Int. J. Aquat. Biol. (2023) 11(2): 131-140 

ISSN: 2322-5270; P-ISSN: 2383-0956

Journal homepage: www.ij-aquaticbiology.com 

© 2023 Iranian Society of Ichthyology 

Original Article 
Length-based fishery status and population dynamics of Spiral Babylon, Babylonia 

spirata, Linnaeus, 1758, stock in the northern waters of the Oman Sea, Sistan and 

Baluchestan Province of Iran 
 

Seyed Ahmad Reza Hashemi1, Mastooreh Doustdar2, Elnaz Erfanifar1 

 
1Offshore Fisheries Research Center, Iranian Fisheries Science and Research Institute, Agricultural Research Education and Extension Organization, Chabahar, Iran. 

2Iranian Fisheries Science and Research Institute, Agricultural Research Education and Extension Organization, Tehran, Iran. 

 

 

 

 

s 

Article history: 

Received 19 June 2022 

Accepted 26 March 2023 

Available online 2 5 April 2023 

Keywords:  

Spiral Babylon  

Exploitation coefficient  

Mortality fishing  

Oman Sea  

Abstract: In the present study, the population characteristics of Spiral Babylon, Babylonia spirata, 
were evaluated by sampling at four sites in the northern Oman Sea, Iran, including Pozm, Konark, 

Beries, Pasabandar from March 2021 to March 2022. A total of 2779 Babylonian snail specimens 

(1489 males, and 1290 females) were measured and about ten percent of the specimens were 

described. The mean length and weight of males and females were 36±5, and 32±5 mm and 14±6, 

10±4 g, respectively. Growth and mortality indices for females and males including infinite length 

(L∞ = 68 and 76 mm), growth coefficient (K = 0.54 and 0.3 (yr-1)), natural mortality (M= 0.7 and 

0.4 (yr-1)), fishing mortality (F = 2.30 and 1.79 (yr-1)), total mortality (Z = 3 and 2.19 (yr-1)) and 

exploitation coefficient (E = 0.77 and 0.82 (yr-1)) were estimated. Based on the LBSPR assessment 

model, estimated to be about 0.3, and a ratio of Pmega<0.1, Lmean/Lopt <1 and Lmean /LF=M<1 show 

considered undesirable. The present study showed that the Spiral Babylon stock has reached 

'overfished' status.  

Introduction 

Population dynamics are driven by changes in the 

abundance or biomass of a population through time 

by a series of life-history traits such as fecundity, 

successful recruitment, growth rates, and mortality 

rates. Estimates of population dynamics can provide 

insights into a harvested marine population species.  

It can indicate how a population arrived at its current 

state and how it might change in the future (Brown 

and Guy, 2007). Recruitment, growth, and mortality 

rates are the primary population dynamics 

parameters that explained the harvestable part of a 

fish population (Brown and Guy, 2007). 

Mollusca is the second phylum in the number of 

living species, after arthropods (Pandian, 2017). Of 

the seven molluscan classes, the 78,000 known 

gastropod species contribute 85% of all mollusks and 

the 60,000 species of prosobranchs contribute a 

significant part of this group (Pandian, 2017).  The 

 
Correspondence: Seyed Ahmad Reza Hashemi                                       DOI: http//doi.org/10.22034/ijab.v11i2.1627 

E-mail: seyedahmad91@gmail.com                                                               DOR: https://dorl.net/dor/20.1001.1.23830956.2023.11.2.6.9 

genus Babylonia (Mollusca: Gastropoda: 

Buccinidae) possesses 11 extant species, two of 

which polytypic with two subspecies  each, and 12 

fossil and extinct species  (Gittenberger, and Goud, 
2014). The three most common species, viz. 

Babylonia areolata, B. japonica and B. spirata, have 

continuous ranges (Gittenberger and Goud, 2014). 

The Oman Sea is a region of the northern Indian 

Ocean that connects the Arabian Sea with the Strait 

of Hormuz, which then runs to the Persian Gulf. It 

borders Iran and Pakistan on the north, Oman on the 

south, and the United Arab Emirates on the west 

(Taghavimotlagh and Shojaei, 2017). The Oman 

Sea, with its unique ecological conditions, hosts a 

wide variety of marine species that provide 

livelihood, employment, and vast economic 

activities for the settlers. Iran has more than 120,000 

fishermen. Therefore, fishing has played a major role 

in creating employment in coastal areas, as well as in 



132 
 

Hashemi et al./ Length-based fishery status and population dynamics of Spiral Babylon 

economic activities for post-harvest operations 

(Taghavimotlagh and Shojaei, 2017). 

Some studies on the biological characteristics of 

shell species in different parts of the world include 

Mohan (2007), Cob et al. (2009), Arrighetti et al. 

(2012), Daza-Guerra et al. (2018), and Matos et al. 

(2020). Recently, the illegal fishing of Babylonia 

spirata in the northern waters of the Oman Sea has 

increased significantly. It is not used for food in the 

region and is caught only for export. There is 

currently no specific management or restriction for 

this species in this area. Despite the economic 

importance of this species, little is known about the 

stock of B. spirata population. Therefore, this study 
investigates several life-history traits and fish 

assessment measures of B. spirata from the Oman 
Sea to provide fundamental information for proper 

identification and management. Hence, first, the 

structure of the population  and growth rate of B.

spirata from the Oman Sea will be studied; then, a 

series of fisheries assessment data will be calculated 

such as mortality and exploitation coefficient to 

evaluate the sustainability of its population in the 

Oman Sea.  

 

Materials and methods 

Sample sites and specimen collection: Based on the 

location in the northern Oman Sea, Iran, four sites 

were selected for sampling B. spirata in the ports of 

Pozem (60˚15´E, 25˚35´N), Konark (60˚28´E, 

25˚60´N), Beris (61˚10´E, 25˚82´N) and Pasabandar 

(61˚25´E, 25˚70´N) (Fig. 1). Samplings were carried 

out monthly from March 2017 to March 2022. 

Samples were collected with the help of fishermen 

using special baskets from subtidal areas (sandy and 

pebble shore) at depths of less than 50 meters. A total 

of 2779 specimens of B. spirata specimens were 

collected (Fig. 2).  

Biometric data: The shell weight and length of all 

specimens were measured.  The total length was 

determined by a biometric ruler with 1 mm precision 

(Fig. 2) and wet weight to the nearest 1 gr. The 

Equation of Wi = a × Li b (1) was used to calculate 

the relationship between the total length and wet 

weight, where, Wi is the shell weight (g), Li is the 

shell length (mm), a is a constant coefficient, and b, 

 Figure 1. Map of Babylonia spirata sampling stations in the northern Oman Sea, Iran. 
 

 

 



133 
 

Int. J. Aquat. Biol. (2023) 11(2): 131-140 

 

an equation power. Equation (2) was used to 

decipher significant differences between the 

calculated b from Equation (1) and b = 3 for a shell 

of similar growth with s.dx, the standard deviation of 

total length natural log, s. dy standard deviation of 

weight natural log,b slope, r2 coefficient of 

determination, and n is sample sizes (Zar, 2010).  

t = [(s.dx)/ (s.dy)] × [(lb-3l)/ (√ (l-r2)] × [√ (n-2)] 

(2) 

Population dynamic parameters of B. spirata: The 

data were pooled monthly from different stations, 

and subsequently grouped into classes of three 

centimeters interval. The ELEFAN method was 

used to analyze the data for growth rates. The 

estimation of L∞, the infinite length was  obtained 

using Equation (3) and the maximum length of 

samples (Lmax) based on Froese and Binohlan (2000) 

Log L∞ = 0.044 + 0.9841 * Log (Lmax) (3) 

The growth rate was obtained by applying the 

ELEFAN method (optimization model), using the 

RStudio software with the TropFishR package 

(Mildenberger et al., 2017).  The optimum value of 

t0 (time that length is zero) was calculated by the 

experimental formula of Pauly equation (Equation 

(4)) with K, the growth factor (Froese and Binohlan, 

2000). 

Log (-t0) = - 0.3922 - 0.2752 Log L∞ - 1.038 Log 

K (4) 

The maximum lifespan was estimated based on 

Froese and Pauly (2017 and the equation of tmax = t0 

+ 3 / K (5). 

Mortality: Natural mortality (M) was calculated 

based on the Brey Equation (6) with, M as the annual 

natural mortality coefficient, Amax as the life 

expectancy based on the year, BMmax as the 

maximum body weight (30*3.81) based on K J (for 

this species the coefficient is 3.81 kJ /g) and T 

(Kelvin) is the average ambient temperature (Brey, 

1999; Mohan, 2007). 

log (M) = 1.672 +( 0.993* log (1/Amax))) – (0.035* 

log (BMmax)) – (300.447/ T+273) 

(6) 

The mean annual temperature of the sea surface 

of the northern Oman Sea was estimated at 26°C or 

299 (K) (Hashemi, 2020). Total mortality (Z) was 

calculated based on the length-converted catch 

curves data. The fishing mortality was estimated 

using Equation (7), with Z, the total mortality, F, the 

fishing mortality, and M the natural mortality. 

F = Z – M (7) 

The exploitation rate (E), which is the ratio of 

fishing mortality to total mortality, was calculated 

using Equation (8) based on Sparre and Venema 

(1998) 

E = F / Z (8) 

The relative yield per recruitment (Y/R) and 

relative biomass per recruitment (B'/R): The 

relative yield per recruitment (Y/R) was estimated 

against the fishing mortality coefficient or 

exploitation rate. In this Equation (9), E is the 

exploitation coefficient, U is the exploitation rate, M 

is the natural mortality coefficient, F is the fishing 

mortality coefficient and Lc (Length at first capture) 

is the same as Lc50 (Gayanilo et al., 2003).  

 

Figure 2. The Babylonia spirata species in the northern Oman Sea, Iran. 
 

 

 



134 
 

Hashemi et al./ Length-based fishery status and population dynamics of Spiral Babylon 

Y' / R = EU M/K ( -3 U / (1 + m) + 3 U2 / (1 + 2m) + 

U3 / (1 + 3m ) with 

U = 1 - (LC / L∞)  ;  m = (1 - E) / ( M / K) = ( K / Z) 

;  E = F / Z (9) 

In addition, the relative biomass per recruitment 

(B'/R) was calculated using the Equation of B' / R= 

Y' / R / F (10). 

Length-based indicators (LBI): In the length-based 

indicators (LBI), Popt is the percentage of fish caught 

at the optimum length for harvest, Pmega, is the 

percentage of mega-spawners (length between 1.1 

Lopt and Lmax on the catch length composition) in the 

catch, and LF=M is values for optimal fishing length 

that were calculated using the formula of 11, 12, and 

13, respectively (Froese and Binohlan, 2000; 

Cousido-Rocha et al., 2022): 

Lopt=Linf (3/(3+M⁄K)) (11) 

LF=M = (0.75Lc+0.25Linf) (12) 

+ %10 (13) optL= megaP 

Length-based spawning potential ratio (LBSPR): 

LBSPR is one of the biological reference points for 

determining the stock status of a species in the 

population. The reference points of LBSPR used are: 

LBSPR 20% is as the limit reference point and 

LBSPR 40% is a target reference point (Hordyk et 

al., 2015) that is calculated as:  

LBSPR =  
∑(1−𝐿𝑋)(𝑀 𝐾⁄ )[(𝐹 𝑀⁄ )+1])𝐿𝑥

𝑏

∑(1−𝐿𝑋)  
 𝐿𝑥

𝑏
𝑀/𝐾    (14) 

Where Lx is fork length, M is natural mortality; k 

is the growth rate, F is fishing mortality, and b is 

exponent usually close to 3. Estimating SPR with 

those functions needs the simple assumptions of 

asymptotic or logistic selectivity and no variation in 

length at age. The F/M ratio can be estimated from 

the length composition of the catch (Hordyk et al., 

2015). The relationship between F/M and SPR is 

asymptotic and determined by the selectivity 

parameters (Carruthers and Hordyk, 2018). 

Statistical analyses: Comparison of population 

dynamic values between male and female lengths, 

and weights were tested by Student’s Test (t-test) 

with paired t-test and independent t-test, 

respectively. The normality of this data was assessed 

using the Kolmogorov–Smirnov test. Data analyses 

were performed using FiSAT II and R Studio 

softwares (1.1.46) with the TropFishR package.  

 

Results 

Length frequency distribution:  The mean ± 

standard deviation of total length (Lm) and total 

weight (Wm) for male (1489 specimens) and female 

(1290 specimens) were 36±6 (18-58 mm), and 32±5 

(19-50mm), and 14±6 (12-63 g), 10±5 (21-50 g), 

respectively. The differences between total length 

and total weight in both sexes were not significant (t 

= 1.45, P= 0.05; t = 1.44, P>0.05, respectively). The 

length (TL) data were categorized into 3-mm groups 

which the highest frequency belonging to individuals 

with 27 to 30 mm (Fig. 3A). 

Length-weight relationship: The relationship 

between length and weight give a=0.0008 and b= 

2.66 (R2 = 0.89) for female, and for male, a=0.008 

and b=2.71 (R2 = 0.88) and for both sexes a=0.0008 

and b=2.70 (R2 = 0.88). The results showed 

significant differences between estimated b and b=3 

at the level of 0.05 (Fig. 3B), which means a negative 

allometric growth pattern for both sexes of B. spirata 

in the northern waters of the Oman Sea.  

 Growth studies: The population dynamic 

parameters of male, female, and both sex of 

B. spirata are detailed in Table 1. Growth parameters 

for both sexes together were L∞ = 71 mm (W∞ = 79 

gr), K = 0.35 (yr-1), and t0= -0.2.  

The growth curve (Fig. 4) highlighted 6 cohorts 

and age groups and the growth performance index as 

Ф = 3.11. There is recruitment throughout the year 

and the highest rate of recruitment is observed in the 

winter and summer season’s time. Based on the 

results, the maximum lifespan of this species was 

nearly 8 years. 

Mortality estimate (M): The natural mortality (M), 

fishing mortality (F), and total mortality (Z) were 

estimated as 0.5 (yr-1), 1.75 (yr-1), and 2.25 (yr-1), 

respectively (Table 1). The exploitation coefficient 

was estimated as 0.78 (yr-1) (Fig. 5). Based on the 

calculated parameters, the von Bertalanffy equations 

for this species (length and weight) in the northern 

Oman Sea, Iran, are as follows. 



135 
 

Int. J. Aquat. Biol. (2023) 11(2): 131-140 

 

  

Figure 3. Length and Frequency (A) as well as Length - weight relationship (B) of total Babylonia spirata in the northern Oman Sea, Iran. 
 

Species (sex) Method L∞ (mm) K ) 1-yr ) ot M F Z E 

(Male) spirata B. ELEFAN  76 0.3 -0.42 0.4 1.79 2.19 0.82 

B. spirata (Female) ELEFAN  68 0.54 -0.24 0.7 2.3 3 0.77 

B. spirata (Total) ELEFAN  71 0.35 -0.37 0.5 1.75 2.25 0.78 
 

Table 1. Comparison of population dynamics values of Babylonia spirata with two methods (Shepherd method and ELEFAN method) (L∞= 
infinite length, K= Growth rate, to= time that length is zero, M= Natural mortality, F= Fishing mortality, Z= Total mortality, E= Exploitation 
rate). 

Figure 4. Growth curve derived from the structure of population of Babylonia spirata from the northern Oman Sea (Iran). The growth curve plot 
shows reconstructed frequencies, with negative and positive values as white and black colored histograms, respectively. The background shading 

shows runs of peaks, with positive peaks in blue, negative peaks in red, and values of zero in white. The different color backgrounds were added 

in order to help visualizing sign and magnitude of the bin values. The sum of all positive peaks is called the “available sum of peaks” (ASP), 

which represents a maximum possible score. The “estimated sum of peaks” (ESP) is the sum of peak values crossed by the growth curves. 
 



136 
 

Hashemi et al./ Length-based fishery status and population dynamics of Spiral Babylon 

Lt = 71 (1 - exp (-0.35 (t + 0.2))) 

Wt = 79 (1 - exp (-0.35 (t + 0.2))) ^ 
2.70 

The relative yield per recruitment (Y/R) and 

relative biomass per recruitment (B'/R): Based on 

length at first capture (Lc = 29 mm), which is 50% 

the probability of catching shell, relative production 

per recruitment and relative biomass per recruitment 

were estimated as Y'/R = 0.009 and B'/R = 0.2, 

respectively. The results demonstrate an exploitation 

rate (U) of 0.70 and the fishing mortality at 

maximum sustainable yield (Fmsy) equaled to 1 for 

this stock (Fig. 5). 

Length-based indicators (LBI) and length-based 

spawning potential ratio (LBSPR): Length-based 

reference point is the percentage of shell caught at 

the optimum length for harvest (Lopt=48 mm, 

Popt=0.10), (LF=M=40 mm, Popt=0.13) and percentage 

of mega-spawners in the catch (Lmega=53 mm, 

Pmega=0.04), respectively. Based on the LBSPR 

assessment model (Fig. 6), Lopt was estimated at 

about 0.3 (0.27-0.33). The ratio of Pmega<0.1, Lmean/ 

Lopt <1 and Lmean /LF=M <1 were estimated. 

A 

B 

Figure 5. Exploitation coefficient curve (A) and Relative Yield per Recruit (Y'/R) and Relative Biomass per Recruit (B'/R), FMSY (B) of Babylonia 
spirata (total) in the northern Oman Sea, Iran (FMSY = Fishing mortality rate of Maximum sustainable yield, F 0.5= fishing mortality rate at which 

the slope of the yield-per-recruit curve is only half the slope of the curve at its origin, F0.1= fishing mortality rate at which the slope of the yield-

per-recruit curve is only one percent the slope of the curve at its origin). 
 

 

 

Figure 6. The LBSPR of Babylonia spirata (total) in the northern Oman Sea, Iran. 
 

 

 



137 
 

Int. J. Aquat. Biol. (2023) 11(2): 131-140 

 

Discussion 

We have managed to study a series of population 

dynamics values of the species B. spirata and this is 

the first time in the study area despite economic and 

ecological importance. Babylonia spirata is one of 

the economically valuable species in the southern 

and northern Oman Sea, Iran.  

Life-history traits of B. spirata: The L-W 

relationship showed negative allometric growth and 

the female was heavier than males in the same length 

group. It seems the growth curve (length and weight) 

of B. spirata (total) slows down after two years and 

allometric growth is common in Gasterpoda (Carare 

and Surugiu, 2020).  The relationship between the 

length and weight of Strombus canarium was 

calculated as a = 0.0000018 and b = 3.2 (R2 = 0.85) 

for males, and for females, a = 0.0000015 and b = 

3.3 (R2 = 0.81) (Cob et al., 2009).  The observed size 

pattern was influenced by latitude and local spatial 

responses to factors such as the substrate changing 

the morphometric variables of the snail. The 

morphological characteristics such as length and 

weight relationship, are greatly affected by latitude, 

and it seems that as the latitude increases, the size of 

the snail becomes larger (Matos et al., 2020). In 

addition, the morphological characteristics of snails 

are influenced by various factors such as 

temperature, available food, intra-group 

competition, substrate, climate, tidal conditions, and 

specific characteristics of habitat  (Matos et al., 

2020). 

The L-W relationship is of great importance in 

fishery assessments (Haimovic and Velasco, 2000). 

According to Marthin (1994), the range of "b" could 

be from 2.5 to 4 and it is believed b = 3 with 

isometric growth. Also, the functional regression b-

value shows the body form, and it is directly related 

to the weight affected by ecological factors such as 

temperature, food supply, spawning conditions, and 

also other factors, such as sex, age, fishing time, and 

area and fishing vessels. 

Population dynamic of B. spirata: Comparisons of 

the population parameters of B. spirata with other 

studies in different parts of the world (Table 2) show 

that the infinite length and growth coefficient of 

males from different species are smaller than 

females. Also, the infinite length and growth 

coefficient of different species changes in various 

regions (Table 2). Differences in the range of the 

infinite length and growth rate are influenced by the 

ecological differences of each region (King, 2007). 

The growth rate is expected to be higher in the 

tropical zones. Higher K-values for species are 

common in tropical waters due to their 

poikilothermic nature and result in a higher 

metabolic rate in relation to high temperate 

(Hashemi, 2020). In general, the difference in the 

infinite length and growth rate in various regions 

might be due to the quantity and quality of food and 

climatic conditions (Bartulovic et al., 2004). Various 

factors can also affect holothurian growth including 

age, sex, season, year, type of feeding, physiological 

conditions, differences in food availability, and 

reproductive period (Lalèyè, 2006). 

References Species/Region L∞ (mm) K (yr-1) M F Z E 

Mohan (2007) B. spirata, India 68 1.08 1.61 4.44 6.05 0.73 

Mohan (2007) B. zeylanica, India 76 1.15 1.65 3.37 5.02 0.67 

Cob et al. (2009) Strombus canarium, Malaysia 
70 (F) 

69 (M) 

1.5 (F) 

1.2 (M) 

0.95 (F) 

0.86 (M 

1.61 (F) 

1.86 (M) 

2.56 (F) 

2.86 (M) 

0.62 (F) 

0.65 (M 

Arrighetti et al. (2012) 
Olivancillaria deshayesiana, 

Argentina 
38 0.14 0.36 0.29 0.65 0.44 

Daza-Guerra et al. (2018) Cittarium pica, Colombia 94 0.32 0.60 1.11 1.71 0.65 

Present study (2022) B. spirata, Oman Sea (Iran) 71 0.35 0.5 1.75 2.25 0.78 

 

Table 2. Comparison of population dynamic values of Babylonia spirata with other studies around the world. For details of the abbreviation 
name of the values see Table 1. 



138 
 

Hashemi et al./ Length-based fishery status and population dynamics of Spiral Babylon 

The natural mortality rate of B. spirata in this 

study was less than that of fishing mortality. The 

ratio of fishing mortality to maximum sustainable 

yield (F/FMSY) was over than one. To recall that if 

the value of F/FMSY is more than 1, it indicates that 

there is overfishing (Arrizabalaga et al., 2012). The 

exploitation coefficient was over 0.5, indicating that 

the amount of capture fisheries was more than the 

optimum level. The exploitation coefficient should 

not be greater than 0.5 or, the fishing mortality 

should not exceed natural mortality otherwise they 

indicate overfishing (King, 2007; Hashemi et al., 

2021; Hashemi and Doustdar, 2022). The most 

important factors affecting the pressure on stocks 

are, first, the amount of catch and harvest of the 

stocks, and second, the environmental factors that 

affect survival and access to the fishery resources 

(Mateus and Estupinan, 2002).  

Length-based indicators (LBI) and length-based 

spawning potential ratio (LBSPR): The Lmean/Lopt 

index has values less than one (about 0.7), which 

means the presence of overfishing, and also 

Lmean/LF=M index has values less than one (about 0.8) 

and Pmega less than 0.1 is calculated which the show 

considered undesirable and index range optimal of 

Lmean/Lopt and Lmean/LF=M is close to one and Pmega is 

about 0.3-0.4 (Cousido-Rocha et al., 2022). 

The LBSPR rate of this species was calculated at 

0.3 (30%). LBSPR gives estimates of spawning 

potential ratio (SPR), where values below 0.2 (≈ 0.5 

B/Bmsy) indicate depletion and values above 0.4 (≈ 

1.0 B/Bmsy) indicate good stock status. The 95% 

confidence limits provided by LBSPR are 

unrealistically narrow, sometimes close to 

deterministic, which partly explains their very low 

matching score.  In conclusion, the LBSPR model is 
a promising tool for the length-based assessment of 

data-limited fish stocks (Hordyk et al., 2015). With 

regards to cheap and simple to collect measurements 

of the length composition of an exploited stock and 

also life-history information, models, and methods 

have been developed in recent years, for instance, the 

length-based spawning potential ratio method 

(LBSPR) (Hordyk et al., 2015). 

Management and conservation of B. spirata in the 

Oman Sea: It is recommended that appropriate 

instructions be established for the harvesting and 

management of this species in the area. This 

parameter is needed for fisheries management and 

conservation of exploited this species. The present 

study showed that these stocks have reached 

'overfished' status. The results of the present study 

could help in the management and sustainable 

exploitation of this species’ stocks.  

 

Acknowledgments 

We would like to thank Prof. Bahmani, the manager 

of the Iranian Fisheries Science Research Institute 

(IFSRI), and very grateful to the experts of the 

Offshore Fisheries Research Center (OFRC, 

Chabahar), for helping with the project work.  

 

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	U = 1 - (LC / L∞)  ;  m = (1 - E) / ( M / K) = ( K / Z) ;  E = F / Z (9)