REINWARDTIA  

2022  21 (2) 

ISSN 0034 – 365 X | E-ISSN 2337 − 8824 | Accredited 158/E/KPT/2021 

 A JOURNAL ON TAXONOMIC BOTANY, PLANT SOCIOLOGY AND ECOLOGY  



 

REINWARDTIA  
 

A JOURNAL ON TAXONOMIC BOTANY,  PLANT SOCIOLOGY AND ECOLOGY  
 

Vol. 21 (2): 43–82, December 23, 2022 
 

Chief Editor 

Kartini Kramadibrata (Mycologist, Herbarium Bogoriense, Indonesia) 

 

Editors 

Dedy Darnaedi (Taxonomist, Herbarium Bogoriense, Indonesia) 

Tukirin Partomihardjo (Ecologist, Herbarium Bogoriense, Indonesia) 

Joeni Setijo Rahajoe (Ecologist, Herbarium Bogoriense, Indonesia) 

Marlina Ardiyani (Taxonomist, Herbarium Bogoriense, Indonesia) 

Himmah Rustiami (Taxonomist, Herbarium Bogoriense, Indonesia)  

Lulut Dwi Sulistyaningsih (Taxonomist, Herbarium Bogoriense, Indonesia) 

Eka Fatmawati Tihurua (Morphologist, Herbarium Bogoriense, Indonesia) 

Topik Hidayat (Taxonomist, Indonesia University of Education, Indonesia) 

Eizi Suzuki (Ecologist, Kagoshima University, Japan) 

Jun Wen (Taxonomist, Smithsonian Natural History Museum,  USA) 

Barry J. Conn (Taxonomist, School of Life and Environmental Sciences, The University of Sydney, Australia) 

Graham Eagleton (Wagstaffe, NSW, Australia) 
 

Layout 

Andri Agus Rahman 

 

Illustrators 

Wahyudi Santoso  

Anne Kusumawaty  
 
 
 

Correspondence on editorial matters and subscriptions for Reinwardtia should be addressed to: 

HERBARIUM BOGORIENSE, NATIONAL RESEARCH AND INNOVATION AGENCY 

CIBINONG SCIENCE CENTER, JLN. RAYA JAKARTA – BOGOR KM 46, 

CIBINONG, BOGOR 16911, INDONESIA 

E-MAIL: reinwardtia@brin.go.id 

http://e-journal.biologi.lipi.go.id/index.php/reinwardtia 
 

 
   
       

Cover images: Top left: Cyrtandra pendula Blume. Top middle: C. rosea Ridl. Top right: C. 
rubriflora P.E.Sm. & H.J.Atkins. Below left: Cyrtandra sp. Below right: C. pauciflora Ridl. 
Photos by S. Barber (C. pendula Blume, C. rosea Ridl., C. pauciflora Ridl.), P. Wilkie (C. ru-
briflora P.E.Sm. & H.J.Atkins), M. Hughes (Cyrtandra sp.).  

http://blog.sivitas.lipi.go.id/ekaf001/


 

The Editors would like to thank all reviewers of volume 21(2): 
 

 

 
 

Daniele Cicuzza, University Brunei Darussalam, Brunei Darussalam. 
Cheng Wei Chen, Academia Sinica, National Taiwan Normal University, Taipei, Taiwan. 

Peter Boyce, University of Florence, Italy. 
Maria Melanie P. Medecilo-Guiang, Central Mindanao University, Philippines. 

Andrew Powling, University of Portsmouth, Portsmouth, UK. 
Ian M. Turner, Singapore Botanic Garden, Singapore. 

Michael Möller, Royal Botanic Garden Edinburgh, Edinburgh, UK. 
Rugayah, National Research and Innovation Agency, Indonesia.