id	sid	tid	token	lemma	pos
iguana-17148	1	1	iguana	iguana	NOUN
iguana-17148	1	2	11.4	11.4	NUM
iguana-17148	1	3	190	190	NUM
iguana-17148	1	4	iguana	iguana	NOUN
iguana-17148	1	5	•	•	NUM
iguana-17148	1	6	volume	volume	NOUN
iguana-17148	1	7	11	11	NUM
iguana-17148	1	8	,	,	PUNCT
iguana-17148	1	9	number	number	NOUN
iguana-17148	1	10	4	4	NUM
iguana-17148	1	11	•	•	NOUN
iguana-17148	1	12	december	december	PROPN
iguana-17148	1	13	2004	2004	NUM
iguana-17148	1	14	wikelski	wikelski	NOUN
iguana-17148	1	15	and	and	CCONJ
iguana-17148	1	16	nelson	nelson	PROPN
iguana-17148	1	17	adult	adult	PROPN
iguana-17148	1	18	marine	marine	ADJ
iguana-17148	1	19	iguana	iguana	NOUN
iguana-17148	1	20	(	(	PUNCT
iguana-17148	1	21	amblyrhynchus	amblyrhynchus	NOUN
iguana-17148	1	22	cristatus	cristatus	PROPN
iguana-17148	1	23	)	)	PUNCT
iguana-17148	1	24	from	from	ADP
iguana-17148	1	25	san	san	PROPN
iguana-17148	1	26	cristobal	cristobal	PROPN
iguana-17148	1	27	island	island	PROPN
iguana-17148	1	28	in	in	ADP
iguana-17148	1	29	the	the	DET
iguana-17148	1	30	galápagos	galápagos	PROPN
iguana-17148	1	31	archipelago	archipelago	PROPN
iguana-17148	1	32	.	.	PUNCT
iguana-17148	2	1	photograph	photograph	NOUN
iguana-17148	2	2	by	by	ADP
iguana-17148	2	3	colette	colette	PROPN
iguana-17148	2	4	adams	adams	PROPN
iguana-17148	2	5	.	.	PUNCT
iguana-17148	3	1	introduction	introduction	NOUN
iguana-17148	3	2	when	when	SCONJ
iguana-17148	3	3	the	the	DET
iguana-17148	3	4	galápagos	galápago	NOUN
iguana-17148	3	5	’	'	PUNCT
iguana-17148	3	6	most	most	ADV
iguana-17148	3	7	famous	famous	ADJ
iguana-17148	3	8	visitor	visitor	NOUN
iguana-17148	3	9	,	,	PUNCT
iguana-17148	3	10	charles	charles	PROPN
iguana-17148	3	11	darwin	darwin	PROPN
iguana-17148	3	12	,	,	PUNCT
iguana-17148	3	13	arrived	arrive	VERB
iguana-17148	3	14	on	on	ADP
iguana-17148	3	15	the	the	DET
iguana-17148	3	16	rocky	rocky	ADJ
iguana-17148	3	17	lava	lava	NOUN
iguana-17148	3	18	shores	shore	NOUN
iguana-17148	4	1	,	,	PUNCT
iguana-17148	4	2	he	he	PRON
iguana-17148	4	3	likened	liken	VERB
iguana-17148	4	4	the	the	DET
iguana-17148	4	5	islands	island	NOUN
iguana-17148	4	6	to	to	ADP
iguana-17148	4	7	the	the	DET
iguana-17148	4	8	entrance	entrance	NOUN
iguana-17148	4	9	of	of	ADP
iguana-17148	4	10	hell	hell	PROPN
iguana-17148	4	11	.	.	PUNCT
iguana-17148	5	1	he	he	PRON
iguana-17148	5	2	found	find	VERB
iguana-17148	5	3	myriads	myriad	NOUN
iguana-17148	5	4	of	of	ADP
iguana-17148	5	5	“	"	PUNCT
iguana-17148	5	6	dirty	dirty	ADJ
iguana-17148	5	7	black	black	ADJ
iguana-17148	5	8	”	"	PUNCT
iguana-17148	5	9	marine	marine	ADJ
iguana-17148	5	10	iguanas	iguana	NOUN
iguana-17148	5	11	(	(	PUNCT
iguana-17148	5	12	darwin	darwin	PROPN
iguana-17148	5	13	1883	1883	NUM
iguana-17148	5	14	)	)	PUNCT
iguana-17148	5	15	.	.	PUNCT
iguana-17148	6	1	young	young	ADJ
iguana-17148	6	2	charles	charles	PROPN
iguana-17148	6	3	found	find	VERB
iguana-17148	6	4	them	they	PRON
iguana-17148	6	5	“	"	PUNCT
iguana-17148	6	6	hideous	hideous	ADJ
iguana-17148	6	7	in	in	ADP
iguana-17148	6	8	appearance	appearance	NOUN
iguana-17148	6	9	,	,	PUNCT
iguana-17148	6	10	sluggish	sluggish	ADJ
iguana-17148	6	11	,	,	PUNCT
iguana-17148	6	12	stupid	stupid	ADJ
iguana-17148	6	13	,	,	PUNCT
iguana-17148	6	14	and	and	CCONJ
iguana-17148	6	15	ugly	ugly	ADJ
iguana-17148	6	16	.	.	PUNCT
iguana-17148	6	17	”	"	PUNCT
iguana-17148	7	1	nevertheless	nevertheless	ADV
iguana-17148	7	2	,	,	PUNCT
iguana-17148	7	3	like	like	ADP
iguana-17148	7	4	every	every	DET
iguana-17148	7	5	modern	modern	ADJ
iguana-17148	7	6	visitor	visitor	NOUN
iguana-17148	7	7	,	,	PUNCT
iguana-17148	7	8	he	he	PRON
iguana-17148	7	9	was	be	AUX
iguana-17148	7	10	fascinated	fascinate	VERB
iguana-17148	7	11	by	by	ADP
iguana-17148	7	12	their	their	PRON
iguana-17148	7	13	sociality	sociality	NOUN
iguana-17148	7	14	and	and	CCONJ
iguana-17148	7	15	their	their	PRON
iguana-17148	7	16	marine	marine	ADJ
iguana-17148	7	17	foraging	forage	VERB
iguana-17148	7	18	style	style	NOUN
iguana-17148	7	19	—	—	PUNCT
iguana-17148	7	20	marine	marine	ADJ
iguana-17148	7	21	iguanas	iguana	NOUN
iguana-17148	7	22	are	be	AUX
iguana-17148	7	23	the	the	DET
iguana-17148	7	24	only	only	ADJ
iguana-17148	7	25	lizards	lizard	NOUN
iguana-17148	7	26	known	know	VERB
iguana-17148	7	27	to	to	PART
iguana-17148	7	28	feed	feed	VERB
iguana-17148	7	29	exclusively	exclusively	ADV
iguana-17148	7	30	on	on	ADP
iguana-17148	7	31	algae	algae	NOUN
iguana-17148	7	32	.	.	PUNCT
iguana-17148	8	1	marine	marine	PROPN
iguana-17148	8	2	iguanas	iguana	NOUN
iguana-17148	8	3	live	live	VERB
iguana-17148	8	4	in	in	ADP
iguana-17148	8	5	dense	dense	ADJ
iguana-17148	8	6	clusters	cluster	NOUN
iguana-17148	8	7	of	of	ADP
iguana-17148	8	8	up	up	ADP
iguana-17148	8	9	to	to	PART
iguana-17148	8	10	8,000	8,000	NUM
iguana-17148	8	11	animals	animal	NOUN
iguana-17148	8	12	per	per	ADP
iguana-17148	8	13	kilometer	kilometer	NOUN
iguana-17148	8	14	of	of	ADP
iguana-17148	8	15	coastline	coastline	NOUN
iguana-17148	8	16	.	.	PUNCT
iguana-17148	9	1	however	however	ADV
iguana-17148	9	2	,	,	PUNCT
iguana-17148	9	3	iguana	iguana	NOUN
iguana-17148	9	4	colonies	colony	NOUN
iguana-17148	9	5	are	be	AUX
iguana-17148	9	6	distributed	distribute	VERB
iguana-17148	9	7	very	very	ADV
iguana-17148	9	8	patchily	patchily	ADV
iguana-17148	9	9	,	,	PUNCT
iguana-17148	9	10	and	and	CCONJ
iguana-17148	9	11	tend	tend	VERB
iguana-17148	9	12	to	to	PART
iguana-17148	9	13	occur	occur	VERB
iguana-17148	9	14	only	only	ADV
iguana-17148	9	15	along	along	ADP
iguana-17148	9	16	the	the	DET
iguana-17148	9	17	southwestern	southwestern	ADJ
iguana-17148	9	18	shores	shore	NOUN
iguana-17148	9	19	of	of	ADP
iguana-17148	9	20	the	the	DET
iguana-17148	9	21	islands	island	NOUN
iguana-17148	9	22	.	.	PUNCT
iguana-17148	10	1	amazingly	amazingly	ADV
iguana-17148	10	2	,	,	PUNCT
iguana-17148	10	3	little	little	ADJ
iguana-17148	10	4	has	have	AUX
iguana-17148	10	5	changed	change	VERB
iguana-17148	10	6	for	for	ADP
iguana-17148	10	7	marine	marine	ADJ
iguana-17148	10	8	iguanas	iguana	NOUN
iguana-17148	10	9	on	on	ADP
iguana-17148	10	10	the	the	DET
iguana-17148	10	11	uninhabited	uninhabited	ADJ
iguana-17148	10	12	islands	island	NOUN
iguana-17148	10	13	since	since	SCONJ
iguana-17148	10	14	darwin	darwin	PROPN
iguana-17148	10	15	’s	’s	PART
iguana-17148	10	16	visit	visit	NOUN
iguana-17148	10	17	more	more	ADJ
iguana-17148	10	18	than	than	ADP
iguana-17148	10	19	170	170	NUM
iguana-17148	10	20	years	year	NOUN
iguana-17148	10	21	ago	ago	ADV
iguana-17148	10	22	.	.	PUNCT
iguana-17148	11	1	however	however	ADV
iguana-17148	11	2	,	,	PUNCT
iguana-17148	11	3	although	although	SCONJ
iguana-17148	11	4	marine	marine	ADJ
iguana-17148	11	5	iguanas	iguana	NOUN
iguana-17148	11	6	still	still	ADV
iguana-17148	11	7	occur	occur	VERB
iguana-17148	11	8	in	in	ADP
iguana-17148	11	9	healthy	healthy	ADJ
iguana-17148	11	10	population	population	NOUN
iguana-17148	11	11	densities	density	NOUN
iguana-17148	11	12	on	on	ADP
iguana-17148	11	13	the	the	DET
iguana-17148	11	14	uninhabited	uninhabited	ADJ
iguana-17148	11	15	islands	island	NOUN
iguana-17148	11	16	,	,	PUNCT
iguana-17148	11	17	they	they	PRON
iguana-17148	11	18	face	face	VERB
iguana-17148	11	19	potentially	potentially	ADV
iguana-17148	11	20	serious	serious	ADJ
iguana-17148	11	21	threats	threat	NOUN
iguana-17148	11	22	on	on	ADP
iguana-17148	11	23	several	several	ADJ
iguana-17148	11	24	inhabited	inhabited	ADJ
iguana-17148	11	25	islands	island	NOUN
iguana-17148	11	26	.	.	PUNCT
iguana-17148	12	1	marine	marine	PROPN
iguana-17148	12	2	iguana	iguana	PROPN
iguana-17148	12	3	natural	natural	PROPN
iguana-17148	12	4	history	history	NOUN
iguana-17148	12	5	marine	marine	PROPN
iguana-17148	12	6	iguanas	iguana	NOUN
iguana-17148	12	7	are	be	AUX
iguana-17148	12	8	endemic	endemic	ADJ
iguana-17148	12	9	to	to	ADP
iguana-17148	12	10	the	the	DET
iguana-17148	12	11	galápagos	galápagos	PROPN
iguana-17148	12	12	archipelago	archipelago	PROPN
iguana-17148	12	13	,	,	PUNCT
iguana-17148	12	14	which	which	PRON
iguana-17148	12	15	belongs	belong	VERB
iguana-17148	12	16	to	to	ADP
iguana-17148	12	17	the	the	DET
iguana-17148	12	18	republic	republic	NOUN
iguana-17148	12	19	of	of	ADP
iguana-17148	12	20	ecuador	ecuador	PROPN
iguana-17148	12	21	.	.	PUNCT
iguana-17148	13	1	they	they	PRON
iguana-17148	13	2	feed	feed	VERB
iguana-17148	13	3	exclusively	exclusively	ADV
iguana-17148	13	4	on	on	ADP
iguana-17148	13	5	marine	marine	ADJ
iguana-17148	13	6	algae	algae	NOUN
iguana-17148	13	7	in	in	ADP
iguana-17148	13	8	the	the	DET
iguana-17148	13	9	rocky	rocky	ADJ
iguana-17148	13	10	intertidal	intertidal	ADJ
iguana-17148	13	11	zone	zone	NOUN
iguana-17148	13	12	(	(	PUNCT
iguana-17148	13	13	darwin	darwin	PROPN
iguana-17148	13	14	1883	1883	NUM
iguana-17148	13	15	,	,	PUNCT
iguana-17148	13	16	carpenter	carpenter	NOUN
iguana-17148	13	17	1966	1966	NUM
iguana-17148	13	18	,	,	PUNCT
iguana-17148	13	19	trillmich	trillmich	NOUN
iguana-17148	13	20	and	and	CCONJ
iguana-17148	13	21	trillmich	trillmich	NOUN
iguana-17148	13	22	1986	1986	NUM
iguana-17148	13	23	,	,	PUNCT
iguana-17148	13	24	wikelski	wikelski	PROPN
iguana-17148	13	25	et	et	PROPN
iguana-17148	13	26	al	al	PROPN
iguana-17148	13	27	.	.	PROPN
iguana-17148	13	28	1993	1993	NUM
iguana-17148	13	29	,	,	PUNCT
iguana-17148	13	30	wikelski	wikelski	NOUN
iguana-17148	13	31	and	and	CCONJ
iguana-17148	13	32	hau	hau	PROPN
iguana-17148	13	33	1995	1995	NUM
iguana-17148	13	34	,	,	PUNCT
iguana-17148	13	35	drent	drent	PROPN
iguana-17148	13	36	et	et	PROPN
iguana-17148	13	37	al	al	PROPN
iguana-17148	13	38	.	.	PROPN
iguana-17148	13	39	1999	1999	NUM
iguana-17148	13	40	)	)	PUNCT
iguana-17148	13	41	.	.	PUNCT
iguana-17148	14	1	marine	marine	PROPN
iguana-17148	14	2	iguanas	iguanas	PROPN
iguana-17148	14	3	possess	possess	VERB
iguana-17148	14	4	an	an	DET
iguana-17148	14	5	internal	internal	ADJ
iguana-17148	14	6	biological	biological	ADJ
iguana-17148	14	7	clock	clock	NOUN
iguana-17148	14	8	that	that	PRON
iguana-17148	14	9	is	be	AUX
iguana-17148	14	10	synchronized	synchronize	VERB
iguana-17148	14	11	to	to	ADP
iguana-17148	14	12	the	the	DET
iguana-17148	14	13	tides	tide	NOUN
iguana-17148	14	14	.	.	PUNCT
iguana-17148	15	1	this	this	DET
iguana-17148	15	2	clock	clock	NOUN
iguana-17148	15	3	cues	cue	VERB
iguana-17148	15	4	them	they	PRON
iguana-17148	15	5	to	to	PART
iguana-17148	15	6	walk	walk	VERB
iguana-17148	15	7	to	to	ADP
iguana-17148	15	8	the	the	DET
iguana-17148	15	9	intertidal	intertidal	ADJ
iguana-17148	15	10	zone	zone	NOUN
iguana-17148	15	11	every	every	DET
iguana-17148	15	12	day	day	NOUN
iguana-17148	15	13	at	at	ADP
iguana-17148	15	14	low	low	ADJ
iguana-17148	15	15	tide	tide	NOUN
iguana-17148	15	16	,	,	PUNCT
iguana-17148	15	17	when	when	SCONJ
iguana-17148	15	18	the	the	DET
iguana-17148	15	19	algae	algae	NOUN
iguana-17148	15	20	are	be	AUX
iguana-17148	15	21	exposed	expose	VERB
iguana-17148	15	22	(	(	PUNCT
iguana-17148	15	23	wikelski	wikelski	NOUN
iguana-17148	15	24	and	and	CCONJ
iguana-17148	15	25	conservation	conservation	NOUN
iguana-17148	15	26	of	of	ADP
iguana-17148	15	27	galápagos	galápagos	PROPN
iguana-17148	15	28	marine	marine	PROPN
iguana-17148	15	29	iguanas	iguana	NOUN
iguana-17148	15	30	(	(	PUNCT
iguana-17148	15	31	amblyrhynchus	amblyrhynchus	NOUN
iguana-17148	15	32	cristatus	cristatus	PROPN
iguana-17148	15	33	)	)	PUNCT
iguana-17148	15	34	martin	martin	PROPN
iguana-17148	15	35	wikelski1	wikelski1	PROPN
iguana-17148	15	36	and	and	CCONJ
iguana-17148	15	37	karin	karin	PROPN
iguana-17148	15	38	nelson2	nelson2	NOUN
iguana-17148	16	1	1department	1department	NUM
iguana-17148	16	2	of	of	ADP
iguana-17148	16	3	ecology	ecology	NOUN
iguana-17148	16	4	and	and	CCONJ
iguana-17148	16	5	evolutionary	evolutionary	ADJ
iguana-17148	16	6	biology	biology	NOUN
iguana-17148	16	7	,	,	PUNCT
iguana-17148	16	8	guyot	guyot	PROPN
iguana-17148	16	9	hall	hall	PROPN
iguana-17148	16	10	303	303	NUM
iguana-17148	16	11	,	,	PUNCT
iguana-17148	16	12	princeton	princeton	PROPN
iguana-17148	16	13	university	university	PROPN
iguana-17148	16	14	,	,	PUNCT
iguana-17148	16	15	princeton	princeton	PROPN
iguana-17148	16	16	,	,	PUNCT
iguana-17148	16	17	nj	nj	PROPN
iguana-17148	16	18	08544	08544	NUM
iguana-17148	16	19	-	-	SYM
iguana-17148	16	20	0001	0001	NUM
iguana-17148	16	21	,	,	PUNCT
iguana-17148	16	22	usa	usa	PROPN
iguana-17148	16	23	(	(	PUNCT
iguana-17148	16	24	wikelski@princeton.edu	wikelski@princeton.edu	PROPN
iguana-17148	16	25	)	)	PUNCT
iguana-17148	17	1	2department	2department	NUM
iguana-17148	17	2	of	of	ADP
iguana-17148	17	3	biology	biology	NOUN
iguana-17148	17	4	,	,	PUNCT
iguana-17148	17	5	triton	triton	PROPN
iguana-17148	17	6	college	college	PROPN
iguana-17148	17	7	,	,	PUNCT
iguana-17148	17	8	2000	2000	NUM
iguana-17148	17	9	fifth	fifth	PROPN
iguana-17148	17	10	avenue	avenue	PROPN
iguana-17148	17	11	,	,	PUNCT
iguana-17148	17	12	river	river	NOUN
iguana-17148	17	13	grove	grove	PROPN
iguana-17148	17	14	,	,	PUNCT
iguana-17148	17	15	il	il	PROPN
iguana-17148	17	16	60171	60171	NUM
iguana-17148	17	17	,	,	PUNCT
iguana-17148	17	18	usa	usa	PROPN
iguana-17148	17	19	(	(	PUNCT
iguana-17148	17	20	knnelsonn@yahoo.com	knnelsonn@yahoo.com	NOUN
iguana-17148	17	21	)	)	PUNCT
iguana-17148	17	22	photographs	photograph	NOUN
iguana-17148	17	23	by	by	ADP
iguana-17148	17	24	martin	martin	PROPN
iguana-17148	17	25	wikelski	wikelski	PROPN
iguana-17148	17	26	except	except	SCONJ
iguana-17148	17	27	where	where	SCONJ
iguana-17148	17	28	indicated	indicate	VERB
iguana-17148	17	29	.	.	PUNCT
iguana-17148	18	1	abstract.—galápagos	abstract.—galápago	NOUN
iguana-17148	18	2	marine	marine	PROPN
iguana-17148	18	3	iguanas	iguana	NOUN
iguana-17148	18	4	are	be	AUX
iguana-17148	18	5	highly	highly	ADV
iguana-17148	18	6	abundant	abundant	ADJ
iguana-17148	18	7	along	along	ADP
iguana-17148	18	8	many	many	ADJ
iguana-17148	18	9	of	of	ADP
iguana-17148	18	10	the	the	DET
iguana-17148	18	11	archipelago	archipelago	PROPN
iguana-17148	18	12	’s	’s	PART
iguana-17148	18	13	shorelines	shoreline	NOUN
iguana-17148	18	14	.	.	PUNCT
iguana-17148	19	1	total	total	ADJ
iguana-17148	19	2	estimated	estimate	VERB
iguana-17148	19	3	population	population	NOUN
iguana-17148	19	4	size	size	NOUN
iguana-17148	19	5	varies	vary	VERB
iguana-17148	19	6	between	between	ADP
iguana-17148	19	7	37,000	37,000	NUM
iguana-17148	19	8	and	and	CCONJ
iguana-17148	19	9	280,000	280,000	NUM
iguana-17148	19	10	individuals	individual	NOUN
iguana-17148	19	11	.	.	PUNCT
iguana-17148	20	1	marine	marine	PROPN
iguana-17148	20	2	iguanas	iguana	NOUN
iguana-17148	20	3	have	have	AUX
iguana-17148	20	4	evolved	evolve	VERB
iguana-17148	20	5	in	in	ADP
iguana-17148	20	6	the	the	DET
iguana-17148	20	7	absence	absence	NOUN
iguana-17148	20	8	of	of	ADP
iguana-17148	20	9	major	major	ADJ
iguana-17148	20	10	predators	predator	NOUN
iguana-17148	20	11	,	,	PUNCT
iguana-17148	20	12	and	and	CCONJ
iguana-17148	20	13	their	their	PRON
iguana-17148	20	14	populations	population	NOUN
iguana-17148	20	15	are	be	AUX
iguana-17148	20	16	regulated	regulate	VERB
iguana-17148	20	17	by	by	ADP
iguana-17148	20	18	cyclically	cyclically	ADV
iguana-17148	20	19	recurring	recur	VERB
iguana-17148	20	20	famine	famine	NOUN
iguana-17148	20	21	(	(	PUNCT
iguana-17148	20	22	el	el	PROPN
iguana-17148	20	23	niño	niño	PROPN
iguana-17148	20	24	)	)	PUNCT
iguana-17148	20	25	and	and	CCONJ
iguana-17148	20	26	feast	feast	NOUN
iguana-17148	20	27	(	(	PUNCT
iguana-17148	20	28	la	la	X
iguana-17148	20	29	niña	niña	NOUN
iguana-17148	20	30	)	)	PUNCT
iguana-17148	20	31	events	event	NOUN
iguana-17148	20	32	.	.	PUNCT
iguana-17148	21	1	population	population	NOUN
iguana-17148	21	2	declines	decline	NOUN
iguana-17148	21	3	are	be	AUX
iguana-17148	21	4	strongly	strongly	ADV
iguana-17148	21	5	density	density	NOUN
iguana-17148	21	6	-	-	PUNCT
iguana-17148	21	7	dependent	dependent	ADJ
iguana-17148	21	8	:	:	PUNCT
iguana-17148	21	9	the	the	PRON
iguana-17148	21	10	higher	high	ADJ
iguana-17148	21	11	the	the	DET
iguana-17148	21	12	population	population	NOUN
iguana-17148	21	13	density	density	NOUN
iguana-17148	21	14	,	,	PUNCT
iguana-17148	21	15	the	the	PRON
iguana-17148	21	16	higher	high	ADJ
iguana-17148	21	17	the	the	DET
iguana-17148	21	18	mortalities	mortality	NOUN
iguana-17148	21	19	during	during	ADP
iguana-17148	21	20	el	el	PROPN
iguana-17148	21	21	niños	niños	PROPN
iguana-17148	21	22	(	(	PUNCT
iguana-17148	21	23	from	from	ADP
iguana-17148	21	24	10–90	10–90	NUM
iguana-17148	21	25	%	%	NOUN
iguana-17148	21	26	)	)	PUNCT
iguana-17148	21	27	.	.	PUNCT
iguana-17148	22	1	recovery	recovery	NOUN
iguana-17148	22	2	after	after	SCONJ
iguana-17148	22	3	el	el	PROPN
iguana-17148	22	4	niños	niños	PROPN
iguana-17148	22	5	is	be	AUX
iguana-17148	22	6	rapid	rapid	ADJ
iguana-17148	22	7	,	,	PUNCT
iguana-17148	22	8	as	as	SCONJ
iguana-17148	22	9	females	female	NOUN
iguana-17148	22	10	compensate	compensate	VERB
iguana-17148	22	11	by	by	ADP
iguana-17148	22	12	reproducing	reproduce	VERB
iguana-17148	22	13	younger	young	ADJ
iguana-17148	22	14	and	and	CCONJ
iguana-17148	22	15	laying	lay	VERB
iguana-17148	22	16	more	more	ADJ
iguana-17148	22	17	eggs	egg	NOUN
iguana-17148	22	18	.	.	PUNCT
iguana-17148	23	1	marine	marine	ADJ
iguana-17148	23	2	iguana	iguana	PROPN
iguana-17148	23	3	morphology	morphology	NOUN
iguana-17148	23	4	differs	differ	VERB
iguana-17148	23	5	between	between	ADP
iguana-17148	23	6	islands	island	NOUN
iguana-17148	23	7	.	.	PUNCT
iguana-17148	24	1	seven	seven	NUM
iguana-17148	24	2	subspecies	subspecie	NOUN
iguana-17148	24	3	have	have	AUX
iguana-17148	24	4	been	be	AUX
iguana-17148	24	5	proposed	propose	VERB
iguana-17148	24	6	,	,	PUNCT
iguana-17148	24	7	although	although	SCONJ
iguana-17148	24	8	only	only	ADV
iguana-17148	24	9	three	three	NUM
iguana-17148	24	10	major	major	ADJ
iguana-17148	24	11	clades	clade	NOUN
iguana-17148	24	12	can	can	AUX
iguana-17148	24	13	be	be	AUX
iguana-17148	24	14	distinguished	distinguish	VERB
iguana-17148	24	15	genetically	genetically	ADV
iguana-17148	24	16	.	.	PUNCT
iguana-17148	25	1	twelve	twelve	NUM
iguana-17148	25	2	populations	population	NOUN
iguana-17148	25	3	(	(	PUNCT
iguana-17148	25	4	approximately	approximately	ADV
iguana-17148	25	5	74	74	NUM
iguana-17148	25	6	%	%	NOUN
iguana-17148	25	7	of	of	ADP
iguana-17148	25	8	all	all	DET
iguana-17148	25	9	marine	marine	ADJ
iguana-17148	25	10	iguanas	iguana	NOUN
iguana-17148	25	11	)	)	PUNCT
iguana-17148	25	12	still	still	ADV
iguana-17148	25	13	live	live	VERB
iguana-17148	25	14	in	in	ADP
iguana-17148	25	15	pristine	pristine	ADJ
iguana-17148	25	16	environments	environment	NOUN
iguana-17148	25	17	,	,	PUNCT
iguana-17148	25	18	whereas	whereas	SCONJ
iguana-17148	25	19	five	five	NUM
iguana-17148	25	20	populations	population	NOUN
iguana-17148	25	21	(	(	PUNCT
iguana-17148	25	22	26	26	NUM
iguana-17148	25	23	%	%	NOUN
iguana-17148	25	24	of	of	ADP
iguana-17148	25	25	all	all	DET
iguana-17148	25	26	marine	marine	ADJ
iguana-17148	25	27	iguanas	iguana	NOUN
iguana-17148	25	28	)	)	PUNCT
iguana-17148	25	29	suffer	suffer	VERB
iguana-17148	25	30	from	from	ADP
iguana-17148	25	31	anthropogenic	anthropogenic	ADJ
iguana-17148	25	32	influences	influence	NOUN
iguana-17148	25	33	.	.	PUNCT
iguana-17148	26	1	major	major	ADJ
iguana-17148	26	2	conservation	conservation	NOUN
iguana-17148	26	3	problems	problem	NOUN
iguana-17148	26	4	arise	arise	VERB
iguana-17148	26	5	from	from	ADP
iguana-17148	26	6	introduced	introduce	VERB
iguana-17148	26	7	predators	predator	NOUN
iguana-17148	26	8	(	(	PUNCT
iguana-17148	26	9	cats	cat	NOUN
iguana-17148	26	10	,	,	PUNCT
iguana-17148	26	11	dogs	dog	NOUN
iguana-17148	26	12	,	,	PUNCT
iguana-17148	26	13	rats	rat	NOUN
iguana-17148	26	14	,	,	PUNCT
iguana-17148	26	15	and	and	CCONJ
iguana-17148	26	16	pigs	pig	NOUN
iguana-17148	26	17	)	)	PUNCT
iguana-17148	26	18	and	and	CCONJ
iguana-17148	26	19	from	from	ADP
iguana-17148	26	20	combinations	combination	NOUN
iguana-17148	26	21	of	of	ADP
iguana-17148	26	22	natural	natural	ADJ
iguana-17148	26	23	events	event	NOUN
iguana-17148	26	24	(	(	PUNCT
iguana-17148	26	25	el	el	PROPN
iguana-17148	26	26	niño	niño	PROPN
iguana-17148	26	27	)	)	PUNCT
iguana-17148	26	28	and	and	CCONJ
iguana-17148	26	29	anthropogenic	anthropogenic	ADJ
iguana-17148	26	30	disasters	disaster	NOUN
iguana-17148	26	31	such	such	ADJ
iguana-17148	26	32	as	as	ADP
iguana-17148	26	33	oil	oil	NOUN
iguana-17148	26	34	spills	spill	NOUN
iguana-17148	26	35	.	.	PUNCT
iguana-17148	27	1	the	the	DET
iguana-17148	27	2	most	most	ADV
iguana-17148	27	3	recent	recent	ADJ
iguana-17148	27	4	oil	oil	NOUN
iguana-17148	27	5	spill	spill	NOUN
iguana-17148	27	6	in	in	ADP
iguana-17148	27	7	2001	2001	NUM
iguana-17148	27	8	killed	kill	VERB
iguana-17148	27	9	62	62	NUM
iguana-17148	27	10	%	%	NOUN
iguana-17148	27	11	of	of	ADP
iguana-17148	27	12	all	all	DET
iguana-17148	27	13	marine	marine	ADJ
iguana-17148	27	14	iguanas	iguana	NOUN
iguana-17148	27	15	on	on	ADP
iguana-17148	27	16	santa	santa	PROPN
iguana-17148	27	17	fe	fe	PROPN
iguana-17148	27	18	island	island	PROPN
iguana-17148	27	19	.	.	PUNCT
iguana-17148	28	1	management	management	NOUN
iguana-17148	28	2	requirements	requirement	NOUN
iguana-17148	28	3	for	for	ADP
iguana-17148	28	4	the	the	DET
iguana-17148	28	5	future	future	NOUN
iguana-17148	28	6	include	include	VERB
iguana-17148	28	7	:	:	PUNCT
iguana-17148	28	8	(	(	PUNCT
iguana-17148	28	9	i	i	NOUN
iguana-17148	28	10	)	)	PUNCT
iguana-17148	28	11	investigating	investigate	VERB
iguana-17148	28	12	population	population	NOUN
iguana-17148	28	13	trends	trend	NOUN
iguana-17148	28	14	in	in	ADP
iguana-17148	28	15	western	western	ADJ
iguana-17148	28	16	isabela	isabela	PROPN
iguana-17148	28	17	and	and	CCONJ
iguana-17148	28	18	san	san	PROPN
iguana-17148	28	19	cristobal	cristobal	PROPN
iguana-17148	28	20	islands	islands	PROPN
iguana-17148	28	21	,	,	PUNCT
iguana-17148	28	22	(	(	PUNCT
iguana-17148	28	23	ii	ii	NOUN
iguana-17148	28	24	)	)	PUNCT
iguana-17148	28	25	investigating	investigate	VERB
iguana-17148	28	26	whether	whether	SCONJ
iguana-17148	28	27	harbor	harbor	NOUN
iguana-17148	28	28	areas	area	NOUN
iguana-17148	28	29	are	be	AUX
iguana-17148	28	30	population	population	NOUN
iguana-17148	28	31	sinks	sink	NOUN
iguana-17148	28	32	because	because	SCONJ
iguana-17148	28	33	of	of	ADP
iguana-17148	28	34	environmental	environmental	ADJ
iguana-17148	28	35	contaminants	contaminant	NOUN
iguana-17148	28	36	,	,	PUNCT
iguana-17148	28	37	(	(	PUNCT
iguana-17148	28	38	iii	iii	X
iguana-17148	28	39	)	)	PUNCT
iguana-17148	28	40	establishing	establish	VERB
iguana-17148	28	41	a	a	DET
iguana-17148	28	42	recovery	recovery	NOUN
iguana-17148	28	43	program	program	NOUN
iguana-17148	28	44	for	for	ADP
iguana-17148	28	45	oil	oil	NOUN
iguana-17148	28	46	-	-	PUNCT
iguana-17148	28	47	contaminated	contaminate	VERB
iguana-17148	28	48	iguanas	iguana	NOUN
iguana-17148	28	49	,	,	PUNCT
iguana-17148	28	50	especially	especially	ADV
iguana-17148	28	51	their	their	PRON
iguana-17148	28	52	reinoculation	reinoculation	NOUN
iguana-17148	28	53	with	with	ADP
iguana-17148	28	54	hindgut	hindgut	NOUN
iguana-17148	28	55	microsymbionts	microsymbiont	NOUN
iguana-17148	28	56	,	,	PUNCT
iguana-17148	28	57	and	and	CCONJ
iguana-17148	28	58	(	(	PUNCT
iguana-17148	28	59	iv	iv	X
iguana-17148	28	60	)	)	PUNCT
iguana-17148	28	61	developing	develop	VERB
iguana-17148	28	62	husbandry	husbandry	NOUN
iguana-17148	28	63	techniques	technique	NOUN
iguana-17148	28	64	and	and	CCONJ
iguana-17148	28	65	establishing	establish	VERB
iguana-17148	28	66	a	a	DET
iguana-17148	28	67	captive	captive	ADJ
iguana-17148	28	68	propagation	propagation	NOUN
iguana-17148	28	69	program	program	NOUN
iguana-17148	28	70	as	as	ADP
iguana-17148	28	71	a	a	DET
iguana-17148	28	72	population	population	NOUN
iguana-17148	28	73	backup	backup	NOUN
iguana-17148	28	74	plan	plan	NOUN
iguana-17148	28	75	(	(	PUNCT
iguana-17148	28	76	marine	marine	ADJ
iguana-17148	28	77	iguanas	iguana	NOUN
iguana-17148	28	78	have	have	AUX
iguana-17148	28	79	not	not	PART
iguana-17148	28	80	been	be	AUX
iguana-17148	28	81	bred	breed	VERB
iguana-17148	28	82	in	in	ADP
iguana-17148	28	83	captivity	captivity	NOUN
iguana-17148	28	84	)	)	PUNCT
iguana-17148	28	85	.	.	PUNCT
iguana-17148	29	1	key	key	ADJ
iguana-17148	29	2	words	word	NOUN
iguana-17148	29	3	:	:	PUNCT
iguana-17148	29	4	marine	marine	ADJ
iguana-17148	29	5	iguanas	iguana	NOUN
iguana-17148	29	6	,	,	PUNCT
iguana-17148	29	7	amblyrhynchus	amblyrhynchus	NOUN
iguana-17148	29	8	cristatus	cristatus	PROPN
iguana-17148	29	9	,	,	PUNCT
iguana-17148	29	10	south	south	PROPN
iguana-17148	29	11	america	america	PROPN
iguana-17148	29	12	,	,	PUNCT
iguana-17148	29	13	galápagos	galápagos	PROPN
iguana-17148	29	14	archipelago	archipelago	PROPN
iguana-17148	29	15	,	,	PUNCT
iguana-17148	29	16	conservation	conservation	NOUN
iguana-17148	29	17	,	,	PUNCT
iguana-17148	29	18	oil	oil	NOUN
iguana-17148	29	19	spill	spill	NOUN
iguana-17148	29	20	,	,	PUNCT
iguana-17148	29	21	feral	feral	ADJ
iguana-17148	29	22	predators	predator	NOUN
iguana-17148	29	23	iguana	iguana	NOUN
iguana-17148	29	24	•	•	NUM
iguana-17148	29	25	volume	volume	NOUN
iguana-17148	29	26	11	11	NUM
iguana-17148	29	27	,	,	PUNCT
iguana-17148	29	28	number	number	NOUN
iguana-17148	29	29	4	4	NUM
iguana-17148	29	30	•	•	NOUN
iguana-17148	29	31	december	december	PROPN
iguana-17148	29	32	2004	2004	NUM
iguana-17148	29	33	191galápagos	191galápagos	PROPN
iguana-17148	29	34	marine	marine	PROPN
iguana-17148	29	35	iguanas	iguana	NOUN
iguana-17148	29	36	�	�	PROPN
iguana-17148	29	37	male	male	ADJ
iguana-17148	29	38	marine	marine	ADJ
iguana-17148	29	39	iguana	iguana	NOUN
iguana-17148	29	40	on	on	ADP
iguana-17148	29	41	seymour	seymour	PROPN
iguana-17148	29	42	norte	norte	PROPN
iguana-17148	29	43	island	island	PROPN
iguana-17148	29	44	eating	eat	VERB
iguana-17148	29	45	saltwort	saltwort	NOUN
iguana-17148	29	46	(	(	PUNCT
iguana-17148	29	47	batis	batis	NOUN
iguana-17148	29	48	maritima	maritima	NOUN
iguana-17148	29	49	)	)	PUNCT
iguana-17148	29	50	on	on	ADP
iguana-17148	29	51	land	land	NOUN
iguana-17148	29	52	.	.	PUNCT
iguana-17148	30	1	some	some	DET
iguana-17148	30	2	individuals	individual	NOUN
iguana-17148	30	3	supplement	supplement	VERB
iguana-17148	30	4	their	their	PRON
iguana-17148	30	5	food	food	NOUN
iguana-17148	30	6	with	with	ADP
iguana-17148	30	7	land	land	NOUN
iguana-17148	30	8	plants	plant	NOUN
iguana-17148	30	9	.	.	PUNCT
iguana-17148	31	1	hau	hau	NOUN
iguana-17148	31	2	1995	1995	NUM
iguana-17148	31	3	)	)	PUNCT
iguana-17148	31	4	.	.	PUNCT
iguana-17148	32	1	the	the	DET
iguana-17148	32	2	largest	large	ADJ
iguana-17148	32	3	iguanas	iguana	NOUN
iguana-17148	32	4	of	of	ADP
iguana-17148	32	5	each	each	DET
iguana-17148	32	6	island	island	NOUN
iguana-17148	32	7	population	population	NOUN
iguana-17148	32	8	also	also	ADV
iguana-17148	32	9	dive	dive	VERB
iguana-17148	32	10	for	for	ADP
iguana-17148	32	11	algae	algae	NOUN
iguana-17148	32	12	(	(	PUNCT
iguana-17148	32	13	2–30	2–30	NOUN
iguana-17148	32	14	m	m	VERB
iguana-17148	32	15	depth	depth	NOUN
iguana-17148	32	16	;	;	PUNCT
iguana-17148	32	17	buttemer	buttemer	NOUN
iguana-17148	32	18	and	and	CCONJ
iguana-17148	32	19	dawson	dawson	PROPN
iguana-17148	32	20	1993	1993	NUM
iguana-17148	32	21	)	)	PUNCT
iguana-17148	32	22	.	.	PUNCT
iguana-17148	33	1	on	on	ADP
iguana-17148	33	2	genovesa	genovesa	PROPN
iguana-17148	33	3	,	,	PUNCT
iguana-17148	33	4	males	male	NOUN
iguana-17148	33	5	with	with	ADP
iguana-17148	33	6	body	body	NOUN
iguana-17148	33	7	mass	mass	NOUN
iguana-17148	33	8	>	>	PROPN
iguana-17148	33	9	500	500	NUM
iguana-17148	33	10	g	g	NOUN
iguana-17148	33	11	are	be	AUX
iguana-17148	33	12	seen	see	VERB
iguana-17148	33	13	diving	diving	NOUN
iguana-17148	33	14	,	,	PUNCT
iguana-17148	33	15	whereas	whereas	SCONJ
iguana-17148	33	16	on	on	ADP
iguana-17148	33	17	fernandina	fernandina	PROPN
iguana-17148	33	18	usually	usually	ADV
iguana-17148	33	19	only	only	ADV
iguana-17148	33	20	males	male	NOUN
iguana-17148	33	21	>	>	X
iguana-17148	33	22	3500	3500	NUM
iguana-17148	33	23	g	g	NOUN
iguana-17148	33	24	dive	dive	NOUN
iguana-17148	33	25	for	for	ADP
iguana-17148	33	26	food	food	NOUN
iguana-17148	33	27	(	(	PUNCT
iguana-17148	33	28	wikelski	wikelski	NOUN
iguana-17148	33	29	and	and	CCONJ
iguana-17148	33	30	trillmich	trillmich	NOUN
iguana-17148	33	31	1994	1994	NUM
iguana-17148	33	32	)	)	PUNCT
iguana-17148	33	33	.	.	PUNCT
iguana-17148	34	1	a	a	DET
iguana-17148	34	2	few	few	ADJ
iguana-17148	34	3	individuals	individual	NOUN
iguana-17148	34	4	supplement	supplement	VERB
iguana-17148	34	5	their	their	PRON
iguana-17148	34	6	food	food	NOUN
iguana-17148	34	7	with	with	ADP
iguana-17148	34	8	land	land	NOUN
iguana-17148	34	9	plants	plant	NOUN
iguana-17148	34	10	,	,	PUNCT
iguana-17148	34	11	in	in	ADP
iguana-17148	34	12	particular	particular	ADJ
iguana-17148	34	13	on	on	ADP
iguana-17148	34	14	seymour	seymour	PROPN
iguana-17148	34	15	norte	norte	PROPN
iguana-17148	34	16	island	island	NOUN
iguana-17148	34	17	.	.	PUNCT
iguana-17148	35	1	only	only	ADV
iguana-17148	35	2	highly	highly	ADV
iguana-17148	35	3	salty	salty	ADJ
iguana-17148	35	4	land	land	NOUN
iguana-17148	35	5	plants	plant	NOUN
iguana-17148	35	6	are	be	AUX
iguana-17148	35	7	ingested	ingest	VERB
iguana-17148	35	8	(	(	PUNCT
iguana-17148	35	9	primarily	primarily	ADV
iguana-17148	35	10	saltwort	saltwort	NOUN
iguana-17148	35	11	,	,	PUNCT
iguana-17148	35	12	batis	batis	NOUN
iguana-17148	35	13	maritima	maritima	NOUN
iguana-17148	35	14	,	,	PUNCT
iguana-17148	35	15	but	but	CCONJ
iguana-17148	35	16	also	also	ADV
iguana-17148	35	17	other	other	ADJ
iguana-17148	35	18	coastal	coastal	ADJ
iguana-17148	35	19	succulents	succulent	NOUN
iguana-17148	35	20	such	such	ADJ
iguana-17148	35	21	as	as	ADP
iguana-17148	35	22	sesuvium	sesuvium	PROPN
iguana-17148	35	23	portulacastrum	portulacastrum	PROPN
iguana-17148	35	24	)	)	PUNCT
iguana-17148	35	25	,	,	PUNCT
iguana-17148	35	26	presumably	presumably	ADV
iguana-17148	35	27	because	because	SCONJ
iguana-17148	35	28	marine	marine	ADJ
iguana-17148	35	29	iguanas	iguana	NOUN
iguana-17148	35	30	possess	possess	VERB
iguana-17148	35	31	very	very	ADV
iguana-17148	35	32	specialized	specialized	ADJ
iguana-17148	35	33	hindgut	hindgut	NOUN
iguana-17148	35	34	micro	micro	NOUN
iguana-17148	35	35	-	-	NOUN
iguana-17148	35	36	symbionts	symbiont	NOUN
iguana-17148	35	37	that	that	PRON
iguana-17148	35	38	help	help	VERB
iguana-17148	35	39	them	they	PRON
iguana-17148	35	40	digest	digest	VERB
iguana-17148	35	41	their	their	PRON
iguana-17148	35	42	food	food	NOUN
iguana-17148	35	43	and	and	CCONJ
iguana-17148	35	44	effectively	effectively	ADV
iguana-17148	35	45	break	break	VERB
iguana-17148	35	46	up	up	ADP
iguana-17148	35	47	cell	cell	NOUN
iguana-17148	35	48	walls	wall	NOUN
iguana-17148	35	49	(	(	PUNCT
iguana-17148	35	50	mackie	mackie	PROPN
iguana-17148	35	51	et	et	PROPN
iguana-17148	35	52	al	al	PROPN
iguana-17148	35	53	.	.	PROPN
iguana-17148	35	54	2003	2003	NUM
iguana-17148	35	55	)	)	PUNCT
iguana-17148	35	56	.	.	PUNCT
iguana-17148	36	1	marine	marine	PROPN
iguana-17148	36	2	iguanas	iguanas	PROPN
iguana-17148	36	3	reproduce	reproduce	VERB
iguana-17148	36	4	once	once	ADV
iguana-17148	36	5	a	a	DET
iguana-17148	36	6	year	year	NOUN
iguana-17148	36	7	during	during	ADP
iguana-17148	36	8	a	a	DET
iguana-17148	36	9	month	month	NOUN
iguana-17148	36	10	-	-	PUNCT
iguana-17148	36	11	long	long	ADJ
iguana-17148	36	12	mating	mating	NOUN
iguana-17148	36	13	season	season	NOUN
iguana-17148	36	14	.	.	PUNCT
iguana-17148	37	1	the	the	DET
iguana-17148	37	2	precise	precise	ADJ
iguana-17148	37	3	timing	timing	NOUN
iguana-17148	37	4	of	of	ADP
iguana-17148	37	5	the	the	DET
iguana-17148	37	6	mating	mating	NOUN
iguana-17148	37	7	season	season	NOUN
iguana-17148	37	8	coincides	coincide	VERB
iguana-17148	37	9	with	with	ADP
iguana-17148	37	10	the	the	DET
iguana-17148	37	11	highest	high	ADJ
iguana-17148	37	12	abundance	abundance	NOUN
iguana-17148	37	13	and	and	CCONJ
iguana-17148	37	14	best	good	ADJ
iguana-17148	37	15	quality	quality	NOUN
iguana-17148	37	16	of	of	ADP
iguana-17148	37	17	food	food	NOUN
iguana-17148	37	18	(	(	PUNCT
iguana-17148	37	19	rubenstein	rubenstein	PROPN
iguana-17148	37	20	and	and	CCONJ
iguana-17148	37	21	wikelski	wikelski	PROPN
iguana-17148	37	22	,	,	PUNCT
iguana-17148	37	23	in	in	ADP
iguana-17148	37	24	preparation	preparation	NOUN
iguana-17148	37	25	)	)	PUNCT
iguana-17148	37	26	.	.	PUNCT
iguana-17148	38	1	because	because	SCONJ
iguana-17148	38	2	the	the	DET
iguana-17148	38	3	nutrient	nutrient	NOUN
iguana-17148	38	4	-	-	PUNCT
iguana-17148	38	5	rich	rich	ADJ
iguana-17148	38	6	upwelling	upwelling	NOUN
iguana-17148	38	7	from	from	ADP
iguana-17148	38	8	the	the	DET
iguana-17148	38	9	cromwell	cromwell	PROPN
iguana-17148	38	10	current	current	NOUN
iguana-17148	38	11	affects	affect	VERB
iguana-17148	38	12	all	all	DET
iguana-17148	38	13	islands	island	NOUN
iguana-17148	38	14	in	in	ADP
iguana-17148	38	15	the	the	DET
iguana-17148	38	16	archipelago	archipelago	NOUN
iguana-17148	38	17	differently	differently	ADV
iguana-17148	38	18	,	,	PUNCT
iguana-17148	38	19	mating	mating	NOUN
iguana-17148	38	20	seasons	season	NOUN
iguana-17148	38	21	occur	occur	VERB
iguana-17148	38	22	at	at	ADP
iguana-17148	38	23	different	different	ADJ
iguana-17148	38	24	times	time	NOUN
iguana-17148	38	25	(	(	PUNCT
iguana-17148	38	26	e.g.	e.g.	ADV
iguana-17148	38	27	,	,	PUNCT
iguana-17148	38	28	december	december	PROPN
iguana-17148	38	29	on	on	ADP
iguana-17148	38	30	santa	santa	PROPN
iguana-17148	38	31	fe	fe	PROPN
iguana-17148	38	32	and	and	CCONJ
iguana-17148	38	33	genovesa	genovesa	PROPN
iguana-17148	38	34	,	,	PUNCT
iguana-17148	38	35	january	january	PROPN
iguana-17148	38	36	on	on	ADP
iguana-17148	38	37	santa	santa	PROPN
iguana-17148	38	38	cruz	cruz	PROPN
iguana-17148	38	39	,	,	PUNCT
iguana-17148	38	40	february	february	PROPN
iguana-17148	38	41	/	/	SYM
iguana-17148	38	42	march	march	PROPN
iguana-17148	38	43	on	on	ADP
iguana-17148	38	44	española	española	PROPN
iguana-17148	38	45	)	)	PUNCT
iguana-17148	38	46	.	.	PUNCT
iguana-17148	39	1	during	during	ADP
iguana-17148	39	2	the	the	DET
iguana-17148	39	3	mating	mating	NOUN
iguana-17148	39	4	season	season	NOUN
iguana-17148	39	5	,	,	PUNCT
iguana-17148	39	6	males	male	NOUN
iguana-17148	39	7	defend	defend	VERB
iguana-17148	39	8	small	small	ADJ
iguana-17148	39	9	territories	territory	NOUN
iguana-17148	39	10	that	that	PRON
iguana-17148	39	11	contain	contain	VERB
iguana-17148	39	12	no	no	DET
iguana-17148	39	13	resources	resource	NOUN
iguana-17148	39	14	other	other	ADJ
iguana-17148	39	15	than	than	ADP
iguana-17148	39	16	the	the	DET
iguana-17148	39	17	males	male	NOUN
iguana-17148	39	18	themselves	themselves	PRON
iguana-17148	39	19	,	,	PUNCT
iguana-17148	39	20	which	which	PRON
iguana-17148	39	21	prompted	prompt	VERB
iguana-17148	39	22	the	the	DET
iguana-17148	39	23	description	description	NOUN
iguana-17148	39	24	of	of	ADP
iguana-17148	39	25	the	the	DET
iguana-17148	39	26	mating	mating	NOUN
iguana-17148	39	27	system	system	NOUN
iguana-17148	39	28	as	as	ADP
iguana-17148	39	29	a	a	DET
iguana-17148	39	30	lek	lek	NOUN
iguana-17148	39	31	,	,	PUNCT
iguana-17148	39	32	or	or	CCONJ
iguana-17148	39	33	mating	mating	NOUN
iguana-17148	39	34	arena	arena	NOUN
iguana-17148	39	35	(	(	PUNCT
iguana-17148	39	36	trillmich	trillmich	NOUN
iguana-17148	39	37	1983	1983	NUM
iguana-17148	39	38	,	,	PUNCT
iguana-17148	39	39	wikelski	wikelski	PROPN
iguana-17148	39	40	et	et	PROPN
iguana-17148	39	41	al	al	PROPN
iguana-17148	39	42	.	.	PROPN
iguana-17148	39	43	1996	1996	NUM
iguana-17148	39	44	)	)	PUNCT
iguana-17148	39	45	.	.	PUNCT
iguana-17148	40	1	male	male	ADJ
iguana-17148	40	2	marine	marine	ADJ
iguana-17148	40	3	iguanas	iguana	NOUN
iguana-17148	40	4	use	use	VERB
iguana-17148	40	5	three	three	NUM
iguana-17148	40	6	different	different	ADJ
iguana-17148	40	7	mating	mating	NOUN
iguana-17148	40	8	strategies	strategy	NOUN
iguana-17148	40	9	.	.	PUNCT
iguana-17148	41	1	the	the	DET
iguana-17148	41	2	largest	large	ADJ
iguana-17148	41	3	males	male	NOUN
iguana-17148	41	4	defend	defend	VERB
iguana-17148	41	5	territories	territory	NOUN
iguana-17148	41	6	and	and	CCONJ
iguana-17148	41	7	court	court	NOUN
iguana-17148	41	8	females	female	NOUN
iguana-17148	41	9	using	use	VERB
iguana-17148	41	10	a	a	DET
iguana-17148	41	11	slow	slow	ADJ
iguana-17148	41	12	,	,	PUNCT
iguana-17148	41	13	stereotyped	stereotype	VERB
iguana-17148	41	14	head	head	NOUN
iguana-17148	41	15	-	-	PUNCT
iguana-17148	41	16	bob	bob	NOUN
iguana-17148	41	17	courtship	courtship	NOUN
iguana-17148	41	18	behavior	behavior	NOUN
iguana-17148	41	19	.	.	PUNCT
iguana-17148	42	1	male	male	ADJ
iguana-17148	42	2	mating	mating	NOUN
iguana-17148	42	3	success	success	NOUN
iguana-17148	42	4	is	be	AUX
iguana-17148	42	5	highly	highly	ADV
iguana-17148	42	6	skewed	skewed	ADJ
iguana-17148	42	7	and	and	CCONJ
iguana-17148	42	8	depends	depend	VERB
iguana-17148	42	9	on	on	ADP
iguana-17148	42	10	body	body	NOUN
iguana-17148	42	11	size	size	NOUN
iguana-17148	42	12	,	,	PUNCT
iguana-17148	42	13	condition	condition	NOUN
iguana-17148	42	14	,	,	PUNCT
iguana-17148	42	15	and	and	CCONJ
iguana-17148	42	16	display	display	NOUN
iguana-17148	42	17	rate	rate	NOUN
iguana-17148	42	18	(	(	PUNCT
iguana-17148	42	19	wikelski	wikelski	NOUN
iguana-17148	42	20	et	et	PROPN
iguana-17148	42	21	al	al	PROPN
iguana-17148	42	22	.	.	PROPN
iguana-17148	42	23	1996	1996	NUM
iguana-17148	42	24	,	,	PUNCT
iguana-17148	42	25	2001	2001	NUM
iguana-17148	42	26	)	)	PUNCT
iguana-17148	42	27	.	.	PUNCT
iguana-17148	43	1	male	male	ADJ
iguana-17148	43	2	territories	territory	NOUN
iguana-17148	43	3	are	be	AUX
iguana-17148	43	4	generally	generally	ADV
iguana-17148	43	5	clustered	cluster	VERB
iguana-17148	43	6	,	,	PUNCT
iguana-17148	43	7	but	but	CCONJ
iguana-17148	43	8	single	single	ADJ
iguana-17148	43	9	territories	territory	NOUN
iguana-17148	43	10	also	also	ADV
iguana-17148	43	11	occur	occur	VERB
iguana-17148	43	12	.	.	PUNCT
iguana-17148	44	1	satellite	satellite	NOUN
iguana-17148	44	2	males	male	NOUN
iguana-17148	44	3	are	be	AUX
iguana-17148	44	4	smaller	small	ADJ
iguana-17148	44	5	than	than	ADP
iguana-17148	44	6	territorial	territorial	ADJ
iguana-17148	44	7	males	male	NOUN
iguana-17148	44	8	and	and	CCONJ
iguana-17148	44	9	roam	roam	VERB
iguana-17148	44	10	around	around	ADP
iguana-17148	44	11	territories	territory	NOUN
iguana-17148	44	12	,	,	PUNCT
iguana-17148	44	13	attempting	attempt	VERB
iguana-17148	44	14	to	to	PART
iguana-17148	44	15	(	(	PUNCT
iguana-17148	44	16	forcibly	forcibly	ADV
iguana-17148	44	17	)	)	PUNCT
iguana-17148	44	18	mate	mate	NOUN
iguana-17148	44	19	with	with	ADP
iguana-17148	44	20	females	female	NOUN
iguana-17148	44	21	.	.	PUNCT
iguana-17148	45	1	“	"	PUNCT
iguana-17148	45	2	sneaker	sneaker	NOUN
iguana-17148	45	3	”	"	PUNCT
iguana-17148	45	4	males	male	NOUN
iguana-17148	45	5	are	be	AUX
iguana-17148	45	6	the	the	DET
iguana-17148	45	7	smallest	small	ADJ
iguana-17148	45	8	males	male	NOUN
iguana-17148	45	9	,	,	PUNCT
iguana-17148	45	10	physically	physically	ADV
iguana-17148	45	11	indistinguishable	indistinguishable	ADJ
iguana-17148	45	12	from	from	ADP
iguana-17148	45	13	females	female	NOUN
iguana-17148	45	14	.	.	PUNCT
iguana-17148	46	1	sneakers	sneaker	NOUN
iguana-17148	46	2	try	try	VERB
iguana-17148	46	3	to	to	PART
iguana-17148	46	4	copulate	copulate	VERB
iguana-17148	46	5	with	with	ADP
iguana-17148	46	6	females	female	NOUN
iguana-17148	46	7	“	"	PUNCT
iguana-17148	46	8	in	in	ADP
iguana-17148	46	9	secret	secret	ADJ
iguana-17148	46	10	”	"	PUNCT
iguana-17148	46	11	on	on	ADP
iguana-17148	46	12	territories	territory	NOUN
iguana-17148	46	13	of	of	ADP
iguana-17148	46	14	large	large	ADJ
iguana-17148	46	15	males	male	NOUN
iguana-17148	46	16	(	(	PUNCT
iguana-17148	46	17	wikelski	wikelski	NOUN
iguana-17148	46	18	and	and	CCONJ
iguana-17148	46	19	bäurle	bäurle	NOUN
iguana-17148	46	20	1996	1996	NUM
iguana-17148	46	21	)	)	PUNCT
iguana-17148	46	22	.	.	PUNCT
iguana-17148	47	1	these	these	DET
iguana-17148	47	2	three	three	NUM
iguana-17148	47	3	mating	mating	NOUN
iguana-17148	47	4	tactics	tactic	NOUN
iguana-17148	47	5	appear	appear	VERB
iguana-17148	47	6	to	to	PART
iguana-17148	47	7	be	be	AUX
iguana-17148	47	8	partially	partially	ADV
iguana-17148	47	9	regulated	regulate	VERB
iguana-17148	47	10	by	by	ADP
iguana-17148	47	11	plasma	plasma	NOUN
iguana-17148	47	12	levels	level	NOUN
iguana-17148	47	13	of	of	ADP
iguana-17148	47	14	testosterone	testosterone	NOUN
iguana-17148	47	15	,	,	PUNCT
iguana-17148	47	16	and	and	CCONJ
iguana-17148	47	17	can	can	AUX
iguana-17148	47	18	be	be	AUX
iguana-17148	47	19	manipulated	manipulate	VERB
iguana-17148	47	20	by	by	ADP
iguana-17148	47	21	hormone	hormone	NOUN
iguana-17148	47	22	administration	administration	NOUN
iguana-17148	47	23	(	(	PUNCT
iguana-17148	47	24	wikelski	wikelski	PROPN
iguana-17148	47	25	et	et	PROPN
iguana-17148	47	26	al	al	PROPN
iguana-17148	47	27	.	.	PROPN
iguana-17148	47	28	,	,	PUNCT
iguana-17148	47	29	submitted	submit	VERB
iguana-17148	47	30	)	)	PUNCT
iguana-17148	47	31	.	.	PUNCT
iguana-17148	48	1	receptive	receptive	ADJ
iguana-17148	48	2	females	female	NOUN
iguana-17148	48	3	generally	generally	ADV
iguana-17148	48	4	copulate	copulate	VERB
iguana-17148	48	5	only	only	ADV
iguana-17148	48	6	once	once	ADV
iguana-17148	48	7	after	after	SCONJ
iguana-17148	48	8	they	they	PRON
iguana-17148	48	9	have	have	AUX
iguana-17148	48	10	selected	select	VERB
iguana-17148	48	11	a	a	DET
iguana-17148	48	12	specific	specific	ADJ
iguana-17148	48	13	male	male	NOUN
iguana-17148	48	14	,	,	PUNCT
iguana-17148	48	15	which	which	PRON
iguana-17148	48	16	they	they	PRON
iguana-17148	48	17	do	do	VERB
iguana-17148	48	18	after	after	ADP
iguana-17148	48	19	long	long	ADJ
iguana-17148	48	20	periods	period	NOUN
iguana-17148	48	21	of	of	ADP
iguana-17148	48	22	mate	mate	NOUN
iguana-17148	48	23	choice	choice	NOUN
iguana-17148	48	24	.	.	PUNCT
iguana-17148	49	1	mate	mate	NOUN
iguana-17148	49	2	choice	choice	NOUN
iguana-17148	49	3	is	be	AUX
iguana-17148	49	4	apparently	apparently	ADV
iguana-17148	49	5	costly	costly	ADJ
iguana-17148	49	6	for	for	ADP
iguana-17148	49	7	marine	marine	ADJ
iguana-17148	49	8	iguanas	iguana	NOUN
iguana-17148	49	9	,	,	PUNCT
iguana-17148	49	10	as	as	SCONJ
iguana-17148	49	11	indicated	indicate	VERB
iguana-17148	49	12	by	by	ADP
iguana-17148	49	13	mass	mass	ADJ
iguana-17148	49	14	loss	loss	NOUN
iguana-17148	49	15	of	of	ADP
iguana-17148	49	16	females	female	NOUN
iguana-17148	49	17	that	that	PRON
iguana-17148	49	18	visit	visit	VERB
iguana-17148	49	19	many	many	ADJ
iguana-17148	49	20	males	male	NOUN
iguana-17148	49	21	,	,	PUNCT
iguana-17148	49	22	compared	compare	VERB
iguana-17148	49	23	to	to	ADP
iguana-17148	49	24	those	those	PRON
iguana-17148	49	25	that	that	PRON
iguana-17148	49	26	visit	visit	VERB
iguana-17148	49	27	only	only	ADV
iguana-17148	49	28	a	a	DET
iguana-17148	49	29	few	few	ADJ
iguana-17148	49	30	males	male	NOUN
iguana-17148	49	31	or	or	CCONJ
iguana-17148	49	32	mate	mate	VERB
iguana-17148	49	33	in	in	ADP
iguana-17148	49	34	low	low	ADJ
iguana-17148	49	35	-	-	PUNCT
iguana-17148	49	36	density	density	NOUN
iguana-17148	49	37	areas	area	NOUN
iguana-17148	49	38	(	(	PUNCT
iguana-17148	49	39	wikelski	wikelski	NOUN
iguana-17148	49	40	et	et	PROPN
iguana-17148	49	41	al	al	PROPN
iguana-17148	49	42	.	.	PROPN
iguana-17148	49	43	2001	2001	NUM
iguana-17148	49	44	)	)	PUNCT
iguana-17148	49	45	.	.	PUNCT
iguana-17148	50	1	females	female	NOUN
iguana-17148	50	2	leave	leave	VERB
iguana-17148	50	3	the	the	DET
iguana-17148	50	4	mating	mating	NOUN
iguana-17148	50	5	area	area	NOUN
iguana-17148	50	6	shortly	shortly	ADV
iguana-17148	50	7	after	after	SCONJ
iguana-17148	50	8	copulation	copulation	NOUN
iguana-17148	50	9	to	to	PART
iguana-17148	50	10	lay	lay	VERB
iguana-17148	50	11	one	one	NUM
iguana-17148	50	12	to	to	ADP
iguana-17148	50	13	six	six	NUM
iguana-17148	50	14	eggs	egg	NOUN
iguana-17148	50	15	in	in	ADP
iguana-17148	50	16	deep	deep	ADJ
iguana-17148	50	17	burrows	burrow	NOUN
iguana-17148	50	18	in	in	ADP
iguana-17148	50	19	sandy	sandy	ADJ
iguana-17148	50	20	areas	area	NOUN
iguana-17148	50	21	.	.	PUNCT
iguana-17148	51	1	eggs	egg	NOUN
iguana-17148	51	2	incubate	incubate	VERB
iguana-17148	51	3	for	for	ADP
iguana-17148	51	4	three	three	NUM
iguana-17148	51	5	months	month	NOUN
iguana-17148	51	6	(	(	PUNCT
iguana-17148	51	7	laurie	laurie	PROPN
iguana-17148	51	8	1990	1990	NUM
iguana-17148	51	9	;	;	PUNCT
iguana-17148	51	10	laurie	laurie	PROPN
iguana-17148	51	11	and	and	CCONJ
iguana-17148	51	12	brown	brown	ADJ
iguana-17148	51	13	1990a	1990a	NOUN
iguana-17148	51	14	,	,	PUNCT
iguana-17148	51	15	1990b	1990b	NUM
iguana-17148	51	16	)	)	PUNCT
iguana-17148	51	17	.	.	PUNCT
iguana-17148	52	1	some	some	DET
iguana-17148	52	2	females	female	NOUN
iguana-17148	52	3	guard	guard	VERB
iguana-17148	52	4	their	their	PRON
iguana-17148	52	5	nests	nest	NOUN
iguana-17148	52	6	for	for	ADP
iguana-17148	52	7	a	a	DET
iguana-17148	52	8	few	few	ADJ
iguana-17148	52	9	days	day	NOUN
iguana-17148	52	10	after	after	ADP
iguana-17148	52	11	egg	egg	NOUN
iguana-17148	52	12	-	-	PUNCT
iguana-17148	52	13	laying	lay	VERB
iguana-17148	52	14	,	,	PUNCT
iguana-17148	52	15	mostly	mostly	ADV
iguana-17148	52	16	to	to	PART
iguana-17148	52	17	defend	defend	VERB
iguana-17148	52	18	against	against	ADP
iguana-17148	52	19	other	other	ADJ
iguana-17148	52	20	females	female	NOUN
iguana-17148	52	21	that	that	PRON
iguana-17148	52	22	try	try	VERB
iguana-17148	52	23	to	to	PART
iguana-17148	52	24	dig	dig	VERB
iguana-17148	52	25	at	at	ADP
iguana-17148	52	26	the	the	DET
iguana-17148	52	27	same	same	ADJ
iguana-17148	52	28	spot	spot	NOUN
iguana-17148	52	29	.	.	PUNCT
iguana-17148	53	1	the	the	DET
iguana-17148	53	2	entire	entire	ADJ
iguana-17148	53	3	clutch	clutch	NOUN
iguana-17148	53	4	amounts	amount	NOUN
iguana-17148	53	5	to	to	ADP
iguana-17148	53	6	about	about	ADP
iguana-17148	53	7	20–28	20–28	NUM
iguana-17148	53	8	%	%	NOUN
iguana-17148	53	9	of	of	ADP
iguana-17148	53	10	a	a	DET
iguana-17148	53	11	female	female	NOUN
iguana-17148	53	12	’s	’s	PART
iguana-17148	53	13	body	body	NOUN
iguana-17148	53	14	mass	mass	NOUN
iguana-17148	53	15	.	.	PUNCT
iguana-17148	54	1	both	both	DET
iguana-17148	54	2	males	male	NOUN
iguana-17148	54	3	and	and	CCONJ
iguana-17148	54	4	females	female	NOUN
iguana-17148	54	5	typically	typically	ADV
iguana-17148	54	6	reproduce	reproduce	VERB
iguana-17148	54	7	every	every	DET
iguana-17148	54	8	other	other	ADJ
iguana-17148	54	9	year	year	NOUN
iguana-17148	54	10	,	,	PUNCT
iguana-17148	54	11	replenishing	replenish	VERB
iguana-17148	54	12	their	their	PRON
iguana-17148	54	13	energy	energy	NOUN
iguana-17148	54	14	reserves	reserve	NOUN
iguana-17148	54	15	in	in	ADP
iguana-17148	54	16	the	the	DET
iguana-17148	54	17	year	year	NOUN
iguana-17148	54	18	they	they	PRON
iguana-17148	54	19	do	do	AUX
iguana-17148	54	20	not	not	PART
iguana-17148	54	21	reproduce	reproduce	VERB
iguana-17148	54	22	.	.	PUNCT
iguana-17148	55	1	however	however	ADV
iguana-17148	55	2	,	,	PUNCT
iguana-17148	55	3	during	during	ADP
iguana-17148	55	4	periods	period	NOUN
iguana-17148	55	5	of	of	ADP
iguana-17148	55	6	food	food	NOUN
iguana-17148	55	7	abundance	abundance	NOUN
iguana-17148	55	8	,	,	PUNCT
iguana-17148	55	9	females	female	NOUN
iguana-17148	55	10	may	may	AUX
iguana-17148	55	11	reproduce	reproduce	VERB
iguana-17148	55	12	annually	annually	ADV
iguana-17148	55	13	(	(	PUNCT
iguana-17148	55	14	laurie	laurie	PROPN
iguana-17148	55	15	1990	1990	NUM
iguana-17148	55	16	)	)	PUNCT
iguana-17148	55	17	.	.	PUNCT
iguana-17148	56	1	marine	marine	PROPN
iguana-17148	56	2	iguanas	iguana	NOUN
iguana-17148	56	3	have	have	VERB
iguana-17148	56	4	only	only	ADV
iguana-17148	56	5	one	one	NUM
iguana-17148	56	6	natural	natural	ADJ
iguana-17148	56	7	predator	predator	NOUN
iguana-17148	56	8	,	,	PUNCT
iguana-17148	56	9	the	the	DET
iguana-17148	56	10	galápagos	galápagos	PROPN
iguana-17148	56	11	hawk	hawk	NOUN
iguana-17148	56	12	(	(	PUNCT
iguana-17148	56	13	buteo	buteo	NOUN
iguana-17148	56	14	galapagoensis	galapagoensis	NOUN
iguana-17148	56	15	)	)	PUNCT
iguana-17148	56	16	,	,	PUNCT
iguana-17148	56	17	which	which	PRON
iguana-17148	56	18	is	be	AUX
iguana-17148	56	19	generally	generally	ADV
iguana-17148	56	20	unable	unable	ADJ
iguana-17148	56	21	to	to	PART
iguana-17148	56	22	prey	prey	VERB
iguana-17148	56	23	on	on	ADP
iguana-17148	56	24	healthy	healthy	ADJ
iguana-17148	56	25	adults	adult	NOUN
iguana-17148	56	26	.	.	PUNCT
iguana-17148	57	1	however	however	ADV
iguana-17148	57	2	,	,	PUNCT
iguana-17148	57	3	hawks	hawk	NOUN
iguana-17148	57	4	can	can	AUX
iguana-17148	57	5	capture	capture	AUX
iguana-17148	57	6	weakened	weaken	VERB
iguana-17148	57	7	adult	adult	NOUN
iguana-17148	57	8	iguanas	iguana	NOUN
iguana-17148	57	9	,	,	PUNCT
iguana-17148	57	10	such	such	ADJ
iguana-17148	57	11	as	as	ADP
iguana-17148	57	12	starving	starve	VERB
iguana-17148	57	13	individuals	individual	NOUN
iguana-17148	57	14	or	or	CCONJ
iguana-17148	57	15	females	female	NOUN
iguana-17148	57	16	exhausted	exhaust	VERB
iguana-17148	57	17	by	by	ADP
iguana-17148	57	18	nesting	nest	VERB
iguana-17148	57	19	.	.	PUNCT
iguana-17148	58	1	they	they	PRON
iguana-17148	58	2	also	also	ADV
iguana-17148	58	3	can	can	AUX
iguana-17148	58	4	learn	learn	VERB
iguana-17148	58	5	to	to	PART
iguana-17148	58	6	capture	capture	VERB
iguana-17148	58	7	juveniles	juvenile	NOUN
iguana-17148	58	8	or	or	CCONJ
iguana-17148	58	9	hatchlings	hatchling	NOUN
iguana-17148	58	10	close	close	ADJ
iguana-17148	58	11	to	to	ADP
iguana-17148	58	12	the	the	DET
iguana-17148	58	13	shoreline	shoreline	NOUN
iguana-17148	58	14	(	(	PUNCT
iguana-17148	58	15	boersma	boersma	PROPN
iguana-17148	58	16	1983	1983	NUM
iguana-17148	58	17	;	;	PUNCT
iguana-17148	58	18	personal	personal	ADJ
iguana-17148	58	19	observation	observation	NOUN
iguana-17148	58	20	,	,	PUNCT
iguana-17148	58	21	santa	santa	PROPN
iguana-17148	58	22	fe	fe	PROPN
iguana-17148	58	23	island	island	PROPN
iguana-17148	58	24	)	)	PUNCT
iguana-17148	58	25	.	.	PUNCT
iguana-17148	59	1	natural	natural	ADJ
iguana-17148	59	2	population	population	NOUN
iguana-17148	59	3	regulation	regulation	NOUN
iguana-17148	59	4	via	via	ADP
iguana-17148	59	5	el	el	PROPN
iguana-17148	59	6	niño	niño	PROPN
iguana-17148	59	7	events	event	NOUN
iguana-17148	59	8	unpredictably	unpredictably	ADV
iguana-17148	59	9	recurring	recur	VERB
iguana-17148	59	10	el	el	PROPN
iguana-17148	59	11	niño	niño	ADJ
iguana-17148	59	12	events	event	NOUN
iguana-17148	59	13	can	can	AUX
iguana-17148	59	14	dramatically	dramatically	ADV
iguana-17148	59	15	reduce	reduce	VERB
iguana-17148	59	16	the	the	DET
iguana-17148	59	17	abundance	abundance	NOUN
iguana-17148	59	18	and	and	CCONJ
iguana-17148	59	19	diversity	diversity	NOUN
iguana-17148	59	20	of	of	ADP
iguana-17148	59	21	marine	marine	ADJ
iguana-17148	59	22	algae	algae	NOUN
iguana-17148	59	23	—	—	PUNCT
iguana-17148	59	24	marine	marine	ADJ
iguana-17148	59	25	iguanas	iguana	NOUN
iguana-17148	59	26	’	'	PUNCT
iguana-17148	59	27	only	only	ADV
iguana-17148	59	28	food	food	NOUN
iguana-17148	59	29	source	source	NOUN
iguana-17148	59	30	—	—	PUNCT
iguana-17148	59	31	and	and	CCONJ
iguana-17148	59	32	can	can	AUX
iguana-17148	59	33	cause	cause	VERB
iguana-17148	59	34	mass	mass	ADJ
iguana-17148	59	35	starvation	starvation	NOUN
iguana-17148	59	36	(	(	PUNCT
iguana-17148	59	37	laurie	laurie	PROPN
iguana-17148	59	38	and	and	CCONJ
iguana-17148	59	39	brown	brown	ADJ
iguana-17148	59	40	1990b	1990b	NUM
iguana-17148	59	41	)	)	PUNCT
iguana-17148	59	42	.	.	PUNCT
iguana-17148	60	1	during	during	ADP
iguana-17148	60	2	el	el	PROPN
iguana-17148	60	3	niño	niño	PROPN
iguana-17148	60	4	events	event	NOUN
iguana-17148	60	5	,	,	PUNCT
iguana-17148	60	6	the	the	DET
iguana-17148	60	7	cold	cold	ADJ
iguana-17148	60	8	,	,	PUNCT
iguana-17148	60	9	nutrient	nutrient	ADJ
iguana-17148	60	10	-	-	PUNCT
iguana-17148	60	11	rich	rich	ADJ
iguana-17148	60	12	upwelling	upwelling	NOUN
iguana-17148	60	13	ceases	cease	VERB
iguana-17148	60	14	and	and	CCONJ
iguana-17148	60	15	warm	warm	ADJ
iguana-17148	60	16	water	water	NOUN
iguana-17148	60	17	from	from	ADP
iguana-17148	60	18	the	the	DET
iguana-17148	60	19	gulf	gulf	PROPN
iguana-17148	60	20	of	of	ADP
iguana-17148	60	21	panama	panama	PROPN
iguana-17148	60	22	flows	flow	VERB
iguana-17148	60	23	toward	toward	ADP
iguana-17148	60	24	the	the	DET
iguana-17148	60	25	galápagos	galápago	NOUN
iguana-17148	60	26	.	.	PUNCT
iguana-17148	61	1	the	the	DET
iguana-17148	61	2	normal	normal	ADJ
iguana-17148	61	3	food	food	NOUN
iguana-17148	61	4	algae	algae	NOUN
iguana-17148	61	5	(	(	PUNCT
iguana-17148	61	6	red	red	ADJ
iguana-17148	61	7	gelidium	gelidium	NOUN
iguana-17148	61	8	and	and	CCONJ
iguana-17148	61	9	192	192	NUM
iguana-17148	61	10	iguana	iguana	NOUN
iguana-17148	61	11	•	•	NUM
iguana-17148	61	12	volume	volume	NOUN
iguana-17148	61	13	11	11	NUM
iguana-17148	61	14	,	,	PUNCT
iguana-17148	61	15	number	number	NOUN
iguana-17148	61	16	4	4	NUM
iguana-17148	61	17	•	•	NOUN
iguana-17148	61	18	december	december	PROPN
iguana-17148	61	19	2004	2004	NUM
iguana-17148	61	20	wikelski	wikelski	NOUN
iguana-17148	61	21	and	and	CCONJ
iguana-17148	61	22	nelson	nelson	PROPN
iguana-17148	61	23	marine	marine	PROPN
iguana-17148	61	24	iguanas	iguana	NOUN
iguana-17148	61	25	in	in	ADP
iguana-17148	61	26	the	the	DET
iguana-17148	61	27	intertidal	intertidal	ADJ
iguana-17148	61	28	zone	zone	NOUN
iguana-17148	61	29	on	on	ADP
iguana-17148	61	30	genovesa	genovesa	PROPN
iguana-17148	61	31	island	island	NOUN
iguana-17148	61	32	during	during	ADP
iguana-17148	61	33	the	the	DET
iguana-17148	61	34	low	low	ADJ
iguana-17148	61	35	tide	tide	NOUN
iguana-17148	61	36	,	,	PUNCT
iguana-17148	61	37	grazing	graze	VERB
iguana-17148	61	38	on	on	ADP
iguana-17148	61	39	green	green	ADJ
iguana-17148	61	40	algae	algae	NOUN
iguana-17148	61	41	(	(	PUNCT
iguana-17148	61	42	mostly	mostly	ADV
iguana-17148	61	43	ulva	ulva	ADJ
iguana-17148	61	44	sp	sp	PROPN
iguana-17148	61	45	.	.	PUNCT
iguana-17148	61	46	)	)	PUNCT
iguana-17148	61	47	.	.	PUNCT
iguana-17148	62	1	this	this	DET
iguana-17148	62	2	population	population	NOUN
iguana-17148	62	3	is	be	AUX
iguana-17148	62	4	the	the	DET
iguana-17148	62	5	smallest	small	ADJ
iguana-17148	62	6	in	in	ADP
iguana-17148	62	7	body	body	NOUN
iguana-17148	62	8	size	size	NOUN
iguana-17148	62	9	and	and	CCONJ
iguana-17148	62	10	males	male	NOUN
iguana-17148	62	11	with	with	ADP
iguana-17148	62	12	body	body	NOUN
iguana-17148	62	13	mass	mass	NOUN
iguana-17148	62	14	as	as	ADV
iguana-17148	62	15	little	little	ADJ
iguana-17148	62	16	as	as	ADP
iguana-17148	62	17	500	500	NUM
iguana-17148	62	18	g	g	NOUN
iguana-17148	62	19	are	be	AUX
iguana-17148	62	20	seen	see	VERB
iguana-17148	62	21	diving	diving	NOUN
iguana-17148	62	22	.	.	PUNCT
iguana-17148	63	1	two	two	NUM
iguana-17148	63	2	sneaker	sneaker	NOUN
iguana-17148	63	3	male	male	ADJ
iguana-17148	63	4	marine	marine	ADJ
iguana-17148	63	5	iguanas	iguana	NOUN
iguana-17148	63	6	on	on	ADP
iguana-17148	63	7	genovesa	genovesa	PROPN
iguana-17148	63	8	island	island	PROPN
iguana-17148	63	9	attempt	attempt	NOUN
iguana-17148	63	10	to	to	PART
iguana-17148	63	11	forcefully	forcefully	ADV
iguana-17148	63	12	copulate	copulate	VERB
iguana-17148	63	13	with	with	ADP
iguana-17148	63	14	a	a	DET
iguana-17148	63	15	female	female	ADJ
iguana-17148	63	16	outside	outside	NOUN
iguana-17148	63	17	of	of	ADP
iguana-17148	63	18	a	a	DET
iguana-17148	63	19	territory	territory	NOUN
iguana-17148	63	20	(	(	PUNCT
iguana-17148	63	21	on	on	ADP
iguana-17148	63	22	sand	sand	NOUN
iguana-17148	63	23	)	)	PUNCT
iguana-17148	63	24	.	.	PUNCT
iguana-17148	64	1	the	the	DET
iguana-17148	64	2	territorial	territorial	ADJ
iguana-17148	64	3	male	male	NOUN
iguana-17148	64	4	(	(	PUNCT
iguana-17148	64	5	right	right	ADJ
iguana-17148	64	6	)	)	PUNCT
iguana-17148	64	7	left	leave	VERB
iguana-17148	64	8	his	his	PRON
iguana-17148	64	9	territory	territory	NOUN
iguana-17148	64	10	and	and	CCONJ
iguana-17148	64	11	interrupted	interrupt	VERB
iguana-17148	64	12	the	the	DET
iguana-17148	64	13	copulation	copulation	NOUN
iguana-17148	64	14	attempt	attempt	NOUN
iguana-17148	64	15	.	.	PUNCT
iguana-17148	65	1	a	a	DET
iguana-17148	65	2	satellite	satellite	NOUN
iguana-17148	65	3	male	male	NOUN
iguana-17148	65	4	marine	marine	ADJ
iguana-17148	65	5	iguana	iguana	NOUN
iguana-17148	65	6	forcefully	forcefully	ADV
iguana-17148	65	7	approaches	approach	VERB
iguana-17148	65	8	a	a	DET
iguana-17148	65	9	female	female	NOUN
iguana-17148	65	10	to	to	PART
iguana-17148	65	11	attempt	attempt	VERB
iguana-17148	65	12	copulation	copulation	NOUN
iguana-17148	65	13	.	.	PUNCT
iguana-17148	66	1	iguana	iguana	NOUN
iguana-17148	66	2	•	•	NUM
iguana-17148	66	3	volume	volume	NOUN
iguana-17148	66	4	11	11	NUM
iguana-17148	66	5	,	,	PUNCT
iguana-17148	66	6	number	number	NOUN
iguana-17148	66	7	4	4	NUM
iguana-17148	66	8	•	•	NOUN
iguana-17148	66	9	december	december	PROPN
iguana-17148	66	10	2004	2004	NUM
iguana-17148	66	11	193galápagos	193galápagos	PROPN
iguana-17148	66	12	marine	marine	ADJ
iguana-17148	66	13	iguanas	iguana	NOUN
iguana-17148	66	14	centroseras	centroseras	PROPN
iguana-17148	66	15	species	species	PROPN
iguana-17148	66	16	or	or	CCONJ
iguana-17148	66	17	green	green	ADJ
iguana-17148	66	18	ulva	ulva	PROPN
iguana-17148	66	19	species	species	PROPN
iguana-17148	66	20	)	)	PUNCT
iguana-17148	66	21	disappear	disappear	VERB
iguana-17148	66	22	when	when	SCONJ
iguana-17148	66	23	water	water	NOUN
iguana-17148	66	24	temperatures	temperature	NOUN
iguana-17148	66	25	become	become	VERB
iguana-17148	66	26	too	too	ADV
iguana-17148	66	27	high	high	ADJ
iguana-17148	66	28	and	and	CCONJ
iguana-17148	66	29	are	be	AUX
iguana-17148	66	30	replaced	replace	VERB
iguana-17148	66	31	by	by	ADP
iguana-17148	66	32	brown	brown	ADJ
iguana-17148	66	33	algae	algae	NOUN
iguana-17148	66	34	.	.	PUNCT
iguana-17148	67	1	however	however	ADV
iguana-17148	67	2	,	,	PUNCT
iguana-17148	67	3	brown	brown	ADJ
iguana-17148	67	4	algae	algae	NOUN
iguana-17148	67	5	are	be	AUX
iguana-17148	67	6	not	not	PART
iguana-17148	67	7	as	as	ADV
iguana-17148	67	8	easily	easily	ADV
iguana-17148	67	9	digested	digest	VERB
iguana-17148	67	10	by	by	ADP
iguana-17148	67	11	the	the	DET
iguana-17148	67	12	marine	marine	ADJ
iguana-17148	67	13	iguanas	iguana	NOUN
iguana-17148	67	14	’	'	PUNCT
iguana-17148	67	15	hindgut	hindgut	NOUN
iguana-17148	67	16	bacteria	bacteria	NOUN
iguana-17148	67	17	,	,	PUNCT
iguana-17148	67	18	and	and	CCONJ
iguana-17148	67	19	they	they	PRON
iguana-17148	67	20	may	may	AUX
iguana-17148	67	21	also	also	ADV
iguana-17148	67	22	be	be	AUX
iguana-17148	67	23	toxic	toxic	ADJ
iguana-17148	67	24	.	.	PUNCT
iguana-17148	68	1	the	the	DET
iguana-17148	68	2	result	result	NOUN
iguana-17148	68	3	is	be	AUX
iguana-17148	68	4	widespread	widespread	ADJ
iguana-17148	68	5	starvation	starvation	NOUN
iguana-17148	68	6	of	of	ADP
iguana-17148	68	7	marine	marine	ADJ
iguana-17148	68	8	iguanas	iguana	NOUN
iguana-17148	68	9	throughout	throughout	ADP
iguana-17148	68	10	the	the	DET
iguana-17148	68	11	archipelago	archipelago	PROPN
iguana-17148	68	12	.	.	PUNCT
iguana-17148	69	1	individual	individual	ADJ
iguana-17148	69	2	animals	animal	NOUN
iguana-17148	69	3	are	be	AUX
iguana-17148	69	4	affected	affect	VERB
iguana-17148	69	5	differently	differently	ADV
iguana-17148	69	6	—	—	PUNCT
iguana-17148	69	7	the	the	DET
iguana-17148	69	8	largest	large	ADJ
iguana-17148	69	9	animals	animal	NOUN
iguana-17148	69	10	starve	starve	VERB
iguana-17148	69	11	first	first	ADV
iguana-17148	69	12	(	(	PUNCT
iguana-17148	69	13	wikelski	wikelski	NOUN
iguana-17148	69	14	and	and	CCONJ
iguana-17148	69	15	trillmich	trillmich	NOUN
iguana-17148	69	16	1997	1997	NUM
iguana-17148	69	17	,	,	PUNCT
iguana-17148	69	18	wikelski	wikelski	PROPN
iguana-17148	69	19	et	et	PROPN
iguana-17148	69	20	al	al	PROPN
iguana-17148	69	21	.	.	PROPN
iguana-17148	69	22	1997	1997	NUM
iguana-17148	69	23	)	)	PUNCT
iguana-17148	69	24	,	,	PUNCT
iguana-17148	69	25	presumably	presumably	ADV
iguana-17148	69	26	because	because	SCONJ
iguana-17148	69	27	they	they	PRON
iguana-17148	69	28	have	have	VERB
iguana-17148	69	29	the	the	DET
iguana-17148	69	30	highest	high	ADJ
iguana-17148	69	31	absolute	absolute	ADJ
iguana-17148	69	32	calorie	calorie	NOUN
iguana-17148	69	33	requirements	requirement	NOUN
iguana-17148	69	34	.	.	PUNCT
iguana-17148	70	1	therefore	therefore	ADV
iguana-17148	70	2	,	,	PUNCT
iguana-17148	70	3	natural	natural	ADJ
iguana-17148	70	4	selection	selection	NOUN
iguana-17148	70	5	favors	favor	VERB
iguana-17148	70	6	smaller	small	ADJ
iguana-17148	70	7	animals	animal	NOUN
iguana-17148	70	8	during	during	ADP
iguana-17148	70	9	food	food	NOUN
iguana-17148	70	10	shortages	shortage	NOUN
iguana-17148	70	11	.	.	PUNCT
iguana-17148	71	1	interestingly	interestingly	ADV
iguana-17148	71	2	,	,	PUNCT
iguana-17148	71	3	marine	marine	ADJ
iguana-17148	71	4	iguanas	iguana	NOUN
iguana-17148	71	5	can	can	AUX
iguana-17148	71	6	shrink	shrink	VERB
iguana-17148	71	7	their	their	PRON
iguana-17148	71	8	body	body	NOUN
iguana-17148	71	9	size	size	NOUN
iguana-17148	71	10	during	during	ADP
iguana-17148	71	11	el	el	PROPN
iguana-17148	71	12	niño	niño	PROPN
iguana-17148	71	13	events	event	NOUN
iguana-17148	71	14	and	and	CCONJ
iguana-17148	71	15	survive	survive	VERB
iguana-17148	71	16	such	such	ADJ
iguana-17148	71	17	conditions	condition	NOUN
iguana-17148	71	18	better	well	ADV
iguana-17148	71	19	(	(	PUNCT
iguana-17148	71	20	wikelski	wikelski	NOUN
iguana-17148	71	21	and	and	CCONJ
iguana-17148	71	22	thom	thom	PROPN
iguana-17148	71	23	2000	2000	NUM
iguana-17148	71	24	)	)	PUNCT
iguana-17148	71	25	.	.	PUNCT
iguana-17148	72	1	how	how	SCONJ
iguana-17148	72	2	such	such	ADJ
iguana-17148	72	3	shrinkage	shrinkage	NOUN
iguana-17148	72	4	is	be	AUX
iguana-17148	72	5	achieved	achieve	VERB
iguana-17148	72	6	physiologically	physiologically	ADV
iguana-17148	72	7	or	or	CCONJ
iguana-17148	72	8	whether	whether	SCONJ
iguana-17148	72	9	and	and	CCONJ
iguana-17148	72	10	to	to	ADP
iguana-17148	72	11	what	what	DET
iguana-17148	72	12	degree	degree	NOUN
iguana-17148	72	13	bone	bone	NOUN
iguana-17148	72	14	loss	loss	NOUN
iguana-17148	72	15	is	be	AUX
iguana-17148	72	16	involved	involve	VERB
iguana-17148	72	17	is	be	AUX
iguana-17148	72	18	not	not	PART
iguana-17148	72	19	clear	clear	ADJ
iguana-17148	72	20	.	.	PUNCT
iguana-17148	73	1	marine	marine	ADJ
iguana-17148	73	2	iguana	iguana	NOUN
iguana-17148	73	3	populations	population	NOUN
iguana-17148	73	4	can	can	AUX
iguana-17148	73	5	crash	crash	VERB
iguana-17148	73	6	dramatically	dramatically	ADV
iguana-17148	73	7	.	.	PUNCT
iguana-17148	74	1	during	during	ADP
iguana-17148	74	2	the	the	DET
iguana-17148	74	3	el	el	PROPN
iguana-17148	74	4	niño	niño	NOUN
iguana-17148	74	5	of	of	ADP
iguana-17148	74	6	1997–1998	1997–1998	NUM
iguana-17148	74	7	,	,	PUNCT
iguana-17148	74	8	about	about	ADV
iguana-17148	74	9	90	90	NUM
iguana-17148	74	10	%	%	NOUN
iguana-17148	74	11	of	of	ADP
iguana-17148	74	12	all	all	DET
iguana-17148	74	13	marine	marine	ADJ
iguana-17148	74	14	iguanas	iguana	NOUN
iguana-17148	74	15	on	on	ADP
iguana-17148	74	16	seymour	seymour	PROPN
iguana-17148	74	17	norte	norte	PROPN
iguana-17148	74	18	island	island	PROPN
iguana-17148	74	19	disappeared	disappear	VERB
iguana-17148	74	20	,	,	PUNCT
iguana-17148	74	21	thus	thus	ADV
iguana-17148	74	22	reducing	reduce	VERB
iguana-17148	74	23	population	population	NOUN
iguana-17148	74	24	size	size	NOUN
iguana-17148	74	25	on	on	ADP
iguana-17148	74	26	this	this	DET
iguana-17148	74	27	island	island	NOUN
iguana-17148	74	28	to	to	ADP
iguana-17148	74	29	less	less	ADJ
iguana-17148	74	30	than	than	ADP
iguana-17148	74	31	150	150	NUM
iguana-17148	74	32	individuals	individual	NOUN
iguana-17148	74	33	(	(	PUNCT
iguana-17148	74	34	wikelski	wikelski	NOUN
iguana-17148	74	35	and	and	CCONJ
iguana-17148	74	36	wrege	wrege	NOUN
iguana-17148	74	37	2000	2000	NUM
iguana-17148	74	38	,	,	PUNCT
iguana-17148	74	39	romero	romero	PROPN
iguana-17148	74	40	and	and	CCONJ
iguana-17148	74	41	wikelski	wikelski	NOUN
iguana-17148	74	42	2001	2001	NUM
iguana-17148	74	43	)	)	PUNCT
iguana-17148	74	44	.	.	PUNCT
iguana-17148	75	1	a	a	DET
iguana-17148	75	2	similar	similar	ADJ
iguana-17148	75	3	situation	situation	NOUN
iguana-17148	75	4	occurred	occur	VERB
iguana-17148	75	5	on	on	ADP
iguana-17148	75	6	genovesa	genovesa	PROPN
iguana-17148	75	7	island	island	NOUN
iguana-17148	75	8	in	in	ADP
iguana-17148	75	9	1991–1994	1991–1994	NUM
iguana-17148	75	10	,	,	PUNCT
iguana-17148	75	11	reducing	reduce	VERB
iguana-17148	75	12	the	the	DET
iguana-17148	75	13	total	total	ADJ
iguana-17148	75	14	population	population	NOUN
iguana-17148	75	15	size	size	NOUN
iguana-17148	75	16	from	from	ADP
iguana-17148	75	17	about	about	ADV
iguana-17148	75	18	15,000	15,000	NUM
iguana-17148	75	19	to	to	PART
iguana-17148	75	20	approximately	approximately	ADV
iguana-17148	75	21	900	900	NUM
iguana-17148	75	22	individuals	individual	NOUN
iguana-17148	75	23	.	.	PUNCT
iguana-17148	76	1	however	however	ADV
iguana-17148	76	2	,	,	PUNCT
iguana-17148	76	3	marine	marine	ADJ
iguana-17148	76	4	iguanas	iguana	NOUN
iguana-17148	76	5	have	have	AUX
iguana-17148	76	6	survived	survive	VERB
iguana-17148	76	7	such	such	ADJ
iguana-17148	76	8	dramatic	dramatic	ADJ
iguana-17148	76	9	natural	natural	ADJ
iguana-17148	76	10	selection	selection	NOUN
iguana-17148	76	11	events	event	NOUN
iguana-17148	76	12	throughout	throughout	ADP
iguana-17148	76	13	their	their	PRON
iguana-17148	76	14	evolutionary	evolutionary	ADJ
iguana-17148	76	15	history	history	NOUN
iguana-17148	76	16	and	and	CCONJ
iguana-17148	76	17	apparently	apparently	ADV
iguana-17148	76	18	adjust	adjust	VERB
iguana-17148	76	19	to	to	ADP
iguana-17148	76	20	such	such	ADJ
iguana-17148	76	21	situations	situation	NOUN
iguana-17148	76	22	.	.	PUNCT
iguana-17148	77	1	as	as	ADV
iguana-17148	77	2	soon	soon	ADV
iguana-17148	77	3	as	as	ADP
iguana-17148	77	4	nutritious	nutritious	ADJ
iguana-17148	77	5	red	red	ADJ
iguana-17148	77	6	and	and	CCONJ
iguana-17148	77	7	green	green	ADJ
iguana-17148	77	8	algae	algae	NOUN
iguana-17148	77	9	reappear	reappear	NOUN
iguana-17148	77	10	after	after	SCONJ
iguana-17148	77	11	an	an	DET
iguana-17148	77	12	el	el	PROPN
iguana-17148	77	13	niño	niño	PROPN
iguana-17148	77	14	ceases	cease	VERB
iguana-17148	77	15	,	,	PUNCT
iguana-17148	77	16	individuals	individual	NOUN
iguana-17148	77	17	face	face	VERB
iguana-17148	77	18	plentiful	plentiful	ADJ
iguana-17148	77	19	intertidal	intertidal	ADJ
iguana-17148	77	20	and	and	CCONJ
iguana-17148	77	21	subtidal	subtidal	ADJ
iguana-17148	77	22	foraging	foraging	NOUN
iguana-17148	77	23	grounds	ground	NOUN
iguana-17148	77	24	.	.	PUNCT
iguana-17148	78	1	iguanas	iguana	NOUN
iguana-17148	78	2	quickly	quickly	ADV
iguana-17148	78	3	replenish	replenish	VERB
iguana-17148	78	4	their	their	PRON
iguana-17148	78	5	fat	fat	ADJ
iguana-17148	78	6	reserves	reserve	NOUN
iguana-17148	78	7	and	and	CCONJ
iguana-17148	78	8	return	return	VERB
iguana-17148	78	9	to	to	ADP
iguana-17148	78	10	good	good	ADJ
iguana-17148	78	11	body	body	NOUN
iguana-17148	78	12	condition	condition	NOUN
iguana-17148	78	13	.	.	PUNCT
iguana-17148	79	1	they	they	PRON
iguana-17148	79	2	reproduce	reproduce	VERB
iguana-17148	79	3	more	more	ADV
iguana-17148	79	4	frequently	frequently	ADV
iguana-17148	79	5	(	(	PUNCT
iguana-17148	79	6	every	every	DET
iguana-17148	79	7	year	year	NOUN
iguana-17148	79	8	)	)	PUNCT
iguana-17148	79	9	,	,	PUNCT
iguana-17148	79	10	at	at	ADP
iguana-17148	79	11	a	a	DET
iguana-17148	79	12	younger	young	ADJ
iguana-17148	79	13	age	age	NOUN
iguana-17148	79	14	(	(	PUNCT
iguana-17148	79	15	mostly	mostly	ADV
iguana-17148	79	16	females	female	NOUN
iguana-17148	79	17	)	)	PUNCT
iguana-17148	79	18	,	,	PUNCT
iguana-17148	79	19	and	and	CCONJ
iguana-17148	79	20	lay	lay	VERB
iguana-17148	79	21	larger	large	ADJ
iguana-17148	79	22	clutches	clutch	NOUN
iguana-17148	79	23	(	(	PUNCT
iguana-17148	79	24	e.g.	e.g.	ADV
iguana-17148	79	25	,	,	PUNCT
iguana-17148	79	26	three	three	NUM
iguana-17148	79	27	instead	instead	ADV
iguana-17148	79	28	of	of	ADP
iguana-17148	79	29	two	two	NUM
iguana-17148	79	30	eggs	egg	NOUN
iguana-17148	79	31	)	)	PUNCT
iguana-17148	79	32	.	.	PUNCT
iguana-17148	80	1	the	the	DET
iguana-17148	80	2	“	"	PUNCT
iguana-17148	80	3	rules	rule	NOUN
iguana-17148	80	4	”	"	PUNCT
iguana-17148	80	5	by	by	ADP
iguana-17148	80	6	which	which	PRON
iguana-17148	80	7	marine	marine	PROPN
iguana-17148	80	8	iguanas	iguana	NOUN
iguana-17148	80	9	determine	determine	VERB
iguana-17148	80	10	whether	whether	SCONJ
iguana-17148	80	11	to	to	PART
iguana-17148	80	12	breed	breed	VERB
iguana-17148	80	13	and	and	CCONJ
iguana-17148	80	14	how	how	SCONJ
iguana-17148	80	15	many	many	ADJ
iguana-17148	80	16	eggs	egg	NOUN
iguana-17148	80	17	to	to	PART
iguana-17148	80	18	lay	lay	VERB
iguana-17148	80	19	are	be	AUX
iguana-17148	80	20	still	still	ADV
iguana-17148	80	21	unclear	unclear	ADJ
iguana-17148	80	22	.	.	PUNCT
iguana-17148	81	1	these	these	DET
iguana-17148	81	2	adaptations	adaptation	NOUN
iguana-17148	81	3	allow	allow	VERB
iguana-17148	81	4	marine	marine	ADJ
iguana-17148	81	5	iguanas	iguana	NOUN
iguana-17148	81	6	to	to	PART
iguana-17148	81	7	increase	increase	VERB
iguana-17148	81	8	their	their	PRON
iguana-17148	81	9	numbers	number	NOUN
iguana-17148	81	10	after	after	SCONJ
iguana-17148	81	11	dramatic	dramatic	ADJ
iguana-17148	81	12	population	population	NOUN
iguana-17148	81	13	declines	decline	VERB
iguana-17148	81	14	,	,	PUNCT
iguana-17148	81	15	such	such	ADJ
iguana-17148	81	16	that	that	SCONJ
iguana-17148	81	17	mortality	mortality	NOUN
iguana-17148	81	18	rates	rate	NOUN
iguana-17148	81	19	of	of	ADP
iguana-17148	81	20	30–50	30–50	NUM
iguana-17148	81	21	%	%	NOUN
iguana-17148	81	22	after	after	ADP
iguana-17148	81	23	an	an	DET
iguana-17148	81	24	el	el	PROPN
iguana-17148	81	25	niño	niño	PROPN
iguana-17148	81	26	event	event	NOUN
iguana-17148	81	27	can	can	AUX
iguana-17148	81	28	be	be	AUX
iguana-17148	81	29	compensated	compensate	VERB
iguana-17148	81	30	within	within	ADP
iguana-17148	81	31	four	four	NUM
iguana-17148	81	32	years	year	NOUN
iguana-17148	81	33	.	.	PUNCT
iguana-17148	82	1	even	even	ADV
iguana-17148	82	2	after	after	ADP
iguana-17148	82	3	enormously	enormously	ADV
iguana-17148	82	4	strong	strong	ADJ
iguana-17148	82	5	population	population	NOUN
iguana-17148	82	6	declines	decline	NOUN
iguana-17148	82	7	(	(	PUNCT
iguana-17148	82	8	90	90	NUM
iguana-17148	82	9	%	%	NOUN
iguana-17148	82	10	)	)	PUNCT
iguana-17148	82	11	,	,	PUNCT
iguana-17148	82	12	marine	marine	PROPN
iguana-17148	82	13	iguanas	iguana	NOUN
iguana-17148	82	14	congregate	congregate	VERB
iguana-17148	82	15	along	along	ADP
iguana-17148	82	16	the	the	DET
iguana-17148	82	17	shoreline	shoreline	NOUN
iguana-17148	82	18	in	in	ADP
iguana-17148	82	19	small	small	ADJ
iguana-17148	82	20	groups	group	NOUN
iguana-17148	82	21	to	to	PART
iguana-17148	82	22	reproduce	reproduce	VERB
iguana-17148	82	23	(	(	PUNCT
iguana-17148	82	24	wikelski	wikelski	NOUN
iguana-17148	82	25	et	et	PROPN
iguana-17148	82	26	al	al	PROPN
iguana-17148	82	27	.	.	PROPN
iguana-17148	82	28	1996	1996	NUM
iguana-17148	82	29	)	)	PUNCT
iguana-17148	82	30	.	.	PUNCT
iguana-17148	83	1	this	this	DET
iguana-17148	83	2	gregariousness	gregariousness	NOUN
iguana-17148	83	3	helps	help	VERB
iguana-17148	83	4	them	they	PRON
iguana-17148	83	5	to	to	PART
iguana-17148	83	6	find	find	VERB
iguana-17148	83	7	each	each	DET
iguana-17148	83	8	other	other	ADJ
iguana-17148	83	9	after	after	SCONJ
iguana-17148	83	10	dramatic	dramatic	ADJ
iguana-17148	83	11	population	population	NOUN
iguana-17148	83	12	declines	decline	VERB
iguana-17148	83	13	that	that	PRON
iguana-17148	83	14	could	could	AUX
iguana-17148	83	15	otherwise	otherwise	ADV
iguana-17148	83	16	cause	cause	VERB
iguana-17148	83	17	individuals	individual	NOUN
iguana-17148	83	18	to	to	PART
iguana-17148	83	19	scatter	scatter	VERB
iguana-17148	83	20	along	along	ADP
iguana-17148	83	21	the	the	DET
iguana-17148	83	22	long	long	ADJ
iguana-17148	83	23	lava	lava	NOUN
iguana-17148	83	24	shores	shore	NOUN
iguana-17148	83	25	of	of	ADP
iguana-17148	83	26	the	the	DET
iguana-17148	83	27	islands	island	NOUN
iguana-17148	83	28	.	.	PUNCT
iguana-17148	84	1	island	island	NOUN
iguana-17148	84	2	populations	population	NOUN
iguana-17148	84	3	and	and	CCONJ
iguana-17148	84	4	threats	threat	NOUN
iguana-17148	84	5	early	early	ADJ
iguana-17148	84	6	naturalists	naturalist	NOUN
iguana-17148	84	7	discovered	discover	VERB
iguana-17148	84	8	that	that	SCONJ
iguana-17148	84	9	not	not	PART
iguana-17148	84	10	all	all	DET
iguana-17148	84	11	marine	marine	ADJ
iguana-17148	84	12	iguana	iguana	NOUN
iguana-17148	84	13	populations	population	NOUN
iguana-17148	84	14	are	be	AUX
iguana-17148	84	15	alike	alike	ADV
iguana-17148	84	16	(	(	PUNCT
iguana-17148	84	17	fig	fig	NOUN
iguana-17148	84	18	.	.	PUNCT
iguana-17148	85	1	1	1	NUM
iguana-17148	85	2	)	)	PUNCT
iguana-17148	85	3	.	.	PUNCT
iguana-17148	86	1	for	for	ADP
iguana-17148	86	2	example	example	NOUN
iguana-17148	86	3	,	,	PUNCT
iguana-17148	86	4	iguanas	iguana	NOUN
iguana-17148	86	5	on	on	ADP
iguana-17148	86	6	genovesa	genovesa	PROPN
iguana-17148	86	7	only	only	ADV
iguana-17148	86	8	grow	grow	VERB
iguana-17148	86	9	up	up	ADP
iguana-17148	86	10	to	to	ADP
iguana-17148	86	11	a	a	DET
iguana-17148	86	12	maximum	maximum	NOUN
iguana-17148	86	13	of	of	ADP
iguana-17148	86	14	900	900	NUM
iguana-17148	86	15	g	g	NOUN
iguana-17148	86	16	(	(	PUNCT
iguana-17148	86	17	subspecies	subspecie	NOUN
iguana-17148	86	18	:	:	PUNCT
iguana-17148	86	19	a.	a.	PROPN
iguana-17148	86	20	c.	c.	PROPN
iguana-17148	86	21	nanus	nanus	PROPN
iguana-17148	86	22	,	,	PUNCT
iguana-17148	86	23	“	"	PUNCT
iguana-17148	86	24	the	the	DET
iguana-17148	86	25	small	small	ADJ
iguana-17148	86	26	ones	one	NOUN
iguana-17148	86	27	”	"	PUNCT
iguana-17148	86	28	)	)	PUNCT
iguana-17148	86	29	.	.	PUNCT
iguana-17148	87	1	animals	animal	NOUN
iguana-17148	87	2	on	on	ADP
iguana-17148	87	3	española	española	PROPN
iguana-17148	87	4	are	be	AUX
iguana-17148	87	5	the	the	DET
iguana-17148	87	6	most	most	ADV
iguana-17148	87	7	brilliant	brilliant	ADJ
iguana-17148	87	8	,	,	PUNCT
iguana-17148	87	9	at	at	ADP
iguana-17148	87	10	least	least	ADJ
iguana-17148	87	11	during	during	ADP
iguana-17148	87	12	the	the	DET
iguana-17148	87	13	mating	mating	NOUN
iguana-17148	87	14	season	season	NOUN
iguana-17148	87	15	,	,	PUNCT
iguana-17148	87	16	when	when	SCONJ
iguana-17148	87	17	they	they	PRON
iguana-17148	87	18	display	display	VERB
iguana-17148	87	19	a	a	DET
iguana-17148	87	20	bright	bright	ADJ
iguana-17148	87	21	red	red	ADJ
iguana-17148	87	22	and	and	CCONJ
iguana-17148	87	23	green	green	ADJ
iguana-17148	87	24	coloration	coloration	NOUN
iguana-17148	87	25	(	(	PUNCT
iguana-17148	87	26	a.	a.	NOUN
iguana-17148	87	27	c.	c.	PROPN
iguana-17148	87	28	venustissimus	venustissimus	PROPN
iguana-17148	87	29	)	)	PUNCT
iguana-17148	87	30	.	.	PUNCT
iguana-17148	88	1	fernandina	fernandina	PROPN
iguana-17148	88	2	iguanas	iguanas	PROPN
iguana-17148	88	3	have	have	VERB
iguana-17148	88	4	especially	especially	ADV
iguana-17148	88	5	elaborate	elaborate	ADJ
iguana-17148	88	6	spines	spine	NOUN
iguana-17148	88	7	(	(	PUNCT
iguana-17148	88	8	a.	a.	NOUN
iguana-17148	88	9	c.	c.	PROPN
iguana-17148	88	10	cristatus	cristatus	PROPN
iguana-17148	88	11	subspecies	subspecie	NOUN
iguana-17148	88	12	)	)	PUNCT
iguana-17148	88	13	.	.	PUNCT
iguana-17148	89	1	although	although	SCONJ
iguana-17148	89	2	these	these	DET
iguana-17148	89	3	populations	population	NOUN
iguana-17148	89	4	differ	differ	VERB
iguana-17148	89	5	very	very	ADV
iguana-17148	89	6	obviously	obviously	ADV
iguana-17148	89	7	on	on	ADP
iguana-17148	89	8	the	the	DET
iguana-17148	89	9	phenotypic	phenotypic	ADJ
iguana-17148	89	10	level	level	NOUN
iguana-17148	89	11	,	,	PUNCT
iguana-17148	89	12	genetic	genetic	ADJ
iguana-17148	89	13	distable	distable	ADJ
iguana-17148	89	14	1	1	NUM
iguana-17148	89	15	.	.	PUNCT
iguana-17148	89	16	rough	rough	ADJ
iguana-17148	89	17	estimate	estimate	NOUN
iguana-17148	89	18	of	of	ADP
iguana-17148	89	19	marine	marine	ADJ
iguana-17148	89	20	iguana	iguana	NOUN
iguana-17148	89	21	population	population	NOUN
iguana-17148	89	22	sizes	size	NOUN
iguana-17148	89	23	on	on	ADP
iguana-17148	89	24	the	the	DET
iguana-17148	89	25	galápagos	galápagos	PROPN
iguana-17148	89	26	archipelago	archipelago	PROPN
iguana-17148	89	27	.	.	PUNCT
iguana-17148	90	1	data	datum	NOUN
iguana-17148	90	2	are	be	AUX
iguana-17148	90	3	based	base	VERB
iguana-17148	90	4	on	on	ADP
iguana-17148	90	5	our	our	PRON
iguana-17148	90	6	own	own	ADJ
iguana-17148	90	7	surveys	survey	NOUN
iguana-17148	90	8	,	,	PUNCT
iguana-17148	90	9	accounts	account	NOUN
iguana-17148	90	10	by	by	ADP
iguana-17148	90	11	galápagos	galápago	NOUN
iguana-17148	90	12	guides	guide	NOUN
iguana-17148	90	13	,	,	PUNCT
iguana-17148	90	14	and	and	CCONJ
iguana-17148	90	15	data	datum	NOUN
iguana-17148	90	16	by	by	ADP
iguana-17148	90	17	andrew	andrew	PROPN
iguana-17148	90	18	laurie	laurie	PROPN
iguana-17148	90	19	(	(	PUNCT
iguana-17148	90	20	unpublished	unpublished	ADJ
iguana-17148	90	21	report	report	NOUN
iguana-17148	90	22	to	to	ADP
iguana-17148	90	23	the	the	DET
iguana-17148	90	24	darwin	darwin	PROPN
iguana-17148	90	25	foundation	foundation	PROPN
iguana-17148	90	26	,	,	PUNCT
iguana-17148	90	27	1981	1981	NUM
iguana-17148	90	28	)	)	PUNCT
iguana-17148	90	29	.	.	PUNCT
iguana-17148	91	1	minimum	minimum	ADJ
iguana-17148	91	2	numbers	number	NOUN
iguana-17148	91	3	indicate	indicate	VERB
iguana-17148	91	4	estimates	estimate	NOUN
iguana-17148	91	5	for	for	ADP
iguana-17148	91	6	total	total	ADJ
iguana-17148	91	7	island	island	NOUN
iguana-17148	91	8	numbers	number	NOUN
iguana-17148	91	9	after	after	ADP
iguana-17148	91	10	a	a	DET
iguana-17148	91	11	strong	strong	ADJ
iguana-17148	91	12	el	el	PROPN
iguana-17148	91	13	niño	niño	PROPN
iguana-17148	91	14	famine	famine	NOUN
iguana-17148	91	15	.	.	PUNCT
iguana-17148	92	1	maximum	maximum	ADJ
iguana-17148	92	2	numbers	number	NOUN
iguana-17148	92	3	indicate	indicate	VERB
iguana-17148	92	4	total	total	ADJ
iguana-17148	92	5	numbers	number	NOUN
iguana-17148	92	6	after	after	ADP
iguana-17148	92	7	several	several	ADJ
iguana-17148	92	8	years	year	NOUN
iguana-17148	92	9	of	of	ADP
iguana-17148	92	10	la	la	X
iguana-17148	92	11	niña	niña	NOUN
iguana-17148	92	12	(	(	PUNCT
iguana-17148	92	13	cold	cold	ADJ
iguana-17148	92	14	,	,	PUNCT
iguana-17148	92	15	nutrient	nutrient	NOUN
iguana-17148	92	16	-	-	PUNCT
iguana-17148	92	17	rich	rich	ADJ
iguana-17148	92	18	)	)	PUNCT
iguana-17148	92	19	conditions	condition	NOUN
iguana-17148	92	20	.	.	PUNCT
iguana-17148	93	1	maximum	maximum	ADJ
iguana-17148	93	2	density	density	NOUN
iguana-17148	93	3	estimates	estimate	NOUN
iguana-17148	93	4	relate	relate	VERB
iguana-17148	93	5	iguana	iguana	NOUN
iguana-17148	93	6	numbers	number	NOUN
iguana-17148	93	7	to	to	ADP
iguana-17148	93	8	the	the	DET
iguana-17148	93	9	total	total	ADJ
iguana-17148	93	10	size	size	NOUN
iguana-17148	93	11	of	of	ADP
iguana-17148	93	12	the	the	DET
iguana-17148	93	13	island	island	NOUN
iguana-17148	93	14	.	.	PUNCT
iguana-17148	94	1	please	please	INTJ
iguana-17148	94	2	note	note	VERB
iguana-17148	94	3	that	that	SCONJ
iguana-17148	94	4	these	these	PRON
iguana-17148	94	5	are	be	AUX
iguana-17148	94	6	only	only	ADV
iguana-17148	94	7	very	very	ADV
iguana-17148	94	8	rough	rough	ADJ
iguana-17148	94	9	estimates	estimate	NOUN
iguana-17148	94	10	.	.	PUNCT
iguana-17148	95	1	island	island	NOUN
iguana-17148	95	2	subspecies	subspecie	NOUN
iguana-17148	95	3	minimum	minimum	ADJ
iguana-17148	95	4	maximum	maximum	ADJ
iguana-17148	95	5	maximum	maximum	ADJ
iguana-17148	95	6	threats	threat	NOUN
iguana-17148	95	7	number	number	NOUN
iguana-17148	95	8	number	number	NOUN
iguana-17148	95	9	density	density	NOUN
iguana-17148	95	10	(	(	PUNCT
iguana-17148	95	11	n	n	CCONJ
iguana-17148	95	12	/	/	SYM
iguana-17148	95	13	km2	km2	PROPN
iguana-17148	95	14	)	)	PUNCT
iguana-17148	95	15	fernandina	fernandina	PROPN
iguana-17148	95	16	cristatus	cristatus	PROPN
iguana-17148	95	17	15,000	15,000	NUM
iguana-17148	95	18	120,000	120,000	NUM
iguana-17148	95	19	187	187	NUM
iguana-17148	95	20	oil	oil	NOUN
iguana-17148	95	21	spill	spill	NOUN
iguana-17148	95	22	isabela	isabela	NOUN
iguana-17148	95	23	albemarlensis	albemarlensis	NOUN
iguana-17148	95	24	5,000	5,000	NUM
iguana-17148	95	25	40,000	40,000	NUM
iguana-17148	95	26	9	9	NUM
iguana-17148	95	27	oil	oil	NOUN
iguana-17148	95	28	spill	spill	NOUN
iguana-17148	95	29	,	,	PUNCT
iguana-17148	95	30	dogs	dog	NOUN
iguana-17148	95	31	,	,	PUNCT
iguana-17148	95	32	cats	cat	NOUN
iguana-17148	95	33	,	,	PUNCT
iguana-17148	95	34	rats	rat	NOUN
iguana-17148	95	35	,	,	PUNCT
iguana-17148	95	36	pigs	pig	NOUN
iguana-17148	95	37	santa	santa	PROPN
iguana-17148	95	38	fe	fe	PROPN
iguana-17148	95	39	3,000	3,000	NUM
iguana-17148	95	40	16,000	16,000	NUM
iguana-17148	95	41	667	667	NUM
iguana-17148	95	42	oil	oil	NOUN
iguana-17148	95	43	spill	spill	NOUN
iguana-17148	95	44	floreana	floreana	NOUN
iguana-17148	95	45	2,000	2,000	NUM
iguana-17148	95	46	16,000	16,000	NUM
iguana-17148	95	47	92	92	NUM
iguana-17148	95	48	oil	oil	NOUN
iguana-17148	95	49	spill	spill	NOUN
iguana-17148	95	50	,	,	PUNCT
iguana-17148	95	51	cats	cat	NOUN
iguana-17148	95	52	,	,	PUNCT
iguana-17148	95	53	rats	rat	NOUN
iguana-17148	95	54	,	,	PUNCT
iguana-17148	95	55	pigs	pig	NOUN
iguana-17148	95	56	santa	santa	PROPN
iguana-17148	95	57	cruz	cruz	PROPN
iguana-17148	95	58	hassi	hassi	PROPN
iguana-17148	95	59	2,000	2,000	NUM
iguana-17148	95	60	13,000	13,000	NUM
iguana-17148	95	61	13	13	NUM
iguana-17148	95	62	oil	oil	NOUN
iguana-17148	95	63	spill	spill	NOUN
iguana-17148	95	64	,	,	PUNCT
iguana-17148	95	65	dogs	dog	NOUN
iguana-17148	95	66	,	,	PUNCT
iguana-17148	95	67	cats	cat	NOUN
iguana-17148	95	68	,	,	PUNCT
iguana-17148	95	69	rats	rat	NOUN
iguana-17148	95	70	,	,	PUNCT
iguana-17148	95	71	pigs	pig	NOUN
iguana-17148	95	72	española	española	PROPN
iguana-17148	95	73	venustissimus	venustissimus	VERB
iguana-17148	95	74	1,700	1,700	NUM
iguana-17148	95	75	21,000	21,000	NUM
iguana-17148	95	76	350	350	NUM
iguana-17148	95	77	oil	oil	NOUN
iguana-17148	95	78	spill	spill	NOUN
iguana-17148	95	79	genovesa	genovesa	PROPN
iguana-17148	95	80	nanus	nanus	PROPN
iguana-17148	95	81	900	900	NUM
iguana-17148	95	82	15,000	15,000	NUM
iguana-17148	95	83	1071	1071	NUM
iguana-17148	95	84	oil	oil	NOUN
iguana-17148	95	85	spill	spill	NOUN
iguana-17148	95	86	marchena	marchena	PROPN
iguana-17148	95	87	1,000	1,000	NUM
iguana-17148	95	88	10,000	10,000	NUM
iguana-17148	95	89	77	77	NUM
iguana-17148	95	90	oil	oil	NOUN
iguana-17148	95	91	spill	spill	NOUN
iguana-17148	95	92	pinta	pinta	PROPN
iguana-17148	95	93	sielmanni	sielmanni	NOUN
iguana-17148	95	94	800	800	NUM
iguana-17148	95	95	6,000	6,000	NUM
iguana-17148	95	96	100	100	NUM
iguana-17148	95	97	oil	oil	NOUN
iguana-17148	95	98	spill	spill	NOUN
iguana-17148	95	99	santiago	santiago	PROPN
iguana-17148	95	100	mertensi	mertensi	PROPN
iguana-17148	95	101	450	450	NUM
iguana-17148	95	102	4,000	4,000	NUM
iguana-17148	95	103	7	7	NUM
iguana-17148	95	104	oil	oil	NOUN
iguana-17148	95	105	spill	spill	NOUN
iguana-17148	95	106	,	,	PUNCT
iguana-17148	95	107	cats	cat	NOUN
iguana-17148	95	108	,	,	PUNCT
iguana-17148	95	109	rats	rat	NOUN
iguana-17148	95	110	wolf	wolf	PROPN
iguana-17148	95	111	400	400	NUM
iguana-17148	95	112	1,500	1,500	NUM
iguana-17148	95	113	1154	1154	NUM
iguana-17148	95	114	oil	oil	NOUN
iguana-17148	95	115	spill	spill	NOUN
iguana-17148	95	116	darwin	darwin	PROPN
iguana-17148	95	117	200	200	NUM
iguana-17148	95	118	800	800	NUM
iguana-17148	95	119	727	727	NUM
iguana-17148	95	120	oil	oil	NOUN
iguana-17148	95	121	spill	spill	NOUN
iguana-17148	95	122	pinzon	pinzon	NOUN
iguana-17148	95	123	200	200	NUM
iguana-17148	95	124	900	900	NUM
iguana-17148	95	125	50	50	NUM
iguana-17148	95	126	oil	oil	NOUN
iguana-17148	95	127	spill	spill	NOUN
iguana-17148	95	128	rabida	rabida	VERB
iguana-17148	95	129	200	200	NUM
iguana-17148	95	130	2,000	2,000	NUM
iguana-17148	95	131	408	408	NUM
iguana-17148	95	132	oil	oil	NOUN
iguana-17148	95	133	spill	spill	NOUN
iguana-17148	95	134	seymour	seymour	PROPN
iguana-17148	95	135	norte	norte	PROPN
iguana-17148	95	136	100	100	NUM
iguana-17148	95	137	1,500	1,500	NUM
iguana-17148	95	138	789	789	NUM
iguana-17148	95	139	oil	oil	NOUN
iguana-17148	95	140	spill	spill	NOUN
iguana-17148	95	141	san	san	PROPN
iguana-17148	95	142	cristobal	cristobal	PROPN
iguana-17148	95	143	50	50	NUM
iguana-17148	95	144	400	400	NUM
iguana-17148	95	145	1	1	NUM
iguana-17148	95	146	oil	oil	NOUN
iguana-17148	95	147	spill	spill	NOUN
iguana-17148	95	148	,	,	PUNCT
iguana-17148	95	149	cats	cat	NOUN
iguana-17148	95	150	,	,	PUNCT
iguana-17148	95	151	rats	rat	NOUN
iguana-17148	95	152	,	,	PUNCT
iguana-17148	95	153	pigs	pig	NOUN
iguana-17148	95	154	remaining	remain	VERB
iguana-17148	95	155	islands	island	NOUN
iguana-17148	95	156	2,000	2,000	NUM
iguana-17148	95	157	10,000	10,000	NUM
iguana-17148	95	158	na	na	ADP
iguana-17148	95	159	oil	oil	NOUN
iguana-17148	95	160	spill	spill	NOUN
iguana-17148	95	161	total	total	NOUN
iguana-17148	95	162	~37,000	~37,000	PUNCT
iguana-17148	96	1	~280,000	~280,000	PUNCT
iguana-17148	96	2	tinction	tinction	NOUN
iguana-17148	96	3	of	of	ADP
iguana-17148	96	4	“	"	PUNCT
iguana-17148	96	5	subspecies	subspecie	NOUN
iguana-17148	96	6	”	"	PUNCT
iguana-17148	96	7	is	be	AUX
iguana-17148	96	8	less	less	ADV
iguana-17148	96	9	clear	clear	ADJ
iguana-17148	96	10	(	(	PUNCT
iguana-17148	96	11	fig	fig	NOUN
iguana-17148	96	12	.	.	PUNCT
iguana-17148	97	1	1	1	NUM
iguana-17148	97	2	;	;	PUNCT
iguana-17148	97	3	rassmann	rassmann	PROPN
iguana-17148	97	4	1997	1997	NUM
iguana-17148	97	5	,	,	PUNCT
iguana-17148	97	6	rassmann	rassmann	PROPN
iguana-17148	97	7	et	et	PROPN
iguana-17148	97	8	al	al	PROPN
iguana-17148	97	9	.	.	PROPN
iguana-17148	97	10	1997	1997	NUM
iguana-17148	97	11	)	)	PUNCT
iguana-17148	97	12	.	.	PUNCT
iguana-17148	98	1	marine	marine	PROPN
iguana-17148	98	2	iguanas	iguana	NOUN
iguana-17148	98	3	reached	reach	VERB
iguana-17148	98	4	the	the	DET
iguana-17148	98	5	galápagos	galápagos	PROPN
iguana-17148	98	6	archipelago	archipelago	PROPN
iguana-17148	98	7	as	as	ADV
iguana-17148	98	8	early	early	ADV
iguana-17148	98	9	as	as	ADP
iguana-17148	98	10	10–15	10–15	NUM
iguana-17148	98	11	million	million	NUM
iguana-17148	98	12	years	year	NOUN
iguana-17148	98	13	ago	ago	ADV
iguana-17148	98	14	.	.	PUNCT
iguana-17148	99	1	they	they	PRON
iguana-17148	99	2	presumably	presumably	ADV
iguana-17148	99	3	arrived	arrive	VERB
iguana-17148	99	4	by	by	ADP
iguana-17148	99	5	“	"	PUNCT
iguana-17148	99	6	leap	leap	NOUN
iguana-17148	99	7	-	-	PUNCT
iguana-17148	99	8	frogging	frogging	NOUN
iguana-17148	99	9	”	"	PUNCT
iguana-17148	99	10	across	across	ADP
iguana-17148	99	11	now	now	ADV
iguana-17148	99	12	-	-	PUNCT
iguana-17148	99	13	sunken	sink	VERB
iguana-17148	99	14	islands	island	NOUN
iguana-17148	99	15	that	that	PRON
iguana-17148	99	16	are	be	AUX
iguana-17148	99	17	only	only	ADV
iguana-17148	99	18	detectable	detectable	ADJ
iguana-17148	99	19	as	as	ADP
iguana-17148	99	20	sea	sea	NOUN
iguana-17148	99	21	mounts	mount	NOUN
iguana-17148	99	22	between	between	ADP
iguana-17148	99	23	the	the	DET
iguana-17148	99	24	present	present	ADJ
iguana-17148	99	25	-	-	PUNCT
iguana-17148	99	26	day	day	NOUN
iguana-17148	99	27	galápagos	galápagos	NOUN
iguana-17148	99	28	islands	island	NOUN
iguana-17148	99	29	and	and	CCONJ
iguana-17148	99	30	the	the	DET
iguana-17148	99	31	south	south	ADJ
iguana-17148	99	32	american	american	PROPN
iguana-17148	99	33	mainland	mainland	PROPN
iguana-17148	99	34	.	.	PUNCT
iguana-17148	100	1	from	from	ADP
iguana-17148	100	2	those	those	DET
iguana-17148	100	3	early	early	ADJ
iguana-17148	100	4	galápagos	galápago	NOUN
iguana-17148	100	5	islands	island	NOUN
iguana-17148	100	6	,	,	PUNCT
iguana-17148	100	7	marine	marine	PROPN
iguana-17148	100	8	iguanas	iguana	NOUN
iguana-17148	100	9	dispersed	disperse	VERB
iguana-17148	100	10	to	to	ADP
iguana-17148	100	11	the	the	DET
iguana-17148	100	12	present	present	ADJ
iguana-17148	100	13	islands	island	NOUN
iguana-17148	100	14	and	and	CCONJ
iguana-17148	100	15	split	split	VERB
iguana-17148	100	16	into	into	ADP
iguana-17148	100	17	northwestern	northwestern	ADJ
iguana-17148	100	18	,	,	PUNCT
iguana-17148	100	19	central	central	ADJ
iguana-17148	100	20	,	,	PUNCT
iguana-17148	100	21	and	and	CCONJ
iguana-17148	100	22	southeastern	southeastern	ADJ
iguana-17148	100	23	clades	clade	NOUN
iguana-17148	100	24	.	.	PUNCT
iguana-17148	101	1	interestingly	interestingly	ADV
iguana-17148	101	2	,	,	PUNCT
iguana-17148	101	3	the	the	DET
iguana-17148	101	4	northwestern	northwestern	ADJ
iguana-17148	101	5	clade	clade	NOUN
iguana-17148	101	6	includes	include	VERB
iguana-17148	101	7	both	both	CCONJ
iguana-17148	101	8	the	the	DET
iguana-17148	101	9	largest	large	ADJ
iguana-17148	101	10	(	(	PUNCT
iguana-17148	101	11	on	on	ADP
iguana-17148	101	12	southwestern	southwestern	ADJ
iguana-17148	101	13	isabela	isabela	NOUN
iguana-17148	101	14	)	)	PUNCT
iguana-17148	101	15	but	but	CCONJ
iguana-17148	101	16	also	also	ADV
iguana-17148	101	17	fairly	fairly	ADV
iguana-17148	101	18	small	small	ADJ
iguana-17148	101	19	iguanas	iguana	NOUN
iguana-17148	101	20	(	(	PUNCT
iguana-17148	101	21	on	on	ADP
iguana-17148	101	22	pinta	pinta	NOUN
iguana-17148	101	23	)	)	PUNCT
iguana-17148	101	24	,	,	PUNCT
iguana-17148	101	25	indicating	indicate	VERB
iguana-17148	101	26	that	that	SCONJ
iguana-17148	101	27	strong	strong	ADJ
iguana-17148	101	28	selection	selection	NOUN
iguana-17148	101	29	on	on	ADP
iguana-17148	101	30	body	body	NOUN
iguana-17148	101	31	size	size	NOUN
iguana-17148	101	32	is	be	AUX
iguana-17148	101	33	present	present	ADJ
iguana-17148	101	34	,	,	PUNCT
iguana-17148	101	35	although	although	SCONJ
iguana-17148	101	36	not	not	PART
iguana-17148	101	37	apparent	apparent	ADJ
iguana-17148	101	38	in	in	ADP
iguana-17148	101	39	the	the	DET
iguana-17148	101	40	genetic	genetic	ADJ
iguana-17148	101	41	markers	marker	NOUN
iguana-17148	101	42	that	that	PRON
iguana-17148	101	43	were	be	AUX
iguana-17148	101	44	investigated	investigate	VERB
iguana-17148	101	45	(	(	PUNCT
iguana-17148	101	46	rassmann	rassmann	PROPN
iguana-17148	101	47	et	et	PROPN
iguana-17148	101	48	al	al	PROPN
iguana-17148	101	49	.	.	PROPN
iguana-17148	101	50	1997	1997	NUM
iguana-17148	101	51	)	)	PUNCT
iguana-17148	101	52	.	.	PUNCT
iguana-17148	102	1	these	these	DET
iguana-17148	102	2	distinct	distinct	ADJ
iguana-17148	102	3	inter	inter	ADJ
iguana-17148	102	4	-	-	ADJ
iguana-17148	102	5	island	island	ADJ
iguana-17148	102	6	differences	difference	NOUN
iguana-17148	102	7	obviously	obviously	ADV
iguana-17148	102	8	indicate	indicate	VERB
iguana-17148	102	9	that	that	SCONJ
iguana-17148	102	10	all	all	DET
iguana-17148	102	11	island	island	NOUN
iguana-17148	102	12	populations	population	NOUN
iguana-17148	102	13	should	should	AUX
iguana-17148	102	14	be	be	AUX
iguana-17148	102	15	conserved	conserve	VERB
iguana-17148	102	16	,	,	PUNCT
iguana-17148	102	17	although	although	SCONJ
iguana-17148	102	18	no	no	DET
iguana-17148	102	19	immediate	immediate	ADJ
iguana-17148	102	20	threat	threat	NOUN
iguana-17148	102	21	exists	exist	VERB
iguana-17148	102	22	to	to	ADP
iguana-17148	102	23	most	most	ADJ
iguana-17148	102	24	island	island	NOUN
iguana-17148	102	25	populations	population	NOUN
iguana-17148	102	26	.	.	PUNCT
iguana-17148	103	1	however	however	ADV
iguana-17148	103	2	,	,	PUNCT
iguana-17148	103	3	even	even	ADV
iguana-17148	103	4	the	the	DET
iguana-17148	103	5	most	most	ADV
iguana-17148	103	6	pristine	pristine	ADJ
iguana-17148	103	7	populations	population	NOUN
iguana-17148	103	8	may	may	AUX
iguana-17148	103	9	suffer	suffer	VERB
iguana-17148	103	10	immediately	immediately	ADV
iguana-17148	103	11	and	and	CCONJ
iguana-17148	103	12	heavily	heavily	ADV
iguana-17148	103	13	from	from	ADP
iguana-17148	103	14	environmental	environmental	ADJ
iguana-17148	103	15	disasters	disaster	NOUN
iguana-17148	103	16	such	such	ADJ
iguana-17148	103	17	as	as	ADP
iguana-17148	103	18	an	an	DET
iguana-17148	103	19	oil	oil	NOUN
iguana-17148	103	20	spill	spill	NOUN
iguana-17148	103	21	,	,	PUNCT
iguana-17148	103	22	even	even	ADV
iguana-17148	103	23	if	if	SCONJ
iguana-17148	103	24	the	the	DET
iguana-17148	103	25	contamination	contamination	NOUN
iguana-17148	103	26	levels	level	NOUN
iguana-17148	103	27	are	be	AUX
iguana-17148	103	28	very	very	ADV
iguana-17148	103	29	minor	minor	ADJ
iguana-17148	103	30	(	(	PUNCT
iguana-17148	103	31	wikelski	wikelski	NOUN
iguana-17148	103	32	et	et	PROPN
iguana-17148	103	33	al	al	PROPN
iguana-17148	103	34	.	.	PROPN
iguana-17148	103	35	,	,	PUNCT
iguana-17148	103	36	in	in	ADP
iguana-17148	103	37	preparation	preparation	NOUN
iguana-17148	103	38	;	;	PUNCT
iguana-17148	103	39	charles	charles	PROPN
iguana-17148	103	40	darwin	darwin	PROPN
iguana-17148	103	41	research	research	PROPN
iguana-17148	103	42	station	station	PROPN
iguana-17148	103	43	,	,	PUNCT
iguana-17148	103	44	unpublished	unpublished	ADJ
iguana-17148	103	45	report	report	NOUN
iguana-17148	103	46	on	on	ADP
iguana-17148	103	47	the	the	DET
iguana-17148	103	48	january	january	PROPN
iguana-17148	103	49	2001	2001	NUM
iguana-17148	103	50	oil	oil	NOUN
iguana-17148	103	51	spill	spill	NOUN
iguana-17148	103	52	)	)	PUNCT
iguana-17148	103	53	.	.	PUNCT
iguana-17148	104	1	on	on	ADP
iguana-17148	104	2	santa	santa	PROPN
iguana-17148	104	3	fe	fe	PROPN
iguana-17148	104	4	,	,	PUNCT
iguana-17148	104	5	approximately	approximately	ADV
iguana-17148	104	6	60	60	NUM
iguana-17148	104	7	%	%	NOUN
iguana-17148	104	8	of	of	ADP
iguana-17148	104	9	the	the	DET
iguana-17148	104	10	entire	entire	ADJ
iguana-17148	104	11	marine	marine	ADJ
iguana-17148	104	12	iguana	iguana	NOUN
iguana-17148	104	13	population	population	NOUN
iguana-17148	104	14	disappeared	disappear	VERB
iguana-17148	104	15	as	as	ADP
iguana-17148	104	16	a	a	DET
iguana-17148	104	17	consequence	consequence	NOUN
iguana-17148	104	18	of	of	ADP
iguana-17148	104	19	low	low	ADJ
iguana-17148	104	20	-	-	PUNCT
iguana-17148	104	21	level	level	NOUN
iguana-17148	104	22	oiling	oiling	NOUN
iguana-17148	104	23	when	when	SCONJ
iguana-17148	104	24	194	194	NUM
iguana-17148	104	25	iguana	iguana	NOUN
iguana-17148	104	26	•	•	NUM
iguana-17148	104	27	volume	volume	NOUN
iguana-17148	104	28	11	11	NUM
iguana-17148	104	29	,	,	PUNCT
iguana-17148	104	30	number	number	NOUN
iguana-17148	104	31	4	4	NUM
iguana-17148	104	32	•	•	NOUN
iguana-17148	104	33	december	december	PROPN
iguana-17148	104	34	2004	2004	NUM
iguana-17148	104	35	wikelski	wikelski	NOUN
iguana-17148	104	36	and	and	CCONJ
iguana-17148	104	37	nelson	nelson	PROPN
iguana-17148	104	38	schematic	schematic	ADJ
iguana-17148	104	39	map	map	NOUN
iguana-17148	104	40	of	of	ADP
iguana-17148	104	41	the	the	DET
iguana-17148	104	42	galápagos	galápagos	PROPN
iguana-17148	104	43	archipelago	archipelago	PROPN
iguana-17148	104	44	and	and	CCONJ
iguana-17148	104	45	the	the	DET
iguana-17148	104	46	different	different	ADJ
iguana-17148	104	47	subspecies	subspecie	NOUN
iguana-17148	104	48	of	of	ADP
iguana-17148	104	49	marine	marine	ADJ
iguana-17148	104	50	iguanas	iguana	NOUN
iguana-17148	104	51	as	as	SCONJ
iguana-17148	104	52	described	describe	VERB
iguana-17148	104	53	by	by	ADP
iguana-17148	104	54	eibl	eibl	NOUN
iguana-17148	104	55	-	-	PUNCT
iguana-17148	104	56	eibesfeldt	eibesfeldt	PROPN
iguana-17148	104	57	(	(	PUNCT
iguana-17148	104	58	1962	1962	NUM
iguana-17148	104	59	;	;	PUNCT
iguana-17148	104	60	inset	inset	NOUN
iguana-17148	104	61	subspecies	subspecie	NOUN
iguana-17148	104	62	names	name	NOUN
iguana-17148	104	63	)	)	PUNCT
iguana-17148	104	64	.	.	PUNCT
iguana-17148	105	1	superimposed	superimpose	VERB
iguana-17148	105	2	(	(	PUNCT
iguana-17148	105	3	ovals	oval	NOUN
iguana-17148	105	4	)	)	PUNCT
iguana-17148	105	5	are	be	AUX
iguana-17148	105	6	the	the	DET
iguana-17148	105	7	three	three	NUM
iguana-17148	105	8	genetically	genetically	ADV
iguana-17148	105	9	similar	similar	ADJ
iguana-17148	105	10	clades	clade	NOUN
iguana-17148	105	11	,	,	PUNCT
iguana-17148	105	12	based	base	VERB
iguana-17148	105	13	on	on	ADP
iguana-17148	105	14	nuclear	nuclear	ADJ
iguana-17148	105	15	and	and	CCONJ
iguana-17148	105	16	mitochondrial	mitochondrial	ADJ
iguana-17148	105	17	genetic	genetic	ADJ
iguana-17148	105	18	analysis	analysis	NOUN
iguana-17148	105	19	(	(	PUNCT
iguana-17148	105	20	rassmann	rassmann	PROPN
iguana-17148	105	21	et	et	PROPN
iguana-17148	105	22	al	al	PROPN
iguana-17148	105	23	.	.	PROPN
iguana-17148	105	24	1997	1997	NUM
iguana-17148	105	25	)	)	PUNCT
iguana-17148	105	26	.	.	PUNCT
iguana-17148	106	1	genetic	genetic	ADJ
iguana-17148	106	2	and	and	CCONJ
iguana-17148	106	3	morphological	morphological	ADJ
iguana-17148	106	4	information	information	NOUN
iguana-17148	106	5	is	be	AUX
iguana-17148	106	6	not	not	PART
iguana-17148	106	7	entirely	entirely	ADV
iguana-17148	106	8	compatible	compatible	ADJ
iguana-17148	106	9	,	,	PUNCT
iguana-17148	106	10	indicated	indicate	VERB
iguana-17148	106	11	,	,	PUNCT
iguana-17148	106	12	for	for	ADP
iguana-17148	106	13	example	example	NOUN
iguana-17148	106	14	,	,	PUNCT
iguana-17148	106	15	by	by	ADP
iguana-17148	106	16	the	the	DET
iguana-17148	106	17	fact	fact	NOUN
iguana-17148	106	18	that	that	SCONJ
iguana-17148	106	19	the	the	DET
iguana-17148	106	20	amblyrhynchus	amblyrhynchus	NOUN
iguana-17148	106	21	cristatus	cristatus	PROPN
iguana-17148	106	22	mertensi	mertensi	PROPN
iguana-17148	106	23	subspecies	subspecie	NOUN
iguana-17148	106	24	is	be	AUX
iguana-17148	106	25	found	find	VERB
iguana-17148	106	26	in	in	ADP
iguana-17148	106	27	two	two	NUM
iguana-17148	106	28	genetically	genetically	ADV
iguana-17148	106	29	distinct	distinct	ADJ
iguana-17148	106	30	clades	clade	NOUN
iguana-17148	106	31	(	(	PUNCT
iguana-17148	106	32	on	on	ADP
iguana-17148	106	33	san	san	PROPN
iguana-17148	106	34	cristobal	cristobal	PROPN
iguana-17148	106	35	and	and	CCONJ
iguana-17148	106	36	santiago	santiago	PROPN
iguana-17148	106	37	islands	island	NOUN
iguana-17148	106	38	)	)	PUNCT
iguana-17148	106	39	.	.	PUNCT
iguana-17148	107	1	iguana	iguana	NOUN
iguana-17148	107	2	•	•	NUM
iguana-17148	107	3	volume	volume	NOUN
iguana-17148	107	4	11	11	NUM
iguana-17148	107	5	,	,	PUNCT
iguana-17148	107	6	number	number	NOUN
iguana-17148	107	7	4	4	NUM
iguana-17148	107	8	•	•	NOUN
iguana-17148	107	9	december	december	PROPN
iguana-17148	107	10	2004	2004	NUM
iguana-17148	107	11	195galápagos	195galápagos	PROPN
iguana-17148	107	12	marine	marine	NOUN
iguana-17148	107	13	iguanas	iguana	VERB
iguana-17148	107	14	the	the	DET
iguana-17148	107	15	oil	oil	NOUN
iguana-17148	107	16	tanker	tanker	NOUN
iguana-17148	107	17	“	"	PUNCT
iguana-17148	107	18	jessica	jessica	PROPN
iguana-17148	107	19	”	"	PUNCT
iguana-17148	107	20	ran	run	VERB
iguana-17148	107	21	ashore	ashore	ADV
iguana-17148	107	22	on	on	ADP
iguana-17148	107	23	nearby	nearby	ADJ
iguana-17148	107	24	san	san	PROPN
iguana-17148	107	25	cristobal	cristobal	PROPN
iguana-17148	107	26	island	island	PROPN
iguana-17148	107	27	and	and	CCONJ
iguana-17148	107	28	spilled	spill	VERB
iguana-17148	107	29	about	about	ADV
iguana-17148	107	30	750,000	750,000	NUM
iguana-17148	107	31	gallons	gallon	NOUN
iguana-17148	107	32	of	of	ADP
iguana-17148	107	33	diesel	diesel	NOUN
iguana-17148	107	34	and	and	CCONJ
iguana-17148	107	35	bunker	bunker	NOUN
iguana-17148	107	36	oil	oil	NOUN
iguana-17148	107	37	.	.	PUNCT
iguana-17148	108	1	consequently	consequently	ADV
iguana-17148	108	2	,	,	PUNCT
iguana-17148	108	3	oil	oil	NOUN
iguana-17148	108	4	contamination	contamination	NOUN
iguana-17148	108	5	could	could	AUX
iguana-17148	108	6	pose	pose	VERB
iguana-17148	108	7	a	a	DET
iguana-17148	108	8	serious	serious	ADJ
iguana-17148	108	9	threat	threat	NOUN
iguana-17148	108	10	to	to	ADP
iguana-17148	108	11	all	all	DET
iguana-17148	108	12	populations	population	NOUN
iguana-17148	108	13	of	of	ADP
iguana-17148	108	14	galápagos	galápago	NOUN
iguana-17148	108	15	marine	marine	NOUN
iguana-17148	108	16	iguanas	iguana	NOUN
iguana-17148	108	17	,	,	PUNCT
iguana-17148	108	18	even	even	ADV
iguana-17148	108	19	those	those	PRON
iguana-17148	108	20	that	that	PRON
iguana-17148	108	21	live	live	VERB
iguana-17148	108	22	far	far	ADV
iguana-17148	108	23	from	from	ADP
iguana-17148	108	24	human	human	ADJ
iguana-17148	108	25	settlements	settlement	NOUN
iguana-17148	108	26	and	and	CCONJ
iguana-17148	108	27	would	would	AUX
iguana-17148	108	28	otherwise	otherwise	ADV
iguana-17148	108	29	be	be	AUX
iguana-17148	108	30	considered	consider	VERB
iguana-17148	108	31	pristine	pristine	ADJ
iguana-17148	108	32	.	.	PUNCT
iguana-17148	109	1	as	as	ADP
iguana-17148	109	2	to	to	ADP
iguana-17148	109	3	the	the	DET
iguana-17148	109	4	specific	specific	ADJ
iguana-17148	109	5	mechanism	mechanism	NOUN
iguana-17148	109	6	of	of	ADP
iguana-17148	109	7	mortality	mortality	NOUN
iguana-17148	109	8	,	,	PUNCT
iguana-17148	109	9	we	we	PRON
iguana-17148	109	10	suggest	suggest	VERB
iguana-17148	109	11	that	that	SCONJ
iguana-17148	109	12	hindgut	hindgut	NOUN
iguana-17148	109	13	endosymbionts	endosymbiont	NOUN
iguana-17148	109	14	are	be	AUX
iguana-17148	109	15	highly	highly	ADV
iguana-17148	109	16	susceptible	susceptible	ADJ
iguana-17148	109	17	to	to	ADP
iguana-17148	109	18	oil	oil	NOUN
iguana-17148	109	19	in	in	ADP
iguana-17148	109	20	their	their	PRON
iguana-17148	109	21	digestive	digestive	ADJ
iguana-17148	109	22	substrate	substrate	NOUN
iguana-17148	109	23	.	.	PUNCT
iguana-17148	110	1	marine	marine	ADJ
iguana-17148	110	2	iguanas	iguana	NOUN
iguana-17148	110	3	need	need	VERB
iguana-17148	110	4	these	these	DET
iguana-17148	110	5	bacteria	bacteria	NOUN
iguana-17148	110	6	to	to	PART
iguana-17148	110	7	digest	digest	VERB
iguana-17148	110	8	algae	algae	NOUN
iguana-17148	110	9	fully	fully	ADV
iguana-17148	110	10	,	,	PUNCT
iguana-17148	110	11	a	a	DET
iguana-17148	110	12	process	process	NOUN
iguana-17148	110	13	that	that	PRON
iguana-17148	110	14	can	can	AUX
iguana-17148	110	15	take	take	VERB
iguana-17148	110	16	up	up	ADP
iguana-17148	110	17	to	to	PART
iguana-17148	110	18	two	two	NUM
iguana-17148	110	19	weeks	week	NOUN
iguana-17148	110	20	.	.	PUNCT
iguana-17148	111	1	these	these	DET
iguana-17148	111	2	specialized	specialized	ADJ
iguana-17148	111	3	and	and	CCONJ
iguana-17148	111	4	highly	highly	ADV
iguana-17148	111	5	effective	effective	ADJ
iguana-17148	111	6	gut	gut	NOUN
iguana-17148	111	7	bacteria	bacteria	NOUN
iguana-17148	111	8	probably	probably	ADV
iguana-17148	111	9	suffered	suffer	VERB
iguana-17148	111	10	heavily	heavily	ADV
iguana-17148	111	11	due	due	ADP
iguana-17148	111	12	to	to	ADP
iguana-17148	111	13	the	the	DET
iguana-17148	111	14	oil	oil	NOUN
iguana-17148	111	15	spill	spill	NOUN
iguana-17148	111	16	,	,	PUNCT
iguana-17148	111	17	which	which	PRON
iguana-17148	111	18	in	in	ADP
iguana-17148	111	19	turn	turn	NOUN
iguana-17148	111	20	caused	cause	VERB
iguana-17148	111	21	failed	fail	VERB
iguana-17148	111	22	digestion	digestion	NOUN
iguana-17148	111	23	and	and	CCONJ
iguana-17148	111	24	widespread	widespread	ADJ
iguana-17148	111	25	starvation	starvation	NOUN
iguana-17148	111	26	among	among	ADP
iguana-17148	111	27	marine	marine	ADJ
iguana-17148	111	28	iguanas	iguana	NOUN
iguana-17148	111	29	.	.	PUNCT
iguana-17148	112	1	on	on	ADP
iguana-17148	112	2	at	at	ADV
iguana-17148	112	3	least	least	ADV
iguana-17148	112	4	five	five	NUM
iguana-17148	112	5	islands	island	NOUN
iguana-17148	112	6	,	,	PUNCT
iguana-17148	112	7	marine	marine	ADJ
iguana-17148	112	8	iguana	iguana	NOUN
iguana-17148	112	9	populations	population	NOUN
iguana-17148	112	10	also	also	ADV
iguana-17148	112	11	are	be	AUX
iguana-17148	112	12	heavily	heavily	ADV
iguana-17148	112	13	impacted	impact	VERB
iguana-17148	112	14	by	by	ADP
iguana-17148	112	15	human	human	NOUN
iguana-17148	112	16	-	-	PUNCT
iguana-17148	112	17	introduced	introduce	VERB
iguana-17148	112	18	predators	predator	NOUN
iguana-17148	112	19	(	(	PUNCT
iguana-17148	112	20	cayot	cayot	NOUN
iguana-17148	112	21	et	et	PROPN
iguana-17148	112	22	al	al	PROPN
iguana-17148	112	23	.	.	PROPN
iguana-17148	112	24	1994	1994	NUM
iguana-17148	112	25	)	)	PUNCT
iguana-17148	112	26	.	.	PUNCT
iguana-17148	113	1	the	the	DET
iguana-17148	113	2	most	most	ADV
iguana-17148	113	3	important	important	ADJ
iguana-17148	113	4	predators	predator	NOUN
iguana-17148	113	5	are	be	AUX
iguana-17148	113	6	feral	feral	ADJ
iguana-17148	113	7	cats	cat	NOUN
iguana-17148	113	8	,	,	PUNCT
iguana-17148	113	9	dogs	dog	NOUN
iguana-17148	113	10	,	,	PUNCT
iguana-17148	113	11	rats	rat	NOUN
iguana-17148	113	12	,	,	PUNCT
iguana-17148	113	13	and	and	CCONJ
iguana-17148	113	14	pigs	pig	NOUN
iguana-17148	113	15	.	.	PUNCT
iguana-17148	114	1	we	we	PRON
iguana-17148	114	2	are	be	AUX
iguana-17148	114	3	currently	currently	ADV
iguana-17148	114	4	uncertain	uncertain	ADJ
iguana-17148	114	5	whether	whether	SCONJ
iguana-17148	114	6	those	those	DET
iguana-17148	114	7	marine	marine	ADJ
iguana-17148	114	8	iguana	iguana	NOUN
iguana-17148	114	9	populations	population	NOUN
iguana-17148	114	10	already	already	ADV
iguana-17148	114	11	affected	affect	VERB
iguana-17148	114	12	are	be	AUX
iguana-17148	114	13	jeopardized	jeopardize	VERB
iguana-17148	114	14	to	to	ADP
iguana-17148	114	15	an	an	DET
iguana-17148	114	16	extent	extent	NOUN
iguana-17148	114	17	that	that	SCONJ
iguana-17148	114	18	their	their	PRON
iguana-17148	114	19	long	long	ADJ
iguana-17148	114	20	-	-	PUNCT
iguana-17148	114	21	term	term	NOUN
iguana-17148	114	22	survival	survival	NOUN
iguana-17148	114	23	is	be	AUX
iguana-17148	114	24	in	in	ADP
iguana-17148	114	25	question	question	NOUN
iguana-17148	114	26	.	.	PUNCT
iguana-17148	115	1	some	some	DET
iguana-17148	115	2	populations	population	NOUN
iguana-17148	115	3	on	on	ADP
iguana-17148	115	4	certain	certain	ADJ
iguana-17148	115	5	parts	part	NOUN
iguana-17148	115	6	of	of	ADP
iguana-17148	115	7	western	western	ADJ
iguana-17148	115	8	isabela	isabela	PROPN
iguana-17148	115	9	island	island	NOUN
iguana-17148	115	10	appear	appear	VERB
iguana-17148	115	11	to	to	PART
iguana-17148	115	12	be	be	AUX
iguana-17148	115	13	in	in	ADP
iguana-17148	115	14	immediate	immediate	ADJ
iguana-17148	115	15	danger	danger	NOUN
iguana-17148	115	16	of	of	ADP
iguana-17148	115	17	extinction	extinction	NOUN
iguana-17148	115	18	.	.	PUNCT
iguana-17148	116	1	during	during	ADP
iguana-17148	116	2	several	several	ADJ
iguana-17148	116	3	research	research	NOUN
iguana-17148	116	4	expeditions	expedition	NOUN
iguana-17148	116	5	,	,	PUNCT
iguana-17148	116	6	we	we	PRON
iguana-17148	116	7	only	only	ADV
iguana-17148	116	8	found	find	VERB
iguana-17148	116	9	adult	adult	NOUN
iguana-17148	116	10	iguanas	iguana	NOUN
iguana-17148	116	11	and	and	CCONJ
iguana-17148	116	12	very	very	ADV
iguana-17148	116	13	few	few	ADJ
iguana-17148	116	14	young	young	ADJ
iguana-17148	116	15	or	or	CCONJ
iguana-17148	116	16	hatchlings	hatchling	NOUN
iguana-17148	116	17	,	,	PUNCT
iguana-17148	116	18	indicating	indicate	VERB
iguana-17148	116	19	almost	almost	ADV
iguana-17148	116	20	no	no	PRON
iguana-17148	116	21	recruitment	recruitment	NOUN
iguana-17148	116	22	,	,	PUNCT
iguana-17148	116	23	at	at	ADP
iguana-17148	116	24	least	least	ADJ
iguana-17148	116	25	not	not	PART
iguana-17148	116	26	along	along	ADP
iguana-17148	116	27	the	the	DET
iguana-17148	116	28	shoreline	shoreline	NOUN
iguana-17148	116	29	of	of	ADP
iguana-17148	116	30	the	the	DET
iguana-17148	116	31	main	main	ADJ
iguana-17148	116	32	island	island	NOUN
iguana-17148	116	33	.	.	PUNCT
iguana-17148	117	1	nevertheless	nevertheless	ADV
iguana-17148	117	2	,	,	PUNCT
iguana-17148	117	3	some	some	DET
iguana-17148	117	4	hatchlings	hatchling	NOUN
iguana-17148	117	5	and	and	CCONJ
iguana-17148	117	6	yearlings	yearling	NOUN
iguana-17148	117	7	were	be	AUX
iguana-17148	117	8	seen	see	VERB
iguana-17148	117	9	on	on	ADP
iguana-17148	117	10	small	small	ADJ
iguana-17148	117	11	offshore	offshore	ADJ
iguana-17148	117	12	islets	islet	NOUN
iguana-17148	117	13	.	.	PUNCT
iguana-17148	118	1	a	a	DET
iguana-17148	118	2	similar	similar	ADJ
iguana-17148	118	3	situation	situation	NOUN
iguana-17148	118	4	applies	apply	VERB
iguana-17148	118	5	to	to	ADP
iguana-17148	118	6	marine	marine	ADJ
iguana-17148	118	7	iguanas	iguana	NOUN
iguana-17148	118	8	on	on	ADP
iguana-17148	118	9	san	san	PROPN
iguana-17148	118	10	cristobal	cristobal	PROPN
iguana-17148	118	11	island	island	PROPN
iguana-17148	118	12	,	,	PUNCT
iguana-17148	118	13	where	where	SCONJ
iguana-17148	118	14	only	only	ADV
iguana-17148	118	15	several	several	ADJ
iguana-17148	118	16	hundred	hundred	NUM
iguana-17148	118	17	iguanas	iguana	NOUN
iguana-17148	118	18	survive	survive	NOUN
iguana-17148	118	19	.	.	PUNCT
iguana-17148	119	1	thus	thus	ADV
iguana-17148	119	2	,	,	PUNCT
iguana-17148	119	3	we	we	PRON
iguana-17148	119	4	would	would	AUX
iguana-17148	119	5	classify	classify	VERB
iguana-17148	119	6	these	these	DET
iguana-17148	119	7	populations	population	NOUN
iguana-17148	119	8	as	as	SCONJ
iguana-17148	119	9	highly	highly	ADV
iguana-17148	119	10	endangered	endanger	VERB
iguana-17148	119	11	,	,	PUNCT
iguana-17148	119	12	and	and	CCONJ
iguana-17148	119	13	recommend	recommend	VERB
iguana-17148	119	14	that	that	SCONJ
iguana-17148	119	15	immediate	immediate	ADJ
iguana-17148	119	16	action	action	NOUN
iguana-17148	119	17	be	be	AUX
iguana-17148	119	18	taken	take	VERB
iguana-17148	119	19	.	.	PUNCT
iguana-17148	120	1	management	management	NOUN
iguana-17148	120	2	needs	need	VERB
iguana-17148	120	3	for	for	ADP
iguana-17148	120	4	the	the	DET
iguana-17148	120	5	future	future	ADJ
iguana-17148	120	6	four	four	NUM
iguana-17148	120	7	major	major	ADJ
iguana-17148	120	8	problems	problem	NOUN
iguana-17148	120	9	exist	exist	VERB
iguana-17148	120	10	for	for	ADP
iguana-17148	120	11	marine	marine	ADJ
iguana-17148	120	12	iguana	iguana	NOUN
iguana-17148	120	13	populations	population	NOUN
iguana-17148	120	14	,	,	PUNCT
iguana-17148	120	15	and	and	CCONJ
iguana-17148	120	16	we	we	PRON
iguana-17148	120	17	recommend	recommend	VERB
iguana-17148	120	18	the	the	DET
iguana-17148	120	19	following	follow	VERB
iguana-17148	120	20	conservation	conservation	NOUN
iguana-17148	120	21	-	-	PUNCT
iguana-17148	120	22	related	relate	VERB
iguana-17148	120	23	research	research	NOUN
iguana-17148	120	24	projects	project	NOUN
iguana-17148	120	25	to	to	PART
iguana-17148	120	26	determine	determine	VERB
iguana-17148	120	27	the	the	DET
iguana-17148	120	28	causes	cause	NOUN
iguana-17148	120	29	of	of	ADP
iguana-17148	120	30	the	the	DET
iguana-17148	120	31	threats	threat	NOUN
iguana-17148	120	32	and	and	CCONJ
iguana-17148	120	33	to	to	PART
iguana-17148	120	34	elaborate	elaborate	VERB
iguana-17148	120	35	possible	possible	ADJ
iguana-17148	120	36	solutions	solution	NOUN
iguana-17148	120	37	:	:	PUNCT
iguana-17148	120	38	(	(	PUNCT
iguana-17148	120	39	i	i	NOUN
iguana-17148	120	40	)	)	PUNCT
iguana-17148	120	41	we	we	PRON
iguana-17148	120	42	need	need	VERB
iguana-17148	120	43	to	to	PART
iguana-17148	120	44	study	study	VERB
iguana-17148	120	45	population	population	NOUN
iguana-17148	120	46	trends	trend	NOUN
iguana-17148	120	47	and	and	CCONJ
iguana-17148	120	48	identify	identify	VERB
iguana-17148	120	49	threats	threat	NOUN
iguana-17148	120	50	to	to	ADP
iguana-17148	120	51	marine	marine	ADJ
iguana-17148	120	52	iguanas	iguana	NOUN
iguana-17148	120	53	at	at	ADP
iguana-17148	120	54	sites	site	NOUN
iguana-17148	120	55	where	where	SCONJ
iguana-17148	120	56	iguanas	iguana	NOUN
iguana-17148	120	57	show	show	VERB
iguana-17148	120	58	no	no	DET
iguana-17148	120	59	apparent	apparent	ADJ
iguana-17148	120	60	recruitment	recruitment	NOUN
iguana-17148	120	61	.	.	PUNCT
iguana-17148	121	1	those	those	DET
iguana-17148	121	2	sites	site	NOUN
iguana-17148	121	3	include	include	VERB
iguana-17148	121	4	western	western	ADJ
iguana-17148	121	5	isabela	isabela	PROPN
iguana-17148	121	6	and	and	CCONJ
iguana-17148	121	7	san	san	PROPN
iguana-17148	121	8	cristobal	cristobal	PROPN
iguana-17148	121	9	islands	island	NOUN
iguana-17148	121	10	as	as	SCONJ
iguana-17148	121	11	mentioned	mention	VERB
iguana-17148	121	12	above	above	ADV
iguana-17148	121	13	.	.	PUNCT
iguana-17148	122	1	we	we	PRON
iguana-17148	122	2	presently	presently	ADV
iguana-17148	122	3	do	do	AUX
iguana-17148	122	4	not	not	PART
iguana-17148	122	5	know	know	VERB
iguana-17148	122	6	where	where	SCONJ
iguana-17148	122	7	and	and	CCONJ
iguana-17148	122	8	when	when	SCONJ
iguana-17148	122	9	reproduction	reproduction	NOUN
iguana-17148	122	10	occurs	occur	VERB
iguana-17148	122	11	,	,	PUNCT
iguana-17148	122	12	which	which	DET
iguana-17148	122	13	predators	predator	NOUN
iguana-17148	122	14	are	be	AUX
iguana-17148	122	15	preventing	prevent	VERB
iguana-17148	122	16	recruitment	recruitment	NOUN
iguana-17148	122	17	of	of	ADP
iguana-17148	122	18	hatchlings	hatchling	NOUN
iguana-17148	122	19	,	,	PUNCT
iguana-17148	122	20	and	and	CCONJ
iguana-17148	122	21	to	to	ADP
iguana-17148	122	22	what	what	DET
iguana-17148	122	23	degree	degree	NOUN
iguana-17148	122	24	big	big	ADJ
iguana-17148	122	25	mammalian	mammalian	ADJ
iguana-17148	122	26	predators	predator	NOUN
iguana-17148	122	27	are	be	AUX
iguana-17148	122	28	responsible	responsible	ADJ
iguana-17148	122	29	for	for	ADP
iguana-17148	122	30	the	the	DET
iguana-17148	122	31	observed	observe	VERB
iguana-17148	122	32	lack	lack	NOUN
iguana-17148	122	33	of	of	ADP
iguana-17148	122	34	recruitment	recruitment	NOUN
iguana-17148	122	35	.	.	PUNCT
iguana-17148	123	1	the	the	DET
iguana-17148	123	2	most	most	ADV
iguana-17148	123	3	desirable	desirable	ADJ
iguana-17148	123	4	conservation	conservation	NOUN
iguana-17148	123	5	project	project	NOUN
iguana-17148	123	6	would	would	AUX
iguana-17148	123	7	be	be	AUX
iguana-17148	123	8	to	to	PART
iguana-17148	123	9	habituate	habituate	VERB
iguana-17148	123	10	feral	feral	ADJ
iguana-17148	123	11	cats	cat	NOUN
iguana-17148	123	12	to	to	ADP
iguana-17148	123	13	the	the	DET
iguana-17148	123	14	presence	presence	NOUN
iguana-17148	123	15	of	of	ADP
iguana-17148	123	16	a	a	DET
iguana-17148	123	17	researcher	researcher	NOUN
iguana-17148	123	18	and	and	CCONJ
iguana-17148	123	19	follow	follow	VERB
iguana-17148	123	20	them	they	PRON
iguana-17148	123	21	along	along	ADP
iguana-17148	123	22	their	their	PRON
iguana-17148	123	23	daily	daily	ADJ
iguana-17148	123	24	activities	activity	NOUN
iguana-17148	123	25	(	(	PUNCT
iguana-17148	123	26	h.	h.	NOUN
iguana-17148	123	27	snell	snell	PROPN
iguana-17148	123	28	,	,	PUNCT
iguana-17148	123	29	personal	personal	ADJ
iguana-17148	123	30	communication	communication	NOUN
iguana-17148	123	31	)	)	PUNCT
iguana-17148	123	32	.	.	PUNCT
iguana-17148	124	1	such	such	ADJ
iguana-17148	124	2	data	datum	NOUN
iguana-17148	124	3	would	would	AUX
iguana-17148	124	4	reveal	reveal	VERB
iguana-17148	124	5	the	the	DET
iguana-17148	124	6	dangers	danger	NOUN
iguana-17148	124	7	that	that	PRON
iguana-17148	124	8	feral	feral	ADJ
iguana-17148	124	9	cats	cat	NOUN
iguana-17148	124	10	pose	pose	VERB
iguana-17148	124	11	to	to	ADP
iguana-17148	124	12	marine	marine	ADJ
iguana-17148	124	13	iguanas	iguana	NOUN
iguana-17148	124	14	and	and	CCONJ
iguana-17148	124	15	would	would	AUX
iguana-17148	124	16	allow	allow	VERB
iguana-17148	124	17	us	we	PRON
iguana-17148	124	18	to	to	PART
iguana-17148	124	19	study	study	VERB
iguana-17148	124	20	the	the	DET
iguana-17148	124	21	impact	impact	NOUN
iguana-17148	124	22	of	of	ADP
iguana-17148	124	23	these	these	DET
iguana-17148	124	24	fierce	fierce	ADJ
iguana-17148	124	25	predators	predator	NOUN
iguana-17148	124	26	on	on	ADP
iguana-17148	124	27	other	other	ADJ
iguana-17148	124	28	endangered	endanger	VERB
iguana-17148	124	29	coastal	coastal	ADJ
iguana-17148	124	30	species	specie	NOUN
iguana-17148	124	31	.	.	PUNCT
iguana-17148	125	1	dog	dog	NOUN
iguana-17148	125	2	predation	predation	NOUN
iguana-17148	125	3	could	could	AUX
iguana-17148	125	4	be	be	AUX
iguana-17148	125	5	a	a	DET
iguana-17148	125	6	problem	problem	NOUN
iguana-17148	125	7	on	on	ADP
iguana-17148	125	8	other	other	ADJ
iguana-17148	125	9	islands	island	NOUN
iguana-17148	125	10	like	like	ADP
iguana-17148	125	11	san	san	PROPN
iguana-17148	125	12	cristobal	cristobal	PROPN
iguana-17148	125	13	,	,	PUNCT
iguana-17148	125	14	and	and	CCONJ
iguana-17148	125	15	this	this	PRON
iguana-17148	125	16	also	also	ADV
iguana-17148	125	17	needs	need	VERB
iguana-17148	125	18	further	further	ADJ
iguana-17148	125	19	investigation	investigation	NOUN
iguana-17148	125	20	(	(	PUNCT
iguana-17148	125	21	kruuk	kruuk	NOUN
iguana-17148	125	22	and	and	CCONJ
iguana-17148	125	23	snell	snell	NOUN
iguana-17148	125	24	1981	1981	NUM
iguana-17148	125	25	)	)	PUNCT
iguana-17148	125	26	.	.	PUNCT
iguana-17148	126	1	(	(	PUNCT
iguana-17148	126	2	ii	ii	X
iguana-17148	126	3	)	)	PUNCT
iguana-17148	126	4	the	the	DET
iguana-17148	126	5	second	second	ADJ
iguana-17148	126	6	important	important	ADJ
iguana-17148	126	7	project	project	NOUN
iguana-17148	126	8	is	be	AUX
iguana-17148	126	9	to	to	PART
iguana-17148	126	10	study	study	VERB
iguana-17148	126	11	the	the	DET
iguana-17148	126	12	impact	impact	NOUN
iguana-17148	126	13	of	of	ADP
iguana-17148	126	14	human	human	ADJ
iguana-17148	126	15	habitation	habitation	NOUN
iguana-17148	126	16	and	and	CCONJ
iguana-17148	126	17	low	low	ADJ
iguana-17148	126	18	-	-	PUNCT
iguana-17148	126	19	level	level	NOUN
iguana-17148	126	20	environmental	environmental	ADJ
iguana-17148	126	21	contamination	contamination	NOUN
iguana-17148	126	22	on	on	ADP
iguana-17148	126	23	marine	marine	ADJ
iguana-17148	126	24	iguanas	iguana	NOUN
iguana-17148	126	25	.	.	PUNCT
iguana-17148	127	1	we	we	PRON
iguana-17148	127	2	are	be	AUX
iguana-17148	127	3	particularly	particularly	ADV
iguana-17148	127	4	concerned	concerned	ADJ
iguana-17148	127	5	about	about	ADP
iguana-17148	127	6	oil	oil	NOUN
iguana-17148	127	7	residues	residue	NOUN
iguana-17148	127	8	from	from	ADP
iguana-17148	127	9	boats	boat	NOUN
iguana-17148	127	10	around	around	ADP
iguana-17148	127	11	human	human	ADJ
iguana-17148	127	12	settlements	settlement	NOUN
iguana-17148	127	13	(	(	PUNCT
iguana-17148	127	14	e.g.	e.g.	ADV
iguana-17148	127	15	,	,	PUNCT
iguana-17148	127	16	the	the	DET
iguana-17148	127	17	towns	town	NOUN
iguana-17148	127	18	of	of	ADP
iguana-17148	127	19	puerto	puerto	PROPN
iguana-17148	127	20	ayora	ayora	PROPN
iguana-17148	127	21	on	on	ADP
iguana-17148	127	22	santa	santa	PROPN
iguana-17148	127	23	cruz	cruz	PROPN
iguana-17148	127	24	island	island	PROPN
iguana-17148	127	25	and	and	CCONJ
iguana-17148	127	26	villamil	villamil	NOUN
iguana-17148	127	27	on	on	ADP
iguana-17148	127	28	isabela	isabela	PROPN
iguana-17148	127	29	island	island	PROPN
iguana-17148	127	30	)	)	PUNCT
iguana-17148	127	31	.	.	PUNCT
iguana-17148	128	1	we	we	PRON
iguana-17148	128	2	currently	currently	ADV
iguana-17148	128	3	do	do	AUX
iguana-17148	128	4	not	not	PART
iguana-17148	128	5	know	know	VERB
iguana-17148	128	6	if	if	SCONJ
iguana-17148	128	7	areas	area	NOUN
iguana-17148	128	8	around	around	ADP
iguana-17148	128	9	human	human	ADJ
iguana-17148	128	10	settlements	settlement	NOUN
iguana-17148	128	11	should	should	AUX
iguana-17148	128	12	be	be	AUX
iguana-17148	128	13	considered	consider	VERB
iguana-17148	128	14	population	population	NOUN
iguana-17148	128	15	sinks	sink	NOUN
iguana-17148	128	16	for	for	ADP
iguana-17148	128	17	marine	marine	ADJ
iguana-17148	128	18	iguanas	iguana	NOUN
iguana-17148	128	19	or	or	CCONJ
iguana-17148	128	20	whether	whether	SCONJ
iguana-17148	128	21	such	such	ADJ
iguana-17148	128	22	areas	area	NOUN
iguana-17148	128	23	still	still	ADV
iguana-17148	128	24	produce	produce	VERB
iguana-17148	128	25	sufficient	sufficient	ADJ
iguana-17148	128	26	recruits	recruit	NOUN
iguana-17148	128	27	to	to	PART
iguana-17148	128	28	be	be	AUX
iguana-17148	128	29	considered	consider	VERB
iguana-17148	128	30	viable	viable	ADJ
iguana-17148	128	31	populations	population	NOUN
iguana-17148	128	32	.	.	PUNCT
iguana-17148	129	1	based	base	VERB
iguana-17148	129	2	on	on	ADP
iguana-17148	129	3	the	the	DET
iguana-17148	129	4	current	current	ADJ
iguana-17148	129	5	information	information	NOUN
iguana-17148	129	6	about	about	ADP
iguana-17148	129	7	the	the	DET
iguana-17148	129	8	effects	effect	NOUN
iguana-17148	129	9	of	of	ADP
iguana-17148	129	10	a	a	DET
iguana-17148	129	11	low	low	ADJ
iguana-17148	129	12	-	-	PUNCT
iguana-17148	129	13	level	level	NOUN
iguana-17148	129	14	oil	oil	NOUN
iguana-17148	129	15	spill	spill	NOUN
iguana-17148	129	16	in	in	ADP
iguana-17148	129	17	2001	2001	NUM
iguana-17148	129	18	,	,	PUNCT
iguana-17148	129	19	marine	marine	ADJ
iguana-17148	129	20	iguanas	iguana	NOUN
iguana-17148	129	21	in	in	ADP
iguana-17148	129	22	oil	oil	NOUN
iguana-17148	129	23	-	-	PUNCT
iguana-17148	129	24	impacted	impact	VERB
iguana-17148	129	25	areas	area	NOUN
iguana-17148	129	26	conceivably	conceivably	ADV
iguana-17148	129	27	will	will	AUX
iguana-17148	129	28	suffer	suffer	VERB
iguana-17148	129	29	long	long	ADJ
iguana-17148	129	30	-	-	PUNCT
iguana-17148	129	31	term	term	NOUN
iguana-17148	129	32	effects	effect	NOUN
iguana-17148	129	33	.	.	PUNCT
iguana-17148	130	1	furthermore	furthermore	ADV
iguana-17148	130	2	,	,	PUNCT
iguana-17148	130	3	our	our	PRON
iguana-17148	130	4	own	own	ADJ
iguana-17148	130	5	observations	observation	NOUN
iguana-17148	130	6	in	in	ADP
iguana-17148	130	7	and	and	CCONJ
iguana-17148	130	8	around	around	ADP
iguana-17148	130	9	puerto	puerto	PROPN
iguana-17148	130	10	ayora	ayora	PROPN
iguana-17148	130	11	(	(	PUNCT
iguana-17148	130	12	the	the	DET
iguana-17148	130	13	main	main	ADJ
iguana-17148	130	14	harbor	harbor	NOUN
iguana-17148	130	15	town	town	NOUN
iguana-17148	130	16	on	on	ADP
iguana-17148	130	17	santa	santa	PROPN
iguana-17148	130	18	cruz	cruz	PROPN
iguana-17148	130	19	island	island	PROPN
iguana-17148	130	20	)	)	PUNCT
iguana-17148	130	21	suggest	suggest	VERB
iguana-17148	130	22	that	that	SCONJ
iguana-17148	130	23	many	many	ADJ
iguana-17148	130	24	of	of	ADP
iguana-17148	130	25	the	the	DET
iguana-17148	130	26	hatchlings	hatchling	NOUN
iguana-17148	130	27	that	that	PRON
iguana-17148	130	28	pass	pass	VERB
iguana-17148	130	29	through	through	ADP
iguana-17148	130	30	the	the	DET
iguana-17148	130	31	town	town	NOUN
iguana-17148	130	32	after	after	ADP
iguana-17148	130	33	hatching	hatch	VERB
iguana-17148	130	34	disappear	disappear	VERB
iguana-17148	130	35	from	from	ADP
iguana-17148	130	36	the	the	DET
iguana-17148	130	37	harbor	harbor	NOUN
iguana-17148	130	38	area	area	NOUN
iguana-17148	130	39	,	,	PUNCT
iguana-17148	130	40	and	and	CCONJ
iguana-17148	130	41	may	may	AUX
iguana-17148	130	42	die	die	VERB
iguana-17148	130	43	due	due	ADP
iguana-17148	130	44	to	to	ADP
iguana-17148	130	45	human	human	NOUN
iguana-17148	130	46	(	(	PUNCT
iguana-17148	130	47	or	or	CCONJ
iguana-17148	130	48	feral	feral	ADJ
iguana-17148	130	49	animal	animal	NOUN
iguana-17148	130	50	)	)	PUNCT
iguana-17148	130	51	actions	action	NOUN
iguana-17148	130	52	.	.	PUNCT
iguana-17148	131	1	a	a	DET
iguana-17148	131	2	long	long	ADJ
iguana-17148	131	3	-	-	PUNCT
iguana-17148	131	4	term	term	NOUN
iguana-17148	131	5	mark	mark	NOUN
iguana-17148	131	6	/	/	SYM
iguana-17148	131	7	recapture	recapture	NOUN
iguana-17148	131	8	program	program	NOUN
iguana-17148	131	9	could	could	AUX
iguana-17148	131	10	resolve	resolve	VERB
iguana-17148	131	11	such	such	ADJ
iguana-17148	131	12	questions	question	NOUN
iguana-17148	131	13	.	.	PUNCT
iguana-17148	132	1	(	(	PUNCT
iguana-17148	132	2	iii	iii	X
iguana-17148	132	3	)	)	PUNCT
iguana-17148	132	4	a	a	DET
iguana-17148	132	5	third	third	ADJ
iguana-17148	132	6	area	area	NOUN
iguana-17148	132	7	of	of	ADP
iguana-17148	132	8	high	high	ADJ
iguana-17148	132	9	interest	interest	NOUN
iguana-17148	132	10	for	for	ADP
iguana-17148	132	11	the	the	DET
iguana-17148	132	12	management	management	NOUN
iguana-17148	132	13	of	of	ADP
iguana-17148	132	14	marine	marine	ADJ
iguana-17148	132	15	iguanas	iguana	NOUN
iguana-17148	132	16	is	be	AUX
iguana-17148	132	17	the	the	DET
iguana-17148	132	18	planning	planning	NOUN
iguana-17148	132	19	and	and	CCONJ
iguana-17148	132	20	establishment	establishment	NOUN
iguana-17148	132	21	of	of	ADP
iguana-17148	132	22	a	a	DET
iguana-17148	132	23	program	program	NOUN
iguana-17148	132	24	for	for	ADP
iguana-17148	132	25	rehabilitation	rehabilitation	NOUN
iguana-17148	132	26	of	of	ADP
iguana-17148	132	27	marine	marine	ADJ
iguana-17148	132	28	iguanas	iguana	NOUN
iguana-17148	132	29	.	.	PUNCT
iguana-17148	133	1	this	this	PRON
iguana-17148	133	2	would	would	AUX
iguana-17148	133	3	include	include	VERB
iguana-17148	133	4	indiadult	indiadult	ADJ
iguana-17148	133	5	female	female	ADJ
iguana-17148	133	6	marine	marine	ADJ
iguana-17148	133	7	iguana	iguana	NOUN
iguana-17148	133	8	from	from	ADP
iguana-17148	133	9	genovesa	genovesa	PROPN
iguana-17148	133	10	island	island	NOUN
iguana-17148	133	11	;	;	PUNCT
iguana-17148	133	12	this	this	DET
iguana-17148	133	13	single	single	ADJ
iguana-17148	133	14	egg	egg	NOUN
iguana-17148	133	15	was	be	AUX
iguana-17148	133	16	laid	lay	VERB
iguana-17148	133	17	about	about	ADV
iguana-17148	133	18	two	two	NUM
iguana-17148	133	19	hours	hour	NOUN
iguana-17148	133	20	prior	prior	ADV
iguana-17148	133	21	to	to	ADP
iguana-17148	133	22	taking	take	VERB
iguana-17148	133	23	this	this	DET
iguana-17148	133	24	picture	picture	NOUN
iguana-17148	133	25	.	.	PUNCT
iguana-17148	134	1	galápagos	galápago	NOUN
iguana-17148	134	2	hawk	hawk	PROPN
iguana-17148	134	3	killing	kill	VERB
iguana-17148	134	4	a	a	DET
iguana-17148	134	5	hatchling	hatchle	VERB
iguana-17148	134	6	marine	marine	ADJ
iguana-17148	134	7	iguana	iguana	NOUN
iguana-17148	134	8	on	on	ADP
iguana-17148	134	9	santa	santa	PROPN
iguana-17148	134	10	fe	fe	PROPN
iguana-17148	134	11	island	island	PROPN
iguana-17148	134	12	.	.	PUNCT
iguana-17148	135	1	viduals	vidual	NOUN
iguana-17148	135	2	that	that	PRON
iguana-17148	135	3	suffered	suffer	VERB
iguana-17148	135	4	low	low	ADJ
iguana-17148	135	5	levels	level	NOUN
iguana-17148	135	6	of	of	ADP
iguana-17148	135	7	environmental	environmental	ADJ
iguana-17148	135	8	contamination	contamination	NOUN
iguana-17148	135	9	.	.	PUNCT
iguana-17148	136	1	hindgut	hindgut	NOUN
iguana-17148	136	2	microbes	microbe	NOUN
iguana-17148	136	3	probably	probably	ADV
iguana-17148	136	4	die	die	VERB
iguana-17148	136	5	during	during	ADP
iguana-17148	136	6	anthropogenic	anthropogenic	ADJ
iguana-17148	136	7	disasters	disaster	NOUN
iguana-17148	136	8	such	such	ADJ
iguana-17148	136	9	as	as	ADP
iguana-17148	136	10	oil	oil	NOUN
iguana-17148	136	11	spills	spill	NOUN
iguana-17148	136	12	,	,	PUNCT
iguana-17148	136	13	and	and	CCONJ
iguana-17148	136	14	marine	marine	PROPN
iguana-17148	136	15	iguanas	iguana	NOUN
iguana-17148	136	16	subsequently	subsequently	ADV
iguana-17148	136	17	starve	starve	VERB
iguana-17148	136	18	.	.	PUNCT
iguana-17148	137	1	such	such	DET
iguana-17148	137	2	a	a	DET
iguana-17148	137	3	situation	situation	NOUN
iguana-17148	137	4	is	be	AUX
iguana-17148	137	5	comparable	comparable	ADJ
iguana-17148	137	6	to	to	ADP
iguana-17148	137	7	what	what	PRON
iguana-17148	137	8	happens	happen	VERB
iguana-17148	137	9	to	to	ADP
iguana-17148	137	10	other	other	ADJ
iguana-17148	137	11	vertebrates	vertebrate	NOUN
iguana-17148	137	12	when	when	SCONJ
iguana-17148	137	13	endogenous	endogenous	ADJ
iguana-17148	137	14	gut	gut	NOUN
iguana-17148	137	15	symbionts	symbiont	NOUN
iguana-17148	137	16	are	be	AUX
iguana-17148	137	17	eliminated	eliminate	VERB
iguana-17148	137	18	,	,	PUNCT
iguana-17148	137	19	for	for	ADP
iguana-17148	137	20	example	example	NOUN
iguana-17148	137	21	,	,	PUNCT
iguana-17148	137	22	by	by	ADP
iguana-17148	137	23	chemical	chemical	ADJ
iguana-17148	137	24	contamination	contamination	NOUN
iguana-17148	137	25	or	or	CCONJ
iguana-17148	137	26	antibiotic	antibiotic	ADJ
iguana-17148	137	27	medication	medication	NOUN
iguana-17148	137	28	.	.	PUNCT
iguana-17148	138	1	in	in	ADP
iguana-17148	138	2	such	such	ADJ
iguana-17148	138	3	situations	situation	NOUN
iguana-17148	138	4	,	,	PUNCT
iguana-17148	138	5	the	the	DET
iguana-17148	138	6	gut	gut	NOUN
iguana-17148	138	7	fauna	fauna	NOUN
iguana-17148	138	8	can	can	AUX
iguana-17148	138	9	be	be	AUX
iguana-17148	138	10	reinoculated	reinoculate	VERB
iguana-17148	138	11	using	use	VERB
iguana-17148	138	12	pills	pill	NOUN
iguana-17148	138	13	containing	contain	VERB
iguana-17148	138	14	spores	spore	NOUN
iguana-17148	138	15	of	of	ADP
iguana-17148	138	16	endogenous	endogenous	ADJ
iguana-17148	138	17	gut	gut	NOUN
iguana-17148	138	18	symbionts	symbiont	NOUN
iguana-17148	138	19	.	.	PUNCT
iguana-17148	139	1	the	the	DET
iguana-17148	139	2	horizontal	horizontal	ADJ
iguana-17148	139	3	transfer	transfer	NOUN
iguana-17148	139	4	of	of	ADP
iguana-17148	139	5	gut	gut	NOUN
iguana-17148	139	6	symbionts	symbiont	NOUN
iguana-17148	139	7	from	from	ADP
iguana-17148	139	8	unaffected	unaffected	ADJ
iguana-17148	139	9	to	to	ADP
iguana-17148	139	10	affected	affected	ADJ
iguana-17148	139	11	individuals	individual	NOUN
iguana-17148	139	12	also	also	ADV
iguana-17148	139	13	is	be	AUX
iguana-17148	139	14	possible	possible	ADJ
iguana-17148	139	15	in	in	ADP
iguana-17148	139	16	marine	marine	ADJ
iguana-17148	139	17	iguanas	iguana	NOUN
iguana-17148	139	18	.	.	PUNCT
iguana-17148	140	1	thus	thus	ADV
iguana-17148	140	2	,	,	PUNCT
iguana-17148	140	3	once	once	SCONJ
iguana-17148	140	4	we	we	PRON
iguana-17148	140	5	have	have	AUX
iguana-17148	140	6	established	establish	VERB
iguana-17148	140	7	how	how	SCONJ
iguana-17148	140	8	many	many	ADJ
iguana-17148	140	9	and	and	CCONJ
iguana-17148	140	10	what	what	DET
iguana-17148	140	11	kind	kind	NOUN
iguana-17148	140	12	of	of	ADP
iguana-17148	140	13	hindgut	hindgut	NOUN
iguana-17148	140	14	endosymbionts	endosymbiont	NOUN
iguana-17148	140	15	marine	marine	PROPN
iguana-17148	140	16	iguanas	iguana	NOUN
iguana-17148	140	17	possess	possess	VERB
iguana-17148	140	18	,	,	PUNCT
iguana-17148	140	19	and	and	CCONJ
iguana-17148	140	20	how	how	SCONJ
iguana-17148	140	21	such	such	ADJ
iguana-17148	140	22	symbionts	symbiont	NOUN
iguana-17148	140	23	work	work	VERB
iguana-17148	140	24	,	,	PUNCT
iguana-17148	140	25	researchers	researcher	NOUN
iguana-17148	140	26	presumably	presumably	ADV
iguana-17148	140	27	could	could	AUX
iguana-17148	140	28	collect	collect	VERB
iguana-17148	140	29	gut	gut	NOUN
iguana-17148	140	30	material	material	NOUN
iguana-17148	140	31	from	from	ADP
iguana-17148	140	32	healthy	healthy	ADJ
iguana-17148	140	33	individuals	individual	NOUN
iguana-17148	140	34	and	and	CCONJ
iguana-17148	140	35	treat	treat	VERB
iguana-17148	140	36	suffering	suffer	VERB
iguana-17148	140	37	individuals	individual	NOUN
iguana-17148	140	38	.	.	PUNCT
iguana-17148	141	1	(	(	PUNCT
iguana-17148	141	2	iv	iv	X
iguana-17148	141	3	)	)	PUNCT
iguana-17148	141	4	the	the	DET
iguana-17148	141	5	fourth	fourth	ADJ
iguana-17148	141	6	and	and	CCONJ
iguana-17148	141	7	potentially	potentially	ADV
iguana-17148	141	8	most	most	ADV
iguana-17148	141	9	difficult	difficult	ADJ
iguana-17148	141	10	research	research	NOUN
iguana-17148	141	11	activity	activity	NOUN
iguana-17148	141	12	for	for	ADP
iguana-17148	141	13	the	the	DET
iguana-17148	141	14	conservation	conservation	NOUN
iguana-17148	141	15	of	of	ADP
iguana-17148	141	16	marine	marine	ADJ
iguana-17148	141	17	iguanas	iguana	NOUN
iguana-17148	141	18	would	would	AUX
iguana-17148	141	19	be	be	AUX
iguana-17148	141	20	to	to	PART
iguana-17148	141	21	try	try	VERB
iguana-17148	141	22	to	to	PART
iguana-17148	141	23	breed	breed	VERB
iguana-17148	141	24	animals	animal	NOUN
iguana-17148	141	25	in	in	ADP
iguana-17148	141	26	captivity	captivity	NOUN
iguana-17148	141	27	.	.	PUNCT
iguana-17148	142	1	astonishingly	astonishingly	ADV
iguana-17148	142	2	,	,	PUNCT
iguana-17148	142	3	this	this	PRON
iguana-17148	142	4	has	have	AUX
iguana-17148	142	5	never	never	ADV
iguana-17148	142	6	been	be	AUX
iguana-17148	142	7	attempted	attempt	VERB
iguana-17148	142	8	,	,	PUNCT
iguana-17148	142	9	and	and	CCONJ
iguana-17148	142	10	we	we	PRON
iguana-17148	142	11	are	be	AUX
iguana-17148	142	12	unsure	unsure	ADJ
iguana-17148	142	13	whether	whether	SCONJ
iguana-17148	142	14	it	it	PRON
iguana-17148	142	15	is	be	AUX
iguana-17148	142	16	even	even	ADV
iguana-17148	142	17	possible	possible	ADJ
iguana-17148	142	18	.	.	PUNCT
iguana-17148	143	1	the	the	DET
iguana-17148	143	2	challenges	challenge	NOUN
iguana-17148	143	3	for	for	ADP
iguana-17148	143	4	holding	hold	VERB
iguana-17148	143	5	and	and	CCONJ
iguana-17148	143	6	breeding	breed	VERB
iguana-17148	143	7	marine	marine	ADJ
iguana-17148	143	8	iguanas	iguana	NOUN
iguana-17148	143	9	in	in	ADP
iguana-17148	143	10	captivity	captivity	NOUN
iguana-17148	143	11	are	be	AUX
iguana-17148	143	12	manifold	manifold	ADJ
iguana-17148	143	13	.	.	PUNCT
iguana-17148	144	1	first	first	ADV
iguana-17148	144	2	,	,	PUNCT
iguana-17148	144	3	marine	marine	ADJ
iguana-17148	144	4	iguanas	iguana	NOUN
iguana-17148	144	5	have	have	VERB
iguana-17148	144	6	very	very	ADV
iguana-17148	144	7	specific	specific	ADJ
iguana-17148	144	8	dietary	dietary	ADJ
iguana-17148	144	9	requirements	requirement	NOUN
iguana-17148	144	10	.	.	PUNCT
iguana-17148	145	1	only	only	ADV
iguana-17148	145	2	certain	certain	ADJ
iguana-17148	145	3	types	type	NOUN
iguana-17148	145	4	of	of	ADP
iguana-17148	145	5	seaweed	seaweed	NOUN
iguana-17148	145	6	are	be	AUX
iguana-17148	145	7	preferred	prefer	VERB
iguana-17148	145	8	and	and	CCONJ
iguana-17148	145	9	eaten	eat	VERB
iguana-17148	145	10	—	—	PUNCT
iguana-17148	145	11	as	as	ADV
iguana-17148	145	12	soon	soon	ADV
iguana-17148	145	13	as	as	ADP
iguana-17148	145	14	the	the	DET
iguana-17148	145	15	natural	natural	ADJ
iguana-17148	145	16	algae	algae	NOUN
iguana-17148	145	17	composition	composition	NOUN
iguana-17148	145	18	changes	change	NOUN
iguana-17148	145	19	,	,	PUNCT
iguana-17148	145	20	we	we	PRON
iguana-17148	145	21	find	find	VERB
iguana-17148	145	22	dramatic	dramatic	ADJ
iguana-17148	145	23	mortality	mortality	NOUN
iguana-17148	145	24	rates	rate	NOUN
iguana-17148	145	25	in	in	ADP
iguana-17148	145	26	the	the	DET
iguana-17148	145	27	wild	wild	NOUN
iguana-17148	145	28	.	.	PUNCT
iguana-17148	146	1	second	second	ADJ
iguana-17148	146	2	,	,	PUNCT
iguana-17148	146	3	although	although	SCONJ
iguana-17148	146	4	early	early	ADJ
iguana-17148	146	5	naturalists	naturalist	NOUN
iguana-17148	146	6	removed	remove	VERB
iguana-17148	146	7	several	several	ADJ
iguana-17148	146	8	marine	marine	ADJ
iguana-17148	146	9	iguanas	iguana	NOUN
iguana-17148	146	10	from	from	ADP
iguana-17148	146	11	the	the	DET
iguana-17148	146	12	galápagos	galápagos	PROPN
iguana-17148	146	13	islands	island	NOUN
iguana-17148	146	14	and	and	CCONJ
iguana-17148	146	15	brought	bring	VERB
iguana-17148	146	16	them	they	PRON
iguana-17148	146	17	into	into	ADP
iguana-17148	146	18	zoos	zoo	NOUN
iguana-17148	146	19	,	,	PUNCT
iguana-17148	146	20	none	none	NOUN
iguana-17148	146	21	of	of	ADP
iguana-17148	146	22	those	those	DET
iguana-17148	146	23	iguanas	iguana	NOUN
iguana-17148	146	24	ever	ever	ADV
iguana-17148	146	25	tried	try	VERB
iguana-17148	146	26	to	to	PART
iguana-17148	146	27	reproduce	reproduce	VERB
iguana-17148	146	28	.	.	PUNCT
iguana-17148	147	1	nevertheless	nevertheless	ADV
iguana-17148	147	2	,	,	PUNCT
iguana-17148	147	3	several	several	ADJ
iguana-17148	147	4	marine	marine	ADJ
iguana-17148	147	5	iguanas	iguana	NOUN
iguana-17148	147	6	were	be	AUX
iguana-17148	147	7	maintained	maintain	VERB
iguana-17148	147	8	for	for	ADP
iguana-17148	147	9	more	more	ADJ
iguana-17148	147	10	than	than	ADP
iguana-17148	147	11	ten	ten	NUM
iguana-17148	147	12	years	year	NOUN
iguana-17148	147	13	.	.	PUNCT
iguana-17148	148	1	marine	marine	PROPN
iguana-17148	148	2	iguanas	iguana	NOUN
iguana-17148	148	3	have	have	VERB
iguana-17148	148	4	a	a	DET
iguana-17148	148	5	different	different	ADJ
iguana-17148	148	6	breeding	breeding	NOUN
iguana-17148	148	7	system	system	NOUN
iguana-17148	148	8	than	than	ADP
iguana-17148	148	9	most	most	ADJ
iguana-17148	148	10	other	other	ADJ
iguana-17148	148	11	iguanids	iguanid	NOUN
iguana-17148	148	12	,	,	PUNCT
iguana-17148	148	13	and	and	CCONJ
iguana-17148	148	14	a	a	DET
iguana-17148	148	15	fairly	fairly	ADV
iguana-17148	148	16	large	large	ADJ
iguana-17148	148	17	colony	colony	NOUN
iguana-17148	148	18	may	may	AUX
iguana-17148	148	19	be	be	AUX
iguana-17148	148	20	required	require	VERB
iguana-17148	148	21	in	in	ADP
iguana-17148	148	22	order	order	NOUN
iguana-17148	148	23	to	to	PART
iguana-17148	148	24	stimulate	stimulate	VERB
iguana-17148	148	25	mating	mating	NOUN
iguana-17148	148	26	activities	activity	NOUN
iguana-17148	148	27	.	.	PUNCT
iguana-17148	149	1	at	at	ADP
iguana-17148	149	2	the	the	DET
iguana-17148	149	3	same	same	ADJ
iguana-17148	149	4	time	time	NOUN
iguana-17148	149	5	,	,	PUNCT
iguana-17148	149	6	we	we	PRON
iguana-17148	149	7	do	do	AUX
iguana-17148	149	8	not	not	PART
iguana-17148	149	9	know	know	VERB
iguana-17148	149	10	how	how	SCONJ
iguana-17148	149	11	hatchling	hatchling	ADJ
iguana-17148	149	12	marine	marine	ADJ
iguana-17148	149	13	iguanas	iguana	NOUN
iguana-17148	149	14	survive	survive	VERB
iguana-17148	149	15	their	their	PRON
iguana-17148	149	16	first	first	ADJ
iguana-17148	149	17	months	month	NOUN
iguana-17148	149	18	,	,	PUNCT
iguana-17148	149	19	how	how	SCONJ
iguana-17148	149	20	they	they	PRON
iguana-17148	149	21	acquire	acquire	VERB
iguana-17148	149	22	their	their	PRON
iguana-17148	149	23	endogenous	endogenous	ADJ
iguana-17148	149	24	hindgut	hindgut	NOUN
iguana-17148	149	25	microsymbionts	microsymbiont	NOUN
iguana-17148	149	26	,	,	PUNCT
iguana-17148	149	27	and	and	CCONJ
iguana-17148	149	28	how	how	SCONJ
iguana-17148	149	29	they	they	PRON
iguana-17148	149	30	become	become	AUX
iguana-17148	149	31	recruited	recruit	VERB
iguana-17148	149	32	into	into	ADP
iguana-17148	149	33	the	the	DET
iguana-17148	149	34	population	population	NOUN
iguana-17148	149	35	.	.	PUNCT
iguana-17148	150	1	marine	marine	PROPN
iguana-17148	150	2	iguanas	iguana	NOUN
iguana-17148	150	3	are	be	AUX
iguana-17148	150	4	the	the	DET
iguana-17148	150	5	only	only	ADJ
iguana-17148	150	6	galápagos	galápago	NOUN
iguana-17148	150	7	animal	animal	NOUN
iguana-17148	150	8	about	about	ADP
iguana-17148	150	9	which	which	PRON
iguana-17148	150	10	we	we	PRON
iguana-17148	150	11	are	be	AUX
iguana-17148	150	12	uncertain	uncertain	ADJ
iguana-17148	150	13	whether	whether	SCONJ
iguana-17148	150	14	they	they	PRON
iguana-17148	150	15	can	can	AUX
iguana-17148	150	16	be	be	AUX
iguana-17148	150	17	kept	keep	VERB
iguana-17148	150	18	and	and	CCONJ
iguana-17148	150	19	bred	breed	VERB
iguana-17148	150	20	in	in	ADP
iguana-17148	150	21	captivity	captivity	NOUN
iguana-17148	150	22	,	,	PUNCT
iguana-17148	150	23	should	should	AUX
iguana-17148	150	24	such	such	DET
iguana-17148	150	25	a	a	DET
iguana-17148	150	26	need	need	NOUN
iguana-17148	150	27	arise	arise	NOUN
iguana-17148	150	28	.	.	PUNCT
iguana-17148	151	1	we	we	PRON
iguana-17148	151	2	would	would	AUX
iguana-17148	151	3	like	like	VERB
iguana-17148	151	4	to	to	PART
iguana-17148	151	5	remind	remind	VERB
iguana-17148	151	6	the	the	DET
iguana-17148	151	7	reader	reader	NOUN
iguana-17148	151	8	critical	critical	ADJ
iguana-17148	151	9	of	of	ADP
iguana-17148	151	10	any	any	DET
iguana-17148	151	11	effort	effort	NOUN
iguana-17148	151	12	to	to	PART
iguana-17148	151	13	remove	remove	VERB
iguana-17148	151	14	individuals	individual	NOUN
iguana-17148	151	15	from	from	ADP
iguana-17148	151	16	the	the	DET
iguana-17148	151	17	wild	wild	NOUN
iguana-17148	151	18	that	that	SCONJ
iguana-17148	151	19	a	a	DET
iguana-17148	151	20	combination	combination	NOUN
iguana-17148	151	21	of	of	ADP
iguana-17148	151	22	a	a	DET
iguana-17148	151	23	strong	strong	ADJ
iguana-17148	151	24	el	el	PROPN
iguana-17148	151	25	niño	niño	PROPN
iguana-17148	151	26	event	event	NOUN
iguana-17148	151	27	and	and	CCONJ
iguana-17148	151	28	even	even	ADV
iguana-17148	151	29	small	small	ADJ
iguana-17148	151	30	-	-	PUNCT
iguana-17148	151	31	scale	scale	NOUN
iguana-17148	151	32	anthropogenic	anthropogenic	ADJ
iguana-17148	151	33	disasters	disaster	NOUN
iguana-17148	151	34	(	(	PUNCT
iguana-17148	151	35	like	like	ADP
iguana-17148	151	36	a	a	DET
iguana-17148	151	37	lowlevel	lowlevel	ADJ
iguana-17148	151	38	environmental	environmental	ADJ
iguana-17148	151	39	contamination	contamination	NOUN
iguana-17148	151	40	)	)	PUNCT
iguana-17148	151	41	could	could	AUX
iguana-17148	151	42	effectively	effectively	ADV
iguana-17148	151	43	wipe	wipe	VERB
iguana-17148	151	44	out	out	ADP
iguana-17148	151	45	an	an	DET
iguana-17148	151	46	entire	entire	ADJ
iguana-17148	151	47	island	island	NOUN
iguana-17148	151	48	population	population	NOUN
iguana-17148	151	49	.	.	PUNCT
iguana-17148	152	1	if	if	SCONJ
iguana-17148	152	2	our	our	PRON
iguana-17148	152	3	aim	aim	NOUN
iguana-17148	152	4	is	be	AUX
iguana-17148	152	5	to	to	PART
iguana-17148	152	6	conserve	conserve	VERB
iguana-17148	152	7	each	each	DET
iguana-17148	152	8	island	island	NOUN
iguana-17148	152	9	population	population	NOUN
iguana-17148	152	10	of	of	ADP
iguana-17148	152	11	marine	marine	ADJ
iguana-17148	152	12	iguanas	iguana	NOUN
iguana-17148	152	13	because	because	SCONJ
iguana-17148	152	14	of	of	ADP
iguana-17148	152	15	their	their	PRON
iguana-17148	152	16	distinctive	distinctive	ADJ
iguana-17148	152	17	differences	difference	NOUN
iguana-17148	152	18	,	,	PUNCT
iguana-17148	152	19	we	we	PRON
iguana-17148	152	20	should	should	AUX
iguana-17148	152	21	seriously	seriously	ADV
iguana-17148	152	22	consider	consider	VERB
iguana-17148	152	23	developing	develop	VERB
iguana-17148	152	24	the	the	DET
iguana-17148	152	25	husbandry	husbandry	ADJ
iguana-17148	152	26	practices	practice	NOUN
iguana-17148	152	27	necessary	necessary	ADJ
iguana-17148	152	28	to	to	AUX
iguana-17148	152	29	rear	rear	ADJ
iguana-17148	152	30	marine	marine	ADJ
iguana-17148	152	31	iguanas	iguana	NOUN
iguana-17148	152	32	in	in	ADP
iguana-17148	152	33	a	a	DET
iguana-17148	152	34	captive	captive	ADJ
iguana-17148	152	35	setting	setting	NOUN
iguana-17148	152	36	.	.	PUNCT
iguana-17148	153	1	acknowledgments	acknowledgment	NOUN
iguana-17148	153	2	we	we	PRON
iguana-17148	153	3	are	be	AUX
iguana-17148	153	4	grateful	grateful	ADJ
iguana-17148	153	5	to	to	ADP
iguana-17148	153	6	all	all	DET
iguana-17148	153	7	friends	friend	NOUN
iguana-17148	153	8	,	,	PUNCT
iguana-17148	153	9	scientists	scientist	NOUN
iguana-17148	153	10	,	,	PUNCT
iguana-17148	153	11	and	and	CCONJ
iguana-17148	153	12	field	field	NOUN
iguana-17148	153	13	assistants	assistant	NOUN
iguana-17148	153	14	for	for	ADP
iguana-17148	153	15	their	their	PRON
iguana-17148	153	16	help	help	NOUN
iguana-17148	153	17	.	.	PUNCT
iguana-17148	154	1	work	work	VERB
iguana-17148	154	2	in	in	ADP
iguana-17148	154	3	the	the	DET
iguana-17148	154	4	galápagos	galápago	NOUN
iguana-17148	154	5	is	be	AUX
iguana-17148	154	6	supported	support	VERB
iguana-17148	154	7	by	by	ADP
iguana-17148	154	8	the	the	DET
iguana-17148	154	9	galápagos	galápagos	PROPN
iguana-17148	154	10	national	national	PROPN
iguana-17148	154	11	park	park	PROPN
iguana-17148	154	12	service	service	NOUN
iguana-17148	154	13	and	and	CCONJ
iguana-17148	154	14	the	the	DET
iguana-17148	154	15	charles	charles	PROPN
iguana-17148	154	16	darwin	darwin	PROPN
iguana-17148	154	17	research	research	PROPN
iguana-17148	154	18	station	station	NOUN
iguana-17148	154	19	.	.	PUNCT
iguana-17148	155	1	this	this	PRON
iguana-17148	155	2	is	be	AUX
iguana-17148	155	3	contribution	contribution	NOUN
iguana-17148	155	4	#	#	SYM
iguana-17148	155	5	392	392	NUM
iguana-17148	155	6	to	to	ADP
iguana-17148	155	7	the	the	DET
iguana-17148	155	8	charles	charles	PROPN
iguana-17148	155	9	darwin	darwin	PROPN
iguana-17148	155	10	foundation	foundation	PROPN
iguana-17148	155	11	.	.	PUNCT
iguana-17148	156	1	this	this	DET
iguana-17148	156	2	work	work	NOUN
iguana-17148	156	3	was	be	AUX
iguana-17148	156	4	supported	support	VERB
iguana-17148	156	5	by	by	ADP
iguana-17148	156	6	the	the	DET
iguana-17148	156	7	national	national	PROPN
iguana-17148	156	8	science	science	PROPN
iguana-17148	156	9	foundation	foundation	PROPN
iguana-17148	156	10	under	under	ADP
iguana-17148	156	11	grant	grant	PROPN
iguana-17148	156	12	ibn-0118069	ibn-0118069	NOUN
iguana-17148	156	13	.	.	PUNCT
iguana-17148	157	1	references	reference	NOUN
iguana-17148	157	2	boersma	boersma	PROPN
iguana-17148	157	3	,	,	PUNCT
iguana-17148	157	4	p.	p.	PROPN
iguana-17148	157	5	d.	d.	PROPN
iguana-17148	157	6	1983	1983	NUM
iguana-17148	157	7	.	.	PUNCT
iguana-17148	158	1	an	an	DET
iguana-17148	158	2	ecological	ecological	ADJ
iguana-17148	158	3	study	study	NOUN
iguana-17148	158	4	of	of	ADP
iguana-17148	158	5	the	the	DET
iguana-17148	158	6	galapagos	galapagos	PROPN
iguana-17148	158	7	marine	marine	PROPN
iguana-17148	158	8	iguana	iguana	PROPN
iguana-17148	158	9	,	,	PUNCT
iguana-17148	158	10	pp	pp	ADV
iguana-17148	158	11	.	.	PUNCT
iguana-17148	159	1	157–176	157–176	NUM
iguana-17148	159	2	.	.	PUNCT
iguana-17148	160	1	in	in	ADP
iguana-17148	160	2	:	:	PUNCT
iguana-17148	160	3	r.	r.	PROPN
iguana-17148	160	4	i.	i.	PROPN
iguana-17148	160	5	bowman	bowman	PROPN
iguana-17148	160	6	,	,	PUNCT
iguana-17148	160	7	m.	m.	NOUN
iguana-17148	160	8	berson	berson	PROPN
iguana-17148	160	9	,	,	PUNCT
iguana-17148	160	10	and	and	CCONJ
iguana-17148	160	11	a.	a.	PROPN
iguana-17148	160	12	e.	e.	PROPN
iguana-17148	160	13	leviton	leviton	PROPN
iguana-17148	160	14	(	(	PUNCT
iguana-17148	160	15	eds	eds	PROPN
iguana-17148	160	16	.	.	PUNCT
iguana-17148	160	17	)	)	PUNCT
iguana-17148	160	18	,	,	PUNCT
iguana-17148	160	19	patterns	pattern	NOUN
iguana-17148	160	20	of	of	ADP
iguana-17148	160	21	evolution	evolution	NOUN
iguana-17148	160	22	in	in	ADP
iguana-17148	160	23	galapagos	galapagos	PROPN
iguana-17148	160	24	organisms	organism	NOUN
iguana-17148	160	25	.	.	PUNCT
iguana-17148	161	1	american	american	PROPN
iguana-17148	161	2	association	association	PROPN
iguana-17148	161	3	for	for	ADP
iguana-17148	161	4	the	the	DET
iguana-17148	161	5	advancement	advancement	NOUN
iguana-17148	161	6	of	of	ADP
iguana-17148	161	7	science	science	NOUN
iguana-17148	161	8	,	,	PUNCT
iguana-17148	161	9	san	san	PROPN
iguana-17148	161	10	francisco	francisco	PROPN
iguana-17148	161	11	,	,	PUNCT
iguana-17148	161	12	california	california	PROPN
iguana-17148	161	13	,	,	PUNCT
iguana-17148	161	14	usa	usa	PROPN
iguana-17148	161	15	.	.	PROPN
iguana-17148	161	16	buttemer	buttemer	PROPN
iguana-17148	161	17	,	,	PUNCT
iguana-17148	161	18	w.	w.	PROPN
iguana-17148	161	19	a.	a.	PROPN
iguana-17148	161	20	and	and	CCONJ
iguana-17148	161	21	w.	w.	PROPN
iguana-17148	161	22	r.	r.	PROPN
iguana-17148	161	23	dawson	dawson	PROPN
iguana-17148	161	24	.	.	PROPN
iguana-17148	161	25	1993	1993	NUM
iguana-17148	161	26	.	.	PUNCT
iguana-17148	162	1	temporal	temporal	ADJ
iguana-17148	162	2	pattern	pattern	NOUN
iguana-17148	162	3	of	of	ADP
iguana-17148	162	4	foraging	forage	VERB
iguana-17148	162	5	and	and	CCONJ
iguana-17148	162	6	microhabitat	microhabitat	NOUN
iguana-17148	162	7	use	use	NOUN
iguana-17148	162	8	by	by	ADP
iguana-17148	162	9	galápagos	galápago	NOUN
iguana-17148	162	10	marine	marine	PROPN
iguana-17148	162	11	iguanas	iguana	NOUN
iguana-17148	162	12	,	,	PUNCT
iguana-17148	162	13	amblyrhynchus	amblyrhynchus	NOUN
iguana-17148	162	14	cristatus	cristatus	PROPN
iguana-17148	162	15	.	.	PUNCT
iguana-17148	163	1	oecologia	oecologia	PROPN
iguana-17148	163	2	96:56–64	96:56–64	PROPN
iguana-17148	163	3	.	.	PUNCT
iguana-17148	163	4	carpenter	carpenter	PROPN
iguana-17148	163	5	,	,	PUNCT
iguana-17148	163	6	c.	c.	PROPN
iguana-17148	163	7	1966	1966	NUM
iguana-17148	163	8	.	.	PUNCT
iguana-17148	164	1	the	the	DET
iguana-17148	164	2	marine	marine	ADJ
iguana-17148	164	3	iguana	iguana	NOUN
iguana-17148	164	4	of	of	ADP
iguana-17148	164	5	the	the	DET
iguana-17148	164	6	galápagos	galápagos	PROPN
iguana-17148	164	7	islands	island	NOUN
iguana-17148	164	8	,	,	PUNCT
iguana-17148	164	9	its	its	PRON
iguana-17148	164	10	behavior	behavior	NOUN
iguana-17148	164	11	and	and	CCONJ
iguana-17148	164	12	ecology	ecology	NOUN
iguana-17148	164	13	.	.	PUNCT
iguana-17148	165	1	proceedings	proceeding	NOUN
iguana-17148	165	2	of	of	ADP
iguana-17148	165	3	the	the	DET
iguana-17148	165	4	california	california	PROPN
iguana-17148	165	5	academy	academy	PROPN
iguana-17148	165	6	of	of	ADP
iguana-17148	165	7	sciences	sciences	PROPN
iguana-17148	165	8	,	,	PUNCT
iguana-17148	165	9	fourth	fourth	ADJ
iguana-17148	165	10	series	series	NOUN
iguana-17148	165	11	34:329–376	34:329–376	PROPN
iguana-17148	165	12	.	.	PUNCT
iguana-17148	166	1	cayot	cayot	PROPN
iguana-17148	166	2	,	,	PUNCT
iguana-17148	166	3	l.	l.	PROPN
iguana-17148	166	4	j.	j.	PROPN
iguana-17148	166	5	,	,	PUNCT
iguana-17148	166	6	k.	k.	PROPN
iguana-17148	166	7	rassmann	rassmann	PROPN
iguana-17148	166	8	,	,	PUNCT
iguana-17148	166	9	and	and	CCONJ
iguana-17148	166	10	f.	f.	PROPN
iguana-17148	166	11	trillmich	trillmich	PROPN
iguana-17148	166	12	.	.	PUNCT
iguana-17148	167	1	1994	1994	NUM
iguana-17148	167	2	.	.	PUNCT
iguana-17148	168	1	are	be	AUX
iguana-17148	168	2	marine	marine	PROPN
iguana-17148	168	3	iguanas	iguana	NOUN
iguana-17148	168	4	endangered	endanger	VERB
iguana-17148	168	5	on	on	ADP
iguana-17148	168	6	islands	island	NOUN
iguana-17148	168	7	with	with	ADP
iguana-17148	168	8	introduced	introduce	VERB
iguana-17148	168	9	predators	predator	NOUN
iguana-17148	168	10	?	?	PUNCT
iguana-17148	169	1	noticias	noticias	PROPN
iguana-17148	169	2	de	de	PROPN
iguana-17148	169	3	galapagos	galapagos	PROPN
iguana-17148	169	4	53:13–15	53:13–15	PROPN
iguana-17148	169	5	.	.	PUNCT
iguana-17148	169	6	darwin	darwin	PROPN
iguana-17148	169	7	,	,	PUNCT
iguana-17148	169	8	c.	c.	PROPN
iguana-17148	169	9	1883	1883	NUM
iguana-17148	169	10	.	.	PUNCT
iguana-17148	170	1	journal	journal	NOUN
iguana-17148	170	2	of	of	ADP
iguana-17148	170	3	researches	research	NOUN
iguana-17148	170	4	.	.	PUNCT
iguana-17148	171	1	new	new	ADJ
iguana-17148	171	2	edition	edition	PROPN
iguana-17148	171	3	.	.	PUNCT
iguana-17148	172	1	appleton	appleton	PROPN
iguana-17148	172	2	,	,	PUNCT
iguana-17148	172	3	new	new	PROPN
iguana-17148	172	4	york	york	PROPN
iguana-17148	172	5	.	.	PUNCT
iguana-17148	173	1	196	196	NUM
iguana-17148	173	2	iguana	iguana	NOUN
iguana-17148	173	3	•	•	NUM
iguana-17148	173	4	volume	volume	NOUN
iguana-17148	173	5	11	11	NUM
iguana-17148	173	6	,	,	PUNCT
iguana-17148	173	7	number	number	NOUN
iguana-17148	173	8	4	4	NUM
iguana-17148	173	9	•	•	NOUN
iguana-17148	173	10	december	december	PROPN
iguana-17148	173	11	2004	2004	NUM
iguana-17148	173	12	wikelski	wikelski	NOUN
iguana-17148	173	13	and	and	CCONJ
iguana-17148	173	14	nelson	nelson	PROPN
iguana-17148	173	15	bachelor	bachelor	PROPN
iguana-17148	173	16	male	male	ADJ
iguana-17148	173	17	marine	marine	ADJ
iguana-17148	173	18	iguanas	iguana	NOUN
iguana-17148	173	19	from	from	ADP
iguana-17148	173	20	fernandina	fernandina	PROPN
iguana-17148	173	21	island	island	PROPN
iguana-17148	173	22	.	.	PUNCT
iguana-17148	174	1	note	note	VERB
iguana-17148	174	2	the	the	DET
iguana-17148	174	3	elaborate	elaborate	ADJ
iguana-17148	174	4	spines	spine	NOUN
iguana-17148	174	5	that	that	PRON
iguana-17148	174	6	are	be	AUX
iguana-17148	174	7	very	very	ADV
iguana-17148	174	8	prominent	prominent	ADJ
iguana-17148	174	9	in	in	ADP
iguana-17148	174	10	this	this	DET
iguana-17148	174	11	population	population	NOUN
iguana-17148	174	12	of	of	ADP
iguana-17148	174	13	large	large	ADJ
iguana-17148	174	14	individuals	individual	NOUN
iguana-17148	174	15	.	.	PUNCT
iguana-17148	175	1	on	on	ADP
iguana-17148	175	2	fernandina	fernandina	PROPN
iguana-17148	175	3	,	,	PUNCT
iguana-17148	175	4	diving	diving	NOUN
iguana-17148	175	5	males	male	NOUN
iguana-17148	175	6	usually	usually	ADV
iguana-17148	175	7	exceed	exceed	VERB
iguana-17148	175	8	3500	3500	NUM
iguana-17148	175	9	g.	g.	NOUN
iguana-17148	175	10	comparing	compare	VERB
iguana-17148	175	11	two	two	NUM
iguana-17148	175	12	marine	marine	ADJ
iguana-17148	175	13	iguanas	iguana	NOUN
iguana-17148	175	14	on	on	ADP
iguana-17148	175	15	santa	santa	PROPN
iguana-17148	175	16	fe	fe	PROPN
iguana-17148	175	17	island	island	NOUN
iguana-17148	175	18	showing	show	VERB
iguana-17148	175	19	how	how	SCONJ
iguana-17148	175	20	much	much	ADJ
iguana-17148	175	21	individual	individual	ADJ
iguana-17148	175	22	iguanas	iguana	NOUN
iguana-17148	175	23	can	can	AUX
iguana-17148	175	24	shrink	shrink	VERB
iguana-17148	175	25	.	.	PUNCT
iguana-17148	176	1	the	the	DET
iguana-17148	176	2	iguana	iguana	NOUN
iguana-17148	176	3	on	on	ADP
iguana-17148	176	4	the	the	DET
iguana-17148	176	5	left	left	NOUN
iguana-17148	176	6	could	could	AUX
iguana-17148	176	7	shrink	shrink	VERB
iguana-17148	176	8	in	in	ADP
iguana-17148	176	9	body	body	NOUN
iguana-17148	176	10	length	length	NOUN
iguana-17148	176	11	to	to	ADP
iguana-17148	176	12	the	the	DET
iguana-17148	176	13	size	size	NOUN
iguana-17148	176	14	of	of	ADP
iguana-17148	176	15	the	the	DET
iguana-17148	176	16	individual	individual	NOUN
iguana-17148	176	17	on	on	ADP
iguana-17148	176	18	the	the	DET
iguana-17148	176	19	right	right	NOUN
iguana-17148	176	20	during	during	ADP
iguana-17148	176	21	an	an	DET
iguana-17148	176	22	el	el	PROPN
iguana-17148	176	23	niño	niño	PROPN
iguana-17148	176	24	.	.	PUNCT
iguana-17148	177	1	drent	drent	PROPN
iguana-17148	177	2	,	,	PUNCT
iguana-17148	177	3	j.	j.	PROPN
iguana-17148	177	4	,	,	PUNCT
iguana-17148	177	5	w.	w.	PROPN
iguana-17148	177	6	van	van	PROPN
iguana-17148	177	7	marken	marken	PROPN
iguana-17148	177	8	lichtenbelt	lichtenbelt	PROPN
iguana-17148	177	9	,	,	PUNCT
iguana-17148	177	10	and	and	CCONJ
iguana-17148	177	11	m.	m.	NOUN
iguana-17148	177	12	wikelski	wikelski	PROPN
iguana-17148	177	13	.	.	PUNCT
iguana-17148	178	1	1999	1999	NUM
iguana-17148	178	2	.	.	PUNCT
iguana-17148	179	1	foraging	forage	VERB
iguana-17148	179	2	mode	mode	NOUN
iguana-17148	179	3	and	and	CCONJ
iguana-17148	179	4	seasonality	seasonality	NOUN
iguana-17148	179	5	effects	effect	NOUN
iguana-17148	179	6	on	on	ADP
iguana-17148	179	7	the	the	DET
iguana-17148	179	8	energetics	energetic	NOUN
iguana-17148	179	9	of	of	ADP
iguana-17148	179	10	the	the	DET
iguana-17148	179	11	marine	marine	ADJ
iguana-17148	179	12	iguana	iguana	NOUN
iguana-17148	179	13	amblyrhynchus	amblyrhynchus	NOUN
iguana-17148	179	14	cristatus	cristatus	PROPN
iguana-17148	179	15	.	.	PUNCT
iguana-17148	180	1	functional	functional	ADJ
iguana-17148	180	2	ecology	ecology	NOUN
iguana-17148	180	3	13:493–499	13:493–499	PROPN
iguana-17148	180	4	.	.	PUNCT
iguana-17148	181	1	eibl	eibl	NOUN
iguana-17148	181	2	-	-	PUNCT
iguana-17148	181	3	eibesfeldt	eibesfeldt	PROPN
iguana-17148	181	4	,	,	PUNCT
iguana-17148	181	5	i.	i.	PROPN
iguana-17148	181	6	1962	1962	NUM
iguana-17148	181	7	.	.	PUNCT
iguana-17148	182	1	neue	neue	PROPN
iguana-17148	182	2	unterarten	unterarten	NOUN
iguana-17148	182	3	der	der	PROPN
iguana-17148	182	4	meerechse	meerechse	NOUN
iguana-17148	182	5	,	,	PUNCT
iguana-17148	182	6	amblyrhynchus	amblyrhynchus	NOUN
iguana-17148	182	7	cristatus	cristatus	PROPN
iguana-17148	182	8	,	,	PUNCT
iguana-17148	182	9	nebst	nebst	PROPN
iguana-17148	182	10	weiteren	weiteren	PROPN
iguana-17148	182	11	angaben	angaben	PROPN
iguana-17148	182	12	zur	zur	PROPN
iguana-17148	182	13	biologie	biologie	PROPN
iguana-17148	182	14	der	der	PROPN
iguana-17148	182	15	art	art	NOUN
iguana-17148	182	16	.	.	PUNCT
iguana-17148	183	1	senckenbergia	senckenbergia	PROPN
iguana-17148	183	2	biologica	biologica	PROPN
iguana-17148	183	3	43:177–199	43:177–199	PROPN
iguana-17148	183	4	.	.	PUNCT
iguana-17148	184	1	kruuk	kruuk	PROPN
iguana-17148	184	2	,	,	PUNCT
iguana-17148	184	3	h.	h.	PROPN
iguana-17148	184	4	and	and	CCONJ
iguana-17148	184	5	h.	h.	PROPN
iguana-17148	184	6	snell	snell	PROPN
iguana-17148	184	7	.	.	PUNCT
iguana-17148	185	1	1981	1981	NUM
iguana-17148	185	2	.	.	PUNCT
iguana-17148	186	1	prey	prey	PROPN
iguana-17148	186	2	selection	selection	NOUN
iguana-17148	186	3	by	by	ADP
iguana-17148	186	4	feral	feral	ADJ
iguana-17148	186	5	dogs	dog	NOUN
iguana-17148	186	6	from	from	ADP
iguana-17148	186	7	a	a	DET
iguana-17148	186	8	population	population	NOUN
iguana-17148	186	9	of	of	ADP
iguana-17148	186	10	marine	marine	ADJ
iguana-17148	186	11	iguanas	iguana	NOUN
iguana-17148	186	12	(	(	PUNCT
iguana-17148	186	13	amblyrhynchus	amblyrhynchus	ADV
iguana-17148	186	14	cristatus	cristatus	PROPN
iguana-17148	186	15	)	)	PUNCT
iguana-17148	186	16	.	.	PUNCT
iguana-17148	187	1	journal	journal	PROPN
iguana-17148	187	2	of	of	ADP
iguana-17148	187	3	applied	applied	ADJ
iguana-17148	187	4	ecology	ecology	NOUN
iguana-17148	187	5	18:197–204	18:197–204	PROPN
iguana-17148	187	6	.	.	PUNCT
iguana-17148	188	1	laurie	laurie	PROPN
iguana-17148	188	2	,	,	PUNCT
iguana-17148	188	3	w.	w.	PROPN
iguana-17148	188	4	1990	1990	NUM
iguana-17148	188	5	.	.	PUNCT
iguana-17148	189	1	population	population	NOUN
iguana-17148	189	2	biology	biology	NOUN
iguana-17148	189	3	of	of	ADP
iguana-17148	189	4	marine	marine	ADJ
iguana-17148	189	5	iguanas	iguana	NOUN
iguana-17148	189	6	(	(	PUNCT
iguana-17148	189	7	amblyrhynchus	amblyrhynchus	ADV
iguana-17148	189	8	cristatus	cristatus	PROPN
iguana-17148	189	9	)	)	PUNCT
iguana-17148	189	10	.	.	PUNCT
iguana-17148	190	1	i.	i.	NOUN
iguana-17148	190	2	changes	change	NOUN
iguana-17148	190	3	in	in	ADP
iguana-17148	190	4	fecundity	fecundity	NOUN
iguana-17148	190	5	related	relate	VERB
iguana-17148	190	6	to	to	ADP
iguana-17148	190	7	a	a	DET
iguana-17148	190	8	population	population	NOUN
iguana-17148	190	9	crash	crash	NOUN
iguana-17148	190	10	.	.	PUNCT
iguana-17148	191	1	journal	journal	NOUN
iguana-17148	191	2	of	of	ADP
iguana-17148	191	3	animal	animal	NOUN
iguana-17148	191	4	ecology	ecology	NOUN
iguana-17148	191	5	59:515–528	59:515–528	NUM
iguana-17148	191	6	.	.	PUNCT
iguana-17148	192	1	laurie	laurie	PROPN
iguana-17148	192	2	,	,	PUNCT
iguana-17148	192	3	w.	w.	PROPN
iguana-17148	192	4	and	and	CCONJ
iguana-17148	192	5	d.	d.	PROPN
iguana-17148	192	6	brown	brown	PROPN
iguana-17148	192	7	.	.	PUNCT
iguana-17148	193	1	1990a	1990a	NOUN
iguana-17148	193	2	.	.	PUNCT
iguana-17148	194	1	population	population	NOUN
iguana-17148	194	2	biology	biology	NOUN
iguana-17148	194	3	of	of	ADP
iguana-17148	194	4	marine	marine	ADJ
iguana-17148	194	5	iguanas	iguana	NOUN
iguana-17148	194	6	(	(	PUNCT
iguana-17148	194	7	amblyrhynchus	amblyrhynchus	ADV
iguana-17148	194	8	cristatus	cristatus	PROPN
iguana-17148	194	9	)	)	PUNCT
iguana-17148	194	10	.	.	PUNCT
iguana-17148	195	1	ii	ii	PROPN
iguana-17148	195	2	.	.	PUNCT
iguana-17148	196	1	changes	change	NOUN
iguana-17148	196	2	in	in	ADP
iguana-17148	196	3	annual	annual	ADJ
iguana-17148	196	4	survival	survival	NOUN
iguana-17148	196	5	rates	rate	NOUN
iguana-17148	196	6	and	and	CCONJ
iguana-17148	196	7	the	the	DET
iguana-17148	196	8	effects	effect	NOUN
iguana-17148	196	9	of	of	ADP
iguana-17148	196	10	size	size	NOUN
iguana-17148	196	11	,	,	PUNCT
iguana-17148	196	12	sex	sex	NOUN
iguana-17148	196	13	,	,	PUNCT
iguana-17148	196	14	age	age	NOUN
iguana-17148	196	15	,	,	PUNCT
iguana-17148	196	16	and	and	CCONJ
iguana-17148	196	17	fecundity	fecundity	NOUN
iguana-17148	196	18	in	in	ADP
iguana-17148	196	19	a	a	DET
iguana-17148	196	20	population	population	NOUN
iguana-17148	196	21	crash	crash	NOUN
iguana-17148	196	22	.	.	PUNCT
iguana-17148	197	1	journal	journal	NOUN
iguana-17148	197	2	of	of	ADP
iguana-17148	197	3	animal	animal	NOUN
iguana-17148	197	4	ecology	ecology	NOUN
iguana-17148	197	5	59:529–544	59:529–544	PROPN
iguana-17148	197	6	.	.	PUNCT
iguana-17148	198	1	laurie	laurie	PROPN
iguana-17148	198	2	,	,	PUNCT
iguana-17148	198	3	w.	w.	PROPN
iguana-17148	198	4	and	and	CCONJ
iguana-17148	198	5	d.	d.	PROPN
iguana-17148	198	6	brown	brown	PROPN
iguana-17148	198	7	.	.	PUNCT
iguana-17148	199	1	1990b	1990b	NUM
iguana-17148	199	2	.	.	PUNCT
iguana-17148	200	1	population	population	NOUN
iguana-17148	200	2	biology	biology	NOUN
iguana-17148	200	3	of	of	ADP
iguana-17148	200	4	marine	marine	ADJ
iguana-17148	200	5	iguanas	iguana	NOUN
iguana-17148	200	6	(	(	PUNCT
iguana-17148	200	7	amblyrhynchus	amblyrhynchus	ADV
iguana-17148	200	8	cristatus	cristatus	PROPN
iguana-17148	200	9	)	)	PUNCT
iguana-17148	200	10	.	.	PUNCT
iguana-17148	201	1	iii	iii	X
iguana-17148	201	2	.	.	PUNCT
iguana-17148	201	3	factors	factor	NOUN
iguana-17148	201	4	affecting	affect	VERB
iguana-17148	201	5	survival	survival	NOUN
iguana-17148	201	6	.	.	PUNCT
iguana-17148	202	1	journal	journal	NOUN
iguana-17148	202	2	of	of	ADP
iguana-17148	202	3	animal	animal	NOUN
iguana-17148	202	4	ecology	ecology	NOUN
iguana-17148	202	5	59:545–568	59:545–568	PROPN
iguana-17148	202	6	.	.	PUNCT
iguana-17148	202	7	mackie	mackie	PROPN
iguana-17148	202	8	,	,	PUNCT
iguana-17148	202	9	r.	r.	PROPN
iguana-17148	202	10	i.	i.	PROPN
iguana-17148	202	11	,	,	PUNCT
iguana-17148	202	12	m.	m.	PROPN
iguana-17148	202	13	rycyk	rycyk	PROPN
iguana-17148	202	14	,	,	PUNCT
iguana-17148	202	15	r.	r.	PROPN
iguana-17148	202	16	l.	l.	PROPN
iguana-17148	202	17	ruemmler	ruemmler	PROPN
iguana-17148	202	18	,	,	PUNCT
iguana-17148	202	19	r.	r.	PROPN
iguana-17148	202	20	i.	i.	PROPN
iguana-17148	202	21	aminov	aminov	PROPN
iguana-17148	202	22	,	,	PUNCT
iguana-17148	202	23	and	and	CCONJ
iguana-17148	202	24	m.	m.	NOUN
iguana-17148	202	25	wikelski	wikelski	NOUN
iguana-17148	202	26	.	.	PUNCT
iguana-17148	203	1	2004	2004	NUM
iguana-17148	203	2	.	.	PUNCT
iguana-17148	204	1	biochemical	biochemical	ADJ
iguana-17148	204	2	and	and	CCONJ
iguana-17148	204	3	microbiological	microbiological	ADJ
iguana-17148	204	4	evidence	evidence	NOUN
iguana-17148	204	5	for	for	ADP
iguana-17148	204	6	fermentative	fermentative	ADJ
iguana-17148	204	7	digestion	digestion	NOUN
iguana-17148	204	8	in	in	ADP
iguana-17148	204	9	free	free	ADJ
iguana-17148	204	10	-	-	PUNCT
iguana-17148	204	11	living	live	VERB
iguana-17148	204	12	land	land	NOUN
iguana-17148	204	13	iguanas	iguana	NOUN
iguana-17148	204	14	(	(	PUNCT
iguana-17148	204	15	conolophus	conolophus	NOUN
iguana-17148	204	16	pallidus	pallidus	NOUN
iguana-17148	204	17	)	)	PUNCT
iguana-17148	204	18	and	and	CCONJ
iguana-17148	204	19	marine	marine	ADJ
iguana-17148	204	20	iguanas	iguana	NOUN
iguana-17148	204	21	(	(	PUNCT
iguana-17148	204	22	amblyrhynchus	amblyrhynchus	NOUN
iguana-17148	204	23	cristatus	cristatus	PROPN
iguana-17148	204	24	)	)	PUNCT
iguana-17148	204	25	on	on	ADP
iguana-17148	204	26	the	the	DET
iguana-17148	204	27	galapagos	galapagos	PROPN
iguana-17148	204	28	archipelago	archipelago	PROPN
iguana-17148	204	29	.	.	PUNCT
iguana-17148	205	1	physiological	physiological	ADJ
iguana-17148	205	2	and	and	CCONJ
iguana-17148	205	3	biochemical	biochemical	ADJ
iguana-17148	205	4	zoology	zoology	NOUN
iguana-17148	205	5	77:127–138	77:127–138	PROPN
iguana-17148	205	6	.	.	PUNCT
iguana-17148	206	1	rassmann	rassmann	PROPN
iguana-17148	206	2	,	,	PUNCT
iguana-17148	206	3	k.	k.	PROPN
iguana-17148	206	4	1997	1997	NUM
iguana-17148	206	5	.	.	PUNCT
iguana-17148	207	1	evolutionary	evolutionary	ADJ
iguana-17148	207	2	age	age	NOUN
iguana-17148	207	3	of	of	ADP
iguana-17148	207	4	the	the	DET
iguana-17148	207	5	galapagos	galapagos	PROPN
iguana-17148	207	6	iguanas	iguanas	PROPN
iguana-17148	207	7	predates	predate	VERB
iguana-17148	207	8	the	the	DET
iguana-17148	207	9	age	age	NOUN
iguana-17148	207	10	of	of	ADP
iguana-17148	207	11	the	the	DET
iguana-17148	207	12	present	present	ADJ
iguana-17148	207	13	galapagos	galapagos	PROPN
iguana-17148	207	14	islands	island	NOUN
iguana-17148	207	15	.	.	PUNCT
iguana-17148	208	1	molecular	molecular	ADJ
iguana-17148	208	2	phylogenetics	phylogenetic	NOUN
iguana-17148	208	3	and	and	CCONJ
iguana-17148	208	4	evolution	evolution	NOUN
iguana-17148	208	5	7:158–172	7:158–172	PROPN
iguana-17148	208	6	.	.	PUNCT
iguana-17148	209	1	rassmann	rassmann	PROPN
iguana-17148	209	2	,	,	PUNCT
iguana-17148	209	3	k.	k.	PROPN
iguana-17148	209	4	,	,	PUNCT
iguana-17148	209	5	d.	d.	PROPN
iguana-17148	209	6	tautz	tautz	PROPN
iguana-17148	209	7	,	,	PUNCT
iguana-17148	209	8	f.	f.	PROPN
iguana-17148	209	9	trillmich	trillmich	PROPN
iguana-17148	209	10	,	,	PUNCT
iguana-17148	209	11	and	and	CCONJ
iguana-17148	209	12	c.	c.	PROPN
iguana-17148	209	13	gliddon	gliddon	NOUN
iguana-17148	209	14	.	.	PUNCT
iguana-17148	210	1	1997	1997	NUM
iguana-17148	210	2	.	.	PUNCT
iguana-17148	211	1	the	the	DET
iguana-17148	211	2	microevolution	microevolution	NOUN
iguana-17148	211	3	of	of	ADP
iguana-17148	211	4	the	the	DET
iguana-17148	211	5	galapagos	galapagos	PROPN
iguana-17148	211	6	marine	marine	PROPN
iguana-17148	211	7	iguana	iguana	NOUN
iguana-17148	211	8	amblyrhynchus	amblyrhynchus	PROPN
iguana-17148	211	9	cristatus	cristatus	PROPN
iguana-17148	211	10	assessed	assess	VERB
iguana-17148	211	11	by	by	ADP
iguana-17148	211	12	nuclear	nuclear	ADJ
iguana-17148	211	13	and	and	CCONJ
iguana-17148	211	14	mitochondrial	mitochondrial	ADJ
iguana-17148	211	15	genetic	genetic	ADJ
iguana-17148	211	16	analyses	analysis	NOUN
iguana-17148	211	17	.	.	PUNCT
iguana-17148	212	1	molecular	molecular	ADJ
iguana-17148	212	2	ecology	ecology	NOUN
iguana-17148	212	3	6:437–452	6:437–452	PROPN
iguana-17148	212	4	.	.	PUNCT
iguana-17148	213	1	romero	romero	PROPN
iguana-17148	213	2	,	,	PUNCT
iguana-17148	213	3	l.	l.	PROPN
iguana-17148	213	4	m.	m.	PROPN
iguana-17148	213	5	and	and	CCONJ
iguana-17148	213	6	m.	m.	NOUN
iguana-17148	213	7	wikelski	wikelski	PROPN
iguana-17148	213	8	,	,	PUNCT
iguana-17148	213	9	m.	m.	NOUN
iguana-17148	213	10	2001	2001	NUM
iguana-17148	213	11	.	.	PUNCT
iguana-17148	214	1	corticosterone	corticosterone	NOUN
iguana-17148	214	2	levels	level	NOUN
iguana-17148	214	3	predict	predict	VERB
iguana-17148	214	4	survival	survival	NOUN
iguana-17148	214	5	rates	rate	NOUN
iguana-17148	214	6	in	in	ADP
iguana-17148	214	7	galápagos	galápagos	PROPN
iguana-17148	214	8	marine	marine	PROPN
iguana-17148	214	9	iguanas	iguana	NOUN
iguana-17148	214	10	.	.	PUNCT
iguana-17148	215	1	proceedings	proceeding	NOUN
iguana-17148	215	2	of	of	ADP
iguana-17148	215	3	the	the	DET
iguana-17148	215	4	national	national	PROPN
iguana-17148	215	5	academy	academy	PROPN
iguana-17148	215	6	of	of	ADP
iguana-17148	215	7	sciences	sciences	PROPN
iguana-17148	215	8	98:7366–7370	98:7366–7370	NUM
iguana-17148	215	9	.	.	PUNCT
iguana-17148	216	1	trillmich	trillmich	NOUN
iguana-17148	216	2	,	,	PUNCT
iguana-17148	216	3	k.	k.	PROPN
iguana-17148	216	4	1983	1983	NUM
iguana-17148	216	5	.	.	PUNCT
iguana-17148	217	1	the	the	DET
iguana-17148	217	2	mating	mating	NOUN
iguana-17148	217	3	system	system	NOUN
iguana-17148	217	4	of	of	ADP
iguana-17148	217	5	the	the	DET
iguana-17148	217	6	marine	marine	PROPN
iguana-17148	217	7	iguana	iguana	PROPN
iguana-17148	217	8	,	,	PUNCT
iguana-17148	217	9	amblyrhynchus	amblyrhynchu	VERB
iguana-17148	217	10	cristatus	cristatus	PROPN
iguana-17148	217	11	.	.	PUNCT
iguana-17148	218	1	zeitschrift	zeitschrift	PROPN
iguana-17148	218	2	für	für	PROPN
iguana-17148	218	3	tierpsychologie	tierpsychologie	PROPN
iguana-17148	218	4	63:141–172	63:141–172	PROPN
iguana-17148	218	5	.	.	PUNCT
iguana-17148	219	1	trillmich	trillmich	NOUN
iguana-17148	219	2	,	,	PUNCT
iguana-17148	219	3	k.	k.	PROPN
iguana-17148	219	4	and	and	CCONJ
iguana-17148	219	5	f.	f.	PROPN
iguana-17148	219	6	trillmich	trillmich	PROPN
iguana-17148	219	7	.	.	PUNCT
iguana-17148	220	1	1986	1986	NUM
iguana-17148	220	2	.	.	PUNCT
iguana-17148	221	1	foraging	forage	VERB
iguana-17148	221	2	strategies	strategy	NOUN
iguana-17148	221	3	of	of	ADP
iguana-17148	221	4	the	the	DET
iguana-17148	221	5	marine	marine	ADJ
iguana-17148	221	6	iguana	iguana	PROPN
iguana-17148	221	7	,	,	PUNCT
iguana-17148	221	8	amblyrhynchus	amblyrhynchu	VERB
iguana-17148	221	9	cristatus	cristatus	NOUN
iguana-17148	221	10	.	.	PUNCT
iguana-17148	222	1	behavioural	behavioural	ADJ
iguana-17148	222	2	ecology	ecology	NOUN
iguana-17148	222	3	and	and	CCONJ
iguana-17148	222	4	sociobiology	sociobiology	NOUN
iguana-17148	222	5	18:259–266	18:259–266	PROPN
iguana-17148	222	6	.	.	PUNCT
iguana-17148	223	1	wikelski	wikelski	PROPN
iguana-17148	223	2	,	,	PUNCT
iguana-17148	223	3	m.	m.	NOUN
iguana-17148	223	4	1999	1999	NUM
iguana-17148	223	5	.	.	PUNCT
iguana-17148	224	1	influences	influence	NOUN
iguana-17148	224	2	of	of	ADP
iguana-17148	224	3	parasites	parasite	NOUN
iguana-17148	224	4	and	and	CCONJ
iguana-17148	224	5	thermoregulation	thermoregulation	NOUN
iguana-17148	224	6	on	on	ADP
iguana-17148	224	7	grouping	group	VERB
iguana-17148	224	8	tendencies	tendency	NOUN
iguana-17148	224	9	in	in	ADP
iguana-17148	224	10	marine	marine	ADJ
iguana-17148	224	11	iguanas	iguana	NOUN
iguana-17148	224	12	.	.	PUNCT
iguana-17148	225	1	behavioral	behavioral	ADJ
iguana-17148	225	2	ecology	ecology	NOUN
iguana-17148	225	3	10:22–29	10:22–29	PROPN
iguana-17148	225	4	.	.	PUNCT
iguana-17148	226	1	wikelski	wikelski	PROPN
iguana-17148	226	2	,	,	PUNCT
iguana-17148	226	3	m.	m.	NOUN
iguana-17148	226	4	and	and	CCONJ
iguana-17148	226	5	s.	s.	PROPN
iguana-17148	226	6	bäurle	bäurle	PROPN
iguana-17148	226	7	.	.	PUNCT
iguana-17148	226	8	1996	1996	NUM
iguana-17148	226	9	.	.	PUNCT
iguana-17148	227	1	pre	pre	ADJ
iguana-17148	227	2	-	-	ADJ
iguana-17148	227	3	copulatory	copulatory	ADJ
iguana-17148	227	4	ejaculation	ejaculation	NOUN
iguana-17148	227	5	solves	solve	NOUN
iguana-17148	227	6	time	time	NOUN
iguana-17148	227	7	constraints	constraint	NOUN
iguana-17148	227	8	during	during	ADP
iguana-17148	227	9	copulation	copulation	NOUN
iguana-17148	227	10	in	in	ADP
iguana-17148	227	11	marine	marine	ADJ
iguana-17148	227	12	iguanas	iguana	NOUN
iguana-17148	227	13	.	.	PUNCT
iguana-17148	228	1	proceedings	proceeding	NOUN
iguana-17148	228	2	of	of	ADP
iguana-17148	228	3	the	the	DET
iguana-17148	228	4	royal	royal	ADJ
iguana-17148	228	5	society	society	NOUN
iguana-17148	228	6	of	of	ADP
iguana-17148	228	7	london	london	PROPN
iguana-17148	228	8	b	b	PROPN
iguana-17148	228	9	263:439–444	263:439–444	NUM
iguana-17148	228	10	.	.	PUNCT
iguana-17148	229	1	wikelski	wikelski	NOUN
iguana-17148	229	2	,	,	PUNCT
iguana-17148	229	3	m.	m.	NOUN
iguana-17148	229	4	,	,	PUNCT
iguana-17148	229	5	c.	c.	NOUN
iguana-17148	229	6	carbone	carbone	NOUN
iguana-17148	229	7	,	,	PUNCT
iguana-17148	229	8	p.	p.	NOUN
iguana-17148	229	9	a.	a.	NOUN
iguana-17148	229	10	bednekoff	bednekoff	PROPN
iguana-17148	229	11	,	,	PUNCT
iguana-17148	229	12	c.	c.	PROPN
iguana-17148	229	13	choudhury	choudhury	PROPN
iguana-17148	229	14	,	,	PUNCT
iguana-17148	229	15	and	and	CCONJ
iguana-17148	229	16	s.	s.	PROPN
iguana-17148	229	17	tebbich	tebbich	PROPN
iguana-17148	229	18	.	.	PUNCT
iguana-17148	230	1	2001	2001	NUM
iguana-17148	230	2	.	.	PUNCT
iguana-17148	231	1	female	female	ADJ
iguana-17148	231	2	choice	choice	NOUN
iguana-17148	231	3	in	in	ADP
iguana-17148	231	4	marine	marine	ADJ
iguana-17148	231	5	iguana	iguana	NOUN
iguana-17148	231	6	leks	lek	NOUN
iguana-17148	231	7	:	:	PUNCT
iguana-17148	231	8	a	a	DET
iguana-17148	231	9	wider	wide	ADJ
iguana-17148	231	10	selection	selection	NOUN
iguana-17148	231	11	of	of	ADP
iguana-17148	231	12	males	male	NOUN
iguana-17148	231	13	obtained	obtain	VERB
iguana-17148	231	14	at	at	ADP
iguana-17148	231	15	a	a	DET
iguana-17148	231	16	cost	cost	NOUN
iguana-17148	231	17	.	.	PUNCT
iguana-17148	232	1	ethology	ethology	NOUN
iguana-17148	232	2	107:623–638	107:623–638	PROPN
iguana-17148	232	3	.	.	PUNCT
iguana-17148	233	1	wikelski	wikelski	PROPN
iguana-17148	233	2	,	,	PUNCT
iguana-17148	233	3	m.	m.	NOUN
iguana-17148	233	4	,	,	PUNCT
iguana-17148	233	5	c.	c.	NOUN
iguana-17148	233	6	carbone	carbone	NOUN
iguana-17148	233	7	,	,	PUNCT
iguana-17148	233	8	and	and	CCONJ
iguana-17148	233	9	f.	f.	PROPN
iguana-17148	233	10	trillmich	trillmich	PROPN
iguana-17148	233	11	.	.	PUNCT
iguana-17148	234	1	1996	1996	NUM
iguana-17148	234	2	.	.	PUNCT
iguana-17148	235	1	lekking	lekke	VERB
iguana-17148	235	2	in	in	ADP
iguana-17148	235	3	marine	marine	ADJ
iguana-17148	235	4	iguanas	iguana	NOUN
iguana-17148	235	5	:	:	PUNCT
iguana-17148	235	6	female	female	ADJ
iguana-17148	235	7	grouping	grouping	NOUN
iguana-17148	235	8	and	and	CCONJ
iguana-17148	235	9	male	male	ADJ
iguana-17148	235	10	reproductive	reproductive	ADJ
iguana-17148	235	11	strategies	strategy	NOUN
iguana-17148	235	12	.	.	PUNCT
iguana-17148	236	1	animal	animal	NOUN
iguana-17148	236	2	behavior	behavior	NOUN
iguana-17148	236	3	52:581–596	52:581–596	PROPN
iguana-17148	236	4	.	.	PUNCT
iguana-17148	237	1	wikelski	wikelski	PROPN
iguana-17148	237	2	,	,	PUNCT
iguana-17148	237	3	m.	m.	NOUN
iguana-17148	237	4	,	,	PUNCT
iguana-17148	237	5	v.	v.	PROPN
iguana-17148	237	6	carillo	carillo	PROPN
iguana-17148	237	7	,	,	PUNCT
iguana-17148	237	8	and	and	CCONJ
iguana-17148	237	9	p.	p.	NOUN
iguana-17148	237	10	trillmich	trillmich	NOUN
iguana-17148	237	11	.	.	PUNCT
iguana-17148	238	1	1997	1997	NUM
iguana-17148	238	2	.	.	PUNCT
iguana-17148	239	1	energy	energy	NOUN
iguana-17148	239	2	limits	limit	NOUN
iguana-17148	239	3	to	to	ADP
iguana-17148	239	4	body	body	NOUN
iguana-17148	239	5	size	size	NOUN
iguana-17148	239	6	in	in	ADP
iguana-17148	239	7	a	a	DET
iguana-17148	239	8	grazing	grazing	NOUN
iguana-17148	239	9	reptile	reptile	NOUN
iguana-17148	239	10	,	,	PUNCT
iguana-17148	239	11	the	the	DET
iguana-17148	239	12	galápagos	galápagos	PROPN
iguana-17148	239	13	marine	marine	PROPN
iguana-17148	239	14	iguana	iguana	PROPN
iguana-17148	239	15	.	.	PUNCT
iguana-17148	240	1	ecology	ecology	NOUN
iguana-17148	240	2	78:2204–2217	78:2204–2217	PROPN
iguana-17148	240	3	.	.	PUNCT
iguana-17148	241	1	wikelski	wikelski	PROPN
iguana-17148	241	2	,	,	PUNCT
iguana-17148	241	3	m.	m.	NOUN
iguana-17148	241	4	,	,	PUNCT
iguana-17148	241	5	b.	b.	PROPN
iguana-17148	241	6	gall	gall	PROPN
iguana-17148	241	7	,	,	PUNCT
iguana-17148	241	8	and	and	CCONJ
iguana-17148	241	9	f.	f.	PROPN
iguana-17148	241	10	trillmich	trillmich	PROPN
iguana-17148	241	11	.	.	PUNCT
iguana-17148	242	1	1993	1993	NUM
iguana-17148	242	2	.	.	PUNCT
iguana-17148	243	1	ontogenetic	ontogenetic	ADJ
iguana-17148	243	2	changes	change	NOUN
iguana-17148	243	3	in	in	ADP
iguana-17148	243	4	food	food	NOUN
iguana-17148	243	5	intake	intake	NOUN
iguana-17148	243	6	and	and	CCONJ
iguana-17148	243	7	digestion	digestion	NOUN
iguana-17148	243	8	rate	rate	NOUN
iguana-17148	243	9	of	of	ADP
iguana-17148	243	10	the	the	DET
iguana-17148	243	11	herbivorous	herbivorous	ADJ
iguana-17148	243	12	marine	marine	ADJ
iguana-17148	243	13	iguana	iguana	NOUN
iguana-17148	243	14	amblyrhynchus	amblyrhynchus	PROPN
iguana-17148	243	15	cristatus	cristatus	PROPN
iguana-17148	243	16	(	(	PUNCT
iguana-17148	243	17	bell	bell	PROPN
iguana-17148	243	18	)	)	PUNCT
iguana-17148	243	19	.	.	PUNCT
iguana-17148	244	1	oecologia	oecologia	PROPN
iguana-17148	244	2	94:373–379	94:373–379	PROPN
iguana-17148	244	3	.	.	PUNCT
iguana-17148	245	1	wikelski	wikelski	PROPN
iguana-17148	245	2	,	,	PUNCT
iguana-17148	245	3	m.	m.	NOUN
iguana-17148	245	4	and	and	CCONJ
iguana-17148	245	5	m.	m.	PROPN
iguana-17148	245	6	hau	hau	PROPN
iguana-17148	245	7	.	.	PROPN
iguana-17148	245	8	1995	1995	NUM
iguana-17148	245	9	.	.	PUNCT
iguana-17148	246	1	is	be	AUX
iguana-17148	246	2	there	there	PRON
iguana-17148	246	3	an	an	DET
iguana-17148	246	4	endogenous	endogenous	ADJ
iguana-17148	246	5	tidal	tidal	ADJ
iguana-17148	246	6	foraging	forage	VERB
iguana-17148	246	7	rhythm	rhythm	NOUN
iguana-17148	246	8	in	in	ADP
iguana-17148	246	9	marine	marine	ADJ
iguana-17148	246	10	iguanas	iguana	NOUN
iguana-17148	246	11	?	?	PUNCT
iguana-17148	247	1	journal	journal	NOUN
iguana-17148	247	2	of	of	ADP
iguana-17148	247	3	biological	biological	ADJ
iguana-17148	247	4	rhythms	rhythm	NOUN
iguana-17148	247	5	10:335–350	10:335–350	PROPN
iguana-17148	247	6	.	.	PUNCT
iguana-17148	248	1	wikelski	wikelski	PROPN
iguana-17148	248	2	,	,	PUNCT
iguana-17148	248	3	m.	m.	NOUN
iguana-17148	248	4	and	and	CCONJ
iguana-17148	248	5	c.	c.	PROPN
iguana-17148	248	6	thom	thom	PROPN
iguana-17148	248	7	.	.	PROPN
iguana-17148	248	8	2000	2000	NUM
iguana-17148	248	9	.	.	PUNCT
iguana-17148	249	1	marine	marine	PROPN
iguana-17148	249	2	iguanas	iguana	NOUN
iguana-17148	249	3	shrink	shrink	VERB
iguana-17148	249	4	to	to	PART
iguana-17148	249	5	survive	survive	VERB
iguana-17148	249	6	el	el	PROPN
iguana-17148	249	7	niño	niño	PROPN
iguana-17148	249	8	.	.	PUNCT
iguana-17148	250	1	nature	nature	PROPN
iguana-17148	251	1	403:37–38	403:37–38	PROPN
iguana-17148	251	2	.	.	PUNCT
iguana-17148	251	3	wikelski	wikelski	PROPN
iguana-17148	251	4	,	,	PUNCT
iguana-17148	251	5	m.	m.	NOUN
iguana-17148	251	6	and	and	CCONJ
iguana-17148	251	7	f.	f.	PROPN
iguana-17148	251	8	trillmich	trillmich	PROPN
iguana-17148	251	9	.	.	PUNCT
iguana-17148	252	1	1994	1994	NUM
iguana-17148	252	2	.	.	PUNCT
iguana-17148	253	1	foraging	forage	VERB
iguana-17148	253	2	strategies	strategy	NOUN
iguana-17148	253	3	of	of	ADP
iguana-17148	253	4	the	the	DET
iguana-17148	253	5	galápagos	galápagos	PROPN
iguana-17148	253	6	marine	marine	PROPN
iguana-17148	253	7	iguana	iguana	NOUN
iguana-17148	253	8	(	(	PUNCT
iguana-17148	253	9	amblyrhynchus	amblyrhynchus	ADV
iguana-17148	253	10	cristatus	cristatus	NOUN
iguana-17148	253	11	):	):	PUNCT
iguana-17148	253	12	adapting	adapt	VERB
iguana-17148	253	13	behavioral	behavioral	ADJ
iguana-17148	253	14	rules	rule	NOUN
iguana-17148	253	15	to	to	ADP
iguana-17148	253	16	ontogenetic	ontogenetic	ADJ
iguana-17148	253	17	size	size	NOUN
iguana-17148	253	18	change	change	NOUN
iguana-17148	253	19	.	.	PUNCT
iguana-17148	254	1	behavior	behavior	NOUN
iguana-17148	254	2	128:255–279	128:255–279	NUM
iguana-17148	254	3	.	.	PUNCT
iguana-17148	255	1	wikelski	wikelski	PROPN
iguana-17148	255	2	,	,	PUNCT
iguana-17148	255	3	m.	m.	NOUN
iguana-17148	255	4	and	and	CCONJ
iguana-17148	255	5	f.	f.	PROPN
iguana-17148	255	6	trillmich	trillmich	PROPN
iguana-17148	255	7	.	.	PUNCT
iguana-17148	256	1	1997	1997	NUM
iguana-17148	256	2	.	.	PUNCT
iguana-17148	257	1	body	body	NOUN
iguana-17148	257	2	size	size	NOUN
iguana-17148	257	3	and	and	CCONJ
iguana-17148	257	4	sexual	sexual	ADJ
iguana-17148	257	5	size	size	NOUN
iguana-17148	257	6	dimorphism	dimorphism	NOUN
iguana-17148	257	7	in	in	ADP
iguana-17148	257	8	marine	marine	ADJ
iguana-17148	257	9	iguanas	iguana	NOUN
iguana-17148	257	10	fluctuate	fluctuate	NOUN
iguana-17148	257	11	as	as	ADP
iguana-17148	257	12	a	a	DET
iguana-17148	257	13	result	result	NOUN
iguana-17148	257	14	of	of	ADP
iguana-17148	257	15	opposing	oppose	VERB
iguana-17148	257	16	natural	natural	ADJ
iguana-17148	257	17	and	and	CCONJ
iguana-17148	257	18	sexual	sexual	ADJ
iguana-17148	257	19	selection	selection	NOUN
iguana-17148	257	20	:	:	PUNCT
iguana-17148	257	21	an	an	DET
iguana-17148	257	22	island	island	NOUN
iguana-17148	257	23	comparison	comparison	NOUN
iguana-17148	257	24	.	.	PUNCT
iguana-17148	258	1	evolution	evolution	PROPN
iguana-17148	258	2	51:922–936	51:922–936	PROPN
iguana-17148	258	3	.	.	PUNCT
iguana-17148	259	1	wikelski	wikelski	PROPN
iguana-17148	259	2	,	,	PUNCT
iguana-17148	259	3	m.	m.	NOUN
iguana-17148	259	4	and	and	CCONJ
iguana-17148	259	5	p.	p.	PROPN
iguana-17148	259	6	h.	h.	PROPN
iguana-17148	259	7	wrege	wrege	PROPN
iguana-17148	259	8	.	.	PUNCT
iguana-17148	260	1	2000	2000	NUM
iguana-17148	260	2	.	.	PUNCT
iguana-17148	261	1	niche	niche	NOUN
iguana-17148	261	2	expansion	expansion	NOUN
iguana-17148	261	3	,	,	PUNCT
iguana-17148	261	4	body	body	NOUN
iguana-17148	261	5	size	size	NOUN
iguana-17148	261	6	,	,	PUNCT
iguana-17148	261	7	and	and	CCONJ
iguana-17148	261	8	survival	survival	NOUN
iguana-17148	261	9	in	in	ADP
iguana-17148	261	10	galápagos	galápagos	PROPN
iguana-17148	261	11	marine	marine	PROPN
iguana-17148	261	12	iguanas	iguana	NOUN
iguana-17148	261	13	.	.	PUNCT
iguana-17148	262	1	oecologia	oecologia	PROPN
iguana-17148	262	2	124:107–115	124:107–115	PROPN
iguana-17148	262	3	.	.	PUNCT
iguana-17148	263	1	author	author	NOUN
iguana-17148	263	2	biographies	biographie	VERB
iguana-17148	263	3	martin	martin	PROPN
iguana-17148	263	4	c.	c.	PROPN
iguana-17148	263	5	wikelski	wikelski	PROPN
iguana-17148	263	6	is	be	AUX
iguana-17148	263	7	an	an	DET
iguana-17148	263	8	assistant	assistant	ADJ
iguana-17148	263	9	professor	professor	NOUN
iguana-17148	263	10	in	in	ADP
iguana-17148	263	11	the	the	DET
iguana-17148	263	12	department	department	NOUN
iguana-17148	263	13	of	of	ADP
iguana-17148	263	14	ecology	ecology	NOUN
iguana-17148	263	15	and	and	CCONJ
iguana-17148	263	16	evolutionary	evolutionary	ADJ
iguana-17148	263	17	biology	biology	NOUN
iguana-17148	263	18	at	at	ADP
iguana-17148	263	19	princeton	princeton	PROPN
iguana-17148	263	20	university	university	PROPN
iguana-17148	263	21	.	.	PUNCT
iguana-17148	264	1	he	he	PRON
iguana-17148	264	2	completed	complete	VERB
iguana-17148	264	3	his	his	PRON
iguana-17148	264	4	masters	master	NOUN
iguana-17148	264	5	and	and	CCONJ
iguana-17148	264	6	ph.d	ph.d	PROPN
iguana-17148	264	7	.	.	PUNCT
iguana-17148	265	1	at	at	ADP
iguana-17148	265	2	the	the	DET
iguana-17148	265	3	german	german	ADJ
iguana-17148	265	4	max	max	PROPN
iguana-17148	265	5	planck	planck	PROPN
iguana-17148	265	6	institute	institute	PROPN
iguana-17148	265	7	and	and	CCONJ
iguana-17148	265	8	the	the	DET
iguana-17148	265	9	university	university	NOUN
iguana-17148	265	10	of	of	ADP
iguana-17148	265	11	bielefeld	bielefeld	PROPN
iguana-17148	265	12	before	before	SCONJ
iguana-17148	265	13	he	he	PRON
iguana-17148	265	14	came	come	VERB
iguana-17148	265	15	to	to	ADP
iguana-17148	265	16	the	the	DET
iguana-17148	265	17	united	united	PROPN
iguana-17148	265	18	states	states	PROPN
iguana-17148	265	19	as	as	ADP
iguana-17148	265	20	a	a	DET
iguana-17148	265	21	postdoctoral	postdoctoral	ADJ
iguana-17148	265	22	researcher	researcher	NOUN
iguana-17148	265	23	working	work	VERB
iguana-17148	265	24	at	at	ADP
iguana-17148	265	25	the	the	DET
iguana-17148	265	26	smithsonian	smithsonian	PROPN
iguana-17148	265	27	tropical	tropical	PROPN
iguana-17148	265	28	research	research	PROPN
iguana-17148	265	29	institute	institute	PROPN
iguana-17148	265	30	and	and	CCONJ
iguana-17148	265	31	the	the	DET
iguana-17148	265	32	university	university	PROPN
iguana-17148	265	33	of	of	ADP
iguana-17148	265	34	washington	washington	PROPN
iguana-17148	265	35	.	.	PUNCT
iguana-17148	266	1	martin	martin	PROPN
iguana-17148	266	2	then	then	ADV
iguana-17148	266	3	held	hold	VERB
iguana-17148	266	4	an	an	DET
iguana-17148	266	5	assistant	assistant	ADJ
iguana-17148	266	6	professorship	professorship	NOUN
iguana-17148	266	7	at	at	ADP
iguana-17148	266	8	the	the	DET
iguana-17148	266	9	university	university	PROPN
iguana-17148	266	10	of	of	ADP
iguana-17148	266	11	illinois	illinois	PROPN
iguana-17148	266	12	at	at	ADP
iguana-17148	266	13	urbana	urbana	PROPN
iguana-17148	266	14	-	-	PUNCT
iguana-17148	266	15	champaign	champaign	PROPN
iguana-17148	266	16	for	for	ADP
iguana-17148	266	17	two	two	NUM
iguana-17148	266	18	years	year	NOUN
iguana-17148	266	19	.	.	PUNCT
iguana-17148	267	1	his	his	PRON
iguana-17148	267	2	research	research	NOUN
iguana-17148	267	3	interest	interest	NOUN
iguana-17148	267	4	is	be	AUX
iguana-17148	267	5	ecological	ecological	ADJ
iguana-17148	267	6	physiology	physiology	NOUN
iguana-17148	267	7	of	of	ADP
iguana-17148	267	8	lizards	lizard	NOUN
iguana-17148	267	9	and	and	CCONJ
iguana-17148	267	10	birds	bird	NOUN
iguana-17148	267	11	.	.	PUNCT
iguana-17148	268	1	he	he	PRON
iguana-17148	268	2	has	have	AUX
iguana-17148	268	3	traveled	travel	VERB
iguana-17148	268	4	extensively	extensively	ADV
iguana-17148	268	5	through	through	ADP
iguana-17148	268	6	south	south	PROPN
iguana-17148	268	7	america	america	PROPN
iguana-17148	268	8	,	,	PUNCT
iguana-17148	268	9	publishing	publish	VERB
iguana-17148	268	10	a	a	DET
iguana-17148	268	11	nature	nature	NOUN
iguana-17148	268	12	travel	travel	NOUN
iguana-17148	268	13	guide	guide	NOUN
iguana-17148	268	14	to	to	ADP
iguana-17148	268	15	venezuela	venezuela	PROPN
iguana-17148	268	16	and	and	CCONJ
iguana-17148	268	17	brazil	brazil	PROPN
iguana-17148	268	18	.	.	PUNCT
iguana-17148	269	1	he	he	PRON
iguana-17148	269	2	continues	continue	VERB
iguana-17148	269	3	work	work	NOUN
iguana-17148	269	4	in	in	ADP
iguana-17148	269	5	panamá	panamá	NOUN
iguana-17148	269	6	and	and	CCONJ
iguana-17148	269	7	the	the	DET
iguana-17148	269	8	galápagos	galápagos	PROPN
iguana-17148	269	9	islands	island	NOUN
iguana-17148	269	10	,	,	PUNCT
iguana-17148	269	11	ecuador	ecuador	PROPN
iguana-17148	269	12	.	.	PUNCT
iguana-17148	270	1	karin	karin	PROPN
iguana-17148	270	2	n.	n.	PROPN
iguana-17148	270	3	nelson	nelson	PROPN
iguana-17148	270	4	completed	complete	VERB
iguana-17148	270	5	her	her	PRON
iguana-17148	270	6	ph.d	ph.d	NOUN
iguana-17148	270	7	.	.	PUNCT
iguana-17148	271	1	in	in	ADP
iguana-17148	271	2	2003	2003	NUM
iguana-17148	271	3	in	in	ADP
iguana-17148	271	4	the	the	DET
iguana-17148	271	5	department	department	NOUN
iguana-17148	271	6	of	of	ADP
iguana-17148	271	7	animal	animal	NOUN
iguana-17148	271	8	biology	biology	NOUN
iguana-17148	271	9	at	at	ADP
iguana-17148	271	10	the	the	DET
iguana-17148	271	11	university	university	PROPN
iguana-17148	271	12	of	of	ADP
iguana-17148	271	13	illinois	illinois	PROPN
iguana-17148	271	14	at	at	ADP
iguana-17148	271	15	urbana	urbana	PROPN
iguana-17148	271	16	-	-	PUNCT
iguana-17148	271	17	champaign	champaign	PROPN
iguana-17148	271	18	.	.	PUNCT
iguana-17148	272	1	before	before	ADP
iguana-17148	272	2	returning	return	VERB
iguana-17148	272	3	to	to	ADP
iguana-17148	272	4	academia	academia	NOUN
iguana-17148	272	5	,	,	PUNCT
iguana-17148	272	6	she	she	PRON
iguana-17148	272	7	served	serve	VERB
iguana-17148	272	8	as	as	ADP
iguana-17148	272	9	editor	editor	NOUN
iguana-17148	272	10	at	at	ADP
iguana-17148	272	11	chicago	chicago	PROPN
iguana-17148	272	12	’s	’s	PART
iguana-17148	272	13	brookfield	brookfield	PROPN
iguana-17148	272	14	zoo	zoo	PROPN
iguana-17148	272	15	for	for	ADP
iguana-17148	272	16	eight	eight	NUM
iguana-17148	272	17	years	year	NOUN
iguana-17148	272	18	.	.	PUNCT
iguana-17148	273	1	during	during	ADP
iguana-17148	273	2	her	her	PRON
iguana-17148	273	3	graduate	graduate	NOUN
iguana-17148	273	4	studies	study	NOUN
iguana-17148	273	5	,	,	PUNCT
iguana-17148	273	6	she	she	PRON
iguana-17148	273	7	investigated	investigate	VERB
iguana-17148	273	8	the	the	DET
iguana-17148	273	9	hormonal	hormonal	ADJ
iguana-17148	273	10	mechanisms	mechanism	NOUN
iguana-17148	273	11	of	of	ADP
iguana-17148	273	12	alternative	alternative	ADJ
iguana-17148	273	13	mating	mating	NOUN
iguana-17148	273	14	strategies	strategy	NOUN
iguana-17148	273	15	in	in	ADP
iguana-17148	273	16	marine	marine	ADJ
iguana-17148	273	17	iguanas	iguana	NOUN
iguana-17148	273	18	.	.	PUNCT
iguana-17148	274	1	iguana	iguana	NOUN
iguana-17148	274	2	•	•	NUM
iguana-17148	274	3	volume	volume	NOUN
iguana-17148	274	4	11	11	NUM
iguana-17148	274	5	,	,	PUNCT
iguana-17148	274	6	number	number	NOUN
iguana-17148	274	7	4	4	NUM
iguana-17148	274	8	•	•	NOUN
iguana-17148	274	9	december	december	PROPN
iguana-17148	274	10	2004	2004	NUM
iguana-17148	274	11	197galápagos	197galápagos	PROPN
iguana-17148	274	12	marine	marine	PROPN
iguana-17148	274	13	iguanas	iguanas	PROPN
iguana-17148	274	14	martin	martin	PROPN
iguana-17148	274	15	wikelski	wikelski	PROPN
iguana-17148	274	16	holding	hold	VERB
iguana-17148	274	17	a	a	DET
iguana-17148	274	18	large	large	ADJ
iguana-17148	274	19	male	male	ADJ
iguana-17148	274	20	marine	marine	ADJ
iguana-17148	274	21	iguana	iguana	NOUN
iguana-17148	274	22	on	on	ADP
iguana-17148	274	23	western	western	ADJ
iguana-17148	274	24	isabela	isabela	PROPN
iguana-17148	274	25	island	island	NOUN
iguana-17148	274	26	.	.	PUNCT
iguana-17148	275	1	dead	dead	ADJ
iguana-17148	275	2	marine	marine	ADJ
iguana-17148	275	3	iguana	iguana	NOUN
iguana-17148	275	4	on	on	ADP
iguana-17148	275	5	fernandina	fernandina	PROPN
iguana-17148	275	6	island	island	PROPN
iguana-17148	275	7	during	during	ADP
iguana-17148	275	8	the	the	DET
iguana-17148	275	9	1998–99	1998–99	PROPN
iguana-17148	275	10	el	el	PROPN
iguana-17148	275	11	niño	niño	PROPN
iguana-17148	275	12	period	period	NOUN
iguana-17148	275	13	.	.	PUNCT
iguana-17148	276	1	the	the	DET
iguana-17148	276	2	likely	likely	ADJ
iguana-17148	276	3	cause	cause	NOUN
iguana-17148	276	4	of	of	ADP
iguana-17148	276	5	death	death	NOUN
iguana-17148	276	6	was	be	AUX
iguana-17148	276	7	starvation	starvation	NOUN
iguana-17148	276	8	.	.	PUNCT