2016 15 (2) A JOURNAL ON TAXONOMIC BOTANY, PLANT SOCIOLOGY AND ECOLOGY ISSN 0034 – 365 X | E-ISSN 2337 − 8824 | Accredited 792/AU3/P2MI-LIPI/04/2016 REINWARDTIA Decem ber 2016 15 (2) : 67 – 135 REINW ARD TIA REINWARDTIA Vol. 15. No. 2. 2016 CONTENTS Page Reinwardtia is a LIPI accredited Journal (792/AU3/P2MI-LIPI/04/2016) http://e-journal.biologi.lipi.go.id/index.php/reinwardtia Herbarium Bogoriense Botany Division Research Center for Biology – Indonesian Institute of Sciences Cibinong Science Center Jln. Raya Jakarta − Bogor, Km 46 Cibinong 16911, P.O. Box 25 Cibinong Indonesia REINWARDTIA Vol. 15. No. 2. 2016 CONTENTS Page ASIH PERWITA DEWI, NUNIK SRI ARIYANTI & EKO BAROTO WALUJO. Diversity of plants used for plaited crafts by the Dayak Iban-Désa in Kabupaten Sintang, Kalimantan Barat, Indonesia .............................................. ..... 67 DIAN LATIFAH, ROBERT A. CONGDON & JOSEPH A. HOLTUM. Growth responses of palm seedlings to different light intensities manipulating canopy gaps with an ecophysiological approach ........................................ ....... 81 ROSIE PRITCHETT, AURORA PHILLIPS, ANI MARDIASTUTI & ANDREW POWLING. Rattan diversity and broad edaphic niches in a tropical rainforest of Buton, Sulawesi, Indonesia ................................................. .................. 99 INGGIT PUJI ASTUTI & RUGAYAH. A new species of Murraya from Cyclops Mountain, Papua, Indonesia ......... 111 DEDEN GIRMANSYAH. A new species of Begonia (Begoniaceae) from Sumbawa, Lesser Sunda Islands, Indonesia ................................................................................................................... ......................................................115 I PUTU GEDE P. DAMAYANTO & ELIZABETH A. WIDJAJA. A new species of Schizostachyum (Poaceae- Bambusoideae) from Sumba Island, Indonesia .............................................................................................................. 119 J. F. VELDKAMP. A revision of Iseilema (Gramineae) in Malesia ............................................................................. 123 MIRAADILA M. I., SHABDIN Z. & MEEKIONG K. Two new species and one new geographical record for Sarawak, Malaysia (Cyperaceae: Mapanioideae) ......................................................................................... ................................. 129 Reinwardtia is a LIPI accredited Journal (792/AU3/P2MI-LIPI/04/2016) http://e-journal.biologi.lipi.go.id/index.php/reinwardtia Herbarium Bogoriense Botany Division Research Center for Biology – Indonesian Institute of Sciences Cibinong Science Center Jln. Raya Jakarta − Bogor, Km 46 Cibinong 16911, P.O. Box 25 Cibinong Indonesia REINWARDTIA A JOURNAL ON TAXONOMIC BOTANY, PLANT SOCIOLOGY AND ECOLOGY Vol. 15 (2): 67 – 135, December 22, 2016 Chief Editor Kartini Kramadibrata (Mycologist, Herbarium Bogoriense, Indonesia) Editors Dedy Darnaedi (Taxonomist, Herbarium Bogoriense, Indonesia) Tukirin Partomihardjo (Ecologist, Herbarium Bogoriense, Indonesia) Joeni Setijo Rahajoe (Ecologist, Herbarium Bogoriense, Indonesia) Marlina Ardiyani (Taxonomist, Herbarium Bogoriense, Indonesia) Himmah Rustiami (Taxonomist, Herbarium Bogoriense, Indonesia) Lulut Dwi Sulistyaningsih (Taxonomist, Herbarium Bogoriense, Indonesia) Topik Hidayat (Taxonomist, Indonesia University of Education, Indonesia) Eizi Suzuki (Ecologist, Kagoshima University, Japan) Jun Wen (Taxonomist, Smithsonian Natural History Museum, USA) Barry J Conn (Taxonomist, School of Life and Environmental Sciences, The University of Sydney, Australia) David G. Frodin (Taxonomist, Royal Botanic Gardens, Kew, United Kingdom) Secretary Rina Munazar Layout Dede Aryanto Illustrators Subari Wahyudi Santoso Anne Kusumawaty Correspondence on editorial matters and subscriptions for Reinwardtia should be addressed to: HERBARIUM BOGORIENSE, BOTANY DIVISION, RESEARCH CENTER FOR BIOLOGY– INDONESIAN INSTITUTE OF SCIENCES CIBINONG SCIENCE CENTER, JLN. RAYA JAKARTA – BOGOR KM 46, CIBINONG 16911, P.O. Box 25 CIBINONG INDONESIA PHONE (+62) 21 8765066; Fax (+62) 21 8765062 E-MAIL: reinwardtia@mail.lipi.go.id http://e-journal.biologi.lipi.go.id/index.php/reinwardtia Cover images: Mapania sembilingensis Miraadila, Shabdin & Meekiong. A. Habit; B. Leaf apex details; C. Sheath margin details; D. Capitate inflorescence; E. Spike; F. Spicoid bract [Drawing by Meekiong, K.]. The Editors would like to thank all reviewers of volume 15(2): David Simpson, Herbarium Kewense, Royal Botanic Gardens, Kew, UK Herwasono Soedjito, Research Center for Biology, Indonesian Institute of Sciences, Bogor, Indonesia Jay H. Bernstein, Robert J. Kibbee Library, Kingsborough Community College, New York, USA Kuswata Kartawinata - Integrative Research Center, The Field Museum, 1400 Lake Shore Drive, Chicago, USA Mark Hughes - Royal Botanic Garden Edinburgh, Edinburgh, Scotland, UK Mien A. Rifai - Akademi Ilmu Pengetahuan Indonesia (AIPI), Indonesia Siti Nur Hidayati - Middle Tennessee State University, Tennessee, USA Soejatmi Dransfield - Herbarium Kewense, Royal Botanic Gardens, Kew, UK Wong Khoon Meng - Singapore Botanic Garden, Singapore REINWARDTIA Vol 15, No 2, pp: 123 − 127 123 A REVISION OF ISEILEMA (GRAMINEAE) IN MALESIA Received 23 August 2016; accepted 05 September 2016 J. F. VELDKAMP Naturalis Biodiversity Center, POB 9517, 2300 RA Leiden, The Netherlands. Email: jef.veldkamp@naturalis.nl ABSTRACT VELDKAMP, J. F. 2016. A revision of Iseilema (Gramineae) in Malesia. Reinwardtia 15(2): 123 – 127. — There are three very rare and localized species in Malesia; one from Java is new. Notes on some other Southeast Asian species are given. Key words: Iseilema, Malesia, Revision. ABSTRAK VELDKAMP, J. F. 2016. Revisi Iseilema (Gramineae) di Malesia. Reinwardtia 15(2): 123 – 127. — Di Malesia ter- dapat tiga jenis yang sangat langka dengan penyebaran terbatas. Satu jenis baru dari Jawa. Catatan mengenai jenis lain dari Asia Tenggara diberikan. Kata kunci: Iseilem a, Malesia, Revisi. INTRODUCTION Iseilema Andersson is a genus with about 20 species, ranging from SE Asia to Australia (13 spp). In Malesia there are three species, all very rare and local. The genus name is derived from “isos” (’ισος, equal) and “eilèma” (’ειλημα, involucre). There has been some confusion about the gender of the name, Andersson had I. arguta and I. prostrata (feminine), but “eilèma” is neuter and therefore the epithets must be neuter as well (Hackel, 1889:681). The similarity to Themeda Forssk. has often been noted and some species have been included in it (or in A nthistiria Naezén, a synonym). For the record, Roberty (1960) reduced all taxa to Themeda section Iseilema with a single species T. prostrata (L.) Roberty with six varieties. Molecular studies by Skendzic et al. (2007:538, 540) found it closely related to Heteropogon Pers. Welker et al. (2015) showed that the relationship with Themeda is consistent, but the two were resolved into different terminal branches, Iseilema being more associated with Bothriochloa Kuntze, Capillipedium Stapf, and Dichanthium Willemet, and Themeda linked to Heteropogon. HISTORY The first species known to Western science was Andropogon prostratus L. (1771: 304) based on a König collection from India (LINN 1211.8), now Iseilema prostratum. Willdenow (1806) renamed it to A nthistiria prostrata. Steudel (1854) described A nthistiria argutum from Burma, and superfluously renamed Andropogon prostratus to Anthistiria linneana because of the supposed homonymy with Anthestiria (!) prostrata Trinius (1832: 321). Trinius, however, cited A nthistiria prostrata (L.) Willd. (1806), based on A ndropogon prostratus L. Andersson (1856) described four species. The lectotype, I. prostratum, was designated by Roberty (1960:99). Hackel (1889) had five species. Over the years others were added, forming a range from Pakistan, Sri Lanka, India, Burma, Thailand, to Laos and Cambodia with centres of speciation in Australia with 12 species (10 in Queensland!) and India with seven. Hackel (1889) mentioned I. laxum as introduced in Mauritius, but it is not mentioned by Hubbard & Vaughan (1940). Roberty (1960) cited this as “Mascareignes”. Roberty (1960) mentioned I. membranaceum Domin and I. vaginiflorum Domin for Hawaii, but the genus is not mentioned in Warren et al. (1990). Ms. Camus (1955) erroneously cited I. minutiflorum Jansen for the Philippines. In Malesia the genus is very rare, of the three species only eight collections were seen, while two are only known from their types. Van Steenis (1936) mentioned the occurrence of I. argutum Nees ex Steud. for Java in much depleted areas that formerly had teak forest, on very desiccated soil and with periodic fire disturbance. This was obviously a misidentification with a species from Burma, only known to me from two collections (K, P). The Java species was recognized by Ohwi (in sched. in BO) as I. javanicum and he is here posthumously regarded as the first author. Jansen (1953a, b) described two more species from Sumba and Wetar. MORPHOLOGY The genus is rather similar to Themeda because of the spatheate and capitulate terminal inflo-rescences with four involucral spikelets. One mailto:jef.veldkamp@naturalis.nl http://www.ipni.org/ipni/idPlantNameSearch.do;jsessionid=12F8375DE9D4BDE12D7F01144BC1876B?id=388952-1&back_page=%2Fipni%2FeditSimplePlantNameSearch.do%3Bjsessionid%3D12F8375DE9D4BDE12D7F01144BC1876B%3Ffind_wholeName%3DAnthistiria%2Bprostrata%26output_format%3 http://www.ipni.org/ipni/idPlantNameSearch.do;jsessionid=12F8375DE9D4BDE12D7F01144BC1876B?id=388952-1&back_page=%2Fipni%2FeditSimplePlantNameSearch.do%3Bjsessionid%3D12F8375DE9D4BDE12D7F01144BC1876B%3Ffind_wholeName%3DAnthistiria%2Bprostrata%26output_format%3 REINWARDTIA 124 [VOL.15 difference is that in Iseilema these capitules would fall off as a unit, but this is also the case in T. gigantea (Cav.) Hack. In the other species of Themeda the spikelets disintegrate. The involucral spikelets are shortly pedicelled, whereas in Themeda they are paired, one sessile, the other subsessile. It has been suggested (Hackel, 1889:679) that these diaspores are dispersed by the wind, while in Themeda the fertile spikelets would be epizoochoric. Some species are provided with small to minute annular structures especially in the inflorescences. These generally have been taken for glands, e.g. by Sprengel (1815: 14, as Cymbopogon glandulosus Spreng. = Iseilema prostratum), Hubbard (1935c), and Uppuluri & Satyavati (1968:668). Andersson (1856) called them “tubercles”. I have seen no field notes on whether they secrete anything or are visited by insects. Iseilema Ander sson Iseilema Andersson, Nova Acta Regiae Soc. Sci. Upsal., III, 2 (1856) 231, 250. — Anthistiria Naezén [unranked] Iseilema (Andersson) Benth. & Hook. f., Gen. Pl. 3 (1883) 1136. —Ischaemum L. sect. Iseilema Roberty, Boissiera 9 (1960) 99. — Lectotype: Iseilema prostratum (L.) Andersson, designated by Roberty [who reduced all taxa to a single species of Themeda Forssk. in Boissiera 9 (1960) 99], followed by Clayton & Renvoize (1986: 360). Annual or perennial. Culms solid. Ligule collar- shaped, membranous. Inflorescence paniculate, spatheate, decompound, spikelet-bearing axes much reduced, clustered in capitules supported by a spatheole, deciduous as a whole, involucre formed by two homogamous involucral pairs of male or sterile subsessile to shortly pedicelled spikelets, fused at base. Involucral spikelets with lower glume 5−9−nerved. Common callus usually hairy. Rachis persistent. Fertile spikelets 1 (rarely 2), pedicelled, female or bisexual; callus absent; lower glumes flat on the back. Awns (when present) stipitate or from a small sinus. Pedicels free from the joints. Pedicelled spikelets 2, vari- ously reduced, male to sterile. x = very variable, probably derived from 10. Distribution. ca. 20 spp. in SE Asia to Australia, three in Malesia, all very local and rare (Java, Sumba, Wetar). Key to the taxa 1a. Plants annual, culm 0.1−0.4 m long, nodes glabrous. Leaf blades 5−16 cm by 0.5−2.6 mm wide. Callus hairs 1.5−2 mm long. Invo- lucral spikelets 4 mm long. Fertile spikelets 3−4 mm long. Awn geniculate i.s., 8−18 mm long. Anthers ca. 2 mm long ………..…….... 2 b. Plants perennial, culm 1.3−2 m long, nodes barbate. Leaf blades 10−60 cm by 5−8 mm wide. Callus hairs 1 mm long. Involucral spikelets 4.5−5 mm long. Fertile spikelets 6−7 mm long. Awn more or less straight i.s., 3−8 mm long. Anthers 1.4−1.5 mm long. — Spathes and glumes of the involucral spikelets on the keels glandular. — Java. .................................................. 1. I. javanicum 2a. Leaf blades distally smooth. Spatheoles and peduncles eglandular. Involucral spikelets with lower glumes glabrous, keels eglandu- lar, purple mottled. Fertile spikelet ca. 4 mm long. Pedicelled spikelets with lower glumes with eglandular keels. Awn 18−20 mm long. — Wetar …….……………… 2. I. maculatum b. Leaf blades distally scabrid. Spatheoles and peduncles glandular. Involucral spikelets with lower glumes distally hairy, keels glan- dular, not mottled. Fertile spikelet 3−3.5 mm long. Pedicelled spikelets with lower glumes with glandular keels. Awn 8−10 mm long. — Sumba ………....…………. 3. I. minutiflorum 1. Iseilema javanicum Ohwi & Veldk., spec. nov. — Type: V an Steenis 8166-d (holo L; BO 1443573, -4; E, K). new § Iseilema argutum auct. non Andersson: Van Steenis, Jubileum Uitg. Trop. Nat. (1936) 111; C. Monod in Backer & Bakh. f., Fl. Java 3 (1968) 616. Plants perennial. Culms 1.3−2 m long, eglandular below nodes. Culm nodes barbate. Leaf blades 10−60 cm by 5−8 mm, margins distally scabrid, apex acute. Spatheoles glandular or not. Peduncles very short, eglandular. Capitule callus hairs ca. 1 mm long. Involucral spikelets pedicels 0.5−1 mm long, longer than broad, glabrous to pilose. Involucral spikelets 4.5−5 mm long, lower glumes not mottled, glabrous, not sulcately nerved, 7−or 9−nerved; lower glumes keels glandular. Fertile spikelet 6−7 mm long; base glabrous or hairy. Awn more or less straight i.s., 3−8 mm long. Pedicelled spikelets well-developed; lower glumes glandular. Anthers 1.4−1.5 mm long. Distribution. Malesia: J ava [Indramayu, Cire- bon, Houtvesterij (Forestry Reserve) Plosokerep]. Habitat. Mar shy gr ass fields, in for mer teak forests, locally abundant, fire-climax savannah on very poor soil with e.g. A ndropogon amboinicus (L.) Merr. [= Sorghum nitidum (Vahl) Pers.], Polytoca bracteata R. Br. [= Polytoca digitata (L. f.) Druce], 0−30 m alt. See Van Steenis (1936) for an extensive description. 2016] 125 VELDKAMP: A revision of Iseilema (Gramineae) In Malesia Specimen examined. J ava: Bernard 6-1925 (BO); Van 3-7-1923, Harreveld sn. (BO); Java, Indramayu, Cirebon, Plosokerep, van Steenis 7483 (BO, L); 8166-d (BO, E, K, L); 6680 (BO, L); teak forest area, 17522 (BO, L). ; van Steenis 8166-d (T) (BO, L); van Steenis 17522 (BO, L). Collector’s notes. Tall, ca. 2 m. Sheath base, culms more or less waxy whitish. Notes. Differ s fr om I. argutum fr om Bur ma as follows: a. Culms 0.1−1 m long, nodes glabrous. Peduncles glandular. Involucral spikelets with lower glumes sulcately 5−nerved, nerved, keels pilose, eglandular. Fertile spikelet 3−5.2 mm long. Awn geniculate i.s. Anthers 2−3 mm long..................................................... I. argutum b. Culms 1.3−2 m long, nodes barbate. Peduncles eglandular. Involucral spikelets with lower glumes not sulcately 7−or 9−nerved, keels glandular. Fertile spikelet 6−7 mm long. Awn more or less straight i.s. Anthers 1.4−1.5 mm long................................................. I. javanicum Slightly different from I. thorelii A. Camus from Continental SE Asia. The differences may be explained by the paucity of material and their development and is summarised as follows: a. Culms 1.3−2 m long. Involucral spikelets with lower glumes glabrous. Awns more or less straight i.s., 3−8 mm long. Anthers 1.4−1.5 mm long. — Java................................... I. javanicum a. Culms 0.6−1.3 m long. Involucral spikelets with lower glumes pilose in lower half. Awns geni-culate i.s., 8−13 mm long. Anthers ca. 2 mm long. — Continental SE Asia........ I. thorelii 2. ISEILEMA MACULATUM Jansen Iseilema maculatum Jansen, Reinwardtia 2 (1953) 302. — Type: Elbert 4685 (holo L, sh. 941.215—87; perhaps FR, K). Plants annual. Culms 0.25−0.45 m long, eglandular below nodes. Culm nodes glabrous. Leaf blades 6−16 cm by 0.5−1.5 mm, distally smooth, apex acute. Spatheoles eglandular. Pedun- cles very short, eglandular. Capitule callus hairs ca. 1.5 mm long. Involucral spikelets pedicels ca. 1 mm long, longer than broad, glabrous. Involucral spikelets ca. 4 mm long; lower glumes purple mottled, glabrous, keels eglandular, not sulcately nerved, 5− or 7−nerved. Fertile spikelet ca. 4 mm long, base glabrous. Awn geniculate i.s, 9–18 mm long. Pedicelled spikelets well-developed; lower glumes eglandular. Anthers ca. 2 mm long. Distribution. Malesia: Lesser Sunda Isl. (Wetar). Habitat. Dr y Eucalyptus savannahs near the coast; up to 50 m alt. Note. Only known fr om the type collection. 3. ISEILEMA MINUTIFLORUM Jansen Iseilema minutiflorum Jansen, Acta Bot. Neerl. 2 (1953) 382, t. 10. — Type: Monod de Froideville 2012 (holo BO1888838; A, K, BRI, L, SING). Plants annual. Culms 0.1–0.3 m long, under the nodes eglandular, nodes glabrous. Leaf blades 3–7 cm by 1.2–2.6 mm, margins distally scabrid, apex acute. Spatheoles glandular. Peduncles very short, minutely glandular. Capitule callus hairs 1.5--2 mm long. Involucral spikelets pedicels ca. 0.4 mm long, longer than broad, pilose. Involucral spikelets ca. 3 mm long. Lower glumes purple mottled, keels pilose in upper half keels glandular (40 x!), not sulcately 5- or 7-nerved. Fertile spikelet 3–3.5 mm long, base hairy. Awn geniculate i.s., 8–10 mm long. Pedicelled spikelets well-developed (fide Jansen, n.v.). Lower glumes glandular. Anthers n.v. Distribution. Malesia: Lesser Sunda Isl. (Sumba). Habitat. In gr ass vegetation on limestone, subjected to annual burning, ca. 50 m alt. In groups on shallow soil, apparently a short living species. Note. Only known fr om the type collection. OTHER SPECIES For the identification of the Malesian species, it was necessary to look more closely at some Continental Asian ones. As in general their descriptions were incomplete, I had to make some of my own, which I think might be useful to other students of the genus. 1. ISEILEMA ARGUTUM (Nees ex Steud.) Andersson Iseilema argutum (Nees) Andersson, Nova Acta Regiae Soc. Sci. Upsal., III, 2 (1856) 252 (“arguta”). — Anthistiria arguta Nees ex Steud., Syn. Pl. Glumac. (1854) 1:401. — [Themeda prostrata (L.) Roberty var. arguta Roberty, Boissiera 9 (1960) 100, nom. inval.] . — Type: W allich 8769 (A kkul Mahmud) (holo P; G, L, fragm.; K, IDC microfiche 7394, ? W). Plants perennial. Culms 0.1−1 m long, with minute dot-like glands or eglandular (with some minute glands: 40 ×!) below nodes. Culm nodes glabrous. REINWARDTIA 126 [VOL.15 Leaf blades 5.5−18 cm by 2−6 mm, margins distally scabrid, apex acute or obtuse-apiculate. Spatheoles glandular. Peduncles very short, glandular. Capitule callus hairs ca. 1.5 mm long. Involucral spikelets pedicels 0.6−1 mm long, longer than broad, glabrous or pilose. Involucral spikelets 4.5−6 mm long; lower glumes not mottled, keels pilose, eglandular, sulcately 5- nerved. Fertile spikelet 3−5.2 mm long, base hairy. Awn geniculate i.s., 4.5−18 mm long. Pedicelled spikelets well-developed or reduced to the pedi- cels; lower glumes eglandular or glandular (margins long hairy). Anthers 2−3 mm long. Distribution. Bur ma: Bago Div., Pyay (“Prome”: Wallich 8769); Magwe Div. (U Thein Lwin 398, K). Kress et al. (2003) also reported Ayeyarwadi, Mandalay, Yangon. Habitat. Fair ly common on black calcar eous stiff clay soil; altitude unknown. Specimen examined. Burma, montes ad Prome, Akkul Ahmad in Wallich 8769 (G, L, fragm., K, P, W; IDC microfiche 7394). Note. Descr iption based on the two collections cited above. 2. ISEILEMA THORELII A. Camus Iseilema thorelii A. Camus, Bull. Mus. Natl. Hist. 24: 540−541. 1918; in Lecomte, Fl. Indo-Chine 7: 365−366, t. 37, f. 6−8. 1922; Schmid, Agron. Trop. (Nogent-sur-Marne) 13: 235, t. 45, f. 3c−e. 1958.— Lectotype: Thorel 2494 (holo P, P 01942662), designat- ed here. ? Iseilema schmidii A. Camus, J. Agric. Trop. & Bot. Appl. 2: 201. 1955; Schmid, Agron. Trop. (Nogent-sur- Marne) 13: 235 (“schmidiana”). 1958. — Type: Schmid 2467 (holo P, not found). Plants perennial. Culms 0.6−1.3 m long, eglandular below nodes. Culm nodes barbate. Leaf blades 20−34 cm by 2.5−6 mm, margins distally scabrid, apex acute. Spatheoles glandular. Peduncles very short, eglandular. Capitule callus hairs ca. 1 mm long. Involucral spikelets pedicels 0.5−1 mm long, longer than broad, pilose. Involucral spikelets 3.5−5.5 mm long; lower glumes not mottled, keels pilose in lower half, glandular, sulcately 5-nerved. Fertile spikelet 4.5−8 mm long, base glabrous or hairy. Awn geniculate i.s., 8−13 mm long. Pedicelled spikelets well-developed; lower glumes glandular. Anthers ca. 2 mm long. Distribution. Continental SE Asia: Laos (Champassak (“Bassac”), Thailand (E: Nakhon Ratchasima; Central: Saraburi; SE: Chanthaburi, Prachin Buri, Sa Kaeo); Vietnam (Prov.: Binh Phuoc, Dak Lac (“Darlac”). Habitat. Open deciduous for est, savannahs, marshy places with Acanthaceae, Impatiens, Ischaemum, Labiatae, 100−700 m alt. Specimen examined. Laos, Champassak (“Bassac”), Thorel 2494 (P). Thailand, Sa Kaeo, Aran Pratet, Put 1981 (K, P) Notes. Notwithstanding Ms. Camus’ (1955) and Schmid’s (1958; “schmidiana”) remarks, the difference from I. schmidii A. Camus (which could not be found in P) is not clear to me. The two species may be distinguished as follows: a. Leaf blades 9−10 cm long, apex acuminate. Involucral spikelets with lower glumes gla- brous. Anthers ca. 2.5 mm long......... I. schmidii b. Leaf blades 20−34 cm long, apex acute. Involucral spikelets with keels of the lower glumes pilose in the lower half. Anthers ca. 2 mm long .............................................. I. thorelii Roberty (1960), apparently based on a syntype in G, equated I. thorelii with I. laxum auct. non Hack. (= I. hackelii U.B. Shrestha & Gandhi). ACKNOWLEDGEMENTS This revision is based on the material in BO and L. The herbaria of BISH, K, MEL, PNH, PTBG, SING were visited and their Directors and Keepers are much thanked for their hospitality and access to their specimens. REFERENCES ANDERSSON, N. J. 1856. Monographiae andropo- gonearum. 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Name of taxon in taxonomic treatment should be presented in the long form that is name of taxon, author’s name, year of publication, abbreviated journal or book title, volume, number and page. Map/line drawing illustration/photograph. Map, line dr awing illustr ation, or photogr aph pr efer ably should be prepared in landscape presentation to occupy two columns. Illustration must be submitted as orig- inal art accompanying, but separated from the manuscript. The illustration should be saved in JPG or GIF format at least 350 pixels. Legends or illustration must be submitted separately at the end of the manu- script. References. Bibliogr aphy, list of liter atur e cited or r efer ences follow the Har var d system as the fol- lowing examples. Journal : KRAENZLIN, F. 1913. Cyrtandraceae novae Philippinenses I. Philipp. J. Sci. 8: 163–179. MAYER, V., MOLLER, M., PERRET, M. & WEBER, A. 2003. Phylogenetic position and generic differentiation of Epithemateae (Gesneriaceae) inferred from plastid DNA sequence data. American J. Bot. 90: 321–329. Proceedings :TEMU, S. T. 1995. Peranan tumbuhan dan ternak dalam upacara adat “Djoka Dju” pada suku Lio, Ende, Flores, Nusa Tenggara Timur. In: NASUTION, E. (Ed.). Prosiding Seminar dan Lokakarya Nasional Etnobotani II. LIPI & Perpustakaan Nasional: 263–268. (In Indonesian). SIMBOLON, H. & MIRMANTO, E. 2000. Checklist of plant species in the peat swamp forests of Central Kalimantan, Indonesia. In: IWAKUMA, T. et al. (Eds.) Proceedings of the International Sym- posium on: Tropical Peatlands. Pp.179-190. Book : RIDLEY, H. N. 1923. Flora of the Malay Peninsula 2. L. Reeve & Co. Ltd, London. Part of Book : BENTHAM, G. 1876. Gesneriaceae. In: BENTHAM, G. & HOOKER, J. D. Genera plantarum 2. Lovell Reeve & Co., London. Pp. 990–1025. Thesis : BAIRD, L. 2002. A Grammar of Kéo: An Austronesian language of East Nusantara. 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Spicoid bract [Drawing by Meekiong, K.]. 2016 15 (2) A JOURNAL ON TAXONOMIC BOTANY, PLANT SOCIOLOGY AND ECOLOGY ISSN 0034 – 365 X | E-ISSN 2337 − 8824 | Accredited 792/AU3/P2MI-LIPI/04/2016 REINWARDTIA Decem ber 2016 15 (2) : 67 – 135 REINW ARD TIA REINWARDTIA Vol. 15. No. 2. 2016 CONTENTS Page Reinwardtia is a LIPI accredited Journal (792/AU3/P2MI-LIPI/04/2016) http://e-journal.biologi.lipi.go.id/index.php/reinwardtia Herbarium Bogoriense Botany Division Research Center for Biology – Indonesian Institute of Sciences Cibinong Science Center Jln. Raya Jakarta − Bogor, Km 46 Cibinong 16911, P.O. Box 25 Cibinong Indonesia REINWARDTIA Vol. 15. No. 2. 2016 CONTENTS Page ASIH PERWITA DEWI, NUNIK SRI ARIYANTI & EKO BAROTO WALUJO. Diversity of plants used for plaited crafts by the Dayak Iban-Désa in Kabupaten Sintang, Kalimantan Barat, Indonesia .............................................. ..... 67 DIAN LATIFAH, ROBERT A. CONGDON & JOSEPH A. HOLTUM. Growth responses of palm seedlings to different light intensities manipulating canopy gaps with an ecophysiological approach ........................................ ....... 81 ROSIE PRITCHETT, AURORA PHILLIPS, ANI MARDIASTUTI & ANDREW POWLING. Rattan diversity and broad edaphic niches in a tropical rainforest of Buton, Sulawesi, Indonesia ................................................. .................. 99 INGGIT PUJI ASTUTI & RUGAYAH. A new species of Murraya from Cyclops Mountain, Papua, Indonesia ......... 111 DEDEN GIRMANSYAH. A new species of Begonia (Begoniaceae) from Sumbawa, Lesser Sunda Islands, Indonesia ................................................................................................................... ......................................................115 I PUTU GEDE P. DAMAYANTO & ELIZABETH A. WIDJAJA. A new species of Schizostachyum (Poaceae- Bambusoideae) from Sumba Island, Indonesia .............................................................................................................. 119 J. F. VELDKAMP. A revision of Iseilema (Gramineae) in Malesia ............................................................................. 123 MIRAADILA M. I., SHABDIN Z. & MEEKIONG K. Two new species and one new geographical record for Sarawak, Malaysia (Cyperaceae: Mapanioideae) ......................................................................................... ................................. 129 Reinwardtia is a LIPI accredited Journal (792/AU3/P2MI-LIPI/04/2016) http://e-journal.biologi.lipi.go.id/index.php/reinwardtia Herbarium Bogoriense Botany Division Research Center for Biology – Indonesian Institute of Sciences Cibinong Science Center Jln. Raya Jakarta − Bogor, Km 46 Cibinong 16911, P.O. Box 25 Cibinong Indonesia 2941-6139-1-SM_1-1 2941-6139-1-SM_2-2 2941-6139-1-SM_3-3 2941-6139-1-SM_4-4 2947-6151-2-PB 2941-6139-1-SM_18-18 2941-6139-1-SM_19-19 2941-6139-1-SM_20-20 2941-6139-1-SM_21-21