REINWARDTIA

Published by Herbarium Bogoriense, Kebun Raya Indonesia
Volume 1, Part 4, pp. 463-466 (1952)

A NEGLECTED INDIAN SPECIES OF CYPERUS

J. H. KERN*

Cyperus alulatus Kern, spec. nov.—Fig. 1

Cyperus iria var. rectangularis Ktikenth. in Engl., Pflanzenr., Heft 101: 152.
1935. — Lectotypus: Duthie 23284 (Herb. Kew.).

Cyperus iria (non L.) sensu Clarke, Illustr. Cyper. pi. H f. 1. 1909, non al. —
Duthie 4480.

Subgen. Cyperus; sect. Iriae Kunth. — Annuus. Culmi erecti, tenues,
triquetri, laeves, 1—7 dm alti, 1—2 mm crassi, inferne paucifoliati. Folia
culmo breviora vel aequilonga, plana, flaccida, longe acuminata, in supe-
riore parte scabra, 1—5 mm lata. Anthela simplex vel subcomposita, laxa.
Bracteae oblique patentes, 2—4 longissimae, foliis consimiles, anthelam
valde superantes. Anthelae radii 3—9, e cladoprophyllis tubulosis ore
oblique truncatis postice cuspidatis vel bidentatis basi bruneis 0.5—1.5 cm
longis emergentes, inaequales, oblique patentes, graciles, compressi, laeves
vel apice hispiduli, ad 16 cm longi, longiores nonnunquam apice pauci-
ramosi; radioli brevissimi ab ochreolo caudato suffulti. Spicae ovatae vel
oblonga-ovatae, laxae vel subdensae, 1—3 cm longae, 8—25(—30) mm latae,
5—20-spiculosae, rhachi flexuosa angulis hispida. Spiculae compressae,
rectangule divaricatae vel imae subreflexae, ovatae usque ad oblongo-linea-
res, 3—12 mm longae, 2—2.5 mm latae, 4—18-f lorae, basi saepe a seta su-
buliformi suffultae. Rhachilla obscure brunea, recta, subexalata, internodiis
0.6—1 mm longis. Glumae membranaceae,1/3—1/2 parte imbricatae, patulae,
concavae, fere orbiculatae, (1.75—)2mm longae ac latae, sub apice emar-
ginato mucronulatae, dorso viridi 7-nerviae, lateribus fulvae purpureo-
notatae uninerviae, carina acuta arcuata superne angustissime alata,
spinuloso-ciliata. Stamina 2; antherae breves, oblongae, interdum lineares;
connectivum in appendicem brevem productum. Stylus fere nullus;
stigmata 3, achenio multo breviora. Achenium ambitu obovatum, trique-
trum, lateribus concavis, basi late stipitatum, apice mucronatum, atro-
bruneum, nitidum, dense puncticulatum, 1.5 mm longum, 0.8—0.9 mm
latum.

TYPUS.—Drummond 24733 (Herb. Bogor.)
Cyperus iria L. differt: rhachi glabri; spiculis 1.5—2 mm latis erecto-

patentibus; glumis minoribus 1.25—1.5 mm longis, dorso 3—5-nervatis,
carina exalulatis laevibus; rhachillae internodiis brevioribus (ca. 0.5 mm) ;
achenio minore, 1—1.25 mm longo.

•Botanist, Herbarium Bogoriense, Kebun Raya Indonesia.

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464 R E I N W A R D T I A [VOL. 1

Cyperus orthostachyus Franch. & Savat. valde quoque similis est;
haec species tamen insignis: foliis latioribus, usque ad 8 mm latis; spiculis
sanguineo-purpurascentibus, 1.5 mm latis, rhachilla subflexuosa, distincte
alata; glumis obsolete 5-nervatis, circa 1.5 mm longis, apice truncato-
rotundatis non productis, carina laevibus rectiusculis exalulatis; stigmati-
bus achenio subaequilongis; achenio minore, 1.25 mm longo.

If Cyperus microiria Steud. and C. orthostachyus Franch. & Savat.
are recognized as specifically distinct from C. iria L. (which, the author
believes, is their correct treatment), specific status should also be granted
to the taxon described above.

Cyperus iria var. rectangularis Kiikenth. was described as follows:

"Radii anthelae saepe patentissimi. Spiculae rectangule patentes 2—2% mm
latae. Squamae longiores 2 mm latae. Nux major 1%—2 mm longa."

This variety is mentioned from a few localities in India. Its identity
with Cyperus alulatus is evident from the specimen Duthie's collector
Inayat 23284, which number is cited by Kiikenthal.

The present author had the opportunity to study several of the num-
bers cited by Clarke (in J. Linn. Soc, Bot. 21:138-139. 1884) under Cyperus
iria. They proved Clarke's conception of C. iria to be fully in accordance
with the current one. Therefore it is remarkable that exactly Duthie 4480
was figured in the "Illustrations of Cyperaceae" as "exemplum typicum"
of Cyperus iria.

Perhaps Haines (Botany of Bihar and Orissa 5: 896. 1924) had this
species in view, when he said: "Glumes [of C.iria] 7— 9-nerved on and
dose to keel; keel sometimes scabrid."

The author is particularly grateful to Mr. E. Nelmes, who searched
for further specimens of the new species in the Herbarium of the Royal
Botanic Gardens, Kew, England, after it had been recognized among
the specimens of Cyperus iria in the Singapore Herbarium. The fragments
from several of the Kew sheets sent to Bogor and the data furnished
by Mr. Nelmes enabled him to amplify the description.

SPECIMENS EXAMINED by Mr. E. Nelmes and/or by the author (the latter ones
designated by !); unless otherwise indicated preserved in the Kew Herbarium.

INDIA and PAKISTAN. W i t h o u t e x a c t 1 o c a 1 i t y, Thomson 279'.— N.W.
H i m a l a y a s . Hazara, Shinkiari, Siran range, Aug. 29, 1899, Duthie's collector
Inayat 23281; Hazara, Abbottabad, 4500 ft., Sept. 9, 1890, Trotter 161G. — K a s h m i r .
Tawi Valley, wet places, 3000 ft., Aug. 25, 1891, Gammie s.n., Chalebagh, rice fields,

EXPLANATION OF FIGURE 1
FIG. 1. Cyperus alutatus Kern: habit; spikelet; part of rhachilla; two glumes, lateral
view; deflorate flower; stamens; nut, not fully ripe; habit 0.7 X, details 14 x. —

After Maries 356.

1952] KERN : On a new species of Cyperus 465

FIG. 1



466 R E I N W A R D T I A [VOL. 1

V. Jacquemont 873. — P u n j a b . Without exact locality, Drummond 24733 (BO, Kew) !,
940, 24730, 24734, 24735, 34732; Rawalpindi, Aug. 1872, Aitchison 129; Chamba,
between Kulel and Musroond, 4000 ft., Sept. 4, 1896, Gammie 18476! Simla, below t h e
cemetary, Aug. 25, 1917, H. H. Rich 681! — U n i t e d P r o v i n c e s . Garhwal, Lobha,
5—6000 ft., Aug. 31, 1885, Duthie 4480; Mussoorie, Dr. Bacon; Dehra Dun, Malapani
Road, 2000 ft., Oct. 1891, Gamble 23192; Dehra, 2000 ft., Aug. 1891, Gamble 23852;
Moradabad, Aug. 1843, ? collector. — C e n t r a l I n d i a . Gwalior, C. Maries 356
(Herb. Singapore) ! — B o m b a y . Poonah, wet fields, V. Jacquemont 344.

REINWARDTIA

Published by Herbarium Bogoriense, Kebun Raya Indonesia
Volume 1, Part 4, pp. 467-481 (1952)

MISCELLANEOUS BOTANICAL NOTES—IV*

C. G. G. J. VAN STEENIS**

SUMMARY

1. In connection with the first record from Malaysia (Atjeh, North Sumatra)
of a species of Schoepfia (Olacaceae), viz. S. fragrans Wall, in Roxb., some notes
on the genus are given, including a key to the species of section Schoepfiopsis (emended)
and to the two Indian species of Schoepfia. The specimens of the Kew and Leyden
Herbaria of these two species are listed. The name Schoepfia ffriffithii Tiegh. is
validly published in the present paper, if this was not done before.

2. Smilax pygmaea Merr. (Liliaceae) is recorded from Atjeh, Sumatra.
3. The first indigenous species of Mivmlus (Scrophulariaceae), M. tenellus Bunge,

is recorded for Malaysia from Atjeh, Sumatra.
4. A new species of Macadamia (Proteaceae) is described from Celebes: Maca-

damia hildebrandii Van Steenis. It belongs to the same genus as the common Australian
bush nut, M. ternifolia, with edible seeds.

5. Some information, additional to a previous paper on Biophytum (Oxalidaceae)
in Malaysia, is given.

• 6. Some records of plants new to Mount Pangrango, West Java, are mentioned.
7. The recent introduction and the present distribution in Malaysia of the

weed Eupatorium odoratum L. (Compositae) is discussed.
8. Some additional records of the liana Hollrungia aurantioides K. Schum.

(Passifloraceae) from New Guinea and from outside this island (Ternate, Moluccas)
are published.

31. Notes on Asiatic species of Schoepfia (Olacaceae)

The identification of a species of Schoepfia collected in the North
Sumatran highlands, in 1934 and 1937, has necessitated the examination
of specimens from continental Asia. Besides the work of Masters (in
Hook, f., Fl. Br. Ind. 1: 581-582. 1875) there are the treatments of Miers
(in J. Linn. Soc. Bot. 17: 70-77. 1878), Van Tieghem (in Bull. Soc. Bot.
Fr. 43: 550-551. 1896), and Valeton (Crit. Overz. Olacin. 123-130. 1886).
Miers and Van Tieghem have split the genus into two, respectively three,
other genera, treated by Engler (Nat. Pfl. Fam. Nachtr. I zum II.-IV.
Teil 145. 1897) as sections. I agree with the latter's view. The sections
appear well defined, but the general characteristics common to all are

*The first paper in this series appeared in Bull. bot. Gdns Buitenzorg1 III 17:
383-411. 1948; the second in Blumea 6: 243-246. 1948; the third in Bull. bot. Gdns
Buitenzorg III 18: 457-461. 1950.

**Director, Flora Malesiana Foundation.

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