 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 7 PaleoDB 56599 CO-26 Rabbit Valley East slightly older than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 56596 CO-22 Rabbit Valley Camarasaurus slightly younger than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 56590 CO-23 Rabbit Valley Iguanodon older than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 11924 Riggs Quarry 12, Redlands CO Mesa Brushy Basin 0 0 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 36576 CO-36 Riggs Quarry 15 CO Mesa Brushy Basin 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 56683 CO-35 Riggs Quarry 13 CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 Split Rock 1 CO Mesa 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 2 Split Rock 2 CO Mesa 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Wolny CO Mesa 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 70031 Cactus Park 3A CO Mesa Not Reported 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 68097 Sinbad Valley CO Mesa Not Reported 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 53022 CO-34 Kirkland site slightly older than 152.0 ± 0.3 Ma CO Mesa Salt Wash 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 11925 Riggs Quarry 14 CO Mesa Salt Wash 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Homestead CO Moffat 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 2 Lily Park CO Moffat 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 2 MF Amphitheater CO Moffat 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 Witherell CO Moffat 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 2 Wolf Creek CO Moffat 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 25141 Calico Gulch - Lily Park CO Moffat Not Reported 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 164830 McElmo Canyon CO Montezuma Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Yellowjacket Canyon CO Montezuma 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 45975 CO-58 Dry Mesa CO Montrose Brushy Basin 1 1 1 1 1 1 1 1 1 0 0 0 0 1 1 0 1 1 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 14 PaleoDB 49622 CO-59 Jones Hole Quarry, Dry Mesa 2 CO Montrose Brushy Basin 0 1 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 49623 CO-60 Potter Creek CO Montrose Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 PaleoDB 53041 CO-66 Sheetz Quarry 1, Uravan CO Montrose Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Agate Basin UT Emery 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 46439 UT-7 Cleveland-Lloyd Dinosaur UT Emery Brushy Basin 1 0 1 1 1 1 1 1 1 0 0 0 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 11 Duke Site UT Emery 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 58551 Emery County sauropod site UT Emery Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Ferron UT Emery 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49626 UT-9 Green River Dinosaur UT Emery Brushy Basin 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 Mussentuchit Quarry UT Emery 0 0 1 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 San Rafael UT Emery 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 64053 UT-8 Sand Bench UT Emery Brushy Basin 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 66 Quarry UT Emery 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 47196 Willow Springs, BYU ESM-163R UT Emery Brushy Basin 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 4 Peterson Site UT Emery 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49614 Hinkle Site UT Grand Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 58516 Mill Canyon UT Grand Brushy Basin 1 0 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 5 PaleoDB 58518 Cisco sauropod site UT Grand Not Reported 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 58517 Floy Junction UT Grand Not Reported 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 104715 La Sal Mountains UT San Juan Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 28360 UT-4 East Canyon ~157 Ma UT San Juan Tidwell 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 PaleoDB 49595 Brush Creek (Marsh Allo) UT Uintah Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Chevron's Morrison North UT Uintah 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Chevron's Morrison South UT Uintah 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 22711 UT-25 Rainbow Park (DNM 94) younger than 151 Ma UT Uintah Brushy Basin 0 0 1 0 1 1 1 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 6 PaleoDB 52110 UT-19 Dinosaur National Monument (DNM 315) 10 m above Quarry Ss) younger than 151 Ma UT Uintah Brushy Basin 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 96826 UT-12 Jensen/Jensen UT Uintah Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 1 PaleoDB 55350 Nielson Gulch, DMN, Marshosaurus UT Uintah Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 57913 UT-26 Rainbow Park microsite younger than 151 Ma UT Uintah Brushy Basin 0 0 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 5 PaleoDB 21852 UT-18 Dinosaur National Monument Carnegie Quarry 150.91 ± 0.43 Ma UT Uintah Brushy Basin 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 9 PaleoDB 58515 UT-13 Utah Field House UT Uintah Not Reported 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 58514 Hanksville-Burpee UT Wayne Brushy Basin 1 1 1 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 7 16 EAST EAST 128 localities Age PDBD # T&P # Locality name State County Member Apatosaurus Barosaurus Diplodocus Galeamopus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Garden of the Gods CO El Paso -- 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 52872 Middle Park CO Grand -- 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 70027 Radium CO Grand -- 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Cleveland Quarry CO Fremont -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 PaleoDB 48127 Cope Quarry 1, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 48128 Cope Quarry II, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 48129 Cope Quarry III, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 48130 Cope Quarry IV, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48146 Cope Quarry V, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48147 Cope Quarry VI, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48148 Cope Quarry VII, Cope's Nipple, Saurian Hill CO Fremont -- 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 48149 CO-1 Cope Quarry VIII, The Fort, Cam Quarry No. 2 CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48153 Cope Quarry XIV CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48158 CO-76 Cope Quarry XV CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48154 CO-71 CopeQuarry XII CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48155 CO-5 Delfs CO Fremont -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 PaleoDB 48163 CO-6 Deweese slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48156 CO-72 Egg Gulch CO Fremont -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 Eric's Tooth CO Fremont 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39377 CO-4 Felch Quarry 2 CO Fremont -- 0 0 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 48164 Greg's Bone - Garden Park slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48170 CO-75 Kenny's Stegosaurus, Garden Park CO Fremont -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48160 CO-7 Kessler's slightly older than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 48162 CO-8 Lindsey Quarry, Garden Park slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 48168 Lucas's Site, Garden Park CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 28366 CO-3 Marsh- Felch Quarry, Garden Park CO Fremont -- 1 0 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0 1 0 0 0 12 PaleoDB 48172 CO-73 Meyer Site 1, Garden Park CO Fremont -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48171 CO-73 Meyer Site 2, Garden Park CO Fremont -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48169 CO-74 Meyer Site 3, Garden Park CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Meyer's Pump House CO Fremont 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Sauropod Quarry CO Fremont 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 PaleoDB 48161 CO-11 Small Quarry, Garden Park slightly older than 152.29 ± 0.27 Ma CO Fremont -- 1 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 48166 CO-09 Valley of Death, Garden Park CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 70026 Wilson Creek CO Fremont -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 66909 Alameda Parkway Morrison dinosaur site CO Jefferson -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39376 Dinosaur Ridge, Lake's Quarry 8 CO Jefferson -- 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 Lakes Quarry 1 CO Jefferson 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 13341 Lakes Quarry 10 CO Jefferson -- 1 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 28382 Lakes Quary 5, Beckwith Quarry CO Jefferson -- 0 0 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 Horsetooth CO Larimer 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 70029 Masonville CO Larimer -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Purgatoire River CO Las Animas 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 70030 Villegreen CO Las Animas -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Webster Park CO Park -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 164787 Gordon-Bramson-Brothers - Snowmass CO Pitkin -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 Los Ochos Mine CO Saguache 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Heeney Site CO Summit 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 AMNH 550 WY Albany 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 55486 WY-42 Bernice WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 55487 WY-44 Bertha between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Albany -- 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 22648 WY-78 Bone Cabin slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 1 1 1 1 1 1 0 0 0 0 0 0 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 10 PaleoDB 49577 WY-79 Bone Cabin Quarry West slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 1 1 0 1 1 1 1 1 0 0 0 0 0 0 0 1 0 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 11 Boris Quarry WY Albany 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 177492 Breakfast Bench WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Brown Quarry A WY Albany 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39372 Brown Quarry B WY Albany 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39373 Brown Quarry C WY Albany -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39374 Brown Quarry D WY Albany -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Brown Quarry G WY Albany 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Cam Bench WY Albany 0 0 0 0 1 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 57902 Carnegie Quarry 4, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 142337 Carnegie Quarry 5, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 142335 Carnegie Quarry A, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 142336 Carnegie Quarry B, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 52815 Carnegie Quarry C, Sheep Creek WY Albany -- 1 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 13352 WY-91 Carnegie Quarry D WY Albany -- 1 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 57897 Carnegie Quarry D(3), Sheep Creek WY Albany -- 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 52816 Carnegie Quarry E, Sheep Creek WY Albany -- 1 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 57898 Carnegie Quarry F, Sheep Creek WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 57899 Carnegie Quarry G, Sheep Creek WY Albany -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 57900 Carnegie Quarry J, Sheep Creek WY Albany -- 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 57901 Carnegie Quarry K, Sheep Creek WY Albany -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48837 Carnegie Quarry N, Sheep Creek WY Albany -- 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 5 Cassiopeia Quarry WY Albany 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 74000 Como Bluff (AMNH 222) Bone Cabin? slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 168056 Como Bluff (AMNH 223) Bone Cabin? slightly older than 152.51 ± 0.47 Ma WY Albany -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Cope's Como 1 WY Albany 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Cope's Como 3 WY Albany 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Cope's Como 4 WY Albany 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 Cope's Como 5 WY Albany 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Convention site WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Drinker Quarry WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 47065 WY-86 East Camarasaurus 1993 (BCQ) slightly older than 152.51 ± 0.47 Ma WY Albany -- 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 3 EP Thompson WY Albany 0 0 1 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 Haystack Quarry WY Albany 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 Jeff P WY Albany 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49628 WY-22 Louise WY Albany -- 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 5 Mummy WY Albany 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 49627 WY-39 Nail Quarry older than 152.51 ± 0.47 Ma WY Albany -- 1 0 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 6 PaleoDB 56654 WY-82 Ninemile Hill very close to 152.51 ± 0.47 Ma WY Albany -- 0 0 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 5 Okie WY Albany 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 49629 op-KAT WY Albany -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Pat M WY Albany 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39368 WY-17 Reed's Quarry 10 slightly younger than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39369 WY-18 Reed's Quarry 11 WY Albany -- 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 39253 WY-46 Reed's Quarry 13 between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Albany -- 0 0 1 0 1 1 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 8 PaleoDB 39371 WY-16 Reed's Quarry 14 WY Albany -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39339 WY-23 Reed's Quarry 2 slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39340 WY-20 Reed's Quarry 3 WY Albany -- 0 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 39347 WY-19 Reed's Quarry 8 WY Albany -- 0 0 1 0 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 5 PaleoDB 39431 Quarry 9 1968-1970 152.51 ± 0.47 Ma WY Albany -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57903 Reed's Quarry R WY Albany -- 1 0 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 55330 Red Mountain WY Albany -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 13212 WY-21 Reed's Quarry 1 WY Albany -- 1 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 39342 WY-40 Reed's Quarry 4 WY Albany -- 0 1 1 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 5 PaleoDB 39346 WY-28 Reed's Quarry 7, Three Trees WY Albany -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 12816 WY-15 Reed's Quarry 9 152.51 ± 0.47 Ma WY Albany -- 0 0 0 0 1 1 1 1 0 1 0 0 0 0 1 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 9 Regina Quarry WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 55483 Stego 99 very close to 152.51 ± 0.47 Ma WY Albany -- 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 Two Thigh WY Albany 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Weege Boys WY Albany 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 75453 WY-92 Zane Quarry, Sheep Creek WY Albany -- 1 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 73998 Aurora Quarry 3 Como Bluff WY Carbon -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 58512 Dyer Ranch Quarry 1 WY Carbon -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 58513 Dyer Ranch Quarry 2 WY Carbon -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 57893 Freezeout Hills Quarry 4 WY Carbon -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 57894 Freezeout Hills Quarry 6 WY Carbon -- 1 0 0 0 1 1 0 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 5 PaleoDB 57895 Freezeout Hills Quarry L WY Carbon -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 57896 Freezeout Hills Quarry O WY Carbon -- 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 55484 Graff Ranch (Matt Quarry) between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Carbon -- 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39337 WY-27 Lakes' 1A, Big Canyon WY Carbon -- 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 38884 WY-83 Meilyn Quarry between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Carbon -- 0 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 39338 WY-30 Quarry 1.5 WY Carbon -- 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 39344 WY-26 Quarry 5 WY Carbon -- 0 0 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 4 PaleoDB 39370 WY-29 Robber's Roost (Reed's Q 12) WY Carbon -- 0 0 1 0 1 1 0 1 0 1 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 7 PaleoDB 74663 Alcova Lake - Dave's othnielia (Cottonwood Creek 1) WY Natrona -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 Allen's Alto WY Natrona 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Cottonwood Creek Quarry 2 WY Natrona 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 Cottonwood Creek Quarry 3 WY Natrona 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 48557 Stegosaurus longispinus type locality, Alcova WY Natrona -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 36 SOUTH SOUTH 26 localities PDBD # T&P 99 # Locality Name Age State County Member Apatosaurus Barosaurus Diplodocus Galeamopus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus PaleoDB 70024 Pinon AZ Not Reported 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49338 OK-1 Stovall's Pit 1, OMNH Quarry 1, Kenton (OMNH V92) OK Cimarron Brushy Basin 1 1 1 1 1 1 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 49598 OK-4 Stovall's Pit 8, Kenton OK Cimarron Brushy Basin 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49596 OK-2 Stovall's Pit 5, Kenton (OMNH) OK Cimarron Not Reported 1 0 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49597 OK-3 Stovall's Pit 6, Kenton OK Cimarron Not Reported 1 0 1 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56421 Stovall's Pit 3A, Kenton OK Cimarron Not Reported 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56422 Stovall's Pit 9, Kenton OK Cimarron Not Reported 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 51990 Hagan Basin, NMMNH locality L-210 NM Sandoval Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38872 Lee Acres, NMMNH L-369 NM Sandoval Brushy Basin 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 119021 McConnell I NM Sandoval Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22717 Peterson Quarry, NMMNH L-3282, ETE Locality 1593 NM Bernalillo Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 68099 NM-3 San Ysidro Camarasaurus, NMMNH L-555, Ridby's Camarasaurus NM Sandoval Brushy Basin 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 San Ysidro 2 NM Sandoval 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38871 NMMNH L-3285 NM Sandoval Brushy Basin 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38867 NMMNH L-3281 NM Union Brushy Basin 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 51992 NM-4 Seismosaurus Quarry, NMMNH locality L-344 NM Sandoval Jackpile Sandstone 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57910 NM-1 Acoma Site NM Cibola Not Reported 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 36265 Suwanee Peak NM Cibola Not Reported 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Boney Canyon NM Cibola 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Concho Springs NM Cibola 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Bull Canyon NM Guadalupe 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Exeter NM Union Not Reported 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 51991 Hagan Basin, NMMNH locality L-211 NM Sandoval Recapture 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Quay County NM Quay 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Blue Peak NM McKinley 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 84308 Section 19 Mine NM McKinley Salt Wash 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 with ages only WEST PDBD # T&P # Locality Name Age State County Member Apatosaurus Barosaurus Diplodocus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Elaphrosaurus Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus PaleoDB 38883 CO-49 Dino Cove 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 1 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 154925 Dinosaur Beach 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 154924 Northern Dinosaur Beach 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 1 0 1 1 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 6 PaleoDB 56586 CO-27 Bollan Stegosaur site - Rabbit Valley slightly older than 152.18 ± 0.29 Ma CO Mesa -- 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 PaleoDB 49652 CO-33 Eriksen Certosaurus FPA 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 22708 CO-33 FPA Quarry 4 - main quarry 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 71476 FPA General Site - Fruitadens, mammals 152.0 ± 0.3 Ma CO Mesa Brushy Basin 1 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 5 PaleoDB 92360 FPA George's "Coelurosaur" Site, LACM Loc. 5576 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 53040 CO-33 Main Callison Quarry, FPA 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 11611 CO-21 Mygatt-Moore 152.18 ± 0.29 Ma CO Mesa Brushy Basin 1 1 1 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 7 PaleoDB 56599 CO-26 Rabbit Valley East slightly older than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 56596 CO-22 Rabbit Valley Camarasaurus slightly younger than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 56590 CO-23 Rabbit Valley Iguanodon older than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 3 PaleoDB 53022 CO-34 Kirkland site slightly older than 152.0 ± 0.3 Ma CO Mesa Salt Wash 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 46440 UT-25 Rainbow Park (DNM 94) younger than 151 Ma UT Uintah Brushy Basin 0 0 1 0 1 1 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 6 PaleoDB 52110 UT-19 Dinosaur National Monument (DNM 315) 10 m above Quarry Ss) younger than 151 Ma UT Uintah Brushy Basin 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 PaleoDB 57913 UT-26 Rainbow Park microsite younger than 151 Ma UT Uintah Brushy Basin 0 0 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 5 PaleoDB 21852 UT-18 Dinosaur National Monument Carnegie Quarry 150.91 ± 0.43 Ma UT Uintah Brushy Basin 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 9 PaleoDB 28360 UT-4 East Canyon ~157 Ma UT San Juan Tidwell 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 EAST PDBD # T&P # Locality name Age State County Member Apatosaurus Barosaurus Diplodocus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Elaphrosaurus Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus Galeamopus PaleoDB 48163 CO-6 Deweese slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48164 Greg's Bone - Garden Park slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48162 CO-8 Lindsey Quarry, Garden Park slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48161 CO-11 Small Quarry, Garden Park slightly older than 152.29 ± 0.27 Ma CO Fremont -- 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48160 CO-7 Kessler's slightly older than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 12816 WY-15 Reed's Quarry 9 152.51 ± 0.47 Ma WY Albany -- 0 0 0 1 1 1 1 0 1 0 0 0 0 0 1 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22648 WY-78 Bone Cabin slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 1 1 1 1 1 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 0 1 PaleoDB 47065 WY-86 East Camarasaurus 1993 (BCQ) slightly older than 152.51 ± 0.47 Ma WY Albany -- 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49627 WY-39 Nail Quarry older than 152.51 ± 0.47 Ma WY Albany -- 1 0 1 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56654 WY-82 Ninemile Hill very close to 152.51 ± 0.47 Ma WY Albany -- 0 0 1 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39368 WY-17 Quarry 10 slightly younger than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39253 WY-46 Quarry 13 between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Albany -- 0 0 1 1 1 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 0 0 PaleoDB 39339 WY-23 Quarry 2 slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39431 Quarry 9 1968-1970 152.51 ± 0.47 Ma WY Albany -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39342 WY-40 Reed's Quarry 4 between 157 and 152.51 ± 0.47 Ma WY Albany -- 0 1 1 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55487 WY-44 Bertha between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Albany -- 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 74000 Como Bluff (AMNH 222) Bone Cabin? slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 168056 Como Bluff (AMNH 223) Bone Cabin? slightly older than 152.51 ± 0.47 Ma WY Albany -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55483 Stego 99 very close to 152.51 ± 0.47 Ma WY Albany -- 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49577 WY-79 Bone Cabin Quarry West slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 1 0 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 PaleoDB 55484 Graff Ranch (Matt Quarry) between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Carbon -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38884 WY-83 Meilyn Quarry between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Carbon -- 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Age clusters WEST Age cluster PDBD # T&P # Locality Name Age State County Member Apatosaurus Barosaurus Diplodocus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Elaphrosaurus Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus younger than 151 PaleoDB 46440 UT-25 Rainbow Park (DNM 94) younger than 151 Ma UT Uintah Brushy Basin 0 0 1 0 1 1 1 0 0 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 52110 UT-19 Dinosaur National Monument (DNM 315) 10 m above Quarry Ss) younger than 151 Ma UT Uintah Brushy Basin 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 57913 UT-26 Rainbow Park microsite younger than 151 Ma UT Uintah Brushy Basin 0 0 0 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 ~151 Ma PaleoDB 38883 CO-49 Dino Cove 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 1 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 154925 Dinosaur Beach 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 154924 Northern Dinosaur Beach 151.43 ± 0.36 Ma CO Gunnison Brushy Basin 1 0 1 1 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 21852 UT-18 Dinosaur National Monument Carnegie Quarry 150.91 ± 0.43 Ma UT Uintah Brushy Basin 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ~ 152 Ma PaleoDB 49652 CO-33 Eriksen Certosaurus FPA 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22708 CO-33 FPA Quarry 4 - main quarry 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 71476 FPA General Site - Fruitadens, mammals 152.0 ± 0.3 Ma CO Mesa Brushy Basin 1 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 92360 FPA George's "Coelurosaur" Site, LACM Loc. 5576 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 53040 CO-33 Main Callison Quarry, FPA 152.0 ± 0.3 Ma CO Mesa Brushy Basin 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 11611 CO-21 Mygatt-Moore 152.18 ± 0.29 Ma CO Mesa Brushy Basin 1 1 1 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 PaleoDB 56599 CO-26 Rabbit Valley East slightly older than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56596 CO-22 Rabbit Valley Camarasaurus slightly younger than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 53022 CO-34 Kirkland site slightly older than 152.0 ± 0.3 Ma CO Mesa Salt Wash 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56586 CO-27 Bollan Stegosaur site - Rabbit Valley slightly older than 152.18 ± 0.29 Ma CO Mesa -- 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56590 CO-23 Rabbit Valley Iguanodon slightly older than 152.18 ± 0.29 Ma CO Mesa Brushy Basin 0 0 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 28360 UT-4 East Canyon ~157 Ma UT San Juan Tidwell 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 EAST Age cluster PDBD # T&P # Locality name Age State County Member Apatosaurus Barosaurus Diplodocus Camarasaurus Stegosaurus Camptosaurus Allosaurus Ceratosaurus Coelurus Saurophaganax Elaphrosaurus Fosterovenator Fruitadens Koparion Ornitholestes Stokesosaurus Tanycolagreus Marshosaurus Torvosaurus Drinker Dryosaurus Nanosaurus Othnielosaurus Uteodon Gargoyleosaurus Hesperosaurus Mymoorapelta Haplocanthosaurus Supersaurus Brachiosaurus Dyslocosaurus Suuwassea Kaatedocus Dystrophaeus Galeamopus ~ 152 Ma PaleoDB 48163 CO-6 Deweese slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48164 Greg's Bone - Garden Park slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48162 CO-8 Lindsey Quarry, Garden Park slightly younger than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48161 CO-11 Small Quarry, Garden Park slightly older than 152.29 ± 0.27 Ma CO Fremont -- 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 48160 CO-7 Kessler's slightly older than 152.29 ± 0.27 Ma CO Fremont -- 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 12816 WY-15 Reed's Quarry 9 152.51 ± 0.47 Ma WY Albany -- 0 0 0 1 1 1 1 0 1 0 0 0 0 0 1 0 0 0 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 22648 WY-78 Bone Cabin slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 1 1 1 1 1 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 0 1 PaleoDB 47065 WY-86 East Camarasaurus 1993 (BCQ) slightly older than 152.51 ± 0.47 Ma WY Albany -- 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49627 WY-39 Nail Quarry older than 152.51 ± 0.47 Ma WY Albany -- 1 0 1 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 56654 WY-82 Ninemile Hill very close to 152.51 ± 0.47 Ma WY Albany -- 0 0 1 1 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39368 WY-17 Quarry 10 slightly younger than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39339 WY-23 Quarry 2 slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39431 Quarry 9 1968-1970 152.51 ± 0.47 Ma WY Albany -- 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 74000 Como Bluff (AMNH 222) Bone Cabin? slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 168056 Como Bluff (AMNH 223) Bone Cabin? slightly older than 152.51 ± 0.47 Ma WY Albany -- 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55483 Stego 99 very close to 152.51 ± 0.47 Ma WY Albany -- 0 0 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 49577 WY-79 Bone Cabin Quarry West slightly older than 152.51 ± 0.47 Ma WY Albany -- 1 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 1 0 0 0 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 older than 152 Ma PaleoDB 39342 WY-40 Reed's Quarry 4 between 157 and 152.51 ± 0.47 Ma WY Albany -- 0 1 1 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 39253 WY-46 Quarry 13 between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Albany -- 0 0 1 1 1 1 1 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 1 0 0 0 0 PaleoDB 55484 Graff Ranch (Matt Quarry) between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Carbon -- 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 55487 WY-44 Bertha between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Albany -- 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 PaleoDB 38884 WY-83 Meilyn Quarry between 157 and 152.51 ± 0.47 Ma, closer to 157 WY Carbon -- 0 0 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0