REINWARDTIA  

2021  20 (1) 

ISSN 0034 – 365 X | E-ISSN 2337 − 8824 | Accredited 10/E/KPT/2019 

 A JOURNAL ON TAXONOMIC BOTANY, PLANT SOCIOLOGY AND ECOLOGY  



 

REINWARDTIA  
 

A JOURNAL ON TAXONOMIC BOTANY,  PLANT SOCIOLOGY AND ECOLOGY  
 

Vol.  20 (1): 1 – 41,  July 07,  2021 
 

Chief Editor 

Kartini Kramadibrata (Mycologist, Herbarium Bogoriense, Indonesia) 

 

Editors 

Dedy Darnaedi (Taxonomist, Herbarium Bogoriense, Indonesia) 

Tukirin Partomihardjo (Ecologist, Herbarium Bogoriense, Indonesia) 

Joeni Setijo Rahajoe (Ecologist, Herbarium Bogoriense, Indonesia) 

Marlina Ardiyani (Taxonomist, Herbarium Bogoriense, Indonesia) 

Himmah Rustiami (Taxonomist, Herbarium Bogoriense, Indonesia)  

Lulut Dwi Sulistyaningsih (Taxonomist, Herbarium Bogoriense, Indonesia) 

Eka Fatmawati Tihurua (Morphologist, Herbarium Bogoriense, Indonesia) 

Topik Hidayat (Taxonomist, Indonesia University of Education, Indonesia) 

Eizi Suzuki (Ecologist, Kagoshima University, Japan) 

Jun Wen (Taxonomist, Smithsonian Natural History Museum,  USA) 

Barry J. Conn (Taxonomist, School of Life and Environmental Sciences, The University of Sydney, Australia) 

Graham Eagleton (Wagstaffe, NSW, Australia) 
 

Layout 

Liana Astuti 

 

Illustrators 

Wahyudi Santoso  

Anne Kusumawaty  
 
 
 

Correspondence on editorial matters and subscriptions for Reinwardtia should be addressed to: 

HERBARIUM BOGORIENSE, BOTANY DIVISION, 

RESEARCH CENTER FOR BIOLOGY– INDONESIAN INSTITUTE OF SCIENCES 

CIBINONG SCIENCE CENTER, JLN. RAYA JAKARTA – BOGOR KM 46, 

CIBINONG 16911,  P.O. Box 25 CIBINONG  

INDONESIA 

PHONE (+62) 21 8765066; Fax (+62) 21 8765062  

E-MAIL: reinwardtia@mail.lipi.go.id 

http://e-journal.biologi.lipi.go.id/index.php/reinwardtia 
 

 
   
       

Cover images: Begonia robii Ardi & Girm. A. Plant habit. B. Stipules. C. Bracts. D. Inflorescence. E. 
Male flower, front view. F. Male flower, side view. G. Female flower, front view. H. Female flower, side 
view. I. Fruit. J. Ovary, cross section of middle part. From Wisnu Ardi WI 761. Photos by W.H. Ardi.  

http://blog.sivitas.lipi.go.id/ekaf001/


 

The Editors would like to thank all reviewers of volume 20(1): 
 

 

 
Andrew J. Henderson, The New York Botanical Garden, New York, USA 

Badal Kumar Datta, Tripura University, Agartala, India  
Chien C. Lee, Kuching, Sarawak, Malaysia 

Daniel C. Thomas, Singapore Botanic Gardens, Singapore 
Ren-Yong Yu, Naturalis Biodiversity Center, Leiden, The Netherlands 

Roderick W. Bouman, Hortus Botanicus, Leiden, The Netherlands  
Rugayah, Herbarium Bogoriense, Bogor, Indonesia 

Shih-Hui Liu, National Sun Yat-Sen University, Kaohsiung, Taiwan