ISSN 1978-3477, eISSN 2087-8575 Volume 10, Number 4, December 2016 I N D O N E S I A Accredited at level “A” until Februari 2019 No.040/P/2014 Patron Siswa Setyahadi, 2020 Chief Editor Debbie S Retnoningrum, 2020 Editorial Board Members Antonius Suwanto, 2020 Brett Neilan, 2020 Dessy Natalia, 2020 Diana E Waturangi, 2020 Endang Purwantini, 2020 Friedhelm Meinhardt, 2016 Managing Editors Is Helianti, 2020 Electronic Editor Iman Rusmana, 2020 Is Helianti, 2020 Business Manager Diana Nurani, 2020 Netty Widyastuti Sigit, 2020 Editorial Office Indonesian Society for Microbiology (Sekretariat PERMI) Room 124/TMC 2 DRN, Puspiptek-Serpong, Tangerang Selatan 15314, Indonesia Phone: +62-21-7560536 ext 7119 Fax: +62-21-7560694 E-mail: microbiology.indonesia@gmail.com URL: http://jurnal.permi.or.id/index.php/mionline Publisher Indonesian Society for Microbiology. Published in March, June, September, and December Subscription Prices for One Year, not including shipping and handling Indonesian Overseas Individual rate (IDR) 150 000,- 200 000,- Institutional rate (institution or library) (IDR) 240 000,- 400 000,- Bank Bank Mandiri Cabang Menara Thamrin, Jakarta, Acc PERMI; Acc No 103-0002080774 Printed by: CV. Istiqom Print GENERAL EXECUTIVE BOARD OF INDONESIAN SOCIETY FOR MICROBIOLOGY 2015-2019 Advisory Board: Prof. Dr. Pratiwi Sudarmono, PhD, Sp.MK; Dr. Mohammad Dimyati; Prof. Dr. Endang Sukara; Prof. dr. Amin Soebandrio, PhD, Sp.MK; Prof. Dr. Ir. Betty Sri Laksmi Jenie, MS; Prof. Dr. Ir. Antonius Suwanto, MSc; Prof. Dr-Eng. Eniya Listiani Dewi, B. Eng, M.Eng; President: Dr. Siswa Setyahadi; Vice President: Prof. Fedik A Rantam, PhD; General Secretary: Diana Nurani, M.Si; Vice General Secretary: Drs. Nuki B Nugroho, M.Si; Treasurer: Dr. Niknik Nurhayati; Dr. Sylva Abraham; Scientific and Publication Committee: Dr. Debbie S. Retnoningrum; Dr. Is Helianti; Dr. Iman Rusmana; Prof. Dr. Marlina, Apt; Prof. Dra. Netty Widyastuti, M.Si; Dr. Ir. Nur Hidayat, MP; Drh. Mahdi Abrar; Dr. Supriyadi; Dr. Astutiati Nurhasanah; Tarwadi, MSc.; Certification Committee: Dr. Ir. Trismilah, M.Si; Dr. Erwahyuni Endah Prabandari; Ir. Dwi Kusuma Indriani, MP.; Dra. Dini Ryandini, M.Si; Dr. Ir. Maman Turjaman, DEA; Dr. Ernawati Giri Rahma; Dra. Harmastini Sukiman, M.Agr; dr. Purwati SpPD, Ph.D; Dr. Wahyu Purbowasito; Organization Advancement and Networking: Dr. Puspita Lisdiyanti; Prof. Dr. Ir. Endang S. Rahayu, MS; Sri Harjati Suhardi, PhD; Lia Yulia Budiarti, MKes; Dr. Chaidir; Dr. Retno Indrawati, drg., M.Si; Alit Pangestu, STP; Promotion and Advocacy Committee: Dra. MG. Isworo Rukmi, M.Kes; Jimmy Hariantono, PhD; Yawarsa; Gianina; Asri Sulfianti, MSi Astutiati Nurhasanah, 2020 Kartini Kramadibrata, 2020 Maggy Thenawijaya Suhartono, 2016 Maria Inge Lusida, 2016 Neung Tiaamroeng, 2020 Norio Kurosawa, 2020 Raija Laiho, 2016 Wellyzar Sjamsuridzal, 2020 Yuan Kun Lee, 2020 Yaya Rukayadi, 2020 Available online at http://jurnal.permi.or.id/index.php/mionline ISSN 1978-3477, eISSN 2087-8575 Vol 10, No 4, December 2017 ACKNOWLEDGMENT VOLUME 10 Siti Khodijah Chaerun, Institut Teknologi Bandung, Triwibowo Yuwono, Universitas Gadjah Mada, Indonesia Valan Arasu, King Saud University, Saudi Arabia Verawat, Biotec National Center for Genetic Engineering and Biotechnology, Thailand Wahono Sumaryono, Badan Pengkajian dan Penerapan Teknologi, Indonesia Wo ro H a s t u t i S a t y a n t i n i , Universitas Airlangga, Indo- nesia Wangsa Tirta Ismaya, Dexa Laboratories of Biomolecular Sciences, Indonesia Wellyzar Sjamsuridzal, Univer- sitas Indonesia, Indonesia Yaya Rukayadi, Universiti Putra Malaysia, Malaysia Yusra, Universitas Bung Hatta, Indonesia Abu Bakar Salleh, Universiti Putra Malaysia, Malaysia Achmad Dinoto, Lembaga Ilmu P e n g e t a h u a n I n d o n e s i a , Indonesia Alucia Anita Artarini, Institut Teknologi Bandung, Indone- sia Amarila Malik, Universitas Indonesia, Indonesia Antonius Suwanto, Institut Pertanian Bogor, Indonesia Astutiati Nurhasanah, Badan Pengkajian dan Penerapan Teknologi, Indonesia Budiasih Wahyuntari, Badan Pengkajian dan Penerapan Teknologi, Indonesia Brett Neilan, University of Newcastle, Australia Catur Riani, Institut Teknologi Bandung, Indonesia D e b b i e S R e t n o n i n g r u m , Institut Teknologi Bandung, Indonesia Dominngus Male, Universitas Patimura, Indonesia Ekowati Chasanah, Department o f M a r i n e A ff a i r s a n d Fisheries, Indonesia Ekachai Chukeatirote, Mae Fah Luang University,Thailand Eny Ida Riyanti, Ministry of Agriculture, IndonesiaElfah- mi, Institut Teknologi Ban- dung, Indonesia E r n a w a t i A r i f i n G i r i - Rachman, Institut Teknolo- gi Bandung, Indonesia Erni Martani, Universitas Gadjah Mada, Indonesia Fernita Puspasari, Institut Teknologi Bandung, Indonesia Hideki Takahashi, Tohoku University, Japan Hiroaki Okamoto, Jichi Medical University, Japan I Made Sudiana, Lembaga Ilmu Pengetahuan Indonesia, Indo- nesia Iman Rusmana, Institut Pertanian Bogor, Indonesia Is Helianti, Badan Pengkajian dan Penerapan Teknologi, Indonesia Jignesh patel, University of Texas Medical Branch at Galveston, USA Kartini Kramadibrata, Lembaga Ilmu Pengetahuan K h o m a i n i H a s a n , I n s t i t u t Teknologi Bandung, Indonesia Mahyudin Abdul Rachman, Badan Pengkajian dan Pene- rapan Teknologi, Indonesia Maria Inge Lusida, Universitas Airlangga, Indonesia Maria Magdalena Coman, Università di Camerino, Italia Maelita Ramdani Moeis, Institut Teknologi Bandung, Indonesia Misri Gozan, Universitas Indone- sia, Indonesia Moritz Müller, Swinburne University of Technology, Malaysia N Venkatesh, St. Joseph’s College of Engineering, OMR Chennai India Nagendra P. Shah, The University of Hong Kong, Hong Kong Neung Tiaamroeng, Suranaree University of Technology, Thailand N i k n i k N u r h a y a t i , B a d a n Pengkajian dan Penerapan Teknologi, Indonesia Oslan Jumadi, Universitas Negri Makasar, Indonesia Puspita Lisdiyanti, Lembaga Ilmu Pengetahuan Indonesia, Indonesia Prasetyawan Yunianto, Badan Pengkajian dan Penerapan Teknologi, Indonesia Priyo Wahyudi, Badan Pengkajian dan Penerapan Teknologi, Indonesia Sabar Pambudi, Badan Pengka- jian dan Penerapan Teknologi, Indonesia Sartaj Alam Syed, The University of Agriculture Peshawar, Pakistan Sathianeson Satheesh, King Abdulaziz University, Saudi Arabia Silva Abraham, Badan Pengkajian dan Penerapan Teknologi, Indonesia Sinosh Skariyachan, Dayananda Sagar Institutions, India Siswa Setyahadi, Badan Pengka- jian dan Penerapan Teknologi a.FRONT COVER 10(4)-DECEMBER