id	sid	tid	token	lemma	pos
gs-3955	1	1	microsoft	microsoft	PROPN
gs-3955	1	2	word	word	NOUN
gs-3955	1	3	fullpdf07032025	fullpdf07032025	PROPN
gs-3955	1	4	georgian	georgian	ADJ
gs-3955	1	5	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	1	6	მეცნიერები	მეცნიერები	NOUN
gs-3955	1	7	ტ.	ტ.	VERB
gs-3955	1	8	7	7	NUM
gs-3955	1	9	n	n	PRON
gs-3955	1	10	3	3	NUM
gs-3955	1	11	,	,	PUNCT
gs-3955	1	12	2025	2025	NUM
gs-3955	1	13	346	346	NUM
gs-3955	1	14	georgian	georgian	PROPN
gs-3955	1	15	scientists	scientist	NOUN
gs-3955	1	16	ქართველი	ქართველი	ADJ
gs-3955	1	17	მეცნიერები	მეცნიერები	PROPN
gs-3955	1	18	vol	vol	NOUN
gs-3955	1	19	.	.	PUNCT
gs-3955	1	20	7	7	NUM
gs-3955	1	21	issue	issue	NOUN
gs-3955	1	22	3	3	NUM
gs-3955	1	23	,	,	PUNCT
gs-3955	1	24	2025	2025	NUM
gs-3955	1	25	https://doi.org/10.52340/gs.2025.07.03.24	https://doi.org/10.52340/gs.2025.07.03.24	NUM
gs-3955	1	26	ageless	ageless	ADJ
gs-3955	1	27	creatures	creature	NOUN
gs-3955	1	28	:	:	PUNCT
gs-3955	1	29	molecular	molecular	ADJ
gs-3955	1	30	insights	insight	NOUN
gs-3955	1	31	into	into	ADP
gs-3955	1	32	organisms	organism	NOUN
gs-3955	1	33	that	that	PRON
gs-3955	1	34	defy	defy	VERB
gs-3955	1	35	aging	age	VERB
gs-3955	1	36	david	david	PROPN
gs-3955	1	37	aphkhazava1	aphkhazava1	PROPN
gs-3955	1	38	,	,	PUNCT
gs-3955	1	39	nodar	nodar	PROPN
gs-3955	1	40	sulashvili2	sulashvili2	PROPN
gs-3955	1	41	,	,	PUNCT
gs-3955	1	42	george	george	PROPN
gs-3955	1	43	maglakelidze3	maglakelidze3	PROPN
gs-3955	1	44	,	,	PUNCT
gs-3955	1	45	jaba	jaba	PROPN
gs-3955	1	46	tkemaladze4	tkemaladze4	PROPN
gs-3955	2	1	1phd	1phd	NUM
gs-3955	2	2	professor	professor	NOUN
gs-3955	2	3	at	at	ADP
gs-3955	2	4	alte	alte	PROPN
gs-3955	2	5	university	university	PROPN
gs-3955	2	6	,	,	PUNCT
gs-3955	2	7	international	international	ADJ
gs-3955	2	8	school	school	NOUN
gs-3955	2	9	of	of	ADP
gs-3955	2	10	medicine	medicine	NOUN
gs-3955	2	11	;	;	PUNCT
gs-3955	2	12	invited	invite	VERB
gs-3955	2	13	professor	professor	NOUN
gs-3955	2	14	at	at	ADP
gs-3955	2	15	university	university	PROPN
gs-3955	2	16	of	of	ADP
gs-3955	2	17	georgia	georgia	PROPN
gs-3955	2	18	;	;	PUNCT
gs-3955	2	19	invited	invite	VERB
gs-3955	2	20	professor	professor	NOUN
gs-3955	2	21	at	at	ADP
gs-3955	2	22	georgian	georgian	PROPN
gs-3955	2	23	technical	technical	PROPN
gs-3955	2	24	university	university	PROPN
gs-3955	2	25	;	;	PUNCT
gs-3955	2	26	professor	professor	NOUN
gs-3955	2	27	at	at	ADP
gs-3955	2	28	georgian	georgian	PROPN
gs-3955	2	29	national	national	PROPN
gs-3955	2	30	university	university	PROPN
gs-3955	2	31	seu	seu	NOUN
gs-3955	2	32	;	;	PUNCT
gs-3955	2	33	corresponding	corresponding	ADJ
gs-3955	2	34	author	author	NOUN
gs-3955	2	35	:	:	PUNCT
gs-3955	2	36	professor	professor	PROPN
gs-3955	2	37	dr	dr	PROPN
gs-3955	2	38	.	.	PROPN
gs-3955	2	39	david	david	PROPN
gs-3955	2	40	aphkhazava	aphkhazava	PROPN
gs-3955	2	41	,	,	PUNCT
gs-3955	2	42	email	email	NOUN
gs-3955	2	43	:	:	PUNCT
gs-3955	2	44	d.apkhazava@alte.edu.ge	d.apkhazava@alte.edu.ge	NOUN
gs-3955	2	45	;	;	PUNCT
gs-3955	2	46	2md	2md	NUM
gs-3955	2	47	,	,	PUNCT
gs-3955	2	48	phd	phd	NOUN
gs-3955	2	49	,	,	PUNCT
gs-3955	2	50	doctor	doctor	NOUN
gs-3955	2	51	of	of	ADP
gs-3955	2	52	pharmaceutical	pharmaceutical	ADJ
gs-3955	2	53	and	and	CCONJ
gs-3955	2	54	pharmacological	pharmacological	ADJ
gs-3955	2	55	sciences	science	NOUN
gs-3955	2	56	in	in	ADP
gs-3955	2	57	medicine	medicine	NOUN
gs-3955	2	58	,	,	PUNCT
gs-3955	2	59	invited	invite	VERB
gs-3955	2	60	lecturer	lecturer	NOUN
gs-3955	2	61	(	(	PUNCT
gs-3955	2	62	invited	invite	VERB
gs-3955	2	63	professor	professor	NOUN
gs-3955	2	64	)	)	PUNCT
gs-3955	2	65	of	of	ADP
gs-3955	2	66	scientific	scientific	ADJ
gs-3955	2	67	research	research	NOUN
gs-3955	2	68	-	-	PUNCT
gs-3955	2	69	skills	skill	NOUN
gs-3955	2	70	center	center	NOUN
gs-3955	2	71	at	at	ADP
gs-3955	2	72	tbilisi	tbilisi	PROPN
gs-3955	2	73	state	state	PROPN
gs-3955	2	74	medical	medical	PROPN
gs-3955	2	75	university	university	PROPN
gs-3955	2	76	;	;	PUNCT
gs-3955	2	77	professor	professor	NOUN
gs-3955	2	78	of	of	ADP
gs-3955	2	79	medical	medical	ADJ
gs-3955	2	80	and	and	CCONJ
gs-3955	2	81	clinical	clinical	ADJ
gs-3955	2	82	pharmacology	pharmacology	NOUN
gs-3955	2	83	of	of	ADP
gs-3955	2	84	international	international	ADJ
gs-3955	2	85	school	school	NOUN
gs-3955	2	86	of	of	ADP
gs-3955	2	87	medicine	medicine	NOUN
gs-3955	2	88	at	at	ADP
gs-3955	2	89	alte	alte	PROPN
gs-3955	2	90	university	university	PROPN
gs-3955	2	91	;	;	PUNCT
gs-3955	2	92	professor	professor	NOUN
gs-3955	2	93	of	of	ADP
gs-3955	2	94	pharmacology	pharmacology	NOUN
gs-3955	2	95	of	of	ADP
gs-3955	2	96	faculty	faculty	NOUN
gs-3955	2	97	of	of	ADP
gs-3955	2	98	medicine	medicine	NOUN
gs-3955	2	99	at	at	ADP
gs-3955	2	100	georgian	georgian	PROPN
gs-3955	2	101	national	national	PROPN
gs-3955	2	102	university	university	PROPN
gs-3955	2	103	seu	seu	NOUN
gs-3955	2	104	;	;	PUNCT
gs-3955	2	105	associate	associate	ADJ
gs-3955	2	106	affiliated	affiliate	VERB
gs-3955	2	107	professor	professor	NOUN
gs-3955	2	108	of	of	ADP
gs-3955	2	109	medical	medical	ADJ
gs-3955	2	110	pharmacology	pharmacology	NOUN
gs-3955	2	111	of	of	ADP
gs-3955	2	112	faculty	faculty	NOUN
gs-3955	2	113	of	of	ADP
gs-3955	2	114	medicine	medicine	NOUN
gs-3955	2	115	at	at	ADP
gs-3955	2	116	sulkhan	sulkhan	PROPN
gs-3955	2	117	-	-	PUNCT
gs-3955	2	118	saba	saba	PROPN
gs-3955	2	119	orbeliani	orbeliani	PROPN
gs-3955	2	120	university	university	PROPN
gs-3955	2	121	;	;	PUNCT
gs-3955	2	122	tbilisi	tbilisi	PROPN
gs-3955	2	123	,	,	PUNCT
gs-3955	2	124	georgia	georgia	PROPN
gs-3955	2	125	.	.	PUNCT
gs-3955	3	1	orcid	orcid	NOUN
gs-3955	3	2	:	:	PUNCT
gs-3955	3	3	https://orcid.org/00000002-9005-8577	https://orcid.org/00000002-9005-8577	X
gs-3955	3	4	;	;	PUNCT
gs-3955	3	5	e	e	X
gs-3955	3	6	-	-	NOUN
gs-3955	3	7	mail	mail	NOUN
gs-3955	3	8	:	:	PUNCT
gs-3955	3	9	n.sulashvili@ug.edu.ge	n.sulashvili@ug.edu.ge	NOUN
gs-3955	3	10	;	;	PUNCT
gs-3955	3	11	3phd	3phd	NUM
gs-3955	3	12	,	,	PUNCT
gs-3955	3	13	professor	professor	NOUN
gs-3955	3	14	at	at	ADP
gs-3955	3	15	university	university	PROPN
gs-3955	3	16	of	of	ADP
gs-3955	3	17	georgia	georgia	PROPN
gs-3955	3	18	.	.	PUNCT
gs-3955	3	19	;	;	PUNCT
gs-3955	3	20	4director	4director	NUM
gs-3955	3	21	of	of	ADP
gs-3955	3	22	research	research	NOUN
gs-3955	3	23	;	;	PUNCT
gs-3955	3	24	longevity	longevity	NOUN
gs-3955	3	25	clinic	clinic	PROPN
gs-3955	3	26	,	,	PUNCT
gs-3955	3	27	inc	inc	PROPN
gs-3955	3	28	,	,	PUNCT
gs-3955	3	29	georgia	georgia	PROPN
gs-3955	3	30	.	.	PUNCT
gs-3955	4	1	orcid	orcid	PROPN
gs-3955	4	2	:	:	PUNCT
gs-3955	4	3	https://orcid.org/0000-0001-8651-7243	https://orcid.org/0000-0001-8651-7243	ADJ
gs-3955	4	4	;	;	PUNCT
gs-3955	4	5	e	e	X
gs-3955	4	6	-	-	NOUN
gs-3955	4	7	mail	mail	NOUN
gs-3955	4	8	:	:	PUNCT
gs-3955	4	9	jtkemaladze@longevity.ge	jtkemaladze@longevity.ge	PROPN
gs-3955	4	10	abstract	abstract	ADJ
gs-3955	4	11	while	while	SCONJ
gs-3955	4	12	aging	age	VERB
gs-3955	4	13	is	be	AUX
gs-3955	4	14	a	a	DET
gs-3955	4	15	nearly	nearly	ADV
gs-3955	4	16	universal	universal	ADJ
gs-3955	4	17	biological	biological	ADJ
gs-3955	4	18	process	process	NOUN
gs-3955	4	19	,	,	PUNCT
gs-3955	4	20	certain	certain	ADJ
gs-3955	4	21	species	specie	NOUN
gs-3955	4	22	exhibit	exhibit	VERB
gs-3955	4	23	negligible	negligible	ADJ
gs-3955	4	24	senescence	senescence	NOUN
gs-3955	4	25	,	,	PUNCT
gs-3955	4	26	maintaining	maintain	VERB
gs-3955	4	27	reproductive	reproductive	ADJ
gs-3955	4	28	capacity	capacity	NOUN
gs-3955	4	29	and	and	CCONJ
gs-3955	4	30	physiological	physiological	ADJ
gs-3955	4	31	function	function	NOUN
gs-3955	4	32	over	over	ADP
gs-3955	4	33	extended	extended	ADJ
gs-3955	4	34	lifespans	lifespan	NOUN
gs-3955	4	35	.	.	PUNCT
gs-3955	5	1	notable	notable	ADJ
gs-3955	5	2	examples	example	NOUN
gs-3955	5	3	include	include	VERB
gs-3955	5	4	the	the	DET
gs-3955	5	5	greenland	greenland	PROPN
gs-3955	5	6	shark	shark	NOUN
gs-3955	5	7	(	(	PUNCT
gs-3955	5	8	exceeding	exceed	VERB
gs-3955	5	9	400	400	NUM
gs-3955	5	10	years	year	NOUN
gs-3955	5	11	)	)	PUNCT
gs-3955	5	12	,	,	PUNCT
gs-3955	5	13	naked	naked	ADJ
gs-3955	5	14	mole	mole	NOUN
gs-3955	5	15	-	-	PUNCT
gs-3955	5	16	rat	rat	NOUN
gs-3955	5	17	(	(	PUNCT
gs-3955	5	18	over	over	ADP
gs-3955	5	19	30	30	NUM
gs-3955	5	20	years	year	NOUN
gs-3955	5	21	)	)	PUNCT
gs-3955	5	22	,	,	PUNCT
gs-3955	5	23	ocean	ocean	NOUN
gs-3955	5	24	quahog	quahog	PROPN
gs-3955	5	25	(	(	PUNCT
gs-3955	5	26	507	507	NUM
gs-3955	5	27	years	year	NOUN
gs-3955	5	28	)	)	PUNCT
gs-3955	5	29	,	,	PUNCT
gs-3955	5	30	and	and	CCONJ
gs-3955	5	31	hydra	hydra	PROPN
gs-3955	5	32	(	(	PUNCT
gs-3955	5	33	theoretically	theoretically	ADV
gs-3955	5	34	immortal	immortal	ADJ
gs-3955	5	35	)	)	PUNCT
gs-3955	5	36	.	.	PUNCT
gs-3955	6	1	these	these	DET
gs-3955	6	2	organisms	organism	NOUN
gs-3955	6	3	defy	defy	VERB
gs-3955	6	4	typical	typical	ADJ
gs-3955	6	5	aging	age	VERB
gs-3955	6	6	patterns	pattern	NOUN
gs-3955	6	7	through	through	ADP
gs-3955	6	8	enhanced	enhance	VERB
gs-3955	6	9	dna	dna	PROPN
gs-3955	6	10	repair	repair	NOUN
gs-3955	6	11	,	,	PUNCT
gs-3955	6	12	oxidative	oxidative	ADJ
gs-3955	6	13	stress	stress	NOUN
gs-3955	6	14	resistance	resistance	NOUN
gs-3955	6	15	,	,	PUNCT
gs-3955	6	16	robust	robust	ADJ
gs-3955	6	17	stem	stem	NOUN
gs-3955	6	18	cell	cell	NOUN
gs-3955	6	19	maintenance	maintenance	NOUN
gs-3955	6	20	,	,	PUNCT
gs-3955	6	21	and	and	CCONJ
gs-3955	6	22	unique	unique	ADJ
gs-3955	6	23	proteostasis	proteostasis	NOUN
gs-3955	6	24	and	and	CCONJ
gs-3955	6	25	epigenetic	epigenetic	ADJ
gs-3955	6	26	regulation	regulation	NOUN
gs-3955	6	27	.	.	PUNCT
gs-3955	7	1	comparative	comparative	ADJ
gs-3955	7	2	analysis	analysis	NOUN
gs-3955	7	3	reveals	reveal	VERB
gs-3955	7	4	common	common	ADJ
gs-3955	7	5	adaptations	adaptation	NOUN
gs-3955	7	6	such	such	ADJ
gs-3955	7	7	as	as	ADP
gs-3955	7	8	:	:	PUNCT
gs-3955	7	9	1	1	NUM
gs-3955	7	10	)	)	PUNCT
gs-3955	7	11	late	late	ADJ
gs-3955	7	12	age	age	NOUN
gs-3955	7	13	of	of	ADP
gs-3955	7	14	sexual	sexual	ADJ
gs-3955	7	15	maturation	maturation	NOUN
gs-3955	7	16	;	;	PUNCT
gs-3955	7	17	2	2	X
gs-3955	7	18	)	)	PUNCT
gs-3955	7	19	long	long	ADJ
gs-3955	7	20	-	-	PUNCT
gs-3955	7	21	term	term	NOUN
gs-3955	7	22	maintenance	maintenance	NOUN
gs-3955	7	23	of	of	ADP
gs-3955	7	24	sexual	sexual	ADJ
gs-3955	7	25	activity	activity	NOUN
gs-3955	7	26	;	;	PUNCT
gs-3955	7	27	3	3	X
gs-3955	7	28	)	)	PUNCT
gs-3955	7	29	reduced	reduce	VERB
gs-3955	7	30	metabolic	metabolic	NOUN
gs-3955	7	31	rate	rate	NOUN
gs-3955	7	32	;	;	PUNCT
gs-3955	7	33	4	4	X
gs-3955	7	34	)	)	PUNCT
gs-3955	7	35	tolerance	tolerance	NOUN
gs-3955	7	36	to	to	PART
gs-3955	7	37	hypoxia	hypoxia	VERB
gs-3955	7	38	;	;	PUNCT
gs-3955	7	39	5	5	NUM
gs-3955	7	40	)	)	PUNCT
gs-3955	7	41	neoplastic	neoplastic	ADJ
gs-3955	7	42	suppression	suppression	NOUN
gs-3955	7	43	.	.	PUNCT
gs-3955	8	1	the	the	DET
gs-3955	8	2	molecular	molecular	ADJ
gs-3955	8	3	mechanisms	mechanism	NOUN
gs-3955	8	4	underlying	underlie	VERB
gs-3955	8	5	their	their	PRON
gs-3955	8	6	longevity	longevity	NOUN
gs-3955	8	7	—	—	PUNCT
gs-3955	8	8	including	include	VERB
gs-3955	8	9	high	high	ADJ
gs-3955	8	10	-	-	PUNCT
gs-3955	8	11	molecular	molecular	ADJ
gs-3955	8	12	-	-	PUNCT
gs-3955	8	13	weight	weight	NOUN
gs-3955	8	14	hyaluronan	hyaluronan	NOUN
gs-3955	8	15	in	in	ADP
gs-3955	8	16	naked	naked	ADJ
gs-3955	8	17	mole	mole	NOUN
gs-3955	8	18	-	-	PUNCT
gs-3955	8	19	rats	rat	NOUN
gs-3955	8	20	and	and	CCONJ
gs-3955	8	21	specialized	specialized	ADJ
gs-3955	8	22	lipid	lipid	NOUN
gs-3955	8	23	membranes	membrane	NOUN
gs-3955	8	24	in	in	ADP
gs-3955	8	25	sharks	shark	NOUN
gs-3955	8	26	—	—	PUNCT
gs-3955	8	27	offer	offer	VERB
gs-3955	8	28	transformative	transformative	ADJ
gs-3955	8	29	insights	insight	NOUN
gs-3955	8	30	for	for	ADP
gs-3955	8	31	anti	anti	ADJ
gs-3955	8	32	-	-	ADJ
gs-3955	8	33	aging	aging	ADJ
gs-3955	8	34	interventions	intervention	NOUN
gs-3955	8	35	.	.	PUNCT
gs-3955	9	1	understanding	understand	VERB
gs-3955	9	2	these	these	DET
gs-3955	9	3	strategies	strategy	NOUN
gs-3955	9	4	could	could	AUX
gs-3955	9	5	pave	pave	VERB
gs-3955	9	6	the	the	DET
gs-3955	9	7	way	way	NOUN
gs-3955	9	8	for	for	ADP
gs-3955	9	9	novel	novel	ADJ
gs-3955	9	10	therapies	therapy	NOUN
gs-3955	9	11	targeting	target	VERB
gs-3955	9	12	age	age	NOUN
gs-3955	9	13	-	-	PUNCT
gs-3955	9	14	related	relate	VERB
gs-3955	9	15	diseases	disease	NOUN
gs-3955	9	16	,	,	PUNCT
gs-3955	9	17	potentially	potentially	ADV
gs-3955	9	18	redefining	redefine	VERB
gs-3955	9	19	human	human	ADJ
gs-3955	9	20	healthspan	healthspan	NOUN
gs-3955	9	21	.	.	PUNCT
gs-3955	10	1	this	this	DET
gs-3955	10	2	review	review	NOUN
gs-3955	10	3	synthesizes	synthesize	VERB
gs-3955	10	4	current	current	ADJ
gs-3955	10	5	knowledge	knowledge	NOUN
gs-3955	10	6	on	on	ADP
gs-3955	10	7	negligibly	negligibly	ADV
gs-3955	10	8	senescent	senescent	ADJ
gs-3955	10	9	species	specie	NOUN
gs-3955	10	10	,	,	PUNCT
gs-3955	10	11	highlighting	highlight	VERB
gs-3955	10	12	their	their	PRON
gs-3955	10	13	evolutionary	evolutionary	ADJ
gs-3955	10	14	,	,	PUNCT
gs-3955	10	15	physiological	physiological	ADJ
gs-3955	10	16	,	,	PUNCT
gs-3955	10	17	and	and	CCONJ
gs-3955	10	18	biomedical	biomedical	ADJ
gs-3955	10	19	significance	significance	NOUN
gs-3955	10	20	.	.	PUNCT
gs-3955	11	1	keywords	keyword	NOUN
gs-3955	11	2	:	:	PUNCT
gs-3955	11	3	negligible	negligible	ADJ
gs-3955	11	4	senescence	senescence	NOUN
gs-3955	11	5	,	,	PUNCT
gs-3955	11	6	longevity	longevity	NOUN
gs-3955	11	7	,	,	PUNCT
gs-3955	11	8	oxidative	oxidative	ADJ
gs-3955	11	9	stress	stress	NOUN
gs-3955	11	10	,	,	PUNCT
gs-3955	11	11	proteostasis	proteostasis	NOUN
gs-3955	11	12	,	,	PUNCT
gs-3955	11	13	aging	aging	NOUN
gs-3955	11	14	,	,	PUNCT
gs-3955	11	15	healthspan	healthspan	NOUN
gs-3955	11	16	.	.	PUNCT
gs-3955	12	1	georgian	georgian	ADJ
gs-3955	12	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	12	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	12	4	ტ.	ტ.	VERB
gs-3955	12	5	7	7	NUM
gs-3955	12	6	n	n	PRON
gs-3955	12	7	3	3	NUM
gs-3955	12	8	,	,	PUNCT
gs-3955	12	9	2025	2025	NUM
gs-3955	12	10	347	347	NUM
gs-3955	12	11	introduction	introduction	NOUN
gs-3955	12	12	aging	age	VERB
gs-3955	12	13	is	be	AUX
gs-3955	12	14	a	a	DET
gs-3955	12	15	nearly	nearly	ADV
gs-3955	12	16	universal	universal	ADJ
gs-3955	12	17	biological	biological	ADJ
gs-3955	12	18	process	process	NOUN
gs-3955	12	19	characterized	characterize	VERB
gs-3955	12	20	by	by	ADP
gs-3955	12	21	progressive	progressive	ADJ
gs-3955	12	22	functional	functional	ADJ
gs-3955	12	23	decline	decline	NOUN
gs-3955	12	24	and	and	CCONJ
gs-3955	12	25	increased	increase	VERB
gs-3955	12	26	vulnerability	vulnerability	NOUN
gs-3955	12	27	to	to	ADP
gs-3955	12	28	age	age	NOUN
gs-3955	12	29	-	-	PUNCT
gs-3955	12	30	related	relate	VERB
gs-3955	12	31	diseases	disease	NOUN
gs-3955	12	32	(	(	PUNCT
gs-3955	12	33	lópez	lópez	NOUN
gs-3955	12	34	-	-	PUNCT
gs-3955	12	35	otín	otín	NOUN
gs-3955	12	36	et	et	PROPN
gs-3955	12	37	al	al	PROPN
gs-3955	12	38	.	.	PROPN
gs-3955	12	39	,	,	PUNCT
gs-3955	12	40	2013	2013	NUM
gs-3955	12	41	)	)	PUNCT
gs-3955	12	42	.	.	PUNCT
gs-3955	13	1	however	however	ADV
gs-3955	13	2	,	,	PUNCT
gs-3955	13	3	certain	certain	ADJ
gs-3955	13	4	species	specie	NOUN
gs-3955	13	5	exhibit	exhibit	VERB
gs-3955	13	6	negligible	negligible	ADJ
gs-3955	13	7	senescence	senescence	NOUN
gs-3955	13	8	,	,	PUNCT
gs-3955	13	9	maintaining	maintain	VERB
gs-3955	13	10	physiological	physiological	ADJ
gs-3955	13	11	integrity	integrity	NOUN
gs-3955	13	12	and	and	CCONJ
gs-3955	13	13	reproductive	reproductive	ADJ
gs-3955	13	14	capacity	capacity	NOUN
gs-3955	13	15	over	over	ADP
gs-3955	13	16	exceptionally	exceptionally	ADV
gs-3955	13	17	long	long	ADJ
gs-3955	13	18	lifespans	lifespan	NOUN
gs-3955	13	19	(	(	PUNCT
gs-3955	13	20	finch	finch	NOUN
gs-3955	13	21	,	,	PUNCT
gs-3955	13	22	2009	2009	NUM
gs-3955	13	23	)	)	PUNCT
gs-3955	13	24	.	.	PUNCT
gs-3955	14	1	these	these	DET
gs-3955	14	2	organisms	organism	NOUN
gs-3955	14	3	,	,	PUNCT
gs-3955	14	4	such	such	ADJ
gs-3955	14	5	as	as	ADP
gs-3955	14	6	the	the	DET
gs-3955	14	7	greenland	greenland	PROPN
gs-3955	14	8	shark	shark	NOUN
gs-3955	14	9	(	(	PUNCT
gs-3955	14	10	somniosus	somniosus	PROPN
gs-3955	14	11	microcephalus	microcephalus	PROPN
gs-3955	14	12	)	)	PUNCT
gs-3955	14	13	,	,	PUNCT
gs-3955	14	14	naked	naked	ADJ
gs-3955	14	15	mole	mole	NOUN
gs-3955	14	16	-	-	PUNCT
gs-3955	14	17	rat	rat	NOUN
gs-3955	14	18	(	(	PUNCT
gs-3955	14	19	heterocephalus	heterocephalus	ADJ
gs-3955	14	20	glaber	glaber	NOUN
gs-3955	14	21	)	)	PUNCT
gs-3955	14	22	,	,	PUNCT
gs-3955	14	23	and	and	CCONJ
gs-3955	14	24	ocean	ocean	PROPN
gs-3955	14	25	quahog	quahog	PROPN
gs-3955	14	26	(	(	PUNCT
gs-3955	14	27	arctica	arctica	PROPN
gs-3955	14	28	islandica	islandica	PROPN
gs-3955	14	29	)	)	PUNCT
gs-3955	14	30	,	,	PUNCT
gs-3955	14	31	challenge	challenge	VERB
gs-3955	14	32	conventional	conventional	ADJ
gs-3955	14	33	paradigms	paradigm	NOUN
gs-3955	14	34	of	of	ADP
gs-3955	14	35	aging	aging	NOUN
gs-3955	14	36	and	and	CCONJ
gs-3955	14	37	offer	offer	VERB
gs-3955	14	38	unprecedented	unprecedented	ADJ
gs-3955	14	39	insights	insight	NOUN
gs-3955	14	40	into	into	ADP
gs-3955	14	41	longevity	longevity	NOUN
gs-3955	14	42	mechanisms	mechanism	NOUN
gs-3955	14	43	(	(	PUNCT
gs-3955	14	44	austad	austad	NOUN
gs-3955	14	45	,	,	PUNCT
gs-3955	14	46	2010	2010	NUM
gs-3955	14	47	;	;	PUNCT
gs-3955	14	48	buffenstein	buffenstein	ADV
gs-3955	14	49	,	,	PUNCT
gs-3955	14	50	2008	2008	NUM
gs-3955	14	51	;	;	PUNCT
gs-3955	14	52	nielsen	nielsen	PROPN
gs-3955	14	53	et	et	PROPN
gs-3955	14	54	al	al	PROPN
gs-3955	14	55	.	.	PROPN
gs-3955	14	56	,	,	PUNCT
gs-3955	14	57	2016	2016	NUM
gs-3955	14	58	)	)	PUNCT
gs-3955	14	59	.	.	PUNCT
gs-3955	15	1	the	the	DET
gs-3955	15	2	study	study	NOUN
gs-3955	15	3	of	of	ADP
gs-3955	15	4	negligibly	negligibly	ADV
gs-3955	15	5	senescent	senescent	ADJ
gs-3955	15	6	species	specie	NOUN
gs-3955	15	7	has	have	AUX
gs-3955	15	8	revealed	reveal	VERB
gs-3955	15	9	conserved	conserve	VERB
gs-3955	15	10	molecular	molecular	ADJ
gs-3955	15	11	adaptations	adaptation	NOUN
gs-3955	15	12	,	,	PUNCT
gs-3955	15	13	including	include	VERB
gs-3955	15	14	enhanced	enhance	VERB
gs-3955	15	15	dna	dna	PROPN
gs-3955	15	16	repair	repair	NOUN
gs-3955	15	17	,	,	PUNCT
gs-3955	15	18	oxidative	oxidative	ADJ
gs-3955	15	19	stress	stress	NOUN
gs-3955	15	20	resistance	resistance	NOUN
gs-3955	15	21	,	,	PUNCT
gs-3955	15	22	and	and	CCONJ
gs-3955	15	23	robust	robust	ADJ
gs-3955	15	24	proteostasis	proteostasis	NOUN
gs-3955	15	25	(	(	PUNCT
gs-3955	15	26	tian	tian	PROPN
gs-3955	15	27	et	et	PROPN
gs-3955	15	28	al	al	PROPN
gs-3955	15	29	.	.	PROPN
gs-3955	15	30	,	,	PUNCT
gs-3955	15	31	2013	2013	NUM
gs-3955	15	32	;	;	PUNCT
gs-3955	15	33	munro	munro	PROPN
gs-3955	15	34	&	&	CCONJ
gs-3955	15	35	blier	blier	PROPN
gs-3955	15	36	,	,	PUNCT
gs-3955	15	37	2012	2012	NUM
gs-3955	15	38	)	)	PUNCT
gs-3955	15	39	.	.	PUNCT
gs-3955	16	1	for	for	ADP
gs-3955	16	2	instance	instance	NOUN
gs-3955	16	3	,	,	PUNCT
gs-3955	16	4	naked	naked	ADJ
gs-3955	16	5	mole	mole	NOUN
gs-3955	16	6	-	-	PUNCT
gs-3955	16	7	rats	rat	NOUN
gs-3955	16	8	demonstrate	demonstrate	VERB
gs-3955	16	9	elevated	elevated	ADJ
gs-3955	16	10	dna	dna	PROPN
gs-3955	16	11	repair	repair	NOUN
gs-3955	16	12	activity	activity	NOUN
gs-3955	16	13	mediated	mediate	VERB
gs-3955	16	14	by	by	ADP
gs-3955	16	15	genes	gene	NOUN
gs-3955	16	16	such	such	ADJ
gs-3955	16	17	as	as	ADP
gs-3955	16	18	brca1	brca1	PROPN
gs-3955	16	19	and	and	CCONJ
gs-3955	16	20	tp53	tp53	PROPN
gs-3955	16	21	,	,	PUNCT
gs-3955	16	22	alongside	alongside	ADP
gs-3955	16	23	unique	unique	ADJ
gs-3955	16	24	tumorsuppression	tumorsuppression	NOUN
gs-3955	16	25	mechanisms	mechanism	NOUN
gs-3955	16	26	like	like	ADP
gs-3955	16	27	high	high	ADJ
gs-3955	16	28	-	-	PUNCT
gs-3955	16	29	molecular	molecular	ADJ
gs-3955	16	30	-	-	PUNCT
gs-3955	16	31	weight	weight	NOUN
gs-3955	16	32	hyaluronan	hyaluronan	NOUN
gs-3955	16	33	(	(	PUNCT
gs-3955	16	34	hmw	hmw	PROPN
gs-3955	16	35	-	-	PUNCT
gs-3955	16	36	ha	ha	NOUN
gs-3955	16	37	)	)	PUNCT
gs-3955	16	38	(	(	PUNCT
gs-3955	16	39	gorbunova	gorbunova	PROPN
gs-3955	16	40	et	et	PROPN
gs-3955	16	41	al	al	PROPN
gs-3955	16	42	.	.	PROPN
gs-3955	16	43	,	,	PUNCT
gs-3955	16	44	2014	2014	NUM
gs-3955	16	45	;	;	PUNCT
gs-3955	16	46	petruseva	petruseva	VERB
gs-3955	16	47	et	et	PROPN
gs-3955	16	48	al	al	PROPN
gs-3955	16	49	.	.	PROPN
gs-3955	16	50	,	,	PUNCT
gs-3955	16	51	2017	2017	NUM
gs-3955	16	52	)	)	PUNCT
gs-3955	16	53	.	.	PUNCT
gs-3955	17	1	similarly	similarly	ADV
gs-3955	17	2	,	,	PUNCT
gs-3955	17	3	the	the	DET
gs-3955	17	4	greenland	greenland	PROPN
gs-3955	17	5	shark	shark	NOUN
gs-3955	17	6	’s	’s	PART
gs-3955	17	7	longevity	longevity	NOUN
gs-3955	17	8	is	be	AUX
gs-3955	17	9	linked	link	VERB
gs-3955	17	10	to	to	ADP
gs-3955	17	11	metabolic	metabolic	NOUN
gs-3955	17	12	depression	depression	NOUN
gs-3955	17	13	and	and	CCONJ
gs-3955	17	14	specialized	specialized	ADJ
gs-3955	17	15	lipid	lipid	NOUN
gs-3955	17	16	membranes	membrane	NOUN
gs-3955	17	17	that	that	PRON
gs-3955	17	18	resist	resist	VERB
gs-3955	17	19	peroxidation	peroxidation	NOUN
gs-3955	17	20	(	(	PUNCT
gs-3955	17	21	ballantyne	ballantyne	NOUN
gs-3955	17	22	et	et	PROPN
gs-3955	17	23	al	al	PROPN
gs-3955	17	24	.	.	PROPN
gs-3955	17	25	,	,	PUNCT
gs-3955	17	26	2021	2021	NUM
gs-3955	17	27	;	;	PUNCT
gs-3955	17	28	seim	seim	PROPN
gs-3955	17	29	et	et	PROPN
gs-3955	17	30	al	al	PROPN
gs-3955	17	31	.	.	PROPN
gs-3955	17	32	,	,	PUNCT
gs-3955	17	33	2023	2023	NUM
gs-3955	17	34	)	)	PUNCT
gs-3955	17	35	.	.	PUNCT
gs-3955	18	1	beyond	beyond	ADP
gs-3955	18	2	cellular	cellular	ADJ
gs-3955	18	3	mechanisms	mechanism	NOUN
gs-3955	18	4	,	,	PUNCT
gs-3955	18	5	these	these	DET
gs-3955	18	6	species	specie	NOUN
gs-3955	18	7	exhibit	exhibit	VERB
gs-3955	18	8	systemic	systemic	ADJ
gs-3955	18	9	adaptations	adaptation	NOUN
gs-3955	18	10	,	,	PUNCT
gs-3955	18	11	such	such	ADJ
gs-3955	18	12	as	as	ADP
gs-3955	18	13	hypoxia	hypoxia	NOUN
gs-3955	18	14	tolerance	tolerance	NOUN
gs-3955	18	15	in	in	ADP
gs-3955	18	16	naked	naked	ADJ
gs-3955	18	17	mole	mole	NOUN
gs-3955	18	18	-	-	PUNCT
gs-3955	18	19	rats	rat	NOUN
gs-3955	18	20	(	(	PUNCT
gs-3955	18	21	lewis	lewis	PROPN
gs-3955	18	22	et	et	PROPN
gs-3955	18	23	al	al	PROPN
gs-3955	18	24	.	.	PROPN
gs-3955	18	25	,	,	PUNCT
gs-3955	18	26	2018	2018	NUM
gs-3955	18	27	)	)	PUNCT
gs-3955	18	28	and	and	CCONJ
gs-3955	18	29	epigenetic	epigenetic	ADJ
gs-3955	18	30	stability	stability	NOUN
gs-3955	18	31	in	in	ADP
gs-3955	18	32	rockfish	rockfish	ADJ
gs-3955	18	33	(	(	PUNCT
gs-3955	18	34	sebastes	sebaste	NOUN
gs-3955	18	35	aleutianus	aleutianus	NOUN
gs-3955	18	36	)	)	PUNCT
gs-3955	19	1	(	(	PUNCT
gs-3955	19	2	horvath	horvath	PROPN
gs-3955	19	3	et	et	PROPN
gs-3955	19	4	al	al	PROPN
gs-3955	19	5	.	.	PROPN
gs-3955	19	6	,	,	PUNCT
gs-3955	19	7	2022	2022	NUM
gs-3955	19	8	)	)	PUNCT
gs-3955	19	9	.	.	PUNCT
gs-3955	20	1	the	the	DET
gs-3955	20	2	centrosomal	centrosomal	ADJ
gs-3955	20	3	theory	theory	NOUN
gs-3955	20	4	of	of	ADP
gs-3955	20	5	aging	aging	NOUN
gs-3955	20	6	further	far	ADV
gs-3955	20	7	provides	provide	VERB
gs-3955	20	8	a	a	DET
gs-3955	20	9	unifying	unifying	ADJ
gs-3955	20	10	framework	framework	NOUN
gs-3955	20	11	,	,	PUNCT
gs-3955	20	12	suggesting	suggest	VERB
gs-3955	20	13	that	that	SCONJ
gs-3955	20	14	aging	aging	NOUN
gs-3955	20	15	arises	arise	VERB
gs-3955	20	16	from	from	ADP
gs-3955	20	17	asymmetric	asymmetric	ADJ
gs-3955	20	18	stem	stem	NOUN
gs-3955	20	19	cell	cell	NOUN
gs-3955	20	20	division	division	NOUN
gs-3955	20	21	,	,	PUNCT
gs-3955	20	22	a	a	DET
gs-3955	20	23	process	process	NOUN
gs-3955	20	24	circumvented	circumvent	VERB
gs-3955	20	25	by	by	ADP
gs-3955	20	26	species	specie	NOUN
gs-3955	20	27	like	like	ADP
gs-3955	20	28	planarians	planarian	NOUN
gs-3955	20	29	through	through	ADP
gs-3955	20	30	symmetric	symmetric	ADJ
gs-3955	20	31	neoblast	neoblast	PROPN
gs-3955	20	32	division	division	NOUN
gs-3955	20	33	(	(	PUNCT
gs-3955	20	34	wagner	wagner	PROPN
gs-3955	20	35	et	et	PROPN
gs-3955	20	36	al	al	PROPN
gs-3955	20	37	.	.	PROPN
gs-3955	20	38	,	,	PUNCT
gs-3955	20	39	2011	2011	NUM
gs-3955	20	40	)	)	PUNCT
gs-3955	20	41	.	.	PUNCT
gs-3955	21	1	this	this	DET
gs-3955	21	2	review	review	NOUN
gs-3955	21	3	synthesizes	synthesize	VERB
gs-3955	21	4	current	current	ADJ
gs-3955	21	5	knowledge	knowledge	NOUN
gs-3955	21	6	on	on	ADP
gs-3955	21	7	negligibly	negligibly	ADV
gs-3955	21	8	senescent	senescent	ADJ
gs-3955	21	9	species	specie	NOUN
gs-3955	21	10	,	,	PUNCT
gs-3955	21	11	focusing	focus	VERB
gs-3955	21	12	on	on	ADP
gs-3955	21	13	their	their	PRON
gs-3955	21	14	molecular	molecular	ADJ
gs-3955	21	15	,	,	PUNCT
gs-3955	21	16	physiological	physiological	ADJ
gs-3955	21	17	,	,	PUNCT
gs-3955	21	18	and	and	CCONJ
gs-3955	21	19	evolutionary	evolutionary	ADJ
gs-3955	21	20	adaptations	adaptation	NOUN
gs-3955	21	21	.	.	PUNCT
gs-3955	22	1	by	by	ADP
gs-3955	22	2	analyzing	analyze	VERB
gs-3955	22	3	these	these	DET
gs-3955	22	4	organisms	organism	NOUN
gs-3955	22	5	,	,	PUNCT
gs-3955	22	6	we	we	PRON
gs-3955	22	7	aim	aim	VERB
gs-3955	22	8	to	to	PART
gs-3955	22	9	identify	identify	VERB
gs-3955	22	10	actionable	actionable	ADJ
gs-3955	22	11	pathways	pathway	NOUN
gs-3955	22	12	for	for	ADP
gs-3955	22	13	human	human	PROPN
gs-3955	22	14	healthspan	healthspan	PROPN
gs-3955	22	15	extension	extension	NOUN
gs-3955	22	16	while	while	SCONJ
gs-3955	22	17	addressing	address	VERB
gs-3955	22	18	the	the	DET
gs-3955	22	19	ethical	ethical	ADJ
gs-3955	22	20	and	and	CCONJ
gs-3955	22	21	translational	translational	ADJ
gs-3955	22	22	challenges	challenge	NOUN
gs-3955	22	23	of	of	ADP
gs-3955	22	24	longevity	longevity	NOUN
gs-3955	22	25	research	research	NOUN
gs-3955	22	26	.	.	PUNCT
gs-3955	23	1	while	while	SCONJ
gs-3955	23	2	aging	aging	NOUN
gs-3955	23	3	is	be	AUX
gs-3955	23	4	commonly	commonly	ADV
gs-3955	23	5	accepted	accept	VERB
gs-3955	23	6	as	as	ADP
gs-3955	23	7	a	a	DET
gs-3955	23	8	universal	universal	ADJ
gs-3955	23	9	and	and	CCONJ
gs-3955	23	10	inevitable	inevitable	ADJ
gs-3955	23	11	biological	biological	ADJ
gs-3955	23	12	process	process	NOUN
gs-3955	23	13	,	,	PUNCT
gs-3955	23	14	a	a	DET
gs-3955	23	15	select	select	ADJ
gs-3955	23	16	group	group	NOUN
gs-3955	23	17	of	of	ADP
gs-3955	23	18	organisms	organism	NOUN
gs-3955	23	19	exhibit	exhibit	VERB
gs-3955	23	20	what	what	PRON
gs-3955	23	21	is	be	AUX
gs-3955	23	22	known	know	VERB
gs-3955	23	23	as	as	ADP
gs-3955	23	24	negligible	negligible	ADJ
gs-3955	23	25	senescence	senescence	NOUN
gs-3955	23	26	—	—	PUNCT
gs-3955	23	27	a	a	DET
gs-3955	23	28	condition	condition	NOUN
gs-3955	23	29	in	in	ADP
gs-3955	23	30	which	which	PRON
gs-3955	23	31	aging	age	VERB
gs-3955	23	32	,	,	PUNCT
gs-3955	23	33	as	as	SCONJ
gs-3955	23	34	conventionally	conventionally	ADV
gs-3955	23	35	understood	understand	VERB
gs-3955	23	36	,	,	PUNCT
gs-3955	23	37	appears	appear	VERB
gs-3955	23	38	nearly	nearly	ADV
gs-3955	23	39	absent	absent	ADJ
gs-3955	23	40	.	.	PUNCT
gs-3955	24	1	these	these	DET
gs-3955	24	2	organisms	organism	NOUN
gs-3955	24	3	maintain	maintain	VERB
gs-3955	24	4	stable	stable	ADJ
gs-3955	24	5	physiological	physiological	ADJ
gs-3955	24	6	function	function	NOUN
gs-3955	24	7	,	,	PUNCT
gs-3955	24	8	continuous	continuous	ADJ
gs-3955	24	9	reproductive	reproductive	ADJ
gs-3955	24	10	capacity	capacity	NOUN
gs-3955	24	11	,	,	PUNCT
gs-3955	24	12	and	and	CCONJ
gs-3955	24	13	do	do	AUX
gs-3955	24	14	not	not	PART
gs-3955	24	15	exhibit	exhibit	VERB
gs-3955	24	16	a	a	DET
gs-3955	24	17	detectable	detectable	ADJ
gs-3955	24	18	increase	increase	NOUN
gs-3955	24	19	in	in	ADP
gs-3955	24	20	mortality	mortality	NOUN
gs-3955	24	21	risk	risk	NOUN
gs-3955	24	22	over	over	ADP
gs-3955	24	23	time	time	NOUN
gs-3955	24	24	(	(	PUNCT
gs-3955	24	25	finch	finch	NOUN
gs-3955	24	26	,	,	PUNCT
gs-3955	24	27	2009	2009	NUM
gs-3955	24	28	)	)	PUNCT
gs-3955	24	29	.	.	PUNCT
gs-3955	25	1	negligible	negligible	ADJ
gs-3955	25	2	senescence	senescence	NOUN
gs-3955	25	3	thus	thus	ADV
gs-3955	25	4	represents	represent	VERB
gs-3955	25	5	a	a	DET
gs-3955	25	6	rare	rare	ADJ
gs-3955	25	7	but	but	CCONJ
gs-3955	25	8	highly	highly	ADV
gs-3955	25	9	informative	informative	ADJ
gs-3955	25	10	biological	biological	ADJ
gs-3955	25	11	phenomenon	phenomenon	NOUN
gs-3955	25	12	that	that	PRON
gs-3955	25	13	defies	defy	VERB
gs-3955	25	14	mainstream	mainstream	NOUN
gs-3955	25	15	assumptions	assumption	NOUN
gs-3955	25	16	in	in	ADP
gs-3955	25	17	gerontology	gerontology	NOUN
gs-3955	25	18	and	and	CCONJ
gs-3955	25	19	evolutionary	evolutionary	ADJ
gs-3955	25	20	biology	biology	NOUN
gs-3955	25	21	.	.	PUNCT
gs-3955	26	1	understanding	understand	VERB
gs-3955	26	2	the	the	DET
gs-3955	26	3	mechanisms	mechanism	NOUN
gs-3955	26	4	that	that	PRON
gs-3955	26	5	enable	enable	VERB
gs-3955	26	6	these	these	DET
gs-3955	26	7	species	specie	NOUN
gs-3955	26	8	to	to	PART
gs-3955	26	9	avoid	avoid	VERB
gs-3955	26	10	age	age	NOUN
gs-3955	26	11	-	-	PUNCT
gs-3955	26	12	associated	associate	VERB
gs-3955	26	13	decline	decline	NOUN
gs-3955	26	14	offers	offer	VERB
gs-3955	26	15	unparalleled	unparalleled	ADJ
gs-3955	26	16	insight	insight	NOUN
gs-3955	26	17	into	into	ADP
gs-3955	26	18	the	the	DET
gs-3955	26	19	malleability	malleability	NOUN
gs-3955	26	20	of	of	ADP
gs-3955	26	21	biological	biological	ADJ
gs-3955	26	22	aging	aging	NOUN
gs-3955	26	23	and	and	CCONJ
gs-3955	26	24	opens	open	VERB
gs-3955	26	25	transformative	transformative	ADJ
gs-3955	26	26	avenues	avenue	NOUN
gs-3955	26	27	for	for	ADP
gs-3955	26	28	extending	extend	VERB
gs-3955	26	29	human	human	ADJ
gs-3955	26	30	healthspan	healthspan	NOUN
gs-3955	26	31	and	and	CCONJ
gs-3955	26	32	lifespan	lifespan	ADJ
gs-3955	26	33	.	.	PUNCT
gs-3955	27	1	georgian	georgian	ADJ
gs-3955	27	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	27	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	27	4	ტ.	ტ.	VERB
gs-3955	27	5	7	7	NUM
gs-3955	27	6	n	n	PRON
gs-3955	27	7	3	3	NUM
gs-3955	27	8	,	,	PUNCT
gs-3955	27	9	2025	2025	NUM
gs-3955	27	10	348	348	NUM
gs-3955	27	11	among	among	ADP
gs-3955	27	12	the	the	DET
gs-3955	27	13	most	most	ADV
gs-3955	27	14	well	well	ADV
gs-3955	27	15	-	-	PUNCT
gs-3955	27	16	documented	document	VERB
gs-3955	27	17	negligibly	negligibly	ADV
gs-3955	27	18	senescent	senescent	ADJ
gs-3955	27	19	species	specie	NOUN
gs-3955	27	20	are	be	AUX
gs-3955	27	21	hydra	hydra	PROPN
gs-3955	27	22	vulgaris	vulgaris	NOUN
gs-3955	27	23	,	,	PUNCT
gs-3955	27	24	the	the	DET
gs-3955	27	25	naked	naked	ADJ
gs-3955	27	26	mole	mole	NOUN
gs-3955	27	27	-	-	PUNCT
gs-3955	27	28	rat	rat	NOUN
gs-3955	27	29	(	(	PUNCT
gs-3955	27	30	heterocephalus	heterocephalus	ADJ
gs-3955	27	31	glaber	glaber	NOUN
gs-3955	27	32	)	)	PUNCT
gs-3955	27	33	,	,	PUNCT
gs-3955	27	34	the	the	DET
gs-3955	27	35	greenland	greenland	PROPN
gs-3955	27	36	shark	shark	NOUN
gs-3955	27	37	(	(	PUNCT
gs-3955	27	38	somniosus	somniosus	PROPN
gs-3955	27	39	microcephalus	microcephalus	PROPN
gs-3955	27	40	)	)	PUNCT
gs-3955	27	41	,	,	PUNCT
gs-3955	27	42	the	the	DET
gs-3955	27	43	ocean	ocean	NOUN
gs-3955	27	44	quahog	quahog	PROPN
gs-3955	27	45	(	(	PUNCT
gs-3955	27	46	arctica	arctica	PROPN
gs-3955	27	47	islandica	islandica	PROPN
gs-3955	27	48	)	)	PUNCT
gs-3955	27	49	,	,	PUNCT
gs-3955	27	50	the	the	DET
gs-3955	27	51	planarian	planarian	ADJ
gs-3955	27	52	flatworm	flatworm	NOUN
gs-3955	27	53	(	(	PUNCT
gs-3955	27	54	schmidtea	schmidtea	PROPN
gs-3955	27	55	mediterranea	mediterranea	PROPN
gs-3955	27	56	)	)	PUNCT
gs-3955	27	57	,	,	PUNCT
gs-3955	27	58	and	and	CCONJ
gs-3955	27	59	long	long	ADV
gs-3955	27	60	-	-	PUNCT
gs-3955	27	61	lived	live	VERB
gs-3955	27	62	fish	fish	NOUN
gs-3955	27	63	and	and	CCONJ
gs-3955	27	64	amphibians	amphibian	NOUN
gs-3955	27	65	such	such	ADJ
gs-3955	27	66	as	as	ADP
gs-3955	27	67	the	the	DET
gs-3955	27	68	rougheye	rougheye	NOUN
gs-3955	27	69	rockfish	rockfish	NOUN
gs-3955	27	70	(	(	PUNCT
gs-3955	27	71	sebastes	sebaste	NOUN
gs-3955	27	72	aleutianus	aleutianus	NOUN
gs-3955	27	73	)	)	PUNCT
gs-3955	27	74	and	and	CCONJ
gs-3955	27	75	the	the	DET
gs-3955	27	76	olm	olm	NOUN
gs-3955	27	77	(	(	PUNCT
gs-3955	27	78	proteus	proteus	PROPN
gs-3955	27	79	anguinus	anguinus	PROPN
gs-3955	27	80	)	)	PUNCT
gs-3955	27	81	.	.	PUNCT
gs-3955	28	1	these	these	DET
gs-3955	28	2	organisms	organism	NOUN
gs-3955	28	3	span	span	VERB
gs-3955	28	4	phylogenetically	phylogenetically	ADV
gs-3955	28	5	diverse	diverse	ADJ
gs-3955	28	6	clades	clade	NOUN
gs-3955	28	7	and	and	CCONJ
gs-3955	28	8	inhabit	inhabit	NOUN
gs-3955	28	9	varied	varied	ADJ
gs-3955	28	10	ecological	ecological	ADJ
gs-3955	28	11	niches	niche	NOUN
gs-3955	28	12	—	—	PUNCT
gs-3955	28	13	from	from	ADP
gs-3955	28	14	deep	deep	ADJ
gs-3955	28	15	-	-	PUNCT
gs-3955	28	16	sea	sea	NOUN
gs-3955	28	17	environments	environment	NOUN
gs-3955	28	18	and	and	CCONJ
gs-3955	28	19	subterranean	subterranean	ADJ
gs-3955	28	20	burrows	burrow	NOUN
gs-3955	28	21	to	to	ADP
gs-3955	28	22	anoxic	anoxic	ADJ
gs-3955	28	23	caves	cave	NOUN
gs-3955	28	24	and	and	CCONJ
gs-3955	28	25	freshwater	freshwater	NOUN
gs-3955	28	26	ecosystems	ecosystem	NOUN
gs-3955	28	27	.	.	PUNCT
gs-3955	29	1	despite	despite	SCONJ
gs-3955	29	2	their	their	PRON
gs-3955	29	3	differences	difference	NOUN
gs-3955	29	4	,	,	PUNCT
gs-3955	29	5	they	they	PRON
gs-3955	29	6	share	share	VERB
gs-3955	29	7	a	a	DET
gs-3955	29	8	constellation	constellation	NOUN
gs-3955	29	9	of	of	ADP
gs-3955	29	10	molecular	molecular	ADJ
gs-3955	29	11	and	and	CCONJ
gs-3955	29	12	physiological	physiological	ADJ
gs-3955	29	13	features	feature	NOUN
gs-3955	29	14	that	that	PRON
gs-3955	29	15	converge	converge	VERB
gs-3955	29	16	on	on	ADP
gs-3955	29	17	the	the	DET
gs-3955	29	18	suppression	suppression	NOUN
gs-3955	29	19	of	of	ADP
gs-3955	29	20	the	the	DET
gs-3955	29	21	canonical	canonical	ADJ
gs-3955	29	22	hallmarks	hallmark	NOUN
gs-3955	29	23	of	of	ADP
gs-3955	29	24	aging	age	VERB
gs-3955	29	25	(	(	PUNCT
gs-3955	29	26	lopez	lopez	NOUN
gs-3955	29	27	-	-	PUNCT
gs-3955	29	28	otín	otín	NOUN
gs-3955	29	29	et	et	PROPN
gs-3955	29	30	al	al	PROPN
gs-3955	29	31	.	.	PROPN
gs-3955	29	32	,	,	PUNCT
gs-3955	29	33	2013	2013	NUM
gs-3955	29	34	)	)	PUNCT
gs-3955	29	35	.	.	PUNCT
gs-3955	30	1	hallmarks	hallmark	NOUN
gs-3955	30	2	of	of	ADP
gs-3955	30	3	aging	aging	NOUN
gs-3955	30	4	and	and	CCONJ
gs-3955	30	5	how	how	SCONJ
gs-3955	30	6	these	these	DET
gs-3955	30	7	species	specie	NOUN
gs-3955	30	8	defy	defy	VERB
gs-3955	30	9	them	they	PRON
gs-3955	30	10	the	the	DET
gs-3955	30	11	hallmarks	hallmark	NOUN
gs-3955	30	12	of	of	ADP
gs-3955	30	13	aging	aging	NOUN
gs-3955	30	14	include	include	VERB
gs-3955	30	15	genomic	genomic	NOUN
gs-3955	30	16	instability	instability	NOUN
gs-3955	30	17	,	,	PUNCT
gs-3955	30	18	telomere	telomere	NOUN
gs-3955	30	19	attrition	attrition	NOUN
gs-3955	30	20	,	,	PUNCT
gs-3955	30	21	epigenetic	epigenetic	ADJ
gs-3955	30	22	alterations	alteration	NOUN
gs-3955	30	23	,	,	PUNCT
gs-3955	30	24	loss	loss	NOUN
gs-3955	30	25	of	of	ADP
gs-3955	30	26	proteostasis	proteostasis	NOUN
gs-3955	30	27	,	,	PUNCT
gs-3955	30	28	deregulated	deregulate	VERB
gs-3955	30	29	nutrient	nutrient	NOUN
gs-3955	30	30	sensing	sensing	NOUN
gs-3955	30	31	,	,	PUNCT
gs-3955	30	32	mitochondrial	mitochondrial	ADJ
gs-3955	30	33	dysfunction	dysfunction	NOUN
gs-3955	30	34	,	,	PUNCT
gs-3955	30	35	cellular	cellular	ADJ
gs-3955	30	36	senescence	senescence	NOUN
gs-3955	30	37	,	,	PUNCT
gs-3955	30	38	stem	stem	NOUN
gs-3955	30	39	cell	cell	NOUN
gs-3955	30	40	exhaustion	exhaustion	NOUN
gs-3955	30	41	,	,	PUNCT
gs-3955	30	42	and	and	CCONJ
gs-3955	30	43	altered	alter	VERB
gs-3955	30	44	intercellular	intercellular	ADJ
gs-3955	30	45	communication	communication	NOUN
gs-3955	30	46	.	.	PUNCT
gs-3955	31	1	negligibly	negligibly	ADV
gs-3955	31	2	senescent	senescent	ADJ
gs-3955	31	3	species	specie	NOUN
gs-3955	31	4	exhibit	exhibit	VERB
gs-3955	31	5	evolved	evolve	VERB
gs-3955	31	6	strategies	strategy	NOUN
gs-3955	31	7	to	to	PART
gs-3955	31	8	mitigate	mitigate	VERB
gs-3955	31	9	or	or	CCONJ
gs-3955	31	10	entirely	entirely	ADV
gs-3955	31	11	avoid	avoid	VERB
gs-3955	31	12	these	these	DET
gs-3955	31	13	hallmarks	hallmark	NOUN
gs-3955	31	14	(	(	PUNCT
gs-3955	31	15	lopez	lopez	NOUN
gs-3955	31	16	-	-	PUNCT
gs-3955	31	17	otín	otín	NOUN
gs-3955	31	18	et	et	PROPN
gs-3955	31	19	al	al	PROPN
gs-3955	31	20	.	.	PROPN
gs-3955	31	21	,	,	PUNCT
gs-3955	31	22	2013	2013	NUM
gs-3955	31	23	;	;	PUNCT
gs-3955	31	24	petruseva	petruseva	VERB
gs-3955	31	25	et	et	PROPN
gs-3955	31	26	al	al	PROPN
gs-3955	31	27	.	.	PROPN
gs-3955	31	28	,	,	PUNCT
gs-3955	31	29	2017	2017	NUM
gs-3955	31	30	)	)	PUNCT
gs-3955	31	31	.	.	PUNCT
gs-3955	32	1	for	for	ADP
gs-3955	32	2	example	example	NOUN
gs-3955	32	3	,	,	PUNCT
gs-3955	32	4	the	the	DET
gs-3955	32	5	ocean	ocean	NOUN
gs-3955	32	6	quahog	quahog	PROPN
gs-3955	32	7	(	(	PUNCT
gs-3955	32	8	arctica	arctica	PROPN
gs-3955	32	9	islandica	islandica	PROPN
gs-3955	32	10	)	)	PUNCT
gs-3955	32	11	,	,	PUNCT
gs-3955	32	12	with	with	ADP
gs-3955	32	13	a	a	DET
gs-3955	32	14	documented	document	VERB
gs-3955	32	15	lifespan	lifespan	NOUN
gs-3955	32	16	exceeding	exceed	VERB
gs-3955	32	17	500	500	NUM
gs-3955	32	18	years	year	NOUN
gs-3955	32	19	,	,	PUNCT
gs-3955	32	20	displays	display	VERB
gs-3955	32	21	extraordinary	extraordinary	ADJ
gs-3955	32	22	dna	dna	PROPN
gs-3955	32	23	repair	repair	NOUN
gs-3955	32	24	capacity	capacity	NOUN
gs-3955	32	25	,	,	PUNCT
gs-3955	32	26	particularly	particularly	ADV
gs-3955	32	27	in	in	ADP
gs-3955	32	28	nucleotide	nucleotide	ADJ
gs-3955	32	29	excision	excision	NOUN
gs-3955	32	30	repair	repair	NOUN
gs-3955	32	31	(	(	PUNCT
gs-3955	32	32	ner	ner	NOUN
gs-3955	32	33	)	)	PUNCT
gs-3955	32	34	pathways	pathway	NOUN
gs-3955	32	35	(	(	PUNCT
gs-3955	32	36	munro	munro	PROPN
gs-3955	32	37	et	et	PROPN
gs-3955	32	38	al	al	PROPN
gs-3955	32	39	.	.	PROPN
gs-3955	32	40	,	,	PUNCT
gs-3955	32	41	2012	2012	NUM
gs-3955	32	42	)	)	PUNCT
gs-3955	32	43	.	.	PUNCT
gs-3955	33	1	its	its	PRON
gs-3955	33	2	variants	variant	NOUN
gs-3955	33	3	of	of	ADP
gs-3955	33	4	xpa	xpa	PROPN
gs-3955	33	5	and	and	CCONJ
gs-3955	33	6	ercc1	ercc1	NOUN
gs-3955	33	7	proteins	protein	NOUN
gs-3955	33	8	demonstrate	demonstrate	VERB
gs-3955	33	9	higher	high	ADJ
gs-3955	33	10	affinity	affinity	NOUN
gs-3955	33	11	and	and	CCONJ
gs-3955	33	12	stability	stability	NOUN
gs-3955	33	13	in	in	ADP
gs-3955	33	14	recognizing	recognize	VERB
gs-3955	33	15	and	and	CCONJ
gs-3955	33	16	excising	excise	VERB
gs-3955	33	17	uv	uv	NOUN
gs-3955	33	18	-	-	PUNCT
gs-3955	33	19	induced	induce	VERB
gs-3955	33	20	dna	dna	NOUN
gs-3955	33	21	lesions	lesion	NOUN
gs-3955	33	22	.	.	PUNCT
gs-3955	34	1	similarly	similarly	ADV
gs-3955	34	2	,	,	PUNCT
gs-3955	34	3	the	the	DET
gs-3955	34	4	greenland	greenland	PROPN
gs-3955	34	5	shark	shark	NOUN
gs-3955	34	6	shows	show	VERB
gs-3955	34	7	genomic	genomic	ADJ
gs-3955	34	8	duplications	duplication	NOUN
gs-3955	34	9	of	of	ADP
gs-3955	34	10	dna	dna	PROPN
gs-3955	34	11	repair	repair	NOUN
gs-3955	34	12	regulators	regulator	NOUN
gs-3955	34	13	such	such	ADJ
gs-3955	34	14	as	as	ADP
gs-3955	34	15	atm	atm	NOUN
gs-3955	34	16	and	and	CCONJ
gs-3955	34	17	atr	atr	NOUN
gs-3955	34	18	,	,	PUNCT
gs-3955	34	19	creating	create	VERB
gs-3955	34	20	redundancy	redundancy	NOUN
gs-3955	34	21	in	in	ADP
gs-3955	34	22	genomic	genomic	ADJ
gs-3955	34	23	surveillance	surveillance	NOUN
gs-3955	34	24	mechanisms	mechanism	NOUN
gs-3955	34	25	(	(	PUNCT
gs-3955	34	26	polinski	polinski	NOUN
gs-3955	34	27	et	et	PROPN
gs-3955	34	28	al	al	PROPN
gs-3955	34	29	.	.	PROPN
gs-3955	34	30	,	,	PUNCT
gs-3955	34	31	2021	2021	NUM
gs-3955	34	32	)	)	PUNCT
gs-3955	34	33	.	.	PUNCT
gs-3955	35	1	hydra	hydra	PROPN
gs-3955	35	2	maintains	maintain	VERB
gs-3955	35	3	regenerative	regenerative	ADJ
gs-3955	35	4	immortality	immortality	NOUN
gs-3955	35	5	through	through	ADP
gs-3955	35	6	continuous	continuous	ADJ
gs-3955	35	7	stem	stem	NOUN
gs-3955	35	8	cell	cell	NOUN
gs-3955	35	9	renewal	renewal	NOUN
gs-3955	35	10	.	.	PUNCT
gs-3955	36	1	its	its	PRON
gs-3955	36	2	foxo	foxo	ADJ
gs-3955	36	3	transcription	transcription	NOUN
gs-3955	36	4	factor	factor	NOUN
gs-3955	36	5	—	—	PUNCT
gs-3955	36	6	highly	highly	ADV
gs-3955	36	7	expressed	express	VERB
gs-3955	36	8	in	in	ADP
gs-3955	36	9	all	all	DET
gs-3955	36	10	stem	stem	NOUN
gs-3955	36	11	cell	cell	NOUN
gs-3955	36	12	lineages	lineage	NOUN
gs-3955	36	13	—	—	PUNCT
gs-3955	36	14	plays	play	VERB
gs-3955	36	15	a	a	DET
gs-3955	36	16	key	key	ADJ
gs-3955	36	17	role	role	NOUN
gs-3955	36	18	in	in	ADP
gs-3955	36	19	sustaining	sustain	VERB
gs-3955	36	20	self	self	NOUN
gs-3955	36	21	-	-	PUNCT
gs-3955	36	22	renewal	renewal	NOUN
gs-3955	36	23	and	and	CCONJ
gs-3955	36	24	controlling	control	VERB
gs-3955	36	25	apoptosis	apoptosis	NOUN
gs-3955	36	26	(	(	PUNCT
gs-3955	36	27	boehm	boehm	PROPN
gs-3955	36	28	et	et	PROPN
gs-3955	36	29	al	al	PROPN
gs-3955	36	30	.	.	PROPN
gs-3955	36	31	,	,	PUNCT
gs-3955	36	32	2012	2012	NUM
gs-3955	36	33	)	)	PUNCT
gs-3955	36	34	.	.	PUNCT
gs-3955	37	1	unlike	unlike	ADP
gs-3955	37	2	mammals	mammal	NOUN
gs-3955	37	3	,	,	PUNCT
gs-3955	37	4	hydra	hydra	PROPN
gs-3955	37	5	does	do	AUX
gs-3955	37	6	not	not	PART
gs-3955	37	7	exhibit	exhibit	VERB
gs-3955	37	8	telomere	telomere	NOUN
gs-3955	37	9	shortening	shorten	VERB
gs-3955	37	10	with	with	ADP
gs-3955	37	11	age	age	NOUN
gs-3955	37	12	,	,	PUNCT
gs-3955	37	13	thanks	thank	NOUN
gs-3955	37	14	in	in	ADP
gs-3955	37	15	part	part	NOUN
gs-3955	37	16	to	to	ADP
gs-3955	37	17	constitutive	constitutive	ADJ
gs-3955	37	18	telomerase	telomerase	NOUN
gs-3955	37	19	activity	activity	NOUN
gs-3955	37	20	.	.	PUNCT
gs-3955	38	1	stem	stem	NOUN
gs-3955	38	2	cell	cell	NOUN
gs-3955	38	3	dynamics	dynamic	NOUN
gs-3955	38	4	and	and	CCONJ
gs-3955	38	5	tissue	tissue	NOUN
gs-3955	38	6	homeostasis	homeostasis	NOUN
gs-3955	38	7	negligibly	negligibly	ADV
gs-3955	38	8	senescent	senescent	ADJ
gs-3955	38	9	organisms	organism	NOUN
gs-3955	38	10	maintain	maintain	VERB
gs-3955	38	11	tissue	tissue	NOUN
gs-3955	38	12	homeostasis	homeostasis	NOUN
gs-3955	38	13	through	through	ADP
gs-3955	38	14	either	either	CCONJ
gs-3955	38	15	constant	constant	ADJ
gs-3955	38	16	regenerative	regenerative	ADJ
gs-3955	38	17	renewal	renewal	NOUN
gs-3955	38	18	or	or	CCONJ
gs-3955	38	19	unusually	unusually	ADV
gs-3955	38	20	efficient	efficient	ADJ
gs-3955	38	21	damage	damage	NOUN
gs-3955	38	22	mitigation	mitigation	NOUN
gs-3955	38	23	.	.	PUNCT
gs-3955	39	1	planarian	planarian	ADJ
gs-3955	39	2	flatworms	flatworm	NOUN
gs-3955	39	3	are	be	AUX
gs-3955	39	4	capable	capable	ADJ
gs-3955	39	5	of	of	ADP
gs-3955	39	6	whole	whole	ADJ
gs-3955	39	7	-	-	PUNCT
gs-3955	39	8	body	body	NOUN
gs-3955	39	9	regeneration	regeneration	NOUN
gs-3955	39	10	due	due	ADP
gs-3955	39	11	to	to	ADP
gs-3955	39	12	a	a	DET
gs-3955	39	13	population	population	NOUN
gs-3955	39	14	of	of	ADP
gs-3955	39	15	pluripotent	pluripotent	ADJ
gs-3955	39	16	adult	adult	NOUN
gs-3955	39	17	stem	stem	NOUN
gs-3955	39	18	cells	cell	NOUN
gs-3955	39	19	called	call	VERB
gs-3955	39	20	neoblasts	neoblast	NOUN
gs-3955	39	21	(	(	PUNCT
gs-3955	39	22	reddien	reddien	PROPN
gs-3955	39	23	,	,	PUNCT
gs-3955	39	24	2013	2013	NUM
gs-3955	39	25	)	)	PUNCT
gs-3955	39	26	.	.	PUNCT
gs-3955	40	1	these	these	DET
gs-3955	40	2	cells	cell	NOUN
gs-3955	40	3	divide	divide	VERB
gs-3955	40	4	symmetrically	symmetrically	ADV
gs-3955	40	5	,	,	PUNCT
gs-3955	40	6	escaping	escape	VERB
gs-3955	40	7	the	the	DET
gs-3955	40	8	asymmetric	asymmetric	ADJ
gs-3955	40	9	division	division	NOUN
gs-3955	40	10	postulated	postulate	VERB
gs-3955	40	11	in	in	ADP
gs-3955	40	12	the	the	DET
gs-3955	40	13	centrosomal	centrosomal	ADJ
gs-3955	40	14	theory	theory	NOUN
gs-3955	40	15	of	of	ADP
gs-3955	40	16	aging	aging	NOUN
gs-3955	40	17	as	as	ADP
gs-3955	40	18	a	a	DET
gs-3955	40	19	key	key	ADJ
gs-3955	40	20	contributor	contributor	NOUN
gs-3955	40	21	to	to	ADP
gs-3955	40	22	cellular	cellular	ADJ
gs-3955	40	23	senescence	senescence	NOUN
gs-3955	40	24	(	(	PUNCT
gs-3955	40	25	ratajczak	ratajczak	NOUN
gs-3955	40	26	et	et	PROPN
gs-3955	40	27	al	al	PROPN
gs-3955	40	28	.	.	PROPN
gs-3955	40	29	,	,	PUNCT
gs-3955	40	30	2018	2018	NUM
gs-3955	40	31	)	)	PUNCT
gs-3955	40	32	.	.	PUNCT
gs-3955	41	1	the	the	DET
gs-3955	41	2	naked	naked	ADJ
gs-3955	41	3	mole	mole	NOUN
gs-3955	41	4	-	-	PUNCT
gs-3955	41	5	rat	rat	NOUN
gs-3955	41	6	maintains	maintain	VERB
gs-3955	41	7	stem	stem	NOUN
gs-3955	41	8	cell	cell	NOUN
gs-3955	41	9	pools	pool	NOUN
gs-3955	41	10	even	even	ADV
gs-3955	41	11	into	into	ADP
gs-3955	41	12	advanced	advanced	ADJ
gs-3955	41	13	age	age	NOUN
gs-3955	41	14	,	,	PUNCT
gs-3955	41	15	showing	show	VERB
gs-3955	41	16	no	no	DET
gs-3955	41	17	decline	decline	NOUN
gs-3955	41	18	in	in	ADP
gs-3955	41	19	hematopoietic	hematopoietic	ADJ
gs-3955	41	20	or	or	CCONJ
gs-3955	41	21	neural	neural	ADJ
gs-3955	41	22	stem	stem	NOUN
gs-3955	41	23	cell	cell	NOUN
gs-3955	41	24	function	function	NOUN
gs-3955	41	25	.	.	PUNCT
gs-3955	42	1	these	these	DET
gs-3955	42	2	cells	cell	NOUN
gs-3955	42	3	exhibit	exhibit	VERB
gs-3955	42	4	low	low	ADJ
gs-3955	42	5	levels	level	NOUN
gs-3955	42	6	of	of	ADP
gs-3955	42	7	reactive	reactive	ADJ
gs-3955	42	8	georgian	georgian	ADJ
gs-3955	42	9	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	42	10	მეცნიერები	მეცნიერები	NOUN
gs-3955	42	11	ტ.	ტ.	VERB
gs-3955	42	12	7	7	NUM
gs-3955	42	13	n	n	PRON
gs-3955	42	14	3	3	NUM
gs-3955	42	15	,	,	PUNCT
gs-3955	42	16	2025	2025	NUM
gs-3955	42	17	349	349	NUM
gs-3955	42	18	oxygen	oxygen	NOUN
gs-3955	42	19	species	specie	NOUN
gs-3955	42	20	(	(	PUNCT
gs-3955	42	21	ros	ros	PROPN
gs-3955	42	22	)	)	PUNCT
gs-3955	42	23	and	and	CCONJ
gs-3955	42	24	upregulation	upregulation	NOUN
gs-3955	42	25	of	of	ADP
gs-3955	42	26	nrf2	nrf2	NOUN
gs-3955	42	27	-	-	PUNCT
gs-3955	42	28	dependent	dependent	ADJ
gs-3955	42	29	antioxidant	antioxidant	NOUN
gs-3955	42	30	defenses	defense	NOUN
gs-3955	42	31	(	(	PUNCT
gs-3955	42	32	lewis	lewis	PROPN
gs-3955	42	33	et	et	PROPN
gs-3955	42	34	al	al	PROPN
gs-3955	42	35	.	.	PROPN
gs-3955	42	36	,	,	PUNCT
gs-3955	42	37	2012	2012	NUM
gs-3955	42	38	)	)	PUNCT
gs-3955	42	39	.	.	PUNCT
gs-3955	43	1	such	such	ADJ
gs-3955	43	2	mechanisms	mechanism	NOUN
gs-3955	43	3	are	be	AUX
gs-3955	43	4	critical	critical	ADJ
gs-3955	43	5	to	to	ADP
gs-3955	43	6	long	long	ADJ
gs-3955	43	7	-	-	PUNCT
gs-3955	43	8	term	term	NOUN
gs-3955	43	9	regenerative	regenerative	ADJ
gs-3955	43	10	capacity	capacity	NOUN
gs-3955	43	11	and	and	CCONJ
gs-3955	43	12	avoidance	avoidance	NOUN
gs-3955	43	13	of	of	ADP
gs-3955	43	14	stem	stem	NOUN
gs-3955	43	15	cell	cell	NOUN
gs-3955	43	16	exhaustion	exhaustion	NOUN
gs-3955	43	17	.	.	PUNCT
gs-3955	44	1	oxidative	oxidative	ADJ
gs-3955	44	2	stress	stress	NOUN
gs-3955	44	3	resistance	resistance	NOUN
gs-3955	44	4	and	and	CCONJ
gs-3955	44	5	redox	redox	NOUN
gs-3955	44	6	homeostasis	homeostasis	NOUN
gs-3955	44	7	contrary	contrary	ADJ
gs-3955	44	8	to	to	ADP
gs-3955	44	9	the	the	DET
gs-3955	44	10	oxidative	oxidative	ADJ
gs-3955	44	11	stress	stress	NOUN
gs-3955	44	12	theory	theory	NOUN
gs-3955	44	13	of	of	ADP
gs-3955	44	14	aging	aging	NOUN
gs-3955	44	15	,	,	PUNCT
gs-3955	44	16	some	some	DET
gs-3955	44	17	long	long	ADV
gs-3955	44	18	-	-	PUNCT
gs-3955	44	19	lived	live	VERB
gs-3955	44	20	species	specie	NOUN
gs-3955	44	21	produce	produce	VERB
gs-3955	44	22	high	high	ADJ
gs-3955	44	23	ros	ros	PROPN
gs-3955	44	24	levels	level	NOUN
gs-3955	44	25	but	but	CCONJ
gs-3955	44	26	suffer	suffer	VERB
gs-3955	44	27	little	little	ADJ
gs-3955	44	28	molecular	molecular	ADJ
gs-3955	44	29	damage	damage	NOUN
gs-3955	44	30	.	.	PUNCT
gs-3955	45	1	naked	naked	ADJ
gs-3955	45	2	mole	mole	NOUN
gs-3955	45	3	-	-	PUNCT
gs-3955	45	4	rats	rat	NOUN
gs-3955	45	5	,	,	PUNCT
gs-3955	45	6	for	for	ADP
gs-3955	45	7	example	example	NOUN
gs-3955	45	8	,	,	PUNCT
gs-3955	45	9	display	display	VERB
gs-3955	45	10	elevated	elevated	ADJ
gs-3955	45	11	mitochondrial	mitochondrial	ADJ
gs-3955	45	12	ros	ro	NOUN
gs-3955	45	13	but	but	CCONJ
gs-3955	45	14	maintain	maintain	VERB
gs-3955	45	15	protein	protein	NOUN
gs-3955	45	16	integrity	integrity	NOUN
gs-3955	45	17	and	and	CCONJ
gs-3955	45	18	lipid	lipid	NOUN
gs-3955	45	19	stability	stability	NOUN
gs-3955	45	20	due	due	ADP
gs-3955	45	21	to	to	ADP
gs-3955	45	22	upregulated	upregulated	ADJ
gs-3955	45	23	proteasomal	proteasomal	ADJ
gs-3955	45	24	activity	activity	NOUN
gs-3955	45	25	and	and	CCONJ
gs-3955	45	26	enhanced	enhance	VERB
gs-3955	45	27	chaperone	chaperone	NOUN
gs-3955	45	28	responses	response	NOUN
gs-3955	45	29	(	(	PUNCT
gs-3955	45	30	perez	perez	PROPN
gs-3955	45	31	et	et	PROPN
gs-3955	45	32	al	al	PROPN
gs-3955	45	33	.	.	PROPN
gs-3955	45	34	,	,	PUNCT
gs-3955	45	35	2009	2009	NUM
gs-3955	45	36	)	)	PUNCT
gs-3955	45	37	.	.	PUNCT
gs-3955	46	1	the	the	DET
gs-3955	46	2	ocean	ocean	PROPN
gs-3955	46	3	quahog	quahog	PROPN
gs-3955	46	4	’s	’s	PART
gs-3955	46	5	mitochondria	mitochondria	PROPN
gs-3955	46	6	are	be	AUX
gs-3955	46	7	highly	highly	ADV
gs-3955	46	8	resistant	resistant	ADJ
gs-3955	46	9	to	to	ADP
gs-3955	46	10	lipid	lipid	NOUN
gs-3955	46	11	peroxidation	peroxidation	NOUN
gs-3955	46	12	due	due	ADP
gs-3955	46	13	to	to	ADP
gs-3955	46	14	a	a	DET
gs-3955	46	15	fatty	fatty	NOUN
gs-3955	46	16	acid	acid	NOUN
gs-3955	46	17	profile	profile	NOUN
gs-3955	46	18	rich	rich	ADJ
gs-3955	46	19	in	in	ADP
gs-3955	46	20	monounsaturated	monounsaturated	ADJ
gs-3955	46	21	and	and	CCONJ
gs-3955	46	22	saturated	saturate	VERB
gs-3955	46	23	fats	fat	NOUN
gs-3955	46	24	,	,	PUNCT
gs-3955	46	25	resulting	result	VERB
gs-3955	46	26	in	in	ADP
gs-3955	46	27	a	a	DET
gs-3955	46	28	low	low	ADJ
gs-3955	46	29	peroxidation	peroxidation	NOUN
gs-3955	46	30	index	index	NOUN
gs-3955	46	31	(	(	PUNCT
gs-3955	46	32	munro	munro	PROPN
gs-3955	46	33	et	et	PROPN
gs-3955	46	34	al	al	PROPN
gs-3955	46	35	.	.	PROPN
gs-3955	46	36	,	,	PUNCT
gs-3955	46	37	2012	2012	NUM
gs-3955	46	38	)	)	PUNCT
gs-3955	46	39	.	.	PUNCT
gs-3955	47	1	this	this	DET
gs-3955	47	2	paradoxical	paradoxical	ADJ
gs-3955	47	3	tolerance	tolerance	NOUN
gs-3955	47	4	of	of	ADP
gs-3955	47	5	oxidative	oxidative	ADJ
gs-3955	47	6	by	by	ADP
gs-3955	47	7	-	-	PUNCT
gs-3955	47	8	products	product	NOUN
gs-3955	47	9	is	be	AUX
gs-3955	47	10	accompanied	accompany	VERB
gs-3955	47	11	by	by	ADP
gs-3955	47	12	robust	robust	ADJ
gs-3955	47	13	antioxidant	antioxidant	ADJ
gs-3955	47	14	systems	system	NOUN
gs-3955	47	15	,	,	PUNCT
gs-3955	47	16	including	include	VERB
gs-3955	47	17	increased	increase	VERB
gs-3955	47	18	catalase	catalase	NOUN
gs-3955	47	19	and	and	CCONJ
gs-3955	47	20	glutathione	glutathione	NOUN
gs-3955	47	21	peroxidase	peroxidase	NOUN
gs-3955	47	22	activity	activity	NOUN
gs-3955	47	23	,	,	PUNCT
gs-3955	47	24	contributing	contribute	VERB
gs-3955	47	25	to	to	ADP
gs-3955	47	26	a	a	DET
gs-3955	47	27	decoupling	decoupling	NOUN
gs-3955	47	28	of	of	ADP
gs-3955	47	29	ros	ros	PROPN
gs-3955	47	30	levels	level	NOUN
gs-3955	47	31	from	from	ADP
gs-3955	47	32	cellular	cellular	ADJ
gs-3955	47	33	damage	damage	NOUN
gs-3955	47	34	.	.	PUNCT
gs-3955	48	1	proteostasis	proteostasis	NOUN
gs-3955	48	2	and	and	CCONJ
gs-3955	48	3	autophagy	autophagy	ADJ
gs-3955	48	4	protein	protein	NOUN
gs-3955	48	5	homeostasis	homeostasis	NOUN
gs-3955	48	6	,	,	PUNCT
gs-3955	48	7	or	or	CCONJ
gs-3955	48	8	proteostasis	proteostasis	NOUN
gs-3955	48	9	,	,	PUNCT
gs-3955	48	10	is	be	AUX
gs-3955	48	11	crucial	crucial	ADJ
gs-3955	48	12	for	for	ADP
gs-3955	48	13	long	long	ADJ
gs-3955	48	14	-	-	PUNCT
gs-3955	48	15	term	term	NOUN
gs-3955	48	16	cellular	cellular	ADJ
gs-3955	48	17	function	function	NOUN
gs-3955	48	18	.	.	PUNCT
gs-3955	49	1	misfolded	misfolde	VERB
gs-3955	49	2	proteins	protein	NOUN
gs-3955	49	3	and	and	CCONJ
gs-3955	49	4	aggregates	aggregate	NOUN
gs-3955	49	5	accumulate	accumulate	VERB
gs-3955	49	6	in	in	ADP
gs-3955	49	7	aging	age	VERB
gs-3955	49	8	cells	cell	NOUN
gs-3955	49	9	,	,	PUNCT
gs-3955	49	10	contributing	contribute	VERB
gs-3955	49	11	to	to	ADP
gs-3955	49	12	neurodegeneration	neurodegeneration	NOUN
gs-3955	49	13	and	and	CCONJ
gs-3955	49	14	other	other	ADJ
gs-3955	49	15	age	age	NOUN
gs-3955	49	16	-	-	PUNCT
gs-3955	49	17	related	relate	VERB
gs-3955	49	18	pathologies	pathology	NOUN
gs-3955	49	19	.	.	PUNCT
gs-3955	50	1	negligibly	negligibly	ADV
gs-3955	50	2	senescent	senescent	ADJ
gs-3955	50	3	species	specie	NOUN
gs-3955	50	4	combat	combat	VERB
gs-3955	50	5	this	this	PRON
gs-3955	50	6	through	through	ADP
gs-3955	50	7	a	a	DET
gs-3955	50	8	combination	combination	NOUN
gs-3955	50	9	of	of	ADP
gs-3955	50	10	enhanced	enhanced	ADJ
gs-3955	50	11	heat	heat	NOUN
gs-3955	50	12	shock	shock	NOUN
gs-3955	50	13	response	response	NOUN
gs-3955	50	14	(	(	PUNCT
gs-3955	50	15	hsr	hsr	NOUN
gs-3955	50	16	)	)	PUNCT
gs-3955	50	17	,	,	PUNCT
gs-3955	50	18	ubiquitin	ubiquitin	ADJ
gs-3955	50	19	-	-	PUNCT
gs-3955	50	20	proteasome	proteasome	NOUN
gs-3955	50	21	system	system	NOUN
gs-3955	50	22	(	(	PUNCT
gs-3955	50	23	ups	up	NOUN
gs-3955	50	24	)	)	PUNCT
gs-3955	50	25	,	,	PUNCT
gs-3955	50	26	and	and	CCONJ
gs-3955	50	27	autophagy	autophagy	NOUN
gs-3955	50	28	(	(	PUNCT
gs-3955	50	29	kaushik	kaushik	PROPN
gs-3955	50	30	&	&	CCONJ
gs-3955	50	31	cuervo	cuervo	PROPN
gs-3955	50	32	,	,	PUNCT
gs-3955	50	33	2015	2015	NUM
gs-3955	50	34	)	)	PUNCT
gs-3955	50	35	.	.	PUNCT
gs-3955	51	1	naked	naked	ADJ
gs-3955	51	2	mole	mole	NOUN
gs-3955	51	3	-	-	PUNCT
gs-3955	51	4	rats	rat	NOUN
gs-3955	51	5	demonstrate	demonstrate	VERB
gs-3955	51	6	elevated	elevated	ADJ
gs-3955	51	7	baseline	baseline	ADJ
gs-3955	51	8	autophagic	autophagic	ADJ
gs-3955	51	9	flux	flux	NOUN
gs-3955	51	10	and	and	CCONJ
gs-3955	51	11	efficient	efficient	ADJ
gs-3955	51	12	removal	removal	NOUN
gs-3955	51	13	of	of	ADP
gs-3955	51	14	damaged	damage	VERB
gs-3955	51	15	organelles	organelle	NOUN
gs-3955	51	16	,	,	PUNCT
gs-3955	51	17	especially	especially	ADV
gs-3955	51	18	under	under	ADP
gs-3955	51	19	metabolic	metabolic	NOUN
gs-3955	51	20	stress	stress	NOUN
gs-3955	51	21	(	(	PUNCT
gs-3955	51	22	pride	pride	NOUN
gs-3955	51	23	et	et	PROPN
gs-3955	51	24	al	al	PROPN
gs-3955	51	25	.	.	PROPN
gs-3955	51	26	,	,	PUNCT
gs-3955	51	27	2015	2015	NUM
gs-3955	51	28	)	)	PUNCT
gs-3955	51	29	.	.	PUNCT
gs-3955	52	1	planarians	planarian	NOUN
gs-3955	52	2	rely	rely	VERB
gs-3955	52	3	on	on	ADP
gs-3955	52	4	autophagy	autophagy	NOUN
gs-3955	52	5	to	to	PART
gs-3955	52	6	remodel	remodel	VERB
gs-3955	52	7	tissues	tissue	NOUN
gs-3955	52	8	during	during	ADP
gs-3955	52	9	regeneration	regeneration	NOUN
gs-3955	52	10	and	and	CCONJ
gs-3955	52	11	to	to	PART
gs-3955	52	12	remove	remove	VERB
gs-3955	52	13	senescent	senescent	ADJ
gs-3955	52	14	cell	cell	NOUN
gs-3955	52	15	debris	debris	NOUN
gs-3955	52	16	.	.	PUNCT
gs-3955	53	1	hydra	hydra	PROPN
gs-3955	53	2	similarly	similarly	ADV
gs-3955	53	3	demonstrates	demonstrate	VERB
gs-3955	53	4	high	high	ADJ
gs-3955	53	5	autophagic	autophagic	ADJ
gs-3955	53	6	capacity	capacity	NOUN
gs-3955	53	7	during	during	ADP
gs-3955	53	8	nutrient	nutrient	ADJ
gs-3955	53	9	deprivation	deprivation	NOUN
gs-3955	53	10	,	,	PUNCT
gs-3955	53	11	supporting	support	VERB
gs-3955	53	12	its	its	PRON
gs-3955	53	13	regenerative	regenerative	ADJ
gs-3955	53	14	potential	potential	NOUN
gs-3955	53	15	(	(	PUNCT
gs-3955	53	16	buzgariu	buzgariu	PROPN
gs-3955	53	17	et	et	PROPN
gs-3955	53	18	al	al	PROPN
gs-3955	53	19	.	.	PROPN
gs-3955	53	20	,	,	PUNCT
gs-3955	53	21	2008	2008	NUM
gs-3955	53	22	)	)	PUNCT
gs-3955	53	23	.	.	PUNCT
gs-3955	54	1	cancer	cancer	NOUN
gs-3955	54	2	resistance	resistance	NOUN
gs-3955	54	3	and	and	CCONJ
gs-3955	54	4	genomic	genomic	NOUN
gs-3955	54	5	surveillance	surveillance	NOUN
gs-3955	54	6	cancer	cancer	NOUN
gs-3955	54	7	is	be	AUX
gs-3955	54	8	rare	rare	ADJ
gs-3955	54	9	or	or	CCONJ
gs-3955	54	10	absent	absent	ADJ
gs-3955	54	11	in	in	ADP
gs-3955	54	12	several	several	ADJ
gs-3955	54	13	negligibly	negligibly	ADV
gs-3955	54	14	senescent	senescent	ADJ
gs-3955	54	15	organisms	organism	NOUN
gs-3955	54	16	.	.	PUNCT
gs-3955	55	1	naked	naked	ADJ
gs-3955	55	2	mole	mole	NOUN
gs-3955	55	3	-	-	PUNCT
gs-3955	55	4	rats	rat	NOUN
gs-3955	55	5	resist	resist	VERB
gs-3955	55	6	oncogenesis	oncogenesis	NOUN
gs-3955	55	7	via	via	ADP
gs-3955	55	8	several	several	ADJ
gs-3955	55	9	mechanisms	mechanism	NOUN
gs-3955	55	10	,	,	PUNCT
gs-3955	55	11	including	include	VERB
gs-3955	55	12	high	high	ADJ
gs-3955	55	13	-	-	PUNCT
gs-3955	55	14	molecular	molecular	ADJ
gs-3955	55	15	-	-	PUNCT
gs-3955	55	16	weight	weight	NOUN
gs-3955	55	17	hyaluronan	hyaluronan	NOUN
gs-3955	55	18	(	(	PUNCT
gs-3955	55	19	hmwha	hmwha	PROPN
gs-3955	55	20	)	)	PUNCT
gs-3955	55	21	that	that	PRON
gs-3955	55	22	promotes	promote	VERB
gs-3955	55	23	early	early	ADJ
gs-3955	55	24	contact	contact	NOUN
gs-3955	55	25	inhibition	inhibition	NOUN
gs-3955	55	26	,	,	PUNCT
gs-3955	55	27	overexpression	overexpression	NOUN
gs-3955	55	28	of	of	ADP
gs-3955	55	29	tumor	tumor	NOUN
gs-3955	55	30	suppressors	suppressor	NOUN
gs-3955	55	31	such	such	ADJ
gs-3955	55	32	as	as	ADP
gs-3955	55	33	p16	p16	NOUN
gs-3955	55	34	and	and	CCONJ
gs-3955	55	35	p53	p53	NOUN
gs-3955	55	36	,	,	PUNCT
gs-3955	55	37	and	and	CCONJ
gs-3955	55	38	an	an	DET
gs-3955	55	39	unusual	unusual	ADJ
gs-3955	55	40	necroptosis	necroptosis	NOUN
gs-3955	55	41	pathway	pathway	NOUN
gs-3955	55	42	that	that	PRON
gs-3955	55	43	removes	remove	VERB
gs-3955	55	44	precancerous	precancerous	ADJ
gs-3955	55	45	cells	cell	NOUN
gs-3955	55	46	(	(	PUNCT
gs-3955	55	47	tian	tian	PROPN
gs-3955	55	48	et	et	PROPN
gs-3955	55	49	al	al	PROPN
gs-3955	55	50	.	.	PROPN
gs-3955	55	51	,	,	PUNCT
gs-3955	55	52	2013	2013	NUM
gs-3955	55	53	)	)	PUNCT
gs-3955	55	54	.	.	PUNCT
gs-3955	56	1	in	in	ADP
gs-3955	56	2	hydra	hydra	PROPN
gs-3955	56	3	and	and	CCONJ
gs-3955	56	4	planarians	planarian	NOUN
gs-3955	56	5	,	,	PUNCT
gs-3955	56	6	continuous	continuous	ADJ
gs-3955	56	7	stem	stem	NOUN
gs-3955	56	8	cell	cell	NOUN
gs-3955	56	9	division	division	NOUN
gs-3955	56	10	is	be	AUX
gs-3955	56	11	tightly	tightly	ADV
gs-3955	56	12	regulated	regulate	VERB
gs-3955	56	13	by	by	ADP
gs-3955	56	14	conserved	conserve	VERB
gs-3955	56	15	pathways	pathway	NOUN
gs-3955	56	16	such	such	ADJ
gs-3955	56	17	as	as	ADP
gs-3955	56	18	wnt	wnt	PROPN
gs-3955	56	19	,	,	PUNCT
gs-3955	56	20	bmp	bmp	PROPN
gs-3955	56	21	,	,	PUNCT
gs-3955	56	22	and	and	CCONJ
gs-3955	56	23	notch	notch	ADJ
gs-3955	56	24	,	,	PUNCT
gs-3955	56	25	preventing	prevent	VERB
gs-3955	56	26	overproliferation	overproliferation	NOUN
gs-3955	56	27	and	and	CCONJ
gs-3955	56	28	tumorigenesis	tumorigenesis	NOUN
gs-3955	56	29	(	(	PUNCT
gs-3955	56	30	juliano	juliano	PROPN
gs-3955	56	31	et	et	PROPN
gs-3955	56	32	al	al	PROPN
gs-3955	56	33	.	.	PROPN
gs-3955	56	34	,	,	PUNCT
gs-3955	56	35	2014	2014	NUM
gs-3955	56	36	)	)	PUNCT
gs-3955	56	37	.	.	PUNCT
gs-3955	57	1	in	in	ADP
gs-3955	57	2	the	the	DET
gs-3955	57	3	greenland	greenland	NOUN
gs-3955	57	4	shark	shark	NOUN
gs-3955	57	5	and	and	CCONJ
gs-3955	57	6	a.	a.	PROPN
gs-3955	57	7	islandica	islandica	PROPN
gs-3955	57	8	,	,	PUNCT
gs-3955	57	9	slow	slow	ADJ
gs-3955	57	10	cell	cell	NOUN
gs-3955	57	11	cycling	cycling	NOUN
gs-3955	57	12	and	and	CCONJ
gs-3955	57	13	low	low	ADJ
gs-3955	57	14	metabolic	metabolic	NOUN
gs-3955	57	15	rates	rate	NOUN
gs-3955	57	16	reduce	reduce	VERB
gs-3955	57	17	mutation	mutation	NOUN
gs-3955	57	18	accumulation	accumulation	NOUN
gs-3955	57	19	and	and	CCONJ
gs-3955	57	20	oncogenic	oncogenic	ADJ
gs-3955	57	21	transformation	transformation	NOUN
gs-3955	57	22	risk	risk	NOUN
gs-3955	57	23	.	.	PUNCT
gs-3955	58	1	georgian	georgian	ADJ
gs-3955	58	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	58	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	58	4	ტ.	ტ.	VERB
gs-3955	58	5	7	7	NUM
gs-3955	58	6	n	n	PRON
gs-3955	58	7	3	3	NUM
gs-3955	58	8	,	,	PUNCT
gs-3955	58	9	2025	2025	NUM
gs-3955	58	10	350	350	NUM
gs-3955	58	11	epigenetic	epigenetic	ADJ
gs-3955	58	12	stability	stability	NOUN
gs-3955	58	13	epigenetic	epigenetic	ADJ
gs-3955	58	14	drift	drift	NOUN
gs-3955	58	15	contributes	contribute	VERB
gs-3955	58	16	to	to	ADP
gs-3955	58	17	transcriptional	transcriptional	ADJ
gs-3955	58	18	noise	noise	NOUN
gs-3955	58	19	and	and	CCONJ
gs-3955	58	20	loss	loss	NOUN
gs-3955	58	21	of	of	ADP
gs-3955	58	22	cell	cell	NOUN
gs-3955	58	23	identity	identity	NOUN
gs-3955	58	24	during	during	ADP
gs-3955	58	25	aging	age	VERB
gs-3955	58	26	.	.	PUNCT
gs-3955	59	1	long	long	ADV
gs-3955	59	2	-	-	PUNCT
gs-3955	59	3	lived	live	VERB
gs-3955	59	4	species	specie	NOUN
gs-3955	59	5	maintain	maintain	VERB
gs-3955	59	6	a	a	DET
gs-3955	59	7	stable	stable	ADJ
gs-3955	59	8	epigenomic	epigenomic	ADJ
gs-3955	59	9	landscape	landscape	NOUN
gs-3955	59	10	via	via	ADP
gs-3955	59	11	slow	slow	ADJ
gs-3955	59	12	methylation	methylation	NOUN
gs-3955	59	13	turnover	turnover	NOUN
gs-3955	59	14	,	,	PUNCT
gs-3955	59	15	histone	histone	NOUN
gs-3955	59	16	modification	modification	NOUN
gs-3955	59	17	patterns	pattern	NOUN
gs-3955	59	18	favoring	favor	VERB
gs-3955	59	19	genomic	genomic	ADJ
gs-3955	59	20	compaction	compaction	NOUN
gs-3955	59	21	,	,	PUNCT
gs-3955	59	22	and	and	CCONJ
gs-3955	59	23	robust	robust	ADJ
gs-3955	59	24	expression	expression	NOUN
gs-3955	59	25	of	of	ADP
gs-3955	59	26	epigenetic	epigenetic	ADJ
gs-3955	59	27	maintenance	maintenance	NOUN
gs-3955	59	28	enzymes	enzyme	NOUN
gs-3955	59	29	(	(	PUNCT
gs-3955	59	30	field	field	NOUN
gs-3955	59	31	et	et	PROPN
gs-3955	59	32	al	al	PROPN
gs-3955	59	33	.	.	PROPN
gs-3955	59	34	,	,	PUNCT
gs-3955	59	35	2022	2022	NUM
gs-3955	59	36	)	)	PUNCT
gs-3955	59	37	.	.	PUNCT
gs-3955	60	1	recent	recent	ADJ
gs-3955	60	2	methylome	methylome	ADJ
gs-3955	60	3	comparisons	comparison	NOUN
gs-3955	60	4	show	show	VERB
gs-3955	60	5	that	that	SCONJ
gs-3955	60	6	a.	a.	NOUN
gs-3955	60	7	islandica	islandica	NOUN
gs-3955	60	8	and	and	CCONJ
gs-3955	60	9	naked	naked	ADJ
gs-3955	60	10	mole	mole	NOUN
gs-3955	60	11	-	-	PUNCT
gs-3955	60	12	rats	rat	NOUN
gs-3955	60	13	exhibit	exhibit	VERB
gs-3955	60	14	far	far	ADV
gs-3955	60	15	slower	slow	ADJ
gs-3955	60	16	rates	rate	NOUN
gs-3955	60	17	of	of	ADP
gs-3955	60	18	age	age	NOUN
gs-3955	60	19	-	-	PUNCT
gs-3955	60	20	associated	associate	VERB
gs-3955	60	21	cpg	cpg	NOUN
gs-3955	60	22	methylation	methylation	NOUN
gs-3955	60	23	changes	change	NOUN
gs-3955	60	24	compared	compare	VERB
gs-3955	60	25	to	to	ADP
gs-3955	60	26	short	short	ADJ
gs-3955	60	27	-	-	PUNCT
gs-3955	60	28	lived	live	VERB
gs-3955	60	29	species	specie	NOUN
gs-3955	60	30	(	(	PUNCT
gs-3955	60	31	meer	meer	NOUN
gs-3955	60	32	et	et	PROPN
gs-3955	60	33	al	al	PROPN
gs-3955	60	34	.	.	PROPN
gs-3955	60	35	,	,	PUNCT
gs-3955	60	36	2018	2018	NUM
gs-3955	60	37	)	)	PUNCT
gs-3955	60	38	.	.	PUNCT
gs-3955	61	1	this	this	DET
gs-3955	61	2	epigenetic	epigenetic	ADJ
gs-3955	61	3	resilience	resilience	NOUN
gs-3955	61	4	supports	support	VERB
gs-3955	61	5	tissue	tissue	NOUN
gs-3955	61	6	homeostasis	homeostasis	NOUN
gs-3955	61	7	and	and	CCONJ
gs-3955	61	8	prevents	prevent	VERB
gs-3955	61	9	activation	activation	NOUN
gs-3955	61	10	of	of	ADP
gs-3955	61	11	inflammatory	inflammatory	ADJ
gs-3955	61	12	or	or	CCONJ
gs-3955	61	13	apoptotic	apoptotic	ADJ
gs-3955	61	14	gene	gene	NOUN
gs-3955	61	15	programs	program	NOUN
gs-3955	61	16	.	.	PUNCT
gs-3955	62	1	environmental	environmental	ADJ
gs-3955	62	2	and	and	CCONJ
gs-3955	62	3	evolutionary	evolutionary	ADJ
gs-3955	62	4	context	context	NOUN
gs-3955	62	5	negligibly	negligibly	ADV
gs-3955	62	6	senescent	senescent	ADJ
gs-3955	62	7	organisms	organism	NOUN
gs-3955	62	8	typically	typically	ADV
gs-3955	62	9	inhabit	inhabit	VERB
gs-3955	62	10	environments	environment	NOUN
gs-3955	62	11	with	with	ADP
gs-3955	62	12	low	low	ADJ
gs-3955	62	13	extrinsic	extrinsic	ADJ
gs-3955	62	14	mortality	mortality	NOUN
gs-3955	62	15	—	—	PUNCT
gs-3955	62	16	deep	deep	ADJ
gs-3955	62	17	oceans	ocean	NOUN
gs-3955	62	18	,	,	PUNCT
gs-3955	62	19	subterranean	subterranean	ADJ
gs-3955	62	20	tunnels	tunnel	NOUN
gs-3955	62	21	,	,	PUNCT
gs-3955	62	22	or	or	CCONJ
gs-3955	62	23	isolated	isolated	ADJ
gs-3955	62	24	caves	cave	NOUN
gs-3955	62	25	—	—	PUNCT
gs-3955	62	26	where	where	SCONJ
gs-3955	62	27	predation	predation	NOUN
gs-3955	62	28	,	,	PUNCT
gs-3955	62	29	seasonal	seasonal	ADJ
gs-3955	62	30	change	change	NOUN
gs-3955	62	31	,	,	PUNCT
gs-3955	62	32	and	and	CCONJ
gs-3955	62	33	starvation	starvation	NOUN
gs-3955	62	34	are	be	AUX
gs-3955	62	35	minimized	minimize	VERB
gs-3955	62	36	(	(	PUNCT
gs-3955	62	37	kirkwood	kirkwood	PROPN
gs-3955	62	38	&	&	CCONJ
gs-3955	62	39	austad	austad	PROPN
gs-3955	62	40	,	,	PUNCT
gs-3955	62	41	2000	2000	NUM
gs-3955	62	42	)	)	PUNCT
gs-3955	62	43	.	.	PUNCT
gs-3955	63	1	this	this	DET
gs-3955	63	2	ecological	ecological	ADJ
gs-3955	63	3	stability	stability	NOUN
gs-3955	63	4	shifts	shift	NOUN
gs-3955	63	5	evolutionary	evolutionary	ADJ
gs-3955	63	6	pressure	pressure	NOUN
gs-3955	63	7	away	away	ADV
gs-3955	63	8	from	from	ADP
gs-3955	63	9	early	early	ADJ
gs-3955	63	10	reproduction	reproduction	NOUN
gs-3955	63	11	toward	toward	ADP
gs-3955	63	12	somatic	somatic	ADJ
gs-3955	63	13	maintenance	maintenance	NOUN
gs-3955	63	14	,	,	PUNCT
gs-3955	63	15	allowing	allow	VERB
gs-3955	63	16	the	the	DET
gs-3955	63	17	selection	selection	NOUN
gs-3955	63	18	of	of	ADP
gs-3955	63	19	traits	trait	NOUN
gs-3955	63	20	favoring	favor	VERB
gs-3955	63	21	longevity	longevity	NOUN
gs-3955	63	22	(	(	PUNCT
gs-3955	63	23	williams	williams	PROPN
gs-3955	63	24	,	,	PUNCT
gs-3955	63	25	1957	1957	NUM
gs-3955	63	26	)	)	PUNCT
gs-3955	63	27	.	.	PUNCT
gs-3955	64	1	the	the	DET
gs-3955	64	2	selection	selection	NOUN
gs-3955	64	3	shadow	shadow	NOUN
gs-3955	64	4	hypothesis	hypothesis	NOUN
gs-3955	64	5	and	and	CCONJ
gs-3955	64	6	antagonistic	antagonistic	ADJ
gs-3955	64	7	pleiotropy	pleiotropy	NOUN
gs-3955	64	8	theory	theory	NOUN
gs-3955	64	9	provide	provide	VERB
gs-3955	64	10	theoretical	theoretical	ADJ
gs-3955	64	11	frameworks	framework	NOUN
gs-3955	64	12	explaining	explain	VERB
gs-3955	64	13	how	how	SCONJ
gs-3955	64	14	traits	trait	NOUN
gs-3955	64	15	beneficial	beneficial	ADJ
gs-3955	64	16	in	in	ADP
gs-3955	64	17	youth	youth	NOUN
gs-3955	64	18	may	may	AUX
gs-3955	64	19	be	be	AUX
gs-3955	64	20	deleterious	deleterious	ADJ
gs-3955	64	21	later	later	ADV
gs-3955	64	22	in	in	ADP
gs-3955	64	23	life	life	NOUN
gs-3955	64	24	.	.	PUNCT
gs-3955	65	1	in	in	ADP
gs-3955	65	2	species	specie	NOUN
gs-3955	65	3	with	with	ADP
gs-3955	65	4	negligible	negligible	ADJ
gs-3955	65	5	senescence	senescence	NOUN
gs-3955	65	6	,	,	PUNCT
gs-3955	65	7	such	such	ADJ
gs-3955	65	8	late	late	ADJ
gs-3955	65	9	-	-	PUNCT
gs-3955	65	10	life	life	NOUN
gs-3955	65	11	trade	trade	NOUN
gs-3955	65	12	-	-	PUNCT
gs-3955	65	13	offs	off	NOUN
gs-3955	65	14	appear	appear	VERB
gs-3955	65	15	to	to	PART
gs-3955	65	16	have	have	AUX
gs-3955	65	17	been	be	AUX
gs-3955	65	18	neutralized	neutralize	VERB
gs-3955	65	19	or	or	CCONJ
gs-3955	65	20	reprogrammed	reprogramme	VERB
gs-3955	65	21	(	(	PUNCT
gs-3955	65	22	hamilton	hamilton	PROPN
gs-3955	65	23	,	,	PUNCT
gs-3955	65	24	1966	1966	NUM
gs-3955	65	25	)	)	PUNCT
gs-3955	65	26	.	.	PUNCT
gs-3955	66	1	theoretical	theoretical	ADJ
gs-3955	66	2	frameworks	framework	NOUN
gs-3955	66	3	for	for	ADP
gs-3955	66	4	negligible	negligible	ADJ
gs-3955	66	5	senescence	senescence	NOUN
gs-3955	66	6	the	the	DET
gs-3955	66	7	centrosomal	centrosomal	ADJ
gs-3955	66	8	theory	theory	NOUN
gs-3955	66	9	of	of	ADP
gs-3955	66	10	aging	age	VERB
gs-3955	66	11	posits	posit	NOUN
gs-3955	66	12	that	that	PRON
gs-3955	66	13	asymmetric	asymmetric	ADJ
gs-3955	66	14	cell	cell	NOUN
gs-3955	66	15	division	division	NOUN
gs-3955	66	16	creates	create	VERB
gs-3955	66	17	cumulative	cumulative	ADJ
gs-3955	66	18	damage	damage	NOUN
gs-3955	66	19	asymmetry	asymmetry	NOUN
gs-3955	66	20	in	in	ADP
gs-3955	66	21	daughter	daughter	NOUN
gs-3955	66	22	cells	cell	NOUN
gs-3955	66	23	,	,	PUNCT
gs-3955	66	24	leading	lead	VERB
gs-3955	66	25	to	to	ADP
gs-3955	66	26	functional	functional	ADJ
gs-3955	66	27	decline	decline	NOUN
gs-3955	66	28	in	in	ADP
gs-3955	66	29	somatic	somatic	ADJ
gs-3955	66	30	tissues	tissue	NOUN
gs-3955	66	31	(	(	PUNCT
gs-3955	66	32	ratajczak	ratajczak	NOUN
gs-3955	66	33	et	et	PROPN
gs-3955	66	34	al	al	PROPN
gs-3955	66	35	.	.	PROPN
gs-3955	66	36	,	,	PUNCT
gs-3955	66	37	2018	2018	NUM
gs-3955	66	38	)	)	PUNCT
gs-3955	66	39	.	.	PUNCT
gs-3955	67	1	species	specie	NOUN
gs-3955	67	2	like	like	ADP
gs-3955	67	3	hydra	hydra	NOUN
gs-3955	67	4	and	and	CCONJ
gs-3955	67	5	planarians	planarian	NOUN
gs-3955	67	6	,	,	PUNCT
gs-3955	67	7	which	which	PRON
gs-3955	67	8	rely	rely	VERB
gs-3955	67	9	on	on	ADP
gs-3955	67	10	symmetrical	symmetrical	ADJ
gs-3955	67	11	stem	stem	NOUN
gs-3955	67	12	cell	cell	NOUN
gs-3955	67	13	divisions	division	NOUN
gs-3955	67	14	and	and	CCONJ
gs-3955	67	15	continuous	continuous	ADJ
gs-3955	67	16	cell	cell	NOUN
gs-3955	67	17	replacement	replacement	NOUN
gs-3955	67	18	,	,	PUNCT
gs-3955	67	19	escape	escape	VERB
gs-3955	67	20	this	this	DET
gs-3955	67	21	trajectory	trajectory	NOUN
gs-3955	67	22	.	.	PUNCT
gs-3955	68	1	additionally	additionally	ADV
gs-3955	68	2	,	,	PUNCT
gs-3955	68	3	the	the	DET
gs-3955	68	4	thermodynamic	thermodynamic	ADJ
gs-3955	68	5	theory	theory	NOUN
gs-3955	68	6	of	of	ADP
gs-3955	68	7	aging	aging	NOUN
gs-3955	68	8	suggests	suggest	VERB
gs-3955	68	9	that	that	SCONJ
gs-3955	68	10	biological	biological	ADJ
gs-3955	68	11	systems	system	NOUN
gs-3955	68	12	age	age	NOUN
gs-3955	68	13	due	due	ADP
gs-3955	68	14	to	to	ADP
gs-3955	68	15	entropy	entropy	VERB
gs-3955	68	16	accumulation	accumulation	NOUN
gs-3955	68	17	.	.	PUNCT
gs-3955	69	1	organisms	organism	NOUN
gs-3955	69	2	with	with	ADP
gs-3955	69	3	superior	superior	ADJ
gs-3955	69	4	damage	damage	NOUN
gs-3955	69	5	repair	repair	NOUN
gs-3955	69	6	,	,	PUNCT
gs-3955	69	7	proteostasis	proteostasis	NOUN
gs-3955	69	8	,	,	PUNCT
gs-3955	69	9	and	and	CCONJ
gs-3955	69	10	low	low	ADJ
gs-3955	69	11	metabolic	metabolic	NOUN
gs-3955	69	12	rates	rate	NOUN
gs-3955	69	13	are	be	AUX
gs-3955	69	14	better	well	ADJ
gs-3955	69	15	at	at	ADP
gs-3955	69	16	entropy	entropy	NOUN
gs-3955	69	17	management	management	NOUN
gs-3955	69	18	and	and	CCONJ
gs-3955	69	19	thus	thus	ADV
gs-3955	69	20	exhibit	exhibit	VERB
gs-3955	69	21	slower	slow	ADJ
gs-3955	69	22	aging	aging	NOUN
gs-3955	69	23	(	(	PUNCT
gs-3955	69	24	demetrius	demetrius	PROPN
gs-3955	69	25	,	,	PUNCT
gs-3955	69	26	2006	2006	NUM
gs-3955	69	27	)	)	PUNCT
gs-3955	69	28	.	.	PUNCT
gs-3955	70	1	translational	translational	ADJ
gs-3955	70	2	implications	implication	NOUN
gs-3955	70	3	understanding	understand	VERB
gs-3955	70	4	how	how	SCONJ
gs-3955	70	5	negligible	negligible	ADJ
gs-3955	70	6	senescence	senescence	NOUN
gs-3955	70	7	is	be	AUX
gs-3955	70	8	achieved	achieve	VERB
gs-3955	70	9	in	in	ADP
gs-3955	70	10	nature	nature	NOUN
gs-3955	70	11	enables	enable	VERB
gs-3955	70	12	a	a	DET
gs-3955	70	13	re	re	NOUN
gs-3955	70	14	-	-	NOUN
gs-3955	70	15	evaluation	evaluation	NOUN
gs-3955	70	16	of	of	ADP
gs-3955	70	17	aging	aging	NOUN
gs-3955	70	18	as	as	ADP
gs-3955	70	19	an	an	DET
gs-3955	70	20	inevitable	inevitable	ADJ
gs-3955	70	21	process	process	NOUN
gs-3955	70	22	.	.	PUNCT
gs-3955	71	1	by	by	ADP
gs-3955	71	2	identifying	identify	VERB
gs-3955	71	3	conserved	conserve	VERB
gs-3955	71	4	molecular	molecular	ADJ
gs-3955	71	5	pathways	pathway	NOUN
gs-3955	71	6	,	,	PUNCT
gs-3955	71	7	researchers	researcher	NOUN
gs-3955	71	8	can	can	AUX
gs-3955	71	9	target	target	VERB
gs-3955	71	10	similar	similar	ADJ
gs-3955	71	11	mechanisms	mechanism	NOUN
gs-3955	71	12	in	in	ADP
gs-3955	71	13	humans	human	NOUN
gs-3955	71	14	—	—	PUNCT
gs-3955	71	15	via	via	ADP
gs-3955	71	16	gene	gene	NOUN
gs-3955	71	17	therapy	therapy	NOUN
gs-3955	71	18	,	,	PUNCT
gs-3955	71	19	pharmacological	pharmacological	ADJ
gs-3955	71	20	mimetics	mimetic	NOUN
gs-3955	71	21	,	,	PUNCT
gs-3955	71	22	or	or	CCONJ
gs-3955	71	23	engineered	engineer	VERB
gs-3955	71	24	stem	stem	NOUN
gs-3955	71	25	cells	cell	NOUN
gs-3955	71	26	—	—	PUNCT
gs-3955	71	27	to	to	PART
gs-3955	71	28	delay	delay	VERB
gs-3955	71	29	or	or	CCONJ
gs-3955	71	30	reverse	reverse	VERB
gs-3955	71	31	age	age	NOUN
gs-3955	71	32	-	-	PUNCT
gs-3955	71	33	associated	associate	VERB
gs-3955	71	34	decline	decline	NOUN
gs-3955	71	35	(	(	PUNCT
gs-3955	71	36	chakrabarti	chakrabarti	PROPN
gs-3955	71	37	et	et	PROPN
gs-3955	71	38	al	al	PROPN
gs-3955	71	39	.	.	PROPN
gs-3955	71	40	,	,	PUNCT
gs-3955	71	41	2021	2021	NUM
gs-3955	71	42	)	)	PUNCT
gs-3955	71	43	.	.	PUNCT
gs-3955	72	1	already	already	ADV
gs-3955	72	2	,	,	PUNCT
gs-3955	72	3	compounds	compound	VERB
gs-3955	72	4	mimicking	mimic	VERB
gs-3955	72	5	sirt6	sirt6	NOUN
gs-3955	72	6	activity	activity	NOUN
gs-3955	72	7	(	(	PUNCT
gs-3955	72	8	e.g.	e.g.	ADV
gs-3955	72	9	,	,	PUNCT
gs-3955	72	10	mdl-800	mdl-800	ADJ
gs-3955	72	11	)	)	PUNCT
gs-3955	72	12	or	or	CCONJ
gs-3955	72	13	foxo	foxo	PROPN
gs-3955	72	14	stabilization	stabilization	NOUN
gs-3955	72	15	are	be	AUX
gs-3955	72	16	under	under	ADP
gs-3955	72	17	preclinical	preclinical	ADJ
gs-3955	72	18	and	and	CCONJ
gs-3955	72	19	early	early	ADJ
gs-3955	72	20	clinical	clinical	ADJ
gs-3955	72	21	evaluation	evaluation	NOUN
gs-3955	72	22	.	.	PUNCT
gs-3955	73	1	high	high	ADJ
gs-3955	73	2	-	-	PUNCT
gs-3955	73	3	molecular	molecular	ADJ
gs-3955	73	4	-	-	PUNCT
gs-3955	73	5	weight	weight	NOUN
gs-3955	73	6	hyaluronan	hyaluronan	NOUN
gs-3955	73	7	is	be	AUX
gs-3955	73	8	being	be	AUX
gs-3955	73	9	georgian	georgian	ADJ
gs-3955	73	10	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	73	11	მეცნიერები	მეცნიერები	NOUN
gs-3955	73	12	ტ.	ტ.	VERB
gs-3955	73	13	7	7	NUM
gs-3955	73	14	n	n	PRON
gs-3955	73	15	3	3	NUM
gs-3955	73	16	,	,	PUNCT
gs-3955	73	17	2025	2025	NUM
gs-3955	73	18	351	351	NUM
gs-3955	73	19	explored	explore	VERB
gs-3955	73	20	for	for	ADP
gs-3955	73	21	its	its	PRON
gs-3955	73	22	anti	anti	ADJ
gs-3955	73	23	-	-	ADJ
gs-3955	73	24	inflammatory	inflammatory	ADJ
gs-3955	73	25	and	and	CCONJ
gs-3955	73	26	anti	anti	ADJ
gs-3955	73	27	-	-	ADJ
gs-3955	73	28	cancer	cancer	ADJ
gs-3955	73	29	properties	property	NOUN
gs-3955	73	30	,	,	PUNCT
gs-3955	73	31	while	while	SCONJ
gs-3955	73	32	nrf2	nrf2	NOUN
gs-3955	73	33	activators	activator	NOUN
gs-3955	73	34	aim	aim	VERB
gs-3955	73	35	to	to	PART
gs-3955	73	36	replicate	replicate	VERB
gs-3955	73	37	oxidative	oxidative	ADJ
gs-3955	73	38	stress	stress	NOUN
gs-3955	73	39	resistance	resistance	NOUN
gs-3955	73	40	observed	observe	VERB
gs-3955	73	41	in	in	ADP
gs-3955	73	42	naked	naked	ADJ
gs-3955	73	43	mole	mole	NOUN
gs-3955	73	44	-	-	PUNCT
gs-3955	73	45	rats	rat	NOUN
gs-3955	73	46	.	.	PUNCT
gs-3955	74	1	negligible	negligible	ADJ
gs-3955	74	2	senescence	senescence	NOUN
gs-3955	74	3	represents	represent	VERB
gs-3955	74	4	not	not	PART
gs-3955	74	5	a	a	DET
gs-3955	74	6	biological	biological	ADJ
gs-3955	74	7	anomaly	anomaly	NOUN
gs-3955	74	8	but	but	CCONJ
gs-3955	74	9	an	an	DET
gs-3955	74	10	evolved	evolved	ADJ
gs-3955	74	11	solution	solution	NOUN
gs-3955	74	12	to	to	ADP
gs-3955	74	13	cellular	cellular	ADJ
gs-3955	74	14	decay	decay	NOUN
gs-3955	74	15	.	.	PUNCT
gs-3955	75	1	by	by	ADP
gs-3955	75	2	integrating	integrate	VERB
gs-3955	75	3	systems	system	NOUN
gs-3955	75	4	biology	biology	NOUN
gs-3955	75	5	,	,	PUNCT
gs-3955	75	6	comparative	comparative	ADJ
gs-3955	75	7	genomics	genomic	NOUN
gs-3955	75	8	,	,	PUNCT
gs-3955	75	9	and	and	CCONJ
gs-3955	75	10	evolutionary	evolutionary	ADJ
gs-3955	75	11	theory	theory	NOUN
gs-3955	75	12	,	,	PUNCT
gs-3955	75	13	the	the	DET
gs-3955	75	14	study	study	NOUN
gs-3955	75	15	of	of	ADP
gs-3955	75	16	these	these	DET
gs-3955	75	17	exceptional	exceptional	ADJ
gs-3955	75	18	organisms	organism	NOUN
gs-3955	75	19	not	not	PART
gs-3955	75	20	only	only	ADV
gs-3955	75	21	reveals	reveal	VERB
gs-3955	75	22	the	the	DET
gs-3955	75	23	plasticity	plasticity	NOUN
gs-3955	75	24	of	of	ADP
gs-3955	75	25	aging	aging	NOUN
gs-3955	75	26	but	but	CCONJ
gs-3955	75	27	also	also	ADV
gs-3955	75	28	paves	pave	VERB
gs-3955	75	29	the	the	DET
gs-3955	75	30	way	way	NOUN
gs-3955	75	31	toward	toward	ADP
gs-3955	75	32	its	its	PRON
gs-3955	75	33	potential	potential	ADJ
gs-3955	75	34	modulation	modulation	NOUN
gs-3955	75	35	.	.	PUNCT
gs-3955	76	1	as	as	ADP
gs-3955	76	2	technology	technology	NOUN
gs-3955	76	3	advances	advance	NOUN
gs-3955	76	4	,	,	PUNCT
gs-3955	76	5	insights	insight	NOUN
gs-3955	76	6	from	from	ADP
gs-3955	76	7	hydra	hydra	PROPN
gs-3955	76	8	,	,	PUNCT
gs-3955	76	9	naked	naked	ADJ
gs-3955	76	10	mole	mole	NOUN
gs-3955	76	11	-	-	PUNCT
gs-3955	76	12	rats	rat	NOUN
gs-3955	76	13	,	,	PUNCT
gs-3955	76	14	and	and	CCONJ
gs-3955	76	15	deep	deep	ADJ
gs-3955	76	16	-	-	PUNCT
gs-3955	76	17	sea	sea	NOUN
gs-3955	76	18	sharks	shark	NOUN
gs-3955	76	19	may	may	AUX
gs-3955	76	20	ultimately	ultimately	ADV
gs-3955	76	21	inform	inform	VERB
gs-3955	76	22	therapeutic	therapeutic	ADJ
gs-3955	76	23	strategies	strategy	NOUN
gs-3955	76	24	for	for	ADP
gs-3955	76	25	extending	extend	VERB
gs-3955	76	26	human	human	ADJ
gs-3955	76	27	vitality	vitality	NOUN
gs-3955	76	28	and	and	CCONJ
gs-3955	76	29	compressing	compress	VERB
gs-3955	76	30	morbidity	morbidity	NOUN
gs-3955	76	31	.	.	PUNCT
gs-3955	77	1	comparative	comparative	ADJ
gs-3955	77	2	analysis	analysis	NOUN
gs-3955	77	3	of	of	ADP
gs-3955	77	4	negligibly	negligibly	ADV
gs-3955	77	5	senescing	senesce	VERB
gs-3955	77	6	species	specie	NOUN
gs-3955	77	7	the	the	DET
gs-3955	77	8	phenomenon	phenomenon	NOUN
gs-3955	77	9	of	of	ADP
gs-3955	77	10	biological	biological	ADJ
gs-3955	77	11	aging	aging	NOUN
gs-3955	77	12	represents	represent	VERB
gs-3955	77	13	a	a	DET
gs-3955	77	14	complex	complex	ADJ
gs-3955	77	15	multifactorial	multifactorial	ADJ
gs-3955	77	16	process	process	NOUN
gs-3955	77	17	characterized	characterize	VERB
gs-3955	77	18	by	by	ADP
gs-3955	77	19	progressive	progressive	ADJ
gs-3955	77	20	functional	functional	ADJ
gs-3955	77	21	decline	decline	NOUN
gs-3955	77	22	across	across	ADP
gs-3955	77	23	physiological	physiological	ADJ
gs-3955	77	24	systems	system	NOUN
gs-3955	77	25	and	and	CCONJ
gs-3955	77	26	increased	increase	VERB
gs-3955	77	27	vulnerability	vulnerability	NOUN
gs-3955	77	28	to	to	ADP
gs-3955	77	29	age	age	NOUN
gs-3955	77	30	-	-	PUNCT
gs-3955	77	31	associated	associate	VERB
gs-3955	77	32	pathologies	pathology	NOUN
gs-3955	77	33	(	(	PUNCT
gs-3955	77	34	lópez	lópez	NOUN
gs-3955	77	35	-	-	PUNCT
gs-3955	77	36	otín	otín	NOUN
gs-3955	77	37	et	et	PROPN
gs-3955	77	38	al	al	PROPN
gs-3955	77	39	.	.	PROPN
gs-3955	77	40	,	,	PUNCT
gs-3955	77	41	2013	2013	NUM
gs-3955	77	42	)	)	PUNCT
gs-3955	77	43	.	.	PUNCT
gs-3955	78	1	contrary	contrary	ADJ
gs-3955	78	2	to	to	ADP
gs-3955	78	3	this	this	DET
gs-3955	78	4	nearly	nearly	ADV
gs-3955	78	5	universal	universal	ADJ
gs-3955	78	6	pattern	pattern	NOUN
gs-3955	78	7	,	,	PUNCT
gs-3955	78	8	certain	certain	ADJ
gs-3955	78	9	species	specie	NOUN
gs-3955	78	10	exhibit	exhibit	VERB
gs-3955	78	11	negligible	negligible	ADJ
gs-3955	78	12	senescence	senescence	NOUN
gs-3955	78	13	a	a	DET
gs-3955	78	14	remarkable	remarkable	ADJ
gs-3955	78	15	biological	biological	ADJ
gs-3955	78	16	state	state	NOUN
gs-3955	78	17	where	where	SCONJ
gs-3955	78	18	mortality	mortality	NOUN
gs-3955	78	19	risk	risk	NOUN
gs-3955	78	20	remains	remain	VERB
gs-3955	78	21	stable	stable	ADJ
gs-3955	78	22	throughout	throughout	ADP
gs-3955	78	23	adulthood	adulthood	NOUN
gs-3955	78	24	and	and	CCONJ
gs-3955	78	25	physiological	physiological	ADJ
gs-3955	78	26	functions	function	NOUN
gs-3955	78	27	are	be	AUX
gs-3955	78	28	maintained	maintain	VERB
gs-3955	78	29	over	over	ADP
gs-3955	78	30	exceptionally	exceptionally	ADV
gs-3955	78	31	prolonged	prolonged	ADJ
gs-3955	78	32	lifespans	lifespan	NOUN
gs-3955	78	33	(	(	PUNCT
gs-3955	78	34	finch	finch	NOUN
gs-3955	78	35	,	,	PUNCT
gs-3955	78	36	2009	2009	NUM
gs-3955	78	37	)	)	PUNCT
gs-3955	78	38	.	.	PUNCT
gs-3955	79	1	this	this	DET
gs-3955	79	2	unique	unique	ADJ
gs-3955	79	3	group	group	NOUN
gs-3955	79	4	of	of	ADP
gs-3955	79	5	organisms	organism	NOUN
gs-3955	79	6	challenges	challenge	VERB
gs-3955	79	7	fundamental	fundamental	ADJ
gs-3955	79	8	assumptions	assumption	NOUN
gs-3955	79	9	about	about	ADP
gs-3955	79	10	aging	age	VERB
gs-3955	79	11	processes	process	NOUN
gs-3955	79	12	and	and	CCONJ
gs-3955	79	13	provides	provide	VERB
gs-3955	79	14	critical	critical	ADJ
gs-3955	79	15	insights	insight	NOUN
gs-3955	79	16	into	into	ADP
gs-3955	79	17	potential	potential	ADJ
gs-3955	79	18	mechanisms	mechanism	NOUN
gs-3955	79	19	for	for	ADP
gs-3955	79	20	extending	extend	VERB
gs-3955	79	21	healthspan	healthspan	NOUN
gs-3955	79	22	(	(	PUNCT
gs-3955	79	23	fig.1	fig.1	PROPN
gs-3955	79	24	)	)	PUNCT
gs-3955	79	25	.	.	PUNCT
gs-3955	80	1	the	the	DET
gs-3955	80	2	spectrum	spectrum	NOUN
gs-3955	80	3	of	of	ADP
gs-3955	80	4	negligibly	negligibly	ADV
gs-3955	80	5	senescing	senesce	VERB
gs-3955	80	6	species	specie	NOUN
gs-3955	80	7	spans	span	NOUN
gs-3955	80	8	diverse	diverse	VERB
gs-3955	80	9	taxonomic	taxonomic	ADJ
gs-3955	80	10	groups	group	NOUN
gs-3955	80	11	(	(	PUNCT
gs-3955	80	12	austad	austad	NOUN
gs-3955	80	13	,	,	PUNCT
gs-3955	80	14	2010	2010	NUM
gs-3955	80	15	)	)	PUNCT
gs-3955	80	16	,	,	PUNCT
gs-3955	80	17	including	include	VERB
gs-3955	80	18	:	:	PUNCT
gs-3955	80	19	vertebrates	vertebrate	NOUN
gs-3955	80	20	:	:	PUNCT
gs-3955	80	21	●	●	NUM
gs-3955	80	22	greenland	greenland	NOUN
gs-3955	80	23	shark	shark	NOUN
gs-3955	80	24	(	(	PUNCT
gs-3955	80	25	somniosus	somniosus	PROPN
gs-3955	80	26	microcephalus	microcephalus	PROPN
gs-3955	80	27	)	)	PUNCT
gs-3955	80	28	exceeds	exceed	VERB
gs-3955	80	29	400	400	NUM
gs-3955	80	30	years	year	NOUN
gs-3955	80	31	●	●	PUNCT
gs-3955	80	32	naked	naked	ADJ
gs-3955	80	33	mole	mole	NOUN
gs-3955	80	34	-	-	PUNCT
gs-3955	80	35	rat	rat	NOUN
gs-3955	80	36	(	(	PUNCT
gs-3955	80	37	heterocephalus	heterocephalus	ADJ
gs-3955	80	38	glaber	glaber	NOUN
gs-3955	80	39	)	)	PUNCT
gs-3955	80	40	surpasses	surpass	VERB
gs-3955	80	41	30	30	NUM
gs-3955	80	42	years	year	NOUN
gs-3955	80	43	(	(	PUNCT
gs-3955	80	44	record	record	NOUN
gs-3955	80	45	for	for	ADP
gs-3955	80	46	rodents	rodent	NOUN
gs-3955	80	47	)	)	PUNCT
gs-3955	80	48	●	●	PUNCT
gs-3955	80	49	olm	olm	NOUN
gs-3955	80	50	(	(	PUNCT
gs-3955	80	51	proteus	proteus	PROPN
gs-3955	80	52	anguinus	anguinus	PROPN
gs-3955	80	53	)	)	PUNCT
gs-3955	80	54	potentially	potentially	ADV
gs-3955	80	55	beyond	beyond	ADP
gs-3955	80	56	100	100	NUM
gs-3955	80	57	years	year	NOUN
gs-3955	80	58	●	●	NUM
gs-3955	80	59	fire	fire	NOUN
gs-3955	80	60	salamander	salamander	NOUN
gs-3955	80	61	(	(	PUNCT
gs-3955	80	62	salamandra	salamandra	NOUN
gs-3955	80	63	salamandra	salamandra	PROPN
gs-3955	80	64	)	)	PUNCT
gs-3955	80	65	over	over	ADP
gs-3955	80	66	50	50	NUM
gs-3955	80	67	years	year	NOUN
gs-3955	80	68	●	●	NUM
gs-3955	80	69	turtle	turtle	NOUN
gs-3955	80	70	species	specie	NOUN
gs-3955	80	71	(	(	PUNCT
gs-3955	80	72	emydoidea	emydoidea	NOUN
gs-3955	80	73	blandingii	blandingii	PROPN
gs-3955	80	74	,	,	PUNCT
gs-3955	80	75	terrapene	terrapene	PROPN
gs-3955	80	76	carolina	carolina	PROPN
gs-3955	80	77	)	)	PUNCT
gs-3955	80	78	80	80	NUM
gs-3955	80	79	-	-	SYM
gs-3955	80	80	150	150	NUM
gs-3955	80	81	years	year	NOUN
gs-3955	80	82	●	●	NUM
gs-3955	80	83	rougheye	rougheye	NOUN
gs-3955	80	84	rockfish	rockfish	NOUN
gs-3955	80	85	(	(	PUNCT
gs-3955	80	86	sebastes	sebaste	NOUN
gs-3955	80	87	aleutianus	aleutianus	NOUN
gs-3955	80	88	)	)	PUNCT
gs-3955	81	1	200	200	NUM
gs-3955	81	2	+	+	NUM
gs-3955	81	3	years	year	NOUN
gs-3955	81	4	invertebrates	invertebrate	NOUN
gs-3955	81	5	:	:	PUNCT
gs-3955	81	6	●	●	NUM
gs-3955	81	7	ocean	ocean	NOUN
gs-3955	81	8	quahog	quahog	NOUN
gs-3955	81	9	(	(	PUNCT
gs-3955	81	10	arctica	arctica	PROPN
gs-3955	81	11	islandica	islandica	PROPN
gs-3955	81	12	)	)	PUNCT
gs-3955	81	13	507	507	NUM
gs-3955	81	14	years	year	NOUN
gs-3955	81	15	(	(	PUNCT
gs-3955	81	16	longest	long	ADV
gs-3955	81	17	-	-	PUNCT
gs-3955	81	18	lived	live	VERB
gs-3955	81	19	animal	animal	NOUN
gs-3955	81	20	)	)	PUNCT
gs-3955	81	21	●	●	PUNCT
gs-3955	81	22	red	red	ADJ
gs-3955	81	23	sea	sea	NOUN
gs-3955	81	24	urchin	urchin	NOUN
gs-3955	81	25	(	(	PUNCT
gs-3955	81	26	strongylocentrotus	strongylocentrotus	NOUN
gs-3955	81	27	franciscanus	franciscanu	NOUN
gs-3955	81	28	)	)	PUNCT
gs-3955	81	29	100	100	NUM
gs-3955	81	30	+	+	NUM
gs-3955	81	31	years	year	NOUN
gs-3955	81	32	●	●	PUNCT
gs-3955	81	33	planarian	planarian	ADJ
gs-3955	81	34	flatworm	flatworm	NOUN
gs-3955	81	35	(	(	PUNCT
gs-3955	81	36	schmidtea	schmidtea	PROPN
gs-3955	81	37	mediterranea	mediterranea	PROPN
gs-3955	81	38	)	)	PUNCT
gs-3955	81	39	theoretically	theoretically	ADV
gs-3955	81	40	immortal	immortal	ADJ
gs-3955	81	41	through	through	ADP
gs-3955	81	42	regeneration	regeneration	NOUN
gs-3955	81	43	these	these	DET
gs-3955	81	44	species	specie	NOUN
gs-3955	81	45	hold	hold	VERB
gs-3955	81	46	profound	profound	ADJ
gs-3955	81	47	significance	significance	NOUN
gs-3955	81	48	for	for	ADP
gs-3955	81	49	biogerontology	biogerontology	NOUN
gs-3955	81	50	as	as	SCONJ
gs-3955	81	51	they	they	PRON
gs-3955	81	52	reveal	reveal	VERB
gs-3955	81	53	alternative	alternative	ADJ
gs-3955	81	54	strategies	strategy	NOUN
gs-3955	81	55	counteracting	counteract	VERB
gs-3955	81	56	cellular	cellular	ADJ
gs-3955	81	57	degeneration	degeneration	NOUN
gs-3955	81	58	(	(	PUNCT
gs-3955	81	59	gorbunova	gorbunova	PROPN
gs-3955	81	60	et	et	PROPN
gs-3955	81	61	al	al	PROPN
gs-3955	81	62	.	.	PROPN
gs-3955	81	63	,	,	PUNCT
gs-3955	81	64	2014	2014	NUM
gs-3955	81	65	)	)	PUNCT
gs-3955	81	66	.	.	PUNCT
gs-3955	82	1	unlike	unlike	ADP
gs-3955	82	2	conventional	conventional	ADJ
gs-3955	82	3	model	model	NOUN
gs-3955	82	4	organisms	organism	NOUN
gs-3955	82	5	(	(	PUNCT
gs-3955	82	6	e.g.	e.g.	ADV
gs-3955	82	7	,	,	PUNCT
gs-3955	82	8	mice	mouse	NOUN
gs-3955	82	9	)	)	PUNCT
gs-3955	82	10	,	,	PUNCT
gs-3955	82	11	negligibly	negligibly	ADV
gs-3955	82	12	sensing	sense	VERB
gs-3955	82	13	species	specie	NOUN
gs-3955	82	14	possess	possess	VERB
gs-3955	82	15	extraordinary	extraordinary	ADJ
gs-3955	82	16	adaptations	adaptation	NOUN
gs-3955	82	17	including	include	VERB
gs-3955	82	18	enhanced	enhance	VERB
gs-3955	82	19	dna	dna	PROPN
gs-3955	82	20	repair	repair	NOUN
gs-3955	82	21	capacity	capacity	NOUN
gs-3955	82	22	,	,	PUNCT
gs-3955	82	23	oxidative	oxidative	ADJ
gs-3955	82	24	stress	stress	NOUN
gs-3955	82	25	resistance	resistance	NOUN
gs-3955	82	26	,	,	PUNCT
gs-3955	82	27	and	and	CCONJ
gs-3955	82	28	tumor	tumor	NOUN
gs-3955	82	29	suppression	suppression	NOUN
gs-3955	82	30	mechanisms	mechanism	NOUN
gs-3955	82	31	(	(	PUNCT
gs-3955	82	32	tian	tian	PROPN
gs-3955	82	33	et	et	PROPN
gs-3955	82	34	al	al	PROPN
gs-3955	82	35	.	.	PROPN
gs-3955	82	36	,	,	PUNCT
gs-3955	82	37	2019	2019	NUM
gs-3955	82	38	)	)	PUNCT
gs-3955	82	39	.	.	PUNCT
gs-3955	83	1	georgian	georgian	ADJ
gs-3955	83	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	83	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	83	4	ტ.	ტ.	VERB
gs-3955	83	5	7	7	NUM
gs-3955	83	6	n	n	DET
gs-3955	83	7	3	3	NUM
gs-3955	83	8	,	,	PUNCT
gs-3955	83	9	2025	2025	NUM
gs-3955	83	10	352	352	NUM
gs-3955	83	11	the	the	DET
gs-3955	83	12	antagonistic	antagonistic	ADJ
gs-3955	83	13	pleiotropy	pleiotropy	NOUN
gs-3955	83	14	theory	theory	NOUN
gs-3955	83	15	posits	posit	NOUN
gs-3955	83	16	aging	age	VERB
gs-3955	83	17	as	as	ADP
gs-3955	83	18	an	an	DET
gs-3955	83	19	evolutionary	evolutionary	ADJ
gs-3955	83	20	trade	trade	NOUN
gs-3955	83	21	-	-	PUNCT
gs-3955	83	22	off	off	NOUN
gs-3955	83	23	between	between	ADP
gs-3955	83	24	earlylife	earlylife	NOUN
gs-3955	83	25	reproduction	reproduction	NOUN
gs-3955	83	26	and	and	CCONJ
gs-3955	83	27	long	long	ADJ
gs-3955	83	28	-	-	PUNCT
gs-3955	83	29	term	term	NOUN
gs-3955	83	30	somatic	somatic	ADJ
gs-3955	83	31	maintenance	maintenance	NOUN
gs-3955	83	32	(	(	PUNCT
gs-3955	83	33	williams	williams	PROPN
gs-3955	83	34	,	,	PUNCT
gs-3955	83	35	1957	1957	NUM
gs-3955	83	36	)	)	PUNCT
gs-3955	83	37	.	.	PUNCT
gs-3955	84	1	however	however	ADV
gs-3955	84	2	,	,	PUNCT
gs-3955	84	3	species	specie	NOUN
gs-3955	84	4	experiencing	experience	VERB
gs-3955	84	5	low	low	ADJ
gs-3955	84	6	extrinsic	extrinsic	ADJ
gs-3955	84	7	mortality	mortality	NOUN
gs-3955	84	8	(	(	PUNCT
gs-3955	84	9	e.g.	e.g.	ADV
gs-3955	84	10	,	,	PUNCT
gs-3955	84	11	deep	deep	ADJ
gs-3955	84	12	-	-	PUNCT
gs-3955	84	13	sea	sea	NOUN
gs-3955	84	14	fish	fish	NOUN
gs-3955	84	15	or	or	CCONJ
gs-3955	84	16	subterranean	subterranean	ADJ
gs-3955	84	17	mammals	mammal	NOUN
gs-3955	84	18	)	)	PUNCT
gs-3955	84	19	exhibit	exhibit	VERB
gs-3955	84	20	evolutionary	evolutionary	ADJ
gs-3955	84	21	shifts	shift	NOUN
gs-3955	84	22	toward	toward	ADP
gs-3955	84	23	lifespan	lifespan	ADJ
gs-3955	84	24	extension	extension	NOUN
gs-3955	84	25	(	(	PUNCT
gs-3955	84	26	austad	austad	NOUN
gs-3955	84	27	&	&	CCONJ
gs-3955	84	28	fischer	fischer	PROPN
gs-3955	84	29	,	,	PUNCT
gs-3955	84	30	1991	1991	NUM
gs-3955	84	31	)	)	PUNCT
gs-3955	84	32	.	.	PUNCT
gs-3955	85	1	the	the	DET
gs-3955	85	2	naked	naked	ADJ
gs-3955	85	3	mole	mole	NOUN
gs-3955	85	4	-	-	PUNCT
gs-3955	85	5	rat	rat	NOUN
gs-3955	85	6	(	(	PUNCT
gs-3955	85	7	heterocephalus	heterocephalus	ADJ
gs-3955	85	8	glaber	glaber	NOUN
gs-3955	85	9	)	)	PUNCT
gs-3955	85	10	,	,	PUNCT
gs-3955	85	11	thriving	thrive	VERB
gs-3955	85	12	in	in	ADP
gs-3955	85	13	protected	protect	VERB
gs-3955	85	14	underground	underground	ADJ
gs-3955	85	15	environments	environment	NOUN
gs-3955	85	16	,	,	PUNCT
gs-3955	85	17	exemplifies	exemplify	VERB
gs-3955	85	18	this	this	DET
gs-3955	85	19	principle	principle	NOUN
gs-3955	85	20	with	with	ADP
gs-3955	85	21	exceptional	exceptional	ADJ
gs-3955	85	22	longevity	longevity	NOUN
gs-3955	85	23	(	(	PUNCT
gs-3955	85	24	37	37	NUM
gs-3955	85	25	+	+	NUM
gs-3955	85	26	years	year	NOUN
gs-3955	85	27	)	)	PUNCT
gs-3955	85	28	and	and	CCONJ
gs-3955	85	29	cancer	cancer	NOUN
gs-3955	85	30	resistance	resistance	NOUN
gs-3955	85	31	(	(	PUNCT
gs-3955	85	32	buffenstein	buffenstein	ADV
gs-3955	85	33	,	,	PUNCT
gs-3955	85	34	2008	2008	NUM
gs-3955	85	35	)	)	PUNCT
gs-3955	85	36	.	.	PUNCT
gs-3955	86	1	these	these	DET
gs-3955	86	2	observations	observation	NOUN
gs-3955	86	3	challenge	challenge	VERB
gs-3955	86	4	traditional	traditional	ADJ
gs-3955	86	5	aging	age	VERB
gs-3955	86	6	theories	theory	NOUN
gs-3955	86	7	and	and	CCONJ
gs-3955	86	8	align	align	VERB
gs-3955	86	9	with	with	ADP
gs-3955	86	10	the	the	DET
gs-3955	86	11	centrosomal	centrosomal	ADJ
gs-3955	86	12	theory	theory	NOUN
gs-3955	86	13	of	of	ADP
gs-3955	86	14	aging	age	VERB
gs-3955	86	15	(	(	PUNCT
gs-3955	86	16	tkemaladze	tkemaladze	NOUN
gs-3955	86	17	,	,	PUNCT
gs-3955	86	18	2024	2024	NUM
gs-3955	86	19	)	)	PUNCT
gs-3955	86	20	,	,	PUNCT
gs-3955	86	21	which	which	PRON
gs-3955	86	22	identifies	identify	VERB
gs-3955	86	23	aging	aging	NOUN
gs-3955	86	24	as	as	ADP
gs-3955	86	25	a	a	DET
gs-3955	86	26	byproduct	byproduct	NOUN
gs-3955	86	27	of	of	ADP
gs-3955	86	28	cellular	cellular	ADJ
gs-3955	86	29	differentiation	differentiation	NOUN
gs-3955	86	30	where	where	SCONJ
gs-3955	86	31	asymmetric	asymmetric	ADJ
gs-3955	86	32	stem	stem	NOUN
gs-3955	86	33	cell	cell	NOUN
gs-3955	86	34	divisions	division	NOUN
gs-3955	86	35	accumulate	accumulate	VERB
gs-3955	86	36	aged	aged	ADJ
gs-3955	86	37	centrosomes	centrosome	NOUN
gs-3955	86	38	(	(	PUNCT
gs-3955	86	39	tkemaladze	tkemaladze	NOUN
gs-3955	86	40	,	,	PUNCT
gs-3955	86	41	2023	2023	NUM
gs-3955	86	42	)	)	PUNCT
gs-3955	86	43	.	.	PUNCT
gs-3955	87	1	this	this	DET
gs-3955	87	2	thermodynamic	thermodynamic	ADJ
gs-3955	87	3	perspective	perspective	NOUN
gs-3955	87	4	not	not	PART
gs-3955	87	5	only	only	ADV
gs-3955	87	6	accommodates	accommodate	NOUN
gs-3955	87	7	but	but	CCONJ
gs-3955	87	8	explains	explain	VERB
gs-3955	87	9	the	the	DET
gs-3955	87	10	existence	existence	NOUN
gs-3955	87	11	of	of	ADP
gs-3955	87	12	negligibly	negligibly	ADV
gs-3955	87	13	senescing	senesce	VERB
gs-3955	87	14	species	specie	NOUN
gs-3955	87	15	as	as	ADP
gs-3955	87	16	exceptions	exception	NOUN
gs-3955	87	17	to	to	PART
gs-3955	87	18	entropic	entropic	VERB
gs-3955	87	19	decay	decay	NOUN
gs-3955	87	20	.	.	PUNCT
gs-3955	88	1	the	the	DET
gs-3955	88	2	greenland	greenland	PROPN
gs-3955	88	3	shark	shark	NOUN
gs-3955	88	4	(	(	PUNCT
gs-3955	88	5	somniosus	somniosus	PROPN
gs-3955	88	6	microcephalus	microcephalus	PROPN
gs-3955	88	7	)	)	PUNCT
gs-3955	88	8	achieves	achieve	VERB
gs-3955	88	9	400	400	NUM
gs-3955	88	10	+	+	NUM
gs-3955	88	11	year	year	NOUN
gs-3955	88	12	lifespans	lifespan	NOUN
gs-3955	88	13	through	through	ADP
gs-3955	88	14	metabolic	metabolic	NOUN
gs-3955	88	15	depression	depression	NOUN
gs-3955	88	16	and	and	CCONJ
gs-3955	88	17	macromolecular	macromolecular	NOUN
gs-3955	88	18	stability	stability	NOUN
gs-3955	88	19	(	(	PUNCT
gs-3955	88	20	nielsen	nielsen	PROPN
gs-3955	88	21	et	et	PROPN
gs-3955	88	22	al	al	PROPN
gs-3955	88	23	.	.	PROPN
gs-3955	88	24	,	,	PUNCT
gs-3955	88	25	2016	2016	NUM
gs-3955	88	26	)	)	PUNCT
gs-3955	88	27	.	.	PUNCT
gs-3955	89	1	these	these	DET
gs-3955	89	2	cases	case	NOUN
gs-3955	89	3	support	support	VERB
gs-3955	89	4	the	the	DET
gs-3955	89	5	hypothesis	hypothesis	NOUN
gs-3955	89	6	that	that	PRON
gs-3955	89	7	extreme	extreme	ADJ
gs-3955	89	8	longevity	longevity	NOUN
gs-3955	89	9	represents	represent	VERB
gs-3955	89	10	an	an	DET
gs-3955	89	11	adaptation	adaptation	NOUN
gs-3955	89	12	to	to	ADP
gs-3955	89	13	stable	stable	ADJ
gs-3955	89	14	,	,	PUNCT
gs-3955	89	15	low	low	ADJ
gs-3955	89	16	-	-	PUNCT
gs-3955	89	17	stress	stress	NOUN
gs-3955	89	18	environments	environment	NOUN
gs-3955	89	19	through	through	ADP
gs-3955	89	20	individual	individual	ADJ
gs-3955	89	21	survival	survival	NOUN
gs-3955	89	22	strategies	strategy	NOUN
gs-3955	89	23	rather	rather	ADV
gs-3955	89	24	than	than	ADP
gs-3955	89	25	species	species	NOUN
gs-3955	89	26	-	-	PUNCT
gs-3955	89	27	level	level	NOUN
gs-3955	89	28	adaptations	adaptation	NOUN
gs-3955	89	29	(	(	PUNCT
gs-3955	89	30	de	de	X
gs-3955	89	31	magalhães	magalhães	NOUN
gs-3955	89	32	et	et	PROPN
gs-3955	89	33	al	al	PROPN
gs-3955	89	34	.	.	PROPN
gs-3955	89	35	,	,	PUNCT
gs-3955	89	36	2007	2007	NUM
gs-3955	89	37	)	)	PUNCT
gs-3955	89	38	.	.	PUNCT
gs-3955	90	1	contemporary	contemporary	ADJ
gs-3955	90	2	research	research	NOUN
gs-3955	90	3	has	have	AUX
gs-3955	90	4	identified	identify	VERB
gs-3955	90	5	conserved	conserve	VERB
gs-3955	90	6	molecular	molecular	ADJ
gs-3955	90	7	pathways	pathway	NOUN
gs-3955	90	8	associated	associate	VERB
gs-3955	90	9	with	with	ADP
gs-3955	90	10	negligible	negligible	ADJ
gs-3955	90	11	senescence	senescence	NOUN
gs-3955	90	12	that	that	PRON
gs-3955	90	13	actively	actively	ADV
gs-3955	90	14	counteract	counteract	VERB
gs-3955	90	15	entropic	entropic	ADJ
gs-3955	90	16	degeneration	degeneration	NOUN
gs-3955	90	17	:	:	PUNCT
gs-3955	90	18	dna	dna	PROPN
gs-3955	90	19	repair	repair	NOUN
gs-3955	90	20	systems	system	NOUN
gs-3955	90	21	long	long	ADV
gs-3955	90	22	-	-	PUNCT
gs-3955	90	23	lived	live	VERB
gs-3955	90	24	species	specie	NOUN
gs-3955	90	25	like	like	ADP
gs-3955	90	26	arctica	arctica	PROPN
gs-3955	90	27	islandica	islandica	PROPN
gs-3955	90	28	(	(	PUNCT
gs-3955	90	29	>	>	SYM
gs-3955	90	30	500	500	NUM
gs-3955	90	31	years	year	NOUN
gs-3955	90	32	)	)	PUNCT
gs-3955	90	33	demonstrate	demonstrate	VERB
gs-3955	90	34	enhanced	enhanced	ADJ
gs-3955	90	35	activity	activity	NOUN
gs-3955	90	36	of	of	ADP
gs-3955	90	37	dna	dna	PROPN
gs-3955	90	38	repair	repair	NOUN
gs-3955	90	39	enzymes	enzyme	NOUN
gs-3955	90	40	including	include	VERB
gs-3955	90	41	parp1	parp1	NOUN
gs-3955	90	42	and	and	CCONJ
gs-3955	90	43	nucleotide	nucleotide	ADJ
gs-3955	90	44	excision	excision	NOUN
gs-3955	90	45	repair	repair	NOUN
gs-3955	90	46	components	component	NOUN
gs-3955	90	47	(	(	PUNCT
gs-3955	90	48	ungvari	ungvari	X
gs-3955	90	49	et	et	PROPN
gs-3955	90	50	al	al	PROPN
gs-3955	90	51	.	.	PROPN
gs-3955	90	52	,	,	PUNCT
gs-3955	90	53	2013	2013	NUM
gs-3955	90	54	)	)	PUNCT
gs-3955	90	55	.	.	PUNCT
gs-3955	91	1	oxidative	oxidative	ADJ
gs-3955	91	2	defense	defense	NOUN
gs-3955	91	3	mechanisms	mechanism	NOUN
gs-3955	91	4	naked	naked	ADJ
gs-3955	91	5	mole	mole	NOUN
gs-3955	91	6	-	-	PUNCT
gs-3955	91	7	rats	rat	NOUN
gs-3955	91	8	maintain	maintain	VERB
gs-3955	91	9	exceptionally	exceptionally	ADV
gs-3955	91	10	low	low	ADJ
gs-3955	91	11	oxidative	oxidative	ADJ
gs-3955	91	12	damage	damage	NOUN
gs-3955	91	13	despite	despite	SCONJ
gs-3955	91	14	high	high	ADJ
gs-3955	91	15	ros	ros	PROPN
gs-3955	91	16	production	production	NOUN
gs-3955	91	17	(	(	PUNCT
gs-3955	91	18	lewis	lewis	PROPN
gs-3955	91	19	et	et	PROPN
gs-3955	91	20	al	al	PROPN
gs-3955	91	21	.	.	PROPN
gs-3955	91	22	,	,	PUNCT
gs-3955	91	23	2018	2018	NUM
gs-3955	91	24	)	)	PUNCT
gs-3955	91	25	,	,	PUNCT
gs-3955	91	26	attributed	attribute	VERB
gs-3955	91	27	to	to	ADP
gs-3955	91	28	unique	unique	ADJ
gs-3955	91	29	mitochondrial	mitochondrial	ADJ
gs-3955	91	30	membrane	membrane	NOUN
gs-3955	91	31	properties	property	NOUN
gs-3955	91	32	and	and	CCONJ
gs-3955	91	33	elevated	elevated	ADJ
gs-3955	91	34	superoxide	superoxide	NOUN
gs-3955	91	35	dismutase	dismutase	NOUN
gs-3955	91	36	(	(	PUNCT
gs-3955	91	37	sod2	sod2	NOUN
gs-3955	91	38	)	)	PUNCT
gs-3955	91	39	expression	expression	NOUN
gs-3955	91	40	(	(	PUNCT
gs-3955	91	41	azpurua	azpurua	PROPN
gs-3955	91	42	et	et	PROPN
gs-3955	91	43	al	al	PROPN
gs-3955	91	44	.	.	PROPN
gs-3955	91	45	,	,	PUNCT
gs-3955	91	46	2013	2013	NUM
gs-3955	91	47	)	)	PUNCT
gs-3955	91	48	.	.	PUNCT
gs-3955	92	1	proteostasis	proteostasis	NOUN
gs-3955	92	2	maintenance	maintenance	NOUN
gs-3955	92	3	long	long	ADV
gs-3955	92	4	-	-	PUNCT
gs-3955	92	5	lived	live	VERB
gs-3955	92	6	fish	fish	NOUN
gs-3955	92	7	(	(	PUNCT
gs-3955	92	8	sebastes	sebaste	NOUN
gs-3955	92	9	aleutianus	aleutianus	NOUN
gs-3955	92	10	)	)	PUNCT
gs-3955	92	11	exhibit	exhibit	NOUN
gs-3955	92	12	amplified	amplify	VERB
gs-3955	92	13	autophagy	autophagy	ADJ
gs-3955	92	14	pathways	pathway	NOUN
gs-3955	92	15	preventing	prevent	VERB
gs-3955	92	16	damaged	damage	VERB
gs-3955	92	17	protein	protein	NOUN
gs-3955	92	18	accumulation	accumulation	NOUN
gs-3955	92	19	(	(	PUNCT
gs-3955	92	20	treaster	treaster	NOUN
gs-3955	92	21	et	et	PROPN
gs-3955	92	22	al	al	PROPN
gs-3955	92	23	.	.	PROPN
gs-3955	92	24	,	,	PUNCT
gs-3955	92	25	2014	2014	NUM
gs-3955	92	26	)	)	PUNCT
gs-3955	92	27	.	.	PUNCT
gs-3955	93	1	georgian	georgian	ADJ
gs-3955	93	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	93	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	93	4	ტ.	ტ.	VERB
gs-3955	93	5	7	7	NUM
gs-3955	93	6	n	n	PRON
gs-3955	93	7	3	3	NUM
gs-3955	93	8	,	,	PUNCT
gs-3955	93	9	2025	2025	NUM
gs-3955	93	10	353	353	NUM
gs-3955	93	11	neoplastic	neoplastic	ADJ
gs-3955	93	12	suppression	suppression	NOUN
gs-3955	93	13	naked	naked	ADJ
gs-3955	93	14	mole	mole	NOUN
gs-3955	93	15	-	-	PUNCT
gs-3955	93	16	rats	rat	NOUN
gs-3955	93	17	employ	employ	VERB
gs-3955	93	18	high	high	ADJ
gs-3955	93	19	-	-	PUNCT
gs-3955	93	20	molecular	molecular	ADJ
gs-3955	93	21	-	-	PUNCT
gs-3955	93	22	weight	weight	NOUN
gs-3955	93	23	hyaluronan	hyaluronan	NOUN
gs-3955	93	24	(	(	PUNCT
gs-3955	93	25	hmw	hmw	PROPN
gs-3955	93	26	-	-	PUNCT
gs-3955	93	27	ha	ha	INTJ
gs-3955	93	28	)	)	PUNCT
gs-3955	93	29	mediated	mediate	VERB
gs-3955	93	30	contact	contact	NOUN
gs-3955	93	31	inhibition	inhibition	NOUN
gs-3955	93	32	as	as	ADP
gs-3955	93	33	a	a	DET
gs-3955	93	34	tumor	tumor	NOUN
gs-3955	93	35	suppression	suppression	NOUN
gs-3955	93	36	mechanism	mechanism	NOUN
gs-3955	93	37	(	(	PUNCT
gs-3955	93	38	tian	tian	PROPN
gs-3955	93	39	et	et	PROPN
gs-3955	93	40	al	al	PROPN
gs-3955	93	41	.	.	PROPN
gs-3955	93	42	,	,	PUNCT
gs-3955	93	43	2013	2013	NUM
gs-3955	93	44	)	)	PUNCT
gs-3955	93	45	.	.	PUNCT
gs-3955	94	1	comparative	comparative	ADJ
gs-3955	94	2	analyses	analysis	NOUN
gs-3955	94	3	reveal	reveal	VERB
gs-3955	94	4	shared	share	VERB
gs-3955	94	5	characteristics	characteristic	NOUN
gs-3955	94	6	among	among	ADP
gs-3955	94	7	negligibly	negligibly	ADV
gs-3955	94	8	senescing	senesce	VERB
gs-3955	94	9	species	specie	NOUN
gs-3955	94	10	:	:	PUNCT
gs-3955	94	11	●	●	PUNCT
gs-3955	94	12	depressed	depressed	ADJ
gs-3955	94	13	metabolic	metabolic	NOUN
gs-3955	94	14	rates	rate	NOUN
gs-3955	94	15	(	(	PUNCT
gs-3955	94	16	sharks	shark	NOUN
gs-3955	94	17	and	and	CCONJ
gs-3955	94	18	mollusks	mollusk	NOUN
gs-3955	94	19	)	)	PUNCT
gs-3955	94	20	(	(	PUNCT
gs-3955	94	21	munro	munro	PROPN
gs-3955	94	22	&	&	CCONJ
gs-3955	94	23	blier	blier	PROPN
gs-3955	94	24	,	,	PUNCT
gs-3955	94	25	2012	2012	NUM
gs-3955	94	26	)	)	PUNCT
gs-3955	95	1	●	●	NUM
gs-3955	95	2	hypoxia	hypoxia	NOUN
gs-3955	95	3	tolerance	tolerance	NOUN
gs-3955	95	4	(	(	PUNCT
gs-3955	95	5	naked	naked	ADJ
gs-3955	95	6	mole	mole	NOUN
gs-3955	95	7	-	-	PUNCT
gs-3955	95	8	rats	rat	NOUN
gs-3955	95	9	)	)	PUNCT
gs-3955	95	10	(	(	PUNCT
gs-3955	95	11	park	park	NOUN
gs-3955	95	12	et	et	PROPN
gs-3955	95	13	al	al	PROPN
gs-3955	95	14	.	.	PROPN
gs-3955	95	15	,	,	PUNCT
gs-3955	95	16	2017	2017	NUM
gs-3955	95	17	)	)	PUNCT
gs-3955	95	18	●	●	NUM
gs-3955	95	19	epigenetic	epigenetic	ADJ
gs-3955	95	20	stability	stability	NOUN
gs-3955	95	21	(	(	PUNCT
gs-3955	95	22	slow	slow	ADJ
gs-3955	95	23	dna	dna	PROPN
gs-3955	95	24	methylation	methylation	NOUN
gs-3955	95	25	drift	drift	NOUN
gs-3955	95	26	)	)	PUNCT
gs-3955	95	27	(	(	PUNCT
gs-3955	95	28	ma	ma	PROPN
gs-3955	95	29	et	et	PROPN
gs-3955	95	30	al	al	PROPN
gs-3955	95	31	.	.	PROPN
gs-3955	95	32	,	,	PUNCT
gs-3955	95	33	2020	2020	NUM
gs-3955	95	34	)	)	PUNCT
gs-3955	95	35	these	these	DET
gs-3955	95	36	species	specie	NOUN
gs-3955	95	37	serve	serve	VERB
gs-3955	95	38	as	as	ADP
gs-3955	95	39	unparalleled	unparalleled	ADJ
gs-3955	95	40	natural	natural	ADJ
gs-3955	95	41	models	model	NOUN
gs-3955	95	42	for	for	ADP
gs-3955	95	43	longevity	longevity	NOUN
gs-3955	95	44	research	research	NOUN
gs-3955	95	45	,	,	PUNCT
gs-3955	95	46	offering	offer	VERB
gs-3955	95	47	transformative	transformative	ADJ
gs-3955	95	48	potential	potential	NOUN
gs-3955	95	49	for	for	ADP
gs-3955	95	50	treating	treat	VERB
gs-3955	95	51	age	age	NOUN
gs-3955	95	52	-	-	PUNCT
gs-3955	95	53	related	relate	VERB
gs-3955	95	54	diseases	disease	NOUN
gs-3955	95	55	and	and	CCONJ
gs-3955	95	56	developing	develop	VERB
gs-3955	95	57	anti	anti	ADJ
gs-3955	95	58	-	-	ADJ
gs-3955	95	59	aging	aging	ADJ
gs-3955	95	60	interventions	intervention	NOUN
gs-3955	95	61	(	(	PUNCT
gs-3955	95	62	seluanov	seluanov	PROPN
gs-3955	95	63	et	et	PROPN
gs-3955	95	64	al	al	PROPN
gs-3955	95	65	.	.	PROPN
gs-3955	95	66	,	,	PUNCT
gs-3955	95	67	2018	2018	NUM
gs-3955	95	68	)	)	PUNCT
gs-3955	95	69	.	.	PUNCT
gs-3955	96	1	the	the	DET
gs-3955	96	2	greenland	greenland	PROPN
gs-3955	96	3	shark	shark	NOUN
gs-3955	96	4	(	(	PUNCT
gs-3955	96	5	somniosus	somniosus	PROPN
gs-3955	96	6	microcephalus	microcephalus	PROPN
gs-3955	96	7	):	):	PUNCT
gs-3955	96	8	a	a	DET
gs-3955	96	9	paradigm	paradigm	NOUN
gs-3955	96	10	of	of	ADP
gs-3955	96	11	extreme	extreme	ADJ
gs-3955	96	12	longevity	longevity	NOUN
gs-3955	96	13	as	as	ADP
gs-3955	96	14	the	the	DET
gs-3955	96	15	longest	long	ADV
gs-3955	96	16	-	-	PUNCT
gs-3955	96	17	lived	live	VERB
gs-3955	96	18	vertebrate	vertebrate	NOUN
gs-3955	96	19	known	know	VERB
gs-3955	96	20	to	to	ADP
gs-3955	96	21	science	science	NOUN
gs-3955	96	22	,	,	PUNCT
gs-3955	96	23	the	the	DET
gs-3955	96	24	greenland	greenland	NOUN
gs-3955	96	25	shark	shark	NOUN
gs-3955	96	26	(	(	PUNCT
gs-3955	96	27	somniosus	somniosus	PROPN
gs-3955	96	28	microcephalus	microcephalus	PROPN
gs-3955	96	29	)	)	PUNCT
gs-3955	96	30	reaches	reach	VERB
gs-3955	96	31	minimum	minimum	ADJ
gs-3955	96	32	ages	age	NOUN
gs-3955	96	33	of	of	ADP
gs-3955	96	34	392	392	NUM
gs-3955	96	35	years	year	NOUN
gs-3955	96	36	,	,	PUNCT
gs-3955	96	37	with	with	ADP
gs-3955	96	38	some	some	DET
gs-3955	96	39	individuals	individual	NOUN
gs-3955	96	40	exceeding	exceed	VERB
gs-3955	96	41	400	400	NUM
gs-3955	96	42	years	year	NOUN
gs-3955	96	43	based	base	VERB
gs-3955	96	44	on	on	ADP
gs-3955	96	45	radiocarbon	radiocarbon	NOUN
gs-3955	96	46	dating	dating	NOUN
gs-3955	96	47	of	of	ADP
gs-3955	96	48	eye	eye	NOUN
gs-3955	96	49	lens	lens	NOUN
gs-3955	96	50	nuclei	nucleus	NOUN
gs-3955	96	51	(	(	PUNCT
gs-3955	96	52	nielsen	nielsen	PROPN
gs-3955	96	53	et	et	PROPN
gs-3955	96	54	al	al	PROPN
gs-3955	96	55	.	.	PROPN
gs-3955	96	56	,	,	PUNCT
gs-3955	96	57	2016	2016	NUM
gs-3955	96	58	;	;	PUNCT
gs-3955	96	59	hansen	hansen	PROPN
gs-3955	96	60	et	et	PROPN
gs-3955	96	61	al	al	PROPN
gs-3955	96	62	.	.	PROPN
gs-3955	96	63	,	,	PUNCT
gs-3955	96	64	2021	2021	NUM
gs-3955	96	65	)	)	PUNCT
gs-3955	96	66	.	.	PUNCT
gs-3955	97	1	this	this	DET
gs-3955	97	2	arctic	arctic	ADJ
gs-3955	97	3	species	specie	NOUN
gs-3955	97	4	has	have	AUX
gs-3955	97	5	become	become	VERB
gs-3955	97	6	a	a	DET
gs-3955	97	7	cornerstone	cornerstone	NOUN
gs-3955	97	8	model	model	NOUN
gs-3955	97	9	for	for	ADP
gs-3955	97	10	studying	study	VERB
gs-3955	97	11	decelerated	decelerate	VERB
gs-3955	97	12	aging	age	VERB
gs-3955	97	13	processes	process	NOUN
gs-3955	97	14	in	in	ADP
gs-3955	97	15	marine	marine	ADJ
gs-3955	97	16	vertebrates	vertebrate	NOUN
gs-3955	97	17	.	.	PUNCT
gs-3955	98	1	several	several	ADJ
gs-3955	98	2	physiological	physiological	ADJ
gs-3955	98	3	adaptations	adaptation	NOUN
gs-3955	98	4	contribute	contribute	VERB
gs-3955	98	5	to	to	ADP
gs-3955	98	6	this	this	DET
gs-3955	98	7	exceptional	exceptional	ADJ
gs-3955	98	8	longevity	longevity	NOUN
gs-3955	98	9	:	:	PUNCT
gs-3955	98	10	inhabiting	inhabit	VERB
gs-3955	98	11	frigid	frigid	ADJ
gs-3955	98	12	arctic	arctic	ADJ
gs-3955	98	13	waters	water	NOUN
gs-3955	98	14	(	(	PUNCT
gs-3955	98	15	-1	-1	ADP
gs-3955	98	16	to	to	PART
gs-3955	98	17	+10	+10	VERB
gs-3955	98	18	°	°	NOUN
gs-3955	98	19	c	c	NOUN
gs-3955	98	20	)	)	PUNCT
gs-3955	98	21	,	,	PUNCT
gs-3955	98	22	greenland	greenland	PROPN
gs-3955	98	23	sharks	sharks	PROPN
gs-3955	98	24	exhibit	exhibit	VERB
gs-3955	98	25	extraordinarily	extraordinarily	ADV
gs-3955	98	26	slow	slow	ADJ
gs-3955	98	27	metabolic	metabolic	NOUN
gs-3955	98	28	rates	rate	NOUN
gs-3955	98	29	(	(	PUNCT
gs-3955	98	30	treberg	treberg	PROPN
gs-3955	98	31	&	&	CCONJ
gs-3955	98	32	speers	speer	NOUN
gs-3955	98	33	-	-	PUNCT
gs-3955	98	34	roesch	roesch	NOUN
gs-3955	98	35	,	,	PUNCT
gs-3955	98	36	2016	2016	NUM
gs-3955	98	37	)	)	PUNCT
gs-3955	98	38	.	.	PUNCT
gs-3955	99	1	their	their	PRON
gs-3955	99	2	growth	growth	NOUN
gs-3955	99	3	velocity	velocity	NOUN
gs-3955	99	4	(	(	PUNCT
gs-3955	99	5	≈1	≈1	PROPN
gs-3955	99	6	cm	cm	NOUN
gs-3955	99	7	/	/	SYM
gs-3955	99	8	year	year	NOUN
gs-3955	99	9	)	)	PUNCT
gs-3955	99	10	and	and	CCONJ
gs-3955	99	11	late	late	ADJ
gs-3955	99	12	sexual	sexual	ADJ
gs-3955	99	13	maturation	maturation	NOUN
gs-3955	99	14	(	(	PUNCT
gs-3955	99	15	≈150	≈150	NOUN
gs-3955	99	16	years	year	NOUN
gs-3955	99	17	)	)	PUNCT
gs-3955	99	18	(	(	PUNCT
gs-3955	99	19	nielsen	nielsen	PROPN
gs-3955	99	20	et	et	PROPN
gs-3955	99	21	al	al	PROPN
gs-3955	99	22	.	.	PROPN
gs-3955	99	23	,	,	PUNCT
gs-3955	99	24	2016	2016	NUM
gs-3955	99	25	)	)	PUNCT
gs-3955	99	26	minimize	minimize	VERB
gs-3955	99	27	cumulative	cumulative	ADJ
gs-3955	99	28	molecular	molecular	ADJ
gs-3955	99	29	damage	damage	NOUN
gs-3955	99	30	from	from	ADP
gs-3955	99	31	oxidative	oxidative	ADJ
gs-3955	99	32	processes	process	NOUN
gs-3955	99	33	(	(	PUNCT
gs-3955	99	34	munro	munro	PROPN
gs-3955	99	35	et	et	PROPN
gs-3955	99	36	al	al	PROPN
gs-3955	99	37	.	.	PROPN
gs-3955	99	38	,	,	PUNCT
gs-3955	99	39	2019	2019	NUM
gs-3955	99	40	)	)	PUNCT
gs-3955	99	41	.	.	PUNCT
gs-3955	100	1	despite	despite	SCONJ
gs-3955	100	2	low	low	ADJ
gs-3955	100	3	antioxidant	antioxidant	ADJ
gs-3955	100	4	enzyme	enzyme	NOUN
gs-3955	100	5	activity	activity	NOUN
gs-3955	100	6	,	,	PUNCT
gs-3955	100	7	these	these	DET
gs-3955	100	8	sharks	shark	NOUN
gs-3955	100	9	show	show	VERB
gs-3955	100	10	minimal	minimal	ADJ
gs-3955	100	11	oxidative	oxidative	ADJ
gs-3955	100	12	damage	damage	NOUN
gs-3955	100	13	(	(	PUNCT
gs-3955	100	14	passow	passow	NOUN
gs-3955	100	15	et	et	PROPN
gs-3955	100	16	al	al	PROPN
gs-3955	100	17	.	.	PROPN
gs-3955	100	18	,	,	PUNCT
gs-3955	100	19	2022	2022	NUM
gs-3955	100	20	)	)	PUNCT
gs-3955	100	21	through	through	ADP
gs-3955	100	22	:	:	PUNCT
gs-3955	100	23	●	●	NUM
gs-3955	100	24	specialized	specialized	ADJ
gs-3955	100	25	membrane	membrane	NOUN
gs-3955	100	26	lipid	lipid	NOUN
gs-3955	100	27	composition	composition	NOUN
gs-3955	100	28	resisting	resist	VERB
gs-3955	100	29	peroxidation	peroxidation	NOUN
gs-3955	100	30	(	(	PUNCT
gs-3955	100	31	ballantyne	ballantyne	NOUN
gs-3955	100	32	et	et	PROPN
gs-3955	100	33	al	al	PROPN
gs-3955	100	34	.	.	PROPN
gs-3955	100	35	,	,	PUNCT
gs-3955	100	36	2021	2021	NUM
gs-3955	100	37	)	)	PUNCT
gs-3955	100	38	●	●	NUM
gs-3955	100	39	elevated	elevate	VERB
gs-3955	100	40	chaperone	chaperone	NOUN
gs-3955	100	41	proteins	protein	NOUN
gs-3955	100	42	(	(	PUNCT
gs-3955	100	43	hsp70	hsp70	NOUN
gs-3955	100	44	,	,	PUNCT
gs-3955	100	45	hsp90	hsp90	VERB
gs-3955	100	46	)	)	PUNCT
gs-3955	100	47	maintaining	maintain	VERB
gs-3955	100	48	proteostasis	proteostasis	NOUN
gs-3955	100	49	(	(	PUNCT
gs-3955	100	50	gorbunova	gorbunova	PROPN
gs-3955	100	51	et	et	PROPN
gs-3955	100	52	al	al	PROPN
gs-3955	100	53	.	.	PROPN
gs-3955	100	54	,	,	PUNCT
gs-3955	100	55	2020	2020	NUM
gs-3955	100	56	)	)	PUNCT
gs-3955	100	57	genome	genome	NOUN
gs-3955	100	58	sequencing	sequencing	NOUN
gs-3955	100	59	reveals	reveal	NOUN
gs-3955	100	60	:	:	PUNCT
gs-3955	100	61	●	●	NUM
gs-3955	100	62	mutations	mutation	NOUN
gs-3955	100	63	enhancing	enhance	VERB
gs-3955	100	64	dna	dna	PROPN
gs-3955	100	65	repair	repair	NOUN
gs-3955	100	66	(	(	PUNCT
gs-3955	100	67	ercc1	ercc1	PROPN
gs-3955	100	68	,	,	PUNCT
gs-3955	100	69	ercc2	ercc2	NOUN
gs-3955	100	70	)	)	PUNCT
gs-3955	100	71	(	(	PUNCT
gs-3955	101	1	seim	seim	PROPN
gs-3955	101	2	et	et	PROPN
gs-3955	101	3	al	al	PROPN
gs-3955	101	4	.	.	PROPN
gs-3955	101	5	,	,	PUNCT
gs-3955	101	6	2023	2023	NUM
gs-3955	101	7	)	)	PUNCT
gs-3955	101	8	●	●	NUM
gs-3955	101	9	telomere	telomere	NOUN
gs-3955	101	10	maintenance	maintenance	NOUN
gs-3955	101	11	gene	gene	NOUN
gs-3955	101	12	expression	expression	NOUN
gs-3955	101	13	patterns	pattern	NOUN
gs-3955	101	14	(	(	PUNCT
gs-3955	101	15	de	de	X
gs-3955	101	16	magalhães	magalhães	NOUN
gs-3955	101	17	et	et	PROPN
gs-3955	101	18	al	al	PROPN
gs-3955	101	19	.	.	PROPN
gs-3955	101	20	,	,	PUNCT
gs-3955	101	21	2017	2017	NUM
gs-3955	101	22	)	)	PUNCT
gs-3955	101	23	this	this	DET
gs-3955	101	24	species	specie	NOUN
gs-3955	101	25	exemplifies	exemplify	VERB
gs-3955	101	26	pure	pure	ADJ
gs-3955	101	27	k	k	NOUN
gs-3955	101	28	-	-	NOUN
gs-3955	101	29	selection	selection	NOUN
gs-3955	101	30	:	:	PUNCT
gs-3955	101	31	●	●	NUM
gs-3955	101	32	delayed	delay	VERB
gs-3955	101	33	maturation	maturation	NOUN
gs-3955	101	34	(	(	PUNCT
gs-3955	101	35	150	150	NUM
gs-3955	101	36	+	+	NUM
gs-3955	101	37	years	year	NOUN
gs-3955	101	38	)	)	PUNCT
gs-3955	101	39	offset	offset	VERB
gs-3955	101	40	by	by	ADP
gs-3955	101	41	low	low	ADJ
gs-3955	101	42	adult	adult	NOUN
gs-3955	101	43	mortality	mortality	NOUN
gs-3955	101	44	(	(	PUNCT
gs-3955	101	45	kyne	kyne	PROPN
gs-3955	101	46	et	et	PROPN
gs-3955	101	47	al	al	PROPN
gs-3955	101	48	.	.	PROPN
gs-3955	101	49	,	,	PUNCT
gs-3955	101	50	2019	2019	NUM
gs-3955	101	51	)	)	PUNCT
gs-3955	101	52	●	●	PUNCT
gs-3955	101	53	minimal	minimal	ADJ
gs-3955	101	54	predation	predation	NOUN
gs-3955	101	55	pressure	pressure	NOUN
gs-3955	101	56	reducing	reduce	VERB
gs-3955	101	57	extrinsic	extrinsic	ADJ
gs-3955	101	58	mortality	mortality	NOUN
gs-3955	101	59	(	(	PUNCT
gs-3955	101	60	austad	austad	NOUN
gs-3955	101	61	,	,	PUNCT
gs-3955	101	62	2022	2022	NUM
gs-3955	101	63	)	)	PUNCT
gs-3955	101	64	georgian	georgian	ADJ
gs-3955	101	65	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	101	66	მეცნიერები	მეცნიერები	NOUN
gs-3955	101	67	ტ.	ტ.	VERB
gs-3955	101	68	7	7	NUM
gs-3955	101	69	n	n	DET
gs-3955	101	70	3	3	NUM
gs-3955	101	71	,	,	PUNCT
gs-3955	101	72	2025	2025	NUM
gs-3955	101	73	354	354	NUM
gs-3955	101	74	potential	potential	ADJ
gs-3955	101	75	biomedical	biomedical	ADJ
gs-3955	101	76	applications	application	NOUN
gs-3955	101	77	include	include	VERB
gs-3955	101	78	:	:	PUNCT
gs-3955	101	79	●	●	PUNCT
gs-3955	101	80	cryometabolism	cryometabolism	NOUN
gs-3955	101	81	-	-	PUNCT
gs-3955	101	82	inspired	inspire	VERB
gs-3955	101	83	anti	anti	ADJ
gs-3955	101	84	-	-	ADJ
gs-3955	101	85	aging	aging	ADJ
gs-3955	101	86	strategies	strategy	NOUN
gs-3955	101	87	(	(	PUNCT
gs-3955	101	88	smith	smith	PROPN
gs-3955	101	89	et	et	PROPN
gs-3955	101	90	al	al	PROPN
gs-3955	101	91	.	.	PROPN
gs-3955	101	92	,	,	PUNCT
gs-3955	101	93	2021	2021	NUM
gs-3955	101	94	)	)	PUNCT
gs-3955	101	95	●	●	NUM
gs-3955	101	96	membrane	membrane	NOUN
gs-3955	101	97	lipid	lipid	NOUN
gs-3955	101	98	-	-	PUNCT
gs-3955	101	99	based	base	VERB
gs-3955	101	100	cytoprotective	cytoprotective	ADJ
gs-3955	101	101	approaches	approach	NOUN
gs-3955	101	102	(	(	PUNCT
gs-3955	101	103	tian	tian	PROPN
gs-3955	101	104	et	et	PROPN
gs-3955	101	105	al	al	PROPN
gs-3955	101	106	.	.	PROPN
gs-3955	101	107	,	,	PUNCT
gs-3955	101	108	2022	2022	NUM
gs-3955	101	109	)	)	PUNCT
gs-3955	101	110	the	the	DET
gs-3955	101	111	naked	naked	ADJ
gs-3955	101	112	mole	mole	NOUN
gs-3955	101	113	-	-	PUNCT
gs-3955	101	114	rat	rat	NOUN
gs-3955	101	115	(	(	PUNCT
gs-3955	101	116	heterocephalus	heterocephalus	ADJ
gs-3955	101	117	glaber	glaber	NOUN
gs-3955	101	118	):	):	PUNCT
gs-3955	101	119	defying	defy	VERB
gs-3955	101	120	mammalian	mammalian	ADJ
gs-3955	101	121	aging	age	VERB
gs-3955	101	122	patterns	pattern	NOUN
gs-3955	101	123	the	the	DET
gs-3955	101	124	naked	naked	ADJ
gs-3955	101	125	mole	mole	NOUN
gs-3955	101	126	-	-	PUNCT
gs-3955	101	127	rat	rat	NOUN
gs-3955	101	128	(	(	PUNCT
gs-3955	101	129	heterocephalus	heterocephalus	ADJ
gs-3955	101	130	glaber	glaber	NOUN
gs-3955	101	131	)	)	PUNCT
gs-3955	101	132	represents	represent	VERB
gs-3955	101	133	a	a	DET
gs-3955	101	134	mammalian	mammalian	ADJ
gs-3955	101	135	anomaly	anomaly	NOUN
gs-3955	101	136	,	,	PUNCT
gs-3955	101	137	achieving	achieve	VERB
gs-3955	101	138	37	37	NUM
gs-3955	101	139	-	-	PUNCT
gs-3955	101	140	year	year	NOUN
gs-3955	101	141	lifespans	lifespan	NOUN
gs-3955	101	142	(	(	PUNCT
gs-3955	101	143	5	5	NUM
gs-3955	101	144	-	-	PUNCT
gs-3955	101	145	10×	10×	NOUN
gs-3955	101	146	longer	long	ADV
gs-3955	101	147	than	than	ADP
gs-3955	101	148	size	size	NOUN
gs-3955	101	149	-	-	PUNCT
gs-3955	101	150	matched	match	VERB
gs-3955	101	151	rodents	rodent	NOUN
gs-3955	101	152	)	)	PUNCT
gs-3955	101	153	while	while	SCONJ
gs-3955	101	154	maintaining	maintain	VERB
gs-3955	101	155	cancer	cancer	NOUN
gs-3955	101	156	resistance	resistance	NOUN
gs-3955	101	157	(	(	PUNCT
gs-3955	101	158	buffenstein	buffenstein	ADV
gs-3955	101	159	,	,	PUNCT
gs-3955	101	160	2008	2008	NUM
gs-3955	101	161	;	;	PUNCT
gs-3955	101	162	seluanov	seluanov	PROPN
gs-3955	101	163	et	et	PROPN
gs-3955	101	164	al	al	PROPN
gs-3955	101	165	.	.	PROPN
gs-3955	101	166	,	,	PUNCT
gs-3955	101	167	2018	2018	NUM
gs-3955	101	168	)	)	PUNCT
gs-3955	101	169	.	.	PUNCT
gs-3955	102	1	this	this	DET
gs-3955	102	2	eusocial	eusocial	ADJ
gs-3955	102	3	subterranean	subterranean	ADJ
gs-3955	102	4	rodent	rodent	NOUN
gs-3955	102	5	has	have	AUX
gs-3955	102	6	revolutionized	revolutionize	VERB
gs-3955	102	7	our	our	PRON
gs-3955	102	8	understanding	understanding	NOUN
gs-3955	102	9	of	of	ADP
gs-3955	102	10	aging	aging	NOUN
gs-3955	102	11	in	in	ADP
gs-3955	102	12	mammals	mammal	NOUN
gs-3955	102	13	.	.	PUNCT
gs-3955	103	1	cancer	cancer	NOUN
gs-3955	103	2	resistance	resistance	NOUN
gs-3955	103	3	mechanisms	mechanism	NOUN
gs-3955	103	4	complete	complete	VERB
gs-3955	103	5	resistance	resistance	NOUN
gs-3955	103	6	to	to	ADP
gs-3955	103	7	spontaneous	spontaneous	ADJ
gs-3955	103	8	and	and	CCONJ
gs-3955	103	9	induced	induced	ADJ
gs-3955	103	10	carcinogenesis	carcinogenesis	NOUN
gs-3955	103	11	(	(	PUNCT
gs-3955	103	12	tian	tian	PROPN
gs-3955	103	13	et	et	PROPN
gs-3955	103	14	al	al	PROPN
gs-3955	103	15	.	.	PROPN
gs-3955	103	16	,	,	PUNCT
gs-3955	103	17	2013	2013	NUM
gs-3955	103	18	)	)	PUNCT
gs-3955	103	19	involves	involve	VERB
gs-3955	103	20	:	:	PUNCT
gs-3955	103	21	●	●	NUM
gs-3955	103	22	high	high	ADJ
gs-3955	103	23	-	-	PUNCT
gs-3955	103	24	molecular	molecular	ADJ
gs-3955	103	25	-	-	PUNCT
gs-3955	103	26	weight	weight	NOUN
gs-3955	103	27	hyaluronan	hyaluronan	NOUN
gs-3955	103	28	(	(	PUNCT
gs-3955	103	29	hmw	hmw	PROPN
gs-3955	103	30	-	-	PUNCT
gs-3955	103	31	ha	ha	INTJ
gs-3955	103	32	)	)	PUNCT
gs-3955	103	33	mediated	mediate	VERB
gs-3955	103	34	contact	contact	NOUN
gs-3955	103	35	inhibition	inhibition	NOUN
gs-3955	103	36	(	(	PUNCT
gs-3955	103	37	gorbunova	gorbunova	PROPN
gs-3955	103	38	et	et	PROPN
gs-3955	103	39	al	al	PROPN
gs-3955	103	40	.	.	PROPN
gs-3955	103	41	,	,	PUNCT
gs-3955	103	42	2014	2014	NUM
gs-3955	103	43	)	)	PUNCT
gs-3955	103	44	●	●	PUNCT
gs-3955	103	45	tumor	tumor	NOUN
gs-3955	103	46	suppressor	suppressor	NOUN
gs-3955	103	47	overexpression	overexpression	NOUN
gs-3955	103	48	(	(	PUNCT
gs-3955	103	49	p16ink4a	p16ink4a	PROPN
gs-3955	103	50	,	,	PUNCT
gs-3955	103	51	p27kip1	p27kip1	PROPN
gs-3955	103	52	)	)	PUNCT
gs-3955	103	53	(	(	PUNCT
gs-3955	103	54	hulbert	hulbert	NOUN
gs-3955	103	55	et	et	PROPN
gs-3955	103	56	al	al	PROPN
gs-3955	103	57	.	.	PROPN
gs-3955	103	58	,	,	PUNCT
gs-3955	103	59	2006	2006	NUM
gs-3955	103	60	)	)	PUNCT
gs-3955	103	61	●	●	PUNCT
gs-3955	103	62	aberrant	aberrant	ADJ
gs-3955	103	63	cell	cell	NOUN
gs-3955	103	64	recognition	recognition	NOUN
gs-3955	103	65	systems	system	NOUN
gs-3955	103	66	(	(	PUNCT
gs-3955	103	67	seluanov	seluanov	PROPN
gs-3955	103	68	et	et	PROPN
gs-3955	103	69	al	al	PROPN
gs-3955	103	70	.	.	PROPN
gs-3955	103	71	,	,	PUNCT
gs-3955	103	72	2009	2009	NUM
gs-3955	103	73	)	)	PUNCT
gs-3955	103	74	oxidative	oxidative	ADJ
gs-3955	103	75	stress	stress	NOUN
gs-3955	103	76	paradox	paradox	NOUN
gs-3955	103	77	despite	despite	SCONJ
gs-3955	103	78	high	high	ADJ
gs-3955	103	79	ros	ros	PROPN
gs-3955	103	80	production	production	NOUN
gs-3955	103	81	,	,	PUNCT
gs-3955	103	82	oxidative	oxidative	ADJ
gs-3955	103	83	damage	damage	NOUN
gs-3955	103	84	remains	remain	VERB
gs-3955	103	85	minimal	minimal	ADJ
gs-3955	103	86	(	(	PUNCT
gs-3955	103	87	lewis	lewis	PROPN
gs-3955	103	88	et	et	PROPN
gs-3955	103	89	al	al	PROPN
gs-3955	103	90	.	.	PROPN
gs-3955	103	91	,	,	PUNCT
gs-3955	103	92	2018	2018	NUM
gs-3955	103	93	)	)	PUNCT
gs-3955	103	94	due	due	ADP
gs-3955	103	95	to	to	ADP
gs-3955	103	96	:	:	PUNCT
gs-3955	103	97	●	●	PUNCT
gs-3955	103	98	unique	unique	ADJ
gs-3955	103	99	cardiolipin	cardiolipin	NOUN
gs-3955	103	100	structures	structure	NOUN
gs-3955	103	101	(	(	PUNCT
gs-3955	103	102	porter	porter	NOUN
gs-3955	103	103	et	et	NOUN
gs-3955	103	104	al	al	PROPN
gs-3955	103	105	.	.	PROPN
gs-3955	103	106	,	,	PUNCT
gs-3955	103	107	2015	2015	NUM
gs-3955	103	108	)	)	PUNCT
gs-3955	104	1	●	●	NUM
gs-3955	104	2	enhanced	enhance	VERB
gs-3955	104	3	dna	dna	PROPN
gs-3955	104	4	repair	repair	NOUN
gs-3955	104	5	capacity	capacity	NOUN
gs-3955	104	6	(	(	PUNCT
gs-3955	104	7	macrae	macrae	PROPN
gs-3955	104	8	et	et	PROPN
gs-3955	104	9	al	al	PROPN
gs-3955	104	10	.	.	PROPN
gs-3955	104	11	,	,	PUNCT
gs-3955	104	12	2015	2015	NUM
gs-3955	104	13	)	)	PUNCT
gs-3955	104	14	●	●	NUM
gs-3955	104	15	ribosomal	ribosomal	NOUN
gs-3955	104	16	biogenesis	biogenesis	NOUN
gs-3955	104	17	modifications	modification	NOUN
gs-3955	104	18	(	(	PUNCT
gs-3955	104	19	azpurua	azpurua	PROPN
gs-3955	104	20	et	et	PROPN
gs-3955	104	21	al	al	PROPN
gs-3955	104	22	.	.	PROPN
gs-3955	104	23	,	,	PUNCT
gs-3955	104	24	2013	2013	NUM
gs-3955	104	25	)	)	PUNCT
gs-3955	104	26	epigenetic	epigenetic	ADJ
gs-3955	104	27	stability	stability	NOUN
gs-3955	104	28	exceptional	exceptional	ADJ
gs-3955	104	29	epigenetic	epigenetic	ADJ
gs-3955	104	30	maintenance	maintenance	NOUN
gs-3955	104	31	features	feature	NOUN
gs-3955	104	32	include	include	VERB
gs-3955	104	33	:	:	PUNCT
gs-3955	104	34	●	●	PUNCT
gs-3955	104	35	slow	slow	ADJ
gs-3955	104	36	methylation	methylation	NOUN
gs-3955	104	37	drift	drift	NOUN
gs-3955	104	38	(	(	PUNCT
gs-3955	104	39	ma	ma	PROPN
gs-3955	104	40	et	et	PROPN
gs-3955	104	41	al	al	PROPN
gs-3955	104	42	.	.	PROPN
gs-3955	104	43	,	,	PUNCT
gs-3955	104	44	2020	2020	NUM
gs-3955	104	45	)	)	PUNCT
gs-3955	104	46	●	●	NUM
gs-3955	104	47	conserved	conserved	ADJ
gs-3955	104	48	aging	age	VERB
gs-3955	104	49	-	-	PUNCT
gs-3955	104	50	related	relate	VERB
gs-3955	104	51	gene	gene	NOUN
gs-3955	104	52	methylation	methylation	NOUN
gs-3955	104	53	(	(	PUNCT
gs-3955	104	54	holtze	holtze	PROPN
gs-3955	104	55	et	et	PROPN
gs-3955	104	56	al	al	PROPN
gs-3955	104	57	.	.	PROPN
gs-3955	104	58	,	,	PUNCT
gs-3955	104	59	2021	2021	NUM
gs-3955	104	60	)	)	PUNCT
gs-3955	104	61	●	●	PUNCT
gs-3955	104	62	dna	dna	NOUN
gs-3955	104	63	methyltransferase	methyltransferase	NOUN
gs-3955	104	64	activity	activity	NOUN
gs-3955	104	65	(	(	PUNCT
gs-3955	104	66	yang	yang	PROPN
gs-3955	104	67	et	et	PROPN
gs-3955	104	68	al	al	PROPN
gs-3955	104	69	.	.	PROPN
gs-3955	104	70	,	,	PUNCT
gs-3955	104	71	2019	2019	NUM
gs-3955	104	72	)	)	PUNCT
gs-3955	104	73	translational	translational	ADJ
gs-3955	104	74	potential	potential	NOUN
gs-3955	104	75	includes	include	VERB
gs-3955	104	76	:	:	PUNCT
gs-3955	104	77	●	●	NUM
gs-3955	104	78	hmw	hmw	PROPN
gs-3955	104	79	-	-	PUNCT
gs-3955	104	80	ha	ha	INTJ
gs-3955	104	81	based	base	VERB
gs-3955	104	82	anticancer	anticancer	NOUN
gs-3955	104	83	therapies	therapy	NOUN
gs-3955	104	84	(	(	PUNCT
gs-3955	104	85	tian	tian	PROPN
gs-3955	104	86	et	et	PROPN
gs-3955	104	87	al	al	PROPN
gs-3955	104	88	.	.	PROPN
gs-3955	104	89	,	,	PUNCT
gs-3955	104	90	2019	2019	NUM
gs-3955	104	91	)	)	PUNCT
gs-3955	104	92	●	●	PUNCT
gs-3955	104	93	mitochondrial	mitochondrial	ADJ
gs-3955	104	94	-	-	PUNCT
gs-3955	104	95	protective	protective	ADJ
gs-3955	104	96	compounds	compound	NOUN
gs-3955	104	97	(	(	PUNCT
gs-3955	104	98	lewis	lewis	PROPN
gs-3955	104	99	et	et	PROPN
gs-3955	104	100	al	al	PROPN
gs-3955	104	101	.	.	PROPN
gs-3955	104	102	,	,	PUNCT
gs-3955	104	103	2020	2020	NUM
gs-3955	104	104	)	)	PUNCT
gs-3955	104	105	●	●	PUNCT
gs-3955	104	106	epigenetic	epigenetic	ADJ
gs-3955	104	107	aging	age	VERB
gs-3955	104	108	interventions	intervention	NOUN
gs-3955	104	109	(	(	PUNCT
gs-3955	104	110	zhang	zhang	X
gs-3955	104	111	et	et	PROPN
gs-3955	104	112	al	al	PROPN
gs-3955	104	113	.	.	PROPN
gs-3955	104	114	,	,	PUNCT
gs-3955	104	115	2021	2021	NUM
gs-3955	104	116	)	)	PUNCT
gs-3955	104	117	this	this	DET
gs-3955	104	118	comprehensive	comprehensive	ADJ
gs-3955	104	119	analysis	analysis	NOUN
gs-3955	104	120	demonstrates	demonstrate	VERB
gs-3955	104	121	how	how	SCONJ
gs-3955	104	122	negligibly	negligibly	ADV
gs-3955	104	123	senescing	senesce	VERB
gs-3955	104	124	species	specie	NOUN
gs-3955	104	125	provide	provide	VERB
gs-3955	104	126	unprecedented	unprecedented	ADJ
gs-3955	104	127	insights	insight	NOUN
gs-3955	104	128	into	into	ADP
gs-3955	104	129	longevity	longevity	NOUN
gs-3955	104	130	mechanisms	mechanism	NOUN
gs-3955	104	131	,	,	PUNCT
gs-3955	104	132	offering	offer	VERB
gs-3955	104	133	novel	novel	ADJ
gs-3955	104	134	avenues	avenue	NOUN
gs-3955	104	135	for	for	ADP
gs-3955	104	136	biomedical	biomedical	ADJ
gs-3955	104	137	innovation	innovation	NOUN
gs-3955	104	138	while	while	SCONJ
gs-3955	104	139	challenging	challenge	VERB
gs-3955	104	140	fundamental	fundamental	ADJ
gs-3955	104	141	paradigms	paradigm	NOUN
gs-3955	104	142	in	in	ADP
gs-3955	104	143	aging	age	VERB
gs-3955	104	144	research	research	NOUN
gs-3955	104	145	.	.	PUNCT
gs-3955	105	1	georgian	georgian	ADJ
gs-3955	105	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	105	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	105	4	ტ.	ტ.	VERB
gs-3955	105	5	7	7	NUM
gs-3955	105	6	n	n	PRON
gs-3955	105	7	3	3	NUM
gs-3955	105	8	,	,	PUNCT
gs-3955	105	9	2025	2025	NUM
gs-3955	105	10	355	355	NUM
gs-3955	105	11	extraordinary	extraordinary	ADJ
gs-3955	105	12	longevity	longevity	NOUN
gs-3955	105	13	in	in	ADP
gs-3955	105	14	cave	cave	NOUN
gs-3955	105	15	-	-	PUNCT
gs-3955	105	16	dwelling	dwell	VERB
gs-3955	105	17	amphibians	amphibian	NOUN
gs-3955	105	18	:	:	PUNCT
gs-3955	105	19	the	the	DET
gs-3955	105	20	olm	olm	NOUN
gs-3955	105	21	(	(	PUNCT
gs-3955	105	22	proteus	proteus	PROPN
gs-3955	105	23	anguinus	anguinus	PROPN
gs-3955	105	24	)	)	PUNCT
gs-3955	105	25	the	the	DET
gs-3955	105	26	olm	olm	NOUN
gs-3955	105	27	or	or	CCONJ
gs-3955	105	28	proteus	proteus	NOUN
gs-3955	105	29	(	(	PUNCT
gs-3955	105	30	proteus	proteus	NOUN
gs-3955	105	31	anguinus	anguinus	PROPN
gs-3955	105	32	)	)	PUNCT
gs-3955	105	33	,	,	PUNCT
gs-3955	105	34	a	a	DET
gs-3955	105	35	remarkable	remarkable	ADJ
gs-3955	105	36	blind	blind	ADJ
gs-3955	105	37	amphibian	amphibian	NOUN
gs-3955	105	38	inhabiting	inhabit	VERB
gs-3955	105	39	subterranean	subterranean	ADJ
gs-3955	105	40	karst	karst	ADJ
gs-3955	105	41	aquifers	aquifer	NOUN
gs-3955	105	42	of	of	ADP
gs-3955	105	43	the	the	DET
gs-3955	105	44	balkan	balkan	PROPN
gs-3955	105	45	peninsula	peninsula	NOUN
gs-3955	105	46	,	,	PUNCT
gs-3955	105	47	stands	stand	VERB
gs-3955	105	48	as	as	ADP
gs-3955	105	49	one	one	NUM
gs-3955	105	50	of	of	ADP
gs-3955	105	51	nature	nature	NOUN
gs-3955	105	52	's	's	PART
gs-3955	105	53	most	most	ADV
gs-3955	105	54	fascinating	fascinating	ADJ
gs-3955	105	55	examples	example	NOUN
gs-3955	105	56	of	of	ADP
gs-3955	105	57	extreme	extreme	ADJ
gs-3955	105	58	longevity	longevity	NOUN
gs-3955	105	59	,	,	PUNCT
gs-3955	105	60	with	with	ADP
gs-3955	105	61	documented	document	VERB
gs-3955	105	62	lifespans	lifespan	NOUN
gs-3955	105	63	exceeding	exceed	VERB
gs-3955	105	64	a	a	DET
gs-3955	105	65	century	century	NOUN
gs-3955	105	66	(	(	PUNCT
gs-3955	105	67	voituron	voituron	PROPN
gs-3955	105	68	et	et	PROPN
gs-3955	105	69	al	al	PROPN
gs-3955	105	70	.	.	PROPN
gs-3955	105	71	,	,	PUNCT
gs-3955	105	72	2011	2011	NUM
gs-3955	105	73	)	)	PUNCT
gs-3955	105	74	.	.	PUNCT
gs-3955	106	1	this	this	DET
gs-3955	106	2	neotenic	neotenic	ADJ
gs-3955	106	3	salamander	salamander	NOUN
gs-3955	106	4	has	have	AUX
gs-3955	106	5	become	become	VERB
gs-3955	106	6	a	a	DET
gs-3955	106	7	focal	focal	ADJ
gs-3955	106	8	point	point	NOUN
gs-3955	106	9	in	in	ADP
gs-3955	106	10	biogerontology	biogerontology	NOUN
gs-3955	106	11	research	research	NOUN
gs-3955	106	12	due	due	ADP
gs-3955	106	13	to	to	ADP
gs-3955	106	14	its	its	PRON
gs-3955	106	15	trifecta	trifecta	NOUN
gs-3955	106	16	of	of	ADP
gs-3955	106	17	biological	biological	ADJ
gs-3955	106	18	marvels	marvel	NOUN
gs-3955	106	19	:	:	PUNCT
gs-3955	106	20	negligible	negligible	ADJ
gs-3955	106	21	senescence	senescence	NOUN
gs-3955	106	22	,	,	PUNCT
gs-3955	106	23	exceptional	exceptional	ADJ
gs-3955	106	24	oxidative	oxidative	ADJ
gs-3955	106	25	stress	stress	NOUN
gs-3955	106	26	resistance	resistance	NOUN
gs-3955	106	27	,	,	PUNCT
gs-3955	106	28	and	and	CCONJ
gs-3955	106	29	unparalleled	unparalleled	ADJ
gs-3955	106	30	regenerative	regenerative	ADJ
gs-3955	106	31	capacities	capacity	NOUN
gs-3955	106	32	(	(	PUNCT
gs-3955	106	33	bulog	bulog	PROPN
gs-3955	106	34	et	et	PROPN
gs-3955	106	35	al	al	PROPN
gs-3955	106	36	.	.	PROPN
gs-3955	106	37	,	,	PUNCT
gs-3955	106	38	2020	2020	NUM
gs-3955	106	39	)	)	PUNCT
gs-3955	106	40	.	.	PUNCT
gs-3955	107	1	metabolic	metabolic	NOUN
gs-3955	107	2	adaptations	adaptation	NOUN
gs-3955	107	3	to	to	ADP
gs-3955	107	4	hypoxic	hypoxic	ADJ
gs-3955	107	5	environments	environment	NOUN
gs-3955	107	6	evolution	evolution	NOUN
gs-3955	107	7	in	in	ADP
gs-3955	107	8	perpetual	perpetual	ADJ
gs-3955	107	9	darkness	darkness	NOUN
gs-3955	107	10	and	and	CCONJ
gs-3955	107	11	oxygen	oxygen	NOUN
gs-3955	107	12	-	-	PUNCT
gs-3955	107	13	deprived	deprive	VERB
gs-3955	107	14	waters	water	NOUN
gs-3955	107	15	has	have	AUX
gs-3955	107	16	sculpted	sculpt	VERB
gs-3955	107	17	unique	unique	ADJ
gs-3955	107	18	metabolic	metabolic	NOUN
gs-3955	107	19	characteristics	characteristic	NOUN
gs-3955	107	20	in	in	ADP
gs-3955	107	21	p.	p.	PROPN
gs-3955	107	22	anguinus	anguinus	PROPN
gs-3955	107	23	:	:	PUNCT
gs-3955	107	24	●	●	NUM
gs-3955	107	25	profound	profound	ADJ
gs-3955	107	26	metabolic	metabolic	NOUN
gs-3955	107	27	depression	depression	NOUN
gs-3955	107	28	,	,	PUNCT
gs-3955	107	29	exhibiting	exhibit	VERB
gs-3955	107	30	30	30	NUM
gs-3955	107	31	-	-	SYM
gs-3955	107	32	50	50	NUM
gs-3955	107	33	%	%	NOUN
gs-3955	107	34	lower	low	ADJ
gs-3955	107	35	metabolic	metabolic	NOUN
gs-3955	107	36	rates	rate	NOUN
gs-3955	107	37	compared	compare	VERB
gs-3955	107	38	to	to	ADP
gs-3955	107	39	surface	surface	NOUN
gs-3955	107	40	-	-	PUNCT
gs-3955	107	41	dwelling	dwell	VERB
gs-3955	107	42	amphibians	amphibian	NOUN
gs-3955	107	43	(	(	PUNCT
gs-3955	107	44	issartel	issartel	PROPN
gs-3955	107	45	et	et	PROPN
gs-3955	107	46	al	al	PROPN
gs-3955	107	47	.	.	PROPN
gs-3955	107	48	,	,	PUNCT
gs-3955	107	49	2009	2009	NUM
gs-3955	107	50	)	)	PUNCT
gs-3955	107	51	●	●	NUM
gs-3955	107	52	enhanced	enhance	VERB
gs-3955	107	53	reperfusion	reperfusion	NOUN
gs-3955	107	54	injury	injury	NOUN
gs-3955	107	55	resistance	resistance	NOUN
gs-3955	107	56	through	through	ADP
gs-3955	107	57	upregulated	upregulated	ADJ
gs-3955	107	58	antioxidant	antioxidant	ADJ
gs-3955	107	59	enzymes	enzyme	NOUN
gs-3955	107	60	(	(	PUNCT
gs-3955	107	61	sod	sod	NOUN
gs-3955	107	62	,	,	PUNCT
gs-3955	107	63	catalase	catalase	NOUN
gs-3955	107	64	)	)	PUNCT
gs-3955	107	65	(	(	PUNCT
gs-3955	107	66	goricar	goricar	NOUN
gs-3955	107	67	et	et	PROPN
gs-3955	107	68	al	al	PROPN
gs-3955	107	69	.	.	PROPN
gs-3955	107	70	,	,	PUNCT
gs-3955	107	71	2021	2021	NUM
gs-3955	107	72	)	)	PUNCT
gs-3955	108	1	●	●	NUM
gs-3955	108	2	anaerobic	anaerobic	NOUN
gs-3955	108	3	metabolic	metabolic	NOUN
gs-3955	108	4	flexibility	flexibility	NOUN
gs-3955	108	5	allowing	allow	VERB
gs-3955	108	6	prolonged	prolonged	ADJ
gs-3955	108	7	survival	survival	NOUN
gs-3955	108	8	without	without	ADP
gs-3955	108	9	oxygen	oxygen	NOUN
gs-3955	108	10	(	(	PUNCT
gs-3955	108	11	voituron	voituron	PROPN
gs-3955	108	12	et	et	PROPN
gs-3955	108	13	al	al	PROPN
gs-3955	108	14	.	.	PROPN
gs-3955	108	15	,	,	PUNCT
gs-3955	108	16	2011	2011	NUM
gs-3955	108	17	)	)	PUNCT
gs-3955	108	18	paradoxical	paradoxical	ADJ
gs-3955	108	19	oxidative	oxidative	ADJ
gs-3955	108	20	stress	stress	NOUN
gs-3955	108	21	resistance	resistance	NOUN
gs-3955	108	22	despite	despite	SCONJ
gs-3955	108	23	relatively	relatively	ADV
gs-3955	108	24	low	low	ADJ
gs-3955	108	25	antioxidant	antioxidant	ADJ
gs-3955	108	26	enzyme	enzyme	NOUN
gs-3955	108	27	levels	level	NOUN
gs-3955	108	28	,	,	PUNCT
gs-3955	108	29	olms	olm	NOUN
gs-3955	108	30	display	display	VERB
gs-3955	108	31	minimal	minimal	ADJ
gs-3955	108	32	oxidative	oxidative	ADJ
gs-3955	108	33	damage	damage	NOUN
gs-3955	108	34	accumulation	accumulation	NOUN
gs-3955	108	35	(	(	PUNCT
gs-3955	108	36	goricar	goricar	NOUN
gs-3955	108	37	et	et	PROPN
gs-3955	108	38	al	al	PROPN
gs-3955	108	39	.	.	PROPN
gs-3955	108	40	,	,	PUNCT
gs-3955	108	41	2021	2021	NUM
gs-3955	108	42	)	)	PUNCT
gs-3955	108	43	,	,	PUNCT
gs-3955	108	44	attributable	attributable	ADJ
gs-3955	108	45	to	to	ADP
gs-3955	108	46	:	:	PUNCT
gs-3955	108	47	●	●	NUM
gs-3955	108	48	specialized	specialized	ADJ
gs-3955	108	49	membrane	membrane	NOUN
gs-3955	108	50	lipid	lipid	NOUN
gs-3955	108	51	composition	composition	NOUN
gs-3955	108	52	conferring	confer	VERB
gs-3955	108	53	peroxidation	peroxidation	NOUN
gs-3955	108	54	resistance	resistance	NOUN
gs-3955	108	55	(	(	PUNCT
gs-3955	108	56	bulog	bulog	PROPN
gs-3955	108	57	et	et	PROPN
gs-3955	108	58	al	al	PROPN
gs-3955	108	59	.	.	PROPN
gs-3955	108	60	,	,	PUNCT
gs-3955	108	61	2020	2020	NUM
gs-3955	108	62	)	)	PUNCT
gs-3955	108	63	●	●	PUNCT
gs-3955	108	64	constitutive	constitutive	ADJ
gs-3955	108	65	activation	activation	NOUN
gs-3955	108	66	of	of	ADP
gs-3955	108	67	nrf2	nrf2	NOUN
gs-3955	108	68	pathway	pathway	NOUN
gs-3955	108	69	,	,	PUNCT
gs-3955	108	70	a	a	DET
gs-3955	108	71	master	master	NOUN
gs-3955	108	72	regulator	regulator	NOUN
gs-3955	108	73	of	of	ADP
gs-3955	108	74	antioxidant	antioxidant	ADJ
gs-3955	108	75	defenses	defense	NOUN
gs-3955	108	76	(	(	PUNCT
gs-3955	108	77	trontelj	trontelj	NOUN
gs-3955	108	78	et	et	PROPN
gs-3955	108	79	al	al	PROPN
gs-3955	108	80	.	.	PROPN
gs-3955	108	81	,	,	PUNCT
gs-3955	108	82	2022	2022	NUM
gs-3955	108	83	)	)	PUNCT
gs-3955	108	84	●	●	PUNCT
gs-3955	108	85	efficient	efficient	ADJ
gs-3955	108	86	mitochondrial	mitochondrial	ADJ
gs-3955	108	87	electron	electron	NOUN
gs-3955	108	88	transport	transport	NOUN
gs-3955	108	89	minimizing	minimize	VERB
gs-3955	108	90	ros	ros	PROPN
gs-3955	108	91	production	production	NOUN
gs-3955	108	92	(	(	PUNCT
gs-3955	108	93	issartel	issartel	PROPN
gs-3955	108	94	et	et	PROPN
gs-3955	108	95	al	al	PROPN
gs-3955	108	96	.	.	PROPN
gs-3955	108	97	,	,	PUNCT
gs-3955	108	98	2009	2009	NUM
gs-3955	108	99	)	)	PUNCT
gs-3955	108	100	regenerative	regenerative	ADJ
gs-3955	108	101	prowess	prowess	NOUN
gs-3955	108	102	p.	p.	NOUN
gs-3955	108	103	anguinus	anguinus	PROPN
gs-3955	108	104	exhibits	exhibit	VERB
gs-3955	108	105	regenerative	regenerative	ADJ
gs-3955	108	106	capabilities	capability	NOUN
gs-3955	108	107	surpassing	surpass	VERB
gs-3955	108	108	most	most	ADJ
gs-3955	108	109	vertebrates	vertebrate	NOUN
gs-3955	108	110	:	:	PUNCT
gs-3955	108	111	●	●	PUNCT
gs-3955	108	112	complete	complete	ADJ
gs-3955	108	113	limb	limb	NOUN
gs-3955	108	114	regeneration	regeneration	NOUN
gs-3955	108	115	with	with	ADP
gs-3955	108	116	perfect	perfect	ADJ
gs-3955	108	117	tissue	tissue	NOUN
gs-3955	108	118	patterning	patterning	NOUN
gs-3955	108	119	and	and	CCONJ
gs-3955	108	120	absence	absence	NOUN
gs-3955	108	121	of	of	ADP
gs-3955	108	122	scarring	scar	VERB
gs-3955	108	123	(	(	PUNCT
gs-3955	108	124	bizjak	bizjak	ADV
gs-3955	108	125	mali	mali	VERB
gs-3955	108	126	et	et	PROPN
gs-3955	108	127	al	al	PROPN
gs-3955	108	128	.	.	PROPN
gs-3955	108	129	,	,	PUNCT
gs-3955	108	130	2019	2019	NUM
gs-3955	108	131	)	)	PUNCT
gs-3955	108	132	●	●	NUM
gs-3955	108	133	visceral	visceral	ADJ
gs-3955	108	134	organ	organ	NOUN
gs-3955	108	135	regeneration	regeneration	NOUN
gs-3955	108	136	,	,	PUNCT
gs-3955	108	137	including	include	VERB
gs-3955	108	138	functional	functional	ADJ
gs-3955	108	139	restoration	restoration	NOUN
gs-3955	108	140	of	of	ADP
gs-3955	108	141	cardiac	cardiac	ADJ
gs-3955	108	142	and	and	CCONJ
gs-3955	108	143	neural	neural	ADJ
gs-3955	108	144	tissues	tissue	NOUN
gs-3955	108	145	(	(	PUNCT
gs-3955	108	146	aphkhazava	aphkhazava	NOUN
gs-3955	108	147	et	et	PROPN
gs-3955	108	148	al	al	PROPN
gs-3955	108	149	.	.	PROPN
gs-3955	108	150	,	,	PUNCT
gs-3955	108	151	2025	2025	NUM
gs-3955	108	152	)	)	PUNCT
gs-3955	108	153	●	●	NUM
gs-3955	108	154	enhanced	enhance	VERB
gs-3955	108	155	wound	wound	NOUN
gs-3955	108	156	healing	healing	NOUN
gs-3955	108	157	through	through	ADP
gs-3955	108	158	rapid	rapid	ADJ
gs-3955	108	159	epithelialization	epithelialization	NOUN
gs-3955	108	160	(	(	PUNCT
gs-3955	108	161	bulog	bulog	PROPN
gs-3955	108	162	et	et	PROPN
gs-3955	108	163	al	al	PROPN
gs-3955	108	164	.	.	PROPN
gs-3955	108	165	,	,	PUNCT
gs-3955	108	166	2020	2020	NUM
gs-3955	108	167	)	)	PUNCT
gs-3955	108	168	genomic	genomic	NOUN
gs-3955	108	169	foundations	foundation	NOUN
gs-3955	108	170	of	of	ADP
gs-3955	108	171	longevity	longevity	NOUN
gs-3955	108	172	genome	genome	NOUN
gs-3955	108	173	sequencing	sequencing	NOUN
gs-3955	108	174	reveals	reveal	VERB
gs-3955	108	175	extraordinary	extraordinary	ADJ
gs-3955	108	176	adaptations	adaptation	NOUN
gs-3955	108	177	:	:	PUNCT
gs-3955	108	178	georgian	georgian	ADJ
gs-3955	108	179	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	108	180	მეცნიერები	მეცნიერები	NOUN
gs-3955	108	181	ტ.	ტ.	VERB
gs-3955	108	182	7	7	NUM
gs-3955	108	183	n	n	DET
gs-3955	108	184	3	3	NUM
gs-3955	108	185	,	,	PUNCT
gs-3955	108	186	2025	2025	NUM
gs-3955	108	187	356	356	NUM
gs-3955	108	188	●	●	NUM
gs-3955	108	189	upregulated	upregulated	ADJ
gs-3955	108	190	dna	dna	PROPN
gs-3955	108	191	repair	repair	NOUN
gs-3955	108	192	genes	gene	NOUN
gs-3955	108	193	(	(	PUNCT
gs-3955	108	194	brca1	brca1	PROPN
gs-3955	108	195	,	,	PUNCT
gs-3955	108	196	rad51	rad51	NOUN
gs-3955	108	197	)	)	PUNCT
gs-3955	108	198	maintaining	maintain	VERB
gs-3955	108	199	genomic	genomic	ADJ
gs-3955	108	200	integrity	integrity	NOUN
gs-3955	108	201	(	(	PUNCT
gs-3955	108	202	goricar	goricar	NOUN
gs-3955	108	203	et	et	PROPN
gs-3955	108	204	al	al	PROPN
gs-3955	108	205	.	.	PROPN
gs-3955	108	206	,	,	PUNCT
gs-3955	108	207	2021	2021	NUM
gs-3955	108	208	)	)	PUNCT
gs-3955	109	1	●	●	SYM
gs-3955	109	2	unique	unique	ADJ
gs-3955	109	3	autophagy	autophagy	NOUN
gs-3955	109	4	-	-	PUNCT
gs-3955	109	5	related	relate	VERB
gs-3955	109	6	variants	variant	NOUN
gs-3955	109	7	(	(	PUNCT
gs-3955	109	8	atg5	atg5	PROPN
gs-3955	109	9	,	,	PUNCT
gs-3955	109	10	becn1	becn1	PROPN
gs-3955	109	11	)	)	PUNCT
gs-3955	109	12	ensuring	ensure	VERB
gs-3955	109	13	cellular	cellular	ADJ
gs-3955	109	14	quality	quality	NOUN
gs-3955	109	15	control	control	NOUN
gs-3955	109	16	(	(	PUNCT
gs-3955	109	17	trontelj	trontelj	NOUN
gs-3955	109	18	et	et	PROPN
gs-3955	109	19	al	al	PROPN
gs-3955	109	20	.	.	PROPN
gs-3955	109	21	,	,	PUNCT
gs-3955	109	22	2022	2022	NUM
gs-3955	109	23	)	)	PUNCT
gs-3955	109	24	●	●	PUNCT
gs-3955	109	25	slow	slow	ADJ
gs-3955	109	26	epigenetic	epigenetic	ADJ
gs-3955	109	27	drift	drift	NOUN
gs-3955	109	28	,	,	PUNCT
gs-3955	109	29	preserving	preserve	VERB
gs-3955	109	30	youthful	youthful	ADJ
gs-3955	109	31	gene	gene	NOUN
gs-3955	109	32	expression	expression	NOUN
gs-3955	109	33	patterns	pattern	NOUN
gs-3955	109	34	(	(	PUNCT
gs-3955	109	35	voituron	voituron	PROPN
gs-3955	109	36	et	et	PROPN
gs-3955	109	37	al	al	PROPN
gs-3955	109	38	.	.	PROPN
gs-3955	109	39	,	,	PUNCT
gs-3955	109	40	2011	2011	NUM
gs-3955	109	41	)	)	PUNCT
gs-3955	109	42	biomedical	biomedical	ADJ
gs-3955	109	43	implications	implication	NOUN
gs-3955	109	44	olm	olm	NOUN
gs-3955	109	45	research	research	NOUN
gs-3955	109	46	offers	offer	VERB
gs-3955	109	47	transformative	transformative	ADJ
gs-3955	109	48	potential	potential	NOUN
gs-3955	109	49	for	for	ADP
gs-3955	109	50	:	:	PUNCT
gs-3955	109	51	●	●	NUM
gs-3955	109	52	regenerative	regenerative	ADJ
gs-3955	109	53	medicine	medicine	NOUN
gs-3955	109	54	through	through	ADP
gs-3955	109	55	conserved	conserved	ADJ
gs-3955	109	56	tissue	tissue	NOUN
gs-3955	109	57	renewal	renewal	NOUN
gs-3955	109	58	mechanisms	mechanism	NOUN
gs-3955	109	59	(	(	PUNCT
gs-3955	109	60	bizjak	bizjak	ADV
gs-3955	109	61	mali	mali	VERB
gs-3955	109	62	et	et	PROPN
gs-3955	109	63	al	al	PROPN
gs-3955	109	64	.	.	PROPN
gs-3955	109	65	,	,	PUNCT
gs-3955	109	66	2019	2019	NUM
gs-3955	109	67	)	)	PUNCT
gs-3955	109	68	●	●	NUM
gs-3955	109	69	hypoxia	hypoxia	NOUN
gs-3955	109	70	therapeutics	therapeutic	NOUN
gs-3955	109	71	mimicking	mimic	VERB
gs-3955	109	72	natural	natural	ADJ
gs-3955	109	73	adaptations	adaptation	NOUN
gs-3955	109	74	(	(	PUNCT
gs-3955	109	75	issartel	issartel	PROPN
gs-3955	109	76	et	et	PROPN
gs-3955	109	77	al	al	PROPN
gs-3955	109	78	.	.	PROPN
gs-3955	109	79	,	,	PUNCT
gs-3955	109	80	2009	2009	NUM
gs-3955	109	81	)	)	PUNCT
gs-3955	109	82	●	●	PUNCT
gs-3955	109	83	epigenetic	epigenetic	ADJ
gs-3955	109	84	interventions	intervention	NOUN
gs-3955	109	85	against	against	ADP
gs-3955	109	86	aging	age	VERB
gs-3955	109	87	(	(	PUNCT
gs-3955	109	88	trontelj	trontelj	NOUN
gs-3955	109	89	et	et	PROPN
gs-3955	109	90	al	al	PROPN
gs-3955	109	91	.	.	PROPN
gs-3955	109	92	,	,	PUNCT
gs-3955	109	93	2022	2022	NUM
gs-3955	109	94	)	)	PUNCT
gs-3955	109	95	the	the	DET
gs-3955	109	96	fire	fire	NOUN
gs-3955	109	97	salamander	salamander	NOUN
gs-3955	109	98	(	(	PUNCT
gs-3955	109	99	salamandra	salamandra	NOUN
gs-3955	109	100	salamandra	salamandra	PROPN
gs-3955	109	101	):	):	PUNCT
gs-3955	109	102	a	a	DET
gs-3955	109	103	model	model	NOUN
gs-3955	109	104	of	of	ADP
gs-3955	109	105	amphibian	amphibian	ADJ
gs-3955	109	106	longevity	longevity	NOUN
gs-3955	109	107	the	the	DET
gs-3955	109	108	fire	fire	NOUN
gs-3955	109	109	salamander	salamander	NOUN
gs-3955	109	110	's	's	PART
gs-3955	109	111	extraordinary	extraordinary	ADJ
gs-3955	109	112	50	50	NUM
gs-3955	109	113	+	+	NUM
gs-3955	109	114	year	year	NOUN
gs-3955	109	115	lifespan	lifespan	NOUN
gs-3955	109	116	in	in	ADP
gs-3955	109	117	natural	natural	ADJ
gs-3955	109	118	habitats	habitat	NOUN
gs-3955	109	119	(	(	PUNCT
gs-3955	109	120	tacutu	tacutu	NOUN
gs-3955	109	121	et	et	PROPN
gs-3955	109	122	al	al	PROPN
gs-3955	109	123	.	.	PROPN
gs-3955	109	124	,	,	PUNCT
gs-3955	109	125	2013	2013	NUM
gs-3955	109	126	)	)	PUNCT
gs-3955	109	127	provides	provide	VERB
gs-3955	109	128	unique	unique	ADJ
gs-3955	109	129	insights	insight	NOUN
gs-3955	109	130	into	into	ADP
gs-3955	109	131	the	the	DET
gs-3955	109	132	interplay	interplay	NOUN
gs-3955	109	133	between	between	ADP
gs-3955	109	134	longevity	longevity	NOUN
gs-3955	109	135	,	,	PUNCT
gs-3955	109	136	regeneration	regeneration	NOUN
gs-3955	109	137	,	,	PUNCT
gs-3955	109	138	and	and	CCONJ
gs-3955	109	139	resistance	resistance	NOUN
gs-3955	109	140	to	to	ADP
gs-3955	109	141	age	age	NOUN
gs-3955	109	142	-	-	PUNCT
gs-3955	109	143	related	relate	VERB
gs-3955	109	144	decline	decline	NOUN
gs-3955	109	145	(	(	PUNCT
gs-3955	109	146	zhang	zhang	X
gs-3955	109	147	et	et	PROPN
gs-3955	109	148	al	al	PROPN
gs-3955	109	149	.	.	PROPN
gs-3955	109	150	,	,	PUNCT
gs-3955	109	151	2022	2022	NUM
gs-3955	109	152	)	)	PUNCT
gs-3955	109	153	.	.	PUNCT
gs-3955	110	1	regenerative	regenerative	ADJ
gs-3955	110	2	capabilities	capability	NOUN
gs-3955	110	3	s.	s.	PROPN
gs-3955	110	4	salamandra	salamandra	PROPN
gs-3955	110	5	displays	display	NOUN
gs-3955	110	6	:	:	PUNCT
gs-3955	110	7	●	●	PUNCT
gs-3955	110	8	complete	complete	ADJ
gs-3955	110	9	limb	limb	NOUN
gs-3955	110	10	regeneration	regeneration	NOUN
gs-3955	110	11	with	with	ADP
gs-3955	110	12	perfect	perfect	ADJ
gs-3955	110	13	musculoskeletal	musculoskeletal	ADJ
gs-3955	110	14	reconstruction	reconstruction	NOUN
gs-3955	110	15	(	(	PUNCT
gs-3955	110	16	joven	joven	PROPN
gs-3955	110	17	et	et	PROPN
gs-3955	110	18	al	al	PROPN
gs-3955	110	19	.	.	PROPN
gs-3955	110	20	,	,	PUNCT
gs-3955	110	21	2019	2019	NUM
gs-3955	110	22	)	)	PUNCT
gs-3955	110	23	●	●	NUM
gs-3955	110	24	cns	cns	NOUN
gs-3955	110	25	regeneration	regeneration	NOUN
gs-3955	110	26	,	,	PUNCT
gs-3955	110	27	including	include	VERB
gs-3955	110	28	functional	functional	ADJ
gs-3955	110	29	spinal	spinal	NOUN
gs-3955	110	30	cord	cord	NOUN
gs-3955	110	31	and	and	CCONJ
gs-3955	110	32	retinal	retinal	ADJ
gs-3955	110	33	repair	repair	NOUN
gs-3955	110	34	(	(	PUNCT
gs-3955	110	35	ferretti	ferretti	PROPN
gs-3955	110	36	&	&	CCONJ
gs-3955	110	37	ghosh	ghosh	PROPN
gs-3955	110	38	,	,	PUNCT
gs-3955	110	39	2021	2021	NUM
gs-3955	110	40	)	)	PUNCT
gs-3955	110	41	●	●	NUM
gs-3955	110	42	scarless	scarless	NOUN
gs-3955	110	43	wound	wound	NOUN
gs-3955	110	44	healing	healing	NOUN
gs-3955	110	45	through	through	ADP
gs-3955	110	46	modified	modify	VERB
gs-3955	110	47	fibroblast	fibroblast	ADJ
gs-3955	110	48	activity	activity	NOUN
gs-3955	110	49	(	(	PUNCT
gs-3955	110	50	seifert	seifert	PROPN
gs-3955	110	51	&	&	CCONJ
gs-3955	110	52	maden	maden	PROPN
gs-3955	110	53	,	,	PUNCT
gs-3955	110	54	2014	2014	NUM
gs-3955	110	55	)	)	PUNCT
gs-3955	110	56	antioxidant	antioxidant	ADJ
gs-3955	110	57	defense	defense	NOUN
gs-3955	110	58	systems	system	NOUN
gs-3955	110	59	remarkable	remarkable	ADJ
gs-3955	110	60	oxidative	oxidative	ADJ
gs-3955	110	61	stress	stress	NOUN
gs-3955	110	62	resistance	resistance	NOUN
gs-3955	110	63	involves	involve	VERB
gs-3955	110	64	:	:	PUNCT
gs-3955	110	65	●	●	NUM
gs-3955	110	66	elevated	elevated	ADJ
gs-3955	110	67	sod	sod	NOUN
gs-3955	110	68	and	and	CCONJ
gs-3955	110	69	catalase	catalase	NOUN
gs-3955	110	70	activity	activity	NOUN
gs-3955	110	71	(	(	PUNCT
gs-3955	110	72	rasch	rasch	PROPN
gs-3955	110	73	et	et	PROPN
gs-3955	110	74	al	al	PROPN
gs-3955	110	75	.	.	PROPN
gs-3955	110	76	,	,	PUNCT
gs-3955	110	77	2020	2020	NUM
gs-3955	110	78	)	)	PUNCT
gs-3955	110	79	●	●	NUM
gs-3955	110	80	high	high	ADJ
gs-3955	110	81	tissue	tissue	NOUN
gs-3955	110	82	glutathione	glutathione	NOUN
gs-3955	110	83	levels	level	NOUN
gs-3955	110	84	(	(	PUNCT
gs-3955	110	85	wells	well	NOUN
gs-3955	110	86	et	et	PROPN
gs-3955	110	87	al	al	PROPN
gs-3955	110	88	.	.	PROPN
gs-3955	110	89	,	,	PUNCT
gs-3955	110	90	2021	2021	NUM
gs-3955	110	91	)	)	PUNCT
gs-3955	110	92	●	●	NUM
gs-3955	110	93	efficient	efficient	ADJ
gs-3955	110	94	dna	dna	PROPN
gs-3955	110	95	repair	repair	NOUN
gs-3955	110	96	mechanisms	mechanism	NOUN
gs-3955	110	97	(	(	PUNCT
gs-3955	110	98	macrae	macrae	NOUN
gs-3955	110	99	et	et	PROPN
gs-3955	110	100	al	al	PROPN
gs-3955	110	101	.	.	PROPN
gs-3955	110	102	,	,	PUNCT
gs-3955	110	103	2015	2015	NUM
gs-3955	110	104	)	)	PUNCT
gs-3955	110	105	metabolic	metabolic	NOUN
gs-3955	110	106	adaptations	adaptation	NOUN
gs-3955	110	107	longevity	longevity	NOUN
gs-3955	110	108	-	-	PUNCT
gs-3955	110	109	promoting	promote	VERB
gs-3955	110	110	features	feature	NOUN
gs-3955	110	111	include	include	VERB
gs-3955	110	112	:	:	PUNCT
gs-3955	110	113	●	●	PUNCT
gs-3955	110	114	seasonal	seasonal	ADJ
gs-3955	110	115	metabolic	metabolic	NOUN
gs-3955	110	116	plasticity	plasticity	NOUN
gs-3955	110	117	(	(	PUNCT
gs-3955	110	118	gao	gao	PROPN
gs-3955	110	119	et	et	PROPN
gs-3955	110	120	al	al	PROPN
gs-3955	110	121	.	.	PROPN
gs-3955	110	122	,	,	PUNCT
gs-3955	110	123	2022	2022	NUM
gs-3955	110	124	)	)	PUNCT
gs-3955	110	125	●	●	NUM
gs-3955	110	126	extended	extend	VERB
gs-3955	110	127	hibernation	hibernation	NOUN
gs-3955	110	128	capacity	capacity	NOUN
gs-3955	110	129	(	(	PUNCT
gs-3955	110	130	storey	storey	NOUN
gs-3955	110	131	&	&	CCONJ
gs-3955	110	132	storey	storey	PROPN
gs-3955	110	133	,	,	PUNCT
gs-3955	110	134	2019	2019	NUM
gs-3955	110	135	)	)	PUNCT
gs-3955	110	136	●	●	NUM
gs-3955	110	137	proteome	proteome	ADJ
gs-3955	110	138	stability	stability	NOUN
gs-3955	110	139	mechanisms	mechanism	NOUN
gs-3955	110	140	(	(	PUNCT
gs-3955	110	141	lópez	lópez	NOUN
gs-3955	110	142	-	-	PUNCT
gs-3955	110	143	otín	otín	NOUN
gs-3955	110	144	et	et	PROPN
gs-3955	110	145	al	al	PROPN
gs-3955	110	146	.	.	PROPN
gs-3955	110	147	,	,	PUNCT
gs-3955	110	148	2021	2021	NUM
gs-3955	110	149	)	)	PUNCT
gs-3955	110	150	georgian	georgian	ADJ
gs-3955	110	151	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	110	152	მეცნიერები	მეცნიერები	NOUN
gs-3955	110	153	ტ.	ტ.	VERB
gs-3955	110	154	7	7	NUM
gs-3955	110	155	n	n	PRON
gs-3955	110	156	3	3	NUM
gs-3955	110	157	,	,	PUNCT
gs-3955	110	158	2025	2025	NUM
gs-3955	110	159	357	357	NUM
gs-3955	110	160	genomic	genomic	ADJ
gs-3955	110	161	insights	insight	NOUN
gs-3955	110	162	sequencing	sequence	VERB
gs-3955	110	163	reveals	reveal	NOUN
gs-3955	110	164	:	:	PUNCT
gs-3955	110	165	●	●	PUNCT
gs-3955	110	166	unique	unique	ADJ
gs-3955	110	167	dna	dna	NOUN
gs-3955	110	168	repair	repair	NOUN
gs-3955	110	169	variants	variant	NOUN
gs-3955	110	170	(	(	PUNCT
gs-3955	110	171	nowoshilow	nowoshilow	PROPN
gs-3955	110	172	et	et	PROPN
gs-3955	110	173	al	al	PROPN
gs-3955	110	174	.	.	PROPN
gs-3955	110	175	,	,	PUNCT
gs-3955	110	176	2018	2018	NUM
gs-3955	110	177	)	)	PUNCT
gs-3955	110	178	●	●	NUM
gs-3955	110	179	autophagy	autophagy	ADJ
gs-3955	110	180	gene	gene	NOUN
gs-3955	110	181	specializations	specialization	NOUN
gs-3955	110	182	(	(	PUNCT
gs-3955	110	183	yun	yun	PROPN
gs-3955	110	184	et	et	PROPN
gs-3955	110	185	al	al	PROPN
gs-3955	110	186	.	.	PROPN
gs-3955	110	187	,	,	PUNCT
gs-3955	110	188	2020	2020	NUM
gs-3955	110	189	)	)	PUNCT
gs-3955	110	190	●	●	NUM
gs-3955	110	191	conserved	conserve	VERB
gs-3955	110	192	telomeric	telomeric	ADJ
gs-3955	110	193	sequences	sequence	NOUN
gs-3955	110	194	(	(	PUNCT
gs-3955	110	195	gorbunova	gorbunova	PROPN
gs-3955	110	196	et	et	PROPN
gs-3955	110	197	al	al	PROPN
gs-3955	110	198	.	.	PROPN
gs-3955	110	199	,	,	PUNCT
gs-3955	110	200	2022	2022	NUM
gs-3955	110	201	)	)	PUNCT
gs-3955	110	202	epigenetic	epigenetic	ADJ
gs-3955	110	203	regulation	regulation	NOUN
gs-3955	110	204	key	key	ADJ
gs-3955	110	205	findings	finding	NOUN
gs-3955	110	206	include	include	VERB
gs-3955	110	207	:	:	PUNCT
gs-3955	110	208	●	●	PUNCT
gs-3955	110	209	dna	dna	PROPN
gs-3955	110	210	methylation	methylation	NOUN
gs-3955	110	211	stability	stability	NOUN
gs-3955	110	212	(	(	PUNCT
gs-3955	110	213	ma	ma	PROPN
gs-3955	110	214	et	et	PROPN
gs-3955	110	215	al	al	PROPN
gs-3955	110	216	.	.	PROPN
gs-3955	110	217	,	,	PUNCT
gs-3955	110	218	2021	2021	NUM
gs-3955	110	219	)	)	PUNCT
gs-3955	110	220	●	●	PUNCT
gs-3955	110	221	regeneration	regeneration	NOUN
gs-3955	110	222	-	-	PUNCT
gs-3955	110	223	associated	associate	VERB
gs-3955	110	224	mirnas	mirna	NOUN
gs-3955	110	225	(	(	PUNCT
gs-3955	110	226	tanaka	tanaka	PROPN
gs-3955	110	227	,	,	PUNCT
gs-3955	110	228	2023	2023	NUM
gs-3955	110	229	)	)	PUNCT
gs-3955	110	230	●	●	PUNCT
gs-3955	110	231	slow	slow	ADJ
gs-3955	110	232	epigenetic	epigenetic	ADJ
gs-3955	110	233	aging	aging	NOUN
gs-3955	110	234	(	(	PUNCT
gs-3955	110	235	horvath	horvath	PROPN
gs-3955	110	236	et	et	PROPN
gs-3955	110	237	al	al	PROPN
gs-3955	110	238	.	.	PROPN
gs-3955	110	239	,	,	PUNCT
gs-3955	110	240	2022	2022	NUM
gs-3955	110	241	)	)	PUNCT
gs-3955	110	242	centenarian	centenarian	ADJ
gs-3955	110	243	chelonians	chelonian	NOUN
gs-3955	110	244	:	:	PUNCT
gs-3955	110	245	emydoidea	emydoidea	NOUN
gs-3955	110	246	blandingii	blandingii	NOUN
gs-3955	110	247	and	and	CCONJ
gs-3955	110	248	terrapene	terrapene	ADV
gs-3955	110	249	carolina	carolina	PROPN
gs-3955	110	250	these	these	DET
gs-3955	110	251	turtle	turtle	NOUN
gs-3955	110	252	species	specie	NOUN
gs-3955	110	253	achieve	achieve	VERB
gs-3955	110	254	80	80	NUM
gs-3955	110	255	-	-	SYM
gs-3955	110	256	150	150	NUM
gs-3955	110	257	year	year	NOUN
gs-3955	110	258	lifespans	lifespan	NOUN
gs-3955	110	259	(	(	PUNCT
gs-3955	110	260	reinke	reinke	PROPN
gs-3955	110	261	et	et	PROPN
gs-3955	110	262	al	al	PROPN
gs-3955	110	263	.	.	PROPN
gs-3955	110	264	,	,	PUNCT
gs-3955	110	265	2022	2022	NUM
gs-3955	110	266	)	)	PUNCT
gs-3955	110	267	through	through	ADP
gs-3955	110	268	remarkable	remarkable	ADJ
gs-3955	110	269	adaptations	adaptation	NOUN
gs-3955	110	270	against	against	ADP
gs-3955	110	271	aging	age	VERB
gs-3955	110	272	(	(	PUNCT
gs-3955	110	273	da	da	PROPN
gs-3955	110	274	silva	silva	PROPN
gs-3955	110	275	et	et	PROPN
gs-3955	110	276	al	al	PROPN
gs-3955	110	277	.	.	PROPN
gs-3955	110	278	,	,	PUNCT
gs-3955	110	279	2020	2020	NUM
gs-3955	110	280	)	)	PUNCT
gs-3955	110	281	.	.	PUNCT
gs-3955	111	1	metabolic	metabolic	NOUN
gs-3955	111	2	innovations	innovation	NOUN
gs-3955	111	3	key	key	ADJ
gs-3955	111	4	features	feature	NOUN
gs-3955	111	5	include	include	VERB
gs-3955	111	6	:	:	PUNCT
gs-3955	111	7	●	●	PUNCT
gs-3955	111	8	extremely	extremely	ADV
gs-3955	111	9	low	low	ADJ
gs-3955	111	10	resting	rest	VERB
gs-3955	111	11	metabolism	metabolism	NOUN
gs-3955	111	12	(	(	PUNCT
gs-3955	111	13	congdon	congdon	PROPN
gs-3955	111	14	et	et	PROPN
gs-3955	111	15	al	al	PROPN
gs-3955	111	16	.	.	PROPN
gs-3955	111	17	,	,	PUNCT
gs-3955	111	18	2019	2019	NUM
gs-3955	111	19	)	)	PUNCT
gs-3955	111	20	●	●	NUM
gs-3955	111	21	prolonged	prolong	VERB
gs-3955	111	22	hibernation	hibernation	NOUN
gs-3955	111	23	capability	capability	NOUN
gs-3955	111	24	(	(	PUNCT
gs-3955	111	25	storey	storey	NOUN
gs-3955	111	26	&	&	CCONJ
gs-3955	111	27	storey	storey	PROPN
gs-3955	111	28	,	,	PUNCT
gs-3955	111	29	2021	2021	NUM
gs-3955	111	30	)	)	PUNCT
gs-3955	111	31	●	●	NUM
gs-3955	111	32	hypoxia	hypoxia	NOUN
gs-3955	111	33	tolerance	tolerance	NOUN
gs-3955	111	34	mechanisms	mechanism	NOUN
gs-3955	111	35	(	(	PUNCT
gs-3955	111	36	milton	milton	PROPN
gs-3955	111	37	&	&	CCONJ
gs-3955	111	38	prentice	prentice	PROPN
gs-3955	111	39	,	,	PUNCT
gs-3955	111	40	2022	2022	NUM
gs-3955	111	41	)	)	PUNCT
gs-3955	111	42	antioxidant	antioxidant	NOUN
gs-3955	111	43	systems	system	NOUN
gs-3955	111	44	unique	unique	ADJ
gs-3955	111	45	protections	protection	NOUN
gs-3955	111	46	involve	involve	VERB
gs-3955	111	47	:	:	PUNCT
gs-3955	111	48	●	●	NUM
gs-3955	111	49	enhanced	enhanced	ADJ
gs-3955	111	50	sod	sod	NOUN
gs-3955	111	51	/	/	SYM
gs-3955	111	52	catalase	catalase	NOUN
gs-3955	111	53	activity	activity	NOUN
gs-3955	111	54	(	(	PUNCT
gs-3955	111	55	lutz	lutz	PROPN
gs-3955	111	56	et	et	PROPN
gs-3955	111	57	al	al	PROPN
gs-3955	111	58	.	.	PROPN
gs-3955	111	59	,	,	PUNCT
gs-3955	111	60	2020	2020	NUM
gs-3955	111	61	)	)	PUNCT
gs-3955	111	62	●	●	NUM
gs-3955	111	63	endogenous	endogenous	ADJ
gs-3955	111	64	antioxidant	antioxidant	ADJ
gs-3955	111	65	reservoirs	reservoir	NOUN
gs-3955	111	66	(	(	PUNCT
gs-3955	111	67	finkel	finkel	PROPN
gs-3955	111	68	&	&	CCONJ
gs-3955	111	69	holbrook	holbrook	PROPN
gs-3955	111	70	,	,	PUNCT
gs-3955	111	71	2021	2021	NUM
gs-3955	111	72	)	)	PUNCT
gs-3955	111	73	●	●	NUM
gs-3955	111	74	advanced	advanced	ADJ
gs-3955	111	75	dna	dna	PROPN
gs-3955	111	76	repair	repair	NOUN
gs-3955	111	77	systems	system	NOUN
gs-3955	111	78	(	(	PUNCT
gs-3955	111	79	gorbunova	gorbunova	PROPN
gs-3955	111	80	et	et	PROPN
gs-3955	111	81	al	al	PROPN
gs-3955	111	82	.	.	PROPN
gs-3955	111	83	,	,	PUNCT
gs-3955	111	84	2022	2022	NUM
gs-3955	111	85	)	)	PUNCT
gs-3955	111	86	immune	immune	ADJ
gs-3955	111	87	adaptations	adaptation	NOUN
gs-3955	111	88	notable	notable	ADJ
gs-3955	111	89	characteristics	characteristic	NOUN
gs-3955	111	90	:	:	PUNCT
gs-3955	111	91	●	●	NUM
gs-3955	111	92	robust	robust	ADJ
gs-3955	111	93	innate	innate	ADJ
gs-3955	111	94	immunity	immunity	NOUN
gs-3955	111	95	(	(	PUNCT
gs-3955	111	96	zimmerman	zimmerman	PROPN
gs-3955	111	97	et	et	PROPN
gs-3955	111	98	al	al	PROPN
gs-3955	111	99	.	.	PROPN
gs-3955	111	100	,	,	PUNCT
gs-3955	111	101	2020	2020	NUM
gs-3955	111	102	)	)	PUNCT
gs-3955	111	103	●	●	NUM
gs-3955	111	104	unique	unique	ADJ
gs-3955	111	105	immune	immune	ADJ
gs-3955	111	106	gene	gene	NOUN
gs-3955	111	107	expression	expression	NOUN
gs-3955	111	108	(	(	PUNCT
gs-3955	111	109	patel	patel	PROPN
gs-3955	111	110	et	et	PROPN
gs-3955	111	111	al	al	PROPN
gs-3955	111	112	.	.	PROPN
gs-3955	111	113	,	,	PUNCT
gs-3955	111	114	2021	2021	NUM
gs-3955	111	115	)	)	PUNCT
gs-3955	111	116	●	●	PUNCT
gs-3955	111	117	anti	anti	ADJ
gs-3955	111	118	-	-	ADJ
gs-3955	111	119	inflammatory	inflammatory	ADJ
gs-3955	111	120	phenotypes	phenotype	NOUN
gs-3955	111	121	(	(	PUNCT
gs-3955	111	122	franceschi	franceschi	PROPN
gs-3955	111	123	et	et	PROPN
gs-3955	111	124	al	al	PROPN
gs-3955	111	125	.	.	PROPN
gs-3955	111	126	,	,	PUNCT
gs-3955	111	127	2022	2022	NUM
gs-3955	111	128	)	)	PUNCT
gs-3955	111	129	genomic	genomic	NOUN
gs-3955	111	130	foundations	foundation	NOUN
gs-3955	111	131	research	research	NOUN
gs-3955	111	132	reveals	reveal	VERB
gs-3955	111	133	:	:	PUNCT
gs-3955	111	134	●	●	PUNCT
gs-3955	111	135	dna	dna	PROPN
gs-3955	111	136	repair	repair	NOUN
gs-3955	111	137	gene	gene	NOUN
gs-3955	111	138	variants	variant	NOUN
gs-3955	111	139	(	(	PUNCT
gs-3955	111	140	tian	tian	PROPN
gs-3955	111	141	et	et	PROPN
gs-3955	111	142	al	al	PROPN
gs-3955	111	143	.	.	PROPN
gs-3955	111	144	,	,	PUNCT
gs-3955	111	145	2022	2022	NUM
gs-3955	111	146	)	)	PUNCT
gs-3955	111	147	georgian	georgian	ADJ
gs-3955	111	148	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	111	149	მეცნიერები	მეცნიერები	NOUN
gs-3955	111	150	ტ.	ტ.	VERB
gs-3955	111	151	7	7	NUM
gs-3955	111	152	n	n	PRON
gs-3955	111	153	3	3	NUM
gs-3955	111	154	,	,	PUNCT
gs-3955	111	155	2025	2025	NUM
gs-3955	111	156	358	358	NUM
gs-3955	111	157	●	●	NUM
gs-3955	111	158	telomerase	telomerase	NOUN
gs-3955	111	159	regulation	regulation	NOUN
gs-3955	111	160	(	(	PUNCT
gs-3955	111	161	shay	shay	PROPN
gs-3955	111	162	&	&	CCONJ
gs-3955	111	163	wright	wright	PROPN
gs-3955	111	164	,	,	PUNCT
gs-3955	111	165	2021	2021	NUM
gs-3955	111	166	)	)	PUNCT
gs-3955	111	167	●	●	NUM
gs-3955	111	168	conserved	conserve	VERB
gs-3955	111	169	longevity	longevity	NOUN
gs-3955	111	170	genes	gene	NOUN
gs-3955	111	171	(	(	PUNCT
gs-3955	111	172	de	de	X
gs-3955	111	173	magalhães	magalhães	NOUN
gs-3955	111	174	,	,	PUNCT
gs-3955	111	175	2023	2023	NUM
gs-3955	111	176	)	)	PUNCT
gs-3955	111	177	epigenetic	epigenetic	ADJ
gs-3955	111	178	features	feature	VERB
gs-3955	111	179	key	key	ADJ
gs-3955	111	180	discoveries	discovery	NOUN
gs-3955	111	181	:	:	PUNCT
gs-3955	111	182	●	●	NUM
gs-3955	111	183	stable	stable	ADJ
gs-3955	111	184	methylation	methylation	NOUN
gs-3955	111	185	patterns	pattern	NOUN
gs-3955	111	186	(	(	PUNCT
gs-3955	111	187	horvath	horvath	PROPN
gs-3955	111	188	et	et	PROPN
gs-3955	111	189	al	al	PROPN
gs-3955	111	190	.	.	PROPN
gs-3955	111	191	,	,	PUNCT
gs-3955	111	192	2022	2022	NUM
gs-3955	111	193	)	)	PUNCT
gs-3955	111	194	●	●	PUNCT
gs-3955	111	195	distinct	distinct	ADJ
gs-3955	111	196	mirna	mirna	NOUN
gs-3955	111	197	profiles	profile	NOUN
gs-3955	111	198	(	(	PUNCT
gs-3955	111	199	smith	smith	PROPN
gs-3955	111	200	et	et	PROPN
gs-3955	111	201	al	al	PROPN
gs-3955	111	202	.	.	PROPN
gs-3955	111	203	,	,	PUNCT
gs-3955	111	204	2023	2023	NUM
gs-3955	111	205	)	)	PUNCT
gs-3955	112	1	●	●	PUNCT
gs-3955	112	2	slow	slow	ADJ
gs-3955	112	3	epigenetic	epigenetic	ADJ
gs-3955	112	4	clocks	clock	NOUN
gs-3955	112	5	(	(	PUNCT
gs-3955	112	6	jones	jones	PROPN
gs-3955	112	7	et	et	PROPN
gs-3955	112	8	al	al	PROPN
gs-3955	112	9	.	.	PROPN
gs-3955	112	10	,	,	PUNCT
gs-3955	112	11	2021	2021	NUM
gs-3955	112	12	)	)	PUNCT
gs-3955	112	13	the	the	DET
gs-3955	112	14	rougheye	rougheye	NOUN
gs-3955	112	15	rockfish	rockfish	NOUN
gs-3955	112	16	(	(	PUNCT
gs-3955	112	17	sebastes	sebaste	NOUN
gs-3955	112	18	aleutianus	aleutianus	NOUN
gs-3955	112	19	):	):	PUNCT
gs-3955	112	20	marine	marine	ADJ
gs-3955	112	21	longevity	longevity	NOUN
gs-3955	112	22	champion	champion	NOUN
gs-3955	112	23	this	this	DET
gs-3955	112	24	fish	fish	NOUN
gs-3955	112	25	species	species	NOUN
gs-3955	112	26	surpasses	surpass	VERB
gs-3955	112	27	200	200	NUM
gs-3955	112	28	years	year	NOUN
gs-3955	112	29	(	(	PUNCT
gs-3955	112	30	munk	munk	NOUN
gs-3955	112	31	,	,	PUNCT
gs-3955	112	32	2001	2001	NUM
gs-3955	112	33	)	)	PUNCT
gs-3955	112	34	while	while	SCONJ
gs-3955	112	35	maintaining	maintain	VERB
gs-3955	112	36	late	late	ADJ
gs-3955	112	37	-	-	PUNCT
gs-3955	112	38	life	life	NOUN
gs-3955	112	39	fertility	fertility	NOUN
gs-3955	112	40	(	(	PUNCT
gs-3955	112	41	love	love	NOUN
gs-3955	112	42	et	et	PROPN
gs-3955	112	43	al	al	PROPN
gs-3955	112	44	.	.	PROPN
gs-3955	112	45	,	,	PUNCT
gs-3955	112	46	2002	2002	NUM
gs-3955	112	47	)	)	PUNCT
gs-3955	112	48	.	.	PUNCT
gs-3955	113	1	metabolic	metabolic	NOUN
gs-3955	113	2	strategies	strategy	NOUN
gs-3955	113	3	unique	unique	ADJ
gs-3955	113	4	adaptations	adaptation	NOUN
gs-3955	113	5	:	:	PUNCT
gs-3955	113	6	●	●	PUNCT
gs-3955	113	7	extremely	extremely	ADV
gs-3955	113	8	low	low	ADJ
gs-3955	113	9	metabolic	metabolic	NOUN
gs-3955	113	10	rates	rate	NOUN
gs-3955	113	11	(	(	PUNCT
gs-3955	113	12	hurst	hurst	PROPN
gs-3955	113	13	et	et	PROPN
gs-3955	113	14	al	al	PROPN
gs-3955	113	15	.	.	PROPN
gs-3955	113	16	,	,	PUNCT
gs-3955	113	17	2010	2010	NUM
gs-3955	113	18	)	)	PUNCT
gs-3955	113	19	●	●	PUNCT
gs-3955	113	20	efficient	efficient	ADJ
gs-3955	113	21	mitochondrial	mitochondrial	ADJ
gs-3955	113	22	respiration	respiration	NOUN
gs-3955	113	23	(	(	PUNCT
gs-3955	113	24	o'brien	o'brien	PROPN
gs-3955	113	25	et	et	PROPN
gs-3955	113	26	al	al	PROPN
gs-3955	113	27	.	.	PROPN
gs-3955	113	28	,	,	PUNCT
gs-3955	113	29	2018	2018	NUM
gs-3955	113	30	)	)	PUNCT
gs-3955	113	31	●	●	PUNCT
gs-3955	113	32	minimal	minimal	ADJ
gs-3955	113	33	ros	ros	PROPN
gs-3955	113	34	production	production	NOUN
gs-3955	113	35	(	(	PUNCT
gs-3955	113	36	treberg	treberg	NOUN
gs-3955	113	37	et	et	PROPN
gs-3955	113	38	al	al	PROPN
gs-3955	113	39	.	.	PROPN
gs-3955	113	40	,	,	PUNCT
gs-3955	113	41	2016	2016	NUM
gs-3955	113	42	)	)	PUNCT
gs-3955	113	43	reproductive	reproductive	ADJ
gs-3955	113	44	longevity	longevity	NOUN
gs-3955	113	45	notable	notable	ADJ
gs-3955	113	46	features	feature	NOUN
gs-3955	113	47	:	:	PUNCT
gs-3955	113	48	●	●	PUNCT
gs-3955	113	49	>	>	SYM
gs-3955	113	50	150	150	NUM
gs-3955	113	51	year	year	NOUN
gs-3955	113	52	fertility	fertility	NOUN
gs-3955	113	53	(	(	PUNCT
gs-3955	113	54	berkeley	berkeley	PROPN
gs-3955	113	55	et	et	PROPN
gs-3955	113	56	al	al	PROPN
gs-3955	113	57	.	.	PROPN
gs-3955	113	58	,	,	PUNCT
gs-3955	113	59	2004	2004	NUM
gs-3955	113	60	)	)	PUNCT
gs-3955	113	61	●	●	PUNCT
gs-3955	113	62	late	late	ADJ
gs-3955	113	63	-	-	PUNCT
gs-3955	113	64	life	life	NOUN
gs-3955	113	65	fecundity	fecundity	NOUN
gs-3955	113	66	(	(	PUNCT
gs-3955	113	67	sogard	sogard	NOUN
gs-3955	113	68	et	et	PROPN
gs-3955	113	69	al	al	PROPN
gs-3955	113	70	.	.	PROPN
gs-3955	113	71	,	,	PUNCT
gs-3955	113	72	2008	2008	NUM
gs-3955	113	73	)	)	PUNCT
gs-3955	113	74	●	●	NUM
gs-3955	113	75	gamete	gamete	ADJ
gs-3955	113	76	quality	quality	NOUN
gs-3955	113	77	maintenance	maintenance	NOUN
gs-3955	113	78	(	(	PUNCT
gs-3955	113	79	hamel	hamel	PROPN
gs-3955	113	80	et	et	PROPN
gs-3955	113	81	al	al	PROPN
gs-3955	113	82	.	.	PROPN
gs-3955	113	83	,	,	PUNCT
gs-3955	113	84	2020	2020	NUM
gs-3955	113	85	)	)	PUNCT
gs-3955	113	86	protective	protective	ADJ
gs-3955	113	87	mechanisms	mechanism	NOUN
gs-3955	113	88	defensive	defensive	ADJ
gs-3955	113	89	systems	system	NOUN
gs-3955	113	90	include	include	VERB
gs-3955	113	91	:	:	PUNCT
gs-3955	113	92	●	●	PUNCT
gs-3955	113	93	potent	potent	ADJ
gs-3955	113	94	antioxidant	antioxidant	NOUN
gs-3955	113	95	defenses	defense	NOUN
gs-3955	113	96	(	(	PUNCT
gs-3955	113	97	gerdes	gerde	NOUN
gs-3955	113	98	et	et	PROPN
gs-3955	113	99	al	al	PROPN
gs-3955	113	100	.	.	PROPN
gs-3955	113	101	,	,	PUNCT
gs-3955	113	102	2019	2019	NUM
gs-3955	113	103	)	)	PUNCT
gs-3955	113	104	●	●	PUNCT
gs-3955	113	105	effective	effective	ADJ
gs-3955	113	106	dna	dna	PROPN
gs-3955	113	107	repair	repair	NOUN
gs-3955	113	108	(	(	PUNCT
gs-3955	113	109	bowen	bowen	PROPN
gs-3955	113	110	et	et	PROPN
gs-3955	113	111	al	al	PROPN
gs-3955	113	112	.	.	PROPN
gs-3955	113	113	,	,	PUNCT
gs-3955	113	114	2021	2021	NUM
gs-3955	113	115	)	)	PUNCT
gs-3955	113	116	●	●	NUM
gs-3955	113	117	proteostasis	proteostasis	NOUN
gs-3955	113	118	maintenance	maintenance	NOUN
gs-3955	113	119	(	(	PUNCT
gs-3955	113	120	perez	perez	PROPN
gs-3955	113	121	-	-	PUNCT
gs-3955	113	122	costas	costas	PROPN
gs-3955	113	123	et	et	PROPN
gs-3955	113	124	al	al	PROPN
gs-3955	113	125	.	.	PROPN
gs-3955	113	126	,	,	PUNCT
gs-3955	113	127	2022	2022	NUM
gs-3955	113	128	)	)	PUNCT
gs-3955	113	129	genomic	genomic	NOUN
gs-3955	113	130	insights	insight	NOUN
gs-3955	113	131	sequencing	sequence	VERB
gs-3955	113	132	shows	show	NOUN
gs-3955	113	133	:	:	PUNCT
gs-3955	113	134	●	●	PUNCT
gs-3955	113	135	dna	dna	PROPN
gs-3955	113	136	repair	repair	NOUN
gs-3955	113	137	variants	variant	NOUN
gs-3955	113	138	(	(	PUNCT
gs-3955	113	139	ercc1	ercc1	PROPN
gs-3955	113	140	,	,	PUNCT
gs-3955	113	141	rad51	rad51	PROPN
gs-3955	113	142	)	)	PUNCT
gs-3955	113	143	(	(	PUNCT
gs-3955	113	144	hyde	hyde	PROPN
gs-3955	113	145	et	et	PROPN
gs-3955	113	146	al	al	PROPN
gs-3955	113	147	.	.	PROPN
gs-3955	113	148	,	,	PUNCT
gs-3955	113	149	2020	2020	NUM
gs-3955	113	150	)	)	PUNCT
gs-3955	113	151	●	●	NUM
gs-3955	113	152	telomerase	telomerase	NOUN
gs-3955	113	153	activity	activity	NOUN
gs-3955	113	154	(	(	PUNCT
gs-3955	113	155	klapper	klapper	NOUN
gs-3955	113	156	et	et	PROPN
gs-3955	113	157	al	al	PROPN
gs-3955	113	158	.	.	PROPN
gs-3955	113	159	,	,	PUNCT
gs-3955	113	160	2021	2021	NUM
gs-3955	113	161	)	)	PUNCT
gs-3955	113	162	●	●	NUM
gs-3955	113	163	longevity	longevity	NOUN
gs-3955	113	164	gene	gene	NOUN
gs-3955	113	165	conservation	conservation	NOUN
gs-3955	113	166	(	(	PUNCT
gs-3955	113	167	tor	tor	NOUN
gs-3955	113	168	,	,	PUNCT
gs-3955	113	169	foxo	foxo	PROPN
gs-3955	113	170	)	)	PUNCT
gs-3955	113	171	(	(	PUNCT
gs-3955	113	172	brunet	brunet	NOUN
gs-3955	113	173	et	et	PROPN
gs-3955	113	174	al	al	PROPN
gs-3955	113	175	.	.	PROPN
gs-3955	113	176	,	,	PUNCT
gs-3955	113	177	2022	2022	NUM
gs-3955	113	178	)	)	PUNCT
gs-3955	113	179	georgian	georgian	ADJ
gs-3955	113	180	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	113	181	მეცნიერები	მეცნიერები	NOUN
gs-3955	113	182	ტ.	ტ.	VERB
gs-3955	113	183	7	7	NUM
gs-3955	113	184	n	n	PRON
gs-3955	113	185	3	3	NUM
gs-3955	113	186	,	,	PUNCT
gs-3955	113	187	2025	2025	NUM
gs-3955	113	188	359	359	NUM
gs-3955	113	189	epigenetic	epigenetic	ADJ
gs-3955	113	190	regulation	regulation	NOUN
gs-3955	113	191	key	key	ADJ
gs-3955	113	192	findings	finding	NOUN
gs-3955	113	193	:	:	PUNCT
gs-3955	113	194	●	●	NUM
gs-3955	113	195	methylation	methylation	NOUN
gs-3955	113	196	stability	stability	NOUN
gs-3955	113	197	(	(	PUNCT
gs-3955	113	198	horvath	horvath	PROPN
gs-3955	113	199	et	et	PROPN
gs-3955	113	200	al	al	PROPN
gs-3955	113	201	.	.	PROPN
gs-3955	113	202	,	,	PUNCT
gs-3955	113	203	2022	2022	NUM
gs-3955	113	204	)	)	PUNCT
gs-3955	113	205	●	●	NUM
gs-3955	113	206	unique	unique	ADJ
gs-3955	113	207	mirna	mirna	NOUN
gs-3955	113	208	profiles	profile	NOUN
gs-3955	113	209	(	(	PUNCT
gs-3955	113	210	smith	smith	PROPN
gs-3955	113	211	et	et	PROPN
gs-3955	113	212	al	al	PROPN
gs-3955	113	213	.	.	PROPN
gs-3955	113	214	,	,	PUNCT
gs-3955	113	215	2023	2023	NUM
gs-3955	113	216	)	)	PUNCT
gs-3955	113	217	●	●	PUNCT
gs-3955	113	218	slow	slow	ADJ
gs-3955	113	219	epigenetic	epigenetic	ADJ
gs-3955	113	220	aging	aging	NOUN
gs-3955	113	221	(	(	PUNCT
gs-3955	113	222	jones	jones	PROPN
gs-3955	113	223	et	et	PROPN
gs-3955	113	224	al	al	PROPN
gs-3955	113	225	.	.	PROPN
gs-3955	113	226	,	,	PUNCT
gs-3955	113	227	2021	2021	NUM
gs-3955	113	228	)	)	PUNCT
gs-3955	113	229	the	the	DET
gs-3955	113	230	ocean	ocean	NOUN
gs-3955	113	231	quahog	quahog	PROPN
gs-3955	113	232	(	(	PUNCT
gs-3955	113	233	arctica	arctica	PROPN
gs-3955	113	234	islandica	islandica	PROPN
gs-3955	113	235	):	):	PUNCT
gs-3955	113	236	methuselah	methuselah	PROPN
gs-3955	113	237	of	of	ADP
gs-3955	113	238	mollusks	mollusk	NOUN
gs-3955	113	239	this	this	DET
gs-3955	113	240	bivalve	bivalve	PROPN
gs-3955	113	241	holds	hold	VERB
gs-3955	113	242	the	the	DET
gs-3955	113	243	animal	animal	NOUN
gs-3955	113	244	longevity	longevity	NOUN
gs-3955	113	245	record	record	NOUN
gs-3955	113	246	at	at	ADP
gs-3955	113	247	507	507	NUM
gs-3955	113	248	years	year	NOUN
gs-3955	113	249	(	(	PUNCT
gs-3955	113	250	butler	butler	PROPN
gs-3955	113	251	et	et	PROPN
gs-3955	113	252	al	al	PROPN
gs-3955	113	253	.	.	PROPN
gs-3955	113	254	,	,	PUNCT
gs-3955	113	255	2013	2013	NUM
gs-3955	113	256	)	)	PUNCT
gs-3955	113	257	with	with	ADP
gs-3955	113	258	minimal	minimal	ADJ
gs-3955	113	259	senescence	senescence	NOUN
gs-3955	113	260	(	(	PUNCT
gs-3955	113	261	abele	abele	PROPN
gs-3955	113	262	et	et	PROPN
gs-3955	113	263	al	al	PROPN
gs-3955	113	264	.	.	PROPN
gs-3955	113	265	,	,	PUNCT
gs-3955	113	266	2008	2008	NUM
gs-3955	113	267	)	)	PUNCT
gs-3955	113	268	.	.	PUNCT
gs-3955	114	1	extreme	extreme	ADJ
gs-3955	114	2	adaptations	adaptation	NOUN
gs-3955	114	3	key	key	ADJ
gs-3955	114	4	features	feature	NOUN
gs-3955	114	5	:	:	PUNCT
gs-3955	114	6	●	●	PUNCT
gs-3955	114	7	ultra	ultra	ADJ
gs-3955	114	8	-	-	ADJ
gs-3955	114	9	low	low	ADJ
gs-3955	114	10	metabolism	metabolism	NOUN
gs-3955	114	11	(	(	PUNCT
gs-3955	114	12	ballesta	ballesta	NOUN
gs-3955	114	13	-	-	PUNCT
gs-3955	114	14	artero	artero	NOUN
gs-3955	114	15	et	et	PROPN
gs-3955	114	16	al	al	PROPN
gs-3955	114	17	.	.	PROPN
gs-3955	114	18	,	,	PUNCT
gs-3955	114	19	2017	2017	NUM
gs-3955	114	20	)	)	PUNCT
gs-3955	114	21	●	●	NUM
gs-3955	114	22	prolonged	prolong	VERB
gs-3955	114	23	anoxia	anoxia	NOUN
gs-3955	114	24	survival	survival	NOUN
gs-3955	114	25	(	(	PUNCT
gs-3955	114	26	strahl	strahl	X
gs-3955	114	27	et	et	PROPN
gs-3955	114	28	al	al	PROPN
gs-3955	114	29	.	.	PROPN
gs-3955	114	30	,	,	PUNCT
gs-3955	114	31	2011	2011	NUM
gs-3955	114	32	)	)	PUNCT
gs-3955	114	33	●	●	PUNCT
gs-3955	114	34	minimal	minimal	ADJ
gs-3955	114	35	ros	ros	NOUN
gs-3955	114	36	generation	generation	NOUN
gs-3955	114	37	(	(	PUNCT
gs-3955	114	38	ungvari	ungvari	X
gs-3955	114	39	et	et	PROPN
gs-3955	114	40	al	al	PROPN
gs-3955	114	41	.	.	PROPN
gs-3955	114	42	,	,	PUNCT
gs-3955	114	43	2013	2013	NUM
gs-3955	114	44	)	)	PUNCT
gs-3955	114	45	cellular	cellular	ADJ
gs-3955	114	46	resilience	resilience	NOUN
gs-3955	114	47	notable	notable	ADJ
gs-3955	114	48	characteristics	characteristic	NOUN
gs-3955	114	49	:	:	PUNCT
gs-3955	114	50	●	●	NUM
gs-3955	114	51	apoptosis	apoptosis	NOUN
gs-3955	114	52	resistance	resistance	NOUN
gs-3955	114	53	(	(	PUNCT
gs-3955	114	54	gruber	gruber	PROPN
gs-3955	114	55	et	et	PROPN
gs-3955	114	56	al	al	PROPN
gs-3955	114	57	.	.	PROPN
gs-3955	114	58	,	,	PUNCT
gs-3955	114	59	2015	2015	NUM
gs-3955	114	60	)	)	PUNCT
gs-3955	114	61	●	●	PUNCT
gs-3955	114	62	dna	dna	NOUN
gs-3955	114	63	repair	repair	NOUN
gs-3955	114	64	efficiency	efficiency	NOUN
gs-3955	114	65	(	(	PUNCT
gs-3955	114	66	sussarellu	sussarellu	NOUN
gs-3955	114	67	et	et	NOUN
gs-3955	114	68	al	al	PROPN
gs-3955	114	69	.	.	PROPN
gs-3955	114	70	,	,	PUNCT
gs-3955	114	71	2020	2020	NUM
gs-3955	114	72	)	)	PUNCT
gs-3955	114	73	●	●	X
gs-3955	114	74	proteome	proteome	ADJ
gs-3955	114	75	stability	stability	NOUN
gs-3955	114	76	(	(	PUNCT
gs-3955	114	77	treaster	treaster	NOUN
gs-3955	114	78	et	et	PROPN
gs-3955	114	79	al	al	PROPN
gs-3955	114	80	.	.	PROPN
gs-3955	114	81	,	,	PUNCT
gs-3955	114	82	2014	2014	NUM
gs-3955	114	83	)	)	PUNCT
gs-3955	114	84	protective	protective	ADJ
gs-3955	114	85	systems	system	NOUN
gs-3955	114	86	defensive	defensive	ADJ
gs-3955	114	87	mechanisms	mechanism	NOUN
gs-3955	114	88	include	include	VERB
gs-3955	114	89	:	:	PUNCT
gs-3955	114	90	●	●	PUNCT
gs-3955	114	91	potent	potent	ADJ
gs-3955	114	92	antioxidant	antioxidant	NOUN
gs-3955	114	93	systems	system	NOUN
gs-3955	114	94	(	(	PUNCT
gs-3955	114	95	philipp	philipp	PROPN
gs-3955	114	96	et	et	PROPN
gs-3955	114	97	al	al	PROPN
gs-3955	114	98	.	.	PROPN
gs-3955	114	99	,	,	PUNCT
gs-3955	114	100	2012	2012	NUM
gs-3955	114	101	)	)	PUNCT
gs-3955	114	102	●	●	NUM
gs-3955	114	103	thermostable	thermostable	NOUN
gs-3955	114	104	proteins	protein	NOUN
gs-3955	114	105	(	(	PUNCT
gs-3955	114	106	abele	abele	PROPN
gs-3955	114	107	et	et	PROPN
gs-3955	114	108	al	al	PROPN
gs-3955	114	109	.	.	PROPN
gs-3955	114	110	,	,	PUNCT
gs-3955	114	111	2010	2010	NUM
gs-3955	114	112	)	)	PUNCT
gs-3955	114	113	●	●	PUNCT
gs-3955	114	114	unique	unique	ADJ
gs-3955	114	115	membrane	membrane	NOUN
gs-3955	114	116	lipids	lipid	NOUN
gs-3955	114	117	(	(	PUNCT
gs-3955	114	118	munro	munro	PROPN
gs-3955	114	119	et	et	PROPN
gs-3955	114	120	al	al	PROPN
gs-3955	114	121	.	.	PROPN
gs-3955	114	122	,	,	PUNCT
gs-3955	114	123	2013	2013	NUM
gs-3955	114	124	)	)	PUNCT
gs-3955	114	125	genomic	genomic	NOUN
gs-3955	114	126	foundations	foundation	NOUN
gs-3955	114	127	research	research	NOUN
gs-3955	114	128	reveals	reveal	VERB
gs-3955	114	129	:	:	PUNCT
gs-3955	114	130	●	●	PUNCT
gs-3955	114	131	dna	dna	PROPN
gs-3955	114	132	repair	repair	NOUN
gs-3955	114	133	variants	variant	NOUN
gs-3955	114	134	(	(	PUNCT
gs-3955	114	135	ercc1	ercc1	PROPN
gs-3955	114	136	,	,	PUNCT
gs-3955	114	137	xpa	xpa	PROPN
gs-3955	114	138	)	)	PUNCT
gs-3955	114	139	(	(	PUNCT
gs-3955	114	140	tian	tian	PROPN
gs-3955	114	141	et	et	PROPN
gs-3955	114	142	al	al	PROPN
gs-3955	114	143	.	.	PROPN
gs-3955	114	144	,	,	PUNCT
gs-3955	114	145	2022	2022	NUM
gs-3955	114	146	)	)	PUNCT
gs-3955	114	147	●	●	NUM
gs-3955	114	148	apoptosis	apoptosis	NOUN
gs-3955	114	149	regulation	regulation	NOUN
gs-3955	114	150	(	(	PUNCT
gs-3955	114	151	bcl-2	bcl-2	NOUN
gs-3955	114	152	family	family	NOUN
gs-3955	114	153	)	)	PUNCT
gs-3955	114	154	(	(	PUNCT
gs-3955	114	155	sussarellu	sussarellu	NOUN
gs-3955	114	156	et	et	NOUN
gs-3955	114	157	al	al	PROPN
gs-3955	114	158	.	.	PROPN
gs-3955	114	159	,	,	PUNCT
gs-3955	114	160	2021	2021	NUM
gs-3955	114	161	)	)	PUNCT
gs-3955	114	162	●	●	NUM
gs-3955	114	163	longevity	longevity	NOUN
gs-3955	114	164	gene	gene	NOUN
gs-3955	114	165	conservation	conservation	NOUN
gs-3955	114	166	(	(	PUNCT
gs-3955	114	167	foxo	foxo	PROPN
gs-3955	114	168	,	,	PUNCT
gs-3955	114	169	sirt	sirt	PROPN
gs-3955	114	170	)	)	PUNCT
gs-3955	114	171	(	(	PUNCT
gs-3955	114	172	ungvari	ungvari	X
gs-3955	114	173	et	et	PROPN
gs-3955	114	174	al	al	PROPN
gs-3955	114	175	.	.	PROPN
gs-3955	114	176	,	,	PUNCT
gs-3955	114	177	2011	2011	NUM
gs-3955	114	178	)	)	PUNCT
gs-3955	114	179	epigenetic	epigenetic	ADJ
gs-3955	114	180	features	feature	VERB
gs-3955	114	181	key	key	ADJ
gs-3955	114	182	discoveries	discovery	NOUN
gs-3955	114	183	:	:	PUNCT
gs-3955	114	184	●	●	NUM
gs-3955	114	185	methylation	methylation	NOUN
gs-3955	114	186	stability	stability	NOUN
gs-3955	114	187	(	(	PUNCT
gs-3955	114	188	jansen	jansen	PROPN
gs-3955	114	189	et	et	PROPN
gs-3955	114	190	al	al	PROPN
gs-3955	114	191	.	.	PROPN
gs-3955	114	192	,	,	PUNCT
gs-3955	114	193	2020	2020	NUM
gs-3955	114	194	)	)	PUNCT
gs-3955	115	1	georgian	georgian	ADJ
gs-3955	115	2	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	115	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	115	4	ტ.	ტ.	VERB
gs-3955	115	5	7	7	NUM
gs-3955	115	6	n	n	DET
gs-3955	115	7	3	3	NUM
gs-3955	115	8	,	,	PUNCT
gs-3955	115	9	2025	2025	NUM
gs-3955	115	10	360	360	NUM
gs-3955	115	11	●	●	NUM
gs-3955	115	12	unique	unique	ADJ
gs-3955	115	13	mirna	mirna	NOUN
gs-3955	115	14	profiles	profile	NOUN
gs-3955	115	15	(	(	PUNCT
gs-3955	115	16	zhang	zhang	X
gs-3955	115	17	et	et	PROPN
gs-3955	115	18	al	al	PROPN
gs-3955	115	19	.	.	PROPN
gs-3955	115	20	,	,	PUNCT
gs-3955	115	21	2023	2023	NUM
gs-3955	115	22	)	)	PUNCT
gs-3955	115	23	●	●	PUNCT
gs-3955	115	24	slow	slow	ADJ
gs-3955	115	25	epigenetic	epigenetic	ADJ
gs-3955	115	26	clocks	clock	NOUN
gs-3955	115	27	(	(	PUNCT
gs-3955	115	28	horvath	horvath	PROPN
gs-3955	115	29	et	et	PROPN
gs-3955	115	30	al	al	PROPN
gs-3955	115	31	.	.	PROPN
gs-3955	115	32	,	,	PUNCT
gs-3955	115	33	2022	2022	NUM
gs-3955	115	34	)	)	PUNCT
gs-3955	115	35	the	the	DET
gs-3955	115	36	giant	giant	ADJ
gs-3955	115	37	red	red	PROPN
gs-3955	115	38	sea	sea	NOUN
gs-3955	115	39	urchin	urchin	NOUN
gs-3955	115	40	(	(	PUNCT
gs-3955	115	41	strongylocentrotus	strongylocentrotus	NOUN
gs-3955	115	42	franciscanus	franciscanu	NOUN
gs-3955	115	43	):	):	PUNCT
gs-3955	115	44	a	a	DET
gs-3955	115	45	marine	marine	ADJ
gs-3955	115	46	model	model	NOUN
gs-3955	115	47	of	of	ADP
gs-3955	115	48	extreme	extreme	ADJ
gs-3955	115	49	longevity	longevity	NOUN
gs-3955	115	50	the	the	DET
gs-3955	115	51	giant	giant	ADJ
gs-3955	115	52	red	red	PROPN
gs-3955	115	53	sea	sea	NOUN
gs-3955	115	54	urchin	urchin	NOUN
gs-3955	115	55	(	(	PUNCT
gs-3955	115	56	strongylocentrotus	strongylocentrotus	NOUN
gs-3955	115	57	franciscanus	franciscanu	NOUN
gs-3955	115	58	)	)	PUNCT
gs-3955	115	59	stands	stand	VERB
gs-3955	115	60	as	as	ADP
gs-3955	115	61	a	a	DET
gs-3955	115	62	remarkable	remarkable	ADJ
gs-3955	115	63	example	example	NOUN
gs-3955	115	64	of	of	ADP
gs-3955	115	65	negligible	negligible	ADJ
gs-3955	115	66	senescence	senescence	NOUN
gs-3955	115	67	among	among	ADP
gs-3955	115	68	marine	marine	ADJ
gs-3955	115	69	invertebrates	invertebrate	NOUN
gs-3955	115	70	,	,	PUNCT
gs-3955	115	71	with	with	ADP
gs-3955	115	72	documented	document	VERB
gs-3955	115	73	lifespans	lifespan	NOUN
gs-3955	115	74	exceeding	exceed	VERB
gs-3955	115	75	a	a	DET
gs-3955	115	76	century	century	NOUN
gs-3955	115	77	(	(	PUNCT
gs-3955	115	78	ebert	ebert	NOUN
gs-3955	115	79	&	&	CCONJ
gs-3955	115	80	southon	southon	PROPN
gs-3955	115	81	,	,	PUNCT
gs-3955	115	82	2003	2003	NUM
gs-3955	115	83	)	)	PUNCT
gs-3955	115	84	.	.	PUNCT
gs-3955	116	1	this	this	DET
gs-3955	116	2	echinoderm	echinoderm	NOUN
gs-3955	116	3	species	specie	NOUN
gs-3955	116	4	has	have	AUX
gs-3955	116	5	emerged	emerge	VERB
gs-3955	116	6	as	as	ADP
gs-3955	116	7	a	a	DET
gs-3955	116	8	particularly	particularly	ADV
gs-3955	116	9	valuable	valuable	ADJ
gs-3955	116	10	model	model	NOUN
gs-3955	116	11	organism	organism	NOUN
gs-3955	116	12	in	in	ADP
gs-3955	116	13	biogerontological	biogerontological	ADJ
gs-3955	116	14	research	research	NOUN
gs-3955	116	15	due	due	ADP
gs-3955	116	16	to	to	ADP
gs-3955	116	17	its	its	PRON
gs-3955	116	18	exceptional	exceptional	ADJ
gs-3955	116	19	combination	combination	NOUN
gs-3955	116	20	of	of	ADP
gs-3955	116	21	extreme	extreme	ADJ
gs-3955	116	22	longevity	longevity	NOUN
gs-3955	116	23	,	,	PUNCT
gs-3955	116	24	maintenance	maintenance	NOUN
gs-3955	116	25	of	of	ADP
gs-3955	116	26	reproductive	reproductive	ADJ
gs-3955	116	27	capacity	capacity	NOUN
gs-3955	116	28	in	in	ADP
gs-3955	116	29	advanced	advanced	ADJ
gs-3955	116	30	age	age	NOUN
gs-3955	116	31	,	,	PUNCT
gs-3955	116	32	and	and	CCONJ
gs-3955	116	33	absence	absence	NOUN
gs-3955	116	34	of	of	ADP
gs-3955	116	35	observable	observable	ADJ
gs-3955	116	36	senescent	senescent	ADJ
gs-3955	116	37	deterioration	deterioration	NOUN
gs-3955	116	38	(	(	PUNCT
gs-3955	116	39	bodnar	bodnar	NOUN
gs-3955	116	40	,	,	PUNCT
gs-3955	116	41	2013	2013	NUM
gs-3955	116	42	)	)	PUNCT
gs-3955	116	43	.	.	PUNCT
gs-3955	117	1	cellular	cellular	ADJ
gs-3955	117	2	and	and	CCONJ
gs-3955	117	3	molecular	molecular	ADJ
gs-3955	117	4	adaptations	adaptation	NOUN
gs-3955	117	5	for	for	ADP
gs-3955	117	6	longevity	longevity	NOUN
gs-3955	117	7	comprehensive	comprehensive	ADJ
gs-3955	117	8	investigations	investigation	NOUN
gs-3955	117	9	have	have	AUX
gs-3955	117	10	revealed	reveal	VERB
gs-3955	117	11	several	several	ADJ
gs-3955	117	12	fundamental	fundamental	ADJ
gs-3955	117	13	biological	biological	ADJ
gs-3955	117	14	adaptations	adaptation	NOUN
gs-3955	117	15	contributing	contribute	VERB
gs-3955	117	16	to	to	ADP
gs-3955	117	17	this	this	DET
gs-3955	117	18	species	specie	NOUN
gs-3955	117	19	'	'	PART
gs-3955	117	20	remarkable	remarkable	ADJ
gs-3955	117	21	lifespan	lifespan	NOUN
gs-3955	117	22	:	:	PUNCT
gs-3955	117	23	cellular	cellular	ADJ
gs-3955	117	24	maintenance	maintenance	NOUN
gs-3955	117	25	mechanisms	mechanism	NOUN
gs-3955	117	26	:	:	PUNCT
gs-3955	117	27	●	●	NUM
gs-3955	117	28	enhanced	enhanced	ADJ
gs-3955	117	29	proliferative	proliferative	ADJ
gs-3955	117	30	capacity	capacity	NOUN
gs-3955	117	31	of	of	ADP
gs-3955	117	32	somatic	somatic	ADJ
gs-3955	117	33	cells	cell	NOUN
gs-3955	117	34	throughout	throughout	ADP
gs-3955	117	35	the	the	DET
gs-3955	117	36	lifespan	lifespan	NOUN
gs-3955	117	37	(	(	PUNCT
gs-3955	117	38	bodnar	bodnar	PROPN
gs-3955	117	39	&	&	CCONJ
gs-3955	117	40	coffman	coffman	PROPN
gs-3955	117	41	,	,	PUNCT
gs-3955	117	42	2016	2016	NUM
gs-3955	117	43	)	)	PUNCT
gs-3955	117	44	●	●	PUNCT
gs-3955	117	45	remarkable	remarkable	ADJ
gs-3955	117	46	resistance	resistance	NOUN
gs-3955	117	47	to	to	ADP
gs-3955	117	48	neoplastic	neoplastic	ADJ
gs-3955	117	49	transformation	transformation	NOUN
gs-3955	117	50	,	,	PUNCT
gs-3955	117	51	with	with	ADP
gs-3955	117	52	virtually	virtually	ADV
gs-3955	117	53	no	no	PRON
gs-3955	117	54	reported	report	VERB
gs-3955	117	55	cases	case	NOUN
gs-3955	117	56	of	of	ADP
gs-3955	117	57	spontaneous	spontaneous	ADJ
gs-3955	117	58	tumors	tumor	NOUN
gs-3955	117	59	(	(	PUNCT
gs-3955	117	60	francis	francis	PROPN
gs-3955	117	61	et	et	PROPN
gs-3955	117	62	al	al	PROPN
gs-3955	117	63	.	.	PROPN
gs-3955	117	64	,	,	PUNCT
gs-3955	117	65	2019	2019	NUM
gs-3955	117	66	)	)	PUNCT
gs-3955	117	67	●	●	NUM
gs-3955	117	68	highly	highly	ADV
gs-3955	117	69	efficient	efficient	ADJ
gs-3955	117	70	dna	dna	PROPN
gs-3955	117	71	repair	repair	NOUN
gs-3955	117	72	systems	system	NOUN
gs-3955	117	73	maintaining	maintain	VERB
gs-3955	117	74	genomic	genomic	ADJ
gs-3955	117	75	integrity	integrity	NOUN
gs-3955	117	76	over	over	ADP
gs-3955	117	77	decades	decade	NOUN
gs-3955	117	78	(	(	PUNCT
gs-3955	117	79	sussarellu	sussarellu	X
gs-3955	117	80	et	et	NOUN
gs-3955	117	81	al	al	PROPN
gs-3955	117	82	.	.	PROPN
gs-3955	117	83	,	,	PUNCT
gs-3955	117	84	2020	2020	NUM
gs-3955	117	85	)	)	PUNCT
gs-3955	117	86	regenerative	regenerative	ADJ
gs-3955	117	87	capabilities	capability	NOUN
gs-3955	117	88	:	:	PUNCT
gs-3955	117	89	●	●	PUNCT
gs-3955	117	90	complete	complete	ADJ
gs-3955	117	91	regeneration	regeneration	NOUN
gs-3955	117	92	of	of	ADP
gs-3955	117	93	spines	spine	NOUN
gs-3955	117	94	and	and	CCONJ
gs-3955	117	95	tube	tube	NOUN
gs-3955	117	96	feet	foot	NOUN
gs-3955	117	97	following	follow	VERB
gs-3955	117	98	injury	injury	NOUN
gs-3955	117	99	(	(	PUNCT
gs-3955	117	100	dubois	dubois	PROPN
gs-3955	117	101	&	&	CCONJ
gs-3955	117	102	ameye	ameye	PROPN
gs-3955	117	103	,	,	PUNCT
gs-3955	117	104	2018	2018	NUM
gs-3955	117	105	)	)	PUNCT
gs-3955	117	106	●	●	NUM
gs-3955	117	107	visceral	visceral	ADJ
gs-3955	117	108	organ	organ	NOUN
gs-3955	117	109	regeneration	regeneration	NOUN
gs-3955	117	110	,	,	PUNCT
gs-3955	117	111	including	include	VERB
gs-3955	117	112	restoration	restoration	NOUN
gs-3955	117	113	of	of	ADP
gs-3955	117	114	digestive	digestive	ADJ
gs-3955	117	115	system	system	NOUN
gs-3955	117	116	components	component	NOUN
gs-3955	117	117	(	(	PUNCT
gs-3955	117	118	garcía	garcía	NOUN
gs-3955	117	119	-	-	PUNCT
gs-3955	117	120	arrarás	arrarás	NOUN
gs-3955	117	121	et	et	PROPN
gs-3955	117	122	al	al	PROPN
gs-3955	117	123	.	.	PROPN
gs-3955	117	124	,	,	PUNCT
gs-3955	117	125	2021	2021	NUM
gs-3955	117	126	)	)	PUNCT
gs-3955	117	127	●	●	NUM
gs-3955	117	128	scar	scar	NOUN
gs-3955	117	129	-	-	PUNCT
gs-3955	117	130	free	free	ADJ
gs-3955	117	131	wound	wound	NOUN
gs-3955	117	132	healing	healing	NOUN
gs-3955	117	133	through	through	ADP
gs-3955	117	134	modified	modify	VERB
gs-3955	117	135	tissue	tissue	NOUN
gs-3955	117	136	remodeling	remodeling	NOUN
gs-3955	117	137	processes	process	NOUN
gs-3955	117	138	(	(	PUNCT
gs-3955	117	139	wilkie	wilkie	PROPN
gs-3955	117	140	et	et	PROPN
gs-3955	117	141	al	al	PROPN
gs-3955	117	142	.	.	PROPN
gs-3955	117	143	,	,	PUNCT
gs-3955	117	144	2022	2022	NUM
gs-3955	117	145	)	)	PUNCT
gs-3955	117	146	protective	protective	ADJ
gs-3955	117	147	systems	system	NOUN
gs-3955	117	148	against	against	ADP
gs-3955	117	149	age	age	NOUN
gs-3955	117	150	-	-	PUNCT
gs-3955	117	151	related	relate	VERB
gs-3955	117	152	damage	damage	NOUN
gs-3955	117	153	the	the	DET
gs-3955	117	154	species	specie	NOUN
gs-3955	117	155	possesses	possess	VERB
gs-3955	117	156	multiple	multiple	ADJ
gs-3955	117	157	overlapping	overlap	VERB
gs-3955	117	158	defense	defense	NOUN
gs-3955	117	159	mechanisms	mechanism	NOUN
gs-3955	117	160	:	:	PUNCT
gs-3955	117	161	oxidative	oxidative	ADJ
gs-3955	117	162	stress	stress	NOUN
gs-3955	117	163	resistance	resistance	NOUN
gs-3955	117	164	:	:	PUNCT
gs-3955	117	165	●	●	NUM
gs-3955	117	166	potent	potent	ADJ
gs-3955	117	167	antioxidant	antioxidant	ADJ
gs-3955	117	168	enzyme	enzyme	NOUN
gs-3955	117	169	systems	system	NOUN
gs-3955	117	170	(	(	PUNCT
gs-3955	117	171	superoxide	superoxide	NOUN
gs-3955	117	172	dismutase	dismutase	NOUN
gs-3955	117	173	,	,	PUNCT
gs-3955	117	174	catalase	catalase	NOUN
gs-3955	117	175	)	)	PUNCT
gs-3955	117	176	(	(	PUNCT
gs-3955	117	177	philipp	philipp	NOUN
gs-3955	117	178	et	et	PROPN
gs-3955	117	179	al	al	PROPN
gs-3955	117	180	.	.	PROPN
gs-3955	117	181	,	,	PUNCT
gs-3955	117	182	2015	2015	NUM
gs-3955	117	183	)	)	PUNCT
gs-3955	117	184	●	●	NUM
gs-3955	117	185	thermostable	thermostable	ADJ
gs-3955	117	186	protein	protein	NOUN
gs-3955	117	187	structures	structure	NOUN
gs-3955	117	188	resistant	resistant	ADJ
gs-3955	117	189	to	to	ADP
gs-3955	117	190	denaturation	denaturation	NOUN
gs-3955	117	191	(	(	PUNCT
gs-3955	117	192	tomanek	tomanek	NOUN
gs-3955	117	193	,	,	PUNCT
gs-3955	117	194	2018	2018	NUM
gs-3955	117	195	)	)	PUNCT
gs-3955	117	196	●	●	NUM
gs-3955	117	197	unique	unique	ADJ
gs-3955	117	198	membrane	membrane	NOUN
gs-3955	117	199	lipid	lipid	NOUN
gs-3955	117	200	composition	composition	NOUN
gs-3955	117	201	conferring	confer	VERB
gs-3955	117	202	peroxidation	peroxidation	NOUN
gs-3955	117	203	resistance	resistance	NOUN
gs-3955	117	204	(	(	PUNCT
gs-3955	117	205	pernet	pernet	NOUN
gs-3955	117	206	et	et	PROPN
gs-3955	117	207	al	al	PROPN
gs-3955	117	208	.	.	PROPN
gs-3955	117	209	,	,	PUNCT
gs-3955	117	210	2020	2020	NUM
gs-3955	117	211	)	)	PUNCT
gs-3955	118	1	georgian	georgian	ADJ
gs-3955	118	2	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	118	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	118	4	ტ.	ტ.	VERB
gs-3955	118	5	7	7	NUM
gs-3955	118	6	n	n	PRON
gs-3955	118	7	3	3	NUM
gs-3955	118	8	,	,	PUNCT
gs-3955	118	9	2025	2025	NUM
gs-3955	118	10	361	361	NUM
gs-3955	118	11	genomic	genomic	ADJ
gs-3955	118	12	foundations	foundation	NOUN
gs-3955	118	13	of	of	ADP
gs-3955	118	14	longevity	longevity	NOUN
gs-3955	118	15	genome	genome	NOUN
gs-3955	118	16	sequencing	sequencing	NOUN
gs-3955	118	17	projects	project	NOUN
gs-3955	118	18	have	have	AUX
gs-3955	118	19	identified	identify	VERB
gs-3955	118	20	:	:	PUNCT
gs-3955	118	21	genetic	genetic	ADJ
gs-3955	118	22	specializations	specialization	NOUN
gs-3955	118	23	:	:	PUNCT
gs-3955	118	24	●	●	PUNCT
gs-3955	118	25	distinct	distinct	ADJ
gs-3955	118	26	variants	variant	NOUN
gs-3955	118	27	in	in	ADP
gs-3955	118	28	dna	dna	PROPN
gs-3955	118	29	repair	repair	NOUN
gs-3955	118	30	genes	gene	NOUN
gs-3955	118	31	(	(	PUNCT
gs-3955	118	32	ercc1	ercc1	PROPN
gs-3955	118	33	,	,	PUNCT
gs-3955	118	34	xpa	xpa	PROPN
gs-3955	118	35	)	)	PUNCT
gs-3955	118	36	(	(	PUNCT
gs-3955	118	37	tian	tian	PROPN
gs-3955	118	38	et	et	PROPN
gs-3955	118	39	al	al	PROPN
gs-3955	118	40	.	.	PROPN
gs-3955	118	41	,	,	PUNCT
gs-3955	118	42	2022	2022	NUM
gs-3955	118	43	)	)	PUNCT
gs-3955	118	44	●	●	NUM
gs-3955	118	45	modified	modify	VERB
gs-3955	118	46	apoptosis	apoptosis	NOUN
gs-3955	118	47	regulation	regulation	NOUN
gs-3955	118	48	through	through	ADP
gs-3955	118	49	bcl-2	bcl-2	VERB
gs-3955	118	50	family	family	NOUN
gs-3955	118	51	genes	gene	NOUN
gs-3955	118	52	(	(	PUNCT
gs-3955	118	53	robertson	robertson	PROPN
gs-3955	118	54	et	et	PROPN
gs-3955	118	55	al	al	PROPN
gs-3955	118	56	.	.	PROPN
gs-3955	118	57	,	,	PUNCT
gs-3955	118	58	2021	2021	NUM
gs-3955	118	59	)	)	PUNCT
gs-3955	118	60	●	●	NUM
gs-3955	118	61	conserved	conserve	VERB
gs-3955	118	62	sequences	sequence	NOUN
gs-3955	118	63	in	in	ADP
gs-3955	118	64	longevity	longevity	NOUN
gs-3955	118	65	-	-	PUNCT
gs-3955	118	66	associated	associate	VERB
gs-3955	118	67	genes	gene	NOUN
gs-3955	118	68	(	(	PUNCT
gs-3955	118	69	foxo	foxo	PROPN
gs-3955	118	70	,	,	PUNCT
gs-3955	118	71	sirt	sirt	PROPN
gs-3955	118	72	pathways	pathway	NOUN
gs-3955	118	73	)	)	PUNCT
gs-3955	118	74	(	(	PUNCT
gs-3955	118	75	ungvari	ungvari	X
gs-3955	118	76	et	et	PROPN
gs-3955	118	77	al	al	PROPN
gs-3955	118	78	.	.	PROPN
gs-3955	118	79	,	,	PUNCT
gs-3955	118	80	2013	2013	NUM
gs-3955	118	81	)	)	PUNCT
gs-3955	118	82	epigenetic	epigenetic	ADJ
gs-3955	118	83	regulation	regulation	NOUN
gs-3955	118	84	:	:	PUNCT
gs-3955	118	85	●	●	PUNCT
gs-3955	118	86	exceptional	exceptional	ADJ
gs-3955	118	87	stability	stability	NOUN
gs-3955	118	88	of	of	ADP
gs-3955	118	89	dna	dna	PROPN
gs-3955	118	90	methylation	methylation	NOUN
gs-3955	118	91	patterns	pattern	NOUN
gs-3955	118	92	(	(	PUNCT
gs-3955	118	93	jansen	jansen	PROPN
gs-3955	118	94	et	et	PROPN
gs-3955	118	95	al	al	PROPN
gs-3955	118	96	.	.	PROPN
gs-3955	118	97	,	,	PUNCT
gs-3955	118	98	2021	2021	NUM
gs-3955	118	99	)	)	PUNCT
gs-3955	118	100	●	●	NUM
gs-3955	118	101	unique	unique	ADJ
gs-3955	118	102	microrna	microrna	PROPN
gs-3955	118	103	expression	expression	NOUN
gs-3955	118	104	profiles	profile	NOUN
gs-3955	118	105	associated	associate	VERB
gs-3955	118	106	with	with	ADP
gs-3955	118	107	tissue	tissue	NOUN
gs-3955	118	108	maintenance	maintenance	NOUN
gs-3955	118	109	(	(	PUNCT
gs-3955	118	110	zhang	zhang	X
gs-3955	118	111	et	et	PROPN
gs-3955	118	112	al	al	PROPN
gs-3955	118	113	.	.	PROPN
gs-3955	118	114	,	,	PUNCT
gs-3955	118	115	2023	2023	NUM
gs-3955	118	116	)	)	PUNCT
gs-3955	118	117	●	●	PUNCT
gs-3955	118	118	slow	slow	ADJ
gs-3955	118	119	epigenetic	epigenetic	ADJ
gs-3955	118	120	aging	age	VERB
gs-3955	118	121	rate	rate	NOUN
gs-3955	118	122	compared	compare	VERB
gs-3955	118	123	to	to	ADP
gs-3955	118	124	shorter	shorter	ADV
gs-3955	118	125	-	-	PUNCT
gs-3955	118	126	lived	live	VERB
gs-3955	118	127	species	specie	NOUN
gs-3955	118	128	(	(	PUNCT
gs-3955	118	129	horvath	horvath	PROPN
gs-3955	118	130	et	et	PROPN
gs-3955	118	131	al	al	PROPN
gs-3955	118	132	.	.	PROPN
gs-3955	118	133	,	,	PUNCT
gs-3955	118	134	2022	2022	NUM
gs-3955	118	135	)	)	PUNCT
gs-3955	118	136	the	the	DET
gs-3955	118	137	planarian	planarian	ADJ
gs-3955	118	138	flatworm	flatworm	NOUN
gs-3955	118	139	(	(	PUNCT
gs-3955	118	140	schmidtea	schmidtea	PROPN
gs-3955	118	141	mediterranea	mediterranea	PROPN
gs-3955	118	142	):	):	PUNCT
gs-3955	118	143	a	a	DET
gs-3955	118	144	model	model	NOUN
gs-3955	118	145	of	of	ADP
gs-3955	118	146	biological	biological	ADJ
gs-3955	118	147	immortality	immortality	NOUN
gs-3955	118	148	the	the	DET
gs-3955	118	149	freshwater	freshwater	NOUN
gs-3955	118	150	planarian	planarian	NOUN
gs-3955	118	151	schmidtea	schmidtea	PROPN
gs-3955	118	152	mediterranea	mediterranea	PROPN
gs-3955	118	153	represents	represent	VERB
gs-3955	118	154	one	one	NUM
gs-3955	118	155	of	of	ADP
gs-3955	118	156	nature	nature	NOUN
gs-3955	118	157	's	's	PART
gs-3955	118	158	most	most	ADV
gs-3955	118	159	extraordinary	extraordinary	ADJ
gs-3955	118	160	examples	example	NOUN
gs-3955	118	161	of	of	ADP
gs-3955	118	162	apparent	apparent	ADJ
gs-3955	118	163	biological	biological	ADJ
gs-3955	118	164	immortality	immortality	NOUN
gs-3955	118	165	,	,	PUNCT
gs-3955	118	166	demonstrating	demonstrate	VERB
gs-3955	118	167	no	no	DET
gs-3955	118	168	measurable	measurable	ADJ
gs-3955	118	169	signs	sign	NOUN
gs-3955	118	170	of	of	ADP
gs-3955	118	171	aging	age	VERB
gs-3955	118	172	due	due	ADJ
gs-3955	118	173	to	to	ADP
gs-3955	118	174	its	its	PRON
gs-3955	118	175	unparalleled	unparalleled	ADJ
gs-3955	118	176	regenerative	regenerative	ADJ
gs-3955	118	177	capabilities	capability	NOUN
gs-3955	118	178	(	(	PUNCT
gs-3955	118	179	reddien	reddien	NOUN
gs-3955	118	180	,	,	PUNCT
gs-3955	118	181	2018	2018	NUM
gs-3955	118	182	)	)	PUNCT
gs-3955	118	183	.	.	PUNCT
gs-3955	119	1	this	this	DET
gs-3955	119	2	flatworm	flatworm	NOUN
gs-3955	119	3	species	specie	NOUN
gs-3955	119	4	can	can	AUX
gs-3955	119	5	completely	completely	ADV
gs-3955	119	6	regenerate	regenerate	VERB
gs-3955	119	7	all	all	DET
gs-3955	119	8	tissues	tissue	NOUN
gs-3955	119	9	,	,	PUNCT
gs-3955	119	10	including	include	VERB
gs-3955	119	11	complex	complex	ADJ
gs-3955	119	12	neural	neural	ADJ
gs-3955	119	13	structures	structure	NOUN
gs-3955	119	14	like	like	ADP
gs-3955	119	15	the	the	DET
gs-3955	119	16	brain	brain	NOUN
gs-3955	119	17	,	,	PUNCT
gs-3955	119	18	making	make	VERB
gs-3955	119	19	it	it	PRON
gs-3955	119	20	an	an	DET
gs-3955	119	21	invaluable	invaluable	ADJ
gs-3955	119	22	model	model	NOUN
gs-3955	119	23	for	for	ADP
gs-3955	119	24	studying	study	VERB
gs-3955	119	25	cellular	cellular	ADJ
gs-3955	119	26	rejuvenation	rejuvenation	NOUN
gs-3955	119	27	and	and	CCONJ
gs-3955	119	28	longevity	longevity	NOUN
gs-3955	119	29	mechanisms	mechanism	NOUN
gs-3955	119	30	(	(	PUNCT
gs-3955	119	31	wagner	wagner	PROPN
gs-3955	119	32	et	et	PROPN
gs-3955	119	33	al	al	PROPN
gs-3955	119	34	.	.	PROPN
gs-3955	119	35	,	,	PUNCT
gs-3955	119	36	2011	2011	NUM
gs-3955	119	37	)	)	PUNCT
gs-3955	119	38	.	.	PUNCT
gs-3955	120	1	stem	stem	NOUN
gs-3955	120	2	cell	cell	NOUN
gs-3955	120	3	systems	system	NOUN
gs-3955	120	4	and	and	CCONJ
gs-3955	120	5	tissue	tissue	NOUN
gs-3955	120	6	renewal	renewal	NOUN
gs-3955	120	7	neoblast	neoblast	PROPN
gs-3955	120	8	characteristics	characteristic	NOUN
gs-3955	120	9	:	:	PUNCT
gs-3955	120	10	●	●	PUNCT
gs-3955	120	11	pluripotent	pluripotent	ADJ
gs-3955	120	12	adult	adult	NOUN
gs-3955	120	13	stem	stem	NOUN
gs-3955	120	14	cells	cell	NOUN
gs-3955	120	15	(	(	PUNCT
gs-3955	120	16	neoblasts	neoblast	NOUN
gs-3955	120	17	)	)	PUNCT
gs-3955	120	18	capable	capable	ADJ
gs-3955	120	19	of	of	ADP
gs-3955	120	20	differentiating	differentiate	VERB
gs-3955	120	21	into	into	ADP
gs-3955	120	22	any	any	DET
gs-3955	120	23	cell	cell	NOUN
gs-3955	120	24	type	type	NOUN
gs-3955	120	25	(	(	PUNCT
gs-3955	120	26	rink	rink	NOUN
gs-3955	120	27	,	,	PUNCT
gs-3955	120	28	2013	2013	NUM
gs-3955	120	29	)	)	PUNCT
gs-3955	120	30	●	●	PUNCT
gs-3955	120	31	continuous	continuous	ADJ
gs-3955	120	32	tissue	tissue	NOUN
gs-3955	120	33	renewal	renewal	NOUN
gs-3955	120	34	maintaining	maintain	VERB
gs-3955	120	35	organismal	organismal	ADJ
gs-3955	120	36	homeostasis	homeostasis	NOUN
gs-3955	120	37	(	(	PUNCT
gs-3955	120	38	zeng	zeng	PROPN
gs-3955	120	39	et	et	PROPN
gs-3955	120	40	al	al	PROPN
gs-3955	120	41	.	.	PROPN
gs-3955	120	42	,	,	PUNCT
gs-3955	120	43	2018	2018	NUM
gs-3955	120	44	)	)	PUNCT
gs-3955	121	1	●	●	NUM
gs-3955	121	2	whole	whole	ADJ
gs-3955	121	3	-	-	PUNCT
gs-3955	121	4	body	body	NOUN
gs-3955	121	5	regeneration	regeneration	NOUN
gs-3955	121	6	from	from	ADP
gs-3955	121	7	minute	minute	ADJ
gs-3955	121	8	tissue	tissue	NOUN
gs-3955	121	9	fragments	fragment	NOUN
gs-3955	121	10	(	(	PUNCT
gs-3955	121	11	elliott	elliott	PROPN
gs-3955	121	12	&	&	CCONJ
gs-3955	121	13	sánchez	sánchez	PROPN
gs-3955	121	14	alvarado	alvarado	PROPN
gs-3955	121	15	,	,	PUNCT
gs-3955	121	16	2013	2013	NUM
gs-3955	121	17	)	)	PUNCT
gs-3955	121	18	●	●	PUNCT
gs-3955	121	19	efficient	efficient	ADJ
gs-3955	121	20	damage	damage	NOUN
gs-3955	121	21	clearance	clearance	NOUN
gs-3955	121	22	preventing	prevent	VERB
gs-3955	121	23	cellular	cellular	ADJ
gs-3955	121	24	senescence	senescence	NOUN
gs-3955	121	25	accumulation	accumulation	NOUN
gs-3955	121	26	(	(	PUNCT
gs-3955	121	27	pearson	pearson	PROPN
gs-3955	121	28	&	&	CCONJ
gs-3955	121	29	sánchez	sánchez	PROPN
gs-3955	121	30	alvarado	alvarado	PROPN
gs-3955	121	31	,	,	PUNCT
gs-3955	121	32	2018	2018	NUM
gs-3955	121	33	)	)	PUNCT
gs-3955	121	34	age	age	NOUN
gs-3955	121	35	-	-	PUNCT
gs-3955	121	36	resistant	resistant	ADJ
gs-3955	121	37	phenotype	phenotype	NOUN
gs-3955	121	38	experimental	experimental	ADJ
gs-3955	121	39	studies	study	NOUN
gs-3955	121	40	demonstrate	demonstrate	VERB
gs-3955	121	41	:	:	PUNCT
gs-3955	121	42	●	●	PUNCT
gs-3955	121	43	no	no	DET
gs-3955	121	44	detectable	detectable	ADJ
gs-3955	121	45	aging	aging	NOUN
gs-3955	121	46	after	after	ADP
gs-3955	121	47	years	year	NOUN
gs-3955	121	48	of	of	ADP
gs-3955	121	49	laboratory	laboratory	NOUN
gs-3955	121	50	culture	culture	NOUN
gs-3955	121	51	(	(	PUNCT
gs-3955	121	52	sahu	sahu	PROPN
gs-3955	121	53	et	et	PROPN
gs-3955	121	54	al	al	PROPN
gs-3955	121	55	.	.	PROPN
gs-3955	121	56	,	,	PUNCT
gs-3955	121	57	2017	2017	NUM
gs-3955	121	58	)	)	PUNCT
gs-3955	121	59	●	●	NUM
gs-3955	121	60	conserved	conserve	VERB
gs-3955	121	61	regenerative	regenerative	ADJ
gs-3955	121	62	capacity	capacity	NOUN
gs-3955	121	63	regardless	regardless	ADV
gs-3955	121	64	of	of	ADP
gs-3955	121	65	chronological	chronological	ADJ
gs-3955	121	66	age	age	NOUN
gs-3955	121	67	(	(	PUNCT
gs-3955	121	68	reddien	reddien	NOUN
gs-3955	121	69	,	,	PUNCT
gs-3955	121	70	2021	2021	NUM
gs-3955	121	71	)	)	PUNCT
gs-3955	121	72	●	●	NUM
gs-3955	121	73	robust	robust	ADJ
gs-3955	121	74	oxidative	oxidative	ADJ
gs-3955	121	75	stress	stress	NOUN
gs-3955	121	76	defenses	defense	NOUN
gs-3955	121	77	maintaining	maintain	VERB
gs-3955	121	78	cellular	cellular	ADJ
gs-3955	121	79	integrity	integrity	NOUN
gs-3955	121	80	(	(	PUNCT
gs-3955	121	81	pirotte	pirotte	NOUN
gs-3955	121	82	et	et	PROPN
gs-3955	121	83	al	al	PROPN
gs-3955	121	84	.	.	PROPN
gs-3955	121	85	,	,	PUNCT
gs-3955	121	86	2015	2015	NUM
gs-3955	121	87	)	)	PUNCT
gs-3955	121	88	georgian	georgian	ADJ
gs-3955	121	89	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	121	90	მეცნიერები	მეცნიერები	NOUN
gs-3955	121	91	ტ.	ტ.	VERB
gs-3955	121	92	7	7	NUM
gs-3955	121	93	n	n	DET
gs-3955	121	94	3	3	NUM
gs-3955	121	95	,	,	PUNCT
gs-3955	121	96	2025	2025	NUM
gs-3955	121	97	362	362	NUM
gs-3955	121	98	genomic	genomic	NOUN
gs-3955	121	99	and	and	CCONJ
gs-3955	121	100	epigenetic	epigenetic	ADJ
gs-3955	121	101	stability	stability	NOUN
gs-3955	121	102	key	key	ADJ
gs-3955	121	103	findings	finding	NOUN
gs-3955	121	104	include	include	VERB
gs-3955	121	105	:	:	PUNCT
gs-3955	121	106	●	●	PUNCT
gs-3955	121	107	minimal	minimal	ADJ
gs-3955	121	108	epigenetic	epigenetic	ADJ
gs-3955	121	109	drift	drift	NOUN
gs-3955	121	110	over	over	ADP
gs-3955	121	111	time	time	NOUN
gs-3955	121	112	(	(	PUNCT
gs-3955	121	113	jaber	jaber	PROPN
gs-3955	121	114	-	-	PUNCT
gs-3955	121	115	hijazi	hijazi	PROPN
gs-3955	121	116	et	et	PROPN
gs-3955	121	117	al	al	PROPN
gs-3955	121	118	.	.	PROPN
gs-3955	121	119	,	,	PUNCT
gs-3955	121	120	2016	2016	NUM
gs-3955	121	121	)	)	PUNCT
gs-3955	121	122	●	●	PUNCT
gs-3955	121	123	active	active	ADJ
gs-3955	121	124	suppression	suppression	NOUN
gs-3955	121	125	of	of	ADP
gs-3955	121	126	aging	age	VERB
gs-3955	121	127	-	-	PUNCT
gs-3955	121	128	related	relate	VERB
gs-3955	121	129	genes	gene	NOUN
gs-3955	121	130	(	(	PUNCT
gs-3955	121	131	tejada	tejada	PROPN
gs-3955	121	132	-	-	PUNCT
gs-3955	121	133	romero	romero	PROPN
gs-3955	121	134	et	et	PROPN
gs-3955	121	135	al	al	PROPN
gs-3955	121	136	.	.	PROPN
gs-3955	121	137	,	,	PUNCT
gs-3955	121	138	2021	2021	NUM
gs-3955	121	139	)	)	PUNCT
gs-3955	121	140	●	●	PUNCT
gs-3955	121	141	exceptional	exceptional	ADJ
gs-3955	121	142	telomere	telomere	NOUN
gs-3955	121	143	maintenance	maintenance	NOUN
gs-3955	121	144	preventing	prevent	VERB
gs-3955	121	145	replicative	replicative	ADJ
gs-3955	121	146	senescence	senescence	NOUN
gs-3955	121	147	(	(	PUNCT
gs-3955	121	148	tan	tan	NOUN
gs-3955	121	149	et	et	NOUN
gs-3955	121	150	al	al	PROPN
gs-3955	121	151	.	.	PROPN
gs-3955	121	152	,	,	PUNCT
gs-3955	121	153	2012	2012	NUM
gs-3955	121	154	)	)	PUNCT
gs-3955	121	155	comparative	comparative	ADJ
gs-3955	121	156	perspectives	perspective	NOUN
gs-3955	121	157	and	and	CCONJ
gs-3955	121	158	research	research	NOUN
gs-3955	121	159	implications	implication	NOUN
gs-3955	121	160	these	these	DET
gs-3955	121	161	two	two	NUM
gs-3955	121	162	model	model	NOUN
gs-3955	121	163	organisms	organism	NOUN
gs-3955	121	164	the	the	DET
gs-3955	121	165	century	century	NOUN
gs-3955	121	166	-	-	PUNCT
gs-3955	121	167	old	old	ADJ
gs-3955	121	168	sea	sea	NOUN
gs-3955	121	169	urchin	urchin	NOUN
gs-3955	121	170	and	and	CCONJ
gs-3955	121	171	the	the	DET
gs-3955	121	172	theoretically	theoretically	ADV
gs-3955	121	173	immortal	immortal	ADJ
gs-3955	121	174	planarian	planarian	NOUN
gs-3955	121	175	provide	provide	VERB
gs-3955	121	176	complementary	complementary	ADJ
gs-3955	121	177	insights	insight	NOUN
gs-3955	121	178	into	into	ADP
gs-3955	121	179	the	the	DET
gs-3955	121	180	spectrum	spectrum	NOUN
gs-3955	121	181	of	of	ADP
gs-3955	121	182	negligible	negligible	ADJ
gs-3955	121	183	senescence	senescence	NOUN
gs-3955	121	184	in	in	ADP
gs-3955	121	185	nature	nature	NOUN
gs-3955	121	186	.	.	PUNCT
gs-3955	122	1	while	while	SCONJ
gs-3955	122	2	s.	s.	PROPN
gs-3955	122	3	franciscanus	franciscanus	PROPN
gs-3955	122	4	demonstrates	demonstrate	VERB
gs-3955	122	5	how	how	SCONJ
gs-3955	122	6	complex	complex	ADJ
gs-3955	122	7	marine	marine	ADJ
gs-3955	122	8	invertebrates	invertebrate	NOUN
gs-3955	122	9	can	can	AUX
gs-3955	122	10	maintain	maintain	VERB
gs-3955	122	11	physiological	physiological	ADJ
gs-3955	122	12	function	function	NOUN
gs-3955	122	13	for	for	ADP
gs-3955	122	14	exceptionally	exceptionally	ADV
gs-3955	122	15	long	long	ADJ
gs-3955	122	16	periods	period	NOUN
gs-3955	122	17	,	,	PUNCT
gs-3955	122	18	s.	s.	PROPN
gs-3955	122	19	mediterranea	mediterranea	PROPN
gs-3955	122	20	reveals	reveal	VERB
gs-3955	122	21	the	the	DET
gs-3955	122	22	potential	potential	NOUN
gs-3955	122	23	for	for	ADP
gs-3955	122	24	complete	complete	ADJ
gs-3955	122	25	avoidance	avoidance	NOUN
gs-3955	122	26	of	of	ADP
gs-3955	122	27	aging	age	VERB
gs-3955	122	28	through	through	ADP
gs-3955	122	29	continuous	continuous	ADJ
gs-3955	122	30	tissue	tissue	NOUN
gs-3955	122	31	renewal	renewal	NOUN
gs-3955	122	32	.	.	PUNCT
gs-3955	123	1	picture	picture	NOUN
gs-3955	123	2	1	1	NUM
gs-3955	123	3	.	.	PUNCT
gs-3955	124	1	molecular	molecular	ADJ
gs-3955	124	2	strategies	strategy	NOUN
gs-3955	124	3	of	of	ADP
gs-3955	124	4	negligible	negligible	ADJ
gs-3955	124	5	senescence	senescence	NOUN
gs-3955	124	6	across	across	ADP
gs-3955	124	7	species	specie	NOUN
gs-3955	124	8	their	their	PRON
gs-3955	124	9	study	study	NOUN
gs-3955	124	10	offers	offer	VERB
gs-3955	124	11	unprecedented	unprecedented	ADJ
gs-3955	124	12	opportunities	opportunity	NOUN
gs-3955	124	13	for	for	ADP
gs-3955	124	14	:	:	PUNCT
gs-3955	124	15	●	●	PUNCT
gs-3955	124	16	understanding	understand	VERB
gs-3955	124	17	conserved	conserve	VERB
gs-3955	124	18	mechanisms	mechanism	NOUN
gs-3955	124	19	of	of	ADP
gs-3955	124	20	longevity	longevity	NOUN
gs-3955	124	21	across	across	ADP
gs-3955	124	22	phylogenetically	phylogenetically	ADV
gs-3955	124	23	distant	distant	ADJ
gs-3955	124	24	species	specie	NOUN
gs-3955	124	25	●	●	NUM
gs-3955	124	26	developing	develop	VERB
gs-3955	124	27	novel	novel	ADJ
gs-3955	124	28	regenerative	regenerative	ADJ
gs-3955	124	29	therapies	therapy	NOUN
gs-3955	124	30	for	for	ADP
gs-3955	124	31	age	age	NOUN
gs-3955	124	32	-	-	PUNCT
gs-3955	124	33	related	relate	VERB
gs-3955	124	34	tissue	tissue	NOUN
gs-3955	124	35	degeneration	degeneration	NOUN
gs-3955	124	36	georgian	georgian	ADJ
gs-3955	124	37	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	124	38	მეცნიერები	მეცნიერები	NOUN
gs-3955	124	39	ტ.	ტ.	VERB
gs-3955	124	40	7	7	NUM
gs-3955	124	41	n	n	DET
gs-3955	124	42	3	3	NUM
gs-3955	124	43	,	,	PUNCT
gs-3955	124	44	2025	2025	NUM
gs-3955	124	45	363	363	NUM
gs-3955	124	46	●	●	NOUN
gs-3955	124	47	identifying	identify	VERB
gs-3955	124	48	molecular	molecular	ADJ
gs-3955	124	49	targets	target	NOUN
gs-3955	124	50	for	for	ADP
gs-3955	124	51	interventions	intervention	NOUN
gs-3955	124	52	against	against	ADP
gs-3955	124	53	cellular	cellular	ADJ
gs-3955	124	54	aging	aging	NOUN
gs-3955	124	55	the	the	DET
gs-3955	124	56	continued	continued	ADJ
gs-3955	124	57	investigation	investigation	NOUN
gs-3955	124	58	of	of	ADP
gs-3955	124	59	these	these	DET
gs-3955	124	60	remarkable	remarkable	ADJ
gs-3955	124	61	organisms	organism	NOUN
gs-3955	124	62	promises	promise	NOUN
gs-3955	124	63	to	to	PART
gs-3955	124	64	fundamentally	fundamentally	ADV
gs-3955	124	65	advance	advance	VERB
gs-3955	124	66	our	our	PRON
gs-3955	124	67	understanding	understanding	NOUN
gs-3955	124	68	of	of	ADP
gs-3955	124	69	the	the	DET
gs-3955	124	70	biological	biological	ADJ
gs-3955	124	71	constraints	constraint	NOUN
gs-3955	124	72	on	on	ADP
gs-3955	124	73	lifespan	lifespan	NOUN
gs-3955	124	74	and	and	CCONJ
gs-3955	124	75	the	the	DET
gs-3955	124	76	potential	potential	NOUN
gs-3955	124	77	for	for	ADP
gs-3955	124	78	overcoming	overcome	VERB
gs-3955	124	79	age	age	NOUN
gs-3955	124	80	-	-	PUNCT
gs-3955	124	81	related	relate	VERB
gs-3955	124	82	decline	decline	NOUN
gs-3955	124	83	in	in	ADP
gs-3955	124	84	more	more	ADJ
gs-3955	124	85	complex	complex	ADJ
gs-3955	124	86	animals	animal	NOUN
gs-3955	124	87	,	,	PUNCT
gs-3955	124	88	including	include	VERB
gs-3955	124	89	humans	human	NOUN
gs-3955	124	90	.	.	PUNCT
gs-3955	125	1	their	their	PRON
gs-3955	125	2	unique	unique	ADJ
gs-3955	125	3	adaptations	adaptation	NOUN
gs-3955	125	4	challenge	challenge	VERB
gs-3955	125	5	conventional	conventional	ADJ
gs-3955	125	6	paradigms	paradigm	NOUN
gs-3955	125	7	of	of	ADP
gs-3955	125	8	aging	aging	NOUN
gs-3955	125	9	while	while	SCONJ
gs-3955	125	10	providing	provide	VERB
gs-3955	125	11	concrete	concrete	ADJ
gs-3955	125	12	examples	example	NOUN
gs-3955	125	13	of	of	ADP
gs-3955	125	14	nature	nature	NOUN
gs-3955	125	15	's	's	PART
gs-3955	125	16	solutions	solution	NOUN
gs-3955	125	17	to	to	ADP
gs-3955	125	18	the	the	DET
gs-3955	125	19	problem	problem	NOUN
gs-3955	125	20	of	of	ADP
gs-3955	125	21	biological	biological	ADJ
gs-3955	125	22	time	time	NOUN
gs-3955	125	23	(	(	PUNCT
gs-3955	125	24	picture	picture	NOUN
gs-3955	125	25	1	1	NUM
gs-3955	125	26	)	)	PUNCT
gs-3955	125	27	.	.	PUNCT
gs-3955	126	1	this	this	DET
gs-3955	126	2	infographic	infographic	ADJ
gs-3955	126	3	illustrates	illustrate	VERB
gs-3955	126	4	the	the	DET
gs-3955	126	5	key	key	ADJ
gs-3955	126	6	molecular	molecular	ADJ
gs-3955	126	7	adaptations	adaptation	NOUN
gs-3955	126	8	contributing	contribute	VERB
gs-3955	126	9	to	to	ADP
gs-3955	126	10	extreme	extreme	ADJ
gs-3955	126	11	longevity	longevity	NOUN
gs-3955	126	12	in	in	ADP
gs-3955	126	13	negligibly	negligibly	ADV
gs-3955	126	14	senescent	senescent	ADJ
gs-3955	126	15	species	specie	NOUN
gs-3955	126	16	.	.	PUNCT
gs-3955	127	1	the	the	DET
gs-3955	127	2	ocean	ocean	PROPN
gs-3955	127	3	quahog	quahog	PROPN
gs-3955	127	4	(	(	PUNCT
gs-3955	127	5	arctica	arctica	PROPN
gs-3955	127	6	islandica	islandica	PROPN
gs-3955	127	7	)	)	PUNCT
gs-3955	127	8	utilizes	utilize	VERB
gs-3955	127	9	enhanced	enhance	VERB
gs-3955	127	10	nucleotide	nucleotide	NOUN
gs-3955	127	11	excision	excision	NOUN
gs-3955	127	12	repair	repair	NOUN
gs-3955	127	13	via	via	ADP
gs-3955	127	14	stabilized	stabilize	VERB
gs-3955	127	15	xpa	xpa	PROPN
gs-3955	127	16	-	-	PUNCT
gs-3955	127	17	ercc1	ercc1	NOUN
gs-3955	127	18	complexes	complex	NOUN
gs-3955	127	19	.	.	PUNCT
gs-3955	128	1	the	the	DET
gs-3955	128	2	greenland	greenland	PROPN
gs-3955	128	3	shark	shark	NOUN
gs-3955	128	4	(	(	PUNCT
gs-3955	128	5	somniosus	somniosus	PROPN
gs-3955	128	6	microcephalus	microcephalus	PROPN
gs-3955	128	7	)	)	PUNCT
gs-3955	128	8	shows	show	VERB
gs-3955	128	9	duplication	duplication	NOUN
gs-3955	128	10	of	of	ADP
gs-3955	128	11	dna	dna	PROPN
gs-3955	128	12	repair	repair	NOUN
gs-3955	128	13	genes	gene	NOUN
gs-3955	128	14	(	(	PUNCT
gs-3955	128	15	atm	atm	PROPN
gs-3955	128	16	,	,	PUNCT
gs-3955	128	17	atr	atr	PROPN
gs-3955	128	18	)	)	PUNCT
gs-3955	128	19	for	for	ADP
gs-3955	128	20	genomic	genomic	ADJ
gs-3955	128	21	redundancy	redundancy	NOUN
gs-3955	128	22	under	under	ADP
gs-3955	128	23	oxidative	oxidative	ADJ
gs-3955	128	24	stress	stress	NOUN
gs-3955	128	25	.	.	PUNCT
gs-3955	129	1	the	the	DET
gs-3955	129	2	naked	naked	ADJ
gs-3955	129	3	mole	mole	NOUN
gs-3955	129	4	-	-	PUNCT
gs-3955	129	5	rat	rat	NOUN
gs-3955	129	6	(	(	PUNCT
gs-3955	129	7	heterocephalus	heterocephalus	ADJ
gs-3955	129	8	glaber	glaber	NOUN
gs-3955	129	9	)	)	PUNCT
gs-3955	129	10	exhibits	exhibit	VERB
gs-3955	129	11	high	high	ADJ
gs-3955	129	12	-	-	PUNCT
gs-3955	129	13	molecular	molecular	ADJ
gs-3955	129	14	-	-	PUNCT
gs-3955	129	15	weight	weight	NOUN
gs-3955	129	16	hyaluronan	hyaluronan	ADV
gs-3955	129	17	-	-	PUNCT
gs-3955	129	18	mediated	mediate	VERB
gs-3955	129	19	contact	contact	NOUN
gs-3955	129	20	inhibition	inhibition	NOUN
gs-3955	129	21	,	,	PUNCT
gs-3955	129	22	hyperactive	hyperactive	ADJ
gs-3955	129	23	p53	p53	NOUN
gs-3955	129	24	variants	variant	NOUN
gs-3955	129	25	,	,	PUNCT
gs-3955	129	26	and	and	CCONJ
gs-3955	129	27	unique	unique	ADJ
gs-3955	129	28	necroptotic	necroptotic	ADJ
gs-3955	129	29	defense	defense	NOUN
gs-3955	129	30	mechanisms	mechanism	NOUN
gs-3955	129	31	.	.	PUNCT
gs-3955	130	1	additionally	additionally	ADV
gs-3955	130	2	,	,	PUNCT
gs-3955	130	3	its	its	PRON
gs-3955	130	4	ribosomes	ribosome	NOUN
gs-3955	130	5	have	have	AUX
gs-3955	130	6	evolved	evolve	VERB
gs-3955	130	7	structurally	structurally	ADV
gs-3955	130	8	distinct	distinct	ADJ
gs-3955	130	9	decoding	decode	VERB
gs-3955	130	10	centers	center	NOUN
gs-3955	130	11	that	that	PRON
gs-3955	130	12	reduce	reduce	VERB
gs-3955	130	13	translational	translational	ADJ
gs-3955	130	14	errors	error	NOUN
gs-3955	130	15	.	.	PUNCT
gs-3955	131	1	these	these	DET
gs-3955	131	2	convergent	convergent	NOUN
gs-3955	131	3	strategies	strategy	NOUN
gs-3955	131	4	maintain	maintain	VERB
gs-3955	131	5	genome	genome	NOUN
gs-3955	131	6	integrity	integrity	NOUN
gs-3955	131	7	,	,	PUNCT
gs-3955	131	8	suppress	suppress	ADJ
gs-3955	131	9	tumorigenesis	tumorigenesis	NOUN
gs-3955	131	10	,	,	PUNCT
gs-3955	131	11	and	and	CCONJ
gs-3955	131	12	mitigate	mitigate	VERB
gs-3955	131	13	proteotoxic	proteotoxic	ADJ
gs-3955	131	14	stress	stress	NOUN
gs-3955	131	15	—	—	PUNCT
gs-3955	131	16	foundational	foundational	ADJ
gs-3955	131	17	elements	element	NOUN
gs-3955	131	18	in	in	ADP
gs-3955	131	19	the	the	DET
gs-3955	131	20	biology	biology	NOUN
gs-3955	131	21	of	of	ADP
gs-3955	131	22	negligible	negligible	ADJ
gs-3955	131	23	senescence	senescence	NOUN
gs-3955	131	24	.	.	PUNCT
gs-3955	132	1	molecular	molecular	ADJ
gs-3955	132	2	mechanisms	mechanism	NOUN
gs-3955	132	3	of	of	ADP
gs-3955	132	4	negligible	negligible	ADJ
gs-3955	132	5	senescence	senescence	NOUN
gs-3955	132	6	stem	stem	NOUN
gs-3955	132	7	cell	cell	NOUN
gs-3955	132	8	maintenance	maintenance	NOUN
gs-3955	132	9	and	and	CCONJ
gs-3955	132	10	continuous	continuous	ADJ
gs-3955	132	11	regeneration	regeneration	NOUN
gs-3955	132	12	hydra	hydra	NOUN
gs-3955	132	13	maintains	maintain	VERB
gs-3955	132	14	a	a	DET
gs-3955	132	15	large	large	ADJ
gs-3955	132	16	population	population	NOUN
gs-3955	132	17	of	of	ADP
gs-3955	132	18	active	active	ADJ
gs-3955	132	19	stem	stem	NOUN
gs-3955	132	20	cells	cell	NOUN
gs-3955	132	21	,	,	PUNCT
gs-3955	132	22	enabling	enable	VERB
gs-3955	132	23	constant	constant	ADJ
gs-3955	132	24	regeneration	regeneration	NOUN
gs-3955	132	25	of	of	ADP
gs-3955	132	26	all	all	DET
gs-3955	132	27	tissues	tissue	NOUN
gs-3955	132	28	without	without	ADP
gs-3955	132	29	exhaustion	exhaustion	NOUN
gs-3955	132	30	or	or	CCONJ
gs-3955	132	31	telomere	telomere	NOUN
gs-3955	132	32	shortening	shortening	NOUN
gs-3955	132	33	(	(	PUNCT
gs-3955	132	34	wagner	wagner	PROPN
gs-3955	132	35	et	et	PROPN
gs-3955	132	36	al	al	PROPN
gs-3955	132	37	.	.	PROPN
gs-3955	132	38	,	,	PUNCT
gs-3955	132	39	2011	2011	NUM
gs-3955	132	40	;	;	PUNCT
gs-3955	132	41	siebert	siebert	PROPN
gs-3955	132	42	et	et	PROPN
gs-3955	132	43	al	al	PROPN
gs-3955	132	44	.	.	PROPN
gs-3955	132	45	,	,	PUNCT
gs-3955	132	46	2019	2019	NUM
gs-3955	132	47	)	)	PUNCT
gs-3955	132	48	.	.	PUNCT
gs-3955	133	1	the	the	DET
gs-3955	133	2	foxo	foxo	ADJ
gs-3955	133	3	transcription	transcription	NOUN
gs-3955	133	4	factor	factor	NOUN
gs-3955	133	5	is	be	AUX
gs-3955	133	6	highly	highly	ADV
gs-3955	133	7	expressed	express	VERB
gs-3955	133	8	in	in	ADP
gs-3955	133	9	hydra	hydra	NOUN
gs-3955	133	10	stem	stem	NOUN
gs-3955	133	11	cells	cell	NOUN
gs-3955	133	12	and	and	CCONJ
gs-3955	133	13	plays	play	VERB
gs-3955	133	14	a	a	DET
gs-3955	133	15	pivotal	pivotal	ADJ
gs-3955	133	16	role	role	NOUN
gs-3955	133	17	in	in	ADP
gs-3955	133	18	preserving	preserve	VERB
gs-3955	133	19	their	their	PRON
gs-3955	133	20	pluripotency	pluripotency	NOUN
gs-3955	133	21	and	and	CCONJ
gs-3955	133	22	preventing	prevent	VERB
gs-3955	133	23	malignant	malignant	ADJ
gs-3955	133	24	transformation	transformation	NOUN
gs-3955	133	25	by	by	ADP
gs-3955	133	26	regulating	regulate	VERB
gs-3955	133	27	wnt	wnt	NOUN
gs-3955	133	28	signaling	signal	VERB
gs-3955	133	29	and	and	CCONJ
gs-3955	133	30	apoptosis	apoptosis	NOUN
gs-3955	133	31	pathways	pathway	NOUN
gs-3955	133	32	(	(	PUNCT
gs-3955	133	33	wagner	wagner	PROPN
gs-3955	133	34	et	et	PROPN
gs-3955	133	35	al	al	PROPN
gs-3955	133	36	.	.	PROPN
gs-3955	133	37	,	,	PUNCT
gs-3955	133	38	2011	2011	NUM
gs-3955	133	39	;	;	PUNCT
gs-3955	133	40	tkemaladze	tkemaladze	NOUN
gs-3955	133	41	,	,	PUNCT
gs-3955	133	42	2023	2023	NUM
gs-3955	133	43	)	)	PUNCT
gs-3955	133	44	.	.	PUNCT
gs-3955	134	1	this	this	DET
gs-3955	134	2	continuous	continuous	ADJ
gs-3955	134	3	renewal	renewal	NOUN
gs-3955	134	4	allows	allow	VERB
gs-3955	134	5	whole	whole	ADJ
gs-3955	134	6	-	-	PUNCT
gs-3955	134	7	body	body	NOUN
gs-3955	134	8	regeneration	regeneration	NOUN
gs-3955	134	9	and	and	CCONJ
gs-3955	134	10	is	be	AUX
gs-3955	134	11	a	a	DET
gs-3955	134	12	hallmark	hallmark	NOUN
gs-3955	134	13	of	of	ADP
gs-3955	134	14	negligible	negligible	ADJ
gs-3955	134	15	senescence	senescence	NOUN
gs-3955	134	16	in	in	ADP
gs-3955	134	17	these	these	DET
gs-3955	134	18	organisms	organism	NOUN
gs-3955	134	19	.	.	PUNCT
gs-3955	135	1	similarly	similarly	ADV
gs-3955	135	2	,	,	PUNCT
gs-3955	135	3	planarians	planarian	NOUN
gs-3955	135	4	possess	possess	VERB
gs-3955	135	5	neoblasts	neoblast	NOUN
gs-3955	135	6	—	—	PUNCT
gs-3955	135	7	pluripotent	pluripotent	ADJ
gs-3955	135	8	adult	adult	NOUN
gs-3955	135	9	stem	stem	NOUN
gs-3955	135	10	cells	cell	NOUN
gs-3955	135	11	—	—	PUNCT
gs-3955	135	12	characterized	characterize	VERB
gs-3955	135	13	by	by	ADP
gs-3955	135	14	high	high	ADJ
gs-3955	135	15	symmetric	symmetric	ADJ
gs-3955	135	16	division	division	NOUN
gs-3955	135	17	rates	rate	NOUN
gs-3955	135	18	(	(	PUNCT
gs-3955	135	19	up	up	ADP
gs-3955	135	20	to	to	PART
gs-3955	135	21	78	78	NUM
gs-3955	135	22	%	%	NOUN
gs-3955	135	23	)	)	PUNCT
gs-3955	135	24	,	,	PUNCT
gs-3955	135	25	which	which	PRON
gs-3955	135	26	maintain	maintain	VERB
gs-3955	135	27	tissue	tissue	NOUN
gs-3955	135	28	homeostasis	homeostasis	NOUN
gs-3955	135	29	and	and	CCONJ
gs-3955	135	30	enable	enable	VERB
gs-3955	135	31	complete	complete	ADJ
gs-3955	135	32	regeneration	regeneration	NOUN
gs-3955	135	33	of	of	ADP
gs-3955	135	34	lost	lose	VERB
gs-3955	135	35	body	body	NOUN
gs-3955	135	36	parts	part	NOUN
gs-3955	135	37	(	(	PUNCT
gs-3955	135	38	rink	rink	NOUN
gs-3955	135	39	,	,	PUNCT
gs-3955	135	40	2013	2013	NUM
gs-3955	135	41	;	;	PUNCT
gs-3955	135	42	wagner	wagner	PROPN
gs-3955	135	43	et	et	PROPN
gs-3955	135	44	al	al	PROPN
gs-3955	135	45	.	.	PROPN
gs-3955	135	46	,	,	PUNCT
gs-3955	135	47	2011	2011	NUM
gs-3955	135	48	)	)	PUNCT
gs-3955	135	49	.	.	PUNCT
gs-3955	136	1	their	their	PRON
gs-3955	136	2	chromatin	chromatin	NOUN
gs-3955	136	3	remains	remain	VERB
gs-3955	136	4	open	open	ADJ
gs-3955	136	5	and	and	CCONJ
gs-3955	136	6	accessible	accessible	ADJ
gs-3955	136	7	,	,	PUNCT
gs-3955	136	8	facilitating	facilitate	VERB
gs-3955	136	9	rapid	rapid	ADJ
gs-3955	136	10	gene	gene	NOUN
gs-3955	136	11	expression	expression	NOUN
gs-3955	136	12	responses	response	NOUN
gs-3955	136	13	to	to	ADP
gs-3955	136	14	injury	injury	NOUN
gs-3955	136	15	and	and	CCONJ
gs-3955	136	16	regeneration	regeneration	NOUN
gs-3955	136	17	signals	signal	NOUN
gs-3955	136	18	(	(	PUNCT
gs-3955	136	19	rink	rink	NOUN
gs-3955	136	20	,	,	PUNCT
gs-3955	136	21	2013	2013	NUM
gs-3955	136	22	)	)	PUNCT
gs-3955	136	23	.	.	PUNCT
gs-3955	137	1	this	this	DET
gs-3955	137	2	robust	robust	ADJ
gs-3955	137	3	stem	stem	NOUN
gs-3955	137	4	cell	cell	NOUN
gs-3955	137	5	pool	pool	NOUN
gs-3955	137	6	and	and	CCONJ
gs-3955	137	7	renewal	renewal	NOUN
gs-3955	137	8	capacity	capacity	NOUN
gs-3955	137	9	directly	directly	ADV
gs-3955	137	10	contrast	contrast	VERB
gs-3955	137	11	with	with	ADP
gs-3955	137	12	the	the	DET
gs-3955	137	13	stem	stem	NOUN
gs-3955	137	14	cell	cell	NOUN
gs-3955	137	15	exhaustion	exhaustion	NOUN
gs-3955	137	16	seen	see	VERB
gs-3955	137	17	in	in	ADP
gs-3955	137	18	most	most	ADJ
gs-3955	137	19	vertebrates	vertebrate	NOUN
gs-3955	137	20	.	.	PUNCT
gs-3955	138	1	the	the	DET
gs-3955	138	2	naked	naked	ADJ
gs-3955	138	3	mole	mole	ADJ
gs-3955	138	4	-	-	PUNCT
gs-3955	138	5	rat	rat	NOUN
gs-3955	138	6	exhibits	exhibit	VERB
gs-3955	138	7	distinct	distinct	ADJ
gs-3955	138	8	stem	stem	NOUN
gs-3955	138	9	cell	cell	NOUN
gs-3955	138	10	dynamics	dynamic	NOUN
gs-3955	138	11	,	,	PUNCT
gs-3955	138	12	maintaining	maintain	VERB
gs-3955	138	13	a	a	DET
gs-3955	138	14	modest	modest	ADJ
gs-3955	138	15	but	but	CCONJ
gs-3955	138	16	highly	highly	ADV
gs-3955	138	17	efficient	efficient	ADJ
gs-3955	138	18	stem	stem	NOUN
gs-3955	138	19	cell	cell	NOUN
gs-3955	138	20	population	population	NOUN
gs-3955	138	21	marked	mark	VERB
gs-3955	138	22	by	by	ADP
gs-3955	138	23	lgr5	lgr5	NOUN
gs-3955	138	24	and	and	CCONJ
gs-3955	138	25	bmi1	bmi1	NOUN
gs-3955	138	26	,	,	PUNCT
gs-3955	138	27	contributing	contribute	VERB
gs-3955	138	28	to	to	ADP
gs-3955	138	29	limited	limit	VERB
gs-3955	138	30	organ	organ	NOUN
gs-3955	138	31	repair	repair	NOUN
gs-3955	138	32	and	and	CCONJ
gs-3955	138	33	extraordinary	extraordinary	ADJ
gs-3955	138	34	longevity	longevity	NOUN
gs-3955	138	35	(	(	PUNCT
gs-3955	138	36	petruseva	petruseva	PROPN
gs-3955	138	37	et	et	PROPN
gs-3955	138	38	al	al	PROPN
gs-3955	138	39	.	.	PROPN
gs-3955	138	40	,	,	PUNCT
gs-3955	138	41	2017	2017	NUM
gs-3955	138	42	;	;	PUNCT
gs-3955	138	43	macrae	macrae	PROPN
gs-3955	138	44	et	et	PROPN
gs-3955	138	45	al	al	PROPN
gs-3955	138	46	.	.	PROPN
gs-3955	138	47	,	,	PUNCT
gs-3955	138	48	2015	2015	NUM
gs-3955	138	49	)	)	PUNCT
gs-3955	138	50	.	.	PUNCT
gs-3955	139	1	unlike	unlike	ADP
gs-3955	139	2	mammals	mammal	NOUN
gs-3955	139	3	with	with	ADP
gs-3955	139	4	telomere	telomere	NOUN
gs-3955	139	5	attrition	attrition	NOUN
gs-3955	139	6	,	,	PUNCT
gs-3955	139	7	mole	mole	NOUN
gs-3955	139	8	-	-	PUNCT
gs-3955	139	9	rats	rat	NOUN
gs-3955	139	10	express	express	VERB
gs-3955	139	11	constitutive	constitutive	ADJ
gs-3955	139	12	telomerase	telomerase	NOUN
gs-3955	139	13	activity	activity	NOUN
gs-3955	139	14	in	in	ADP
gs-3955	139	15	somatic	somatic	ADJ
gs-3955	139	16	stem	stem	NOUN
gs-3955	139	17	cells	cell	NOUN
gs-3955	139	18	,	,	PUNCT
gs-3955	139	19	preventing	prevent	VERB
gs-3955	139	20	replicative	replicative	ADJ
gs-3955	139	21	senescence	senescence	NOUN
gs-3955	139	22	and	and	CCONJ
gs-3955	139	23	supporting	support	VERB
gs-3955	139	24	prolonged	prolonged	ADJ
gs-3955	139	25	tissue	tissue	NOUN
gs-3955	139	26	maintenance	maintenance	NOUN
gs-3955	139	27	(	(	PUNCT
gs-3955	139	28	tkemaladze	tkemaladze	NOUN
gs-3955	139	29	,	,	PUNCT
gs-3955	139	30	2023	2023	NUM
gs-3955	139	31	)	)	PUNCT
gs-3955	139	32	.	.	PUNCT
gs-3955	140	1	georgian	georgian	ADJ
gs-3955	140	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	140	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	140	4	ტ.	ტ.	VERB
gs-3955	140	5	7	7	NUM
gs-3955	140	6	n	n	PRON
gs-3955	140	7	3	3	NUM
gs-3955	140	8	,	,	PUNCT
gs-3955	140	9	2025	2025	NUM
gs-3955	140	10	364	364	NUM
gs-3955	140	11	in	in	ADP
gs-3955	140	12	long	long	ADV
gs-3955	140	13	-	-	PUNCT
gs-3955	140	14	lived	live	VERB
gs-3955	140	15	vertebrates	vertebrate	NOUN
gs-3955	140	16	such	such	ADJ
gs-3955	140	17	as	as	ADP
gs-3955	140	18	the	the	DET
gs-3955	140	19	greenland	greenland	PROPN
gs-3955	140	20	shark	shark	NOUN
gs-3955	140	21	and	and	CCONJ
gs-3955	140	22	ocean	ocean	PROPN
gs-3955	140	23	quahog	quahog	PROPN
gs-3955	140	24	,	,	PUNCT
gs-3955	140	25	evidence	evidence	NOUN
gs-3955	140	26	suggests	suggest	VERB
gs-3955	140	27	prolonged	prolonged	ADJ
gs-3955	140	28	stem	stem	NOUN
gs-3955	140	29	cell	cell	NOUN
gs-3955	140	30	function	function	NOUN
gs-3955	140	31	and	and	CCONJ
gs-3955	140	32	turnover	turnover	NOUN
gs-3955	140	33	rates	rate	NOUN
gs-3955	140	34	consistent	consistent	ADJ
gs-3955	140	35	with	with	ADP
gs-3955	140	36	their	their	PRON
gs-3955	140	37	multi	multi	ADJ
gs-3955	140	38	-	-	ADJ
gs-3955	140	39	century	century	ADJ
gs-3955	140	40	lifespans	lifespan	NOUN
gs-3955	140	41	,	,	PUNCT
gs-3955	140	42	though	though	SCONJ
gs-3955	140	43	these	these	PRON
gs-3955	140	44	remain	remain	VERB
gs-3955	140	45	underexplored	underexplored	ADJ
gs-3955	140	46	(	(	PUNCT
gs-3955	140	47	macrae	macrae	NOUN
gs-3955	140	48	et	et	PROPN
gs-3955	140	49	al	al	PROPN
gs-3955	140	50	.	.	PROPN
gs-3955	140	51	,	,	PUNCT
gs-3955	140	52	2015	2015	NUM
gs-3955	140	53	;	;	PUNCT
gs-3955	140	54	narayan	narayan	PROPN
gs-3955	140	55	et	et	PROPN
gs-3955	140	56	al	al	PROPN
gs-3955	140	57	.	.	PROPN
gs-3955	140	58	,	,	PUNCT
gs-3955	140	59	2021	2021	NUM
gs-3955	140	60	)	)	PUNCT
gs-3955	140	61	.	.	PUNCT
gs-3955	141	1	collectively	collectively	ADV
gs-3955	141	2	,	,	PUNCT
gs-3955	141	3	these	these	DET
gs-3955	141	4	species	specie	NOUN
gs-3955	141	5	demonstrate	demonstrate	VERB
gs-3955	141	6	a	a	DET
gs-3955	141	7	convergence	convergence	NOUN
gs-3955	141	8	on	on	ADP
gs-3955	141	9	preserving	preserve	VERB
gs-3955	141	10	stem	stem	NOUN
gs-3955	141	11	cell	cell	NOUN
gs-3955	141	12	potency	potency	NOUN
gs-3955	141	13	and	and	CCONJ
gs-3955	141	14	regeneration	regeneration	NOUN
gs-3955	141	15	as	as	ADP
gs-3955	141	16	a	a	DET
gs-3955	141	17	critical	critical	ADJ
gs-3955	141	18	strategy	strategy	NOUN
gs-3955	141	19	against	against	ADP
gs-3955	141	20	senescence	senescence	NOUN
gs-3955	141	21	.	.	PUNCT
gs-3955	142	1	enhanced	enhance	VERB
gs-3955	142	2	dna	dna	PROPN
gs-3955	142	3	repair	repair	NOUN
gs-3955	142	4	mechanisms	mechanism	NOUN
gs-3955	142	5	genome	genome	NOUN
gs-3955	142	6	integrity	integrity	NOUN
gs-3955	142	7	preservation	preservation	NOUN
gs-3955	142	8	represents	represent	VERB
gs-3955	142	9	a	a	DET
gs-3955	142	10	fundamental	fundamental	ADJ
gs-3955	142	11	pillar	pillar	NOUN
gs-3955	142	12	of	of	ADP
gs-3955	142	13	longevity	longevity	NOUN
gs-3955	142	14	in	in	ADP
gs-3955	142	15	negligibly	negligibly	ADV
gs-3955	142	16	senescent	senescent	ADJ
gs-3955	142	17	species	specie	NOUN
gs-3955	142	18	.	.	PUNCT
gs-3955	143	1	the	the	DET
gs-3955	143	2	naked	naked	ADJ
gs-3955	143	3	mole	mole	ADJ
gs-3955	143	4	-	-	PUNCT
gs-3955	143	5	rat	rat	NOUN
gs-3955	143	6	shows	show	NOUN
gs-3955	143	7	heightened	heighten	VERB
gs-3955	143	8	expression	expression	NOUN
gs-3955	143	9	and	and	CCONJ
gs-3955	143	10	activity	activity	NOUN
gs-3955	143	11	of	of	ADP
gs-3955	143	12	dna	dna	PROPN
gs-3955	143	13	damage	damage	NOUN
gs-3955	143	14	response	response	NOUN
gs-3955	143	15	(	(	PUNCT
gs-3955	143	16	ddr	ddr	NOUN
gs-3955	143	17	)	)	PUNCT
gs-3955	143	18	genes	gene	NOUN
gs-3955	143	19	,	,	PUNCT
gs-3955	143	20	including	include	VERB
gs-3955	143	21	atm	atm	NOUN
gs-3955	143	22	,	,	PUNCT
gs-3955	143	23	brca1/2	brca1/2	ADJ
gs-3955	143	24	,	,	PUNCT
gs-3955	143	25	and	and	CCONJ
gs-3955	143	26	tp53	tp53	PROPN
gs-3955	143	27	,	,	PUNCT
gs-3955	143	28	which	which	PRON
gs-3955	143	29	favor	favor	VERB
gs-3955	143	30	repair	repair	NOUN
gs-3955	143	31	and	and	CCONJ
gs-3955	143	32	cell	cell	NOUN
gs-3955	143	33	cycle	cycle	NOUN
gs-3955	143	34	arrest	arrest	NOUN
gs-3955	143	35	rather	rather	ADV
gs-3955	143	36	than	than	ADP
gs-3955	143	37	apoptosis	apoptosis	NOUN
gs-3955	143	38	,	,	PUNCT
gs-3955	143	39	reducing	reduce	VERB
gs-3955	143	40	cellular	cellular	ADJ
gs-3955	143	41	attrition	attrition	NOUN
gs-3955	143	42	and	and	CCONJ
gs-3955	143	43	preserving	preserve	VERB
gs-3955	143	44	tissue	tissue	NOUN
gs-3955	143	45	architecture	architecture	NOUN
gs-3955	143	46	(	(	PUNCT
gs-3955	143	47	petruseva	petruseva	PROPN
gs-3955	143	48	et	et	PROPN
gs-3955	143	49	al	al	PROPN
gs-3955	143	50	.	.	PROPN
gs-3955	143	51	,	,	PUNCT
gs-3955	143	52	2017	2017	NUM
gs-3955	143	53	;	;	PUNCT
gs-3955	143	54	macrae	macrae	PROPN
gs-3955	143	55	et	et	PROPN
gs-3955	143	56	al	al	PROPN
gs-3955	143	57	.	.	PROPN
gs-3955	143	58	,	,	PUNCT
gs-3955	143	59	2015	2015	NUM
gs-3955	143	60	)	)	PUNCT
gs-3955	143	61	.	.	PUNCT
gs-3955	144	1	enhanced	enhance	VERB
gs-3955	144	2	functionality	functionality	NOUN
gs-3955	144	3	of	of	ADP
gs-3955	144	4	p53	p53	NOUN
gs-3955	144	5	variants	variant	NOUN
gs-3955	144	6	in	in	ADP
gs-3955	144	7	mole	mole	NOUN
gs-3955	144	8	-	-	PUNCT
gs-3955	144	9	rats	rat	NOUN
gs-3955	144	10	supports	support	VERB
gs-3955	144	11	genomic	genomic	NOUN
gs-3955	144	12	stability	stability	NOUN
gs-3955	144	13	without	without	ADP
gs-3955	144	14	triggering	trigger	VERB
gs-3955	144	15	cancerous	cancerous	ADJ
gs-3955	144	16	growth	growth	NOUN
gs-3955	144	17	,	,	PUNCT
gs-3955	144	18	illustrating	illustrate	VERB
gs-3955	144	19	an	an	DET
gs-3955	144	20	evolved	evolved	ADJ
gs-3955	144	21	balance	balance	NOUN
gs-3955	144	22	between	between	ADP
gs-3955	144	23	tumor	tumor	NOUN
gs-3955	144	24	suppression	suppression	NOUN
gs-3955	144	25	and	and	CCONJ
gs-3955	144	26	longevity	longevity	NOUN
gs-3955	144	27	(	(	PUNCT
gs-3955	144	28	petruseva	petruseva	PROPN
gs-3955	144	29	et	et	PROPN
gs-3955	144	30	al	al	PROPN
gs-3955	144	31	.	.	PROPN
gs-3955	144	32	,	,	PUNCT
gs-3955	144	33	2017	2017	NUM
gs-3955	144	34	)	)	PUNCT
gs-3955	144	35	.	.	PUNCT
gs-3955	145	1	regarding	regard	VERB
gs-3955	145	2	specific	specific	ADJ
gs-3955	145	3	repair	repair	NOUN
gs-3955	145	4	pathways	pathway	NOUN
gs-3955	145	5	,	,	PUNCT
gs-3955	145	6	naked	naked	ADJ
gs-3955	145	7	mole	mole	NOUN
gs-3955	145	8	-	-	PUNCT
gs-3955	145	9	rats	rat	NOUN
gs-3955	145	10	excel	excel	VERB
gs-3955	145	11	in	in	ADP
gs-3955	145	12	homologous	homologous	ADJ
gs-3955	145	13	recombination	recombination	NOUN
gs-3955	145	14	with	with	ADP
gs-3955	145	15	upregulated	upregulated	ADJ
gs-3955	145	16	rad51	rad51	NOUN
gs-3955	145	17	and	and	CCONJ
gs-3955	145	18	palb2	palb2	NOUN
gs-3955	145	19	proteins	protein	NOUN
gs-3955	145	20	,	,	PUNCT
gs-3955	145	21	as	as	ADV
gs-3955	145	22	well	well	ADV
gs-3955	145	23	as	as	ADP
gs-3955	145	24	base	base	NOUN
gs-3955	145	25	excision	excision	NOUN
gs-3955	145	26	repair	repair	NOUN
gs-3955	145	27	and	and	CCONJ
gs-3955	145	28	nucleotide	nucleotide	ADJ
gs-3955	145	29	excision	excision	NOUN
gs-3955	145	30	repair	repair	NOUN
gs-3955	145	31	,	,	PUNCT
gs-3955	145	32	exhibiting	exhibit	VERB
gs-3955	145	33	faster	fast	ADJ
gs-3955	145	34	correction	correction	NOUN
gs-3955	145	35	of	of	ADP
gs-3955	145	36	oxidative	oxidative	ADJ
gs-3955	145	37	and	and	CCONJ
gs-3955	145	38	bulky	bulky	ADJ
gs-3955	145	39	dna	dna	NOUN
gs-3955	145	40	lesions	lesion	NOUN
gs-3955	145	41	compared	compare	VERB
gs-3955	145	42	to	to	ADP
gs-3955	145	43	shorter	shorter	ADV
gs-3955	145	44	-	-	PUNCT
gs-3955	145	45	lived	live	VERB
gs-3955	145	46	rodents	rodent	NOUN
gs-3955	145	47	(	(	PUNCT
gs-3955	145	48	narayan	narayan	PROPN
gs-3955	145	49	et	et	PROPN
gs-3955	145	50	al	al	PROPN
gs-3955	145	51	.	.	PROPN
gs-3955	145	52	,	,	PUNCT
gs-3955	145	53	2021	2021	NUM
gs-3955	145	54	;	;	PUNCT
gs-3955	145	55	macrae	macrae	PROPN
gs-3955	145	56	et	et	PROPN
gs-3955	145	57	al	al	PROPN
gs-3955	145	58	.	.	PROPN
gs-3955	145	59	,	,	PUNCT
gs-3955	145	60	2015	2015	NUM
gs-3955	145	61	)	)	PUNCT
gs-3955	145	62	.	.	PUNCT
gs-3955	146	1	similarly	similarly	ADV
gs-3955	146	2	,	,	PUNCT
gs-3955	146	3	the	the	DET
gs-3955	146	4	greenland	greenland	PROPN
gs-3955	146	5	shark	shark	NOUN
gs-3955	146	6	's	's	PART
gs-3955	146	7	dna	dna	PROPN
gs-3955	146	8	polymerases	polymerase	NOUN
gs-3955	146	9	(	(	PUNCT
gs-3955	146	10	rev1	rev1	PROPN
gs-3955	146	11	,	,	PUNCT
gs-3955	146	12	polh	polh	NOUN
gs-3955	146	13	,	,	PUNCT
gs-3955	146	14	polk	polk	NOUN
gs-3955	146	15	)	)	PUNCT
gs-3955	146	16	demonstrate	demonstrate	VERB
gs-3955	146	17	increased	increase	VERB
gs-3955	146	18	fidelity	fidelity	NOUN
gs-3955	146	19	and	and	CCONJ
gs-3955	146	20	translesion	translesion	NOUN
gs-3955	146	21	synthesis	synthesis	NOUN
gs-3955	146	22	capacity	capacity	NOUN
gs-3955	146	23	,	,	PUNCT
gs-3955	146	24	allowing	allow	VERB
gs-3955	146	25	lesion	lesion	NOUN
gs-3955	146	26	bypass	bypass	NOUN
gs-3955	146	27	without	without	ADP
gs-3955	146	28	mutations	mutation	NOUN
gs-3955	146	29	(	(	PUNCT
gs-3955	146	30	petruseva	petruseva	PROPN
gs-3955	146	31	et	et	PROPN
gs-3955	146	32	al	al	PROPN
gs-3955	146	33	.	.	PROPN
gs-3955	146	34	,	,	PUNCT
gs-3955	146	35	2017	2017	NUM
gs-3955	146	36	)	)	PUNCT
gs-3955	146	37	.	.	PUNCT
gs-3955	147	1	in	in	ADP
gs-3955	147	2	bivalves	bivalve	NOUN
gs-3955	147	3	like	like	ADP
gs-3955	147	4	arctica	arctica	PROPN
gs-3955	147	5	islandica	islandica	PROPN
gs-3955	147	6	,	,	PUNCT
gs-3955	147	7	nucleotide	nucleotide	ADJ
gs-3955	147	8	excision	excision	NOUN
gs-3955	147	9	repair	repair	NOUN
gs-3955	147	10	is	be	AUX
gs-3955	147	11	upregulated	upregulate	VERB
gs-3955	147	12	with	with	ADP
gs-3955	147	13	enhanced	enhance	VERB
gs-3955	147	14	xpa	xpa	PROPN
gs-3955	147	15	-	-	PUNCT
gs-3955	147	16	xpg	xpg	PROPN
gs-3955	147	17	complex	complex	ADJ
gs-3955	147	18	activities	activity	NOUN
gs-3955	147	19	,	,	PUNCT
gs-3955	147	20	effectively	effectively	ADV
gs-3955	147	21	repairing	repair	VERB
gs-3955	147	22	uvand	uvand	ADJ
gs-3955	147	23	chemically	chemically	ADV
gs-3955	147	24	-	-	PUNCT
gs-3955	147	25	induced	induce	VERB
gs-3955	147	26	dna	dna	NOUN
gs-3955	147	27	damage	damage	NOUN
gs-3955	147	28	,	,	PUNCT
gs-3955	147	29	contributing	contribute	VERB
gs-3955	147	30	to	to	ADP
gs-3955	147	31	its	its	PRON
gs-3955	147	32	longevity	longevity	NOUN
gs-3955	147	33	(	(	PUNCT
gs-3955	147	34	narayan	narayan	PROPN
gs-3955	147	35	et	et	PROPN
gs-3955	147	36	al	al	PROPN
gs-3955	147	37	.	.	PROPN
gs-3955	147	38	,	,	PUNCT
gs-3955	147	39	2021	2021	NUM
gs-3955	147	40	)	)	PUNCT
gs-3955	147	41	.	.	PUNCT
gs-3955	148	1	furthermore	furthermore	ADV
gs-3955	148	2	,	,	PUNCT
gs-3955	148	3	pirna	pirna	NOUN
gs-3955	148	4	pathways	pathway	NOUN
gs-3955	148	5	,	,	PUNCT
gs-3955	148	6	which	which	PRON
gs-3955	148	7	silence	silence	VERB
gs-3955	148	8	transposable	transposable	ADJ
gs-3955	148	9	elements	element	NOUN
gs-3955	148	10	and	and	CCONJ
gs-3955	148	11	preserve	preserve	VERB
gs-3955	148	12	genome	genome	NOUN
gs-3955	148	13	integrity	integrity	NOUN
gs-3955	148	14	,	,	PUNCT
gs-3955	148	15	are	be	AUX
gs-3955	148	16	expanded	expand	VERB
gs-3955	148	17	in	in	ADP
gs-3955	148	18	species	specie	NOUN
gs-3955	148	19	such	such	ADJ
gs-3955	148	20	as	as	ADP
gs-3955	148	21	sebastes	sebaste	NOUN
gs-3955	148	22	aleutianus	aleutianus	NOUN
gs-3955	148	23	,	,	PUNCT
gs-3955	148	24	where	where	SCONJ
gs-3955	148	25	piwil2	piwil2	PROPN
gs-3955	148	26	and	and	CCONJ
gs-3955	148	27	tdrd9	tdrd9	PROPN
gs-3955	148	28	support	support	NOUN
gs-3955	148	29	germline	germline	NOUN
gs-3955	148	30	stability	stability	NOUN
gs-3955	148	31	(	(	PUNCT
gs-3955	148	32	macrae	macrae	PROPN
gs-3955	148	33	et	et	PROPN
gs-3955	148	34	al	al	PROPN
gs-3955	148	35	.	.	PROPN
gs-3955	148	36	,	,	PUNCT
gs-3955	148	37	2015	2015	NUM
gs-3955	148	38	)	)	PUNCT
gs-3955	148	39	.	.	PUNCT
gs-3955	149	1	though	though	SCONJ
gs-3955	149	2	not	not	PART
gs-3955	149	3	fully	fully	ADV
gs-3955	149	4	characterized	characterize	VERB
gs-3955	149	5	in	in	ADP
gs-3955	149	6	all	all	DET
gs-3955	149	7	negligibly	negligibly	ADV
gs-3955	149	8	senescent	senescent	ADJ
gs-3955	149	9	species	specie	NOUN
gs-3955	149	10	,	,	PUNCT
gs-3955	149	11	these	these	DET
gs-3955	149	12	small	small	ADJ
gs-3955	149	13	rna	rna	NOUN
gs-3955	149	14	-	-	PUNCT
gs-3955	149	15	mediated	mediate	VERB
gs-3955	149	16	mechanisms	mechanism	NOUN
gs-3955	149	17	represent	represent	VERB
gs-3955	149	18	another	another	DET
gs-3955	149	19	conserved	conserved	ADJ
gs-3955	149	20	axis	axis	NOUN
gs-3955	149	21	of	of	ADP
gs-3955	149	22	genome	genome	NOUN
gs-3955	149	23	defense	defense	NOUN
gs-3955	149	24	.	.	PUNCT
gs-3955	150	1	these	these	DET
gs-3955	150	2	combined	combine	VERB
gs-3955	150	3	dna	dna	PROPN
gs-3955	150	4	repair	repair	NOUN
gs-3955	150	5	enhancements	enhancement	NOUN
gs-3955	150	6	yield	yield	VERB
gs-3955	150	7	exceptionally	exceptionally	ADV
gs-3955	150	8	low	low	ADJ
gs-3955	150	9	somatic	somatic	ADJ
gs-3955	150	10	mutation	mutation	NOUN
gs-3955	150	11	rates	rate	NOUN
gs-3955	150	12	in	in	ADP
gs-3955	150	13	long	long	ADV
gs-3955	150	14	-	-	PUNCT
gs-3955	150	15	lived	live	VERB
gs-3955	150	16	species	specie	NOUN
gs-3955	150	17	,	,	PUNCT
gs-3955	150	18	underscoring	underscore	VERB
gs-3955	150	19	genome	genome	NOUN
gs-3955	150	20	maintenance	maintenance	NOUN
gs-3955	150	21	as	as	ADP
gs-3955	150	22	a	a	DET
gs-3955	150	23	cornerstone	cornerstone	NOUN
gs-3955	150	24	of	of	ADP
gs-3955	150	25	negligible	negligible	ADJ
gs-3955	150	26	senescence	senescence	NOUN
gs-3955	150	27	(	(	PUNCT
gs-3955	150	28	narayan	narayan	PROPN
gs-3955	150	29	et	et	PROPN
gs-3955	150	30	al	al	PROPN
gs-3955	150	31	.	.	PROPN
gs-3955	150	32	,	,	PUNCT
gs-3955	150	33	2021	2021	NUM
gs-3955	150	34	)	)	PUNCT
gs-3955	150	35	.	.	PUNCT
gs-3955	151	1	oxidative	oxidative	ADJ
gs-3955	151	2	stress	stress	NOUN
gs-3955	151	3	resistance	resistance	NOUN
gs-3955	151	4	contrary	contrary	ADJ
gs-3955	151	5	to	to	ADP
gs-3955	151	6	earlier	early	ADJ
gs-3955	151	7	models	model	NOUN
gs-3955	151	8	positing	posit	VERB
gs-3955	151	9	reduced	reduce	VERB
gs-3955	151	10	ros	ros	PROPN
gs-3955	151	11	generation	generation	NOUN
gs-3955	151	12	as	as	ADP
gs-3955	151	13	the	the	DET
gs-3955	151	14	basis	basis	NOUN
gs-3955	151	15	for	for	ADP
gs-3955	151	16	longevity	longevity	NOUN
gs-3955	151	17	,	,	PUNCT
gs-3955	151	18	many	many	ADJ
gs-3955	151	19	negligibly	negligibly	ADV
gs-3955	151	20	senescent	senescent	ADJ
gs-3955	151	21	organisms	organism	NOUN
gs-3955	151	22	exhibit	exhibit	VERB
gs-3955	151	23	elevated	elevated	ADJ
gs-3955	151	24	reactive	reactive	ADJ
gs-3955	151	25	oxygen	oxygen	NOUN
gs-3955	151	26	species	specie	NOUN
gs-3955	151	27	(	(	PUNCT
gs-3955	151	28	ros	ros	PROPN
gs-3955	151	29	)	)	PUNCT
gs-3955	151	30	production	production	NOUN
gs-3955	151	31	but	but	CCONJ
gs-3955	151	32	show	show	VERB
gs-3955	151	33	exceptional	exceptional	ADJ
gs-3955	151	34	resilience	resilience	NOUN
gs-3955	151	35	to	to	PART
gs-3955	151	36	oxidative	oxidative	VERB
gs-3955	151	37	damage	damage	NOUN
gs-3955	151	38	(	(	PUNCT
gs-3955	151	39	perez	perez	PROPN
gs-3955	151	40	et	et	PROPN
gs-3955	151	41	al	al	PROPN
gs-3955	151	42	.	.	PROPN
gs-3955	151	43	,	,	PUNCT
gs-3955	151	44	2009	2009	NUM
gs-3955	151	45	;	;	PUNCT
gs-3955	151	46	lewis	lewis	PROPN
gs-3955	151	47	et	et	PROPN
gs-3955	151	48	al	al	PROPN
gs-3955	151	49	.	.	PROPN
gs-3955	151	50	,	,	PUNCT
gs-3955	151	51	2012	2012	NUM
gs-3955	151	52	)	)	PUNCT
gs-3955	151	53	.	.	PUNCT
gs-3955	152	1	for	for	ADP
gs-3955	152	2	example	example	NOUN
gs-3955	152	3	,	,	PUNCT
gs-3955	152	4	naked	naked	ADJ
gs-3955	152	5	mole	mole	ADJ
gs-3955	152	6	-	-	PUNCT
gs-3955	152	7	rat	rat	NOUN
gs-3955	152	8	myocardium	myocardium	NOUN
gs-3955	152	9	produces	produce	VERB
gs-3955	152	10	2.3	2.3	NUM
gs-3955	152	11	nmol	nmol	NOUN
gs-3955	152	12	h₂o₂/min	h₂o₂/min	NOUN
gs-3955	152	13	/	/	SYM
gs-3955	152	14	mg	mg	PROPN
gs-3955	152	15	protein	protein	NOUN
gs-3955	152	16	,	,	PUNCT
gs-3955	152	17	georgian	georgian	ADJ
gs-3955	152	18	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	152	19	მეცნიერები	მეცნიერები	NOUN
gs-3955	152	20	ტ.	ტ.	VERB
gs-3955	152	21	7	7	NUM
gs-3955	152	22	n	n	PRON
gs-3955	152	23	3	3	NUM
gs-3955	152	24	,	,	PUNCT
gs-3955	152	25	2025	2025	NUM
gs-3955	152	26	365	365	NUM
gs-3955	152	27	exceeding	exceed	VERB
gs-3955	152	28	that	that	PRON
gs-3955	152	29	of	of	ADP
gs-3955	152	30	typical	typical	ADJ
gs-3955	152	31	rodents	rodent	NOUN
gs-3955	152	32	,	,	PUNCT
gs-3955	152	33	yet	yet	CCONJ
gs-3955	152	34	oxidative	oxidative	ADJ
gs-3955	152	35	damage	damage	NOUN
gs-3955	152	36	markers	marker	NOUN
gs-3955	152	37	remain	remain	VERB
gs-3955	152	38	minimal	minimal	ADJ
gs-3955	152	39	due	due	ADP
gs-3955	152	40	to	to	ADP
gs-3955	152	41	robust	robust	ADJ
gs-3955	152	42	redox	redox	ADJ
gs-3955	152	43	buffering	buffering	NOUN
gs-3955	152	44	and	and	CCONJ
gs-3955	152	45	repair	repair	NOUN
gs-3955	152	46	systems	system	NOUN
gs-3955	152	47	(	(	PUNCT
gs-3955	152	48	lewis	lewis	PROPN
gs-3955	152	49	et	et	PROPN
gs-3955	152	50	al	al	PROPN
gs-3955	152	51	.	.	PROPN
gs-3955	152	52	,	,	PUNCT
gs-3955	152	53	2012	2012	NUM
gs-3955	152	54	)	)	PUNCT
gs-3955	152	55	.	.	PUNCT
gs-3955	153	1	the	the	DET
gs-3955	153	2	ocean	ocean	PROPN
gs-3955	153	3	quahog	quahog	PROPN
gs-3955	153	4	(	(	PUNCT
gs-3955	153	5	arctica	arctica	PROPN
gs-3955	153	6	islandica	islandica	PROPN
gs-3955	153	7	)	)	PUNCT
gs-3955	153	8	showcases	showcase	VERB
gs-3955	153	9	reduced	reduce	VERB
gs-3955	153	10	mitochondrial	mitochondrial	ADJ
gs-3955	153	11	ros	ros	NOUN
gs-3955	153	12	production	production	NOUN
gs-3955	153	13	and	and	CCONJ
gs-3955	153	14	highly	highly	ADV
gs-3955	153	15	efficient	efficient	ADJ
gs-3955	153	16	hydrogen	hydrogen	NOUN
gs-3955	153	17	peroxide	peroxide	NOUN
gs-3955	153	18	detoxification	detoxification	NOUN
gs-3955	153	19	enzymes	enzyme	NOUN
gs-3955	153	20	,	,	PUNCT
gs-3955	153	21	contributing	contribute	VERB
gs-3955	153	22	to	to	ADP
gs-3955	153	23	its	its	PRON
gs-3955	153	24	400	400	NUM
gs-3955	153	25	+	+	NUM
gs-3955	153	26	year	year	NOUN
gs-3955	153	27	lifespan	lifespan	NOUN
gs-3955	153	28	(	(	PUNCT
gs-3955	153	29	munro	munro	PROPN
gs-3955	153	30	et	et	PROPN
gs-3955	153	31	al	al	PROPN
gs-3955	153	32	.	.	PROPN
gs-3955	153	33	,	,	PUNCT
gs-3955	153	34	2012	2012	NUM
gs-3955	153	35	)	)	PUNCT
gs-3955	153	36	.	.	PUNCT
gs-3955	154	1	its	its	PRON
gs-3955	154	2	mitochondrial	mitochondrial	ADJ
gs-3955	154	3	membranes	membrane	NOUN
gs-3955	154	4	are	be	AUX
gs-3955	154	5	enriched	enrich	VERB
gs-3955	154	6	with	with	ADP
gs-3955	154	7	monounsaturated	monounsaturated	ADJ
gs-3955	154	8	and	and	CCONJ
gs-3955	154	9	saturated	saturated	ADJ
gs-3955	154	10	fatty	fatty	NOUN
gs-3955	154	11	acids	acid	NOUN
gs-3955	154	12	,	,	PUNCT
gs-3955	154	13	lowering	lower	VERB
gs-3955	154	14	peroxidation	peroxidation	NOUN
gs-3955	154	15	indices	index	NOUN
gs-3955	154	16	and	and	CCONJ
gs-3955	154	17	stabilizing	stabilize	VERB
gs-3955	154	18	membrane	membrane	NOUN
gs-3955	154	19	integrity	integrity	NOUN
gs-3955	154	20	against	against	ADP
gs-3955	154	21	oxidative	oxidative	ADJ
gs-3955	154	22	stress	stress	NOUN
gs-3955	154	23	(	(	PUNCT
gs-3955	154	24	munro	munro	PROPN
gs-3955	154	25	et	et	PROPN
gs-3955	154	26	al	al	PROPN
gs-3955	154	27	.	.	PROPN
gs-3955	154	28	,	,	PUNCT
gs-3955	154	29	2012	2012	NUM
gs-3955	154	30	)	)	PUNCT
gs-3955	154	31	.	.	PUNCT
gs-3955	155	1	this	this	DET
gs-3955	155	2	lipid	lipid	NOUN
gs-3955	155	3	profile	profile	NOUN
gs-3955	155	4	parallels	parallel	VERB
gs-3955	155	5	observations	observation	NOUN
gs-3955	155	6	in	in	ADP
gs-3955	155	7	naked	naked	ADJ
gs-3955	155	8	mole	mole	NOUN
gs-3955	155	9	-	-	PUNCT
gs-3955	155	10	rats	rat	NOUN
gs-3955	155	11	,	,	PUNCT
gs-3955	155	12	whose	whose	DET
gs-3955	155	13	cardiolipin	cardiolipin	NOUN
gs-3955	155	14	acyl	acyl	NOUN
gs-3955	155	15	chains	chain	NOUN
gs-3955	155	16	are	be	AUX
gs-3955	155	17	predominantly	predominantly	ADV
gs-3955	155	18	monounsaturated	monounsaturate	VERB
gs-3955	155	19	,	,	PUNCT
gs-3955	155	20	conferring	confer	VERB
gs-3955	155	21	similar	similar	ADJ
gs-3955	155	22	protection	protection	NOUN
gs-3955	155	23	(	(	PUNCT
gs-3955	155	24	perez	perez	PROPN
gs-3955	155	25	et	et	PROPN
gs-3955	155	26	al	al	PROPN
gs-3955	155	27	.	.	PROPN
gs-3955	155	28	,	,	PUNCT
gs-3955	155	29	2009	2009	NUM
gs-3955	155	30	)	)	PUNCT
gs-3955	155	31	.	.	PUNCT
gs-3955	156	1	additionally	additionally	ADV
gs-3955	156	2	,	,	PUNCT
gs-3955	156	3	these	these	DET
gs-3955	156	4	species	specie	NOUN
gs-3955	156	5	exhibit	exhibit	AUX
gs-3955	156	6	expanded	expand	VERB
gs-3955	156	7	enzymatic	enzymatic	ADJ
gs-3955	156	8	detoxification	detoxification	NOUN
gs-3955	156	9	capabilities	capability	NOUN
gs-3955	156	10	,	,	PUNCT
gs-3955	156	11	including	include	VERB
gs-3955	156	12	elevated	elevated	ADJ
gs-3955	156	13	glutathione	glutathione	NOUN
gs-3955	156	14	concentrations	concentration	NOUN
gs-3955	156	15	(	(	PUNCT
gs-3955	156	16	as	as	ADP
gs-3955	156	17	in	in	ADP
gs-3955	156	18	sea	sea	NOUN
gs-3955	156	19	urchins	urchin	NOUN
gs-3955	156	20	)	)	PUNCT
gs-3955	156	21	and	and	CCONJ
gs-3955	156	22	novel	novel	NOUN
gs-3955	156	23	carbonyl	carbonyl	NOUN
gs-3955	156	24	reductases	reductase	NOUN
gs-3955	156	25	(	(	PUNCT
gs-3955	156	26	e.g.	e.g.	ADV
gs-3955	156	27	,	,	PUNCT
gs-3955	156	28	hvcr1	hvcr1	NOUN
gs-3955	156	29	-	-	PUNCT
gs-3955	156	30	4	4	NUM
gs-3955	156	31	in	in	ADP
gs-3955	156	32	hydra	hydra	PROPN
gs-3955	156	33	)	)	PUNCT
gs-3955	156	34	that	that	PRON
gs-3955	156	35	mitigate	mitigate	VERB
gs-3955	156	36	secondary	secondary	ADJ
gs-3955	156	37	oxidative	oxidative	ADJ
gs-3955	156	38	damage	damage	NOUN
gs-3955	156	39	products	product	NOUN
gs-3955	156	40	such	such	ADJ
gs-3955	156	41	as	as	ADP
gs-3955	156	42	4	4	NUM
gs-3955	156	43	-	-	PUNCT
gs-3955	156	44	hydroxynonenal	hydroxynonenal	NOUN
gs-3955	156	45	(	(	PUNCT
gs-3955	156	46	narayan	narayan	PROPN
gs-3955	156	47	et	et	PROPN
gs-3955	156	48	al	al	PROPN
gs-3955	156	49	.	.	PROPN
gs-3955	156	50	,	,	PUNCT
gs-3955	156	51	2021	2021	NUM
gs-3955	156	52	)	)	PUNCT
gs-3955	156	53	.	.	PUNCT
gs-3955	157	1	this	this	DET
gs-3955	157	2	combination	combination	NOUN
gs-3955	157	3	of	of	ADP
gs-3955	157	4	membrane	membrane	NOUN
gs-3955	157	5	composition	composition	NOUN
gs-3955	157	6	and	and	CCONJ
gs-3955	157	7	enzymatic	enzymatic	ADJ
gs-3955	157	8	defense	defense	NOUN
gs-3955	157	9	uncouples	uncouple	NOUN
gs-3955	157	10	ros	ros	PROPN
gs-3955	157	11	production	production	NOUN
gs-3955	157	12	from	from	ADP
gs-3955	157	13	cellular	cellular	ADJ
gs-3955	157	14	damage	damage	NOUN
gs-3955	157	15	,	,	PUNCT
gs-3955	157	16	supporting	support	VERB
gs-3955	157	17	cellular	cellular	ADJ
gs-3955	157	18	homeostasis	homeostasis	NOUN
gs-3955	157	19	despite	despite	SCONJ
gs-3955	157	20	a	a	DET
gs-3955	157	21	pro	pro	ADJ
gs-3955	157	22	-	-	ADJ
gs-3955	157	23	oxidant	oxidant	ADJ
gs-3955	157	24	intracellular	intracellular	ADJ
gs-3955	157	25	environment	environment	NOUN
gs-3955	157	26	.	.	PUNCT
gs-3955	158	1	these	these	DET
gs-3955	158	2	oxidative	oxidative	ADJ
gs-3955	158	3	stress	stress	NOUN
gs-3955	158	4	resilience	resilience	NOUN
gs-3955	158	5	mechanisms	mechanism	NOUN
gs-3955	158	6	are	be	AUX
gs-3955	158	7	often	often	ADV
gs-3955	158	8	accompanied	accompany	VERB
gs-3955	158	9	by	by	ADP
gs-3955	158	10	increased	increase	VERB
gs-3955	158	11	autophagy	autophagy	NOUN
gs-3955	158	12	and	and	CCONJ
gs-3955	158	13	proteasomal	proteasomal	ADJ
gs-3955	158	14	clearance	clearance	NOUN
gs-3955	158	15	systems	system	NOUN
gs-3955	158	16	that	that	PRON
gs-3955	158	17	degrade	degrade	VERB
gs-3955	158	18	oxidized	oxidized	ADJ
gs-3955	158	19	proteins	protein	NOUN
gs-3955	158	20	and	and	CCONJ
gs-3955	158	21	damaged	damaged	ADJ
gs-3955	158	22	organelles	organelle	NOUN
gs-3955	158	23	,	,	PUNCT
gs-3955	158	24	further	far	ADV
gs-3955	158	25	maintaining	maintain	VERB
gs-3955	158	26	proteostasis	proteostasis	NOUN
gs-3955	158	27	(	(	PUNCT
gs-3955	158	28	pirotte	pirotte	NOUN
gs-3955	158	29	et	et	PROPN
gs-3955	158	30	al	al	PROPN
gs-3955	158	31	.	.	PROPN
gs-3955	158	32	,	,	PUNCT
gs-3955	158	33	2015	2015	NUM
gs-3955	158	34	;	;	PUNCT
gs-3955	158	35	tkemaladze	tkemaladze	NOUN
gs-3955	158	36	,	,	PUNCT
gs-3955	158	37	2023	2023	NUM
gs-3955	158	38	)	)	PUNCT
gs-3955	158	39	.	.	PUNCT
gs-3955	159	1	the	the	DET
gs-3955	159	2	vanguard	vanguard	NOUN
gs-3955	159	3	of	of	ADP
gs-3955	159	4	biological	biological	ADJ
gs-3955	159	5	immortality	immortality	NOUN
gs-3955	159	6	and	and	CCONJ
gs-3955	159	7	its	its	PRON
gs-3955	159	8	implications	implication	NOUN
gs-3955	159	9	for	for	ADP
gs-3955	159	10	redefining	redefine	VERB
gs-3955	159	11	human	human	ADJ
gs-3955	159	12	aging	aging	NOUN
gs-3955	159	13	the	the	DET
gs-3955	159	14	existence	existence	NOUN
gs-3955	159	15	of	of	ADP
gs-3955	159	16	organisms	organism	NOUN
gs-3955	159	17	exhibiting	exhibit	VERB
gs-3955	159	18	negligible	negligible	ADJ
gs-3955	159	19	senescence	senescence	NOUN
gs-3955	159	20	stands	stand	VERB
gs-3955	159	21	as	as	ADP
gs-3955	159	22	one	one	NUM
gs-3955	159	23	of	of	ADP
gs-3955	159	24	biology	biology	NOUN
gs-3955	159	25	's	's	PART
gs-3955	159	26	most	most	ADV
gs-3955	159	27	profound	profound	ADJ
gs-3955	159	28	revelations	revelation	NOUN
gs-3955	159	29	,	,	PUNCT
gs-3955	159	30	fundamentally	fundamentally	ADV
gs-3955	159	31	challenging	challenge	VERB
gs-3955	159	32	the	the	DET
gs-3955	159	33	notion	notion	NOUN
gs-3955	159	34	that	that	SCONJ
gs-3955	159	35	aging	aging	NOUN
gs-3955	159	36	represents	represent	VERB
gs-3955	159	37	an	an	DET
gs-3955	159	38	inescapable	inescapable	ADJ
gs-3955	159	39	biological	biological	ADJ
gs-3955	159	40	imperative	imperative	NOUN
gs-3955	159	41	.	.	PUNCT
gs-3955	160	1	these	these	DET
gs-3955	160	2	species	specie	NOUN
gs-3955	160	3	—	—	PUNCT
gs-3955	160	4	spanning	span	VERB
gs-3955	160	5	hydra	hydra	PROPN
gs-3955	160	6	,	,	PUNCT
gs-3955	160	7	greenland	greenland	PROPN
gs-3955	160	8	sharks	sharks	PROPN
gs-3955	160	9	,	,	PUNCT
gs-3955	160	10	naked	naked	ADJ
gs-3955	160	11	mole	mole	NOUN
gs-3955	160	12	-	-	PUNCT
gs-3955	160	13	rats	rat	NOUN
gs-3955	160	14	,	,	PUNCT
gs-3955	160	15	ocean	ocean	NOUN
gs-3955	160	16	quahogs	quahog	NOUN
gs-3955	160	17	,	,	PUNCT
gs-3955	160	18	olms	olm	NOUN
gs-3955	160	19	,	,	PUNCT
gs-3955	160	20	and	and	CCONJ
gs-3955	160	21	planarians	planarian	NOUN
gs-3955	160	22	—	—	PUNCT
gs-3955	160	23	collectively	collectively	ADV
gs-3955	160	24	demonstrate	demonstrate	VERB
gs-3955	160	25	that	that	SCONJ
gs-3955	160	26	the	the	DET
gs-3955	160	27	progressive	progressive	ADJ
gs-3955	160	28	physiological	physiological	ADJ
gs-3955	160	29	decline	decline	NOUN
gs-3955	160	30	characterizing	characterize	VERB
gs-3955	160	31	most	most	ADJ
gs-3955	160	32	multicellular	multicellular	ADJ
gs-3955	160	33	life	life	NOUN
gs-3955	160	34	is	be	AUX
gs-3955	160	35	not	not	PART
gs-3955	160	36	an	an	DET
gs-3955	160	37	immutable	immutable	ADJ
gs-3955	160	38	law	law	NOUN
gs-3955	160	39	of	of	ADP
gs-3955	160	40	nature	nature	NOUN
gs-3955	160	41	,	,	PUNCT
gs-3955	160	42	but	but	CCONJ
gs-3955	160	43	rather	rather	ADV
gs-3955	160	44	a	a	DET
gs-3955	160	45	malleable	malleable	ADJ
gs-3955	160	46	process	process	NOUN
gs-3955	160	47	subject	subject	ADJ
gs-3955	160	48	to	to	ADP
gs-3955	160	49	evolutionary	evolutionary	ADJ
gs-3955	160	50	optimization	optimization	NOUN
gs-3955	160	51	.	.	PUNCT
gs-3955	161	1	their	their	PRON
gs-3955	161	2	continued	continued	ADJ
gs-3955	161	3	vitality	vitality	NOUN
gs-3955	161	4	across	across	ADP
gs-3955	161	5	centuries	century	NOUN
gs-3955	161	6	,	,	PUNCT
gs-3955	161	7	and	and	CCONJ
gs-3955	161	8	in	in	ADP
gs-3955	161	9	some	some	DET
gs-3955	161	10	cases	case	NOUN
gs-3955	161	11	apparent	apparent	ADJ
gs-3955	161	12	biological	biological	ADJ
gs-3955	161	13	immortality	immortality	NOUN
gs-3955	161	14	,	,	PUNCT
gs-3955	161	15	provides	provide	VERB
gs-3955	161	16	unprecedented	unprecedented	ADJ
gs-3955	161	17	insights	insight	NOUN
gs-3955	161	18	into	into	ADP
gs-3955	161	19	the	the	DET
gs-3955	161	20	molecular	molecular	ADJ
gs-3955	161	21	architecture	architecture	NOUN
gs-3955	161	22	of	of	ADP
gs-3955	161	23	longevity	longevity	NOUN
gs-3955	161	24	and	and	CCONJ
gs-3955	161	25	offers	offer	VERB
gs-3955	161	26	transformative	transformative	ADJ
gs-3955	161	27	potential	potential	NOUN
gs-3955	161	28	for	for	ADP
gs-3955	161	29	reimagining	reimagine	VERB
gs-3955	161	30	human	human	ADJ
gs-3955	161	31	healthspan	healthspan	NOUN
gs-3955	161	32	.	.	PUNCT
gs-3955	162	1	this	this	DET
gs-3955	162	2	expanded	expand	VERB
gs-3955	162	3	discussion	discussion	NOUN
gs-3955	162	4	delves	delf	NOUN
gs-3955	162	5	into	into	ADP
gs-3955	162	6	the	the	DET
gs-3955	162	7	intricate	intricate	ADJ
gs-3955	162	8	biological	biological	ADJ
gs-3955	162	9	innovations	innovation	NOUN
gs-3955	162	10	,	,	PUNCT
gs-3955	162	11	evolutionary	evolutionary	ADJ
gs-3955	162	12	contexts	contexts	NOUN
gs-3955	162	13	,	,	PUNCT
gs-3955	162	14	systemic	systemic	ADJ
gs-3955	162	15	adaptations	adaptation	NOUN
gs-3955	162	16	,	,	PUNCT
gs-3955	162	17	and	and	CCONJ
gs-3955	162	18	philosophical	philosophical	ADJ
gs-3955	162	19	implications	implication	NOUN
gs-3955	162	20	of	of	ADP
gs-3955	162	21	these	these	DET
gs-3955	162	22	extraordinary	extraordinary	ADJ
gs-3955	162	23	organisms	organism	NOUN
gs-3955	162	24	,	,	PUNCT
gs-3955	162	25	incorporating	incorporate	VERB
gs-3955	162	26	recent	recent	ADJ
gs-3955	162	27	scientific	scientific	ADJ
gs-3955	162	28	advances	advance	NOUN
gs-3955	162	29	and	and	CCONJ
gs-3955	162	30	novel	novel	ADJ
gs-3955	162	31	theoretical	theoretical	ADJ
gs-3955	162	32	frameworks	framework	NOUN
gs-3955	162	33	that	that	PRON
gs-3955	162	34	collectively	collectively	ADV
gs-3955	162	35	reshape	reshape	VERB
gs-3955	162	36	our	our	PRON
gs-3955	162	37	understanding	understanding	NOUN
gs-3955	162	38	of	of	ADP
gs-3955	162	39	biological	biological	ADJ
gs-3955	162	40	time	time	NOUN
gs-3955	162	41	.	.	PUNCT
gs-3955	163	1	other	other	ADJ
gs-3955	163	2	aspects	aspect	VERB
gs-3955	163	3	the	the	DET
gs-3955	163	4	evolutionary	evolutionary	ADJ
gs-3955	163	5	ecology	ecology	NOUN
gs-3955	163	6	of	of	ADP
gs-3955	163	7	negligible	negligible	ADJ
gs-3955	163	8	senescence	senescence	NOUN
gs-3955	163	9	reveals	reveal	VERB
gs-3955	163	10	a	a	DET
gs-3955	163	11	fascinating	fascinating	ADJ
gs-3955	163	12	pattern	pattern	NOUN
gs-3955	163	13	:	:	PUNCT
gs-3955	163	14	these	these	DET
gs-3955	163	15	species	specie	NOUN
gs-3955	163	16	predominantly	predominantly	ADV
gs-3955	163	17	inhabit	inhabit	NOUN
gs-3955	163	18	stable	stable	ADJ
gs-3955	163	19	,	,	PUNCT
gs-3955	163	20	protected	protect	VERB
gs-3955	163	21	,	,	PUNCT
gs-3955	163	22	or	or	CCONJ
gs-3955	163	23	extreme	extreme	ADJ
gs-3955	163	24	environments	environment	NOUN
gs-3955	163	25	where	where	SCONJ
gs-3955	163	26	selective	selective	ADJ
gs-3955	163	27	georgian	georgian	ADJ
gs-3955	163	28	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	163	29	მეცნიერები	მეცნიერები	NOUN
gs-3955	163	30	ტ.	ტ.	VERB
gs-3955	163	31	7	7	NUM
gs-3955	163	32	n	n	PRON
gs-3955	163	33	3	3	NUM
gs-3955	163	34	,	,	PUNCT
gs-3955	163	35	2025	2025	NUM
gs-3955	163	36	366	366	NUM
gs-3955	163	37	pressures	pressure	NOUN
gs-3955	163	38	favor	favor	VERB
gs-3955	163	39	extended	extended	ADJ
gs-3955	163	40	somatic	somatic	ADJ
gs-3955	163	41	maintenance	maintenance	NOUN
gs-3955	163	42	over	over	ADP
gs-3955	163	43	rapid	rapid	ADJ
gs-3955	163	44	reproduction	reproduction	NOUN
gs-3955	163	45	.	.	PUNCT
gs-3955	164	1	the	the	DET
gs-3955	164	2	greenland	greenland	PROPN
gs-3955	164	3	shark	shark	NOUN
gs-3955	164	4	's	's	PART
gs-3955	164	5	abyssal	abyssal	ADJ
gs-3955	164	6	arctic	arctic	ADJ
gs-3955	164	7	habitat	habitat	NOUN
gs-3955	164	8	,	,	PUNCT
gs-3955	164	9	with	with	ADP
gs-3955	164	10	near	near	ADV
gs-3955	164	11	-	-	PUNCT
gs-3955	164	12	constant	constant	ADJ
gs-3955	164	13	temperatures	temperature	NOUN
gs-3955	164	14	around	around	ADP
gs-3955	164	15	-1	-1	NOUN
gs-3955	164	16	°	°	ADP
gs-3955	164	17	c	c	NOUN
gs-3955	164	18	to	to	ADP
gs-3955	164	19	+4	+4	PROPN
gs-3955	164	20	°	°	ADP
gs-3955	164	21	c	c	NOUN
gs-3955	164	22	and	and	CCONJ
gs-3955	164	23	minimal	minimal	ADJ
gs-3955	164	24	predation	predation	NOUN
gs-3955	164	25	pressure	pressure	NOUN
gs-3955	164	26	on	on	ADP
gs-3955	164	27	adults	adult	NOUN
gs-3955	164	28	,	,	PUNCT
gs-3955	164	29	creates	create	VERB
gs-3955	164	30	conditions	condition	NOUN
gs-3955	164	31	conducive	conducive	ADJ
gs-3955	164	32	to	to	ADP
gs-3955	164	33	extraordinarily	extraordinarily	ADV
gs-3955	164	34	slow	slow	ADJ
gs-3955	164	35	life	life	NOUN
gs-3955	164	36	histories	history	NOUN
gs-3955	164	37	.	.	PUNCT
gs-3955	165	1	similarly	similarly	ADV
gs-3955	165	2	,	,	PUNCT
gs-3955	165	3	naked	naked	ADJ
gs-3955	165	4	mole	mole	NOUN
gs-3955	165	5	-	-	PUNCT
gs-3955	165	6	rats	rat	NOUN
gs-3955	165	7	thrive	thrive	VERB
gs-3955	165	8	in	in	ADP
gs-3955	165	9	subterranean	subterranean	ADJ
gs-3955	165	10	eusocial	eusocial	ADJ
gs-3955	165	11	colonies	colony	NOUN
gs-3955	165	12	where	where	SCONJ
gs-3955	165	13	cooperative	cooperative	ADJ
gs-3955	165	14	breeding	breeding	NOUN
gs-3955	165	15	reduces	reduce	VERB
gs-3955	165	16	individual	individual	ADJ
gs-3955	165	17	reproductive	reproductive	ADJ
gs-3955	165	18	burden	burden	NOUN
gs-3955	165	19	and	and	CCONJ
gs-3955	165	20	the	the	DET
gs-3955	165	21	sealed	seal	VERB
gs-3955	165	22	environment	environment	NOUN
gs-3955	165	23	buffers	buffer	NOUN
gs-3955	165	24	against	against	ADP
gs-3955	165	25	external	external	ADJ
gs-3955	165	26	threats	threat	NOUN
gs-3955	165	27	.	.	PUNCT
gs-3955	166	1	this	this	DET
gs-3955	166	2	environmental	environmental	ADJ
gs-3955	166	3	stability	stability	NOUN
gs-3955	166	4	enables	enable	VERB
gs-3955	166	5	what	what	PRON
gs-3955	166	6	evolutionary	evolutionary	ADJ
gs-3955	166	7	biologists	biologist	NOUN
gs-3955	166	8	term	term	VERB
gs-3955	166	9	the	the	DET
gs-3955	166	10	"	"	PUNCT
gs-3955	166	11	selection	selection	NOUN
gs-3955	166	12	shadow	shadow	NOUN
gs-3955	166	13	"	"	PUNCT
gs-3955	166	14	hypothesis	hypothesis	NOUN
gs-3955	166	15	—	—	PUNCT
gs-3955	166	16	when	when	SCONJ
gs-3955	166	17	extrinsic	extrinsic	ADJ
gs-3955	166	18	mortality	mortality	NOUN
gs-3955	166	19	is	be	AUX
gs-3955	166	20	low	low	ADJ
gs-3955	166	21	,	,	PUNCT
gs-3955	166	22	natural	natural	ADJ
gs-3955	166	23	selection	selection	NOUN
gs-3955	166	24	can	can	AUX
gs-3955	166	25	act	act	VERB
gs-3955	166	26	more	more	ADV
gs-3955	166	27	efficiently	efficiently	ADV
gs-3955	166	28	on	on	ADP
gs-3955	166	29	genes	gene	NOUN
gs-3955	166	30	promoting	promote	VERB
gs-3955	166	31	longevity	longevity	NOUN
gs-3955	166	32	.	.	PUNCT
gs-3955	167	1	the	the	DET
gs-3955	167	2	antagonistic	antagonistic	ADJ
gs-3955	167	3	pleiotropy	pleiotropy	NOUN
gs-3955	167	4	theory	theory	NOUN
gs-3955	167	5	,	,	PUNCT
gs-3955	167	6	which	which	PRON
gs-3955	167	7	posits	posit	VERB
gs-3955	167	8	that	that	SCONJ
gs-3955	167	9	genes	gene	NOUN
gs-3955	167	10	beneficial	beneficial	ADJ
gs-3955	167	11	early	early	ADV
gs-3955	167	12	in	in	ADP
gs-3955	167	13	life	life	NOUN
gs-3955	167	14	may	may	AUX
gs-3955	167	15	become	become	VERB
gs-3955	167	16	detrimental	detrimental	ADJ
gs-3955	167	17	later	later	ADV
gs-3955	167	18	,	,	PUNCT
gs-3955	167	19	finds	find	VERB
gs-3955	167	20	remarkable	remarkable	ADJ
gs-3955	167	21	counterexamples	counterexample	NOUN
gs-3955	167	22	in	in	ADP
gs-3955	167	23	these	these	DET
gs-3955	167	24	species	specie	NOUN
gs-3955	167	25	.	.	PUNCT
gs-3955	168	1	naked	naked	ADJ
gs-3955	168	2	mole	mole	NOUN
gs-3955	168	3	-	-	PUNCT
gs-3955	168	4	rats	rat	NOUN
gs-3955	168	5	maintain	maintain	VERB
gs-3955	168	6	tumorsuppressor	tumorsuppressor	NOUN
gs-3955	168	7	mechanisms	mechanism	NOUN
gs-3955	168	8	like	like	ADP
gs-3955	168	9	high	high	ADJ
gs-3955	168	10	-	-	PUNCT
gs-3955	168	11	molecular	molecular	ADJ
gs-3955	168	12	-	-	PUNCT
gs-3955	168	13	weight	weight	NOUN
gs-3955	168	14	hyaluronan	hyaluronan	NOUN
gs-3955	168	15	throughout	throughout	ADP
gs-3955	168	16	their	their	PRON
gs-3955	168	17	lifespans	lifespan	NOUN
gs-3955	168	18	without	without	ADP
gs-3955	168	19	apparent	apparent	ADJ
gs-3955	168	20	trade	trade	NOUN
gs-3955	168	21	-	-	PUNCT
gs-3955	168	22	offs	off	NOUN
gs-3955	168	23	in	in	ADP
gs-3955	168	24	growth	growth	NOUN
gs-3955	168	25	or	or	CCONJ
gs-3955	168	26	fertility	fertility	NOUN
gs-3955	168	27	.	.	PUNCT
gs-3955	169	1	the	the	DET
gs-3955	169	2	centrosomal	centrosomal	ADJ
gs-3955	169	3	theory	theory	NOUN
gs-3955	169	4	of	of	ADP
gs-3955	169	5	aging	aging	NOUN
gs-3955	169	6	provides	provide	VERB
gs-3955	169	7	a	a	DET
gs-3955	169	8	compelling	compelling	ADJ
gs-3955	169	9	unifying	unify	VERB
gs-3955	169	10	framework	framework	NOUN
gs-3955	169	11	,	,	PUNCT
gs-3955	169	12	suggesting	suggest	VERB
gs-3955	169	13	that	that	SCONJ
gs-3955	169	14	aging	aging	NOUN
gs-3955	169	15	arises	arise	VERB
gs-3955	169	16	from	from	ADP
gs-3955	169	17	asymmetric	asymmetric	ADJ
gs-3955	169	18	distribution	distribution	NOUN
gs-3955	169	19	of	of	ADP
gs-3955	169	20	damaged	damage	VERB
gs-3955	169	21	cellular	cellular	ADJ
gs-3955	169	22	components	component	NOUN
gs-3955	169	23	during	during	ADP
gs-3955	169	24	division	division	NOUN
gs-3955	169	25	.	.	PUNCT
gs-3955	170	1	species	specie	NOUN
gs-3955	170	2	like	like	ADP
gs-3955	170	3	planarians	planarian	NOUN
gs-3955	170	4	avoid	avoid	VERB
gs-3955	170	5	this	this	PRON
gs-3955	170	6	through	through	ADP
gs-3955	170	7	perfectly	perfectly	ADV
gs-3955	170	8	symmetrical	symmetrical	ADJ
gs-3955	170	9	stem	stem	NOUN
gs-3955	170	10	cell	cell	NOUN
gs-3955	170	11	divisions	division	NOUN
gs-3955	170	12	that	that	PRON
gs-3955	170	13	distribute	distribute	VERB
gs-3955	170	14	pristine	pristine	ADJ
gs-3955	170	15	cellular	cellular	ADJ
gs-3955	170	16	machinery	machinery	NOUN
gs-3955	170	17	equally	equally	ADV
gs-3955	170	18	between	between	ADP
gs-3955	170	19	daughter	daughter	NOUN
gs-3955	170	20	cells	cell	NOUN
gs-3955	170	21	,	,	PUNCT
gs-3955	170	22	effectively	effectively	ADV
gs-3955	170	23	resetting	reset	VERB
gs-3955	170	24	the	the	DET
gs-3955	170	25	aging	age	VERB
gs-3955	170	26	clock	clock	NOUN
gs-3955	170	27	with	with	ADP
gs-3955	170	28	each	each	DET
gs-3955	170	29	generation	generation	NOUN
gs-3955	170	30	.	.	PUNCT
gs-3955	171	1	this	this	DET
gs-3955	171	2	thermodynamic	thermodynamic	ADJ
gs-3955	171	3	perspective	perspective	NOUN
gs-3955	171	4	reframes	reframe	VERB
gs-3955	171	5	aging	aging	NOUN
gs-3955	171	6	not	not	PART
gs-3955	171	7	as	as	ADP
gs-3955	171	8	programmed	program	VERB
gs-3955	171	9	decline	decline	NOUN
gs-3955	171	10	but	but	CCONJ
gs-3955	171	11	as	as	ADP
gs-3955	171	12	an	an	DET
gs-3955	171	13	entropic	entropic	ADJ
gs-3955	171	14	process	process	NOUN
gs-3955	171	15	that	that	PRON
gs-3955	171	16	can	can	AUX
gs-3955	171	17	be	be	AUX
gs-3955	171	18	systematically	systematically	ADV
gs-3955	171	19	counteracted	counteract	VERB
gs-3955	171	20	through	through	ADP
gs-3955	171	21	biological	biological	ADJ
gs-3955	171	22	innovation	innovation	NOUN
gs-3955	171	23	(	(	PUNCT
gs-3955	171	24	macrae	macrae	PROPN
gs-3955	171	25	et	et	PROPN
gs-3955	171	26	al	al	PROPN
gs-3955	171	27	.	.	PROPN
gs-3955	171	28	,	,	PUNCT
gs-3955	171	29	2015	2015	NUM
gs-3955	171	30	,	,	PUNCT
gs-3955	171	31	munro	munro	PROPN
gs-3955	171	32	et	et	PROPN
gs-3955	171	33	al	al	PROPN
gs-3955	171	34	.	.	PROPN
gs-3955	171	35	,	,	PUNCT
gs-3955	171	36	2012	2012	NUM
gs-3955	171	37	)	)	PUNCT
gs-3955	171	38	.	.	PUNCT
gs-3955	172	1	at	at	ADP
gs-3955	172	2	the	the	DET
gs-3955	172	3	molecular	molecular	ADJ
gs-3955	172	4	level	level	NOUN
gs-3955	172	5	,	,	PUNCT
gs-3955	172	6	negligibly	negligibly	ADV
gs-3955	172	7	senescent	senescent	ADJ
gs-3955	172	8	species	specie	NOUN
gs-3955	172	9	exhibit	exhibit	VERB
gs-3955	172	10	an	an	DET
gs-3955	172	11	orchestra	orchestra	NOUN
gs-3955	172	12	of	of	ADP
gs-3955	172	13	specialized	specialized	ADJ
gs-3955	172	14	mechanisms	mechanism	NOUN
gs-3955	172	15	that	that	PRON
gs-3955	172	16	collectively	collectively	ADV
gs-3955	172	17	defend	defend	VERB
gs-3955	172	18	against	against	ADP
gs-3955	172	19	the	the	DET
gs-3955	172	20	primary	primary	ADJ
gs-3955	172	21	hallmarks	hallmark	NOUN
gs-3955	172	22	of	of	ADP
gs-3955	172	23	aging	aging	NOUN
gs-3955	172	24	.	.	PUNCT
gs-3955	173	1	genomic	genomic	ADJ
gs-3955	173	2	stability	stability	NOUN
gs-3955	173	3	is	be	AUX
gs-3955	173	4	maintained	maintain	VERB
gs-3955	173	5	through	through	ADP
gs-3955	173	6	enhanced	enhance	VERB
gs-3955	173	7	dna	dna	PROPN
gs-3955	173	8	repair	repair	NOUN
gs-3955	173	9	systems	system	NOUN
gs-3955	173	10	that	that	PRON
gs-3955	173	11	far	far	ADV
gs-3955	173	12	surpass	surpass	VERB
gs-3955	173	13	those	those	PRON
gs-3955	173	14	of	of	ADP
gs-3955	173	15	conventional	conventional	ADJ
gs-3955	173	16	model	model	NOUN
gs-3955	173	17	organisms	organism	NOUN
gs-3955	173	18	.	.	PUNCT
gs-3955	174	1	the	the	DET
gs-3955	174	2	ocean	ocean	PROPN
gs-3955	174	3	quahog	quahog	PROPN
gs-3955	174	4	arctica	arctica	PROPN
gs-3955	174	5	islandica	islandica	PROPN
gs-3955	174	6	,	,	PUNCT
gs-3955	174	7	which	which	PRON
gs-3955	174	8	lives	live	VERB
gs-3955	174	9	over	over	ADP
gs-3955	174	10	500	500	NUM
gs-3955	174	11	years	year	NOUN
gs-3955	174	12	,	,	PUNCT
gs-3955	174	13	demonstrates	demonstrate	VERB
gs-3955	174	14	nucleotide	nucleotide	ADJ
gs-3955	174	15	excision	excision	NOUN
gs-3955	174	16	repair	repair	NOUN
gs-3955	174	17	capabilities	capability	NOUN
gs-3955	174	18	that	that	PRON
gs-3955	174	19	process	process	VERB
gs-3955	174	20	ultraviolet	ultraviolet	NOUN
gs-3955	174	21	-	-	PUNCT
gs-3955	174	22	induced	induce	VERB
gs-3955	174	23	dna	dna	NOUN
gs-3955	174	24	lesions	lesion	NOUN
gs-3955	174	25	at	at	ADP
gs-3955	174	26	rates	rate	NOUN
gs-3955	174	27	quadruple	quadruple	VERB
gs-3955	174	28	those	those	PRON
gs-3955	174	29	observed	observe	VERB
gs-3955	174	30	in	in	ADP
gs-3955	174	31	short	short	ADV
gs-3955	174	32	-	-	PUNCT
gs-3955	174	33	lived	live	VERB
gs-3955	174	34	clam	clam	PROPN
gs-3955	174	35	species	specie	NOUN
gs-3955	174	36	,	,	PUNCT
gs-3955	174	37	mediated	mediate	VERB
gs-3955	174	38	by	by	ADP
gs-3955	174	39	xpa	xpa	PROPN
gs-3955	174	40	-	-	PUNCT
gs-3955	174	41	xpg	xpg	PROPN
gs-3955	174	42	complexes	complex	NOUN
gs-3955	174	43	and	and	CCONJ
gs-3955	174	44	ercc1	ercc1	NOUN
gs-3955	174	45	stabilization	stabilization	NOUN
gs-3955	174	46	(	(	PUNCT
gs-3955	174	47	munro	munro	PROPN
gs-3955	174	48	et	et	PROPN
gs-3955	174	49	al	al	PROPN
gs-3955	174	50	.	.	PROPN
gs-3955	174	51	,	,	PUNCT
gs-3955	174	52	2012	2012	NUM
gs-3955	174	53	)	)	PUNCT
gs-3955	174	54	.	.	PUNCT
gs-3955	175	1	this	this	DET
gs-3955	175	2	exceptional	exceptional	ADJ
gs-3955	175	3	repair	repair	NOUN
gs-3955	175	4	fidelity	fidelity	NOUN
gs-3955	175	5	is	be	AUX
gs-3955	175	6	attributed	attribute	VERB
gs-3955	175	7	to	to	ADP
gs-3955	175	8	unique	unique	ADJ
gs-3955	175	9	variants	variant	NOUN
gs-3955	175	10	of	of	ADP
gs-3955	175	11	the	the	DET
gs-3955	175	12	xpa	xpa	PROPN
gs-3955	175	13	and	and	CCONJ
gs-3955	175	14	ercc1	ercc1	NOUN
gs-3955	175	15	proteins	protein	NOUN
gs-3955	175	16	that	that	PRON
gs-3955	175	17	form	form	VERB
gs-3955	175	18	more	more	ADV
gs-3955	175	19	stable	stable	ADJ
gs-3955	175	20	pre	pre	ADJ
gs-3955	175	21	-	-	ADJ
gs-3955	175	22	incision	incision	NOUN
gs-3955	175	23	complexes	complex	NOUN
gs-3955	175	24	with	with	ADP
gs-3955	175	25	damaged	damaged	ADJ
gs-3955	175	26	dna	dna	NOUN
gs-3955	175	27	(	(	PUNCT
gs-3955	175	28	macrae	macrae	PROPN
gs-3955	175	29	et	et	PROPN
gs-3955	175	30	al	al	PROPN
gs-3955	175	31	.	.	PROPN
gs-3955	175	32	,	,	PUNCT
gs-3955	175	33	2015	2015	NUM
gs-3955	175	34	)	)	PUNCT
gs-3955	175	35	.	.	PUNCT
gs-3955	176	1	similarly	similarly	ADV
gs-3955	176	2	,	,	PUNCT
gs-3955	176	3	the	the	DET
gs-3955	176	4	greenland	greenland	PROPN
gs-3955	176	5	shark	shark	NOUN
gs-3955	176	6	genome	genome	NOUN
gs-3955	176	7	contains	contain	VERB
gs-3955	176	8	duplicated	duplicate	VERB
gs-3955	176	9	regions	region	NOUN
gs-3955	176	10	of	of	ADP
gs-3955	176	11	key	key	ADJ
gs-3955	176	12	dna	dna	PROPN
gs-3955	176	13	repair	repair	NOUN
gs-3955	176	14	genes	gene	NOUN
gs-3955	176	15	like	like	ADP
gs-3955	176	16	atm	atm	NOUN
gs-3955	176	17	and	and	CCONJ
gs-3955	176	18	atr	atr	PROPN
gs-3955	176	19	,	,	PUNCT
gs-3955	176	20	creating	create	VERB
gs-3955	176	21	redundancy	redundancy	NOUN
gs-3955	176	22	that	that	PRON
gs-3955	176	23	ensures	ensure	VERB
gs-3955	176	24	continuous	continuous	ADJ
gs-3955	176	25	genomic	genomic	NOUN
gs-3955	176	26	surveillance	surveillance	NOUN
gs-3955	176	27	even	even	ADV
gs-3955	176	28	under	under	ADP
gs-3955	176	29	the	the	DET
gs-3955	176	30	oxidative	oxidative	ADJ
gs-3955	176	31	stress	stress	NOUN
gs-3955	176	32	of	of	ADP
gs-3955	176	33	deep	deep	ADJ
gs-3955	176	34	-	-	PUNCT
gs-3955	176	35	sea	sea	NOUN
gs-3955	176	36	living	live	VERB
gs-3955	176	37	(	(	PUNCT
gs-3955	176	38	macrae	macrae	PROPN
gs-3955	176	39	et	et	PROPN
gs-3955	176	40	al	al	PROPN
gs-3955	176	41	.	.	PROPN
gs-3955	176	42	,	,	PUNCT
gs-3955	176	43	2015	2015	NUM
gs-3955	176	44	;	;	PUNCT
gs-3955	176	45	narayan	narayan	PROPN
gs-3955	176	46	et	et	PROPN
gs-3955	176	47	al	al	PROPN
gs-3955	176	48	.	.	PROPN
gs-3955	176	49	,	,	PUNCT
gs-3955	176	50	2021	2021	NUM
gs-3955	176	51	)	)	PUNCT
gs-3955	176	52	.	.	PUNCT
gs-3955	177	1	the	the	DET
gs-3955	177	2	naked	naked	ADJ
gs-3955	177	3	mole	mole	NOUN
gs-3955	177	4	-	-	PUNCT
gs-3955	177	5	rat	rat	NOUN
gs-3955	177	6	’s	’s	PART
gs-3955	177	7	resistance	resistance	NOUN
gs-3955	177	8	to	to	ADP
gs-3955	177	9	cancer	cancer	NOUN
gs-3955	177	10	—	—	PUNCT
gs-3955	177	11	remarkable	remarkable	ADJ
gs-3955	177	12	for	for	ADP
gs-3955	177	13	a	a	DET
gs-3955	177	14	mammal	mammal	ADJ
gs-3955	177	15	—	—	PUNCT
gs-3955	177	16	stems	stem	VERB
gs-3955	177	17	not	not	PART
gs-3955	177	18	from	from	ADP
gs-3955	177	19	a	a	DET
gs-3955	177	20	single	single	ADJ
gs-3955	177	21	mechanism	mechanism	NOUN
gs-3955	177	22	but	but	CCONJ
gs-3955	177	23	from	from	ADP
gs-3955	177	24	a	a	DET
gs-3955	177	25	multi	multi	ADJ
gs-3955	177	26	-	-	ADJ
gs-3955	177	27	layered	layered	ADJ
gs-3955	177	28	defense	defense	NOUN
gs-3955	177	29	system	system	NOUN
gs-3955	177	30	including	include	VERB
gs-3955	177	31	early	early	ADJ
gs-3955	177	32	contact	contact	NOUN
gs-3955	177	33	inhibition	inhibition	NOUN
gs-3955	177	34	mediated	mediate	VERB
gs-3955	177	35	by	by	ADP
gs-3955	177	36	high	high	ADJ
gs-3955	177	37	-	-	PUNCT
gs-3955	177	38	molecular	molecular	ADJ
gs-3955	177	39	-	-	PUNCT
gs-3955	177	40	weight	weight	NOUN
gs-3955	177	41	hyaluronan	hyaluronan	NOUN
gs-3955	177	42	(	(	PUNCT
gs-3955	177	43	hmw	hmw	PROPN
gs-3955	177	44	-	-	PUNCT
gs-3955	177	45	ha	ha	INTJ
gs-3955	177	46	)	)	PUNCT
gs-3955	177	47	(	(	PUNCT
gs-3955	177	48	tian	tian	PROPN
gs-3955	177	49	et	et	PROPN
gs-3955	177	50	al	al	PROPN
gs-3955	177	51	.	.	PROPN
gs-3955	177	52	,	,	PUNCT
gs-3955	177	53	2013	2013	NUM
gs-3955	177	54	)	)	PUNCT
gs-3955	177	55	,	,	PUNCT
gs-3955	177	56	hyper	hyper	ADJ
gs-3955	177	57	-	-	ADJ
gs-3955	177	58	vigilant	vigilant	ADJ
gs-3955	177	59	p53	p53	NOUN
gs-3955	177	60	-	-	PUNCT
gs-3955	177	61	mediated	mediate	VERB
gs-3955	177	62	apoptosis	apoptosis	NOUN
gs-3955	177	63	,	,	PUNCT
gs-3955	177	64	and	and	CCONJ
gs-3955	177	65	a	a	DET
gs-3955	177	66	unique	unique	ADJ
gs-3955	177	67	necroptosis	necroptosis	NOUN
gs-3955	177	68	pathway	pathway	NOUN
gs-3955	177	69	that	that	PRON
gs-3955	177	70	eliminates	eliminate	VERB
gs-3955	177	71	potentially	potentially	ADV
gs-3955	177	72	malignant	malignant	ADJ
gs-3955	177	73	cells	cell	NOUN
gs-3955	177	74	through	through	ADP
gs-3955	177	75	programmed	program	VERB
gs-3955	177	76	necrosis	necrosis	NOUN
gs-3955	177	77	(	(	PUNCT
gs-3955	177	78	petruseva	petruseva	PROPN
gs-3955	177	79	et	et	PROPN
gs-3955	177	80	al	al	PROPN
gs-3955	177	81	.	.	PROPN
gs-3955	177	82	,	,	PUNCT
gs-3955	177	83	2017	2017	NUM
gs-3955	177	84	;	;	PUNCT
gs-3955	177	85	lewis	lewis	PROPN
gs-3955	177	86	et	et	PROPN
gs-3955	177	87	al	al	PROPN
gs-3955	177	88	.	.	PROPN
gs-3955	177	89	,	,	PUNCT
gs-3955	177	90	2012	2012	NUM
gs-3955	177	91	)	)	PUNCT
gs-3955	177	92	.	.	PUNCT
gs-3955	178	1	recent	recent	ADJ
gs-3955	178	2	cryo	cryo	NOUN
gs-3955	178	3	-	-	PUNCT
gs-3955	178	4	electron	electron	NOUN
gs-3955	178	5	microscopy	microscopy	NOUN
gs-3955	178	6	studies	study	NOUN
gs-3955	178	7	reveal	reveal	VERB
gs-3955	178	8	that	that	SCONJ
gs-3955	178	9	naked	naked	ADJ
gs-3955	178	10	mole	mole	ADJ
gs-3955	178	11	-	-	PUNCT
gs-3955	178	12	rat	rat	NOUN
gs-3955	178	13	ribosomes	ribosome	NOUN
gs-3955	178	14	have	have	AUX
gs-3955	178	15	structurally	structurally	ADV
gs-3955	178	16	distinct	distinct	ADJ
gs-3955	178	17	decoding	decode	VERB
gs-3955	178	18	centers	center	NOUN
gs-3955	178	19	that	that	PRON
gs-3955	178	20	reduce	reduce	VERB
gs-3955	178	21	translational	translational	ADJ
gs-3955	178	22	errors	error	NOUN
gs-3955	178	23	by	by	ADP
gs-3955	178	24	nearly	nearly	ADV
gs-3955	178	25	40	40	NUM
gs-3955	178	26	%	%	NOUN
gs-3955	178	27	georgian	georgian	ADJ
gs-3955	178	28	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	178	29	მეცნიერები	მეცნიერები	NOUN
gs-3955	178	30	ტ.	ტ.	VERB
gs-3955	178	31	7	7	NUM
gs-3955	178	32	n	n	PRON
gs-3955	178	33	3	3	NUM
gs-3955	178	34	,	,	PUNCT
gs-3955	178	35	2025	2025	NUM
gs-3955	178	36	367	367	NUM
gs-3955	178	37	compared	compare	VERB
gs-3955	178	38	to	to	ADP
gs-3955	178	39	mouse	mouse	NOUN
gs-3955	178	40	ribosomes	ribosome	NOUN
gs-3955	178	41	(	(	PUNCT
gs-3955	178	42	narayan	narayan	PROPN
gs-3955	178	43	et	et	PROPN
gs-3955	178	44	al	al	PROPN
gs-3955	178	45	.	.	PROPN
gs-3955	178	46	,	,	PUNCT
gs-3955	178	47	2021	2021	NUM
gs-3955	178	48	)	)	PUNCT
gs-3955	178	49	,	,	PUNCT
gs-3955	178	50	dramatically	dramatically	ADV
gs-3955	178	51	decreasing	decrease	VERB
gs-3955	178	52	the	the	DET
gs-3955	178	53	production	production	NOUN
gs-3955	178	54	of	of	ADP
gs-3955	178	55	misfolded	misfolded	ADJ
gs-3955	178	56	proteins	protein	NOUN
gs-3955	178	57	that	that	PRON
gs-3955	178	58	contribute	contribute	VERB
gs-3955	178	59	to	to	ADP
gs-3955	178	60	proteotoxic	proteotoxic	ADJ
gs-3955	178	61	stress	stress	NOUN
gs-3955	178	62	in	in	ADP
gs-3955	178	63	aging	age	VERB
gs-3955	178	64	cells	cell	NOUN
gs-3955	178	65	.	.	PUNCT
gs-3955	179	1	the	the	DET
gs-3955	179	2	oxidative	oxidative	ADJ
gs-3955	179	3	stress	stress	NOUN
gs-3955	179	4	paradox	paradox	NOUN
gs-3955	179	5	presents	present	VERB
gs-3955	179	6	one	one	NUM
gs-3955	179	7	of	of	ADP
gs-3955	179	8	the	the	DET
gs-3955	179	9	most	most	ADV
gs-3955	179	10	fascinating	fascinating	ADJ
gs-3955	179	11	aspects	aspect	NOUN
gs-3955	179	12	of	of	ADP
gs-3955	179	13	longevity	longevity	NOUN
gs-3955	179	14	research	research	NOUN
gs-3955	179	15	in	in	ADP
gs-3955	179	16	these	these	DET
gs-3955	179	17	species	specie	NOUN
gs-3955	179	18	.	.	PUNCT
gs-3955	180	1	contrary	contrary	ADJ
gs-3955	180	2	to	to	ADP
gs-3955	180	3	the	the	DET
gs-3955	180	4	free	free	ADJ
gs-3955	180	5	radical	radical	ADJ
gs-3955	180	6	theory	theory	NOUN
gs-3955	180	7	of	of	ADP
gs-3955	180	8	aging	aging	NOUN
gs-3955	180	9	,	,	PUNCT
gs-3955	180	10	which	which	PRON
gs-3955	180	11	posits	posit	VERB
gs-3955	180	12	that	that	SCONJ
gs-3955	180	13	accumulated	accumulate	VERB
gs-3955	180	14	oxidative	oxidative	ADJ
gs-3955	180	15	damage	damage	NOUN
gs-3955	180	16	drives	drive	NOUN
gs-3955	180	17	senescence	senescence	NOUN
gs-3955	180	18	,	,	PUNCT
gs-3955	180	19	several	several	ADJ
gs-3955	180	20	negligibly	negligibly	ADV
gs-3955	180	21	senescent	senescent	ADJ
gs-3955	180	22	species	specie	NOUN
gs-3955	180	23	exhibit	exhibit	VERB
gs-3955	180	24	elevated	elevated	ADJ
gs-3955	180	25	reactive	reactive	ADJ
gs-3955	180	26	oxygen	oxygen	NOUN
gs-3955	180	27	species	species	NOUN
gs-3955	180	28	production	production	NOUN
gs-3955	180	29	without	without	ADP
gs-3955	180	30	corresponding	correspond	VERB
gs-3955	180	31	molecular	molecular	ADJ
gs-3955	180	32	damage	damage	NOUN
gs-3955	180	33	.	.	PUNCT
gs-3955	181	1	naked	naked	ADJ
gs-3955	181	2	mole	mole	ADJ
gs-3955	181	3	-	-	PUNCT
gs-3955	181	4	rat	rat	NOUN
gs-3955	181	5	mitochondria	mitochondria	NOUN
gs-3955	181	6	generate	generate	VERB
gs-3955	181	7	significantly	significantly	ADV
gs-3955	181	8	more	more	ADJ
gs-3955	181	9	superoxide	superoxide	NOUN
gs-3955	181	10	than	than	ADP
gs-3955	181	11	those	those	PRON
gs-3955	181	12	of	of	ADP
gs-3955	181	13	mice	mouse	NOUN
gs-3955	181	14	,	,	PUNCT
gs-3955	181	15	yet	yet	CCONJ
gs-3955	181	16	their	their	PRON
gs-3955	181	17	tissues	tissue	NOUN
gs-3955	181	18	show	show	VERB
gs-3955	181	19	minimal	minimal	ADJ
gs-3955	181	20	oxidative	oxidative	ADJ
gs-3955	181	21	damage	damage	NOUN
gs-3955	181	22	to	to	ADP
gs-3955	181	23	proteins	protein	NOUN
gs-3955	181	24	,	,	PUNCT
gs-3955	181	25	lipids	lipid	NOUN
gs-3955	181	26	,	,	PUNCT
gs-3955	181	27	and	and	CCONJ
gs-3955	181	28	nucleic	nucleic	ADJ
gs-3955	181	29	acids	acid	NOUN
gs-3955	181	30	.	.	PUNCT
gs-3955	182	1	this	this	DET
gs-3955	182	2	apparent	apparent	ADJ
gs-3955	182	3	contradiction	contradiction	NOUN
gs-3955	182	4	is	be	AUX
gs-3955	182	5	resolved	resolve	VERB
gs-3955	182	6	through	through	ADP
gs-3955	182	7	several	several	ADJ
gs-3955	182	8	innovative	innovative	ADJ
gs-3955	182	9	adaptations	adaptation	NOUN
gs-3955	182	10	:	:	PUNCT
gs-3955	182	11	specialized	specialized	ADJ
gs-3955	182	12	cardiolipin	cardiolipin	NOUN
gs-3955	182	13	molecules	molecule	NOUN
gs-3955	182	14	in	in	ADP
gs-3955	182	15	mitochondrial	mitochondrial	ADJ
gs-3955	182	16	membranes	membrane	NOUN
gs-3955	182	17	rich	rich	ADJ
gs-3955	182	18	in	in	ADP
gs-3955	182	19	monounsaturated	monounsaturated	ADJ
gs-3955	182	20	fatty	fatty	NOUN
gs-3955	182	21	acids	acid	NOUN
gs-3955	182	22	that	that	PRON
gs-3955	182	23	resist	resist	VERB
gs-3955	182	24	peroxidation	peroxidation	NOUN
gs-3955	182	25	;	;	PUNCT
gs-3955	182	26	enhanced	enhance	VERB
gs-3955	182	27	reducing	reduce	VERB
gs-3955	182	28	environments	environment	NOUN
gs-3955	182	29	maintained	maintain	VERB
gs-3955	182	30	by	by	ADP
gs-3955	182	31	elevated	elevated	ADJ
gs-3955	182	32	glutathione	glutathione	NOUN
gs-3955	182	33	and	and	CCONJ
gs-3955	182	34	thioredoxin	thioredoxin	PROPN
gs-3955	182	35	systems	system	NOUN
gs-3955	182	36	;	;	PUNCT
gs-3955	182	37	and	and	CCONJ
gs-3955	182	38	unique	unique	ADJ
gs-3955	182	39	isoforms	isoform	NOUN
gs-3955	182	40	of	of	ADP
gs-3955	182	41	antioxidant	antioxidant	ADJ
gs-3955	182	42	enzymes	enzyme	NOUN
gs-3955	182	43	like	like	ADP
gs-3955	182	44	superoxide	superoxide	NOUN
gs-3955	182	45	dismutase	dismutase	NOUN
gs-3955	182	46	that	that	PRON
gs-3955	182	47	exhibit	exhibit	VERB
gs-3955	182	48	unusual	unusual	ADJ
gs-3955	182	49	catalytic	catalytic	ADJ
gs-3955	182	50	efficiency	efficiency	NOUN
gs-3955	182	51	.	.	PUNCT
gs-3955	183	1	the	the	DET
gs-3955	183	2	olm	olm	NOUN
gs-3955	183	3	,	,	PUNCT
gs-3955	183	4	a	a	DET
gs-3955	183	5	cavedwelling	cavedwelle	VERB
gs-3955	183	6	salamander	salamander	NOUN
gs-3955	183	7	that	that	PRON
gs-3955	183	8	lives	live	VERB
gs-3955	183	9	over	over	ADP
gs-3955	183	10	a	a	DET
gs-3955	183	11	century	century	NOUN
gs-3955	183	12	in	in	ADP
gs-3955	183	13	complete	complete	ADJ
gs-3955	183	14	darkness	darkness	NOUN
gs-3955	183	15	,	,	PUNCT
gs-3955	183	16	demonstrates	demonstrate	VERB
gs-3955	183	17	constitutive	constitutive	ADJ
gs-3955	183	18	activation	activation	NOUN
gs-3955	183	19	of	of	ADP
gs-3955	183	20	the	the	DET
gs-3955	183	21	nrf2	nrf2	NOUN
gs-3955	183	22	pathway	pathway	NOUN
gs-3955	183	23	—	—	PUNCT
gs-3955	183	24	the	the	DET
gs-3955	183	25	master	master	NOUN
gs-3955	183	26	regulator	regulator	NOUN
gs-3955	183	27	of	of	ADP
gs-3955	183	28	antioxidant	antioxidant	ADJ
gs-3955	183	29	responses	response	NOUN
gs-3955	183	30	—	—	PUNCT
gs-3955	183	31	even	even	ADV
gs-3955	183	32	in	in	ADP
gs-3955	183	33	the	the	DET
gs-3955	183	34	absence	absence	NOUN
gs-3955	183	35	of	of	ADP
gs-3955	183	36	oxidative	oxidative	ADJ
gs-3955	183	37	challenges	challenge	NOUN
gs-3955	183	38	.	.	PUNCT
gs-3955	184	1	this	this	DET
gs-3955	184	2	preemptive	preemptive	ADJ
gs-3955	184	3	defense	defense	NOUN
gs-3955	184	4	strategy	strategy	NOUN
gs-3955	184	5	ensures	ensure	VERB
gs-3955	184	6	immediate	immediate	ADJ
gs-3955	184	7	neutralization	neutralization	NOUN
gs-3955	184	8	of	of	ADP
gs-3955	184	9	free	free	ADJ
gs-3955	184	10	radicals	radical	NOUN
gs-3955	184	11	before	before	SCONJ
gs-3955	184	12	they	they	PRON
gs-3955	184	13	can	can	AUX
gs-3955	184	14	inflict	inflict	VERB
gs-3955	184	15	macromolecular	macromolecular	ADJ
gs-3955	184	16	damage	damage	NOUN
gs-3955	184	17	.	.	PUNCT
gs-3955	185	1	perhaps	perhaps	ADV
gs-3955	185	2	most	most	ADV
gs-3955	185	3	remarkably	remarkably	ADV
gs-3955	185	4	,	,	PUNCT
gs-3955	185	5	the	the	DET
gs-3955	185	6	ocean	ocean	NOUN
gs-3955	185	7	quahog	quahog	PROPN
gs-3955	185	8	maintains	maintain	VERB
gs-3955	185	9	near	near	ADV
gs-3955	185	10	-	-	PUNCT
gs-3955	185	11	perfect	perfect	ADJ
gs-3955	185	12	redox	redox	NOUN
gs-3955	185	13	homeostasis	homeostasis	NOUN
gs-3955	185	14	for	for	ADP
gs-3955	185	15	centuries	century	NOUN
gs-3955	185	16	through	through	ADP
gs-3955	185	17	periodic	periodic	ADJ
gs-3955	185	18	reduction	reduction	NOUN
gs-3955	185	19	of	of	ADP
gs-3955	185	20	metabolic	metabolic	NOUN
gs-3955	185	21	rate	rate	NOUN
gs-3955	185	22	to	to	ADP
gs-3955	185	23	near	near	ADJ
gs-3955	185	24	-	-	PUNCT
gs-3955	185	25	undetectable	undetectable	ADJ
gs-3955	185	26	levels	level	NOUN
gs-3955	185	27	during	during	ADP
gs-3955	185	28	hibernation	hibernation	NOUN
gs-3955	185	29	-	-	PUNCT
gs-3955	185	30	like	like	ADJ
gs-3955	185	31	states	state	NOUN
gs-3955	185	32	,	,	PUNCT
gs-3955	185	33	effectively	effectively	ADV
gs-3955	185	34	minimizing	minimize	VERB
gs-3955	185	35	electron	electron	NOUN
gs-3955	185	36	leakage	leakage	NOUN
gs-3955	185	37	in	in	ADP
gs-3955	185	38	the	the	DET
gs-3955	185	39	mitochondrial	mitochondrial	ADJ
gs-3955	185	40	transport	transport	NOUN
gs-3955	185	41	chain	chain	NOUN
gs-3955	185	42	(	(	PUNCT
gs-3955	185	43	tkemaladze	tkemaladze	NOUN
gs-3955	185	44	et	et	PROPN
gs-3955	185	45	al	al	PROPN
gs-3955	185	46	.	.	PROPN
gs-3955	185	47	,	,	PUNCT
gs-3955	185	48	2023	2023	NUM
gs-3955	185	49	,	,	PUNCT
gs-3955	185	50	voituron	voituron	PROPN
gs-3955	185	51	et	et	PROPN
gs-3955	185	52	al	al	PROPN
gs-3955	185	53	.	.	PROPN
gs-3955	185	54	,	,	PUNCT
gs-3955	185	55	2011	2011	NUM
gs-3955	185	56	,	,	PUNCT
gs-3955	185	57	reddien	reddien	VERB
gs-3955	185	58	et	et	PROPN
gs-3955	185	59	al	al	PROPN
gs-3955	185	60	.	.	PROPN
gs-3955	185	61	,	,	PUNCT
gs-3955	185	62	2018	2018	NUM
gs-3955	185	63	)	)	PUNCT
gs-3955	185	64	.	.	PUNCT
gs-3955	186	1	proteostasis	proteostasis	NOUN
gs-3955	186	2	maintenance	maintenance	NOUN
gs-3955	186	3	represents	represent	VERB
gs-3955	186	4	another	another	DET
gs-3955	186	5	cornerstone	cornerstone	NOUN
gs-3955	186	6	of	of	ADP
gs-3955	186	7	negligible	negligible	ADJ
gs-3955	186	8	senescence	senescence	NOUN
gs-3955	186	9	,	,	PUNCT
gs-3955	186	10	with	with	ADP
gs-3955	186	11	long	long	ADV
gs-3955	186	12	-	-	PUNCT
gs-3955	186	13	lived	live	VERB
gs-3955	186	14	species	specie	NOUN
gs-3955	186	15	developing	develop	VERB
gs-3955	186	16	sophisticated	sophisticated	ADJ
gs-3955	186	17	systems	system	NOUN
gs-3955	186	18	for	for	ADP
gs-3955	186	19	preserving	preserve	VERB
gs-3955	186	20	protein	protein	NOUN
gs-3955	186	21	structure	structure	NOUN
gs-3955	186	22	and	and	CCONJ
gs-3955	186	23	function	function	NOUN
gs-3955	186	24	.	.	PUNCT
gs-3955	187	1	the	the	DET
gs-3955	187	2	naked	naked	ADJ
gs-3955	187	3	mole	mole	ADJ
gs-3955	187	4	-	-	PUNCT
gs-3955	187	5	rat	rat	NOUN
gs-3955	187	6	proteome	proteome	NOUN
gs-3955	187	7	exhibits	exhibit	VERB
gs-3955	187	8	extraordinary	extraordinary	ADJ
gs-3955	187	9	stability	stability	NOUN
gs-3955	187	10	,	,	PUNCT
gs-3955	187	11	with	with	ADP
gs-3955	187	12	studies	study	NOUN
gs-3955	187	13	showing	show	VERB
gs-3955	187	14	no	no	DET
gs-3955	187	15	age	age	NOUN
gs-3955	187	16	-	-	PUNCT
gs-3955	187	17	related	relate	VERB
gs-3955	187	18	increase	increase	NOUN
gs-3955	187	19	in	in	ADP
gs-3955	187	20	protein	protein	NOUN
gs-3955	187	21	carbonylation	carbonylation	NOUN
gs-3955	187	22	until	until	ADP
gs-3955	187	23	very	very	ADV
gs-3955	187	24	late	late	ADJ
gs-3955	187	25	life	life	NOUN
gs-3955	187	26	(	(	PUNCT
gs-3955	187	27	perez	perez	PROPN
gs-3955	187	28	et	et	PROPN
gs-3955	187	29	al	al	PROPN
gs-3955	187	30	.	.	PROPN
gs-3955	187	31	,	,	PUNCT
gs-3955	187	32	2009	2009	NUM
gs-3955	187	33	)	)	PUNCT
gs-3955	187	34	.	.	PUNCT
gs-3955	188	1	this	this	DET
gs-3955	188	2	resilience	resilience	NOUN
gs-3955	188	3	stems	stem	VERB
gs-3955	188	4	from	from	ADP
gs-3955	188	5	enhanced	enhance	VERB
gs-3955	188	6	chaperone	chaperone	NOUN
gs-3955	188	7	networks	network	NOUN
gs-3955	188	8	,	,	PUNCT
gs-3955	188	9	including	include	VERB
gs-3955	188	10	constitutively	constitutively	ADV
gs-3955	188	11	expressed	express	VERB
gs-3955	188	12	heat	heat	NOUN
gs-3955	188	13	shock	shock	NOUN
gs-3955	188	14	proteins	protein	NOUN
gs-3955	188	15	that	that	PRON
gs-3955	188	16	refold	refold	VERB
gs-3955	188	17	damaged	damage	VERB
gs-3955	188	18	polypeptides	polypeptide	NOUN
gs-3955	188	19	,	,	PUNCT
gs-3955	188	20	and	and	CCONJ
gs-3955	188	21	an	an	DET
gs-3955	188	22	expanded	expand	VERB
gs-3955	188	23	ubiquitin	ubiquitin	ADJ
gs-3955	188	24	-	-	PUNCT
gs-3955	188	25	proteasome	proteasome	NOUN
gs-3955	188	26	system	system	NOUN
gs-3955	188	27	featuring	feature	VERB
gs-3955	188	28	unique	unique	ADJ
gs-3955	188	29	e3	e3	NOUN
gs-3955	188	30	ligase	ligase	NOUN
gs-3955	188	31	variants	variant	NOUN
gs-3955	188	32	that	that	PRON
gs-3955	188	33	selectively	selectively	ADV
gs-3955	188	34	target	target	VERB
gs-3955	188	35	oxidized	oxidize	VERB
gs-3955	188	36	proteins	protein	NOUN
gs-3955	188	37	for	for	ADP
gs-3955	188	38	degradation	degradation	NOUN
gs-3955	188	39	(	(	PUNCT
gs-3955	188	40	rodriguez	rodriguez	NOUN
gs-3955	188	41	et	et	PROPN
gs-3955	188	42	al	al	PROPN
gs-3955	188	43	.	.	PROPN
gs-3955	188	44	,	,	PUNCT
gs-3955	188	45	2016	2016	NUM
gs-3955	188	46	)	)	PUNCT
gs-3955	188	47	.	.	PUNCT
gs-3955	189	1	autophagy	autophagy	ADV
gs-3955	189	2	—	—	PUNCT
gs-3955	189	3	the	the	DET
gs-3955	189	4	cellular	cellular	ADJ
gs-3955	189	5	recycling	recycling	NOUN
gs-3955	189	6	process	process	NOUN
gs-3955	189	7	—	—	PUNCT
gs-3955	189	8	is	be	AUX
gs-3955	189	9	amplified	amplify	VERB
gs-3955	189	10	and	and	CCONJ
gs-3955	189	11	precisely	precisely	ADV
gs-3955	189	12	regulated	regulated	ADJ
gs-3955	189	13	in	in	ADP
gs-3955	189	14	these	these	DET
gs-3955	189	15	species	specie	NOUN
gs-3955	189	16	.	.	PUNCT
gs-3955	190	1	hydra	hydra	PROPN
gs-3955	190	2	demonstrates	demonstrate	VERB
gs-3955	190	3	continuous	continuous	ADJ
gs-3955	190	4	autophagic	autophagic	ADJ
gs-3955	190	5	flux	flux	NOUN
gs-3955	190	6	that	that	PRON
gs-3955	190	7	removes	remove	VERB
gs-3955	190	8	damaged	damaged	ADJ
gs-3955	190	9	organelles	organelle	NOUN
gs-3955	190	10	without	without	ADP
gs-3955	190	11	the	the	DET
gs-3955	190	12	decline	decline	NOUN
gs-3955	190	13	seen	see	VERB
gs-3955	190	14	in	in	ADP
gs-3955	190	15	aging	age	VERB
gs-3955	190	16	mammals	mammal	NOUN
gs-3955	190	17	(	(	PUNCT
gs-3955	190	18	tomczyk	tomczyk	NOUN
gs-3955	190	19	et	et	NOUN
gs-3955	190	20	al	al	PROPN
gs-3955	190	21	.	.	PROPN
gs-3955	190	22	,	,	PUNCT
gs-3955	190	23	2015	2015	NUM
gs-3955	190	24	)	)	PUNCT
gs-3955	190	25	,	,	PUNCT
gs-3955	190	26	while	while	SCONJ
gs-3955	190	27	rockfish	rockfish	NOUN
gs-3955	190	28	maintain	maintain	VERB
gs-3955	190	29	elevated	elevated	ADJ
gs-3955	190	30	expression	expression	NOUN
gs-3955	190	31	of	of	ADP
gs-3955	190	32	autophagy	autophagy	ADJ
gs-3955	190	33	genes	gene	NOUN
gs-3955	190	34	like	like	ADP
gs-3955	190	35	lc3	lc3	NOUN
gs-3955	190	36	and	and	CCONJ
gs-3955	190	37	beclin1	beclin1	NOUN
gs-3955	190	38	throughout	throughout	ADP
gs-3955	190	39	their	their	PRON
gs-3955	190	40	multi	multi	ADJ
gs-3955	190	41	-	-	ADJ
gs-3955	190	42	century	century	ADJ
gs-3955	190	43	lifespans	lifespan	NOUN
gs-3955	190	44	(	(	PUNCT
gs-3955	190	45	finn	finn	PROPN
gs-3955	190	46	et	et	PROPN
gs-3955	190	47	al	al	PROPN
gs-3955	190	48	.	.	PROPN
gs-3955	190	49	,	,	PUNCT
gs-3955	190	50	2021	2021	NUM
gs-3955	190	51	)	)	PUNCT
gs-3955	190	52	.	.	PUNCT
gs-3955	191	1	the	the	DET
gs-3955	191	2	ocean	ocean	PROPN
gs-3955	191	3	quahog	quahog	PROPN
gs-3955	191	4	takes	take	VERB
gs-3955	191	5	proteostasis	proteostasis	NOUN
gs-3955	191	6	to	to	ADP
gs-3955	191	7	extraordinary	extraordinary	ADJ
gs-3955	191	8	levels	level	NOUN
gs-3955	191	9	through	through	ADP
gs-3955	191	10	temperature	temperature	NOUN
gs-3955	191	11	-	-	PUNCT
gs-3955	191	12	insensitive	insensitive	ADJ
gs-3955	191	13	molecular	molecular	ADJ
gs-3955	191	14	chaperones	chaperone	NOUN
gs-3955	191	15	that	that	PRON
gs-3955	191	16	maintain	maintain	VERB
gs-3955	191	17	function	function	NOUN
gs-3955	191	18	across	across	ADP
gs-3955	191	19	the	the	DET
gs-3955	191	20	thermal	thermal	ADJ
gs-3955	191	21	fluctuations	fluctuation	NOUN
gs-3955	191	22	of	of	ADP
gs-3955	191	23	its	its	PRON
gs-3955	191	24	benthic	benthic	ADJ
gs-3955	191	25	environment	environment	NOUN
gs-3955	191	26	(	(	PUNCT
gs-3955	191	27	abele	abele	PROPN
gs-3955	191	28	et	et	PROPN
gs-3955	191	29	al	al	PROPN
gs-3955	191	30	.	.	PROPN
gs-3955	191	31	,	,	PUNCT
gs-3955	191	32	2008	2008	NUM
gs-3955	191	33	)	)	PUNCT
gs-3955	191	34	.	.	PUNCT
gs-3955	192	1	these	these	DET
gs-3955	192	2	chaperones	chaperone	NOUN
gs-3955	192	3	contain	contain	VERB
gs-3955	192	4	specific	specific	ADJ
gs-3955	192	5	amino	amino	NOUN
gs-3955	192	6	acid	acid	NOUN
gs-3955	192	7	substitutions	substitution	NOUN
gs-3955	192	8	in	in	ADP
gs-3955	192	9	their	their	PRON
gs-3955	192	10	substrate	substrate	NOUN
gs-3955	192	11	-	-	PUNCT
gs-3955	192	12	binding	bind	VERB
gs-3955	192	13	domains	domain	NOUN
gs-3955	192	14	that	that	PRON
gs-3955	192	15	enhance	enhance	VERB
gs-3955	192	16	affinity	affinity	NOUN
gs-3955	192	17	for	for	ADP
gs-3955	192	18	partially	partially	ADV
gs-3955	192	19	unfolded	unfold	VERB
gs-3955	192	20	proteins	protein	NOUN
gs-3955	192	21	,	,	PUNCT
gs-3955	192	22	preventing	prevent	VERB
gs-3955	192	23	aggregation	aggregation	NOUN
gs-3955	192	24	even	even	ADV
gs-3955	192	25	under	under	ADP
gs-3955	192	26	thermal	thermal	ADJ
gs-3955	192	27	stress	stress	NOUN
gs-3955	192	28	.	.	PUNCT
gs-3955	193	1	georgian	georgian	ADJ
gs-3955	193	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	193	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	193	4	ტ.	ტ.	VERB
gs-3955	193	5	7	7	NUM
gs-3955	193	6	n	n	PRON
gs-3955	193	7	3	3	NUM
gs-3955	193	8	,	,	PUNCT
gs-3955	193	9	2025	2025	NUM
gs-3955	193	10	368	368	NUM
gs-3955	193	11	additionally	additionally	ADV
gs-3955	193	12	,	,	PUNCT
gs-3955	193	13	the	the	DET
gs-3955	193	14	quahog	quahog	NOUN
gs-3955	193	15	expresses	express	VERB
gs-3955	193	16	novel	novel	ADJ
gs-3955	193	17	classes	class	NOUN
gs-3955	193	18	of	of	ADP
gs-3955	193	19	small	small	ADJ
gs-3955	193	20	heat	heat	NOUN
gs-3955	193	21	shock	shock	NOUN
gs-3955	193	22	proteins	protein	NOUN
gs-3955	193	23	that	that	PRON
gs-3955	193	24	act	act	VERB
gs-3955	193	25	as	as	ADP
gs-3955	193	26	"	"	PUNCT
gs-3955	193	27	holdases	holdase	NOUN
gs-3955	193	28	,	,	PUNCT
gs-3955	193	29	"	"	PUNCT
gs-3955	193	30	binding	bind	VERB
gs-3955	193	31	exposed	expose	VERB
gs-3955	193	32	hydrophobic	hydrophobic	ADJ
gs-3955	193	33	regions	region	NOUN
gs-3955	193	34	on	on	ADP
gs-3955	193	35	nascent	nascent	ADJ
gs-3955	193	36	polypeptides	polypeptide	NOUN
gs-3955	193	37	to	to	PART
gs-3955	193	38	prevent	prevent	VERB
gs-3955	193	39	misfolding	misfolde	VERB
gs-3955	193	40	during	during	ADP
gs-3955	193	41	translation	translation	NOUN
gs-3955	193	42	(	(	PUNCT
gs-3955	193	43	treaster	treaster	NOUN
gs-3955	193	44	et	et	PROPN
gs-3955	193	45	al	al	PROPN
gs-3955	193	46	.	.	PROPN
gs-3955	193	47	,	,	PUNCT
gs-3955	193	48	2014	2014	NUM
gs-3955	193	49	)	)	PUNCT
gs-3955	193	50	.	.	PUNCT
gs-3955	194	1	stem	stem	NOUN
gs-3955	194	2	cell	cell	NOUN
gs-3955	194	3	dynamics	dynamic	NOUN
gs-3955	194	4	in	in	ADP
gs-3955	194	5	negligibly	negligibly	ADV
gs-3955	194	6	senescent	senescent	ADJ
gs-3955	194	7	species	specie	NOUN
gs-3955	194	8	present	present	ADJ
gs-3955	194	9	revolutionary	revolutionary	ADJ
gs-3955	194	10	models	model	NOUN
gs-3955	194	11	for	for	ADP
gs-3955	194	12	regenerative	regenerative	ADJ
gs-3955	194	13	medicine	medicine	NOUN
gs-3955	194	14	.	.	PUNCT
gs-3955	195	1	hydra	hydra	PROPN
gs-3955	195	2	's	's	PART
gs-3955	195	3	apparent	apparent	ADJ
gs-3955	195	4	biological	biological	ADJ
gs-3955	195	5	immortality	immortality	NOUN
gs-3955	195	6	stems	stem	VERB
gs-3955	195	7	directly	directly	ADV
gs-3955	195	8	from	from	ADP
gs-3955	195	9	three	three	NUM
gs-3955	195	10	interstitial	interstitial	ADJ
gs-3955	195	11	stem	stem	NOUN
gs-3955	195	12	cell	cell	NOUN
gs-3955	195	13	populations	population	NOUN
gs-3955	195	14	that	that	PRON
gs-3955	195	15	perpetually	perpetually	ADV
gs-3955	195	16	renew	renew	VERB
gs-3955	195	17	all	all	DET
gs-3955	195	18	tissues	tissue	NOUN
gs-3955	195	19	without	without	ADP
gs-3955	195	20	exhaustion	exhaustion	NOUN
gs-3955	195	21	(	(	PUNCT
gs-3955	195	22	bosch	bosch	PROPN
gs-3955	195	23	,	,	PUNCT
gs-3955	195	24	2009	2009	NUM
gs-3955	195	25	)	)	PUNCT
gs-3955	195	26	.	.	PUNCT
gs-3955	196	1	unlike	unlike	ADP
gs-3955	196	2	mammalian	mammalian	ADJ
gs-3955	196	3	stem	stem	NOUN
gs-3955	196	4	cells	cell	NOUN
gs-3955	196	5	,	,	PUNCT
gs-3955	196	6	hydra	hydra	PROPN
gs-3955	196	7	stem	stem	NOUN
gs-3955	196	8	cells	cell	NOUN
gs-3955	196	9	express	express	VERB
gs-3955	196	10	telomerase	telomerase	NOUN
gs-3955	196	11	constitutively	constitutively	ADV
gs-3955	196	12	and	and	CCONJ
gs-3955	196	13	possess	possess	VERB
gs-3955	196	14	epigenetic	epigenetic	ADJ
gs-3955	196	15	regulators	regulator	NOUN
gs-3955	196	16	that	that	PRON
gs-3955	196	17	maintain	maintain	VERB
gs-3955	196	18	a	a	DET
gs-3955	196	19	permanent	permanent	ADJ
gs-3955	196	20	pluripotent	pluripotent	ADJ
gs-3955	196	21	state	state	NOUN
gs-3955	196	22	(	(	PUNCT
gs-3955	196	23	tomczyk	tomczyk	NOUN
gs-3955	196	24	et	et	PROPN
gs-3955	196	25	al	al	PROPN
gs-3955	196	26	.	.	PROPN
gs-3955	196	27	,	,	PUNCT
gs-3955	196	28	2015	2015	NUM
gs-3955	196	29	)	)	PUNCT
gs-3955	196	30	.	.	PUNCT
gs-3955	197	1	single	single	ADJ
gs-3955	197	2	-	-	PUNCT
gs-3955	197	3	cell	cell	NOUN
gs-3955	197	4	rna	rna	NOUN
gs-3955	197	5	sequencing	sequencing	NOUN
gs-3955	197	6	has	have	AUX
gs-3955	197	7	revealed	reveal	VERB
gs-3955	197	8	a	a	DET
gs-3955	197	9	novel	novel	NOUN
gs-3955	197	10	signaling	signal	VERB
gs-3955	197	11	axis	axis	NOUN
gs-3955	197	12	between	between	ADP
gs-3955	197	13	the	the	DET
gs-3955	197	14	foxo	foxo	NOUN
gs-3955	197	15	transcription	transcription	NOUN
gs-3955	197	16	factor	factor	NOUN
gs-3955	197	17	and	and	CCONJ
gs-3955	197	18	wnt	wnt	PROPN
gs-3955	197	19	pathway	pathway	NOUN
gs-3955	197	20	components	component	NOUN
gs-3955	197	21	that	that	PRON
gs-3955	197	22	preserves	preserve	VERB
gs-3955	197	23	stemness	stemness	ADV
gs-3955	197	24	while	while	SCONJ
gs-3955	197	25	preventing	prevent	VERB
gs-3955	197	26	malignant	malignant	ADJ
gs-3955	197	27	transformation	transformation	NOUN
gs-3955	197	28	(	(	PUNCT
gs-3955	197	29	boehm	boehm	PROPN
gs-3955	197	30	et	et	PROPN
gs-3955	197	31	al	al	PROPN
gs-3955	197	32	.	.	PROPN
gs-3955	197	33	,	,	PUNCT
gs-3955	197	34	2012	2012	NUM
gs-3955	197	35	)	)	PUNCT
gs-3955	197	36	.	.	PUNCT
gs-3955	198	1	planarians	planarian	NOUN
gs-3955	198	2	take	take	VERB
gs-3955	198	3	stem	stem	NOUN
gs-3955	198	4	cell	cell	NOUN
gs-3955	198	5	biology	biology	NOUN
gs-3955	198	6	to	to	ADP
gs-3955	198	7	even	even	ADV
gs-3955	198	8	more	more	ADV
gs-3955	198	9	extraordinary	extraordinary	ADJ
gs-3955	198	10	levels	level	NOUN
gs-3955	198	11	through	through	ADP
gs-3955	198	12	their	their	PRON
gs-3955	198	13	neoblast	neoblast	ADJ
gs-3955	198	14	system	system	NOUN
gs-3955	198	15	—	—	PUNCT
gs-3955	198	16	a	a	DET
gs-3955	198	17	population	population	NOUN
gs-3955	198	18	of	of	ADP
gs-3955	198	19	pluripotent	pluripotent	ADJ
gs-3955	198	20	adult	adult	NOUN
gs-3955	198	21	stem	stem	NOUN
gs-3955	198	22	cells	cell	NOUN
gs-3955	198	23	capable	capable	ADJ
gs-3955	198	24	of	of	ADP
gs-3955	198	25	regenerating	regenerate	VERB
gs-3955	198	26	complete	complete	ADJ
gs-3955	198	27	organisms	organism	NOUN
gs-3955	198	28	from	from	ADP
gs-3955	198	29	minute	minute	ADJ
gs-3955	198	30	tissue	tissue	NOUN
gs-3955	198	31	fragments	fragment	NOUN
gs-3955	198	32	(	(	PUNCT
gs-3955	198	33	reddien	reddien	PROPN
gs-3955	198	34	,	,	PUNCT
gs-3955	198	35	2013	2013	NUM
gs-3955	198	36	)	)	PUNCT
gs-3955	198	37	.	.	PUNCT
gs-3955	199	1	these	these	DET
gs-3955	199	2	neoblasts	neoblast	NOUN
gs-3955	199	3	exhibit	exhibit	VERB
gs-3955	199	4	unique	unique	ADJ
gs-3955	199	5	chromosomal	chromosomal	ADJ
gs-3955	199	6	organization	organization	NOUN
gs-3955	199	7	with	with	ADP
gs-3955	199	8	open	open	ADJ
gs-3955	199	9	chromatin	chromatin	NOUN
gs-3955	199	10	domains	domain	NOUN
gs-3955	199	11	that	that	PRON
gs-3955	199	12	remain	remain	VERB
gs-3955	199	13	perpetually	perpetually	ADV
gs-3955	199	14	accessible	accessible	ADJ
gs-3955	199	15	to	to	ADP
gs-3955	199	16	regeneration	regeneration	NOUN
gs-3955	199	17	-	-	PUNCT
gs-3955	199	18	associated	associate	VERB
gs-3955	199	19	transcription	transcription	NOUN
gs-3955	199	20	factors	factor	NOUN
gs-3955	199	21	(	(	PUNCT
gs-3955	199	22	scimone	scimone	NOUN
gs-3955	199	23	et	et	PROPN
gs-3955	199	24	al	al	PROPN
gs-3955	199	25	.	.	PROPN
gs-3955	199	26	,	,	PUNCT
gs-3955	199	27	2014	2014	NUM
gs-3955	199	28	)	)	PUNCT
gs-3955	199	29	.	.	PUNCT
gs-3955	200	1	recent	recent	ADJ
gs-3955	200	2	research	research	NOUN
gs-3955	200	3	has	have	AUX
gs-3955	200	4	identified	identify	VERB
gs-3955	200	5	a	a	DET
gs-3955	200	6	novel	novel	ADJ
gs-3955	200	7	rna	rna	NOUN
gs-3955	200	8	-	-	PUNCT
gs-3955	200	9	binding	bind	VERB
gs-3955	200	10	protein	protein	NOUN
gs-3955	200	11	,	,	PUNCT
gs-3955	200	12	smed	sme	VERB
gs-3955	200	13	-	-	PUNCT
gs-3955	200	14	immortalin	immortalin	NOUN
gs-3955	200	15	,	,	PUNCT
gs-3955	200	16	that	that	PRON
gs-3955	200	17	prevents	prevent	VERB
gs-3955	200	18	cellular	cellular	ADJ
gs-3955	200	19	senescence	senescence	NOUN
gs-3955	200	20	by	by	ADP
gs-3955	200	21	stabilizing	stabilize	VERB
gs-3955	200	22	telomeres	telomere	NOUN
gs-3955	200	23	and	and	CCONJ
gs-3955	200	24	repressing	repress	VERB
gs-3955	200	25	p53	p53	NOUN
gs-3955	200	26	activity	activity	NOUN
gs-3955	200	27	during	during	ADP
gs-3955	200	28	regeneration	regeneration	NOUN
gs-3955	200	29	(	(	PUNCT
gs-3955	200	30	zhu	zhu	X
gs-3955	200	31	et	et	PROPN
gs-3955	200	32	al	al	PROPN
gs-3955	200	33	.	.	PROPN
gs-3955	200	34	,	,	PUNCT
gs-3955	200	35	2018	2018	NUM
gs-3955	200	36	)	)	PUNCT
gs-3955	200	37	.	.	PUNCT
gs-3955	201	1	even	even	ADV
gs-3955	201	2	in	in	ADP
gs-3955	201	3	vertebrates	vertebrate	NOUN
gs-3955	201	4	,	,	PUNCT
gs-3955	201	5	long	long	ADV
gs-3955	201	6	-	-	PUNCT
gs-3955	201	7	lived	live	VERB
gs-3955	201	8	species	specie	NOUN
gs-3955	201	9	demonstrate	demonstrate	VERB
gs-3955	201	10	remarkable	remarkable	ADJ
gs-3955	201	11	stem	stem	NOUN
gs-3955	201	12	cell	cell	NOUN
gs-3955	201	13	adaptations	adaptation	NOUN
gs-3955	201	14	.	.	PUNCT
gs-3955	202	1	the	the	DET
gs-3955	202	2	fire	fire	NOUN
gs-3955	202	3	salamander	salamander	NOUN
gs-3955	202	4	maintains	maintain	VERB
gs-3955	202	5	active	active	ADJ
gs-3955	202	6	neural	neural	ADJ
gs-3955	202	7	stem	stem	NOUN
gs-3955	202	8	cells	cell	NOUN
gs-3955	202	9	throughout	throughout	ADP
gs-3955	202	10	its	its	PRON
gs-3955	202	11	50	50	NUM
gs-3955	202	12	-	-	PUNCT
gs-3955	202	13	year	year	NOUN
gs-3955	202	14	lifespan	lifespan	NOUN
gs-3955	202	15	that	that	PRON
gs-3955	202	16	enable	enable	VERB
gs-3955	202	17	complete	complete	ADJ
gs-3955	202	18	spinal	spinal	ADJ
gs-3955	202	19	cord	cord	NOUN
gs-3955	202	20	regeneration	regeneration	NOUN
gs-3955	202	21	after	after	ADP
gs-3955	202	22	injury	injury	NOUN
gs-3955	202	23	(	(	PUNCT
gs-3955	202	24	joven	joven	ADJ
gs-3955	202	25	and	and	CCONJ
gs-3955	202	26	simon	simon	PROPN
gs-3955	202	27	,	,	PUNCT
gs-3955	202	28	2018	2018	NUM
gs-3955	202	29	)	)	PUNCT
gs-3955	202	30	,	,	PUNCT
gs-3955	202	31	while	while	SCONJ
gs-3955	202	32	greenland	greenland	PROPN
gs-3955	202	33	sharks	sharks	PROPN
gs-3955	202	34	show	show	VERB
gs-3955	202	35	no	no	DET
gs-3955	202	36	decline	decline	NOUN
gs-3955	202	37	in	in	ADP
gs-3955	202	38	hepatic	hepatic	ADJ
gs-3955	202	39	stem	stem	NOUN
gs-3955	202	40	cell	cell	NOUN
gs-3955	202	41	populations	population	NOUN
gs-3955	202	42	across	across	ADP
gs-3955	202	43	centuries	century	NOUN
gs-3955	202	44	of	of	ADP
gs-3955	202	45	life	life	NOUN
gs-3955	202	46	(	(	PUNCT
gs-3955	202	47	nielsen	nielsen	PROPN
gs-3955	202	48	et	et	PROPN
gs-3955	202	49	al	al	PROPN
gs-3955	202	50	.	.	PROPN
gs-3955	202	51	,	,	PUNCT
gs-3955	202	52	2016	2016	NUM
gs-3955	202	53	)	)	PUNCT
gs-3955	202	54	.	.	PUNCT
gs-3955	203	1	these	these	DET
gs-3955	203	2	organisms	organism	NOUN
gs-3955	203	3	collectively	collectively	ADV
gs-3955	203	4	challenge	challenge	VERB
gs-3955	203	5	the	the	DET
gs-3955	203	6	dogma	dogma	NOUN
gs-3955	203	7	of	of	ADP
gs-3955	203	8	inevitable	inevitable	ADJ
gs-3955	203	9	stem	stem	NOUN
gs-3955	203	10	cell	cell	NOUN
gs-3955	203	11	exhaustion	exhaustion	NOUN
gs-3955	203	12	and	and	CCONJ
gs-3955	203	13	provide	provide	VERB
gs-3955	203	14	templates	template	NOUN
gs-3955	203	15	for	for	ADP
gs-3955	203	16	engineering	engineer	VERB
gs-3955	203	17	similar	similar	ADJ
gs-3955	203	18	resilience	resilience	NOUN
gs-3955	203	19	in	in	ADP
gs-3955	203	20	human	human	ADJ
gs-3955	203	21	tissues	tissue	NOUN
gs-3955	203	22	.	.	PUNCT
gs-3955	204	1	the	the	DET
gs-3955	204	2	neuroendocrine	neuroendocrine	ADJ
gs-3955	204	3	and	and	CCONJ
gs-3955	204	4	immune	immune	ADJ
gs-3955	204	5	adaptations	adaptation	NOUN
gs-3955	204	6	of	of	ADP
gs-3955	204	7	negligibly	negligibly	ADV
gs-3955	204	8	senescent	senescent	ADJ
gs-3955	204	9	species	specie	NOUN
gs-3955	204	10	offer	offer	VERB
gs-3955	204	11	equally	equally	ADV
gs-3955	204	12	compelling	compelling	ADJ
gs-3955	204	13	insights	insight	NOUN
gs-3955	204	14	.	.	PUNCT
gs-3955	205	1	naked	naked	ADJ
gs-3955	205	2	mole	mole	NOUN
gs-3955	205	3	-	-	PUNCT
gs-3955	205	4	rats	rat	NOUN
gs-3955	205	5	exhibit	exhibit	VERB
gs-3955	205	6	a	a	DET
gs-3955	205	7	unique	unique	ADJ
gs-3955	205	8	glucocorticoid	glucocorticoid	ADJ
gs-3955	205	9	profile	profile	NOUN
gs-3955	205	10	with	with	ADP
gs-3955	205	11	elevated	elevated	ADJ
gs-3955	205	12	corticosterone	corticosterone	NOUN
gs-3955	205	13	levels	level	NOUN
gs-3955	205	14	that	that	PRON
gs-3955	205	15	paradoxically	paradoxically	ADV
gs-3955	205	16	do	do	AUX
gs-3955	205	17	not	not	PART
gs-3955	205	18	induce	induce	VERB
gs-3955	205	19	the	the	DET
gs-3955	205	20	immunosuppressive	immunosuppressive	ADJ
gs-3955	205	21	and	and	CCONJ
gs-3955	205	22	catabolic	catabolic	ADJ
gs-3955	205	23	effects	effect	NOUN
gs-3955	205	24	seen	see	VERB
gs-3955	205	25	in	in	ADP
gs-3955	205	26	other	other	ADJ
gs-3955	205	27	mammals	mammal	NOUN
gs-3955	205	28	(	(	PUNCT
gs-3955	205	29	edrey	edrey	PROPN
gs-3955	205	30	et	et	PROPN
gs-3955	205	31	al	al	PROPN
gs-3955	205	32	.	.	PROPN
gs-3955	205	33	,	,	PUNCT
gs-3955	205	34	2011	2011	NUM
gs-3955	205	35	)	)	PUNCT
gs-3955	205	36	.	.	PUNCT
gs-3955	206	1	this	this	DET
gs-3955	206	2	resilience	resilience	NOUN
gs-3955	206	3	stems	stem	VERB
gs-3955	206	4	from	from	ADP
gs-3955	206	5	a	a	DET
gs-3955	206	6	modified	modify	VERB
gs-3955	206	7	glucocorticoid	glucocorticoid	ADJ
gs-3955	206	8	receptor	receptor	NOUN
gs-3955	206	9	with	with	ADP
gs-3955	206	10	reduced	reduce	VERB
gs-3955	206	11	ligand	ligand	NOUN
gs-3955	206	12	-	-	PUNCT
gs-3955	206	13	binding	bind	VERB
gs-3955	206	14	affinity	affinity	NOUN
gs-3955	206	15	and	and	CCONJ
gs-3955	206	16	preferential	preferential	ADJ
gs-3955	206	17	formation	formation	NOUN
gs-3955	206	18	of	of	ADP
gs-3955	206	19	non	non	ADJ
gs-3955	206	20	-	-	ADJ
gs-3955	206	21	transactivating	transactivating	ADJ
gs-3955	206	22	dimers	dimer	NOUN
gs-3955	206	23	(	(	PUNCT
gs-3955	206	24	turturro	turturro	PROPN
gs-3955	206	25	et	et	PROPN
gs-3955	206	26	al	al	PROPN
gs-3955	206	27	.	.	PROPN
gs-3955	206	28	,	,	PUNCT
gs-3955	206	29	2007	2007	NUM
gs-3955	206	30	)	)	PUNCT
gs-3955	206	31	.	.	PUNCT
gs-3955	207	1	similarly	similarly	ADV
gs-3955	207	2	,	,	PUNCT
gs-3955	207	3	their	their	PRON
gs-3955	207	4	immune	immune	ADJ
gs-3955	207	5	system	system	NOUN
gs-3955	207	6	features	feature	VERB
gs-3955	207	7	an	an	DET
gs-3955	207	8	expanded	expand	VERB
gs-3955	207	9	population	population	NOUN
gs-3955	207	10	of	of	ADP
gs-3955	207	11	natural	natural	ADJ
gs-3955	207	12	killer	killer	NOUN
gs-3955	207	13	cells	cell	NOUN
gs-3955	207	14	and	and	CCONJ
gs-3955	207	15	cytotoxic	cytotoxic	ADJ
gs-3955	207	16	t	t	NOUN
gs-3955	207	17	cells	cell	NOUN
gs-3955	207	18	that	that	PRON
gs-3955	207	19	provide	provide	VERB
gs-3955	207	20	robust	robust	ADJ
gs-3955	207	21	surveillance	surveillance	NOUN
gs-3955	207	22	against	against	ADP
gs-3955	207	23	pathogens	pathogen	NOUN
gs-3955	207	24	without	without	ADP
gs-3955	207	25	the	the	DET
gs-3955	207	26	inflammatory	inflammatory	ADJ
gs-3955	207	27	overreactions	overreaction	NOUN
gs-3955	207	28	that	that	PRON
gs-3955	207	29	drive	drive	VERB
gs-3955	207	30	mammalian	mammalian	NOUN
gs-3955	207	31	inflammaging	inflammage	VERB
gs-3955	207	32	(	(	PUNCT
gs-3955	207	33	hilton	hilton	PROPN
gs-3955	207	34	et	et	PROPN
gs-3955	207	35	al	al	PROPN
gs-3955	207	36	.	.	PROPN
gs-3955	207	37	,	,	PUNCT
gs-3955	207	38	2019	2019	NUM
gs-3955	207	39	)	)	PUNCT
gs-3955	207	40	.	.	PUNCT
gs-3955	208	1	the	the	DET
gs-3955	208	2	greenland	greenland	PROPN
gs-3955	208	3	shark	shark	NOUN
gs-3955	208	4	's	's	PART
gs-3955	208	5	immune	immune	ADJ
gs-3955	208	6	system	system	NOUN
gs-3955	208	7	remains	remain	VERB
gs-3955	208	8	functionally	functionally	ADV
gs-3955	208	9	competent	competent	ADJ
gs-3955	208	10	across	across	ADP
gs-3955	208	11	centuries	century	NOUN
gs-3955	208	12	due	due	ADP
gs-3955	208	13	to	to	ADP
gs-3955	208	14	a	a	DET
gs-3955	208	15	diverse	diverse	ADJ
gs-3955	208	16	repertoire	repertoire	NOUN
gs-3955	208	17	of	of	ADP
gs-3955	208	18	immunoglobulin	immunoglobulin	NOUN
gs-3955	208	19	new	new	ADJ
gs-3955	208	20	antigen	antigen	ADJ
gs-3955	208	21	receptors	receptor	NOUN
gs-3955	208	22	that	that	PRON
gs-3955	208	23	recognize	recognize	VERB
gs-3955	208	24	pathogens	pathogen	NOUN
gs-3955	208	25	with	with	ADP
gs-3955	208	26	remarkable	remarkable	ADJ
gs-3955	208	27	specificity	specificity	NOUN
gs-3955	208	28	(	(	PUNCT
gs-3955	208	29	nielsen	nielsen	PROPN
gs-3955	208	30	et	et	PROPN
gs-3955	208	31	al	al	PROPN
gs-3955	208	32	.	.	PROPN
gs-3955	208	33	,	,	PUNCT
gs-3955	208	34	2016	2016	NUM
gs-3955	208	35	)	)	PUNCT
gs-3955	208	36	.	.	PUNCT
gs-3955	209	1	olms	olm	NOUN
gs-3955	209	2	,	,	PUNCT
gs-3955	209	3	living	live	VERB
gs-3955	209	4	in	in	ADP
gs-3955	209	5	oxygen	oxygen	NOUN
gs-3955	209	6	-	-	PUNCT
gs-3955	209	7	poor	poor	ADJ
gs-3955	209	8	cave	cave	NOUN
gs-3955	209	9	waters	water	NOUN
gs-3955	209	10	,	,	PUNCT
gs-3955	209	11	have	have	AUX
gs-3955	209	12	evolved	evolve	VERB
gs-3955	209	13	a	a	DET
gs-3955	209	14	unique	unique	ADJ
gs-3955	209	15	neuroendocrine	neuroendocrine	ADJ
gs-3955	209	16	response	response	NOUN
gs-3955	209	17	to	to	ADP
gs-3955	209	18	hypoxia	hypoxia	NOUN
gs-3955	209	19	involving	involve	VERB
gs-3955	209	20	melatonin	melatonin	NOUN
gs-3955	209	21	amplification	amplification	NOUN
gs-3955	209	22	and	and	CCONJ
gs-3955	209	23	adenosine	adenosine	NOUN
gs-3955	209	24	signaling	signal	VERB
gs-3955	209	25	that	that	PRON
gs-3955	209	26	prevents	prevent	VERB
gs-3955	209	27	neuronal	neuronal	ADJ
gs-3955	209	28	apoptosis	apoptosis	NOUN
gs-3955	209	29	during	during	ADP
gs-3955	209	30	prolonged	prolonged	ADJ
gs-3955	209	31	oxygen	oxygen	NOUN
gs-3955	209	32	deprivation	deprivation	NOUN
gs-3955	209	33	georgian	georgian	ADJ
gs-3955	209	34	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	209	35	მეცნიერები	მეცნიერები	NOUN
gs-3955	209	36	ტ.	ტ.	VERB
gs-3955	209	37	7	7	NUM
gs-3955	209	38	n	n	PRON
gs-3955	209	39	3	3	NUM
gs-3955	209	40	,	,	PUNCT
gs-3955	209	41	2025	2025	NUM
gs-3955	209	42	369	369	NUM
gs-3955	209	43	(	(	PUNCT
gs-3955	209	44	hervant	hervant	VERB
gs-3955	209	45	et	et	PROPN
gs-3955	209	46	al	al	PROPN
gs-3955	209	47	.	.	PROPN
gs-3955	209	48	,	,	PUNCT
gs-3955	209	49	2001	2001	NUM
gs-3955	209	50	)	)	PUNCT
gs-3955	209	51	.	.	PUNCT
gs-3955	210	1	these	these	DET
gs-3955	210	2	systemic	systemic	ADJ
gs-3955	210	3	adaptations	adaptation	NOUN
gs-3955	210	4	demonstrate	demonstrate	VERB
gs-3955	210	5	that	that	SCONJ
gs-3955	210	6	longevity	longevity	NOUN
gs-3955	210	7	requires	require	VERB
gs-3955	210	8	organism	organism	NOUN
gs-3955	210	9	-	-	PUNCT
gs-3955	210	10	wide	wide	ADJ
gs-3955	210	11	integration	integration	NOUN
gs-3955	210	12	of	of	ADP
gs-3955	210	13	defenses	defense	NOUN
gs-3955	210	14	rather	rather	ADV
gs-3955	210	15	than	than	ADP
gs-3955	210	16	isolated	isolated	ADJ
gs-3955	210	17	cellular	cellular	ADJ
gs-3955	210	18	mechanisms	mechanism	NOUN
gs-3955	210	19	.	.	PUNCT
gs-3955	211	1	translating	translate	VERB
gs-3955	211	2	these	these	DET
gs-3955	211	3	biological	biological	ADJ
gs-3955	211	4	innovations	innovation	NOUN
gs-3955	211	5	into	into	ADP
gs-3955	211	6	human	human	ADJ
gs-3955	211	7	applications	application	NOUN
gs-3955	211	8	presents	present	VERB
gs-3955	211	9	both	both	CCONJ
gs-3955	211	10	unprecedented	unprecedented	ADJ
gs-3955	211	11	opportunities	opportunity	NOUN
gs-3955	211	12	and	and	CCONJ
gs-3955	211	13	formidable	formidable	ADJ
gs-3955	211	14	challenges	challenge	NOUN
gs-3955	211	15	.	.	PUNCT
gs-3955	212	1	the	the	DET
gs-3955	212	2	naked	naked	ADJ
gs-3955	212	3	mole	mole	NOUN
gs-3955	212	4	-	-	PUNCT
gs-3955	212	5	rat	rat	NOUN
gs-3955	212	6	's	's	PART
gs-3955	212	7	highmolecular	highmolecular	ADJ
gs-3955	212	8	-	-	PUNCT
gs-3955	212	9	weight	weight	NOUN
gs-3955	212	10	hyaluronan	hyaluronan	NOUN
gs-3955	212	11	has	have	AUX
gs-3955	212	12	inspired	inspire	VERB
gs-3955	212	13	novel	novel	ADJ
gs-3955	212	14	biomaterials	biomaterial	NOUN
gs-3955	212	15	for	for	ADP
gs-3955	212	16	preventing	prevent	VERB
gs-3955	212	17	postoperative	postoperative	ADJ
gs-3955	212	18	adhesions	adhesion	NOUN
gs-3955	212	19	and	and	CCONJ
gs-3955	212	20	is	be	AUX
gs-3955	212	21	entering	enter	VERB
gs-3955	212	22	clinical	clinical	ADJ
gs-3955	212	23	trials	trial	NOUN
gs-3955	212	24	for	for	ADP
gs-3955	212	25	osteoarthritis	osteoarthritis	NOUN
gs-3955	212	26	treatment	treatment	NOUN
gs-3955	212	27	(	(	PUNCT
gs-3955	212	28	tian	tian	PROPN
gs-3955	212	29	et	et	PROPN
gs-3955	212	30	al	al	PROPN
gs-3955	212	31	.	.	PROPN
gs-3955	212	32	,	,	PUNCT
gs-3955	212	33	2013	2013	NUM
gs-3955	212	34	)	)	PUNCT
gs-3955	212	35	.	.	PUNCT
gs-3955	213	1	compounds	compound	VERB
gs-3955	213	2	that	that	PRON
gs-3955	213	3	activate	activate	VERB
gs-3955	213	4	sirt6	sirt6	NOUN
gs-3955	213	5	—	—	PUNCT
gs-3955	213	6	a	a	DET
gs-3955	213	7	dna	dna	PROPN
gs-3955	213	8	repair	repair	NOUN
gs-3955	213	9	enzyme	enzyme	NOUN
gs-3955	213	10	hyperactive	hyperactive	ADJ
gs-3955	213	11	in	in	ADP
gs-3955	213	12	long	long	ADV
gs-3955	213	13	-	-	PUNCT
gs-3955	213	14	lived	live	VERB
gs-3955	213	15	sharks	shark	NOUN
gs-3955	213	16	—	—	PUNCT
gs-3955	213	17	show	show	VERB
gs-3955	213	18	promise	promise	NOUN
gs-3955	213	19	in	in	ADP
gs-3955	213	20	preclinical	preclinical	ADJ
gs-3955	213	21	models	model	NOUN
gs-3955	213	22	for	for	ADP
gs-3955	213	23	reducing	reduce	VERB
gs-3955	213	24	radiation	radiation	NOUN
gs-3955	213	25	-	-	PUNCT
gs-3955	213	26	induced	induce	VERB
gs-3955	213	27	genomic	genomic	NOUN
gs-3955	213	28	instability	instability	NOUN
gs-3955	213	29	(	(	PUNCT
gs-3955	213	30	beerman	beerman	NOUN
gs-3955	213	31	et	et	PROPN
gs-3955	213	32	al	al	PROPN
gs-3955	213	33	.	.	PROPN
gs-3955	213	34	,	,	PUNCT
gs-3955	213	35	2014	2014	NUM
gs-3955	213	36	)	)	PUNCT
gs-3955	213	37	.	.	PUNCT
gs-3955	214	1	however	however	ADV
gs-3955	214	2	,	,	PUNCT
gs-3955	214	3	significant	significant	ADJ
gs-3955	214	4	barriers	barrier	NOUN
gs-3955	214	5	exist	exist	VERB
gs-3955	214	6	:	:	PUNCT
gs-3955	214	7	the	the	DET
gs-3955	214	8	pleiotropic	pleiotropic	ADJ
gs-3955	214	9	nature	nature	NOUN
gs-3955	214	10	of	of	ADP
gs-3955	214	11	longevity	longevity	NOUN
gs-3955	214	12	genes	gene	NOUN
gs-3955	214	13	means	mean	VERB
gs-3955	214	14	that	that	SCONJ
gs-3955	214	15	manipulating	manipulate	VERB
gs-3955	214	16	single	single	ADJ
gs-3955	214	17	pathways	pathway	NOUN
gs-3955	214	18	may	may	AUX
gs-3955	214	19	have	have	VERB
gs-3955	214	20	unforeseen	unforeseen	ADJ
gs-3955	214	21	consequences	consequence	NOUN
gs-3955	214	22	(	(	PUNCT
gs-3955	214	23	gems	gem	NOUN
gs-3955	214	24	and	and	CCONJ
gs-3955	214	25	partridge	partridge	NOUN
gs-3955	214	26	,	,	PUNCT
gs-3955	214	27	2013	2013	NUM
gs-3955	214	28	)	)	PUNCT
gs-3955	214	29	;	;	PUNCT
gs-3955	214	30	the	the	DET
gs-3955	214	31	metabolic	metabolic	NOUN
gs-3955	214	32	depression	depression	NOUN
gs-3955	214	33	seen	see	VERB
gs-3955	214	34	in	in	ADP
gs-3955	214	35	many	many	ADJ
gs-3955	214	36	longlived	longlived	ADJ
gs-3955	214	37	species	species	NOUN
gs-3955	214	38	conflicts	conflict	NOUN
gs-3955	214	39	with	with	ADP
gs-3955	214	40	human	human	ADJ
gs-3955	214	41	cognitive	cognitive	ADJ
gs-3955	214	42	demands	demand	NOUN
gs-3955	214	43	(	(	PUNCT
gs-3955	214	44	austad	austad	NOUN
gs-3955	214	45	,	,	PUNCT
gs-3955	214	46	2010	2010	NUM
gs-3955	214	47	)	)	PUNCT
gs-3955	214	48	;	;	PUNCT
gs-3955	214	49	and	and	CCONJ
gs-3955	214	50	the	the	DET
gs-3955	214	51	evolutionary	evolutionary	ADJ
gs-3955	214	52	divergence	divergence	NOUN
gs-3955	214	53	between	between	ADP
gs-3955	214	54	these	these	DET
gs-3955	214	55	species	specie	NOUN
gs-3955	214	56	and	and	CCONJ
gs-3955	214	57	humans	human	NOUN
gs-3955	214	58	complicates	complicate	VERB
gs-3955	214	59	direct	direct	ADJ
gs-3955	214	60	therapeutic	therapeutic	ADJ
gs-3955	214	61	transfer	transfer	NOUN
gs-3955	214	62	(	(	PUNCT
gs-3955	214	63	de	de	X
gs-3955	214	64	magalhães	magalhães	NOUN
gs-3955	214	65	,	,	PUNCT
gs-3955	214	66	2014	2014	NUM
gs-3955	214	67	)	)	PUNCT
gs-3955	214	68	.	.	PUNCT
gs-3955	215	1	emerging	emerge	VERB
gs-3955	215	2	technologies	technology	NOUN
gs-3955	215	3	offer	offer	VERB
gs-3955	215	4	potential	potential	ADJ
gs-3955	215	5	solutions	solution	NOUN
gs-3955	215	6	.	.	PUNCT
gs-3955	216	1	crispr	crispr	NOUN
gs-3955	216	2	-	-	PUNCT
gs-3955	216	3	based	base	VERB
gs-3955	216	4	gene	gene	NOUN
gs-3955	216	5	drives	drive	NOUN
gs-3955	216	6	could	could	AUX
gs-3955	216	7	theoretically	theoretically	ADV
gs-3955	216	8	introduce	introduce	VERB
gs-3955	216	9	beneficial	beneficial	ADJ
gs-3955	216	10	genetic	genetic	ADJ
gs-3955	216	11	variants	variant	NOUN
gs-3955	216	12	from	from	ADP
gs-3955	216	13	long	long	ADV
gs-3955	216	14	-	-	PUNCT
gs-3955	216	15	lived	live	VERB
gs-3955	216	16	species	specie	NOUN
gs-3955	216	17	into	into	ADP
gs-3955	216	18	human	human	ADJ
gs-3955	216	19	stem	stem	NOUN
gs-3955	216	20	cells	cell	NOUN
gs-3955	216	21	(	(	PUNCT
gs-3955	216	22	grunwald	grunwald	PROPN
gs-3955	216	23	et	et	PROPN
gs-3955	216	24	al	al	PROPN
gs-3955	216	25	.	.	PROPN
gs-3955	216	26	,	,	PUNCT
gs-3955	216	27	2019	2019	NUM
gs-3955	216	28	)	)	PUNCT
gs-3955	216	29	,	,	PUNCT
gs-3955	216	30	while	while	SCONJ
gs-3955	216	31	machine	machine	NOUN
gs-3955	216	32	learning	learn	VERB
gs-3955	216	33	algorithms	algorithm	NOUN
gs-3955	216	34	are	be	AUX
gs-3955	216	35	now	now	ADV
gs-3955	216	36	identifying	identify	VERB
gs-3955	216	37	small	small	ADJ
gs-3955	216	38	molecules	molecule	NOUN
gs-3955	216	39	that	that	PRON
gs-3955	216	40	mimic	mimic	VERB
gs-3955	216	41	the	the	DET
gs-3955	216	42	effects	effect	NOUN
gs-3955	216	43	of	of	ADP
gs-3955	216	44	longevity	longevity	NOUN
gs-3955	216	45	mutations	mutation	NOUN
gs-3955	216	46	without	without	ADP
gs-3955	216	47	genetic	genetic	ADJ
gs-3955	216	48	modification	modification	NOUN
gs-3955	216	49	(	(	PUNCT
gs-3955	216	50	zhavoronkov	zhavoronkov	NOUN
gs-3955	216	51	et	et	NOUN
gs-3955	216	52	al	al	PROPN
gs-3955	216	53	.	.	PROPN
gs-3955	216	54	,	,	PUNCT
gs-3955	216	55	2019	2019	NUM
gs-3955	216	56	)	)	PUNCT
gs-3955	216	57	.	.	PUNCT
gs-3955	217	1	synthetic	synthetic	ADJ
gs-3955	217	2	biology	biology	NOUN
gs-3955	217	3	approaches	approach	NOUN
gs-3955	217	4	are	be	AUX
gs-3955	217	5	creating	create	VERB
gs-3955	217	6	artificial	artificial	ADJ
gs-3955	217	7	gene	gene	NOUN
gs-3955	217	8	circuits	circuit	NOUN
gs-3955	217	9	that	that	PRON
gs-3955	217	10	replicate	replicate	VERB
gs-3955	217	11	the	the	DET
gs-3955	217	12	foxo	foxo	NOUN
gs-3955	217	13	–	–	PUNCT
gs-3955	217	14	sirt	sirt	PROPN
gs-3955	217	15	–	–	PUNCT
gs-3955	217	16	ampk	ampk	NOUN
gs-3955	217	17	signaling	signal	VERB
gs-3955	217	18	networks	network	NOUN
gs-3955	217	19	from	from	ADP
gs-3955	217	20	hydra	hydra	PROPN
gs-3955	217	21	and	and	CCONJ
gs-3955	217	22	naked	naked	ADJ
gs-3955	217	23	mole	mole	NOUN
gs-3955	217	24	-	-	PUNCT
gs-3955	217	25	rats	rat	NOUN
gs-3955	217	26	in	in	ADP
gs-3955	217	27	human	human	ADJ
gs-3955	217	28	cells	cell	NOUN
gs-3955	217	29	(	(	PUNCT
gs-3955	217	30	kenyon	kenyon	PROPN
gs-3955	217	31	,	,	PUNCT
gs-3955	217	32	2010	2010	NUM
gs-3955	217	33	)	)	PUNCT
gs-3955	217	34	.	.	PUNCT
gs-3955	218	1	perhaps	perhaps	ADV
gs-3955	218	2	most	most	ADV
gs-3955	218	3	promising	promising	ADJ
gs-3955	218	4	are	be	AUX
gs-3955	218	5	lipid	lipid	NOUN
gs-3955	218	6	nanoparticle	nanoparticle	NOUN
gs-3955	218	7	delivery	delivery	NOUN
gs-3955	218	8	systems	system	NOUN
gs-3955	218	9	carrying	carry	VERB
gs-3955	218	10	modified	modified	ADJ
gs-3955	218	11	mrna	mrna	NOUN
gs-3955	218	12	that	that	PRON
gs-3955	218	13	temporarily	temporarily	ADV
gs-3955	218	14	express	express	VERB
gs-3955	218	15	sharkderived	sharkderived	ADJ
gs-3955	218	16	dna	dna	PROPN
gs-3955	218	17	repair	repair	NOUN
gs-3955	218	18	enzymes	enzyme	NOUN
gs-3955	218	19	or	or	CCONJ
gs-3955	218	20	clam	clam	NOUN
gs-3955	218	21	-	-	PUNCT
gs-3955	218	22	derived	derive	VERB
gs-3955	218	23	chaperone	chaperone	NOUN
gs-3955	218	24	proteins	protein	NOUN
gs-3955	218	25	in	in	ADP
gs-3955	218	26	human	human	ADJ
gs-3955	218	27	tissues	tissue	NOUN
gs-3955	218	28	(	(	PUNCT
gs-3955	218	29	hou	hou	NOUN
gs-3955	218	30	et	et	PROPN
gs-3955	218	31	al	al	PROPN
gs-3955	218	32	.	.	PROPN
gs-3955	218	33	,	,	PUNCT
gs-3955	218	34	2021	2021	NUM
gs-3955	218	35	)	)	PUNCT
gs-3955	218	36	.	.	PUNCT
gs-3955	219	1	the	the	DET
gs-3955	219	2	ethical	ethical	ADJ
gs-3955	219	3	dimensions	dimension	NOUN
gs-3955	219	4	of	of	ADP
gs-3955	219	5	extending	extend	VERB
gs-3955	219	6	human	human	ADJ
gs-3955	219	7	healthspan	healthspan	NOUN
gs-3955	219	8	through	through	ADP
gs-3955	219	9	these	these	DET
gs-3955	219	10	discoveries	discovery	NOUN
gs-3955	219	11	warrant	warrant	VERB
gs-3955	219	12	careful	careful	ADJ
gs-3955	219	13	consideration	consideration	NOUN
gs-3955	219	14	.	.	PUNCT
gs-3955	220	1	while	while	SCONJ
gs-3955	220	2	eliminating	eliminate	VERB
gs-3955	220	3	debilitating	debilitate	VERB
gs-3955	220	4	age	age	NOUN
gs-3955	220	5	-	-	PUNCT
gs-3955	220	6	related	relate	VERB
gs-3955	220	7	diseases	disease	NOUN
gs-3955	220	8	represents	represent	VERB
gs-3955	220	9	an	an	DET
gs-3955	220	10	unambiguous	unambiguous	ADJ
gs-3955	220	11	good	good	ADJ
gs-3955	220	12	,	,	PUNCT
gs-3955	220	13	significant	significant	ADJ
gs-3955	220	14	lifespan	lifespan	NOUN
gs-3955	220	15	extension	extension	NOUN
gs-3955	220	16	could	could	AUX
gs-3955	220	17	exacerbate	exacerbate	VERB
gs-3955	220	18	social	social	ADJ
gs-3955	220	19	inequalities	inequality	NOUN
gs-3955	220	20	if	if	SCONJ
gs-3955	220	21	available	available	ADJ
gs-3955	220	22	only	only	ADV
gs-3955	220	23	to	to	ADP
gs-3955	220	24	economic	economic	ADJ
gs-3955	220	25	elites	elite	NOUN
gs-3955	220	26	(	(	PUNCT
gs-3955	220	27	austad	austad	NOUN
gs-3955	220	28	,	,	PUNCT
gs-3955	220	29	2010	2010	NUM
gs-3955	220	30	)	)	PUNCT
gs-3955	220	31	.	.	PUNCT
gs-3955	221	1	demographic	demographic	ADJ
gs-3955	221	2	shifts	shift	NOUN
gs-3955	221	3	toward	toward	ADP
gs-3955	221	4	older	old	ADJ
gs-3955	221	5	populations	population	NOUN
gs-3955	221	6	would	would	AUX
gs-3955	221	7	require	require	VERB
gs-3955	221	8	reimagining	reimagine	VERB
gs-3955	221	9	social	social	ADJ
gs-3955	221	10	structures	structure	NOUN
gs-3955	221	11	,	,	PUNCT
gs-3955	221	12	retirement	retirement	NOUN
gs-3955	221	13	systems	system	NOUN
gs-3955	221	14	,	,	PUNCT
gs-3955	221	15	and	and	CCONJ
gs-3955	221	16	intergenerational	intergenerational	ADJ
gs-3955	221	17	resource	resource	NOUN
gs-3955	221	18	allocation	allocation	NOUN
gs-3955	221	19	(	(	PUNCT
gs-3955	221	20	finch	finch	NOUN
gs-3955	221	21	,	,	PUNCT
gs-3955	221	22	2009	2009	NUM
gs-3955	221	23	)	)	PUNCT
gs-3955	221	24	.	.	PUNCT
gs-3955	222	1	philosophically	philosophically	ADV
gs-3955	222	2	,	,	PUNCT
gs-3955	222	3	these	these	DET
gs-3955	222	4	organisms	organism	NOUN
gs-3955	222	5	challenge	challenge	VERB
gs-3955	222	6	our	our	PRON
gs-3955	222	7	very	very	ADJ
gs-3955	222	8	conception	conception	NOUN
gs-3955	222	9	of	of	ADP
gs-3955	222	10	what	what	PRON
gs-3955	222	11	it	it	PRON
gs-3955	222	12	means	mean	VERB
gs-3955	222	13	to	to	PART
gs-3955	222	14	be	be	AUX
gs-3955	222	15	human	human	ADJ
gs-3955	222	16	in	in	ADP
gs-3955	222	17	a	a	DET
gs-3955	222	18	potentially	potentially	ADV
gs-3955	222	19	post	post	ADJ
gs-3955	222	20	-	-	ADJ
gs-3955	222	21	aging	age	VERB
gs-3955	222	22	future	future	NOUN
gs-3955	222	23	(	(	PUNCT
gs-3955	222	24	lópez	lópez	NOUN
gs-3955	222	25	-	-	PUNCT
gs-3955	222	26	otín	otín	NOUN
gs-3955	222	27	et	et	PROPN
gs-3955	222	28	al	al	PROPN
gs-3955	222	29	.	.	PROPN
gs-3955	222	30	,	,	PUNCT
gs-3955	222	31	2013	2013	NUM
gs-3955	222	32	)	)	PUNCT
gs-3955	222	33	.	.	PUNCT
gs-3955	223	1	if	if	SCONJ
gs-3955	223	2	biological	biological	ADJ
gs-3955	223	3	constraints	constraint	NOUN
gs-3955	223	4	on	on	ADP
gs-3955	223	5	lifespan	lifespan	NOUN
gs-3955	223	6	are	be	AUX
gs-3955	223	7	indeed	indeed	ADV
gs-3955	223	8	mutable	mutable	ADJ
gs-3955	223	9	,	,	PUNCT
gs-3955	223	10	how	how	SCONJ
gs-3955	223	11	might	might	AUX
gs-3955	223	12	our	our	PRON
gs-3955	223	13	relationship	relationship	NOUN
gs-3955	223	14	to	to	ADP
gs-3955	223	15	time	time	NOUN
gs-3955	223	16	,	,	PUNCT
gs-3955	223	17	ambition	ambition	NOUN
gs-3955	223	18	,	,	PUNCT
gs-3955	223	19	and	and	CCONJ
gs-3955	223	20	legacy	legacy	NOUN
gs-3955	223	21	transform	transform	NOUN
gs-3955	223	22	?	?	PUNCT
gs-3955	224	1	the	the	DET
gs-3955	224	2	naked	naked	ADJ
gs-3955	224	3	mole	mole	NOUN
gs-3955	224	4	-	-	PUNCT
gs-3955	224	5	rat	rat	NOUN
gs-3955	224	6	’s	’s	PART
gs-3955	224	7	social	social	ADJ
gs-3955	224	8	structure	structure	NOUN
gs-3955	224	9	suggests	suggest	VERB
gs-3955	224	10	that	that	SCONJ
gs-3955	224	11	cooperative	cooperative	ADJ
gs-3955	224	12	social	social	ADJ
gs-3955	224	13	systems	system	NOUN
gs-3955	224	14	may	may	AUX
gs-3955	224	15	be	be	AUX
gs-3955	224	16	integral	integral	ADJ
gs-3955	224	17	to	to	ADP
gs-3955	224	18	extreme	extreme	ADJ
gs-3955	224	19	longevity	longevity	NOUN
gs-3955	224	20	—	—	PUNCT
gs-3955	224	21	a	a	DET
gs-3955	224	22	finding	finding	NOUN
gs-3955	224	23	with	with	ADP
gs-3955	224	24	profound	profound	ADJ
gs-3955	224	25	implications	implication	NOUN
gs-3955	224	26	for	for	ADP
gs-3955	224	27	human	human	ADJ
gs-3955	224	28	societal	societal	ADJ
gs-3955	224	29	organization	organization	NOUN
gs-3955	224	30	(	(	PUNCT
gs-3955	224	31	buffenstein	buffenstein	ADV
gs-3955	224	32	,	,	PUNCT
gs-3955	224	33	2008	2008	NUM
gs-3955	224	34	)	)	PUNCT
gs-3955	224	35	.	.	PUNCT
gs-3955	225	1	as	as	SCONJ
gs-3955	225	2	we	we	PRON
gs-3955	225	3	decode	decode	VERB
gs-3955	225	4	the	the	DET
gs-3955	225	5	biological	biological	ADJ
gs-3955	225	6	blueprints	blueprint	NOUN
gs-3955	225	7	of	of	ADP
gs-3955	225	8	these	these	DET
gs-3955	225	9	species	specie	NOUN
gs-3955	225	10	,	,	PUNCT
gs-3955	225	11	we	we	PRON
gs-3955	225	12	must	must	AUX
gs-3955	225	13	simultaneously	simultaneously	ADV
gs-3955	225	14	develop	develop	VERB
gs-3955	225	15	ethical	ethical	ADJ
gs-3955	225	16	frameworks	framework	NOUN
gs-3955	225	17	that	that	PRON
gs-3955	225	18	ensure	ensure	VERB
gs-3955	225	19	equitable	equitable	ADJ
gs-3955	225	20	access	access	NOUN
gs-3955	225	21	to	to	ADP
gs-3955	225	22	longevity	longevity	NOUN
gs-3955	225	23	interventions	intervention	NOUN
gs-3955	225	24	and	and	CCONJ
gs-3955	225	25	preserve	preserve	VERB
gs-3955	225	26	the	the	DET
gs-3955	225	27	human	human	ADJ
gs-3955	225	28	experience	experience	NOUN
gs-3955	225	29	amidst	amidst	ADP
gs-3955	225	30	extended	extended	ADJ
gs-3955	225	31	vitality	vitality	NOUN
gs-3955	225	32	(	(	PUNCT
gs-3955	225	33	austad	austad	NOUN
gs-3955	225	34	,	,	PUNCT
gs-3955	225	35	2022	2022	NUM
gs-3955	225	36	)	)	PUNCT
gs-3955	225	37	.	.	PUNCT
gs-3955	226	1	recent	recent	ADJ
gs-3955	226	2	technological	technological	ADJ
gs-3955	226	3	advances	advance	NOUN
gs-3955	226	4	are	be	AUX
gs-3955	226	5	accelerating	accelerate	VERB
gs-3955	226	6	discovery	discovery	NOUN
gs-3955	226	7	in	in	ADP
gs-3955	226	8	this	this	DET
gs-3955	226	9	field	field	NOUN
gs-3955	226	10	at	at	ADP
gs-3955	226	11	an	an	DET
gs-3955	226	12	unprecedented	unprecedented	ADJ
gs-3955	226	13	pace	pace	NOUN
gs-3955	226	14	.	.	PUNCT
gs-3955	227	1	quantum	quantum	NOUN
gs-3955	227	2	-	-	PUNCT
gs-3955	227	3	enhanced	enhance	VERB
gs-3955	227	4	magnetic	magnetic	ADJ
gs-3955	227	5	resonance	resonance	NOUN
gs-3955	227	6	imaging	imaging	NOUN
gs-3955	227	7	now	now	ADV
gs-3955	227	8	allows	allow	VERB
gs-3955	227	9	non	non	ADJ
gs-3955	227	10	-	-	ADJ
gs-3955	227	11	invasive	invasive	ADJ
gs-3955	227	12	study	study	NOUN
gs-3955	227	13	of	of	ADP
gs-3955	227	14	georgian	georgian	PROPN
gs-3955	227	15	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	227	16	მეცნიერები	მეცნიერები	NOUN
gs-3955	227	17	ტ.	ტ.	VERB
gs-3955	227	18	7	7	NUM
gs-3955	227	19	n	n	PRON
gs-3955	227	20	3	3	NUM
gs-3955	227	21	,	,	PUNCT
gs-3955	227	22	2025	2025	NUM
gs-3955	227	23	370	370	NUM
gs-3955	227	24	metabolic	metabolic	NOUN
gs-3955	227	25	processes	process	NOUN
gs-3955	227	26	in	in	ADP
gs-3955	227	27	living	live	VERB
gs-3955	227	28	greenland	greenland	PROPN
gs-3955	227	29	sharks	shark	NOUN
gs-3955	227	30	without	without	ADP
gs-3955	227	31	capture	capture	NOUN
gs-3955	227	32	or	or	CCONJ
gs-3955	227	33	disturbance	disturbance	NOUN
gs-3955	227	34	(	(	PUNCT
gs-3955	227	35	hansen	hansen	PROPN
gs-3955	227	36	et	et	PROPN
gs-3955	227	37	al	al	PROPN
gs-3955	227	38	.	.	PROPN
gs-3955	227	39	,	,	PUNCT
gs-3955	227	40	2021	2021	NUM
gs-3955	227	41	)	)	PUNCT
gs-3955	227	42	.	.	PUNCT
gs-3955	228	1	single	single	ADJ
gs-3955	228	2	-	-	PUNCT
gs-3955	228	3	molecule	molecule	NOUN
gs-3955	228	4	real	real	ADJ
gs-3955	228	5	-	-	PUNCT
gs-3955	228	6	time	time	NOUN
gs-3955	228	7	sequencing	sequencing	NOUN
gs-3955	228	8	provides	provide	VERB
gs-3955	228	9	complete	complete	ADJ
gs-3955	228	10	epigenetic	epigenetic	ADJ
gs-3955	228	11	maps	map	NOUN
gs-3955	228	12	of	of	ADP
gs-3955	228	13	century	century	NOUN
gs-3955	228	14	-	-	PUNCT
gs-3955	228	15	old	old	ADJ
gs-3955	228	16	rockfish	rockfish	ADJ
gs-3955	228	17	tissues	tissue	NOUN
gs-3955	228	18	,	,	PUNCT
gs-3955	228	19	revealing	reveal	VERB
gs-3955	228	20	methylation	methylation	NOUN
gs-3955	228	21	patterns	pattern	NOUN
gs-3955	228	22	that	that	PRON
gs-3955	228	23	maintain	maintain	VERB
gs-3955	228	24	youthful	youthful	ADJ
gs-3955	228	25	gene	gene	NOUN
gs-3955	228	26	expression	expression	NOUN
gs-3955	228	27	(	(	PUNCT
gs-3955	228	28	horvath	horvath	PROPN
gs-3955	228	29	et	et	PROPN
gs-3955	228	30	al	al	PROPN
gs-3955	228	31	.	.	PROPN
gs-3955	228	32	,	,	PUNCT
gs-3955	228	33	2022	2022	NUM
gs-3955	228	34	)	)	PUNCT
gs-3955	228	35	.	.	PUNCT
gs-3955	229	1	artificial	artificial	ADJ
gs-3955	229	2	intelligence	intelligence	NOUN
gs-3955	229	3	systems	system	NOUN
gs-3955	229	4	trained	train	VERB
gs-3955	229	5	on	on	ADP
gs-3955	229	6	multi	multi	ADJ
gs-3955	229	7	-	-	ADJ
gs-3955	229	8	omic	omic	ADJ
gs-3955	229	9	data	datum	NOUN
gs-3955	229	10	from	from	ADP
gs-3955	229	11	long	long	ADV
gs-3955	229	12	-	-	PUNCT
gs-3955	229	13	lived	live	VERB
gs-3955	229	14	species	specie	NOUN
gs-3955	229	15	are	be	AUX
gs-3955	229	16	identifying	identify	VERB
gs-3955	229	17	previously	previously	ADV
gs-3955	229	18	unrecognized	unrecognize	VERB
gs-3955	229	19	longevity	longevity	NOUN
gs-3955	229	20	signatures	signature	NOUN
gs-3955	229	21	,	,	PUNCT
gs-3955	229	22	such	such	ADJ
gs-3955	229	23	as	as	ADP
gs-3955	229	24	conserved	conserve	VERB
gs-3955	229	25	microrna	microrna	PROPN
gs-3955	229	26	clusters	cluster	NOUN
gs-3955	229	27	that	that	PRON
gs-3955	229	28	coordinate	coordinate	VERB
gs-3955	229	29	stress	stress	NOUN
gs-3955	229	30	response	response	NOUN
gs-3955	229	31	pathways	pathway	NOUN
gs-3955	229	32	across	across	ADP
gs-3955	229	33	phylogeny	phylogeny	NOUN
gs-3955	229	34	(	(	PUNCT
gs-3955	229	35	zhang	zhang	X
gs-3955	229	36	et	et	PROPN
gs-3955	229	37	al	al	PROPN
gs-3955	229	38	.	.	PROPN
gs-3955	229	39	,	,	PUNCT
gs-3955	229	40	2023	2023	NUM
gs-3955	229	41	)	)	PUNCT
gs-3955	229	42	.	.	PUNCT
gs-3955	230	1	organoid	organoid	PROPN
gs-3955	230	2	systems	system	NOUN
gs-3955	230	3	derived	derive	VERB
gs-3955	230	4	from	from	ADP
gs-3955	230	5	naked	naked	ADJ
gs-3955	230	6	mole	mole	ADJ
gs-3955	230	7	-	-	PUNCT
gs-3955	230	8	rat	rat	NOUN
gs-3955	230	9	tissues	tissue	NOUN
gs-3955	230	10	permit	permit	VERB
gs-3955	230	11	highthroughput	highthroughput	NOUN
gs-3955	230	12	screening	screening	NOUN
gs-3955	230	13	of	of	ADP
gs-3955	230	14	compounds	compound	NOUN
gs-3955	230	15	that	that	PRON
gs-3955	230	16	enhance	enhance	VERB
gs-3955	230	17	dna	dna	PROPN
gs-3955	230	18	repair	repair	NOUN
gs-3955	230	19	fidelity	fidelity	NOUN
gs-3955	230	20	or	or	CCONJ
gs-3955	230	21	proteostasis	proteostasis	NOUN
gs-3955	230	22	(	(	PUNCT
gs-3955	230	23	macrae	macrae	PROPN
gs-3955	230	24	et	et	PROPN
gs-3955	230	25	al	al	PROPN
gs-3955	230	26	.	.	PROPN
gs-3955	230	27	,	,	PUNCT
gs-3955	230	28	2015	2015	NUM
gs-3955	230	29	)	)	PUNCT
gs-3955	230	30	.	.	PUNCT
gs-3955	231	1	perhaps	perhaps	ADV
gs-3955	231	2	most	most	ADV
gs-3955	231	3	revolutionary	revolutionary	ADJ
gs-3955	231	4	is	be	AUX
gs-3955	231	5	the	the	DET
gs-3955	231	6	development	development	NOUN
gs-3955	231	7	of	of	ADP
gs-3955	231	8	interspecies	interspecie	NOUN
gs-3955	231	9	chimeric	chimeric	ADJ
gs-3955	231	10	models	model	NOUN
gs-3955	231	11	,	,	PUNCT
gs-3955	231	12	where	where	SCONJ
gs-3955	231	13	human	human	ADJ
gs-3955	231	14	stem	stem	NOUN
gs-3955	231	15	cells	cell	NOUN
gs-3955	231	16	are	be	AUX
gs-3955	231	17	introduced	introduce	VERB
gs-3955	231	18	into	into	ADP
gs-3955	231	19	planarian	planarian	ADJ
gs-3955	231	20	or	or	CCONJ
gs-3955	231	21	hydra	hydra	NOUN
gs-3955	231	22	regeneration	regeneration	NOUN
gs-3955	231	23	systems	system	NOUN
gs-3955	231	24	to	to	PART
gs-3955	231	25	determine	determine	VERB
gs-3955	231	26	whether	whether	SCONJ
gs-3955	231	27	their	their	PRON
gs-3955	231	28	immortality	immortality	NOUN
gs-3955	231	29	pathways	pathway	NOUN
gs-3955	231	30	can	can	AUX
gs-3955	231	31	be	be	AUX
gs-3955	231	32	activated	activate	VERB
gs-3955	231	33	in	in	ADP
gs-3955	231	34	human	human	ADJ
gs-3955	231	35	cellular	cellular	ADJ
gs-3955	231	36	contexts	context	NOUN
gs-3955	231	37	(	(	PUNCT
gs-3955	231	38	rink	rink	NOUN
gs-3955	231	39	,	,	PUNCT
gs-3955	231	40	2013	2013	NUM
gs-3955	231	41	;	;	PUNCT
gs-3955	231	42	pearson	pearson	PROPN
gs-3955	231	43	&	&	CCONJ
gs-3955	231	44	sánchez	sánchez	PROPN
gs-3955	231	45	alvarado	alvarado	PROPN
gs-3955	231	46	,	,	PUNCT
gs-3955	231	47	2018	2018	NUM
gs-3955	231	48	)	)	PUNCT
gs-3955	231	49	.	.	PUNCT
gs-3955	232	1	the	the	DET
gs-3955	232	2	study	study	NOUN
gs-3955	232	3	of	of	ADP
gs-3955	232	4	negligibly	negligibly	ADV
gs-3955	232	5	senescent	senescent	ADJ
gs-3955	232	6	organisms	organism	NOUN
gs-3955	232	7	ultimately	ultimately	ADV
gs-3955	232	8	transcends	transcend	VERB
gs-3955	232	9	biomedical	biomedical	ADJ
gs-3955	232	10	implications	implication	NOUN
gs-3955	232	11	to	to	PART
gs-3955	232	12	challenge	challenge	VERB
gs-3955	232	13	our	our	PRON
gs-3955	232	14	fundamental	fundamental	ADJ
gs-3955	232	15	understanding	understanding	NOUN
gs-3955	232	16	of	of	ADP
gs-3955	232	17	life	life	NOUN
gs-3955	232	18	's	's	PART
gs-3955	232	19	temporal	temporal	ADJ
gs-3955	232	20	boundaries	boundary	NOUN
gs-3955	232	21	.	.	PUNCT
gs-3955	233	1	these	these	DET
gs-3955	233	2	species	specie	NOUN
gs-3955	233	3	demonstrate	demonstrate	VERB
gs-3955	233	4	that	that	SCONJ
gs-3955	233	5	biological	biological	ADJ
gs-3955	233	6	time	time	NOUN
gs-3955	233	7	is	be	AUX
gs-3955	233	8	not	not	PART
gs-3955	233	9	an	an	DET
gs-3955	233	10	inevitable	inevitable	ADJ
gs-3955	233	11	linear	linear	NOUN
gs-3955	233	12	progression	progression	NOUN
gs-3955	233	13	toward	toward	ADP
gs-3955	233	14	disorder	disorder	NOUN
gs-3955	233	15	,	,	PUNCT
gs-3955	233	16	but	but	CCONJ
gs-3955	233	17	rather	rather	ADV
gs-3955	233	18	a	a	DET
gs-3955	233	19	flexible	flexible	ADJ
gs-3955	233	20	dimension	dimension	NOUN
gs-3955	233	21	that	that	PRON
gs-3955	233	22	can	can	AUX
gs-3955	233	23	be	be	AUX
gs-3955	233	24	shaped	shape	VERB
gs-3955	233	25	through	through	ADP
gs-3955	233	26	evolutionary	evolutionary	ADJ
gs-3955	233	27	innovation	innovation	NOUN
gs-3955	233	28	and	and	CCONJ
gs-3955	233	29	molecular	molecular	ADJ
gs-3955	233	30	engineering	engineering	NOUN
gs-3955	233	31	(	(	PUNCT
gs-3955	233	32	finch	finch	NOUN
gs-3955	233	33	,	,	PUNCT
gs-3955	233	34	2009	2009	NUM
gs-3955	233	35	;	;	PUNCT
gs-3955	233	36	austad	austad	NOUN
gs-3955	233	37	,	,	PUNCT
gs-3955	233	38	2022	2022	NUM
gs-3955	233	39	)	)	PUNCT
gs-3955	233	40	.	.	PUNCT
gs-3955	234	1	their	their	PRON
gs-3955	234	2	existence	existence	NOUN
gs-3955	234	3	suggests	suggest	VERB
gs-3955	234	4	that	that	SCONJ
gs-3955	234	5	aging	aging	NOUN
gs-3955	234	6	may	may	AUX
gs-3955	234	7	be	be	AUX
gs-3955	234	8	less	less	ADJ
gs-3955	234	9	an	an	DET
gs-3955	234	10	intrinsic	intrinsic	ADJ
gs-3955	234	11	property	property	NOUN
gs-3955	234	12	of	of	ADP
gs-3955	234	13	complex	complex	ADJ
gs-3955	234	14	life	life	NOUN
gs-3955	234	15	than	than	ADP
gs-3955	234	16	an	an	DET
gs-3955	234	17	evolutionary	evolutionary	ADJ
gs-3955	234	18	compromise	compromise	NOUN
gs-3955	234	19	—	—	PUNCT
gs-3955	234	20	one	one	NOUN
gs-3955	234	21	that	that	PRON
gs-3955	234	22	can	can	AUX
gs-3955	234	23	be	be	AUX
gs-3955	234	24	systematically	systematically	ADV
gs-3955	234	25	overcome	overcome	VERB
gs-3955	234	26	(	(	PUNCT
gs-3955	234	27	lópez	lópez	NOUN
gs-3955	234	28	-	-	PUNCT
gs-3955	234	29	otín	otín	NOUN
gs-3955	234	30	et	et	PROPN
gs-3955	234	31	al	al	PROPN
gs-3955	234	32	.	.	PROPN
gs-3955	234	33	,	,	PUNCT
gs-3955	234	34	2013	2013	NUM
gs-3955	234	35	)	)	PUNCT
gs-3955	234	36	.	.	PUNCT
gs-3955	235	1	as	as	SCONJ
gs-3955	235	2	research	research	NOUN
gs-3955	235	3	progresses	progress	NOUN
gs-3955	235	4	,	,	PUNCT
gs-3955	235	5	we	we	PRON
gs-3955	235	6	stand	stand	VERB
gs-3955	235	7	at	at	ADP
gs-3955	235	8	the	the	DET
gs-3955	235	9	threshold	threshold	NOUN
gs-3955	235	10	of	of	ADP
gs-3955	235	11	a	a	DET
gs-3955	235	12	new	new	ADJ
gs-3955	235	13	paradigm	paradigm	NOUN
gs-3955	235	14	in	in	ADP
gs-3955	235	15	which	which	PRON
gs-3955	235	16	the	the	DET
gs-3955	235	17	molecular	molecular	ADJ
gs-3955	235	18	strategies	strategy	NOUN
gs-3955	235	19	of	of	ADP
gs-3955	235	20	earth	earth	NOUN
gs-3955	235	21	's	's	PART
gs-3955	235	22	most	most	ADV
gs-3955	235	23	enduring	endure	VERB
gs-3955	235	24	organisms	organism	NOUN
gs-3955	235	25	illuminate	illuminate	VERB
gs-3955	235	26	pathways	pathway	NOUN
gs-3955	235	27	to	to	PART
gs-3955	235	28	extend	extend	VERB
gs-3955	235	29	human	human	ADJ
gs-3955	235	30	vitality	vitality	NOUN
gs-3955	235	31	(	(	PUNCT
gs-3955	235	32	de	de	X
gs-3955	235	33	magalhães	magalhães	NOUN
gs-3955	235	34	,	,	PUNCT
gs-3955	235	35	2023	2023	NUM
gs-3955	235	36	;	;	PUNCT
gs-3955	235	37	buffenstein	buffenstein	ADV
gs-3955	235	38	,	,	PUNCT
gs-3955	235	39	2008	2008	NUM
gs-3955	235	40	)	)	PUNCT
gs-3955	235	41	.	.	PUNCT
gs-3955	236	1	the	the	DET
gs-3955	236	2	scientific	scientific	ADJ
gs-3955	236	3	journey	journey	NOUN
gs-3955	236	4	from	from	ADP
gs-3955	236	5	observing	observe	VERB
gs-3955	236	6	these	these	DET
gs-3955	236	7	biological	biological	ADJ
gs-3955	236	8	marvels	marvel	NOUN
gs-3955	236	9	to	to	PART
gs-3955	236	10	applying	apply	VERB
gs-3955	236	11	their	their	PRON
gs-3955	236	12	secrets	secret	NOUN
gs-3955	236	13	represents	represent	VERB
gs-3955	236	14	one	one	NUM
gs-3955	236	15	of	of	ADP
gs-3955	236	16	the	the	DET
gs-3955	236	17	most	most	ADV
gs-3955	236	18	profound	profound	ADJ
gs-3955	236	19	quests	quest	NOUN
gs-3955	236	20	in	in	ADP
gs-3955	236	21	modern	modern	ADJ
gs-3955	236	22	science	science	NOUN
gs-3955	236	23	—	—	PUNCT
gs-3955	236	24	a	a	DET
gs-3955	236	25	convergence	convergence	NOUN
gs-3955	236	26	of	of	ADP
gs-3955	236	27	evolutionary	evolutionary	ADJ
gs-3955	236	28	biology	biology	NOUN
gs-3955	236	29	,	,	PUNCT
gs-3955	236	30	molecular	molecular	ADJ
gs-3955	236	31	genetics	genetic	NOUN
gs-3955	236	32	,	,	PUNCT
gs-3955	236	33	and	and	CCONJ
gs-3955	236	34	systems	system	NOUN
gs-3955	236	35	medicine	medicine	NOUN
gs-3955	236	36	that	that	PRON
gs-3955	236	37	promises	promise	VERB
gs-3955	236	38	to	to	PART
gs-3955	236	39	redefine	redefine	VERB
gs-3955	236	40	what	what	PRON
gs-3955	236	41	it	it	PRON
gs-3955	236	42	means	mean	VERB
gs-3955	236	43	to	to	PART
gs-3955	236	44	grow	grow	VERB
gs-3955	236	45	older	old	ADJ
gs-3955	236	46	.	.	PUNCT
gs-3955	237	1	in	in	ADP
gs-3955	237	2	decoding	decode	VERB
gs-3955	237	3	how	how	SCONJ
gs-3955	237	4	a	a	DET
gs-3955	237	5	shark	shark	NOUN
gs-3955	237	6	's	's	PART
gs-3955	237	7	cartilage	cartilage	NOUN
gs-3955	237	8	resists	resist	VERB
gs-3955	237	9	centuries	century	NOUN
gs-3955	237	10	of	of	ADP
gs-3955	237	11	oxidative	oxidative	ADJ
gs-3955	237	12	stress	stress	NOUN
gs-3955	237	13	(	(	PUNCT
gs-3955	237	14	passow	passow	NOUN
gs-3955	237	15	et	et	PROPN
gs-3955	237	16	al	al	PROPN
gs-3955	237	17	.	.	PROPN
gs-3955	237	18	,	,	PUNCT
gs-3955	237	19	2022	2022	NUM
gs-3955	237	20	)	)	PUNCT
gs-3955	237	21	,	,	PUNCT
gs-3955	237	22	how	how	SCONJ
gs-3955	237	23	a	a	DET
gs-3955	237	24	hydra	hydra	NOUN
gs-3955	237	25	's	's	PART
gs-3955	237	26	stem	stem	NOUN
gs-3955	237	27	cells	cell	NOUN
gs-3955	237	28	remain	remain	VERB
gs-3955	237	29	perpetually	perpetually	ADV
gs-3955	237	30	youthful	youthful	ADJ
gs-3955	237	31	(	(	PUNCT
gs-3955	237	32	wagner	wagner	PROPN
gs-3955	237	33	et	et	PROPN
gs-3955	237	34	al	al	PROPN
gs-3955	237	35	.	.	PROPN
gs-3955	237	36	,	,	PUNCT
gs-3955	237	37	2011	2011	NUM
gs-3955	237	38	)	)	PUNCT
gs-3955	237	39	,	,	PUNCT
gs-3955	237	40	or	or	CCONJ
gs-3955	237	41	how	how	SCONJ
gs-3955	237	42	a	a	DET
gs-3955	237	43	naked	naked	ADJ
gs-3955	237	44	mole	mole	NOUN
gs-3955	237	45	-	-	PUNCT
gs-3955	237	46	rat	rat	NOUN
gs-3955	237	47	's	's	PART
gs-3955	237	48	proteins	protein	NOUN
gs-3955	237	49	defy	defy	VERB
gs-3955	237	50	structural	structural	ADJ
gs-3955	237	51	decay	decay	NOUN
gs-3955	237	52	(	(	PUNCT
gs-3955	237	53	perez	perez	PROPN
gs-3955	237	54	et	et	PROPN
gs-3955	237	55	al	al	PROPN
gs-3955	237	56	.	.	PROPN
gs-3955	237	57	,	,	PUNCT
gs-3955	237	58	2009	2009	NUM
gs-3955	237	59	)	)	PUNCT
gs-3955	237	60	,	,	PUNCT
gs-3955	237	61	we	we	PRON
gs-3955	237	62	gain	gain	VERB
gs-3955	237	63	not	not	PART
gs-3955	237	64	just	just	ADV
gs-3955	237	65	incremental	incremental	ADJ
gs-3955	237	66	knowledge	knowledge	NOUN
gs-3955	237	67	but	but	CCONJ
gs-3955	237	68	revolutionary	revolutionary	ADJ
gs-3955	237	69	insights	insight	NOUN
gs-3955	237	70	into	into	ADP
gs-3955	237	71	life	life	NOUN
gs-3955	237	72	's	's	PART
gs-3955	237	73	capacity	capacity	NOUN
gs-3955	237	74	to	to	PART
gs-3955	237	75	preserve	preserve	VERB
gs-3955	237	76	itself	itself	PRON
gs-3955	237	77	against	against	ADP
gs-3955	237	78	time	time	NOUN
gs-3955	237	79	's	's	PART
gs-3955	237	80	arrow	arrow	NOUN
gs-3955	237	81	.	.	PUNCT
gs-3955	238	1	these	these	DET
gs-3955	238	2	organisms	organism	NOUN
gs-3955	238	3	are	be	AUX
gs-3955	238	4	nature	nature	NOUN
gs-3955	238	5	's	's	PART
gs-3955	238	6	masterclass	masterclass	NOUN
gs-3955	238	7	in	in	ADP
gs-3955	238	8	durability	durability	NOUN
gs-3955	238	9	,	,	PUNCT
gs-3955	238	10	and	and	CCONJ
gs-3955	238	11	their	their	PRON
gs-3955	238	12	lessons	lesson	NOUN
gs-3955	238	13	may	may	AUX
gs-3955	238	14	ultimately	ultimately	ADV
gs-3955	238	15	liberate	liberate	VERB
gs-3955	238	16	humanity	humanity	NOUN
gs-3955	238	17	from	from	ADP
gs-3955	238	18	its	its	PRON
gs-3955	238	19	most	most	ADV
gs-3955	238	20	universal	universal	ADJ
gs-3955	238	21	affliction	affliction	NOUN
gs-3955	238	22	.	.	PUNCT
gs-3955	239	1	redefining	redefine	VERB
gs-3955	239	2	the	the	DET
gs-3955	239	3	boundaries	boundary	NOUN
gs-3955	239	4	of	of	ADP
gs-3955	239	5	biological	biological	ADJ
gs-3955	239	6	aging	aging	NOUN
gs-3955	239	7	through	through	ADP
gs-3955	239	8	negligibly	negligibly	ADV
gs-3955	239	9	senescent	senescent	ADJ
gs-3955	239	10	species	specie	NOUN
gs-3955	239	11	the	the	DET
gs-3955	239	12	phenomenon	phenomenon	NOUN
gs-3955	239	13	of	of	ADP
gs-3955	239	14	negligible	negligible	ADJ
gs-3955	239	15	senescence	senescence	NOUN
gs-3955	239	16	represents	represent	VERB
gs-3955	239	17	one	one	NUM
gs-3955	239	18	of	of	ADP
gs-3955	239	19	the	the	DET
gs-3955	239	20	most	most	ADV
gs-3955	239	21	profound	profound	ADJ
gs-3955	239	22	challenges	challenge	NOUN
gs-3955	239	23	to	to	ADP
gs-3955	239	24	fundamental	fundamental	ADJ
gs-3955	239	25	biological	biological	ADJ
gs-3955	239	26	paradigms	paradigm	NOUN
gs-3955	239	27	.	.	PUNCT
gs-3955	240	1	while	while	SCONJ
gs-3955	240	2	aging	age	VERB
gs-3955	240	3	—	—	PUNCT
gs-3955	240	4	characterized	characterize	VERB
gs-3955	240	5	by	by	ADP
gs-3955	240	6	progressive	progressive	ADJ
gs-3955	240	7	functional	functional	ADJ
gs-3955	240	8	decline	decline	NOUN
gs-3955	240	9	,	,	PUNCT
gs-3955	240	10	accumulation	accumulation	NOUN
gs-3955	240	11	of	of	ADP
gs-3955	240	12	molecular	molecular	ADJ
gs-3955	240	13	damage	damage	NOUN
gs-3955	240	14	,	,	PUNCT
gs-3955	240	15	and	and	CCONJ
gs-3955	240	16	increased	increase	VERB
gs-3955	240	17	mortality	mortality	NOUN
gs-3955	240	18	risk	risk	NOUN
gs-3955	240	19	—	—	PUNCT
gs-3955	240	20	manifests	manifest	NOUN
gs-3955	240	21	universally	universally	ADV
gs-3955	240	22	across	across	ADP
gs-3955	240	23	most	most	ADJ
gs-3955	240	24	multicellular	multicellular	ADJ
gs-3955	240	25	organisms	organism	NOUN
gs-3955	240	26	,	,	PUNCT
gs-3955	240	27	a	a	DET
gs-3955	240	28	select	select	ADJ
gs-3955	240	29	group	group	NOUN
gs-3955	240	30	of	of	ADP
gs-3955	240	31	species	specie	NOUN
gs-3955	240	32	defies	defy	VERB
gs-3955	240	33	this	this	DET
gs-3955	240	34	trajectory	trajectory	NOUN
gs-3955	240	35	.	.	PUNCT
gs-3955	241	1	these	these	DET
gs-3955	241	2	biological	biological	ADJ
gs-3955	241	3	outliers	outlier	NOUN
gs-3955	241	4	,	,	PUNCT
gs-3955	241	5	spanning	span	VERB
gs-3955	241	6	cnidarians	cnidarian	NOUN
gs-3955	241	7	,	,	PUNCT
gs-3955	241	8	elasmobranchs	elasmobranch	NOUN
gs-3955	241	9	,	,	PUNCT
gs-3955	241	10	rodents	rodent	NOUN
gs-3955	241	11	,	,	PUNCT
gs-3955	241	12	bivalves	bivalve	NOUN
gs-3955	241	13	,	,	PUNCT
gs-3955	241	14	and	and	CCONJ
gs-3955	241	15	amphibians	amphibian	NOUN
gs-3955	241	16	,	,	PUNCT
gs-3955	241	17	maintain	maintain	VERB
gs-3955	241	18	physiological	physiological	ADJ
gs-3955	241	19	integrity	integrity	NOUN
gs-3955	241	20	,	,	PUNCT
gs-3955	241	21	reproductive	reproductive	ADJ
gs-3955	241	22	capacity	capacity	NOUN
gs-3955	241	23	,	,	PUNCT
gs-3955	241	24	and	and	CCONJ
gs-3955	241	25	negligible	negligible	ADJ
gs-3955	241	26	georgian	georgian	ADJ
gs-3955	241	27	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	241	28	მეცნიერები	მეცნიერები	NOUN
gs-3955	241	29	ტ.	ტ.	VERB
gs-3955	241	30	7	7	NUM
gs-3955	241	31	n	n	PRON
gs-3955	241	32	3	3	NUM
gs-3955	241	33	,	,	PUNCT
gs-3955	241	34	2025	2025	NUM
gs-3955	241	35	371	371	NUM
gs-3955	241	36	mortality	mortality	NOUN
gs-3955	241	37	rate	rate	NOUN
gs-3955	241	38	increases	increase	NOUN
gs-3955	241	39	for	for	ADP
gs-3955	241	40	periods	period	NOUN
gs-3955	241	41	vastly	vastly	ADV
gs-3955	241	42	exceeding	exceed	VERB
gs-3955	241	43	related	related	ADJ
gs-3955	241	44	species	specie	NOUN
gs-3955	241	45	(	(	PUNCT
gs-3955	241	46	finch	finch	NOUN
gs-3955	241	47	,	,	PUNCT
gs-3955	241	48	2009	2009	NUM
gs-3955	241	49	;	;	PUNCT
gs-3955	241	50	buffenstein	buffenstein	ADV
gs-3955	241	51	,	,	PUNCT
gs-3955	241	52	2008	2008	NUM
gs-3955	241	53	;	;	PUNCT
gs-3955	241	54	de	de	X
gs-3955	241	55	magalhães	magalhães	NOUN
gs-3955	241	56	,	,	PUNCT
gs-3955	241	57	2023	2023	NUM
gs-3955	241	58	)	)	PUNCT
gs-3955	241	59	.	.	PUNCT
gs-3955	242	1	their	their	PRON
gs-3955	242	2	existence	existence	NOUN
gs-3955	242	3	compels	compel	VERB
gs-3955	242	4	a	a	DET
gs-3955	242	5	radical	radical	ADJ
gs-3955	242	6	reconceptualization	reconceptualization	NOUN
gs-3955	242	7	of	of	ADP
gs-3955	242	8	aging	aging	NOUN
gs-3955	242	9	not	not	PART
gs-3955	242	10	as	as	ADP
gs-3955	242	11	an	an	DET
gs-3955	242	12	inevitable	inevitable	ADJ
gs-3955	242	13	thermodynamic	thermodynamic	ADJ
gs-3955	242	14	destiny	destiny	NOUN
gs-3955	242	15	,	,	PUNCT
gs-3955	242	16	but	but	CCONJ
gs-3955	242	17	as	as	ADP
gs-3955	242	18	a	a	DET
gs-3955	242	19	malleable	malleable	ADJ
gs-3955	242	20	biological	biological	ADJ
gs-3955	242	21	process	process	NOUN
gs-3955	242	22	subject	subject	ADJ
gs-3955	242	23	to	to	ADP
gs-3955	242	24	evolutionary	evolutionary	ADJ
gs-3955	242	25	optimization	optimization	NOUN
gs-3955	242	26	(	(	PUNCT
gs-3955	242	27	lópez	lópez	NOUN
gs-3955	242	28	-	-	PUNCT
gs-3955	242	29	otín	otín	NOUN
gs-3955	242	30	et	et	PROPN
gs-3955	242	31	al	al	PROPN
gs-3955	242	32	.	.	PROPN
gs-3955	242	33	,	,	PUNCT
gs-3955	242	34	2013	2013	NUM
gs-3955	242	35	;	;	PUNCT
gs-3955	242	36	austad	austad	NOUN
gs-3955	242	37	,	,	PUNCT
gs-3955	242	38	2022	2022	NUM
gs-3955	242	39	)	)	PUNCT
gs-3955	242	40	.	.	PUNCT
gs-3955	243	1	this	this	DET
gs-3955	243	2	expanded	expand	VERB
gs-3955	243	3	discussion	discussion	NOUN
gs-3955	243	4	synthesizes	synthesize	VERB
gs-3955	243	5	molecular	molecular	ADJ
gs-3955	243	6	mechanisms	mechanism	NOUN
gs-3955	243	7	,	,	PUNCT
gs-3955	243	8	evolutionary	evolutionary	ADJ
gs-3955	243	9	drivers	driver	NOUN
gs-3955	243	10	,	,	PUNCT
gs-3955	243	11	systems	system	NOUN
gs-3955	243	12	-	-	PUNCT
gs-3955	243	13	level	level	NOUN
gs-3955	243	14	adaptations	adaptation	NOUN
gs-3955	243	15	,	,	PUNCT
gs-3955	243	16	and	and	CCONJ
gs-3955	243	17	translational	translational	ADJ
gs-3955	243	18	implications	implication	NOUN
gs-3955	243	19	revealed	reveal	VERB
gs-3955	243	20	by	by	ADP
gs-3955	243	21	these	these	DET
gs-3955	243	22	extraordinary	extraordinary	ADJ
gs-3955	243	23	organisms	organism	NOUN
gs-3955	243	24	,	,	PUNCT
gs-3955	243	25	incorporating	incorporate	VERB
gs-3955	243	26	novel	novel	ADJ
gs-3955	243	27	frameworks	framework	NOUN
gs-3955	243	28	and	and	CCONJ
gs-3955	243	29	recent	recent	ADJ
gs-3955	243	30	research	research	NOUN
gs-3955	243	31	frontiers	frontier	NOUN
gs-3955	243	32	.	.	PUNCT
gs-3955	244	1	thermodynamic	thermodynamic	ADJ
gs-3955	244	2	and	and	CCONJ
gs-3955	244	3	evolutionary	evolutionary	ADJ
gs-3955	244	4	foundations	foundation	NOUN
gs-3955	244	5	of	of	ADP
gs-3955	244	6	negligible	negligible	ADJ
gs-3955	244	7	senescence	senescence	NOUN
gs-3955	244	8	:	:	PUNCT
gs-3955	244	9	entropy	entropy	VERB
gs-3955	244	10	defiance	defiance	NOUN
gs-3955	244	11	in	in	ADP
gs-3955	244	12	biological	biological	ADJ
gs-3955	244	13	systems	system	NOUN
gs-3955	244	14	the	the	DET
gs-3955	244	15	second	second	ADJ
gs-3955	244	16	law	law	NOUN
gs-3955	244	17	of	of	ADP
gs-3955	244	18	thermodynamics	thermodynamic	NOUN
gs-3955	244	19	predicts	predict	VERB
gs-3955	244	20	inevitable	inevitable	ADJ
gs-3955	244	21	energy	energy	NOUN
gs-3955	244	22	dissipation	dissipation	NOUN
gs-3955	244	23	and	and	CCONJ
gs-3955	244	24	system	system	NOUN
gs-3955	244	25	disorder	disorder	NOUN
gs-3955	244	26	.	.	PUNCT
gs-3955	245	1	aging	age	VERB
gs-3955	245	2	aligns	align	VERB
gs-3955	245	3	with	with	ADP
gs-3955	245	4	this	this	DET
gs-3955	245	5	entropic	entropic	ADJ
gs-3955	245	6	progression	progression	NOUN
gs-3955	245	7	through	through	ADP
gs-3955	245	8	:	:	PUNCT
gs-3955	245	9	●	●	NUM
gs-3955	245	10	cumulative	cumulative	ADJ
gs-3955	245	11	dna	dna	NOUN
gs-3955	245	12	mutations	mutation	NOUN
gs-3955	245	13	(	(	PUNCT
gs-3955	245	14	1.7	1.7	NUM
gs-3955	245	15	×	×	NOUN
gs-3955	245	16	10⁻⁹	10⁻⁹	NUM
gs-3955	245	17	mutations	mutation	NOUN
gs-3955	245	18	/	/	SYM
gs-3955	245	19	base	base	NOUN
gs-3955	245	20	/	/	SYM
gs-3955	245	21	generation	generation	NOUN
gs-3955	245	22	in	in	ADP
gs-3955	245	23	humans	human	NOUN
gs-3955	245	24	)	)	PUNCT
gs-3955	245	25	(	(	PUNCT
gs-3955	245	26	tacutu	tacutu	NOUN
gs-3955	245	27	et	et	PROPN
gs-3955	245	28	al	al	PROPN
gs-3955	245	29	.	.	PROPN
gs-3955	245	30	,	,	PUNCT
gs-3955	245	31	2013	2013	NUM
gs-3955	245	32	)	)	PUNCT
gs-3955	245	33	●	●	NUM
gs-3955	245	34	protein	protein	NOUN
gs-3955	245	35	misfolding	misfolding	NOUN
gs-3955	245	36	(	(	PUNCT
gs-3955	245	37	amyloid	amyloid	NOUN
gs-3955	245	38	-	-	PUNCT
gs-3955	245	39	β	β	NOUN
gs-3955	245	40	accumulation	accumulation	NOUN
gs-3955	245	41	rate	rate	NOUN
gs-3955	245	42	:	:	PUNCT
gs-3955	245	43	0.5	0.5	NUM
gs-3955	245	44	-	-	SYM
gs-3955	245	45	2	2	NUM
gs-3955	245	46	%	%	NOUN
gs-3955	245	47	annually	annually	ADV
gs-3955	245	48	)	)	PUNCT
gs-3955	245	49	(	(	PUNCT
gs-3955	245	50	perez	perez	PROPN
gs-3955	245	51	et	et	PROPN
gs-3955	245	52	al	al	PROPN
gs-3955	245	53	.	.	PROPN
gs-3955	245	54	,	,	PUNCT
gs-3955	245	55	2009	2009	NUM
gs-3955	245	56	)	)	PUNCT
gs-3955	245	57	●	●	PUNCT
gs-3955	245	58	mitochondrial	mitochondrial	ADJ
gs-3955	245	59	membrane	membrane	NOUN
gs-3955	245	60	depolarization	depolarization	NOUN
gs-3955	245	61	(	(	PUNCT
gs-3955	245	62	1.5	1.5	NUM
gs-3955	245	63	%	%	NOUN
gs-3955	245	64	efficiency	efficiency	NOUN
gs-3955	245	65	loss	loss	NOUN
gs-3955	245	66	/	/	SYM
gs-3955	245	67	decade	decade	NOUN
gs-3955	245	68	)	)	PUNCT
gs-3955	245	69	(	(	PUNCT
gs-3955	245	70	abele	abele	PROPN
gs-3955	245	71	et	et	PROPN
gs-3955	245	72	al	al	PROPN
gs-3955	245	73	.	.	PROPN
gs-3955	245	74	,	,	PUNCT
gs-3955	245	75	2008	2008	NUM
gs-3955	245	76	)	)	PUNCT
gs-3955	245	77	●	●	PUNCT
gs-3955	245	78	epigenetic	epigenetic	ADJ
gs-3955	245	79	drift	drift	NOUN
gs-3955	245	80	(	(	PUNCT
gs-3955	245	81	5	5	NUM
gs-3955	245	82	-	-	SYM
gs-3955	245	83	15	15	NUM
gs-3955	245	84	%	%	NOUN
gs-3955	245	85	methylation	methylation	NOUN
gs-3955	245	86	variance	variance	NOUN
gs-3955	245	87	/	/	SYM
gs-3955	245	88	decade	decade	NOUN
gs-3955	245	89	)	)	PUNCT
gs-3955	245	90	(	(	PUNCT
gs-3955	245	91	horvath	horvath	PROPN
gs-3955	245	92	et	et	PROPN
gs-3955	245	93	al	al	PROPN
gs-3955	245	94	.	.	PROPN
gs-3955	245	95	,	,	PUNCT
gs-3955	245	96	2022	2022	NUM
gs-3955	245	97	)	)	PUNCT
gs-3955	245	98	negligibly	negligibly	ADV
gs-3955	245	99	senescent	senescent	ADJ
gs-3955	245	100	species	specie	NOUN
gs-3955	245	101	deploy	deploy	VERB
gs-3955	245	102	counter	counter	ADJ
gs-3955	245	103	-	-	ADJ
gs-3955	245	104	entropic	entropic	ADJ
gs-3955	245	105	strategies	strategy	NOUN
gs-3955	245	106	:	:	PUNCT
gs-3955	245	107	●	●	PUNCT
gs-3955	245	108	energy	energy	NOUN
gs-3955	245	109	redistribution	redistribution	NOUN
gs-3955	245	110	:	:	PUNCT
gs-3955	245	111	greenland	greenland	PROPN
gs-3955	245	112	sharks	sharks	PROPN
gs-3955	245	113	(	(	PUNCT
gs-3955	245	114	somniosus	somniosus	PROPN
gs-3955	245	115	microcephalus	microcephalus	PROPN
gs-3955	245	116	)	)	PUNCT
gs-3955	245	117	reduce	reduce	VERB
gs-3955	245	118	basal	basal	ADJ
gs-3955	245	119	metabolic	metabolic	NOUN
gs-3955	245	120	rates	rate	NOUN
gs-3955	245	121	to	to	ADP
gs-3955	245	122	17	17	NUM
gs-3955	245	123	-	-	SYM
gs-3955	245	124	21	21	NUM
gs-3955	245	125	µmol	µmol	ADP
gs-3955	245	126	o₂/kg	o₂/kg	PROPN
gs-3955	245	127	/	/	SYM
gs-3955	245	128	min	min	PROPN
gs-3955	245	129	(	(	PUNCT
gs-3955	245	130	vs.	vs.	X
gs-3955	245	131	150	150	NUM
gs-3955	245	132	-	-	SYM
gs-3955	245	133	300	300	NUM
gs-3955	245	134	in	in	ADP
gs-3955	245	135	teleosts	teleost	NOUN
gs-3955	245	136	)	)	PUNCT
gs-3955	245	137	,	,	PUNCT
gs-3955	245	138	minimizing	minimize	VERB
gs-3955	245	139	metabolic	metabolic	NOUN
gs-3955	245	140	byproducts	byproduct	NOUN
gs-3955	245	141	(	(	PUNCT
gs-3955	245	142	hansen	hansen	PROPN
gs-3955	245	143	et	et	PROPN
gs-3955	245	144	al	al	PROPN
gs-3955	245	145	.	.	PROPN
gs-3955	245	146	,	,	PUNCT
gs-3955	245	147	2021	2021	NUM
gs-3955	245	148	;	;	PUNCT
gs-3955	245	149	treberg	treberg	PROPN
gs-3955	245	150	&	&	CCONJ
gs-3955	245	151	speers	speer	NOUN
gs-3955	245	152	-	-	PUNCT
gs-3955	245	153	roesch	roesch	NOUN
gs-3955	245	154	,	,	PUNCT
gs-3955	245	155	2016	2016	NUM
gs-3955	245	156	)	)	PUNCT
gs-3955	245	157	●	●	NUM
gs-3955	245	158	structural	structural	ADJ
gs-3955	245	159	optimization	optimization	NOUN
gs-3955	245	160	:	:	PUNCT
gs-3955	246	1	arctica	arctica	PROPN
gs-3955	246	2	islandica	islandica	PROPN
gs-3955	246	3	incorporates	incorporate	VERB
gs-3955	246	4	tetradecanoic	tetradecanoic	ADJ
gs-3955	246	5	(	(	PUNCT
gs-3955	246	6	25	25	NUM
gs-3955	246	7	%	%	NOUN
gs-3955	246	8	)	)	PUNCT
gs-3955	246	9	and	and	CCONJ
gs-3955	246	10	hexadecenoic	hexadecenoic	ADJ
gs-3955	246	11	(	(	PUNCT
gs-3955	246	12	40	40	NUM
gs-3955	246	13	%	%	NOUN
gs-3955	246	14	)	)	PUNCT
gs-3955	246	15	fatty	fatty	NOUN
gs-3955	246	16	acids	acid	NOUN
gs-3955	246	17	into	into	ADP
gs-3955	246	18	mitochondrial	mitochondrial	ADJ
gs-3955	246	19	membranes	membrane	NOUN
gs-3955	246	20	,	,	PUNCT
gs-3955	246	21	reducing	reduce	VERB
gs-3955	246	22	peroxidation	peroxidation	NOUN
gs-3955	246	23	index	index	NOUN
gs-3955	246	24	to	to	ADP
gs-3955	246	25	35	35	NUM
gs-3955	246	26	(	(	PUNCT
gs-3955	246	27	vs.	vs.	X
gs-3955	246	28	85	85	NUM
gs-3955	246	29	-	-	SYM
gs-3955	246	30	110	110	NUM
gs-3955	246	31	in	in	ADP
gs-3955	246	32	short	short	ADJ
gs-3955	246	33	-	-	PUNCT
gs-3955	246	34	lived	live	VERB
gs-3955	246	35	bivalves	bivalve	NOUN
gs-3955	246	36	)	)	PUNCT
gs-3955	246	37	(	(	PUNCT
gs-3955	246	38	munro	munro	PROPN
gs-3955	246	39	&	&	CCONJ
gs-3955	246	40	blier	blier	PROPN
gs-3955	246	41	,	,	PUNCT
gs-3955	246	42	2012	2012	NUM
gs-3955	246	43	;	;	PUNCT
gs-3955	246	44	ballantyne	ballantyne	PROPN
gs-3955	246	45	et	et	PROPN
gs-3955	246	46	al	al	PROPN
gs-3955	246	47	.	.	PROPN
gs-3955	246	48	,	,	PUNCT
gs-3955	246	49	2021	2021	NUM
gs-3955	246	50	)	)	PUNCT
gs-3955	247	1	●	●	NUM
gs-3955	247	2	information	information	NOUN
gs-3955	247	3	preservation	preservation	NOUN
gs-3955	247	4	:	:	PUNCT
gs-3955	247	5	naked	naked	ADJ
gs-3955	247	6	mole	mole	NOUN
gs-3955	247	7	-	-	PUNCT
gs-3955	247	8	rats	rat	NOUN
gs-3955	247	9	(	(	PUNCT
gs-3955	247	10	heterocephalus	heterocephalus	ADP
gs-3955	247	11	glaber	glaber	NOUN
gs-3955	247	12	)	)	PUNCT
gs-3955	247	13	maintain	maintain	VERB
gs-3955	247	14	telomeric	telomeric	ADJ
gs-3955	247	15	reserves	reserve	NOUN
gs-3955	247	16	>	>	PUNCT
gs-3955	247	17	20	20	NUM
gs-3955	247	18	kb	kb	PROPN
gs-3955	247	19	through	through	ADP
gs-3955	247	20	constitutive	constitutive	ADJ
gs-3955	247	21	telomerase	telomerase	NOUN
gs-3955	247	22	expression	expression	NOUN
gs-3955	247	23	,	,	PUNCT
gs-3955	247	24	contrasting	contrast	VERB
gs-3955	247	25	with	with	ADP
gs-3955	247	26	humans	human	NOUN
gs-3955	247	27	'	'	PART
gs-3955	247	28	50	50	NUM
gs-3955	247	29	-	-	PUNCT
gs-3955	247	30	200	200	NUM
gs-3955	247	31	bp	bp	PROPN
gs-3955	247	32	/	/	SYM
gs-3955	247	33	year	year	NOUN
gs-3955	247	34	attrition	attrition	NOUN
gs-3955	247	35	(	(	PUNCT
gs-3955	247	36	azpurua	azpurua	PROPN
gs-3955	247	37	et	et	PROPN
gs-3955	247	38	al	al	PROPN
gs-3955	247	39	.	.	PROPN
gs-3955	247	40	,	,	PUNCT
gs-3955	247	41	2013	2013	NUM
gs-3955	247	42	;	;	PUNCT
gs-3955	247	43	macrae	macrae	PROPN
gs-3955	247	44	et	et	PROPN
gs-3955	247	45	al	al	PROPN
gs-3955	247	46	.	.	PROPN
gs-3955	247	47	,	,	PUNCT
gs-3955	247	48	2015	2015	NUM
gs-3955	247	49	)	)	PUNCT
gs-3955	247	50	georgian	georgian	ADJ
gs-3955	247	51	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	247	52	მეცნიერები	მეცნიერები	NOUN
gs-3955	247	53	ტ.	ტ.	VERB
gs-3955	247	54	7	7	NUM
gs-3955	247	55	n	n	DET
gs-3955	247	56	3	3	NUM
gs-3955	247	57	,	,	PUNCT
gs-3955	247	58	2025	2025	NUM
gs-3955	247	59	372	372	NUM
gs-3955	247	60	evolutionary	evolutionary	ADJ
gs-3955	247	61	ecology	ecology	NOUN
gs-3955	247	62	of	of	ADP
gs-3955	247	63	extreme	extreme	ADJ
gs-3955	247	64	longevity	longevity	NOUN
gs-3955	247	65	table	table	NOUN
gs-3955	247	66	1	1	NUM
gs-3955	247	67	.	.	PUNCT
gs-3955	247	68	negligible	negligible	ADJ
gs-3955	247	69	senescence	senescence	NOUN
gs-3955	247	70	emerges	emerge	VERB
gs-3955	247	71	under	under	ADP
gs-3955	247	72	specific	specific	ADJ
gs-3955	247	73	selective	selective	ADJ
gs-3955	247	74	pressures	pressure	NOUN
gs-3955	247	75	:	:	PUNCT
gs-3955	247	76	selective	selective	ADJ
gs-3955	247	77	pressure	pressure	NOUN
gs-3955	247	78	species	specie	NOUN
gs-3955	247	79	response	response	NOUN
gs-3955	247	80	example	example	NOUN
gs-3955	247	81	organism	organism	NOUN
gs-3955	247	82	lifespan	lifespan	NOUN
gs-3955	247	83	increase	increase	VERB
gs-3955	247	84	low	low	ADJ
gs-3955	247	85	predation	predation	NOUN
gs-3955	247	86	mortality(nielsen	mortality(nielsen	PROPN
gs-3955	247	87	et	et	PROPN
gs-3955	247	88	al	al	PROPN
gs-3955	247	89	.	.	PROPN
gs-3955	247	90	,	,	PUNCT
gs-3955	247	91	2016	2016	NUM
gs-3955	247	92	;	;	PUNCT
gs-3955	247	93	kyne	kyne	PROPN
gs-3955	247	94	et	et	PROPN
gs-3955	247	95	al	al	PROPN
gs-3955	247	96	.	.	PROPN
gs-3955	247	97	,	,	PUNCT
gs-3955	247	98	2019	2019	NUM
gs-3955	247	99	)	)	PUNCT
gs-3955	247	100	delayed	delay	VERB
gs-3955	247	101	maturation	maturation	NOUN
gs-3955	247	102	greenland	greenland	NOUN
gs-3955	247	103	shark	shark	NOUN
gs-3955	247	104	150	150	NUM
gs-3955	247	105	-	-	PUNCT
gs-3955	247	106	year	year	NOUN
gs-3955	247	107	maturation	maturation	NOUN
gs-3955	247	108	stable	stable	ADJ
gs-3955	247	109	niches	niche	NOUN
gs-3955	247	110	(	(	PUNCT
gs-3955	247	111	issartel	issartel	PROPN
gs-3955	247	112	et	et	PROPN
gs-3955	247	113	al	al	PROPN
gs-3955	247	114	.	.	PROPN
gs-3955	247	115	,	,	PUNCT
gs-3955	247	116	2009	2009	NUM
gs-3955	247	117	;	;	PUNCT
gs-3955	247	118	bulog	bulog	PROPN
gs-3955	247	119	et	et	PROPN
gs-3955	247	120	al	al	PROPN
gs-3955	247	121	.	.	PROPN
gs-3955	247	122	,	,	PUNCT
gs-3955	247	123	2020	2020	NUM
gs-3955	247	124	)	)	PUNCT
gs-3955	247	125	reduced	reduce	VERB
gs-3955	247	126	fecundity	fecundity	NOUN
gs-3955	247	127	olm	olm	NOUN
gs-3955	247	128	(	(	PUNCT
gs-3955	247	129	proteus	proteus	PROPN
gs-3955	247	130	anguinus	anguinus	PROPN
gs-3955	247	131	)	)	PUNCT
gs-3955	247	132	biennial	biennial	ADJ
gs-3955	247	133	reproduction	reproduction	NOUN
gs-3955	247	134	hypoxia	hypoxia	NOUN
gs-3955	247	135	/	/	SYM
gs-3955	247	136	stress	stress	NOUN
gs-3955	247	137	exposure	exposure	NOUN
gs-3955	247	138	(	(	PUNCT
gs-3955	247	139	buffenstein	buffenstein	ADV
gs-3955	247	140	,	,	PUNCT
gs-3955	247	141	2005	2005	NUM
gs-3955	247	142	;	;	PUNCT
gs-3955	247	143	dammann	dammann	PROPN
gs-3955	247	144	et	et	PROPN
gs-3955	247	145	al	al	PROPN
gs-3955	247	146	.	.	PROPN
gs-3955	247	147	,	,	PUNCT
gs-3955	247	148	2011	2011	NUM
gs-3955	247	149	)	)	PUNCT
gs-3955	247	150	enhanced	enhance	VERB
gs-3955	247	151	stress	stress	NOUN
gs-3955	247	152	resistance	resistance	NOUN
gs-3955	247	153	naked	naked	ADJ
gs-3955	247	154	mole	mole	ADJ
gs-3955	247	155	-	-	PUNCT
gs-3955	247	156	rat	rat	NOUN
gs-3955	247	157	30x	30x	NOUN
gs-3955	247	158	rodent	rodent	NOUN
gs-3955	247	159	norm	norm	NOUN
gs-3955	247	160	resource	resource	NOUN
gs-3955	247	161	scarcity	scarcity	NOUN
gs-3955	247	162	(	(	PUNCT
gs-3955	247	163	munro	munro	PROPN
gs-3955	247	164	&	&	CCONJ
gs-3955	247	165	blier	blier	PROPN
gs-3955	247	166	,	,	PUNCT
gs-3955	247	167	2012	2012	NUM
gs-3955	247	168	;	;	PUNCT
gs-3955	247	169	ballestaartero	ballestaartero	NOUN
gs-3955	247	170	et	et	PROPN
gs-3955	247	171	al	al	PROPN
gs-3955	247	172	.	.	PROPN
gs-3955	247	173	,	,	PUNCT
gs-3955	247	174	2017	2017	NUM
gs-3955	247	175	)	)	PUNCT
gs-3955	247	176	metabolic	metabolic	NOUN
gs-3955	247	177	depression	depression	NOUN
gs-3955	247	178	ocean	ocean	NOUN
gs-3955	247	179	quahog	quahog	PROPN
gs-3955	247	180	500	500	NUM
gs-3955	247	181	-	-	PUNCT
gs-3955	247	182	year	year	NOUN
gs-3955	247	183	anoxia	anoxia	NOUN
gs-3955	247	184	tolerance	tolerance	NOUN
gs-3955	247	185	the	the	DET
gs-3955	247	186	centrosomal	centrosomal	ADJ
gs-3955	247	187	theory	theory	NOUN
gs-3955	247	188	of	of	ADP
gs-3955	247	189	aging	age	VERB
gs-3955	247	190	(	(	PUNCT
gs-3955	247	191	tkemaladze	tkemaladze	NOUN
gs-3955	247	192	,	,	PUNCT
gs-3955	247	193	2023	2023	NUM
gs-3955	247	194	,	,	PUNCT
gs-3955	247	195	2024	2024	NUM
gs-3955	247	196	)	)	PUNCT
gs-3955	247	197	provides	provide	VERB
gs-3955	247	198	a	a	DET
gs-3955	247	199	unifying	unifying	ADJ
gs-3955	247	200	framework	framework	NOUN
gs-3955	247	201	:	:	PUNCT
gs-3955	247	202	asymmetric	asymmetric	ADJ
gs-3955	247	203	stem	stem	NOUN
gs-3955	247	204	cell	cell	NOUN
gs-3955	247	205	division	division	NOUN
gs-3955	247	206	distributes	distribute	VERB
gs-3955	247	207	"	"	PUNCT
gs-3955	247	208	aged	aged	ADJ
gs-3955	247	209	"	"	PUNCT
gs-3955	247	210	centrosomes	centrosome	NOUN
gs-3955	247	211	carrying	carry	VERB
gs-3955	247	212	cumulative	cumulative	ADJ
gs-3955	247	213	damage	damage	NOUN
gs-3955	247	214	.	.	PUNCT
gs-3955	248	1	species	specie	NOUN
gs-3955	248	2	like	like	ADP
gs-3955	248	3	planarians	planarian	NOUN
gs-3955	248	4	(	(	PUNCT
gs-3955	248	5	schmidtea	schmidtea	NOUN
gs-3955	248	6	mediterranea	mediterranea	PROPN
gs-3955	248	7	)	)	PUNCT
gs-3955	248	8	avoid	avoid	VERB
gs-3955	248	9	this	this	PRON
gs-3955	248	10	through	through	ADP
gs-3955	248	11	:	:	PUNCT
gs-3955	248	12	●	●	NUM
gs-3955	248	13	symmetric	symmetric	ADJ
gs-3955	248	14	neoblast	neoblast	NOUN
gs-3955	248	15	divisions	division	NOUN
gs-3955	248	16	(	(	PUNCT
gs-3955	248	17	wagner	wagner	PROPN
gs-3955	248	18	et	et	PROPN
gs-3955	248	19	al	al	PROPN
gs-3955	248	20	.	.	PROPN
gs-3955	248	21	,	,	PUNCT
gs-3955	248	22	2011	2011	NUM
gs-3955	248	23	;	;	PUNCT
gs-3955	248	24	rink	rink	NOUN
gs-3955	248	25	,	,	PUNCT
gs-3955	248	26	2013	2013	NUM
gs-3955	248	27	)	)	PUNCT
gs-3955	248	28	●	●	PUNCT
gs-3955	248	29	centrosomal	centrosomal	ADJ
gs-3955	248	30	rejuvenation	rejuvenation	NOUN
gs-3955	248	31	via	via	ADP
gs-3955	248	32	tubg1/2	tubg1/2	PROPN
gs-3955	248	33	recycling	recycling	NOUN
gs-3955	248	34	(	(	PUNCT
gs-3955	248	35	chichinadze	chichinadze	NOUN
gs-3955	248	36	et	et	PROPN
gs-3955	248	37	al	al	PROPN
gs-3955	248	38	.	.	PROPN
gs-3955	248	39	,	,	PUNCT
gs-3955	248	40	2013	2013	NUM
gs-3955	248	41	;	;	PUNCT
gs-3955	248	42	tkemaladze	tkemaladze	NOUN
gs-3955	248	43	,	,	PUNCT
gs-3955	248	44	2023	2023	NUM
gs-3955	248	45	)	)	PUNCT
gs-3955	248	46	●	●	NUM
gs-3955	248	47	selective	selective	ADJ
gs-3955	248	48	autophagy	autophagy	NOUN
gs-3955	248	49	of	of	ADP
gs-3955	248	50	damaged	damage	VERB
gs-3955	248	51	centrioles	centriole	NOUN
gs-3955	248	52	(	(	PUNCT
gs-3955	248	53	cep164	cep164	NOUN
gs-3955	248	54	degradation	degradation	NOUN
gs-3955	248	55	)	)	PUNCT
gs-3955	248	56	(	(	PUNCT
gs-3955	248	57	pirotte	pirotte	NOUN
gs-3955	248	58	et	et	PROPN
gs-3955	248	59	al	al	PROPN
gs-3955	248	60	.	.	PROPN
gs-3955	248	61	,	,	PUNCT
gs-3955	248	62	2015	2015	NUM
gs-3955	248	63	;	;	PUNCT
gs-3955	248	64	tkemaladze	tkemaladze	NOUN
gs-3955	248	65	,	,	PUNCT
gs-3955	248	66	2023	2023	NUM
gs-3955	248	67	)	)	PUNCT
gs-3955	248	68	georgian	georgian	ADJ
gs-3955	248	69	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	248	70	მეცნიერები	მეცნიერები	NOUN
gs-3955	248	71	ტ.	ტ.	VERB
gs-3955	248	72	7	7	NUM
gs-3955	248	73	n	n	DET
gs-3955	248	74	3	3	NUM
gs-3955	248	75	,	,	PUNCT
gs-3955	248	76	2025	2025	NUM
gs-3955	248	77	373	373	NUM
gs-3955	248	78	beyond	beyond	ADP
gs-3955	248	79	canonical	canonical	ADJ
gs-3955	248	80	pathways	pathway	NOUN
gs-3955	248	81	genomic	genomic	NOUN
gs-3955	248	82	guardianship	guardianship	NOUN
gs-3955	248	83	systems	system	NOUN
gs-3955	248	84	table	table	NOUN
gs-3955	248	85	2	2	NUM
gs-3955	248	86	:	:	PUNCT
gs-3955	248	87	dna	dna	NOUN
gs-3955	248	88	repair	repair	NOUN
gs-3955	248	89	mechanisms	mechanism	NOUN
gs-3955	248	90	in	in	ADP
gs-3955	248	91	long	long	ADV
gs-3955	248	92	-	-	PUNCT
gs-3955	248	93	lived	live	VERB
gs-3955	248	94	species	species	NOUN
gs-3955	248	95	mechanism	mechanism	NOUN
gs-3955	248	96	species	species	NOUN
gs-3955	248	97	enhancement	enhancement	NOUN
gs-3955	248	98	factor	factor	NOUN
gs-3955	248	99	key	key	ADJ
gs-3955	248	100	regulators	regulator	NOUN
gs-3955	248	101	nucleotide	nucleotide	ADJ
gs-3955	248	102	excision	excision	NOUN
gs-3955	248	103	repair	repair	NOUN
gs-3955	248	104	(	(	PUNCT
gs-3955	248	105	sussarellu	sussarellu	NOUN
gs-3955	248	106	et	et	NOUN
gs-3955	248	107	al	al	PROPN
gs-3955	248	108	.	.	PROPN
gs-3955	248	109	,	,	PUNCT
gs-3955	248	110	2020	2020	NUM
gs-3955	248	111	;	;	PUNCT
gs-3955	248	112	philipp	philipp	PROPN
gs-3955	248	113	et	et	PROPN
gs-3955	248	114	al	al	PROPN
gs-3955	248	115	.	.	PROPN
gs-3955	248	116	,	,	PUNCT
gs-3955	248	117	2012	2012	NUM
gs-3955	248	118	)	)	PUNCT
gs-3955	249	1	arctica	arctica	PROPN
gs-3955	249	2	islandica	islandica	PROPN
gs-3955	249	3	4.7x	4.7x	PROPN
gs-3955	249	4	xpa	xpa	PROPN
gs-3955	249	5	-	-	PUNCT
gs-3955	249	6	xpg	xpg	PROPN
gs-3955	249	7	complex	complex	NOUN
gs-3955	249	8	,	,	PUNCT
gs-3955	249	9	csa	csa	PROPN
gs-3955	249	10	/	/	SYM
gs-3955	249	11	csb	csb	PROPN
gs-3955	249	12	homologous	homologous	ADJ
gs-3955	249	13	recombination	recombination	NOUN
gs-3955	249	14	(	(	PUNCT
gs-3955	249	15	macrae	macrae	NOUN
gs-3955	249	16	et	et	PROPN
gs-3955	249	17	al	al	PROPN
gs-3955	249	18	.	.	PROPN
gs-3955	249	19	,	,	PUNCT
gs-3955	249	20	2015	2015	NUM
gs-3955	249	21	;	;	PUNCT
gs-3955	249	22	yang	yang	PROPN
gs-3955	249	23	et	et	PROPN
gs-3955	249	24	al	al	PROPN
gs-3955	249	25	.	.	PROPN
gs-3955	249	26	,	,	PUNCT
gs-3955	249	27	2019	2019	NUM
gs-3955	249	28	)	)	PUNCT
gs-3955	249	29	naked	naked	ADJ
gs-3955	249	30	molerat	molerat	NOUN
gs-3955	249	31	3.2x	3.2x	NUM
gs-3955	249	32	rad51	rad51	PROPN
gs-3955	249	33	,	,	PUNCT
gs-3955	249	34	brca2	brca2	PROPN
gs-3955	249	35	,	,	PUNCT
gs-3955	249	36	palb2	palb2	NOUN
gs-3955	249	37	translesion	translesion	NOUN
gs-3955	249	38	synthesis	synthesis	NOUN
gs-3955	249	39	(	(	PUNCT
gs-3955	249	40	seim	seim	PROPN
gs-3955	249	41	et	et	PROPN
gs-3955	249	42	al	al	PROPN
gs-3955	249	43	.	.	PROPN
gs-3955	249	44	,	,	PUNCT
gs-3955	249	45	2023	2023	NUM
gs-3955	249	46	)	)	PUNCT
gs-3955	249	47	greenland	greenland	NOUN
gs-3955	249	48	shark	shark	NOUN
gs-3955	249	49	2.8x	2.8x	NUM
gs-3955	249	50	rev1	rev1	NOUN
gs-3955	249	51	,	,	PUNCT
gs-3955	249	52	polh	polh	NOUN
gs-3955	249	53	,	,	PUNCT
gs-3955	249	54	polk	polk	NOUN
gs-3955	249	55	pirna	pirna	NOUN
gs-3955	249	56	silencing	silence	VERB
gs-3955	249	57	(	(	PUNCT
gs-3955	249	58	hyde	hyde	PROPN
gs-3955	249	59	et	et	PROPN
gs-3955	249	60	al	al	PROPN
gs-3955	249	61	.	.	PROPN
gs-3955	249	62	,	,	PUNCT
gs-3955	249	63	2020	2020	NUM
gs-3955	249	64	;	;	PUNCT
gs-3955	249	65	bowen	bowen	PROPN
gs-3955	249	66	et	et	PROPN
gs-3955	249	67	al	al	PROPN
gs-3955	249	68	.	.	PROPN
gs-3955	249	69	,	,	PUNCT
gs-3955	249	70	2021	2021	NUM
gs-3955	249	71	)	)	PUNCT
gs-3955	249	72	sebastes	sebaste	VERB
gs-3955	249	73	aleutianus	aleutianus	NOUN
gs-3955	249	74	5.1x	5.1x	PROPN
gs-3955	249	75	piwil2	piwil2	PROPN
gs-3955	249	76	,	,	PUNCT
gs-3955	249	77	tdrd9	tdrd9	PROPN
gs-3955	249	78	,	,	PUNCT
gs-3955	249	79	mov10l1	mov10l1	AUX
gs-3955	249	80	novel	novel	ADJ
gs-3955	249	81	findings	finding	NOUN
gs-3955	249	82	reveal	reveal	VERB
gs-3955	249	83	:	:	PUNCT
gs-3955	249	84	●	●	NUM
gs-3955	249	85	shark	shark	NOUN
gs-3955	249	86	-	-	PUNCT
gs-3955	249	87	specific	specific	ADJ
gs-3955	249	88	dna	dna	NOUN
gs-3955	249	89	polymerases	polymerase	NOUN
gs-3955	249	90	(	(	PUNCT
gs-3955	249	91	polq	polq	PRON
gs-3955	249	92	-	-	PUNCT
gs-3955	249	93	κ	κ	NOUN
gs-3955	249	94	variants	variant	NOUN
gs-3955	249	95	)	)	PUNCT
gs-3955	249	96	with	with	ADP
gs-3955	249	97	99.8	99.8	NUM
gs-3955	249	98	%	%	NOUN
gs-3955	249	99	fidelity	fidelity	NOUN
gs-3955	249	100	during	during	ADP
gs-3955	249	101	oxidative	oxidative	ADJ
gs-3955	249	102	stress	stress	NOUN
gs-3955	249	103	(	(	PUNCT
gs-3955	249	104	seim	seim	PROPN
gs-3955	249	105	et	et	PROPN
gs-3955	249	106	al	al	PROPN
gs-3955	249	107	.	.	PROPN
gs-3955	249	108	,	,	PUNCT
gs-3955	249	109	2023	2023	NUM
gs-3955	249	110	)	)	PUNCT
gs-3955	249	111	●	●	PUNCT
gs-3955	249	112	mole	mole	NOUN
gs-3955	249	113	-	-	PUNCT
gs-3955	249	114	rat	rat	NOUN
gs-3955	249	115	histone	histone	NOUN
gs-3955	249	116	variants	variant	NOUN
gs-3955	249	117	(	(	PUNCT
gs-3955	249	118	h2a.z.7	h2a.z.7	ADV
gs-3955	249	119	)	)	PUNCT
gs-3955	249	120	promoting	promote	VERB
gs-3955	249	121	chromatin	chromatin	NOUN
gs-3955	249	122	relaxation	relaxation	NOUN
gs-3955	249	123	at	at	ADP
gs-3955	249	124	damage	damage	NOUN
gs-3955	249	125	sites	site	NOUN
gs-3955	249	126	(	(	PUNCT
gs-3955	249	127	yang	yang	PROPN
gs-3955	249	128	et	et	PROPN
gs-3955	249	129	al	al	PROPN
gs-3955	249	130	.	.	PROPN
gs-3955	249	131	,	,	PUNCT
gs-3955	249	132	2019	2019	NUM
gs-3955	249	133	;	;	PUNCT
gs-3955	249	134	macrae	macrae	PROPN
gs-3955	249	135	et	et	PROPN
gs-3955	249	136	al	al	PROPN
gs-3955	249	137	.	.	PROPN
gs-3955	249	138	,	,	PUNCT
gs-3955	249	139	2015	2015	NUM
gs-3955	249	140	)	)	PUNCT
gs-3955	249	141	●	●	PUNCT
gs-3955	249	142	bivalve	bivalve	ADJ
gs-3955	249	143	error	error	NOUN
gs-3955	249	144	-	-	PUNCT
gs-3955	249	145	free	free	ADJ
gs-3955	249	146	translesion	translesion	NOUN
gs-3955	249	147	synthesis	synthesis	NOUN
gs-3955	249	148	via	via	ADP
gs-3955	249	149	rev3l	rev3l	NOUN
gs-3955	249	150	isoforms	isoform	NOUN
gs-3955	249	151	lacking	lack	VERB
gs-3955	249	152	exon	exon	NOUN
gs-3955	249	153	12	12	NUM
gs-3955	249	154	(	(	PUNCT
gs-3955	249	155	sussarellu	sussarellu	NOUN
gs-3955	249	156	et	et	NOUN
gs-3955	249	157	al	al	PROPN
gs-3955	249	158	.	.	PROPN
gs-3955	249	159	,	,	PUNCT
gs-3955	249	160	2020	2020	NUM
gs-3955	249	161	)	)	PUNCT
gs-3955	249	162	the	the	DET
gs-3955	249	163	oxidative	oxidative	ADJ
gs-3955	249	164	stress	stress	NOUN
gs-3955	249	165	paradox	paradox	NOUN
gs-3955	249	166	contrary	contrary	ADJ
gs-3955	249	167	to	to	ADP
gs-3955	249	168	the	the	DET
gs-3955	249	169	free	free	ADJ
gs-3955	249	170	radical	radical	ADJ
gs-3955	249	171	theory	theory	NOUN
gs-3955	249	172	,	,	PUNCT
gs-3955	249	173	several	several	ADJ
gs-3955	249	174	long	long	ADV
gs-3955	249	175	-	-	PUNCT
gs-3955	249	176	lived	live	VERB
gs-3955	249	177	species	specie	NOUN
gs-3955	249	178	exhibit	exhibit	VERB
gs-3955	249	179	elevated	elevate	VERB
gs-3955	249	180	ros	ros	PROPN
gs-3955	249	181	production	production	NOUN
gs-3955	249	182	:	:	PUNCT
gs-3955	249	183	georgian	georgian	ADJ
gs-3955	249	184	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	249	185	მეცნიერები	მეცნიერები	NOUN
gs-3955	249	186	ტ.	ტ.	VERB
gs-3955	249	187	7	7	NUM
gs-3955	249	188	n	n	DET
gs-3955	249	189	3	3	NUM
gs-3955	249	190	,	,	PUNCT
gs-3955	249	191	2025	2025	NUM
gs-3955	249	192	374	374	NUM
gs-3955	249	193	●	●	NUM
gs-3955	249	194	naked	naked	ADJ
gs-3955	249	195	mole	mole	ADJ
gs-3955	249	196	-	-	PUNCT
gs-3955	249	197	rat	rat	NOUN
gs-3955	249	198	myocardium	myocardium	NOUN
gs-3955	249	199	:	:	PUNCT
gs-3955	249	200	2.3	2.3	NUM
gs-3955	249	201	nmol	nmol	NOUN
gs-3955	249	202	h₂o₂/min	h₂o₂/min	NOUN
gs-3955	249	203	/	/	SYM
gs-3955	249	204	mg	mg	PROPN
gs-3955	249	205	protein	protein	NOUN
gs-3955	249	206	(	(	PUNCT
gs-3955	249	207	vs.	vs.	X
gs-3955	249	208	0.8	0.8	NUM
gs-3955	249	209	in	in	ADP
gs-3955	249	210	mice	mouse	NOUN
gs-3955	249	211	)	)	PUNCT
gs-3955	249	212	(	(	PUNCT
gs-3955	249	213	lewis	lewis	PROPN
gs-3955	249	214	et	et	PROPN
gs-3955	249	215	al	al	PROPN
gs-3955	249	216	.	.	PROPN
gs-3955	249	217	,	,	PUNCT
gs-3955	249	218	2018	2018	NUM
gs-3955	249	219	;	;	PUNCT
gs-3955	250	1	perez	perez	PROPN
gs-3955	250	2	et	et	PROPN
gs-3955	250	3	al	al	PROPN
gs-3955	250	4	.	.	PROPN
gs-3955	250	5	,	,	PUNCT
gs-3955	250	6	2009	2009	NUM
gs-3955	250	7	)	)	PUNCT
gs-3955	251	1	●	●	NUM
gs-3955	251	2	olm	olm	NOUN
gs-3955	251	3	neural	neural	ADJ
gs-3955	251	4	tissue	tissue	NOUN
gs-3955	251	5	:	:	PUNCT
gs-3955	251	6	superoxide	superoxide	NOUN
gs-3955	251	7	at	at	ADP
gs-3955	251	8	15.7	15.7	NUM
gs-3955	251	9	µm	µm	NOUN
gs-3955	251	10	(	(	PUNCT
gs-3955	251	11	vs.	vs.	ADP
gs-3955	251	12	8.2	8.2	NUM
gs-3955	251	13	µm	µm	NOUN
gs-3955	251	14	in	in	ADP
gs-3955	251	15	newts	newts	PROPN
gs-3955	251	16	)	)	PUNCT
gs-3955	251	17	(	(	PUNCT
gs-3955	251	18	goricar	goricar	NOUN
gs-3955	251	19	et	et	PROPN
gs-3955	251	20	al	al	PROPN
gs-3955	251	21	.	.	PROPN
gs-3955	251	22	,	,	PUNCT
gs-3955	251	23	2021	2021	NUM
gs-3955	251	24	)	)	PUNCT
gs-3955	251	25	their	their	PRON
gs-3955	251	26	resilience	resilience	NOUN
gs-3955	251	27	derives	derive	VERB
gs-3955	251	28	from	from	ADP
gs-3955	251	29	:	:	PUNCT
gs-3955	251	30	●	●	NUM
gs-3955	251	31	structural	structural	ADJ
gs-3955	251	32	resilience	resilience	NOUN
gs-3955	251	33	:	:	PUNCT
gs-3955	251	34	cardiolipin	cardiolipin	NOUN
gs-3955	251	35	acyl	acyl	NOUN
gs-3955	251	36	chains	chain	NOUN
gs-3955	251	37	in	in	ADP
gs-3955	251	38	mole	mole	NOUN
gs-3955	251	39	-	-	PUNCT
gs-3955	251	40	rats	rat	NOUN
gs-3955	251	41	contain	contain	VERB
gs-3955	251	42	82	82	NUM
gs-3955	251	43	%	%	NOUN
gs-3955	251	44	monounsaturates	monounsaturate	NOUN
gs-3955	251	45	(	(	PUNCT
gs-3955	251	46	vs.	vs.	X
gs-3955	251	47	43	43	NUM
gs-3955	251	48	%	%	NOUN
gs-3955	251	49	polyunsaturates	polyunsaturate	NOUN
gs-3955	251	50	in	in	ADP
gs-3955	251	51	mice	mouse	NOUN
gs-3955	251	52	)	)	PUNCT
gs-3955	251	53	,	,	PUNCT
gs-3955	251	54	reducing	reduce	VERB
gs-3955	251	55	peroxidation	peroxidation	NOUN
gs-3955	251	56	susceptibility	susceptibility	NOUN
gs-3955	251	57	(	(	PUNCT
gs-3955	251	58	perez	perez	PROPN
gs-3955	251	59	et	et	PROPN
gs-3955	251	60	al	al	PROPN
gs-3955	251	61	.	.	PROPN
gs-3955	251	62	,	,	PUNCT
gs-3955	251	63	2009	2009	NUM
gs-3955	251	64	)	)	PUNCT
gs-3955	251	65	●	●	NUM
gs-3955	251	66	redox	redox	ADJ
gs-3955	251	67	buffering	buffering	NOUN
gs-3955	251	68	:	:	PUNCT
gs-3955	251	69	giant	giant	ADJ
gs-3955	251	70	red	red	PROPN
gs-3955	251	71	sea	sea	NOUN
gs-3955	251	72	urchins	urchin	NOUN
gs-3955	251	73	(	(	PUNCT
gs-3955	251	74	strongylocentrotus	strongylocentrotus	NOUN
gs-3955	251	75	franciscanus	franciscanu	NOUN
gs-3955	251	76	)	)	PUNCT
gs-3955	251	77	maintain	maintain	VERB
gs-3955	251	78	glutathione	glutathione	NOUN
gs-3955	251	79	at	at	ADP
gs-3955	251	80	12.3	12.3	NUM
gs-3955	251	81	mm	mm	NOUN
gs-3955	251	82	(	(	PUNCT
gs-3955	251	83	vs.	vs.	ADP
gs-3955	251	84	2.1	2.1	NUM
gs-3955	251	85	mm	mm	NOUN
gs-3955	251	86	in	in	ADP
gs-3955	251	87	short	short	ADJ
gs-3955	251	88	-	-	PUNCT
gs-3955	251	89	spined	spin	VERB
gs-3955	251	90	urchins	urchin	NOUN
gs-3955	251	91	)	)	PUNCT
gs-3955	251	92	(	(	PUNCT
gs-3955	251	93	francis	francis	PROPN
gs-3955	251	94	et	et	PROPN
gs-3955	251	95	al	al	PROPN
gs-3955	251	96	.	.	PROPN
gs-3955	251	97	,	,	PUNCT
gs-3955	251	98	2019	2019	NUM
gs-3955	251	99	)	)	PUNCT
gs-3955	251	100	●	●	NUM
gs-3955	251	101	damage	damage	NOUN
gs-3955	251	102	decoupling	decoupling	NOUN
gs-3955	251	103	:	:	PUNCT
gs-3955	251	104	hydra	hydra	PROPN
gs-3955	251	105	expresses	express	VERB
gs-3955	251	106	novel	novel	ADJ
gs-3955	251	107	carbonyl	carbonyl	NOUN
gs-3955	251	108	reductases	reductase	NOUN
gs-3955	251	109	(	(	PUNCT
gs-3955	251	110	hvcr1	hvcr1	NOUN
gs-3955	251	111	-	-	PUNCT
gs-3955	251	112	4	4	NUM
gs-3955	251	113	)	)	PUNCT
gs-3955	251	114	that	that	PRON
gs-3955	251	115	detoxify	detoxify	VERB
gs-3955	251	116	4	4	NUM
gs-3955	251	117	-	-	PUNCT
gs-3955	251	118	hydroxynonenal	hydroxynonenal	NOUN
gs-3955	251	119	at	at	ADP
gs-3955	251	120	340	340	NUM
gs-3955	251	121	nmol	nmol	NOUN
gs-3955	251	122	/	/	SYM
gs-3955	251	123	min	min	NOUN
gs-3955	251	124	/	/	SYM
gs-3955	251	125	mg	mg	ADJ
gs-3955	251	126	protein	protein	NOUN
gs-3955	251	127	(	(	PUNCT
gs-3955	251	128	abele	abele	PROPN
gs-3955	251	129	et	et	PROPN
gs-3955	251	130	al	al	PROPN
gs-3955	251	131	.	.	PROPN
gs-3955	251	132	,	,	PUNCT
gs-3955	251	133	2008	2008	NUM
gs-3955	251	134	;	;	PUNCT
gs-3955	251	135	elliott	elliott	PROPN
gs-3955	251	136	&	&	CCONJ
gs-3955	251	137	sánchez	sánchez	PROPN
gs-3955	251	138	alvarado	alvarado	PROPN
gs-3955	251	139	,	,	PUNCT
gs-3955	251	140	2013	2013	NUM
gs-3955	251	141	)	)	PUNCT
gs-3955	251	142	proteostatic	proteostatic	NOUN
gs-3955	251	143	excellence	excellence	NOUN
gs-3955	251	144	long	long	ADV
gs-3955	251	145	-	-	PUNCT
gs-3955	251	146	lived	live	VERB
gs-3955	251	147	species	species	NOUN
gs-3955	251	148	exhibit	exhibit	VERB
gs-3955	251	149	three	three	NUM
gs-3955	251	150	-	-	PUNCT
gs-3955	251	151	tiered	tiere	VERB
gs-3955	251	152	protein	protein	NOUN
gs-3955	251	153	maintenance	maintenance	NOUN
gs-3955	251	154	:	:	PUNCT
gs-3955	252	1	folding	folding	NOUN
gs-3955	252	2	:	:	PUNCT
gs-3955	252	3	enhanced	enhanced	ADJ
gs-3955	252	4	chaperonin	chaperonin	PROPN
gs-3955	252	5	systems	system	NOUN
gs-3955	252	6	(	(	PUNCT
gs-3955	252	7	cct	cct	NOUN
gs-3955	252	8	-	-	PUNCT
gs-3955	252	9	ζ	ζ	NOUN
gs-3955	252	10	in	in	ADP
gs-3955	252	11	sharks	shark	NOUN
gs-3955	252	12	with	with	ADP
gs-3955	252	13	atpase	atpase	PROPN
gs-3955	252	14	efficiency	efficiency	NOUN
gs-3955	252	15	1.8×	1.8×	NUM
gs-3955	252	16	mammalian	mammalian	NOUN
gs-3955	252	17	)	)	PUNCT
gs-3955	252	18	(	(	PUNCT
gs-3955	252	19	ballantyne	ballantyne	PROPN
gs-3955	252	20	et	et	PROPN
gs-3955	252	21	al	al	PROPN
gs-3955	252	22	.	.	PROPN
gs-3955	252	23	,	,	PUNCT
gs-3955	252	24	2021	2021	NUM
gs-3955	252	25	)	)	PUNCT
gs-3955	252	26	surveillance	surveillance	NOUN
gs-3955	252	27	:	:	PUNCT
gs-3955	252	28	ubiquitin	ubiquitin	ADJ
gs-3955	252	29	-	-	PUNCT
gs-3955	252	30	independent	independent	ADJ
gs-3955	252	31	degradation	degradation	NOUN
gs-3955	252	32	via	via	ADP
gs-3955	252	33	20s	20	NOUN
gs-3955	252	34	proteasomes	proteasome	NOUN
gs-3955	252	35	(	(	PUNCT
gs-3955	252	36	constitutive	constitutive	ADJ
gs-3955	252	37	activation	activation	NOUN
gs-3955	252	38	in	in	ADP
gs-3955	252	39	hydra	hydra	PROPN
gs-3955	252	40	)	)	PUNCT
gs-3955	252	41	(	(	PUNCT
gs-3955	252	42	elliott	elliott	PROPN
gs-3955	252	43	&	&	CCONJ
gs-3955	252	44	sánchez	sánchez	PROPN
gs-3955	252	45	alvarado	alvarado	PROPN
gs-3955	252	46	,	,	PUNCT
gs-3955	252	47	2013	2013	NUM
gs-3955	252	48	)	)	PUNCT
gs-3955	252	49	clearance	clearance	NOUN
gs-3955	252	50	:	:	PUNCT
gs-3955	252	51	selective	selective	ADJ
gs-3955	252	52	autophagy	autophagy	NOUN
gs-3955	252	53	:	:	PUNCT
gs-3955	252	54	●	●	PUNCT
gs-3955	252	55	mitophagy	mitophagy	NOUN
gs-3955	252	56	:	:	PUNCT
gs-3955	252	57	pink1	pink1	NOUN
gs-3955	252	58	/	/	SYM
gs-3955	252	59	parkin	parkin	NOUN
gs-3955	252	60	-	-	PUNCT
gs-3955	252	61	independent	independent	ADJ
gs-3955	252	62	pathways	pathway	NOUN
gs-3955	252	63	in	in	ADP
gs-3955	252	64	naked	naked	ADJ
gs-3955	252	65	mole	mole	NOUN
gs-3955	252	66	-	-	PUNCT
gs-3955	252	67	rats	rat	NOUN
gs-3955	252	68	(	(	PUNCT
gs-3955	252	69	buffenstein	buffenstein	ADV
gs-3955	252	70	,	,	PUNCT
gs-3955	252	71	2008	2008	NUM
gs-3955	252	72	;	;	PUNCT
gs-3955	252	73	lewis	lewis	PROPN
gs-3955	252	74	et	et	PROPN
gs-3955	252	75	al	al	PROPN
gs-3955	252	76	.	.	PROPN
gs-3955	252	77	,	,	PUNCT
gs-3955	252	78	2018	2018	NUM
gs-3955	252	79	)	)	PUNCT
gs-3955	253	1	●	●	PUNCT
gs-3955	253	2	aggrephagy	aggrephagy	ADJ
gs-3955	253	3	:	:	PUNCT
gs-3955	253	4	p62	p62	NOUN
gs-3955	253	5	/	/	SYM
gs-3955	253	6	sqstm1	sqstm1	NOUN
gs-3955	253	7	variants	variant	NOUN
gs-3955	253	8	with	with	ADP
gs-3955	253	9	expanded	expand	VERB
gs-3955	253	10	lir	lir	NOUN
gs-3955	253	11	domains	domain	NOUN
gs-3955	253	12	in	in	ADP
gs-3955	253	13	rockfish	rockfish	PROPN
gs-3955	253	14	(	(	PUNCT
gs-3955	253	15	hamel	hamel	PROPN
gs-3955	253	16	et	et	PROPN
gs-3955	253	17	al	al	PROPN
gs-3955	253	18	.	.	PROPN
gs-3955	253	19	,	,	PUNCT
gs-3955	253	20	2020	2020	NUM
gs-3955	253	21	)	)	PUNCT
gs-3955	254	1	arctica	arctica	PROPN
gs-3955	254	2	islandica	islandica	PROPN
gs-3955	254	3	achieves	achieve	VERB
gs-3955	254	4	near	near	ADV
gs-3955	254	5	-	-	PUNCT
gs-3955	254	6	perfect	perfect	ADJ
gs-3955	254	7	proteostasis	proteostasis	NOUN
gs-3955	254	8	through	through	ADP
gs-3955	254	9	:	:	PUNCT
gs-3955	254	10	●	●	PUNCT
gs-3955	254	11	temperature	temperature	NOUN
gs-3955	254	12	-	-	PUNCT
gs-3955	254	13	independent	independent	ADJ
gs-3955	254	14	chaperones	chaperone	NOUN
gs-3955	254	15	(	(	PUNCT
gs-3955	254	16	hsp90α	hsp90α	PROPN
gs-3955	254	17	-	-	ADJ
gs-3955	254	18	t36v	t36v	ADJ
gs-3955	254	19	mutant	mutant	NOUN
gs-3955	254	20	)	)	PUNCT
gs-3955	254	21	(	(	PUNCT
gs-3955	254	22	munro	munro	PROPN
gs-3955	254	23	&	&	CCONJ
gs-3955	254	24	blier	blier	PROPN
gs-3955	254	25	,	,	PUNCT
gs-3955	254	26	2012	2012	NUM
gs-3955	254	27	)	)	PUNCT
gs-3955	254	28	●	●	NUM
gs-3955	254	29	asparagine	asparagine	NOUN
gs-3955	254	30	endopeptidase	endopeptidase	NOUN
gs-3955	254	31	preventing	prevent	VERB
gs-3955	254	32	cross	cross	ADJ
gs-3955	254	33	-	-	ADJ
gs-3955	254	34	linking	linking	ADJ
gs-3955	254	35	(	(	PUNCT
gs-3955	254	36	gruber	gruber	PROPN
gs-3955	254	37	et	et	PROPN
gs-3955	254	38	al	al	PROPN
gs-3955	254	39	.	.	PROPN
gs-3955	254	40	,	,	PUNCT
gs-3955	254	41	2015	2015	NUM
gs-3955	254	42	)	)	PUNCT
gs-3955	254	43	●	●	PUNCT
gs-3955	254	44	liquid	liquid	NOUN
gs-3955	254	45	-	-	PUNCT
gs-3955	254	46	liquid	liquid	NOUN
gs-3955	254	47	phase	phase	NOUN
gs-3955	254	48	separation	separation	NOUN
gs-3955	254	49	regulation	regulation	NOUN
gs-3955	254	50	(	(	PUNCT
gs-3955	254	51	tdp-43	tdp-43	NOUN
gs-3955	254	52	phosphorylation	phosphorylation	NOUN
gs-3955	254	53	resistance	resistance	NOUN
gs-3955	254	54	)	)	PUNCT
gs-3955	254	55	(	(	PUNCT
gs-3955	254	56	ballesta	ballesta	NOUN
gs-3955	254	57	-	-	PUNCT
gs-3955	254	58	artero	artero	NOUN
gs-3955	254	59	et	et	PROPN
gs-3955	254	60	al	al	PROPN
gs-3955	254	61	.	.	PROPN
gs-3955	254	62	,	,	PUNCT
gs-3955	254	63	2017	2017	NUM
gs-3955	254	64	)	)	PUNCT
gs-3955	254	65	georgian	georgian	ADJ
gs-3955	254	66	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	254	67	მეცნიერები	მეცნიერები	NOUN
gs-3955	254	68	ტ.	ტ.	VERB
gs-3955	254	69	7	7	NUM
gs-3955	254	70	n	n	PRON
gs-3955	254	71	3	3	NUM
gs-3955	254	72	,	,	PUNCT
gs-3955	254	73	2025	2025	NUM
gs-3955	254	74	375	375	NUM
gs-3955	254	75	stem	stem	NOUN
gs-3955	254	76	cell	cell	NOUN
gs-3955	254	77	immortality	immortality	NOUN
gs-3955	254	78	systems	system	NOUN
gs-3955	254	79	table	table	NOUN
gs-3955	254	80	3	3	NUM
gs-3955	254	81	:	:	PUNCT
gs-3955	254	82	stem	stem	NOUN
gs-3955	254	83	cell	cell	NOUN
gs-3955	254	84	maintenance	maintenance	NOUN
gs-3955	254	85	across	across	ADP
gs-3955	254	86	species	specie	NOUN
gs-3955	254	87	species	specie	NOUN
gs-3955	254	88	stem	stem	NOUN
gs-3955	254	89	cell	cell	NOUN
gs-3955	254	90	pool	pool	NOUN
gs-3955	254	91	turnover	turnover	NOUN
gs-3955	254	92	rate	rate	NOUN
gs-3955	254	93	key	key	ADJ
gs-3955	254	94	regulators	regulator	NOUN
gs-3955	254	95	regenerative	regenerative	ADJ
gs-3955	254	96	capacity	capacity	NOUN
gs-3955	254	97	hydra	hydra	NOUN
gs-3955	254	98	vulgaris	vulgaris	NOUN
gs-3955	254	99	(	(	PUNCT
gs-3955	254	100	bosch	bosch	PROPN
gs-3955	254	101	,	,	PUNCT
gs-3955	254	102	2009	2009	NUM
gs-3955	254	103	;	;	PUNCT
gs-3955	254	104	chichinadze	chichinadze	NOUN
gs-3955	254	105	et	et	PROPN
gs-3955	254	106	al	al	PROPN
gs-3955	254	107	.	.	PROPN
gs-3955	254	108	,	,	PUNCT
gs-3955	254	109	2013	2013	NUM
gs-3955	254	110	)	)	PUNCT
gs-3955	254	111	60	60	NUM
gs-3955	254	112	-	-	SYM
gs-3955	254	113	70	70	NUM
gs-3955	254	114	%	%	NOUN
gs-3955	254	115	3	3	NUM
gs-3955	254	116	-	-	SYM
gs-3955	254	117	4	4	NUM
gs-3955	254	118	days	day	NOUN
gs-3955	254	119	foxo	foxo	NOUN
gs-3955	254	120	,	,	PUNCT
gs-3955	254	121	wnt3	wnt3	PROPN
gs-3955	254	122	,	,	PUNCT
gs-3955	254	123	alk	alk	VERB
gs-3955	254	124	whole	whole	ADJ
gs-3955	254	125	-	-	PUNCT
gs-3955	254	126	body	body	NOUN
gs-3955	254	127	renewal	renewal	NOUN
gs-3955	254	128	planarians	planarian	NOUN
gs-3955	254	129	(	(	PUNCT
gs-3955	254	130	wagner	wagner	PROPN
gs-3955	254	131	et	et	PROPN
gs-3955	254	132	al	al	PROPN
gs-3955	254	133	.	.	PROPN
gs-3955	254	134	,	,	PUNCT
gs-3955	254	135	2011	2011	NUM
gs-3955	254	136	;	;	PUNCT
gs-3955	254	137	rink	rink	NOUN
gs-3955	254	138	,	,	PUNCT
gs-3955	254	139	2013	2013	NUM
gs-3955	254	140	)	)	PUNCT
gs-3955	254	141	25	25	NUM
gs-3955	254	142	-	-	SYM
gs-3955	254	143	30	30	NUM
gs-3955	254	144	%	%	NOUN
gs-3955	254	145	7	7	NUM
gs-3955	254	146	days	day	NOUN
gs-3955	254	147	piwi	piwi	NOUN
gs-3955	254	148	,	,	PUNCT
gs-3955	254	149	smedwi	smedwi	NOUN
gs-3955	254	150	,	,	PUNCT
gs-3955	254	151	nb.21.7e	nb.21.7e	PROPN
gs-3955	254	152	complete	complete	ADJ
gs-3955	254	153	regeneration	regeneration	NOUN
gs-3955	254	154	naked	naked	ADJ
gs-3955	254	155	molerat	molerat	NOUN
gs-3955	254	156	(	(	PUNCT
gs-3955	254	157	buffenstein	buffenstein	ADV
gs-3955	254	158	,	,	PUNCT
gs-3955	254	159	2008	2008	NUM
gs-3955	254	160	;	;	PUNCT
gs-3955	254	161	yang	yang	PROPN
gs-3955	254	162	et	et	PROPN
gs-3955	254	163	al	al	PROPN
gs-3955	254	164	.	.	PROPN
gs-3955	254	165	,	,	PUNCT
gs-3955	254	166	2019	2019	NUM
gs-3955	254	167	)	)	PUNCT
gs-3955	254	168	8	8	NUM
gs-3955	254	169	-	-	SYM
gs-3955	254	170	12	12	NUM
gs-3955	254	171	%	%	NOUN
gs-3955	254	172	28	28	NUM
gs-3955	254	173	days	day	NOUN
gs-3955	254	174	lgr5	lgr5	PROPN
gs-3955	254	175	,	,	PUNCT
gs-3955	254	176	bmi1	bmi1	NOUN
gs-3955	254	177	,	,	PUNCT
gs-3955	254	178	sox9	sox9	PROPN
gs-3955	254	179	limited	limit	VERB
gs-3955	254	180	organ	organ	NOUN
gs-3955	254	181	repair	repair	NOUN
gs-3955	254	182	fire	fire	NOUN
gs-3955	254	183	salamander	salamander	NOUN
gs-3955	254	184	(	(	PUNCT
gs-3955	254	185	joven	joven	PROPN
gs-3955	254	186	&	&	CCONJ
gs-3955	254	187	simon	simon	PROPN
gs-3955	254	188	,	,	PUNCT
gs-3955	254	189	2021	2021	NUM
gs-3955	254	190	;	;	PUNCT
gs-3955	254	191	zhang	zhang	PROPN
gs-3955	254	192	et	et	PROPN
gs-3955	254	193	al	al	PROPN
gs-3955	254	194	.	.	PROPN
gs-3955	254	195	,	,	PUNCT
gs-3955	254	196	2022	2022	NUM
gs-3955	254	197	)	)	PUNCT
gs-3955	254	198	5	5	NUM
gs-3955	254	199	-	-	SYM
gs-3955	254	200	8	8	NUM
gs-3955	254	201	%	%	NOUN
gs-3955	254	202	60	60	NUM
gs-3955	254	203	days	day	NOUN
gs-3955	254	204	msx1	msx1	PROPN
gs-3955	254	205	,	,	PUNCT
gs-3955	254	206	prrx1	prrx1	PROPN
gs-3955	254	207	,	,	PUNCT
gs-3955	254	208	fgf20	fgf20	ADJ
gs-3955	254	209	limb	limb	NOUN
gs-3955	254	210	/	/	SYM
gs-3955	254	211	cns	cns	NOUN
gs-3955	254	212	regeneration	regeneration	NOUN
gs-3955	254	213	novel	novel	NOUN
gs-3955	254	214	mechanisms	mechanism	NOUN
gs-3955	254	215	include	include	VERB
gs-3955	254	216	:	:	PUNCT
gs-3955	254	217	●	●	NUM
gs-3955	254	218	telomeric	telomeric	ADJ
gs-3955	254	219	epigenetics	epigenetic	NOUN
gs-3955	254	220	:	:	PUNCT
gs-3955	254	221	sharks	shark	NOUN
gs-3955	254	222	maintain	maintain	VERB
gs-3955	254	223	telomeric	telomeric	ADJ
gs-3955	254	224	heterochromatin	heterochromatin	NOUN
gs-3955	254	225	via	via	ADP
gs-3955	254	226	trf2suv39h1	trf2suv39h1	PROPN
gs-3955	254	227	complexes	complex	NOUN
gs-3955	254	228	(	(	PUNCT
gs-3955	254	229	seim	seim	PROPN
gs-3955	254	230	et	et	PROPN
gs-3955	254	231	al	al	PROPN
gs-3955	254	232	.	.	PROPN
gs-3955	254	233	,	,	PUNCT
gs-3955	254	234	2023	2023	NUM
gs-3955	254	235	;	;	PUNCT
gs-3955	254	236	de	de	X
gs-3955	254	237	magalhães	magalhães	NOUN
gs-3955	254	238	,	,	PUNCT
gs-3955	254	239	2023	2023	NUM
gs-3955	254	240	)	)	PUNCT
gs-3955	254	241	●	●	PUNCT
gs-3955	254	242	metabolic	metabolic	NOUN
gs-3955	254	243	priming	priming	NOUN
gs-3955	254	244	:	:	PUNCT
gs-3955	254	245	olm	olm	NOUN
gs-3955	254	246	stem	stem	NOUN
gs-3955	254	247	cells	cell	NOUN
gs-3955	254	248	utilize	utilize	VERB
gs-3955	254	249	ketone	ketone	NOUN
gs-3955	254	250	bodies	body	NOUN
gs-3955	254	251	(	(	PUNCT
gs-3955	254	252	β	β	NOUN
gs-3955	254	253	-	-	NOUN
gs-3955	254	254	hydroxybutyrate	hydroxybutyrate	NOUN
gs-3955	254	255	)	)	PUNCT
gs-3955	254	256	to	to	PART
gs-3955	254	257	maintain	maintain	VERB
gs-3955	254	258	pluripotency	pluripotency	NOUN
gs-3955	254	259	(	(	PUNCT
gs-3955	254	260	voituron	voituron	PROPN
gs-3955	254	261	et	et	PROPN
gs-3955	254	262	al	al	PROPN
gs-3955	254	263	.	.	PROPN
gs-3955	254	264	,	,	PUNCT
gs-3955	254	265	2011	2011	NUM
gs-3955	254	266	;	;	PUNCT
gs-3955	254	267	bulog	bulog	PROPN
gs-3955	254	268	et	et	PROPN
gs-3955	254	269	al	al	PROPN
gs-3955	254	270	.	.	PROPN
gs-3955	254	271	,	,	PUNCT
gs-3955	254	272	2020	2020	NUM
gs-3955	254	273	)	)	PUNCT
gs-3955	255	1	georgian	georgian	ADJ
gs-3955	255	2	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	255	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	255	4	ტ.	ტ.	VERB
gs-3955	255	5	7	7	NUM
gs-3955	255	6	n	n	DET
gs-3955	255	7	3	3	NUM
gs-3955	255	8	,	,	PUNCT
gs-3955	255	9	2025	2025	NUM
gs-3955	255	10	376	376	NUM
gs-3955	255	11	●	●	NUM
gs-3955	255	12	symmetric	symmetric	ADJ
gs-3955	255	13	division	division	NOUN
gs-3955	255	14	bias	bias	NOUN
gs-3955	255	15	:	:	PUNCT
gs-3955	255	16	planarian	planarian	NOUN
gs-3955	255	17	neoblasts	neoblast	NOUN
gs-3955	255	18	exhibit	exhibit	VERB
gs-3955	255	19	78	78	NUM
gs-3955	255	20	%	%	NOUN
gs-3955	255	21	symmetric	symmetric	ADJ
gs-3955	255	22	divisions	division	NOUN
gs-3955	255	23	(	(	PUNCT
gs-3955	255	24	vs.	vs.	ADP
gs-3955	255	25	35	35	NUM
gs-3955	255	26	%	%	NOUN
gs-3955	255	27	in	in	ADP
gs-3955	255	28	mammalian	mammalian	PROPN
gs-3955	255	29	scs	scs	PROPN
gs-3955	255	30	)	)	PUNCT
gs-3955	255	31	(	(	PUNCT
gs-3955	255	32	wagner	wagner	PROPN
gs-3955	255	33	et	et	PROPN
gs-3955	255	34	al	al	PROPN
gs-3955	255	35	.	.	PROPN
gs-3955	255	36	,	,	PUNCT
gs-3955	255	37	2011	2011	NUM
gs-3955	255	38	;	;	PUNCT
gs-3955	255	39	rink	rink	NOUN
gs-3955	255	40	,	,	PUNCT
gs-3955	255	41	2013	2013	NUM
gs-3955	255	42	)	)	PUNCT
gs-3955	255	43	systemic	systemic	ADJ
gs-3955	255	44	adaptations	adaptation	NOUN
gs-3955	255	45	:	:	PUNCT
gs-3955	255	46	beyond	beyond	ADP
gs-3955	255	47	molecular	molecular	ADJ
gs-3955	255	48	biology	biology	NOUN
gs-3955	255	49	neuroendocrine	neuroendocrine	NOUN
gs-3955	255	50	regulation	regulation	NOUN
gs-3955	255	51	long	long	ADV
gs-3955	255	52	-	-	PUNCT
gs-3955	255	53	lived	live	VERB
gs-3955	255	54	species	species	NOUN
gs-3955	255	55	exhibit	exhibit	VERB
gs-3955	255	56	optimized	optimize	VERB
gs-3955	255	57	stress	stress	NOUN
gs-3955	255	58	response	response	NOUN
gs-3955	255	59	systems	system	NOUN
gs-3955	255	60	:	:	PUNCT
gs-3955	255	61	●	●	NUM
gs-3955	255	62	glucocorticoid	glucocorticoid	ADJ
gs-3955	255	63	resistance	resistance	NOUN
gs-3955	255	64	:	:	PUNCT
gs-3955	255	65	greenland	greenland	PROPN
gs-3955	255	66	sharks	sharks	PROPN
gs-3955	255	67	show	show	VERB
gs-3955	255	68	10	10	NUM
gs-3955	255	69	-	-	ADJ
gs-3955	255	70	fold	fold	ADJ
gs-3955	255	71	higher	high	ADJ
gs-3955	255	72	grβ	grβ	NOUN
gs-3955	255	73	:	:	PUNCT
gs-3955	255	74	grα	grα	NOUN
gs-3955	255	75	ratio	ratio	NOUN
gs-3955	255	76	,	,	PUNCT
gs-3955	255	77	buffering	buffer	VERB
gs-3955	255	78	cortisol	cortisol	ADJ
gs-3955	255	79	effects	effect	NOUN
gs-3955	255	80	(	(	PUNCT
gs-3955	255	81	passow	passow	NOUN
gs-3955	255	82	et	et	PROPN
gs-3955	255	83	al	al	PROPN
gs-3955	255	84	.	.	PROPN
gs-3955	255	85	,	,	PUNCT
gs-3955	255	86	2022	2022	NUM
gs-3955	255	87	)	)	PUNCT
gs-3955	256	1	●	●	NUM
gs-3955	256	2	insulin	insulin	NOUN
gs-3955	256	3	signaling	signal	VERB
gs-3955	256	4	tuning	tuning	NOUN
gs-3955	256	5	:	:	PUNCT
gs-3955	256	6	naked	naked	ADJ
gs-3955	256	7	mole	mole	ADJ
gs-3955	256	8	-	-	PUNCT
gs-3955	256	9	rat	rat	NOUN
gs-3955	256	10	insulin	insulin	NOUN
gs-3955	256	11	receptors	receptor	NOUN
gs-3955	256	12	have	have	VERB
gs-3955	256	13	kd	kd	NOUN
gs-3955	256	14	=	=	SYM
gs-3955	256	15	3.7	3.7	NUM
gs-3955	256	16	nm	nm	NOUN
gs-3955	256	17	(	(	PUNCT
gs-3955	256	18	vs.	vs.	X
gs-3955	256	19	0.8	0.8	NUM
gs-3955	256	20	nm	nm	NOUN
gs-3955	256	21	in	in	ADP
gs-3955	256	22	mice	mouse	NOUN
gs-3955	256	23	)	)	PUNCT
gs-3955	256	24	,	,	PUNCT
gs-3955	256	25	preventing	prevent	VERB
gs-3955	256	26	hypoglycemia	hypoglycemia	NOUN
gs-3955	256	27	(	(	PUNCT
gs-3955	256	28	buffenstein	buffenstein	ADV
gs-3955	256	29	,	,	PUNCT
gs-3955	256	30	2008	2008	NUM
gs-3955	256	31	;	;	PUNCT
gs-3955	256	32	lewis	lewis	PROPN
gs-3955	256	33	et	et	PROPN
gs-3955	256	34	al	al	PROPN
gs-3955	256	35	.	.	PROPN
gs-3955	256	36	,	,	PUNCT
gs-3955	256	37	2018	2018	NUM
gs-3955	256	38	)	)	PUNCT
gs-3955	256	39	●	●	NUM
gs-3955	256	40	melatonin	melatonin	NOUN
gs-3955	256	41	amplification	amplification	NOUN
gs-3955	256	42	:	:	PUNCT
gs-3955	256	43	blind	blind	ADJ
gs-3955	256	44	cave	cave	NOUN
gs-3955	256	45	tetras	tetra	NOUN
gs-3955	256	46	(	(	PUNCT
gs-3955	256	47	astyanax	astyanax	NOUN
gs-3955	256	48	mexicanus	mexicanus	NOUN
gs-3955	256	49	)	)	PUNCT
gs-3955	256	50	produce	produce	VERB
gs-3955	256	51	melatonin	melatonin	NOUN
gs-3955	256	52	at	at	ADP
gs-3955	256	53	450	450	NUM
gs-3955	256	54	pg	pg	NOUN
gs-3955	256	55	/	/	SYM
gs-3955	256	56	ml	ml	PROPN
gs-3955	257	1	(	(	PUNCT
gs-3955	257	2	vs.	vs.	X
gs-3955	257	3	80	80	NUM
gs-3955	257	4	pg	pg	NOUN
gs-3955	257	5	/	/	SYM
gs-3955	257	6	ml	ml	X
gs-3955	257	7	in	in	ADP
gs-3955	257	8	surface	surface	NOUN
gs-3955	257	9	fish	fish	NOUN
gs-3955	257	10	)	)	PUNCT
gs-3955	257	11	(	(	PUNCT
gs-3955	257	12	bulog	bulog	NOUN
gs-3955	257	13	et	et	PROPN
gs-3955	257	14	al	al	PROPN
gs-3955	257	15	.	.	PROPN
gs-3955	257	16	,	,	PUNCT
gs-3955	257	17	2020	2020	NUM
gs-3955	257	18	)	)	PUNCT
gs-3955	257	19	immune	immune	ADJ
gs-3955	257	20	system	system	NOUN
gs-3955	257	21	longevity	longevity	NOUN
gs-3955	257	22	unique	unique	ADJ
gs-3955	257	23	adaptations	adaptation	NOUN
gs-3955	257	24	prevent	prevent	VERB
gs-3955	257	25	inflammaging	inflammage	VERB
gs-3955	257	26	:	:	PUNCT
gs-3955	257	27	●	●	NUM
gs-3955	257	28	expanded	expand	VERB
gs-3955	257	29	innate	innate	ADJ
gs-3955	257	30	repertoire	repertoire	NOUN
gs-3955	257	31	:	:	PUNCT
gs-3955	257	32	turtles	turtle	NOUN
gs-3955	257	33	express	express	VERB
gs-3955	257	34	43	43	NUM
gs-3955	257	35	tlr	tlr	PROPN
gs-3955	257	36	variants	variant	NOUN
gs-3955	257	37	(	(	PUNCT
gs-3955	257	38	vs.	vs.	ADP
gs-3955	257	39	10	10	NUM
gs-3955	257	40	in	in	ADP
gs-3955	257	41	mammals	mammal	NOUN
gs-3955	257	42	)	)	PUNCT
gs-3955	257	43	(	(	PUNCT
gs-3955	257	44	franceschi	franceschi	PROPN
gs-3955	257	45	et	et	PROPN
gs-3955	257	46	al	al	PROPN
gs-3955	257	47	.	.	PROPN
gs-3955	257	48	,	,	PUNCT
gs-3955	257	49	2022	2022	NUM
gs-3955	257	50	;	;	PUNCT
gs-3955	257	51	da	da	PROPN
gs-3955	257	52	silva	silva	PROPN
gs-3955	257	53	et	et	PROPN
gs-3955	257	54	al	al	PROPN
gs-3955	257	55	.	.	PROPN
gs-3955	257	56	,	,	PUNCT
gs-3955	257	57	2020	2020	NUM
gs-3955	257	58	)	)	PUNCT
gs-3955	257	59	●	●	PUNCT
gs-3955	257	60	regulatory	regulatory	ADJ
gs-3955	257	61	t	t	NOUN
gs-3955	257	62	-	-	PUNCT
gs-3955	257	63	cell	cell	NOUN
gs-3955	257	64	dominance	dominance	NOUN
gs-3955	257	65	:	:	PUNCT
gs-3955	257	66	naked	naked	ADJ
gs-3955	257	67	mole	mole	NOUN
gs-3955	257	68	-	-	PUNCT
gs-3955	257	69	rats	rat	NOUN
gs-3955	257	70	maintain	maintain	VERB
gs-3955	257	71	35	35	NUM
gs-3955	257	72	%	%	NOUN
gs-3955	257	73	tregs	treg	NOUN
gs-3955	257	74	in	in	ADP
gs-3955	257	75	circulation	circulation	NOUN
gs-3955	257	76	(	(	PUNCT
gs-3955	257	77	vs.	vs.	X
gs-3955	257	78	5	5	NUM
gs-3955	257	79	-	-	SYM
gs-3955	257	80	10	10	NUM
gs-3955	257	81	%	%	NOUN
gs-3955	257	82	in	in	ADP
gs-3955	257	83	mice	mouse	NOUN
gs-3955	257	84	)	)	PUNCT
gs-3955	258	1	(	(	PUNCT
gs-3955	258	2	lewis	lewis	PROPN
gs-3955	258	3	et	et	PROPN
gs-3955	258	4	al	al	PROPN
gs-3955	258	5	.	.	PROPN
gs-3955	258	6	,	,	PUNCT
gs-3955	258	7	2018	2018	NUM
gs-3955	258	8	;	;	PUNCT
gs-3955	258	9	buffenstein	buffenstein	ADV
gs-3955	258	10	,	,	PUNCT
gs-3955	258	11	2005	2005	NUM
gs-3955	258	12	)	)	PUNCT
gs-3955	258	13	●	●	PUNCT
gs-3955	258	14	non	non	ADJ
gs-3955	258	15	-	-	ADJ
gs-3955	258	16	classical	classical	ADJ
gs-3955	258	17	mhc	mhc	NOUN
gs-3955	258	18	:	:	PUNCT
gs-3955	258	19	sharks	shark	NOUN
gs-3955	258	20	utilize	utilize	VERB
gs-3955	258	21	ignar	ignar	NOUN
gs-3955	258	22	-	-	PUNCT
gs-3955	258	23	based	base	VERB
gs-3955	258	24	antigen	antigen	ADJ
gs-3955	258	25	recognition	recognition	NOUN
gs-3955	258	26	with	with	ADP
gs-3955	258	27	10	10	NUM
gs-3955	258	28	^	^	SYM
gs-3955	258	29	14	14	NUM
gs-3955	258	30	diversity	diversity	NOUN
gs-3955	258	31	(	(	PUNCT
gs-3955	258	32	kyne	kyne	PROPN
gs-3955	258	33	et	et	PROPN
gs-3955	258	34	al	al	PROPN
gs-3955	258	35	.	.	PROPN
gs-3955	258	36	,	,	PUNCT
gs-3955	258	37	2019	2019	NUM
gs-3955	258	38	)	)	PUNCT
gs-3955	258	39	metabolic	metabolic	NOUN
gs-3955	258	40	architecture	architecture	NOUN
gs-3955	258	41	convergent	convergent	NOUN
gs-3955	258	42	strategies	strategy	NOUN
gs-3955	258	43	include	include	VERB
gs-3955	258	44	:	:	PUNCT
gs-3955	258	45	●	●	NUM
gs-3955	258	46	mitochondrial	mitochondrial	ADJ
gs-3955	258	47	uncoupling	uncoupling	NOUN
gs-3955	258	48	:	:	PUNCT
gs-3955	258	49	fire	fire	NOUN
gs-3955	258	50	salamanders	salamander	NOUN
gs-3955	258	51	(	(	PUNCT
gs-3955	258	52	salamandra	salamandra	NOUN
gs-3955	258	53	salamandra	salamandra	NOUN
gs-3955	258	54	)	)	PUNCT
gs-3955	258	55	express	express	VERB
gs-3955	258	56	ucp1	ucp1	NOUN
gs-3955	258	57	in	in	ADP
gs-3955	258	58	brain	brain	NOUN
gs-3955	258	59	(	(	PUNCT
gs-3955	258	60	δψm	δψm	X
gs-3955	258	61	=	=	SYM
gs-3955	258	62	110	110	NUM
gs-3955	258	63	mv	mv	NOUN
gs-3955	258	64	vs.	vs.	ADP
gs-3955	258	65	140	140	NUM
gs-3955	258	66	mv	mv	PROPN
gs-3955	258	67	in	in	ADP
gs-3955	258	68	frogs	frog	NOUN
gs-3955	258	69	)	)	PUNCT
gs-3955	258	70	(	(	PUNCT
gs-3955	258	71	zhang	zhang	X
gs-3955	258	72	et	et	PROPN
gs-3955	258	73	al	al	PROPN
gs-3955	258	74	.	.	PROPN
gs-3955	258	75	,	,	PUNCT
gs-3955	258	76	2022	2022	NUM
gs-3955	258	77	;	;	PUNCT
gs-3955	258	78	joven	joven	PROPN
gs-3955	258	79	&	&	CCONJ
gs-3955	258	80	simon	simon	PROPN
gs-3955	258	81	,	,	PUNCT
gs-3955	258	82	2021	2021	NUM
gs-3955	258	83	)	)	PUNCT
gs-3955	258	84	●	●	NUM
gs-3955	258	85	hypometabolic	hypometabolic	ADJ
gs-3955	258	86	hibernation	hibernation	NOUN
gs-3955	258	87	:	:	PUNCT
gs-3955	258	88	blanding	blanding	PROPN
gs-3955	258	89	's	's	PART
gs-3955	258	90	turtles	turtle	NOUN
gs-3955	258	91	(	(	PUNCT
gs-3955	258	92	emydoidea	emydoidea	NOUN
gs-3955	258	93	blandingii	blandingii	NOUN
gs-3955	258	94	)	)	PUNCT
gs-3955	258	95	reduce	reduce	VERB
gs-3955	258	96	metabolic	metabolic	NOUN
gs-3955	258	97	rate	rate	NOUN
gs-3955	258	98	to	to	ADP
gs-3955	258	99	4	4	NUM
gs-3955	258	100	%	%	NOUN
gs-3955	258	101	of	of	ADP
gs-3955	258	102	basal	basal	NOUN
gs-3955	258	103	for	for	ADP
gs-3955	258	104	9	9	NUM
gs-3955	258	105	months	month	NOUN
gs-3955	258	106	(	(	PUNCT
gs-3955	258	107	storey	storey	NOUN
gs-3955	258	108	&	&	CCONJ
gs-3955	258	109	storey	storey	PROPN
gs-3955	258	110	,	,	PUNCT
gs-3955	258	111	2021	2021	NUM
gs-3955	258	112	)	)	PUNCT
gs-3955	258	113	●	●	NUM
gs-3955	258	114	anaerobic	anaerobic	NOUN
gs-3955	258	115	flexibility	flexibility	NOUN
gs-3955	258	116	:	:	PUNCT
gs-3955	258	117	olms	olm	NOUN
gs-3955	258	118	survive	survive	VERB
gs-3955	258	119	7	7	NUM
gs-3955	258	120	years	year	NOUN
gs-3955	258	121	without	without	ADP
gs-3955	258	122	oxygen	oxygen	NOUN
gs-3955	258	123	via	via	ADP
gs-3955	258	124	:	:	PUNCT
gs-3955	258	125	○	○	ADJ
gs-3955	258	126	succinate	succinate	NOUN
gs-3955	258	127	-	-	PUNCT
gs-3955	258	128	propionate	propionate	NOUN
gs-3955	258	129	fermentation	fermentation	NOUN
gs-3955	258	130	○	○	ADJ
gs-3955	258	131	alanine	alanine	NOUN
gs-3955	258	132	accumulation	accumulation	NOUN
gs-3955	258	133	(	(	PUNCT
gs-3955	258	134	up	up	ADP
gs-3955	258	135	to	to	PART
gs-3955	258	136	45	45	NUM
gs-3955	258	137	mm	mm	NOUN
gs-3955	258	138	)	)	PUNCT
gs-3955	258	139	○	○	ADJ
gs-3955	258	140	metabolic	metabolic	NOUN
gs-3955	258	141	depression	depression	NOUN
gs-3955	258	142	to	to	ADP
gs-3955	258	143	0.3	0.3	NUM
gs-3955	258	144	µl	µl	ADP
gs-3955	258	145	o₂/g	o₂/g	ADJ
gs-3955	258	146	/	/	SYM
gs-3955	258	147	h	h	NOUN
gs-3955	258	148	(	(	PUNCT
gs-3955	258	149	voituron	voituron	PROPN
gs-3955	258	150	et	et	PROPN
gs-3955	258	151	al	al	PROPN
gs-3955	258	152	.	.	PROPN
gs-3955	258	153	,	,	PUNCT
gs-3955	258	154	2011	2011	NUM
gs-3955	258	155	;	;	PUNCT
gs-3955	258	156	bulog	bulog	PROPN
gs-3955	258	157	et	et	PROPN
gs-3955	258	158	al	al	PROPN
gs-3955	258	159	.	.	PROPN
gs-3955	258	160	,	,	PUNCT
gs-3955	258	161	2020	2020	NUM
gs-3955	258	162	)	)	PUNCT
gs-3955	258	163	georgian	georgian	ADJ
gs-3955	258	164	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	258	165	მეცნიერები	მეცნიერები	NOUN
gs-3955	258	166	ტ.	ტ.	VERB
gs-3955	258	167	7	7	NUM
gs-3955	258	168	n	n	DET
gs-3955	258	169	3	3	NUM
gs-3955	258	170	,	,	PUNCT
gs-3955	258	171	2025	2025	NUM
gs-3955	258	172	377	377	NUM
gs-3955	258	173	translational	translational	ADJ
gs-3955	258	174	applications	application	NOUN
gs-3955	258	175	:	:	PUNCT
gs-3955	258	176	bridging	bridge	VERB
gs-3955	258	177	species	specie	NOUN
gs-3955	258	178	barriers	barrier	VERB
gs-3955	258	179	cancer	cancer	NOUN
gs-3955	258	180	resistance	resistance	NOUN
gs-3955	258	181	mechanisms	mechanism	NOUN
gs-3955	258	182	table	table	VERB
gs-3955	258	183	4	4	NUM
gs-3955	258	184	.	.	PUNCT
gs-3955	258	185	anti	anti	ADJ
gs-3955	258	186	-	-	ADJ
gs-3955	258	187	cancer	cancer	ADJ
gs-3955	258	188	adaptations	adaptation	NOUN
gs-3955	258	189	with	with	ADP
gs-3955	258	190	therapeutic	therapeutic	ADJ
gs-3955	258	191	potential	potential	ADJ
gs-3955	258	192	mechanism	mechanism	NOUN
gs-3955	258	193	species	specie	NOUN
gs-3955	258	194	key	key	ADJ
gs-3955	258	195	molecule	molecule	NOUN
gs-3955	258	196	human	human	ADJ
gs-3955	258	197	application	application	NOUN
gs-3955	258	198	status	status	NOUN
gs-3955	258	199	contact	contact	NOUN
gs-3955	258	200	inhibition	inhibition	NOUN
gs-3955	258	201	(	(	PUNCT
gs-3955	258	202	tian	tian	PROPN
gs-3955	258	203	et	et	PROPN
gs-3955	258	204	al	al	PROPN
gs-3955	258	205	.	.	PROPN
gs-3955	258	206	,	,	PUNCT
gs-3955	258	207	2013	2013	NUM
gs-3955	258	208	;	;	PUNCT
gs-3955	258	209	seluanov	seluanov	PROPN
gs-3955	258	210	et	et	PROPN
gs-3955	258	211	al	al	PROPN
gs-3955	258	212	.	.	PROPN
gs-3955	258	213	,	,	PUNCT
gs-3955	258	214	2009	2009	NUM
gs-3955	258	215	)	)	PUNCT
gs-3955	258	216	naked	naked	ADJ
gs-3955	258	217	mole	mole	ADJ
gs-3955	258	218	-	-	PUNCT
gs-3955	258	219	rat	rat	NOUN
gs-3955	258	220	hmw	hmw	PROPN
gs-3955	258	221	-	-	PUNCT
gs-3955	258	222	ha	ha	INTJ
gs-3955	258	223	(	(	PUNCT
gs-3955	258	224	>	>	NOUN
gs-3955	258	225	6,000	6,000	NUM
gs-3955	258	226	kda	kda	NOUN
gs-3955	258	227	)	)	PUNCT
gs-3955	258	228	phase	phase	NOUN
gs-3955	258	229	ii	ii	NOUN
gs-3955	258	230	trials	trial	NOUN
gs-3955	258	231	(	(	PUNCT
gs-3955	258	232	hmw	hmw	PROPN
gs-3955	258	233	-	-	PUNCT
gs-3955	258	234	ha	ha	INTJ
gs-3955	258	235	gels	gel	NOUN
gs-3955	258	236	)	)	PUNCT
gs-3955	258	237	p53	p53	NOUN
gs-3955	258	238	variant	variant	NOUN
gs-3955	258	239	function	function	NOUN
gs-3955	258	240	(	(	PUNCT
gs-3955	258	241	abegglen	abegglen	VERB
gs-3955	258	242	et	et	PROPN
gs-3955	258	243	al	al	PROPN
gs-3955	258	244	.	.	PROPN
gs-3955	258	245	,	,	PUNCT
gs-3955	258	246	2015	2015	NUM
gs-3955	258	247	;	;	PUNCT
gs-3955	258	248	sulak	sulak	PROPN
gs-3955	258	249	et	et	PROPN
gs-3955	258	250	al	al	PROPN
gs-3955	258	251	.	.	PROPN
gs-3955	258	252	,	,	PUNCT
gs-3955	258	253	2016	2016	NUM
gs-3955	258	254	)	)	PUNCT
gs-3955	258	255	elephants	elephant	VERB
gs-3955	258	256	tp53	tp53	PROPN
gs-3955	258	257	retrogenes	retrogene	NOUN
gs-3955	258	258	(	(	PUNCT
gs-3955	258	259	20	20	NUM
gs-3955	258	260	copies	copy	NOUN
gs-3955	258	261	)	)	PUNCT
gs-3955	258	262	gene	gene	NOUN
gs-3955	258	263	therapy	therapy	NOUN
gs-3955	258	264	development	development	NOUN
gs-3955	258	265	senescence	senescence	PROPN
gs-3955	258	266	evasion(zhang	evasion(zhang	PROPN
gs-3955	258	267	et	et	PROPN
gs-3955	258	268	al	al	PROPN
gs-3955	258	269	.	.	PROPN
gs-3955	258	270	,	,	PUNCT
gs-3955	258	271	2013	2013	NUM
gs-3955	258	272	;	;	PUNCT
gs-3955	258	273	foley	foley	PROPN
gs-3955	258	274	et	et	PROPN
gs-3955	258	275	al	al	PROPN
gs-3955	258	276	.	.	PROPN
gs-3955	258	277	,	,	PUNCT
gs-3955	258	278	2018	2018	NUM
gs-3955	258	279	)	)	PUNCT
gs-3955	258	280	bats	bat	NOUN
gs-3955	258	281	atm	atm	VERB
gs-3955	258	282	hyperphosphorylation	hyperphosphorylation	NOUN
gs-3955	258	283	small	small	ADJ
gs-3955	258	284	molecule	molecule	NOUN
gs-3955	258	285	screening	screen	VERB
gs-3955	258	286	hypoxic	hypoxic	ADV
gs-3955	258	287	tumor	tumor	NOUN
gs-3955	258	288	suppression	suppression	NOUN
gs-3955	258	289	(	(	PUNCT
gs-3955	258	290	voituron	voituron	PROPN
gs-3955	258	291	et	et	PROPN
gs-3955	258	292	al	al	PROPN
gs-3955	258	293	.	.	PROPN
gs-3955	258	294	,	,	PUNCT
gs-3955	258	295	2011	2011	NUM
gs-3955	258	296	;	;	PUNCT
gs-3955	258	297	bulog	bulog	PROPN
gs-3955	258	298	et	et	PROPN
gs-3955	258	299	al	al	PROPN
gs-3955	258	300	.	.	PROPN
gs-3955	258	301	,	,	PUNCT
gs-3955	258	302	2020	2020	NUM
gs-3955	258	303	)	)	PUNCT
gs-3955	258	304	olm	olm	NOUN
gs-3955	258	305	hif-1α	hif-1α	NOUN
gs-3955	258	306	inhibitory	inhibitory	ADJ
gs-3955	258	307	peptide	peptide	NOUN
gs-3955	258	308	preclinical	preclinical	ADJ
gs-3955	258	309	testing	testing	NOUN
gs-3955	258	310	pharmacological	pharmacological	NOUN
gs-3955	258	311	targets	target	NOUN
gs-3955	258	312	emerging	emerge	VERB
gs-3955	258	313	therapeutic	therapeutic	ADJ
gs-3955	258	314	strategies	strategy	NOUN
gs-3955	258	315	include	include	VERB
gs-3955	258	316	:	:	PUNCT
gs-3955	258	317	●	●	NUM
gs-3955	258	318	sirt6	sirt6	NOUN
gs-3955	258	319	activators	activator	NOUN
gs-3955	258	320	:	:	PUNCT
gs-3955	258	321	mimicking	mimic	VERB
gs-3955	258	322	shark	shark	NOUN
gs-3955	258	323	dna	dna	PROPN
gs-3955	258	324	repair	repair	NOUN
gs-3955	258	325	(	(	PUNCT
gs-3955	258	326	mdl-800	mdl-800	ADJ
gs-3955	258	327	series	series	NOUN
gs-3955	258	328	in	in	ADP
gs-3955	258	329	phase	phase	NOUN
gs-3955	258	330	i	i	PROPN
gs-3955	258	331	)	)	PUNCT
gs-3955	258	332	(	(	PUNCT
gs-3955	258	333	kumar	kumar	PROPN
gs-3955	258	334	et	et	PROPN
gs-3955	258	335	al	al	PROPN
gs-3955	258	336	.	.	PROPN
gs-3955	258	337	,	,	PUNCT
gs-3955	258	338	2021	2021	NUM
gs-3955	258	339	;	;	PUNCT
gs-3955	258	340	zhang	zhang	PROPN
gs-3955	258	341	et	et	PROPN
gs-3955	258	342	al	al	PROPN
gs-3955	258	343	.	.	PROPN
gs-3955	258	344	,	,	PUNCT
gs-3955	258	345	2023	2023	NUM
gs-3955	258	346	)	)	PUNCT
gs-3955	258	347	●	●	NUM
gs-3955	258	348	foxo	foxo	NOUN
gs-3955	258	349	stabilizers	stabilizer	NOUN
gs-3955	258	350	:	:	PUNCT
gs-3955	258	351	hydra	hydra	NOUN
gs-3955	258	352	-	-	PUNCT
gs-3955	258	353	inspired	inspire	VERB
gs-3955	258	354	peptides	peptide	NOUN
gs-3955	258	355	(	(	PUNCT
gs-3955	258	356	foxo	foxo	NOUN
gs-3955	258	357	-	-	PUNCT
gs-3955	258	358	stab	stab	NOUN
gs-3955	258	359	peptides	peptide	NOUN
gs-3955	258	360	)	)	PUNCT
gs-3955	258	361	(	(	PUNCT
gs-3955	258	362	siebert	siebert	PROPN
gs-3955	258	363	et	et	PROPN
gs-3955	258	364	al	al	PROPN
gs-3955	258	365	.	.	PROPN
gs-3955	258	366	,	,	PUNCT
gs-3955	258	367	2019	2019	NUM
gs-3955	258	368	;	;	PUNCT
gs-3955	258	369	wang	wang	PROPN
gs-3955	258	370	et	et	PROPN
gs-3955	258	371	al	al	PROPN
gs-3955	258	372	.	.	PROPN
gs-3955	258	373	,	,	PUNCT
gs-3955	258	374	2022	2022	NUM
gs-3955	258	375	)	)	PUNCT
gs-3955	258	376	georgian	georgian	ADJ
gs-3955	258	377	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	258	378	მეცნიერები	მეცნიერები	NOUN
gs-3955	258	379	ტ.	ტ.	VERB
gs-3955	258	380	7	7	NUM
gs-3955	258	381	n	n	DET
gs-3955	258	382	3	3	NUM
gs-3955	258	383	,	,	PUNCT
gs-3955	258	384	2025	2025	NUM
gs-3955	258	385	378	378	NUM
gs-3955	258	386	●	●	NUM
gs-3955	258	387	membrane	membrane	NOUN
gs-3955	258	388	lipid	lipid	NOUN
gs-3955	258	389	emulators	emulator	NOUN
gs-3955	258	390	:	:	PUNCT
gs-3955	258	391	synthetic	synthetic	ADJ
gs-3955	258	392	alkylglycerols	alkylglycerol	NOUN
gs-3955	258	393	based	base	VERB
gs-3955	258	394	on	on	ADP
gs-3955	258	395	shark	shark	NOUN
gs-3955	258	396	lipids	lipid	NOUN
gs-3955	258	397	(	(	PUNCT
gs-3955	258	398	alg101	alg101	PROPN
gs-3955	258	399	)	)	PUNCT
gs-3955	258	400	(	(	PUNCT
gs-3955	258	401	martinez	martinez	PROPN
gs-3955	258	402	et	et	PROPN
gs-3955	258	403	al	al	PROPN
gs-3955	258	404	.	.	PROPN
gs-3955	258	405	,	,	PUNCT
gs-3955	258	406	2020	2020	NUM
gs-3955	258	407	)	)	PUNCT
gs-3955	258	408	challenges	challenge	NOUN
gs-3955	258	409	in	in	ADP
gs-3955	258	410	translation	translation	NOUN
gs-3955	258	411	key	key	ADJ
gs-3955	258	412	barriers	barrier	NOUN
gs-3955	258	413	remain	remain	VERB
gs-3955	258	414	:	:	PUNCT
gs-3955	258	415	●	●	PUNCT
gs-3955	258	416	evolutionary	evolutionary	ADJ
gs-3955	258	417	divergence	divergence	NOUN
gs-3955	258	418	:	:	PUNCT
gs-3955	259	1	human	human	ADJ
gs-3955	259	2	telomerase	telomerase	NOUN
gs-3955	259	3	reactivation	reactivation	NOUN
gs-3955	259	4	risks	risk	VERB
gs-3955	259	5	oncogenesis	oncogenesis	VERB
gs-3955	259	6	absent	absent	ADJ
gs-3955	259	7	mole	mole	NOUN
gs-3955	259	8	-	-	PUNCT
gs-3955	259	9	rat	rat	NOUN
gs-3955	259	10	's	's	PART
gs-3955	259	11	concurrent	concurrent	ADJ
gs-3955	259	12	cancer	cancer	NOUN
gs-3955	259	13	resistance	resistance	NOUN
gs-3955	259	14	(	(	PUNCT
gs-3955	259	15	seluanov	seluanov	PROPN
gs-3955	259	16	et	et	PROPN
gs-3955	259	17	al	al	PROPN
gs-3955	259	18	.	.	PROPN
gs-3955	259	19	,	,	PUNCT
gs-3955	259	20	2018	2018	NUM
gs-3955	259	21	;	;	PUNCT
gs-3955	259	22	tian	tian	PROPN
gs-3955	259	23	et	et	PROPN
gs-3955	259	24	al	al	PROPN
gs-3955	259	25	.	.	PROPN
gs-3955	259	26	,	,	PUNCT
gs-3955	259	27	2013	2013	NUM
gs-3955	259	28	)	)	PUNCT
gs-3955	259	29	●	●	PUNCT
gs-3955	259	30	metabolic	metabolic	NOUN
gs-3955	259	31	trade	trade	NOUN
gs-3955	259	32	-	-	PUNCT
gs-3955	259	33	offs	off	NOUN
gs-3955	259	34	:	:	PUNCT
gs-3955	259	35	mimicking	mimic	VERB
gs-3955	259	36	shark	shark	NOUN
gs-3955	259	37	metabolic	metabolic	NOUN
gs-3955	259	38	depression	depression	NOUN
gs-3955	259	39	could	could	AUX
gs-3955	259	40	impair	impair	VERB
gs-3955	259	41	cognition	cognition	NOUN
gs-3955	259	42	(	(	PUNCT
gs-3955	259	43	cheng	cheng	PROPN
gs-3955	259	44	et	et	PROPN
gs-3955	259	45	al	al	PROPN
gs-3955	259	46	.	.	PROPN
gs-3955	259	47	,	,	PUNCT
gs-3955	259	48	2019	2019	NUM
gs-3955	259	49	)	)	PUNCT
gs-3955	259	50	●	●	NUM
gs-3955	259	51	developmental	developmental	ADJ
gs-3955	259	52	constraints	constraint	NOUN
gs-3955	259	53	:	:	PUNCT
gs-3955	259	54	planarian	planarian	ADJ
gs-3955	259	55	-	-	PUNCT
gs-3955	259	56	like	like	ADJ
gs-3955	259	57	regeneration	regeneration	NOUN
gs-3955	259	58	requires	require	VERB
gs-3955	259	59	embryonic	embryonic	ADJ
gs-3955	259	60	-	-	PUNCT
gs-3955	259	61	level	level	NOUN
gs-3955	259	62	plasticity	plasticity	NOUN
gs-3955	259	63	unattainable	unattainable	ADJ
gs-3955	259	64	in	in	ADP
gs-3955	259	65	adults	adult	NOUN
gs-3955	259	66	(	(	PUNCT
gs-3955	259	67	wagner	wagner	PROPN
gs-3955	259	68	et	et	PROPN
gs-3955	259	69	al	al	PROPN
gs-3955	259	70	.	.	PROPN
gs-3955	259	71	,	,	PUNCT
gs-3955	259	72	2011	2011	NUM
gs-3955	259	73	;	;	PUNCT
gs-3955	259	74	rink	rink	NOUN
gs-3955	259	75	,	,	PUNCT
gs-3955	259	76	2013	2013	NUM
gs-3955	259	77	)	)	PUNCT
gs-3955	259	78	future	future	ADJ
gs-3955	259	79	research	research	NOUN
gs-3955	259	80	frontiers	frontier	NOUN
gs-3955	259	81	integrated	integrate	VERB
gs-3955	259	82	"	"	PUNCT
gs-3955	259	83	longevomics	longevomic	NOUN
gs-3955	259	84	"	"	PUNCT
gs-3955	259	85	next	next	ADJ
gs-3955	259	86	-	-	PUNCT
gs-3955	259	87	generation	generation	NOUN
gs-3955	259	88	approaches	approach	NOUN
gs-3955	259	89	include	include	VERB
gs-3955	259	90	:	:	PUNCT
gs-3955	259	91	●	●	NUM
gs-3955	259	92	single	single	ADJ
gs-3955	259	93	-	-	PUNCT
gs-3955	259	94	cell	cell	NOUN
gs-3955	259	95	atlases	atlas	NOUN
gs-3955	259	96	:	:	PUNCT
gs-3955	259	97	spatiotemporal	spatiotemporal	ADJ
gs-3955	259	98	mapping	mapping	NOUN
gs-3955	259	99	of	of	ADP
gs-3955	259	100	500	500	NUM
gs-3955	259	101	-	-	PUNCT
gs-3955	259	102	year	year	NOUN
gs-3955	259	103	-	-	PUNCT
gs-3955	259	104	old	old	ADJ
gs-3955	259	105	arctica	arctica	PROPN
gs-3955	259	106	islandica	islandica	PROPN
gs-3955	259	107	hemocytes	hemocyte	VERB
gs-3955	259	108	(	(	PUNCT
gs-3955	259	109	smith	smith	PROPN
gs-3955	259	110	et	et	PROPN
gs-3955	259	111	al	al	PROPN
gs-3955	259	112	.	.	PROPN
gs-3955	259	113	,	,	PUNCT
gs-3955	259	114	2022	2022	NUM
gs-3955	259	115	)	)	PUNCT
gs-3955	259	116	●	●	NUM
gs-3955	259	117	quantum	quantum	NOUN
gs-3955	259	118	biology	biology	NOUN
gs-3955	259	119	probes	probe	NOUN
gs-3955	259	120	:	:	PUNCT
gs-3955	259	121	assessing	assess	VERB
gs-3955	259	122	electron	electron	NOUN
gs-3955	259	123	tunneling	tunneling	NOUN
gs-3955	259	124	in	in	ADP
gs-3955	259	125	shark	shark	NOUN
gs-3955	259	126	mitochondrial	mitochondrial	ADJ
gs-3955	259	127	complexes	complex	NOUN
gs-3955	259	128	(	(	PUNCT
gs-3955	259	129	jones	jones	PROPN
gs-3955	259	130	et	et	PROPN
gs-3955	259	131	al	al	PROPN
gs-3955	259	132	.	.	PROPN
gs-3955	259	133	,	,	PUNCT
gs-3955	259	134	2021	2021	NUM
gs-3955	259	135	)	)	PUNCT
gs-3955	259	136	●	●	PUNCT
gs-3955	259	137	cryo	cryo	NOUN
gs-3955	259	138	-	-	PUNCT
gs-3955	259	139	electron	electron	NOUN
gs-3955	259	140	tomography	tomography	NOUN
gs-3955	259	141	:	:	PUNCT
gs-3955	259	142	visualizing	visualize	VERB
gs-3955	259	143	naked	naked	ADJ
gs-3955	259	144	mole	mole	ADJ
gs-3955	259	145	-	-	PUNCT
gs-3955	259	146	rat	rat	NOUN
gs-3955	259	147	ribosome	ribosome	NOUN
gs-3955	259	148	error	error	NOUN
gs-3955	259	149	-	-	PUNCT
gs-3955	259	150	correction	correction	NOUN
gs-3955	259	151	(	(	PUNCT
gs-3955	259	152	liu	liu	PROPN
gs-3955	259	153	et	et	PROPN
gs-3955	259	154	al	al	PROPN
gs-3955	259	155	.	.	PROPN
gs-3955	259	156	,	,	PUNCT
gs-3955	259	157	2023	2023	NUM
gs-3955	259	158	)	)	PUNCT
gs-3955	259	159	synthetic	synthetic	ADJ
gs-3955	259	160	negligible	negligible	ADJ
gs-3955	259	161	senescence	senescence	NOUN
gs-3955	259	162	engineering	engineering	NOUN
gs-3955	259	163	longevity	longevity	NOUN
gs-3955	259	164	through	through	ADP
gs-3955	259	165	:	:	PUNCT
gs-3955	259	166	●	●	NUM
gs-3955	259	167	chimeric	chimeric	ADJ
gs-3955	259	168	models	model	NOUN
gs-3955	259	169	:	:	PUNCT
gs-3955	259	170	humanized	humanize	VERB
gs-3955	259	171	mole	mole	ADJ
gs-3955	259	172	-	-	PUNCT
gs-3955	259	173	rat	rat	NOUN
gs-3955	259	174	immune	immune	ADJ
gs-3955	259	175	systems	system	NOUN
gs-3955	259	176	in	in	ADP
gs-3955	259	177	primates	primate	NOUN
gs-3955	259	178	(	(	PUNCT
gs-3955	259	179	garcia	garcia	PROPN
gs-3955	259	180	et	et	PROPN
gs-3955	259	181	al	al	PROPN
gs-3955	259	182	.	.	PROPN
gs-3955	259	183	,	,	PUNCT
gs-3955	259	184	2023	2023	NUM
gs-3955	259	185	)	)	PUNCT
gs-3955	259	186	●	●	NUM
gs-3955	259	187	gene	gene	NOUN
gs-3955	259	188	circuitry	circuitry	NOUN
gs-3955	259	189	:	:	PUNCT
gs-3955	259	190	synthetic	synthetic	ADJ
gs-3955	259	191	foxo	foxo	PROPN
gs-3955	259	192	-	-	PUNCT
gs-3955	259	193	hif	hif	PROPN
gs-3955	259	194	-	-	PUNCT
gs-3955	259	195	sirt	sirt	PROPN
gs-3955	259	196	networks	network	NOUN
gs-3955	259	197	(	(	PUNCT
gs-3955	259	198	lee	lee	PROPN
gs-3955	259	199	et	et	PROPN
gs-3955	259	200	al	al	PROPN
gs-3955	259	201	.	.	PROPN
gs-3955	259	202	,	,	PUNCT
gs-3955	259	203	2022	2022	NUM
gs-3955	259	204	)	)	PUNCT
gs-3955	259	205	●	●	PUNCT
gs-3955	259	206	mitochondrial	mitochondrial	ADJ
gs-3955	259	207	transplantation	transplantation	NOUN
gs-3955	259	208	:	:	PUNCT
gs-3955	259	209	shark	shark	NOUN
gs-3955	259	210	-	-	PUNCT
gs-3955	259	211	derived	derive	VERB
gs-3955	259	212	organelles	organelle	NOUN
gs-3955	259	213	in	in	ADP
gs-3955	259	214	mammalian	mammalian	ADJ
gs-3955	259	215	cells	cell	NOUN
gs-3955	259	216	(	(	PUNCT
gs-3955	259	217	patel	patel	PROPN
gs-3955	259	218	et	et	PROPN
gs-3955	259	219	al	al	PROPN
gs-3955	259	220	.	.	PROPN
gs-3955	259	221	,	,	PUNCT
gs-3955	259	222	2024	2024	NUM
gs-3955	259	223	)	)	PUNCT
gs-3955	259	224	ethical	ethical	ADJ
gs-3955	259	225	frameworks	framework	NOUN
gs-3955	259	226	emerging	emerge	VERB
gs-3955	259	227	considerations	consideration	NOUN
gs-3955	259	228	:	:	PUNCT
gs-3955	259	229	●	●	NUM
gs-3955	259	230	longevity	longevity	NOUN
gs-3955	259	231	equity	equity	NOUN
gs-3955	259	232	:	:	PUNCT
gs-3955	259	233	ensuring	ensure	VERB
gs-3955	259	234	access	access	NOUN
gs-3955	259	235	beyond	beyond	ADP
gs-3955	259	236	socioeconomic	socioeconomic	ADJ
gs-3955	259	237	elites	elite	NOUN
gs-3955	259	238	(	(	PUNCT
gs-3955	259	239	nguyen	nguyen	NOUN
gs-3955	259	240	et	et	PROPN
gs-3955	259	241	al	al	PROPN
gs-3955	259	242	.	.	PROPN
gs-3955	259	243	,	,	PUNCT
gs-3955	259	244	2022	2022	NUM
gs-3955	259	245	)	)	PUNCT
gs-3955	259	246	●	●	PUNCT
gs-3955	259	247	ecological	ecological	ADJ
gs-3955	259	248	impact	impact	NOUN
gs-3955	259	249	:	:	PUNCT
gs-3955	259	250	population	population	NOUN
gs-3955	259	251	controls	control	VERB
gs-3955	259	252	for	for	ADP
gs-3955	259	253	extended	extended	ADJ
gs-3955	259	254	human	human	ADJ
gs-3955	259	255	lifespans	lifespan	NOUN
gs-3955	259	256	(	(	PUNCT
gs-3955	259	257	rodriguez	rodriguez	PROPN
gs-3955	259	258	&	&	CCONJ
gs-3955	259	259	kim	kim	PROPN
gs-3955	259	260	,	,	PUNCT
gs-3955	259	261	2023	2023	NUM
gs-3955	259	262	)	)	PUNCT
gs-3955	259	263	●	●	NUM
gs-3955	259	264	identity	identity	NOUN
gs-3955	259	265	continuity	continuity	NOUN
gs-3955	259	266	:	:	PUNCT
gs-3955	259	267	psychological	psychological	ADJ
gs-3955	259	268	implications	implication	NOUN
gs-3955	259	269	of	of	ADP
gs-3955	259	270	150	150	NUM
gs-3955	259	271	-	-	PUNCT
gs-3955	259	272	year+	year+	NOUN
gs-3955	259	273	lifespans	lifespan	NOUN
gs-3955	259	274	(	(	PUNCT
gs-3955	259	275	martin	martin	PROPN
gs-3955	259	276	&	&	CCONJ
gs-3955	259	277	davies	davy	NOUN
gs-3955	259	278	,	,	PUNCT
gs-3955	259	279	2021	2021	NUM
gs-3955	259	280	)	)	PUNCT
gs-3955	259	281	georgian	georgian	ADJ
gs-3955	259	282	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	259	283	მეცნიერები	მეცნიერები	NOUN
gs-3955	259	284	ტ.	ტ.	VERB
gs-3955	259	285	7	7	NUM
gs-3955	259	286	n	n	PRON
gs-3955	259	287	3	3	NUM
gs-3955	259	288	,	,	PUNCT
gs-3955	259	289	2025	2025	NUM
gs-3955	259	290	379	379	NUM
gs-3955	259	291	multifaceted	multifaceted	ADJ
gs-3955	259	292	roles	role	NOUN
gs-3955	259	293	of	of	ADP
gs-3955	259	294	autophagy	autophagy	NOUN
gs-3955	259	295	in	in	ADP
gs-3955	259	296	negligibly	negligibly	ADV
gs-3955	259	297	senescent	senescent	ADJ
gs-3955	259	298	organisms	organism	NOUN
gs-3955	259	299	autophagy	autophagy	VERB
gs-3955	259	300	is	be	AUX
gs-3955	259	301	a	a	DET
gs-3955	259	302	fundamental	fundamental	ADJ
gs-3955	259	303	cellular	cellular	ADJ
gs-3955	259	304	process	process	NOUN
gs-3955	259	305	conserved	conserve	VERB
gs-3955	259	306	across	across	ADP
gs-3955	259	307	eukaryotes	eukaryote	NOUN
gs-3955	259	308	,	,	PUNCT
gs-3955	259	309	essential	essential	ADJ
gs-3955	259	310	not	not	PART
gs-3955	259	311	only	only	ADV
gs-3955	259	312	for	for	ADP
gs-3955	259	313	longevity	longevity	NOUN
gs-3955	259	314	but	but	CCONJ
gs-3955	259	315	also	also	ADV
gs-3955	259	316	for	for	ADP
gs-3955	259	317	development	development	NOUN
gs-3955	259	318	,	,	PUNCT
gs-3955	259	319	stress	stress	NOUN
gs-3955	259	320	adaptation	adaptation	NOUN
gs-3955	259	321	,	,	PUNCT
gs-3955	259	322	and	and	CCONJ
gs-3955	259	323	tissue	tissue	NOUN
gs-3955	259	324	regeneration	regeneration	NOUN
gs-3955	259	325	.	.	PUNCT
gs-3955	260	1	in	in	ADP
gs-3955	260	2	negligibly	negligibly	ADV
gs-3955	260	3	senescent	senescent	ADJ
gs-3955	260	4	species	specie	NOUN
gs-3955	260	5	such	such	ADJ
gs-3955	260	6	as	as	ADP
gs-3955	260	7	hydra	hydra	PROPN
gs-3955	260	8	vulgaris	vulgaris	PROPN
gs-3955	260	9	,	,	PUNCT
gs-3955	260	10	arctica	arctica	PROPN
gs-3955	260	11	islandica	islandica	PROPN
gs-3955	260	12	,	,	PUNCT
gs-3955	261	1	naked	naked	ADJ
gs-3955	261	2	mole	mole	NOUN
gs-3955	261	3	-	-	PUNCT
gs-3955	261	4	rats	rat	NOUN
gs-3955	261	5	(	(	PUNCT
gs-3955	261	6	heterocephalus	heterocephalus	ADJ
gs-3955	261	7	glaber	glaber	NOUN
gs-3955	261	8	)	)	PUNCT
gs-3955	261	9	,	,	PUNCT
gs-3955	261	10	and	and	CCONJ
gs-3955	261	11	rockfish	rockfish	ADJ
gs-3955	261	12	(	(	PUNCT
gs-3955	261	13	sebastes	sebaste	VERB
gs-3955	261	14	spp	spp	NOUN
gs-3955	261	15	.	.	PUNCT
gs-3955	261	16	)	)	PUNCT
gs-3955	262	1	,	,	PUNCT
gs-3955	262	2	autophagy	autophagy	NOUN
gs-3955	262	3	fulfills	fulfill	VERB
gs-3955	262	4	diverse	diverse	ADJ
gs-3955	262	5	biological	biological	ADJ
gs-3955	262	6	functions	function	NOUN
gs-3955	262	7	crucial	crucial	ADJ
gs-3955	262	8	for	for	ADP
gs-3955	262	9	their	their	PRON
gs-3955	262	10	exceptional	exceptional	ADJ
gs-3955	262	11	survival	survival	NOUN
gs-3955	262	12	strategies	strategy	NOUN
gs-3955	262	13	.	.	PUNCT
gs-3955	263	1	autophagy	autophagy	VERB
gs-3955	263	2	in	in	ADP
gs-3955	263	3	development	development	NOUN
gs-3955	263	4	and	and	CCONJ
gs-3955	263	5	regeneration	regeneration	NOUN
gs-3955	263	6	in	in	ADP
gs-3955	263	7	hydra	hydra	PROPN
gs-3955	263	8	,	,	PUNCT
gs-3955	263	9	autophagy	autophagy	NOUN
gs-3955	263	10	plays	play	VERB
gs-3955	263	11	an	an	DET
gs-3955	263	12	indispensable	indispensable	ADJ
gs-3955	263	13	role	role	NOUN
gs-3955	263	14	during	during	ADP
gs-3955	263	15	morphogenesis	morphogenesis	NOUN
gs-3955	263	16	and	and	CCONJ
gs-3955	263	17	regeneration	regeneration	NOUN
gs-3955	263	18	.	.	PUNCT
gs-3955	264	1	continuous	continuous	ADJ
gs-3955	264	2	autophagic	autophagic	PROPN
gs-3955	264	3	flux	flux	NOUN
gs-3955	264	4	supports	support	VERB
gs-3955	264	5	the	the	DET
gs-3955	264	6	high	high	ADJ
gs-3955	264	7	proliferative	proliferative	ADJ
gs-3955	264	8	activity	activity	NOUN
gs-3955	264	9	of	of	ADP
gs-3955	264	10	interstitial	interstitial	ADJ
gs-3955	264	11	stem	stem	NOUN
gs-3955	264	12	cells	cell	NOUN
gs-3955	264	13	,	,	PUNCT
gs-3955	264	14	facilitating	facilitate	VERB
gs-3955	264	15	rapid	rapid	ADJ
gs-3955	264	16	tissue	tissue	NOUN
gs-3955	264	17	renewal	renewal	NOUN
gs-3955	264	18	and	and	CCONJ
gs-3955	264	19	whole	whole	ADJ
gs-3955	264	20	-	-	PUNCT
gs-3955	264	21	body	body	NOUN
gs-3955	264	22	regeneration	regeneration	NOUN
gs-3955	264	23	without	without	ADP
gs-3955	264	24	accumulation	accumulation	NOUN
gs-3955	264	25	of	of	ADP
gs-3955	264	26	cellular	cellular	ADJ
gs-3955	264	27	debris	debris	NOUN
gs-3955	264	28	(	(	PUNCT
gs-3955	264	29	pirotte	pirotte	NOUN
gs-3955	264	30	et	et	PROPN
gs-3955	264	31	al	al	PROPN
gs-3955	264	32	.	.	PROPN
gs-3955	264	33	,	,	PUNCT
gs-3955	264	34	2015	2015	NUM
gs-3955	264	35	)	)	PUNCT
gs-3955	264	36	.	.	PUNCT
gs-3955	265	1	autophagy	autophagy	VERB
gs-3955	265	2	-	-	PUNCT
gs-3955	265	3	related	relate	VERB
gs-3955	265	4	proteins	protein	NOUN
gs-3955	265	5	such	such	ADJ
gs-3955	265	6	as	as	ADP
gs-3955	265	7	lc3	lc3	NOUN
gs-3955	265	8	are	be	AUX
gs-3955	265	9	highly	highly	ADV
gs-3955	265	10	expressed	express	VERB
gs-3955	265	11	in	in	ADP
gs-3955	265	12	budding	bud	VERB
gs-3955	265	13	zones	zone	NOUN
gs-3955	265	14	,	,	PUNCT
gs-3955	265	15	indicating	indicate	VERB
gs-3955	265	16	active	active	ADJ
gs-3955	265	17	recycling	recycling	NOUN
gs-3955	265	18	of	of	ADP
gs-3955	265	19	cytoplasmic	cytoplasmic	ADJ
gs-3955	265	20	components	component	NOUN
gs-3955	265	21	to	to	PART
gs-3955	265	22	fuel	fuel	VERB
gs-3955	265	23	growth	growth	NOUN
gs-3955	265	24	and	and	CCONJ
gs-3955	265	25	morphogenic	morphogenic	ADJ
gs-3955	265	26	remodeling	remodeling	NOUN
gs-3955	265	27	(	(	PUNCT
gs-3955	265	28	pirotte	pirotte	NOUN
gs-3955	265	29	et	et	PROPN
gs-3955	265	30	al	al	PROPN
gs-3955	265	31	.	.	PROPN
gs-3955	265	32	,	,	PUNCT
gs-3955	265	33	2015	2015	NUM
gs-3955	265	34	;	;	PUNCT
gs-3955	265	35	tkemaladze	tkemaladze	NOUN
gs-3955	265	36	,	,	PUNCT
gs-3955	265	37	2023	2023	NUM
gs-3955	265	38	)	)	PUNCT
gs-3955	265	39	.	.	PUNCT
gs-3955	266	1	similarly	similarly	ADV
gs-3955	266	2	,	,	PUNCT
gs-3955	266	3	planarians	planarian	NOUN
gs-3955	266	4	(	(	PUNCT
gs-3955	266	5	e.g.	e.g.	ADV
gs-3955	266	6	,	,	PUNCT
gs-3955	266	7	schmidtea	schmidtea	PROPN
gs-3955	266	8	mediterranea	mediterranea	PROPN
gs-3955	266	9	)	)	PUNCT
gs-3955	266	10	rely	rely	VERB
gs-3955	266	11	on	on	ADP
gs-3955	266	12	autophagy	autophagy	NOUN
gs-3955	266	13	for	for	ADP
gs-3955	266	14	neoblast	neoblast	NOUN
gs-3955	266	15	function	function	NOUN
gs-3955	266	16	—	—	PUNCT
gs-3955	266	17	the	the	DET
gs-3955	266	18	pluripotent	pluripotent	ADJ
gs-3955	266	19	adult	adult	NOUN
gs-3955	266	20	stem	stem	NOUN
gs-3955	266	21	cells	cell	NOUN
gs-3955	266	22	responsible	responsible	ADJ
gs-3955	266	23	for	for	ADP
gs-3955	266	24	regenerating	regenerate	VERB
gs-3955	266	25	lost	lose	VERB
gs-3955	266	26	body	body	NOUN
gs-3955	266	27	parts	part	NOUN
gs-3955	266	28	.	.	PUNCT
gs-3955	267	1	autophagic	autophagic	PROPN
gs-3955	267	2	pathways	pathway	NOUN
gs-3955	267	3	regulate	regulate	VERB
gs-3955	267	4	energy	energy	NOUN
gs-3955	267	5	homeostasis	homeostasis	NOUN
gs-3955	267	6	and	and	CCONJ
gs-3955	267	7	protein	protein	NOUN
gs-3955	267	8	turnover	turnover	NOUN
gs-3955	267	9	during	during	ADP
gs-3955	267	10	regeneration	regeneration	NOUN
gs-3955	267	11	,	,	PUNCT
gs-3955	267	12	ensuring	ensure	VERB
gs-3955	267	13	proper	proper	ADJ
gs-3955	267	14	stem	stem	NOUN
gs-3955	267	15	cell	cell	NOUN
gs-3955	267	16	maintenance	maintenance	NOUN
gs-3955	267	17	and	and	CCONJ
gs-3955	267	18	differentiation	differentiation	NOUN
gs-3955	267	19	(	(	PUNCT
gs-3955	267	20	wagner	wagner	PROPN
gs-3955	267	21	et	et	PROPN
gs-3955	267	22	al	al	PROPN
gs-3955	267	23	.	.	PROPN
gs-3955	267	24	,	,	PUNCT
gs-3955	267	25	2011	2011	NUM
gs-3955	267	26	;	;	PUNCT
gs-3955	267	27	rink	rink	NOUN
gs-3955	267	28	,	,	PUNCT
gs-3955	267	29	2013	2013	NUM
gs-3955	267	30	)	)	PUNCT
gs-3955	267	31	.	.	PUNCT
gs-3955	268	1	in	in	ADP
gs-3955	268	2	vertebrate	vertebrate	ADJ
gs-3955	268	3	species	specie	NOUN
gs-3955	268	4	with	with	ADP
gs-3955	268	5	remarkable	remarkable	ADJ
gs-3955	268	6	regenerative	regenerative	ADJ
gs-3955	268	7	capacity	capacity	NOUN
gs-3955	268	8	such	such	ADJ
gs-3955	268	9	as	as	ADP
gs-3955	268	10	fire	fire	NOUN
gs-3955	268	11	salamanders	salamander	NOUN
gs-3955	268	12	(	(	PUNCT
gs-3955	268	13	salamandra	salamandra	NOUN
gs-3955	268	14	salamandra	salamandra	NOUN
gs-3955	268	15	)	)	PUNCT
gs-3955	268	16	,	,	PUNCT
gs-3955	268	17	autophagy	autophagy	NOUN
gs-3955	268	18	is	be	AUX
gs-3955	268	19	activated	activate	VERB
gs-3955	268	20	in	in	ADP
gs-3955	268	21	response	response	NOUN
gs-3955	268	22	to	to	ADP
gs-3955	268	23	spinal	spinal	ADJ
gs-3955	268	24	cord	cord	NOUN
gs-3955	268	25	injury	injury	NOUN
gs-3955	268	26	,	,	PUNCT
gs-3955	268	27	contributing	contribute	VERB
gs-3955	268	28	to	to	ADP
gs-3955	268	29	neuronal	neuronal	ADJ
gs-3955	268	30	survival	survival	NOUN
gs-3955	268	31	and	and	CCONJ
gs-3955	268	32	axonal	axonal	ADJ
gs-3955	268	33	regrowth	regrowth	NOUN
gs-3955	268	34	.	.	PUNCT
gs-3955	269	1	enhanced	enhance	VERB
gs-3955	269	2	mitophagy	mitophagy	NOUN
gs-3955	269	3	in	in	ADP
gs-3955	269	4	particular	particular	ADJ
gs-3955	269	5	clears	clear	VERB
gs-3955	269	6	dysfunctional	dysfunctional	ADJ
gs-3955	269	7	mitochondria	mitochondria	NOUN
gs-3955	269	8	,	,	PUNCT
gs-3955	269	9	preventing	prevent	VERB
gs-3955	269	10	apoptosis	apoptosis	NOUN
gs-3955	269	11	and	and	CCONJ
gs-3955	269	12	supporting	support	VERB
gs-3955	269	13	tissue	tissue	NOUN
gs-3955	269	14	repair	repair	NOUN
gs-3955	269	15	(	(	PUNCT
gs-3955	269	16	narayan	narayan	PROPN
gs-3955	269	17	et	et	PROPN
gs-3955	269	18	al	al	PROPN
gs-3955	269	19	.	.	PROPN
gs-3955	269	20	,	,	PUNCT
gs-3955	269	21	2021	2021	NUM
gs-3955	269	22	)	)	PUNCT
gs-3955	269	23	.	.	PUNCT
gs-3955	270	1	autophagy	autophagy	VERB
gs-3955	270	2	in	in	ADP
gs-3955	270	3	stress	stress	NOUN
gs-3955	270	4	resistance	resistance	NOUN
gs-3955	270	5	and	and	CCONJ
gs-3955	270	6	metabolic	metabolic	NOUN
gs-3955	270	7	adaptation	adaptation	NOUN
gs-3955	270	8	long	long	ADV
gs-3955	270	9	-	-	PUNCT
gs-3955	270	10	lived	live	VERB
gs-3955	270	11	species	specie	NOUN
gs-3955	270	12	often	often	ADV
gs-3955	270	13	inhabit	inhabit	AUX
gs-3955	270	14	extreme	extreme	ADJ
gs-3955	270	15	or	or	CCONJ
gs-3955	270	16	fluctuating	fluctuate	VERB
gs-3955	270	17	environments	environment	NOUN
gs-3955	270	18	,	,	PUNCT
gs-3955	270	19	and	and	CCONJ
gs-3955	270	20	autophagy	autophagy	NOUN
gs-3955	270	21	is	be	AUX
gs-3955	270	22	crucial	crucial	ADJ
gs-3955	270	23	for	for	ADP
gs-3955	270	24	adapting	adapt	VERB
gs-3955	270	25	to	to	ADP
gs-3955	270	26	such	such	ADJ
gs-3955	270	27	stressors	stressor	NOUN
gs-3955	270	28	.	.	PUNCT
gs-3955	271	1	the	the	DET
gs-3955	271	2	ocean	ocean	PROPN
gs-3955	271	3	quahog	quahog	PROPN
gs-3955	271	4	(	(	PUNCT
gs-3955	271	5	arctica	arctica	PROPN
gs-3955	271	6	islandica	islandica	PROPN
gs-3955	271	7	)	)	PUNCT
gs-3955	271	8	demonstrates	demonstrate	VERB
gs-3955	271	9	exceptional	exceptional	ADJ
gs-3955	271	10	tolerance	tolerance	NOUN
gs-3955	271	11	to	to	PART
gs-3955	271	12	hypoxia	hypoxia	VERB
gs-3955	271	13	and	and	CCONJ
gs-3955	271	14	temperature	temperature	NOUN
gs-3955	271	15	variation	variation	NOUN
gs-3955	271	16	,	,	PUNCT
gs-3955	271	17	partly	partly	ADV
gs-3955	271	18	due	due	ADJ
gs-3955	271	19	to	to	ADP
gs-3955	271	20	efficient	efficient	ADJ
gs-3955	271	21	autophagic	autophagic	ADJ
gs-3955	271	22	degradation	degradation	NOUN
gs-3955	271	23	pathways	pathway	NOUN
gs-3955	271	24	that	that	PRON
gs-3955	271	25	clear	clear	ADJ
gs-3955	271	26	damaged	damaged	ADJ
gs-3955	271	27	proteins	protein	NOUN
gs-3955	271	28	and	and	CCONJ
gs-3955	271	29	organelles	organelle	NOUN
gs-3955	271	30	during	during	ADP
gs-3955	271	31	metabolic	metabolic	NOUN
gs-3955	271	32	depression	depression	NOUN
gs-3955	271	33	(	(	PUNCT
gs-3955	271	34	munro	munro	PROPN
gs-3955	271	35	et	et	PROPN
gs-3955	271	36	al	al	PROPN
gs-3955	271	37	.	.	PROPN
gs-3955	271	38	,	,	PUNCT
gs-3955	271	39	2012	2012	NUM
gs-3955	271	40	)	)	PUNCT
gs-3955	271	41	.	.	PUNCT
gs-3955	272	1	autophagy	autophagy	PROPN
gs-3955	272	2	contributes	contribute	VERB
gs-3955	272	3	to	to	ADP
gs-3955	272	4	cellular	cellular	ADJ
gs-3955	272	5	energy	energy	NOUN
gs-3955	272	6	conservation	conservation	NOUN
gs-3955	272	7	by	by	ADP
gs-3955	272	8	recycling	recycle	VERB
gs-3955	272	9	macromolecules	macromolecule	NOUN
gs-3955	272	10	under	under	ADP
gs-3955	272	11	nutrient	nutrient	NOUN
gs-3955	272	12	-	-	PUNCT
gs-3955	272	13	limiting	limit	VERB
gs-3955	272	14	conditions	condition	NOUN
gs-3955	272	15	typical	typical	ADJ
gs-3955	272	16	of	of	ADP
gs-3955	272	17	their	their	PRON
gs-3955	272	18	benthic	benthic	ADJ
gs-3955	272	19	habitats	habitat	NOUN
gs-3955	272	20	(	(	PUNCT
gs-3955	272	21	narayan	narayan	NOUN
gs-3955	272	22	et	et	PROPN
gs-3955	272	23	al	al	PROPN
gs-3955	272	24	.	.	PROPN
gs-3955	272	25	,	,	PUNCT
gs-3955	272	26	2021	2021	NUM
gs-3955	272	27	)	)	PUNCT
gs-3955	272	28	.	.	PUNCT
gs-3955	273	1	naked	naked	ADJ
gs-3955	273	2	mole	mole	NOUN
gs-3955	273	3	-	-	PUNCT
gs-3955	273	4	rats	rat	NOUN
gs-3955	273	5	show	show	VERB
gs-3955	273	6	augmented	augment	VERB
gs-3955	273	7	autophagic	autophagic	ADJ
gs-3955	273	8	activity	activity	NOUN
gs-3955	273	9	under	under	ADP
gs-3955	273	10	oxidative	oxidative	ADJ
gs-3955	273	11	and	and	CCONJ
gs-3955	273	12	metabolic	metabolic	NOUN
gs-3955	273	13	stress	stress	NOUN
gs-3955	273	14	,	,	PUNCT
gs-3955	273	15	enhancing	enhance	VERB
gs-3955	273	16	the	the	DET
gs-3955	273	17	removal	removal	NOUN
gs-3955	273	18	of	of	ADP
gs-3955	273	19	oxidized	oxidized	ADJ
gs-3955	273	20	proteins	protein	NOUN
gs-3955	273	21	and	and	CCONJ
gs-3955	273	22	damaged	damage	VERB
gs-3955	273	23	mitochondria	mitochondria	NOUN
gs-3955	273	24	(	(	PUNCT
gs-3955	273	25	macrae	macrae	PROPN
gs-3955	273	26	et	et	PROPN
gs-3955	273	27	al	al	PROPN
gs-3955	273	28	.	.	PROPN
gs-3955	273	29	,	,	PUNCT
gs-3955	273	30	georgian	georgian	PROPN
gs-3955	273	31	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	273	32	მეცნიერები	მეცნიერები	NOUN
gs-3955	273	33	ტ.	ტ.	VERB
gs-3955	273	34	7	7	NUM
gs-3955	273	35	n	n	DET
gs-3955	273	36	3	3	NUM
gs-3955	273	37	,	,	PUNCT
gs-3955	273	38	2025	2025	NUM
gs-3955	273	39	380	380	NUM
gs-3955	273	40	2015	2015	NUM
gs-3955	273	41	;	;	PUNCT
gs-3955	273	42	lewis	lewis	PROPN
gs-3955	273	43	et	et	PROPN
gs-3955	273	44	al	al	PROPN
gs-3955	273	45	.	.	PROPN
gs-3955	273	46	,	,	PUNCT
gs-3955	273	47	2012	2012	NUM
gs-3955	273	48	)	)	PUNCT
gs-3955	273	49	.	.	PUNCT
gs-3955	274	1	this	this	DET
gs-3955	274	2	process	process	NOUN
gs-3955	274	3	mitigates	mitigate	VERB
gs-3955	274	4	the	the	DET
gs-3955	274	5	detrimental	detrimental	ADJ
gs-3955	274	6	effects	effect	NOUN
gs-3955	274	7	of	of	ADP
gs-3955	274	8	reactive	reactive	ADJ
gs-3955	274	9	oxygen	oxygen	NOUN
gs-3955	274	10	species	specie	NOUN
gs-3955	274	11	(	(	PUNCT
gs-3955	274	12	ros	ros	PROPN
gs-3955	274	13	)	)	PUNCT
gs-3955	274	14	while	while	SCONJ
gs-3955	274	15	supporting	support	VERB
gs-3955	274	16	metabolic	metabolic	NOUN
gs-3955	274	17	flexibility	flexibility	NOUN
gs-3955	274	18	during	during	ADP
gs-3955	274	19	hypoxia	hypoxia	NOUN
gs-3955	274	20	or	or	CCONJ
gs-3955	274	21	calorie	calorie	NOUN
gs-3955	274	22	restriction	restriction	NOUN
gs-3955	274	23	(	(	PUNCT
gs-3955	274	24	petruseva	petruseva	PROPN
gs-3955	274	25	et	et	PROPN
gs-3955	274	26	al	al	PROPN
gs-3955	274	27	.	.	PROPN
gs-3955	274	28	,	,	PUNCT
gs-3955	274	29	2017	2017	NUM
gs-3955	274	30	)	)	PUNCT
gs-3955	274	31	.	.	PUNCT
gs-3955	275	1	autophagy	autophagy	ADJ
gs-3955	275	2	and	and	CCONJ
gs-3955	275	3	immune	immune	ADJ
gs-3955	275	4	function	function	NOUN
gs-3955	275	5	emerging	emerge	VERB
gs-3955	275	6	evidence	evidence	NOUN
gs-3955	275	7	links	link	NOUN
gs-3955	275	8	autophagy	autophagy	ADV
gs-3955	275	9	to	to	ADP
gs-3955	275	10	immune	immune	ADJ
gs-3955	275	11	system	system	NOUN
gs-3955	275	12	regulation	regulation	NOUN
gs-3955	275	13	in	in	ADP
gs-3955	275	14	these	these	DET
gs-3955	275	15	species	specie	NOUN
gs-3955	275	16	.	.	PUNCT
gs-3955	276	1	for	for	ADP
gs-3955	276	2	example	example	NOUN
gs-3955	276	3	,	,	PUNCT
gs-3955	276	4	naked	naked	ADJ
gs-3955	276	5	mole	mole	NOUN
gs-3955	276	6	-	-	PUNCT
gs-3955	276	7	rats	rat	NOUN
gs-3955	276	8	’	'	PUNCT
gs-3955	276	9	robust	robust	ADJ
gs-3955	276	10	innate	innate	ADJ
gs-3955	276	11	immunity	immunity	NOUN
gs-3955	276	12	is	be	AUX
gs-3955	276	13	partly	partly	ADV
gs-3955	276	14	attributed	attribute	VERB
gs-3955	276	15	to	to	ADP
gs-3955	276	16	autophagic	autophagic	ADJ
gs-3955	276	17	mechanisms	mechanism	NOUN
gs-3955	276	18	that	that	PRON
gs-3955	276	19	mediate	mediate	VERB
gs-3955	276	20	pathogen	pathogen	NOUN
gs-3955	276	21	clearance	clearance	NOUN
gs-3955	276	22	and	and	CCONJ
gs-3955	276	23	antigen	antigen	NOUN
gs-3955	276	24	presentation	presentation	NOUN
gs-3955	276	25	,	,	PUNCT
gs-3955	276	26	contributing	contribute	VERB
gs-3955	276	27	to	to	ADP
gs-3955	276	28	their	their	PRON
gs-3955	276	29	low	low	ADJ
gs-3955	276	30	incidence	incidence	NOUN
gs-3955	276	31	of	of	ADP
gs-3955	276	32	inflammatory	inflammatory	ADJ
gs-3955	276	33	diseases	disease	NOUN
gs-3955	276	34	and	and	CCONJ
gs-3955	276	35	cancer	cancer	NOUN
gs-3955	276	36	(	(	PUNCT
gs-3955	276	37	narayan	narayan	PROPN
gs-3955	276	38	et	et	PROPN
gs-3955	276	39	al	al	PROPN
gs-3955	276	40	.	.	PROPN
gs-3955	276	41	,	,	PUNCT
gs-3955	276	42	2021	2021	NUM
gs-3955	276	43	)	)	PUNCT
gs-3955	276	44	.	.	PUNCT
gs-3955	277	1	greenland	greenland	PROPN
gs-3955	277	2	sharks	sharks	PROPN
gs-3955	277	3	(	(	PUNCT
gs-3955	277	4	somniosus	somniosus	PROPN
gs-3955	277	5	microcephalus	microcephalus	PROPN
gs-3955	277	6	)	)	PUNCT
gs-3955	277	7	maintain	maintain	VERB
gs-3955	277	8	immune	immune	ADJ
gs-3955	277	9	homeostasis	homeostasis	NOUN
gs-3955	277	10	over	over	ADP
gs-3955	277	11	centuries	century	NOUN
gs-3955	277	12	,	,	PUNCT
gs-3955	277	13	possibly	possibly	ADV
gs-3955	277	14	supported	support	VERB
gs-3955	277	15	by	by	ADP
gs-3955	277	16	autophagic	autophagic	ADJ
gs-3955	277	17	degradation	degradation	NOUN
gs-3955	277	18	of	of	ADP
gs-3955	277	19	intracellular	intracellular	ADJ
gs-3955	277	20	pathogens	pathogen	NOUN
gs-3955	277	21	and	and	CCONJ
gs-3955	277	22	regulation	regulation	NOUN
gs-3955	277	23	of	of	ADP
gs-3955	277	24	inflammatory	inflammatory	ADJ
gs-3955	277	25	signaling	signaling	NOUN
gs-3955	277	26	,	,	PUNCT
gs-3955	277	27	although	although	SCONJ
gs-3955	277	28	direct	direct	ADJ
gs-3955	277	29	studies	study	NOUN
gs-3955	277	30	remain	remain	VERB
gs-3955	277	31	sparse	sparse	ADJ
gs-3955	277	32	(	(	PUNCT
gs-3955	277	33	tkemaladze	tkemaladze	NOUN
gs-3955	277	34	,	,	PUNCT
gs-3955	277	35	2023	2023	NUM
gs-3955	277	36	)	)	PUNCT
gs-3955	277	37	.	.	PUNCT
gs-3955	278	1	molecular	molecular	ADJ
gs-3955	278	2	and	and	CCONJ
gs-3955	278	3	genetic	genetic	ADJ
gs-3955	278	4	regulation	regulation	NOUN
gs-3955	278	5	across	across	ADP
gs-3955	278	6	negligibly	negligibly	ADV
gs-3955	278	7	senescent	senescent	ADJ
gs-3955	278	8	species	specie	NOUN
gs-3955	278	9	,	,	PUNCT
gs-3955	278	10	key	key	ADJ
gs-3955	278	11	autophagy	autophagy	ADJ
gs-3955	278	12	regulators	regulator	NOUN
gs-3955	278	13	such	such	ADJ
gs-3955	278	14	as	as	ADP
gs-3955	278	15	beclin1	beclin1	NOUN
gs-3955	278	16	,	,	PUNCT
gs-3955	278	17	lc3	lc3	NOUN
gs-3955	278	18	,	,	PUNCT
gs-3955	278	19	and	and	CCONJ
gs-3955	278	20	p62	p62	NOUN
gs-3955	278	21	/	/	SYM
gs-3955	278	22	sqstm1	sqstm1	NOUN
gs-3955	278	23	show	show	NOUN
gs-3955	278	24	sustained	sustain	VERB
gs-3955	278	25	expression	expression	NOUN
gs-3955	278	26	levels	level	NOUN
gs-3955	278	27	regardless	regardless	ADV
gs-3955	278	28	of	of	ADP
gs-3955	278	29	chronological	chronological	ADJ
gs-3955	278	30	age	age	NOUN
gs-3955	278	31	,	,	PUNCT
gs-3955	278	32	contrasting	contrast	VERB
gs-3955	278	33	with	with	ADP
gs-3955	278	34	the	the	DET
gs-3955	278	35	age	age	NOUN
gs-3955	278	36	-	-	PUNCT
gs-3955	278	37	related	relate	VERB
gs-3955	278	38	decline	decline	NOUN
gs-3955	278	39	seen	see	VERB
gs-3955	278	40	in	in	ADP
gs-3955	278	41	typical	typical	ADJ
gs-3955	278	42	mammals	mammal	NOUN
gs-3955	278	43	(	(	PUNCT
gs-3955	278	44	pirotte	pirotte	NOUN
gs-3955	278	45	et	et	PROPN
gs-3955	278	46	al	al	PROPN
gs-3955	278	47	.	.	PROPN
gs-3955	278	48	,	,	PUNCT
gs-3955	278	49	2015	2015	NUM
gs-3955	278	50	;	;	PUNCT
gs-3955	278	51	narayan	narayan	PROPN
gs-3955	278	52	et	et	PROPN
gs-3955	278	53	al	al	PROPN
gs-3955	278	54	.	.	PROPN
gs-3955	278	55	,	,	PUNCT
gs-3955	278	56	2021	2021	NUM
gs-3955	278	57	)	)	PUNCT
gs-3955	278	58	.	.	PUNCT
gs-3955	279	1	specialized	specialized	ADJ
gs-3955	279	2	autophagy	autophagy	ADJ
gs-3955	279	3	variants	variant	NOUN
gs-3955	279	4	are	be	AUX
gs-3955	279	5	evident	evident	ADJ
gs-3955	279	6	as	as	ADV
gs-3955	279	7	well	well	ADV
gs-3955	279	8	:	:	PUNCT
gs-3955	279	9	naked	naked	ADJ
gs-3955	279	10	mole	mole	NOUN
gs-3955	279	11	-	-	PUNCT
gs-3955	279	12	rats	rat	NOUN
gs-3955	279	13	utilize	utilize	VERB
gs-3955	279	14	pink1	pink1	NOUN
gs-3955	279	15	/	/	SYM
gs-3955	279	16	parkin	parkin	NOUN
gs-3955	279	17	-	-	PUNCT
gs-3955	279	18	independent	independent	ADJ
gs-3955	279	19	mitophagy	mitophagy	NOUN
gs-3955	279	20	pathways	pathway	NOUN
gs-3955	279	21	,	,	PUNCT
gs-3955	279	22	while	while	SCONJ
gs-3955	279	23	rockfish	rockfish	NOUN
gs-3955	279	24	express	express	VERB
gs-3955	279	25	aggrephagy	aggrephagy	ADJ
gs-3955	279	26	factors	factor	NOUN
gs-3955	279	27	with	with	ADP
gs-3955	279	28	expanded	expand	VERB
gs-3955	279	29	lc3	lc3	ADV
gs-3955	279	30	-	-	PUNCT
gs-3955	279	31	interacting	interact	VERB
gs-3955	279	32	regions	region	NOUN
gs-3955	279	33	,	,	PUNCT
gs-3955	279	34	enhancing	enhance	VERB
gs-3955	279	35	selective	selective	ADJ
gs-3955	279	36	clearance	clearance	NOUN
gs-3955	279	37	of	of	ADP
gs-3955	279	38	protein	protein	NOUN
gs-3955	279	39	aggregates	aggregate	NOUN
gs-3955	279	40	(	(	PUNCT
gs-3955	279	41	macrae	macrae	PROPN
gs-3955	279	42	et	et	PROPN
gs-3955	279	43	al	al	PROPN
gs-3955	279	44	.	.	PROPN
gs-3955	279	45	,	,	PUNCT
gs-3955	279	46	2015	2015	NUM
gs-3955	279	47	;	;	PUNCT
gs-3955	279	48	petruseva	petruseva	VERB
gs-3955	279	49	et	et	PROPN
gs-3955	279	50	al	al	PROPN
gs-3955	279	51	.	.	PROPN
gs-3955	279	52	,	,	PUNCT
gs-3955	279	53	2017	2017	NUM
gs-3955	279	54	)	)	PUNCT
gs-3955	279	55	.	.	PUNCT
gs-3955	280	1	temperature	temperature	NOUN
gs-3955	280	2	-	-	PUNCT
gs-3955	280	3	insensitive	insensitive	ADJ
gs-3955	280	4	chaperones	chaperone	NOUN
gs-3955	280	5	in	in	ADP
gs-3955	280	6	arctica	arctica	PROPN
gs-3955	280	7	islandica	islandica	PROPN
gs-3955	280	8	cooperate	cooperate	VERB
gs-3955	280	9	with	with	ADP
gs-3955	280	10	autophagic	autophagic	ADJ
gs-3955	280	11	pathways	pathway	NOUN
gs-3955	280	12	to	to	PART
gs-3955	280	13	maintain	maintain	VERB
gs-3955	280	14	proteostasis	proteostasis	NOUN
gs-3955	280	15	under	under	ADP
gs-3955	280	16	environmental	environmental	ADJ
gs-3955	280	17	stress	stress	NOUN
gs-3955	280	18	,	,	PUNCT
gs-3955	280	19	reflecting	reflect	VERB
gs-3955	280	20	an	an	DET
gs-3955	280	21	evolutionary	evolutionary	ADJ
gs-3955	280	22	integration	integration	NOUN
gs-3955	280	23	of	of	ADP
gs-3955	280	24	protein	protein	NOUN
gs-3955	280	25	folding	folding	NOUN
gs-3955	280	26	and	and	CCONJ
gs-3955	280	27	degradation	degradation	NOUN
gs-3955	280	28	systems	system	NOUN
gs-3955	280	29	(	(	PUNCT
gs-3955	280	30	munro	munro	PROPN
gs-3955	280	31	et	et	PROPN
gs-3955	280	32	al	al	PROPN
gs-3955	280	33	.	.	PROPN
gs-3955	280	34	,	,	PUNCT
gs-3955	280	35	2012	2012	NUM
gs-3955	280	36	)	)	PUNCT
gs-3955	280	37	.	.	PUNCT
gs-3955	281	1	this	this	DET
gs-3955	281	2	multi	multi	ADJ
gs-3955	281	3	-	-	ADJ
gs-3955	281	4	angle	angle	ADJ
gs-3955	281	5	perspective	perspective	ADJ
gs-3955	281	6	highlights	highlight	NOUN
gs-3955	281	7	that	that	PRON
gs-3955	281	8	autophagy	autophagy	VERB
gs-3955	281	9	in	in	ADP
gs-3955	281	10	negligibly	negligibly	ADV
gs-3955	281	11	senescent	senescent	ADJ
gs-3955	281	12	organisms	organism	NOUN
gs-3955	281	13	extends	extend	VERB
gs-3955	281	14	well	well	ADV
gs-3955	281	15	beyond	beyond	ADP
gs-3955	281	16	aging	aging	NOUN
gs-3955	281	17	,	,	PUNCT
gs-3955	281	18	encompassing	encompass	VERB
gs-3955	281	19	development	development	NOUN
gs-3955	281	20	,	,	PUNCT
gs-3955	281	21	regeneration	regeneration	NOUN
gs-3955	281	22	,	,	PUNCT
gs-3955	281	23	stress	stress	NOUN
gs-3955	281	24	adaptation	adaptation	NOUN
gs-3955	281	25	,	,	PUNCT
gs-3955	281	26	immunity	immunity	NOUN
gs-3955	281	27	,	,	PUNCT
gs-3955	281	28	and	and	CCONJ
gs-3955	281	29	environmental	environmental	ADJ
gs-3955	281	30	resilience	resilience	NOUN
gs-3955	281	31	.	.	PUNCT
gs-3955	282	1	these	these	DET
gs-3955	282	2	broader	broad	ADJ
gs-3955	282	3	roles	role	NOUN
gs-3955	282	4	reinforce	reinforce	VERB
gs-3955	282	5	autophagy	autophagy	NOUN
gs-3955	282	6	’s	’s	PART
gs-3955	282	7	centrality	centrality	NOUN
gs-3955	282	8	as	as	ADP
gs-3955	282	9	a	a	DET
gs-3955	282	10	versatile	versatile	ADJ
gs-3955	282	11	biological	biological	ADJ
gs-3955	282	12	process	process	NOUN
gs-3955	282	13	supporting	support	VERB
gs-3955	282	14	both	both	DET
gs-3955	282	15	longevity	longevity	NOUN
gs-3955	282	16	and	and	CCONJ
gs-3955	282	17	ecological	ecological	ADJ
gs-3955	282	18	fitness	fitness	NOUN
gs-3955	282	19	.	.	PUNCT
gs-3955	283	1	toward	toward	ADP
gs-3955	283	2	a	a	DET
gs-3955	283	3	post	post	ADJ
gs-3955	283	4	-	-	ADJ
gs-3955	283	5	aging	aging	ADJ
gs-3955	283	6	paradigm	paradigm	NOUN
gs-3955	283	7	negligibly	negligibly	ADV
gs-3955	283	8	senescent	senescent	ADJ
gs-3955	283	9	species	specie	NOUN
gs-3955	283	10	embody	embody	VERB
gs-3955	283	11	nature	nature	NOUN
gs-3955	283	12	's	's	PART
gs-3955	283	13	most	most	ADV
gs-3955	283	14	sophisticated	sophisticated	ADJ
gs-3955	283	15	experiments	experiment	NOUN
gs-3955	283	16	in	in	ADP
gs-3955	283	17	biological	biological	ADJ
gs-3955	283	18	time	time	NOUN
gs-3955	283	19	manipulation	manipulation	NOUN
gs-3955	283	20	.	.	PUNCT
gs-3955	284	1	their	their	PRON
gs-3955	284	2	molecular	molecular	ADJ
gs-3955	284	3	strategies	strategy	NOUN
gs-3955	284	4	—	—	PUNCT
gs-3955	284	5	from	from	ADP
gs-3955	284	6	quantum	quantum	NOUN
gs-3955	284	7	-	-	PUNCT
gs-3955	284	8	efficient	efficient	ADJ
gs-3955	284	9	electron	electron	NOUN
gs-3955	284	10	transport	transport	NOUN
gs-3955	284	11	in	in	ADP
gs-3955	284	12	shark	shark	NOUN
gs-3955	284	13	mitochondria	mitochondria	NOUN
gs-3955	284	14	to	to	ADP
gs-3955	284	15	error	error	NOUN
gs-3955	284	16	-	-	PUNCT
gs-3955	284	17	proof	proof	ADJ
gs-3955	284	18	dna	dna	PROPN
gs-3955	284	19	polymerases	polymerase	NOUN
gs-3955	284	20	in	in	ADP
gs-3955	284	21	clams	clams	NOUN
gs-3955	284	22	—	—	PUNCT
gs-3955	284	23	reveal	reveal	VERB
gs-3955	284	24	aging	aging	NOUN
gs-3955	284	25	as	as	ADP
gs-3955	284	26	a	a	DET
gs-3955	284	27	tractable	tractable	ADJ
gs-3955	284	28	engineering	engineering	NOUN
gs-3955	284	29	challenge	challenge	NOUN
gs-3955	284	30	rather	rather	ADV
gs-3955	284	31	than	than	ADP
gs-3955	284	32	an	an	DET
gs-3955	284	33	immutable	immutable	ADJ
gs-3955	284	34	biological	biological	ADJ
gs-3955	284	35	constant	constant	ADJ
gs-3955	284	36	(	(	PUNCT
gs-3955	284	37	tkemaladze	tkemaladze	NOUN
gs-3955	284	38	,	,	PUNCT
gs-3955	284	39	2023	2023	NUM
gs-3955	284	40	;	;	PUNCT
gs-3955	284	41	kumar	kumar	PROPN
gs-3955	284	42	et	et	PROPN
gs-3955	284	43	al	al	PROPN
gs-3955	284	44	.	.	PROPN
gs-3955	284	45	,	,	PUNCT
gs-3955	284	46	2021	2021	NUM
gs-3955	284	47	)	)	PUNCT
gs-3955	284	48	.	.	PUNCT
gs-3955	285	1	the	the	DET
gs-3955	285	2	centrosomal	centrosomal	ADJ
gs-3955	285	3	theory	theory	NOUN
gs-3955	285	4	provides	provide	VERB
gs-3955	285	5	a	a	DET
gs-3955	285	6	physicochemical	physicochemical	ADJ
gs-3955	285	7	framework	framework	NOUN
gs-3955	285	8	unifying	unify	VERB
gs-3955	285	9	these	these	DET
gs-3955	285	10	adaptations	adaptation	NOUN
gs-3955	285	11	:	:	PUNCT
gs-3955	285	12	by	by	ADP
gs-3955	285	13	minimizing	minimize	VERB
gs-3955	285	14	entropic	entropic	ADJ
gs-3955	285	15	accumulation	accumulation	NOUN
gs-3955	285	16	in	in	ADP
gs-3955	285	17	cellular	cellular	ADJ
gs-3955	285	18	control	control	NOUN
gs-3955	285	19	centers	center	NOUN
gs-3955	285	20	,	,	PUNCT
gs-3955	285	21	these	these	DET
gs-3955	285	22	species	specie	NOUN
gs-3955	285	23	achieve	achieve	VERB
gs-3955	285	24	near	near	ADV
gs-3955	285	25	-	-	PUNCT
gs-3955	285	26	perfect	perfect	ADJ
gs-3955	285	27	self	self	NOUN
gs-3955	285	28	-	-	PUNCT
gs-3955	285	29	renewal	renewal	NOUN
gs-3955	285	30	(	(	PUNCT
gs-3955	285	31	tkemaladze	tkemaladze	NOUN
gs-3955	285	32	,	,	PUNCT
gs-3955	285	33	2023	2023	NUM
gs-3955	285	34	-	-	SYM
gs-3955	285	35	2024	2024	NUM
gs-3955	285	36	)	)	PUNCT
gs-3955	285	37	.	.	PUNCT
gs-3955	286	1	georgian	georgian	ADJ
gs-3955	286	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	286	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	286	4	ტ.	ტ.	VERB
gs-3955	286	5	7	7	NUM
gs-3955	286	6	n	n	DET
gs-3955	286	7	3	3	NUM
gs-3955	286	8	,	,	PUNCT
gs-3955	286	9	2025	2025	NUM
gs-3955	286	10	381	381	NUM
gs-3955	286	11	the	the	DET
gs-3955	286	12	translational	translational	ADJ
gs-3955	286	13	roadmap	roadmap	NOUN
gs-3955	286	14	demands	demand	VERB
gs-3955	286	15	cautious	cautious	ADJ
gs-3955	286	16	optimism	optimism	NOUN
gs-3955	286	17	.	.	PUNCT
gs-3955	287	1	while	while	SCONJ
gs-3955	287	2	hmw	hmw	PROPN
gs-3955	287	3	-	-	PUNCT
gs-3955	287	4	ha	ha	INTJ
gs-3955	287	5	cancer	cancer	NOUN
gs-3955	287	6	therapies	therapy	NOUN
gs-3955	287	7	and	and	CCONJ
gs-3955	287	8	sirt6	sirt6	NOUN
gs-3955	287	9	activators	activator	NOUN
gs-3955	287	10	show	show	VERB
gs-3955	287	11	promise	promise	NOUN
gs-3955	287	12	,	,	PUNCT
gs-3955	287	13	true	true	ADJ
gs-3955	287	14	healthspan	healthspan	PROPN
gs-3955	287	15	extension	extension	NOUN
gs-3955	287	16	requires	require	VERB
gs-3955	287	17	systems	system	NOUN
gs-3955	287	18	-	-	PUNCT
gs-3955	287	19	level	level	NOUN
gs-3955	287	20	recalibration	recalibration	NOUN
gs-3955	287	21	of	of	ADP
gs-3955	287	22	human	human	ADJ
gs-3955	287	23	biology	biology	NOUN
gs-3955	287	24	(	(	PUNCT
gs-3955	287	25	seluanov	seluanov	PROPN
gs-3955	287	26	et	et	PROPN
gs-3955	287	27	al	al	PROPN
gs-3955	287	28	.	.	PROPN
gs-3955	287	29	,	,	PUNCT
gs-3955	287	30	2009	2009	NUM
gs-3955	287	31	;	;	PUNCT
gs-3955	287	32	kumar	kumar	PROPN
gs-3955	287	33	et	et	PROPN
gs-3955	287	34	al	al	PROPN
gs-3955	287	35	.	.	PROPN
gs-3955	287	36	,	,	PUNCT
gs-3955	287	37	2021	2021	NUM
gs-3955	287	38	)	)	PUNCT
gs-3955	287	39	.	.	PUNCT
gs-3955	288	1	the	the	DET
gs-3955	288	2	next	next	ADJ
gs-3955	288	3	decade	decade	NOUN
gs-3955	288	4	will	will	AUX
gs-3955	288	5	witness	witness	VERB
gs-3955	288	6	convergence	convergence	NOUN
gs-3955	288	7	of	of	ADP
gs-3955	288	8	comparative	comparative	ADJ
gs-3955	288	9	genomics	genomic	NOUN
gs-3955	288	10	,	,	PUNCT
gs-3955	288	11	synthetic	synthetic	ADJ
gs-3955	288	12	biology	biology	NOUN
gs-3955	288	13	,	,	PUNCT
gs-3955	288	14	and	and	CCONJ
gs-3955	288	15	quantum	quantum	ADJ
gs-3955	288	16	biophysics	biophysic	NOUN
gs-3955	288	17	to	to	PART
gs-3955	288	18	decode	decode	VERB
gs-3955	288	19	these	these	DET
gs-3955	288	20	species	specie	NOUN
gs-3955	288	21	'	'	PART
gs-3955	288	22	blueprints	blueprint	NOUN
gs-3955	288	23	(	(	PUNCT
gs-3955	288	24	garcia	garcia	PROPN
gs-3955	288	25	et	et	PROPN
gs-3955	288	26	al	al	PROPN
gs-3955	288	27	.	.	PROPN
gs-3955	288	28	,	,	PUNCT
gs-3955	288	29	2023	2023	NUM
gs-3955	288	30	;	;	PUNCT
gs-3955	288	31	jones	jones	PROPN
gs-3955	288	32	et	et	PROPN
gs-3955	288	33	al	al	PROPN
gs-3955	288	34	.	.	PROPN
gs-3955	288	35	,	,	PUNCT
gs-3955	288	36	2021	2021	NUM
gs-3955	288	37	)	)	PUNCT
gs-3955	288	38	.	.	PUNCT
gs-3955	289	1	as	as	SCONJ
gs-3955	289	2	we	we	PRON
gs-3955	289	3	stand	stand	VERB
gs-3955	289	4	at	at	ADP
gs-3955	289	5	this	this	DET
gs-3955	289	6	frontier	frontier	NOUN
gs-3955	289	7	,	,	PUNCT
gs-3955	289	8	we	we	PRON
gs-3955	289	9	must	must	AUX
gs-3955	289	10	recall	recall	VERB
gs-3955	289	11	that	that	PRON
gs-3955	289	12	hydra	hydra	PROPN
gs-3955	289	13	's	's	PART
gs-3955	289	14	immortality	immortality	NOUN
gs-3955	289	15	emerged	emerge	VERB
gs-3955	289	16	through	through	ADP
gs-3955	289	17	500	500	NUM
gs-3955	289	18	million	million	NUM
gs-3955	289	19	years	year	NOUN
gs-3955	289	20	of	of	ADP
gs-3955	289	21	evolution	evolution	NOUN
gs-3955	289	22	—	—	PUNCT
gs-3955	289	23	our	our	PRON
gs-3955	289	24	ambition	ambition	NOUN
gs-3955	289	25	is	be	AUX
gs-3955	289	26	to	to	PART
gs-3955	289	27	achieve	achieve	VERB
gs-3955	289	28	similar	similar	ADJ
gs-3955	289	29	mastery	mastery	NOUN
gs-3955	289	30	within	within	ADP
gs-3955	289	31	mere	mere	ADJ
gs-3955	289	32	decades	decade	NOUN
gs-3955	289	33	(	(	PUNCT
gs-3955	289	34	siebert	siebert	PROPN
gs-3955	289	35	et	et	PROPN
gs-3955	289	36	al	al	PROPN
gs-3955	289	37	.	.	PROPN
gs-3955	289	38	,	,	PUNCT
gs-3955	289	39	2019	2019	NUM
gs-3955	289	40	)	)	PUNCT
gs-3955	289	41	.	.	PUNCT
gs-3955	290	1	the	the	DET
gs-3955	290	2	organisms	organism	NOUN
gs-3955	290	3	described	describe	VERB
gs-3955	290	4	herein	herein	NOUN
gs-3955	290	5	are	be	AUX
gs-3955	290	6	not	not	PART
gs-3955	290	7	mere	mere	ADJ
gs-3955	290	8	biological	biological	ADJ
gs-3955	290	9	curiosities	curiosity	NOUN
gs-3955	290	10	;	;	PUNCT
gs-3955	290	11	they	they	PRON
gs-3955	290	12	are	be	AUX
gs-3955	290	13	living	live	VERB
gs-3955	290	14	proofs	proof	NOUN
gs-3955	290	15	of	of	ADP
gs-3955	290	16	concept	concept	NOUN
gs-3955	290	17	that	that	SCONJ
gs-3955	290	18	aging	aging	NOUN
gs-3955	290	19	is	be	AUX
gs-3955	290	20	not	not	PART
gs-3955	290	21	the	the	DET
gs-3955	290	22	price	price	NOUN
gs-3955	290	23	of	of	ADP
gs-3955	290	24	complexity	complexity	NOUN
gs-3955	290	25	,	,	PUNCT
gs-3955	290	26	but	but	CCONJ
gs-3955	290	27	a	a	DET
gs-3955	290	28	solvable	solvable	ADJ
gs-3955	290	29	problem	problem	NOUN
gs-3955	290	30	in	in	ADP
gs-3955	290	31	cellular	cellular	ADJ
gs-3955	290	32	information	information	NOUN
gs-3955	290	33	management	management	NOUN
gs-3955	290	34	(	(	PUNCT
gs-3955	290	35	tkemaladze	tkemaladze	NOUN
gs-3955	290	36	,	,	PUNCT
gs-3955	290	37	2023	2023	NUM
gs-3955	290	38	)	)	PUNCT
gs-3955	290	39	.	.	PUNCT
gs-3955	291	1	their	their	PRON
gs-3955	291	2	continued	continue	VERB
gs-3955	291	3	study	study	NOUN
gs-3955	291	4	promises	promise	VERB
gs-3955	291	5	not	not	PART
gs-3955	291	6	just	just	ADV
gs-3955	291	7	incremental	incremental	ADJ
gs-3955	291	8	health	health	NOUN
gs-3955	291	9	improvements	improvement	NOUN
gs-3955	291	10	,	,	PUNCT
gs-3955	291	11	but	but	CCONJ
gs-3955	291	12	a	a	DET
gs-3955	291	13	fundamental	fundamental	ADJ
gs-3955	291	14	redefinition	redefinition	NOUN
gs-3955	291	15	of	of	ADP
gs-3955	291	16	the	the	DET
gs-3955	291	17	human	human	ADJ
gs-3955	291	18	biological	biological	ADJ
gs-3955	291	19	trajectory	trajectory	NOUN
gs-3955	291	20	(	(	PUNCT
gs-3955	291	21	lee	lee	PROPN
gs-3955	291	22	et	et	PROPN
gs-3955	291	23	al	al	PROPN
gs-3955	291	24	.	.	PROPN
gs-3955	291	25	,	,	PUNCT
gs-3955	291	26	2022	2022	NUM
gs-3955	291	27	;	;	PUNCT
gs-3955	291	28	patel	patel	PROPN
gs-3955	291	29	et	et	PROPN
gs-3955	291	30	al	al	PROPN
gs-3955	291	31	.	.	PROPN
gs-3955	291	32	,	,	PUNCT
gs-3955	291	33	2024	2024	NUM
gs-3955	291	34	)	)	PUNCT
gs-3955	291	35	.	.	PUNCT
gs-3955	292	1	discussion	discussion	NOUN
gs-3955	292	2	the	the	DET
gs-3955	292	3	study	study	NOUN
gs-3955	292	4	of	of	ADP
gs-3955	292	5	negligibly	negligibly	ADV
gs-3955	292	6	senescent	senescent	ADJ
gs-3955	292	7	organisms	organism	NOUN
gs-3955	292	8	reveals	reveal	VERB
gs-3955	292	9	that	that	SCONJ
gs-3955	292	10	aging	aging	NOUN
gs-3955	292	11	is	be	AUX
gs-3955	292	12	not	not	PART
gs-3955	292	13	inevitable	inevitable	ADJ
gs-3955	292	14	and	and	CCONJ
gs-3955	292	15	may	may	AUX
gs-3955	292	16	be	be	AUX
gs-3955	292	17	modifiable	modifiable	ADJ
gs-3955	292	18	.	.	PUNCT
gs-3955	293	1	common	common	ADJ
gs-3955	293	2	themes	theme	NOUN
gs-3955	293	3	include	include	VERB
gs-3955	293	4	sustained	sustain	VERB
gs-3955	293	5	stem	stem	NOUN
gs-3955	293	6	cell	cell	NOUN
gs-3955	293	7	function	function	NOUN
gs-3955	293	8	,	,	PUNCT
gs-3955	293	9	reduced	reduce	VERB
gs-3955	293	10	or	or	CCONJ
gs-3955	293	11	efficiently	efficiently	ADV
gs-3955	293	12	repaired	repair	VERB
gs-3955	293	13	molecular	molecular	ADJ
gs-3955	293	14	damage	damage	NOUN
gs-3955	293	15	,	,	PUNCT
gs-3955	293	16	and	and	CCONJ
gs-3955	293	17	preserved	preserve	VERB
gs-3955	293	18	genomic	genomic	NOUN
gs-3955	293	19	and	and	CCONJ
gs-3955	293	20	proteomic	proteomic	ADJ
gs-3955	293	21	integrity	integrity	NOUN
gs-3955	293	22	.	.	PUNCT
gs-3955	294	1	these	these	DET
gs-3955	294	2	species	specie	NOUN
gs-3955	294	3	show	show	VERB
gs-3955	294	4	that	that	SCONJ
gs-3955	294	5	extreme	extreme	ADJ
gs-3955	294	6	longevity	longevity	NOUN
gs-3955	294	7	results	result	VERB
gs-3955	294	8	not	not	PART
gs-3955	294	9	from	from	ADP
gs-3955	294	10	a	a	DET
gs-3955	294	11	single	single	ADJ
gs-3955	294	12	factor	factor	NOUN
gs-3955	294	13	but	but	CCONJ
gs-3955	294	14	from	from	ADP
gs-3955	294	15	an	an	DET
gs-3955	294	16	orchestrated	orchestrated	ADJ
gs-3955	294	17	network	network	NOUN
gs-3955	294	18	of	of	ADP
gs-3955	294	19	molecular	molecular	ADJ
gs-3955	294	20	safeguards	safeguard	NOUN
gs-3955	294	21	.	.	PUNCT
gs-3955	295	1	for	for	ADP
gs-3955	295	2	example	example	NOUN
gs-3955	295	3	,	,	PUNCT
gs-3955	295	4	tp53	tp53	PROPN
gs-3955	295	5	’s	’s	PART
gs-3955	295	6	modified	modify	VERB
gs-3955	295	7	functionality	functionality	NOUN
gs-3955	295	8	in	in	ADP
gs-3955	295	9	naked	naked	ADJ
gs-3955	295	10	mole	mole	NOUN
gs-3955	295	11	-	-	PUNCT
gs-3955	295	12	rats	rat	NOUN
gs-3955	295	13	illustrates	illustrate	VERB
gs-3955	295	14	how	how	SCONJ
gs-3955	295	15	a	a	DET
gs-3955	295	16	single	single	ADJ
gs-3955	295	17	gene	gene	NOUN
gs-3955	295	18	may	may	AUX
gs-3955	295	19	be	be	AUX
gs-3955	295	20	tuned	tune	VERB
gs-3955	295	21	to	to	PART
gs-3955	295	22	prioritize	prioritize	VERB
gs-3955	295	23	maintenance	maintenance	NOUN
gs-3955	295	24	over	over	ADP
gs-3955	295	25	programmed	program	VERB
gs-3955	295	26	cell	cell	NOUN
gs-3955	295	27	death	death	NOUN
gs-3955	295	28	(	(	PUNCT
gs-3955	295	29	lewis	lewis	PROPN
gs-3955	295	30	et	et	PROPN
gs-3955	295	31	al	al	PROPN
gs-3955	295	32	.	.	PROPN
gs-3955	295	33	,	,	PUNCT
gs-3955	295	34	2013	2013	NUM
gs-3955	295	35	)	)	PUNCT
gs-3955	295	36	.	.	PUNCT
gs-3955	296	1	similarly	similarly	ADV
gs-3955	296	2	,	,	PUNCT
gs-3955	296	3	oxidative	oxidative	ADJ
gs-3955	296	4	stress	stress	NOUN
gs-3955	296	5	resistance	resistance	NOUN
gs-3955	296	6	in	in	ADP
gs-3955	296	7	a.	a.	PROPN
gs-3955	296	8	islandica	islandica	PROPN
gs-3955	296	9	stems	stem	VERB
gs-3955	296	10	from	from	ADP
gs-3955	296	11	both	both	CCONJ
gs-3955	296	12	mitochondrial	mitochondrial	ADJ
gs-3955	296	13	adaptations	adaptation	NOUN
gs-3955	296	14	and	and	CCONJ
gs-3955	296	15	membrane	membrane	NOUN
gs-3955	296	16	stability	stability	NOUN
gs-3955	296	17	(	(	PUNCT
gs-3955	296	18	macrae	macrae	PROPN
gs-3955	296	19	et	et	PROPN
gs-3955	296	20	al	al	PROPN
gs-3955	296	21	.	.	PROPN
gs-3955	296	22	,	,	PUNCT
gs-3955	296	23	2015	2015	NUM
gs-3955	296	24	)	)	PUNCT
gs-3955	296	25	.	.	PUNCT
gs-3955	297	1	research	research	NOUN
gs-3955	297	2	on	on	ADP
gs-3955	297	3	these	these	DET
gs-3955	297	4	species	specie	NOUN
gs-3955	297	5	is	be	AUX
gs-3955	297	6	ongoing	ongoing	ADJ
gs-3955	297	7	but	but	CCONJ
gs-3955	297	8	already	already	ADV
gs-3955	297	9	highlights	highlight	VERB
gs-3955	297	10	potential	potential	ADJ
gs-3955	297	11	avenues	avenue	NOUN
gs-3955	297	12	for	for	ADP
gs-3955	297	13	human	human	ADJ
gs-3955	297	14	anti	anti	ADJ
gs-3955	297	15	-	-	ADJ
gs-3955	297	16	aging	aging	ADJ
gs-3955	297	17	therapies	therapy	NOUN
gs-3955	297	18	.	.	PUNCT
gs-3955	298	1	foxo	foxo	PROPN
gs-3955	298	2	signaling	signal	VERB
gs-3955	298	3	,	,	PUNCT
gs-3955	298	4	telomerase	telomerase	NOUN
gs-3955	298	5	regulation	regulation	NOUN
gs-3955	298	6	,	,	PUNCT
gs-3955	298	7	and	and	CCONJ
gs-3955	298	8	autophagy	autophagy	ADJ
gs-3955	298	9	enhancement	enhancement	NOUN
gs-3955	298	10	are	be	AUX
gs-3955	298	11	among	among	ADP
gs-3955	298	12	the	the	DET
gs-3955	298	13	promising	promising	ADJ
gs-3955	298	14	interventions	intervention	NOUN
gs-3955	298	15	.	.	PUNCT
gs-3955	299	1	while	while	SCONJ
gs-3955	299	2	the	the	DET
gs-3955	299	3	translation	translation	NOUN
gs-3955	299	4	to	to	ADP
gs-3955	299	5	humans	human	NOUN
gs-3955	299	6	is	be	AUX
gs-3955	299	7	complex	complex	ADJ
gs-3955	299	8	,	,	PUNCT
gs-3955	299	9	further	far	ADV
gs-3955	299	10	comparative	comparative	ADJ
gs-3955	299	11	genomic	genomic	NOUN
gs-3955	299	12	and	and	CCONJ
gs-3955	299	13	proteomic	proteomic	ADJ
gs-3955	299	14	studies	study	NOUN
gs-3955	299	15	will	will	AUX
gs-3955	299	16	clarify	clarify	VERB
gs-3955	299	17	how	how	SCONJ
gs-3955	299	18	these	these	DET
gs-3955	299	19	pathways	pathway	NOUN
gs-3955	299	20	operate	operate	VERB
gs-3955	299	21	in	in	ADP
gs-3955	299	22	long	long	ADV
gs-3955	299	23	-	-	PUNCT
gs-3955	299	24	lived	live	VERB
gs-3955	299	25	species	specie	NOUN
gs-3955	299	26	(	(	PUNCT
gs-3955	299	27	munro	munro	PROPN
gs-3955	299	28	et	et	PROPN
gs-3955	299	29	al	al	PROPN
gs-3955	299	30	.	.	PROPN
gs-3955	299	31	,	,	PUNCT
gs-3955	299	32	2012	2012	NUM
gs-3955	299	33	)	)	PUNCT
gs-3955	299	34	.	.	PUNCT
gs-3955	300	1	negligibly	negligibly	ADV
gs-3955	300	2	senescent	senescent	ADJ
gs-3955	300	3	species	specie	NOUN
gs-3955	300	4	like	like	ADP
gs-3955	300	5	hydra	hydra	PROPN
gs-3955	300	6	,	,	PUNCT
gs-3955	300	7	sharks	shark	NOUN
gs-3955	300	8	,	,	PUNCT
gs-3955	300	9	and	and	CCONJ
gs-3955	300	10	naked	naked	ADJ
gs-3955	300	11	mole	mole	NOUN
gs-3955	300	12	-	-	PUNCT
gs-3955	300	13	rats	rat	NOUN
gs-3955	300	14	defy	defy	VERB
gs-3955	300	15	the	the	DET
gs-3955	300	16	typical	typical	ADJ
gs-3955	300	17	rules	rule	NOUN
gs-3955	300	18	of	of	ADP
gs-3955	300	19	aging	aging	NOUN
gs-3955	300	20	.	.	PUNCT
gs-3955	301	1	through	through	ADP
gs-3955	301	2	a	a	DET
gs-3955	301	3	combination	combination	NOUN
gs-3955	301	4	of	of	ADP
gs-3955	301	5	enhanced	enhanced	ADJ
gs-3955	301	6	regeneration	regeneration	NOUN
gs-3955	301	7	,	,	PUNCT
gs-3955	301	8	molecular	molecular	ADJ
gs-3955	301	9	repair	repair	NOUN
gs-3955	301	10	systems	system	NOUN
gs-3955	301	11	,	,	PUNCT
gs-3955	301	12	and	and	CCONJ
gs-3955	301	13	environmental	environmental	ADJ
gs-3955	301	14	adaptations	adaptation	NOUN
gs-3955	301	15	,	,	PUNCT
gs-3955	301	16	these	these	DET
gs-3955	301	17	organisms	organism	NOUN
gs-3955	301	18	remain	remain	VERB
gs-3955	301	19	functionally	functionally	ADV
gs-3955	301	20	youthful	youthful	ADJ
gs-3955	301	21	across	across	ADP
gs-3955	301	22	their	their	PRON
gs-3955	301	23	lifespans	lifespan	NOUN
gs-3955	301	24	.	.	PUNCT
gs-3955	302	1	decoding	decode	VERB
gs-3955	302	2	the	the	DET
gs-3955	302	3	molecular	molecular	ADJ
gs-3955	302	4	blueprints	blueprint	NOUN
gs-3955	302	5	of	of	ADP
gs-3955	302	6	these	these	DET
gs-3955	302	7	"	"	PUNCT
gs-3955	302	8	immortal	immortal	ADJ
gs-3955	302	9	"	"	PUNCT
gs-3955	302	10	organisms	organism	NOUN
gs-3955	302	11	could	could	AUX
gs-3955	302	12	revolutionize	revolutionize	VERB
gs-3955	302	13	our	our	PRON
gs-3955	302	14	understanding	understanding	NOUN
gs-3955	302	15	of	of	ADP
gs-3955	302	16	aging	aging	NOUN
gs-3955	302	17	and	and	CCONJ
gs-3955	302	18	inform	inform	VERB
gs-3955	302	19	the	the	DET
gs-3955	302	20	development	development	NOUN
gs-3955	302	21	of	of	ADP
gs-3955	302	22	novel	novel	ADJ
gs-3955	302	23	therapies	therapy	NOUN
gs-3955	302	24	aimed	aim	VERB
gs-3955	302	25	at	at	ADP
gs-3955	302	26	promoting	promote	VERB
gs-3955	302	27	longevity	longevity	NOUN
gs-3955	302	28	and	and	CCONJ
gs-3955	302	29	age	age	NOUN
gs-3955	302	30	-	-	PUNCT
gs-3955	302	31	related	relate	VERB
gs-3955	302	32	disease	disease	NOUN
gs-3955	302	33	resistance	resistance	NOUN
gs-3955	302	34	in	in	ADP
gs-3955	302	35	humans	human	NOUN
gs-3955	302	36	.	.	PUNCT
gs-3955	303	1	emulating	emulate	VERB
gs-3955	303	2	their	their	PRON
gs-3955	303	3	stress	stress	ADJ
gs-3955	303	4	response	response	NOUN
gs-3955	303	5	mechanisms	mechanism	NOUN
gs-3955	303	6	,	,	PUNCT
gs-3955	303	7	dna	dna	PROPN
gs-3955	303	8	repair	repair	NOUN
gs-3955	303	9	efficiency	efficiency	NOUN
gs-3955	303	10	,	,	PUNCT
gs-3955	303	11	and	and	CCONJ
gs-3955	303	12	epigenetic	epigenetic	ADJ
gs-3955	303	13	stability	stability	NOUN
gs-3955	303	14	holds	hold	VERB
gs-3955	303	15	immense	immense	ADJ
gs-3955	303	16	promise	promise	NOUN
gs-3955	303	17	for	for	ADP
gs-3955	303	18	enhancing	enhance	VERB
gs-3955	303	19	the	the	DET
gs-3955	303	20	human	human	ADJ
gs-3955	303	21	healthspan	healthspan	PROPN
gs-3955	303	22	(	(	PUNCT
gs-3955	303	23	narayan	narayan	PROPN
gs-3955	303	24	et	et	PROPN
gs-3955	303	25	al	al	PROPN
gs-3955	303	26	.	.	PROPN
gs-3955	303	27	,	,	PUNCT
gs-3955	303	28	2021	2021	NUM
gs-3955	303	29	)	)	PUNCT
gs-3955	303	30	.	.	PUNCT
gs-3955	304	1	the	the	DET
gs-3955	304	2	study	study	NOUN
gs-3955	304	3	of	of	ADP
gs-3955	304	4	negligibly	negligibly	ADV
gs-3955	304	5	senescing	senesce	VERB
gs-3955	304	6	species	specie	NOUN
gs-3955	304	7	fundamentally	fundamentally	ADV
gs-3955	304	8	challenges	challenge	VERB
gs-3955	304	9	the	the	DET
gs-3955	304	10	paradigm	paradigm	NOUN
gs-3955	304	11	that	that	SCONJ
gs-3955	304	12	aging	aging	NOUN
gs-3955	304	13	is	be	AUX
gs-3955	304	14	an	an	DET
gs-3955	304	15	inescapable	inescapable	ADJ
gs-3955	304	16	biological	biological	ADJ
gs-3955	304	17	inevitability	inevitability	NOUN
gs-3955	304	18	.	.	PUNCT
gs-3955	305	1	across	across	ADP
gs-3955	305	2	phylogenetically	phylogenetically	ADV
gs-3955	305	3	diverse	diverse	ADJ
gs-3955	305	4	organisms	organism	NOUN
gs-3955	305	5	—	—	PUNCT
gs-3955	305	6	from	from	ADP
gs-3955	305	7	georgian	georgian	ADJ
gs-3955	305	8	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	305	9	მეცნიერები	მეცნიერები	NOUN
gs-3955	305	10	ტ.	ტ.	VERB
gs-3955	305	11	7	7	NUM
gs-3955	305	12	n	n	PRON
gs-3955	305	13	3	3	NUM
gs-3955	305	14	,	,	PUNCT
gs-3955	305	15	2025	2025	NUM
gs-3955	305	16	382	382	NUM
gs-3955	305	17	the	the	DET
gs-3955	305	18	greenland	greenland	NOUN
gs-3955	305	19	shark	shark	NOUN
gs-3955	305	20	and	and	CCONJ
gs-3955	305	21	naked	naked	ADJ
gs-3955	305	22	mole	mole	NOUN
gs-3955	305	23	-	-	PUNCT
gs-3955	305	24	rat	rat	NOUN
gs-3955	305	25	to	to	ADP
gs-3955	305	26	hydra	hydra	PROPN
gs-3955	305	27	and	and	CCONJ
gs-3955	305	28	the	the	DET
gs-3955	305	29	ocean	ocean	NOUN
gs-3955	305	30	quahog	quahog	PROPN
gs-3955	305	31	—	—	PUNCT
gs-3955	305	32	a	a	DET
gs-3955	305	33	convergence	convergence	NOUN
gs-3955	305	34	of	of	ADP
gs-3955	305	35	molecular	molecular	ADJ
gs-3955	305	36	and	and	CCONJ
gs-3955	305	37	physiological	physiological	ADJ
gs-3955	305	38	adaptations	adaptation	NOUN
gs-3955	305	39	enables	enable	VERB
gs-3955	305	40	the	the	DET
gs-3955	305	41	decoupling	decoupling	NOUN
gs-3955	305	42	of	of	ADP
gs-3955	305	43	chronological	chronological	ADJ
gs-3955	305	44	age	age	NOUN
gs-3955	305	45	from	from	ADP
gs-3955	305	46	functional	functional	ADJ
gs-3955	305	47	decline	decline	NOUN
gs-3955	305	48	.	.	PUNCT
gs-3955	306	1	these	these	DET
gs-3955	306	2	species	specie	NOUN
gs-3955	306	3	collectively	collectively	ADV
gs-3955	306	4	demonstrate	demonstrate	VERB
gs-3955	306	5	that	that	SCONJ
gs-3955	306	6	sustained	sustain	VERB
gs-3955	306	7	cellular	cellular	ADJ
gs-3955	306	8	integrity	integrity	NOUN
gs-3955	306	9	,	,	PUNCT
gs-3955	306	10	reproductive	reproductive	ADJ
gs-3955	306	11	capacity	capacity	NOUN
gs-3955	306	12	,	,	PUNCT
gs-3955	306	13	and	and	CCONJ
gs-3955	306	14	physiological	physiological	ADJ
gs-3955	306	15	resilience	resilience	NOUN
gs-3955	306	16	are	be	AUX
gs-3955	306	17	achievable	achievable	ADJ
gs-3955	306	18	through	through	ADP
gs-3955	306	19	evolutionary	evolutionary	ADJ
gs-3955	306	20	innovations	innovation	NOUN
gs-3955	306	21	that	that	PRON
gs-3955	306	22	counteract	counteract	VERB
gs-3955	306	23	entropic	entropic	ADJ
gs-3955	306	24	degeneration	degeneration	NOUN
gs-3955	306	25	(	(	PUNCT
gs-3955	306	26	perez	perez	PROPN
gs-3955	306	27	et	et	PROPN
gs-3955	306	28	al	al	PROPN
gs-3955	306	29	.	.	PROPN
gs-3955	306	30	,	,	PUNCT
gs-3955	306	31	2009	2009	NUM
gs-3955	306	32	)	)	PUNCT
gs-3955	306	33	.	.	PUNCT
gs-3955	307	1	a	a	DET
gs-3955	307	2	central	central	ADJ
gs-3955	307	3	theme	theme	NOUN
gs-3955	307	4	unifying	unify	VERB
gs-3955	307	5	these	these	DET
gs-3955	307	6	organisms	organism	NOUN
gs-3955	307	7	is	be	AUX
gs-3955	307	8	the	the	DET
gs-3955	307	9	orchestrated	orchestrated	ADJ
gs-3955	307	10	enhancement	enhancement	NOUN
gs-3955	307	11	of	of	ADP
gs-3955	307	12	genomic	genomic	ADJ
gs-3955	307	13	stability	stability	NOUN
gs-3955	307	14	.	.	PUNCT
gs-3955	308	1	enhanced	enhance	VERB
gs-3955	308	2	dna	dna	PROPN
gs-3955	308	3	repair	repair	NOUN
gs-3955	308	4	mechanisms	mechanism	NOUN
gs-3955	308	5	,	,	PUNCT
gs-3955	308	6	including	include	VERB
gs-3955	308	7	elevated	elevated	ADJ
gs-3955	308	8	activity	activity	NOUN
gs-3955	308	9	in	in	ADP
gs-3955	308	10	homologous	homologous	ADJ
gs-3955	308	11	recombination	recombination	NOUN
gs-3955	308	12	,	,	PUNCT
gs-3955	308	13	nucleotide	nucleotide	ADJ
gs-3955	308	14	excision	excision	NOUN
gs-3955	308	15	repair	repair	NOUN
gs-3955	308	16	,	,	PUNCT
gs-3955	308	17	and	and	CCONJ
gs-3955	308	18	base	base	NOUN
gs-3955	308	19	excision	excision	NOUN
gs-3955	308	20	repair	repair	NOUN
gs-3955	308	21	pathways	pathway	NOUN
gs-3955	308	22	,	,	PUNCT
gs-3955	308	23	are	be	AUX
gs-3955	308	24	consistently	consistently	ADV
gs-3955	308	25	observed	observe	VERB
gs-3955	308	26	.	.	PUNCT
gs-3955	309	1	for	for	ADP
gs-3955	309	2	instance	instance	NOUN
gs-3955	309	3	,	,	PUNCT
gs-3955	309	4	naked	naked	ADJ
gs-3955	309	5	mole	mole	NOUN
gs-3955	309	6	-	-	PUNCT
gs-3955	309	7	rats	rat	NOUN
gs-3955	309	8	exhibit	exhibit	VERB
gs-3955	309	9	upregulated	upregulated	ADJ
gs-3955	309	10	expression	expression	NOUN
gs-3955	309	11	of	of	ADP
gs-3955	309	12	atm	atm	PROPN
gs-3955	309	13	,	,	PUNCT
gs-3955	309	14	brca1/2	brca1/2	ADJ
gs-3955	309	15	,	,	PUNCT
gs-3955	309	16	and	and	CCONJ
gs-3955	309	17	tp53	tp53	PROPN
gs-3955	309	18	,	,	PUNCT
gs-3955	309	19	favoring	favor	VERB
gs-3955	309	20	dna	dna	PROPN
gs-3955	309	21	repair	repair	NOUN
gs-3955	309	22	over	over	ADP
gs-3955	309	23	apoptosis	apoptosis	NOUN
gs-3955	309	24	,	,	PUNCT
gs-3955	309	25	while	while	SCONJ
gs-3955	309	26	greenland	greenland	PROPN
gs-3955	309	27	sharks	sharks	PROPN
gs-3955	309	28	possess	possess	VERB
gs-3955	309	29	mutations	mutation	NOUN
gs-3955	309	30	in	in	ADP
gs-3955	309	31	ercc1/2	ercc1/2	NOUN
gs-3955	309	32	genes	gene	NOUN
gs-3955	309	33	.	.	PUNCT
gs-3955	310	1	this	this	DET
gs-3955	310	2	genomic	genomic	ADJ
gs-3955	310	3	vigilance	vigilance	NOUN
gs-3955	310	4	minimizes	minimize	VERB
gs-3955	310	5	somatic	somatic	ADJ
gs-3955	310	6	mutation	mutation	NOUN
gs-3955	310	7	accumulation	accumulation	NOUN
gs-3955	310	8	and	and	CCONJ
gs-3955	310	9	is	be	AUX
gs-3955	310	10	further	far	ADV
gs-3955	310	11	reinforced	reinforce	VERB
gs-3955	310	12	in	in	ADP
gs-3955	310	13	invertebrates	invertebrate	NOUN
gs-3955	310	14	like	like	ADP
gs-3955	310	15	arctica	arctica	PROPN
gs-3955	310	16	islandica	islandica	PROPN
gs-3955	310	17	through	through	ADP
gs-3955	310	18	efficient	efficient	ADJ
gs-3955	310	19	lesion	lesion	NOUN
gs-3955	310	20	repair	repair	NOUN
gs-3955	310	21	.	.	PUNCT
gs-3955	311	1	such	such	ADJ
gs-3955	311	2	mechanisms	mechanism	NOUN
gs-3955	311	3	operate	operate	VERB
gs-3955	311	4	synergistically	synergistically	ADV
gs-3955	311	5	with	with	ADP
gs-3955	311	6	epigenetic	epigenetic	ADJ
gs-3955	311	7	stability	stability	NOUN
gs-3955	311	8	,	,	PUNCT
gs-3955	311	9	where	where	SCONJ
gs-3955	311	10	slow	slow	ADJ
gs-3955	311	11	methylation	methylation	NOUN
gs-3955	311	12	drift	drift	NOUN
gs-3955	311	13	and	and	CCONJ
gs-3955	311	14	conserved	conserve	VERB
gs-3955	311	15	mirna	mirna	PROPN
gs-3955	311	16	profiles	profile	NOUN
gs-3955	311	17	preserve	preserve	VERB
gs-3955	311	18	youthful	youthful	ADJ
gs-3955	311	19	transcriptional	transcriptional	ADJ
gs-3955	311	20	programs	program	NOUN
gs-3955	311	21	across	across	ADP
gs-3955	311	22	taxa	taxa	NOUN
gs-3955	311	23	(	(	PUNCT
gs-3955	311	24	petruseva	petruseva	PROPN
gs-3955	311	25	et	et	PROPN
gs-3955	311	26	al	al	PROPN
gs-3955	311	27	.	.	PROPN
gs-3955	311	28	,	,	PUNCT
gs-3955	311	29	2017	2017	NUM
gs-3955	311	30	)	)	PUNCT
gs-3955	311	31	.	.	PUNCT
gs-3955	312	1	equally	equally	ADV
gs-3955	312	2	critical	critical	ADJ
gs-3955	312	3	is	be	AUX
gs-3955	312	4	the	the	DET
gs-3955	312	5	mastery	mastery	NOUN
gs-3955	312	6	of	of	ADP
gs-3955	312	7	oxidative	oxidative	ADJ
gs-3955	312	8	stress	stress	NOUN
gs-3955	312	9	management	management	NOUN
gs-3955	312	10	.	.	PUNCT
gs-3955	313	1	long	long	ADV
gs-3955	313	2	-	-	PUNCT
gs-3955	313	3	lived	live	VERB
gs-3955	313	4	species	specie	NOUN
gs-3955	313	5	deploy	deploy	VERB
gs-3955	313	6	divergent	divergent	ADJ
gs-3955	313	7	strategies	strategy	NOUN
gs-3955	313	8	:	:	PUNCT
gs-3955	313	9	arctica	arctica	PROPN
gs-3955	313	10	islandica	islandica	PROPN
gs-3955	313	11	reduces	reduce	VERB
gs-3955	313	12	mitochondrial	mitochondrial	ADJ
gs-3955	313	13	ros	ros	PROPN
gs-3955	313	14	production	production	NOUN
gs-3955	313	15	via	via	ADP
gs-3955	313	16	specialized	specialized	ADJ
gs-3955	313	17	membrane	membrane	NOUN
gs-3955	313	18	lipids	lipid	NOUN
gs-3955	313	19	with	with	ADP
gs-3955	313	20	low	low	ADJ
gs-3955	313	21	peroxidation	peroxidation	NOUN
gs-3955	313	22	indices	index	NOUN
gs-3955	313	23	,	,	PUNCT
gs-3955	313	24	whereas	whereas	SCONJ
gs-3955	313	25	naked	naked	ADJ
gs-3955	313	26	mole	mole	NOUN
gs-3955	313	27	-	-	PUNCT
gs-3955	313	28	rats	rat	NOUN
gs-3955	313	29	tolerate	tolerate	VERB
gs-3955	313	30	high	high	ADJ
gs-3955	313	31	ros	ros	PROPN
gs-3955	313	32	levels	level	NOUN
gs-3955	313	33	without	without	ADP
gs-3955	313	34	incurring	incur	VERB
gs-3955	313	35	macromolecular	macromolecular	ADJ
gs-3955	313	36	damage	damage	NOUN
gs-3955	313	37	due	due	ADP
gs-3955	313	38	to	to	ADP
gs-3955	313	39	unique	unique	ADJ
gs-3955	313	40	cardiolipin	cardiolipin	NOUN
gs-3955	313	41	structures	structure	NOUN
gs-3955	313	42	and	and	CCONJ
gs-3955	313	43	ribosomal	ribosomal	NOUN
gs-3955	313	44	modifications	modification	NOUN
gs-3955	313	45	.	.	PUNCT
gs-3955	314	1	similarly	similarly	ADV
gs-3955	314	2	,	,	PUNCT
gs-3955	314	3	the	the	DET
gs-3955	314	4	olm	olm	NOUN
gs-3955	314	5	(	(	PUNCT
gs-3955	314	6	proteus	proteus	PROPN
gs-3955	314	7	anguinus	anguinus	PROPN
gs-3955	314	8	)	)	PUNCT
gs-3955	314	9	leverages	leverage	VERB
gs-3955	314	10	constitutive	constitutive	ADJ
gs-3955	314	11	nrf2	nrf2	NOUN
gs-3955	314	12	pathway	pathway	NOUN
gs-3955	314	13	activation	activation	NOUN
gs-3955	314	14	for	for	ADP
gs-3955	314	15	antioxidant	antioxidant	ADJ
gs-3955	314	16	defense	defense	NOUN
gs-3955	314	17	,	,	PUNCT
gs-3955	314	18	despite	despite	SCONJ
gs-3955	314	19	low	low	ADJ
gs-3955	314	20	enzyme	enzyme	NOUN
gs-3955	314	21	levels	level	NOUN
gs-3955	314	22	.	.	PUNCT
gs-3955	315	1	this	this	DET
gs-3955	315	2	plasticity	plasticity	NOUN
gs-3955	315	3	underscores	underscore	VERB
gs-3955	315	4	that	that	PRON
gs-3955	315	5	oxidative	oxidative	ADJ
gs-3955	315	6	resistance	resistance	NOUN
gs-3955	315	7	is	be	AUX
gs-3955	315	8	not	not	PART
gs-3955	315	9	merely	merely	ADV
gs-3955	315	10	a	a	DET
gs-3955	315	11	reduction	reduction	NOUN
gs-3955	315	12	in	in	ADP
gs-3955	315	13	ros	ros	PROPN
gs-3955	315	14	but	but	CCONJ
gs-3955	315	15	a	a	DET
gs-3955	315	16	systemic	systemic	ADJ
gs-3955	315	17	recalibration	recalibration	NOUN
gs-3955	315	18	of	of	ADP
gs-3955	315	19	damage	damage	NOUN
gs-3955	315	20	tolerance	tolerance	NOUN
gs-3955	315	21	(	(	PUNCT
gs-3955	315	22	strahlendorf	strahlendorf	PROPN
gs-3955	315	23	et	et	PROPN
gs-3955	315	24	al	al	PROPN
gs-3955	315	25	.	.	PROPN
gs-3955	315	26	,	,	PUNCT
gs-3955	315	27	2008	2008	NUM
gs-3955	315	28	,	,	PUNCT
gs-3955	315	29	nielsen	nielsen	PROPN
gs-3955	315	30	et	et	PROPN
gs-3955	315	31	al	al	PROPN
gs-3955	315	32	.	.	PROPN
gs-3955	315	33	2016	2016	NUM
gs-3955	315	34	)	)	PUNCT
gs-3955	315	35	.	.	PUNCT
gs-3955	316	1	proteostasis	proteostasis	NOUN
gs-3955	316	2	and	and	CCONJ
gs-3955	316	3	regenerative	regenerative	ADJ
gs-3955	316	4	capacity	capacity	NOUN
gs-3955	316	5	further	far	ADV
gs-3955	316	6	underpin	underpin	VERB
gs-3955	316	7	negligible	negligible	ADJ
gs-3955	316	8	senescence	senescence	NOUN
gs-3955	316	9	.	.	PUNCT
gs-3955	317	1	autophagy	autophagy	ADJ
gs-3955	317	2	pathways	pathway	NOUN
gs-3955	317	3	are	be	AUX
gs-3955	317	4	amplified	amplify	VERB
gs-3955	317	5	in	in	ADP
gs-3955	317	6	rockfish	rockfish	ADJ
gs-3955	317	7	(	(	PUNCT
gs-3955	317	8	sebastes	sebaste	NOUN
gs-3955	317	9	aleutianus	aleutianus	NOUN
gs-3955	317	10	)	)	PUNCT
gs-3955	317	11	and	and	CCONJ
gs-3955	317	12	hydra	hydra	PROPN
gs-3955	317	13	,	,	PUNCT
gs-3955	317	14	ensuring	ensure	VERB
gs-3955	317	15	efficient	efficient	ADJ
gs-3955	317	16	clearance	clearance	NOUN
gs-3955	317	17	of	of	ADP
gs-3955	317	18	misfolded	misfolde	VERB
gs-3955	317	19	proteins	protein	NOUN
gs-3955	317	20	.	.	PUNCT
gs-3955	318	1	stem	stem	NOUN
gs-3955	318	2	cell	cell	NOUN
gs-3955	318	3	systems	system	NOUN
gs-3955	318	4	are	be	AUX
gs-3955	318	5	paramount	paramount	ADJ
gs-3955	318	6	—	—	PUNCT
gs-3955	318	7	hydra	hydra	PROPN
gs-3955	318	8	’s	’s	PART
gs-3955	318	9	perpetually	perpetually	ADV
gs-3955	318	10	active	active	ADJ
gs-3955	318	11	stem	stem	NOUN
gs-3955	318	12	cells	cell	NOUN
gs-3955	318	13	(	(	PUNCT
gs-3955	318	14	governed	govern	VERB
gs-3955	318	15	by	by	ADP
gs-3955	318	16	foxo	foxo	ADJ
gs-3955	318	17	expression	expression	NOUN
gs-3955	318	18	)	)	PUNCT
gs-3955	318	19	enable	enable	VERB
gs-3955	318	20	whole	whole	ADJ
gs-3955	318	21	-	-	PUNCT
gs-3955	318	22	body	body	NOUN
gs-3955	318	23	renewal	renewal	NOUN
gs-3955	318	24	,	,	PUNCT
gs-3955	318	25	while	while	SCONJ
gs-3955	318	26	planarians	planarian	NOUN
gs-3955	318	27	achieve	achieve	VERB
gs-3955	318	28	theoretical	theoretical	ADJ
gs-3955	318	29	immortality	immortality	NOUN
gs-3955	318	30	through	through	ADP
gs-3955	318	31	neoblast	neoblast	NOUN
gs-3955	318	32	-	-	PUNCT
gs-3955	318	33	mediated	mediate	VERB
gs-3955	318	34	regeneration	regeneration	NOUN
gs-3955	318	35	.	.	PUNCT
gs-3955	319	1	in	in	ADP
gs-3955	319	2	vertebrates	vertebrate	NOUN
gs-3955	319	3	,	,	PUNCT
gs-3955	319	4	salamanders	salamander	NOUN
gs-3955	319	5	and	and	CCONJ
gs-3955	319	6	sea	sea	NOUN
gs-3955	319	7	urchins	urchin	NOUN
gs-3955	319	8	exhibit	exhibit	VERB
gs-3955	319	9	unparalleled	unparalleled	ADJ
gs-3955	319	10	tissue	tissue	NOUN
gs-3955	319	11	regeneration	regeneration	NOUN
gs-3955	319	12	,	,	PUNCT
gs-3955	319	13	from	from	ADP
gs-3955	319	14	limb	limb	NOUN
gs-3955	319	15	reconstruction	reconstruction	NOUN
gs-3955	319	16	to	to	ADP
gs-3955	319	17	neural	neural	ADJ
gs-3955	319	18	repair	repair	NOUN
gs-3955	319	19	,	,	PUNCT
gs-3955	319	20	minimizing	minimize	VERB
gs-3955	319	21	scar	scar	NOUN
gs-3955	319	22	-	-	PUNCT
gs-3955	319	23	associated	associate	VERB
gs-3955	319	24	dysfunction	dysfunction	NOUN
gs-3955	319	25	(	(	PUNCT
gs-3955	319	26	buffenstein	buffenstein	ADV
gs-3955	319	27	,	,	PUNCT
gs-3955	319	28	et	et	PROPN
gs-3955	319	29	al	al	PROPN
gs-3955	319	30	.	.	PROPN
gs-3955	319	31	,	,	PUNCT
gs-3955	319	32	2008	2008	NUM
gs-3955	319	33	,	,	PUNCT
gs-3955	319	34	butler	butler	PROPN
gs-3955	319	35	,	,	PUNCT
gs-3955	319	36	et	et	PROPN
gs-3955	319	37	al	al	PROPN
gs-3955	319	38	.	.	PROPN
gs-3955	319	39	,	,	PUNCT
gs-3955	319	40	2013	2013	NUM
gs-3955	319	41	)	)	PUNCT
gs-3955	319	42	.	.	PUNCT
gs-3955	320	1	the	the	DET
gs-3955	320	2	evolutionary	evolutionary	ADJ
gs-3955	320	3	context	context	NOUN
gs-3955	320	4	of	of	ADP
gs-3955	320	5	these	these	DET
gs-3955	320	6	adaptations	adaptation	NOUN
gs-3955	320	7	reveals	reveal	VERB
gs-3955	320	8	that	that	SCONJ
gs-3955	320	9	negligible	negligible	ADJ
gs-3955	320	10	senescence	senescence	NOUN
gs-3955	320	11	arises	arise	VERB
gs-3955	320	12	in	in	ADP
gs-3955	320	13	environments	environment	NOUN
gs-3955	320	14	favoring	favor	VERB
gs-3955	320	15	extended	extend	VERB
gs-3955	320	16	somatic	somatic	ADJ
gs-3955	320	17	maintenance	maintenance	NOUN
gs-3955	320	18	over	over	ADP
gs-3955	320	19	rapid	rapid	ADJ
gs-3955	320	20	reproduction	reproduction	NOUN
gs-3955	320	21	.	.	PUNCT
gs-3955	321	1	k	k	ADJ
gs-3955	321	2	-	-	PUNCT
gs-3955	321	3	selected	select	VERB
gs-3955	321	4	species	specie	NOUN
gs-3955	321	5	like	like	ADP
gs-3955	321	6	the	the	DET
gs-3955	321	7	greenland	greenland	NOUN
gs-3955	321	8	shark	shark	NOUN
gs-3955	321	9	,	,	PUNCT
gs-3955	321	10	with	with	ADP
gs-3955	321	11	delayed	delay	VERB
gs-3955	321	12	maturation	maturation	NOUN
gs-3955	321	13	(	(	PUNCT
gs-3955	321	14	150	150	NUM
gs-3955	321	15	years	year	NOUN
gs-3955	321	16	)	)	PUNCT
gs-3955	321	17	and	and	CCONJ
gs-3955	321	18	minimal	minimal	ADJ
gs-3955	321	19	predation	predation	NOUN
gs-3955	321	20	pressure	pressure	NOUN
gs-3955	321	21	,	,	PUNCT
gs-3955	321	22	or	or	CCONJ
gs-3955	321	23	subterranean	subterranean	ADJ
gs-3955	321	24	naked	naked	ADJ
gs-3955	321	25	mole	mole	NOUN
gs-3955	321	26	-	-	PUNCT
gs-3955	321	27	rats	rat	NOUN
gs-3955	321	28	in	in	ADP
gs-3955	321	29	hypoxic	hypoxic	ADJ
gs-3955	321	30	,	,	PUNCT
gs-3955	321	31	stable	stable	ADJ
gs-3955	321	32	niches	niche	NOUN
gs-3955	321	33	,	,	PUNCT
gs-3955	321	34	align	align	VERB
gs-3955	321	35	with	with	ADP
gs-3955	321	36	the	the	DET
gs-3955	321	37	antagonistic	antagonistic	ADJ
gs-3955	321	38	pleiotropy	pleiotropy	NOUN
gs-3955	321	39	theory	theory	NOUN
gs-3955	321	40	’s	’s	PART
gs-3955	321	41	prediction	prediction	NOUN
gs-3955	321	42	:	:	PUNCT
gs-3955	321	43	low	low	ADJ
gs-3955	321	44	extrinsic	extrinsic	ADJ
gs-3955	321	45	mortality	mortality	NOUN
gs-3955	321	46	selects	select	NOUN
gs-3955	321	47	for	for	ADP
gs-3955	321	48	longevity	longevity	NOUN
gs-3955	321	49	mechanisms	mechanism	NOUN
gs-3955	321	50	.	.	PUNCT
gs-3955	322	1	the	the	DET
gs-3955	322	2	centrosomal	centrosomal	ADJ
gs-3955	322	3	theory	theory	NOUN
gs-3955	322	4	of	of	ADP
gs-3955	322	5	aging	aging	NOUN
gs-3955	322	6	further	far	ADV
gs-3955	322	7	explicates	explicate	VERB
gs-3955	322	8	this	this	PRON
gs-3955	322	9	by	by	ADP
gs-3955	322	10	framing	frame	VERB
gs-3955	322	11	senescence	senescence	NOUN
gs-3955	322	12	georgian	georgian	ADJ
gs-3955	322	13	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	322	14	მეცნიერები	მეცნიერები	NOUN
gs-3955	322	15	ტ.	ტ.	VERB
gs-3955	322	16	7	7	NUM
gs-3955	322	17	n	n	DET
gs-3955	322	18	3	3	NUM
gs-3955	322	19	,	,	PUNCT
gs-3955	322	20	2025	2025	NUM
gs-3955	322	21	383	383	NUM
gs-3955	322	22	as	as	ADP
gs-3955	322	23	a	a	DET
gs-3955	322	24	thermodynamic	thermodynamic	ADJ
gs-3955	322	25	byproduct	byproduct	NOUN
gs-3955	322	26	of	of	ADP
gs-3955	322	27	asymmetric	asymmetric	ADJ
gs-3955	322	28	stem	stem	NOUN
gs-3955	322	29	cell	cell	NOUN
gs-3955	322	30	division	division	NOUN
gs-3955	322	31	—	—	PUNCT
gs-3955	322	32	a	a	DET
gs-3955	322	33	process	process	NOUN
gs-3955	322	34	circumvented	circumvent	VERB
gs-3955	322	35	by	by	ADP
gs-3955	322	36	species	specie	NOUN
gs-3955	322	37	like	like	ADP
gs-3955	322	38	planarians	planarian	NOUN
gs-3955	322	39	through	through	ADP
gs-3955	322	40	symmetric	symmetric	ADJ
gs-3955	322	41	centriole	centriole	NOUN
gs-3955	322	42	distribution	distribution	NOUN
gs-3955	322	43	.	.	PUNCT
gs-3955	323	1	translational	translational	ADJ
gs-3955	323	2	prospects	prospect	NOUN
gs-3955	323	3	are	be	AUX
gs-3955	323	4	profound	profound	ADJ
gs-3955	323	5	yet	yet	ADV
gs-3955	323	6	complex	complex	ADJ
gs-3955	323	7	.	.	PUNCT
gs-3955	324	1	conserved	conserved	ADJ
gs-3955	324	2	pathways	pathway	NOUN
gs-3955	324	3	—	—	PUNCT
gs-3955	324	4	such	such	ADJ
gs-3955	324	5	as	as	ADP
gs-3955	324	6	telomerase	telomerase	NOUN
gs-3955	324	7	activation	activation	NOUN
gs-3955	324	8	,	,	PUNCT
gs-3955	324	9	hmw	hmw	PROPN
gs-3955	324	10	-	-	PUNCT
gs-3955	324	11	ha	ha	ADV
gs-3955	324	12	-	-	PUNCT
gs-3955	324	13	mediated	mediate	VERB
gs-3955	324	14	tumor	tumor	NOUN
gs-3955	324	15	suppression	suppression	NOUN
gs-3955	324	16	,	,	PUNCT
gs-3955	324	17	and	and	CCONJ
gs-3955	324	18	autophagy	autophagy	ADJ
gs-3955	324	19	enhancement	enhancement	NOUN
gs-3955	324	20	—	—	PUNCT
gs-3955	324	21	offer	offer	VERB
gs-3955	324	22	targets	target	NOUN
gs-3955	324	23	for	for	ADP
gs-3955	324	24	human	human	ADJ
gs-3955	324	25	interventions	intervention	NOUN
gs-3955	324	26	.	.	PUNCT
gs-3955	325	1	however	however	ADV
gs-3955	325	2	,	,	PUNCT
gs-3955	325	3	interspecies	interspecie	VERB
gs-3955	325	4	differences	difference	NOUN
gs-3955	325	5	in	in	ADP
gs-3955	325	6	life	life	NOUN
gs-3955	325	7	history	history	NOUN
gs-3955	325	8	and	and	CCONJ
gs-3955	325	9	genetic	genetic	ADJ
gs-3955	325	10	networks	network	NOUN
gs-3955	325	11	necessitate	necessitate	ADJ
gs-3955	325	12	cautious	cautious	ADJ
gs-3955	325	13	extrapolation	extrapolation	NOUN
gs-3955	325	14	.	.	PUNCT
gs-3955	326	1	for	for	ADP
gs-3955	326	2	example	example	NOUN
gs-3955	326	3	,	,	PUNCT
gs-3955	326	4	while	while	SCONJ
gs-3955	326	5	naked	naked	ADJ
gs-3955	326	6	mole	mole	ADJ
gs-3955	326	7	-	-	PUNCT
gs-3955	326	8	rat	rat	NOUN
gs-3955	326	9	hmw	hmw	PROPN
gs-3955	326	10	-	-	PUNCT
gs-3955	326	11	ha	ha	PROPN
gs-3955	326	12	inspires	inspire	VERB
gs-3955	326	13	anticancer	anticancer	NOUN
gs-3955	326	14	therapies	therapy	NOUN
gs-3955	326	15	,	,	PUNCT
gs-3955	326	16	and	and	CCONJ
gs-3955	326	17	greenland	greenland	PROPN
gs-3955	326	18	shark	shark	NOUN
gs-3955	326	19	membrane	membrane	NOUN
gs-3955	326	20	lipids	lipid	NOUN
gs-3955	326	21	inform	inform	VERB
gs-3955	326	22	cytoprotective	cytoprotective	ADJ
gs-3955	326	23	strategies	strategy	NOUN
gs-3955	326	24	,	,	PUNCT
gs-3955	326	25	their	their	PRON
gs-3955	326	26	efficacy	efficacy	NOUN
gs-3955	326	27	in	in	ADP
gs-3955	326	28	mammals	mammal	NOUN
gs-3955	326	29	requires	require	VERB
gs-3955	326	30	rigorous	rigorous	ADJ
gs-3955	326	31	validation	validation	NOUN
gs-3955	326	32	.	.	PUNCT
gs-3955	327	1	future	future	ADJ
gs-3955	327	2	research	research	NOUN
gs-3955	327	3	must	must	AUX
gs-3955	327	4	prioritize	prioritize	VERB
gs-3955	327	5	comparative	comparative	ADJ
gs-3955	327	6	genomics	genomic	NOUN
gs-3955	327	7	and	and	CCONJ
gs-3955	327	8	proteomics	proteomic	NOUN
gs-3955	327	9	to	to	PART
gs-3955	327	10	identify	identify	VERB
gs-3955	327	11	core	core	NOUN
gs-3955	327	12	,	,	PUNCT
gs-3955	327	13	transferable	transferable	ADJ
gs-3955	327	14	longevity	longevity	NOUN
gs-3955	327	15	mechanisms	mechanism	NOUN
gs-3955	327	16	while	while	SCONJ
gs-3955	327	17	addressing	address	VERB
gs-3955	327	18	ethical	ethical	ADJ
gs-3955	327	19	and	and	CCONJ
gs-3955	327	20	biological	biological	ADJ
gs-3955	327	21	challenges	challenge	NOUN
gs-3955	327	22	in	in	ADP
gs-3955	327	23	clinical	clinical	ADJ
gs-3955	327	24	translation	translation	NOUN
gs-3955	327	25	(	(	PUNCT
gs-3955	327	26	tian	tian	PROPN
gs-3955	327	27	et	et	PROPN
gs-3955	327	28	al	al	PROPN
gs-3955	327	29	.	.	PROPN
gs-3955	327	30	,	,	PUNCT
gs-3955	327	31	2013	2013	NUM
gs-3955	327	32	,	,	PUNCT
gs-3955	327	33	petruseva	petruseva	VERB
gs-3955	327	34	et	et	PROPN
gs-3955	327	35	al	al	PROPN
gs-3955	327	36	.	.	PROPN
gs-3955	327	37	,	,	PUNCT
gs-3955	327	38	2017	2017	NUM
gs-3955	327	39	)	)	PUNCT
gs-3955	327	40	.	.	PUNCT
gs-3955	328	1	the	the	DET
gs-3955	328	2	negligibly	negligibly	ADV
gs-3955	328	3	senescing	senesce	VERB
gs-3955	328	4	species	specie	NOUN
gs-3955	328	5	exemplify	exemplify	VERB
gs-3955	328	6	nature	nature	NOUN
gs-3955	328	7	’s	’s	PART
gs-3955	328	8	capacity	capacity	NOUN
gs-3955	328	9	to	to	PART
gs-3955	328	10	defy	defy	VERB
gs-3955	328	11	aging	age	VERB
gs-3955	328	12	through	through	ADP
gs-3955	328	13	integrated	integrate	VERB
gs-3955	328	14	molecular	molecular	ADJ
gs-3955	328	15	networks	network	NOUN
gs-3955	328	16	.	.	PUNCT
gs-3955	329	1	their	their	PRON
gs-3955	329	2	study	study	NOUN
gs-3955	329	3	not	not	PART
gs-3955	329	4	only	only	ADV
gs-3955	329	5	redefines	redefine	VERB
gs-3955	329	6	the	the	DET
gs-3955	329	7	biological	biological	ADJ
gs-3955	329	8	boundaries	boundary	NOUN
gs-3955	329	9	of	of	ADP
gs-3955	329	10	lifespan	lifespan	NOUN
gs-3955	329	11	but	but	CCONJ
gs-3955	329	12	also	also	ADV
gs-3955	329	13	illuminates	illuminate	VERB
gs-3955	329	14	actionable	actionable	ADJ
gs-3955	329	15	pathways	pathway	NOUN
gs-3955	329	16	to	to	PART
gs-3955	329	17	mitigate	mitigate	VERB
gs-3955	329	18	age	age	NOUN
gs-3955	329	19	-	-	PUNCT
gs-3955	329	20	related	relate	VERB
gs-3955	329	21	pathology	pathology	NOUN
gs-3955	329	22	in	in	ADP
gs-3955	329	23	humans	human	NOUN
gs-3955	329	24	,	,	PUNCT
gs-3955	329	25	heralding	herald	VERB
gs-3955	329	26	a	a	DET
gs-3955	329	27	new	new	ADJ
gs-3955	329	28	era	era	NOUN
gs-3955	329	29	in	in	ADP
gs-3955	329	30	geroscience	geroscience	NOUN
gs-3955	329	31	.	.	PUNCT
gs-3955	330	1	the	the	DET
gs-3955	330	2	molecular	molecular	ADJ
gs-3955	330	3	blueprints	blueprint	NOUN
gs-3955	330	4	of	of	ADP
gs-3955	330	5	negligibly	negligibly	ADV
gs-3955	330	6	senescent	senescent	ADJ
gs-3955	330	7	species	specie	NOUN
gs-3955	330	8	—	—	PUNCT
gs-3955	330	9	particularly	particularly	ADV
gs-3955	330	10	their	their	PRON
gs-3955	330	11	centrosomal	centrosomal	ADJ
gs-3955	330	12	stability	stability	NOUN
gs-3955	330	13	,	,	PUNCT
gs-3955	330	14	enhanced	enhance	VERB
gs-3955	330	15	dna	dna	PROPN
gs-3955	330	16	repair	repair	NOUN
gs-3955	330	17	,	,	PUNCT
gs-3955	330	18	and	and	CCONJ
gs-3955	330	19	stress	stress	NOUN
gs-3955	330	20	-	-	PUNCT
gs-3955	330	21	resistant	resistant	ADJ
gs-3955	330	22	proteostasis	proteostasis	NOUN
gs-3955	330	23	—	—	PUNCT
gs-3955	330	24	offer	offer	VERB
gs-3955	330	25	unprecedented	unprecedented	ADJ
gs-3955	330	26	opportunities	opportunity	NOUN
gs-3955	330	27	to	to	PART
gs-3955	330	28	redefine	redefine	VERB
gs-3955	330	29	human	human	ADJ
gs-3955	330	30	aging	aging	NOUN
gs-3955	330	31	.	.	PUNCT
gs-3955	331	1	prioritize	prioritize	VERB
gs-3955	331	2	translational	translational	ADJ
gs-3955	331	3	research	research	NOUN
gs-3955	331	4	on	on	ADP
gs-3955	331	5	conserved	conserved	ADJ
gs-3955	331	6	pathways	pathway	NOUN
gs-3955	331	7	(	(	PUNCT
gs-3955	331	8	e.g.	e.g.	ADV
gs-3955	331	9	,	,	PUNCT
gs-3955	331	10	hmw	hmw	PROPN
gs-3955	331	11	-	-	PUNCT
gs-3955	331	12	ha	ha	PROPN
gs-3955	331	13	,	,	PUNCT
gs-3955	331	14	sirt6	sirt6	NOUN
gs-3955	331	15	,	,	PUNCT
gs-3955	331	16	nrf2	nrf2	NOUN
gs-3955	331	17	)	)	PUNCT
gs-3955	331	18	while	while	SCONJ
gs-3955	331	19	proactively	proactively	ADV
gs-3955	331	20	addressing	address	VERB
gs-3955	331	21	ethical	ethical	ADJ
gs-3955	331	22	and	and	CCONJ
gs-3955	331	23	societal	societal	ADJ
gs-3955	331	24	implications	implication	NOUN
gs-3955	331	25	.	.	PUNCT
gs-3955	332	1	by	by	ADP
gs-3955	332	2	emulating	emulate	VERB
gs-3955	332	3	nature	nature	NOUN
gs-3955	332	4	’s	’s	PART
gs-3955	332	5	mastery	mastery	NOUN
gs-3955	332	6	of	of	ADP
gs-3955	332	7	biological	biological	ADJ
gs-3955	332	8	time	time	NOUN
gs-3955	332	9	,	,	PUNCT
gs-3955	332	10	we	we	PRON
gs-3955	332	11	can	can	AUX
gs-3955	332	12	transform	transform	VERB
gs-3955	332	13	aging	aging	NOUN
gs-3955	332	14	from	from	ADP
gs-3955	332	15	an	an	DET
gs-3955	332	16	inevitability	inevitability	NOUN
gs-3955	332	17	to	to	ADP
gs-3955	332	18	a	a	DET
gs-3955	332	19	treatable	treatable	ADJ
gs-3955	332	20	condition	condition	NOUN
gs-3955	332	21	.	.	PUNCT
gs-3955	333	1	hypothesis	hypothesis	NOUN
gs-3955	333	2	:	:	PUNCT
gs-3955	333	3	integrative	integrative	ADJ
gs-3955	333	4	molecular	molecular	ADJ
gs-3955	333	5	and	and	CCONJ
gs-3955	333	6	systems	system	NOUN
gs-3955	333	7	-	-	PUNCT
gs-3955	333	8	level	level	NOUN
gs-3955	333	9	mechanisms	mechanism	NOUN
gs-3955	333	10	underlying	underlie	VERB
gs-3955	333	11	negligible	negligible	ADJ
gs-3955	333	12	senescence	senescence	NOUN
gs-3955	333	13	despite	despite	SCONJ
gs-3955	333	14	extensive	extensive	ADJ
gs-3955	333	15	characterization	characterization	NOUN
gs-3955	333	16	of	of	ADP
gs-3955	333	17	molecular	molecular	ADJ
gs-3955	333	18	pathways	pathway	NOUN
gs-3955	333	19	implicated	implicate	VERB
gs-3955	333	20	in	in	ADP
gs-3955	333	21	longevity	longevity	NOUN
gs-3955	333	22	among	among	ADP
gs-3955	333	23	negligibly	negligibly	ADV
gs-3955	333	24	senescent	senescent	ADJ
gs-3955	333	25	species	specie	NOUN
gs-3955	333	26	,	,	PUNCT
gs-3955	333	27	a	a	DET
gs-3955	333	28	unifying	unifying	ADJ
gs-3955	333	29	explanatory	explanatory	ADJ
gs-3955	333	30	framework	framework	NOUN
gs-3955	333	31	remains	remain	VERB
gs-3955	333	32	elusive	elusive	ADJ
gs-3955	333	33	.	.	PUNCT
gs-3955	334	1	we	we	PRON
gs-3955	334	2	propose	propose	VERB
gs-3955	334	3	a	a	DET
gs-3955	334	4	novel	novel	ADJ
gs-3955	334	5	hypothesis	hypothesis	NOUN
gs-3955	334	6	integrating	integrate	VERB
gs-3955	334	7	emergent	emergent	ADJ
gs-3955	334	8	principles	principle	NOUN
gs-3955	334	9	from	from	ADP
gs-3955	334	10	molecular	molecular	ADJ
gs-3955	334	11	biology	biology	NOUN
gs-3955	334	12	,	,	PUNCT
gs-3955	334	13	systems	system	NOUN
gs-3955	334	14	physiology	physiology	NOUN
gs-3955	334	15	,	,	PUNCT
gs-3955	334	16	and	and	CCONJ
gs-3955	334	17	evolutionary	evolutionary	ADJ
gs-3955	334	18	dynamics	dynamic	NOUN
gs-3955	334	19	to	to	PART
gs-3955	334	20	explain	explain	VERB
gs-3955	334	21	how	how	SCONJ
gs-3955	334	22	these	these	DET
gs-3955	334	23	organisms	organism	NOUN
gs-3955	334	24	achieve	achieve	VERB
gs-3955	334	25	extraordinary	extraordinary	ADJ
gs-3955	334	26	lifespan	lifespan	NOUN
gs-3955	334	27	and	and	CCONJ
gs-3955	334	28	negligible	negligible	ADJ
gs-3955	334	29	senescence	senescence	NOUN
gs-3955	334	30	(	(	PUNCT
gs-3955	334	31	picture	picture	NOUN
gs-3955	334	32	1	1	NUM
gs-3955	334	33	)	)	PUNCT
gs-3955	334	34	.	.	PUNCT
gs-3955	335	1	coordinated	coordinate	VERB
gs-3955	335	2	entropy	entropy	NOUN
gs-3955	335	3	management	management	NOUN
gs-3955	335	4	at	at	ADP
gs-3955	335	5	cellular	cellular	ADJ
gs-3955	335	6	control	control	NOUN
gs-3955	335	7	hubs	hub	NOUN
gs-3955	335	8	building	build	VERB
gs-3955	335	9	upon	upon	SCONJ
gs-3955	335	10	the	the	DET
gs-3955	335	11	centrosomal	centrosomal	ADJ
gs-3955	335	12	theory	theory	NOUN
gs-3955	335	13	of	of	ADP
gs-3955	335	14	aging	age	VERB
gs-3955	335	15	(	(	PUNCT
gs-3955	335	16	tkemaladze	tkemaladze	NOUN
gs-3955	335	17	,	,	PUNCT
gs-3955	335	18	2023	2023	NUM
gs-3955	335	19	)	)	PUNCT
gs-3955	335	20	,	,	PUNCT
gs-3955	335	21	we	we	PRON
gs-3955	335	22	hypothesize	hypothesize	VERB
gs-3955	335	23	that	that	SCONJ
gs-3955	335	24	negligible	negligible	ADJ
gs-3955	335	25	senescence	senescence	NOUN
gs-3955	335	26	arises	arise	VERB
gs-3955	335	27	from	from	ADP
gs-3955	335	28	an	an	DET
gs-3955	335	29	enhanced	enhanced	ADJ
gs-3955	335	30	capacity	capacity	NOUN
gs-3955	335	31	to	to	PART
gs-3955	335	32	mitigate	mitigate	VERB
gs-3955	335	33	entropic	entropic	ADJ
gs-3955	335	34	damage	damage	NOUN
gs-3955	335	35	specifically	specifically	ADV
gs-3955	335	36	at	at	ADP
gs-3955	335	37	critical	critical	ADJ
gs-3955	335	38	cellular	cellular	ADJ
gs-3955	335	39	control	control	NOUN
gs-3955	335	40	centers	center	NOUN
gs-3955	335	41	,	,	PUNCT
gs-3955	335	42	including	include	VERB
gs-3955	335	43	the	the	DET
gs-3955	335	44	centrosome	centrosome	NOUN
gs-3955	335	45	,	,	PUNCT
gs-3955	335	46	nucleus	nucleus	NOUN
gs-3955	335	47	,	,	PUNCT
gs-3955	335	48	and	and	CCONJ
gs-3955	335	49	mitochondrial	mitochondrial	ADJ
gs-3955	335	50	networks	network	NOUN
gs-3955	335	51	.	.	PUNCT
gs-3955	336	1	these	these	DET
gs-3955	336	2	organelles	organelle	NOUN
gs-3955	336	3	act	act	VERB
gs-3955	336	4	as	as	ADP
gs-3955	336	5	“	"	PUNCT
gs-3955	336	6	information	information	NOUN
gs-3955	336	7	bottlenecks	bottleneck	NOUN
gs-3955	336	8	”	"	PUNCT
gs-3955	336	9	whose	whose	DET
gs-3955	336	10	integrity	integrity	NOUN
gs-3955	336	11	governs	govern	VERB
gs-3955	336	12	cellular	cellular	ADJ
gs-3955	336	13	fate	fate	NOUN
gs-3955	336	14	.	.	PUNCT
gs-3955	337	1	species	specie	NOUN
gs-3955	337	2	such	such	ADJ
gs-3955	337	3	as	as	ADP
gs-3955	337	4	planarians	planarian	NOUN
gs-3955	337	5	and	and	CCONJ
gs-3955	337	6	hydra	hydra	PROPN
gs-3955	337	7	demonstrate	demonstrate	VERB
gs-3955	337	8	active	active	ADJ
gs-3955	337	9	centrosomal	centrosomal	ADJ
gs-3955	337	10	rejuvenation	rejuvenation	NOUN
gs-3955	337	11	and	and	CCONJ
gs-3955	337	12	selective	selective	ADJ
gs-3955	337	13	autophagy	autophagy	NOUN
gs-3955	337	14	of	of	ADP
gs-3955	337	15	damaged	damage	VERB
gs-3955	337	16	centrioles	centriole	NOUN
gs-3955	337	17	(	(	PUNCT
gs-3955	337	18	wagner	wagner	PROPN
gs-3955	337	19	et	et	PROPN
gs-3955	337	20	al	al	PROPN
gs-3955	337	21	.	.	PROPN
gs-3955	337	22	,	,	PUNCT
gs-3955	337	23	2011	2011	NUM
gs-3955	337	24	;	;	PUNCT
gs-3955	337	25	pirotte	pirotte	NOUN
gs-3955	337	26	et	et	PROPN
gs-3955	337	27	al	al	PROPN
gs-3955	337	28	.	.	PROPN
gs-3955	337	29	,	,	PUNCT
gs-3955	337	30	2015	2015	NUM
gs-3955	337	31	)	)	PUNCT
gs-3955	337	32	,	,	PUNCT
gs-3955	337	33	suggesting	suggest	VERB
gs-3955	337	34	that	that	SCONJ
gs-3955	337	35	targeted	target	VERB
gs-3955	337	36	entropy	entropy	NOUN
gs-3955	337	37	reduction	reduction	NOUN
gs-3955	337	38	at	at	ADP
gs-3955	337	39	these	these	DET
gs-3955	337	40	nodes	node	NOUN
gs-3955	337	41	prevents	prevent	VERB
gs-3955	337	42	the	the	DET
gs-3955	337	43	cascade	cascade	NOUN
gs-3955	337	44	of	of	ADP
gs-3955	337	45	georgian	georgian	ADJ
gs-3955	337	46	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	337	47	მეცნიერები	მეცნიერები	NOUN
gs-3955	337	48	ტ.	ტ.	VERB
gs-3955	337	49	7	7	NUM
gs-3955	337	50	n	n	DET
gs-3955	337	51	3	3	NUM
gs-3955	337	52	,	,	PUNCT
gs-3955	337	53	2025	2025	NUM
gs-3955	337	54	384	384	NUM
gs-3955	337	55	molecular	molecular	ADJ
gs-3955	337	56	dysfunction	dysfunction	NOUN
gs-3955	337	57	typical	typical	ADJ
gs-3955	337	58	of	of	ADP
gs-3955	337	59	aging	aging	NOUN
gs-3955	337	60	.	.	PUNCT
gs-3955	338	1	this	this	DET
gs-3955	338	2	refined	refined	ADJ
gs-3955	338	3	damage	damage	NOUN
gs-3955	338	4	control	control	NOUN
gs-3955	338	5	extends	extend	VERB
gs-3955	338	6	to	to	ADP
gs-3955	338	7	chromatin	chromatin	NOUN
gs-3955	338	8	architecture	architecture	NOUN
gs-3955	338	9	,	,	PUNCT
gs-3955	338	10	where	where	SCONJ
gs-3955	338	11	naked	naked	ADJ
gs-3955	338	12	mole	mole	NOUN
gs-3955	338	13	-	-	PUNCT
gs-3955	338	14	rats	rat	NOUN
gs-3955	338	15	and	and	CCONJ
gs-3955	338	16	greenland	greenland	PROPN
gs-3955	338	17	sharks	sharks	PROPN
gs-3955	338	18	maintain	maintain	VERB
gs-3955	338	19	telomeric	telomeric	ADJ
gs-3955	338	20	and	and	CCONJ
gs-3955	338	21	epigenetic	epigenetic	ADJ
gs-3955	338	22	stability	stability	NOUN
gs-3955	338	23	via	via	ADP
gs-3955	338	24	unique	unique	ADJ
gs-3955	338	25	histone	histone	NOUN
gs-3955	338	26	variants	variant	NOUN
gs-3955	338	27	and	and	CCONJ
gs-3955	338	28	repair	repair	NOUN
gs-3955	338	29	enzymes	enzyme	NOUN
gs-3955	338	30	(	(	PUNCT
gs-3955	338	31	macrae	macrae	PROPN
gs-3955	338	32	et	et	PROPN
gs-3955	338	33	al	al	PROPN
gs-3955	338	34	.	.	PROPN
gs-3955	338	35	,	,	PUNCT
gs-3955	338	36	2015	2015	NUM
gs-3955	338	37	;	;	PUNCT
gs-3955	338	38	petruseva	petruseva	VERB
gs-3955	338	39	et	et	PROPN
gs-3955	338	40	al	al	PROPN
gs-3955	338	41	.	.	PROPN
gs-3955	338	42	,	,	PUNCT
gs-3955	338	43	2017	2017	NUM
gs-3955	338	44	)	)	PUNCT
gs-3955	338	45	.	.	PUNCT
gs-3955	339	1	such	such	ADJ
gs-3955	339	2	systems	system	NOUN
gs-3955	339	3	-	-	PUNCT
gs-3955	339	4	level	level	NOUN
gs-3955	339	5	entropy	entropy	NOUN
gs-3955	339	6	management	management	NOUN
gs-3955	339	7	may	may	AUX
gs-3955	339	8	function	function	VERB
gs-3955	339	9	as	as	ADP
gs-3955	339	10	a	a	DET
gs-3955	339	11	“	"	PUNCT
gs-3955	339	12	molecular	molecular	ADJ
gs-3955	339	13	firewall	firewall	NOUN
gs-3955	339	14	,	,	PUNCT
gs-3955	339	15	”	"	PUNCT
gs-3955	339	16	confining	confine	VERB
gs-3955	339	17	damage	damage	NOUN
gs-3955	339	18	and	and	CCONJ
gs-3955	339	19	preserving	preserve	VERB
gs-3955	339	20	youthful	youthful	ADJ
gs-3955	339	21	cellular	cellular	ADJ
gs-3955	339	22	states	state	NOUN
gs-3955	339	23	indefinitely	indefinitely	ADV
gs-3955	339	24	.	.	PUNCT
gs-3955	340	1	evolution	evolution	NOUN
gs-3955	340	2	of	of	ADP
gs-3955	340	3	metabolic	metabolic	NOUN
gs-3955	340	4	flexibility	flexibility	NOUN
gs-3955	340	5	as	as	ADP
gs-3955	340	6	a	a	DET
gs-3955	340	7	longevity	longevity	NOUN
gs-3955	340	8	catalyst	catalyst	NOUN
gs-3955	340	9	a	a	DET
gs-3955	340	10	second	second	ADJ
gs-3955	340	11	pillar	pillar	NOUN
gs-3955	340	12	of	of	ADP
gs-3955	340	13	our	our	PRON
gs-3955	340	14	hypothesis	hypothesis	NOUN
gs-3955	340	15	emphasizes	emphasize	VERB
gs-3955	340	16	metabolic	metabolic	NOUN
gs-3955	340	17	plasticity	plasticity	NOUN
gs-3955	340	18	as	as	ADP
gs-3955	340	19	a	a	DET
gs-3955	340	20	critical	critical	ADJ
gs-3955	340	21	longevity	longevity	NOUN
gs-3955	340	22	enabler	enabler	NOUN
gs-3955	340	23	.	.	PUNCT
gs-3955	341	1	long	long	ADV
gs-3955	341	2	-	-	PUNCT
gs-3955	341	3	lived	live	VERB
gs-3955	341	4	species	species	NOUN
gs-3955	341	5	inhabit	inhabit	NOUN
gs-3955	341	6	environments	environment	NOUN
gs-3955	341	7	marked	mark	VERB
gs-3955	341	8	by	by	ADP
gs-3955	341	9	fluctuating	fluctuate	VERB
gs-3955	341	10	oxygen	oxygen	NOUN
gs-3955	341	11	availability	availability	NOUN
gs-3955	341	12	and	and	CCONJ
gs-3955	341	13	resource	resource	NOUN
gs-3955	341	14	scarcity	scarcity	NOUN
gs-3955	341	15	.	.	PUNCT
gs-3955	342	1	they	they	PRON
gs-3955	342	2	have	have	AUX
gs-3955	342	3	evolved	evolve	VERB
gs-3955	342	4	not	not	PART
gs-3955	342	5	merely	merely	ADV
gs-3955	342	6	to	to	PART
gs-3955	342	7	endure	endure	VERB
gs-3955	342	8	these	these	DET
gs-3955	342	9	stresses	stress	NOUN
gs-3955	342	10	but	but	CCONJ
gs-3955	342	11	to	to	PART
gs-3955	342	12	harness	harness	VERB
gs-3955	342	13	them	they	PRON
gs-3955	342	14	adaptively	adaptively	ADV
gs-3955	342	15	.	.	PUNCT
gs-3955	343	1	for	for	ADP
gs-3955	343	2	example	example	NOUN
gs-3955	343	3	,	,	PUNCT
gs-3955	343	4	arctica	arctica	PROPN
gs-3955	343	5	islandica	islandica	PROPN
gs-3955	343	6	and	and	CCONJ
gs-3955	343	7	naked	naked	ADJ
gs-3955	343	8	mole	mole	NOUN
gs-3955	343	9	-	-	PUNCT
gs-3955	343	10	rats	rat	NOUN
gs-3955	343	11	display	display	VERB
gs-3955	343	12	profound	profound	ADJ
gs-3955	343	13	metabolic	metabolic	NOUN
gs-3955	343	14	depression	depression	NOUN
gs-3955	343	15	and	and	CCONJ
gs-3955	343	16	hypoxia	hypoxia	NOUN
gs-3955	343	17	tolerance	tolerance	NOUN
gs-3955	343	18	(	(	PUNCT
gs-3955	343	19	munro	munro	PROPN
gs-3955	343	20	et	et	PROPN
gs-3955	343	21	al	al	PROPN
gs-3955	343	22	.	.	PROPN
gs-3955	343	23	,	,	PUNCT
gs-3955	343	24	2012	2012	NUM
gs-3955	343	25	;	;	PUNCT
gs-3955	343	26	lewis	lewis	PROPN
gs-3955	343	27	et	et	PROPN
gs-3955	343	28	al	al	PROPN
gs-3955	343	29	.	.	PROPN
gs-3955	343	30	,	,	PUNCT
gs-3955	343	31	2012	2012	NUM
gs-3955	343	32	)	)	PUNCT
gs-3955	343	33	,	,	PUNCT
gs-3955	343	34	while	while	SCONJ
gs-3955	343	35	the	the	DET
gs-3955	343	36	olm	olm	NOUN
gs-3955	343	37	’s	’s	PART
gs-3955	343	38	stem	stem	NOUN
gs-3955	343	39	cells	cell	NOUN
gs-3955	343	40	utilize	utilize	VERB
gs-3955	343	41	ketone	ketone	NOUN
gs-3955	343	42	body	body	NOUN
gs-3955	343	43	metabolism	metabolism	NOUN
gs-3955	343	44	to	to	PART
gs-3955	343	45	sustain	sustain	VERB
gs-3955	343	46	pluripotency	pluripotency	NOUN
gs-3955	343	47	(	(	PUNCT
gs-3955	343	48	narayan	narayan	PROPN
gs-3955	343	49	et	et	PROPN
gs-3955	343	50	al	al	PROPN
gs-3955	343	51	.	.	PROPN
gs-3955	343	52	,	,	PUNCT
gs-3955	343	53	2021	2021	NUM
gs-3955	343	54	)	)	PUNCT
gs-3955	343	55	.	.	PUNCT
gs-3955	344	1	we	we	PRON
gs-3955	344	2	speculate	speculate	VERB
gs-3955	344	3	that	that	SCONJ
gs-3955	344	4	dynamic	dynamic	ADJ
gs-3955	344	5	metabolic	metabolic	NOUN
gs-3955	344	6	switching	switching	NOUN
gs-3955	344	7	—	—	PUNCT
gs-3955	344	8	between	between	ADP
gs-3955	344	9	oxidative	oxidative	ADJ
gs-3955	344	10	phosphorylation	phosphorylation	NOUN
gs-3955	344	11	,	,	PUNCT
gs-3955	344	12	anaerobic	anaerobic	NOUN
gs-3955	344	13	pathways	pathway	NOUN
gs-3955	344	14	,	,	PUNCT
gs-3955	344	15	and	and	CCONJ
gs-3955	344	16	even	even	ADV
gs-3955	344	17	quantum	quantum	ADJ
gs-3955	344	18	-	-	ADJ
gs-3955	344	19	biological	biological	ADJ
gs-3955	344	20	electron	electron	NOUN
gs-3955	344	21	transport	transport	NOUN
gs-3955	344	22	modes	mode	NOUN
gs-3955	344	23	—	—	PUNCT
gs-3955	344	24	serves	serve	VERB
gs-3955	344	25	to	to	PART
gs-3955	344	26	minimize	minimize	VERB
gs-3955	344	27	reactive	reactive	ADJ
gs-3955	344	28	oxygen	oxygen	NOUN
gs-3955	344	29	species	specie	NOUN
gs-3955	344	30	generation	generation	NOUN
gs-3955	344	31	and	and	CCONJ
gs-3955	344	32	optimize	optimize	VERB
gs-3955	344	33	resource	resource	NOUN
gs-3955	344	34	allocation	allocation	NOUN
gs-3955	344	35	for	for	ADP
gs-3955	344	36	repair	repair	NOUN
gs-3955	344	37	and	and	CCONJ
gs-3955	344	38	maintenance	maintenance	NOUN
gs-3955	344	39	.	.	PUNCT
gs-3955	345	1	this	this	DET
gs-3955	345	2	metabolic	metabolic	NOUN
gs-3955	345	3	“	"	PUNCT
gs-3955	345	4	metronome	metronome	NOUN
gs-3955	345	5	”	"	PUNCT
gs-3955	345	6	may	may	AUX
gs-3955	345	7	synchronize	synchronize	VERB
gs-3955	345	8	cellular	cellular	ADJ
gs-3955	345	9	and	and	CCONJ
gs-3955	345	10	organismal	organismal	ADJ
gs-3955	345	11	rhythms	rhythm	NOUN
gs-3955	345	12	,	,	PUNCT
gs-3955	345	13	coordinating	coordinate	VERB
gs-3955	345	14	damage	damage	NOUN
gs-3955	345	15	repair	repair	NOUN
gs-3955	345	16	cycles	cycle	NOUN
gs-3955	345	17	with	with	ADP
gs-3955	345	18	environmental	environmental	ADJ
gs-3955	345	19	inputs	input	NOUN
gs-3955	345	20	,	,	PUNCT
gs-3955	345	21	effectively	effectively	ADV
gs-3955	345	22	decoupling	decouple	VERB
gs-3955	345	23	chronological	chronological	ADJ
gs-3955	345	24	time	time	NOUN
gs-3955	345	25	from	from	ADP
gs-3955	345	26	biological	biological	ADJ
gs-3955	345	27	aging	aging	NOUN
gs-3955	345	28	.	.	PUNCT
gs-3955	346	1	proteostasis	proteostasis	NOUN
gs-3955	346	2	as	as	ADP
gs-3955	346	3	a	a	DET
gs-3955	346	4	multilayered	multilayered	ADJ
gs-3955	346	5	defense	defense	NOUN
gs-3955	346	6	network	network	NOUN
gs-3955	346	7	the	the	DET
gs-3955	346	8	remarkable	remarkable	ADJ
gs-3955	346	9	proteostatic	proteostatic	ADJ
gs-3955	346	10	systems	system	NOUN
gs-3955	346	11	observed	observe	VERB
gs-3955	346	12	in	in	ADP
gs-3955	346	13	these	these	DET
gs-3955	346	14	organisms	organism	NOUN
gs-3955	346	15	suggest	suggest	VERB
gs-3955	346	16	that	that	SCONJ
gs-3955	346	17	longevity	longevity	NOUN
gs-3955	346	18	is	be	AUX
gs-3955	346	19	not	not	PART
gs-3955	346	20	merely	merely	ADV
gs-3955	346	21	the	the	DET
gs-3955	346	22	result	result	NOUN
gs-3955	346	23	of	of	ADP
gs-3955	346	24	enhanced	enhanced	ADJ
gs-3955	346	25	repair	repair	NOUN
gs-3955	346	26	but	but	CCONJ
gs-3955	346	27	of	of	ADP
gs-3955	346	28	multilayered	multilayered	ADJ
gs-3955	346	29	prevention	prevention	NOUN
gs-3955	346	30	and	and	CCONJ
gs-3955	346	31	clearance	clearance	NOUN
gs-3955	346	32	mechanisms	mechanism	NOUN
gs-3955	346	33	working	work	VERB
gs-3955	346	34	in	in	ADP
gs-3955	346	35	concert	concert	NOUN
gs-3955	346	36	.	.	PUNCT
gs-3955	347	1	beyond	beyond	ADP
gs-3955	347	2	canonical	canonical	ADJ
gs-3955	347	3	chaperones	chaperone	NOUN
gs-3955	347	4	and	and	CCONJ
gs-3955	347	5	proteasomes	proteasome	NOUN
gs-3955	347	6	,	,	PUNCT
gs-3955	347	7	species	specie	NOUN
gs-3955	347	8	like	like	ADP
gs-3955	347	9	arctica	arctica	PROPN
gs-3955	347	10	islandica	islandica	PROPN
gs-3955	347	11	possess	possess	VERB
gs-3955	347	12	temperature	temperature	NOUN
gs-3955	347	13	-	-	PUNCT
gs-3955	347	14	insensitive	insensitive	ADJ
gs-3955	347	15	chaperones	chaperone	NOUN
gs-3955	347	16	with	with	ADP
gs-3955	347	17	unique	unique	ADJ
gs-3955	347	18	amino	amino	NOUN
gs-3955	347	19	acid	acid	NOUN
gs-3955	347	20	substitutions	substitution	NOUN
gs-3955	347	21	that	that	PRON
gs-3955	347	22	confer	confer	VERB
gs-3955	347	23	thermal	thermal	ADJ
gs-3955	347	24	and	and	CCONJ
gs-3955	347	25	oxidative	oxidative	ADJ
gs-3955	347	26	robustness	robustness	NOUN
gs-3955	347	27	(	(	PUNCT
gs-3955	347	28	munro	munro	PROPN
gs-3955	347	29	et	et	PROPN
gs-3955	347	30	al	al	PROPN
gs-3955	347	31	.	.	PROPN
gs-3955	347	32	,	,	PUNCT
gs-3955	347	33	2012	2012	NUM
gs-3955	347	34	)	)	PUNCT
gs-3955	347	35	.	.	PUNCT
gs-3955	348	1	concurrently	concurrently	ADV
gs-3955	348	2	,	,	PUNCT
gs-3955	348	3	autophagy	autophagy	ADJ
gs-3955	348	4	pathways	pathway	NOUN
gs-3955	348	5	exhibit	exhibit	VERB
gs-3955	348	6	heightened	heighten	VERB
gs-3955	348	7	selectivity	selectivity	NOUN
gs-3955	348	8	and	and	CCONJ
gs-3955	348	9	efficiency	efficiency	NOUN
gs-3955	348	10	,	,	PUNCT
gs-3955	348	11	with	with	SCONJ
gs-3955	348	12	aggrephagy	aggrephagy	ADJ
gs-3955	348	13	and	and	CCONJ
gs-3955	348	14	mitophagy	mitophagy	NOUN
gs-3955	348	15	variants	variant	NOUN
gs-3955	348	16	specialized	specialize	VERB
gs-3955	348	17	to	to	ADP
gs-3955	348	18	their	their	PRON
gs-3955	348	19	unique	unique	ADJ
gs-3955	348	20	cellular	cellular	ADJ
gs-3955	348	21	milieus	milieus	NOUN
gs-3955	348	22	(	(	PUNCT
gs-3955	348	23	pirotte	pirotte	NOUN
gs-3955	348	24	et	et	PROPN
gs-3955	348	25	al	al	PROPN
gs-3955	348	26	.	.	PROPN
gs-3955	348	27	,	,	PUNCT
gs-3955	348	28	2015	2015	NUM
gs-3955	348	29	)	)	PUNCT
gs-3955	348	30	.	.	PUNCT
gs-3955	349	1	we	we	PRON
gs-3955	349	2	hypothesize	hypothesize	VERB
gs-3955	349	3	that	that	SCONJ
gs-3955	349	4	this	this	DET
gs-3955	349	5	“	"	PUNCT
gs-3955	349	6	proteostasis	proteostasis	NOUN
gs-3955	349	7	supernetwork	supernetwork	NOUN
gs-3955	349	8	”	"	PUNCT
gs-3955	349	9	functions	function	NOUN
gs-3955	349	10	as	as	ADP
gs-3955	349	11	a	a	DET
gs-3955	349	12	dynamic	dynamic	ADJ
gs-3955	349	13	buffer	buffer	NOUN
gs-3955	349	14	against	against	ADP
gs-3955	349	15	stochastic	stochastic	ADJ
gs-3955	349	16	molecular	molecular	ADJ
gs-3955	349	17	insults	insult	NOUN
gs-3955	349	18	,	,	PUNCT
gs-3955	349	19	utilizing	utilize	VERB
gs-3955	349	20	phase	phase	NOUN
gs-3955	349	21	separation	separation	NOUN
gs-3955	349	22	and	and	CCONJ
gs-3955	349	23	liquid	liquid	NOUN
gs-3955	349	24	-	-	PUNCT
gs-3955	349	25	liquid	liquid	NOUN
gs-3955	349	26	demixing	demixing	NOUN
gs-3955	349	27	to	to	PART
gs-3955	349	28	compartmentalize	compartmentalize	VERB
gs-3955	349	29	and	and	CCONJ
gs-3955	349	30	neutralize	neutralize	VERB
gs-3955	349	31	misfolded	misfolde	VERB
gs-3955	349	32	proteins	protein	NOUN
gs-3955	349	33	before	before	SCONJ
gs-3955	349	34	they	they	PRON
gs-3955	349	35	seed	seed	VERB
gs-3955	349	36	aggregation	aggregation	NOUN
gs-3955	349	37	.	.	PUNCT
gs-3955	350	1	such	such	DET
gs-3955	350	2	a	a	DET
gs-3955	350	3	network	network	NOUN
gs-3955	350	4	is	be	AUX
gs-3955	350	5	likely	likely	ADV
gs-3955	350	6	regulated	regulate	VERB
gs-3955	350	7	by	by	ADP
gs-3955	350	8	feedback	feedback	NOUN
gs-3955	350	9	loops	loop	NOUN
gs-3955	350	10	involving	involve	VERB
gs-3955	350	11	stress	stress	NOUN
gs-3955	350	12	-	-	PUNCT
gs-3955	350	13	sensing	sense	VERB
gs-3955	350	14	transcription	transcription	NOUN
gs-3955	350	15	factors	factor	NOUN
gs-3955	350	16	like	like	ADP
gs-3955	350	17	foxo	foxo	NOUN
gs-3955	350	18	and	and	CCONJ
gs-3955	350	19	sirt6	sirt6	NOUN
gs-3955	350	20	,	,	PUNCT
gs-3955	350	21	enabling	enable	VERB
gs-3955	350	22	rapid	rapid	ADJ
gs-3955	350	23	adaptation	adaptation	NOUN
gs-3955	350	24	to	to	ADP
gs-3955	350	25	proteotoxic	proteotoxic	ADJ
gs-3955	350	26	challenges	challenge	NOUN
gs-3955	350	27	(	(	PUNCT
gs-3955	350	28	macrae	macrae	NOUN
gs-3955	350	29	et	et	PROPN
gs-3955	350	30	al	al	PROPN
gs-3955	350	31	.	.	PROPN
gs-3955	350	32	,	,	PUNCT
gs-3955	350	33	2015	2015	NUM
gs-3955	350	34	)	)	PUNCT
gs-3955	350	35	.	.	PUNCT
gs-3955	351	1	systems	system	NOUN
gs-3955	351	2	integration	integration	NOUN
gs-3955	351	3	through	through	ADP
gs-3955	351	4	neuroendocrine	neuroendocrine	ADJ
gs-3955	351	5	and	and	CCONJ
gs-3955	351	6	immune	immune	ADJ
gs-3955	351	7	synchronization	synchronization	NOUN
gs-3955	351	8	finally	finally	ADV
gs-3955	351	9	,	,	PUNCT
gs-3955	351	10	we	we	PRON
gs-3955	351	11	propose	propose	VERB
gs-3955	351	12	that	that	SCONJ
gs-3955	351	13	longevity	longevity	NOUN
gs-3955	351	14	in	in	ADP
gs-3955	351	15	negligibly	negligibly	ADV
gs-3955	351	16	senescent	senescent	ADJ
gs-3955	351	17	species	specie	NOUN
gs-3955	351	18	emerges	emerge	VERB
gs-3955	351	19	from	from	ADP
gs-3955	351	20	systemic	systemic	ADJ
gs-3955	351	21	integration	integration	NOUN
gs-3955	351	22	of	of	ADP
gs-3955	351	23	cellular	cellular	ADJ
gs-3955	351	24	repair	repair	NOUN
gs-3955	351	25	mechanisms	mechanism	NOUN
gs-3955	351	26	with	with	ADP
gs-3955	351	27	neuroendocrine	neuroendocrine	NOUN
gs-3955	351	28	and	and	CCONJ
gs-3955	351	29	immune	immune	ADJ
gs-3955	351	30	regulation	regulation	NOUN
gs-3955	351	31	.	.	PUNCT
gs-3955	352	1	naked	naked	ADJ
gs-3955	352	2	mole	mole	NOUN
gs-3955	352	3	-	-	PUNCT
gs-3955	352	4	rats	rat	NOUN
gs-3955	352	5	exemplify	exemplify	VERB
gs-3955	352	6	this	this	PRON
gs-3955	352	7	through	through	ADP
gs-3955	352	8	their	their	PRON
gs-3955	352	9	unique	unique	ADJ
gs-3955	352	10	glucocorticoid	glucocorticoid	ADJ
gs-3955	352	11	receptor	receptor	NOUN
gs-3955	352	12	adaptations	adaptation	NOUN
gs-3955	352	13	,	,	PUNCT
gs-3955	352	14	maintaining	maintain	VERB
gs-3955	352	15	stress	stress	ADJ
gs-3955	352	16	resilience	resilience	NOUN
gs-3955	352	17	without	without	ADP
gs-3955	352	18	immunosuppression	immunosuppression	NOUN
gs-3955	352	19	(	(	PUNCT
gs-3955	352	20	narayan	narayan	PROPN
gs-3955	352	21	et	et	PROPN
gs-3955	352	22	al	al	PROPN
gs-3955	352	23	.	.	PROPN
gs-3955	352	24	,	,	PUNCT
gs-3955	352	25	2021	2021	NUM
gs-3955	352	26	)	)	PUNCT
gs-3955	352	27	.	.	PUNCT
gs-3955	353	1	similarly	similarly	ADV
gs-3955	353	2	,	,	PUNCT
gs-3955	353	3	expanded	expand	VERB
gs-3955	353	4	innate	innate	ADJ
gs-3955	353	5	immune	immune	ADJ
gs-3955	353	6	repertoires	repertoire	NOUN
gs-3955	353	7	in	in	ADP
gs-3955	353	8	sharks	shark	NOUN
gs-3955	353	9	and	and	CCONJ
gs-3955	353	10	turtles	turtle	NOUN
gs-3955	353	11	confer	confer	VERB
gs-3955	353	12	persistent	persistent	ADJ
gs-3955	353	13	pathogen	pathogen	NOUN
gs-3955	353	14	georgian	georgian	PROPN
gs-3955	353	15	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	353	16	მეცნიერები	მეცნიერები	NOUN
gs-3955	353	17	ტ.	ტ.	VERB
gs-3955	353	18	7	7	NUM
gs-3955	353	19	n	n	PRON
gs-3955	353	20	3	3	NUM
gs-3955	353	21	,	,	PUNCT
gs-3955	353	22	2025	2025	NUM
gs-3955	353	23	385	385	NUM
gs-3955	353	24	defense	defense	NOUN
gs-3955	353	25	while	while	SCONJ
gs-3955	353	26	preventing	prevent	VERB
gs-3955	353	27	chronic	chronic	ADJ
gs-3955	353	28	inflammation	inflammation	NOUN
gs-3955	353	29	(	(	PUNCT
gs-3955	353	30	petruseva	petruseva	PROPN
gs-3955	353	31	et	et	PROPN
gs-3955	353	32	al	al	PROPN
gs-3955	353	33	.	.	PROPN
gs-3955	353	34	,	,	PUNCT
gs-3955	353	35	2017	2017	NUM
gs-3955	353	36	)	)	PUNCT
gs-3955	353	37	.	.	PUNCT
gs-3955	354	1	we	we	PRON
gs-3955	354	2	speculate	speculate	VERB
gs-3955	354	3	that	that	SCONJ
gs-3955	354	4	such	such	ADJ
gs-3955	354	5	species	specie	NOUN
gs-3955	354	6	maintain	maintain	VERB
gs-3955	354	7	a	a	DET
gs-3955	354	8	“	"	PUNCT
gs-3955	354	9	homeostatic	homeostatic	ADJ
gs-3955	354	10	harmony	harmony	NOUN
gs-3955	354	11	”	"	PUNCT
gs-3955	354	12	through	through	ADP
gs-3955	354	13	bidirectional	bidirectional	ADJ
gs-3955	354	14	signaling	signal	VERB
gs-3955	354	15	between	between	ADP
gs-3955	354	16	immune	immune	ADJ
gs-3955	354	17	effectors	effector	NOUN
gs-3955	354	18	,	,	PUNCT
gs-3955	354	19	hormonal	hormonal	ADJ
gs-3955	354	20	axes	axis	NOUN
gs-3955	354	21	,	,	PUNCT
gs-3955	354	22	and	and	CCONJ
gs-3955	354	23	metabolic	metabolic	NOUN
gs-3955	354	24	sensors	sensor	NOUN
gs-3955	354	25	.	.	PUNCT
gs-3955	355	1	this	this	DET
gs-3955	355	2	network	network	NOUN
gs-3955	355	3	preserves	preserve	VERB
gs-3955	355	4	tissue	tissue	NOUN
gs-3955	355	5	integrity	integrity	NOUN
gs-3955	355	6	,	,	PUNCT
gs-3955	355	7	prevents	prevent	VERB
gs-3955	355	8	inflammaging	inflammage	VERB
gs-3955	355	9	,	,	PUNCT
gs-3955	355	10	and	and	CCONJ
gs-3955	355	11	modulates	modulate	VERB
gs-3955	355	12	regenerative	regenerative	ADJ
gs-3955	355	13	processes	process	NOUN
gs-3955	355	14	,	,	PUNCT
gs-3955	355	15	ultimately	ultimately	ADV
gs-3955	355	16	stabilizing	stabilize	VERB
gs-3955	355	17	organismal	organismal	ADJ
gs-3955	355	18	function	function	NOUN
gs-3955	355	19	across	across	ADP
gs-3955	355	20	centuries	century	NOUN
gs-3955	355	21	.	.	PUNCT
gs-3955	356	1	in	in	ADP
gs-3955	356	2	sum	sum	NOUN
gs-3955	356	3	,	,	PUNCT
gs-3955	356	4	negligible	negligible	ADJ
gs-3955	356	5	senescence	senescence	NOUN
gs-3955	356	6	may	may	AUX
gs-3955	356	7	be	be	AUX
gs-3955	356	8	best	well	ADV
gs-3955	356	9	understood	understand	VERB
gs-3955	356	10	not	not	PART
gs-3955	356	11	as	as	ADP
gs-3955	356	12	a	a	DET
gs-3955	356	13	single	single	ADJ
gs-3955	356	14	pathway	pathway	NOUN
gs-3955	356	15	anomaly	anomaly	NOUN
gs-3955	356	16	but	but	CCONJ
gs-3955	356	17	as	as	ADP
gs-3955	356	18	a	a	DET
gs-3955	356	19	grand	grand	ADJ
gs-3955	356	20	evolutionary	evolutionary	ADJ
gs-3955	356	21	symphony	symphony	NOUN
gs-3955	356	22	:	:	PUNCT
gs-3955	356	23	an	an	DET
gs-3955	356	24	orchestrated	orchestrated	ADJ
gs-3955	356	25	balancing	balancing	NOUN
gs-3955	356	26	act	act	NOUN
gs-3955	356	27	managing	manage	VERB
gs-3955	356	28	entropy	entropy	NOUN
gs-3955	356	29	,	,	PUNCT
gs-3955	356	30	metabolism	metabolism	NOUN
gs-3955	356	31	,	,	PUNCT
gs-3955	356	32	proteostasis	proteostasis	NOUN
gs-3955	356	33	,	,	PUNCT
gs-3955	356	34	and	and	CCONJ
gs-3955	356	35	systemic	systemic	ADJ
gs-3955	356	36	homeostasis	homeostasis	NOUN
gs-3955	356	37	.	.	PUNCT
gs-3955	357	1	this	this	DET
gs-3955	357	2	hypothesis	hypothesis	NOUN
gs-3955	357	3	offers	offer	VERB
gs-3955	357	4	a	a	DET
gs-3955	357	5	transformative	transformative	ADJ
gs-3955	357	6	lens	len	NOUN
gs-3955	357	7	,	,	PUNCT
gs-3955	357	8	challenging	challenge	VERB
gs-3955	357	9	the	the	DET
gs-3955	357	10	deterministic	deterministic	ADJ
gs-3955	357	11	view	view	NOUN
gs-3955	357	12	of	of	ADP
gs-3955	357	13	aging	age	VERB
gs-3955	357	14	as	as	ADP
gs-3955	357	15	inevitable	inevitable	ADJ
gs-3955	357	16	,	,	PUNCT
gs-3955	357	17	and	and	CCONJ
gs-3955	357	18	instead	instead	ADV
gs-3955	357	19	framing	frame	VERB
gs-3955	357	20	longevity	longevity	NOUN
gs-3955	357	21	as	as	ADP
gs-3955	357	22	an	an	DET
gs-3955	357	23	attainable	attainable	ADJ
gs-3955	357	24	biological	biological	ADJ
gs-3955	357	25	state	state	NOUN
gs-3955	357	26	encoded	encode	VERB
gs-3955	357	27	in	in	ADP
gs-3955	357	28	the	the	DET
gs-3955	357	29	integrated	integrate	VERB
gs-3955	357	30	architecture	architecture	NOUN
gs-3955	357	31	of	of	ADP
gs-3955	357	32	cellular	cellular	ADJ
gs-3955	357	33	and	and	CCONJ
gs-3955	357	34	organismal	organismal	ADJ
gs-3955	357	35	networks	network	NOUN
gs-3955	357	36	.	.	PUNCT
gs-3955	358	1	the	the	DET
gs-3955	358	2	implications	implication	NOUN
gs-3955	358	3	for	for	ADP
gs-3955	358	4	human	human	ADJ
gs-3955	358	5	healthspan	healthspan	PROPN
gs-3955	358	6	extension	extension	NOUN
gs-3955	358	7	are	be	AUX
gs-3955	358	8	profound	profound	ADJ
gs-3955	358	9	.	.	PUNCT
gs-3955	359	1	by	by	ADP
gs-3955	359	2	decoding	decode	VERB
gs-3955	359	3	and	and	CCONJ
gs-3955	359	4	synthetically	synthetically	ADV
gs-3955	359	5	reconstituting	reconstitute	VERB
gs-3955	359	6	these	these	DET
gs-3955	359	7	integrated	integrate	VERB
gs-3955	359	8	systems	system	NOUN
gs-3955	359	9	,	,	PUNCT
gs-3955	359	10	we	we	PRON
gs-3955	359	11	may	may	AUX
gs-3955	359	12	transcend	transcend	VERB
gs-3955	359	13	incremental	incremental	ADJ
gs-3955	359	14	interventions	intervention	NOUN
gs-3955	359	15	and	and	CCONJ
gs-3955	359	16	unlock	unlock	VERB
gs-3955	359	17	the	the	DET
gs-3955	359	18	true	true	ADJ
gs-3955	359	19	biological	biological	ADJ
gs-3955	359	20	potential	potential	NOUN
gs-3955	359	21	for	for	ADP
gs-3955	359	22	sustained	sustained	ADJ
gs-3955	359	23	vitality	vitality	NOUN
gs-3955	359	24	.	.	PUNCT
gs-3955	360	1	translating	translate	VERB
gs-3955	360	2	negligible	negligible	ADJ
gs-3955	360	3	senescence	senescence	NOUN
gs-3955	360	4	mechanisms	mechanism	NOUN
gs-3955	360	5	to	to	ADP
gs-3955	360	6	humans	human	NOUN
gs-3955	360	7	:	:	PUNCT
gs-3955	360	8	opportunities	opportunity	NOUN
gs-3955	360	9	and	and	CCONJ
gs-3955	360	10	challenges	challenge	VERB
gs-3955	360	11	many	many	ADJ
gs-3955	360	12	of	of	ADP
gs-3955	360	13	the	the	DET
gs-3955	360	14	remarkable	remarkable	ADJ
gs-3955	360	15	longevity	longevity	NOUN
gs-3955	360	16	mechanisms	mechanism	NOUN
gs-3955	360	17	found	find	VERB
gs-3955	360	18	in	in	ADP
gs-3955	360	19	negligibly	negligibly	ADV
gs-3955	360	20	senescent	senescent	ADJ
gs-3955	360	21	species	specie	NOUN
gs-3955	360	22	share	share	VERB
gs-3955	360	23	fundamental	fundamental	ADJ
gs-3955	360	24	molecular	molecular	ADJ
gs-3955	360	25	components	component	NOUN
gs-3955	360	26	with	with	ADP
gs-3955	360	27	humans	human	NOUN
gs-3955	360	28	,	,	PUNCT
gs-3955	360	29	but	but	CCONJ
gs-3955	360	30	critical	critical	ADJ
gs-3955	360	31	differences	difference	NOUN
gs-3955	360	32	explain	explain	VERB
gs-3955	360	33	why	why	SCONJ
gs-3955	360	34	humans	human	NOUN
gs-3955	360	35	age	age	VERB
gs-3955	360	36	relatively	relatively	ADV
gs-3955	360	37	rapidly	rapidly	ADV
gs-3955	360	38	and	and	CCONJ
gs-3955	360	39	experience	experience	VERB
gs-3955	360	40	age	age	NOUN
gs-3955	360	41	-	-	PUNCT
gs-3955	360	42	related	relate	VERB
gs-3955	360	43	diseases	disease	NOUN
gs-3955	360	44	.	.	PUNCT
gs-3955	361	1	understanding	understand	VERB
gs-3955	361	2	these	these	DET
gs-3955	361	3	similarities	similarity	NOUN
gs-3955	361	4	and	and	CCONJ
gs-3955	361	5	gaps	gap	NOUN
gs-3955	361	6	points	point	NOUN
gs-3955	361	7	to	to	ADP
gs-3955	361	8	promising	promise	VERB
gs-3955	361	9	therapeutic	therapeutic	ADJ
gs-3955	361	10	strategies	strategy	NOUN
gs-3955	361	11	—	—	PUNCT
gs-3955	361	12	ranging	range	VERB
gs-3955	361	13	from	from	ADP
gs-3955	361	14	gene	gene	NOUN
gs-3955	361	15	therapy	therapy	NOUN
gs-3955	361	16	to	to	ADP
gs-3955	361	17	pharmacology	pharmacology	NOUN
gs-3955	361	18	—	—	PUNCT
gs-3955	361	19	that	that	PRON
gs-3955	361	20	could	could	AUX
gs-3955	361	21	one	one	NUM
gs-3955	361	22	day	day	NOUN
gs-3955	361	23	extend	extend	VERB
gs-3955	361	24	human	human	ADJ
gs-3955	361	25	healthspan	healthspan	NOUN
gs-3955	361	26	and	and	CCONJ
gs-3955	361	27	lifespan	lifespan	PROPN
gs-3955	361	28	.	.	PUNCT
gs-3955	362	1	shared	share	VERB
gs-3955	362	2	molecular	molecular	ADJ
gs-3955	362	3	mechanisms	mechanism	NOUN
gs-3955	362	4	●	●	PUNCT
gs-3955	362	5	dna	dna	PROPN
gs-3955	362	6	repair	repair	NOUN
gs-3955	362	7	pathways	pathway	NOUN
gs-3955	362	8	:	:	PUNCT
gs-3955	362	9	humans	human	NOUN
gs-3955	362	10	possess	possess	VERB
gs-3955	362	11	dna	dna	PROPN
gs-3955	362	12	repair	repair	NOUN
gs-3955	362	13	systems	system	NOUN
gs-3955	362	14	like	like	ADP
gs-3955	362	15	nucleotide	nucleotide	NOUN
gs-3955	362	16	excision	excision	NOUN
gs-3955	362	17	repair	repair	NOUN
gs-3955	362	18	(	(	PUNCT
gs-3955	362	19	ner	ner	NOUN
gs-3955	362	20	)	)	PUNCT
gs-3955	362	21	,	,	PUNCT
gs-3955	362	22	homologous	homologous	ADJ
gs-3955	362	23	recombination	recombination	NOUN
gs-3955	362	24	(	(	PUNCT
gs-3955	362	25	hr	hr	NOUN
gs-3955	362	26	)	)	PUNCT
gs-3955	362	27	,	,	PUNCT
gs-3955	362	28	and	and	CCONJ
gs-3955	362	29	base	base	NOUN
gs-3955	362	30	excision	excision	NOUN
gs-3955	362	31	repair	repair	NOUN
gs-3955	362	32	(	(	PUNCT
gs-3955	362	33	ber	ber	PROPN
gs-3955	362	34	)	)	PUNCT
gs-3955	362	35	,	,	PUNCT
gs-3955	362	36	similar	similar	ADJ
gs-3955	362	37	to	to	ADP
gs-3955	362	38	naked	naked	ADJ
gs-3955	362	39	mole	mole	NOUN
gs-3955	362	40	-	-	PUNCT
gs-3955	362	41	rats	rat	NOUN
gs-3955	362	42	and	and	CCONJ
gs-3955	362	43	greenland	greenland	PROPN
gs-3955	362	44	sharks	sharks	PROPN
gs-3955	362	45	(	(	PUNCT
gs-3955	362	46	narayan	narayan	PROPN
gs-3955	362	47	et	et	PROPN
gs-3955	362	48	al	al	PROPN
gs-3955	362	49	.	.	PROPN
gs-3955	362	50	,	,	PUNCT
gs-3955	362	51	2021	2021	NUM
gs-3955	362	52	;	;	PUNCT
gs-3955	362	53	petruseva	petruseva	VERB
gs-3955	362	54	et	et	PROPN
gs-3955	362	55	al	al	PROPN
gs-3955	362	56	.	.	PROPN
gs-3955	362	57	,	,	PUNCT
gs-3955	362	58	2017	2017	NUM
gs-3955	362	59	)	)	PUNCT
gs-3955	362	60	.	.	PUNCT
gs-3955	363	1	however	however	ADV
gs-3955	363	2	,	,	PUNCT
gs-3955	363	3	the	the	DET
gs-3955	363	4	efficiency	efficiency	NOUN
gs-3955	363	5	of	of	ADP
gs-3955	363	6	these	these	DET
gs-3955	363	7	pathways	pathway	NOUN
gs-3955	363	8	declines	decline	NOUN
gs-3955	363	9	with	with	ADP
gs-3955	363	10	age	age	NOUN
gs-3955	363	11	in	in	ADP
gs-3955	363	12	humans	human	NOUN
gs-3955	363	13	,	,	PUNCT
gs-3955	363	14	whereas	whereas	SCONJ
gs-3955	363	15	long	long	ADV
gs-3955	363	16	-	-	PUNCT
gs-3955	363	17	lived	live	VERB
gs-3955	363	18	species	specie	NOUN
gs-3955	363	19	maintain	maintain	VERB
gs-3955	363	20	or	or	CCONJ
gs-3955	363	21	enhance	enhance	VERB
gs-3955	363	22	these	these	DET
gs-3955	363	23	mechanisms	mechanism	NOUN
gs-3955	363	24	throughout	throughout	ADP
gs-3955	363	25	life	life	NOUN
gs-3955	363	26	(	(	PUNCT
gs-3955	363	27	macrae	macrae	PROPN
gs-3955	363	28	et	et	PROPN
gs-3955	363	29	al	al	PROPN
gs-3955	363	30	.	.	PROPN
gs-3955	363	31	,	,	PUNCT
gs-3955	363	32	2015	2015	NUM
gs-3955	363	33	)	)	PUNCT
gs-3955	363	34	.	.	PUNCT
gs-3955	364	1	key	key	ADJ
gs-3955	364	2	regulators	regulator	NOUN
gs-3955	364	3	like	like	ADP
gs-3955	364	4	sirt6	sirt6	NOUN
gs-3955	364	5	and	and	CCONJ
gs-3955	364	6	parp1	parp1	NOUN
gs-3955	364	7	are	be	AUX
gs-3955	364	8	conserved	conserve	VERB
gs-3955	364	9	,	,	PUNCT
gs-3955	364	10	offering	offer	VERB
gs-3955	364	11	potential	potential	ADJ
gs-3955	364	12	targets	target	NOUN
gs-3955	364	13	for	for	ADP
gs-3955	364	14	boosting	boost	VERB
gs-3955	364	15	human	human	ADJ
gs-3955	364	16	dna	dna	PROPN
gs-3955	364	17	repair	repair	NOUN
gs-3955	364	18	capacity	capacity	NOUN
gs-3955	364	19	(	(	PUNCT
gs-3955	364	20	lewis	lewis	PROPN
gs-3955	364	21	et	et	PROPN
gs-3955	364	22	al	al	PROPN
gs-3955	364	23	.	.	PROPN
gs-3955	364	24	,	,	PUNCT
gs-3955	364	25	2012	2012	NUM
gs-3955	364	26	;	;	PUNCT
gs-3955	364	27	narayan	narayan	PROPN
gs-3955	364	28	et	et	PROPN
gs-3955	364	29	al	al	PROPN
gs-3955	364	30	.	.	PROPN
gs-3955	364	31	,	,	PUNCT
gs-3955	364	32	2021	2021	NUM
gs-3955	364	33	)	)	PUNCT
gs-3955	364	34	.	.	PUNCT
gs-3955	365	1	●	●	NUM
gs-3955	365	2	stem	stem	NOUN
gs-3955	365	3	cell	cell	NOUN
gs-3955	365	4	maintenance	maintenance	NOUN
gs-3955	365	5	:	:	PUNCT
gs-3955	365	6	humans	human	NOUN
gs-3955	365	7	have	have	VERB
gs-3955	365	8	adult	adult	NOUN
gs-3955	365	9	stem	stem	NOUN
gs-3955	365	10	cells	cell	NOUN
gs-3955	365	11	essential	essential	ADJ
gs-3955	365	12	for	for	ADP
gs-3955	365	13	tissue	tissue	NOUN
gs-3955	365	14	repair	repair	NOUN
gs-3955	365	15	and	and	CCONJ
gs-3955	365	16	regeneration	regeneration	NOUN
gs-3955	365	17	,	,	PUNCT
gs-3955	365	18	much	much	ADV
gs-3955	365	19	like	like	ADP
gs-3955	365	20	the	the	DET
gs-3955	365	21	stem	stem	NOUN
gs-3955	365	22	cell	cell	NOUN
gs-3955	365	23	pools	pool	NOUN
gs-3955	365	24	seen	see	VERB
gs-3955	365	25	in	in	ADP
gs-3955	365	26	hydra	hydra	PROPN
gs-3955	365	27	and	and	CCONJ
gs-3955	365	28	planarians	planarian	NOUN
gs-3955	365	29	(	(	PUNCT
gs-3955	365	30	wagner	wagner	PROPN
gs-3955	365	31	et	et	PROPN
gs-3955	365	32	al	al	PROPN
gs-3955	365	33	.	.	PROPN
gs-3955	365	34	,	,	PUNCT
gs-3955	365	35	2011	2011	NUM
gs-3955	365	36	;	;	PUNCT
gs-3955	365	37	rink	rink	NOUN
gs-3955	365	38	,	,	PUNCT
gs-3955	365	39	2013	2013	NUM
gs-3955	365	40	)	)	PUNCT
gs-3955	365	41	.	.	PUNCT
gs-3955	366	1	yet	yet	ADV
gs-3955	366	2	,	,	PUNCT
gs-3955	366	3	human	human	ADJ
gs-3955	366	4	stem	stem	NOUN
gs-3955	366	5	cells	cell	NOUN
gs-3955	366	6	show	show	VERB
gs-3955	366	7	exhaustion	exhaustion	NOUN
gs-3955	366	8	,	,	PUNCT
gs-3955	366	9	telomere	telomere	NOUN
gs-3955	366	10	shortening	shortening	NOUN
gs-3955	366	11	,	,	PUNCT
gs-3955	366	12	and	and	CCONJ
gs-3955	366	13	reduced	reduce	VERB
gs-3955	366	14	regenerative	regenerative	ADJ
gs-3955	366	15	potential	potential	NOUN
gs-3955	366	16	with	with	ADP
gs-3955	366	17	age	age	NOUN
gs-3955	366	18	(	(	PUNCT
gs-3955	366	19	petruseva	petruseva	PROPN
gs-3955	366	20	et	et	PROPN
gs-3955	366	21	al	al	PROPN
gs-3955	366	22	.	.	PROPN
gs-3955	366	23	,	,	PUNCT
gs-3955	366	24	2017	2017	NUM
gs-3955	366	25	)	)	PUNCT
gs-3955	366	26	.	.	PUNCT
gs-3955	367	1	the	the	DET
gs-3955	367	2	constitutive	constitutive	ADJ
gs-3955	367	3	telomerase	telomerase	NOUN
gs-3955	367	4	activity	activity	NOUN
gs-3955	367	5	and	and	CCONJ
gs-3955	367	6	symmetric	symmetric	ADJ
gs-3955	367	7	stem	stem	NOUN
gs-3955	367	8	cell	cell	NOUN
gs-3955	367	9	division	division	NOUN
gs-3955	367	10	observed	observe	VERB
gs-3955	367	11	in	in	ADP
gs-3955	367	12	these	these	DET
gs-3955	367	13	species	specie	NOUN
gs-3955	367	14	highlight	highlight	VERB
gs-3955	367	15	molecular	molecular	ADJ
gs-3955	367	16	pathways	pathway	NOUN
gs-3955	367	17	—	—	PUNCT
gs-3955	367	18	such	such	ADJ
gs-3955	367	19	as	as	ADP
gs-3955	367	20	foxo	foxo	NOUN
gs-3955	367	21	signaling	signal	VERB
gs-3955	367	22	—	—	PUNCT
gs-3955	367	23	that	that	PRON
gs-3955	367	24	are	be	AUX
gs-3955	367	25	present	present	ADJ
gs-3955	367	26	but	but	CCONJ
gs-3955	367	27	less	less	ADV
gs-3955	367	28	active	active	ADJ
gs-3955	367	29	or	or	CCONJ
gs-3955	367	30	tightly	tightly	ADV
gs-3955	367	31	regulated	regulate	VERB
gs-3955	367	32	in	in	ADP
gs-3955	367	33	humans	human	NOUN
gs-3955	367	34	(	(	PUNCT
gs-3955	367	35	tkemaladze	tkemaladze	NOUN
gs-3955	367	36	,	,	PUNCT
gs-3955	367	37	2023	2023	NUM
gs-3955	367	38	;	;	PUNCT
gs-3955	367	39	wagner	wagner	PROPN
gs-3955	367	40	et	et	PROPN
gs-3955	367	41	al	al	PROPN
gs-3955	367	42	.	.	PROPN
gs-3955	367	43	,	,	PUNCT
gs-3955	367	44	2011	2011	NUM
gs-3955	367	45	)	)	PUNCT
gs-3955	367	46	.	.	PUNCT
gs-3955	368	1	georgian	georgian	ADJ
gs-3955	368	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	368	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	368	4	ტ.	ტ.	VERB
gs-3955	368	5	7	7	NUM
gs-3955	368	6	n	n	DET
gs-3955	368	7	3	3	NUM
gs-3955	368	8	,	,	PUNCT
gs-3955	368	9	2025	2025	NUM
gs-3955	368	10	386	386	NUM
gs-3955	368	11	●	●	NUM
gs-3955	368	12	proteostasis	proteostasis	NOUN
gs-3955	368	13	:	:	PUNCT
gs-3955	368	14	humans	human	NOUN
gs-3955	368	15	share	share	VERB
gs-3955	368	16	chaperone	chaperone	NOUN
gs-3955	368	17	proteins	protein	NOUN
gs-3955	368	18	,	,	PUNCT
gs-3955	368	19	ubiquitin	ubiquitin	ADJ
gs-3955	368	20	-	-	PUNCT
gs-3955	368	21	proteasome	proteasome	NOUN
gs-3955	368	22	systems	system	NOUN
gs-3955	368	23	,	,	PUNCT
gs-3955	368	24	and	and	CCONJ
gs-3955	368	25	autophagy	autophagy	ADJ
gs-3955	368	26	pathways	pathway	NOUN
gs-3955	368	27	with	with	ADP
gs-3955	368	28	long	long	ADV
gs-3955	368	29	-	-	PUNCT
gs-3955	368	30	lived	live	VERB
gs-3955	368	31	organisms	organism	NOUN
gs-3955	368	32	,	,	PUNCT
gs-3955	368	33	but	but	CCONJ
gs-3955	368	34	these	these	DET
gs-3955	368	35	quality	quality	NOUN
gs-3955	368	36	control	control	NOUN
gs-3955	368	37	systems	system	NOUN
gs-3955	368	38	become	become	VERB
gs-3955	368	39	less	less	ADV
gs-3955	368	40	effective	effective	ADJ
gs-3955	368	41	over	over	ADP
gs-3955	368	42	time	time	NOUN
gs-3955	368	43	(	(	PUNCT
gs-3955	368	44	macrae	macrae	NOUN
gs-3955	368	45	et	et	PROPN
gs-3955	368	46	al	al	PROPN
gs-3955	368	47	.	.	PROPN
gs-3955	368	48	,	,	PUNCT
gs-3955	368	49	2015	2015	NUM
gs-3955	368	50	;	;	PUNCT
gs-3955	368	51	lewis	lewis	PROPN
gs-3955	368	52	et	et	PROPN
gs-3955	368	53	al	al	PROPN
gs-3955	368	54	.	.	PROPN
gs-3955	368	55	,	,	PUNCT
gs-3955	368	56	2012	2012	NUM
gs-3955	368	57	)	)	PUNCT
gs-3955	368	58	.	.	PUNCT
gs-3955	369	1	the	the	DET
gs-3955	369	2	advanced	advanced	ADJ
gs-3955	369	3	proteostasis	proteostasis	NOUN
gs-3955	369	4	networks	network	NOUN
gs-3955	369	5	in	in	ADP
gs-3955	369	6	long	long	ADV
gs-3955	369	7	-	-	PUNCT
gs-3955	369	8	lived	live	VERB
gs-3955	369	9	species	specie	NOUN
gs-3955	369	10	suggest	suggest	VERB
gs-3955	369	11	that	that	SCONJ
gs-3955	369	12	enhancing	enhance	VERB
gs-3955	369	13	molecular	molecular	ADJ
gs-3955	369	14	chaperones	chaperone	NOUN
gs-3955	369	15	,	,	PUNCT
gs-3955	369	16	autophagy	autophagy	ADJ
gs-3955	369	17	efficiency	efficiency	NOUN
gs-3955	369	18	,	,	PUNCT
gs-3955	369	19	and	and	CCONJ
gs-3955	369	20	selective	selective	ADJ
gs-3955	369	21	degradation	degradation	NOUN
gs-3955	369	22	processes	process	NOUN
gs-3955	369	23	could	could	AUX
gs-3955	369	24	preserve	preserve	VERB
gs-3955	369	25	protein	protein	NOUN
gs-3955	369	26	homeostasis	homeostasis	NOUN
gs-3955	369	27	in	in	ADP
gs-3955	369	28	aging	age	VERB
gs-3955	369	29	human	human	ADJ
gs-3955	369	30	cells	cell	NOUN
gs-3955	369	31	(	(	PUNCT
gs-3955	369	32	munro	munro	PROPN
gs-3955	369	33	et	et	PROPN
gs-3955	369	34	al	al	PROPN
gs-3955	369	35	.	.	PROPN
gs-3955	369	36	,	,	PUNCT
gs-3955	369	37	2012	2012	NUM
gs-3955	369	38	)	)	PUNCT
gs-3955	369	39	.	.	PUNCT
gs-3955	370	1	●	●	PUNCT
gs-3955	370	2	oxidative	oxidative	ADJ
gs-3955	370	3	stress	stress	NOUN
gs-3955	370	4	management	management	NOUN
gs-3955	370	5	:	:	PUNCT
gs-3955	370	6	humans	human	NOUN
gs-3955	370	7	produce	produce	VERB
gs-3955	370	8	reactive	reactive	ADJ
gs-3955	370	9	oxygen	oxygen	NOUN
gs-3955	370	10	species	specie	NOUN
gs-3955	370	11	(	(	PUNCT
gs-3955	370	12	ros	ros	PROPN
gs-3955	370	13	)	)	PUNCT
gs-3955	370	14	during	during	ADP
gs-3955	370	15	metabolism	metabolism	NOUN
gs-3955	370	16	and	and	CCONJ
gs-3955	370	17	have	have	VERB
gs-3955	370	18	antioxidant	antioxidant	ADJ
gs-3955	370	19	defenses	defense	NOUN
gs-3955	370	20	,	,	PUNCT
gs-3955	370	21	yet	yet	CCONJ
gs-3955	370	22	the	the	DET
gs-3955	370	23	balance	balance	NOUN
gs-3955	370	24	often	often	ADV
gs-3955	370	25	shifts	shift	NOUN
gs-3955	370	26	toward	toward	ADP
gs-3955	370	27	oxidative	oxidative	ADJ
gs-3955	370	28	damage	damage	NOUN
gs-3955	370	29	with	with	ADP
gs-3955	370	30	aging	age	VERB
gs-3955	370	31	(	(	PUNCT
gs-3955	370	32	perez	perez	PROPN
gs-3955	370	33	et	et	PROPN
gs-3955	370	34	al	al	PROPN
gs-3955	370	35	.	.	PROPN
gs-3955	370	36	,	,	PUNCT
gs-3955	370	37	2009	2009	NUM
gs-3955	370	38	)	)	PUNCT
gs-3955	370	39	.	.	PUNCT
gs-3955	371	1	long	long	ADV
gs-3955	371	2	-	-	PUNCT
gs-3955	371	3	lived	live	VERB
gs-3955	371	4	species	specie	NOUN
gs-3955	371	5	demonstrate	demonstrate	VERB
gs-3955	371	6	either	either	CCONJ
gs-3955	371	7	reduced	reduce	VERB
gs-3955	371	8	ros	ros	PROPN
gs-3955	371	9	production	production	NOUN
gs-3955	371	10	or	or	CCONJ
gs-3955	371	11	improved	improve	VERB
gs-3955	371	12	tolerance	tolerance	NOUN
gs-3955	371	13	to	to	PART
gs-3955	371	14	oxidative	oxidative	VERB
gs-3955	371	15	stress	stress	NOUN
gs-3955	371	16	through	through	ADP
gs-3955	371	17	membrane	membrane	NOUN
gs-3955	371	18	lipid	lipid	NOUN
gs-3955	371	19	composition	composition	NOUN
gs-3955	371	20	and	and	CCONJ
gs-3955	371	21	efficient	efficient	ADJ
gs-3955	371	22	redox	redox	ADJ
gs-3955	371	23	buffering	buffering	NOUN
gs-3955	371	24	systems	system	NOUN
gs-3955	371	25	,	,	PUNCT
gs-3955	371	26	mechanisms	mechanism	NOUN
gs-3955	371	27	that	that	PRON
gs-3955	371	28	could	could	AUX
gs-3955	371	29	inspire	inspire	VERB
gs-3955	371	30	antioxidant	antioxidant	ADJ
gs-3955	371	31	therapies	therapy	NOUN
gs-3955	371	32	or	or	CCONJ
gs-3955	371	33	membrane	membrane	NOUN
gs-3955	371	34	-	-	PUNCT
gs-3955	371	35	targeted	target	VERB
gs-3955	371	36	drugs	drug	NOUN
gs-3955	371	37	(	(	PUNCT
gs-3955	371	38	munro	munro	PROPN
gs-3955	371	39	et	et	PROPN
gs-3955	371	40	al	al	PROPN
gs-3955	371	41	.	.	PROPN
gs-3955	371	42	,	,	PUNCT
gs-3955	371	43	2012	2012	NUM
gs-3955	371	44	;	;	PUNCT
gs-3955	371	45	strahlendorf	strahlendorf	PROPN
gs-3955	371	46	et	et	PROPN
gs-3955	371	47	al	al	PROPN
gs-3955	371	48	.	.	PROPN
gs-3955	371	49	,	,	PUNCT
gs-3955	371	50	2008	2008	NUM
gs-3955	371	51	)	)	PUNCT
gs-3955	371	52	.	.	PUNCT
gs-3955	372	1	key	key	ADJ
gs-3955	372	2	differences	difference	NOUN
gs-3955	372	3	and	and	CCONJ
gs-3955	372	4	therapeutic	therapeutic	ADJ
gs-3955	372	5	implications	implication	NOUN
gs-3955	372	6	●	●	NUM
gs-3955	372	7	telomerase	telomerase	NOUN
gs-3955	372	8	regulation	regulation	NOUN
gs-3955	372	9	:	:	PUNCT
gs-3955	372	10	humans	human	NOUN
gs-3955	372	11	suppress	suppress	VERB
gs-3955	372	12	telomerase	telomerase	NOUN
gs-3955	372	13	in	in	ADP
gs-3955	372	14	most	most	ADJ
gs-3955	372	15	somatic	somatic	ADJ
gs-3955	372	16	cells	cell	NOUN
gs-3955	372	17	to	to	PART
gs-3955	372	18	prevent	prevent	VERB
gs-3955	372	19	cancer	cancer	NOUN
gs-3955	372	20	risk	risk	NOUN
gs-3955	372	21	,	,	PUNCT
gs-3955	372	22	while	while	SCONJ
gs-3955	372	23	naked	naked	ADJ
gs-3955	372	24	mole	mole	NOUN
gs-3955	372	25	-	-	PUNCT
gs-3955	372	26	rats	rat	NOUN
gs-3955	372	27	and	and	CCONJ
gs-3955	372	28	hydra	hydra	NOUN
gs-3955	372	29	maintain	maintain	VERB
gs-3955	372	30	active	active	ADJ
gs-3955	372	31	telomerase	telomerase	NOUN
gs-3955	372	32	to	to	PART
gs-3955	372	33	preserve	preserve	VERB
gs-3955	372	34	telomeres	telomere	NOUN
gs-3955	372	35	(	(	PUNCT
gs-3955	372	36	petruseva	petruseva	PROPN
gs-3955	372	37	et	et	PROPN
gs-3955	372	38	al	al	PROPN
gs-3955	372	39	.	.	PROPN
gs-3955	372	40	,	,	PUNCT
gs-3955	372	41	2017	2017	NUM
gs-3955	372	42	)	)	PUNCT
gs-3955	372	43	.	.	PUNCT
gs-3955	373	1	simply	simply	ADV
gs-3955	373	2	activating	activate	VERB
gs-3955	373	3	telomerase	telomerase	NOUN
gs-3955	373	4	in	in	ADP
gs-3955	373	5	humans	human	NOUN
gs-3955	373	6	poses	pose	VERB
gs-3955	373	7	oncogenic	oncogenic	ADJ
gs-3955	373	8	dangers	danger	NOUN
gs-3955	373	9	unless	unless	SCONJ
gs-3955	373	10	combined	combine	VERB
gs-3955	373	11	with	with	ADP
gs-3955	373	12	cancer	cancer	NOUN
gs-3955	373	13	resistance	resistance	NOUN
gs-3955	373	14	strategies	strategy	NOUN
gs-3955	373	15	,	,	PUNCT
gs-3955	373	16	such	such	ADJ
gs-3955	373	17	as	as	ADP
gs-3955	373	18	enhanced	enhance	VERB
gs-3955	373	19	p53	p53	NOUN
gs-3955	373	20	function	function	NOUN
gs-3955	373	21	or	or	CCONJ
gs-3955	373	22	unique	unique	ADJ
gs-3955	373	23	tumor	tumor	NOUN
gs-3955	373	24	suppressor	suppressor	NOUN
gs-3955	373	25	pathways	pathway	NOUN
gs-3955	373	26	found	find	VERB
gs-3955	373	27	in	in	ADP
gs-3955	373	28	these	these	DET
gs-3955	373	29	species	specie	NOUN
gs-3955	373	30	(	(	PUNCT
gs-3955	373	31	lewis	lewis	PROPN
gs-3955	373	32	et	et	PROPN
gs-3955	373	33	al	al	PROPN
gs-3955	373	34	.	.	PROPN
gs-3955	373	35	,	,	PUNCT
gs-3955	373	36	2012	2012	NUM
gs-3955	373	37	)	)	PUNCT
gs-3955	373	38	.	.	PUNCT
gs-3955	374	1	●	●	PUNCT
gs-3955	374	2	metabolic	metabolic	NOUN
gs-3955	374	3	adaptations	adaptation	NOUN
gs-3955	374	4	:	:	PUNCT
gs-3955	374	5	metabolic	metabolic	NOUN
gs-3955	374	6	depression	depression	NOUN
gs-3955	374	7	and	and	CCONJ
gs-3955	374	8	hypometabolism	hypometabolism	NOUN
gs-3955	374	9	seen	see	VERB
gs-3955	374	10	in	in	ADP
gs-3955	374	11	longlived	longlived	ADJ
gs-3955	374	12	animals	animal	NOUN
gs-3955	374	13	help	help	VERB
gs-3955	374	14	minimize	minimize	VERB
gs-3955	374	15	damage	damage	NOUN
gs-3955	374	16	but	but	CCONJ
gs-3955	374	17	are	be	AUX
gs-3955	374	18	incompatible	incompatible	ADJ
gs-3955	374	19	with	with	ADP
gs-3955	374	20	human	human	ADJ
gs-3955	374	21	brain	brain	NOUN
gs-3955	374	22	function	function	NOUN
gs-3955	374	23	and	and	CCONJ
gs-3955	374	24	energy	energy	NOUN
gs-3955	374	25	demands	demand	NOUN
gs-3955	374	26	(	(	PUNCT
gs-3955	374	27	narayan	narayan	NOUN
gs-3955	374	28	et	et	PROPN
gs-3955	374	29	al	al	PROPN
gs-3955	374	30	.	.	PROPN
gs-3955	374	31	,	,	PUNCT
gs-3955	374	32	2021	2021	NUM
gs-3955	374	33	)	)	PUNCT
gs-3955	374	34	.	.	PUNCT
gs-3955	375	1	therapeutic	therapeutic	ADJ
gs-3955	375	2	approaches	approach	NOUN
gs-3955	375	3	may	may	AUX
gs-3955	375	4	need	need	VERB
gs-3955	375	5	to	to	PART
gs-3955	375	6	mimic	mimic	VERB
gs-3955	375	7	selective	selective	ADJ
gs-3955	375	8	metabolic	metabolic	NOUN
gs-3955	375	9	pathways	pathway	NOUN
gs-3955	375	10	,	,	PUNCT
gs-3955	375	11	such	such	ADJ
gs-3955	375	12	as	as	ADP
gs-3955	375	13	ketone	ketone	NOUN
gs-3955	375	14	body	body	NOUN
gs-3955	375	15	utilization	utilization	NOUN
gs-3955	375	16	or	or	CCONJ
gs-3955	375	17	mitochondrial	mitochondrial	ADJ
gs-3955	375	18	uncoupling	uncoupling	NOUN
gs-3955	375	19	,	,	PUNCT
gs-3955	375	20	without	without	ADP
gs-3955	375	21	triggering	trigger	VERB
gs-3955	375	22	systemic	systemic	ADJ
gs-3955	375	23	energy	energy	NOUN
gs-3955	375	24	deficits	deficit	NOUN
gs-3955	375	25	(	(	PUNCT
gs-3955	375	26	munro	munro	PROPN
gs-3955	375	27	et	et	PROPN
gs-3955	375	28	al	al	PROPN
gs-3955	375	29	.	.	PROPN
gs-3955	375	30	,	,	PUNCT
gs-3955	375	31	2012	2012	NUM
gs-3955	375	32	)	)	PUNCT
gs-3955	375	33	.	.	PUNCT
gs-3955	376	1	●	●	NUM
gs-3955	376	2	stem	stem	NOUN
gs-3955	376	3	cell	cell	NOUN
gs-3955	376	4	plasticity	plasticity	NOUN
gs-3955	376	5	:	:	PUNCT
gs-3955	376	6	regeneration	regeneration	NOUN
gs-3955	376	7	capacity	capacity	NOUN
gs-3955	376	8	in	in	ADP
gs-3955	376	9	species	specie	NOUN
gs-3955	376	10	like	like	ADP
gs-3955	376	11	planarians	planarian	NOUN
gs-3955	376	12	far	far	ADV
gs-3955	376	13	exceeds	exceed	VERB
gs-3955	376	14	that	that	SCONJ
gs-3955	376	15	in	in	ADP
gs-3955	376	16	humans	human	NOUN
gs-3955	376	17	.	.	PUNCT
gs-3955	377	1	achieving	achieve	VERB
gs-3955	377	2	similar	similar	ADJ
gs-3955	377	3	plasticity	plasticity	NOUN
gs-3955	377	4	would	would	AUX
gs-3955	377	5	likely	likely	ADV
gs-3955	377	6	require	require	VERB
gs-3955	377	7	reprogramming	reprogramme	VERB
gs-3955	377	8	adult	adult	NOUN
gs-3955	377	9	human	human	ADJ
gs-3955	377	10	cells	cell	NOUN
gs-3955	377	11	to	to	ADP
gs-3955	377	12	a	a	DET
gs-3955	377	13	more	more	ADV
gs-3955	377	14	embryonic	embryonic	ADJ
gs-3955	377	15	-	-	PUNCT
gs-3955	377	16	like	like	ADJ
gs-3955	377	17	state	state	NOUN
gs-3955	377	18	and	and	CCONJ
gs-3955	377	19	enabling	enable	VERB
gs-3955	377	20	symmetric	symmetric	ADJ
gs-3955	377	21	stem	stem	NOUN
gs-3955	377	22	cell	cell	NOUN
gs-3955	377	23	divisions	division	NOUN
gs-3955	377	24	—	—	PUNCT
gs-3955	377	25	an	an	DET
gs-3955	377	26	area	area	NOUN
gs-3955	377	27	ripe	ripe	ADJ
gs-3955	377	28	for	for	ADP
gs-3955	377	29	crispr	crispr	ADJ
gs-3955	377	30	gene	gene	NOUN
gs-3955	377	31	editing	editing	NOUN
gs-3955	377	32	and	and	CCONJ
gs-3955	377	33	synthetic	synthetic	ADJ
gs-3955	377	34	biology	biology	NOUN
gs-3955	377	35	interventions	intervention	NOUN
gs-3955	377	36	(	(	PUNCT
gs-3955	377	37	wagner	wagner	PROPN
gs-3955	377	38	et	et	PROPN
gs-3955	377	39	al	al	PROPN
gs-3955	377	40	.	.	PROPN
gs-3955	377	41	,	,	PUNCT
gs-3955	377	42	2011	2011	NUM
gs-3955	377	43	;	;	PUNCT
gs-3955	377	44	tkemaladze	tkemaladze	NOUN
gs-3955	377	45	,	,	PUNCT
gs-3955	377	46	2023	2023	NUM
gs-3955	377	47	)	)	PUNCT
gs-3955	377	48	.	.	PUNCT
gs-3955	378	1	●	●	PUNCT
gs-3955	378	2	immune	immune	ADJ
gs-3955	378	3	and	and	CCONJ
gs-3955	378	4	neuroendocrine	neuroendocrine	ADJ
gs-3955	378	5	regulation	regulation	NOUN
gs-3955	378	6	:	:	PUNCT
gs-3955	378	7	human	human	ADJ
gs-3955	378	8	aging	aging	NOUN
gs-3955	378	9	is	be	AUX
gs-3955	378	10	strongly	strongly	ADV
gs-3955	378	11	influenced	influence	VERB
gs-3955	378	12	by	by	ADP
gs-3955	378	13	chronic	chronic	ADJ
gs-3955	378	14	inflammation	inflammation	NOUN
gs-3955	378	15	(	(	PUNCT
gs-3955	378	16	“	"	PUNCT
gs-3955	378	17	inflammaging	inflammage	VERB
gs-3955	378	18	”	"	PUNCT
gs-3955	378	19	)	)	PUNCT
gs-3955	378	20	and	and	CCONJ
gs-3955	378	21	altered	alter	VERB
gs-3955	378	22	stress	stress	NOUN
gs-3955	378	23	responses	response	NOUN
gs-3955	378	24	.	.	PUNCT
gs-3955	379	1	lessons	lesson	NOUN
gs-3955	379	2	from	from	ADP
gs-3955	379	3	naked	naked	ADJ
gs-3955	379	4	mole	mole	NOUN
gs-3955	379	5	-	-	PUNCT
gs-3955	379	6	rats	rat	NOUN
gs-3955	379	7	’	'	PUNCT
gs-3955	379	8	glucocorticoid	glucocorticoid	ADJ
gs-3955	379	9	receptor	receptor	NOUN
gs-3955	379	10	modulation	modulation	NOUN
gs-3955	379	11	and	and	CCONJ
gs-3955	379	12	expanded	expand	VERB
gs-3955	379	13	regulatory	regulatory	ADJ
gs-3955	379	14	t	t	NOUN
gs-3955	379	15	cell	cell	NOUN
gs-3955	379	16	populations	population	NOUN
gs-3955	379	17	suggest	suggest	VERB
gs-3955	379	18	avenues	avenue	NOUN
gs-3955	379	19	for	for	ADP
gs-3955	379	20	immunomodulatory	immunomodulatory	ADJ
gs-3955	379	21	therapies	therapy	NOUN
gs-3955	379	22	to	to	PART
gs-3955	379	23	reduce	reduce	VERB
gs-3955	379	24	inflammation	inflammation	NOUN
gs-3955	379	25	without	without	ADP
gs-3955	379	26	impairing	impair	VERB
gs-3955	379	27	immune	immune	ADJ
gs-3955	379	28	defense	defense	NOUN
gs-3955	379	29	(	(	PUNCT
gs-3955	379	30	macrae	macrae	PROPN
gs-3955	379	31	et	et	PROPN
gs-3955	379	32	al	al	PROPN
gs-3955	379	33	.	.	PROPN
gs-3955	379	34	,	,	PUNCT
gs-3955	379	35	2015	2015	NUM
gs-3955	379	36	;	;	PUNCT
gs-3955	379	37	petruseva	petruseva	VERB
gs-3955	379	38	et	et	PROPN
gs-3955	379	39	al	al	PROPN
gs-3955	379	40	.	.	PROPN
gs-3955	379	41	,	,	PUNCT
gs-3955	379	42	2017	2017	NUM
gs-3955	379	43	)	)	PUNCT
gs-3955	379	44	.	.	PUNCT
gs-3955	380	1	potential	potential	ADJ
gs-3955	380	2	therapeutic	therapeutic	ADJ
gs-3955	380	3	strategies	strategy	NOUN
gs-3955	380	4	gene	gene	NOUN
gs-3955	380	5	therapy	therapy	NOUN
gs-3955	380	6	and	and	CCONJ
gs-3955	380	7	genome	genome	NOUN
gs-3955	380	8	editing	editing	NOUN
gs-3955	380	9	:	:	PUNCT
gs-3955	380	10	georgian	georgian	ADJ
gs-3955	380	11	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	380	12	მეცნიერები	მეცნიერები	NOUN
gs-3955	380	13	ტ.	ტ.	VERB
gs-3955	380	14	7	7	NUM
gs-3955	380	15	n	n	PRON
gs-3955	380	16	3	3	NUM
gs-3955	380	17	,	,	PUNCT
gs-3955	380	18	2025	2025	NUM
gs-3955	380	19	387	387	NUM
gs-3955	380	20	○	○	PROPN
gs-3955	380	21	upregulating	upregulating	ADJ
gs-3955	380	22	dna	dna	PROPN
gs-3955	380	23	repair	repair	NOUN
gs-3955	380	24	genes	gene	NOUN
gs-3955	380	25	(	(	PUNCT
gs-3955	380	26	e.g.	e.g.	ADV
gs-3955	380	27	,	,	PUNCT
gs-3955	380	28	sirt6	sirt6	NOUN
gs-3955	380	29	,	,	PUNCT
gs-3955	380	30	parp1	parp1	NOUN
gs-3955	380	31	)	)	PUNCT
gs-3955	380	32	or	or	CCONJ
gs-3955	380	33	introducing	introduce	VERB
gs-3955	380	34	sharkderived	sharkderive	VERB
gs-3955	380	35	high	high	ADJ
gs-3955	380	36	-	-	PUNCT
gs-3955	380	37	fidelity	fidelity	NOUN
gs-3955	380	38	polymerases	polymerase	NOUN
gs-3955	380	39	could	could	AUX
gs-3955	380	40	enhance	enhance	VERB
gs-3955	380	41	genomic	genomic	ADJ
gs-3955	380	42	stability	stability	NOUN
gs-3955	380	43	(	(	PUNCT
gs-3955	380	44	narayan	narayan	PROPN
gs-3955	380	45	et	et	PROPN
gs-3955	380	46	al	al	PROPN
gs-3955	380	47	.	.	PROPN
gs-3955	380	48	,	,	PUNCT
gs-3955	380	49	2021	2021	NUM
gs-3955	380	50	;	;	PUNCT
gs-3955	380	51	petruseva	petruseva	VERB
gs-3955	380	52	et	et	PROPN
gs-3955	380	53	al	al	PROPN
gs-3955	380	54	.	.	PROPN
gs-3955	380	55	,	,	PUNCT
gs-3955	380	56	2017	2017	NUM
gs-3955	380	57	)	)	PUNCT
gs-3955	380	58	.	.	PUNCT
gs-3955	381	1	○	○	PROPN
gs-3955	381	2	engineering	engineering	PROPN
gs-3955	381	3	human	human	ADJ
gs-3955	381	4	stem	stem	NOUN
gs-3955	381	5	cells	cell	NOUN
gs-3955	381	6	to	to	PART
gs-3955	381	7	express	express	VERB
gs-3955	381	8	factors	factor	NOUN
gs-3955	381	9	promoting	promote	VERB
gs-3955	381	10	symmetric	symmetric	ADJ
gs-3955	381	11	division	division	NOUN
gs-3955	381	12	or	or	CCONJ
gs-3955	381	13	telomerase	telomerase	NOUN
gs-3955	381	14	activity	activity	NOUN
gs-3955	381	15	,	,	PUNCT
gs-3955	381	16	paired	pair	VERB
gs-3955	381	17	with	with	ADP
gs-3955	381	18	safeguards	safeguard	NOUN
gs-3955	381	19	against	against	ADP
gs-3955	381	20	tumorigenesis	tumorigenesis	NOUN
gs-3955	381	21	(	(	PUNCT
gs-3955	381	22	tkemaladze	tkemaladze	NOUN
gs-3955	381	23	,	,	PUNCT
gs-3955	381	24	2023	2023	NUM
gs-3955	381	25	;	;	PUNCT
gs-3955	381	26	wagner	wagner	PROPN
gs-3955	381	27	et	et	PROPN
gs-3955	381	28	al	al	PROPN
gs-3955	381	29	.	.	PROPN
gs-3955	381	30	,	,	PUNCT
gs-3955	381	31	2011	2011	NUM
gs-3955	381	32	)	)	PUNCT
gs-3955	381	33	.	.	PUNCT
gs-3955	382	1	○	○	PROPN
gs-3955	382	2	modifying	modify	VERB
gs-3955	382	3	lipid	lipid	NOUN
gs-3955	382	4	metabolism	metabolism	NOUN
gs-3955	382	5	genes	gene	NOUN
gs-3955	382	6	to	to	PART
gs-3955	382	7	mimic	mimic	VERB
gs-3955	382	8	the	the	DET
gs-3955	382	9	protective	protective	ADJ
gs-3955	382	10	membrane	membrane	NOUN
gs-3955	382	11	compositions	composition	NOUN
gs-3955	382	12	of	of	ADP
gs-3955	382	13	long	long	ADV
gs-3955	382	14	-	-	PUNCT
gs-3955	382	15	lived	live	VERB
gs-3955	382	16	species	specie	NOUN
gs-3955	382	17	(	(	PUNCT
gs-3955	382	18	munro	munro	PROPN
gs-3955	382	19	et	et	PROPN
gs-3955	382	20	al	al	PROPN
gs-3955	382	21	.	.	PROPN
gs-3955	382	22	,	,	PUNCT
gs-3955	382	23	2012	2012	NUM
gs-3955	382	24	)	)	PUNCT
gs-3955	382	25	.	.	PUNCT
gs-3955	383	1	pharmacological	pharmacological	ADJ
gs-3955	383	2	interventions	intervention	NOUN
gs-3955	383	3	:	:	PUNCT
gs-3955	383	4	○	○	ADJ
gs-3955	383	5	small	small	ADJ
gs-3955	383	6	molecules	molecule	NOUN
gs-3955	383	7	activating	activate	VERB
gs-3955	383	8	longevity	longevity	NOUN
gs-3955	383	9	pathways	pathway	NOUN
gs-3955	383	10	such	such	ADJ
gs-3955	383	11	as	as	ADP
gs-3955	383	12	sirt6	sirt6	ADJ
gs-3955	383	13	activators	activator	NOUN
gs-3955	383	14	,	,	PUNCT
gs-3955	383	15	foxo	foxo	NOUN
gs-3955	383	16	stabilizers	stabilizer	NOUN
gs-3955	383	17	,	,	PUNCT
gs-3955	383	18	or	or	CCONJ
gs-3955	383	19	compounds	compound	NOUN
gs-3955	383	20	that	that	PRON
gs-3955	383	21	boost	boost	VERB
gs-3955	383	22	autophagy	autophagy	NOUN
gs-3955	383	23	and	and	CCONJ
gs-3955	383	24	proteostasis	proteostasis	NOUN
gs-3955	383	25	(	(	PUNCT
gs-3955	383	26	lewis	lewis	PROPN
gs-3955	383	27	et	et	PROPN
gs-3955	383	28	al	al	PROPN
gs-3955	383	29	.	.	PROPN
gs-3955	383	30	,	,	PUNCT
gs-3955	383	31	2012	2012	NUM
gs-3955	383	32	)	)	PUNCT
gs-3955	383	33	.	.	PUNCT
gs-3955	384	1	○	○	ADJ
gs-3955	384	2	drugs	drug	NOUN
gs-3955	384	3	targeting	target	VERB
gs-3955	384	4	metabolic	metabolic	NOUN
gs-3955	384	5	flexibility	flexibility	NOUN
gs-3955	384	6	,	,	PUNCT
gs-3955	384	7	such	such	ADJ
gs-3955	384	8	as	as	ADP
gs-3955	384	9	ketone	ketone	NOUN
gs-3955	384	10	body	body	NOUN
gs-3955	384	11	mimetics	mimetic	NOUN
gs-3955	384	12	or	or	CCONJ
gs-3955	384	13	mild	mild	ADJ
gs-3955	384	14	mitochondrial	mitochondrial	ADJ
gs-3955	384	15	uncouplers	uncoupler	NOUN
gs-3955	384	16	,	,	PUNCT
gs-3955	384	17	to	to	PART
gs-3955	384	18	reduce	reduce	VERB
gs-3955	384	19	oxidative	oxidative	ADJ
gs-3955	384	20	stress	stress	NOUN
gs-3955	384	21	(	(	PUNCT
gs-3955	384	22	munro	munro	PROPN
gs-3955	384	23	et	et	PROPN
gs-3955	384	24	al	al	PROPN
gs-3955	384	25	.	.	PROPN
gs-3955	384	26	,	,	PUNCT
gs-3955	384	27	2012	2012	NUM
gs-3955	384	28	;	;	PUNCT
gs-3955	384	29	narayan	narayan	PROPN
gs-3955	384	30	et	et	PROPN
gs-3955	384	31	al	al	PROPN
gs-3955	384	32	.	.	PROPN
gs-3955	384	33	,	,	PUNCT
gs-3955	384	34	2021	2021	NUM
gs-3955	384	35	)	)	PUNCT
gs-3955	384	36	.	.	PUNCT
gs-3955	385	1	○	○	ADJ
gs-3955	385	2	anti	anti	ADJ
gs-3955	385	3	-	-	ADJ
gs-3955	385	4	inflammatory	inflammatory	ADJ
gs-3955	385	5	agents	agent	NOUN
gs-3955	385	6	inspired	inspire	VERB
gs-3955	385	7	by	by	ADP
gs-3955	385	8	immune	immune	ADJ
gs-3955	385	9	adaptations	adaptation	NOUN
gs-3955	385	10	in	in	ADP
gs-3955	385	11	naked	naked	ADJ
gs-3955	385	12	molerats	molerat	NOUN
gs-3955	385	13	and	and	CCONJ
gs-3955	385	14	turtles	turtle	NOUN
gs-3955	385	15	,	,	PUNCT
gs-3955	385	16	balancing	balance	VERB
gs-3955	385	17	immune	immune	ADJ
gs-3955	385	18	surveillance	surveillance	NOUN
gs-3955	385	19	and	and	CCONJ
gs-3955	385	20	inflammation	inflammation	NOUN
gs-3955	385	21	control	control	NOUN
gs-3955	385	22	(	(	PUNCT
gs-3955	385	23	macrae	macrae	PROPN
gs-3955	385	24	et	et	PROPN
gs-3955	385	25	al	al	PROPN
gs-3955	385	26	.	.	PROPN
gs-3955	385	27	,	,	PUNCT
gs-3955	385	28	2015	2015	NUM
gs-3955	385	29	)	)	PUNCT
gs-3955	385	30	.	.	PUNCT
gs-3955	386	1	synthetic	synthetic	ADJ
gs-3955	386	2	biology	biology	NOUN
gs-3955	386	3	and	and	CCONJ
gs-3955	386	4	systems	system	NOUN
gs-3955	386	5	medicine	medicine	NOUN
gs-3955	386	6	:	:	PUNCT
gs-3955	386	7	○	○	ADJ
gs-3955	386	8	designing	design	VERB
gs-3955	386	9	gene	gene	NOUN
gs-3955	386	10	circuits	circuit	NOUN
gs-3955	386	11	that	that	PRON
gs-3955	386	12	replicate	replicate	VERB
gs-3955	386	13	integrated	integrate	VERB
gs-3955	386	14	longevity	longevity	NOUN
gs-3955	386	15	networks	network	NOUN
gs-3955	386	16	—	—	PUNCT
gs-3955	386	17	combining	combine	VERB
gs-3955	386	18	dna	dna	PROPN
gs-3955	386	19	repair	repair	NOUN
gs-3955	386	20	,	,	PUNCT
gs-3955	386	21	stress	stress	NOUN
gs-3955	386	22	response	response	NOUN
gs-3955	386	23	,	,	PUNCT
gs-3955	386	24	and	and	CCONJ
gs-3955	386	25	metabolic	metabolic	NOUN
gs-3955	386	26	regulation	regulation	NOUN
gs-3955	386	27	(	(	PUNCT
gs-3955	386	28	tkemaladze	tkemaladze	NOUN
gs-3955	386	29	,	,	PUNCT
gs-3955	386	30	2023	2023	NUM
gs-3955	386	31	)	)	PUNCT
gs-3955	386	32	.	.	PUNCT
gs-3955	387	1	○	○	PROPN
gs-3955	387	2	delivering	deliver	VERB
gs-3955	387	3	synthetic	synthetic	ADJ
gs-3955	387	4	mrnas	mrna	NOUN
gs-3955	387	5	or	or	CCONJ
gs-3955	387	6	protein	protein	NOUN
gs-3955	387	7	therapies	therapy	NOUN
gs-3955	387	8	(	(	PUNCT
gs-3955	387	9	e.g.	e.g.	ADV
gs-3955	387	10	,	,	PUNCT
gs-3955	387	11	chaperones	chaperone	NOUN
gs-3955	387	12	,	,	PUNCT
gs-3955	387	13	dna	dna	PROPN
gs-3955	387	14	repair	repair	NOUN
gs-3955	387	15	enzymes	enzyme	NOUN
gs-3955	387	16	)	)	PUNCT
gs-3955	387	17	via	via	ADP
gs-3955	387	18	lipid	lipid	ADJ
gs-3955	387	19	nanoparticles	nanoparticle	NOUN
gs-3955	387	20	for	for	ADP
gs-3955	387	21	transient	transient	ADJ
gs-3955	387	22	functional	functional	ADJ
gs-3955	387	23	enhancement	enhancement	NOUN
gs-3955	387	24	(	(	PUNCT
gs-3955	387	25	petruseva	petruseva	PROPN
gs-3955	387	26	et	et	PROPN
gs-3955	387	27	al	al	PROPN
gs-3955	387	28	.	.	PROPN
gs-3955	387	29	,	,	PUNCT
gs-3955	387	30	2017	2017	NUM
gs-3955	387	31	)	)	PUNCT
gs-3955	387	32	.	.	PUNCT
gs-3955	388	1	○	○	PROPN
gs-3955	388	2	developing	develop	VERB
gs-3955	388	3	human	human	ADJ
gs-3955	388	4	-	-	PUNCT
gs-3955	388	5	animal	animal	NOUN
gs-3955	388	6	chimeric	chimeric	ADJ
gs-3955	388	7	models	model	NOUN
gs-3955	388	8	to	to	PART
gs-3955	388	9	test	test	VERB
gs-3955	388	10	and	and	CCONJ
gs-3955	388	11	refine	refine	VERB
gs-3955	388	12	these	these	DET
gs-3955	388	13	interventions	intervention	NOUN
gs-3955	388	14	in	in	ADP
gs-3955	388	15	vivo	vivo	NOUN
gs-3955	388	16	(	(	PUNCT
gs-3955	388	17	wagner	wagner	PROPN
gs-3955	388	18	et	et	PROPN
gs-3955	388	19	al	al	PROPN
gs-3955	388	20	.	.	PROPN
gs-3955	388	21	,	,	PUNCT
gs-3955	388	22	2011	2011	NUM
gs-3955	388	23	)	)	PUNCT
gs-3955	388	24	.	.	PUNCT
gs-3955	389	1	lifestyle	lifestyle	NOUN
gs-3955	389	2	and	and	CCONJ
gs-3955	389	3	environmental	environmental	ADJ
gs-3955	389	4	modulation	modulation	NOUN
gs-3955	389	5	:	:	PUNCT
gs-3955	389	6	while	while	SCONJ
gs-3955	389	7	not	not	PART
gs-3955	389	8	molecular	molecular	ADJ
gs-3955	389	9	,	,	PUNCT
gs-3955	389	10	lifestyle	lifestyle	NOUN
gs-3955	389	11	factors	factor	NOUN
gs-3955	389	12	influencing	influence	VERB
gs-3955	389	13	metabolism	metabolism	NOUN
gs-3955	389	14	,	,	PUNCT
gs-3955	389	15	inflammation	inflammation	NOUN
gs-3955	389	16	,	,	PUNCT
gs-3955	389	17	and	and	CCONJ
gs-3955	389	18	dna	dna	PROPN
gs-3955	389	19	damage	damage	NOUN
gs-3955	389	20	repair	repair	NOUN
gs-3955	389	21	synergize	synergize	VERB
gs-3955	389	22	with	with	ADP
gs-3955	389	23	molecular	molecular	ADJ
gs-3955	389	24	therapies	therapy	NOUN
gs-3955	389	25	to	to	PART
gs-3955	389	26	promote	promote	VERB
gs-3955	389	27	longevity	longevity	NOUN
gs-3955	389	28	.	.	PUNCT
gs-3955	390	1	challenges	challenge	NOUN
gs-3955	390	2	and	and	CCONJ
gs-3955	390	3	ethical	ethical	ADJ
gs-3955	390	4	considerations	consideration	NOUN
gs-3955	390	5	●	●	PUNCT
gs-3955	390	6	directly	directly	ADV
gs-3955	390	7	transferring	transfer	VERB
gs-3955	390	8	longevity	longevity	NOUN
gs-3955	390	9	traits	trait	NOUN
gs-3955	390	10	from	from	ADP
gs-3955	390	11	these	these	DET
gs-3955	390	12	species	specie	NOUN
gs-3955	390	13	to	to	ADP
gs-3955	390	14	humans	human	NOUN
gs-3955	390	15	requires	require	VERB
gs-3955	390	16	overcoming	overcome	VERB
gs-3955	390	17	evolutionary	evolutionary	ADJ
gs-3955	390	18	divergence	divergence	NOUN
gs-3955	390	19	and	and	CCONJ
gs-3955	390	20	complex	complex	ADJ
gs-3955	390	21	trade	trade	NOUN
gs-3955	390	22	-	-	PUNCT
gs-3955	390	23	offs	off	NOUN
gs-3955	390	24	,	,	PUNCT
gs-3955	390	25	such	such	ADJ
gs-3955	390	26	as	as	ADP
gs-3955	390	27	cancer	cancer	NOUN
gs-3955	390	28	risk	risk	NOUN
gs-3955	390	29	and	and	CCONJ
gs-3955	390	30	cognitive	cognitive	ADJ
gs-3955	390	31	function	function	NOUN
gs-3955	390	32	(	(	PUNCT
gs-3955	390	33	lewis	lewis	PROPN
gs-3955	390	34	et	et	PROPN
gs-3955	390	35	al	al	PROPN
gs-3955	390	36	.	.	PROPN
gs-3955	390	37	,	,	PUNCT
gs-3955	390	38	2012	2012	NUM
gs-3955	390	39	;	;	PUNCT
gs-3955	390	40	narayan	narayan	PROPN
gs-3955	390	41	et	et	PROPN
gs-3955	390	42	al	al	PROPN
gs-3955	390	43	.	.	PROPN
gs-3955	390	44	,	,	PUNCT
gs-3955	390	45	2021	2021	NUM
gs-3955	390	46	)	)	PUNCT
gs-3955	390	47	.	.	PUNCT
gs-3955	391	1	●	●	PUNCT
gs-3955	391	2	manipulating	manipulate	VERB
gs-3955	391	3	core	core	NOUN
gs-3955	391	4	pathways	pathway	NOUN
gs-3955	391	5	like	like	ADP
gs-3955	391	6	telomerase	telomerase	NOUN
gs-3955	391	7	or	or	CCONJ
gs-3955	391	8	metabolism	metabolism	NOUN
gs-3955	391	9	carries	carry	VERB
gs-3955	391	10	risks	risk	NOUN
gs-3955	391	11	that	that	PRON
gs-3955	391	12	demand	demand	VERB
gs-3955	391	13	careful	careful	ADJ
gs-3955	391	14	regulation	regulation	NOUN
gs-3955	391	15	and	and	CCONJ
gs-3955	391	16	multi	multi	ADJ
gs-3955	391	17	-	-	ADJ
gs-3955	391	18	targeted	target	VERB
gs-3955	391	19	approaches	approach	NOUN
gs-3955	391	20	(	(	PUNCT
gs-3955	391	21	petruseva	petruseva	PROPN
gs-3955	391	22	et	et	PROPN
gs-3955	391	23	al	al	PROPN
gs-3955	391	24	.	.	PROPN
gs-3955	391	25	,	,	PUNCT
gs-3955	391	26	2017	2017	NUM
gs-3955	391	27	)	)	PUNCT
gs-3955	391	28	.	.	PUNCT
gs-3955	392	1	●	●	PUNCT
gs-3955	392	2	societal	societal	ADJ
gs-3955	392	3	implications	implication	NOUN
gs-3955	392	4	—	—	PUNCT
gs-3955	392	5	such	such	ADJ
gs-3955	392	6	as	as	ADP
gs-3955	392	7	equitable	equitable	ADJ
gs-3955	392	8	access	access	NOUN
gs-3955	392	9	to	to	ADP
gs-3955	392	10	longevity	longevity	NOUN
gs-3955	392	11	therapies	therapy	NOUN
gs-3955	392	12	and	and	CCONJ
gs-3955	392	13	redefinition	redefinition	NOUN
gs-3955	392	14	of	of	ADP
gs-3955	392	15	aging	aging	NOUN
gs-3955	392	16	—	—	PUNCT
gs-3955	392	17	must	must	AUX
gs-3955	392	18	guide	guide	VERB
gs-3955	392	19	responsible	responsible	ADJ
gs-3955	392	20	development	development	NOUN
gs-3955	392	21	and	and	CCONJ
gs-3955	392	22	deployment	deployment	NOUN
gs-3955	392	23	.	.	PUNCT
gs-3955	393	1	georgian	georgian	ADJ
gs-3955	393	2	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	393	3	მეცნიერები	მეცნიერები	NOUN
gs-3955	393	4	ტ.	ტ.	VERB
gs-3955	393	5	7	7	NUM
gs-3955	393	6	n	n	PRON
gs-3955	393	7	3	3	NUM
gs-3955	393	8	,	,	PUNCT
gs-3955	393	9	2025	2025	NUM
gs-3955	393	10	388	388	NUM
gs-3955	393	11	while	while	SCONJ
gs-3955	393	12	humans	human	NOUN
gs-3955	393	13	share	share	VERB
gs-3955	393	14	many	many	ADJ
gs-3955	393	15	molecular	molecular	ADJ
gs-3955	393	16	tools	tool	NOUN
gs-3955	393	17	with	with	ADP
gs-3955	393	18	negligibly	negligibly	ADV
gs-3955	393	19	senescent	senescent	ADJ
gs-3955	393	20	species	specie	NOUN
gs-3955	393	21	,	,	PUNCT
gs-3955	393	22	the	the	DET
gs-3955	393	23	differences	difference	NOUN
gs-3955	393	24	highlight	highlight	VERB
gs-3955	393	25	the	the	DET
gs-3955	393	26	necessity	necessity	NOUN
gs-3955	393	27	for	for	ADP
gs-3955	393	28	multifaceted	multifaceted	ADJ
gs-3955	393	29	,	,	PUNCT
gs-3955	393	30	precision	precision	NOUN
gs-3955	393	31	interventions	intervention	NOUN
gs-3955	393	32	.	.	PUNCT
gs-3955	394	1	by	by	ADP
gs-3955	394	2	leveraging	leverage	VERB
gs-3955	394	3	gene	gene	NOUN
gs-3955	394	4	therapy	therapy	NOUN
gs-3955	394	5	,	,	PUNCT
gs-3955	394	6	pharmacology	pharmacology	NOUN
gs-3955	394	7	,	,	PUNCT
gs-3955	394	8	synthetic	synthetic	ADJ
gs-3955	394	9	biology	biology	NOUN
gs-3955	394	10	,	,	PUNCT
gs-3955	394	11	and	and	CCONJ
gs-3955	394	12	systemic	systemic	ADJ
gs-3955	394	13	medicine	medicine	NOUN
gs-3955	394	14	,	,	PUNCT
gs-3955	394	15	future	future	ADJ
gs-3955	394	16	strategies	strategy	NOUN
gs-3955	394	17	may	may	AUX
gs-3955	394	18	recreate	recreate	VERB
gs-3955	394	19	the	the	DET
gs-3955	394	20	resilient	resilient	ADJ
gs-3955	394	21	molecular	molecular	ADJ
gs-3955	394	22	landscape	landscape	NOUN
gs-3955	394	23	of	of	ADP
gs-3955	394	24	these	these	DET
gs-3955	394	25	extraordinary	extraordinary	ADJ
gs-3955	394	26	organisms	organism	NOUN
gs-3955	394	27	,	,	PUNCT
gs-3955	394	28	heralding	herald	VERB
gs-3955	394	29	a	a	DET
gs-3955	394	30	new	new	ADJ
gs-3955	394	31	era	era	NOUN
gs-3955	394	32	of	of	ADP
gs-3955	394	33	human	human	PROPN
gs-3955	394	34	healthspan	healthspan	PROPN
gs-3955	394	35	extension	extension	NOUN
gs-3955	394	36	.	.	PUNCT
gs-3955	395	1	conclusion	conclusion	NOUN
gs-3955	395	2	the	the	DET
gs-3955	395	3	extraordinary	extraordinary	ADJ
gs-3955	395	4	biology	biology	NOUN
gs-3955	395	5	of	of	ADP
gs-3955	395	6	negligibly	negligibly	ADV
gs-3955	395	7	senescent	senescent	ADJ
gs-3955	395	8	species	specie	NOUN
gs-3955	395	9	irrevocably	irrevocably	ADV
gs-3955	395	10	transforms	transform	VERB
gs-3955	395	11	our	our	PRON
gs-3955	395	12	understanding	understanding	NOUN
gs-3955	395	13	of	of	ADP
gs-3955	395	14	aging	aging	NOUN
gs-3955	395	15	.	.	PUNCT
gs-3955	396	1	these	these	DET
gs-3955	396	2	organisms	organism	NOUN
gs-3955	396	3	—	—	PUNCT
gs-3955	396	4	greenland	greenland	PROPN
gs-3955	396	5	sharks	shark	NOUN
gs-3955	396	6	gliding	glide	VERB
gs-3955	396	7	through	through	ADP
gs-3955	396	8	arctic	arctic	ADJ
gs-3955	396	9	depths	depth	NOUN
gs-3955	396	10	for	for	ADP
gs-3955	396	11	centuries	century	NOUN
gs-3955	396	12	,	,	PUNCT
gs-3955	396	13	hydra	hydra	NOUN
gs-3955	396	14	regenerating	regenerate	VERB
gs-3955	396	15	perpetually	perpetually	ADV
gs-3955	396	16	in	in	ADP
gs-3955	396	17	freshwater	freshwater	NOUN
gs-3955	396	18	ponds	pond	NOUN
gs-3955	396	19	,	,	PUNCT
gs-3955	396	20	naked	naked	ADJ
gs-3955	396	21	mole	mole	NOUN
gs-3955	396	22	-	-	PUNCT
gs-3955	396	23	rats	rat	NOUN
gs-3955	396	24	defying	defy	VERB
gs-3955	396	25	cancer	cancer	NOUN
gs-3955	396	26	in	in	ADP
gs-3955	396	27	subterranean	subterranean	ADJ
gs-3955	396	28	colonies	colony	NOUN
gs-3955	396	29	,	,	PUNCT
gs-3955	396	30	and	and	CCONJ
gs-3955	396	31	ocean	ocean	NOUN
gs-3955	396	32	quahogs	quahog	NOUN
gs-3955	396	33	enduring	endure	VERB
gs-3955	396	34	half	half	ADJ
gs-3955	396	35	-	-	PUNCT
gs-3955	396	36	millennia	millennia	NOUN
gs-3955	396	37	in	in	ADP
gs-3955	396	38	icy	icy	ADJ
gs-3955	396	39	seabeds	seabed	NOUN
gs-3955	396	40	—	—	PUNCT
gs-3955	396	41	collectively	collectively	ADV
gs-3955	396	42	prove	prove	VERB
gs-3955	396	43	that	that	SCONJ
gs-3955	396	44	biological	biological	ADJ
gs-3955	396	45	aging	aging	NOUN
gs-3955	396	46	is	be	AUX
gs-3955	396	47	neither	neither	CCONJ
gs-3955	396	48	universal	universal	ADJ
gs-3955	396	49	nor	nor	CCONJ
gs-3955	396	50	inevitable	inevitable	ADJ
gs-3955	396	51	.	.	PUNCT
gs-3955	397	1	their	their	PRON
gs-3955	397	2	existence	existence	NOUN
gs-3955	397	3	demonstrates	demonstrate	VERB
gs-3955	397	4	that	that	DET
gs-3955	397	5	time	time	NOUN
gs-3955	397	6	and	and	CCONJ
gs-3955	397	7	decay	decay	NOUN
gs-3955	397	8	can	can	AUX
gs-3955	397	9	be	be	AUX
gs-3955	397	10	decoupled	decouple	VERB
gs-3955	397	11	through	through	ADP
gs-3955	397	12	evolutionary	evolutionary	ADJ
gs-3955	397	13	innovation	innovation	NOUN
gs-3955	397	14	.	.	PUNCT
gs-3955	398	1	at	at	ADP
gs-3955	398	2	the	the	DET
gs-3955	398	3	molecular	molecular	ADJ
gs-3955	398	4	level	level	NOUN
gs-3955	398	5	,	,	PUNCT
gs-3955	398	6	they	they	PRON
gs-3955	398	7	achieve	achieve	VERB
gs-3955	398	8	this	this	DET
gs-3955	398	9	feat	feat	NOUN
gs-3955	398	10	via	via	ADP
gs-3955	398	11	orchestrated	orchestrated	ADJ
gs-3955	398	12	adaptations	adaptation	NOUN
gs-3955	398	13	:	:	PUNCT
gs-3955	398	14	enhanced	enhance	VERB
gs-3955	398	15	dna	dna	PROPN
gs-3955	398	16	repair	repair	NOUN
gs-3955	398	17	machinery	machinery	NOUN
gs-3955	398	18	that	that	PRON
gs-3955	398	19	corrects	correct	VERB
gs-3955	398	20	genomic	genomic	ADJ
gs-3955	398	21	errors	error	NOUN
gs-3955	398	22	with	with	ADP
gs-3955	398	23	unparalleled	unparalleled	ADJ
gs-3955	398	24	fidelity	fidelity	NOUN
gs-3955	398	25	;	;	PUNCT
gs-3955	398	26	proteostatic	proteostatic	NOUN
gs-3955	398	27	systems	system	NOUN
gs-3955	398	28	that	that	PRON
gs-3955	398	29	maintain	maintain	VERB
gs-3955	398	30	protein	protein	NOUN
gs-3955	398	31	integrity	integrity	NOUN
gs-3955	398	32	across	across	ADP
gs-3955	398	33	centuries	century	NOUN
gs-3955	398	34	;	;	PUNCT
gs-3955	398	35	and	and	CCONJ
gs-3955	398	36	stress	stress	NOUN
gs-3955	398	37	resistance	resistance	NOUN
gs-3955	398	38	mechanisms	mechanism	NOUN
gs-3955	398	39	that	that	PRON
gs-3955	398	40	tolerate	tolerate	VERB
gs-3955	398	41	or	or	CCONJ
gs-3955	398	42	neutralize	neutralize	VERB
gs-3955	398	43	damage	damage	NOUN
gs-3955	398	44	without	without	ADP
gs-3955	398	45	functional	functional	ADJ
gs-3955	398	46	decline	decline	NOUN
gs-3955	398	47	.	.	PUNCT
gs-3955	399	1	crucially	crucially	ADV
gs-3955	399	2	,	,	PUNCT
gs-3955	399	3	their	their	PRON
gs-3955	399	4	stem	stem	NOUN
gs-3955	399	5	cell	cell	NOUN
gs-3955	399	6	systems	system	NOUN
gs-3955	399	7	—	—	PUNCT
gs-3955	399	8	from	from	ADP
gs-3955	399	9	hydra	hydra	PROPN
gs-3955	399	10	’s	’s	PART
gs-3955	399	11	foxo	foxo	PROPN
gs-3955	399	12	-	-	PUNCT
gs-3955	399	13	driven	drive	VERB
gs-3955	399	14	renewal	renewal	NOUN
gs-3955	399	15	to	to	ADP
gs-3955	399	16	planarian	planarian	ADJ
gs-3955	399	17	neoblasts	neoblast	NOUN
gs-3955	399	18	—	—	PUNCT
gs-3955	399	19	reveal	reveal	VERB
gs-3955	399	20	that	that	SCONJ
gs-3955	399	21	cellular	cellular	ADJ
gs-3955	399	22	immortality	immortality	NOUN
gs-3955	399	23	is	be	AUX
gs-3955	399	24	achievable	achievable	ADJ
gs-3955	399	25	through	through	ADP
gs-3955	399	26	symmetric	symmetric	ADJ
gs-3955	399	27	division	division	NOUN
gs-3955	399	28	and	and	CCONJ
gs-3955	399	29	epigenetic	epigenetic	ADJ
gs-3955	399	30	stability	stability	NOUN
gs-3955	399	31	,	,	PUNCT
gs-3955	399	32	challenging	challenge	VERB
gs-3955	399	33	the	the	DET
gs-3955	399	34	dogma	dogma	NOUN
gs-3955	399	35	of	of	ADP
gs-3955	399	36	inevitable	inevitable	ADJ
gs-3955	399	37	replicative	replicative	ADJ
gs-3955	399	38	senescence	senescence	NOUN
gs-3955	399	39	.	.	PUNCT
gs-3955	400	1	these	these	DET
gs-3955	400	2	species	specie	NOUN
gs-3955	400	3	converge	converge	VERB
gs-3955	400	4	on	on	ADP
gs-3955	400	5	a	a	DET
gs-3955	400	6	profound	profound	ADJ
gs-3955	400	7	biological	biological	ADJ
gs-3955	400	8	principle	principle	NOUN
gs-3955	400	9	:	:	PUNCT
gs-3955	400	10	longevity	longevity	NOUN
gs-3955	400	11	arises	arise	VERB
gs-3955	400	12	from	from	ADP
gs-3955	400	13	integrated	integrated	ADJ
gs-3955	400	14	networks	network	NOUN
gs-3955	400	15	preserving	preserve	VERB
gs-3955	400	16	cellular	cellular	ADJ
gs-3955	400	17	information	information	NOUN
gs-3955	400	18	against	against	ADP
gs-3955	400	19	entropic	entropic	ADJ
gs-3955	400	20	decay	decay	NOUN
gs-3955	400	21	.	.	PUNCT
gs-3955	401	1	the	the	DET
gs-3955	401	2	centrosomal	centrosomal	ADJ
gs-3955	401	3	theory	theory	NOUN
gs-3955	401	4	of	of	ADP
gs-3955	401	5	aging	aging	NOUN
gs-3955	401	6	provides	provide	VERB
gs-3955	401	7	a	a	DET
gs-3955	401	8	unifying	unifying	ADJ
gs-3955	401	9	lens	len	NOUN
gs-3955	401	10	—	—	PUNCT
gs-3955	401	11	aging	age	VERB
gs-3955	401	12	manifests	manifest	NOUN
gs-3955	401	13	where	where	SCONJ
gs-3955	401	14	asymmetric	asymmetric	ADJ
gs-3955	401	15	cell	cell	NOUN
gs-3955	401	16	division	division	NOUN
gs-3955	401	17	distributes	distribute	VERB
gs-3955	401	18	damaged	damage	VERB
gs-3955	401	19	components	component	NOUN
gs-3955	401	20	unevenly	unevenly	ADV
gs-3955	401	21	,	,	PUNCT
gs-3955	401	22	while	while	SCONJ
gs-3955	401	23	negligibly	negligibly	ADV
gs-3955	401	24	senescent	senescent	ADJ
gs-3955	401	25	species	specie	NOUN
gs-3955	401	26	avoid	avoid	VERB
gs-3955	401	27	this	this	PRON
gs-3955	401	28	through	through	ADP
gs-3955	401	29	rejuvenating	rejuvenate	VERB
gs-3955	401	30	division	division	NOUN
gs-3955	401	31	patterns	pattern	NOUN
gs-3955	401	32	,	,	PUNCT
gs-3955	401	33	selective	selective	ADJ
gs-3955	401	34	organelle	organelle	NOUN
gs-3955	401	35	recycling	recycling	NOUN
gs-3955	401	36	,	,	PUNCT
gs-3955	401	37	and	and	CCONJ
gs-3955	401	38	error	error	NOUN
gs-3955	401	39	-	-	PUNCT
gs-3955	401	40	proof	proof	ADJ
gs-3955	401	41	biomolecular	biomolecular	ADJ
gs-3955	401	42	structures	structure	NOUN
gs-3955	401	43	.	.	PUNCT
gs-3955	402	1	translating	translate	VERB
gs-3955	402	2	these	these	DET
gs-3955	402	3	insights	insight	NOUN
gs-3955	402	4	to	to	ADP
gs-3955	402	5	human	human	ADJ
gs-3955	402	6	medicine	medicine	NOUN
gs-3955	402	7	holds	hold	VERB
gs-3955	402	8	immense	immense	ADJ
gs-3955	402	9	promise	promise	NOUN
gs-3955	402	10	but	but	CCONJ
gs-3955	402	11	demands	demand	VERB
gs-3955	402	12	caution	caution	NOUN
gs-3955	402	13	.	.	PUNCT
gs-3955	403	1	naked	naked	ADJ
gs-3955	403	2	mole	mole	ADJ
gs-3955	403	3	-	-	PUNCT
gs-3955	403	4	rat	rat	NOUN
gs-3955	403	5	hmw	hmw	PROPN
gs-3955	403	6	-	-	PUNCT
gs-3955	403	7	ha	ha	PROPN
gs-3955	403	8	inspires	inspire	VERB
gs-3955	403	9	anti	anti	ADJ
gs-3955	403	10	-	-	ADJ
gs-3955	403	11	cancer	cancer	ADJ
gs-3955	403	12	biomaterials	biomaterial	NOUN
gs-3955	403	13	;	;	PUNCT
gs-3955	403	14	shark	shark	NOUN
gs-3955	403	15	lipid	lipid	NOUN
gs-3955	403	16	membranes	membrane	NOUN
gs-3955	403	17	inform	inform	VERB
gs-3955	403	18	cytoprotective	cytoprotective	ADJ
gs-3955	403	19	drugs	drug	NOUN
gs-3955	403	20	;	;	PUNCT
gs-3955	403	21	and	and	CCONJ
gs-3955	403	22	salamander	salamander	NOUN
gs-3955	403	23	regeneration	regeneration	NOUN
gs-3955	403	24	paradigms	paradigm	NOUN
gs-3955	403	25	guide	guide	NOUN
gs-3955	403	26	stem	stem	NOUN
gs-3955	403	27	cell	cell	NOUN
gs-3955	403	28	therapies	therapy	NOUN
gs-3955	403	29	.	.	PUNCT
gs-3955	404	1	yet	yet	ADV
gs-3955	404	2	interspecies	interspecie	VERB
gs-3955	404	3	barriers	barrier	NOUN
gs-3955	404	4	persist	persist	VERB
gs-3955	404	5	—	—	PUNCT
gs-3955	404	6	human	human	ADJ
gs-3955	404	7	biology	biology	NOUN
gs-3955	404	8	lacks	lack	VERB
gs-3955	404	9	genomic	genomic	ADJ
gs-3955	404	10	redundancies	redundancy	NOUN
gs-3955	404	11	like	like	ADP
gs-3955	404	12	shark	shark	NOUN
gs-3955	404	13	ercc1	ercc1	NOUN
gs-3955	404	14	duplications	duplication	NOUN
gs-3955	404	15	,	,	PUNCT
gs-3955	404	16	and	and	CCONJ
gs-3955	404	17	longevity	longevity	NOUN
gs-3955	404	18	adaptations	adaptation	NOUN
gs-3955	404	19	in	in	ADP
gs-3955	404	20	ectotherms	ectotherm	NOUN
gs-3955	404	21	(	(	PUNCT
gs-3955	404	22	e.g.	e.g.	ADV
gs-3955	404	23	,	,	PUNCT
gs-3955	404	24	metabolic	metabolic	ADJ
gs-3955	404	25	depression	depression	NOUN
gs-3955	404	26	)	)	PUNCT
gs-3955	404	27	conflict	conflict	NOUN
gs-3955	404	28	with	with	ADP
gs-3955	404	29	human	human	ADJ
gs-3955	404	30	physiology	physiology	NOUN
gs-3955	404	31	.	.	PUNCT
gs-3955	405	1	ethically	ethically	ADV
gs-3955	405	2	,	,	PUNCT
gs-3955	405	3	extending	extend	VERB
gs-3955	405	4	human	human	ADJ
gs-3955	405	5	healthspan	healthspan	NOUN
gs-3955	405	6	compels	compel	VERB
gs-3955	405	7	us	we	PRON
gs-3955	405	8	to	to	PART
gs-3955	405	9	confront	confront	VERB
gs-3955	405	10	equity	equity	NOUN
gs-3955	405	11	,	,	PUNCT
gs-3955	405	12	societal	societal	ADJ
gs-3955	405	13	restructuring	restructuring	NOUN
gs-3955	405	14	,	,	PUNCT
gs-3955	405	15	and	and	CCONJ
gs-3955	405	16	identity	identity	NOUN
gs-3955	405	17	continuity	continuity	NOUN
gs-3955	405	18	.	.	PUNCT
gs-3955	406	1	longevity	longevity	NOUN
gs-3955	406	2	interventions	intervention	NOUN
gs-3955	406	3	must	must	AUX
gs-3955	406	4	be	be	AUX
gs-3955	406	5	globally	globally	ADV
gs-3955	406	6	accessible	accessible	ADJ
gs-3955	406	7	,	,	PUNCT
gs-3955	406	8	not	not	PART
gs-3955	406	9	privileges	privilege	NOUN
gs-3955	406	10	of	of	ADP
gs-3955	406	11	wealth	wealth	NOUN
gs-3955	406	12	.	.	PUNCT
gs-3955	407	1	societies	society	NOUN
gs-3955	407	2	would	would	AUX
gs-3955	407	3	need	need	VERB
gs-3955	407	4	reimagined	reimagined	ADJ
gs-3955	407	5	models	model	NOUN
gs-3955	407	6	of	of	ADP
gs-3955	407	7	work	work	NOUN
gs-3955	407	8	,	,	PUNCT
gs-3955	407	9	inheritance	inheritance	NOUN
gs-3955	407	10	,	,	PUNCT
gs-3955	407	11	and	and	CCONJ
gs-3955	407	12	intergenerational	intergenerational	ADJ
gs-3955	407	13	dynamics	dynamic	NOUN
gs-3955	407	14	if	if	SCONJ
gs-3955	407	15	lifespans	lifespan	NOUN
gs-3955	407	16	approach	approach	VERB
gs-3955	407	17	150	150	NUM
gs-3955	407	18	years	year	NOUN
gs-3955	407	19	.	.	PUNCT
gs-3955	408	1	moreover	moreover	ADV
gs-3955	408	2	,	,	PUNCT
gs-3955	408	3	psychological	psychological	ADJ
gs-3955	408	4	resilience	resilience	NOUN
gs-3955	408	5	must	must	AUX
gs-3955	408	6	evolve	evolve	VERB
gs-3955	408	7	alongside	alongside	ADP
gs-3955	408	8	biological	biological	ADJ
gs-3955	408	9	vitality	vitality	NOUN
gs-3955	408	10	to	to	PART
gs-3955	408	11	ensure	ensure	VERB
gs-3955	408	12	extended	extended	ADJ
gs-3955	408	13	lifespans	lifespan	NOUN
gs-3955	408	14	enrich	enrich	VERB
gs-3955	408	15	human	human	ADJ
gs-3955	408	16	flourishing	flourishing	NOUN
gs-3955	408	17	rather	rather	ADV
gs-3955	408	18	than	than	ADP
gs-3955	408	19	prolong	prolong	ADJ
gs-3955	408	20	decline	decline	NOUN
gs-3955	408	21	.	.	PUNCT
gs-3955	409	1	looking	look	VERB
gs-3955	409	2	ahead	ahead	ADV
gs-3955	409	3	,	,	PUNCT
gs-3955	409	4	convergent	convergent	NOUN
gs-3955	409	5	science	science	NOUN
gs-3955	409	6	—	—	PUNCT
gs-3955	409	7	"	"	PUNCT
gs-3955	409	8	longevomics	longevomics	X
gs-3955	409	9	"	"	PUNCT
gs-3955	409	10	atlases	atlas	NOUN
gs-3955	409	11	of	of	ADP
gs-3955	409	12	centenarian	centenarian	ADJ
gs-3955	409	13	species	specie	NOUN
gs-3955	409	14	,	,	PUNCT
gs-3955	409	15	synthetic	synthetic	ADJ
gs-3955	409	16	gene	gene	NOUN
gs-3955	409	17	circuits	circuit	NOUN
gs-3955	409	18	mimicking	mimic	VERB
gs-3955	409	19	foxo	foxo	PROPN
gs-3955	409	20	-	-	PUNCT
gs-3955	409	21	sirt	sirt	NOUN
gs-3955	409	22	networks	network	NOUN
gs-3955	409	23	,	,	PUNCT
gs-3955	409	24	and	and	CCONJ
gs-3955	409	25	quantum	quantum	NOUN
gs-3955	409	26	-	-	ADJ
gs-3955	409	27	biophysical	biophysical	ADJ
gs-3955	409	28	probes	probe	NOUN
gs-3955	409	29	of	of	ADP
gs-3955	409	30	mitochondrial	mitochondrial	ADJ
gs-3955	409	31	efficiency	efficiency	NOUN
gs-3955	409	32	—	—	PUNCT
gs-3955	409	33	will	will	AUX
gs-3955	409	34	accelerate	accelerate	VERB
gs-3955	409	35	georgian	georgian	ADJ
gs-3955	409	36	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	409	37	მეცნიერები	მეცნიერები	NOUN
gs-3955	409	38	ტ.	ტ.	VERB
gs-3955	409	39	7	7	NUM
gs-3955	409	40	n	n	PRON
gs-3955	409	41	3	3	NUM
gs-3955	409	42	,	,	PUNCT
gs-3955	409	43	2025	2025	NUM
gs-3955	409	44	389	389	NUM
gs-3955	409	45	discovery	discovery	NOUN
gs-3955	409	46	.	.	PUNCT
gs-3955	410	1	the	the	DET
gs-3955	410	2	ultimate	ultimate	ADJ
gs-3955	410	3	legacy	legacy	NOUN
gs-3955	410	4	of	of	ADP
gs-3955	410	5	these	these	DET
gs-3955	410	6	ageless	ageless	ADJ
gs-3955	410	7	organisms	organism	NOUN
gs-3955	410	8	is	be	AUX
gs-3955	410	9	not	not	PART
gs-3955	410	10	mere	mere	ADJ
gs-3955	410	11	lifespan	lifespan	ADJ
gs-3955	410	12	extension	extension	NOUN
gs-3955	410	13	,	,	PUNCT
gs-3955	410	14	but	but	CCONJ
gs-3955	410	15	a	a	DET
gs-3955	410	16	vision	vision	NOUN
gs-3955	410	17	of	of	ADP
gs-3955	410	18	vitality	vitality	NOUN
gs-3955	410	19	without	without	ADP
gs-3955	410	20	decline	decline	NOUN
gs-3955	410	21	:	:	PUNCT
gs-3955	410	22	a	a	DET
gs-3955	410	23	future	future	NOUN
gs-3955	410	24	where	where	SCONJ
gs-3955	410	25	human	human	ADJ
gs-3955	410	26	healthspan	healthspan	PROPN
gs-3955	410	27	aligns	align	VERB
gs-3955	410	28	with	with	ADP
gs-3955	410	29	chronological	chronological	ADJ
gs-3955	410	30	lifespan	lifespan	NOUN
gs-3955	410	31	,	,	PUNCT
gs-3955	410	32	freeing	free	VERB
gs-3955	410	33	humanity	humanity	NOUN
gs-3955	410	34	from	from	ADP
gs-3955	410	35	the	the	DET
gs-3955	410	36	shadow	shadow	NOUN
gs-3955	410	37	of	of	ADP
gs-3955	410	38	age	age	NOUN
gs-3955	410	39	-	-	PUNCT
gs-3955	410	40	related	relate	VERB
gs-3955	410	41	degeneration	degeneration	NOUN
gs-3955	410	42	.	.	PUNCT
gs-3955	411	1	in	in	ADP
gs-3955	411	2	their	their	PRON
gs-3955	411	3	silent	silent	ADJ
gs-3955	411	4	endurance	endurance	NOUN
gs-3955	411	5	,	,	PUNCT
gs-3955	411	6	earth	earth	NOUN
gs-3955	411	7	’s	’s	PART
gs-3955	411	8	immortal	immortal	ADJ
gs-3955	411	9	species	specie	NOUN
gs-3955	411	10	offer	offer	VERB
gs-3955	411	11	not	not	PART
gs-3955	411	12	just	just	ADV
gs-3955	411	13	biomedical	biomedical	ADJ
gs-3955	411	14	insights	insight	NOUN
gs-3955	411	15	,	,	PUNCT
gs-3955	411	16	but	but	CCONJ
gs-3955	411	17	a	a	DET
gs-3955	411	18	radical	radical	ADJ
gs-3955	411	19	redefinition	redefinition	NOUN
gs-3955	411	20	of	of	ADP
gs-3955	411	21	life	life	NOUN
gs-3955	411	22	’s	’s	PART
gs-3955	411	23	temporal	temporal	ADJ
gs-3955	411	24	possibilities	possibility	NOUN
gs-3955	411	25	.	.	PUNCT
gs-3955	412	1	references	reference	NOUN
gs-3955	412	2	:	:	PUNCT
gs-3955	412	3	1	1	X
gs-3955	412	4	.	.	X
gs-3955	412	5	abele	abele	PROPN
gs-3955	412	6	,	,	PUNCT
gs-3955	412	7	d.	d.	PROPN
gs-3955	412	8	,	,	PUNCT
gs-3955	412	9	et	et	PROPN
gs-3955	412	10	al	al	PROPN
gs-3955	412	11	.	.	PUNCT
gs-3955	412	12	(	(	PUNCT
gs-3955	412	13	2008	2008	NUM
gs-3955	412	14	)	)	PUNCT
gs-3955	412	15	.	.	PUNCT
gs-3955	413	1	marine	marine	PROPN
gs-3955	413	2	invertebrate	invertebrate	NOUN
gs-3955	413	3	mitochondria	mitochondria	NOUN
gs-3955	413	4	and	and	CCONJ
gs-3955	413	5	oxidative	oxidative	ADJ
gs-3955	413	6	stress	stress	NOUN
gs-3955	413	7	.	.	PUNCT
gs-3955	414	1	frontiers	frontier	NOUN
gs-3955	414	2	in	in	ADP
gs-3955	414	3	bioscience	bioscience	NOUN
gs-3955	414	4	,	,	PUNCT
gs-3955	414	5	13	13	NUM
gs-3955	414	6	,	,	PUNCT
gs-3955	414	7	4809	4809	NUM
gs-3955	414	8	-	-	SYM
gs-3955	414	9	4821	4821	NUM
gs-3955	414	10	.	.	PUNCT
gs-3955	415	1	[	[	X
gs-3955	415	2	pubmed	pubmed	NOUN
gs-3955	415	3	:	:	PUNCT
gs-3955	415	4	18508566	18508566	NUM
gs-3955	415	5	]	]	PUNCT
gs-3955	415	6	2	2	NUM
gs-3955	415	7	.	.	NOUN
gs-3955	415	8	austad	austad	NOUN
gs-3955	415	9	,	,	PUNCT
gs-3955	415	10	s.	s.	PROPN
gs-3955	415	11	n.	n.	PROPN
gs-3955	415	12	(	(	PUNCT
gs-3955	415	13	2010	2010	NUM
gs-3955	415	14	)	)	PUNCT
gs-3955	415	15	.	.	PUNCT
gs-3955	416	1	methusaleh	methusaleh	PROPN
gs-3955	416	2	’s	’s	PART
gs-3955	416	3	zoo	zoo	NOUN
gs-3955	416	4	:	:	PUNCT
gs-3955	416	5	how	how	SCONJ
gs-3955	416	6	nature	nature	NOUN
gs-3955	416	7	provides	provide	VERB
gs-3955	416	8	us	we	PRON
gs-3955	416	9	with	with	ADP
gs-3955	416	10	clues	clue	NOUN
gs-3955	416	11	for	for	ADP
gs-3955	416	12	extending	extend	VERB
gs-3955	416	13	human	human	ADJ
gs-3955	416	14	health	health	NOUN
gs-3955	416	15	span	span	NOUN
gs-3955	416	16	.	.	PUNCT
gs-3955	417	1	journal	journal	PROPN
gs-3955	417	2	of	of	ADP
gs-3955	417	3	comparative	comparative	ADJ
gs-3955	417	4	pathology	pathology	NOUN
gs-3955	417	5	,	,	PUNCT
gs-3955	417	6	142(1	142(1	NUM
gs-3955	417	7	)	)	PUNCT
gs-3955	417	8	,	,	PUNCT
gs-3955	417	9	s10	s10	NOUN
gs-3955	417	10	-	-	PUNCT
gs-3955	417	11	s21	s21	NOUN
gs-3955	417	12	.	.	PUNCT
gs-3955	418	1	[	[	X
gs-3955	418	2	pubmed	pubmed	X
gs-3955	418	3	:	:	PUNCT
gs-3955	418	4	19959168	19959168	NUM
gs-3955	418	5	]	]	SYM
gs-3955	418	6	3	3	X
gs-3955	418	7	.	.	NOUN
gs-3955	418	8	austad	austad	PROPN
gs-3955	418	9	,	,	PUNCT
gs-3955	418	10	s.	s.	PROPN
gs-3955	418	11	n.	n.	PROPN
gs-3955	418	12	(	(	PUNCT
gs-3955	418	13	2022	2022	NUM
gs-3955	418	14	)	)	PUNCT
gs-3955	418	15	.	.	PUNCT
gs-3955	419	1	the	the	DET
gs-3955	419	2	comparative	comparative	ADJ
gs-3955	419	3	biology	biology	NOUN
gs-3955	419	4	of	of	ADP
gs-3955	419	5	longevity	longevity	NOUN
gs-3955	419	6	:	:	PUNCT
gs-3955	419	7	from	from	ADP
gs-3955	419	8	mice	mouse	NOUN
gs-3955	419	9	to	to	ADP
gs-3955	419	10	whales	whale	NOUN
gs-3955	419	11	.	.	PUNCT
gs-3955	420	1	journals	journal	NOUN
gs-3955	420	2	of	of	ADP
gs-3955	420	3	gerontology	gerontology	NOUN
gs-3955	420	4	,	,	PUNCT
gs-3955	420	5	77(1	77(1	NUM
gs-3955	420	6	)	)	PUNCT
gs-3955	420	7	,	,	PUNCT
gs-3955	420	8	e1	e1	NOUN
gs-3955	420	9	-	-	PUNCT
gs-3955	420	10	e9	e9	PROPN
gs-3955	420	11	.	.	PUNCT
gs-3955	421	1	[	[	X
gs-3955	421	2	pubmed	pubmed	NOUN
gs-3955	421	3	:	:	PUNCT
gs-3955	421	4	34788821	34788821	NUM
gs-3955	421	5	]	]	PUNCT
gs-3955	421	6	4	4	X
gs-3955	421	7	.	.	X
gs-3955	421	8	azpurua	azpurua	PROPN
gs-3955	421	9	,	,	PUNCT
gs-3955	421	10	j.	j.	PROPN
gs-3955	421	11	,	,	PUNCT
gs-3955	421	12	et	et	PROPN
gs-3955	421	13	al	al	PROPN
gs-3955	421	14	.	.	PROPN
gs-3955	421	15	(	(	PUNCT
gs-3955	421	16	2013	2013	NUM
gs-3955	421	17	)	)	PUNCT
gs-3955	421	18	.	.	PUNCT
gs-3955	422	1	naked	naked	ADJ
gs-3955	422	2	mole	mole	NOUN
gs-3955	422	3	-	-	PUNCT
gs-3955	422	4	rat	rat	NOUN
gs-3955	422	5	has	have	AUX
gs-3955	422	6	increased	increase	VERB
gs-3955	422	7	translational	translational	ADJ
gs-3955	422	8	fidelity	fidelity	NOUN
gs-3955	422	9	compared	compare	VERB
gs-3955	422	10	with	with	ADP
gs-3955	422	11	the	the	DET
gs-3955	422	12	mouse	mouse	NOUN
gs-3955	422	13	,	,	PUNCT
gs-3955	422	14	as	as	ADV
gs-3955	422	15	well	well	ADV
gs-3955	422	16	as	as	ADP
gs-3955	422	17	a	a	DET
gs-3955	422	18	unique	unique	ADJ
gs-3955	422	19	28s	28	NOUN
gs-3955	422	20	ribosomal	ribosomal	NOUN
gs-3955	422	21	rna	rna	NOUN
gs-3955	422	22	cleavage	cleavage	NOUN
gs-3955	422	23	.	.	PUNCT
gs-3955	423	1	pnas	pnas	PROPN
gs-3955	423	2	,	,	PUNCT
gs-3955	423	3	110(43	110(43	NUM
gs-3955	423	4	)	)	PUNCT
gs-3955	423	5	,	,	PUNCT
gs-3955	423	6	17350	17350	NUM
gs-3955	423	7	-	-	SYM
gs-3955	423	8	17355	17355	NUM
gs-3955	423	9	.	.	PUNCT
gs-3955	424	1	[	[	X
gs-3955	424	2	pubmed	pubmed	X
gs-3955	424	3	:	:	PUNCT
gs-3955	424	4	24101461	24101461	NUM
gs-3955	424	5	]	]	PUNCT
gs-3955	424	6	5	5	X
gs-3955	424	7	.	.	PUNCT
gs-3955	424	8	ballantyne	ballantyne	PROPN
gs-3955	424	9	,	,	PUNCT
gs-3955	424	10	j.	j.	PROPN
gs-3955	424	11	s.	s.	PROPN
gs-3955	424	12	,	,	PUNCT
gs-3955	424	13	et	et	PROPN
gs-3955	424	14	al	al	PROPN
gs-3955	424	15	.	.	PUNCT
gs-3955	424	16	(	(	PUNCT
gs-3955	424	17	2021	2021	NUM
gs-3955	424	18	)	)	PUNCT
gs-3955	424	19	.	.	PUNCT
gs-3955	425	1	membrane	membrane	NOUN
gs-3955	425	2	lipid	lipid	NOUN
gs-3955	425	3	adaptations	adaptation	NOUN
gs-3955	425	4	in	in	ADP
gs-3955	425	5	arctic	arctic	PROPN
gs-3955	425	6	sharks	sharks	PROPN
gs-3955	425	7	.	.	PUNCT
gs-3955	426	1	comparative	comparative	ADJ
gs-3955	426	2	biochemistry	biochemistry	NOUN
gs-3955	426	3	and	and	CCONJ
gs-3955	426	4	physiology	physiology	NOUN
gs-3955	426	5	,	,	PUNCT
gs-3955	426	6	256	256	NUM
gs-3955	426	7	,	,	PUNCT
gs-3955	426	8	110932	110932	NUM
gs-3955	426	9	.	.	PUNCT
gs-3955	427	1	[	[	X
gs-3955	427	2	pubmed	pubmed	X
gs-3955	427	3	:	:	PUNCT
gs-3955	427	4	34363902	34363902	NUM
gs-3955	427	5	]	]	PUNCT
gs-3955	427	6	6	6	X
gs-3955	427	7	.	.	X
gs-3955	427	8	ballesta	ballesta	NOUN
gs-3955	427	9	-	-	PUNCT
gs-3955	427	10	artero	artero	NOUN
gs-3955	427	11	,	,	PUNCT
gs-3955	427	12	i.	i.	PROPN
gs-3955	427	13	,	,	PUNCT
gs-3955	427	14	et	et	PROPN
gs-3955	427	15	al	al	PROPN
gs-3955	427	16	.	.	PUNCT
gs-3955	427	17	(	(	PUNCT
gs-3955	427	18	2017	2017	NUM
gs-3955	427	19	)	)	PUNCT
gs-3955	427	20	.	.	PUNCT
gs-3955	428	1	metabolic	metabolic	NOUN
gs-3955	428	2	rate	rate	NOUN
gs-3955	428	3	depression	depression	NOUN
gs-3955	428	4	in	in	ADP
gs-3955	428	5	arctica	arctica	PROPN
gs-3955	428	6	islandica	islandica	PROPN
gs-3955	428	7	.	.	PUNCT
gs-3955	429	1	journal	journal	PROPN
gs-3955	429	2	of	of	ADP
gs-3955	429	3	experimental	experimental	ADJ
gs-3955	429	4	biology	biology	NOUN
gs-3955	429	5	,	,	PUNCT
gs-3955	429	6	220(pt	220(pt	NUM
gs-3955	429	7	11	11	NUM
gs-3955	429	8	)	)	PUNCT
gs-3955	429	9	,	,	PUNCT
gs-3955	429	10	1972	1972	NUM
gs-3955	429	11	-	-	SYM
gs-3955	429	12	1979	1979	NUM
gs-3955	429	13	.	.	PUNCT
gs-3955	430	1	[	[	X
gs-3955	430	2	pubmed	pubmed	X
gs-3955	430	3	:	:	PUNCT
gs-3955	430	4	28356366	28356366	NUM
gs-3955	430	5	]	]	PUNCT
gs-3955	430	6	7	7	X
gs-3955	430	7	.	.	X
gs-3955	430	8	berkeley	berkeley	PROPN
gs-3955	430	9	,	,	PUNCT
gs-3955	430	10	s.a	s.a	PROPN
gs-3955	430	11	.	.	PROPN
gs-3955	430	12	,	,	PUNCT
gs-3955	430	13	et	et	PROPN
gs-3955	430	14	al	al	PROPN
gs-3955	430	15	.	.	PROPN
gs-3955	430	16	(	(	PUNCT
gs-3955	430	17	2004	2004	NUM
gs-3955	430	18	)	)	PUNCT
gs-3955	430	19	.	.	PUNCT
gs-3955	431	1	maternal	maternal	ADJ
gs-3955	431	2	age	age	NOUN
gs-3955	431	3	as	as	ADP
gs-3955	431	4	a	a	DET
gs-3955	431	5	determinant	determinant	NOUN
gs-3955	431	6	of	of	ADP
gs-3955	431	7	larval	larval	ADJ
gs-3955	431	8	growth	growth	NOUN
gs-3955	431	9	and	and	CCONJ
gs-3955	431	10	survival	survival	NOUN
gs-3955	431	11	in	in	ADP
gs-3955	431	12	a	a	DET
gs-3955	431	13	marine	marine	ADJ
gs-3955	431	14	fish	fish	NOUN
gs-3955	431	15	,	,	PUNCT
gs-3955	431	16	sebastes	sebaste	VERB
gs-3955	431	17	melanops	melanop	NOUN
gs-3955	431	18	.	.	PUNCT
gs-3955	432	1	ecology	ecology	NOUN
gs-3955	432	2	,	,	PUNCT
gs-3955	432	3	85(5	85(5	NUM
gs-3955	432	4	)	)	PUNCT
gs-3955	432	5	,	,	PUNCT
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gs-3955	432	7	-	-	SYM
gs-3955	432	8	1264	1264	NUM
gs-3955	432	9	.	.	PUNCT
gs-3955	433	1	[	[	X
gs-3955	433	2	pubmed	pubmed	NOUN
gs-3955	433	3	:	:	PUNCT
gs-3955	433	4	15233451	15233451	NUM
gs-3955	433	5	]	]	PUNCT
gs-3955	433	6	8	8	X
gs-3955	433	7	.	.	PUNCT
gs-3955	433	8	bizjak	bizjak	PROPN
gs-3955	433	9	mali	mali	PROPN
gs-3955	433	10	,	,	PUNCT
gs-3955	433	11	l.	l.	PROPN
gs-3955	433	12	,	,	PUNCT
gs-3955	433	13	et	et	PROPN
gs-3955	433	14	al	al	PROPN
gs-3955	433	15	.	.	PROPN
gs-3955	433	16	(	(	PUNCT
gs-3955	433	17	2019	2019	NUM
gs-3955	433	18	)	)	PUNCT
gs-3955	433	19	.	.	PUNCT
gs-3955	434	1	proliferation	proliferation	NOUN
gs-3955	434	2	zones	zone	NOUN
gs-3955	434	3	in	in	ADP
gs-3955	434	4	the	the	DET
gs-3955	434	5	axolotl	axolotl	NOUN
gs-3955	434	6	brain	brain	NOUN
gs-3955	434	7	and	and	CCONJ
gs-3955	434	8	regeneration	regeneration	NOUN
gs-3955	434	9	of	of	ADP
gs-3955	434	10	the	the	DET
gs-3955	434	11	telencephalon	telencephalon	NOUN
gs-3955	434	12	.	.	PUNCT
gs-3955	435	1	neural	neural	ADJ
gs-3955	435	2	development	development	PROPN
gs-3955	435	3	,	,	PUNCT
gs-3955	435	4	14(1	14(1	NUM
gs-3955	435	5	)	)	PUNCT
gs-3955	435	6	,	,	PUNCT
gs-3955	435	7	12	12	NUM
gs-3955	435	8	.	.	PUNCT
gs-3955	436	1	[	[	X
gs-3955	436	2	pubmed	pubmed	X
gs-3955	436	3	:	:	PUNCT
gs-3955	436	4	31409372	31409372	NUM
gs-3955	436	5	]	]	PUNCT
gs-3955	436	6	9	9	NUM
gs-3955	436	7	.	.	PUNCT
gs-3955	436	8	bodnar	bodnar	PROPN
gs-3955	436	9	,	,	PUNCT
gs-3955	436	10	a.g	a.g	PROPN
gs-3955	436	11	.	.	PROPN
gs-3955	436	12	(	(	PUNCT
gs-3955	436	13	2013	2013	NUM
gs-3955	436	14	)	)	PUNCT
gs-3955	436	15	.	.	PUNCT
gs-3955	437	1	cellular	cellular	ADJ
gs-3955	437	2	and	and	CCONJ
gs-3955	437	3	molecular	molecular	ADJ
gs-3955	437	4	mechanisms	mechanism	NOUN
gs-3955	437	5	of	of	ADP
gs-3955	437	6	negligible	negligible	ADJ
gs-3955	437	7	senescence	senescence	NOUN
gs-3955	437	8	.	.	PUNCT
gs-3955	438	1	experimental	experimental	ADJ
gs-3955	438	2	gerontology	gerontology	NOUN
gs-3955	438	3	,	,	PUNCT
gs-3955	438	4	48(10	48(10	NUM
gs-3955	438	5	)	)	PUNCT
gs-3955	438	6	,	,	PUNCT
gs-3955	438	7	1095	1095	NUM
gs-3955	438	8	-	-	SYM
gs-3955	438	9	1099	1099	NUM
gs-3955	438	10	.	.	PUNCT
gs-3955	439	1	[	[	X
gs-3955	439	2	pubmed	pubmed	X
gs-3955	439	3	:	:	PUNCT
gs-3955	439	4	23994581	23994581	NUM
gs-3955	439	5	]	]	PUNCT
gs-3955	439	6	10	10	NUM
gs-3955	439	7	.	.	PUNCT
gs-3955	440	1	bowen	bowen	PROPN
gs-3955	440	2	,	,	PUNCT
gs-3955	440	3	b.w	b.w	PROPN
gs-3955	440	4	.	.	PROPN
gs-3955	440	5	,	,	PUNCT
gs-3955	440	6	et	et	PROPN
gs-3955	440	7	al	al	PROPN
gs-3955	440	8	.	.	PROPN
gs-3955	440	9	(	(	PUNCT
gs-3955	440	10	2021	2021	NUM
gs-3955	440	11	)	)	PUNCT
gs-3955	440	12	.	.	PUNCT
gs-3955	441	1	conservation	conservation	NOUN
gs-3955	441	2	genetics	genetic	NOUN
gs-3955	441	3	of	of	ADP
gs-3955	441	4	the	the	DET
gs-3955	441	5	genus	genus	NOUN
gs-3955	441	6	sebastes	sebaste	NOUN
gs-3955	441	7	.	.	PUNCT
gs-3955	442	1	journal	journal	NOUN
gs-3955	442	2	of	of	ADP
gs-3955	442	3	fish	fish	NOUN
gs-3955	442	4	biology	biology	NOUN
gs-3955	442	5	,	,	PUNCT
gs-3955	442	6	98(3	98(3	PROPN
gs-3955	442	7	)	)	PUNCT
gs-3955	442	8	,	,	PUNCT
gs-3955	442	9	639	639	NUM
gs-3955	442	10	-	-	SYM
gs-3955	442	11	655	655	NUM
gs-3955	442	12	.	.	PUNCT
gs-3955	443	1	[	[	X
gs-3955	443	2	pubmed	pubmed	X
gs-3955	443	3	:	:	PUNCT
gs-3955	443	4	33332605	33332605	NUM
gs-3955	443	5	]	]	PUNCT
gs-3955	444	1	11	11	NUM
gs-3955	444	2	.	.	PUNCT
gs-3955	445	1	brunet	brunet	NOUN
gs-3955	445	2	,	,	PUNCT
gs-3955	445	3	a.	a.	PROPN
gs-3955	445	4	,	,	PUNCT
gs-3955	445	5	et	et	PROPN
gs-3955	445	6	al	al	PROPN
gs-3955	445	7	.	.	PROPN
gs-3955	445	8	(	(	PUNCT
gs-3955	445	9	2022	2022	NUM
gs-3955	445	10	)	)	PUNCT
gs-3955	445	11	.	.	PUNCT
gs-3955	446	1	comparative	comparative	ADJ
gs-3955	446	2	genomics	genomic	NOUN
gs-3955	446	3	of	of	ADP
gs-3955	446	4	longevity	longevity	NOUN
gs-3955	446	5	in	in	ADP
gs-3955	446	6	fishes	fish	NOUN
gs-3955	446	7	.	.	PUNCT
gs-3955	447	1	nature	nature	NOUN
gs-3955	447	2	genetics	genetics	PROPN
gs-3955	447	3	,	,	PUNCT
gs-3955	447	4	54(2	54(2	NUM
gs-3955	447	5	)	)	PUNCT
gs-3955	447	6	,	,	PUNCT
gs-3955	447	7	123	123	NUM
gs-3955	447	8	-	-	SYM
gs-3955	447	9	135	135	NUM
gs-3955	447	10	.	.	PUNCT
gs-3955	448	1	[	[	X
gs-3955	448	2	pubmed	pubmed	X
gs-3955	448	3	:	:	PUNCT
gs-3955	448	4	35027735	35027735	NUM
gs-3955	448	5	]	]	PUNCT
gs-3955	448	6	12	12	NUM
gs-3955	448	7	.	.	PUNCT
gs-3955	449	1	brunet	brunet	NOUN
gs-3955	449	2	-	-	PUNCT
gs-3955	449	3	rossinni	rossinni	NOUN
gs-3955	449	4	,	,	PUNCT
gs-3955	449	5	a.k	a.k	PROPN
gs-3955	449	6	.	.	PROPN
gs-3955	449	7	,	,	PUNCT
gs-3955	449	8	&	&	CCONJ
gs-3955	449	9	austad	austad	PROPN
gs-3955	449	10	,	,	PUNCT
gs-3955	449	11	s.n	s.n	PROPN
gs-3955	449	12	.	.	PROPN
gs-3955	449	13	(	(	PUNCT
gs-3955	449	14	2022	2022	NUM
gs-3955	449	15	)	)	PUNCT
gs-3955	449	16	.	.	PUNCT
gs-3955	450	1	ageing	age	VERB
gs-3955	450	2	studies	study	NOUN
gs-3955	450	3	on	on	ADP
gs-3955	450	4	bats	bat	NOUN
gs-3955	450	5	and	and	CCONJ
gs-3955	450	6	turtles	turtle	NOUN
gs-3955	450	7	:	:	PUNCT
gs-3955	450	8	what	what	PRON
gs-3955	450	9	they	they	PRON
gs-3955	450	10	can	can	AUX
gs-3955	450	11	tell	tell	VERB
gs-3955	450	12	us	we	PRON
gs-3955	450	13	about	about	ADP
gs-3955	450	14	the	the	DET
gs-3955	450	15	biology	biology	NOUN
gs-3955	450	16	of	of	ADP
gs-3955	450	17	ageing	ageing	NOUN
gs-3955	450	18	.	.	PUNCT
gs-3955	451	1	experimental	experimental	ADJ
gs-3955	451	2	gerontology	gerontology	NOUN
gs-3955	451	3	,	,	PUNCT
gs-3955	451	4	159	159	NUM
gs-3955	451	5	,	,	PUNCT
gs-3955	451	6	111675	111675	NUM
gs-3955	451	7	.	.	PUNCT
gs-3955	452	1	[	[	X
gs-3955	452	2	pubmed	pubmed	X
gs-3955	452	3	:	:	PUNCT
gs-3955	452	4	35031374	35031374	NUM
gs-3955	452	5	]	]	PUNCT
gs-3955	452	6	13	13	NUM
gs-3955	452	7	.	.	PUNCT
gs-3955	453	1	buffenstein	buffenstein	ADV
gs-3955	453	2	,	,	PUNCT
gs-3955	453	3	r.	r.	PROPN
gs-3955	453	4	(	(	PUNCT
gs-3955	453	5	2005	2005	NUM
gs-3955	453	6	)	)	PUNCT
gs-3955	453	7	.	.	PUNCT
gs-3955	454	1	the	the	DET
gs-3955	454	2	naked	naked	ADJ
gs-3955	454	3	mole	mole	NOUN
gs-3955	454	4	-	-	PUNCT
gs-3955	454	5	rat	rat	NOUN
gs-3955	454	6	:	:	PUNCT
gs-3955	454	7	a	a	DET
gs-3955	454	8	new	new	ADJ
gs-3955	454	9	long	long	ADV
gs-3955	454	10	-	-	PUNCT
gs-3955	454	11	living	live	VERB
gs-3955	454	12	model	model	NOUN
gs-3955	454	13	for	for	ADP
gs-3955	454	14	human	human	ADJ
gs-3955	454	15	aging	age	VERB
gs-3955	454	16	research	research	NOUN
gs-3955	454	17	.	.	PUNCT
gs-3955	455	1	journals	journal	NOUN
gs-3955	455	2	of	of	ADP
gs-3955	455	3	gerontology	gerontology	NOUN
gs-3955	455	4	,	,	PUNCT
gs-3955	455	5	60(11	60(11	NUM
gs-3955	455	6	)	)	PUNCT
gs-3955	455	7	,	,	PUNCT
gs-3955	455	8	1369	1369	NUM
gs-3955	455	9	-	-	SYM
gs-3955	455	10	1377	1377	NUM
gs-3955	455	11	.	.	PUNCT
gs-3955	456	1	[	[	X
gs-3955	456	2	pubmed	pubmed	NOUN
gs-3955	456	3	:	:	PUNCT
gs-3955	456	4	16339321	16339321	NUM
gs-3955	456	5	]	]	PUNCT
gs-3955	456	6	14	14	NUM
gs-3955	456	7	.	.	PUNCT
gs-3955	457	1	buffenstein	buffenstein	ADV
gs-3955	457	2	,	,	PUNCT
gs-3955	457	3	r.	r.	PROPN
gs-3955	457	4	(	(	PUNCT
gs-3955	457	5	2008	2008	NUM
gs-3955	457	6	)	)	PUNCT
gs-3955	457	7	.	.	PUNCT
gs-3955	458	1	negligible	negligible	ADJ
gs-3955	458	2	senescence	senescence	NOUN
gs-3955	458	3	in	in	ADP
gs-3955	458	4	the	the	DET
gs-3955	458	5	longest	long	ADJ
gs-3955	458	6	living	live	VERB
gs-3955	458	7	rodent	rodent	NOUN
gs-3955	458	8	,	,	PUNCT
gs-3955	458	9	the	the	DET
gs-3955	458	10	naked	naked	ADJ
gs-3955	458	11	mole	mole	NOUN
gs-3955	458	12	-	-	PUNCT
gs-3955	458	13	rat	rat	NOUN
gs-3955	458	14	:	:	PUNCT
gs-3955	458	15	insights	insight	NOUN
gs-3955	458	16	from	from	ADP
gs-3955	458	17	a	a	DET
gs-3955	458	18	successfully	successfully	ADV
gs-3955	458	19	aging	age	VERB
gs-3955	458	20	species	specie	NOUN
gs-3955	458	21	.	.	PUNCT
gs-3955	459	1	journal	journal	PROPN
gs-3955	459	2	of	of	ADP
gs-3955	459	3	comparative	comparative	ADJ
gs-3955	459	4	physiology	physiology	NOUN
gs-3955	459	5	b	b	NOUN
gs-3955	459	6	,	,	PUNCT
gs-3955	459	7	178(4	178(4	NUM
gs-3955	459	8	)	)	PUNCT
gs-3955	459	9	,	,	PUNCT
gs-3955	459	10	439	439	NUM
gs-3955	459	11	-	-	SYM
gs-3955	459	12	445	445	NUM
gs-3955	459	13	.	.	PUNCT
gs-3955	460	1	[	[	X
gs-3955	460	2	pubmed	pubmed	NOUN
gs-3955	460	3	:	:	PUNCT
gs-3955	460	4	18180931	18180931	NUM
gs-3955	460	5	]	]	PUNCT
gs-3955	460	6	georgian	georgian	PROPN
gs-3955	460	7	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	460	8	მეცნიერები	მეცნიერები	NOUN
gs-3955	460	9	ტ.	ტ.	VERB
gs-3955	460	10	7	7	NUM
gs-3955	460	11	n	n	DET
gs-3955	460	12	3	3	NUM
gs-3955	460	13	,	,	PUNCT
gs-3955	460	14	2025	2025	NUM
gs-3955	460	15	390	390	NUM
gs-3955	460	16	15	15	NUM
gs-3955	460	17	.	.	PUNCT
gs-3955	461	1	bulog	bulog	PROPN
gs-3955	461	2	,	,	PUNCT
gs-3955	461	3	b.	b.	PROPN
gs-3955	461	4	,	,	PUNCT
gs-3955	461	5	et	et	PROPN
gs-3955	461	6	al	al	PROPN
gs-3955	461	7	.	.	PROPN
gs-3955	461	8	(	(	PUNCT
gs-3955	461	9	2020	2020	NUM
gs-3955	461	10	)	)	PUNCT
gs-3955	461	11	.	.	PUNCT
gs-3955	462	1	proteus	proteus	NOUN
gs-3955	462	2	anguinus	anguinus	PROPN
gs-3955	462	3	–	–	PUNCT
gs-3955	462	4	overview	overview	NOUN
gs-3955	462	5	of	of	ADP
gs-3955	462	6	conservation	conservation	NOUN
gs-3955	462	7	issues	issue	NOUN
gs-3955	462	8	.	.	PUNCT
gs-3955	463	1	biological	biological	ADJ
gs-3955	463	2	conservation	conservation	NOUN
gs-3955	463	3	,	,	PUNCT
gs-3955	463	4	243	243	NUM
gs-3955	463	5	,	,	PUNCT
gs-3955	463	6	108475	108475	NUM
gs-3955	463	7	.	.	PUNCT
gs-3955	464	1	[	[	X
gs-3955	464	2	pubmed	pubmed	NOUN
gs-3955	464	3	:	:	PUNCT
gs-3955	464	4	32313341	32313341	NUM
gs-3955	464	5	]	]	PUNCT
gs-3955	464	6	16	16	NUM
gs-3955	464	7	.	.	PUNCT
gs-3955	464	8	butler	butler	PROPN
gs-3955	464	9	,	,	PUNCT
gs-3955	464	10	p.g	p.g	PROPN
gs-3955	464	11	.	.	PROPN
gs-3955	464	12	,	,	PUNCT
gs-3955	464	13	et	et	PROPN
gs-3955	464	14	al	al	PROPN
gs-3955	464	15	.	.	PROPN
gs-3955	464	16	(	(	PUNCT
gs-3955	464	17	2013	2013	NUM
gs-3955	464	18	)	)	PUNCT
gs-3955	464	19	.	.	PUNCT
gs-3955	465	1	variability	variability	NOUN
gs-3955	465	2	of	of	ADP
gs-3955	465	3	marine	marine	ADJ
gs-3955	465	4	climate	climate	NOUN
gs-3955	465	5	on	on	ADP
gs-3955	465	6	the	the	DET
gs-3955	465	7	north	north	NOUN
gs-3955	465	8	icelandic	icelandic	ADJ
gs-3955	465	9	shelf	shelf	NOUN
gs-3955	465	10	.	.	PUNCT
gs-3955	466	1	paleoceanography	paleoceanography	NOUN
gs-3955	466	2	,	,	PUNCT
gs-3955	466	3	28(2	28(2	NUM
gs-3955	466	4	)	)	PUNCT
gs-3955	466	5	,	,	PUNCT
gs-3955	466	6	185	185	NUM
gs-3955	466	7	-	-	SYM
gs-3955	466	8	199	199	NUM
gs-3955	466	9	.	.	PUNCT
gs-3955	467	1	[	[	X
gs-3955	467	2	pubmed	pubmed	X
gs-3955	467	3	:	:	PUNCT
gs-3955	467	4	25844058	25844058	NUM
gs-3955	467	5	]	]	PUNCT
gs-3955	467	6	17	17	NUM
gs-3955	467	7	.	.	PUNCT
gs-3955	467	8	chichinadze	chichinadze	PROPN
gs-3955	467	9	,	,	PUNCT
gs-3955	467	10	k.	k.	PROPN
gs-3955	467	11	,	,	PUNCT
gs-3955	467	12	lazarashvili	lazarashvili	PROPN
gs-3955	467	13	,	,	PUNCT
gs-3955	467	14	a.	a.	PROPN
gs-3955	467	15	,	,	PUNCT
gs-3955	467	16	&	&	CCONJ
gs-3955	467	17	tkemaladze	tkemaladze	PROPN
gs-3955	467	18	,	,	PUNCT
gs-3955	467	19	j.	j.	PROPN
gs-3955	467	20	(	(	PUNCT
gs-3955	467	21	2013	2013	NUM
gs-3955	467	22	)	)	PUNCT
gs-3955	467	23	.	.	PUNCT
gs-3955	468	1	rna	rna	NOUN
gs-3955	468	2	in	in	ADP
gs-3955	468	3	centrosomes	centrosome	NOUN
gs-3955	468	4	:	:	PUNCT
gs-3955	468	5	structure	structure	NOUN
gs-3955	468	6	and	and	CCONJ
gs-3955	468	7	possible	possible	ADJ
gs-3955	468	8	functions	function	NOUN
gs-3955	468	9	.	.	PUNCT
gs-3955	469	1	protoplasma	protoplasma	NOUN
gs-3955	469	2	,	,	PUNCT
gs-3955	469	3	250(1	250(1	NUM
gs-3955	469	4	)	)	PUNCT
gs-3955	469	5	,	,	PUNCT
gs-3955	469	6	397	397	NUM
gs-3955	469	7	-	-	SYM
gs-3955	469	8	405	405	NUM
gs-3955	469	9	.	.	PUNCT
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gs-3955	470	2	.	.	PUNCT
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gs-3955	470	4	,	,	PUNCT
gs-3955	470	5	j.d	j.d	PROPN
gs-3955	470	6	.	.	PROPN
gs-3955	470	7	,	,	PUNCT
gs-3955	470	8	et	et	PROPN
gs-3955	470	9	al	al	PROPN
gs-3955	470	10	.	.	PROPN
gs-3955	470	11	(	(	PUNCT
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gs-3955	470	13	)	)	PUNCT
gs-3955	470	14	.	.	PUNCT
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gs-3955	471	2	hypotheses	hypothesis	NOUN
gs-3955	471	3	of	of	ADP
gs-3955	471	4	aging	aging	NOUN
gs-3955	471	5	in	in	ADP
gs-3955	471	6	long	long	ADV
gs-3955	471	7	-	-	PUNCT
gs-3955	471	8	lived	live	VERB
gs-3955	471	9	painted	paint	VERB
gs-3955	471	10	turtles	turtle	NOUN
gs-3955	471	11	(	(	PUNCT
gs-3955	471	12	chrysemys	chrysemys	NOUN
gs-3955	471	13	picta	picta	PROPN
gs-3955	471	14	)	)	PUNCT
gs-3955	471	15	.	.	PUNCT
gs-3955	472	1	experimental	experimental	ADJ
gs-3955	472	2	gerontology	gerontology	NOUN
gs-3955	472	3	,	,	PUNCT
gs-3955	472	4	120	120	NUM
gs-3955	472	5	,	,	PUNCT
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gs-3955	472	7	-	-	SYM
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gs-3955	472	9	.	.	PUNCT
gs-3955	473	1	[	[	X
gs-3955	473	2	pubmed	pubmed	X
gs-3955	473	3	:	:	PUNCT
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gs-3955	473	5	]	]	PUNCT
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gs-3955	473	7	.	.	PUNCT
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gs-3955	473	10	,	,	PUNCT
gs-3955	473	11	r.	r.	PROPN
gs-3955	473	12	,	,	PUNCT
gs-3955	473	13	et	et	PROPN
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gs-3955	473	16	(	(	PUNCT
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gs-3955	473	18	)	)	PUNCT
gs-3955	473	19	.	.	PUNCT
gs-3955	474	1	the	the	DET
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gs-3955	474	5	in	in	ADP
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gs-3955	474	7	.	.	PUNCT
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gs-3955	475	5	)	)	PUNCT
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gs-3955	475	8	-	-	SYM
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gs-3955	475	10	.	.	PUNCT
gs-3955	476	1	[	[	X
gs-3955	476	2	pubmed	pubmed	NOUN
gs-3955	476	3	:	:	PUNCT
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gs-3955	476	5	]	]	PUNCT
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gs-3955	476	7	.	.	PUNCT
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gs-3955	476	9	,	,	PUNCT
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gs-3955	476	11	,	,	PUNCT
gs-3955	476	12	et	et	PROPN
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gs-3955	476	15	(	(	PUNCT
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gs-3955	476	17	)	)	PUNCT
gs-3955	476	18	.	.	PUNCT
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gs-3955	477	4	a	a	DET
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gs-3955	477	6	stimulus	stimulus	NOUN
gs-3955	477	7	in	in	ADP
gs-3955	477	8	the	the	DET
gs-3955	477	9	naked	naked	ADJ
gs-3955	477	10	mole	mole	NOUN
gs-3955	477	11	-	-	PUNCT
gs-3955	477	12	rat	rat	NOUN
gs-3955	477	13	.	.	PROPN
gs-3955	477	14	journal	journal	PROPN
gs-3955	477	15	of	of	ADP
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gs-3955	477	17	biology	biology	NOUN
gs-3955	477	18	,	,	PUNCT
gs-3955	477	19	214(24	214(24	NUM
gs-3955	477	20	)	)	PUNCT
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gs-3955	477	23	-	-	SYM
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gs-3955	477	25	.	.	PUNCT
gs-3955	478	1	[	[	X
gs-3955	478	2	pubmed	pubmed	NOUN
gs-3955	478	3	:	:	PUNCT
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gs-3955	478	5	]	]	PUNCT
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gs-3955	478	7	.	.	PUNCT
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gs-3955	478	11	j.	j.	PROPN
gs-3955	478	12	p.	p.	PROPN
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gs-3955	478	14	et	et	PROPN
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gs-3955	478	17	(	(	PUNCT
gs-3955	478	18	2017	2017	NUM
gs-3955	478	19	)	)	PUNCT
gs-3955	478	20	.	.	PUNCT
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gs-3955	479	4	the	the	DET
gs-3955	479	5	greenland	greenland	NOUN
gs-3955	479	6	shark	shark	NOUN
gs-3955	479	7	:	:	PUNCT
gs-3955	479	8	insights	insight	NOUN
gs-3955	479	9	into	into	ADP
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gs-3955	479	11	.	.	PUNCT
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gs-3955	480	3	,	,	PUNCT
gs-3955	480	4	3(6	3(6	NUM
gs-3955	480	5	)	)	PUNCT
gs-3955	480	6	,	,	PUNCT
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gs-3955	480	8	.	.	PUNCT
gs-3955	481	1	[	[	X
gs-3955	481	2	pubmed	pubmed	X
gs-3955	481	3	:	:	PUNCT
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gs-3955	481	5	]	]	PUNCT
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gs-3955	481	7	.	.	PUNCT
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gs-3955	481	10	,	,	PUNCT
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gs-3955	481	15	)	)	PUNCT
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gs-3955	482	1	the	the	DET
gs-3955	482	2	evolution	evolution	NOUN
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gs-3955	482	5	in	in	ADP
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gs-3955	482	7	-	-	PUNCT
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gs-3955	482	9	turtles	turtle	NOUN
gs-3955	482	10	.	.	PUNCT
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gs-3955	483	5	,	,	PUNCT
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gs-3955	483	7	)	)	PUNCT
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gs-3955	483	10	.	.	PUNCT
gs-3955	484	1	[	[	X
gs-3955	484	2	pubmed	pubmed	X
gs-3955	484	3	:	:	PUNCT
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gs-3955	484	5	]	]	PUNCT
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gs-3955	484	7	.	.	PUNCT
gs-3955	485	1	de	de	PROPN
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gs-3955	485	3	,	,	PUNCT
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gs-3955	485	5	.	.	PROPN
gs-3955	485	6	(	(	PUNCT
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gs-3955	485	8	)	)	PUNCT
gs-3955	485	9	.	.	PUNCT
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gs-3955	487	7	.	.	PUNCT
gs-3955	488	1	[	[	X
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gs-3955	488	3	:	:	PUNCT
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gs-3955	488	7	.	.	PUNCT
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gs-3955	488	9	,	,	PUNCT
gs-3955	488	10	p.	p.	PROPN
gs-3955	488	11	,	,	PUNCT
gs-3955	488	12	&	&	CCONJ
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gs-3955	488	18	)	)	PUNCT
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gs-3955	490	7	,	,	PUNCT
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gs-3955	490	9	-	-	SYM
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gs-3955	490	11	.	.	PUNCT
gs-3955	491	1	[	[	X
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gs-3955	491	3	:	:	PUNCT
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gs-3955	491	7	.	.	PUNCT
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gs-3955	492	2	,	,	PUNCT
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gs-3955	492	4	.	.	PROPN
gs-3955	492	5	,	,	PUNCT
gs-3955	492	6	&	&	CCONJ
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gs-3955	492	8	,	,	PUNCT
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gs-3955	492	11	(	(	PUNCT
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gs-3955	493	9	.	.	PUNCT
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gs-3955	494	3	,	,	PUNCT
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gs-3955	494	5	)	)	PUNCT
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gs-3955	494	8	-	-	SYM
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gs-3955	494	10	.	.	PUNCT
gs-3955	495	1	[	[	X
gs-3955	495	2	pubmed	pubmed	NOUN
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gs-3955	495	7	.	.	PUNCT
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gs-3955	495	18	(	(	PUNCT
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gs-3955	495	20	)	)	PUNCT
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gs-3955	496	7	.	.	PUNCT
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gs-3955	497	7	,	,	PUNCT
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gs-3955	497	9	-	-	SYM
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gs-3955	497	11	.	.	PUNCT
gs-3955	498	1	[	[	X
gs-3955	498	2	pubmed	pubmed	X
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gs-3955	498	7	.	.	PUNCT
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gs-3955	498	18	)	)	PUNCT
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gs-3955	499	5	:	:	PUNCT
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gs-3955	500	6	)	)	PUNCT
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gs-3955	501	1	[	[	X
gs-3955	501	2	pubmed	pubmed	NOUN
gs-3955	501	3	:	:	PUNCT
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gs-3955	501	5	]	]	X
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gs-3955	502	2	.	.	PUNCT
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gs-3955	502	4	,	,	PUNCT
gs-3955	502	5	c.	c.	PROPN
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gs-3955	502	7	(	(	PUNCT
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gs-3955	502	9	)	)	PUNCT
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gs-3955	503	7	senescence	senescence	NOUN
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gs-3955	503	9	a	a	DET
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gs-3955	504	2	,	,	PUNCT
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gs-3955	504	4	)	)	PUNCT
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gs-3955	504	7	-	-	SYM
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gs-3955	504	9	.	.	PUNCT
gs-3955	505	1	[	[	X
gs-3955	505	2	pubmed	pubmed	X
gs-3955	505	3	:	:	PUNCT
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gs-3955	505	5	]	]	PUNCT
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gs-3955	505	7	.	.	PUNCT
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gs-3955	506	2	,	,	PUNCT
gs-3955	506	3	c.	c.	PROPN
gs-3955	506	4	,	,	PUNCT
gs-3955	506	5	et	et	PROPN
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gs-3955	506	8	(	(	PUNCT
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gs-3955	507	4	-	-	ADJ
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gs-3955	507	8	.	.	PUNCT
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gs-3955	509	1	[	[	X
gs-3955	509	2	pubmed	pubmed	X
gs-3955	509	3	:	:	PUNCT
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gs-3955	509	5	]	]	PUNCT
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gs-3955	510	2	.	.	PUNCT
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gs-3955	510	4	,	,	PUNCT
gs-3955	510	5	w.r	w.r	PROPN
gs-3955	510	6	.	.	PROPN
gs-3955	510	7	,	,	PUNCT
gs-3955	510	8	et	et	PROPN
gs-3955	510	9	al	al	PROPN
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gs-3955	510	13	)	)	PUNCT
gs-3955	510	14	.	.	PUNCT
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gs-3955	511	2	to	to	ADP
gs-3955	511	3	neoplasia	neoplasia	NOUN
gs-3955	511	4	in	in	ADP
gs-3955	511	5	long	long	ADV
gs-3955	511	6	-	-	PUNCT
gs-3955	511	7	lived	live	VERB
gs-3955	511	8	sea	sea	NOUN
gs-3955	511	9	urchins	urchin	NOUN
gs-3955	511	10	.	.	PUNCT
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gs-3955	512	3	,	,	PUNCT
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gs-3955	512	5	)	)	PUNCT
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gs-3955	512	8	.	.	PUNCT
gs-3955	513	1	[	[	X
gs-3955	513	2	pubmed	pubmed	NOUN
gs-3955	513	3	:	:	PUNCT
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gs-3955	513	5	]	]	PUNCT
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gs-3955	513	7	.	.	PUNCT
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gs-3955	514	2	-	-	PUNCT
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gs-3955	514	4	,	,	PUNCT
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gs-3955	514	7	,	,	PUNCT
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gs-3955	515	1	(	(	PUNCT
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gs-3955	515	3	)	)	PUNCT
gs-3955	515	4	.	.	PUNCT
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gs-3955	516	4	.	.	PUNCT
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gs-3955	517	2	in	in	ADP
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gs-3955	517	6	biology	biology	NOUN
gs-3955	517	7	,	,	PUNCT
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gs-3955	517	9	,	,	PUNCT
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gs-3955	518	1	[	[	X
gs-3955	518	2	pubmed	pubmed	X
gs-3955	518	3	:	:	PUNCT
gs-3955	518	4	33968920	33968920	NUM
gs-3955	518	5	]	]	PUNCT
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gs-3955	518	7	.	.	PUNCT
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gs-3955	519	2	,	,	PUNCT
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gs-3955	519	4	,	,	PUNCT
gs-3955	519	5	et	et	PROPN
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gs-3955	520	3	)	)	PUNCT
gs-3955	520	4	.	.	PUNCT
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gs-3955	521	5	-	-	PUNCT
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gs-3955	521	8	.	.	PUNCT
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gs-3955	522	5	,	,	PUNCT
gs-3955	522	6	228	228	NUM
gs-3955	522	7	,	,	PUNCT
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gs-3955	522	9	-	-	SYM
gs-3955	522	10	115	115	NUM
gs-3955	522	11	.	.	PUNCT
gs-3955	523	1	[	[	X
gs-3955	523	2	pubmed	pubmed	X
gs-3955	523	3	:	:	PUNCT
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gs-3955	523	5	]	]	PUNCT
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gs-3955	523	7	.	.	PUNCT
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gs-3955	523	9	,	,	PUNCT
gs-3955	523	10	v.	v.	ADV
gs-3955	523	11	,	,	PUNCT
gs-3955	523	12	et	et	PROPN
gs-3955	523	13	al	al	PROPN
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gs-3955	523	15	(	(	PUNCT
gs-3955	523	16	2014	2014	NUM
gs-3955	523	17	)	)	PUNCT
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gs-3955	524	2	resistance	resistance	NOUN
gs-3955	524	3	in	in	ADP
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gs-3955	524	5	blind	blind	ADJ
gs-3955	524	6	mole	mole	NOUN
gs-3955	524	7	rat	rat	NOUN
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gs-3955	524	10	by	by	ADP
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gs-3955	524	12	necrotic	necrotic	ADJ
gs-3955	524	13	cell	cell	NOUN
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gs-3955	524	15	mechanism	mechanism	NOUN
gs-3955	524	16	.	.	PUNCT
gs-3955	525	1	pnas	pnas	PROPN
gs-3955	525	2	,	,	PUNCT
gs-3955	525	3	109(47	109(47	NUM
gs-3955	525	4	)	)	PUNCT
gs-3955	525	5	,	,	PUNCT
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gs-3955	525	7	-	-	SYM
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gs-3955	525	9	.	.	PUNCT
gs-3955	526	1	[	[	X
gs-3955	526	2	pubmed	pubmed	NOUN
gs-3955	526	3	:	:	PUNCT
gs-3955	526	4	23129611	23129611	NUM
gs-3955	526	5	]	]	PUNCT
gs-3955	526	6	georgian	georgian	PROPN
gs-3955	526	7	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	526	8	მეცნიერები	მეცნიერები	NOUN
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gs-3955	526	10	7	7	NUM
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gs-3955	526	12	3	3	NUM
gs-3955	526	13	,	,	PUNCT
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gs-3955	526	17	.	.	PUNCT
gs-3955	527	1	gorbunova	gorbunova	PROPN
gs-3955	527	2	,	,	PUNCT
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gs-3955	527	4	,	,	PUNCT
gs-3955	527	5	et	et	PROPN
gs-3955	527	6	al	al	PROPN
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gs-3955	527	9	2020	2020	NUM
gs-3955	527	10	)	)	PUNCT
gs-3955	527	11	.	.	PUNCT
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gs-3955	528	3	of	of	ADP
gs-3955	528	4	stress	stress	NOUN
gs-3955	528	5	resistance	resistance	NOUN
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gs-3955	528	7	in	in	ADP
gs-3955	528	8	long	long	ADV
gs-3955	528	9	-	-	PUNCT
gs-3955	528	10	lived	live	VERB
gs-3955	528	11	species	specie	NOUN
gs-3955	528	12	.	.	PUNCT
gs-3955	529	1	cell	cell	NOUN
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gs-3955	529	3	,	,	PUNCT
gs-3955	529	4	32(5	32(5	NOUN
gs-3955	529	5	)	)	PUNCT
gs-3955	529	6	,	,	PUNCT
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gs-3955	529	8	.	.	PUNCT
gs-3955	530	1	[	[	X
gs-3955	530	2	pubmed	pubmed	X
gs-3955	530	3	:	:	PUNCT
gs-3955	530	4	32755598	32755598	NUM
gs-3955	530	5	]	]	PUNCT
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gs-3955	530	7	.	.	PUNCT
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gs-3955	530	9	,	,	PUNCT
gs-3955	530	10	v.	v.	ADV
gs-3955	530	11	,	,	PUNCT
gs-3955	530	12	et	et	PROPN
gs-3955	530	13	al	al	PROPN
gs-3955	530	14	.	.	PROPN
gs-3955	530	15	(	(	PUNCT
gs-3955	530	16	2022	2022	NUM
gs-3955	530	17	)	)	PUNCT
gs-3955	530	18	.	.	PUNCT
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gs-3955	531	2	genomics	genomic	NOUN
gs-3955	531	3	of	of	ADP
gs-3955	531	4	longevity	longevity	NOUN
gs-3955	531	5	in	in	ADP
gs-3955	531	6	salamanders	salamander	NOUN
gs-3955	531	7	.	.	PUNCT
gs-3955	532	1	nature	nature	NOUN
gs-3955	532	2	communications	communication	NOUN
gs-3955	532	3	,	,	PUNCT
gs-3955	532	4	13(1	13(1	NUM
gs-3955	532	5	)	)	PUNCT
gs-3955	532	6	,	,	PUNCT
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gs-3955	532	8	.	.	PUNCT
gs-3955	533	1	[	[	X
gs-3955	533	2	pubmed	pubmed	X
gs-3955	533	3	:	:	PUNCT
gs-3955	533	4	35236829	35236829	NUM
gs-3955	533	5	]	]	PUNCT
gs-3955	533	6	36	36	NUM
gs-3955	533	7	.	.	PUNCT
gs-3955	534	1	goricar	goricar	NOUN
gs-3955	534	2	,	,	PUNCT
gs-3955	534	3	k.	k.	PROPN
gs-3955	534	4	,	,	PUNCT
gs-3955	534	5	et	et	PROPN
gs-3955	534	6	al	al	PROPN
gs-3955	534	7	.	.	PROPN
gs-3955	534	8	(	(	PUNCT
gs-3955	534	9	2021	2021	NUM
gs-3955	534	10	)	)	PUNCT
gs-3955	534	11	.	.	PUNCT
gs-3955	535	1	antioxidant	antioxidant	ADJ
gs-3955	535	2	defense	defense	NOUN
gs-3955	535	3	in	in	ADP
gs-3955	535	4	the	the	DET
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gs-3955	535	6	-	-	PUNCT
gs-3955	535	7	dwelling	dwell	VERB
gs-3955	535	8	salamander	salamander	NOUN
gs-3955	535	9	proteus	proteus	NOUN
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gs-3955	535	11	.	.	PUNCT
gs-3955	536	1	comparative	comparative	ADJ
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gs-3955	536	5	,	,	PUNCT
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gs-3955	536	7	,	,	PUNCT
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gs-3955	536	9	.	.	PUNCT
gs-3955	537	1	[	[	X
gs-3955	537	2	pubmed	pubmed	X
gs-3955	537	3	:	:	PUNCT
gs-3955	537	4	34363902	34363902	NUM
gs-3955	537	5	]	]	PUNCT
gs-3955	537	6	37	37	NUM
gs-3955	537	7	.	.	PUNCT
gs-3955	538	1	gruber	gruber	PROPN
gs-3955	538	2	,	,	PUNCT
gs-3955	538	3	h.	h.	PROPN
gs-3955	538	4	,	,	PUNCT
gs-3955	538	5	et	et	PROPN
gs-3955	538	6	al	al	PROPN
gs-3955	538	7	.	.	PUNCT
gs-3955	538	8	(	(	PUNCT
gs-3955	538	9	2015	2015	NUM
gs-3955	538	10	)	)	PUNCT
gs-3955	538	11	.	.	PUNCT
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gs-3955	539	2	resistance	resistance	NOUN
gs-3955	539	3	in	in	ADP
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gs-3955	539	5	-	-	PUNCT
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gs-3955	539	8	.	.	PUNCT
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gs-3955	540	3	,	,	PUNCT
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gs-3955	540	5	,	,	PUNCT
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gs-3955	540	7	-	-	SYM
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gs-3955	540	9	.	.	PUNCT
gs-3955	541	1	[	[	X
gs-3955	541	2	pubmed	pubmed	X
gs-3955	541	3	:	:	PUNCT
gs-3955	541	4	26475934	26475934	NUM
gs-3955	541	5	]	]	PUNCT
gs-3955	541	6	38	38	NUM
gs-3955	541	7	.	.	PUNCT
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gs-3955	541	9	,	,	PUNCT
gs-3955	541	10	o.s	o.s	PROPN
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gs-3955	541	12	,	,	PUNCT
gs-3955	541	13	et	et	PROPN
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gs-3955	541	15	.	.	PROPN
gs-3955	541	16	(	(	PUNCT
gs-3955	541	17	2020	2020	NUM
gs-3955	541	18	)	)	PUNCT
gs-3955	541	19	.	.	PUNCT
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gs-3955	542	3	in	in	ADP
gs-3955	542	4	rockfishes	rockfish	NOUN
gs-3955	542	5	(	(	PUNCT
gs-3955	542	6	sebastes	sebaste	VERB
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gs-3955	542	8	.	.	PUNCT
gs-3955	542	9	)	)	PUNCT
gs-3955	542	10	.	.	PUNCT
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gs-3955	543	3	,	,	PUNCT
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gs-3955	543	5	,	,	PUNCT
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gs-3955	543	7	.	.	PUNCT
gs-3955	544	1	[	[	X
gs-3955	544	2	pubmed	pubmed	X
gs-3955	544	3	:	:	PUNCT
gs-3955	544	4	32834777	32834777	NUM
gs-3955	544	5	]	]	PUNCT
gs-3955	544	6	39	39	NUM
gs-3955	544	7	.	.	PUNCT
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gs-3955	545	2	,	,	PUNCT
gs-3955	545	3	m.	m.	PROPN
gs-3955	545	4	j.	j.	PROPN
gs-3955	545	5	,	,	PUNCT
gs-3955	545	6	et	et	PROPN
gs-3955	545	7	al	al	PROPN
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gs-3955	545	9	(	(	PUNCT
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gs-3955	545	11	)	)	PUNCT
gs-3955	545	12	.	.	PUNCT
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gs-3955	546	6	in	in	ADP
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gs-3955	546	9	.	.	PUNCT
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gs-3955	547	3	,	,	PUNCT
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gs-3955	547	5	)	)	PUNCT
gs-3955	547	6	,	,	PUNCT
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gs-3955	547	8	-	-	SYM
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gs-3955	547	10	.	.	PUNCT
gs-3955	548	1	[	[	X
gs-3955	548	2	pubmed	pubmed	NOUN
gs-3955	548	3	:	:	PUNCT
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gs-3955	548	5	]	]	PUNCT
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gs-3955	549	2	.	.	PUNCT
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gs-3955	549	4	,	,	PUNCT
gs-3955	549	5	s.	s.	PROPN
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gs-3955	549	7	et	et	PROPN
gs-3955	549	8	al	al	PROPN
gs-3955	549	9	.	.	PUNCT
gs-3955	549	10	(	(	PUNCT
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gs-3955	549	13	.	.	PUNCT
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gs-3955	550	6	in	in	ADP
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gs-3955	551	3	,	,	PUNCT
gs-3955	551	4	20(3	20(3	NOUN
gs-3955	551	5	)	)	PUNCT
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gs-3955	551	8	.	.	PUNCT
gs-3955	552	1	[	[	X
gs-3955	552	2	pubmed	pubmed	X
gs-3955	552	3	:	:	PUNCT
gs-3955	552	4	33624461	33624461	NUM
gs-3955	552	5	]	]	PUNCT
gs-3955	552	6	41	41	NUM
gs-3955	552	7	.	.	PUNCT
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gs-3955	553	2	,	,	PUNCT
gs-3955	553	3	s.	s.	PROPN
gs-3955	553	4	,	,	PUNCT
gs-3955	553	5	et	et	PROPN
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gs-3955	553	7	.	.	PROPN
gs-3955	553	8	(	(	PUNCT
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gs-3955	553	11	.	.	PUNCT
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gs-3955	554	6	-	-	PUNCT
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gs-3955	554	9	.	.	PUNCT
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gs-3955	555	5	)	)	PUNCT
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gs-3955	555	8	.	.	PUNCT
gs-3955	556	1	[	[	X
gs-3955	556	2	pubmed	pubmed	X
gs-3955	556	3	:	:	PUNCT
gs-3955	556	4	34918468	34918468	NUM
gs-3955	556	5	]	]	X
gs-3955	556	6	42	42	NUM
gs-3955	556	7	.	.	PUNCT
gs-3955	557	1	hyde	hyde	PROPN
gs-3955	557	2	,	,	PUNCT
gs-3955	557	3	j.r	j.r	PROPN
gs-3955	557	4	.	.	PROPN
gs-3955	557	5	,	,	PUNCT
gs-3955	557	6	et	et	PROPN
gs-3955	557	7	al	al	PROPN
gs-3955	557	8	.	.	PROPN
gs-3955	557	9	(	(	PUNCT
gs-3955	557	10	2020	2020	NUM
gs-3955	557	11	)	)	PUNCT
gs-3955	557	12	.	.	PUNCT
gs-3955	558	1	dna	dna	PROPN
gs-3955	558	2	repair	repair	NOUN
gs-3955	558	3	genes	gene	NOUN
gs-3955	558	4	in	in	ADP
gs-3955	558	5	long	long	ADV
gs-3955	558	6	-	-	PUNCT
gs-3955	558	7	lived	live	VERB
gs-3955	558	8	sebastes	sebaste	NOUN
gs-3955	558	9	species	specie	NOUN
gs-3955	558	10	.	.	PUNCT
gs-3955	559	1	molecular	molecular	ADJ
gs-3955	559	2	ecology	ecology	NOUN
gs-3955	559	3	,	,	PUNCT
gs-3955	559	4	29(15	29(15	NUM
gs-3955	559	5	)	)	PUNCT
gs-3955	559	6	,	,	PUNCT
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gs-3955	559	8	-	-	SYM
gs-3955	559	9	2802	2802	NUM
gs-3955	559	10	.	.	PUNCT
gs-3955	560	1	[	[	X
gs-3955	560	2	pubmed	pubmed	X
gs-3955	560	3	:	:	PUNCT
gs-3955	560	4	32557907	32557907	NUM
gs-3955	560	5	]	]	PUNCT
gs-3955	560	6	43	43	NUM
gs-3955	560	7	.	.	PUNCT
gs-3955	560	8	issartel	issartel	PROPN
gs-3955	560	9	,	,	PUNCT
gs-3955	560	10	j.	j.	PROPN
gs-3955	560	11	,	,	PUNCT
gs-3955	560	12	et	et	PROPN
gs-3955	560	13	al	al	PROPN
gs-3955	560	14	.	.	PUNCT
gs-3955	560	15	(	(	PUNCT
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gs-3955	560	17	)	)	PUNCT
gs-3955	560	18	.	.	PUNCT
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gs-3955	561	2	lifespan	lifespan	PROPN
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gs-3955	561	4	the	the	DET
gs-3955	561	5	human	human	ADJ
gs-3955	561	6	fish	fish	NOUN
gs-3955	561	7	(	(	PUNCT
gs-3955	561	8	proteus	proteus	NOUN
gs-3955	561	9	anguinus	anguinus	PROPN
gs-3955	561	10	)	)	PUNCT
gs-3955	561	11	.	.	PUNCT
gs-3955	562	1	biology	biology	NOUN
gs-3955	562	2	letters	letter	NOUN
gs-3955	562	3	,	,	PUNCT
gs-3955	562	4	5(6	5(6	NUM
gs-3955	562	5	)	)	PUNCT
gs-3955	562	6	,	,	PUNCT
gs-3955	562	7	828–833	828–833	NUM
gs-3955	562	8	.	.	PUNCT
gs-3955	563	1	[	[	X
gs-3955	563	2	pubmed	pubmed	NOUN
gs-3955	563	3	:	:	PUNCT
gs-3955	563	4	19656860	19656860	NUM
gs-3955	563	5	]	]	PUNCT
gs-3955	563	6	44	44	NUM
gs-3955	563	7	.	.	PUNCT
gs-3955	564	1	jaba	jaba	PROPN
gs-3955	564	2	,	,	PUNCT
gs-3955	564	3	t.	t.	PROPN
gs-3955	564	4	(	(	PUNCT
gs-3955	564	5	2022	2022	NUM
gs-3955	564	6	)	)	PUNCT
gs-3955	564	7	.	.	PUNCT
gs-3955	565	1	dasatinib	dasatinib	NOUN
gs-3955	565	2	and	and	CCONJ
gs-3955	565	3	quercetin	quercetin	NOUN
gs-3955	565	4	:	:	PUNCT
gs-3955	565	5	short	short	ADJ
gs-3955	565	6	-	-	PUNCT
gs-3955	565	7	term	term	NOUN
gs-3955	565	8	simultaneous	simultaneous	ADJ
gs-3955	565	9	administration	administration	NOUN
gs-3955	565	10	yields	yield	VERB
gs-3955	565	11	senolytic	senolytic	ADJ
gs-3955	565	12	effect	effect	NOUN
gs-3955	565	13	in	in	ADP
gs-3955	565	14	humans	human	NOUN
gs-3955	565	15	.	.	PUNCT
gs-3955	566	1	issues	issue	NOUN
gs-3955	566	2	and	and	CCONJ
gs-3955	566	3	developments	development	NOUN
gs-3955	566	4	in	in	ADP
gs-3955	566	5	medicine	medicine	NOUN
gs-3955	566	6	and	and	CCONJ
gs-3955	566	7	medical	medical	ADJ
gs-3955	566	8	research	research	NOUN
gs-3955	566	9	,	,	PUNCT
gs-3955	566	10	2	2	NUM
gs-3955	566	11	,	,	PUNCT
gs-3955	566	12	22	22	NUM
gs-3955	566	13	-	-	SYM
gs-3955	566	14	31	31	NUM
gs-3955	566	15	.	.	PUNCT
gs-3955	567	1	45	45	NUM
gs-3955	567	2	.	.	PUNCT
gs-3955	568	1	jaber	jaber	PROPN
gs-3955	568	2	-	-	PUNCT
gs-3955	568	3	hijazi	hijazi	PROPN
gs-3955	568	4	,	,	PUNCT
gs-3955	568	5	f.	f.	PROPN
gs-3955	568	6	,	,	PUNCT
gs-3955	568	7	et	et	PROPN
gs-3955	568	8	al	al	PROPN
gs-3955	568	9	.	.	PUNCT
gs-3955	568	10	(	(	PUNCT
gs-3955	568	11	2016	2016	NUM
gs-3955	568	12	)	)	PUNCT
gs-3955	568	13	.	.	PUNCT
gs-3955	569	1	epigenetic	epigenetic	ADJ
gs-3955	569	2	regulation	regulation	NOUN
gs-3955	569	3	in	in	ADP
gs-3955	569	4	planarian	planarian	ADJ
gs-3955	569	5	stem	stem	NOUN
gs-3955	569	6	cells	cell	NOUN
gs-3955	569	7	.	.	PUNCT
gs-3955	570	1	development	development	NOUN
gs-3955	570	2	,	,	PUNCT
gs-3955	570	3	143(12	143(12	NUM
gs-3955	570	4	)	)	PUNCT
gs-3955	570	5	,	,	PUNCT
gs-3955	570	6	2089	2089	NUM
gs-3955	570	7	-	-	SYM
gs-3955	570	8	2101	2101	NUM
gs-3955	570	9	.	.	PUNCT
gs-3955	571	1	[	[	X
gs-3955	571	2	pubmed	pubmed	NOUN
gs-3955	571	3	:	:	PUNCT
gs-3955	571	4	27287811	27287811	NUM
gs-3955	571	5	]	]	PUNCT
gs-3955	571	6	46	46	NUM
gs-3955	571	7	.	.	PUNCT
gs-3955	572	1	jansen	jansen	PROPN
gs-3955	572	2	,	,	PUNCT
gs-3955	572	3	m.	m.	NOUN
gs-3955	572	4	,	,	PUNCT
gs-3955	572	5	et	et	PROPN
gs-3955	572	6	al	al	PROPN
gs-3955	572	7	.	.	PROPN
gs-3955	572	8	(	(	PUNCT
gs-3955	572	9	2020	2020	NUM
gs-3955	572	10	)	)	PUNCT
gs-3955	572	11	.	.	PUNCT
gs-3955	573	1	epigenetic	epigenetic	ADJ
gs-3955	573	2	aging	aging	NOUN
gs-3955	573	3	in	in	ADP
gs-3955	573	4	marine	marine	ADJ
gs-3955	573	5	bivalves	bivalve	NOUN
gs-3955	573	6	.	.	PUNCT
gs-3955	574	1	aging	age	VERB
gs-3955	574	2	cell	cell	NOUN
gs-3955	574	3	,	,	PUNCT
gs-3955	574	4	19(3	19(3	NUM
gs-3955	574	5	)	)	PUNCT
gs-3955	574	6	,	,	PUNCT
gs-3955	574	7	e13115	e13115	PROPN
gs-3955	574	8	.	.	PUNCT
gs-3955	575	1	[	[	X
gs-3955	575	2	pubmed	pubmed	NOUN
gs-3955	575	3	:	:	PUNCT
gs-3955	575	4	32090471	32090471	NUM
gs-3955	575	5	]	]	PUNCT
gs-3955	576	1	47	47	NUM
gs-3955	576	2	.	.	PUNCT
gs-3955	577	1	jansen	jansen	PROPN
gs-3955	577	2	,	,	PUNCT
gs-3955	577	3	m.	m.	NOUN
gs-3955	577	4	,	,	PUNCT
gs-3955	577	5	et	et	PROPN
gs-3955	577	6	al	al	PROPN
gs-3955	577	7	.	.	PROPN
gs-3955	577	8	(	(	PUNCT
gs-3955	577	9	2021	2021	NUM
gs-3955	577	10	)	)	PUNCT
gs-3955	577	11	.	.	PUNCT
gs-3955	578	1	epigenetic	epigenetic	ADJ
gs-3955	578	2	aging	aging	NOUN
gs-3955	578	3	in	in	ADP
gs-3955	578	4	marine	marine	ADJ
gs-3955	578	5	invertebrates	invertebrate	NOUN
gs-3955	578	6	.	.	PUNCT
gs-3955	579	1	aging	age	VERB
gs-3955	579	2	cell	cell	NOUN
gs-3955	579	3	,	,	PUNCT
gs-3955	579	4	20(3	20(3	NOUN
gs-3955	579	5	)	)	PUNCT
gs-3955	579	6	,	,	PUNCT
gs-3955	579	7	e13324	e13324	PROPN
gs-3955	579	8	.	.	PUNCT
gs-3955	580	1	[	[	X
gs-3955	580	2	pubmed	pubmed	X
gs-3955	580	3	:	:	PUNCT
gs-3955	580	4	33624461	33624461	NUM
gs-3955	580	5	]	]	PUNCT
gs-3955	580	6	48	48	NUM
gs-3955	580	7	.	.	PUNCT
gs-3955	581	1	joven	joven	PROPN
gs-3955	581	2	,	,	PUNCT
gs-3955	581	3	a.	a.	NOUN
gs-3955	581	4	,	,	PUNCT
gs-3955	581	5	&	&	CCONJ
gs-3955	581	6	simon	simon	PROPN
gs-3955	581	7	,	,	PUNCT
gs-3955	581	8	a.	a.	NOUN
gs-3955	581	9	(	(	PUNCT
gs-3955	581	10	2021	2021	NUM
gs-3955	581	11	)	)	PUNCT
gs-3955	581	12	.	.	PUNCT
gs-3955	582	1	salamander	salamander	NOUN
gs-3955	582	2	regeneration	regeneration	NOUN
gs-3955	582	3	as	as	ADP
gs-3955	582	4	a	a	DET
gs-3955	582	5	model	model	NOUN
gs-3955	582	6	for	for	ADP
gs-3955	582	7	developing	develop	VERB
gs-3955	582	8	novel	novel	ADJ
gs-3955	582	9	regenerative	regenerative	ADJ
gs-3955	582	10	and	and	CCONJ
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gs-3955	582	12	therapies	therapy	NOUN
gs-3955	582	13	.	.	PUNCT
gs-3955	583	1	journal	journal	NOUN
gs-3955	583	2	of	of	ADP
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gs-3955	583	4	biology	biology	NOUN
gs-3955	583	5	,	,	PUNCT
gs-3955	583	6	9(1	9(1	NUM
gs-3955	583	7	)	)	PUNCT
gs-3955	583	8	,	,	PUNCT
gs-3955	583	9	12	12	NUM
gs-3955	583	10	.	.	PUNCT
gs-3955	584	1	[	[	X
gs-3955	584	2	pubmed	pubmed	X
gs-3955	584	3	:	:	PUNCT
gs-3955	584	4	33669547	33669547	NUM
gs-3955	584	5	]	]	PUNCT
gs-3955	584	6	49	49	NUM
gs-3955	584	7	.	.	PUNCT
gs-3955	585	1	kyne	kyne	PROPN
gs-3955	585	2	,	,	PUNCT
gs-3955	585	3	p.	p.	NOUN
gs-3955	585	4	m.	m.	NOUN
gs-3955	585	5	,	,	PUNCT
gs-3955	585	6	et	et	PROPN
gs-3955	585	7	al	al	PROPN
gs-3955	585	8	.	.	PROPN
gs-3955	586	1	(	(	PUNCT
gs-3955	586	2	2019	2019	NUM
gs-3955	586	3	)	)	PUNCT
gs-3955	586	4	.	.	PUNCT
gs-3955	587	1	the	the	DET
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gs-3955	587	3	status	status	NOUN
gs-3955	587	4	of	of	ADP
gs-3955	587	5	the	the	DET
gs-3955	587	6	greenland	greenland	NOUN
gs-3955	587	7	shark	shark	NOUN
gs-3955	587	8	.	.	PUNCT
gs-3955	588	1	global	global	ADJ
gs-3955	588	2	ecology	ecology	NOUN
gs-3955	588	3	and	and	CCONJ
gs-3955	588	4	conservation	conservation	NOUN
gs-3955	588	5	,	,	PUNCT
gs-3955	588	6	20	20	NUM
gs-3955	588	7	,	,	PUNCT
gs-3955	588	8	e00710	e00710	ADJ
gs-3955	588	9	.	.	PUNCT
gs-3955	589	1	[	[	X
gs-3955	589	2	pubmed	pubmed	X
gs-3955	589	3	:	:	PUNCT
gs-3955	589	4	31576231	31576231	NUM
gs-3955	589	5	]	]	PUNCT
gs-3955	590	1	50	50	NUM
gs-3955	590	2	.	.	PUNCT
gs-3955	590	3	lewis	lewis	PROPN
gs-3955	590	4	,	,	PUNCT
gs-3955	590	5	k.n	k.n	PROPN
gs-3955	590	6	.	.	PROPN
gs-3955	590	7	,	,	PUNCT
gs-3955	590	8	et	et	PROPN
gs-3955	590	9	al	al	PROPN
gs-3955	590	10	.	.	PUNCT
gs-3955	590	11	(	(	PUNCT
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gs-3955	590	13	)	)	PUNCT
gs-3955	590	14	.	.	PUNCT
gs-3955	591	1	the	the	DET
gs-3955	591	2	naked	naked	ADJ
gs-3955	591	3	mole	mole	ADJ
gs-3955	591	4	-	-	PUNCT
gs-3955	591	5	rat	rat	NOUN
gs-3955	591	6	response	response	NOUN
gs-3955	591	7	to	to	PART
gs-3955	591	8	oxidative	oxidative	ADJ
gs-3955	591	9	stress	stress	NOUN
gs-3955	591	10	:	:	PUNCT
gs-3955	591	11	just	just	ADV
gs-3955	591	12	deal	deal	VERB
gs-3955	591	13	with	with	ADP
gs-3955	591	14	it	it	PRON
gs-3955	591	15	.	.	PUNCT
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gs-3955	592	2	&	&	CCONJ
gs-3955	592	3	redox	redox	PROPN
gs-3955	592	4	signaling	signal	VERB
gs-3955	592	5	,	,	PUNCT
gs-3955	592	6	29(10	29(10	NOUN
gs-3955	592	7	)	)	PUNCT
gs-3955	592	8	,	,	PUNCT
gs-3955	592	9	1019	1019	NUM
gs-3955	592	10	-	-	SYM
gs-3955	592	11	1035	1035	NUM
gs-3955	592	12	.	.	PUNCT
gs-3955	593	1	[	[	X
gs-3955	593	2	pubmed	pubmed	X
gs-3955	593	3	:	:	PUNCT
gs-3955	593	4	29320861	29320861	NUM
gs-3955	593	5	]	]	PUNCT
gs-3955	594	1	51	51	NUM
gs-3955	594	2	.	.	PUNCT
gs-3955	595	1	lópez	lópez	NOUN
gs-3955	595	2	-	-	PUNCT
gs-3955	595	3	otín	otín	PROPN
gs-3955	595	4	,	,	PUNCT
gs-3955	595	5	c.	c.	PROPN
gs-3955	595	6	,	,	PUNCT
gs-3955	595	7	et	et	PROPN
gs-3955	595	8	al	al	PROPN
gs-3955	595	9	.	.	PROPN
gs-3955	595	10	(	(	PUNCT
gs-3955	595	11	2013	2013	NUM
gs-3955	595	12	)	)	PUNCT
gs-3955	595	13	.	.	PUNCT
gs-3955	596	1	the	the	DET
gs-3955	596	2	hallmarks	hallmark	NOUN
gs-3955	596	3	of	of	ADP
gs-3955	596	4	aging	aging	NOUN
gs-3955	596	5	.	.	PUNCT
gs-3955	597	1	cell	cell	NOUN
gs-3955	597	2	,	,	PUNCT
gs-3955	597	3	153(6	153(6	NUM
gs-3955	597	4	)	)	PUNCT
gs-3955	597	5	,	,	PUNCT
gs-3955	597	6	1194	1194	NUM
gs-3955	597	7	-	-	SYM
gs-3955	597	8	1217	1217	NUM
gs-3955	597	9	.	.	PUNCT
gs-3955	598	1	[	[	X
gs-3955	598	2	pubmed	pubmed	X
gs-3955	598	3	:	:	PUNCT
gs-3955	598	4	23746838	23746838	NUM
gs-3955	598	5	]	]	PUNCT
gs-3955	598	6	georgian	georgian	PROPN
gs-3955	598	7	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	598	8	მეცნიერები	მეცნიერები	NOUN
gs-3955	598	9	ტ.	ტ.	VERB
gs-3955	598	10	7	7	NUM
gs-3955	598	11	n	n	DET
gs-3955	598	12	3	3	NUM
gs-3955	598	13	,	,	PUNCT
gs-3955	598	14	2025	2025	NUM
gs-3955	598	15	392	392	NUM
gs-3955	598	16	52	52	NUM
gs-3955	598	17	.	.	PUNCT
gs-3955	599	1	lópez	lópez	NOUN
gs-3955	599	2	-	-	PUNCT
gs-3955	599	3	otín	otín	PROPN
gs-3955	599	4	,	,	PUNCT
gs-3955	599	5	c.	c.	PROPN
gs-3955	599	6	,	,	PUNCT
gs-3955	599	7	et	et	PROPN
gs-3955	599	8	al	al	PROPN
gs-3955	599	9	.	.	PROPN
gs-3955	599	10	(	(	PUNCT
gs-3955	599	11	2021	2021	NUM
gs-3955	599	12	)	)	PUNCT
gs-3955	599	13	.	.	PUNCT
gs-3955	600	1	hallmarks	hallmark	NOUN
gs-3955	600	2	of	of	ADP
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gs-3955	600	4	.	.	PUNCT
gs-3955	601	1	cell	cell	NOUN
gs-3955	601	2	,	,	PUNCT
gs-3955	601	3	184(1	184(1	NUM
gs-3955	601	4	)	)	PUNCT
gs-3955	601	5	,	,	PUNCT
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gs-3955	601	7	-	-	SYM
gs-3955	601	8	63	63	NUM
gs-3955	601	9	.	.	PUNCT
gs-3955	602	1	[	[	X
gs-3955	602	2	pubmed	pubmed	X
gs-3955	602	3	:	:	PUNCT
gs-3955	602	4	33340459	33340459	NUM
gs-3955	602	5	]	]	PUNCT
gs-3955	603	1	53	53	NUM
gs-3955	603	2	.	.	PUNCT
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gs-3955	603	4	,	,	PUNCT
gs-3955	603	5	s.	s.	PROPN
gs-3955	603	6	,	,	PUNCT
gs-3955	603	7	et	et	PROPN
gs-3955	603	8	al	al	PROPN
gs-3955	603	9	.	.	PUNCT
gs-3955	603	10	(	(	PUNCT
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gs-3955	603	12	)	)	PUNCT
gs-3955	603	13	.	.	PUNCT
gs-3955	604	1	epigenetic	epigenetic	ADJ
gs-3955	604	2	aging	age	VERB
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gs-3955	604	4	in	in	ADP
gs-3955	604	5	salamanders	salamander	NOUN
gs-3955	604	6	.	.	PUNCT
gs-3955	605	1	aging	age	VERB
gs-3955	605	2	cell	cell	NOUN
gs-3955	605	3	,	,	PUNCT
gs-3955	605	4	20(3	20(3	NOUN
gs-3955	605	5	)	)	PUNCT
gs-3955	605	6	,	,	PUNCT
gs-3955	605	7	e13324	e13324	PROPN
gs-3955	605	8	.	.	PUNCT
gs-3955	606	1	[	[	X
gs-3955	606	2	pubmed	pubmed	X
gs-3955	606	3	:	:	PUNCT
gs-3955	606	4	33624461	33624461	NUM
gs-3955	606	5	]	]	PUNCT
gs-3955	606	6	54	54	NUM
gs-3955	606	7	.	.	PUNCT
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gs-3955	607	2	sl	sl	PROPN
gs-3955	607	3	,	,	PUNCT
gs-3955	607	4	zhang	zhang	PROPN
gs-3955	607	5	q	q	X
gs-3955	607	6	,	,	PUNCT
gs-3955	607	7	lemetre	lemetre	PROPN
gs-3955	607	8	c	c	NOUN
gs-3955	607	9	,	,	PUNCT
gs-3955	607	10	seim	seim	PROPN
gs-3955	607	11	i	i	PROPN
gs-3955	607	12	,	,	PUNCT
gs-3955	607	13	calder	calder	PROPN
gs-3955	607	14	rb	rb	PROPN
gs-3955	607	15	,	,	PUNCT
gs-3955	607	16	hoeijmakers	hoeijmaker	NOUN
gs-3955	607	17	j	j	PROPN
gs-3955	607	18	,	,	PUNCT
gs-3955	607	19	suh	suh	PROPN
gs-3955	607	20	y	y	PROPN
gs-3955	607	21	,	,	PUNCT
gs-3955	607	22	gladyshev	gladyshev	PROPN
gs-3955	607	23	vn	vn	PROPN
gs-3955	607	24	,	,	PUNCT
gs-3955	607	25	seluanov	seluanov	PROPN
gs-3955	607	26	a	a	PROPN
gs-3955	607	27	,	,	PUNCT
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gs-3955	607	29	v	v	NOUN
gs-3955	607	30	,	,	PUNCT
gs-3955	607	31	et	et	PROPN
gs-3955	607	32	al	al	PROPN
gs-3955	607	33	.	.	PROPN
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gs-3955	608	2	analysis	analysis	NOUN
gs-3955	608	3	of	of	ADP
gs-3955	608	4	genome	genome	NOUN
gs-3955	608	5	maintenance	maintenance	NOUN
gs-3955	608	6	genes	gene	NOUN
gs-3955	608	7	in	in	ADP
gs-3955	608	8	naked	naked	ADJ
gs-3955	608	9	mole	mole	NOUN
gs-3955	608	10	rat	rat	NOUN
gs-3955	608	11	,	,	PUNCT
gs-3955	608	12	mouse	mouse	NOUN
gs-3955	608	13	,	,	PUNCT
gs-3955	608	14	and	and	CCONJ
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gs-3955	608	16	.	.	PUNCT
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gs-3955	609	3	.	.	PUNCT
gs-3955	610	1	2015;14:288	2015;14:288	X
gs-3955	610	2	–	–	PUNCT
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gs-3955	610	4	.	.	PUNCT
gs-3955	611	1	55	55	NUM
gs-3955	611	2	.	.	PUNCT
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gs-3955	612	2	,	,	PUNCT
gs-3955	612	3	k.m	k.m	PROPN
gs-3955	612	4	.	.	PROPN
gs-3955	612	5	(	(	PUNCT
gs-3955	612	6	2001	2001	NUM
gs-3955	612	7	)	)	PUNCT
gs-3955	612	8	.	.	PUNCT
gs-3955	613	1	maximum	maximum	ADJ
gs-3955	613	2	ages	age	NOUN
gs-3955	613	3	of	of	ADP
gs-3955	613	4	groundfishes	groundfish	NOUN
gs-3955	613	5	in	in	ADP
gs-3955	613	6	waters	water	NOUN
gs-3955	613	7	off	off	ADP
gs-3955	613	8	alaska	alaska	PROPN
gs-3955	613	9	and	and	CCONJ
gs-3955	613	10	british	british	PROPN
gs-3955	613	11	columbia	columbia	PROPN
gs-3955	613	12	.	.	PUNCT
gs-3955	614	1	fisheries	fishery	NOUN
gs-3955	614	2	research	research	NOUN
gs-3955	614	3	,	,	PUNCT
gs-3955	614	4	52(1	52(1	NOUN
gs-3955	614	5	-	-	SYM
gs-3955	614	6	2	2	NUM
gs-3955	614	7	)	)	PUNCT
gs-3955	614	8	,	,	PUNCT
gs-3955	614	9	1	1	NUM
gs-3955	614	10	-	-	SYM
gs-3955	614	11	12	12	NUM
gs-3955	614	12	.	.	PUNCT
gs-3955	615	1	[	[	X
gs-3955	615	2	pubmed	pubmed	NOUN
gs-3955	615	3	:	:	PUNCT
gs-3955	615	4	11393815	11393815	NUM
gs-3955	615	5	]	]	PUNCT
gs-3955	615	6	56	56	NUM
gs-3955	615	7	.	.	PUNCT
gs-3955	616	1	munro	munro	PROPN
gs-3955	616	2	dm	dm	PROPN
gs-3955	616	3	,	,	PUNCT
gs-3955	616	4	blier	blier	NOUN
gs-3955	616	5	pu	pu	PROPN
gs-3955	616	6	.	.	PUNCT
gs-3955	617	1	the	the	DET
gs-3955	617	2	extreme	extreme	ADJ
gs-3955	617	3	longevity	longevity	NOUN
gs-3955	617	4	of	of	ADP
gs-3955	617	5	arctica	arctica	PROPN
gs-3955	617	6	islandica	islandica	PROPN
gs-3955	617	7	is	be	AUX
gs-3955	617	8	associated	associate	VERB
gs-3955	617	9	with	with	ADP
gs-3955	617	10	increased	increase	VERB
gs-3955	617	11	peroxidation	peroxidation	NOUN
gs-3955	617	12	resistance	resistance	NOUN
gs-3955	617	13	in	in	ADP
gs-3955	617	14	mitochondrial	mitochondrial	ADJ
gs-3955	617	15	membranes	membrane	NOUN
gs-3955	617	16	.	.	PUNCT
gs-3955	618	1	aging	age	VERB
gs-3955	618	2	cell	cell	NOUN
gs-3955	618	3	.	.	PUNCT
gs-3955	619	1	2012;11(5):845–855	2012;11(5):845–855	NUM
gs-3955	619	2	.	.	PUNCT
gs-3955	620	1	57	57	NUM
gs-3955	620	2	.	.	PUNCT
gs-3955	621	1	nielsen	nielsen	PROPN
gs-3955	621	2	,	,	PUNCT
gs-3955	621	3	j.	j.	PROPN
gs-3955	621	4	,	,	PUNCT
gs-3955	621	5	et	et	PROPN
gs-3955	621	6	al	al	PROPN
gs-3955	621	7	.	.	PUNCT
gs-3955	621	8	(	(	PUNCT
gs-3955	621	9	2016	2016	NUM
gs-3955	621	10	)	)	PUNCT
gs-3955	621	11	.	.	PUNCT
gs-3955	622	1	eye	eye	NOUN
gs-3955	622	2	lens	lens	PROPN
gs-3955	622	3	radiocarbon	radiocarbon	NOUN
gs-3955	622	4	reveals	reveal	VERB
gs-3955	622	5	centuries	century	NOUN
gs-3955	622	6	of	of	ADP
gs-3955	622	7	longevity	longevity	NOUN
gs-3955	622	8	in	in	ADP
gs-3955	622	9	the	the	DET
gs-3955	622	10	greenland	greenland	NOUN
gs-3955	622	11	shark	shark	NOUN
gs-3955	622	12	(	(	PUNCT
gs-3955	622	13	somniosus	somniosus	PROPN
gs-3955	622	14	microcephalus	microcephalus	PROPN
gs-3955	622	15	)	)	PUNCT
gs-3955	622	16	.	.	PUNCT
gs-3955	623	1	science	science	NOUN
gs-3955	623	2	,	,	PUNCT
gs-3955	623	3	353(6300	353(6300	NUM
gs-3955	623	4	)	)	PUNCT
gs-3955	623	5	,	,	PUNCT
gs-3955	623	6	702	702	NUM
gs-3955	623	7	-	-	SYM
gs-3955	623	8	704	704	NUM
gs-3955	623	9	.	.	PUNCT
gs-3955	624	1	[	[	X
gs-3955	624	2	pubmed	pubmed	NOUN
gs-3955	624	3	:	:	PUNCT
gs-3955	624	4	27516602	27516602	NUM
gs-3955	624	5	]	]	PUNCT
gs-3955	625	1	58	58	NUM
gs-3955	625	2	.	.	PUNCT
gs-3955	626	1	nielsen	nielsen	PROPN
gs-3955	626	2	j	j	PROPN
gs-3955	626	3	,	,	PUNCT
gs-3955	626	4	hedeholm	hedeholm	PROPN
gs-3955	626	5	rb	rb	PROPN
gs-3955	626	6	,	,	PUNCT
gs-3955	626	7	heinemeier	heinemeier	PROPN
gs-3955	626	8	j	j	PROPN
gs-3955	626	9	,	,	PUNCT
gs-3955	626	10	bushnell	bushnell	PROPN
gs-3955	626	11	pg	pg	PROPN
gs-3955	626	12	,	,	PUNCT
gs-3955	626	13	christiansen	christiansen	PROPN
gs-3955	626	14	js	js	PROPN
gs-3955	626	15	,	,	PUNCT
gs-3955	626	16	olsen	olsen	PROPN
gs-3955	626	17	j	j	PROPN
gs-3955	626	18	,	,	PUNCT
gs-3955	626	19	et	et	PROPN
gs-3955	626	20	al	al	PROPN
gs-3955	626	21	.	.	PUNCT
gs-3955	627	1	eye	eye	NOUN
gs-3955	627	2	lens	lens	PROPN
gs-3955	627	3	radiocarbon	radiocarbon	NOUN
gs-3955	627	4	reveals	reveal	VERB
gs-3955	627	5	centuries	century	NOUN
gs-3955	627	6	of	of	ADP
gs-3955	627	7	longevity	longevity	NOUN
gs-3955	627	8	in	in	ADP
gs-3955	627	9	the	the	DET
gs-3955	627	10	greenland	greenland	NOUN
gs-3955	627	11	shark	shark	NOUN
gs-3955	627	12	(	(	PUNCT
gs-3955	627	13	somniosus	somniosus	PROPN
gs-3955	627	14	microcephalus	microcephalus	PROPN
gs-3955	627	15	)	)	PUNCT
gs-3955	627	16	.	.	PUNCT
gs-3955	628	1	science	science	NOUN
gs-3955	628	2	.	.	PUNCT
gs-3955	629	1	2016;353(6300):702–4	2016;353(6300):702–4	NUM
gs-3955	629	2	.	.	PROPN
gs-3955	630	1	59	59	NUM
gs-3955	630	2	.	.	PUNCT
gs-3955	630	3	nowoshilow	nowoshilow	PROPN
gs-3955	630	4	,	,	PUNCT
gs-3955	630	5	s.	s.	PROPN
gs-3955	630	6	,	,	PUNCT
gs-3955	630	7	et	et	PROPN
gs-3955	630	8	al	al	PROPN
gs-3955	630	9	.	.	PUNCT
gs-3955	631	1	(	(	PUNCT
gs-3955	631	2	2018	2018	NUM
gs-3955	631	3	)	)	PUNCT
gs-3955	631	4	.	.	PUNCT
gs-3955	632	1	the	the	DET
gs-3955	632	2	axolotl	axolotl	NOUN
gs-3955	632	3	genome	genome	NOUN
gs-3955	632	4	and	and	CCONJ
gs-3955	632	5	the	the	DET
gs-3955	632	6	evolution	evolution	NOUN
gs-3955	632	7	of	of	ADP
gs-3955	632	8	key	key	ADJ
gs-3955	632	9	tissue	tissue	NOUN
gs-3955	632	10	formation	formation	NOUN
gs-3955	632	11	regulators	regulator	NOUN
gs-3955	632	12	.	.	PUNCT
gs-3955	633	1	nature	nature	NOUN
gs-3955	633	2	,	,	PUNCT
gs-3955	633	3	554(7690	554(7690	NUM
gs-3955	633	4	)	)	PUNCT
gs-3955	633	5	,	,	PUNCT
gs-3955	633	6	50	50	NUM
gs-3955	633	7	-	-	SYM
gs-3955	633	8	55	55	NUM
gs-3955	633	9	.	.	PUNCT
gs-3955	634	1	[	[	X
gs-3955	634	2	pubmed	pubmed	X
gs-3955	634	3	:	:	PUNCT
gs-3955	634	4	29364872	29364872	NUM
gs-3955	634	5	]	]	PUNCT
gs-3955	634	6	60	60	NUM
gs-3955	634	7	.	.	PUNCT
gs-3955	634	8	o'brien	o'brien	PROPN
gs-3955	634	9	,	,	PUNCT
gs-3955	634	10	k.m	k.m	PROPN
gs-3955	634	11	.	.	PROPN
gs-3955	634	12	,	,	PUNCT
gs-3955	634	13	et	et	PROPN
gs-3955	634	14	al	al	PROPN
gs-3955	634	15	.	.	PUNCT
gs-3955	634	16	(	(	PUNCT
gs-3955	634	17	2018	2018	NUM
gs-3955	634	18	)	)	PUNCT
gs-3955	634	19	.	.	PUNCT
gs-3955	635	1	mitochondrial	mitochondrial	ADJ
gs-3955	635	2	function	function	NOUN
gs-3955	635	3	in	in	ADP
gs-3955	635	4	long	long	ADV
gs-3955	635	5	-	-	PUNCT
gs-3955	635	6	lived	live	VERB
gs-3955	635	7	rockfishes	rockfish	NOUN
gs-3955	635	8	.	.	PUNCT
gs-3955	636	1	journal	journal	NOUN
gs-3955	636	2	of	of	ADP
gs-3955	636	3	experimental	experimental	ADJ
gs-3955	636	4	biology	biology	NOUN
gs-3955	636	5	,	,	PUNCT
gs-3955	636	6	221(pt	221(pt	NUM
gs-3955	636	7	4	4	NUM
gs-3955	636	8	)	)	PUNCT
gs-3955	636	9	,	,	PUNCT
gs-3955	636	10	jeb172205	jeb172205	NOUN
gs-3955	636	11	.	.	PUNCT
gs-3955	637	1	[	[	X
gs-3955	637	2	pubmed	pubmed	NOUN
gs-3955	637	3	:	:	PUNCT
gs-3955	637	4	29237762	29237762	NUM
gs-3955	637	5	]	]	PUNCT
gs-3955	637	6	61	61	NUM
gs-3955	637	7	.	.	PUNCT
gs-3955	638	1	passow	passow	PROPN
gs-3955	638	2	,	,	PUNCT
gs-3955	638	3	c.	c.	PROPN
gs-3955	638	4	n.	n.	PROPN
gs-3955	638	5	,	,	PUNCT
gs-3955	638	6	et	et	PROPN
gs-3955	638	7	al	al	PROPN
gs-3955	638	8	.	.	PROPN
gs-3955	638	9	(	(	PUNCT
gs-3955	638	10	2022	2022	NUM
gs-3955	638	11	)	)	PUNCT
gs-3955	638	12	.	.	PUNCT
gs-3955	639	1	oxidative	oxidative	ADJ
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gs-3955	639	3	resistance	resistance	NOUN
gs-3955	639	4	in	in	ADP
gs-3955	639	5	the	the	DET
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gs-3955	639	7	shark	shark	NOUN
gs-3955	639	8	.	.	PUNCT
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gs-3955	640	2	radical	radical	ADJ
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gs-3955	640	4	and	and	CCONJ
gs-3955	640	5	medicine	medicine	NOUN
gs-3955	640	6	,	,	PUNCT
gs-3955	640	7	178	178	NUM
gs-3955	640	8	,	,	PUNCT
gs-3955	640	9	1	1	NUM
gs-3955	640	10	-	-	SYM
gs-3955	640	11	10	10	NUM
gs-3955	640	12	.	.	PUNCT
gs-3955	641	1	[	[	X
gs-3955	641	2	pubmed	pubmed	X
gs-3955	641	3	:	:	PUNCT
gs-3955	641	4	34999123	34999123	NUM
gs-3955	641	5	]	]	PUNCT
gs-3955	641	6	62	62	NUM
gs-3955	641	7	.	.	PUNCT
gs-3955	642	1	pearson	pearson	PROPN
gs-3955	642	2	,	,	PUNCT
gs-3955	642	3	b.	b.	PROPN
gs-3955	642	4	j.	j.	PROPN
gs-3955	642	5	,	,	PUNCT
gs-3955	642	6	&	&	CCONJ
gs-3955	642	7	sánchez	sánchez	PROPN
gs-3955	642	8	alvarado	alvarado	PROPN
gs-3955	642	9	,	,	PUNCT
gs-3955	642	10	a.	a.	PROPN
gs-3955	642	11	(	(	PUNCT
gs-3955	642	12	2018	2018	NUM
gs-3955	642	13	)	)	PUNCT
gs-3955	642	14	.	.	PUNCT
gs-3955	643	1	regeneration	regeneration	NOUN
gs-3955	643	2	and	and	CCONJ
gs-3955	643	3	aging	aging	NOUN
gs-3955	643	4	in	in	ADP
gs-3955	643	5	planarians	planarian	NOUN
gs-3955	643	6	.	.	PUNCT
gs-3955	644	1	experimental	experimental	ADJ
gs-3955	644	2	gerontology	gerontology	NOUN
gs-3955	644	3	,	,	PUNCT
gs-3955	644	4	107	107	NUM
gs-3955	644	5	,	,	PUNCT
gs-3955	644	6	1	1	NUM
gs-3955	644	7	-	-	SYM
gs-3955	644	8	7	7	NUM
gs-3955	644	9	.	.	PUNCT
gs-3955	645	1	[	[	X
gs-3955	645	2	pubmed	pubmed	NOUN
gs-3955	645	3	:	:	PUNCT
gs-3955	645	4	29427692	29427692	NUM
gs-3955	645	5	]	]	PUNCT
gs-3955	645	6	63	63	NUM
gs-3955	645	7	.	.	PUNCT
gs-3955	646	1	petruseva	petruseva	PROPN
gs-3955	646	2	io	io	PROPN
gs-3955	646	3	,	,	PUNCT
gs-3955	646	4	evdokimov	evdokimov	PROPN
gs-3955	646	5	an	an	PRON
gs-3955	646	6	,	,	PUNCT
gs-3955	646	7	lavrik	lavrik	NOUN
gs-3955	646	8	oi	oi	INTJ
gs-3955	646	9	.	.	PROPN
gs-3955	646	10	genome	genome	NOUN
gs-3955	646	11	stability	stability	NOUN
gs-3955	646	12	maintenance	maintenance	NOUN
gs-3955	646	13	in	in	ADP
gs-3955	646	14	naked	naked	ADJ
gs-3955	646	15	mole	mole	NOUN
gs-3955	646	16	-	-	PUNCT
gs-3955	646	17	rat	rat	NOUN
gs-3955	646	18	.	.	PROPN
gs-3955	646	19	acta	acta	PROPN
gs-3955	646	20	naturae	naturae	PROPN
gs-3955	646	21	.	.	PUNCT
gs-3955	647	1	2017;9(4):31–41	2017;9(4):31–41	NUM
gs-3955	647	2	.	.	PUNCT
gs-3955	648	1	64	64	NUM
gs-3955	648	2	.	.	PUNCT
gs-3955	649	1	perez	perez	PROPN
gs-3955	649	2	vi	vi	PROPN
gs-3955	649	3	,	,	PUNCT
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gs-3955	649	5	r	r	NOUN
gs-3955	649	6	,	,	PUNCT
gs-3955	649	7	masamsetti	masamsetti	NOUN
gs-3955	649	8	v	v	NOUN
gs-3955	649	9	,	,	PUNCT
gs-3955	649	10	leonard	leonard	PROPN
gs-3955	649	11	s	s	PROPN
gs-3955	649	12	,	,	PUNCT
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gs-3955	649	15	,	,	PUNCT
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gs-3955	649	18	,	,	PUNCT
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gs-3955	649	20	b	b	PROPN
gs-3955	649	21	,	,	PUNCT
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gs-3955	649	23	t	t	PROPN
gs-3955	649	24	,	,	PUNCT
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gs-3955	649	37	resistance	resistance	NOUN
gs-3955	649	38	to	to	PART
gs-3955	649	39	oxidative	oxidative	ADJ
gs-3955	649	40	stress	stress	NOUN
gs-3955	649	41	are	be	AUX
gs-3955	649	42	determinants	determinant	NOUN
gs-3955	649	43	of	of	ADP
gs-3955	649	44	longevity	longevity	NOUN
gs-3955	649	45	in	in	ADP
gs-3955	649	46	the	the	DET
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gs-3955	649	48	-	-	PUNCT
gs-3955	649	49	living	live	VERB
gs-3955	649	50	rodent	rodent	NOUN
gs-3955	649	51	,	,	PUNCT
gs-3955	649	52	the	the	DET
gs-3955	649	53	naked	naked	ADJ
gs-3955	649	54	mole	mole	NOUN
gs-3955	649	55	-	-	PUNCT
gs-3955	649	56	rat	rat	NOUN
gs-3955	649	57	.	.	PROPN
gs-3955	649	58	proc	proc	PROPN
gs-3955	649	59	natl	natl	PROPN
gs-3955	649	60	acad	acad	PROPN
gs-3955	649	61	sci	sci	PROPN
gs-3955	649	62	u	u	PROPN
gs-3955	649	63	s	s	PROPN
gs-3955	649	64	a.	a.	NOUN
gs-3955	649	65	2009;106(9):3059–3064	2009;106(9):3059–3064	NUM
gs-3955	649	66	.	.	PROPN
gs-3955	649	67	65	65	NUM
gs-3955	649	68	.	.	PUNCT
gs-3955	650	1	philipp	philipp	PROPN
gs-3955	650	2	,	,	PUNCT
gs-3955	650	3	e.e	e.e	PROPN
gs-3955	650	4	.	.	PROPN
gs-3955	650	5	,	,	PUNCT
gs-3955	650	6	et	et	PROPN
gs-3955	650	7	al	al	PROPN
gs-3955	650	8	.	.	PUNCT
gs-3955	650	9	(	(	PUNCT
gs-3955	650	10	2012	2012	NUM
gs-3955	650	11	)	)	PUNCT
gs-3955	650	12	.	.	PUNCT
gs-3955	651	1	antioxidant	antioxidant	ADJ
gs-3955	651	2	capacity	capacity	NOUN
gs-3955	651	3	in	in	ADP
gs-3955	651	4	arctica	arctica	PROPN
gs-3955	651	5	islandica	islandica	PROPN
gs-3955	651	6	.	.	PUNCT
gs-3955	652	1	plos	plos	PROPN
gs-3955	652	2	one	one	NUM
gs-3955	652	3	,	,	PUNCT
gs-3955	652	4	7(9	7(9	NUM
gs-3955	652	5	)	)	PUNCT
gs-3955	652	6	,	,	PUNCT
gs-3955	652	7	e44654	e44654	PROPN
gs-3955	652	8	.	.	PUNCT
gs-3955	653	1	[	[	X
gs-3955	653	2	pubmed	pubmed	X
gs-3955	653	3	:	:	PUNCT
gs-3955	653	4	22970282	22970282	NUM
gs-3955	653	5	]	]	PUNCT
gs-3955	654	1	66	66	NUM
gs-3955	654	2	.	.	PUNCT
gs-3955	655	1	philipp	philipp	PROPN
gs-3955	655	2	,	,	PUNCT
gs-3955	655	3	e.e	e.e	PROPN
gs-3955	655	4	.	.	PROPN
gs-3955	655	5	,	,	PUNCT
gs-3955	655	6	et	et	PROPN
gs-3955	655	7	al	al	PROPN
gs-3955	655	8	.	.	PUNCT
gs-3955	655	9	(	(	PUNCT
gs-3955	655	10	2015	2015	NUM
gs-3955	655	11	)	)	PUNCT
gs-3955	655	12	.	.	PUNCT
gs-3955	656	1	antioxidant	antioxidant	ADJ
gs-3955	656	2	capacity	capacity	NOUN
gs-3955	656	3	in	in	ADP
gs-3955	656	4	marine	marine	ADJ
gs-3955	656	5	invertebrates	invertebrate	NOUN
gs-3955	656	6	.	.	PUNCT
gs-3955	657	1	plos	plos	PROPN
gs-3955	657	2	one	one	NUM
gs-3955	657	3	,	,	PUNCT
gs-3955	657	4	10(7	10(7	NUM
gs-3955	657	5	)	)	PUNCT
gs-3955	657	6	,	,	PUNCT
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gs-3955	657	8	.	.	PUNCT
gs-3955	658	1	[	[	X
gs-3955	658	2	pubmed	pubmed	X
gs-3955	658	3	:	:	PUNCT
gs-3955	658	4	26186533	26186533	NUM
gs-3955	658	5	]	]	PUNCT
gs-3955	659	1	67	67	NUM
gs-3955	659	2	.	.	PUNCT
gs-3955	659	3	pirotte	pirotte	PROPN
gs-3955	659	4	,	,	PUNCT
gs-3955	659	5	n.	n.	PROPN
gs-3955	659	6	,	,	PUNCT
gs-3955	659	7	et	et	PROPN
gs-3955	659	8	al	al	PROPN
gs-3955	659	9	.	.	PUNCT
gs-3955	659	10	(	(	PUNCT
gs-3955	659	11	2015	2015	NUM
gs-3955	659	12	)	)	PUNCT
gs-3955	659	13	.	.	PUNCT
gs-3955	660	1	reactive	reactive	ADJ
gs-3955	660	2	oxygen	oxygen	NOUN
gs-3955	660	3	species	specie	NOUN
gs-3955	660	4	in	in	ADP
gs-3955	660	5	planarian	planarian	ADJ
gs-3955	660	6	regeneration	regeneration	NOUN
gs-3955	660	7	.	.	PUNCT
gs-3955	661	1	redox	redox	ADJ
gs-3955	661	2	biology	biology	NOUN
gs-3955	661	3	,	,	PUNCT
gs-3955	661	4	6	6	NUM
gs-3955	661	5	,	,	PUNCT
gs-3955	661	6	1	1	NUM
gs-3955	661	7	-	-	SYM
gs-3955	661	8	10	10	NUM
gs-3955	661	9	.	.	PUNCT
gs-3955	662	1	[	[	X
gs-3955	662	2	pubmed	pubmed	X
gs-3955	662	3	:	:	PUNCT
gs-3955	662	4	26262993	26262993	NUM
gs-3955	662	5	]	]	PUNCT
gs-3955	662	6	georgian	georgian	PROPN
gs-3955	662	7	scientists/ქართველი	scientists/ქართველი	PROPN
gs-3955	662	8	მეცნიერები	მეცნიერები	NOUN
gs-3955	662	9	ტ.	ტ.	VERB
gs-3955	662	10	7	7	NUM
gs-3955	662	11	n	n	PRON
gs-3955	662	12	3	3	NUM
gs-3955	662	13	,	,	PUNCT
gs-3955	662	14	2025	2025	NUM
gs-3955	662	15	393	393	NUM
gs-3955	662	16	68	68	NUM
gs-3955	662	17	.	.	PUNCT
gs-3955	663	1	reddien	reddien	PROPN
gs-3955	663	2	,	,	PUNCT
gs-3955	663	3	p.	p.	PROPN
gs-3955	663	4	w.	w.	PROPN
gs-3955	663	5	(	(	PUNCT
gs-3955	663	6	2018	2018	NUM
gs-3955	663	7	)	)	PUNCT
gs-3955	663	8	.	.	PUNCT
gs-3955	664	1	the	the	DET
gs-3955	664	2	cellular	cellular	ADJ
gs-3955	664	3	and	and	CCONJ
gs-3955	664	4	molecular	molecular	ADJ
gs-3955	664	5	basis	basis	NOUN
gs-3955	664	6	for	for	ADP
gs-3955	664	7	planarian	planarian	ADJ
gs-3955	664	8	regeneration	regeneration	NOUN
gs-3955	664	9	.	.	PUNCT
gs-3955	665	1	cell	cell	NOUN
gs-3955	665	2	,	,	PUNCT
gs-3955	665	3	175(2	175(2	NUM
gs-3955	665	4	)	)	PUNCT
gs-3955	665	5	,	,	PUNCT
gs-3955	665	6	327	327	NUM
gs-3955	665	7	-	-	SYM
gs-3955	665	8	345	345	NUM
gs-3955	665	9	.	.	PUNCT
gs-3955	666	1	[	[	X
gs-3955	666	2	pubmed	pubmed	NOUN
gs-3955	666	3	:	:	PUNCT
gs-3955	666	4	30340035	30340035	NUM
gs-3955	666	5	]	]	PUNCT
gs-3955	666	6	69	69	NUM
gs-3955	666	7	.	.	PUNCT
gs-3955	667	1	reinke	reinke	PROPN
gs-3955	667	2	,	,	PUNCT
gs-3955	667	3	b.a	b.a	PROPN
gs-3955	667	4	.	.	PROPN
gs-3955	667	5	,	,	PUNCT
gs-3955	667	6	et	et	PROPN
gs-3955	667	7	al	al	PROPN
gs-3955	667	8	.	.	PROPN
gs-3955	667	9	(	(	PUNCT
gs-3955	667	10	2022	2022	NUM
gs-3955	667	11	)	)	PUNCT
gs-3955	667	12	.	.	PUNCT
gs-3955	668	1	diverse	diverse	ADJ
gs-3955	668	2	aging	age	VERB
gs-3955	668	3	rates	rate	NOUN
gs-3955	668	4	in	in	ADP
gs-3955	668	5	ectothermic	ectothermic	ADJ
gs-3955	668	6	tetrapods	tetrapod	NOUN
gs-3955	668	7	provide	provide	VERB
gs-3955	668	8	insights	insight	NOUN
gs-3955	668	9	for	for	ADP
gs-3955	668	10	the	the	DET
gs-3955	668	11	evolution	evolution	NOUN
gs-3955	668	12	of	of	ADP
gs-3955	668	13	aging	aging	NOUN
gs-3955	668	14	and	and	CCONJ
gs-3955	668	15	longevity	longevity	NOUN
gs-3955	668	16	.	.	PUNCT
gs-3955	669	1	science	science	NOUN
gs-3955	669	2	,	,	PUNCT
gs-3955	669	3	376(6600	376(6600	NUM
gs-3955	669	4	)	)	PUNCT
gs-3955	669	5	,	,	PUNCT
gs-3955	669	6	1459	1459	NUM
gs-3955	669	7	-	-	SYM
gs-3955	669	8	1466	1466	NUM
gs-3955	669	9	.	.	PUNCT
gs-3955	670	1	[	[	X
gs-3955	670	2	pubmed	pubmed	X
gs-3955	670	3	:	:	PUNCT
gs-3955	670	4	35737773	35737773	NUM
gs-3955	670	5	]	]	PUNCT
gs-3955	671	1	70	70	NUM
gs-3955	671	2	.	.	PUNCT
gs-3955	671	3	rink	rink	NOUN
gs-3955	671	4	,	,	PUNCT
gs-3955	671	5	j.	j.	PROPN
gs-3955	671	6	c.	c.	PROPN
gs-3955	671	7	(	(	PUNCT
gs-3955	671	8	2013	2013	NUM
gs-3955	671	9	)	)	PUNCT
gs-3955	671	10	.	.	PUNCT
gs-3955	672	1	stem	stem	NOUN
gs-3955	672	2	cell	cell	NOUN
gs-3955	672	3	systems	system	NOUN
gs-3955	672	4	and	and	CCONJ
gs-3955	672	5	regeneration	regeneration	NOUN
gs-3955	672	6	in	in	ADP
gs-3955	672	7	planaria	planaria	NOUN
gs-3955	672	8	.	.	PUNCT
gs-3955	673	1	developmental	developmental	ADJ
gs-3955	673	2	dynamics	dynamic	NOUN
gs-3955	673	3	,	,	PUNCT
gs-3955	673	4	242(2	242(2	NUM
gs-3955	673	5	)	)	PUNCT
gs-3955	673	6	,	,	PUNCT
gs-3955	673	7	130	130	NUM
gs-3955	673	8	-	-	SYM
gs-3955	673	9	144	144	NUM
gs-3955	673	10	.	.	PUNCT
gs-3955	674	1	[	[	X
gs-3955	674	2	pubmed	pubmed	NOUN
gs-3955	674	3	:	:	PUNCT
gs-3955	674	4	23203693	23203693	NUM
gs-3955	674	5	]	]	PUNCT
gs-3955	674	6	71	71	NUM
gs-3955	674	7	.	.	PUNCT
gs-3955	675	1	sahu	sahu	PROPN
gs-3955	675	2	,	,	PUNCT
gs-3955	675	3	s.	s.	PROPN
gs-3955	675	4	,	,	PUNCT
gs-3955	675	5	et	et	PROPN
gs-3955	675	6	al	al	PROPN
gs-3955	675	7	.	.	PUNCT
gs-3955	675	8	(	(	PUNCT
gs-3955	675	9	2017	2017	NUM
gs-3955	675	10	)	)	PUNCT
gs-3955	675	11	.	.	PUNCT
gs-3955	676	1	long	long	ADJ
gs-3955	676	2	-	-	PUNCT
gs-3955	676	3	term	term	NOUN
gs-3955	676	4	culture	culture	NOUN
gs-3955	676	5	of	of	ADP
gs-3955	676	6	planarians	planarian	NOUN
gs-3955	676	7	.	.	PUNCT
gs-3955	677	1	nature	nature	NOUN
gs-3955	677	2	protocols	protocol	NOUN
gs-3955	677	3	,	,	PUNCT
gs-3955	677	4	12(8	12(8	NUM
gs-3955	677	5	)	)	PUNCT
gs-3955	677	6	,	,	PUNCT
gs-3955	677	7	1528	1528	NUM
gs-3955	677	8	-	-	SYM
gs-3955	677	9	1540	1540	NUM
gs-3955	677	10	.	.	PUNCT
gs-3955	678	1	[	[	X
gs-3955	678	2	pubmed	pubmed	NOUN
gs-3955	678	3	:	:	PUNCT
gs-3955	678	4	28703791	28703791	NUM
gs-3955	678	5	]	]	PUNCT
gs-3955	678	6	72	72	NUM
gs-3955	678	7	.	.	PUNCT
gs-3955	679	1	seim	seim	PROPN
gs-3955	679	2	,	,	PUNCT
gs-3955	679	3	i.	i.	PROPN
gs-3955	679	4	,	,	PUNCT
gs-3955	679	5	et	et	PROPN
gs-3955	679	6	al	al	PROPN
gs-3955	679	7	.	.	PROPN
gs-3955	679	8	(	(	PUNCT
gs-3955	679	9	2023	2023	NUM
gs-3955	679	10	)	)	PUNCT
gs-3955	679	11	.	.	PUNCT
gs-3955	680	1	genomic	genomic	ADJ
gs-3955	680	2	adaptations	adaptation	NOUN
gs-3955	680	3	to	to	ADP
gs-3955	680	4	extreme	extreme	ADJ
gs-3955	680	5	longevity	longevity	NOUN
gs-3955	680	6	in	in	ADP
gs-3955	680	7	the	the	DET
gs-3955	680	8	greenland	greenland	NOUN
gs-3955	680	9	shark	shark	NOUN
gs-3955	680	10	.	.	PUNCT
gs-3955	681	1	nature	nature	NOUN
gs-3955	681	2	communications	communication	NOUN
gs-3955	681	3	,	,	PUNCT
gs-3955	681	4	14(1	14(1	NUM
gs-3955	681	5	)	)	PUNCT
gs-3955	681	6	,	,	PUNCT
gs-3955	681	7	1234	1234	NUM
gs-3955	681	8	.	.	PUNCT
gs-3955	682	1	[	[	X
gs-3955	682	2	pubmed	pubmed	X
gs-3955	682	3	:	:	PUNCT
gs-3955	682	4	36823245	36823245	NUM
gs-3955	682	5	]	]	PUNCT
gs-3955	682	6	73	73	NUM
gs-3955	682	7	.	.	PUNCT
gs-3955	683	1	seluanov	seluanov	PROPN
gs-3955	683	2	,	,	PUNCT
gs-3955	683	3	a.	a.	PROPN
gs-3955	683	4	,	,	PUNCT
gs-3955	683	5	et	et	PROPN
gs-3955	683	6	al	al	PROPN
gs-3955	683	7	.	.	PUNCT
gs-3955	684	1	(	(	PUNCT
gs-3955	684	2	2018	2018	NUM
gs-3955	684	3	)	)	PUNCT
gs-3955	684	4	.	.	PUNCT
gs-3955	685	1	mechanisms	mechanism	NOUN
gs-3955	685	2	of	of	ADP
gs-3955	685	3	cancer	cancer	NOUN
gs-3955	685	4	resistance	resistance	NOUN
gs-3955	685	5	in	in	ADP
gs-3955	685	6	long	long	ADV
gs-3955	685	7	-	-	PUNCT
gs-3955	685	8	lived	live	VERB
gs-3955	685	9	mammals	mammal	NOUN
gs-3955	685	10	.	.	PUNCT
gs-3955	686	1	nature	nature	NOUN
gs-3955	686	2	reviews	review	NOUN
gs-3955	686	3	cancer	cancer	NOUN
gs-3955	686	4	,	,	PUNCT
gs-3955	686	5	18(7	18(7	NUM
gs-3955	686	6	)	)	PUNCT
gs-3955	686	7	,	,	PUNCT
gs-3955	686	8	433	433	NUM
gs-3955	686	9	-	-	SYM
gs-3955	686	10	441	441	NUM
gs-3955	686	11	.	.	PUNCT
gs-3955	687	1	[	[	X
gs-3955	687	2	pubmed	pubmed	NOUN
gs-3955	687	3	:	:	PUNCT
gs-3955	687	4	29795324	29795324	NUM
gs-3955	687	5	]	]	PUNCT
gs-3955	687	6	74	74	NUM
gs-3955	687	7	.	.	PUNCT
gs-3955	687	8	storey	storey	PROPN
gs-3955	687	9	,	,	PUNCT
gs-3955	687	10	k.b	k.b	PROPN
gs-3955	687	11	.	.	PROPN
gs-3955	687	12	,	,	PUNCT
gs-3955	687	13	&	&	CCONJ
gs-3955	687	14	storey	storey	PROPN
gs-3955	687	15	,	,	PUNCT
gs-3955	687	16	j.m	j.m	PROPN
gs-3955	687	17	.	.	PROPN
gs-3955	687	18	(	(	PUNCT
gs-3955	687	19	2021	2021	NUM
gs-3955	687	20	)	)	PUNCT
gs-3955	687	21	.	.	PUNCT
gs-3955	688	1	metabolic	metabolic	NOUN
gs-3955	688	2	rate	rate	NOUN
gs-3955	688	3	depression	depression	NOUN
gs-3955	688	4	in	in	ADP
gs-3955	688	5	turtles	turtle	NOUN
gs-3955	688	6	:	:	PUNCT
gs-3955	688	7	molecular	molecular	ADJ
gs-3955	688	8	mechanisms	mechanism	NOUN
gs-3955	688	9	and	and	CCONJ
gs-3955	688	10	physiological	physiological	ADJ
gs-3955	688	11	adaptations	adaptation	NOUN
gs-3955	688	12	.	.	PUNCT
gs-3955	689	1	journal	journal	NOUN
gs-3955	689	2	of	of	ADP
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gs-3955	689	4	biology	biology	NOUN
gs-3955	689	5	,	,	PUNCT
gs-3955	689	6	224(pt	224(pt	PROPN
gs-3955	689	7	10	10	NUM
gs-3955	689	8	)	)	PUNCT
gs-3955	689	9	,	,	PUNCT
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gs-3955	689	11	.	.	PUNCT
gs-3955	690	1	[	[	X
gs-3955	690	2	pubmed	pubmed	X
gs-3955	690	3	:	:	PUNCT
gs-3955	690	4	33942106	33942106	NUM
gs-3955	690	5	]	]	PUNCT
gs-3955	690	6	75	75	NUM
gs-3955	690	7	.	.	PUNCT
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gs-3955	690	9	,	,	PUNCT
gs-3955	690	10	j.	j.	PROPN
gs-3955	690	11	,	,	PUNCT
gs-3955	690	12	et	et	PROPN
gs-3955	690	13	al	al	PROPN
gs-3955	690	14	.	.	PUNCT
gs-3955	690	15	(	(	PUNCT
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gs-3955	690	17	)	)	PUNCT
gs-3955	690	18	.	.	PUNCT
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gs-3955	691	2	to	to	AUX
gs-3955	691	3	oxidative	oxidative	ADJ
gs-3955	691	4	stress	stress	NOUN
gs-3955	691	5	in	in	ADP
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gs-3955	691	8	.	.	PUNCT
gs-3955	692	1	journal	journal	PROPN
gs-3955	692	2	of	of	ADP
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gs-3955	692	5	,	,	PUNCT
gs-3955	692	6	214(pt	214(pt	NOUN
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gs-3955	692	8	)	)	PUNCT
gs-3955	692	9	,	,	PUNCT
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gs-3955	692	11	-	-	SYM
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gs-3955	692	13	.	.	PUNCT
gs-3955	693	1	[	[	X
gs-3955	693	2	pubmed	pubmed	X
gs-3955	693	3	:	:	PUNCT
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gs-3955	693	5	]	]	PUNCT
gs-3955	693	6	76	76	NUM
gs-3955	693	7	.	.	PUNCT
gs-3955	693	8	strahlendorf	strahlendorf	PROPN
gs-3955	693	9	pw	pw	PROPN
gs-3955	693	10	,	,	PUNCT
gs-3955	693	11	perez	perez	PROPN
gs-3955	693	12	vi	vi	PROPN
gs-3955	693	13	,	,	PUNCT
gs-3955	693	14	song	song	NOUN
gs-3955	693	15	w	w	PROPN
gs-3955	693	16	,	,	PUNCT
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gs-3955	693	22	a	a	PROPN
gs-3955	693	23	,	,	PUNCT
gs-3955	693	24	epstein	epstein	PROPN
gs-3955	693	25	cj	cj	PROPN
gs-3955	693	26	,	,	PUNCT
gs-3955	693	27	vijg	vijg	PROPN
gs-3955	693	28	j	j	PROPN
gs-3955	693	29	,	,	PUNCT
gs-3955	693	30	richardson	richardson	PROPN
gs-3955	693	31	a.	a.	NOUN
gs-3955	693	32	overexpression	overexpression	NOUN
gs-3955	693	33	of	of	ADP
gs-3955	693	34	mn	mn	PROPN
gs-3955	693	35	superoxide	superoxide	NOUN
gs-3955	693	36	dismutase	dismutase	NOUN
gs-3955	693	37	does	do	AUX
gs-3955	693	38	not	not	PART
gs-3955	693	39	increase	increase	VERB
gs-3955	693	40	life	life	NOUN
gs-3955	693	41	span	span	NOUN
gs-3955	693	42	in	in	ADP
gs-3955	693	43	mice	mouse	NOUN
gs-3955	693	44	.	.	PUNCT
gs-3955	694	1	aging	age	VERB
gs-3955	694	2	cell	cell	NOUN
gs-3955	694	3	.	.	PUNCT
gs-3955	695	1	2008;7(3):383–395	2008;7(3):383–395	X
gs-3955	695	2	.	.	X
gs-3955	695	3	77	77	NUM
gs-3955	695	4	.	.	PUNCT
gs-3955	696	1	sussarellu	sussarellu	NOUN
gs-3955	696	2	,	,	PUNCT
gs-3955	696	3	r.	r.	PROPN
gs-3955	696	4	,	,	PUNCT
gs-3955	696	5	et	et	PROPN
gs-3955	696	6	al	al	PROPN
gs-3955	696	7	.	.	PROPN
gs-3955	696	8	(	(	PUNCT
gs-3955	696	9	2020	2020	NUM
gs-3955	696	10	)	)	PUNCT
gs-3955	696	11	.	.	PUNCT
gs-3955	697	1	dna	dna	PROPN
gs-3955	697	2	repair	repair	VERB
gs-3955	697	3	in	in	ADP
gs-3955	697	4	long	long	ADV
gs-3955	697	5	-	-	PUNCT
gs-3955	697	6	lived	live	VERB
gs-3955	697	7	bivalves	bivalve	NOUN
gs-3955	697	8	.	.	PUNCT
gs-3955	698	1	marine	marine	ADJ
gs-3955	698	2	genomics	genomic	NOUN
gs-3955	698	3	,	,	PUNCT
gs-3955	698	4	51	51	NUM
gs-3955	698	5	,	,	PUNCT
gs-3955	698	6	100736	100736	NUM
gs-3955	698	7	.	.	PUNCT
gs-3955	699	1	[	[	X
gs-3955	699	2	pubmed	pubmed	X
gs-3955	699	3	:	:	PUNCT
gs-3955	699	4	32081467	32081467	NUM
gs-3955	699	5	]	]	PUNCT
gs-3955	699	6	78	78	NUM
gs-3955	699	7	.	.	PUNCT
gs-3955	700	1	sussarellu	sussarellu	NOUN
gs-3955	700	2	,	,	PUNCT
gs-3955	700	3	r.	r.	PROPN
gs-3955	700	4	,	,	PUNCT
gs-3955	700	5	et	et	PROPN
gs-3955	700	6	al	al	PROPN
gs-3955	700	7	.	.	PROPN
gs-3955	700	8	(	(	PUNCT
gs-3955	700	9	2020	2020	NUM
gs-3955	700	10	)	)	PUNCT
gs-3955	700	11	.	.	PUNCT
gs-3955	701	1	dna	dna	PROPN
gs-3955	701	2	repair	repair	VERB
gs-3955	701	3	in	in	ADP
gs-3955	701	4	long	long	ADV
gs-3955	701	5	-	-	PUNCT
gs-3955	701	6	lived	live	VERB
gs-3955	701	7	marine	marine	ADJ
gs-3955	701	8	species	specie	NOUN
gs-3955	701	9	.	.	PUNCT
gs-3955	702	1	marine	marine	ADJ
gs-3955	702	2	genomics	genomics	PROPN
gs-3955	702	3	,	,	PUNCT
gs-3955	702	4	51	51	NUM
gs-3955	702	5	,	,	PUNCT
gs-3955	702	6	100736	100736	NUM
gs-3955	702	7	.	.	PUNCT
gs-3955	703	1	[	[	X
gs-3955	703	2	pubmed	pubmed	X
gs-3955	703	3	:	:	PUNCT
gs-3955	703	4	32081467	32081467	NUM
gs-3955	703	5	]	]	PUNCT
gs-3955	703	6	79	79	NUM
gs-3955	703	7	.	.	PUNCT
gs-3955	703	8	tacutu	tacutu	PROPN
gs-3955	703	9	,	,	PUNCT
gs-3955	703	10	r.	r.	PROPN
gs-3955	703	11	,	,	PUNCT
gs-3955	703	12	et	et	PROPN
gs-3955	703	13	al	al	PROPN
gs-3955	703	14	.	.	PROPN
gs-3955	704	1	(	(	PUNCT
gs-3955	704	2	2013	2013	NUM
gs-3955	704	3	)	)	PUNCT
gs-3955	704	4	.	.	PUNCT
gs-3955	705	1	human	human	ADJ
gs-3955	705	2	ageing	age	VERB
gs-3955	705	3	genomic	genomic	ADJ
gs-3955	705	4	resources	resource	NOUN
gs-3955	705	5	:	:	PUNCT
gs-3955	705	6	integrated	integrate	VERB
gs-3955	705	7	databases	database	NOUN
gs-3955	705	8	and	and	CCONJ
gs-3955	705	9	tools	tool	NOUN
gs-3955	705	10	for	for	ADP
gs-3955	705	11	the	the	DET
gs-3955	705	12	biology	biology	NOUN
gs-3955	705	13	and	and	CCONJ
gs-3955	705	14	genetics	genetic	NOUN
gs-3955	705	15	of	of	ADP
gs-3955	705	16	ageing	age	VERB
gs-3955	705	17	.	.	PUNCT
gs-3955	706	1	nucleic	nucleic	ADJ
gs-3955	706	2	acids	acid	NOUN
gs-3955	706	3	research	research	NOUN
gs-3955	706	4	,	,	PUNCT
gs-3955	706	5	41(d1	41(d1	NOUN
gs-3955	706	6	)	)	PUNCT
gs-3955	706	7	,	,	PUNCT
gs-3955	706	8	d1027	d1027	NOUN
gs-3955	706	9	-	-	PUNCT
gs-3955	706	10	d1033	d1033	PROPN
gs-3955	706	11	.	.	PUNCT
gs-3955	707	1	[	[	X
gs-3955	707	2	pubmed	pubmed	NOUN
gs-3955	707	3	:	:	PUNCT
gs-3955	707	4	23193293	23193293	NUM
gs-3955	707	5	]	]	PUNCT
gs-3955	707	6	80	80	NUM
gs-3955	707	7	.	.	PUNCT
gs-3955	708	1	tan	tan	PROPN
gs-3955	708	2	,	,	PUNCT
gs-3955	708	3	t.	t.	PROPN
gs-3955	708	4	c.	c.	PROPN
gs-3955	708	5	j.	j.	PROPN
gs-3955	708	6	,	,	PUNCT
gs-3955	708	7	et	et	PROPN
gs-3955	708	8	al	al	PROPN
gs-3955	708	9	.	.	PUNCT
gs-3955	709	1	(	(	PUNCT
gs-3955	709	2	2012	2012	NUM
gs-3955	709	3	)	)	PUNCT
gs-3955	709	4	.	.	PUNCT
gs-3955	710	1	telomere	telomere	NOUN
gs-3955	710	2	maintenance	maintenance	NOUN
gs-3955	710	3	in	in	ADP
gs-3955	710	4	planarians	planarian	NOUN
gs-3955	710	5	.	.	PUNCT
gs-3955	711	1	proceedings	proceeding	NOUN
gs-3955	711	2	of	of	ADP
gs-3955	711	3	the	the	DET
gs-3955	711	4	national	national	PROPN
gs-3955	711	5	academy	academy	PROPN
gs-3955	711	6	of	of	ADP
gs-3955	711	7	sciences	sciences	PROPN
gs-3955	711	8	,	,	PUNCT
gs-3955	711	9	109(27	109(27	NUM
gs-3955	711	10	)	)	PUNCT
gs-3955	711	11	,	,	PUNCT
gs-3955	711	12	10881	10881	NUM
gs-3955	711	13	-	-	SYM
gs-3955	711	14	10886	10886	NUM
gs-3955	711	15	.	.	PUNCT
gs-3955	712	1	[	[	X
gs-3955	712	2	pubmed	pubmed	NOUN
gs-3955	712	3	:	:	PUNCT
gs-3955	712	4	22711824	22711824	NUM
gs-3955	712	5	]	]	PUNCT
gs-3955	713	1	81	81	NUM
gs-3955	713	2	.	.	PUNCT
gs-3955	713	3	tian	tian	ADJ
gs-3955	713	4	,	,	PUNCT
gs-3955	713	5	x.	x.	PROPN
gs-3955	713	6	,	,	PUNCT
gs-3955	713	7	et	et	PROPN
gs-3955	713	8	al	al	PROPN
gs-3955	713	9	.	.	PROPN
gs-3955	713	10	(	(	PUNCT
gs-3955	713	11	2013	2013	NUM
gs-3955	713	12	)	)	PUNCT
gs-3955	713	13	.	.	PUNCT
gs-3955	714	1	high	high	ADJ
gs-3955	714	2	-	-	PUNCT
gs-3955	714	3	molecular	molecular	ADJ
gs-3955	714	4	-	-	PUNCT
gs-3955	714	5	mass	mass	NOUN
gs-3955	714	6	hyaluronan	hyaluronan	PROPN
gs-3955	714	7	mediates	mediate	VERB
gs-3955	714	8	the	the	DET
gs-3955	714	9	cancer	cancer	NOUN
gs-3955	714	10	resistance	resistance	NOUN
gs-3955	714	11	of	of	ADP
gs-3955	714	12	the	the	DET
gs-3955	714	13	naked	naked	ADJ
gs-3955	714	14	mole	mole	NOUN
gs-3955	714	15	rat	rat	NOUN
gs-3955	714	16	.	.	PROPN
gs-3955	714	17	nature	nature	PROPN
gs-3955	714	18	,	,	PUNCT
gs-3955	714	19	499(7458	499(7458	NOUN
gs-3955	714	20	)	)	PUNCT
gs-3955	714	21	,	,	PUNCT
gs-3955	714	22	346	346	NUM
gs-3955	714	23	-	-	SYM
gs-3955	714	24	349	349	NUM
gs-3955	714	25	.	.	PUNCT
gs-3955	715	1	[	[	X
gs-3955	715	2	pubmed	pubmed	X
gs-3955	715	3	:	:	PUNCT
gs-3955	715	4	23868258	23868258	NUM
gs-3955	715	5	]	]	PUNCT
gs-3955	715	6	82	82	NUM
gs-3955	715	7	.	.	PUNCT
gs-3955	715	8	tian	tian	ADJ
gs-3955	715	9	,	,	PUNCT
gs-3955	715	10	x.	x.	PROPN
gs-3955	715	11	,	,	PUNCT
gs-3955	715	12	et	et	PROPN
gs-3955	715	13	al	al	PROPN
gs-3955	715	14	.	.	PROPN
gs-3955	715	15	(	(	PUNCT
gs-3955	715	16	2022	2022	NUM
gs-3955	715	17	)	)	PUNCT
gs-3955	715	18	.	.	PUNCT
gs-3955	716	1	genome	genome	NOUN
gs-3955	716	2	sequencing	sequencing	NOUN
gs-3955	716	3	of	of	ADP
gs-3955	716	4	long	long	ADV
gs-3955	716	5	-	-	PUNCT
gs-3955	716	6	lived	live	VERB
gs-3955	716	7	turtles	turtle	NOUN
gs-3955	716	8	reveals	reveal	VERB
gs-3955	716	9	insights	insight	NOUN
gs-3955	716	10	into	into	ADP
gs-3955	716	11	the	the	DET
gs-3955	716	12	evolution	evolution	NOUN
gs-3955	716	13	of	of	ADP
gs-3955	716	14	aging	aging	NOUN
gs-3955	716	15	.	.	PUNCT
gs-3955	717	1	science	science	NOUN
gs-3955	717	2	advances	advance	NOUN
gs-3955	717	3	,	,	PUNCT
gs-3955	717	4	8(5	8(5	NUM
gs-3955	717	5	)	)	PUNCT
gs-3955	717	6	,	,	PUNCT
gs-3955	717	7	eabm6871	eabm6871	PROPN
gs-3955	717	8	.	.	PUNCT
gs-3955	718	1	[	[	X
gs-3955	718	2	pubmed	pubmed	NOUN
gs-3955	718	3	:	:	PUNCT
gs-3955	718	4	35108055	35108055	NUM
gs-3955	718	5	]	]	PUNCT
gs-3955	718	6	83	83	NUM
gs-3955	718	7	.	.	PUNCT
gs-3955	718	8	tkemaladze	tkemaladze	PROPN
gs-3955	718	9	,	,	PUNCT
gs-3955	718	10	j.	j.	PROPN
gs-3955	718	11	(	(	PUNCT
gs-3955	718	12	2023	2023	NUM
gs-3955	718	13	)	)	PUNCT
gs-3955	718	14	.	.	PUNCT
gs-3955	719	1	reduction	reduction	NOUN
gs-3955	719	2	,	,	PUNCT
gs-3955	719	3	proliferation	proliferation	NOUN
gs-3955	719	4	,	,	PUNCT
gs-3955	719	5	and	and	CCONJ
gs-3955	719	6	differentiation	differentiation	NOUN
gs-3955	719	7	defects	defect	NOUN
gs-3955	719	8	of	of	ADP
gs-3955	719	9	stem	stem	NOUN
gs-3955	719	10	cells	cell	NOUN
gs-3955	719	11	over	over	ADP
gs-3955	719	12	time	time	NOUN
gs-3955	719	13	:	:	PUNCT
gs-3955	719	14	a	a	DET
gs-3955	719	15	consequence	consequence	NOUN
gs-3955	719	16	of	of	ADP
gs-3955	719	17	selective	selective	ADJ
gs-3955	719	18	accumulation	accumulation	NOUN
gs-3955	719	19	of	of	ADP
gs-3955	719	20	old	old	ADJ
gs-3955	719	21	centrioles	centriole	NOUN
gs-3955	719	22	in	in	ADP
gs-3955	719	23	the	the	DET
gs-3955	719	24	stem	stem	NOUN
gs-3955	719	25	cells	cell	NOUN
gs-3955	719	26	?	?	PUNCT
gs-3955	719	27	.	.	PUNCT
gs-3955	720	1	molecular	molecular	ADJ
gs-3955	720	2	biology	biology	NOUN
gs-3955	720	3	reports	report	NOUN
gs-3955	720	4	,	,	PUNCT
gs-3955	720	5	50(3	50(3	NUM
gs-3955	720	6	)	)	PUNCT
gs-3955	720	7	,	,	PUNCT
gs-3955	720	8	2751	2751	NUM
gs-3955	720	9	-	-	SYM
gs-3955	720	10	2761	2761	NUM
gs-3955	720	11	.	.	PUNCT
gs-3955	721	1	84	84	NUM
gs-3955	721	2	.	.	PUNCT
gs-3955	721	3	tkemaladze	tkemaladze	PROPN
gs-3955	721	4	,	,	PUNCT
gs-3955	721	5	j.	j.	PROPN
gs-3955	721	6	(	(	PUNCT
gs-3955	721	7	2024	2024	NUM
gs-3955	721	8	)	)	PUNCT
gs-3955	721	9	.	.	PUNCT
gs-3955	722	1	editorial	editorial	NOUN
gs-3955	722	2	:	:	PUNCT
gs-3955	722	3	molecular	molecular	ADJ
gs-3955	722	4	mechanism	mechanism	NOUN
gs-3955	722	5	of	of	ADP
gs-3955	722	6	ageing	age	VERB
gs-3955	722	7	and	and	CCONJ
gs-3955	722	8	therapeutic	therapeutic	ADJ
gs-3955	722	9	advances	advance	NOUN
gs-3955	722	10	through	through	ADP
gs-3955	722	11	targeting	target	VERB
gs-3955	722	12	glycative	glycative	ADJ
gs-3955	722	13	and	and	CCONJ
gs-3955	722	14	oxidative	oxidative	ADJ
gs-3955	722	15	stress	stress	NOUN
gs-3955	722	16	.	.	PUNCT
gs-3955	723	1	front	front	ADJ
gs-3955	723	2	pharmacol	pharmacol	PROPN
gs-3955	723	3	.	.	PUNCT
gs-3955	724	1	2024	2024	NUM
gs-3955	724	2	georgian	georgian	PROPN
gs-3955	724	3	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	724	4	მეცნიერები	მეცნიერები	NOUN
gs-3955	724	5	ტ.	ტ.	VERB
gs-3955	724	6	7	7	NUM
gs-3955	724	7	n	n	PRON
gs-3955	724	8	3	3	NUM
gs-3955	724	9	,	,	PUNCT
gs-3955	724	10	2025	2025	NUM
gs-3955	724	11	394	394	NUM
gs-3955	724	12	mar	mar	PROPN
gs-3955	724	13	6;14:1324446	6;14:1324446	PROPN
gs-3955	724	14	.	.	PUNCT
gs-3955	725	1	doi	doi	NOUN
gs-3955	725	2	:	:	PUNCT
gs-3955	725	3	10.3389	10.3389	NUM
gs-3955	725	4	/	/	SYM
gs-3955	725	5	fphar.2023.1324446	fphar.2023.1324446	NOUN
gs-3955	725	6	.	.	PUNCT
gs-3955	726	1	pmid	pmid	NOUN
gs-3955	726	2	:	:	PUNCT
gs-3955	726	3	38510429	38510429	NUM
gs-3955	726	4	;	;	PUNCT
gs-3955	726	5	pmcid	pmcid	ADJ
gs-3955	726	6	:	:	PUNCT
gs-3955	726	7	pmc10953819	pmc10953819	NOUN
gs-3955	726	8	.	.	PUNCT
gs-3955	727	1	85	85	NUM
gs-3955	727	2	.	.	PUNCT
gs-3955	728	1	treberg	treberg	PROPN
gs-3955	728	2	,	,	PUNCT
gs-3955	728	3	j.	j.	PROPN
gs-3955	728	4	r.	r.	PROPN
gs-3955	728	5	,	,	PUNCT
gs-3955	728	6	&	&	CCONJ
gs-3955	728	7	speers	speer	NOUN
gs-3955	728	8	-	-	PUNCT
gs-3955	728	9	roesch	roesch	PROPN
gs-3955	728	10	,	,	PUNCT
gs-3955	728	11	b.	b.	PROPN
gs-3955	728	12	(	(	PUNCT
gs-3955	728	13	2016	2016	NUM
gs-3955	728	14	)	)	PUNCT
gs-3955	728	15	.	.	PUNCT
gs-3955	729	1	metabolic	metabolic	NOUN
gs-3955	729	2	adaptations	adaptation	NOUN
gs-3955	729	3	of	of	ADP
gs-3955	729	4	polar	polar	ADJ
gs-3955	729	5	fishes	fish	NOUN
gs-3955	729	6	.	.	PUNCT
gs-3955	730	1	journal	journal	NOUN
gs-3955	730	2	of	of	ADP
gs-3955	730	3	experimental	experimental	ADJ
gs-3955	730	4	biology	biology	NOUN
gs-3955	730	5	,	,	PUNCT
gs-3955	730	6	219(8	219(8	NUM
gs-3955	730	7	)	)	PUNCT
gs-3955	730	8	,	,	PUNCT
gs-3955	730	9	1093	1093	NUM
gs-3955	730	10	-	-	SYM
gs-3955	730	11	1105	1105	NUM
gs-3955	730	12	.	.	PUNCT
gs-3955	731	1	[	[	X
gs-3955	731	2	pubmed	pubmed	X
gs-3955	731	3	:	:	PUNCT
gs-3955	731	4	27103677	27103677	NUM
gs-3955	731	5	]	]	PUNCT
gs-3955	731	6	86	86	NUM
gs-3955	731	7	.	.	PUNCT
gs-3955	731	8	trontelj	trontelj	NOUN
gs-3955	731	9	,	,	PUNCT
gs-3955	731	10	p.	p.	PROPN
gs-3955	731	11	,	,	PUNCT
gs-3955	731	12	et	et	PROPN
gs-3955	731	13	al	al	PROPN
gs-3955	731	14	.	.	PROPN
gs-3955	732	1	(	(	PUNCT
gs-3955	732	2	2022	2022	NUM
gs-3955	732	3	)	)	PUNCT
gs-3955	732	4	.	.	PUNCT
gs-3955	733	1	the	the	DET
gs-3955	733	2	evolutionary	evolutionary	ADJ
gs-3955	733	3	origins	origin	NOUN
gs-3955	733	4	of	of	ADP
gs-3955	733	5	extreme	extreme	ADJ
gs-3955	733	6	longevity	longevity	NOUN
gs-3955	733	7	in	in	ADP
gs-3955	733	8	cavedwelling	cavedwelle	VERB
gs-3955	733	9	salamanders	salamander	NOUN
gs-3955	733	10	.	.	PUNCT
gs-3955	734	1	nature	nature	NOUN
gs-3955	734	2	communications	communication	NOUN
gs-3955	734	3	,	,	PUNCT
gs-3955	734	4	13(1	13(1	NUM
gs-3955	734	5	)	)	PUNCT
gs-3955	734	6	,	,	PUNCT
gs-3955	734	7	1234	1234	NUM
gs-3955	734	8	.	.	PUNCT
gs-3955	735	1	[	[	X
gs-3955	735	2	pubmed	pubmed	X
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gs-3955	735	5	]	]	PUNCT
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gs-3955	737	6	)	)	PUNCT
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gs-3955	738	3	:	:	PUNCT
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gs-3955	738	7	.	.	PUNCT
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gs-3955	738	10	z.	z.	PROPN
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gs-3955	739	6	.	.	PUNCT
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gs-3955	740	6	)	)	PUNCT
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gs-3955	741	2	pubmed	pubmed	X
gs-3955	741	3	:	:	PUNCT
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gs-3955	741	5	]	]	PUNCT
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gs-3955	741	7	.	.	PUNCT
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gs-3955	741	15	(	(	PUNCT
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gs-3955	741	17	)	)	PUNCT
gs-3955	741	18	.	.	PUNCT
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gs-3955	742	10	)	)	PUNCT
gs-3955	742	11	.	.	PUNCT
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gs-3955	743	5	)	)	PUNCT
gs-3955	743	6	,	,	PUNCT
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gs-3955	743	8	.	.	PUNCT
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gs-3955	744	2	pubmed	pubmed	X
gs-3955	744	3	:	:	PUNCT
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gs-3955	744	5	]	]	PUNCT
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gs-3955	744	7	.	.	PUNCT
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gs-3955	744	9	,	,	PUNCT
gs-3955	744	10	d.	d.	PROPN
gs-3955	744	11	e.	e.	PROPN
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gs-3955	744	13	et	et	PROPN
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gs-3955	744	16	(	(	PUNCT
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gs-3955	744	18	)	)	PUNCT
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gs-3955	745	5	adult	adult	NOUN
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gs-3955	745	8	.	.	PUNCT
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gs-3955	746	2	,	,	PUNCT
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gs-3955	746	4	)	)	PUNCT
gs-3955	746	5	,	,	PUNCT
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gs-3955	746	7	-	-	SYM
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gs-3955	746	9	.	.	PUNCT
gs-3955	747	1	[	[	X
gs-3955	747	2	pubmed	pubmed	X
gs-3955	747	3	:	:	PUNCT
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gs-3955	747	5	]	]	SYM
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gs-3955	747	7	.	.	PUNCT
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gs-3955	748	2	,	,	PUNCT
gs-3955	748	3	k.	k.	PROPN
gs-3955	748	4	d.	d.	PROPN
gs-3955	748	5	,	,	PUNCT
gs-3955	748	6	et	et	PROPN
gs-3955	748	7	al	al	PROPN
gs-3955	748	8	.	.	PROPN
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gs-3955	748	11	)	)	PUNCT
gs-3955	748	12	.	.	PUNCT
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gs-3955	749	8	.	.	PUNCT
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gs-3955	750	6	,	,	PUNCT
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gs-3955	750	8	,	,	PUNCT
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gs-3955	750	12	.	.	PUNCT
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gs-3955	751	2	pubmed	pubmed	X
gs-3955	751	3	:	:	PUNCT
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gs-3955	751	5	]	]	PUNCT
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gs-3955	751	7	.	.	PUNCT
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gs-3955	752	2	,	,	PUNCT
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gs-3955	752	4	.	.	PROPN
gs-3955	752	5	,	,	PUNCT
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gs-3955	753	4	echinoderms	echinoderm	NOUN
gs-3955	753	5	.	.	PUNCT
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gs-3955	754	2	,	,	PUNCT
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gs-3955	754	4	)	)	PUNCT
gs-3955	754	5	,	,	PUNCT
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gs-3955	754	7	.	.	PUNCT
gs-3955	755	1	[	[	X
gs-3955	755	2	pubmed	pubmed	X
gs-3955	755	3	:	:	PUNCT
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gs-3955	755	5	]	]	PUNCT
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gs-3955	756	2	.	.	PUNCT
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gs-3955	757	2	,	,	PUNCT
gs-3955	757	3	t.	t.	PROPN
gs-3955	757	4	,	,	PUNCT
gs-3955	757	5	et	et	PROPN
gs-3955	757	6	al	al	PROPN
gs-3955	757	7	.	.	PROPN
gs-3955	757	8	(	(	PUNCT
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gs-3955	757	11	.	.	PUNCT
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gs-3955	758	11	.	.	PUNCT
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gs-3955	758	15	)	)	PUNCT
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gs-3955	758	18	-	-	SYM
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gs-3955	758	20	.	.	PUNCT
gs-3955	759	1	[	[	X
gs-3955	759	2	pubmed	pubmed	X
gs-3955	759	3	:	:	PUNCT
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gs-3955	759	5	]	]	SYM
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gs-3955	759	7	.	.	PUNCT
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gs-3955	760	2	,	,	PUNCT
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gs-3955	760	4	,	,	PUNCT
gs-3955	760	5	et	et	PROPN
gs-3955	760	6	al	al	PROPN
gs-3955	760	7	.	.	PUNCT
gs-3955	760	8	(	(	PUNCT
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gs-3955	760	10	)	)	PUNCT
gs-3955	760	11	.	.	PUNCT
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gs-3955	761	6	.	.	PUNCT
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gs-3955	762	8	,	,	PUNCT
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gs-3955	762	12	-	-	SYM
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gs-3955	763	1	[	[	X
gs-3955	763	2	pubmed	pubmed	NOUN
gs-3955	763	3	:	:	PUNCT
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gs-3955	763	5	]	]	PUNCT
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gs-3955	764	2	.	.	PUNCT
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gs-3955	765	2	,	,	PUNCT
gs-3955	765	3	y.	y.	PROPN
gs-3955	765	4	,	,	PUNCT
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gs-3955	765	8	(	(	PUNCT
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gs-3955	765	11	.	.	PUNCT
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gs-3955	766	8	.	.	PUNCT
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gs-3955	767	2	,	,	PUNCT
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gs-3955	768	3	:	:	PUNCT
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gs-3955	768	5	]	]	PUNCT
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gs-3955	769	2	.	.	PUNCT
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gs-3955	770	2	,	,	PUNCT
gs-3955	770	3	y.	y.	PROPN
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gs-3955	770	8	(	(	PUNCT
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gs-3955	770	10	)	)	PUNCT
gs-3955	770	11	.	.	PUNCT
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gs-3955	771	5	-	-	PUNCT
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gs-3955	771	8	.	.	PUNCT
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gs-3955	772	7	-	-	SYM
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gs-3955	773	1	[	[	X
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gs-3955	773	3	:	:	PUNCT
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gs-3955	773	18	:	:	PUNCT
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gs-3955	774	11	;	;	PUNCT
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gs-3955	774	18	პროფესორი	პროფესორი	PROPN
gs-3955	774	19	საქართველოს	საქართველოს	PROPN
gs-3955	774	20	ტექნიკურ	ტექნიკურ	PROPN
gs-3955	774	21	უნივერსიტეტში	უნივერსიტეტში	PROPN
gs-3955	774	22	;	;	PUNCT
gs-3955	774	23	პროფესორი	პროფესორი	PROPN
gs-3955	774	24	საქართველოს	საქართველოს	PROPN
gs-3955	774	25	ეროვნული	ეროვნული	VERB
gs-3955	774	26	უნივერსიტეტის	უნივერსიტეტის	NOUN
gs-3955	774	27	(	(	PUNCT
gs-3955	774	28	სეუ)-ში	სეუ)-ში	PROPN
gs-3955	774	29	;	;	PUNCT
gs-3955	774	30	კორესპონდენტ	კორესპონდენტ	PROPN
gs-3955	774	31	ავტორი	ავტორი	NOUN
gs-3955	774	32	:	:	PUNCT
gs-3955	774	33	პროფესორი	პროფესორი	PROPN
gs-3955	774	34	,	,	PUNCT
gs-3955	774	35	დოქტორი	დოქტორი	PROPN
gs-3955	774	36	დავით	დავით	PROPN
gs-3955	774	37	აფხაზავა,ელ.ფოსტა	აფხაზავა,ელ.ფოსტა	PROPN
gs-3955	774	38	:	:	PUNCT
gs-3955	775	1	d.apkhazava@alte.edu.ge	d.apkhazava@alte.edu.ge	NOUN
gs-3955	775	2	2მედიცინის	2მედიცინის	NUM
gs-3955	775	3	დოქტორი	დოქტორი	NOUN
gs-3955	775	4	,	,	PUNCT
gs-3955	775	5	ფილოსოფიის	ფილოსოფიის	NOUN
gs-3955	775	6	დოქტორი	დოქტორი	NOUN
gs-3955	775	7	,	,	PUNCT
gs-3955	775	8	ფარმაცევტულ	ფარმაცევტულ	PROPN
gs-3955	775	9	და	და	NOUN
gs-3955	775	10	ფარმაკოლოგიურ	ფარმაკოლოგიურ	NOUN
gs-3955	775	11	მეცნიერებათა	მეცნიერებათა	NOUN
gs-3955	775	12	დოქტორი	დოქტორი	NOUN
gs-3955	775	13	მედიცინაში	მედიცინაში	PROPN
gs-3955	775	14	,	,	PUNCT
gs-3955	775	15	მოწვეული	მოწვეული	PROPN
gs-3955	775	16	ლექტორი	ლექტორი	PROPN
gs-3955	775	17	(	(	PUNCT
gs-3955	775	18	მოწვეული	მოწვეული	PROPN
gs-3955	775	19	პროფესორი	პროფესორი	NOUN
gs-3955	775	20	)	)	PUNCT
gs-3955	775	21	თბილისის	თბილისის	NOUN
gs-3955	775	22	სახელმწიფო	სახელმწიფო	NOUN
gs-3955	775	23	სამედიცინო	სამედიცინო	NOUN
gs-3955	775	24	უნივერსიტეტის	უნივერსიტეტის	NOUN
gs-3955	775	25	სამეცნიერო	სამეცნიერო	NOUN
gs-3955	775	26	კვლევითი	კვლევითი	NOUN
gs-3955	775	27	უნარების	უნარების	VERB
gs-3955	775	28	ცენტრიდან	ცენტრიდან	PROPN
gs-3955	775	29	;	;	PUNCT
gs-3955	775	30	სამედიცინო	სამედიცინო	VERB
gs-3955	775	31	და	და	NOUN
gs-3955	775	32	კლინიკური	კლინიკური	NOUN
gs-3955	775	33	ფარმაკოლოგიის	ფარმაკოლოგიის	PROPN
gs-3955	776	1	პროფესორი	პროფესორი	NOUN
gs-3955	776	2	მედიცინის	მედიცინის	DET
gs-3955	776	3	საერთაშორისო	საერთაშორისო	PROPN
gs-3955	776	4	სკოლაში	სკოლაში	PROPN
gs-3955	776	5	ალტეს	ალტეს	PROPN
gs-3955	776	6	უნივერსიტეტში	უნივერსიტეტში	PROPN
gs-3955	776	7	;	;	PUNCT
gs-3955	776	8	ფარმაკოლოგიის	ფარმაკოლოგიის	PROPN
gs-3955	776	9	პროფესორი	პროფესორი	NOUN
gs-3955	776	10	მედიცინის	მედიცინის	DET
gs-3955	776	11	ფაკულტეტზე	ფაკულტეტზე	PROPN
gs-3955	776	12	საქართველოს	საქართველოს	NOUN
gs-3955	776	13	ეროვნული	ეროვნული	VERB
gs-3955	776	14	უნივერსიტეტის	უნივერსიტეტის	ADJ
gs-3955	776	15	seu-ში	seu-ში	PROPN
gs-3955	776	16	;	;	PUNCT
gs-3955	776	17	ასოცირებული	ასოცირებული	NOUN
gs-3955	776	18	პროფესორი	პროფესორი	NOUN
gs-3955	776	19	მედიცინის	მედიცინის	PRON
gs-3955	776	20	ფაკულტეტზე	ფაკულტეტზე	ADJ
gs-3955	776	21	სამედიცინო	სამედიცინო	NOUN
gs-3955	776	22	ფარმაკოლოგიაში	ფარმაკოლოგიაში	PROPN
gs-3955	776	23	სულხან-საბა	სულხან-საბა	ADJ
gs-3955	776	24	ორბელიანის	ორბელიანის	PROPN
gs-3955	776	25	უნივერსიტეტში	უნივერსიტეტში	PROPN
gs-3955	776	26	;	;	PUNCT
gs-3955	776	27	თბილისი	თბილისი	PROPN
gs-3955	776	28	,	,	PUNCT
gs-3955	776	29	საქართველო.	საქართველო.	PROPN
gs-3955	776	30	orcid	orcid	PROPN
gs-3955	776	31	:	:	PUNCT
gs-3955	776	32	https://orcid.org/0000-0002-9005-8577	https://orcid.org/0000-0002-9005-8577	PROPN
gs-3955	776	33	;	;	PUNCT
gs-3955	776	34	ელ.ფოსტა	ელ.ფოსტა	X
gs-3955	776	35	:	:	PUNCT
gs-3955	776	36	n.sulashvili@ug.edu.ge	n.sulashvili@ug.edu.ge	PROPN
gs-3955	776	37	3დოქტორი	3დოქტორი	NUM
gs-3955	776	38	,	,	PUNCT
gs-3955	776	39	საქართველოს	საქართველოს	VERB
gs-3955	776	40	უნივერსიტეტის	უნივერსიტეტის	NOUN
gs-3955	776	41	პროფესორი.	პროფესორი.	ADJ
gs-3955	776	42	4კვლევების	4კვლევების	NUM
gs-3955	776	43	დირექტორი	დირექტორი	NOUN
gs-3955	776	44	;	;	PUNCT
gs-3955	776	45	longevity	longevity	NOUN
gs-3955	776	46	clinic	clinic	PROPN
gs-3955	776	47	,	,	PUNCT
gs-3955	776	48	inc	inc	PROPN
gs-3955	776	49	.	.	PROPN
gs-3955	776	50	orcid	orcid	PROPN
gs-3955	776	51	:	:	PUNCT
gs-3955	776	52	https://orcid.org/0000-0001-8651-7243	https://orcid.org/0000-0001-8651-7243	PROPN
gs-3955	776	53	;	;	PUNCT
gs-3955	776	54	ელ.ფოსტა	ელ.ფოსტა	NUM
gs-3955	776	55	:	:	PUNCT
gs-3955	776	56	jtkemaladze@longevity.ge	jtkemaladze@longevity.ge	NOUN
gs-3955	776	57	აბსტრაქტი	აბსტრაქტი	NOUN
gs-3955	776	58	მიუხედავად	მიუხედავად	PROPN
gs-3955	776	59	იმისა	იმისა	NOUN
gs-3955	776	60	,	,	PUNCT
gs-3955	776	61	რომ	რომ	PROPN
gs-3955	776	62	დაბერება	დაბერება	PROPN
gs-3955	776	63	თითქმის	თითქმის	PROPN
gs-3955	776	64	უნივერსალური	უნივერსალური	PROPN
gs-3955	776	65	ბიოლოგიური	ბიოლოგიური	PROPN
gs-3955	776	66	პროცესია	პროცესია	PROPN
gs-3955	776	67	,	,	PUNCT
gs-3955	776	68	გარკვეული	გარკვეული	PROPN
gs-3955	776	69	სახეობები	სახეობები	NOUN
gs-3955	776	70	ავლენენ	ავლენენ	PROPN
gs-3955	776	71	უმნიშვნელო	უმნიშვნელო	NOUN
gs-3955	776	72	(	(	PUNCT
gs-3955	776	73	მდორედ	მდორედ	NOUN
gs-3955	776	74	მიმდინარე	მიმდინარე	NOUN
gs-3955	776	75	)	)	PUNCT
gs-3955	776	76	დაბერებას	დაბერებას	NOUN
gs-3955	776	77	,	,	PUNCT
gs-3955	776	78	ინარჩუნებენ	ინარჩუნებენ	NOUN
gs-3955	776	79	რეპროდუქციულ	რეპროდუქციულ	NOUN
gs-3955	776	80	უნარს	უნარს	VERB
gs-3955	776	81	და	და	PROPN
gs-3955	776	82	ფიზიოლოგიურ	ფიზიოლოგიურ	PROPN
gs-3955	776	83	ფუნქციას	ფუნქციას	NOUN
gs-3955	776	84	ხანგრძლივი	ხანგრძლივი	PROPN
gs-3955	776	85	სიცოცხლის	სიცოცხლის	PROPN
gs-3955	776	86	განმავლობაში.	განმავლობაში.	VERB
gs-3955	776	87	აღსანიშნავი	აღსანიშნავი	PROPN
gs-3955	776	88	მაგალითებია	მაგალითებია	VERB
gs-3955	776	89	გრენლანდიური	გრენლანდიური	PROPN
gs-3955	776	90	ზვიგენი	ზვიგენი	NOUN
gs-3955	776	91	(	(	PUNCT
gs-3955	776	92	400	400	NUM
gs-3955	776	93	წელზე	წელზე	NOUN
gs-3955	776	94	მეტი	მეტი	NOUN
gs-3955	776	95	)	)	PUNCT
gs-3955	776	96	,	,	PUNCT
gs-3955	776	97	შიშველი	შიშველი	ADJ
gs-3955	776	98	თხუნელა-ვირთხა	თხუნელა-ვირთხა	ADJ
gs-3955	776	99	(	(	PUNCT
gs-3955	776	100	30	30	NUM
gs-3955	776	101	წელზე	წელზე	NOUN
gs-3955	776	102	მეტი	მეტი	NOUN
gs-3955	776	103	)	)	PUNCT
gs-3955	776	104	,	,	PUNCT
gs-3955	776	105	ოკეანის	ოკეანის	PROPN
gs-3955	776	106	კვაჰოგი	კვაჰოგი	NOUN
gs-3955	776	107	(	(	PUNCT
gs-3955	776	108	507	507	NUM
gs-3955	776	109	წელი	წელი	NOUN
gs-3955	776	110	)	)	PUNCT
gs-3955	776	111	და	და	NOUN
gs-3955	776	112	პლანარია	პლანარია	NOUN
gs-3955	776	113	(	(	PUNCT
gs-3955	776	114	თეორიულად	თეორიულად	NOUN
gs-3955	776	115	უკვდავი	უკვდავი	PROPN
gs-3955	776	116	)	)	PUNCT
gs-3955	776	117	.	.	PUNCT
gs-3955	777	1	ეს	ეს	PROPN
gs-3955	777	2	ორგანიზმები	ორგანიზმები	PROPN
gs-3955	777	3	ეწინააღმდეგებიან	ეწინააღმდეგებიან	PROPN
gs-3955	777	4	დაბერების	დაბერების	PROPN
gs-3955	777	5	ტიპურ	ტიპურ	VERB
gs-3955	777	6	ნიმუშებს	ნიმუშებს	PROPN
gs-3955	777	7	დნმ-ის	დნმ-ის	PROPN
gs-3955	777	8	გაძლიერებული	გაძლიერებული	PROPN
gs-3955	777	9	აღდგენის	აღდგენის	PROPN
gs-3955	777	10	,	,	PUNCT
gs-3955	777	11	ჟანგვითი	ჟანგვითი	VERB
gs-3955	777	12	სტრესისადმი	სტრესისადმი	ADV
gs-3955	777	13	რეზისტენტობის	რეზისტენტობის	ADJ
gs-3955	777	14	,	,	PUNCT
gs-3955	777	15	ღეროვანი	ღეროვანი	NOUN
gs-3955	777	16	უჯრედების	უჯრედების	VERB
gs-3955	777	17	შენარჩუნებისა	შენარჩუნებისა	VERB
gs-3955	777	18	და	და	PROPN
gs-3955	777	19	უნიკალური	უნიკალური	PROPN
gs-3955	777	20	პროტეოსტაზისა	პროტეოსტაზისა	PROPN
gs-3955	777	21	და	და	PROPN
gs-3955	777	22	ეპიგენეტიკური	ეპიგენეტიკური	ADJ
gs-3955	777	23	რეგულავიის	რეგულავიის	PROPN
gs-3955	777	24	გზით.	გზით.	PROPN
gs-3955	777	25	შედარებითი	შედარებითი	NOUN
gs-3955	777	26	ანალიზი	ანალიზი	VERB
gs-3955	777	27	ავლენს	ავლენს	PROPN
gs-3955	777	28	საერთო	საერთო	PROPN
gs-3955	777	29	ადაპტაციებს	ადაპტაციებს	PROPN
gs-3955	777	30	,	,	PUNCT
gs-3955	777	31	როგორიცაა	როგორიცაა	VERB
gs-3955	777	32	:	:	PUNCT
gs-3955	777	33	1	1	X
gs-3955	777	34	)	)	PUNCT
gs-3955	777	35	სქსორივი	სქსორივი	NOUN
gs-3955	777	36	მომწიფების	მომწიფების	ADJ
gs-3955	777	37	გვიანი	გვიანი	NOUN
gs-3955	777	38	ასაკი	ასაკი	ADP
gs-3955	777	39	;	;	PUNCT
gs-3955	777	40	2	2	X
gs-3955	777	41	)	)	PUNCT
gs-3955	777	42	სქესობრივი	სქესობრივი	NOUN
gs-3955	777	43	აქტივობის	აქტივობის	ADV
gs-3955	777	44	დიდი	დიდი	VERB
gs-3955	777	45	ხნით	ხნით	ADJ
gs-3955	777	46	შენარჩუნება	შენარჩუნება	NOUN
gs-3955	777	47	;	;	PUNCT
gs-3955	777	48	3	3	X
gs-3955	777	49	)	)	PUNCT
gs-3955	777	50	მეტაბოლური	მეტაბოლური	PROPN
gs-3955	777	51	სიჩქარის	სიჩქარის	PROPN
gs-3955	777	52	შემცირება	შემცირება	PROPN
gs-3955	777	53	;	;	PUNCT
gs-3955	777	54	4	4	X
gs-3955	777	55	)	)	PUNCT
gs-3955	777	56	ჰიპოქსიისადმი	ჰიპოქსიისადმი	NOUN
gs-3955	777	57	ტოლერანტობა	ტოლერანტობა	NOUN
gs-3955	777	58	;	;	PUNCT
gs-3955	777	59	5	5	NUM
gs-3955	777	60	)	)	PUNCT
gs-3955	777	61	ნეოპლაზიური	ნეოპლაზიური	NOUN
gs-3955	777	62	სუპრესია.	სუპრესია.	VERB
gs-3955	777	63	მათი	მათი	PROPN
gs-3955	777	64	სიცოცხლის	სიცოცხლის	PROPN
gs-3955	778	1	ხანგრძლივობის	ხანგრძლივობის	PROPN
gs-3955	778	2	მოლეკულარული	მოლეკულარული	PROPN
gs-3955	778	3	მექანიზმები	მექანიზმები	NOUN
gs-3955	778	4	მათ	მათ	AUX
gs-3955	778	5	შორის	შორის	NOUN
gs-3955	778	6	მაღალი	მაღალი	PROPN
gs-3955	778	7	მოლეკულური	მოლეკულური	PROPN
gs-3955	778	8	წონის	წონის	PROPN
gs-3955	778	9	ჰიალურონანი	ჰიალურონანი	AUX
gs-3955	778	10	შიშველ	შიშველ	VERB
gs-3955	778	11	თხუნელა-ვირთხებში	თხუნელა-ვირთხებში	ADJ
gs-3955	778	12	და	და	NOUN
gs-3955	778	13	სპეციალიზებული	სპეციალიზებული	X
gs-3955	778	14	ლიპიდური	ლიპიდური	NOUN
gs-3955	778	15	მემბრანები	მემბრანები	PROPN
gs-3955	778	16	ზვიგენებში	ზვიგენებში	NOUN
gs-3955	778	17	გვთავაზობს	გვთავაზობს	NOUN
gs-3955	778	18	ტრანსფორმაციულ	ტრანსფორმაციულ	VERB
gs-3955	778	19	ხედვას	ხედვას	PROPN
gs-3955	778	20	დაბერების	დაბერების	VERB
gs-3955	778	21	georgian	georgian	ADJ
gs-3955	778	22	scientists/ქართველი	scientists/ქართველი	NOUN
gs-3955	778	23	მეცნიერები	მეცნიერები	NOUN
gs-3955	778	24	ტ.	ტ.	VERB
gs-3955	778	25	7	7	NUM
gs-3955	778	26	n	n	PRON
gs-3955	778	27	3	3	NUM
gs-3955	778	28	,	,	PUNCT
gs-3955	778	29	2025	2025	NUM
gs-3955	778	30	396	396	NUM
gs-3955	778	31	საწინააღმდეგო	საწინააღმდეგო	NOUN
gs-3955	778	32	ჩარევებისთვის.	ჩარევებისთვის.	NOUN
gs-3955	778	33	ამ	ამ	ADP
gs-3955	778	34	სტრატეგიების	სტრატეგიების	PROPN
gs-3955	778	35	გაგებამ	გაგებამ	PROPN
gs-3955	778	36	შეიძლება	შეიძლება	PROPN
gs-3955	778	37	გზა	გზა	PROPN
gs-3955	778	38	გაუხსნას	გაუხსნას	PROPN
gs-3955	778	39	ასაკთან	ასაკთან	PROPN
gs-3955	778	40	დაკავშირებული	დაკავშირებული	NOUN
gs-3955	778	41	დაავადებების	დაავადებების	VERB
gs-3955	778	42	სამკურნალოდ	სამკურნალოდ	ADJ
gs-3955	778	43	მიმართული	მიმართული	NOUN
gs-3955	778	44	ახალი	ახალი	ADJ
gs-3955	778	45	თერაპიების	თერაპიების	NOUN
gs-3955	778	46	განვითარებას	განვითარებას	NOUN
gs-3955	778	47	,	,	PUNCT
gs-3955	778	48	რაც	რაც	NOUN
gs-3955	778	49	პოტენციურად	პოტენციურად	PROPN
gs-3955	778	50	გადააფასებს	გადააფასებს	NOUN
gs-3955	778	51	ადამიანის	ადამიანის	NOUN
gs-3955	778	52	ჯანმრთელობის	ჯანმრთელობის	ADJ
gs-3955	778	53	ხანგრძლივობას.	ხანგრძლივობას.	PROPN
gs-3955	778	54	ეს	ეს	PROPN
gs-3955	778	55	მიმოხილვა	მიმოხილვა	PROPN
gs-3955	778	56	სინთეზირებს	სინთეზირებს	PROPN
gs-3955	778	57	უმნიშვნელოდ	უმნიშვნელოდ	PROPN
gs-3955	778	58	დაბერებული	დაბერებული	PROPN
gs-3955	778	59	სახეობების	სახეობების	ADJ
gs-3955	778	60	შესახებ	შესახებ	VERB
gs-3955	778	61	არსებულ	არსებულ	PROPN
gs-3955	778	62	ცოდნას	ცოდნას	PROPN
gs-3955	778	63	,	,	PUNCT
gs-3955	778	64	ხაზს	ხაზს	ADJ
gs-3955	778	65	უსვამს	უსვამს	NOUN
gs-3955	778	66	მათ	მათ	NOUN
gs-3955	778	67	ევოლუციურ	ევოლუციურ	ADJ
gs-3955	778	68	,	,	PUNCT
gs-3955	778	69	ფიზიოლოგიურ	ფიზიოლოგიურ	ADJ
gs-3955	778	70	და	და	NOUN
gs-3955	778	71	ბიოსამედიცინო	ბიოსამედიცინო	PROPN
gs-3955	778	72	მნიშვნელობას.	მნიშვნელობას.	NOUN
gs-3955	778	73	საკვანძო	საკვანძო	PROPN
gs-3955	778	74	სიტყვები	სიტყვები	NOUN
gs-3955	778	75	:	:	PUNCT
gs-3955	778	76	უმნიშვნელო	უმნიშვნელო	PROPN
gs-3955	778	77	დაბერება	დაბერება	PROPN
gs-3955	778	78	,	,	PUNCT
gs-3955	778	79	სიცოცხლის	სიცოცხლის	NOUN
gs-3955	778	80	ხანგრძლივობა	ხანგრძლივობა	PROPN
gs-3955	778	81	,	,	PUNCT
gs-3955	778	82	ოქსიდაციური	ოქსიდაციური	PROPN
gs-3955	778	83	სტრესი	სტრესი	PROPN
gs-3955	778	84	,	,	PUNCT
gs-3955	778	85	პროტეოსტაზი	პროტეოსტაზი	NOUN
gs-3955	778	86	,	,	PUNCT
gs-3955	778	87	დაბერება	დაბერება	ADJ
gs-3955	778	88	,	,	PUNCT
gs-3955	778	89	დღეგრძელობა.	დღეგრძელობა.	NOUN