REINWARDT1A

Vol. 10, Part 3, pp. 381 - 382 (1987)

ANEW STATUS FOE DESMODIUM OVALIFOLIUM (PAPILIONACEAE)

RUGAYAH
Herbarium Bogoriense, Bogor, Indonesia

ABSTRACT

Based on differences in anatomical and morphological characters, the
new combination Desmodium heterocarpon (L.) DC. subsp. heterocarpon
var. ovalifolium (Wall, ex Prain) Rugayah is proposed to accommodate
some specimens formerly included in D. heterocarpon var. heterocarpon,

ABSTRAK

Berdasarkan perbedaan ciri anatomi dan morfologinya, kombinasi
baru Desmodium heterocarpon (L.) DC. subsp. heterocarpon var. ovali-
folium (Wall, ex Prain) Rugayah diusulkan untuk mewadahi beberapa
spesimen yang sebelumnya dimasukkan dalam D. heterocarpon var.
heterocarpon.

In a recent study on the D. heterocarpon (L.) DC. complex (Rugayah
1986), some specimens were found to fit with the concept of D. ovalifolium
(Wall, ex Prain) Gagnep. which some recent authors (van Meeuwen, van
Steenis & Stemmerik 1961; Knaap-van Meeuwen 1962; Ohashi 1973) would
include in D. heterocarpon var. heterocarpon. Based on anatomical and mor-
phological evidences they are considered to belong to a separate taxonomic
entity, as can be recognized from the key presented below.

1 a. Stem and rachis strigosely-hairy, pods hairy at the margin only; the upper leaf
epidermis thicker than the lower one var. strigosum

b. Stem and rachis with hooked hairs, pods evenly hairy; the upper leaf epidermis
as thick as the lower one 2

2 a. Flowers loose, 3-4 mm long, pedicel 4-7 mm; leaflets obovate or elliptical; papilla
on both lower and upper epidermis, epidermal cells slightly sinuous; stomata 13,75-
16,25 X 10 - 12,5 pm var. heterocarpon

b. Flowers dense, more than 4 mm long, pedicel 2-3 mm; leaflets broadly elliptical
to almost orbicular; papilla on the lower epidermis only, epidermal cells very
sinuous; stomata 16,25 - 21,25 X 12,5 • 13,75 /µm var. ovalifolium

Consequently a new combination is proposed to accommodate this tax-
on as a variety of D. heterocarpon subsp. heterocarpon.

3 8 1



382 KKINWARDTFA [.Vol. 10

Desmodium heterocarpon (L.) DC. subsp. heterocarpon var. ovalifolium
(Wall, ex Prain) Rugayah, comb. nov.

Desmodium ovalifolium Wall. cat. no. 5730, nom. nud. —D. polycarpum DC. var.
ovalifolium (Wall, ex) Prain in King, J. As. Soc. Beng. 66 (2) : 141. 1897 (basionym).
—D. ovalifolium (Wall, ex Prain) Gagnep., FI. Indo-China 2 : 587. 1920.

DISTRIBUTION : Siam, Indo-China, Malay Peninsula, Sumatra, Borneo,
Celebes and Java.

SPECIMENS EXAMINED. Siam : Holttum 9714. - Indo-China : Squires 104. -
Malay Peninsula : Henderson 38116. — Sumatra : Bunnemeijer 1361, 1592, 2274, 2307,
2376, 5774; Kostermans 473;L'drzing 11626; Ouwehand 300; Rutten Kooistra 57; Teys-
mann s.n.; de Voogd 137, 152; van der Voort 3; van Steenis 1146. — Borneo : Jaheri s.n.;
J, & M.S. Clemens 205 78; Mondih 281; Tandom 2818. Celebes : Kjellberg 1 746— Java.:
Backer 1890, 4242, 4609, 9026, 32042; Bakhuizen v/d Brink f. 810; Huitema s.n.; Karta
186.

REFERENCES

KNAAP-VAN MEEUWEN. M.S. 1962. Preliminary revisions of some genera of Malaysian
Papilionaceae V — A census of the genus Desmodium. In Reinwardtia 6 (3) : 239 —
276.

OHASHI, H. 1973. The Asiatic species of Desmodium and its allied genera (Leguminosae).
In Ginkgoana 1 : 4 — 318.

KUGAYAH. 1986. Penelaahan kembali status taksonomi Desmodium ovalifolium. In
Berita Biologi 3 (5) : 203 - 208.

VAN MEEUWEN. M.S., VAN STEENIS, C.G.G.J. & STEMMERIK. J. 1961. Preliminary revisi-
ons of some genera of Malaysian Papilionaceae II. In Reinwardtia 6 (1) : 85 — 108.




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