Journal of the Oklahoma Native Plant Society, Volume 2, Number 1, December 2002 ISSN 1536-7738 Oklahoma Native Plant Record Journal of the Oklahoma Native Plant Society Volume 2, Number 1, December 2002 Oklahom a N ative Plant Record Journal of the Oklahoma Native Plant Society Vol. 2 No. 1 December 2002 ISSN 1536-7738 Managing Editor, Sheila A. Strawn Technical Editor, Patricia Folley Technical Advisor, Bruce Hoagland CD-ROM Producer, Chadwick Cox Website: http://www.usao.edu/~onps/ The purpose of the ONPS is to encourage the study, protection, propagation, appreciation and use of the native plants of Oklahoma. Membership in ONPS shall be open to any person who supports the aims of the Society. ONPS offers individual, student, family, and life membership. Officers and Board Members President: Patricia Folley Vice-president: Chadwick Cox Conservation Chair: open Secretary: Maurita Nations Publicity Co-chairs: Treasurer: Mary Korthase Ruth Boyd & Betty Culpepper Board Members: Marketing Chair: Lawrence Magrath Berlin Heck Photo Contest: Paul Reimer Iris McPherson Ann Long Award Chair: Paul Reimer Sue Amstutz Harriet Barclay Award Chair: James Elder Constance Taylor Paul Reimer ONPS Service Award Chair: Sue Amstutz Lawrence Magrath Newsletter Editor: Chadwick Cox Librarian: Bonnie Winchester Northeast Chapter Chair: James Elder Website Manager: Chadwick Cox Central Chapter Chair: Judy Jordan Cross-timbers Chapter Chair: Ronald Tyrl Cover: Gaillardia pulchella (Indian Blanket) Historian: Lynn Allen Photo by Paul Buck, April 25, 1981. Winner of the first ONPS Photo Contest in 1988. Articles (c) The Authors Journal compilation (c) Oklahoma Native Plant Society Except where otherwise noted, this work is licensed under a Creative Commons Attribution- NonCommercial-ShareAlike4.0 International License, https://creativecommons.org/licenses/ by-nc-sa/4.0/, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly attributed, not used for commercial purposes, and, if transformed, the resulting work is redistributed under the same or similar license to this one. https://doi.org/10.22488/okstate.17.100009 2 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Oklahoma Native Plant Record Volume 2, Number 1 Table of Contents Forward ...................................................................................................... 3 Ms. Patricia Folley, ONPS President Vascular Plants of the Wichita Mountains ............................................ 4 Dr. Paul Buck Floristic List for Comanche County, Oklahoma ................................ 22 Dr. Bruce W. Hoagland Schoenoplectus hallii and S. saximontanus ................................................... 54 Wichita Mountains Wildlife Refuge Survey 2000 Dr. Lawrence K. Magrath Pontotoc Ridge Floristic Survey: 1999 ................................................. 64 Dr. Forrest L. Johnson Ms. Patricia Folley (ed.) ONPS Editorial ......................................................................................... 82 Water, Soil, and Plant Diversity in Oklahoma Dr. Sheila A. Strawn Oklahoma Native Plant Record 3 Volume 2, Number 1, December 2002 Foreword Continuing our goal of publishing historical and current floristic surveys of the different areas of Oklahoma, this 2002 edition features Comanche County and the Wichita Mountains of southwestern Oklahoma. These beautiful pink granite remains of ancient high mountains are completely contained within our state and have an interesting history from the viewpoints of Native Americans and the later European settlers. Several decades ago a young botanist named Paul Buck spent summers sleeping on concrete picnic tables at the Wichita Wildlife Refuge while making the best-known specific survey of the flora of that site. The resulting list of plants is still in use by visitors to the refuge and others interested in the plants of that region. We have published that survey for you here, for use whenever you visit the Refuge. More recently, the U.S. Army at Ft. Sill, which adjoins the refuge, contracted with the Oklahoma Biological Survey to make a similar study of the flora of the military reservation. From voucher specimens taken during those surveys and from many other botanists who visited the area for the love of its wild beauty, the Oklahoma Natural Heritage Inventory has compiled a floristic list for the whole of Comanche County. Dr. Bruce Hoagland, at the Oklahoma Biological Survey, has provided that list. We hope it will inspire today’s botanists to continue to monitor extant species on the Refuge and surrounding areas. The Wichita Wildlife Refuge was established by President William McKinley in 1901, with 57,120 acres of land. Currently, about 22,500 acres are open to the public. The Refuge is a part of the National Wildlife Refuge system, and maintained by the federal government. A visitors’ center provides assistance, educational material, and a library and gift shop. But a remarkable organization of volunteers manages the prodigious task of everyday contact with the many visitors who enter the Refuge every day of the year. By planned tours, educational programs, and hours of labor maintaining trails, the Friends of the Wichitas have made the Refuge a comfortable place for people without reducing the quality of the environment for wildlife. A sister organization of ONPS, the Friends and we share many members. Through the years Oklahoma Native Plant Society volunteers, especially Dr. Buck, have helped staff the field trips. In the process, some of us who might not have made the trip to Comanche County quite so often before, have come to love it as part of our ‘home state’. Next year, we hope to feature another, and very different section of this varied state: the Ozarks. All who have, or know about, material on that area that might be suitable for this publication, should contact our editor, Sheila Strawn. Also planned for future issues are lists of the plants reported at specific sites by our intrepid teams of field-trippers. Help us make this publication grow! Patricia Folley President, ONPS August, 2002 4 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 VASCULAR PLANTS OF THE WICHITA MOUNTAINS WILDLIFE REFUGE (1977) Paul Buck Department of Life Sciences The University of Tulsa Tulsa, Oklahoma Introduction This list is an effort to update a popular list of the vascular plants of the WichitaMountains Wildlife Refuge prepared by the author approximately ten years ago. The earlier work was compiled from lists published by Eskew (1938), Osborn and Allen(1949) and from studies of several regional herbaria. The changes over the previous list reflect new species found during field activities in the Refuge through the years,additions reported in the literature and modifications of scientific names as published in floras, keys, and manuals. Common names are always a problem,frequently changing several times within relatively small geographic areas. In an effort to avoid this difficulty the common names have been taken from A Guide to Plant Names, (Rechenthin, 1954) which was prepared for use by the regional Soil Conservation Service. The very few not listed in this publication were obtained from Fernald (1950)and the updated version of Britton and Brown (Gleason 1952). The asterisks (*) indicate species that are questionable as present inhabitants of the Refuge for a couple of reasons: (1) Many were reported in the area by earlier workers but recent literature indicates their ranges do not extend into the Wichita Mountains system. It might be inferred these were misidentified,and in several cases where the specimens were located, this proved to be the case. Buck, P. https://doi.org/10.22488/okstate.17.100011 (2) During the 1930’s thousands of seedlings of non-native woody species were introduced and planted (example: Mahonia aquifolium, Barberry; Spiraea arguta, Spirea; Hibiscus syriacus, Rose of Sharon; Lagerstroemia indica, Crape myrtle, etc.). Most of these were subsequently moved to other locations within the southwest. Probably none of those left in the Refuge survived the competition of the native flora. The identity of the maple within the Refuge has been a subject of debate for many years. Opinions have been divided between the bigtooth maple (Acer grandidentatum) and sugar maple (A. saccharum). The writer felt, when the earlier list was prepared, the Wichita maple was the latter and deleted the bigtooth maple from the list. Dent(1969) made a study of the sugar maples of Oklahoma and concluded the Caddo Canyon and Wichita Mountains groups are both Acer saccharum. References Cited Dent, Thomas Curtis 1969 Relationships of two isolated groups of sugar maple in central Oklahoma to eastern and western species. Unpublished PhD Dissertation, University ofOklahoma, Norman. Eskew, Cletis T. 1938 The flowering plants of the Wichita Mountains Wildlife Refuge, Oklahoma. Amer. Mid. Nat. 20:695-703. Fernald, M.L. 1950 Gray’s Manual of Botany. American Book Company, New York. Oklahoma Native Plant Record 5 Volume 2, Number 1, December 2002 Buck, P. Gleason, Henry A. 1952 The New Britton and Brown Illustrated Flora of the Northeastern United States and Adjacent Canada. The New York Botanical Garden, New York. Osborn, B. and P. F. Allan 1949 Vegetation of an abandoned prairie-dog town in tall-grass prairie. Ecology 30:322-332. Rechenthin, C.A. 1954 A Guide to Plant Names in Texas, Oklahoma, Louisiana, Arkansas. United States Department of Agriculture, Soil Conservation Service. Species List Equisetum laevigatum EQUISETACEAE Kansas horsetail Selaginella peruviana SELAGINELLACEAE Sheldon selaginella Selaginella rupestris Rock selaginella Isoetes butleri ISOETACEAE Butler quillwort Isoetes melanopoda Blackfoot quillwort Asplenium trichomanes POLYPODIACEAE Spleenwort Cheilanthes eatoni Eaton lipfern Cheilanthes fendleri* Fendler lipfern Cheilanthes lanosa Hairy lipfern Cheilanthes lindheimeri Lindheimer lipfern Cheilanthes vestita Lipfern Cheilanthes wootoni* Wooton lipfern Dryopteris marginalis Standley’s lipfern Notholaena standleyi Standley’s lipfern Pellaea atropurpurea Purple cliffbrake Pellaea ternifolia Wright’s cliffbrake Woodsia obtuse Woodsia Pilularia americana MARSILEACEAE American pillwort Marsilea mucronata Pepper wort Juniperus virginiana PINACEAE Eastern Red Cedar Pinus echinata* Shortleaf pine Pinus edulis* Nut pine Pinus ponderosa* Western Yellow pine Pinus taeda* Loblolly pine Thuja orientalis* CUPRESSACEAE Oriental Arbor-Vitae Typha angustifolia TYPHACEAE Narrow leaf cattail Typha domingensis Cattail Typha latifolia Cattail ZOSTERACEAE Potamogeton amplifolius Largeleaf pondweed Potamogeton diversifolius Waterthread pondweed Potamogeton nodosus Longleaf pondweed Potamogeton pusillus Baby pondweed NAJADACEAE Najas guadalupensis Southern naiad ALISMATACEAE Echinodorus cordifolius Erect burhead Sagittaria calycina Longlobe arrowhead Sagittarria latifolia Arrowhead Agropyron repens* POACEAE Couch grass Agrostis ellottiana 6 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Buck, P. Elliott bentgrass Agrostis hymenalis Winter bentgrass Alopecurus caroliniana Carolina foxtail Andropogon barbinodis Cane bluestem Andropogon gerardi Big bluestam Andropogon saccharoides Silver bluestem Andropogon scoparius Little bluestem Aristida dichotoma var. dichotoma Churchmouse threeawn Aristida dichotoma var. curtissii Curtis threeawn Aristida longiseta Red threeawn Aristida longespica Slimspike threeawn Aristida oligantha Oldfield threeawn Aristida purpurea Purple threeawn Aristida wrightii Wright threeawn Bouteloua curtipendula Sideoats grama Bouteloua gracilis Blue grama Bouteloua hirsute Hairy grama Bouteloua rigidiseta Texas grama Bromus japonicus Japanese brome Bromus arvensis* Field brome Bromus purgans Canada brome Bromus tectorum Cheatgrass Bromus uniloides Rescuegraas Buchloe dactyloides Buffalo grass Cenchrus pauciflorus Mat sandbur Chloris verticillata Tumble windmillgrass Chloris virgata Showy chloris Cynodon dactylon Bermuda grass Dactylis glomerata Orchard grass Digitaria sanguinalis Hairy crabgrass Diplachne fascicularis Bearded spranglegrass Echinochloa crusgalli Barnyard grass Elusine indica Goosegrass Elymus canadensis Canada wildrye Elymus virginicus Virginia wildrye Eragrostis capillaris Lacegrass Eragrostis curtipedicellata Gummy lovegrass Eragrostis hypnoides Teal lovegrass Eragrostis intermedia Plains lovegrass Eragrostis cilianensis Stinkgrass Eragrostis oxylepsis var. oxylepis Red lovegrass Eragrostis oxylepsis var. beyrichii Wichita lovegrass Eragrostis pectinacea Carolina lovegrass Eragrostis pilosa India lovegrass Eragrostis reptans Creeping lovegrass Eragrostis sessilispica Tumble lovegrass Eragrostis spectabilis Purple lovegrass Eragrostis trichodes Sand lovegrass Eriochloa contracta Prairie cutgrass Festuca octoflora Sixweeks fescue Festuca pratensis Suiter fescue Festuca versuta Texas fescue Hordeum pusillum Little barley Leersia oryzoides Rice grass Leptochloa dubia Oklahoma Native Plant Record 7 Volume 2, Number 1, December 2002 Buck, P. Green sprangletop Leptoloma cognatum Fall witchgrass Limnodea arkansana Ozark grass Manisuris cylindrica Carolina jointtail Melica nitens Threeflower melic Muhlenbergia capillaris Hairawn muhly Muhlenbergia frondosa Wirestem muhly Muhlenbergia racemosa Green muhly Panicum agrostoides Redtop panicum Panicum anceps Beaked panicum Panicum capillare Witchgrass Panicum depauperatum Poor panic grass Panicum dichotomiflorum Fall panicum Panicum lanuginosum Wooly panicum Panicum linearifolium Slimleaf panicum Panicum malacophyllum Shortleaf panicum Panicum obtusum Vine-mesquite Panicum obligosanthes var. scribnerianum Scribner panicum Panicum perlongum Elongate panic grass Panicum philadelphicum Philadelphia witchgrass Panicum virgatum Switchgrass Paspalum pubiflorum var. publiflorum Hairyseed paspalum Paspalum publiflorum var. glabrum Smoothseed paspalum Paspalum setaceum Fringeleaf paspalum Phalaris caroliniana Carolina canary grass Poa annua Annual bluegrass Poa arachnifera Texas bluegrass Poa compressa Canada bluegrass Schedonnardus paniculatus Tumblegrass Setaria geniculata Knotroot bristlegrass Setaria viridis Green bristlegrass Sorgastrum nutans Indiangrass Sorghum halapense Johnson grass Spartina pectinata var. suttiei Prairie cordgrass Sphenopholis obtusata Prairie wedgescale Sporobolus airoides Alkali Sacaton Sporobolus asper Tall dropseed Sporobolus cryptandrus Sand dropseed Sporobolus pyramidatus Whorled dropseed Sporobolus vaginiflorus var. vaginiflorus Poverty dropseed Sporobolus vaginiflorus var. neglectus Puffsheath dropseed Trichachne californica Arizona cottontop Tridens albescens White tridens Tridens flavus Purpletop Tridens muticus var. muticus Slim tridens Tridens muticus var. elongatus Rough tridens Tridens pilosus Hairy tridens Tridens strictus Longspike tridens Tripsacum dactyloides Eastern gamagrass Uniola latifolia Broadleaf uniola Bulbostylis capillaris CYPERACEAE Hairsedge Carex amphibola Narrowleaf sedge Carex annectens Yellowfruit sedge 8 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Buck, P. Carex austrina Southern sedge Carex blanda Woodland sedge Carex festucacea Fescue sedge Carex frankii Frank sedge Carex gravida var. lunellina Lunnell sedge Carex joori Hummock sedge Carex microrhyncha Littlesnout sedge Carex praegracilis Slender sedge Carex stricta var. elongata Emory sedge Carex vulpinoidea Fox sedge Cyperus acuminatus Taperleaf flatsedge Cyperus aristatus Bearded flatsedge Cyperus erythrorhizos Redroot flatsedge Cyperus esculentus Chufa Cyperus filiculmis Slender flatsedge Cyperus odoratus Fragrant flatsedge Cyperus ovularis Globe flatsedge Cyperus schweinitzii Schweinitz flatsedge Cyperus setigerus Bristled spike flatsedge Cyperus strigosus False nut sedge Cyperus virens Green flatsedge Eleocharis compressa Flatstem spikesedge Eleocharis engelmanii Englemann spikesedge Eleocharis macrostachya Longspike spikesedge Eleocharis montevidensis Sand spikesedge Eleocharis obtusa Blunt spikesedge Eleocharis parvula Dwarf spikesedge Eleocharis quadrangulata Squarestem spikesedge Eleocharis tenuis Slender spikesedge Fimbristylis spadicea Plains fimbry Fimbristylis vahlii Vahl fimbry Fuirena simplex Western umbrella sedge Hemicarpha micrantha Hemicarpha Scirpus acutus Hardstem bulrush Scirpus americanus American bulrush Scirpus atrovirens Green bulrush Scirpus lacustris Softstem bulrush Scirpus lineatus Rusty bulrush Scirpus paludosis Alkali bulrush Scirpus pauciflora Fewflower nutrush Arisaema dracontium ARACEAE Dragonroot or Jack-in-the-Pulpit Lemna minor LEMNACEAE Duckweed Commelina erecta COMMELINACEAE var. angustifolia Narrowleaf dayflower Commelina virginica Virginia dayflower Tradescantia hirsutiflora Hairyflower spiderwort Tradescantia occidentalis Prairie spiderwort Tradescantia ohiensis Smoothstalk spiderwort Tradescantia tharpii Tharp spiderwort Heteranthera limosa PONTEDERIACEAE Blue mudplantain Juncos acuminatus JUNCACEAE Knotleaf rush Oklahoma Native Plant Record 9 Volume 2, Number 1, December 2002 Buck, P. Juncus balticus Baltic rush Juncus interior Inland rush Juncus marginatus Grassleaf rush Juncus tenuis var. tenuis Poverty rush Juncus tenuis var. dudleyi Dudley rush Juncus torreyiTorrey rush Allium canadense LILIACEAE Canada garlic Allium drummondii Drummond onion Allium stellatum Prairie onion Androstephium coeruleum Blue funnellily Camassia scilloides Atlantic camass Erythronium americanum Fawn lily Nothoscordum bivalve Yellow false garlic Smilax bona-nox Saw greenbrier Smilax herbacea Carrionflower Smilax rotundifolia Greenbrier Smilax tamnoides Bristly greenbrier Yucca glauca Small soapweed Agave lata AMARYLLIDACEAE Agave Zephyranthes brazosensis Evening start or Rain lily Nemastylis geminiflora IRIDACEAE Prairie iris Sisyrinchium angustifolium Blueeye grass Spiranthes cernua ORCHIDACEAE Nodding ladiestresses Populus deltoids SALICACEAE Cottonwood Salix amygdaloides Peachleaf willow Salix caroliniana Ward's willow Salix interior Sandbar willow Salix matsudana* Willow Salix nigra Black willow Carya cordiformis JUGLANDACEAE Bitternut hickory Carya illinoensis Pecan Juglans nigra Black walnut Juglans rupestris Little walnut Quercus falcate FAGACEAE Southern red oak Quercus macrocarpa Bur oak Quercus marilandica Blackjack oak Quercus mohriana Mohrs oak Quercus muhlenbergii Chinquapin oak Quercus rubus var. borealis Red oak Quercus shumardii var. shumardii Shumard’s oak Quercus shumardii var. schneckii Schneck's oak Quercus stellata Post oak Quercus velutina Black oak Quercus virginiana Live oak Celtis laevigata ULMACEAE Sugar hackberry Celtis occidentalis Hackberry Celtis reticulata Netleaf hackberry 10 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Buck, P. Ulmus americanus American elm Ulmus pumila* Chinese elm Ulmus rubra Slippery elm Maclura pomifera MORACEAE Osage orange Morus microphylla Texas mulberry Morus rubra Red mulberry Morus tatarica* Russian mulberry Boehmeria cylindrica URTICACEAE Smallspike falsenettle Parietaria pensylvanica Pennsylvania pellitory Comandra pallida SANTALACEAE Western comandra LORANTHACEAE Phoradendron serotinum Mistletoe ARISTOLOCHIACEAE Aristolochia tomentosa Wooly dutchmanpipe Eriogonum annum POLYGONACEAE Annual wild buckwheat Eriogonum longifolium Longleaf wild buckwheat Polygonum aviculare Prostrate knotweed Polygonum bicorne Longstype smartweed Polygonum coccinium Bigroot smartweed Polygonum convolvulus Dullseed cornbind Polygonum hydropiper Marshpepper smartweed Polygonum hydropiperoides var. hydropiperoides Swamp smartweed Polygonum hydropiperoides var. opelousanum Opelousas swamp smartweed Polygonum lapathifolium Curlytop smartweed Polygonum pensylvanicum Pennsylvania smartweed Polygonum punctatum Dotted smartweed Polygonum ramosissimum Bushy knotweed Polygonum scandens Hedge cornbind Polygonum tenue Pleatleaf knotweed Rumex altissimus Pale dock Rumex crispus Curly dock Chenopodium album CHENOPODIACEAE Lambsquarters Chenopodium hybridum var. gigantospermum Bigseed goosefoot Chenopodium leptophyllum Slimleaf goosefoot Chenopodium standleyanum Standley goosefoot Salsola Kali var. tenuifolia Russian thistle Monolepis nutalliana Nuttall monolepis Alternanthera repens AMARANTHAECEAE Mat chaff flower Amaranthus graecizans Tumbleweed Amaranthus hybridus Slim amaranth Amaranthus palmeri Palmer amaranth Amaranthus retroflexus Redroot amaranth Amaranthus tamarascinus Waterhemp Brayulinea densa* Cottonflower Froelichia floridana Florida snakecotton Floelichia gracilis Slender snakecotton Gossypianthus lanuginosus Woolly cottonflower Gossypianthus tenuiflorus Lanceleaf cottonflower Oklahoma Native Plant Record 11 Volume 2, Number 1, December 2002 Buck, P. Mirabilis albida NYCTAGINACEAE Pale umbrella-wort Mirabilis inearis Narrowleaf umbrella-wort Mirabilis nyctaginea Heartleaf umbrella-wort Phytolacca americana PHYTOLACCACEAE Pokeberry Mollugo verticillata AIZOACEAE Green carpetweed Claytonia virginica PORTULACACEAE Virginia spring beauty Portulaca oleracea Purslane Portulaca parvla Shaggy purslane Portulaca umbraticola Wingpod purslane Talinum calycinum Rockpink Talinum parvifolorum Prairie fameflower Talinum rugospermum* Roughseed fameflower Arenaria patula CARYOPHYLLACEAE Pitcher’s sandwort Arenaria stricta var. texana Texas sandwort Cerastium brachypodum Shortstalk chickweed Sagina decumbens Trailing pearlwort Silene antirrhina Sleepy catchfly Paronychia jamesii ILLECEBRACEAE James nailwort Paronychia virginica Virginia nailwort Ceratophyllum demersum CERATOPHYLLACEAE Coontail RANUNCULACEAE Anemone caroliniana Carolina anemone Anemone decapetala Tenpetal anemone Clematis pitcheri Pitcher clematis Delphinium virescens Plains larkspur BERBERIDACEAE Mahonia aquifolium* Barberry Cocculus carolinus MENISPERMACEAE Carolina snailseed Menispermum canadense Moonseed Argemone polyanthemos PAPAVERACEAE Prickle poppy Corydalis aurea FUMARIACEAE var. occidentalis Goldon corydalis Cleome serrulata CAPPARIDACEAE Bee spider flower Cleomella angustifolia Narrowleaf rhombopod Polanisia dodecandra Roughseed clammywort CRUCIFERAE Arabis missouriensis Green rockcress Capsella bursa–pastoris Shepherdspurse Descurainia pinnata Pinnate tansymustard Dithyrea wislizenii Wislizen's spectacle pod Draba brachycarpa Shortpod draba Draba cuneifolia Wedgeleaf draba Draba reptans Carolina draba Erysimum capitatum Plains erysimum Lepidium oblongum Veiny pepperwort Lepidium virginicum Virginia pepperwort Lesquerella auriculata Earleaf bladderpod 12 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Buck, P. Lesquerella gordonii Gordon bladderpod Lesquerella gracilis Lax bladderpod Lesquerella ovalifolia var. alba Roundleaf bladderpod Nasturtium officinale Watercress Rorippa islandica Bog marshcress Rorippa sessilifolia Stalkless yellowcress Sibara virginica Virginia rockcress Sedum nuttallianum CRASSULACEAE Yellow stonecrop Sedum pulchellum Texas stonecrop Ribes odoratum SAXIFRAGACEAE Clove current Argimonia parviflora ROSACEAE Mayflower grovebur Amelanchier arborea Service berry Crataegus crus-galli Cockspur hawthorn Crataegus engelmannii Engelmann hawthorn Crataegus reverchonii Reverchon hawthorn Geum canadense White avens Geum virginiana* Bristly avens Potentilla arguta Cinquefoil Prunus americana American plum Prunus angustifolia Chickasaw plum Prunus armeniaca* Apricot Prunus persica Peach Prunus gracilis Oklahoma plum Prunus mexicana Mexican plum Prunus virginiana Chokecherry Rosa foliolosa* Leafy rose Rosa multiflora* Multiflora rose Rubus aboriginum Northern dewberry Rubus idaeus Red raspberry Rubus louisanus Louisiana blackberry Rubus occidentalis Blackcap raspberry Rubus trivialis Lowbush blackberry Spiraea arguta* Spirea Spirea cantoniensis* Spirea Spirea vanhouttei* Spirea Acacia angustissima LEGUMINOSAE Prairie acacia Amorpha canescens Leadplant Amorpha fruticosa Indigobush amorpha Amorpha microphylla* Smallleaf amorpha Amorpha nana* Fragrant false indigo Apios americana Groundnut Astragalus crassicarpus Groundplum milkvetch Astragalus lindheimeri Lindheimer milkvetch Astragalus plattensis Platte milkvetch Baptisia australis var. minor Blue wild indigo Baptisia leucophaea Plains wild indigo Baptisia sphaerocarpa Golden wild indigo Cassia fasciculate Showy partridgepea Cassia marilandica Wild senna Cercis canadensis Redbud Clitoria mariana Atlantic pigeonwings Oklahoma Native Plant Record 13 Volume 2, Number 1, December 2002 Buck, P. Colutea arborescens* Bladder senna Crotalaria sagittalis Arrow crotalaria Dalea aurea Siltop aurea Dalea enneandra Bigtop dalea Dalea purpurea Purple prairie-clover Desmanthus illinoensis Illinois bundleflower Desmodium ciliare Littleleaf tickclover Desmodium glutinosum Sticky tickclover Desmodium nudiflorum Barestem tickclover Desmodium paniculatum Panicled tickclover Desmodium sessilisovium Sessile tickclover Galactia volubilis Downy milkpea Gleditsia triacanthos Honeylocust Glycyrrhiza lepidota American licorice Gymnocladus dioica Kentucky coffee tree Hoffmanseggia densiflora Indian rushpea Indigofera miniata var. leptosepala Coast indigo Krameria secundiflora Trailing ratany Lespedeza capitata Roundhead lespedeza Lespedeza intermedia Intermediate lespedeza Lespedeza procumbens Trailing lespedeza Lespedeza virginica Slender lespedeza Lotus americanus Deervetch Melilotus alba White sweet clover Melilotus officianalis Yellow sweet clover Neptunea lutea Yellow neptunea Oxytropis lamberti Stemless loco Petalostemum candidum White prairie clover Petalostemum multiflorum Roundtop prairie clover Petalostemum purpureum Purple prairie clover Petalostemum tenuifolium Slimleaf prairie clover Prosopis juliflora Mesquite Psoralea cuspidata Tallbread scurfpea Psoralea esculenta Indian breadroot Psoralea lineariflolia Slimleaf scurfpea Psoralea tenuiflora Wild alfalfa Robinia pseudoacacia Black locust Shrankia uncinata Catclaw sensitive brier Strophostyles helvola Trailing wildbean Strophostyles leiosperma Slickseed wildbean Stylosanthes riparia Endbeak pencilflower Trifolium reflexum Buffalo clover Vicia ludoviciana Louisiana vetch Vicia sparsifolia Stiffleaf vetch GERANIACEAE Geranium carolinianum Carolina geranium Erodium cicutarium Storksbill OXALIDACEAE Oxalis stricta Oxalis Oxalis violacea Violet woodsorrel LINACEAE Linum hudsonioides Tufted flax Linum rigidum Stiffstem flax Linum sulcatum Grooved flax ZYGOPHYLLACEAE Tribulus terrestris 14 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Buck, P. Puncturevine RUTACEAE Ptelea trifoliate Wafer ash SIMAROUBACEAE Ailanthus altissima* Tree-of-Heaven POLYGALACEAE Polygala alba White milkwort Polygala verticillata Whorled milkwort EUPHORBIACEAE Acalypha gracilens Slender copperleaf Acalypha ostryaefolia Hophornbeam copperleaf Croton capitatus Woolly croton Croton glandulosus Tropic croton Croton lindheimerianus Threeseed croton Croton monanthogynus Oneseed croton Croton texensis Texas croton Euphorbia commutata Tinted euphorbia Euphorbia corollata Flowering spurge Euphorbia dentata Toothed euphorbia Euphorbia glyptosperma Ridgeseed euphorbia Euphorbia hexagona Six-angle euphorbia Euphorbia marginata Snow-on-the-mountain Euphorbia missourica Missouri euphorbia Euphorbia nutans Spotted euphorbia Euphorbia obtusata Roughpor euphorbia Euphorbia prostrata Prostrate euphorbia Euphorbia spathulata Warty euphorbia Euphorbia supina Milk purslane Phyllanthus caroliniensis Carolina leafflower Phyllanthus polygonoides Knotweed leafflower Stillingia sylvatica Queen's delight Tragia ramose Branching noseburn Callitriche heterophylla CALLITRICHACEAE Larger waterstar wort ANACARDIACEAE Rhus aromatica Skunkbush Rhus copallina Flameleaf sumac Rhus glabra Smooth sumac Rhus toxicodendron Poison ivy CELASTRACEAE Celastrus scandens American Bittersweet Euonymus atropurpureus Eastern wahoo Euonymus europaeus* European spindle-tree Acer saccharinum* ACERACEAE Silver maple Acer saccharum Sugar maple HIPPOSCASTANACEAE Aesculus glabra Ohio buckeye SAPINDACEAE Sapindus drummondii* Western soapberry RHAMNACEAE Ceanothus herbaceus var. pubescens Fuzzy ceanothus Rhamnus frangula* Alder buckthorn Ampelopsis cordata VITACEAE Heartleaf ampelopsis Cissus incisa Ivy treebine Parthenocissus quinquefolia Virginia creeper Oklahoma Native Plant Record 15 Volume 2, Number 1, December 2002 Buck, P. Vitis aestivalis Summer grape Vitis cinerea Sweet grape Vitis palmata Cat grape Vitis rotundifolia Muscadine grape Vitis rupestris Sand grape Vitis vulpine Riverbank grape MALVACEAE Callirhoe involucrata Low poppymallow Callirhoe leiocarpa Tall poppymallow Hibiscus syriacus* Rose of Sharon Sphaeralcea coccinea Scarlet mallow GUTTIFERAE Hypericum drummondii Nits and lice Hypericum mutilum Dwarf St. John's wort TAMARICACEAE Tamarix hispida* Kashgar tamarisk Tamarix odessana* Tamarisk Tamarix pentandra* Tamarisk Lechea tenuifolia CISTACEAE Narrowleaf pinwheel VIOLACEAE Hybanthus verticillatus Whorled nodviolet Viola missouriensis Missouri violet Viola papilionacea Butterfly violet Viola rafinesquii Johnny-jump-up LOASACEAE Mentzelia oligosperma Chickthief Mentzelia stricta Sandlily CACTACEAE Echinocereus baileyi Wichita pin cushion Echinocereus reichenbachii Lace echinocereus Opuntia compressa Prickly pear LYTHRACEAE Ammannia auriculata Earleaf ammannia Ammannia coccinea Purple ammannia Lagerstroemia indica* Crape myrtle Lythrum alatum Winged lythrum Gaura biennis var. pitcheri ONAGRACEAE Pitcher gaura Gaura coccinea Scarlet gaura Gaura parviflora Smallflower gaura Gaura sinuata Wavyleaf gaura Guara suffulta Roadside gaura Gaura tripetala var. triangulata Three petal gaura Jussiaea decurrens Wingleaf water primrose Jussiaea peploides var. glabrescens Smooth water primrose Ludwigia alternifolia bushy boxseed Ludwigia palustris Marsh boxseed Oenothera biennis Evening primrose Oenothera heterophylla Varileaf evening primrose Oenothera jamesii Trumpet evening primrose Oenothera laciniata var. laciniata Cutleaf evening primrose Oenothera laciniata var. grandiflora Largeflowered cutleaf evening primrose Oenothera linifolia Threadleaf sundrops 16 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Buck, P. Oenothera macrocarpa var.macrocarpa Ozark sundrops Oenothera macrocarpa var. incana Gray sundrops Oenothera missouriensis Missouri evening primrose Oenothera serrulata Halfshrub sundrops Oenothera speciosa Showy sundrops Oenothera triloba Stemless evening primrose Stenosiphon virgatus False gaura HALORAGACEAE Myriophyllum brasiliensis Parrot feather Myriophyllum exalbescens Parrot feather UMBELLIFERAE Ammoselinum butleri Butler sandparsley Ammoselinum popei Plains sandparlsey Chaerophyllum tainturieri var. dasycarpum Hairyfruit chervil Cicuta maculata Spotted water hemlock Daucus carota* Wild carrot Daucus pusillus Southwestern carrot Eryngium diffusum Bushy eryngo Eryngium leavenworthii Leavenworth eryngo Limnosciadum pinnatum Arkansas dogshade Lomatium daucifolium Carrotleaf lomatium Polytaenia nuttallii Prairie parsley Ptilimnium nuttallii Nuttall mockbishop Sanicula canadensis Canada sanic1e Spermolepis divaricata Forked scaleseed Spermolepis echinata Bristly scaleseed CORNACEAE Cornus drummondii Roughleaf dogwood Cornus florida* Flowering dogwood PRIMULACEAE Androsace occidentalis Western rockjasmine Dodecatheon meadia Shooting star Samolus parviflorus Thinleaf brookweed SAPOTACEAE Bumelia lanuginosa Woollybucket bumelia EBENACEAE Diospyros virginiana Persimmon OLEACEAE Forsythia suspensa var. fortunei* Golden bells Fraxinus americana White ash Fraxinus pennsylvanica var. pennsylvanica Red ash Fraxinus pennsylvanica var. subintegerrima Green ash Fraxinus velutina* Arizona ash Jasminum nudiflorum* Jasmine Ligustrum amurense* Privet Ligustrum vulgare* Privet Sabatia angularis GENTIANACEAE Squarestem rosegentium Sabatia campestris Prairie rosegentian Amsonia ciliata var. texana APOCYNACEAE Texas slimpod Amsonia tabernaemontana Willow slimpod Apocynum cannabinium var. glaberrimum Hemp dogbane Oklahoma Native Plant Record 17 Volume 2, Number 1, December 2002 Buck, P. Asclepias asperula ASCLEPIADACEAE var. decumbens Trailing milkweed Asclepias latifolia Broadleaf milkweed Asclepias pumila Plains milkweed Asclepias stenophylla Slimleaf milkweed Asclepias sullivantii Sullivant milkweed Asclepias tuberosa Butterfly milkweed Asclepias verticillata Whorled milkweed Asclepias viridiflora Green acerates Asclepias viridis Green antelope horn Convolvulus arvensis CONVOLVULACEAE Small bindweed Convolvulus incanus Gray bindweed Cuscuta coryli Hazel dodder Cuscuta gronovii Gronovius dodder Cuscuta indecora Showy dodder Cuscuta cuspidata Cusp dodder Evolvulus nuttallianus Silver evolvulus Ipomoea shumardiana Narrowleaf morning glory Gilia rubra POLEMONIACEAE Texasplume Phlox pilosa Downy phlox HYDROPHYLLACEAE Nama hispidum Rough nama Phacelia congesta Spike phacelia Phacelia hirsuta Hairy phacelia BORAGINACEAE Lithospermum arvense Corn Gromwell Lithospermum incisum Narrowleaf Gromwell Myosotis macrosperma Southern forget-me-not Myosotis verna Forget-me-not Onosmodium molle Marbleseed VERBENACEAE Phyla cuneifolia Wedgeleaf fogfruit Phyla incisa Sawtooth fogfruit Verbena bipinnatifida Dakota verbena Verbena bracteata Bigbract verbena Verbena canadensis Rose verbena Verbena pumila Pink verbena Verbena stricta Woolly verbena Verbena urticifolia White verbena Vitex agnus-castus* Chaste tree LABIATAE Hedeoma drummondii Drummond hedeoma Hedeoma hispida Rough hedeoma Lamium amplexicaule Henbit Lycopus americanus American bugleweed Monarda citriodora Lemon beebalm Monarda clinopodioides Basil beebalm Monarda fistulosa Shrubby beebalm Monarda punctata Spotted beebalm Nepeta cataria Catnip Salvia azurea var. grandiflora Azure sage Salvia reflexa Lanceleaf sage Scutellaria drummondii 18 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Buck, P. Drummond skullcap Scutellaria resinosa Resindot skullcap Scutellaria wrightii Wright skullcap Teucrium canadense American germander Teucrium laciniatum Cutleaf germander Trichostema brachiatum Fluxweed SOLANACEAE Physalis lobata Purple groundcherry Physalis virginiana var. sonorae Virginia groundcherry Physalis viscosa var. cinerascens Beach groundcherry Physalis viscosa var. mollis Field groundcherry Solanum carolinense Carolina horsenettle Solanum citrullifolium* Watermelon leaved solanum Solanum elaegnifolium Silverleaf nightshade Solanum rostratum Buffalobur Solanum torreyi Torrey horsenettle SCROPHULARIACEAE Bacopa rotundifolia Disc waterhyssop Castilleja citrina Citron paintbrush Castilleja sessiliflora Downy paintbrush Collinsia violacea Violet collinsia Gratiola virginiana Virginia hedgehyssop Leucospora multifida Narrowleaf conobea Linaria canadensis var. texana Oldfield toadflax Lindernia anagallidea C1asping False-pimpernel Lindernia dubia Yellowseed false-pimpernel Penstemon albidus White penstemon Penstemon cobea Cobaea penstemon Penstemon oklahomensis Oklahoma penstemon Penstemon tubaeflorus Tube panstemon Scrophularia lanceolata Narrowleaf figwort Veronica arvensis Speedwell Veronica peregrina Purslane speedwell Verbascum thapsus Flannel mullein Catalpa speciosa BIGNONIACEAE Catalpa Chilopsis linearis* Desert willow MARTYNIACEAE Proboscidea louisianica Unicorn plant LENTIBULARIACEAE Utricularia biflora Bladderwort Utricularia vulgaris Bladderwort ACANTHACEAE Justicia americana Justicia Ruellia humilis Low ruellia Ruellia pedunculata Stalked ruellia PHRYMACEAE Phryma leptostachya Lopseed PLANTAGINACEAE Plantago aristata Bottlebrush plantain Plantago elongata Elongate plantain Plantago purshii var. purshii Slender plantain Plantago purshii var. spinulosa Bristlebract plantain Plantago rhodosperma Redseed plantain Plantago virginica Paleseed plantain Plantago wrightiana Wright plantain Oklahoma Native Plant Record 19 Volume 2, Number 1, December 2002 Buck, P. RUBIACEAE Cephalanthus occidentalis Buttonbush Diodia teres var. setifera Rough buttonweed Galium aparine Catchweed bedstraw Galium pilosum Hairy bedstraw Galium texense Texas bedstraw Galium triflorum Fragrant bedstraw Galium virgatum Southwest bedstraw Hedyotis crassifolia Tiny bluet Hedyotis nigricans Narrowleaf bluet Hedyotis purpurea var. longifolia Purple bluet Lonicera japonica* CAPRIFOLIACEAE Honeysuckle Lonicera tatarica* Tatarian honeysuckle Symphoricarpos orbiculatus Coralberry Viburnum prunifolium var. ferrugineum Rusty blackhaw Valerianella radiata VALERIANACEAE var. radiata Beaked cornsalad Valerianella radiata var. parviflora Narrowcell cornsalad Cucurbita foetidissima CUCURBITACEAE Buffalo gourd Cyclanthera dissectra Cutleaf cyclanthera Ibervillea lindheimeri Lindheimer globeberry Melothria pendula Drooping me1onette CAMPANULACEAE Specularia biflora Small venus-looking glass Specularia leptocarpa Slimpod venus-looking glass Specularia perfoliata Clasping venus-looking glass LOBELIACEAE Lobelia appendiculata Earflower lobelia Lobelia cardinalis Cardinal flower Lobelia spicata Palespike lobelia COMPOSITAE Achillea lanulosa Western yarrow Achillea millefolium Yarrow Agoseris cuspidate Wavyleaf agoseris Ambrosia artemisiifolia Ragweed Ambrosia psilostachys Western ragweed Ambrosia trifida Giant ragweed Antennaria plantaginifolia Plantain-leaf pussy toes Aphanostephus pilosus Hairy dozedaisy Aphanostephus ramosissimus Plains dozedaisy Aphanostephus skirrhobasis Arkansas dozedaisy Artemisia carruthii Carruth sagewort Artemisia filifolia Figleaf sagewort Artemisia glauca False terragon sagewort Artemisia ludoviciana var. ludoviciana Louisiana sagewort Artemisia ludoviciana var. mexicana American sagewort Artemisia serrata* Sawtooth sagewort Aster communtatus* Cluster aster Aster ericoides Heath aster Aster fendleri Fendler aster Aster oblongifolius Aromatic aster 20 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Buck, P. Aster patens Skydrop aster Aster subulatus Awl shaped aster Baccharis salicina Willow groundsel-tree Berlandiera texana Texas greeneyes Bidens cernua Nodding beggarticks Bidens frondosa Devil’s beggarticks Centaurea americana Basketflower Chaetopappa asteroides Least daisy Chrysopsis pilosa Soft goldaster Chrysopsis villosa var. villosa Hairy goldaster Chrysopsis villosa var. canescens Hairy goldaster Chrysopsis villosa var. stenophylla Narrowleaf goldaster Cirsium ochrocentrum Yellowspine thistle Cirsium undulatum Wavyleaf thistle Conyza canadensis Horseweed fleabane Conyza ramoisissima Conyza Coreopsis grandiflora var. longipes Bigflower coreopsis Coreopsis lanceolata var. villosa Lanceleaf coreopsis Coreopsis tinctoria Plains coreopsis Dyssodia tagetioides Marigold dogweed Echinacea angustifolia Black sampson Echinacea pallida Pale purple coneflower Eclipta alba Yerba de tajo Englemannia pinnatifida Engelmann daisy Erigeron bellidiastrum Western fleabane Erigeron divergens Spreading fleabane Erigeron strigosus Prairie fleabane Eupatorium serotinum Late eupatorium Evax multicaulis Manystem evax Evax prolifera Bighead evax Gaillardia pulchella Rosering gaillardia Gaillardia suavis Sweet gaillardia Gnaphalium chilense Cottonbatting cudweed Gnaphalium purpureum Purple cudweed Gnaphalium wrightii Wright cudweed Grindelia squarrosa var. squarrosa Curlycup gumweed Grindelia squarrosa var. nuda Rayless gumweed Gutierrezia dracunculoides Broom weed Gutierrezia sarothrae* Broom snakeweed Haplopappus ciliatus Wax goldenweed Haplopappus spinulosis Cutleaf goldonweed Helenium amarum var. amarum Bitter sneezeweed Helenium amarum var. badium Basin sneezeweed Helenium autumnale Sneezeweed Helenium flexuosum Flexuous sneezeweed Helenium microcephalum Smallhead sneezeweed Helianthus annuus Annual sunflower Helianthus hirsutus Hairy sunflower Helianthus laetiflorus Stiff sunflower Helianthus maximilani Maximilian sunflower Helianthus petiolaris Prairie sunflower Hieracium longipilum Longbeard hawkseed Hymenopappus scabiosaeus Flattop woollywhite Oklahoma Native Plant Record 21 Volume 2, Number 1, December 2002 Buck, P. Hymenopappus tenuifolius Chalkhill woollywhite Hymenoxys linearifolia Fineleaf actinea Hymenoxys scaposa var. linearis Plains actinea Iva ciliata Seacoast sumpweed Krigia dandelion Tuber dwarf dandelion Krigia oppositifolia Weedy dwarf dandelion Kuhnia eupatorioides var. corymbulosa Plains kuhnia Lactuca canadensis Canada lettuce Lactuca pulchella* Chickory lettuce Lactuca scariola Prickly lettuce Liatris aspera Rough gayfeather Liatris punctata Dotted gayfeather Liatris scariosa Tall gayfeathcr Palafoxia texana Texas palafoxia Pluchea camphorata Camphor pluchea Pyrrhopappus carolinianus Carolina false-dandelion Pyrrhopappus scaposus Tuber false-dandelion Ratisida columnifera Upright prairie coneflower Rudbeckia hirta Blackeyed susan Santolina chamaecyparissus* Lavender cotton Senecio plattensis Prairie groundsel Senecio riddellii Riddell groundsel Silphium asteriscus Sand rosinweed Silphium laciniatum Compass plant Solidago bootii* Boot goldenrod Solidago gigantea var. leiophylla November goldenrod Solidago missouriensis var. fasciculate Missouri goldenrod Solidago mollis Ashy goldenrod Solidago petiolaris Downy goldenrod Solidago radula Rough goldenrod Taraxacum officinale Dandelion Thelesperma ambiguum Colorado greenthread Thelesperma filifolium Plains greenthread Thelesperma megapotamicum Rayless greenthread Townsendia excapa Stemless Townsendia Tragopogon major Yellow goat's beard Verbescina virginica White crownbeard Vernonia baldwinii Baldwin ironweed Vernonia missurica Missouri ironweed Xanthisma texanum Texas sleepydaisyXanthium strumarium Cocklebur Editor’s note: This paper, written in 1977, was originally distributed by the Wichita Mountains Wildlife Refuge as an informational handout but never published. Scientific names that were originally underlined have been italicized and species epithets derived from a person’s name are not capitalized as they were in the original work to abide with current taxonomic practice. The author emphasizess that there is a great need to update this twenty-five year old floristic list and, together with WMWR, encourages new research on this site. 22 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Floristic List for Comanche County, Oklahoma Bruce Hoagland Oklahoma Biological Survey/Department of Geography University of Oklahoma Norman, OK 73019 Floristic lists are vital tools for conservation planning, research, and wildflower appreciation. The diverse physical geography of Comanche County contributes to the high degree of plant diversity and habitats. The county is probably best known for the Wichita Mountains, which are composed of Cambrian gabbro and granites. To the north of the Wichita Mountains is the Slick Hill, an extensive outcropping of limestone. The surface geology of most of the county is composed of Permian sandstones. Land use in the county includes row crop agriculture and pastureland. INTRODUCTION The floristic list presented here was generated form the Atlas of the Flora of Oklahoma database. The database contains records for specimens deposited in Oklahoma herbaria. Voucher specimens for plants on the list can be found at one of the following herbaria: Oklahoma State University, Cameron University, University of Science and Arts of Oklahoma, Southeastern Oklahoma State University, and the Bebb Herbarium at the University of Oklahoma. Taxonomy and common names follow the United States Department of Agricultures Plants Database (www.plants.gov.us). There are 1,137 species and subspecific taxa, from the 116 families listed. There is a total of 3,560 specimens current recorded in the database. Of those records, Hoagland, B.W. https://doi.org/10.22488/okstate.17.100012 2,009 list the Wichita Mountains as the place of collection and 1,661 list the Wichita Mountains Wildlife Refuge specifically. Other locations in the Wichitas include Medicine Park and Meers. There are 836 records listed for Fort Sill, which includes the Wichita Mountains and a portion of Permian sandstone. The earliest collections in the database are dated 2 July 1903 and were collected by A. Van Vleet in the Wichita Mountains. These specimens are deposited at the Bebb Herbarium. Botanists with large numbers of collections in the database include F. L. Johnson and R. Thompson (790 records for specimens collected as part of florisitic inventory of Fort Sill), F. B. McMurry (498 records, all from the Wichita Mountains National Wildlife Refuge), B. Osborn (262 records), and U.T. Waterfall (121 records). Oklahoma Native Plant Record 23 Volume 2, Number 1, December 2002 Hoagland, B.W. Floristic List for Comanche County, Oklahoma Ferns and allies Aspleniaceae Asplenium platyneuron (L.) B.S.P. ebony spleenwort Asplenium resiliens Kunze blackstem spleenwort Asplenium trichomanes L. maidenhair spleenwort Dryopteridaceae Dryopteris marginalis (L.) gray marginal woodfern Woodsia obtusa (Spreng.) Torr. bluntlobe cliff fern Equisetaceae Equisetum hyemale L. scouringrush horsetail Equisetum hyemale L. var. affine (Engelm.) A.A. Eat. scouringrush horsetail Equisetum laevigatum A. Braun smooth horsetail Equisetum X ferrissii Clute (pro sp.) Marsileaceae Marsilea vestita Hook. & Grev. hairy waterclover Pilularia americana A. Braun American pillwort Ophioglossaceae Ophioglossum engelmannii Prantl limestone adderstongue Ophioglossum vulgatum L. southern adderstongue Pteridaceae Cheilanthes eatonii Baker Eaton's lipfern Cheilanthes lanosa (Michx.) D.C. Eat. hairy lipfern Cheilanthes lindheimeri Hook. fairyswords Cheilanthes tomentosa Link woolly lipfern Cheilanthes wootonii Maxon beaded lipfern Notholaena sinuata (Lag. ex Sw.) Kaulfuss Notholaena standleyi Maxon star cloak fern Pellaea atropurpurea (L.) Link purple cliffbrake Pellaea wrightiana Hook. Wright's cliffbrake Selaginellaceae Selaginella peruviana (Milde) Hieron. Peruvian spikemoss Selaginella rupestris (L.) Spring northern selaginella Gymnosperms Cupressaceae Juniperus virginiana L. eastern redcedar Thuja orientalis L. oriental arborvitae Angiosperms Monocotyledonae Agavaceae Manfreda virginica (L.) Salisb. ex Rose false aloe Yucca glauca Nutt. soapweed yucca Alismataceae Echinodorus berteroi (Spreng.) Fassett upright burrhead Sagittaria brevirostra Mackenzie & Bush Shortbeak arrowhead Sagittaria calycina Engelm. hooded arrowhead Sagittaria calycina Engelm. var. calycina hooded arrowhead 24 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Sagittaria latifolia Willd. broadleaf arrowhead Araceae Arisaema dracontium (L.) Schott green dragon Callitrichaceae Callitriche heterophylla Pursh twoheaded water-starwort Ceratophyllaceae Ceratophyllum demersum L. coon's tail Commelinaceae Commelina erecta L. whitemouth dayflower Commelina erecta L. var. angustifolia (Michx.) Fern. whitemouth dayflower Commelina erecta L. var. deamiana Fern. whitemouth dayflower Commelina virginica L. Virginia dayflower Tradescantia ernestiana E.S. Anderson & Woods. Ernest's spiderwort Tradescantia occidentalis (Britt.) Smyth prairie spiderwort Tradescantia occidentalis (Britt.) Smyth var. occidentalis prairie spiderwort Tradescantia ohiensis Raf. bluejacket Tradescantia tharpii E.S. Anderson & Woods. Tharp's spiderwort Hydrocharitaceae Egeria densa Planch. Brazilian waterweed Liliaceae Allium canadense L. meadow garlic Allium canadense L. var. fraseri Ownbey Fraser meadow garlic Allium canadense L. var. hyacinthoides (Bush) Ownbey & Aase hyacinth meadow garlic Allium canadense L. var. mobilense (Regel) Ownbey meadow garlic Allium drummondii Regel Drummond's onion Allium stellatum Nutt. ex Ker-Gawl. autumn onion Androstephium coeruleum (Scheele) Greene [orthographic variant] Camassia angusta (Engelm. & Gray) Blank. prairie camas Camassia scilloides (Raf.) Cory Atlantic camas Cooperia drummondii Herbert evening rainlily Erythronium albidum Nutt. white fawnlily Erythronium mesochoreum Knerr midland fawnlily Oklahoma Native Plant Record 25 Volume 2, Number 1, December 2002 Hoagland, B.W. Erythronium rostratum W. Wolf yellow troutlily Nolina texana S. Wats. Texas sacahuista Nothoscordum bivalve (L.) Britt. crowpoison Polygonatum biflorum (Walt.) Ell. smooth Solomon's seal Polygonatum biflorum (Walt.) Ell. var. commutatum (J.A. & J.H. Schultes) Morong smooth Solomon's seal Zigadenus nuttallii (Gray) S. Wats. Nuttall's deathcamas Najadaceae Najas guadalupensis (Spreng.) Magnus southern waternymph Orchidaceae Spiranthes cernua (L.) L.C. Rich. nodding ladies'-tresses Spiranthes magnicamporum Sheviak Great Plains ladies'-tresses Spiranthes vernalis Engelm. & Gray spring ladies'-tresses Poaceae Aegilops cylindrica Host jointed goatgrass Agropyron smithii Rydb. Agrostis hyemalis (Walt.) B.S.P. winter bentgrass Alopecurus carolinianus Walt. Carolina foxtail Andropogon gerardii Vitman big bluestem Aristida curtissii (Gray ex S. Wats. & Coult.) Nash Aristida dichotoma Michx. churchmouse threeawn Aristida dichotoma Michx. var. curtissii Gray ex S. Wats. & Coult. Curtis' threeawn Aristida longispica Poir. var. longispica slimspike threeawn Aristida oligantha Michx. prairie threeawn Aristida purpurascens Poir. arrowfeather threeawn Aristida purpurea Nutt. var. fendleriana (Steud.) Vasey Fendler's threeawn Aristida purpurea Nutt. var. longiseta (Steud.) Vasey Fendler threeawn Bothriochloa barbinodis (Lag.) Herter cane bluestem Bothriochloa ischaemum (L.) Keng yellow bluestem Bothriochloa saccharoides (Sw.) Rydb. silver bluestem Bouteloua gracilis (Willd. ex Kunth) Lag. 26 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. ex Griffiths blue grama Bouteloua hirsuta Lag. hairy grama Bouteloua pectinata Featherly Bouteloua rigidiseta (Steud.) A.S. Hitchc. Texas grama Bromus catharticus Vahl rescuegrass Bromus commutatus Schrad. meadow brome Bromus japonicus Thunb. ex Murr. Japanese brome Bromus pubescens Muhl. ex Willd. hairy woodland brome Bromus tectorum L. cheatgrass Buchloe dactyloides (Nutt.) Engelm. buffalograss Cenchrus incertus M.A. Curtis Cenchrus longispinus (Hack.) Fern. mat sandbur Chasmanthium latifolium (Michx.) Yates Indian woodoats Chloris verticillata Nutt. tumble windmill grass Chloris virgata Sw. feather fingergrass Cinna arundinacea L. sweet woodreed Cynodon dactylon (L.) Pers. Bermudagrass Dactylis glomerata L. orchardgrass Dichanthelium acuminatum (Sw.) Gould & C.A. Clark tapered rosette grass Dichanthelium acuminatum (Sw.) Gould & C.A. Clark var. fasciculatum (Torr.) Freckmann western panicgrass Dichanthelium acuminatum (Sw.) Gould & C.A. Clark var. lindheimeri (Nash) Gould & C.A. Clark Lindheimer panicgrass Dichanthelium linearifolium (Scribn. ex Nash) Gould slimleaf panicgrass Dichanthelium malacophyllum (Nash) Gould softleaf rosette grass Dichanthelium oligosanthes (J.A. Schultes) Gould Heller's rosette grass Dichanthelium oligosanthes (J.A. Schultes) Gould var. scribnerianum (Nash) Gould Scribner's rosette grass Digitaria californica (Benth.) Henr. Arizona cottontop Digitaria cognata (J.A. Schultes) Pilger Carolina crabgrass Digitaria ischaemum (Schreb.) Schreb. ex Muhl. smooth crabgrass Digitaria sanguinalis (L.) Scop. hairy crabgrass Echinochloa colona (L.) Link jungle rice Echinochloa crus-galli (L.) Beauv. barnyardgrass Echinochloa crus-galli (L.) Beauv. barnyardgrass Eleusine indica (L.) Gaertn. Indian goosegrass Eragrostis barrelieri Daveau Mediterranean lovegrass Oklahoma Native Plant Record 27 Volume 2, Number 1, December 2002 Hoagland, B.W. Eragrostis capillaris (L.) Nees lace grass Eragrostis cilianensis (All.) Vign. ex Janchen stinkgrass Eragrostis curtipedicellata Buckl. gummy lovegrass Eragrostis curvula (Schrad.) Nees weeping lovegrass Eragrostis hypnoides (Lam.) B.S.P. teal lovegrass Eragrostis intermedia A.S. Hitchc. plains lovegrass Eragrostis pectinacea (Michx.) Nees ex Steud. tufted lovegrass Eragrostis pectinacea (Michx.) Nees ex Steud. var. pectinacea tufted lovegrass Eragrostis pilosa (L.) Beauv. Indian lovegrass Eragrostis reptans (Michx.) Nees Eragrostis secundiflora J. Presl red lovegrass Eragrostis sessilispica Buckl. tumble lovegrass Eragrostis spectabilis (Pursh) Steud. purple lovegrass Eragrostis trichodes (Nutt.) Wood sand lovegrass Eriochloa contracta A.S. Hitchc. prairie cupgrass Eriochloa sericea (Scheele) Munro ex Vasey Texas cupgrass Erioneuron pilosum (Buckl.) Nash hairy woollygrass Festuca octoflora Walt. Festuca subverticillata (Pers.) Alexeev nodding fescue Leersia oryzoides (L.) Sw. rice cutgrass Leersia virginica Willd. whitegrass Leptochloa fascicularis (Lam.) Gray Leptochloa mucronata (Michx.) Kunth Melica nitens (Scribn.) Nutt. ex Piper threeflower melicgrass Muhlenbergia racemosa (Michx.) B.S.P. marsh muhly Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. rock muhly Panicum anceps Michx beaked panicgrass Panicum capillare L. witchgrass Panicum dichotomiflorum Michx. fall panicgrass Panicum hallii Vasey Hall's panicgrass Panicum hillmanii Chase Hillman's panicgrass Panicum obtusum Kunth vine mesquite Panicum virgatum L. switchgrass Pascopyrum smithii (Rydb.) A. Löve western wheatgrass Paspalidium geminatum (Forsk.) Stapf Egyptian panicgrass Paspalum distichum L. knotgrass Paspalum pubiflorum Rupr. ex Fourn. hairyseed paspalum Paspalum pubiflorum Rupr. ex Fourn. var. glabrum Vasey ex Scribn. Paspalum setaceum Michx. thin paspalum Paspalum setaceum Michx. var. stramineum (Nash) D. Banks Phalaris caroliniana Walt. Carolina canarygrass Poa annua L. annual bluegrass 28 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Poa arachnifera Torr. Texas bluegrass Poa compressa L. Canada bluegrass Schedonnardus paniculatus (Nutt.) Trel. tumblegrass Schizachyrium scoparium (Michx.) Nash little bluestem Setaria glauca (L.) Beauv. Sphenopholis obtusata (Michx.) Scribn. prairie wedgescale Sporobolus airoides (Torr.) Torr. alkali sacaton Sporobolus asper (Michx.) Kunth Sporobolus asper (Michx.) Kunth var. drummondii (Trin.) Vasey Sporobolus clandestinus (Biehler) A.S. Hitchc. rough dropseed Sporobolus cryptandrus (Torr.) Gray sand dropseed Sporobolus neglectus Nash puffsheath dropseed Sporobolus ozarkanus Fern. Sporobolus pyramidatus (Lam.) A.S. Hitchc. Sporobolus vaginiflorus (Torr. ex Gray) Wood poverty dropseed Tripsacum dactyloides (L.) L. eastern gamagrass Vulpia octoflora (Walt.) Rydb. sixweeks fescue Potamogetonaceae Potamogeton amplifolius Tuckerman largeleaf pondweed Potamogeton crispus L. curly pondweed Potamogeton diversifolius Raf. waterthread pondweed Potamogeton illinoensis Morong Illinois pondweed Potamogeton nodosus Poir. longleaf pondweed Potamogeton pusillus L. small pondweed Potamogeton pusillus L. ssp. pusillus small pondweed Potamogeton pusillus L. var. tenuissimus Mert. & Koch Typhaceae Typha angustifolia L. narrowleaf cattail Typha domingensis Pers. southern cattail Typha latifolia L. broadleaf cattail Zannichelliaceae Zannichellia palustris L. horned pondweed Dicotyledonae Acanthaceae Dicliptera brachiata (Pursh) Spreng. branched foldwing Justicia americana (L.) Vahl American water-willow Ruellia ciliosa Pursh. Ruellia humilis Nutt. fringeleaf wild petunia Ruellia humilis Nutt. var. expansa Fern. Ruellia humilis Nutt. var. longiflora (Gray) Fern. Aceraceae Acer negundo L. boxelder Acer negundo L. var. interius (Britt.) Sarg. boxelder Acer negundo L. var. texanum Pax boxelder Acer saccharinum L. silver maple Oklahoma Native Plant Record 29 Volume 2, Number 1, December 2002 Hoagland, B.W. Acer saccharum Marsh. sugar maple Amaranthaceae Amaranthus albus L. prostrate pigweed Amaranthus arenicola I.M. Johnston sandhill amaranth Amaranthus graecizans L. Amaranthus hybridus L. slim amaranth Amaranthus retroflexus L. redroot amaranth Amaranthus rudis Sauer tall amaranth Froelichia floridana (Nutt.) Moq. plains snakecotton Froelichia gracilis (Hook.) Moq. slender snakecotton Gossypianthus lanuginosus (Poir.) Moq. woolly cottonflower Gossypianthus lanuginosus (Poir.) Moq. var. lanuginosus woolly cottonflower Iresine rhizomatosa Standl. Juda's bush Anacardiaceae Rhus aromatica Ait. fragrant sumac Rhus aromatica Ait. var. serotina (Greene) Rehd. fragrant sumac Rhus glabra L. smooth sumac Rhus trilobata Nutt. var. trilobata skunkbush sumac Toxicodendron radicans (L.) Kuntze eastern poison ivy Apiaceae Ammoselinum popei Torr. & Gray plains sandparsley Chaerophyllum procumbens (L.) Crantz spreading chervil Chaerophyllum tainturieri Hook. hairyfruit chervil Chaerophyllum tainturieri Hook. var. dasycarpum Hook. ex S. Wats. Chaerophyllum tainturieri Hook. var. tainturieri Chaerophyllum texanum Coult. & Rose Cicuta maculata L. spotted water hemlock Conium maculatum L. poison hemlock Daucus carota L. Queen Anne's lace Daucus pusillus Michx. American wild carrot Eryngium hookerii Walp. Hooker's eryngo Eryngium leavenworthii Torr. & Gray Leavenworth's eryngo Limnosciadium pinnatum (DC.) Mathias & Constance tansy dogshade Lomatium daucifolium (Torr. & Gray) Coult. & Rose Lomatium foeniculaceum (Nutt.) Coult. & Rose ssp. daucifolium (Torr. & Gray) Theobald desert biscuitroot Polytaenia nuttallii DC. Nutall’s prairie parsley Sanicula canadensis L. Canadian blacksnakeroot Spermolepis divaricata (Walt.) Raf. ex Ser. roughfruit scaleseed Spermolepis echinata (Nutt. ex DC.) Heller bristly scaleseed Spermolepis inermis (Nutt. ex DC.) Mathias & Constance Red River scaleseed Spermolepis patens (Nutt. ex DC.) B.L. Robins. 30 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Torilis nodosa (L.) Gaertn. knotted hedgeparsley Apocynaceae Amsonia ciliata Walt. fringed bluestar Amsonia ciliata Walt. var. texana (Gray) Coult. Texas bluestar Amsonia illustris Woods. Ozark bluestar Apocynum cannabinum L. Indianhemp Apocynum cannabinum L. var. glaberrimum A. DC. Asclepiadaceae Asclepias asperula (Dcne.) Woods. spider milkweed Asclepias asperula (Dcne.) Woods. ssp. capricornu (Woods.) Woods. antelopehorns Asclepias asperula (Dcne.) Woods. var. decumbens (Nutt.) Shinners Asclepias engelmanniana Woods. Engelmann's milkweed Asclepias pumila (Gray) Vail plains milkweed Asclepias stenophylla Gray slimleaf milkweed Asclepias sullivantii Engelm. ex Gray prairie milkweed Asclepias tuberosa L. butterfly milkweed Asclepias tuberosa L. ssp. interior Woods. butterfly milkweed Asclepias verticillata L. whorled milkweed Asclepias viridiflora Raf. green comet milkweed Ascleias viridis Walt. green antelopehorn Asclepiodora decumbens (Nutt.) Gray Matelea biflora (Raf.) Woods. star milkvine Asteraceae Achillea millefolium L. common yarrow Achillea millefolium L. var. occidentalis DC. western yarrow Ambrosia artemisiifolia L. annual ragweed Ambrosia artemisiifolia L. var. elatior (L.) Descourtils annual ragweed Ambrosia psilostachya DC. Cuman ragweed Ambrosia trifida L. great ragweed Ambrosia trifida L. var. texana Scheele Texan great ragweed Amphiachyris dracunculoides (DC.) Nutt. prairie broomweed Antennaria parlinii Fern. ssp. fallax (Greene) Bayer & Stebbins Parlin's pussytoes Antennaria plantaginifolia (L.) Richards. woman's tobacco Aphanostephus pilosus Buckl. hairy dozedaisy Aphanostephus skirrhobasis (DC.) Trel. Arkansas dozedaisy Artemisia carruthii Wood ex Carruth. Carruth's sagewort Artemisia dracunculus L. tarragon Artemisia filifolia Torr. sand sagebrush Artemisia ludoviciana Nutt. white sagebrush Oklahoma Native Plant Record 31 Volume 2, Number 1, December 2002 Hoagland, B.W. Artemisia ludoviciana Nutt. ssp. ludoviciana white sagebrush Artemisia ludoviciana Nutt. ssp. mexicana (Willd. ex Spreng.) Keck white sagebrush Artemisia neomexicana Greene ex Rydb. Aster ericoides L. Aster oblongifolius Nutt. Aster oblongifolius Nutt. var. rigidulus Gray Aster ontarionis Wieg. Aster patens Ait. Aster patens Ait. var. patentissimus (Lindl. ex DC.) Torr. & Gray Aster subulatus Michx. Aster subulatus Michx. var. ligulatus Shinners Astranthium integrifolium (Michx.) Nutt. ssp. ciliatum (Raf.) DeJong entireleaf western daisy Baccharis salicina Torr. & Gray Great Plains false willow Bidens bipinnata L. Spanish needles Bidens frondosa L. devil's beggartick Brickellia eupatorioides (L.) Shinners var. chlorolepis (Woot. & Standl.) B.L. Turner false boneset Brickellia eupatorioides (L.) Shinners var. corymbulosa (Torr. & Gray) Shinners false boneset Centaurea americana Nutt. American star-thistle Chaetopappa asteroides Nutt. ex DC. Arkansas leastdaisy Chrysopsis berlandieri Greene Chrysopsis pilosa Nutt. soft goldenaster Chrysopsis stenophylla (Gray) Greene Chrysopsis villosa (Pursh) Nutt. ex DC. Chrysopsis villosa (Pursh) Nutt. ex DC. var. stenophylla (Gray) Gray Cirsium altissimum (L.) Hill tall thistle Cirsium ochrocentrum Gray yellowspine thistle Cirsium texanum Buckl. Texas thistle Cirsium undulatum (Nutt.) Spreng. wavyleaf thistle Cirsium undulatum (Nutt.) Spreng. var. megacephalum (Gray) Fern. Conyza canadensis (L.) Cronq. Canadian horseweed Conyza canadensis (L.) Cronq. var. canadensis Canadian horseweed Conyza ramosissima Cronq. dwarf horseweed Coreopsis grandiflora Hogg ex Sweet largeflower tickseed Coreopsis grandiflora Hogg ex Sweet var. harveyana (Gray) Sherff largeflower tickseed Coreopsis grandiflora Hogg ex Sweet var. longipes (Hook.) Torr. & Gray largeflower tickseed Coreopsis tinctoria Nutt. golden tickseed Coreopsis tinctoria Nutt. var. tinctoria golden tickseed Dracopis amplexicaulis (Vahl) Cass. clasping coneflower Dysodiopsis tagetoides (Torr. & Gray) Rydb. false dogfennel 32 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Dyssodia tagetoides Torr. & Gray Echinacea angustifolia DC. blacksamson echinacea Echinacea purpurea (L.) Moench eastern purple coneflower Eclipta prostrata (L.) L. false daisy Elephantopus carolinianus Raeusch. Carolina elephantsfoot Engelmannia pinnatifida Gray ex Nutt. Erigeron canadensis L. Erigeron philadelphicus L. Philadelphia fleabane Erigeron ramosus (Walt.) B.S.P. Erigeron strigosus Muhl. ex Willd. prairie fleabane Erigeron strigosus Muhl. ex Willd. var. strigosus prairie fleabane Erigeron tenuis Torr. & Gray slenderleaf fleabane Evax prolifera Nutt. ex DC. bighead pygmycudweed Filago prolifera (Nutt. ex DC.) Britt., non Pomel Gaillardia lanceolata Michx. Gaillardia lutea Greene Gaillardia pulchella Foug. firewheel Gaillardia suavis (Gray & Engelm.) Britt. & Rusby perfumeballs Gnaphalium obtusifolium L. Gnaphalium wrightii Gray Grindelia nuda Wood var. nuda curlytop gumweed Grindelia squarrosa (Pursh) Dunal curlycup gumweed Grindelia squarrosa (Pursh) Dunal var. nuda (Wood) Gray Gutierrezia dracunculoides (DC.) Blake Gutierrezia sarothrae (Pursh) Britt. & Rusby broom snakeweed Gutierrezia texana (DC.) Torr. & Gray var. texana Texas snakeweed Haplopappus ciliatus (Nutt.) DC. Haplopappus divaricatus (Nutt.) Gray Haplopappus spinulosus (Pursh) DC. Haplopappus spinulosus (Pursh) DC. ssp. australis (Greene) Hall Haplopappus spinulosus (Pursh) DC. ssp. cotulus (Small) Hall Haplopappus spinulosus (Pursh) DC. ssp. glaberrimus (Rydb.) Hall Haplopappus spinulosus (Pursh) DC. var. turbinellus (Rydb.) Blake Helenium amarum (Raf.) H. Rock yellowdicks Helenium amarum (Raf.) H. Rock var. badium (Gray ex S. Wats.) Waterfall yellowdicks Helenium badium (Gray ex S. Wats.) Greene Helenium microcephalum DC. smallhead sneezeweed Helianthus annuus L. common sunflower Helianthus hirsutus Raf. hairy sunflower Helianthus maximiliani Schrad. Maximilian sunflower Helianthus orgyalis DC. Helianthus petiolaris Nutt. prairie sunflower Helianthus salicifolius A. Dietr. willowleaf sunflower Heterotheca canescens (DC.) Shinners hoary false goldenaster Heterotheca latifolia Buckl. Oklahoma Native Plant Record 33 Volume 2, Number 1, December 2002 Hoagland, B.W. Heterotheca latifolia Buckl. var. macgregoris Wagenkn. Heterotheca stenophylla (Gray) Shinners stiffleaf false goldenaster Heterotheca villosa (Pursh) Shinners hairy false goldenaster Heterotheca villosa var. villosa (Pursh) Shinners hairy goldenaster Hieracium longipilum Torr. hairy hawkweed Hymenopappus corymbosus Torr. & Gray Hymenopappus scabiosaeus L'Hér. Carolina woollywhite Hymenopappus scabiosaeus L'Hér. var. corymbosus (Torr. & Gray) B.L. Turner Carolina woollywhite Hymenopappus tenuifolius Pursh Chalk Hill hymenopappus Hymenoxys linearifolia Hook. Iva annua L. annual marshelder Iva annua L. var. annua annual marshelder Krigia biflora (Walt.) Blake twoflower dwarfdandelion Krigia caespitosa (Raf.) Chambers weedy dwarfdandelion Krigia cespitosa (Raf.) Chambers [orthographic variant] Krigia dandelion (L.) Nutt. potato dwarfdandelion Krigia occidentalis Nutt. western dwarfdandelion Kuhnia eupatorioides L. false boneset Kuhnia eupatorioides L. var. corymbulosa Torr. & Gray Kuhnia glutosina Ell. false boneset Lactuca canadensis L. Canada lettuce Lactuca canadensis L. var. latifolia Kuntze Lactuca ludoviciana (Nutt.) Riddell biannual lettuce Lactuca scariola L. var. integrata Gren. & Godr. Lactuca serriola L. prickly lettuce Liatris aspera Michx. tall blazing star Liatris punctata Hook. dotted blazing star Liatris punctata Hook. var. nebraskana Gaiser Nebraska blazing star Liatris scabra (Greene) K. Schum. Liatris squarrosa (L.) Michx. scaly blazing star Lindheimera texana Gray & Engelm. Texas yellowstar Machaeranthera annua (Rydb.) Shinners Machaeranthera spinulosa (Greene) Cory hoary tansyaster Packera plattensis (Nutt.) W.A. Weber & A. Löve prairie groundsel Packera tampicana (DC.) C. Jeffrey Great Plains ragwort Palafoxia rosea (Bush) Cory rosy palafox Palafoxia rosea (Bush) Cory var. rosea rosy palafox Pluchea camphorata (L.) DC. camphor pluchea Pluchea odorata (L.) Cass. var. odorata sweetscent Prenanthes altissima L. tall rattlesnakeroot Prionopsis ciliata (Nutt.) Nutt. Pyrrhopappus grandiflorus (Nutt.) Nutt. tuberous desert-chicory 34 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Rudbeckia hirta L. blackeyed Susan Rudbeckia hirta L. var. pulcherrima Farw. blackeyed Susan Rudbeckia hirta L. var. serotina (Nutt.) Core Rudbeckia serotina Nutt. Senecio plattensis Nutt. Silphium asperrimum Hook. Silphium asteriscus L. starry rosinweed Silphium laciniatum L. compassplant Solidago arguta Ait. Atlantic goldenrod Solidago canadensis L. var. gilvocanescens Rydb. shorthair goldenrod Solidago delicatula Small Solidago gigantea Ait. giant goldenrod Solidago missouriensis Nutt. Missouri goldenrod Solidago missouriensis Nutt. var. fasciculata Holz. Missouri goldenrod Solidago mollis Bartl. velvety goldenrod Solidago nemoralis Ait. gray goldenrod Solidago nemoralis Ait. ssp. longipetiolata (Mackenzie & Bush) G.W. Douglas Solidago petiolaris Ait. downy ragged goldenrod Solidago petiolaris Ait. var. petiolaris downy ragged goldenrod Solidago rigida L. Solidago speciosa Nutt. var. rigidiuscula Torr. & Gray showy goldenrod Solidago ulmifolia Muhl. ex Willd. var. microphylla Gray elmleaf goldenrod Sonchus asper (L.) Hill spiny sowthistle Taraxacum officinale G.H. Weber ex Wiggers common dandelion Tetraneuris acaulis (Pursh) Greene stemless four-nerve daisy Tetraneuris linearifolia (Hook.) Greene fineleaf fournerved daisy Thelesperma filifolium (Hook.) Gray stiff greenthread Thelesperma filifolium (Hook.) Gray var. filifolium stiff greenthread Thelesperma intermedium Rydb. Thelesperma trifidum (Poir.) Britt. Townsendia exscapa (Richards.) Porter stemless Townsend daisy Tragopogon dubius Scop. yellow salsify Verbesina alternifolia (L.) Britt. ex Kearney wingstem Verbesina encelioides (Cav.) Benth. & Hook. f. ex Gray ssp. encelioides golden crownbeard Verbesina virginica L. white crownbeard Vernonia baldwinii Torr. Baldwin's ironweed Vernonia baldwinii Torr. ssp. interior (Small) Faust interior ironweed Vernonia interior Small Vernonia missurica Raf. Missouri ironweed Xanthisma texanum DC. Texas sleepydaisy Xanthisma texanum DC. ssp. drummondii Oklahoma Native Plant Record 35 Volume 2, Number 1, December 2002 Hoagland, B.W. (Torr. & Gray) Semple Drummond's sleepydaisy Xanthisma texanum DC. var. drummondii (Torr. & Gray) Gray Xanthium strumarium L. rough cockleburr Xanthium strumarium L. var. canadense (P. Mill.) Torr. & Gray Canada Cocklebur Bignoniaceae Campsis radicans (L.) Seem. ex Bureau trumpet creeper Catalpa bignonioides Walt. southern catalpa Catalpa speciosa (Warder) Warder ex Engelm. northern catalpa Boraginaceae Heliotropium tenellum (Nutt.) Torr. pasture heliotrope Lithospermum incisum Lehm. narrowleaf stoneseed Myosotis verna Nutt. spring forget-me-not Onosmodium molle Michx. ssp. occidentale (Mackenzie) Cochrane western marbleseed Brassicaceae Arabis fendleri (S. Wats.) Greene Fendler's rockcress Arabis laevigata (Muhl. ex Willd.) Poir. smooth rockcress Arabis missouriensis Greene green rockcress Camelina microcarpa DC. littlepod false flax Capsella bursa-pastoris (L.) Medik. shepherd's purse Chorispora tenella (Pallas) DC. crossflower Descurainia pinnata (Walt.) Britt. ssp. brachycarpa (Richards.) Detling western tansymustard Draba brachycarpa Nutt. ex Torr. & Gray shortpod draba Draba cuneifolia Nutt. ex Torr. & Gray wedgeleaf draba Draba cuneifolia Nutt. ex Torr. & Gray var. cuneifolia wedgeleaf draba Draba cuneifolia Nutt. ex Torr. & Gray var. helleri (Small) O.E. Schulz Draba platycarpa Torr. & Gray broadpod draba Draba reptans (Lam.) Fern. Carolina draba Erysimum asperum (Nutt.) DC. Erysimum asperum (Nutt.) DC. var. arkansanum (Nutt.) Gray Erysimum capitatum (Dougl. ex Hook.) Greene sanddune wallflower Erysimum capitatum (Dougl. ex Hook.) Greene var. capitatum sanddune wallflower Erysimum repandum L. spreading wallflower Lepidium austrinum Small southern pepperwort Lepidium densiflorum Schrad. common pepperweed Lepidium densiflorum Schrad. var. densiflorum common pepperweed Lepidium oblongum Small veiny pepperweed Lepidium virginicum L. Virginia pepperweed 36 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Lepidium virginicum L. var. medium (Greene) C.L. Hitchc. medium pepperweed Lesquerella auriculata (Engelm. & Gray) S. Wats. earleaf bladderpod Lesquerella gordonii (Gray) S. Wats. Gordon's bladderpod Lesquerella gracilis (Hook.) S. Wats. spreading bladderpod Lesquerella gracilis (Hook.) S. Wats. ssp. nuttallii (Torr. & Gray) Rollins & Shaw Nuttall's bladderpod Lesquerella ovalifolia Rydb. ex Britt. roundleaf bladderpod Lesquerella ovalifolia Rydb. ex Britt. ssp. alba (Goodman) Rollins & Shaw roundleaf bladderpod Nasturtium officinale Ait. f. Rorippa nasturtium-aquaticum (L.) Hayek watercress Rorippa palustris (L.) Bess. bog yellowcress Rorippa sessiliflora (Nutt.) A.S.Hitchc. stalkless yellowcress Sibara virginica (L.) RollinsVirginia winged rockcress Cactaceae Echinocereus baileyi Rose Echinocereus reichenbachii (Terscheck ex Walp.) Haage f. lace hedgehog cactus Echinocereus reichenbachii (Terscheck ex Walp.) Haage f. var. albispinus (Lahman) L. Benson Echinocereus reichenbachii (Terscheck ex Walp.) Haage f. var. baileyi (Rose) N.P. Taylor Bailey's hedgehog cactus Opuntia engelmannii Salm-Dyck cactus apple Opuntia humifusa (Raf.) Raf. devil's-tongue Campanulaceae Lobelia appendiculata A. DC. pale lobelia Lobelia cardinalis L. cardinalflower Lobelia cardinalis L. ssp. graminea (Lam.) McVaugh Triodanis leptocarpa (Nutt.) Nieuwl. slimpod Venus' looking-glass Triodanis perfoliata (L.) Nieuwl. var. biflora (Ruiz & Pavón) Bradley clasping Venus' looking-glass Capparaceae Cleome serrulata Pursh Rocky Mountain beeplant Cleomella angustifolia Torr. narrowleaf rhombopod Polanisia dodecandra (L.) DC. redwhisker clammyweed Polanisia dodecandra (L.) DC. ssp.trachysperma (Torr. & Gray) Iltis sandyseed clammyweed Caprifoliaceae Lonicera albiflora Torr. & Gray western white honeysuckle Lonicera japonica Thunb. Japanese honeysuckle Sambucus nigra L. ssp. Canadensis (L.) R. Bolli common elderberry Symphoricarpos orbiculatus Moench coralberry Viburnum rufidulum Raf. rusty blackhaw Oklahoma Native Plant Record 37 Volume 2, Number 1, December 2002 Hoagland, B.W. Caryophyllaceae Cerastium brachypodum (Engelm. ex Gray) B.L. Robins. shortstalk chickweed Cerastium glomeratum Thuill. sticky chickweed Minuartia michauxii (Fenzl) Farw. Michaux's stitchwort Minuartia michauxii (Fenzl) Farw. var. texana (B.L. Robins.) Mattf. Texas stitchwort Minuartia patula (Michx.) Mattf. pitcher's stitchwort Minuartia patula var. patula (Michx.) Mattf. Paronychia jamesii Torr. & Gray James' nailwort Paronychia virginica Spreng. yellow nailwort Sagina decumbens (Ell.) Torr. & Gray trailing pearlwort Silene antirrhina L. sleepy silene Celastraceae Celastrus scandens L. American bittersweet Euonymus atropurpurea Jacq. eastern wahoo Chenopodiaceae Chenopodium album L. lambsquarters Chenopodium ambrosioides L. Mexican tea Chenopodium leptophyllum (Moq.) Nutt. ex S. Wats. narrowleaf goosefoot Chenopodium pratericola Rydb. desert goosefoot Chenopodium simplex (Torr.) Raf. mapleleaf goosefoot Chenopodium standleyanum Aellen Standley's goosefoot Kochia scoparia (L.) Schrad. Mexican-fireweed Salsola kali L. prickly Russian thistle Salsola kali L. ssp. tragus (L.) Celak. Cistaceae Lechea tenuifolia Michx. narrowleaf pinweed Clusiaceae Hypericum drummondii (Grev. & Hook.) Torr. & Gray nits and lice Hypericum mutilum L. dwarf St. Johnswort Convolvulaceae Convolvulus arvensis L. field bindweed Convolvulus equitans Benth. Texas bindweed Convolvulus incanus auct. non Vahl Evolvulus nuttallianus J.A. Schultes shaggy dwarf morning-glory Evolvulus pilosus Nutt. Ipomoea pandurata (L.) G.F.W. Mey. man of the earth Ipomoea shumardiana (Torr.) Shinners narrowleaf morning-glory Cornaceae Cornus drummondii C.A. Mey. roughleaf dogwood Crassulaceae Sedum nuttallianum Raf. yellow stonecrop Cucurbitaceae Cucurbita foetidissima Kunth Missouri gourd Cyclanthera dissecta (Torr. & Gray) Arn. cutleaf cyclanthera 38 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Ibervillea lindheimeri (Gray) Greene Lindheimer's globeberry Melothria pendula L. Guadeloupe cucumber Cuscutaceae Cuscuta glabrior (Engelm.) Yuncker var. pubescens (Engelm.) Yuncker Cuscuta gronovii Willd. ex J.A. Schultes scaldweed Cuscuta indecora Choisy bigseed alfalfa dodder Cuscuta indecora Choisy var. indecora bigseed alfalfa dodder Cuscuta pentagona Engelm. fiveangled dodder Cuscuta pentagona Engelm. var. pentagona fiveangled dodder Cyperaceae Bolboschoenus maritimus (L.) Palla Bulbostylis capillaris (L.) Kunth ex C.B. Clarke densetuft hairsedge Bulbostylis capillaris (L.) Kunth ex C.B. Clarke ssp. capillaries densetuft hairsedge Carex albicans Willd. ex Spreng. var. albicans whitetinge sedge Carex austrina Mackenzie southern sedge Carex blanda Dewey eastern woodland sedge Carex brevior (Dewey) Mackenzie shortbeak sedge Carex emoryi Dewey Emory's sedge Carex fissa Mackenzie hammock sedge Carex frankii Kunth Frank's sedge Carex granularis Muhl. ex Willd. limestone meadow sedge Carex gravida Bailey heavy sedge Carex gravida Bailey var. lunelliana (Mackenzie) F.J. Herm. heavy sedge Carex grisea Wahlenb. Carex joorii Bailey cypress swamp sedge Carex muehlenbergii Schkuhr ex Willd. var. enervis Boott Muhlenberg's sedge Carex muehlenbergii Schkuhr ex Willd. var. muehlenbergii Muhlenberg's sedge Carex tetrastachya Scheele Britton's sedge Carex umbellata Schkukr ex Willd. parasol sedge Carex vulpinoidea Michx. fox sedge Cyperus acuminatus Torr. & Hook. ex Torr. tapertip flatsedge Cyperus echinatus (L.) Wood globe flatsedge Cyperus erythrorhizos Muhl. redroot flatsedge Cyperus esculentus L. chufa flatsedge Cyperus ferax L.C. Rich. Cyperus filiculmis auct. non Vahl Cyperus hallii Britt. Cyperus lupulinus (Spreng.) Marcks ssp. lupulinus Great Plains flatsedge Cyperus odoratus L. fragrant flatsedge Cyperus ovularis (Michx.) Torr. Cyperus pseudovegetus Steud. marsh flatsedge Cyperus schweinitzii Torr. Schweinitz's flatsedge Cyperus setigerus Torr. & Hook. lean flatsedge Cyperus squarrosus L. bearded flatsedge Oklahoma Native Plant Record 39 Volume 2, Number 1, December 2002 Hoagland, B.W. Cyperus strigosus L. strawcolored flatsedge Eleocharis acutisquamata Buckl. sharpscale spikerush Eleocharis atropurpurea (Retz.) J. &. K. Presl. purple spikerush Eleocharis compressa Sullivant flatstem spikerush Eleocharis engelmannii Steud. Engelmann's spikerush Eleocharis erythropoda Steud. bald spikerush Eleocharis montevidensis Kunth sand spikerush Eleocharis obtusa (Willd.) J.A. Schultes blunt spikerush Eleocharis palustris (L.) Roemer & J.A. Schultes common spikerush Eleocharis parvula (Roemer & J.A. Schultes) Link ex Bluff, Nees & Schauer dwarf spikerush Eleocharis quadrangulata (Michx.) Roemer & J.A. Schultes squarestem spikerush Fimbristylis puberula (Michx.) Vahl hairy fimbry Fimbristylis puberula (Michx.) Vahl var. puberula hairy fimbry Fimbristylis spadicea auct. non (L.) Vahl. Fimbristylis vahlii (Lam.) Link Vahl's fimbry Fuirena simplex Vahl western umbrella-sedge Hemicarpha aristulata (Coville) Smyth Hemicarpha drummondii Nees Hemicarpha micrantha (Vahl) Pax Lipocarpha aristulata (Coville) G. Tucker awned halfchaff sedge Lipocarpha drummondii (Nees) G. Tucker Drummond's halfchaff sedge Lipocarpha micrantha (Vahl) G. Tucker smallflower halfchaff sedge Schoenoplectus acutus (Muhl. ex Bigelow) A.& D. Löve var. acutus hardstem bulrush Schoenoplectus americanus (Pers.) Volk. ex Schinz & R. Keller chairmaker's bulrush Schoenoplectus hallii (Gray) S.G. Sm. Hall's bulrush Schoenoplectus pungens (Vahl) Palla common threesquare Schoenoplectus tabernaemontani (K.C. Gmel.) Palla softstem bulrush Scirpus americanus Pers. Scirpus atrovirens Willd. green bulrush Scirpus lineatus Michx. rusty bulrush, drooping bulrush Scirpus pallidus (Britt.) Fern. cloaked bulrush Scirpus pendulus Muhl. rufous bulrush Scleria ciliata Michx. fringed nutrush Scleria pauciflora Muhl. ex Willd. fewflower nutrush Ebenaceae Diospyros virginiana L. common persimmon Elatinaceae Bergia texana (Hook.) Seub. ex Walp. Texas bergia 40 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Euphorbiaceae Acalypha gracilens Gray var. monococca Engelm. ex Gray Acalypha monococca (Engelm. ex Gray) L. Mill. &Gandhi slender threeseed mercury Acalypha ostryifolia Riddell pineland threeseed mercury Acalypha virginica L. Virginia threeseed mercury Argythamnia mercurialina (Nutt.) Muell.-Arg. tall silverbush Argythamnia mercurialina (Nutt.) Muell.-Arg. var. mercurialina tall silverbush Chamaesyce maculata (L.) Small spotted sandmat Chamaesyce missurica (Raf.) Shinners prairie sandmat Chamaesyce nutans (Lag.) Small eyebane Chamaesyce prostrata (Ait.) Small prostrate sandmat Chamaesyce serpens (Kunth) Small matted sandmat Cnidoscolus texanus (Muell.-Arg.) Small Texas bullnettle Croton capitatus Michx. hogwort Croton capitatus Michx. var. capitatus hogwort Croton capitatus Michx. var. lindheimeri (Engelm. & Gray) Muell.-Arg. Lindheimer's hogwort Croton glandulosus L. vente conmigo Croton glandulosus L. var. septentrionalis Muell.-Arg. vente conmigo Croton lindheimerianus Scheele threeseed croton Croton monanthogynus Michx. prairie tea Croton texensis (Klotzsch) Muell.-Arg. Texas croton Euphorbia chamaesyce auct. non L. Euphorbia corollata L. flowering spurge Euphorbia cyathophora Murr. fire on the mountain Euphorbia dentata Michx. toothed spurge Euphorbia lata Engelm. Euphorbia longicruris Scheele wedgeleaf spurge Euphorbia maculata L. Euphorbia marginata Pursh snow on the mountain Euphorbia missurica Raf. Euphorbia nutans Lag. Euphorbia serpens Kunth Euphorbia spathulata Lam. warty spurge Euphorbia supina Raf. Phyllanthus abnormis Baill. Drummond's leaf-flower Phyllanthus abnormis Baill. var. abnormis Drummond's leaf-flower Phyllanthus caroliniensis Walt. Carolina leaf-flower Phyllanthus polygonoides Nutt. ex Spreng. smartweed leaf-flower Stillingia sylvatica Garden ex L. queen's-delight Tragia ramosa Torr. branched noseburn Fabaceae Acacia angustissima (P. Mill.) Kuntze var. hirta (Nutt.) B.L. Robins. prairie acacia Oklahoma Native Plant Record 41 Volume 2, Number 1, December 2002 Hoagland, B.W. Amorpha canescens Pursh leadplant Amorpha fruticosa L. desert false indigo Apios americana Medik. groundnut Astragalus caryocarpus Ker-Gawl. Astragalus crassicarpus Nutt. var. crassicarpus groundplum milkvetch Astragalus crassicarpus Nutt. var. trichocalyx (Nutt.) Barneby groundplum milkvetch Astragalus distortus Torr. & Gray Ozark milkvetch Astragalus lindheimeri Engelm. ex Gray Lindheimer's milkvetch Astragalus nuttallianus DC. var. nuttallianus smallflowered milkvetch Astragalus plattensis Nutt. Platte River milkvetch Baptisia australis (L.) R. Br. ex Ait. f. blue wild indigo Baptisia australis (L.) R. Br. ex Ait. f. var. minor (Lehm.) Fern. blue wild indigo Baptisia bracteata Muhl. ex Ell. longbract wild indigo Baptisia bracteata Muhl. ex Ell. var. glabrescens (Larisey) Isely Baptisia bracteata Muhl. ex Ell. var. leucophaea (Nutt.) Kartesz & Gandhi longbract wild indigo Baptisia leucophaea Nutt. Baptisia sphaerocarpa Nutt. yellow wild indigo Baptisia Vent. wild indigo Cassia fasciculata Michx. Cassia marilandica L. Cercis canadensis L. eastern redbud Chamaecrista fasciculata (Michx.) Greene sleepingplant Clitoria mariana L. Atlantic pigeonwings Crotalaria sagittalis L. arrowhead rattlebox Dalea aurea Nutt. ex Pursh golden prairie clover Dalea candida Michx. ex Willd. white prairie clover Dalea candida Michx. ex Willd. var. candida white prairie clover Dalea candida Michx. ex Willd. var. oligophylla (Torr.) Shinners white prairie clover Dalea enneandra Nutt. nineanther prairie clover Dalea frutescens Gray black prairie clover Dalea multiflora (Nutt.) Shinners roundhead prairie clover Dalea purpurea Vent. violet prairie clover Dalea purpurea Vent. var. purpurea violet prairie clover Dalea tenuifolia (Gray) Shinners slimleaf prairie clover Dalea villosa (Nutt.) Spreng. var. villosa silky prairie clover Desmanthus illinoensis (Michx.) MacM. ex B.L. Robins. & Fern. prairie bundleflower Desmanthus leptolobus Torr. & Gray slenderlobe bundleflower Desmodium canescens (L.) DC. hoary ticktrefoil Desmodium ciliare (Muhl. ex Willd.) DC. hairy small-leaf ticktrefoil Desmodium dillenii Darl. p.p. Desmodium glutinosum (Muhl. ex Willd.) Wood pointedleaf ticktrefoil 42 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Desmodium illinoense Gray Illinois ticktrefoil Desmodium nudiflorum (L.) DC. nakedflower ticktrefoil Desmodium paniculatum (L.) DC. panicledleaf ticktrefoil Desmodium paniculatum (L.) DC. var. paniculatum panicledleaf ticktrefoil Desmodium sessilifolium (Torr.) Torr. & Gray sessileleaf ticktrefoil Desmodium tweedyi Britt. Tweedy's ticktrefoil Desmodium viridiflorum (L.) DC. velvetleaf ticktrefoil Galactia volubilis (L.) Britt. downy milkpea Gleditsia triacanthos L. honeylocust Gymnocladus dioicus (L.) K. Koch Kentucky coffeetree Hoffmannseggia glauca (Ortega) Eifert Indian rushpea Hosackia americana (Nutt.) Piper Indigofera miniata Ortega coastal indigo Indigofera miniata Ortega var. leptosepala (Nutt. ex Torr. & Gray) B.L. Turner western indigo Lespedeza capitata Michx. roundhead lespedeza Lespedeza intermedia sensu Clewell, 1966 Lespedeza procumbens Michx. trailing lespedeza Lespedeza repens (L.) W. Bart. creeping lespedeza Lespedeza virginica (L.) Britt. slender lespedeza Lotus purshianus F. E. & E. S. Clements Lotus unifoliolatus (Hook.) Benth. var. unifoliolatus American bird's-foot trefoil Lupinus texensis Hook. Texas lupine Medicago minima (L.) L. Bur medick Medicago sativa L. alfalfa Melilotus albus Melilotus officinalis (L.) Lam. yellow sweetclover Mimosa nuttallii (DC.) B.L. Turner Nuttall's sensitive-briar Neptunia lutea (Leavenworth) Benth. yellow puff Oxytropis lambertii Pursh purple locoweed Oxytropis lambertii Pursh var. articulata (Greene) Barneby purple locoweed Oxytropis lambertii Pursh var. lambertii purple locoweed Pediomelum cuspidatum (Pursh) Rydb. largebract Indian breadroot Pediomelum esculentum (Pursh) Rydb. large Indian breadroot Pediomelum linearifolium (Torr. & Gray) J. Grimes narrowleaf Indian breadroot Petalostemon occidentalis (Heller) Fern. Petalostemon purpureus (Vent.) Rydb. Prosopis glandulosa Torr. honey mesquite Prosopis glandulosa Torr. var. glandulosa honey mesquite Prosopis juliflora (Sw.) DC. var. glandulosa (Torr.) Cockerell Psoralea argophylla Pursh Oklahoma Native Plant Record 43 Volume 2, Number 1, December 2002 Hoagland, B.W. Psoralea cuspidata Pursh Psoralea esculenta Pursh Psoralea linearifolia Torr. & Gray Psoralea tenuiflora Pursh Psoralidium lanceolatum (Pursh) Rydb. lemon scurfpea Psoralidium tenuiflorum (Pursh) Rydb. slimflower scurfpea Robinia pseudoacacia L. black locust Schrankia uncinata Willd. Senna marilandica (L.) Link Maryland senna Sophora affinis Torr. & Gray Eve's necklacepod Strophostyles helvula (L.) Ell. trailing fuzzybean Strophostyles leiosperma (Torr. & Gray) Piper slickseed fuzzybean Tephrosia onobrychoides Nutt. multibloom hoarypea Tephrosia virginiana (L.) Pers. Virginia tephrosia Trifolium dubium Sibthorp suckling clover Trifolium reflexum L. buffalo clover Trifolium repens L. white clover Vicia americana Muhl. ex Willd. ssp. minor (Hook.) C.R. Gunn mat vetch Vicia caroliniana Walt. Carolina vetch Vicia ludoviciana Nutt. Louisiana vetch Vicia ludoviciana Nutt. ssp. ludoviciana Louisiana vetch Vicia ludoviciana Nutt. var. texana (Torr. & Gray) Shinners Vicia texana (Torr. & Gray) Small Fagaceae Quercus buckleyi Nixon & Dorr Buckley oak Quercus fusiformis Small plateau oak Quercus macrocarpa Michx. bur oak Quercus marilandica Muench. blackjack oak Quercus mohriana Buckl. ex Rydb. Mohr oak Quercus muehlenbergii Engelm. chinkapin oak Quercus shumardii Buckl. Shumard's oak Quercus stellata Wangenh. post oak Quercus velutina Lam. black oak Fumariaceae Corydalis aurea Willd. scrambled eggs Corydalis aurea Willd. ssp. occidentalis (Engelm. ex Gray) G.B. Ownbey Corydalis curvisiliqua Engelm. curvepod fumewort Corydalis curvisiliqua Engelm. ssp. grandibracteata (Fedde) G.B. Ownbey curvepod fumewort Corydalis micrantha (Engelm. ex Gray) Gray smallflower fumewort Corydalis micrantha (Engelm. ex Gray) Gray ssp. micrantha smallflower fumewort Gentianaceae Centaurium texense (Griseb.) Fern. Lady Bird's centaury Eustoma russellianum (Hook.) G. Don 44 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Sabatia campestris Nutt. Texas star Geraniaceae Erodium cicutarium (L.) L'Hér. ex Ait. redstem stork's bill Geranium carolinianum L. Carolina geranium Grossulariaceae Ribes aureum Pursh var. villosum DC. golden currant Haloragaceae Myriophyllum aquaticum (Vell.) Verdc. parrot feather watermilfoil Myriophyllum heterophyllum Michx. twoleaf watermilfoil Myriophyllum pinnatum (Walt.) B.S.P. cutleaf watermilfoil Myriophyllum sibiricum Komarov shortspike watermilfoil Myriophyllum spicatum L. spike watermilfoil Myriophyllum spicatum L. var. exalbescens (Fern.) Jepson Hippocastanaceae Aesculus glabra Willd. Ohio buckeye Aesculus glabra Willd. var. arguta (Buckl.) B.L. Robins. Ohio buckeye Hydrophyllaceae Ellisia nyctelea (L.) L. Aunt Lucy Nama hispidum Gray bristly nama Nemophila phacelioides Nutt. largeflower baby blue eyes Phacelia congesta Hook. caterpillars Phacelia strictiflora (Engelm. & Gray) Gray prairie phacelia Phacelia strictiflora (Engelm. & Gray) Gray var. lundelliana Constance Lundell's phacelia Iridaceae Nemastylis geminiflora Nutt. prairie pleatleaf Sisyrinchium angustifolium P. Mill. narrowleaf blue-eyed grass Sisyrinchium campestre Bickn. prairie blue-eyed grass Isoetaceae Isoetes melanopoda Gay & Durieu ex Durieu blackfoot quillwort Juglandaceae Carya cordiformis (Wangenh.) K. Koch bitternut hickory Carya illinoinensis (Wangenh.) K. Koch pecan Juglans microcarpa Berl. little walnut Juglans nigra L. black walnut Juncaceae Juncus acuminatus Michx. tapertip rush Juncus aristulatus Michx. Juncus brachycarpus Engelm. whiteroot rush Juncus diffusissimus Buckl. slimpod rush Juncus dudleyi Wieg. Dudley's rush Juncus interior Wieg. inland rush Juncus marginatus Rostk. grassleaf rush Oklahoma Native Plant Record 45 Volume 2, Number 1, December 2002 Hoagland, B.W. Juncus marginatus Rostk. var. setosus Coville Juncus nodatus Coville stout rush Juncus scirpoides Lam. needlepod rush Juncus secundus Beauv. ex Poir. lopsided rush Juncus tenuis Willd. poverty rush Juncus torreyi Coville Torrey's rush Krameriaceae Krameria lanceolata Torr. trailing krameria Lamiaceae Hedeoma hispida Pursh rough false pennyroyal Hedeoma reverchonii (Gray) Gray Reverchon's false pennyroyal Lamium amplexicaule L. henbit deadnettle Lycopus americanus Muhl. ex W. Bart. American water horehound Lycopus americanus Muhl. ex W. Bart. var. scabrifolius Fern. Mentha spicata L. spearmint Monarda citriodora Cerv. ex Lag. lemon beebalm Monarda clinopodioides Gray basil beebalm Monarda dispersa Small Monarda fistulosa L. wild bergamot Monarda mollis L. Monarda pectinata Nutt. pony beebalm Monarda punctata L. spotted beebalm Nepeta cataria L. catnip Prunella vulgaris L. common selfheal Salvia azurea Michx. ex Lam. azure blue sage Scutellaria drummondii Benth. Drummond's skullcap Scutellaria ovata Hill heartleaf skullcap Scutellaria wrightii Gray Wright's skullcap Teucrium canadense L. Canada germander Teucrium canadense L. var. occidentale (Gray) McClintock & Epling western germander Teucrium canadense L. var. virginicum (L.) Eat. Teucrium laciniatum Torr. lacy germander Lentibulariaceae Utricularia gibba L. humped bladderwort Linaceae Linum berlandieri Hook. var. berlandieri Berlandier's yellow flax Linum hudsonioides Planch. Texas flax Linum imbricatum (Raf.) Shinners tufted flax Linum lewisii Pursh prairie flax Linum pratense (J.B.S. Norton) Small meadow flax Linum rigidum Pursh stiffstem flax Linum rigidum Pursh var. berlandieri (Hook.) Torr. & Gray Linum rigidum Pursh var. rigidum stiffstem flax 46 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Linum sulcatum Riddell grooved flax Linum sulcatum Riddell var. sulcatum grooved flax Loasaceae Mentzelia albescens (Gillies & Arn.) Griseb. wavyleaf blazingstar Mentzelia decapetala (Pursh ex Sims) Urban & Gilg ex Gilg tenpetal blazingstar Mentzelia oligosperma Nutt. ex Sims chickenthief Mentzelia reverchonii (Urban & Gilg) H.J. Thompson & Zavortink Reverchon's blazingstar Lythraceae Ammannia auriculata Willd. eared redstem Ammannia coccinea Rottb. valley redstem Ammannia robusta Heer & Regel grand redstem Lythrum alatum Pursh winged lythrum Lythrum alatum Pursh var. lanceolatum (Ell.) Torr. & Gray ex Rothrock Rotala ramosior (L.) Koehne lowland rotala Malvaceae Abutilon incanum (Link) Sweet pelotazo Callirhoe alcaeoides (Michx.) Gray light poppymallow Callirhoe digitata Nutt. winecup Callirhoe involucrata (Torr. & Gray) Gray purple poppymallow Callirhoe involucrata (Torr. & Gray) Gray var. involucrata purple poppymallow Callirhoe leiocarpa R.F. Martin tall poppymallow Callirhoe pedata (Nutt. ex Hook.) Gray palmleaf poppymallow Hibiscus trionum L. flower of an hour Sida spinosa L. prickly fanpetals Sphaeralcea coccinea (Nutt.) Rydb. scarlet globemallow Sphaeralcea coccinea (Nutt.)Rydb. ssp. coccinea scarlet globemallow Menispermaceae Cocculus carolinus (L.) DC. Carolina coralbead Menispermum canadense L. common moonseed Molluginaceae Mollugo verticillata L. green carpetweed Moraceae Maclura pomifera (Raf.) Schneid. osage orange Morus alba L. white mulberry Morus rubra L. red mulberry Nelumbonaceae Nelumbo lutea Willd. American lotus Nyctaginaceae Mirabilis albida (Walt.) Heimerl white four o'clock Mirabilis glabra (S. Wats.) Standl. smooth four o'clock Mirabilis hirsuta (Pursh) MacM. hairy four o'clock Oklahoma Native Plant Record 47 Volume 2, Number 1, December 2002 Hoagland, B.W. Mirabilis linearis (Pursh) Heimerl narrowleaf four o'clock Mirabilis nyctaginea (Michx.) MacM. heartleaf four o'clock Nymphaeaceae Nymphaea odorata Ait. American white waterlily Oleaceae Fraxinus pennsylvanica Marsh. green ash Onagraceae Calylophus berlandieri Spach Berlandier's sundrops Calylophus berlandieri Spach ssp. pinifolius (Engelm. ex Gray) Towner Berlandier's sundrops Calylophus hartwegii (Benth.) Raven ssp. pubescens (Gray) Towner & Raven Hartweg's sundrops Calylophus serrulatus (Nutt.) Raven yellow sundrops Gaura coccinea Nutt. ex Pursh scarlet beeblossom Gaura longiflora Spach longflower beeblossom Gaura parviflora Dougl. ex Lehm. Gaura sinuata Nutt. ex Ser. wavyleaf beeblossom Gaura suffulta Engelm. ex Gray kisses Gaura suffulta Engelm. ex Gray ssp. suffulta kisses Gaura triangulata Buckl. prairie beeblossom Jussiaea diffusa auct. non Forsk. Jussiaea repens L. var. glabrescens Kuntze Ludwigia alternifolia L. seedbox Ludwigia decurrens Walt. wingleaf primrose-willow Ludwigia palustris (L.) Ell. marsh seedbox Ludwigia peploides (Kunth) Raven ssp. glabrescens (Kunze) Raven floating primrose-willow Ludwigia repens J.R. Forst. creeping primrose-willow Oenothera biennis L. common evening-primrose Oenothera grandis (Britt.) Smyth showy evening-primrose Oenothera laciniata Hill cutleaf evening-primrose Oenothera linifolia Nutt. threadleaf evening-primrose Oenothera macrocarpa Nutt. ssp. incana (Gray) Wagner bigfruit evening-primrose Oenothera macrocarpa Nutt. ssp. macrocarpa bigfruit evening-primrose Oenothera speciosa Nutt. pinkladies Oenothera triloba Nutt. stemless evening-primrose Stenosiphon linifolius (Nutt. ex James) Heynh. false gaura Oxalidaceae Oxalis corniculata L. creeping woodsorrel Oxalis dillenii Jacq. Oxalis stricta L. common yellow oxalis Oxalis violacea L. violet woodsorrel Papaveraceae Argemone polyanthemos (Fedde) G.B. Ownbey crested prickly poppy Papaver dubium L. blindeyes 48 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Passifloraceae Passiflora lutea L. yellow passionflower Pedaliaceae Proboscidea louisianica (P. Mill.) Thellung ram's horn Phytolaccaceae Phytolacca americana L. American pokeweed Rivina humilis L. rougeplant Plantaginaceae Plantago aristata Michx. largebracted plantain Plantago elongata Pursh prairie plantain Plantago major L. common plantain Plantago patagonica Jacq. woolly plantain Plantago pusilla Nutt. dwarf plantain Plantago rhodosperma Dcne. redseed plantain Plantago rugelii Dcne. blackseed plantain Plantago virginica L. Virginia plantain Plantago wrightiana Dcne. Wright's plantain Platanaceae Platanus occidentalis L. American sycamore Polemoniaceae Gilia aggregata (Pursh) Spreng. ssp. formosissima (Greene) Wherry Gilia rubra (L.) Heller Ipomopsis rubra (L.) Wherry standing-cypress Phlox longipilosa Waterfall longhair phlox Polygalaceae Polygala alba Nutt. white milkwort Polygala verticillata L. whorled milkwort Polygala verticillata L. var. isocycla Fern. whorled milkwort Polygonaceae Eriogonum annuum Nutt. annual buckwheat Eriogonum longifolium Nutt. longleaf buckwheat Eriogonum longifolium Nutt. var. longifolium longleaf buckwheat Persicaria punctata (Ell.) Small Polygonum amphibium L. var. emersum Michx. longroot smartweed Polygonum aviculare L. prostrate knotweed Polygonum bicorne Raf. Polygonum convolvulus L. black bindweed Polygonum hydropiper L. marshpepper knotweed Polygonum hydropiperoides Michx. swamp smartweed Polygonum lapathifolium L. curlytop knotweed Polygonum pensylvanicum L. Pennsylvania smartweed Polygonum persicaria L. spotted ladysthumb Polygonum punctatum Ell. dotted smartweed Polygonum ramosissimum Michx. bushy knotweed Polygonum ramosissimum Michx. var. prolificum Small bushy knotweed Oklahoma Native Plant Record 49 Volume 2, Number 1, December 2002 Hoagland, B.W. Polygonum scandens L. climbing false buckwheat Polygonum striatulum B.L. Robins. striped knotweed Polygonum tenue Michx. pleatleaf knotweed Polygonum virginianum L. jumpseed Rumex altissimus Wood pale dock Rumex crispus L. curly dock Pontederiaceae Heteranthera limosa (Sw.) Willd. blue mudplantain Pontederia cordata L. pickerelweed Portulacaceae Claytonia virginica L. Virginia springbeauty Portulaca pilosa L. kiss me quick Portulaca umbraticola Kunth ssp. lanceolata (Engelm.) Matthews & Ketron Talinum calycinum Engelm. largeflower fameflower Talinum parviflorum Nutt. sunbright Primulaceae Androsace occidentalis Pursh western rockjasmine Dodecatheon meadia L. pride of Ohio Dodecatheon meadia L. ssp. meadia pride of Ohio Samolus floribundus Kunth Samolus parviflorus Raf. Samolus valerandi L. ssp. parviflorus (Raf.) Hultén seaside brookweed Ranunculaceae Anemone berlandieri Pritz. tenpetal thimbleweed Anemone caroliniana Walt. Carolina anemone Anemone decapetala auct. non Ard. Clematis pitcheri Torr. & Gray bluebill Clematis versicolor Small ex Rydb. pale leather flower Consolida ajacis (L.) Schur doubtful knight's-spur Delphinium carolinianum Walt. Carolina larkspur Delphinium carolinianum Walt. ssp. virescens (Nutt.) Brooks Carolina larkspur Myosurus minimus L. tiny mousetail Ranunculus abortivus L. littleleaf buttercup Ranunculus longirostris Godr. longbeak buttercup Ranunculus sceleratus L. cursed buttercup Rhamnaceae Ceanothus herbaceus Raf. Jersey tea Rosaceae Agrimonia parviflora Ait. harvestlice Crataegus crus-galli L. cockspur hawthorn Crataegus engelmannii Sarg. Engelmann's hawthorn Crataegus mollis Scheele Arnold hawthorn Crataegus reverchonii Sarg. Reverchon's hawthorn Crataegus viridis L. green hawthorn Crataegus viridis L. var. viridis green hawthorn 50 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Fragaria virginiana Duchesne Virginia strawberry Geum canadense Jacq. white avens Geum canadense Jacq. var. camporum (Rydb.) Fern. & Weatherby white avens Potentilla arguta Pursh tall cinquefoil Potentilla arguta Pursh ssp. arguta tall cinquefoil Prunus angustifolia Marsh. Chickasaw plum Prunus gracilis Engelm. & Gray Oklahoma plum Prunus hortulana Bailey hortulan plum Prunus mexicana S. Wats. Mexican plum Prunus persica (L.) Batsch peach Prunus rivularis Scheele creek plum Prunus virginiana L. chokecherry Rosa foliolosa Nutt. ex Torr. & Gray white prairie rose Rosa multiflora Thunb. ex Murr. multiflora rose Rubus aboriginum Rydb. garden dewberry Rubus allegheniensis Porter Allegheny blackberry Rubus frondosus Bigelow yankee blackberry Rubus idaeus L. ssp. strigosus (Michx.) Focke grayleaf red raspberry Rubus ostryafolius Rydb. highbush blackberry Rubus trivialis Michx. southern dewberry Sanguisorba annua (Nutt. ex Hook.) Torr. & Gray prairie burnet Rubiaceae Cephalanthus occidentalis L. common buttonbush Diodia teres Walt. poorjoe Galium aparine L. stickywilly Galium circaezans Michx. var. hypomalacum Fern. licorice bedstraw Galium obtusum Bigelow bluntleaf bedstraw Galium pilosum Ait. hairy bedstraw Galium pilosum Ait. var. pilosum hairy bedstraw Galium texense Gray Texas bedstraw Galium virgatum Nutt. southwestern bedstraw Hedyotis nigricans (Lam.) Fosberg diamondflowers Hedyotis nigricans (Lam.) Fosberg var. nigricans diamondflowers Houstonia longifolia var. longifolia Gaertn. Houstonia purpurea L. var. purpurea Venus' pride Houstonia pusilla Schoepf tiny bluet Sherardia arvensis L. blue fieldmadder Rutaceae Ptelea trifoliata L. common hoptree Ptelea trifoliata L. ssp. polyadenia (Greene) V. Bailey pallid hoptree Ptelea trifoliata L. ssp. trifoliata var. trifoliate common hoptree Oklahoma Native Plant Record 51 Volume 2, Number 1, December 2002 Hoagland, B.W. Ptelea trifoliata ssp. trifoliata L. var. mollis Torr. & Gray common hoptree Salicaceae Populus deltoides Bartr. ex Marsh. eastern cottonwood Salix caroliniana Michx. coastal plain willow Salix nigra Marsh. black willow Santalaceae Comandra umbellata (L.) Nutt. ssp. pallida (A. DC.) Piehl pale bastard toadflax Sapindaceae Cardiospermum halicacabum L. love in a puff Sapindus drummondii Hook. & Arn. Sapindus saponaria L. var. drummondii (Hook. & Arn.) L.Benson western soapberry Sapotaceae Sideroxylon lanuginosum Michx. gum bully Sideroxylon lanuginosum Michx. ssp. rigidum (Gray) T.D. Pennington Scrophulariaceae Agalinis heterophylla (Nutt.) Small ex Britt. prairie false foxglove Bacopa rotundifolia (Michx.) Wettst. disk waterhyssop Castilleja coccinea (L.) Spreng. scarlet Indian paintbrush Castilleja purpurea (Nutt.) G. Don downy Indian paintbrush Castilleja purpurea (Nutt.) G. Don var. citrina (Pennell) Shinners prairie Indian paintbrush Collinsia verna Nutt. spring blue eyed Mary Collinsia violacea Nutt. violet blue eyed Mary Gratiola sphaerocarpa Ell. Gratiola virginiana L. roundfruit hedgehyssop Leucospora multifida (Michx.) Nutt. narrowleaf paleseed Lindernia anagallidea (Michx.) Pennell Lindernia dubia (L.) Pennell yellowseed false pimpernel Macuillamia rotundifolia (Michx.) Raf. Mimulus glabratus Kunth roundleaf monkeyflower Mimulus glabratus Kunth var. jamesii (Torr. & Gray ex Benth.) Gray James' monkeyflower Nuttallanthus canadensis (L.) D.A. Sutton Canada toadflax Nuttallanthus texanus (Scheele) D.A. Sutton Texas toadflax Penstemon albidus Nutt. white penstemon Penstemon australis Small ssp. laxiflorus (Pennell) Bennett Penstemon cobaea Nutt. cobaea beardtongue Penstemon fendleri Torr. & Gray Fendler's penstemon Penstemon oklahomensis Pennell Oklahoma beardtongue Scrophularia lanceolata Pursh lanceleaf figwort Veronica arvensis L. corn speedwell Veronica peregrina L. neckweed Veronica peregrina L. ssp. xalapensis (Kunth) Pennell hairy purslane speedwell 52 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Hoagland, B.W. Simaroubaceae Ailanthus altissima (P. Mill.) Swingle tree of heaven Smilacaceae Smilax bona-nox L. saw greenbrier Smilax rotundifolia L. roundleaf greenbrier Smilax tamnoides L. bristly greenbrier Solanaceae Datura meteloides auct. p.p. non Dunal Physalis angulata L. cutleaf groundcherry Physalis cinerascens (Dunal) A.S. Hitchc. var. cinerascens smallflower groundcherry Physalis heterophylla Nees clammy groundcherry Physalis lobata Torr. Physalis longifolia Nutt. longleaf groundcherry Physalis mollis Nutt. field groundcherry Physalis mollis Nutt. var. mollis field groundcherry Physalis pubescens L. var. integrifolia (Dunal) Waterfall husk tomato Physalis pumila Nutt. dwarf groundcherry Physalis viscosa L. ssp. mollis (Nutt.) Waterfall Quincula lobata (Torr.) Raf. Chinese lantern Solanum citrullifolium A. Braun watermelon nightshade Solanum dimidiatum Raf. western horsenettle Solanum elaeagnifolium Cav. silverleaf nightshade Solanum nigrum auct. non L. Solanum ptychanthum Dunal West Indian nightshade Solanum rostratum Dunal buffalobur nightshade Tamaricaceae Tamarix gallica L. salt cedar Ulmaceae Celtis laevigata Willd. sugarberry Celtis laevigata Willd. var. reticulata (Torr.) L. Benson netleaf hackberry Celtis laevigata Willd. var. texana Sarg. Texan sugarberry Ulmus americana L. American elm Ulmus rubra Muhl. slippery elm Urticaceae Boehmeria cylindrica (L.) Sw. smallspike false nettle Boehmeria cylindrica (L.) Sw. var. drummondiana (Weddell) Weddell Parietaria pensylvanica Muhl. ex Willd. Pennsylvania pellitory Parietaria pensylvanica Muhl. ex Willd. var. obtusa (Rydb. ex Small) Shinners Valerianaceae Valerianella radiata (L.) Dufr. beaked cornsalad Verbenaceae Glandularia bipinnatifida (Nutt.) Nutt. var. bipinnatifida Dakota mock vervain Glandularia canadensis (L.) Nutt. rose mock vervain Oklahoma Native Plant Record 53 Volume 2, Number 1, December 2002 Hoagland, B.W. Glandularia pumila (Rydb.) Umber pink mock vervain Lippia lanceolata Michx. Phryma leptostachya L. American lopseed Phyla incisa Small Phyla lanceolata (Michx.) Greene lanceleaf fogfruit Verbena bipinnatifida Nutt. Verbena bracteata Lag. & Rodr. bigbract verbena Verbena canadensis (L.) Britt. Verbena pumila Rydb. Verbena stricta Vent. hoary verbena Verbena urticifolia L. white vervain Violaceae Hybanthus verticillatus (Ortega) Baill. babyslippers Viola bicolor Pursh field pansy Viola missouriensis Greene Viola retusa Greene Viola sororia Willd. common blue violet Viscaceae Phoradendron flavescens Nutt. ex Engelm. Phoradendron leucarpum (Raf.) Reveal & M.C. Johnston oak mistletoe Vitaceae Ampelopsis cordata Michx. heartleaf peppervine Cissus incisa auct. non Des Moulins Cissus trifoliata (L.) L. sorrelvine Parthenocissus quinquefolia (L.) Planch. Virginia creeper Parthenocissus vitacea (Knerr) A.S. Hitchc. woodbine Vitis cinerea (Engelm.) Millard graybark grape Vitis cinerea (Engelm.) Millard var. cinerea graybark grape Vitis riparia Michx. riverbank grape Vitis rupestris Scheele sand grape Vitis vulpina L. frost grape Zygophyllaceae Tribulus terrestris L. Puncturevine 54 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Schoenoplectus hallii and S. saximontanus 2000 Wichita Mountain Wildlife Refuge Survey Dr. Lawrence K. Magrath Curator-USAO (OCLA) Herbarium Chickasha, OK 73018-5358 A survey to determine locations of populations of Schoenoplectus hallii and S. saximontanus was conducted at Wichita Mountains Wildlife Refuge in August and September 2000. One or both species were found at 20 of the 134 locations surveyed. A distinctive terminal achene character was found specifically that the transverse ridges of S. hallii appeared to be rounded and S. saximontanus appeared to be rounded with a projecting narrow wing. Basal macroachenes have not yet been properly described but are borne singly at the base of each culm and are about 3-4 times larger than the terminal achenes. It is speculated that amphicarpy may be related to grazing pressure, the basal macroachene being produced even if the upper portion is consumed, as a response to grazing. Both species are grazed/disturbed by bison, elk, and longhorns on the Refuge. Introduction A survey to determine locations of populations of Schoenoplectus hallii (A. Gray) S.G. Smith (Hall’s bulrush) and S. saximontanus (Fernald) J. Raynal (Rocky Mountain bulrush) was conducted on the Wichita Mountains Wildlife Refuge during late August through September 2000. The survey team members were myself, Sam Waldstein, Refuge Manager; Chip Kimball, Range Biologist; and Bob Timberman, Biology Technician. Sites selected for observation were areas similar to the Jed Johnson Dam habitat, which is the site of the original collection of S. hallii made in 1987. Selection of sites to be surveyed was done by WMWR team members. The three Sundays spent in the field were some of the hottest of the season with temperatures ranging from 100o to 110oF. General Observations The presence of Schoenoplectus halllii and S. saximontanus at the various sites sampled. represent, for the most part, a response to the drawdown of water levels in the various lakes and ponds. The plants seem to occur mainly on Magrath, L.K. https://doi.org/10.22488/okstate.17.100013 the drawdown mud, sand, or gravel flats. However in some places they occur in shallow water up to a depth of about a foot [30.5cm]. They seem to compete with perennial emergent plants and with most emergent annuals. In addition to the 36 sites that I personally examined, WMWR staff examined an additional 98 sites with similar habitat and found no Schoenoplectus present. Schoenoplectus occurred in only 20 of the 134 sites (14.93%). Of those 134 sites, 4 had both species present (3%), 14 had S. halllii present (10.45%) and 10 had S. saximontanus present (7.46%) (see Table). Based on the WMWR observations, it is strongly advised that the adjacent area of Fort Sill should be inventoried since these two species are, most likely, present there. However, Rahmona Thompson who has conducted plant surveys at Fort Sill has not found either species at this time (pers. comm. 2002). At several of the sites plants had been uprooted as a result of trampling by bison and other animals in the mud flat areas. The number of uprooted plants ranged between 0.0% and 1.0% at those Oklahoma Native Plant Record 55 Volume 2, Number 1, December 2002 Magrath, L.K. locations, but in general was less than .05%. While this obviously does cause some damage to populations in localized areas the damage appears to be negligible and it is even possible that it may be an important transport method of matured achenes to previously uncolonized areas. This could readily be accomplished by mud containing seeds or inflorescences with seeds adhering to the hooves, hair, or skin of the lower part of the animals’ legs. There appear to be three useful characters associated with the achenes that can be used to separate the two species: style branching, achene cross section, and transverse ridges. S. hallii S. saximontanus Style Branches mostly bifid trifid Achene cross section unequally biconvex (1 of 2 sides may be flat) Trigonus (3-sided) Transverse ridges rounded mostly rounded with narrow wing To my knowledge, “Transverse ridges” has not been mentioned in the literature on these two taxa. McKenzie (1988) does not mention this character in his status assessment report on S. hallii, nor does Yatskievych (1999) in Steyermark’s Flora of Missouri. Dr. Marian Smith at Southern Illinois University is working on both terminal achenes and basal macroachenes using Scanning Electron Microscopy and more precise measuring techniques (pers. comm. 2001). So this may be a new useful character to separate these two taxa. (see Figures). McKenzie (1998) reports that “heavy grazing has been noted at sites in Kansas, Missouri, and Wisconsin but it is not known whether this disturbance negatively impacts the species…” It is my hypothesis that amphicarpy may well be a response to heavy grazing pressure disturbance by native grazing animals such as bison, elk, and deer prior to the introduction of exotic grazers such as cattle, horses and sheep by European settlers. It would be a way that the plant could insure seed production even if it is heavily grazed and the terminal achenes damaged or destroyed. The observation that the plant produces only a very limited number of basal macroachenes (one per culm) with significantly more food reserves per achene would seem to support this interpretation. The smaller, more numerous terminal achenes would offer a relatively easy method for seeds to be transported farther distances and to new sites while the macroachenes would provide an excellent way to persist in presently occupied sites. The number and identification of the two species at sites where both occur could be somewhat problematic. S.G. Smith and A. E. Schuyler independently identified a hybrid from WMWR that was collected by M. Smith and McKenzie July 28, 2002 as the first documented, putative hybrid of the two species ever recorded. [USA Oklahoma: Comanche Co., WMWR, 28 July 02, P. McKenzie 2028 pers., WIS, MO] (pers. comm. McKenzie 2002; Smith 2002). There obviously needs to be further research done to confirm or deny this hypothesis as well as possible hybridization between the two species at the Refuge. Voucher Specimens of Schoenoplectus hallii and S. saximontanus resulting from this study are kept in the USAO Herbarium (OCLA) at the University of Science and arts of Oklahoma in Chickasha, Oklahoma. 56 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Magrath, L.K. FIG. (a) and (b) Fruit and inflorescence of Schoenoplectus saximontanus (Steyermark’s Flora of Missouri, 1999. Used by permission). FIG. (e), (f), and (g) Fruit, inflorescence, and habit of Schoenoplectus hallii (Steyermark’s Flora of Missouri, 1999. Used by permission). Oklahoma Native Plant Record 57 Volume 2, Number 1, December 2002 Magrath, L.K. TABLE Schoenoplectus occurrence in thirty-six survey sites August and September 2000 (based on achene characteristics of the two species) Site # Description & Date abundance/# collected/voucher# S. hallii S. saximontanu 1-Pond 27-Aug Draw-down mudflat around pond Some plants grazed, some uprooted Some dried out on shore Few floating in water still alive 0 scat/8/21254 2-Pond 27-Aug Draw-down mudflat 0 0 3-Pond 27 Aug Draw-down mudflat 0 0 4-Corral area 27 Aug Stream with draw-down mudflat Utricularia observered 0 0 5-Kiowa Lake 27 Aug Draw-down mud flat around pond Some plants grazed, some uprooted Some dried out on shore Few floating in water still alive 0 scattered/4/21259 6-Wing Pasture west of creek 27 Aug Draw-down mud flat around pond Some plants grazed 0 rare/12/21261 7-Buford Lake 27 Aug Draw-down mud flats Utricularia and Nelumbo abundant 0 0 58 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Magrath, L.K. 8-Quanah Parker Lake 27 Aug Near Nature Center Draw-down mud flat around pond scat/6/21263 0 9-New Pond by Crater Lake 27 Aug South of Visitor Center Draw-down mud flat 0 0 10-Pond at Sulphur Trap Corrals 27 Aug East of Visitor Center Draw-down mud flat 0 0 11-Pond in Sulphur Trap 27 Aug North of Visitor Center Draw-down mud flat around pond 0 0 12-Jed Johnson Lake 18 Aug 8 identifiable plants. Not collected Plants just coming into bloom Original site for the original Collections for WMWR rare/0/- 0 27 Aug 8 identifiable plants No collections made rare/0/- 0 13-Crater Lake 27 Aug Draw-down mud flat Apparently too much perennial vegetation present 0 0 14-West Gate Pond 3 Sep Draw-down mud flat Apparently too much perennial vegetation present 0 0 15-Comanche Lake 18 Aug Draw-down mud flat scat/2/21237 0 Oklahoma Native Plant Record 59 Volume 2, Number 1, December 2002 Magrath, L.K. 3 Sep Draw-down mud flat Mixed populations scat-loc com/24/21272 scat-locom/83/27271 16-Grama Lake near dam & Gram Flat 18 Aug Draw-down mud flat Several hundred plants Numerous basal rosettes in shallow water (20-30 cm) 0 scat/3/21236 3 Sep Mixed populations on draw-down Several hundred plants Numerous submerged basal rosettes in shallow water (20-30 cm) scat-loc com/48/21274 scat-loc com/15/21273 10 Sep Revisit & complete walk-around Both present on draw-down mud flat Several thousand plants Dominant plant in a few places Numerous basal rosettes present in shallows in several areas scat-loc abd/211/21216 scat/99/21322 17-Hollis Lake 3 Sep Apparently too much perennial vegetation in draw-down zone 0 0 18-Pond 0.5 mile west of Hollis 3 Sep com-loc abd/300+/21278 0 19-Pond 3 Sep Draw-down mudflat 0 0 20-Pond 3 Sep Draw-down mudflat 0 0 21-Boggy Flats 18 August 2000 Draw-down mudflat around pond Some plants grazed or uprooted Some dried out on shore Few floating in water still alive First located by Sam Waldenstein on Aug 17, 2000. 0 loc abd/x/21231 60 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Magrath, L.K. Sun 3 September 2000 Draw-down mud flats on ponds 0 loc abd/5/21280 22-Pond southwest of Grace Mountain 3 Sep Draw-down mud flat Water-clover fern present 0 0 23-Cut Throat Lake 3 Sep Draw-down mud flat Water-clover fern present Beautiful clear water and bass 0 0 24-Northwest Corner of Pinchot Loop 3 Sep Draw-down mud flat around pond 0 0 25-Barow Pit east side of Pinchot Loop 3 Sep Draw-down mud flat around pond 0 0 26-Straight east of site 25 3 Sep Draw-down mud flat around pond Bacopa present scat/8/21287 0 27-Medicine Tank 3 Sep Draw-down mud flat around pond Colonial Bryozoan Pectinatella magnifica Leidy in shallow water by dam Identification by Dr. Mike Mather, USAO common/12/21290 scat/3/21289 28-West Gate Road, Buffalo Gap 10 sep Draw-down mud flat around pond Bacopa present 0 scat/11/21303 29-Winter Valley at end of Wing Fence at Hot Trap 10 Sep Draw-down mud flat around pond Mixed collection not discovered until laboratory observations were made. rare/3/21304 in part* rare/6/21304 in part* Oklahoma Native Plant Record 61 Volume 2, Number 1, December 2002 Magrath, L.K. 30-Winter Valley southeast fo Road 10 Sep Draw-down mudflat around pond Bacopa present scat/2/21305 0 31-North end of Research 10 Sep Draw-down mud flat around pond Bacopa present 0 scat/14/21306 32-Pond in Exhibition Pasture 10 Sep Pond in Exhibition Pasture Draw-down mud flat one plant/0/0 0 33 Ingram House Pond 10 Sep Draw-down mud flat Several hundred plants Bacopa present scat-loc com/19/21309 0 34-Quanah Parker 10 Sep Near dam Draw-down mud flat Several thousand? plants scat-loc com/54/21210 0 35-Quanah Parker 10 Sep SE of Environmental Center Draw-down mud flat Several hundred plants scat-loc com/7/21313 scat-loc com/25/21311 36-Elmer Thomas Lake 10 Sep Draw-down mud flat along north shore scat-loc abd/46/21314 0 Key to abundance descriptors rare = fewer than 10 plants at site scat = scattered, a few plants occurring over several square meters common = many plants occurring over several square meters abundant = large numbers of plants (often the local dominant plant) loc com = many plants in a small area, but may be scattered over a large area loc abd = large numbers of plants in small area, but may be scattered over a large area 62 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Magrath, L.K. References McKenzie, Paul M. 1998. Hall’s Bulrush (Schoenoplectus hallii) Status Assessment. U.S. Fish and Wildlife Service. Columbia, MO. Yatskievych, George. 1999. Steyermark’s Flora of Missouri, Vol 1, rev. ed. Missouri Dept. of Conservation in cooperation with the Missouri Botanical Garden Press. St. Louis, MO. Editor’s Note: Exact locations of individual sites were determined by GPS and are recorded in the WMWR database. However, that information and access to the sites is strictly limited and permission must be requested from the refuge manager. Oklahoma Native Plant Record 63 Volume 2, Number 1, December 2002 Magrath, L.K. Wichita Mountains Wildlife Refuge, Comanche County Echinocerus baileyi on Elk Mountain Trail (photos by Sheila Strawn) 64 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 The Nature Conservency’s Pontatoc Ridge Preserve, Johnston County Entrance to Pontotoc Ridge Preserve (photo by Patricia Folley) Wildflower community on exposed limestone formation at Pontotoc Ridge Preserve (photo by Patricia Folley) Johnson, F.L. https://doi.org/10.22488/okstate.17.100014 Oklahoma Native Plant Record 65 Volume 2, Number 1, December 2002 Johnson, F.L. https://doi.org/10.22488/okstate.17.100014 Pontotoc Ridge Floristic Survey: 1999 transcribed with later additions Forrest L. Johnson Oklahoma Biological Survey Patricia Folley (ed.) President, Oklahoma Native Plant Society In 1997, the Oklahoma Biological Survey and the Oklahoma Native Plant Society undertook an unusual joint project: the surveying of the plant resources of a preserve recently acquired by The Nature Conservancy. In part this was because of the deep attachment of some of the participants for this nearby and very interesting place. During the early exploration of the “Smith Ranch Site”, as it was called, amateurs and interns were brought in to call out the names of plants as they walked, or in some cases, rode past in trucks. As a rapid assessment, that was probably a good thing, but it certainly wasn’t scientific, and none of those names called could be verified for publication. The Biological Survey allotted the necessary manpower and equipment to conduct a survey structured after the ones being done for several military bases in the area. Frequent, disciplined visits were made to view the diverse habitats as the plants thereon developed over the course of the growing year. One or two specimens of each species was taken, processed, identified and preserved in a public herbarium for the use of future researchers. The herbarium chosen was the Robert Bebb Herbarium (OKL) at the University of Oklahoma, host institution for the Biological Survey. 2002 INTRODUCTION Forrest Johnson of the Biological Survey, Patricia Folley of the Oklahoma Native Plant Society and a co-editor of the Flora of Oklahoma Project, and two fieldassistants, Newell McCarty and Deborah Benesh from the University of Oklahoma’s Bebb Herbarium carried out the fieldwork for the Survey. Almost all processing of specimens, including identification, was done by the late Dr. Johnson. Folley made the identifications of the specimens of sedges, and Benesh and Johnson maintained the databaseand prepared labels. Eventually, over 500 specimens were mounted and placed in the herbarium. Also included in the survey is Plantago rugelii, which was collected by Patricia Folley in 1993 when the site was still known as “Smith Ranch”. Since the survey Folley and others have added a few species, similarlydocumented, to those surveyed in 1997. They are identified by a later collection date in the inventory. No natural living place can ever be fully documented and there are still species known to exist in that preserve that have not yet been found in flowering or fruiting condition. It will no doubt provide interesting work for our successors. After the original introduction was written, an infestation of Lespedeza cuneata (serecia lespedeza) proliferated in the old pastures and a program of removal was begun. The research team is indebted to the kindness and assistance so generously contributed by TNC’s on-site managers, Jim and Hollie Erwin. 66 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 INTRODUCTION In the growing season (March- October) of 1997, The Nature Conservancy’s Pontotoc Ridge Preserve was visited approximately every two weeks by a team of botanists from the Oklahoma Native Plant Society and the Oklahoma Biological Survey for the purpose of preparing an inventory of the plant species on the Preserve. Two to four persons on each visit examined several sites on the Preserve. The number and location of sites varied with the condition of the primitive roads on the property. When roads were muddy, we collected plants at three or four sites along the west side of the Preserve. When the roads were dry enough, we visited several widely separated sites spread over the whole property. At each collecting site members of the team walked separate paths through the area. Any plant in flower or fruit that was not on a list of the plants previously collected or which could not be positively identified was collected. Some trees and shrubs that are usually identified by their leaves, buds, etc., were collected without flowers or fruits. Each specimen was pressed, dried, and fumigated to eliminate insects. Then they were stored in the Robert Bebb Herbarium at the University of Oklahoma. The specimens were identified with the keys and descriptions in Flora of the Great Plains (Great Plains Flora Association, 1986, University Press of Kansas, Lawrence) and Keys to the Flora of Oklahoma, (U. T. Waterfall, 1973, published by the author, Stillwater, OK), then confirmed by comparison with known specimens in the Bebb Herbarium. The specimens from Pontotoc Ridge will remain in the Bebb Herbarium and become part of its permanent collection. Several plant names have been changed since the publication of Keys to the Flora of Oklahoma and Flora of the Great Plains. Names in our list names comply with the PLANTS online database: plants.usda.gov/ plants/qurymenu.html , maintained by the U.S. Department of Agriculture, Natural Resources Conservation Service. We included Johnson, F.L. both the old and new names for a few plants that have had recent name changes. The older name is listed in brackets below the new name. Two status and 17 habit categories are defined by the National List of Scientific Plant Names (1982, U.S.D.A. Soil Conservation Service Publication SCS-TP-159, Washington, D.C.) They are applied as appropriate for each species. The categories and representative symbols used to define status categories are: (N) native and (I) introduced. Habit is represented by one or more of the following: (A) annual; (B) biennial; (P) perennial; (F) herbaceous; (S) shrub; (T) tree; (G) grasslike; (H) partly woody; (W) woody; ($) succulent; (V) vine; (Z) submersed; (E) emergent; (L) floating; (@) tree epiphyte; (+) parasitic; and (_) saprophytic. This information is determined by examining the specimens and consulting the designated floras. These symbols are combined into a convenient status/habit code for each species. For example, NPG = native perennial grasslike; NT = native tree, etc. In the 1997 growing season, we collected 399 species in 261 genera and 79 families. About a third of the species found were in the three common Great Plains families: Asteraceae (composites); Poaceae (grasses) and Fabaceae (legumes). Only about 6 percent of the species are exotic and none of the exotic species is conspicuous or seems to present a threat to the native vegetation. No truly rare plants were found, although Penstemon oklahomensis (S3) and Echinocereus riechenbachii (S2) are present. Based on previous experience, we are likely to have found about 80% of the total plant species present at Pontotoc Ridge. There are probably about 500 plant species in the area. Some perennial plant species do not flower every year, and some annuals vary greatly in abundance from year to year. Continued investigation will result in more species being found. Several field trips to Pontotoc Ridge are planned in 1998. Oklahoma Native Plant Record 67 Volume 2, Number 1, December 2002 Johnson, F.L. Pontotoc Ridge Floristic Survey Acanthaceae (acanthus) Ruellia humilis Nutt. NPF 29 May, 10 Sep 1997 Low ruellia infrequent, prairie Ruellia strepens L. NPF 29 May 1997 Limestone ruellia Aceraceae (maple) Acer negundo L. NT 18 Jul 1997 Box elder infrequent, floodplain forest Agavaceae (yucca, agave family) Yucca glauca Nuttall NS (not yet collected) Common yucca infrequent, uplands Anacardiaceae (sumac, poison ivy) Rhus aromatica Ait. NS 28 Mar, 18 Jul 1997 Aromatic sumac common, rocky upland Rhus copallinum L. NS 18 Jul 1997 Winged sumac infrequent, rockly upland Rhus glabra L. NS 18 Jun 1997 Smooth sumac common, rocky upland Apiaceae (carrot, celery, etc.) *Ammoselinum popei, see Chaerophyllum Bifora americana (DC.) Watson NAF 14 May 1997 Prairie bishop’s weed infrequent, rocky upland Chaerophyllum procumbens (L.) Crantz NAF 15 Apr 1997 Wild chervil common, disturbed area Chaerophyllum taintureiri Hook. NAF 15 Apr, 14 May 1997 Wild chervil common, disturbed area, mowed meadow Cryptotaenia canadensis (L.) DC. NPF 29 May 1997 Honewort common, floodplain forest Daucus pusillus Michaux NAF 18 Jun 1997 Rattlesnake weed common, disturbed area Eryngium leavenworthii T. & G. NAF 10 Sep 1997 Rattlesnake master common, rocky upland Eryngium yuccifolium Michaux var. synchaetum Gray ex C & Rose NPF 31 July 1997 Button snakeroot common, prairie Lomatium foeniculaceum (Nutt.) C.&R. var. foeniculaceum NPF 28 March 1997 Wild parsley common, rocky upland Osmorhiza longistylis (Torr.) DC. NPF 30 April 1997 Anise root common, floodplain forest Polytaenia nuttallii (Nutt.) DC. NPF 14 May 1997 Prairie parsley common, prairie Sanicula canadensis L NPF 30 April, 29 May 1997 Snakeroot common, floodplain forest Torilis arvensis (Huds.) Link IAF 18 June 1997 Hedge parsley common, disturbed area Trepocarpus aethusae Nuttall NAF 18 June 1997 White nymph infrequent, rocky upland Zizia aurea (L.) Koch NPF 14 May 1997 Golden alexanders infrequent, floodplain forest Apocynaceae (dogbane) Amsonia ciliata Walt. NPF 10 April 1996 Fringed bluestars infrequent, rocky upland Apocynum cannabinum L. var. cannabinum NPF 29 May 1997 Indian hemp infrequent, prairie Araceae (arum) Arisaema dracontium (L.) Schott NPF (not yet colleted) Green dragon infrequent, floodplain forest Asclepiadaceae (milkweed) Asclepias asperula (Dcne.) Woodson var. decumbens (Nutt.) Shinners NPF 30 Apr, 14 May 1997 Spider antelopehorn common, rocky upland Asclepias stenophylla (Gray) NPF 18 June 1997 Narrowleaf milkweed infrequent, prairie 68 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Johnson, F.L. Asclepias tuberosa L. NPF 18 June 1997 Butterfly milkweed common, prairie Asclepias viridis Walt. NPF 29 May 1997 Green milkweed common, prairie Matelia biflora (Raf.) Woodson NPV 14 May 1997 Twoflower milkvine infrequent, rocky upland Matelia gonocarpa (Walt.) Shinners NPV 18 July 1997 Climbing milkweed infrequent, disturbed area Aspleniaceae (cliff ferns) Woodsia obtusa (Spreng.) Torr. NPF 29 May 1997 Blunt-lobed cliff fern infrequent, Cross Timbers Asteraceae (asters, daisies, sunflowers) Achillea millefolium L. NPF 14 May 1997 Yarrow common, prairie Ambrosia artemisiifolia L. NAF 10 Sept 1997 Common ragweed common, disturbed areas Ambrosia bidentata Michaux NAF 10 Sept 1997 Southern ragweed common disturbed area Ambrosia psilostachya DC. NPF 8 October 1997 Western ragweed infrequent, disturbed areas Ambrosia trifida L. NAF 20 August 1997 Giant ragweed common, disturbed areas Amphiachyris dracunculoides (DC.) Shinners NAF 10 Sept. 1997 Broomweed infrequent, disturbed area Artemisia ludoviciana Nuttall var. mexicana (Willd.) Fernald NPF 10 Sept, 8 October 1997 White sage, watersage infrequent, disturbed area, prairie Aster drummondii Lindl var. parviceps (Shinners) A.G. Jones NPF 8 October 1997 Blue wood aster infrequent, disturbed areas Aster ericoides L. NPF 8 October 1997 Heath aster common, disturbed area Aster oolentangensis Riddell NPF 8 October 1997 Blue aster infrequent, disturbed area Aster patens Ait. NPF 8 October 1997 Spreading aster infrequent, disturbed area, rocky upland Aster prealtus Poir. NPF 8 October 1997 Willowleaf aster infrequent, disturbed area Astranthium integrifolium(Michx.)Nutt. NAF 15 April, 30 April 1997 Western daisy common, disturbed area Brickellia eupatorioides (L.) Raf. NPF False snakeweed uncommon, disturbed area Centaurea americana Nuttall NAF 18 June 1997 Basketflower common, rocky upland Chaetopappa asteroides Nutt. Ex DC. NAF 30 April 1997 Least daisy infrequent, disturbed area Chrysopsis pilosa Nuttall NAF 31 July, 8 Oct. 1997 Golden aster infrequent, disturbed area, prairie Cirsium altissimum (L.) Spreng. NPF 31 July, 10 Sept 1997 Tall thistle infrequent, prairie Cirsium undulatum (Nutt.) Spreng. NPF 18 June 1997 Wavy-leaf thistle common, disturbed area Conyza canadensis (L.) Cronq. NAF 20 August 1997 Horseweed common, disturbed area Echinacea atrorubens Nuttall NPF Rose coneflower common, rocky upland Eclipta prostrata (L.) L. NAF 10 Sept 1997 Yerba de tajo infrequent, pond margin Elephantopus carolinianus Willd. NPF 10 Sept 1997 Elephant’s foot common, floodplain forest Engelmannia pinnatifida Gray ex Nutt. NPF 14 May 1997 Engelmann’s daisy common, rocky upland, prairie Erigeron annuus (L.) Pers. NAF 14 May 1997 Oklahoma Native Plant Record 69 Volume 2, Number 1, December 2002 Johnson, F.L. Daisy fleabane common, prairie Erigeron philadelphicus L. NAF 15 April 1997 Philadelphia fleabane common, riparian Eupatorium serotinum Michaux NPF 20 August 1997 Late boneset infrequent, disturbed areas Gaillardia aestivalis (Walt.) Rock NAF 10 Sept 1997 Summer gaillardia infrequent, prairie Gaillardia pulchella Fouq. NAF 9 May, 18 June 1997 Indian blankets, firewheels common, prairie, rocky upland Gamochaeta purpurea (L.) Cabrera NPF 30 April 1997 Purple cudweed infrequent, disturbed area Gnaphalium obtusifolium L. NPF 10 Sept 1997 Sweet everlasting infrequent, prairie Helenium amarum (Raf.) H. Rock NAF 18 Jun 1997 Bitterweed common, disturbed area Helianthus annuus L. NAF 18 July 1997 Common sunflower infrequent, disturbed area Helianthus hirsutus Raf. NPF 18 July 1997 Hairy or rough sunflower infrequent, disturbed area Helianthus mollis Lam. NPF 31 July 1997 Ashy or soft sunflower common, prairie Heliopsis helianthoides (L.) Sweet NPF 31 July 1997 False sunflower common, floodplain forest Heterotheca latifolia Buckl. NPF 10 Sept 1997 Golden aster common, prairie Hieracium longipilum Torrey NPF 19 Sept 1997 Longbeard hawkweed infrequent, prairie Hymenopappus scabieoseus L’Her. NPF 14 May 1997 Old plainsmen common, rocky upland Hymenoxys linearifolia Hook. NAF 15 April – 15 May 1997 Narrowleaf hymenoxys common, rocky upland, disturbed area Iva annua L. NAF 10 Sept 1997 Annual false ragweed, sumpweed common, disturbed area Krigia caespitosa (Raf.) Chambers NAF 14 May 1997 Common dwarf dandelion infrequent, prairie Krigia dandelion (L.) Nuttall NPF 14 May 1997 Potato dandelion common, prairie Lactuca ludoviciana (Nutt.) DC. NAF 18 Jun 1997 Western wild lettuce infrequent, disturbed area Liatris punctata Hook., var. nebraskana Geiser NPF 10 Sept 1997 Dotted gayfeather abundant, prairie Liatris pycnostachya Michaux NPF 31 July 1997 Button snakeroot common, prairie Liatris squarrosa (L.) Michaux NPF 31 July 1997 Gayfeather common, prairie Lindheimera texana Gray & Engelm NAF 30 April, 11 June 1997 Texas yellow star infrequent, rocky upland, prairie Marshallia caespitosa Nutt. Ex DC. NPF 14 May, 18 June 1997 Barbara’s buttons infrequent, prairie, rocky upland Pluchea odorata (L.) Cass. NAF 8 October 1997 Purple camphorweed common, pond margin Prionopsis ciliata Nuttall NAF 10 Sep 1997 Wax goldenweed infrequent, disturbed area Pyrrhopappus grandiflorus Nuttall NPF 30 April 1997 False dandelion common, disturbed area Pyrrhopappus pauciflorus DC. NAF 14 May 1997 Geiser’s false dandelion common, prairie Ratibida columnifera Woot. & Standl. NPF 18 June 1997 Yellow coneflower common, disturbed area Rudbeckia hirta L. NABPF 18 June 1997 Blackeyed Susan common, disturbed area 70 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Johnson, F.L. Rudbeckia missouriensis Engelm. NPF 31 July 1997 Missouri coneflower common, prairie Rudbeckia triloba L. NPF 31 July 1997 Browneyed Susan infrequent, floodplain forest Senecio plattensis Nuttall NPF 28 March 1997 Prairie groundsel common, Cross timbers, floodplain Silphium laciniatum L. NPF 31 July 1997 Compass plant common, prairie Solidago arguta Ait. NPF 31 July 1997 Boott’s woodland goldenrod infrequent, floodplain forest Solidago canadensis L. var. Gilvocanescens Rydb. NPF 8 October 1997 Prairie goldenrod common, rocky upland Solidago nemoralis Ait. NPF 10 Sept. 1997 Oldfield goldenrod infrequent, prairie Solidago rigida L. NPF 19 Sept. 1997 Stiff prairie goldenrod infrequent, prairie Thelesperma filifolia (Hook.) Gray NPF 29 May 1997 Greenthread common, prairie Verbesina virginica L. NPF 28 Sept 1999 Virginia crownbeard uncommon, floodplain forest Vernonia baldwinii Torrey NPF 31 July 1997 Western ironweed common, prairie Xanthium strumarium L. NAF 10 Sept. 1997 Cocklebur common, pond margin Berberidaceae (barberry family) Podophyllum peltatum L. NPF 10 Apr 1996, 15 April 1997 Mayapple abundant, Cross timbers Boraginaceae (borage, heliotrope family) Heliotropium tenellum (Nutt.) Torrey NAF 18 June, 31 July 1997 Pasture heliotrope infrequent, prairie, rocky upland Lithospermum incisum Lehm. NPF 28 March, 18 June 1997 Narrowleaf puccoon infrequent, rocky upland Myosotis macrosperma Engelmann NPF 30 April 1997 Spring forget-me-not common, disturbed area Myositis verna Nuttall NAF 15 April 1997 Early scorpiongrass common, disturbed area Brassicaceae (mustard family) Capsella bursa-pastoris (L.) Medik IAF 30 April 1997 Shepherd’s purse infrequent, distubed area Draba brachycarpa Nutt. Ex T.&G. NAF 28 Feb. 1997 Whitlow infrequent, mowed meadow Draba cuneifolia Nutt. Ex T. & G. Whitlow NAF 28 March 1997 infrequent, disturbed area Lepidium densiflorum Schrad. IAF 30 April 1997 Peppergrass common, disturbed area Lepidium virginicum L. NAF 30 April 1997 Virginia peppergrass infrequent, disturbed area Lesquerella gracilis (Hook.) Wats. var. repandum (Nutt.) Payson NAF 15 April 1997 Spreading bladderpod common, disturbed area Lesquerella ovalifolia Rydb. var. alba Goodman NPF 15 April 1997 Oval-leaf or white bladderpod abundant, rocky upland Rorippa nasturtium-aquaticum (L.) Hayek IAE 15 April 1997 Water cress common, aquatic Cactaceae (Cactus family) Echinocereus riechenbachii (Tersch.) Hogue NP$ 12 May 2001 Lace or pincushion cactus infrequent, rocky upland Opuntia engelmannii Salm-Dyck var. lindheimeri (Engl.) Parfit. & Pink. NP$ 11 June 1997 Texas or limestone prickly-pear infrequent, rocky upland Callitrichaceae (waterweed family) Callitriche heterophylla Pursh NZF 14 May 1997 Oklahoma Native Plant Record 71 Volume 2, Number 1, December 2002 Johnson, F.L. Water starwort common, aquatic Campanulaceae (bellflower family) Lobelia appendiculata DC. NPF 29 May 1997 Earflower lobelia common, prairie Lobelia siphilitica L. var. ludoviciana DC. NPF 10 Sept. 1997 Blue cardinal flower uncommon, seeps Triodanis leptocarpa (Nutt.) Niew. NAF 29 May 1997 Western Venus’ looking-glass common, prairie Triodanis perfoliata (L.) Niew. NAF 14 May 1997 Venus’ looking glass infrequent, prairie Caprifoliaceae (honeysuckle family) Symphoricarpos orbiculatus Moensch NS (in fruit) 10 Sept. 1997 Coralberry, buckbrush common, cross timbers Viburnum rufidulum Raf. NST 15 April 1997 Rusty blackhaw infrequent, cross timbers Caryophyllaceae (Pink, carnation family) Arenaria patula Michaux NAF 28 March, 30 April 1997 Sandwort infrequent, rocky upland Arenaria stricta Michx, var. texana Rydberg NAF 30 April, 29 May 1997 rock sandwort common, rocky upland Paronychia jamesii T. & G. NPF 20 August 1997 James’ nailwort common, rocky upland Celastraceae (bittersweet family) Celastrus scandens L. NWV (in fruit) 8 October 1997 Climbing bittersweet infrequent, disturbed area Cistaceae (pinweed family) Lechea villosa Raf. NPF 20 August 1997 Pinweed infrequent, disturbed area Clusiaceae (St. John’s Wort family) Hypericum drummondii T. & G. NAF 20 August 1997 Nits-and-lice infrequent, rocky upland Hypericum sphaerocarpon Michx. NPF 18 June 1997 Roundfruit St. John’s Wort infrequent, rocky upland Commelinaceae (spiderwort & dayflower family) Tradescantia ohiensis Raf. NPF 14 May 1997 Ohio spiderwort infrequent, prairie Tradescantia tharpii Anders & Woods NPF 30 April 1997 Tharp’s spiderwort common, rocky upland Convolvulaceae (morning glory family) Convolvulus equitans Benth. NAVF 29 May, 18 June 1997 Texas bindweed common, disturbed area Evolvulus nuttallianus R. & S. NPF 30 April 1997 Hairy evolvulus infrequent, rocky upland Cornaceae (dogwood family) Cornus drummondii Meyer NS 14 May 1997 Rough-leaf dogwood common, rocky upland Cornus florida L. NT 15 April 1997 Flowering dogwood infrequent, cross-timbers Crassulaceae (stonecrop family) Sedum nuttallianum Rf. NAF 29 May 1997 Yellow stonecrop infrequent, seep Sedum pulchellum Michaux NAF 14 May 1997 Pink stonecrop, widow’s cross infrequent, wet meadow Cyperaceae (sedge family) Carex festucacea Schkuhr NPG 30 April 1997 None infrequent, disturbed area Carex flaccosperma Dewey NPG None infrequent, floodplain forest Carex frankii Kunth NPG 29 May to 31 July 1997 None common, floodplain forest Carex gravida Bailey NPG 15 April 1997 72 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Johnson, F.L. Heavy sedge common, rocky upland Carex hirsutella Mackenzie NPG Hairy sedge common, disturbed area Carex leavenworthii Dewey NPG 30 April 1997 None infrequent, disturbed area Carex retroflexa Muhlenberg NPG None common, rocky ground Carex vulpinoidea Michaux NPG 30 April 1997 None infrequent, aquatic Cyperus filiculmis Vahl NPG 18 June 1997 Slender flatsedge common, prairie Cyperus odoratus L. NAG 10 Sept. 1997 Fragrant flatsedge infrequent, pond margin Cyperus ovularis (Michaux) Torrey NPG 18 June 1997 Globe flatsedge common, disturbed area Eleocharis compressa Sulliv. NPG 14 May 1997 Weakstem spikerush common, riparian Eleocharis engelmannii Steudel NPG 10 Sept 1997 Engelmann’s spikerush common, pond margin Eleocharis parvula (R.&S.) Link NAG 10 Sept. 1997 Dwarf spikesedge common, pond margin Fimbristylis puberula (Michx) Vahl NPG 30 April 1997 None infrequent, rocky upland Fuirena simplex Vahl NPG 31 July 1997 Umbrella sedge common, seep Rhynchospora nivea Boeckl. NPG 8 Oct 1998 White-top sedge infrequent, seep Scirpus koilolepis (Steud.) Gleason NAG None common, pond margin Scirpus pendulus Muhelenberg NPG 31 July 1997 None common, floodplain forest Scleria pauciflora Michx. Ex Willd. NPG 31 July 1997 Few-flower nutrush infrequent, prairie Ebenaceae (Ebony family) Diospyros virginiana L. NT 29 May 1997 Persimmon infrequent, crosstimbers Euphorbiaceae (spurge family) Acalypha gracilens Gray NAF 18 July 1997 Three-seeded mercury infrequent, rocky upland Argythamnia mercurialina Muell. NPF 11 June 1997 Wild mercury infrequent, forest margin Chamaesyce missurica (Raf.) Shinners NAF 20 August 1997 [Euphorbia missurica Lag.] Missouri spurge common, disturbed areas Chamaesyce nutans Lag.) Small NAF 20 Aug. – 10 Sept. 1997 [Euphorbia nutans Lag.] Eyebane common, disturbed areas Croton capitatus Michaux NAF 10 Sept. 1997 Wooly croton common, disturbed area Croton monanthogynus Michaux NAF 18 July – 20 Aug. 1997 One-seed croton common, disturbed area Euphorbia corollata L. NAF 10 Sept. 1997 Flowering spurge infrequent, prairie Euphorbia dentata Michaux NAF 18 July 1997 Toothed spurge infrequent, rocky upland Euphorbia hexagona Nuttall NAF 8 October 1996 6-angle spurge infrequent, rocky upland Euphorbia spathulata Lam. NAF 30 April 1997 Warty spurge infrequent, rocky upland Phyllanthus polygonoides Nuttall NPF 29 May 1997 Knotweed leaf-flower common, prairie Stillingia sylvatica Graden ex L. NPF 29 May 1997 Queen’s delight infrequent, prairie Oklahoma Native Plant Record 73 Volume 2, Number 1, December 2002 Johnson, F.L. Tragia ramosa Torrey NPF 29 May 1997 Nettle-leaf, noseburn common, prairie Fabaceae (legume, bean and pea family) [includes Caesalpinaceae and Mimosaceae] Albizia julibrissin Durazz. IT 18 July 1997 Mimosa infrequent, disturbed area Amorpha canescens Pursh NS 31 July 1997 Leadplant infrequent, prairie Amorpha fruticosa L. NS 10 Sept. 1997 False indigo common, pond margin Astragalus nuttallianus DC. NAF 30 April 1997 Nuttall’s milkvetch infrequent, rocky upland Baptisia australis (L.) R. Br. NPF 30 April 1997 Blue wild indigo common, rocky upland Cercis canadensis L. NT 13-28 March, 1997 Redbud common, Crosstimbers, rocky upland Chamaecrista fasciculata (Michx.) NAF 18 July 1997 Greene Partridge pea frequent, disturbed area, prairie Dalea aurea Nuttall ex Pursh NPF 18 July 1997 Golden prairie clover infrequent, rocky upland Dalea candida Willdenow NPF 18 June 1997 White prairie clover common, rocky upland Dalea enneandra Nuttall NPF 31 July 1997 Nine-anther prairieclover infrequent, prairie Dalea purpurea Vent. NPF 18 June 1997 Purple prairieclover common, rocky upland Desmanthus illinoinsis (Michx.) NPF 18 July 1997 MacM. Bundleflower common, disturbed area Desmodium glutinosum (Muhl.) Wood NPF 18 June 1997 Large-flowered tickclover common, floodplain forest Desmodium paniculatum (L.) DC. NPF 10 Sept. 1997 Panicled tickclover common, floodplain forest Gleditsia triacanthos L. NT 31 July 1997 Honey locust common, mowed meadow Lespedeza cuneata G. Don IPF 20 August 1997 Sericea lespedeza common, disturbed area Lespedeza stipulacea Maxim. IAF 20 August, 1997 Korean clover common, disturbed area Lespedeza stuevei Nuttall NPF 10 Sept 1997 Stueve’s lespedeza common, prairie Mimosa nuttallii (DC.) B.L. Turner NWV 14 May 1997 [Schrankia nuttalii DC.] Sensitive brier common, rocky upland Pediomelum cuspidatum (Pursh) NPF 14 May 1997 Rydb. [Psoralea cuspidata Pursh] Tallbread scurfpea infrequent, rocky upland Pediomelum esculentum (Pursh) NPF 14 May 1997 Rydb. [Psoralea esculenta Pursh] Prairie turnip common, rocky upland Psoralidum tenuifolium (Pursh) Rydb. NPF 14 – 29 May 1997 [Psoralea tenuifolia Pursh] Wild alfalfa common, prairie, rocky upland Stylosanthes biflora (L.) B.S.P. NPF 31 July 1997 Pencil flower infrequent, prairie Tephrosia virginica (L.) Pers. NPF 31 July 1997 Goat’s rue infrequent, prairie Trifolium dubium Sibth. IAF 15 – 30 April 1997 Small hop-clover common, mowed meadow Trifolim repens L. IPF 30 April 1997 White clover common, disturbed area Vicia sativa L. IAF 15 April 1997 Common vetch common, disturbed area 74 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Johnson, F.L. Fagaceae (oak, beech, chestnut family) Quercus macrocarpa Michaux NT 18 July 1997 Burr oak infrequent, floodplain forest Quercus marilandica Muench. NT 18 July 1997 Blackjack common, crosstimbers Quercus muehlenbergii Engelm. NT 14 May 1997 Chinquapin oak common, floodplain forest Quercus stellata Wang. NT 18 July 1997 Post oak dominant, crosstimbers Quercus velutina Lam. NT 8 October 1997 Black oak infrequent, crosstimbers Gentianaceae (gentian family) Sabatia campestris Nuttall NAF 11 June 1997 Prairie rose, pink gentian infrequent, prairie Geraniaceae (geranium family) Geranium carolinianum L. NAF 30 April 1997 Carolina crane’sbill infrequent, disturbed area Grossulariaceae (gooseberry family) Ribes aureum Pursh NS 30 April 1997 Golden or buffalo current infrequent, rocky upland Iridaceae (iris family) Nemastylis geminiflora Nuttall NPF 30 April 1997 Prairie iris Sisyrinchium angustifolium Miller NPF 30 April 1997 Blue-eyed grass infrequent, mowed meadow Sisyrinchium campestre Bicknell NPF 15 April 1997 Prairie blue-eyed grass infrequent, rocky upland Juglandaceae (walnuts, hickory family) Carya cordiformis (Wang.) Koch NT 18 July 1997 Bitternut hickory infrequent, floodplain forest Carya illinoensis (Wang.) Koch NT 31 July 1997 Pecan common, ravine Carya texana Buckley NT 18 July 1997 Black hickory infrequent, crosstimbers Juglans nigra L. NT 31 July 1997 Black walnut common, floodplain forest Juncaceae (rush family) Juncus brachycarpus Engelmann NPG 18 June 1997 Whiteroot rush common, disturbed area Juncus marginatus Rostk. NPG 18 June 1997 Grassleaf rush infrequent, prairie Juncus tenuis Willdenow NPG 18 June 1997 Slender rush common, disturbed area Juncus validus Coville NPG 31 July 1997 Bulrush common, seep Krameriaceae (ratany family) Krameria lanceolata Torrey NPF 29 May 1997 Trailing ratany infrequent, prairie Lamiaceae (mint family) Hedeoma drummondii Benth. NPF 20 august 1997 False pennyroyal common, disturbed area, rocky upland Mondarda citriodora Cerv. Ex Lagasca NAF 29 May 1997 Lemon beebalm infrequent, prairie Monarda fistulosa L. NPF 22 June 1997 Wild bergamot infrequent, mowed meadow Prunella vulgaris L. NPF 29 May – 31 July 1997 Heal-all common, floodplain forest Pycnanthemum tenuifolium Schrad. NPF 10 Sept. 1997 Narrowleaf mountain mint common, prairie Salvia azurea Lam. NPF 10 Sept. 1997 Blue sage common, prairie Satureja calamintha (L.) Scheele NPF 14-29 May 1997 Ozark savory abundant, prairie, rocky upland Oklahoma Native Plant Record 75 Volume 2, Number 1, December 2002 Johnson, F.L. Scutellaria parvula Michaux NPF 30 April – 29 May 1997 Small skullcap infrequent, prairie, rocky upland Teucrium canadense L. NPF 31 July 1997 American germander common, seep Trichostema brachiatum L. NAF 20 August 1997 Blue curls infrequent, rocky upland Lemnaceae (duckweed family) Wolffia columbiana Karst NPL 8 October 1997 Watermeal common, aquatic Liliaceae (lily family) Allium canadense L. NPF 30 April 1997 Wild onion/garlic common, rocky upland Allium drummondii Regel NPF 15 April 1997 Drummond’s wild onion common, rocky upland Allium stellatum Nuttal NPF 8 October 1997 Autumn wild onion common, rocky upland Androstephium coerulium (Scheele) Torr. NPF 28 March 1997 Blue funnel lily common, rocky upland Camassia scilloides (Raf.) Cory NPF 14 May 1997 Eastern camass infrequent, rocky upland Cooperia drummondii Herb. NPF 10 Sept 1997 Rain lily infrequent, disturbed area Erythronium albidum Nuttall NPF 13 March 1997 White dogtooth violet, fawn lily common, rocky upland Lythraceae (loosestrife family) Lythrum alatum Pursh var. lanceolatum (Ell.) T. & G NPF 31 July – 10 Sept. 1997 Tall loosestrife infrequent, wet prairie Rotala ramosior (L.) Koehne var. interior Fern. & Grisc. NAF 10 Sept. 1997 Toothcup common, pond margin Malvaceae (mallow, hibiscus, cotton family) Callirhoe alcaeoides (Michx.) Gray NPF 30 April 1997 Pink poppy mallow infrequent, disturbed area Callirhoe digitata Nuttall var. stipulata Waterfall ssp glabrescens (Kuntze) Raven NPF 29 May 1997 Fringed poppy mallow infrequent, prairie Callirhoe involucrata (Nutt.) Gray NPF 29 May 1997 Winecup, cowboy rose common, prairie Sida procumbens Sw. NPF 31 July – 10 Sept. 1997 Spreading sida common, rocky upland Menispermaceae (moonseed family) Cocculus carolinus (L.) DC. NWV 29 May 1997 Carolina snailseed infrequent, floodplain forest Moraceae (mulberry, fig family) Maclura pomifera (Raf.) Schneider NT 14 May 1997 Bois d’arc, osage orange common, rocky upland Morus rubra L. NT 31 July 1997 Red mulberry common, floodplain forest Oleaceae (olive, privet, ash family) Forestiera pubescens Nuttall NS 28 Feb – 13 Mar 1997 Elbowbush abundant, rocky upland Fraxinus americana L., var. Americana NT 30 April 1997 White ash infrequent, rocky upland Fraxinus pennsylvanica Marsh. NT 31 July 1997 Green ash common, floodplain forest Ligustrum sinense Lour. IS 29 May 1997 Privet infrequent, disturbed area Onagraceae (evening primrose family) Calylophus serrulatus (Nutt.) Raven NPF 14, 29 May 1997 Sundrops, plains yellow primrose common, prairie, rocky upland Gaura longiflora Spach. NAF 20 Aug, 10 Sept. 1997 Common, prairie Gaura parviflora Dougl. Ex Lehm 76 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Johnson, F.L. NAF 31 July 1997 Velvety gaura common, disturbed area Gaura suffulta Engelm. Ex Gray NAF 14 May – 18 Jun 1997 Beeblossom common, rocky upland, prairie Ludwigia peploides (H.B.K.) Raven NAE 31 July – 10 Sept. 1997 Water primrose, water pennies common pond margin Oenothera linifolia Nuttall NAF 14 May 1997 Flax-leaf evening primrose common, rocky upland, prairie Oenothera macrocarpa Nuttall NPF 14 May 1997 Large-fruited evening primrose infrequent, rocky upland Oenothera speciosa Nuttall NPF 29 May 1997 Showy evening primrose common, prairie Stenosiphon linifolius (Nutt.) Heynh. NPF 10 Sept. 1997 Stenosiphon common, prairie Ophioglossaceae (grape fern, adder’s-tongue family) Botrychium virginicum (L.) Sw. NPF 15 April – 29 May 1997 Virginia grape-fern common, crosstimbers Ophioglossum engelmannii Prandtl NPF 30 April 1997 Limestone adder’s tongue common, rocky upland Orchidaceae (orchid family) Spiranthes cernua (L.) L.C. Rich. NPF 8 October 1997 Nodding ladies’-tresses infrequent, rocky upland Spiranthes lacera Raffinesque NPF 10 Sept. 1997 Slender ladies’-tresses infrequent, prairie Spiranthes magnicamporum Sheviak NPF Great plains ladies’-tresses infrequent, seep Spiranthes ovalis Lindl. NPF October ladies’-tresses infrequent, prairie Orobanchaceae (broomrape family) Orobanche uniflora L. NPF(one, no voucher) 29 April 2000 One-flowered broomrape infrequent, floodplain forest Oxalidaceae (oxalis family) Oxalis corniculata L. NPF 30 April 1997 Yellow wood-sorrel common, disturbed area Oxalis volacea L. NPF 15 April 1997 Violet wood-sorrel common, floodplain forest Passifloraceae (passionflower family) Passiflora incarnata L. NPF 2 August 2001 Blue passionflower infrequent, prairie Passiflora lutea L. NPF (not yet found, but predicted) Yellow passionvine infrequent, floodplain forest Phytolaccaceae (pokeweed family) Phytolacca americana L. NPF 18 June 1997 Pokeweed infrequent, disturbed area Plantaginaceae (plantain family) Plantago aristata Michaux NAF 29 May 1997 Bottlebrush plantain infrequent, prairie Plantago lanceolata L. IPF 29 May 1997 English plantain infrequent, prairie Plantago patagonica Jacq. NAF 29 May 1997 Wooly plantain infrequent, prairie Plantago pusilla Nuttall NAF April 1999 Dwarf plantain infrequent, disturbed area Plantago rugelii Dcne. NPF 16 July 1993 Blackseed plantain infrequent, prairie Plantago virginica L. NAF 29 May 1997 Paleseed plantain common, prairie Platanaceae (sycamore family) Platanus occidentalis L. NT 31 July 1997 Sycamore common, floodplain forest Poaceae (grass family) Agrostis hyemalis (Walt.) B.S.P. NPG 29 May 1997 Ticklegrass Oklahoma Native Plant Record 77 Volume 2, Number 1, December 2002 Johnson, F.L. common, prairie Aira elegantissima Schur. IAG 14 May 1997 Hairgrass infrequent, rocky upland Andropogon gerardii Vitman NPG 20 August 1997 Big bluestem infrequent, rocky upland Aristida dichotoma Michaux NAG 8 October 1997 Churchmouse threeawn infrequent, disturbed area Aristida oligantha Michaux NAG 10 Sept. 1997 Oldfield threeawn common, disturbed area Aristida purpurea Nuttall NPG 14 May 1997 Purple threeawn common, rocky upland Bothriochloa saccharoides (Sw) Rydb NPG 18 June 1997 Silvertop bluestem common, disturbed area Buteloua curtipendula (Michx.) Torrey NPG 18 July 1997 Sideoats grama infrequent, rocky upland Bouteloua hirsuta Lag. NPG 20 August 1997 Hairy grama common, rocky upland Bouteloua pectinata Featherly NPG 20 August 1997 Tall grama common, rocky upland, prairie Bromus catharticus Vahl IAG 30 April 1997 Rescue grass infrequent, rocky upland, disturbed area Bromus pubescens Muhl. Ex Willd. NPG 14 May 1997 Canada brome common, floodplain forest Buchloe dactyloides (Nutt.) Engelm NPG 14 May 1997 Buffalo grass infrequent, prairie Chasmanthium latifolium (Michx) Yates NAG 18 July – 10 Sept. 1997 Inland sea oats common, floodplain forest, Crosstimbers Coelorachis cylindrica (Michx.) Nash NPG 29 May, 18 June 1997 Mousetail common, disturbed area, prairie Cynodon dactylon (L.) Pers. IPG 29 May 1997 Bermuda grass common, disturbed area Dactylis glomerata L. IPG 30 April - 29 May 1997 Orchard grass common, disturbed area Dicanthelium acuminatum (Sw.) Gould & Clark NPG 29 May & 10 Sept. 1997 Early panicum common, disturbed area, prairie Dicanthelium boscii (Poir.) Gould & Clark NPG 14 May 1997 Bosc’s panic common, floodplain forest Dicanthelium oligosanthes Schult.) Gould var. oligosanthes NPG 14 – 29 May 1997 Small panicgrass infrequent, disturbed area, rocky upland, prairie Dicanthelium sphaerocarpon (Ell.) Gould NPG 14 May 1997 Leafy panicum common, prairie Digitata cognata (Schult.) Pilger NPG 10 Sept. 1997 [Leptoloma cognatum (Schult.) Chase] Fall witchgrass common, floodplain forest Echinochloa crusgalli (L.) Beauv. IAG 20 Aug – 10 Sept. 1997 Barnyard grass infrequent, pond margin Eleusine indica (L.) Gaertn. IAG 18 July 1997 Goosegrass infrequent, disturbed area Elymus virginicus L. NPG 18 June 1997 Virginia wild rye common, disturbed area Erioneuron pilosum (Buckl.) Nash NPG 30 April 1997 Hairy tridens infrequent, disturbed area Festuca arundinacea Schreb. IPG 30 April – 14 May 1997 Meadow fescue infrequent, disturbed area, floodplain forest Hordeum pusillum Nuttall NAG 29 May 1997 Little barley infrequent, disturbed area Koelaria macrantha (Ledeb.) J.A. Schultes NPG 14 May 1997 Junegrass infrequent, rocky upland Leptochloa fusca (L.) Kunth NPG 10 Sept. 1997 Bearded sprangletop infrequent, pond margin Lolium perenne L. IPG 29 May 1997 Perennial ryegrass 78 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Johnson, F.L. common, prairie Muhelbergia reverchonii Vasey & Scrib. NPG 20 Aug 1999 Seep muhly infrequent, rocky upland Muhlenbergia schreberei J.F. Gmel. NPG Nimblewill infrequent, floodplain forest Nassella leucotricha (T.& R.) Barkw. NPG 14 May 1997 [Stipa leucotrichaTrin & Rupr.] Texas wintergrass common, prairie Panicum anceps Michaux NPG 31 July 1997 Beaked panic common, floodplain forest Panicum capillare L. NAG 18 July 1997 Witchgrass infrequent, rocky upland Panicum virgatum L. NPG 10 Sept. 1997 Switchgrass common, prairie Paspalum dilatatum Poir. IPG 18 June 1997 Dallis grass common, disturbed area Phalaris caroliniana Walt. NAG 14 May 1997 Maygrass infrequent, prairie Poa annua L. IAG 15 April 1997 Annual bluegrass common, disturbed area Poa pratensis L. IPG 30 April 1997 Kentucky bluegrass common, disturbed area Schizachyrium scoparium (Michx.) Nash NPG 20 August 1997 Little bluestem common, rocky upland Setaria geniculata (Lam.) Beauv. NPG 18 June – 30 July 1997 Knotroot foxtail common, disturbed area Sorghastrum nutans (L.) Nash NPG 10 Sept. 1999 Indiangrass infrequent, prairie Sorghum halapense (L.) Pers. IPG 18 June 1997 Johnson grass common, disturbed area Sporobolus compositus (Poir) Merr. [Sporobolus asper (Michx.) Kunth NAG 28 Sept. 1999 Dropseed infrequent, disturbed area Sporobolus vaginiflorus (T.ex G.) Wood NAG 10 Sept. 1997 Poverty dropseed common, disturbed area Tridens flavus (L.) Hitchcock NPG 31 July 1997 Purpletop common, floodplain forest Tridens strictus (Nutt.) Nash NPG 10 Sept. 1997 Longspike tridens infrequent, pond margin Tripsacum dactyloides L. NPG 18 June 1997 Gama grass infrequent, prairie Vulpia octoflora (Walt.) Rydb. IAG 8 October 1997 Poverty grass common, disturbed area Polygalaceae (milkwort family) Polygala incarnata L. NAF 31 July 1997 Pink milkwort infrequent, prairie Polygonaceae (buckwheat, knotweed family) Eriogonum longifolium Nuttall NPF 20 August – 8 Oct. 1997 Longleaf buckwheat common, rocky upland, prairie Polygonum hydropiperoides Michx. NPF 8 October 1997 Mild water-pepper infrequent, pond margin Polygonum lapathifolium L. NAF 10 Sept. 1997 Pale smartweed infrequent, pond margin Polygonum persicaria L. NAF 20 August 1997 Lady’s thumb common, disturbed area Polygonum punctatum L. NAF 20 August 1997 Water smartweed infrequent, pond margin Rumex crispus L. IPF 29 May 1997 Curly dock common, riparian Rumex hastatulus Baldw. NAF 18 June 1997 Heartwing sorrel common, disturbed area Portulacaceae (portulaca family) Claytonia virginica L. NPF 28 March 1997 Spring beauty common, rocky upland Oklahoma Native Plant Record 79 Volume 2, Number 1, December 2002 Johnson, F.L. Primulaceae (primrose family) Dodecatheon meadia L. var. brachycarpa (Small) Fassett NPF 30 April 1997, 16 July 1993 Shooting star common, rocky upland Samolus parviflorus Raf NAEF 14 May 1997 Smallflower brookweed common, aquatic Pteridaceae (bracken fern) Pellaea atropurpurea (L.) Link. NPF 29 May 1997 Purple cliff brake Ranunculaceae (buttercup, larkspur family) Anemone berlandieri Pritz NPF 28 March, 14 May 1997 Ten-petal windflower infrequent, rocky upland, prairie Delphinium carolinianum Walt NPF 14 - 29 May 1997 Ssp. Virescens (Nutt.) Brooks Plains larkspur common, prairie Ranunculus hispidus Michaux NPF 30 April 1997 Bristly buttercup infrequent, disturbed area Rhamnaceae (buckthorn family) Berchemia scandens (Hill) K. Koch NWV 14 May 1997 Rattan vine common, disturbed area, forest edges Ceanothus herbaceus Raffinesque NS 30 April 1997 New Jersey tea infrequent, rocky upland Frangula caroliniana (Walt.) Gray NT 29 May 1997 [Rhamnus caroliniana Walt.] Indian cherry common, rocky upland, Crosstimbers, forest Rosaceae (rose family) Agrimonia pubescens Wallr. NPF 20 August 1997 Downy agrimony infrequent, floodplain forest Agrimonia rostellata Wallr. NPF 28 Sept. 1999 Beaked agrimony infrequent, floodplain forest Crataegus crus-gallii L. NST 30 April 1999 Cockspur hawthorn infrequent, rocky upland Crataetus mollis (T.& G.) Scheele NT 10 Sept. 1997 Downy haw infrequent, rocky upland Fragaria virginica P. Mill. var. illinoensis (Prince) Gray NPF 15 April 1997 Wild strawberry infrequent, crosstimbers Geum canadense Jacq. NPF 29 May 1997 White avens infrequent, crosstimbers Prunus angustifolia Marsh. NS 13 March 1997 Chickasaw plum infrequent, rocky upland Prunus mexicana Wats. NT 13 March 1997 Mexican plum common, crosstimbers Prunus serotina Ehrh. NT 18 July 1997 Wild cherry infrequent, crosstimbers Pyrus communis L. IT 28 March 1997 Common pear infrequent, rocky upland Rosa foliolosa Nuttall NPF 18 June 1997 Prairie rose infrequent, rocky upland Rosa carolina L. NS 14 May 1997 Swamp rose common, pond margins Rosa setigera Michaux NS 18 June 1997 Climbing prairie rose common, rocky upland Rubus flagellaris Willdenow NS 15 April 1997 Northern dewberry common, disturbed area Rubiaceae (madder, coffee family) Cephalanthus occidentalis L. NS 31 July 1997 Buttonbush infrequent, seep Diodia teres Walt. NAF 20 August 1997 Poor-Joe infrequent, rocky upland Galium circaezans Michaux NPF 14 – 29 May 1997 Weedy bedstraw infrequent, floodplain forest Galium virgatum Nuttall NAF 14 May 1997 Southwest bedstraw infrequent, rocky upland 80 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Johnson, F.L. Hedyotis crassifolia Raffinesque NAF 13 March 1997 Prairie bluets, Quaker ladies common, rocky upland Rutaceae (citrus family) Ptelea trifoliata L. NST(fruit) 10 Sept. 1997 Hop tree, wafer ash infrequent, rocky upland Zanthoxylum americanum Miller NS 30 April 1997 Prickly ash common, crosstimbers Zanthoxylum clava-herculis L. NT (no flrs) 10 Sept. 1997 Hercules’ club infrequent, old homesite Salicaceae (willow, cottonwood family) Populus nigra L. NT Not collected: need specimen Cottonwood infrequent, floodplain forest Salix nigra Marsh NT 30 April 1997 Black willow infrequent, riparian, pond margin Sapotaceae (sapodilla family) Sideroxylon lanuginosum Michaux NT 18 July 1997 [Bumelia lanuginosa (Mix.) Pers Chittamwood common, rocky upland Scrophulariaceae (penstemon family) Agalinis fasciculata (Ell.) Raf. NAF 10 Sept. 1997 Beach gerardia abundant, disturbed area Castilleja indivisa Engelmann NAF 15 April 1997 Indian paintbrush common, rocky upland Nuttallanthus texanus (Scheele) D.A.Sutton NAF 15 April 1997 [Linaria texana Scheele] Texas toadflax common, rocky upland Penstemon cobaea Nuttall NPF 14 May 1997 Large beardtongue common, rocky upland Penstemon oklahomensis Pennell NPF 14 May 1997 Oklahoma penstemon infrequent, prairie Veronica arvensis L. IAF 15 April 1997 Common speedwell infrequent, mowed meadow Veronica peregrina L. NAF 15 April 1997 Purslane speedwell common, mowed meadow Smilacaceae (greenbrier family) Smilax bona-nox L. NWV 14 May 1997 Greenbrier, cat’s claw common, rocky upland Smilax rotundifolia L. NWV 30 April 1997 Greenbrier infrequent, floodplain forest Solanaceae (nightshade, potato family) Physalis angulata L. NAF 18 June 1997 Cutleaf ground cherry common, disturbed area Solanum carolinense L. NPF 18 June 1997 Carolina horsenettle infrequent, disturbed area Solanum dimidiatum Raffinesque NAF 18 June 1997 Horsenettle common, disturbed area Solanum rostratum Dunal NAF 18 June 1997 Buffalo bur infrequent, disturbed area Typhaceae (cattail family) Typha angustifolia L. NPEF 18 June 1997 Narrowleaf cattail infrequent, aquatic Ulmaceae (elm, hackberry family) Celtis laevigata Willdenow NT 31 July 1997 Sugarberry common, floodplain forest Celtis reticulata Torrey NT 18 July 1997 Roughleaf hackberry common, rocky upland Ulmus alata Michaux NT 28 Feb 1997 Winged elm common, rocky upland, crosstimbers Ulmus americana L. NT 13 March 1997 American elm infrequent, floodplain forest, rocky upland Ulmus rubra Muhlenberg NT 28 Feb 1997 Slippery elm common, crosstimbers Oklahoma Native Plant Record 81 Volume 2, Number 1, December 2002 Johnson, F.L. Urticaceae (nettle family) Parietaria pennsylvanica Muhl. NAF 29 May 1997 Pennsylvania pellitory common, floodplain forest Valerianaceae (valerian family) Valerianella radiata Dufr. NAF 30 April 1997 Beaked cornsalad infrequent, rocky upland Verbenaceae (vervain family)\ Glandularia bipinnatifida Nuttall NAF 15 April 1997 [Verbena bipinnatifida Nuttall] Sweet William common, rocky upland Verbena stricta Ventenat NPF 18 June – 31 July 1997 Wooly vervain common, disturbed area Verbena urticifolia L. NAF 18 June 1997 White vervain infrequent, floodplain forest Violaceae (violet family) Viola bicolor Pursh NAF 13 March 1997 Johnny-jump-up common, mowed meadow Viola sororia Willdenow NPF 28 March 1997 Butterfly violet common, floodplain forest Viscaceae (mistletoe family) Phoradendron leucarpum (Raf.) Reveal & Johnston NP%@ Vitaceae (grape family) Ampelopsis cordata Michaux NWV 18 July 1997 Raccoon grape infrequent, crosstimbers pond margin, disturbed area Cissus incisa (Nutt.) Des Moulins NWV 29 May 1997 Possum grape infrequent, rocky upland Parthenocissus quinquefolia (L.) Planch NWV 31 July 1997 Virginia creeper common, ravine, floodplain forest, crosstimbers Vitis aestivalis Michaux NWV 18 July 1997 Pigeon grape infrequent, disturbed S 13 March 1997 Mistletoe infrequent, crosstimbers 82 Oklahoma Native Plant Record Volume 2, Number 1, December 2002 Editorial Water, Soil, and Plant Diversity in Oklahoma Sheila Strawn In 1963 John Shed and William Penfound did a study on legume distribution. They found a general relationship between soils and vegetation types. Sand deposits generally became forests while limestone and clay deposits generally became prairie (Proc. Okla. Acad. Sci. 44:2-6). But soil-to-plant relationships are not always that simple. Tropographic effects such as slope and solar and wind exposure as well as precipitation rates can override the characteristics of the various soils making it difficult to determine specific soil types based on vegetation. In 1964 Dr. Paul Buck wrote "Relationships of the Woody Vegetation of the Wichita Mountains Wildlife Refuge to Geological Formations and Soil Types (Ecology 45:2). He had found that when it rains, soil type and topographic features, together, control the movement of water. Granite formations at the Refuge erode to make a cobbly soil at the foot of the mountains that drains quickly, making it difficult for some plants to become established. Different amounts and patterns of precipitation can also interact with soil type producing gradients of moisture availability for plants with different moisture requirements. Some species can live only at certain positions across the moisture gradient. Others are more tolerant of variable moisture availability. Sugar Maple is able to dominate some of the more mesic, cobbly patches near streams, while Black-jack Oak dominates the less mesic, cobbly upland patches in this newly formed soil. In places like the Carolinas bountiful precipitation on limestone soils can actually make soil very poor in nutrients. Organic acids from the abundant vegetation react with calcium, sodium, and magnesium ions in the soil The abundant rainfall then leaches them from the soil and washes them into the groundwater below the reach of roots. However, this is not usually the case in Oklahoma. Oklahoma actually loses more water to evaporation than it gets in precipitation. Rivers and groundwater from the Rocky Mountains make up the difference. Here, precipitation is not sufficient to leach all the minerals from our limestone soils. Dense stands of forbs and grasses grow in the Tallgrass Prairie Preserve where these mineral-rich formations are not leached by excess precipitation. Southwest of the Refuge the precipitation rate is so low that much of the water evaporates before it gets very deep into the soil. In those soils, the calcium ions accumulate at the lowest level of water penetration, sometimes, just inches from the surface and form a calcium carbonate. Known as "caliche", this layer restricts root growth, so fewer species grow on caliche. Gypswn, mostly calcium sulfate, was formed long ago in a similar manner when the seawater evaporated from the plains. In Oklahoma Mesquite and mixed grass are associated with “gyp soil”. Soil and water relations alone do not explain why plants grow where they do in Oklahoma. Other variables like humus, micro-organisms, nematodes, grazing animals, fire, ice, windstorms, and human influence all work on a smaller scale and are responsible for much of the physical heterogeneity and resultant biodiversity. This small-scale heterogeneity is what actually accounts for our having more species than other continental United States, except California, Florida, and Texas. Ironically, plant diversity in Oklahoma has been woefully understudied. With the promise of improved medical treatments based on genetic engineering, we need to know where species are in Oklahoma that might be beneficial. We are, therefore, obliged to preserve as much of our diversity as we can. But we will need to save it on the landscape level, because it is our diverse landscape that supports our diverse biota. Indeed, researching and preservmg Oklahoma's landscape-to-species relationships is worth researching and preserving twice as much elsewhere. ONPS Strawn, S. https://doi.org/10.22488/okstate.17.100015 Editorial Policies and Practices Oklahoma Native Plant Record is published annually by Oklahoma Native Plant Society. Submission for publication in the journal is open to all. Manuscripts will be accepted on topics related to Oklahoma's regional botany, including historical research reports, current research articles, site record species lists, and descriptions of new or important species sightings in Oklahoma. Oklahoma's environmental gradients of human impact, climate, and elevation make us a prime target for research on habitat edges, species ranges, and edge species, but articles of other themes may be included as well. Important works overlooked by journals of broader geographic regions will also be considered for publication here. Papers must not have been published previously or accepted for submission elsewhere and should represent research conducted in accordance with accepted procedures and scientific ethics. Submission of the article implies the granting of copyright permission to Oklahoma Native Plant Society. Manuscripts will be reviewed for content and appropriateness by at least two reviewers. The title page should state the affiliation and complete addresses of all authors and telephone numbers for the corresponding author. Research and technical papers should include a one-paragraph abstract of not more than 250 words. It should concisely state the goals, principal results, and major conclusions of the paper. All references, figures, and tables should be cited in the text. Common names should be referenced to a scientific name. Abbreviations of authorities for scientific names should follow Authors of Plant Names (Brummitt and Powell, 1992). Titles of periodicals should be abbreviated following Botanico-Peridoicum-Huntianum and its supplement - except in historic publications when original format will be used. Authors with access to IBM-compatible microcomputers are encouraged to send a copy of the manuscript on diskette in rtf (rich text format). If the manuscript is typed, manuscripts should be double-spaced on 8 1/2 X 11 inch paper with minimum one-inch margins and should be submitted in duplicate. Diskette or hardcopy manuscripts should be sent to the managing editor at the ONPS address on the back cover by June 1. Non-Profit Oklahoma Native Plant Society Organization c/o Tulsa Garden Center U.S. Postage Paid 2435 South Peoria Tulsa, OK Tulsa, Oklahoma 74114 Permit No. 357 In this issue of Oklahoma Native Plant Record Volume 2, Number 1, December 2002: 4 Vascular Plants of theWichita Mountains. Dr. PaulBuck 22 Floristic List for Comanche County, Oklahoma. Dr. Bruce W. Hoagland 54 Schoenoplectus hallii and S. saximontanus Wichita Mountains Wildlife Refuge Survey 2000. Dr. Lawrence K. Magrath 65 Pontotoc Ridge Floristic Survey: 1999. Dr. Forest Johnson, Ms. Patricia Folley (ed.) 82 ONPS Editorial: Water, Soil, and Plant Diversity in Oklahoma Oklahoma Native Plant Record, Journal of the Oklahoma Native Plant Society, Volume 2, Number 1, December 2002 Title Page Table of Contents Foreward Vascular Plants of the Wichita Mountains by Dr. Paul Buck Floristic List for Comanche County, Oklahoma by Dr. Bruce W. Hoagland Schoenoplectus hallii and S. saximontanus - Wichita Mountains Wildlife Refuge Survey 2000 by Dr. Lawrence K. Magrath Pontotoc Ridge Floristic Survey: 1999 by Dr. Forrest L. Johnson and Ms. Patricia Folley (ed.) ONPS Editorial - Water, Soil, and Plant Diversity in Oklahoma by Dr. Sheila A. Strawn Editorial Policies and Practices Back page index