Bulletin STATE OF ILLINOIS DEPARTMENT OF REGISTRATION AND EDUCATION DIVISION OF THE NATURAL HISTORY SURVEY THEODORE H. FRISON, Acling Chief Vol. XIX. BULLETIN Article II. Records of Spring Migration of Birds at Urbana, Illinois, 1903-1922 BY FRANK SMITH PRINTED BY AUTHORITY OF THE STATE OF ILLINOIS URBANA, ILLINOIS OCTOBER, 1930 STATE OP ILLINOIS DEPARTMENT OF REGISTRATION AND EDUCATION M. F. Walsh, Director BOARD OF NATURAL RESOURCES AND CONSERVATION W. F. Walsh, Chairman WiLLLVM Trelkask. BiolOQy Henry C. Cowlks. Forestry EnsoN S. Bastix. Geology William A. Noyks. Chemistry John W. Ai.miim). Engineering CiiABLEf; M. TiioMrsox. Beprrsenting the Picsidcnt of the Vniversity of Illinois STATE NATURAL HISTORY SURVEY DIVISION Theodoke H. Fkison. Acting Chief H. C. Oestekling, Editor SoiiNEPP & Barnes, Printers Spri.ngfiki.d. III. 4IS4S—1500 VOLUMK XIX. Article II. RECORDS OF SPRING MIGRATION OF BIRDS AT URBANA. ILLINOIS 1903-1922* FRANK SMITH The chief object in view in the preparation of this paper has been to make available to those interested certain data connected with spring bird migration activities in East Central Illinois in tlie vicinity of Urbana. Historical.—A regular course of instruction at the University of Illinois involving field work in ornithology was given by the writer in the spring of 1903 and annually thereafter for more than "iO years. Although he had been interested in birds in a general way for a num- ber of years, their study had not been his major field of research and he was led to offer such a course, in part, because of the growing in- terest in bird study in some schools of the state and hence the need for an opportunity for the preparation of persons desiring lo take charge of such work. During the first year records of the numbers of the various kinds of birds seen on each of the field trips were made and filed, but without any very definite plans for their further use. .\t a meeting of the American Association for the Advancement of Science in December, 1903, the writer learned from Dr. C. C. Adams, then Curator of the Museum of the University of ]\Iichigan, of the observa- tion by N. A. Wood that a study of weather maps during the spring migration in the Ann Arbor region enabled one to predict with a con- siderable flegree of certainty the "bird waves" when tmusually large numbers of arrivals of species and individuals are recorded. ,\ desire to learn whether there might be such a definite correlation in I'lastern Illinois between the migration activities and certain kinds of weather conditions led to definite [jlans for daily observation trips throughout the spring migration season and including as wide a range of habitats as feasible. Especial effort was made to have the dates when rejiresen- tatives of the various migratory species were first seen correspond as • Contributions from the Zoolosital I.,.il)oratory of the University of IllinoLs, No. 380. fl05] 106 Jl.LlNOIS NaTL'UAI, HiSTOKY SLUVEY BlLLKTIN closely as possible with the dates of actual first arrival. The coopera- tion of competent and interested observers who made records in dif- ferent areas in the vicinity aided niateriallv in accomplishing this re- sult. .\mong these collaborators at various times were advanced stu- dents who were working on problems in which such observations were a more or less important part of their work. Included in the list of those who were especially helpful were A. O. Gross and T. E. Mussel- man, who have since contributed to our knowledge of Illinois ornitho- logy, and F. C. Gates, well known for his work in plant ecclog}". Since the correlation between migration flights and certain kind^ of weather conditions was the object of study, special ettort was made to obtain records of the first appearance of birds of the various species in the particular areas available for daily observation rather than in more remote areas which might be more attractive to birds of some species and perhaps have earlier arrivals, but to which it was not prac- ticable to make daily trips. It is quite probable that in wooded areas along the banks of larger streams with bottom-land ponds or lakes and with marshy areas, the dates of arrival may have been somewhat earlier for some species of birds than the dates given in the following table. Alore favorable conditions attracting larger numbers of in- dividuals of any particular kind are quite likely to result in earlier records for the observer who can make daily trips to such areas. During the period of I!iu:5-Ui22 there were extensive changes in the environmental conditions of some of the areas studied which had a marked effect on their bird faunas.* With some species a marked ad- verse change in environment has been accompanied bv a reduction from a large number of records per annum to a much smaller number. This has sometimes resulted in the average of the "first seen" records for the later years being several days or even a week or more later than that for the years when the species was abundantly represented. In a few cases of this kind it has seemed advisable to omit some of such abnormally late records when determining the median date of the records for use in the following talnilar list. Areas in i^'liich records leere luude.—The areas most thoroughly sttidied were necessarily those readily accessible from the university campus, since the records were made chiefly by those in charge of sections of the classes in ornithologv while doing field work, or liy those making early-morning trips. In cither case it was not feasible to spend ver}- much time in journeying to jilaces too far removed. One inipnrtant source of records was an artificially forested area of about 1 .J acres, locally known as the l-"iiri'siry. which is a part of the campus. DiUMng the first lew )ears of ihe jieriod discussed. Spki.ng Migration of Birds at Uruaxa 101 the roadway on the east side of this area was bordered on each side b\ hedges, and the land on the east side was used by the owners chiefly for orchards and pasture. Shrubber)- and brush heaps were common, and the character of the environment made the region very attractive to many of the migrant species. In the Forestry itself there was con- siderable shrubbery and occasional brush heaps. With the passing of the years and the great increase in the develop- ment of the university and the growth of the neighboring city, there was brought about the disappearance of hedges, brush heaps, and much of the shrubber)- : the pastures were replaced by dooryards, and in the Forestry itself the trees were trimmed high and the forest floor cleaned up in such a manner as to make it far less attractive to many of the ground-feeding species. A natural result of such changes has been that most of the birds of some of these species have sought more attractive localities and the records of dates when they were first seen have been less reliable for indicating the time of their first arrival in the region. A cemetery lying south of the university campus is another definite area which was regularly visited and which supplied a quite important share of the records. A very considerable number of large trees scattered through the area attracted a good many kinds of birds, while some hedges and considerable shrubbery attracted others. In the earlier years many of the burial lots were surrounded by hedges and contained much shrubbery, but the tendency in recent years has been to eliminate the hedges and most of the shruljbery, with resulting diminution in the numbers of some species of birds attracted by such an environment. One very important area regularly visited contains what is locally known as Crystal Lake. It was the chief source of records for species having a preference for stream, lake, and marsh environments. It is also the area which has sutTered most, from the ornithnlogist's point of view, because of changes affecting the environmental conditions. The region about Urbana and the University of Illinois has a slightly higher altitude than most of the adjacent territory and is without natural lakes, while the streams are mostly ditches and merely the beginnings of water-courses that l:)ecome rivers elsewhere. ( )ne of these streams known locally as Salt Fork, has its origin a few miles north of Urbana and flows through the northeastern part of the city, then has an easterly course and is one of the tributaries of the Vermilion River. .\t some time in the earlier history of Urljana, a dam several feet high had been built across the stream in the northeastern part of the city, which re- sulted in the formation of an artificial lake of several acres in extent 108 Illinois Natiral History Sukvev Billetin (Crystal Lake) and also produced some tracts with marsh-like condi- tions. The banks of the lake were adapted to the needs of shore birds, and considerable numbers of coots, ducks, and other water birds were also regular visitors during the migration seasons. The stream above the lake had a meandering course through some wooded tracts, and the patches of willows and shrubbery along its banks made habitats that were attractive to birds of quite a large number of species. Extensive changes made in this area in IfiOS greatly altered the environmental conditions, with resulting changes in the bird life. For the more effective drainage of farm lands north of the city, an exten- sive drainage ditch was dug that replaced the old stream bed. but had a much more direct course and had steep, bare banks manv feet in height. The part of the old stream bed where the lake had been formed was not included in the course of the ditch, but with the aid of the dredge a long curved excavation was made in the area that had been covered by the lake, which had been included in a city park. This sub- stitute for the former lake received its water supply from the draina,ge ditch and had steep banks which were later mostly covered by shrub- bery. The low marsh-like tracts and muddy flats attractive to shore birds were eliminated, and although the area has been successfully de- veloped for park purposes, there is little to attract shore and water birds and several other kinds formerly fnund in the region. Genera! considerations.—In the accumulation of the following data it soon became obvious that it is sometimes difficult to be certain of accuracy in determining the "first seen'" dates. Frequently one may not have an opportunity for a sufficiently careful detailed study of a s]5ecimen because of intervening brush or foliage, or because of its timidity and hasty departure. In making use of data furnished by other observers one cannot always be sure that they had sufficient knowledge to allow accurate identification of all kinds seen or that they listed only those which furnished an ojiportunitv for an adequate study. In the preparation of the following data it has been the usual practice to omit the records tiiat were considered doubtful. In a study of the bird fauna of a region like the one under con- sideration, one must expect to deal with some species of which speci- mens are rarely seen, and not infreiiuently there will be but one or two records in a season and for some s]iecies but one or two records for the entire ]ieriod, Such records may be of but little use in an at- tem])t tt) determine the earliest and latest limits of a normal migration ])eriod for the species. The lack of suitable environments for many species of water liirds accounts for the ]iaucity of records for some species, although thev may be common in more faxorable localities in the same county. Sprin'o Migration of Biriis at Uruana 109 Irregularities in the migration activities of birds of some species have been noticed that are similar to those recorded in other regions. Several species normallv rated as summer residents have been repre- sented by a few individuals which have stayed in the region during one or more winters. The Mourning Dove, Red-headed Woodpecker. Red- winged Blackbird, Robin, Bluebird, and others are examples. The Song Sparrow is re|iresented throughout the year although presumably all of the individuals are migratory. Its summer and winter ranges overlap in this region, and hence it may l>e listed as a permanent resi- dent, although the majority of the individuals seen are annual visitants, wdiile a few are summer residents and a few others are winter residents. The Mockingbird seems to have been altering its winter range some- what. February IS, ]!!()(), and March ".^S, 11)08, are the only records of its being seen earlier than April during any year preceding the winter of 1917/1018. Specimens were seen that winter and in each succeed- ing winter except that of 1918/1919. They were frequently seen in midwinter feeding in hedges and asparagus beds of city dooryards and gardens. Lists of recorded species and z'urieties.—Of the 22'^ different kinds on our records, the names of all but eight may Ije found in the tabular list which follows. Four species of especially irregular occurrence and not in that list include the Snowy Owl and (iolden Eagle, of each of which sjiecimens were taken in Novemljer of two dififerent years ; the Bald Eagle, seen in March of 1905 and 1930; and the Duck Hawk, of which a specimen was taken in May of 1908. F. C. Gates made records of four other species as follows: Whistling Swan, April 23, 1908; Little Blue Heron, May 17, 1908; Sandhill Crane, May 11, 1910; and Semipalmated Sandpiper, April 25, ]!)()8. For the convenience of those who may be interested in other than migrant species, the names of all birds regularly seen have been included in the list. The recorfls for most of the migratory species that are listed in the following table include: (column i) the earliest date record of any one of the years; (column 2) a median "first seen" date record, which is the median date for those recognized as summer residents and an- nual visitants; (column j;) a median "last seen" record for winter resi- dents and annual visitants; and (column 4) a latest spring record for any one of the years. The month is indicated by a Roman numeral and the day Ijy an Araljic numeral. The number of years in which any given species was recorded (column f,j is i)reccded in some instances by a lesser number which indicates the number of years of which "first seen" records were used in determining the median date. This lesser number occurs with some species which were represented during the 110 Ir.Lixois Natikai, History Survey BtLLEXix winters of some years by a few specimens and also with certain other kinds which in some years had but one or few records and these late in the season. Symbols indicate the migratory group to which each species belongs (cohiinn 6), as follows: A. V. indicates annual visitants that spend the summers further north and the winters further south ; W. R., winter residents ; S. K.. summer residents ; and P. R., permanent residents, including non-migratory forms and also certain migratory forms of which the summer and winter ranges overlap in the region and hence representatives are normally present at all times of the year even though the individuals are migratory. Interrogation marks after such symbols indicate uncertainty concerning the actual status of the species in this locality. Conflicting data in different papers dealing with the seasonal distribution of birds in the state are frequently found. A paper by Forbes and Gross (1923) has been especially helpful as a source of distribution data. During some years our data included no records made after June 1st., and hence the data for 'last seen'' records are less complete and reliable than those used for the "first seen" records. Records of the V'eery and its western variety, the \\'illow Thrush, are combined in the records of most years. This is due in part to the difficulty of ready distinction between the two forms in the held, and in part to the fact that in the earlier years of the period it was not realized that both forms were represented in the region. Similarly, the two species of Chickadees were not carefully distinguished in the earlier years of the period. Apparently the Carolina Chickadee is the one commonly found breeding and the other is common chiefly in the winter season. No attempt was made to separate the records of the House Wren and its variety, the Western House Wren : nor those of the Water Thrush and its variety, Grinnell's \\ater Thrush. For each of these species the records are listed under the name of the form which seems to be most commonly represented. The writer has thought best to follow the example of Lynds Jones (1914) and use the median dates of "first seen" records rather than the average dates since thev are less affected by extremes in earliest or latest records. The names of the species are listed in the same order as that followed in the American Ornithologists" L'nion (.\.0. U.) Check List, by N. A. Wood (1907) in his report on bird migration in Michigan, and by Forbes and dross (1923). A comparison of the median dates of our "first seen" records with those of Hess (1910), which were obtained a few miles farther .south and in the same county, reveals that in most cases the dates here listed are several days earlier than those of Hess. This is jiresumably due Sprung MKiHAXioN of Birds at Urisaxa 111 to his lack of opportunity for daily observation trips and to his lack of sufficient aid from other observers, as intimated in his iiajier (p. 20). For the convenience of those who are interested in the general subject of bird migration but who have not yet gained a detailed knowledge of the various phases of the subject or of the extensive literature dealing with them, it has seemed advisable to refer briefly to some facts which have a direct relation to the times at which various kinds of birds may be expected to appear in a region. Some kinds, including the Bluebird and the Meadowlark, usually appear in Central Illinois in February and often so early that their lives are in peril be- cause of interference with their food supply by snow storms and freez- ing weather that may prevail after their arrival. The majority of birds that migrate arrive later in the season, and a good many kinds are not to be seen until Way. The time of arrival is largely dependent on the location of the region in which the birds of any particular species spend the winter. Birds that winter in Southern Illinois or but little further south may be expected to arrive weeks before those that winter in the states along the Gulf Coast and months before those that winter in South Arnerica. Birds of about seven-eighths oi the species common in Illinois make annual journeys back and forth between their summer and winter homes, and birds of nearly one-half of the siiecies have the areas oc- cupied as summer and winter homes entirely separated, and hence tliey are annual visitants in the intervening territory. The l)irds of about one-third of our Illinois species journey beyond the boundaries of the United States to reach their winter homes. A few kinds winter in the West Indies, some in Mexico, others in Central America, while birds of about one-fifth of our species extend their journeys into South America. Included among these are some of the thrushes, warblers, swallows, flycatchers, cuckoos, and sandpipers. Hennit thrushes, phoebes, and myrtle warblers that winter in the states be- tween Illinois and the Gulf of Mexico have much shorter distances to travel in spring than do some of the other thrushes, flycatchers, and warblers that winter in South America, and it is only natural that they should arrive in Illinois much earlier than do those that have much longer distances to travel. A study of the daily records made during the spring migration periods under consideration shows a great lack of uniformity in the amount of migration activity on successive nights. On some mornings we have found new arrivals of as many as 15 or 20 species not ])revi- ously seen that season, and such movements are often preceded and followed by nights of but little migratory activity. Such extensive 112 Illi.nois Natikai. Histoicy Sikvey Bilif.tix movements, or "bird waves," are not due to food conditions, but seem to be closely correlated with certain weather conditions. A careful study of the weather maps during the time when records were being made revealed that the greatest migratory activity in spring occurred at times when the weatlier maps showed an area of low barometric pressure approaching from the west, with the south winds and rising temperatures which normally accompany such movements. Definite illustrations of such correlation will be found in two papers by the writer ( lltK and l!t'^l). In the latter of these two papers (Illinois Birds as Travellers) there are also memoranda on winter homes of certain kinds of Illinois birds and on the routes followed in the journeys to and from them. In the use of this latter paper much confusion ma> be avoided by the correction of an unfortimate error made in publica- tion : the reference in the text to Fig. 1 should be to Fig. 3. and the one to Fig. 3 should be to Fig. 1. LITERATURE CITED. Forbes. S. A., and Ghdss. A. 0. 192.3. On the Numbers and Local Distribution of Illinois Land Birds of the Open Country in Winter. Spring, and Fall. //(. Kat. Hi.st. Suit. BuL. Vol. 14. pp. 397-4.53. Hess. I. E. 1910. One Hundred Breeding Birds of an Illinois Ten-mile Radius. Thr Auk. Vol. 27, pp. 19-32. Jones, Lyxds. 1914. Nineteen Years of Bird Migration at Oberlin. U'if.vow Bui. No. 89, pp. 198-205. Smith. F. 1917. The Correlation between the Migratory Flights of Birds and Certain Accompanying Meteorological Conditions. Wilson Bui. No. 98, pp. 32-35. 1921. Illinois Birds as Travelers. lU. .^uiil. I'uh. Iii.str. Cin\ No. 151. "Arbor and Bird Days." pp. 20-29. Wood. N. A. 1907. Twenty-five Years of Bird Migration at Ann Arbor. Michigan. Mich. Acail. Sci., Stii Ann. Rrpt.. pp. 151-156. Table No. 1. Spring MKiitATiox of Birds at Uiii!A.\A 113 114 Ili.ixois Natural History Survey Bulletin . Spring Migration of Birds at Urmana 115 Common name of species Earli- est date record Median of "first seen" records Median of "lastl seen" records Latest spring record Num- Migra- ber of I tory years ! group Orchard Oriole Baltimore Oriole Rusty Blackbird Bronzed Crackle Purple Finch English Sparrow Crossbill White-winged Crossbill Redpoll Goldfinch Pine Siskin Snow Bunting Lapland Longspur Smith's Longspur Vesper Sparrow Savannah Sparrow Grasshopper Sparrow Henslow's Sparrow Leconte's Sparrow Lark Sparrow White-crowned Sparrow White-throated Sparrow Tree Sparrow Chipping Sparrow Clay-colored Sparrow Field Sparrow Slate-colored Junco Montana Junco Bachman's Sparrow Song Sparrow Lincoln's Sparrow Swamp Sparrow Fox Sparrow Towhee Cardinal Rose-breasted Grosbeak Indigo Bunting Dickcissel Scarlet Tanager Summer Tanager Purple Martin Cliff Swallow Barn Swallow Tree Swallow Bank Swallow Rough-winged Swallow Bohemian Waxwing Cedar Waxwing Migrant Shrike Red-eyed Vireo Philadelphia Vireo Warbling Vireo Yellow-throated Vireo lV-25 116 Ilh.nois Natlkal History Survey Bulletin Sl'HlXG lIlGRATIOX OF BiRDS AT UhHAXA 117 Z D d Common name of species Earli- est date record Median of "first seen" records Mediani of "last! seen" records Latest spring record Num- ber of years Migra- tory group 721 721a 722 724 725 726 727 72S 731 735 736 748 749 751 755 756 756a 757 75Sa 759b 761 766 House Wren and | Western House Wren [ Winter Wren Short-billed Marsh Wren Long-billed Marsh Wren Brown Creeper White-breasted Nuthatch Red-breasted Nuthatch Tufted Titmouse Chickadee Carolina Chickadee IGolden-crowned Kinglet Ruby-crowned Kinglet Blue-gray Gnatcatcher iWood Thrush .Veery and | Willow Thrush j Gray-cheeked Thrush Olive-backed Thrush Hermit Thrush Robin Bluebird ni-26