id	sid	tid	token	lemma	pos
iguana-17918	1	1	iguana	iguana	NOUN
iguana-17918	1	2	b&w	b&w	PROPN
iguana-17918	1	3	text	text	NOUN
iguana-17918	1	4	iguana	iguana	NOUN
iguana-17918	1	5	•	•	NUM
iguana-17918	1	6	volume	volume	NOUN
iguana-17918	1	7	15	15	NUM
iguana-17918	1	8	,	,	PUNCT
iguana-17918	1	9	number	number	NOUN
iguana-17918	1	10	4	4	NUM
iguana-17918	1	11	•	•	NOUN
iguana-17918	1	12	december	december	PROPN
iguana-17918	1	13	2008	2008	NUM
iguana-17918	2	1	245conservation	245conservation	PROPN
iguana-17918	2	2	research	research	NOUN
iguana-17918	2	3	reports	report	VERB
iguana-17918	2	4	new	new	ADJ
iguana-17918	2	5	pacific	pacific	ADJ
iguana-17918	2	6	iguana	iguana	NOUN
iguana-17918	2	7	the	the	DET
iguana-17918	2	8	pacific	pacific	PROPN
iguana-17918	2	9	iguanas	iguana	NOUN
iguana-17918	2	10	of	of	ADP
iguana-17918	2	11	the	the	DET
iguana-17918	2	12	fijian	fijian	ADJ
iguana-17918	2	13	and	and	CCONJ
iguana-17918	2	14	tongan	tongan	PROPN
iguana-17918	2	15	archipelagos	archipelago	NOUN
iguana-17918	2	16	are	be	AUX
iguana-17918	2	17	a	a	DET
iguana-17918	2	18	biogeographic	biogeographic	ADJ
iguana-17918	2	19	enigma	enigma	NOUN
iguana-17918	2	20	in	in	SCONJ
iguana-17918	2	21	that	that	SCONJ
iguana-17918	2	22	their	their	PRON
iguana-17918	2	23	closest	close	ADJ
iguana-17918	2	24	relatives	relative	NOUN
iguana-17918	2	25	are	be	AUX
iguana-17918	2	26	found	find	VERB
iguana-17918	2	27	only	only	ADV
iguana-17918	2	28	in	in	ADP
iguana-17918	2	29	the	the	DET
iguana-17918	2	30	western	western	ADJ
iguana-17918	2	31	hemisphere	hemisphere	NOUN
iguana-17918	2	32	.	.	PUNCT
iguana-17918	3	1	these	these	DET
iguana-17918	3	2	iguanas	iguana	NOUN
iguana-17918	3	3	were	be	AUX
iguana-17918	3	4	thought	think	VERB
iguana-17918	3	5	to	to	PART
iguana-17918	3	6	comprise	comprise	VERB
iguana-17918	3	7	two	two	NUM
iguana-17918	3	8	genera	genera	NOUN
iguana-17918	3	9	and	and	CCONJ
iguana-17918	3	10	four	four	NUM
iguana-17918	3	11	species	specie	NOUN
iguana-17918	3	12	of	of	ADP
iguana-17918	3	13	extinct	extinct	ADJ
iguana-17918	3	14	and	and	CCONJ
iguana-17918	3	15	extant	extant	ADJ
iguana-17918	3	16	taxa	taxa	NOUN
iguana-17918	3	17	.	.	PUNCT
iguana-17918	4	1	the	the	DET
iguana-17918	4	2	two	two	NUM
iguana-17918	4	3	currently	currently	ADV
iguana-17918	4	4	recognized	recognize	VERB
iguana-17918	4	5	extant	extant	ADJ
iguana-17918	4	6	species	specie	NOUN
iguana-17918	4	7	,	,	PUNCT
iguana-17918	4	8	brachylophus	brachylophus	NOUN
iguana-17918	4	9	fasciatus	fasciatus	NOUN
iguana-17918	4	10	,	,	PUNCT
iguana-17918	4	11	from	from	ADP
iguana-17918	4	12	fiji	fiji	PROPN
iguana-17918	4	13	,	,	PUNCT
iguana-17918	4	14	tonga	tonga	PROPN
iguana-17918	4	15	,	,	PUNCT
iguana-17918	4	16	and	and	CCONJ
iguana-17918	4	17	vanuatu	vanuatu	NOUN
iguana-17918	4	18	,	,	PUNCT
iguana-17918	4	19	and	and	CCONJ
iguana-17918	4	20	brachylophus	brachylophus	NOUN
iguana-17918	4	21	vitiensis	vitiensis	NOUN
iguana-17918	4	22	,	,	PUNCT
iguana-17918	4	23	from	from	ADP
iguana-17918	4	24	western	western	PROPN
iguana-17918	4	25	fiji	fiji	PROPN
iguana-17918	4	26	,	,	PUNCT
iguana-17918	4	27	are	be	AUX
iguana-17918	4	28	of	of	ADP
iguana-17918	4	29	considerable	considerable	ADJ
iguana-17918	4	30	conservation	conservation	NOUN
iguana-17918	4	31	concern	concern	NOUN
iguana-17918	4	32	,	,	PUNCT
iguana-17918	4	33	with	with	ADP
iguana-17918	4	34	b.	b.	PROPN
iguana-17918	4	35	vitiensis	vitiensis	PROPN
iguana-17918	4	36	listed	list	VERB
iguana-17918	4	37	as	as	ADP
iguana-17918	4	38	critically	critically	ADV
iguana-17918	4	39	endangered	endanger	VERB
iguana-17918	4	40	.	.	PUNCT
iguana-17918	5	1	in	in	ADP
iguana-17918	5	2	a	a	DET
iguana-17918	5	3	recent	recent	ADJ
iguana-17918	5	4	molecular	molecular	ADJ
iguana-17918	5	5	study	study	NOUN
iguana-17918	5	6	,	,	PUNCT
iguana-17918	5	7	keogh	keogh	PROPN
iguana-17918	5	8	et	et	PROPN
iguana-17918	5	9	al	al	PROPN
iguana-17918	5	10	.	.	PROPN
iguana-17918	6	1	(	(	PUNCT
iguana-17918	6	2	2008	2008	NUM
iguana-17918	6	3	.	.	PUNCT
iguana-17918	7	1	philosophical	philosophical	ADJ
iguana-17918	7	2	transactions	transaction	NOUN
iguana-17918	7	3	of	of	ADP
iguana-17918	7	4	the	the	DET
iguana-17918	7	5	royal	royal	ADJ
iguana-17918	7	6	society	society	PROPN
iguana-17918	7	7	b	b	PROPN
iguana-17918	7	8	(	(	PUNCT
iguana-17918	7	9	biological	biological	ADJ
iguana-17918	7	10	sciences	science	NOUN
iguana-17918	7	11	)	)	PUNCT
iguana-17918	7	12	363:3413–3426	363:3413–3426	NUM
iguana-17918	7	13	)	)	PUNCT
iguana-17918	7	14	showed	show	VERB
iguana-17918	7	15	that	that	SCONJ
iguana-17918	7	16	brachylophus	brachylophus	NOUN
iguana-17918	7	17	comprised	comprise	VERB
iguana-17918	7	18	three	three	NUM
iguana-17918	7	19	evolutionarily	evolutionarily	ADV
iguana-17918	7	20	significant	significant	ADJ
iguana-17918	7	21	units	unit	NOUN
iguana-17918	7	22	.	.	PUNCT
iguana-17918	8	1	to	to	PART
iguana-17918	8	2	test	test	VERB
iguana-17918	8	3	these	these	DET
iguana-17918	8	4	conclusions	conclusion	NOUN
iguana-17918	8	5	and	and	CCONJ
iguana-17918	8	6	to	to	PART
iguana-17918	8	7	reevaluate	reevaluate	VERB
iguana-17918	8	8	the	the	DET
iguana-17918	8	9	phylogenetic	phylogenetic	ADJ
iguana-17918	8	10	and	and	CCONJ
iguana-17918	8	11	biogeographic	biogeographic	ADJ
iguana-17918	8	12	relationships	relationship	NOUN
iguana-17918	8	13	within	within	ADP
iguana-17918	8	14	brachylophus	brachylophus	NOUN
iguana-17918	8	15	,	,	PUNCT
iguana-17918	8	16	the	the	DET
iguana-17918	8	17	authors	author	NOUN
iguana-17918	8	18	generated	generate	VERB
iguana-17918	8	19	a	a	DET
iguana-17918	8	20	mitochondrial	mitochondrial	ADJ
iguana-17918	8	21	dna	dna	NOUN
iguana-17918	8	22	dataset	dataset	VERB
iguana-17918	8	23	for	for	ADP
iguana-17918	8	24	61	61	NUM
iguana-17918	8	25	individuals	individual	NOUN
iguana-17918	8	26	from	from	ADP
iguana-17918	8	27	13	13	NUM
iguana-17918	8	28	islands	island	NOUN
iguana-17918	8	29	,	,	PUNCT
iguana-17918	8	30	representing	represent	VERB
iguana-17918	8	31	both	both	PRON
iguana-17918	8	32	currently	currently	ADV
iguana-17918	8	33	recognized	recognize	VERB
iguana-17918	8	34	species	specie	NOUN
iguana-17918	8	35	of	of	ADP
iguana-17918	8	36	brachylophus	brachylophus	NOUN
iguana-17918	8	37	.	.	PUNCT
iguana-17918	9	1	the	the	DET
iguana-17918	9	2	data	datum	NOUN
iguana-17918	9	3	rejected	reject	VERB
iguana-17918	9	4	the	the	DET
iguana-17918	9	5	monophyly	monophyly	NOUN
iguana-17918	9	6	of	of	ADP
iguana-17918	9	7	specimens	specimen	NOUN
iguana-17918	9	8	previously	previously	ADV
iguana-17918	9	9	believed	believe	VERB
iguana-17918	9	10	to	to	PART
iguana-17918	9	11	comprise	comprise	VERB
iguana-17918	9	12	b.	b.	PROPN
iguana-17918	9	13	fasciatus	fasciatus	PROPN
iguana-17918	9	14	,	,	PUNCT
iguana-17918	9	15	instead	instead	ADV
iguana-17918	9	16	demonstrating	demonstrate	VERB
iguana-17918	9	17	that	that	SCONJ
iguana-17918	9	18	brachylophus	brachylophus	NOUN
iguana-17918	9	19	includes	include	VERB
iguana-17918	9	20	three	three	NUM
iguana-17918	9	21	distinct	distinct	ADJ
iguana-17918	9	22	species	specie	NOUN
iguana-17918	9	23	:	:	PUNCT
iguana-17918	9	24	b.	b.	PROPN
iguana-17918	9	25	fasciatus	fasciatus	PROPN
iguana-17918	9	26	from	from	ADP
iguana-17918	9	27	the	the	DET
iguana-17918	9	28	lau	lau	PROPN
iguana-17918	9	29	group	group	NOUN
iguana-17918	9	30	of	of	ADP
iguana-17918	9	31	fiji	fiji	PROPN
iguana-17918	9	32	and	and	CCONJ
iguana-17918	9	33	tonga	tonga	PROPN
iguana-17918	9	34	,	,	PUNCT
iguana-17918	9	35	a	a	DET
iguana-17918	9	36	new	new	ADJ
iguana-17918	9	37	species	specie	NOUN
iguana-17918	9	38	,	,	PUNCT
iguana-17918	9	39	b.	b.	PROPN
iguana-17918	9	40	bulabula	bulabula	PROPN
iguana-17918	9	41	,	,	PUNCT
iguana-17918	9	42	from	from	ADP
iguana-17918	9	43	the	the	DET
iguana-17918	9	44	central	central	ADJ
iguana-17918	9	45	regions	region	NOUN
iguana-17918	9	46	of	of	ADP
iguana-17918	9	47	fiji	fiji	PROPN
iguana-17918	9	48	,	,	PUNCT
iguana-17918	9	49	populations	population	NOUN
iguana-17918	9	50	which	which	PRON
iguana-17918	9	51	until	until	ADP
iguana-17918	9	52	now	now	ADV
iguana-17918	9	53	were	be	AUX
iguana-17918	9	54	considered	consider	VERB
iguana-17918	9	55	to	to	PART
iguana-17918	9	56	be	be	AUX
iguana-17918	9	57	b.	b.	PROPN
iguana-17918	9	58	fasciatus	fasciatus	PROPN
iguana-17918	9	59	or	or	CCONJ
iguana-17918	9	60	b.	b.	PROPN
iguana-17918	9	61	vitiensis	vitiensis	PROPN
iguana-17918	9	62	.	.	PUNCT
iguana-17918	10	1	these	these	DET
iguana-17918	10	2	molecular	molecular	ADJ
iguana-17918	10	3	and	and	CCONJ
iguana-17918	10	4	taxonomic	taxonomic	ADJ
iguana-17918	10	5	results	result	NOUN
iguana-17918	10	6	have	have	VERB
iguana-17918	10	7	important	important	ADJ
iguana-17918	10	8	implications	implication	NOUN
iguana-17918	10	9	for	for	ADP
iguana-17918	10	10	future	future	ADJ
iguana-17918	10	11	conservation	conservation	NOUN
iguana-17918	10	12	initiatives	initiative	NOUN
iguana-17918	10	13	for	for	ADP
iguana-17918	10	14	pacific	pacific	ADJ
iguana-17918	10	15	iguanas	iguana	NOUN
iguana-17918	10	16	.	.	PUNCT
iguana-17918	11	1	emerging	emerge	VERB
iguana-17918	11	2	threats	threat	NOUN
iguana-17918	11	3	to	to	ADP
iguana-17918	11	4	long	long	ADV
iguana-17918	11	5	-	-	PUNCT
iguana-17918	11	6	lived	live	VERB
iguana-17918	11	7	vertebrates	vertebrate	NOUN
iguana-17918	11	8	persistent	persistent	ADJ
iguana-17918	11	9	contaminants	contaminant	NOUN
iguana-17918	11	10	are	be	AUX
iguana-17918	11	11	ubiquitous	ubiquitous	ADJ
iguana-17918	11	12	in	in	ADP
iguana-17918	11	13	the	the	DET
iguana-17918	11	14	environment	environment	NOUN
iguana-17918	11	15	,	,	PUNCT
iguana-17918	11	16	often	often	ADV
iguana-17918	11	17	present	present	ADJ
iguana-17918	11	18	at	at	ADP
iguana-17918	11	19	concentrations	concentration	NOUN
iguana-17918	11	20	that	that	PRON
iguana-17918	11	21	may	may	AUX
iguana-17918	11	22	jeopardize	jeopardize	VERB
iguana-17918	11	23	reproductive	reproductive	ADJ
iguana-17918	11	24	fitness	fitness	NOUN
iguana-17918	11	25	only	only	ADV
iguana-17918	11	26	after	after	ADP
iguana-17918	11	27	long	long	ADJ
iguana-17918	11	28	periods	period	NOUN
iguana-17918	11	29	of	of	ADP
iguana-17918	11	30	exposure	exposure	NOUN
iguana-17918	11	31	.	.	PUNCT
iguana-17918	12	1	as	as	SCONJ
iguana-17918	12	2	the	the	DET
iguana-17918	12	3	duration	duration	NOUN
iguana-17918	12	4	of	of	ADP
iguana-17918	12	5	exposure	exposure	NOUN
iguana-17918	12	6	is	be	AUX
iguana-17918	12	7	largely	largely	ADV
iguana-17918	12	8	regulated	regulate	VERB
iguana-17918	12	9	by	by	ADP
iguana-17918	12	10	life	life	NOUN
iguana-17918	12	11	span	span	NOUN
iguana-17918	12	12	,	,	PUNCT
iguana-17918	12	13	long	long	ADV
iguana-17918	12	14	-	-	PUNCT
iguana-17918	12	15	lived	live	VERB
iguana-17918	12	16	species	specie	NOUN
iguana-17918	12	17	of	of	ADP
iguana-17918	12	18	high	high	ADJ
iguana-17918	12	19	trophic	trophic	ADJ
iguana-17918	12	20	status	status	NOUN
iguana-17918	12	21	,	,	PUNCT
iguana-17918	12	22	such	such	ADJ
iguana-17918	12	23	as	as	ADP
iguana-17918	12	24	many	many	ADJ
iguana-17918	12	25	reptiles	reptile	NOUN
iguana-17918	12	26	,	,	PUNCT
iguana-17918	12	27	birds	bird	NOUN
iguana-17918	12	28	,	,	PUNCT
iguana-17918	12	29	and	and	CCONJ
iguana-17918	12	30	mammals	mammal	NOUN
iguana-17918	12	31	,	,	PUNCT
iguana-17918	12	32	may	may	AUX
iguana-17918	12	33	be	be	AUX
iguana-17918	12	34	at	at	ADP
iguana-17918	12	35	risk	risk	NOUN
iguana-17918	12	36	of	of	ADP
iguana-17918	12	37	reduced	reduce	VERB
iguana-17918	12	38	fitness	fitness	NOUN
iguana-17918	12	39	and	and	CCONJ
iguana-17918	12	40	population	population	NOUN
iguana-17918	12	41	decline	decline	NOUN
iguana-17918	12	42	.	.	PUNCT
iguana-17918	13	1	rowe	rowe	PROPN
iguana-17918	13	2	(	(	PUNCT
iguana-17918	13	3	2008	2008	NUM
iguana-17918	13	4	.	.	PUNCT
iguana-17918	14	1	bioscience	bioscience	NOUN
iguana-17918	14	2	58:623–631	58:623–631	NUM
iguana-17918	14	3	)	)	PUNCT
iguana-17918	14	4	suggested	suggest	VERB
iguana-17918	14	5	that	that	SCONJ
iguana-17918	14	6	delayed	delay	VERB
iguana-17918	14	7	maturation	maturation	NOUN
iguana-17918	14	8	and	and	CCONJ
iguana-17918	14	9	iteroparity	iteroparity	NOUN
iguana-17918	14	10	(	(	PUNCT
iguana-17918	14	11	repeated	repeat	VERB
iguana-17918	14	12	production	production	NOUN
iguana-17918	14	13	of	of	ADP
iguana-17918	14	14	offspring	offspring	NOUN
iguana-17918	14	15	at	at	ADP
iguana-17918	14	16	intervals	interval	NOUN
iguana-17918	14	17	throughout	throughout	ADP
iguana-17918	14	18	the	the	DET
iguana-17918	14	19	life	life	NOUN
iguana-17918	14	20	cycle	cycle	NOUN
iguana-17918	14	21	)	)	PUNCT
iguana-17918	14	22	confer	confer	VERB
iguana-17918	14	23	the	the	DET
iguana-17918	14	24	potential	potential	NOUN
iguana-17918	14	25	for	for	ADP
iguana-17918	14	26	cumulative	cumulative	ADJ
iguana-17918	14	27	effects	effect	NOUN
iguana-17918	14	28	to	to	PART
iguana-17918	14	29	be	be	AUX
iguana-17918	14	30	expressed	express	VERB
iguana-17918	14	31	prior	prior	ADV
iguana-17918	14	32	to	to	ADP
iguana-17918	14	33	reproduction	reproduction	NOUN
iguana-17918	14	34	,	,	PUNCT
iguana-17918	14	35	and	and	CCONJ
iguana-17918	14	36	large	large	ADJ
iguana-17918	14	37	parental	parental	ADJ
iguana-17918	14	38	investments	investment	NOUN
iguana-17918	14	39	in	in	ADP
iguana-17918	14	40	yolk	yolk	NOUN
iguana-17918	14	41	and	and	CCONJ
iguana-17918	14	42	milk	milk	NOUN
iguana-17918	14	43	may	may	AUX
iguana-17918	14	44	threaten	threaten	VERB
iguana-17918	14	45	offspring	offspring	NOUN
iguana-17918	14	46	because	because	SCONJ
iguana-17918	14	47	of	of	ADP
iguana-17918	14	48	exposure	exposure	NOUN
iguana-17918	14	49	during	during	ADP
iguana-17918	14	50	critical	critical	ADJ
iguana-17918	14	51	developmental	developmental	ADJ
iguana-17918	14	52	periods	period	NOUN
iguana-17918	14	53	.	.	PUNCT
iguana-17918	15	1	long	long	ADJ
iguana-17918	15	2	generation	generation	NOUN
iguana-17918	15	3	times	time	NOUN
iguana-17918	15	4	may	may	AUX
iguana-17918	15	5	delay	delay	VERB
iguana-17918	15	6	emergence	emergence	NOUN
iguana-17918	15	7	of	of	ADP
iguana-17918	15	8	obvious	obvious	ADJ
iguana-17918	15	9	effects	effect	NOUN
iguana-17918	15	10	on	on	ADP
iguana-17918	15	11	populations	population	NOUN
iguana-17918	15	12	,	,	PUNCT
iguana-17918	15	13	perhaps	perhaps	ADV
iguana-17918	15	14	eluding	elude	VERB
iguana-17918	15	15	early	early	ADJ
iguana-17918	15	16	intervention	intervention	NOUN
iguana-17918	15	17	,	,	PUNCT
iguana-17918	15	18	while	while	SCONJ
iguana-17918	15	19	constraining	constrain	VERB
iguana-17918	15	20	rates	rate	NOUN
iguana-17918	15	21	at	at	ADP
iguana-17918	15	22	which	which	PRON
iguana-17918	15	23	populations	population	NOUN
iguana-17918	15	24	may	may	AUX
iguana-17918	15	25	recover	recover	VERB
iguana-17918	15	26	if	if	SCONJ
iguana-17918	15	27	conditions	condition	NOUN
iguana-17918	15	28	subsequently	subsequently	ADV
iguana-17918	15	29	improve	improve	VERB
iguana-17918	15	30	.	.	PUNCT
iguana-17918	16	1	life	life	NOUN
iguana-17918	16	2	history	history	NOUN
iguana-17918	16	3	theory	theory	NOUN
iguana-17918	16	4	thus	thus	ADV
iguana-17918	16	5	suggests	suggest	VERB
iguana-17918	16	6	that	that	SCONJ
iguana-17918	16	7	the	the	DET
iguana-17918	16	8	suite	suite	NOUN
iguana-17918	16	9	of	of	ADP
iguana-17918	16	10	traits	trait	NOUN
iguana-17918	16	11	that	that	PRON
iguana-17918	16	12	optimized	optimize	VERB
iguana-17918	16	13	reproductive	reproductive	ADJ
iguana-17918	16	14	fitness	fitness	NOUN
iguana-17918	16	15	throughout	throughout	ADP
iguana-17918	16	16	long	long	ADV
iguana-17918	16	17	-	-	PUNCT
iguana-17918	16	18	lived	live	VERB
iguana-17918	16	19	species	specie	NOUN
iguana-17918	16	20	’	'	PUNCT
iguana-17918	16	21	evolutionary	evolutionary	ADJ
iguana-17918	16	22	histories	history	NOUN
iguana-17918	16	23	may	may	AUX
iguana-17918	16	24	ultimately	ultimately	ADV
iguana-17918	16	25	put	put	VERB
iguana-17918	16	26	them	they	PRON
iguana-17918	16	27	in	in	ADP
iguana-17918	16	28	peril	peril	NOUN
iguana-17918	16	29	in	in	ADP
iguana-17918	16	30	the	the	DET
iguana-17918	16	31	modern	modern	ADJ
iguana-17918	16	32	anthropogenically	anthropogenically	ADV
iguana-17918	16	33	altered	altered	ADJ
iguana-17918	16	34	environment	environment	NOUN
iguana-17918	16	35	.	.	PUNCT
iguana-17918	17	1	frogs	frog	NOUN
iguana-17918	17	2	with	with	ADP
iguana-17918	17	3	disease	disease	NOUN
iguana-17918	17	4	-	-	PUNCT
iguana-17918	17	5	resistance	resistance	NOUN
iguana-17918	17	6	genes	gene	NOUN
iguana-17918	17	7	may	may	AUX
iguana-17918	17	8	escape	escape	VERB
iguana-17918	17	9	extinction	extinction	NOUN
iguana-17918	17	10	as	as	SCONJ
iguana-17918	17	11	frog	frog	ADJ
iguana-17918	17	12	populations	population	NOUN
iguana-17918	17	13	die	die	VERB
iguana-17918	17	14	off	off	ADP
iguana-17918	17	15	around	around	ADP
iguana-17918	17	16	the	the	DET
iguana-17918	17	17	world	world	NOUN
iguana-17918	17	18	,	,	PUNCT
iguana-17918	17	19	researchers	researcher	NOUN
iguana-17918	17	20	have	have	AUX
iguana-17918	17	21	identified	identify	VERB
iguana-17918	17	22	certain	certain	ADJ
iguana-17918	17	23	genes	gene	NOUN
iguana-17918	17	24	that	that	PRON
iguana-17918	17	25	can	can	AUX
iguana-17918	17	26	help	help	VERB
iguana-17918	17	27	the	the	DET
iguana-17918	17	28	amphibians	amphibian	NOUN
iguana-17918	17	29	develop	develop	VERB
iguana-17918	17	30	resistance	resistance	NOUN
iguana-17918	17	31	to	to	ADP
iguana-17918	17	32	harmful	harmful	ADJ
iguana-17918	17	33	bacteria	bacteria	NOUN
iguana-17918	17	34	and	and	CCONJ
iguana-17918	17	35	disease	disease	NOUN
iguana-17918	17	36	.	.	PUNCT
iguana-17918	18	1	the	the	DET
iguana-17918	18	2	discovery	discovery	NOUN
iguana-17918	18	3	may	may	AUX
iguana-17918	18	4	provide	provide	VERB
iguana-17918	18	5	new	new	ADJ
iguana-17918	18	6	strategies	strategy	NOUN
iguana-17918	18	7	to	to	PART
iguana-17918	18	8	protect	protect	VERB
iguana-17918	18	9	frog	frog	ADJ
iguana-17918	18	10	populations	population	NOUN
iguana-17918	18	11	in	in	ADP
iguana-17918	18	12	the	the	DET
iguana-17918	18	13	wild	wild	NOUN
iguana-17918	18	14	.	.	PUNCT
iguana-17918	19	1	new	new	ADJ
iguana-17918	19	2	research	research	NOUN
iguana-17918	19	3	examines	examine	VERB
iguana-17918	19	4	how	how	SCONJ
iguana-17918	19	5	genes	gene	NOUN
iguana-17918	19	6	encoding	encode	VERB
iguana-17918	19	7	the	the	DET
iguana-17918	19	8	major	major	ADJ
iguana-17918	19	9	histocompatibility	histocompatibility	NOUN
iguana-17918	19	10	(	(	PUNCT
iguana-17918	19	11	mhc	mhc	NOUN
iguana-17918	19	12	)	)	PUNCT
iguana-17918	19	13	complex	complex	NOUN
iguana-17918	19	14	affect	affect	VERB
iguana-17918	19	15	the	the	DET
iguana-17918	19	16	ability	ability	NOUN
iguana-17918	19	17	of	of	ADP
iguana-17918	19	18	frogs	frog	NOUN
iguana-17918	19	19	to	to	PART
iguana-17918	19	20	resist	resist	VERB
iguana-17918	19	21	infection	infection	NOUN
iguana-17918	19	22	by	by	ADP
iguana-17918	19	23	a	a	DET
iguana-17918	19	24	bacterium	bacterium	NOUN
iguana-17918	19	25	that	that	PRON
iguana-17918	19	26	is	be	AUX
iguana-17918	19	27	commonly	commonly	ADV
iguana-17918	19	28	associated	associate	VERB
iguana-17918	19	29	with	with	ADP
iguana-17918	19	30	frog	frog	ADJ
iguana-17918	19	31	population	population	NOUN
iguana-17918	19	32	declines	decline	VERB
iguana-17918	19	33	.	.	PUNCT
iguana-17918	20	1	“	"	PUNCT
iguana-17918	20	2	in	in	ADP
iguana-17918	20	3	the	the	DET
iguana-17918	20	4	short	short	ADJ
iguana-17918	20	5	term	term	NOUN
iguana-17918	20	6	,	,	PUNCT
iguana-17918	20	7	captive	captive	ADJ
iguana-17918	20	8	management	management	NOUN
iguana-17918	20	9	of	of	ADP
iguana-17918	20	10	frogs	frog	NOUN
iguana-17918	20	11	with	with	ADP
iguana-17918	20	12	complementary	complementary	ADJ
iguana-17918	20	13	diseaseresistance	diseaseresistance	NOUN
iguana-17918	20	14	genes	gene	NOUN
iguana-17918	20	15	may	may	AUX
iguana-17918	20	16	offer	offer	VERB
iguana-17918	20	17	the	the	DET
iguana-17918	20	18	best	good	ADJ
iguana-17918	20	19	hope	hope	NOUN
iguana-17918	20	20	for	for	ADP
iguana-17918	20	21	saving	save	VERB
iguana-17918	20	22	species	specie	NOUN
iguana-17918	20	23	from	from	ADP
iguana-17918	20	24	extinction	extinction	NOUN
iguana-17918	20	25	,	,	PUNCT
iguana-17918	20	26	”	"	PUNCT
iguana-17918	20	27	says	say	VERB
iguana-17918	20	28	bruce	bruce	PROPN
iguana-17918	20	29	waldman	waldman	PROPN
iguana-17918	20	30	,	,	PUNCT
iguana-17918	20	31	a	a	DET
iguana-17918	20	32	biologist	biologist	NOUN
iguana-17918	20	33	at	at	ADP
iguana-17918	20	34	lincoln	lincoln	PROPN
iguana-17918	20	35	university	university	PROPN
iguana-17918	20	36	in	in	ADP
iguana-17918	20	37	new	new	PROPN
iguana-17918	20	38	zealand	zealand	PROPN
iguana-17918	20	39	and	and	CCONJ
iguana-17918	20	40	one	one	NUM
iguana-17918	20	41	of	of	ADP
iguana-17918	20	42	the	the	DET
iguana-17918	20	43	paper	paper	NOUN
iguana-17918	20	44	’s	’s	PART
iguana-17918	20	45	authors	author	NOUN
iguana-17918	20	46	.	.	PUNCT
iguana-17918	21	1	“	"	PUNCT
iguana-17918	21	2	management	management	NOUN
iguana-17918	21	3	practices	practice	NOUN
iguana-17918	21	4	that	that	PRON
iguana-17918	21	5	maintain	maintain	VERB
iguana-17918	21	6	or	or	CCONJ
iguana-17918	21	7	enhance	enhance	VERB
iguana-17918	21	8	diversity	diversity	NOUN
iguana-17918	21	9	in	in	ADP
iguana-17918	21	10	mhc	mhc	PROPN
iguana-17918	21	11	genes	gene	NOUN
iguana-17918	21	12	may	may	AUX
iguana-17918	21	13	prove	prove	VERB
iguana-17918	21	14	the	the	DET
iguana-17918	21	15	key	key	NOUN
iguana-17918	21	16	to	to	ADP
iguana-17918	21	17	safeguarding	safeguard	VERB
iguana-17918	21	18	frog	frog	ADJ
iguana-17918	21	19	populations	population	NOUN
iguana-17918	21	20	in	in	ADP
iguana-17918	21	21	the	the	DET
iguana-17918	21	22	wild	wild	NOUN
iguana-17918	21	23	.	.	PUNCT
iguana-17918	21	24	”	"	PUNCT
iguana-17918	22	1	“	"	PUNCT
iguana-17918	22	2	massive	massive	ADJ
iguana-17918	22	3	die	die	NOUN
iguana-17918	22	4	-	-	PUNCT
iguana-17918	22	5	offs	off	NOUN
iguana-17918	22	6	of	of	ADP
iguana-17918	22	7	frogs	frog	NOUN
iguana-17918	22	8	may	may	AUX
iguana-17918	22	9	indicate	indicate	VERB
iguana-17918	22	10	environmental	environmental	ADJ
iguana-17918	22	11	problems	problem	NOUN
iguana-17918	22	12	that	that	PRON
iguana-17918	22	13	ultimately	ultimately	ADV
iguana-17918	22	14	will	will	AUX
iguana-17918	22	15	affect	affect	VERB
iguana-17918	22	16	other	other	ADJ
iguana-17918	22	17	species	specie	NOUN
iguana-17918	22	18	,	,	PUNCT
iguana-17918	22	19	including	include	VERB
iguana-17918	22	20	humans	human	NOUN
iguana-17918	22	21	,	,	PUNCT
iguana-17918	22	22	”	"	PUNCT
iguana-17918	22	23	waldman	waldman	PROPN
iguana-17918	22	24	says	say	VERB
iguana-17918	22	25	.	.	PUNCT
iguana-17918	23	1	“	"	PUNCT
iguana-17918	23	2	but	but	CCONJ
iguana-17918	23	3	,	,	PUNCT
iguana-17918	23	4	despite	despite	SCONJ
iguana-17918	23	5	the	the	DET
iguana-17918	23	6	concern	concern	NOUN
iguana-17918	23	7	,	,	PUNCT
iguana-17918	23	8	little	little	ADJ
iguana-17918	23	9	is	be	AUX
iguana-17918	23	10	known	know	VERB
iguana-17918	23	11	about	about	ADP
iguana-17918	23	12	factors	factor	NOUN
iguana-17918	23	13	that	that	PRON
iguana-17918	23	14	make	make	VERB
iguana-17918	23	15	individuals	individual	NOUN
iguana-17918	23	16	susceptible	susceptible	ADJ
iguana-17918	23	17	to	to	ADP
iguana-17918	23	18	disease	disease	NOUN
iguana-17918	23	19	.	.	PUNCT
iguana-17918	23	20	”	"	PUNCT
iguana-17918	24	1	c	c	NOUN
iguana-17918	24	2	o	o	NOUN
iguana-17918	24	3	n	n	X
iguana-17918	24	4	s	s	X
iguana-17918	24	5	e	e	NOUN
iguana-17918	24	6	r	r	NOUN
iguana-17918	24	7	v	v	PROPN
iguana-17918	24	8	a	a	DET
iguana-17918	24	9	t	t	NOUN
iguana-17918	25	1	i	i	PRON
iguana-17918	25	2	o	o	VERB
iguana-17918	26	1	n	n	CCONJ
iguana-17918	26	2	r	r	NOUN
iguana-17918	26	3	e	e	NOUN
iguana-17918	26	4	s	s	X
iguana-17918	26	5	e	e	X
iguana-17918	26	6	a	a	DET
iguana-17918	26	7	r	r	NOUN
iguana-17918	27	1	c	c	NOUN
iguana-17918	27	2	h	h	NOUN
iguana-17918	27	3	r	r	NOUN
iguana-17918	27	4	e	e	NOUN
iguana-17918	27	5	p	p	NOUN
iguana-17918	27	6	o	o	NOUN
iguana-17918	27	7	r	r	NOUN
iguana-17918	27	8	t	t	NOUN
iguana-17918	27	9	s	s	NOUN
iguana-17918	27	10	populations	population	NOUN
iguana-17918	27	11	from	from	ADP
iguana-17918	27	12	central	central	ADJ
iguana-17918	27	13	regions	region	NOUN
iguana-17918	27	14	of	of	ADP
iguana-17918	27	15	fiji	fiji	PROPN
iguana-17918	27	16	,	,	PUNCT
iguana-17918	27	17	formerly	formerly	ADV
iguana-17918	27	18	assigned	assign	VERB
iguana-17918	27	19	to	to	AUX
iguana-17918	27	20	brachylophus	brachylophus	VERB
iguana-17918	27	21	fasciatus	fasciatus	NOUN
iguana-17918	27	22	,	,	PUNCT
iguana-17918	27	23	were	be	AUX
iguana-17918	27	24	shown	show	VERB
iguana-17918	27	25	to	to	PART
iguana-17918	27	26	represent	represent	VERB
iguana-17918	27	27	a	a	DET
iguana-17918	27	28	new	new	ADJ
iguana-17918	27	29	species	specie	NOUN
iguana-17918	27	30	,	,	PUNCT
iguana-17918	27	31	b.	b.	PROPN
iguana-17918	27	32	bulabula	bulabula	PROPN
iguana-17918	27	33	.	.	PUNCT
iguana-17918	28	1	pe	pe	VERB
iguana-17918	29	1	te	te	ADP
iguana-17918	29	2	r	r	NOUN
iguana-17918	29	3	h	h	NOUN
iguana-17918	30	1	a	a	DET
iguana-17918	30	2	r	r	NOUN
iguana-17918	30	3	lo	lo	PROPN
iguana-17918	30	4	w	w	NOUN
iguana-17918	30	5	top	top	ADJ
iguana-17918	30	6	predators	predator	NOUN
iguana-17918	30	7	such	such	ADJ
iguana-17918	30	8	as	as	ADP
iguana-17918	30	9	nile	nile	PROPN
iguana-17918	30	10	monitors	monitor	NOUN
iguana-17918	30	11	(	(	PUNCT
iguana-17918	30	12	varanus	varanus	NOUN
iguana-17918	30	13	niloticus	niloticus	PROPN
iguana-17918	30	14	)	)	PUNCT
iguana-17918	30	15	may	may	AUX
iguana-17918	30	16	be	be	AUX
iguana-17918	30	17	at	at	ADP
iguana-17918	30	18	risk	risk	NOUN
iguana-17918	30	19	of	of	ADP
iguana-17918	30	20	reduced	reduce	VERB
iguana-17918	30	21	fitness	fitness	NOUN
iguana-17918	30	22	and	and	CCONJ
iguana-17918	30	23	population	population	NOUN
iguana-17918	30	24	declines	decline	VERB
iguana-17918	30	25	due	due	ADJ
iguana-17918	30	26	to	to	ADP
iguana-17918	30	27	cumulative	cumulative	ADJ
iguana-17918	30	28	effects	effect	NOUN
iguana-17918	30	29	of	of	ADP
iguana-17918	30	30	persistent	persistent	ADJ
iguana-17918	30	31	environmental	environmental	ADJ
iguana-17918	30	32	contaminants	contaminant	NOUN
iguana-17918	30	33	to	to	PART
iguana-17918	30	34	which	which	PRON
iguana-17918	30	35	they	they	PRON
iguana-17918	30	36	are	be	AUX
iguana-17918	30	37	exposed	expose	VERB
iguana-17918	30	38	over	over	ADP
iguana-17918	30	39	long	long	ADJ
iguana-17918	30	40	life	life	NOUN
iguana-17918	30	41	spans	span	NOUN
iguana-17918	30	42	.	.	PUNCT
iguana-17918	31	1	b	b	X
iguana-17918	31	2	r	r	NOUN
iguana-17918	31	3	u	u	NOUN
iguana-17918	31	4	c	c	NOUN
iguana-17918	31	5	e	e	NOUN
iguana-17918	31	6	b	b	PROPN
iguana-17918	32	1	lo	lo	NOUN
iguana-17918	32	2	c	c	NOUN
iguana-17918	33	1	k	k	PROPN
iguana-17918	33	2	246	246	NUM
iguana-17918	33	3	iguana	iguana	NOUN
iguana-17918	33	4	•	•	NUM
iguana-17918	33	5	volume	volume	NOUN
iguana-17918	33	6	15	15	NUM
iguana-17918	33	7	,	,	PUNCT
iguana-17918	33	8	number	number	NOUN
iguana-17918	33	9	4	4	NUM
iguana-17918	33	10	•	•	NOUN
iguana-17918	33	11	december	december	PROPN
iguana-17918	33	12	2008	2008	NUM
iguana-17918	33	13	conservation	conservation	NOUN
iguana-17918	33	14	research	research	NOUN
iguana-17918	33	15	reports	report	NOUN
iguana-17918	33	16	barribeau	barribeau	VERB
iguana-17918	33	17	et	et	PROPN
iguana-17918	33	18	al	al	PROPN
iguana-17918	33	19	.	.	PUNCT
iguana-17918	34	1	(	(	PUNCT
iguana-17918	34	2	plos	plos	PROPN
iguana-17918	34	3	one	one	NUM
iguana-17918	34	4	3	3	NUM
iguana-17918	34	5	:	:	PUNCT
iguana-17918	34	6	e2692	e2692	NOUN
iguana-17918	34	7	,	,	PUNCT
iguana-17918	34	8	<	<	X
iguana-17918	34	9	www.plosone.org/article/info	www.plosone.org/article/info	NOUN
iguana-17918	34	10	:	:	PUNCT
iguana-17918	34	11	doi/10.1371	doi/10.1371	NOUN
iguana-17918	34	12	/	/	SYM
iguana-17918	34	13	journal.pone.0002692	journal.pone.0002692	PROPN
iguana-17918	34	14	>	>	NOUN
iguana-17918	34	15	)	)	PUNCT
iguana-17918	34	16	exposed	expose	VERB
iguana-17918	34	17	african	african	ADJ
iguana-17918	34	18	clawed	claw	VERB
iguana-17918	34	19	frog	frog	ADJ
iguana-17918	34	20	tadpoles	tadpole	NOUN
iguana-17918	34	21	to	to	ADP
iguana-17918	34	22	several	several	ADJ
iguana-17918	34	23	doses	dose	NOUN
iguana-17918	34	24	of	of	ADP
iguana-17918	34	25	the	the	DET
iguana-17918	34	26	bacterium	bacterium	NOUN
iguana-17918	34	27	aeromonas	aeromonas	PROPN
iguana-17918	34	28	hydrophila	hydrophila	VERB
iguana-17918	34	29	and	and	CCONJ
iguana-17918	34	30	examined	examine	VERB
iguana-17918	34	31	the	the	DET
iguana-17918	34	32	number	number	NOUN
iguana-17918	34	33	of	of	ADP
iguana-17918	34	34	tadpoles	tadpole	NOUN
iguana-17918	34	35	that	that	PRON
iguana-17918	34	36	survived	survive	VERB
iguana-17918	34	37	and	and	CCONJ
iguana-17918	34	38	measured	measure	VERB
iguana-17918	34	39	how	how	SCONJ
iguana-17918	34	40	fast	fast	ADV
iguana-17918	34	41	they	they	PRON
iguana-17918	34	42	grew	grow	VERB
iguana-17918	34	43	.	.	PUNCT
iguana-17918	35	1	certain	certain	ADJ
iguana-17918	35	2	genes	gene	NOUN
iguana-17918	35	3	allowed	allow	VERB
iguana-17918	35	4	tadpoles	tadpole	NOUN
iguana-17918	35	5	to	to	PART
iguana-17918	35	6	survive	survive	VERB
iguana-17918	35	7	bacterial	bacterial	ADJ
iguana-17918	35	8	infection	infection	NOUN
iguana-17918	35	9	but	but	CCONJ
iguana-17918	35	10	at	at	ADP
iguana-17918	35	11	a	a	DET
iguana-17918	35	12	cost	cost	NOUN
iguana-17918	35	13	,	,	PUNCT
iguana-17918	35	14	as	as	SCONJ
iguana-17918	35	15	these	these	DET
iguana-17918	35	16	tadpoles	tadpole	NOUN
iguana-17918	35	17	sometimes	sometimes	ADV
iguana-17918	35	18	grew	grow	VERB
iguana-17918	35	19	more	more	ADV
iguana-17918	35	20	slowly	slowly	ADV
iguana-17918	35	21	.	.	PUNCT
iguana-17918	36	1	among	among	ADP
iguana-17918	36	2	siblings	sibling	NOUN
iguana-17918	36	3	,	,	PUNCT
iguana-17918	36	4	patterns	pattern	NOUN
iguana-17918	36	5	of	of	ADP
iguana-17918	36	6	disease	disease	NOUN
iguana-17918	36	7	resistance	resistance	NOUN
iguana-17918	36	8	corresponded	correspond	VERB
iguana-17918	36	9	to	to	ADP
iguana-17918	36	10	tadpoles	tadpole	NOUN
iguana-17918	36	11	’	'	PUNCT
iguana-17918	36	12	mhc	mhc	PROPN
iguana-17918	36	13	genes	gene	NOUN
iguana-17918	36	14	rather	rather	ADV
iguana-17918	36	15	than	than	ADP
iguana-17918	36	16	other	other	ADJ
iguana-17918	36	17	genes	gene	NOUN
iguana-17918	36	18	that	that	PRON
iguana-17918	36	19	they	they	PRON
iguana-17918	36	20	shared	share	VERB
iguana-17918	36	21	,	,	PUNCT
iguana-17918	36	22	demonstrating	demonstrate	VERB
iguana-17918	36	23	that	that	SCONJ
iguana-17918	36	24	the	the	DET
iguana-17918	36	25	mhc	mhc	PROPN
iguana-17918	36	26	genes	gene	NOUN
iguana-17918	36	27	conferred	confer	VERB
iguana-17918	36	28	immunity	immunity	NOUN
iguana-17918	36	29	.	.	PUNCT
iguana-17918	37	1	programs	program	NOUN
iguana-17918	37	2	currently	currently	ADV
iguana-17918	37	3	are	be	AUX
iguana-17918	37	4	underway	underway	ADJ
iguana-17918	37	5	to	to	PART
iguana-17918	37	6	rescue	rescue	NOUN
iguana-17918	37	7	frogs	frog	NOUN
iguana-17918	37	8	from	from	ADP
iguana-17918	37	9	declining	decline	VERB
iguana-17918	37	10	wild	wild	ADJ
iguana-17918	37	11	populations	population	NOUN
iguana-17918	37	12	and	and	CCONJ
iguana-17918	37	13	breed	breed	VERB
iguana-17918	37	14	them	they	PRON
iguana-17918	37	15	in	in	ADP
iguana-17918	37	16	captivity	captivity	NOUN
iguana-17918	37	17	to	to	PART
iguana-17918	37	18	ensure	ensure	VERB
iguana-17918	37	19	that	that	SCONJ
iguana-17918	37	20	species	specie	NOUN
iguana-17918	37	21	are	be	AUX
iguana-17918	37	22	not	not	PART
iguana-17918	37	23	lost	lose	VERB
iguana-17918	37	24	to	to	ADP
iguana-17918	37	25	extinction	extinction	NOUN
iguana-17918	37	26	.	.	PUNCT
iguana-17918	38	1	this	this	DET
iguana-17918	38	2	study	study	NOUN
iguana-17918	38	3	suggests	suggest	VERB
iguana-17918	38	4	that	that	SCONJ
iguana-17918	38	5	selective	selective	ADJ
iguana-17918	38	6	breeding	breeding	NOUN
iguana-17918	38	7	of	of	ADP
iguana-17918	38	8	individuals	individual	NOUN
iguana-17918	38	9	with	with	ADP
iguana-17918	38	10	known	know	VERB
iguana-17918	38	11	disease	disease	NOUN
iguana-17918	38	12	-	-	PUNCT
iguana-17918	38	13	resistance	resistance	NOUN
iguana-17918	38	14	genes	gene	NOUN
iguana-17918	38	15	might	might	AUX
iguana-17918	38	16	produce	produce	VERB
iguana-17918	38	17	frogs	frog	NOUN
iguana-17918	38	18	that	that	PRON
iguana-17918	38	19	can	can	AUX
iguana-17918	38	20	survive	survive	VERB
iguana-17918	38	21	infection	infection	NOUN
iguana-17918	38	22	by	by	ADP
iguana-17918	38	23	pathogens	pathogen	NOUN
iguana-17918	38	24	,	,	PUNCT
iguana-17918	38	25	even	even	ADV
iguana-17918	38	26	after	after	SCONJ
iguana-17918	38	27	the	the	DET
iguana-17918	38	28	frogs	frog	NOUN
iguana-17918	38	29	are	be	AUX
iguana-17918	38	30	reintroduced	reintroduce	VERB
iguana-17918	38	31	into	into	ADP
iguana-17918	38	32	the	the	DET
iguana-17918	38	33	wild	wild	NOUN
iguana-17918	38	34	.	.	PUNCT
iguana-17918	39	1	the	the	DET
iguana-17918	39	2	research	research	NOUN
iguana-17918	39	3	team	team	NOUN
iguana-17918	39	4	studied	study	VERB
iguana-17918	39	5	the	the	DET
iguana-17918	39	6	african	african	PROPN
iguana-17918	39	7	clawed	claw	VERB
iguana-17918	39	8	frog	frog	NOUN
iguana-17918	39	9	because	because	SCONJ
iguana-17918	39	10	its	its	PRON
iguana-17918	39	11	immune	immune	ADJ
iguana-17918	39	12	system	system	NOUN
iguana-17918	39	13	already	already	ADV
iguana-17918	39	14	had	have	AUX
iguana-17918	39	15	been	be	AUX
iguana-17918	39	16	well	well	ADV
iguana-17918	39	17	characterized	characterize	VERB
iguana-17918	39	18	,	,	PUNCT
iguana-17918	39	19	but	but	CCONJ
iguana-17918	39	20	as	as	SCONJ
iguana-17918	39	21	most	most	ADJ
iguana-17918	39	22	frogs	frog	NOUN
iguana-17918	39	23	and	and	CCONJ
iguana-17918	39	24	toads	toad	NOUN
iguana-17918	39	25	have	have	VERB
iguana-17918	39	26	similar	similar	ADJ
iguana-17918	39	27	immune	immune	ADJ
iguana-17918	39	28	systems	system	NOUN
iguana-17918	39	29	,	,	PUNCT
iguana-17918	39	30	they	they	PRON
iguana-17918	39	31	believe	believe	VERB
iguana-17918	39	32	that	that	SCONJ
iguana-17918	39	33	their	their	PRON
iguana-17918	39	34	results	result	NOUN
iguana-17918	39	35	will	will	AUX
iguana-17918	39	36	be	be	AUX
iguana-17918	39	37	generally	generally	ADV
iguana-17918	39	38	applicable	applicable	ADJ
iguana-17918	39	39	to	to	ADP
iguana-17918	39	40	all	all	DET
iguana-17918	39	41	threatened	threaten	VERB
iguana-17918	39	42	and	and	CCONJ
iguana-17918	39	43	endangered	endangered	ADJ
iguana-17918	39	44	amphibians	amphibian	NOUN
iguana-17918	39	45	.	.	PUNCT
iguana-17918	40	1	new	new	ADJ
iguana-17918	40	2	approach	approach	NOUN
iguana-17918	40	3	to	to	ADP
iguana-17918	40	4	conserving	conserve	VERB
iguana-17918	40	5	amphibians	amphibian	NOUN
iguana-17918	40	6	the	the	DET
iguana-17918	40	7	global	global	ADJ
iguana-17918	40	8	extinction	extinction	NOUN
iguana-17918	40	9	crisis	crisis	NOUN
iguana-17918	40	10	demands	demand	VERB
iguana-17918	40	11	immediate	immediate	ADJ
iguana-17918	40	12	action	action	NOUN
iguana-17918	40	13	to	to	PART
iguana-17918	40	14	conserve	conserve	VERB
iguana-17918	40	15	species	specie	NOUN
iguana-17918	40	16	at	at	ADP
iguana-17918	40	17	risk	risk	NOUN
iguana-17918	40	18	.	.	PUNCT
iguana-17918	41	1	however	however	ADV
iguana-17918	41	2	,	,	PUNCT
iguana-17918	41	3	if	if	SCONJ
iguana-17918	41	4	entire	entire	ADJ
iguana-17918	41	5	groups	group	NOUN
iguana-17918	41	6	such	such	ADJ
iguana-17918	41	7	as	as	ADP
iguana-17918	41	8	superfamilies	superfamily	NOUN
iguana-17918	41	9	are	be	AUX
iguana-17918	41	10	at	at	ADP
iguana-17918	41	11	risk	risk	NOUN
iguana-17918	41	12	due	due	ADP
iguana-17918	41	13	to	to	ADP
iguana-17918	41	14	shared	share	VERB
iguana-17918	41	15	evolutionary	evolutionary	ADJ
iguana-17918	41	16	history	history	NOUN
iguana-17918	41	17	,	,	PUNCT
iguana-17918	41	18	a	a	DET
iguana-17918	41	19	shift	shift	NOUN
iguana-17918	41	20	towards	towards	ADP
iguana-17918	41	21	conserving	conserve	VERB
iguana-17918	41	22	such	such	ADJ
iguana-17918	41	23	groups	group	NOUN
iguana-17918	41	24	rather	rather	ADV
iguana-17918	41	25	than	than	ADP
iguana-17918	41	26	individual	individual	ADJ
iguana-17918	41	27	species	specie	NOUN
iguana-17918	41	28	may	may	AUX
iguana-17918	41	29	be	be	AUX
iguana-17918	41	30	needed	need	VERB
iguana-17918	41	31	.	.	PUNCT
iguana-17918	42	1	using	use	VERB
iguana-17918	42	2	phylogenetic	phylogenetic	ADJ
iguana-17918	42	3	autocorrelation	autocorrelation	NOUN
iguana-17918	42	4	analysis	analysis	NOUN
iguana-17918	42	5	,	,	PUNCT
iguana-17918	42	6	corey	corey	PROPN
iguana-17918	42	7	and	and	CCONJ
iguana-17918	42	8	waite	waite	PROPN
iguana-17918	42	9	(	(	PUNCT
iguana-17918	42	10	2008	2008	NUM
iguana-17918	42	11	.	.	PUNCT
iguana-17918	43	1	diversity	diversity	NOUN
iguana-17918	43	2	and	and	CCONJ
iguana-17918	43	3	distributions	distribution	NOUN
iguana-17918	43	4	14:614–629	14:614–629	NUM
iguana-17918	43	5	)	)	PUNCT
iguana-17918	43	6	demonstrated	demonstrate	VERB
iguana-17918	43	7	that	that	SCONJ
iguana-17918	43	8	multiple	multiple	ADJ
iguana-17918	43	9	kinds	kind	NOUN
iguana-17918	43	10	of	of	ADP
iguana-17918	43	11	extinction	extinction	NOUN
iguana-17918	43	12	threat	threat	NOUN
iguana-17918	43	13	clump	clump	NOUN
iguana-17918	43	14	within	within	ADP
iguana-17918	43	15	the	the	DET
iguana-17918	43	16	amphibian	amphibian	ADJ
iguana-17918	43	17	tree	tree	NOUN
iguana-17918	43	18	of	of	ADP
iguana-17918	43	19	life	life	NOUN
iguana-17918	43	20	.	.	PUNCT
iguana-17918	44	1	their	their	PRON
iguana-17918	44	2	study	study	NOUN
iguana-17918	44	3	provided	provide	VERB
iguana-17918	44	4	insight	insight	NOUN
iguana-17918	44	5	into	into	ADP
iguana-17918	44	6	how	how	SCONJ
iguana-17918	44	7	these	these	DET
iguana-17918	44	8	threats	threat	NOUN
iguana-17918	44	9	might	might	AUX
iguana-17918	44	10	collectively	collectively	ADV
iguana-17918	44	11	influence	influence	VERB
iguana-17918	44	12	the	the	DET
iguana-17918	44	13	extinction	extinction	NOUN
iguana-17918	44	14	risk	risk	NOUN
iguana-17918	44	15	of	of	ADP
iguana-17918	44	16	whole	whole	ADJ
iguana-17918	44	17	groups	group	NOUN
iguana-17918	44	18	,	,	PUNCT
iguana-17918	44	19	consistent	consistent	ADJ
iguana-17918	44	20	with	with	ADP
iguana-17918	44	21	the	the	DET
iguana-17918	44	22	supposition	supposition	NOUN
iguana-17918	44	23	that	that	SCONJ
iguana-17918	44	24	related	related	ADJ
iguana-17918	44	25	species	specie	NOUN
iguana-17918	44	26	,	,	PUNCT
iguana-17918	44	27	with	with	ADP
iguana-17918	44	28	similar	similar	ADJ
iguana-17918	44	29	traits	trait	NOUN
iguana-17918	44	30	,	,	PUNCT
iguana-17918	44	31	share	share	VERB
iguana-17918	44	32	an	an	DET
iguana-17918	44	33	intrinsic	intrinsic	ADJ
iguana-17918	44	34	vulnerability	vulnerability	NOUN
iguana-17918	44	35	to	to	ADP
iguana-17918	44	36	common	common	ADJ
iguana-17918	44	37	kinds	kind	NOUN
iguana-17918	44	38	of	of	ADP
iguana-17918	44	39	threat	threat	NOUN
iguana-17918	44	40	.	.	PUNCT
iguana-17918	45	1	most	most	ADV
iguana-17918	45	2	strikingly	strikingly	ADV
iguana-17918	45	3	,	,	PUNCT
iguana-17918	45	4	they	they	PRON
iguana-17918	45	5	found	find	VERB
iguana-17918	45	6	a	a	DET
iguana-17918	45	7	significant	significant	ADJ
iguana-17918	45	8	concentration	concentration	NOUN
iguana-17918	45	9	of	of	ADP
iguana-17918	45	10	‘	'	PUNCT
iguana-17918	45	11	enigmatic	enigmatic	ADJ
iguana-17918	45	12	’	'	PUNCT
iguana-17918	45	13	decline	decline	NOUN
iguana-17918	45	14	and	and	CCONJ
iguana-17918	45	15	critically	critically	ADV
iguana-17918	45	16	endangered	endanger	VERB
iguana-17918	45	17	status	status	NOUN
iguana-17918	45	18	within	within	ADP
iguana-17918	45	19	families	family	NOUN
iguana-17918	45	20	of	of	ADP
iguana-17918	45	21	the	the	DET
iguana-17918	45	22	hyloid	hyloid	ADJ
iguana-17918	45	23	frogs	frog	NOUN
iguana-17918	45	24	.	.	PUNCT
iguana-17918	46	1	this	this	DET
iguana-17918	46	2	phylogenetic	phylogenetic	ADJ
iguana-17918	46	3	clumping	clumping	NOUN
iguana-17918	46	4	of	of	ADP
iguana-17918	46	5	risk	risk	NOUN
iguana-17918	46	6	is	be	AUX
iguana-17918	46	7	also	also	ADV
iguana-17918	46	8	geographically	geographically	ADV
iguana-17918	46	9	concentrated	concentrate	VERB
iguana-17918	46	10	,	,	PUNCT
iguana-17918	46	11	with	with	ADP
iguana-17918	46	12	most	most	ADJ
iguana-17918	46	13	threats	threat	NOUN
iguana-17918	46	14	found	find	VERB
iguana-17918	46	15	in	in	ADP
iguana-17918	46	16	central	central	ADJ
iguana-17918	46	17	and	and	CCONJ
iguana-17918	46	18	south	south	PROPN
iguana-17918	46	19	america	america	PROPN
iguana-17918	46	20	,	,	PUNCT
iguana-17918	46	21	and	and	CCONJ
iguana-17918	46	22	australia	australia	PROPN
iguana-17918	46	23	,	,	PUNCT
iguana-17918	46	24	coinciding	coincide	VERB
iguana-17918	46	25	with	with	ADP
iguana-17918	46	26	reported	report	VERB
iguana-17918	46	27	outbreaks	outbreak	NOUN
iguana-17918	46	28	of	of	ADP
iguana-17918	46	29	chytridiomycosis	chytridiomycosis	NOUN
iguana-17918	46	30	.	.	PUNCT
iguana-17918	47	1	they	they	PRON
iguana-17918	47	2	speculated	speculate	VERB
iguana-17918	47	3	that	that	SCONJ
iguana-17918	47	4	the	the	DET
iguana-17918	47	5	phylogenetic	phylogenetic	ADJ
iguana-17918	47	6	clumping	clumping	NOUN
iguana-17918	47	7	of	of	ADP
iguana-17918	47	8	threat	threat	NOUN
iguana-17918	47	9	represents	represent	VERB
iguana-17918	47	10	,	,	PUNCT
iguana-17918	47	11	in	in	ADP
iguana-17918	47	12	part	part	NOUN
iguana-17918	47	13	,	,	PUNCT
iguana-17918	47	14	shared	share	VERB
iguana-17918	47	15	extinction	extinction	NOUN
iguana-17918	47	16	proneness	proneness	NOUN
iguana-17918	47	17	due	due	ADP
iguana-17918	47	18	to	to	ADP
iguana-17918	47	19	shared	share	VERB
iguana-17918	47	20	evolutionary	evolutionary	ADJ
iguana-17918	47	21	history	history	NOUN
iguana-17918	47	22	.	.	PUNCT
iguana-17918	48	1	however	however	ADV
iguana-17918	48	2	,	,	PUNCT
iguana-17918	48	3	even	even	ADV
iguana-17918	48	4	if	if	SCONJ
iguana-17918	48	5	the	the	DET
iguana-17918	48	6	phylogenetic	phylogenetic	ADJ
iguana-17918	48	7	clumping	clumping	NOUN
iguana-17918	48	8	of	of	ADP
iguana-17918	48	9	threat	threat	NOUN
iguana-17918	48	10	were	be	AUX
iguana-17918	48	11	simply	simply	ADV
iguana-17918	48	12	a	a	DET
iguana-17918	48	13	by	by	ADP
iguana-17918	48	14	-	-	PUNCT
iguana-17918	48	15	product	product	NOUN
iguana-17918	48	16	of	of	ADP
iguana-17918	48	17	shared	share	VERB
iguana-17918	48	18	geography	geography	NOUN
iguana-17918	48	19	,	,	PUNCT
iguana-17918	48	20	this	this	DET
iguana-17918	48	21	concordance	concordance	NOUN
iguana-17918	48	22	between	between	ADP
iguana-17918	48	23	phylogenetic	phylogenetic	ADJ
iguana-17918	48	24	and	and	CCONJ
iguana-17918	48	25	geographical	geographical	ADJ
iguana-17918	48	26	patterns	pattern	NOUN
iguana-17918	48	27	represents	represent	VERB
iguana-17918	48	28	a	a	DET
iguana-17918	48	29	prime	prime	ADJ
iguana-17918	48	30	opportunity	opportunity	NOUN
iguana-17918	48	31	.	.	PUNCT
iguana-17918	49	1	where	where	SCONJ
iguana-17918	49	2	practical	practical	ADJ
iguana-17918	49	3	,	,	PUNCT
iguana-17918	49	4	conservation	conservation	NOUN
iguana-17918	49	5	plans	plan	NOUN
iguana-17918	49	6	should	should	AUX
iguana-17918	49	7	focus	focus	VERB
iguana-17918	49	8	on	on	ADP
iguana-17918	49	9	biogeographical	biogeographical	ADJ
iguana-17918	49	10	regions	region	NOUN
iguana-17918	49	11	where	where	SCONJ
iguana-17918	49	12	threatened	threaten	VERB
iguana-17918	49	13	groups	group	NOUN
iguana-17918	49	14	occur	occur	VERB
iguana-17918	49	15	,	,	PUNCT
iguana-17918	49	16	thereby	thereby	ADV
iguana-17918	49	17	improving	improve	VERB
iguana-17918	49	18	the	the	DET
iguana-17918	49	19	capacity	capacity	NOUN
iguana-17918	49	20	for	for	ADP
iguana-17918	49	21	conserving	conserve	VERB
iguana-17918	49	22	species	specie	NOUN
iguana-17918	49	23	.	.	PUNCT
iguana-17918	50	1	this	this	DET
iguana-17918	50	2	approach	approach	NOUN
iguana-17918	50	3	could	could	AUX
iguana-17918	50	4	outperform	outperform	VERB
iguana-17918	50	5	the	the	DET
iguana-17918	50	6	usual	usual	ADJ
iguana-17918	50	7	triage	triage	NOUN
iguana-17918	50	8	approach	approach	NOUN
iguana-17918	50	9	of	of	ADP
iguana-17918	50	10	saving	save	VERB
iguana-17918	50	11	individual	individual	ADJ
iguana-17918	50	12	species	specie	NOUN
iguana-17918	50	13	after	after	SCONJ
iguana-17918	50	14	they	they	PRON
iguana-17918	50	15	have	have	AUX
iguana-17918	50	16	become	become	VERB
iguana-17918	50	17	critically	critically	ADV
iguana-17918	50	18	endangered	endanger	VERB
iguana-17918	50	19	.	.	PUNCT
iguana-17918	51	1	surveying	survey	VERB
iguana-17918	51	2	herpetofaunal	herpetofaunal	ADJ
iguana-17918	51	3	mortality	mortality	NOUN
iguana-17918	51	4	on	on	ADP
iguana-17918	51	5	rural	rural	ADJ
iguana-17918	51	6	highways	highway	NOUN
iguana-17918	51	7	road	road	NOUN
iguana-17918	51	8	mortality	mortality	NOUN
iguana-17918	51	9	can	can	AUX
iguana-17918	51	10	contribute	contribute	VERB
iguana-17918	51	11	to	to	ADP
iguana-17918	51	12	local	local	ADJ
iguana-17918	51	13	and	and	CCONJ
iguana-17918	51	14	regional	regional	ADJ
iguana-17918	51	15	declines	decline	NOUN
iguana-17918	51	16	in	in	ADP
iguana-17918	51	17	amphibian	amphibian	ADJ
iguana-17918	51	18	and	and	CCONJ
iguana-17918	51	19	reptilian	reptilian	ADJ
iguana-17918	51	20	populations	population	NOUN
iguana-17918	51	21	.	.	PUNCT
iguana-17918	52	1	consequently	consequently	ADV
iguana-17918	52	2	,	,	PUNCT
iguana-17918	52	3	accurately	accurately	ADV
iguana-17918	52	4	and	and	CCONJ
iguana-17918	52	5	efficiently	efficiently	ADV
iguana-17918	52	6	identifying	identify	VERB
iguana-17918	52	7	hotspots	hotspot	NOUN
iguana-17918	52	8	of	of	ADP
iguana-17918	52	9	road	road	NOUN
iguana-17918	52	10	-	-	PUNCT
iguana-17918	52	11	mortality	mortality	NOUN
iguana-17918	52	12	is	be	AUX
iguana-17918	52	13	necessary	necessary	ADJ
iguana-17918	52	14	for	for	ADP
iguana-17918	52	15	hazard	hazard	NOUN
iguana-17918	52	16	assessment	assessment	NOUN
iguana-17918	52	17	and	and	CCONJ
iguana-17918	52	18	mitigation	mitigation	NOUN
iguana-17918	52	19	.	.	PUNCT
iguana-17918	53	1	in	in	ADP
iguana-17918	53	2	2002	2002	NUM
iguana-17918	53	3	,	,	PUNCT
iguana-17918	53	4	langen	langen	PROPN
iguana-17918	53	5	et	et	PROPN
iguana-17918	53	6	al	al	PROPN
iguana-17918	53	7	.	.	PUNCT
iguana-17918	54	1	(	(	PUNCT
iguana-17918	54	2	2007	2007	NUM
iguana-17918	54	3	.	.	PUNCT
iguana-17918	55	1	journal	journal	NOUN
iguana-17918	55	2	of	of	ADP
iguana-17918	55	3	wildlife	wildlife	NOUN
iguana-17918	55	4	management	management	NOUN
iguana-17918	55	5	71:1361–1368	71:1361–1368	NUM
iguana-17918	55	6	)	)	PUNCT
iguana-17918	55	7	conducted	conduct	VERB
iguana-17918	55	8	walking	walk	VERB
iguana-17918	55	9	and	and	CCONJ
iguana-17918	55	10	driving	drive	VERB
iguana-17918	55	11	surveys	survey	NOUN
iguana-17918	55	12	throughout	throughout	ADP
iguana-17918	55	13	an	an	DET
iguana-17918	55	14	extensive	extensive	ADJ
iguana-17918	55	15	rural	rural	ADJ
iguana-17918	55	16	highway	highway	NOUN
iguana-17918	55	17	network	network	NOUN
iguana-17918	55	18	in	in	ADP
iguana-17918	55	19	northern	northern	ADJ
iguana-17918	55	20	new	new	PROPN
iguana-17918	55	21	york	york	PROPN
iguana-17918	55	22	to	to	PART
iguana-17918	55	23	evaluate	evaluate	VERB
iguana-17918	55	24	survey	survey	NOUN
iguana-17918	55	25	methods	method	NOUN
iguana-17918	55	26	and	and	CCONJ
iguana-17918	55	27	to	to	PART
iguana-17918	55	28	quantify	quantify	VERB
iguana-17918	55	29	spatial	spatial	ADJ
iguana-17918	55	30	and	and	CCONJ
iguana-17918	55	31	temporal	temporal	ADJ
iguana-17918	55	32	patterns	pattern	NOUN
iguana-17918	55	33	of	of	ADP
iguana-17918	55	34	herpetofaunal	herpetofaunal	ADJ
iguana-17918	55	35	road	road	NOUN
iguana-17918	55	36	-	-	PUNCT
iguana-17918	55	37	mortality	mortality	NOUN
iguana-17918	55	38	.	.	PUNCT
iguana-17918	56	1	in	in	ADP
iguana-17918	56	2	2004	2004	NUM
iguana-17918	56	3	,	,	PUNCT
iguana-17918	56	4	they	they	PRON
iguana-17918	56	5	repeated	repeat	VERB
iguana-17918	56	6	the	the	DET
iguana-17918	56	7	surveys	survey	NOUN
iguana-17918	56	8	at	at	ADP
iguana-17918	56	9	a	a	DET
iguana-17918	56	10	subset	subset	NOUN
iguana-17918	56	11	of	of	ADP
iguana-17918	56	12	locations	location	NOUN
iguana-17918	56	13	to	to	PART
iguana-17918	56	14	quantify	quantify	VERB
iguana-17918	56	15	interannual	interannual	ADJ
iguana-17918	56	16	repeatability	repeatability	NOUN
iguana-17918	56	17	.	.	PUNCT
iguana-17918	57	1	reptilian	reptilian	ADJ
iguana-17918	57	2	and	and	CCONJ
iguana-17918	57	3	amphibian	amphibian	ADJ
iguana-17918	57	4	species	specie	NOUN
iguana-17918	57	5	had	have	VERB
iguana-17918	57	6	different	different	ADJ
iguana-17918	57	7	peak	peak	NOUN
iguana-17918	57	8	periods	period	NOUN
iguana-17918	57	9	of	of	ADP
iguana-17918	57	10	road	road	NOUN
iguana-17918	57	11	-	-	PUNCT
iguana-17918	57	12	mortality	mortality	NOUN
iguana-17918	57	13	because	because	SCONJ
iguana-17918	57	14	they	they	PRON
iguana-17918	57	15	differed	differ	VERB
iguana-17918	57	16	in	in	ADP
iguana-17918	57	17	the	the	DET
iguana-17918	57	18	causes	cause	NOUN
iguana-17918	57	19	of	of	ADP
iguana-17918	57	20	movements	movement	NOUN
iguana-17918	57	21	that	that	PRON
iguana-17918	57	22	resulted	result	VERB
iguana-17918	57	23	in	in	ADP
iguana-17918	57	24	crossings	crossing	NOUN
iguana-17918	57	25	.	.	PUNCT
iguana-17918	58	1	spatial	spatial	ADJ
iguana-17918	58	2	locations	location	NOUN
iguana-17918	58	3	of	of	ADP
iguana-17918	58	4	herpetofaunal	herpetofaunal	ADJ
iguana-17918	58	5	roadmortality	roadmortality	NOUN
iguana-17918	58	6	were	be	AUX
iguana-17918	58	7	concentrated	concentrate	VERB
iguana-17918	58	8	at	at	ADP
iguana-17918	58	9	a	a	DET
iguana-17918	58	10	limited	limited	ADJ
iguana-17918	58	11	number	number	NOUN
iguana-17918	58	12	of	of	ADP
iguana-17918	58	13	hotspots	hotspot	NOUN
iguana-17918	58	14	.	.	PUNCT
iguana-17918	59	1	hotspots	hotspot	NOUN
iguana-17918	59	2	overlapped	overlap	VERB
iguana-17918	59	3	across	across	ADP
iguana-17918	59	4	species	specie	NOUN
iguana-17918	59	5	and	and	CCONJ
iguana-17918	59	6	were	be	AUX
iguana-17918	59	7	located	locate	VERB
iguana-17918	59	8	at	at	ADP
iguana-17918	59	9	consistent	consistent	ADJ
iguana-17918	59	10	locations	location	NOUN
iguana-17918	59	11	across	across	ADP
iguana-17918	59	12	years	year	NOUN
iguana-17918	59	13	.	.	PUNCT
iguana-17918	60	1	results	result	NOUN
iguana-17918	60	2	of	of	ADP
iguana-17918	60	3	walking	walk	VERB
iguana-17918	60	4	and	and	CCONJ
iguana-17918	60	5	driving	driving	NOUN
iguana-17918	60	6	surveys	survey	NOUN
iguana-17918	60	7	were	be	AUX
iguana-17918	60	8	highly	highly	ADV
iguana-17918	60	9	repeatable	repeatable	ADJ
iguana-17918	60	10	among	among	ADP
iguana-17918	60	11	survey	survey	NOUN
iguana-17918	60	12	teams	team	NOUN
iguana-17918	60	13	,	,	PUNCT
iguana-17918	60	14	but	but	CCONJ
iguana-17918	60	15	driving	drive	VERB
iguana-17918	60	16	surveys	survey	NOUN
iguana-17918	60	17	underestimated	underestimate	VERB
iguana-17918	60	18	the	the	DET
iguana-17918	60	19	density	density	NOUN
iguana-17918	60	20	of	of	ADP
iguana-17918	60	21	road	road	NOUN
iguana-17918	60	22	-	-	PUNCT
iguana-17918	60	23	mortality	mortality	NOUN
iguana-17918	60	24	because	because	SCONJ
iguana-17918	60	25	many	many	ADJ
iguana-17918	60	26	animals	animal	NOUN
iguana-17918	60	27	were	be	AUX
iguana-17918	60	28	missed	miss	VERB
iguana-17918	60	29	.	.	PUNCT
iguana-17918	61	1	detection	detection	NOUN
iguana-17918	61	2	failure	failure	NOUN
iguana-17918	61	3	was	be	AUX
iguana-17918	61	4	higher	high	ADJ
iguana-17918	61	5	in	in	ADP
iguana-17918	61	6	some	some	DET
iguana-17918	61	7	taxa	taxa	NOUN
iguana-17918	61	8	(	(	PUNCT
iguana-17918	61	9	e.g.	e.g.	ADV
iguana-17918	61	10	,	,	PUNCT
iguana-17918	61	11	frogs	frog	NOUN
iguana-17918	61	12	)	)	PUNCT
iguana-17918	61	13	than	than	ADP
iguana-17918	61	14	others	other	NOUN
iguana-17918	61	15	(	(	PUNCT
iguana-17918	61	16	e.g.	e.g.	ADV
iguana-17918	61	17	,	,	PUNCT
iguana-17918	61	18	turtles	turtle	NOUN
iguana-17918	61	19	)	)	PUNCT
iguana-17918	61	20	.	.	PUNCT
iguana-17918	62	1	their	their	PRON
iguana-17918	62	2	results	result	NOUN
iguana-17918	62	3	indicated	indicate	VERB
iguana-17918	62	4	that	that	SCONJ
iguana-17918	62	5	designing	design	VERB
iguana-17918	62	6	a	a	DET
iguana-17918	62	7	valid	valid	ADJ
iguana-17918	62	8	,	,	PUNCT
iguana-17918	62	9	efficient	efficient	ADJ
iguana-17918	62	10	methodology	methodology	NOUN
iguana-17918	62	11	for	for	ADP
iguana-17918	62	12	locating	locate	VERB
iguana-17918	62	13	hotspots	hotspot	NOUN
iguana-17918	62	14	of	of	ADP
iguana-17918	62	15	reptilian	reptilian	ADJ
iguana-17918	62	16	and	and	CCONJ
iguana-17918	62	17	amphibian	amphibian	ADJ
iguana-17918	62	18	road	road	NOUN
iguana-17918	62	19	-	-	PUNCT
iguana-17918	62	20	mortality	mortality	NOUN
iguana-17918	62	21	along	along	ADP
iguana-17918	62	22	a	a	DET
iguana-17918	62	23	road	road	NOUN
iguana-17918	62	24	network	network	NOUN
iguana-17918	62	25	is	be	AUX
iguana-17918	62	26	possible	possible	ADJ
iguana-17918	62	27	,	,	PUNCT
iguana-17918	62	28	thus	thus	ADV
iguana-17918	62	29	pinpointing	pinpoint	VERB
iguana-17918	62	30	priority	priority	NOUN
iguana-17918	62	31	sites	site	NOUN
iguana-17918	62	32	for	for	ADP
iguana-17918	62	33	mitigation	mitigation	NOUN
iguana-17918	62	34	.	.	PUNCT
iguana-17918	63	1	a	a	DET
iguana-17918	63	2	study	study	NOUN
iguana-17918	63	3	of	of	ADP
iguana-17918	63	4	tadpoles	tadpole	NOUN
iguana-17918	63	5	of	of	ADP
iguana-17918	63	6	african	african	ADJ
iguana-17918	63	7	clawed	claw	VERB
iguana-17918	63	8	frogs	frog	NOUN
iguana-17918	63	9	(	(	PUNCT
iguana-17918	63	10	xenopus	xenopus	PROPN
iguana-17918	63	11	laevis	laevis	NOUN
iguana-17918	63	12	)	)	PUNCT
iguana-17918	63	13	indicated	indicate	VERB
iguana-17918	63	14	that	that	SCONJ
iguana-17918	63	15	some	some	DET
iguana-17918	63	16	individuals	individual	NOUN
iguana-17918	63	17	with	with	ADP
iguana-17918	63	18	disease	disease	NOUN
iguana-17918	63	19	resistant	resistant	ADJ
iguana-17918	63	20	genes	gene	NOUN
iguana-17918	63	21	grew	grow	VERB
iguana-17918	63	22	more	more	ADV
iguana-17918	63	23	slowly	slowly	ADV
iguana-17918	63	24	.	.	PUNCT
iguana-17918	64	1	er	er	INTJ
iguana-17918	64	2	ic	ic	PROPN
iguana-17918	64	3	is	be	AUX
iguana-17918	64	4	se	se	X
iguana-17918	64	5	lé	lé	PROPN
iguana-17918	64	6	e	e	PROPN
iguana-17918	64	7	a	a	DET
iguana-17918	64	8	clumping	clumping	NOUN
iguana-17918	64	9	of	of	ADP
iguana-17918	64	10	threats	threat	NOUN
iguana-17918	64	11	to	to	ADP
iguana-17918	64	12	groups	group	NOUN
iguana-17918	64	13	of	of	ADP
iguana-17918	64	14	frogs	frog	NOUN
iguana-17918	64	15	,	,	PUNCT
iguana-17918	64	16	especially	especially	ADV
iguana-17918	64	17	the	the	DET
iguana-17918	64	18	hyloid	hyloid	ADJ
iguana-17918	64	19	frogs	frog	NOUN
iguana-17918	64	20	such	such	ADJ
iguana-17918	64	21	as	as	ADP
iguana-17918	64	22	this	this	DET
iguana-17918	64	23	redeyed	redeye	VERB
iguana-17918	64	24	treefrog	treefrog	NOUN
iguana-17918	64	25	(	(	PUNCT
iguana-17918	64	26	agalychnis	agalychnis	PROPN
iguana-17918	64	27	callidryas	callidryas	NOUN
iguana-17918	64	28	)	)	PUNCT
iguana-17918	64	29	,	,	PUNCT
iguana-17918	64	30	might	might	AUX
iguana-17918	64	31	represent	represent	VERB
iguana-17918	64	32	,	,	PUNCT
iguana-17918	64	33	at	at	ADP
iguana-17918	64	34	least	least	ADJ
iguana-17918	64	35	in	in	ADP
iguana-17918	64	36	part	part	NOUN
iguana-17918	64	37	,	,	PUNCT
iguana-17918	64	38	a	a	DET
iguana-17918	64	39	shared	share	VERB
iguana-17918	64	40	extinction	extinction	NOUN
iguana-17918	64	41	proneness	proneness	NOUN
iguana-17918	64	42	due	due	ADJ
iguana-17918	64	43	to	to	ADP
iguana-17918	64	44	shared	share	VERB
iguana-17918	64	45	evolutionary	evolutionary	ADJ
iguana-17918	64	46	history	history	NOUN
iguana-17918	64	47	.	.	PUNCT
iguana-17918	65	1	reptilian	reptilian	ADJ
iguana-17918	65	2	and	and	CCONJ
iguana-17918	65	3	amphibian	amphibian	ADJ
iguana-17918	65	4	species	specie	NOUN
iguana-17918	65	5	had	have	VERB
iguana-17918	65	6	different	different	ADJ
iguana-17918	65	7	peak	peak	NOUN
iguana-17918	65	8	periods	period	NOUN
iguana-17918	65	9	of	of	ADP
iguana-17918	65	10	road	road	NOUN
iguana-17918	65	11	-	-	PUNCT
iguana-17918	65	12	mortality	mortality	NOUN
iguana-17918	65	13	because	because	SCONJ
iguana-17918	65	14	they	they	PRON
iguana-17918	65	15	differed	differ	VERB
iguana-17918	65	16	in	in	ADP
iguana-17918	65	17	the	the	DET
iguana-17918	65	18	causes	cause	NOUN
iguana-17918	65	19	of	of	ADP
iguana-17918	65	20	movements	movement	NOUN
iguana-17918	65	21	that	that	PRON
iguana-17918	65	22	resulted	result	VERB
iguana-17918	65	23	in	in	ADP
iguana-17918	65	24	crossings	crossing	NOUN
iguana-17918	65	25	.	.	PUNCT
iguana-17918	66	1	spatial	spatial	ADJ
iguana-17918	66	2	locations	location	NOUN
iguana-17918	66	3	of	of	ADP
iguana-17918	66	4	herpetofaunal	herpetofaunal	ADJ
iguana-17918	66	5	road	road	NOUN
iguana-17918	66	6	-	-	PUNCT
iguana-17918	66	7	mortality	mortality	NOUN
iguana-17918	66	8	were	be	AUX
iguana-17918	66	9	concentrated	concentrate	VERB
iguana-17918	66	10	at	at	ADP
iguana-17918	66	11	a	a	DET
iguana-17918	66	12	limited	limited	ADJ
iguana-17918	66	13	number	number	NOUN
iguana-17918	66	14	of	of	ADP
iguana-17918	66	15	hotspots	hotspot	NOUN
iguana-17918	66	16	,	,	PUNCT
iguana-17918	66	17	such	such	ADJ
iguana-17918	66	18	as	as	ADP
iguana-17918	66	19	those	those	PRON
iguana-17918	66	20	used	use	VERB
iguana-17918	66	21	by	by	ADP
iguana-17918	66	22	female	female	ADJ
iguana-17918	66	23	turtles	turtle	NOUN
iguana-17918	66	24	moving	move	VERB
iguana-17918	66	25	to	to	ADP
iguana-17918	66	26	nesting	nesting	ADJ
iguana-17918	66	27	areas	area	NOUN
iguana-17918	66	28	.	.	PUNCT
iguana-17918	67	1	c	c	NOUN
iguana-17918	67	2	h	h	NOUN
iguana-17918	67	3	u	u	NOUN
iguana-17918	68	1	c	c	PROPN
iguana-17918	68	2	k	k	PROPN
iguana-17918	68	3	s	s	PROPN
iguana-17918	68	4	c	c	NOUN
iguana-17918	68	5	h	h	NOUN
iguana-17918	68	6	a	a	DET
iguana-17918	68	7	ff	ff	INTJ
iguana-17918	68	8	er	er	INTJ