A JOURNAL ON TAXONOMIC BOTANY, PLANT SOCIOLOGY AND ECOLOGY REINWARDTIA A JOURNAL ON TAXONOMIC BOTANY, PLANT SOCIOLOGY AND ECOLOGY Vol. 13(3): 221 — 3 1 5 , April 11, 2012 Chief Editor KARTINI KRAMADIBRATA Editors DEDYDARNAEDI (INDONESIA) TUKTRIN PARTOMIHARDJO (INDONESIA) JOENI SETIJO RAHAJOE (INDONESIA) TEGUHTRIONO (INDONESIA) MARLINAARDIYANI (INDONESIA) EIZI SUZUKI (JAPAN) JUN WEN (UNITED STATE OF AMERICA) Managing editor HIMMAH RUSTIAMI Secretary ENDANG TRI UTAMI Lay out DEDEN SUMIRATHIDAYAT Illustrators SUBARI WAHYUDI SANTOSO ANNE KUSUMAWAIY Reviewers BRYAN SIMON (AUSTRALIA), EVE J. LUCAS (UNITED KINGDOM), J.F.VELDKAMP (NETHERLANDS), LAUR- ENCE SKOG (USA), PIETER BAAS (NETHERLANDS), RUTH KIEW (MALAYSIA), ROBERT J. SORENG (USA), HE- LENA DUISTERMAAT (NETHERLANDS), LYN A. CRAVEN (AUSTRALIA), RUGAYAH (INDONESIA), MARK HUGHES (UNITED KINGDOM), MARTIN CALLMANDER (USA), PETER C. VAN WELZEN (NETHERLANDS), WAYNE TAKEUCHI (USA), NOBUYUKI FUKUOKA (JAPAN). Correspondence on editorial matters and subscriptions for Reinwardtia should be addressed to: HERBARIUM BOGORIENSE, BOTANY DIVISION, RESEARCH CENTER FOR BIOLOGY-LIPI, CIBINONG 16911, INDONESIA E-mail: reinwardtia@mail.lipi.go.id REINWARDTIA Vol 13, No 3, pp: 235 − 240 235 young floral buds, persistent and conspicuous bracteoles, and a seed coat which adheres to the pericarp in Eugenia. Recent molecular evidence supports a scenario in which these two genera are in fact independent lineages (Biffin, 2005; Widodo, 2010). Direct observation and comparison of all type specimens cited below, as well as of extensive material of Eugenia from the New World, allows the following nomenclatural transfers to be made with confidence. SYZYGIUM Gaertn. Syzygium Gaertn., Fruct. Sem. Pl. 1(1788) 166, t. 33, f. 1, nom. cons. –Eusyzygium Miq., Fl. Ned. Ind. 1, 1(1855) 447, nom. inval. – Lectotype: Syzygium caryophyllaeum Gaertn. (typ. cons.), Ceylon, designated by McVaugh (1956). [= Syzygium caryophyllatum (L.) Alston, pro specim. Ceylon. Eugenia auct. non L. (1753); M.R. Hend., Gard. Bull. Singapore 12 (1949)16; Kochummen, Tree Fl. Malaya 3 (1978)172. Jambosa Adans., Fam. Pl. 2 (1763) 88, 564, orth. cons. (‖Jambos‖, orth. rej.); Rumph. ex DC., Prodr. 3 (1828) 286, isonym. – Eugenia L. subgen. Jambosa (Adans.) Benth. in Benth. & Hook. f., Gen. Pl. 1(1865) 718. – Eugenia L. sect. Jambosa (Adans.) F. Muell., Fragm. Phytogr. Austral. 5 (1865), nom. inval.; Boerlage, Handl. Fl. Ned. Indië 1, 2 (1890) 497. – Eugenia L. sect. Eujambosa Nied. in Engl. & Prantl., Nat. Pflanzenfam. 3, 7 (1893)83, nom. inval. – Type: Jambosa vulgaris DC., INTRODUCTION Syzygium is an entirely Old World genus. In Indonesia and its surrounding areas, many Syzygium species were originally described in Eugenia L. or Jambosa Adans. The generic circumscriptions of Eugenia, Jambosa, Syzygium and some minor satellite genera has had a troubled history with authors frequently transferring species between genera. Alston (1931) reduced Jambosa Adans. (1763) to the synonymy of Syzygium and the former name is now treated as a nomen rejiciendum. The generic concepts of Niedenzu (1893) and Diels (1922) were largely followed by Merrill and Perry (1939) and Ashton (1981) who included in Syzygium not only those Old World species with calyptrate petals (Syzygium s.s.) but also most of the other Old World species with free petals (Jambosa). Generally, this concept has been adopted by the majority of subsequent taxonomists, with the exception of Henderson (1949), and Kochummen (1978), who treated all species as Eugenia s.l., creating significant taxonomic uncertainty. Taxonomic confusion in Eugenia and Syzygium resulted from the considerable overlap of macro- and micro-morphological characters. Currently it is clear that these genera are significantly different. Detailed analysis by Schmid (1972) showed the differences between Eugenia and Syzygium included the evident pubescence of the NEW NOMENCLATURE IN SYZYGIUM (MYRTACEAE) FROM INDONESIA AND ITS VICINITIES Received September 29, 2010; accepted August 01, 2011 PUDJI WIDODO Departemen Biologi, Institut Pertanian Bogor, Bogor, Indonesia. Permanent Address: Fakultas Biologi, Universitas Jenderal Soedirman, Jl. Dr. Soeparno 63, Purwokerto 53122, Indonesia. Email: pwidodo@unsoed.ac.id ABSTRACT WIDODO, P. 2012. New nomenclature in Syzygium (Myrtaceae) from Indonesia and its vicinities. Reinwardtia 13(3): 235−240. ― Current generic concepts in Myrtaceae suggest that nearly all Old World taxa originally described in Eugenia L. and Jambosa Adans. should be accommodated within Syzygium P. Br. ex Gaertn. Six new combinations and a new name in Syzygium are proposed. Complete synonymy and typifications are given. Keywords: Eugenia s.l., Jambosa, Myrtaceae, new combinations, Syzygium, taxonomy. ABSTRAK WIDODO, P. 2012. Tatanama baru pada Syzygium (Myrtaceae) dari Indonesia dan kawasan sekitarnya. Reinwardtia 13 (3): 235−240. ― Konsep generik pada jambu-jambuan (Myrtaceae) saat ini menunjukkan bahwa hampir semua taksa dari Dunia Lama yang semula dipertelakan dalam Eugenia L. dan Jambosa Adans. harus dimasukkan ke dalam Syzygium P. Br. ex Gaertn. Enam kombinasi baru dan satu nama baru dalam Syzygium diusulkan. Sinonimi dan tipifikasi lengkap disajikan. Kata kunci: Eugenia s.l., Jambosa, Myrtaceae, kombinasi baru, Syzygium, taksonomi. REINWARDTIA 236 [VOL.13 nom. superfl. [= Jambosa jambos (L.) Millsp. (typ. cons.). = Syzygium jambos (L.) Alston] 1. Syzygium biniflorum (Ridl.) Widodo comb. nov. Basionym: Eugenia biniflora Ridl., Bull. Misc. Inform. Kew (1925) 80. – Type: Sumatra, Lubok Tandai, 0 m asl. Cybrootes 7103 (Holotype: K!). Small tree, glabrous. Twigs terete and compressed, canaliculated. Leaves elliptic- lanceolate, acuminate cuspidate, 12.5–15 cm long, 4.5–5 cm wide; greenish brown above and yellow- ish brown beneath when dry; leaf base long attenuate or cuneate, subcoriaceous, apex shortly acuminate; venation horizontally paralleled, 30 pairs of veins; petiole slender, 1.5–2 cm long, dark brown to black when dry; midrib depressed, elevated beneath; 12.5–15 cm long, 4.5–5 cm wide; petiole 1 cm long; intramarginal vein 1 mm from margin. Inflorescence a few flowered cyme, usually two per leaf axis and subterminal; peduncle ca. 2 cm long, terete and compressed, canaliculate, dark brown when dry; bracts ovate to acute, persistent, patent, 2 mm long; 1 or 2 flowers per cyme; pseudostipe very short or none; calyxes obconic, 4 mm long; lobes short rounded-ovate; petals calyptrate; stamens up to 20. Notes. This is a small tree in undergrowth of mature forests. It is remarkable for the numerous parallel nerves, and the very few flowers (one or two) in each axillary inflorescence. 2. Syzygium celebicum (Blume) Widodo comb. nov. ― Fig. 1. Basionym: Jambosa celebica Blume, Mus. Bot. 1 (1850) 107. – Eugenia celebica (Blume) Merr., Interpr. Herb. Amboin. (1917) 397. – Type: Celebes, Tondano Herb. Lugd. Bat. Sheet No 898.203- 317 & 318 (Holotype: L!). A tree, height unknown. Twigs terete and compressed below nodes, drying yellowish. Leaves opposite, elliptic-oblong, 10–19 cm long, 3.8–8.25 cm wide; dark purplish brown above and paler below when dry; petiole slender, 0.5–10 mm long, canaliculate above, convex beneath, black when dry; leaf base cuneate, apex acuminate; major lateral veins 6–9 pairs, nerves confluent curved in the intramarginal vein; coriaceous without glandular dots; intramarginal vein 2–5 mm from margin. Inflorescence axillary; peduncle terete, short, drying yellowish; few flowered, subsessile; pseudostipe very short or none; calyx tube subcampanulate, the limb disk-shaped or circular, short, 4-lobes, subequal; corolla subincurved. Stamen numerous, filament filiform. Ovary 2-locules, multiovule. 3. Syzygium horsfieldii (Miq.) Widodo comb. nov. Basionym: Jambosa horsfieldii Miq., Fl. Ned. Ind. 1(1) (1855)420. – Eugenia horsfieldii K. et V. – Type: Java, Surakarta, Horsfield T No 6 (Holotype: K!). ― Fig. 2. A tree, size unknown. Twigs slightly 4-angled, and compressed below nodes, pale brown or grayish when dry. Leaves subopposite, elliptic or elliptic- oblong, acuminate, 10–15.2 long, 4.5–7 cm wide; drying brown above and pale brown below; petiole slender, 4–7 mm long, black when dry, subtereted when young, canaliculated when young, leaf base acute; elliptic or elliptic-oblong; base obtuse– rounded, apex acuminate, glandular dotted, punctuate; midrib pale brown when dry; major lateral veins 6–10 pairs; intramarginal veins 3–5 mm from margin. Inflorescence terminal, few flow- ered (or solitary?); pedicel short (ca. 25 mm long); pseudostipe very short or none; calyxes short, calyx tube campanulate, base short, conspicuously constricted to peduncle junction; lobe 4, inequal. 4. Syzygium korthalsii Widodo, nom. nov. [non S. lanceolatum (Lam.) Wight & Arn., 1834]. ― Fig. 3. Basionym: Jambosa lanceolata Korth. ex Miq., Fl. Ned. Ind. 1(1) (1855) 426; Ned. Kruidk. Arch. 1 (1846) 199. – Jambosa korthalsii Blume, Mus. Bot. Lugd.-Bat. 1 (1849)101, nom. superfl. – Lectotype: Sumatra, Gunung Malintang, Korthals s.n. (Holotype: L, sh. no. 898.203- 346 designated here). A tree, height and diameter not seen. All parts glabrous. Twigs usually 4-angled to winged, with smooth and whitish pale brown bark. Leaves opposite, relatively very long compared to width, lanceolate-linear, 30–45 cm by 2.5–5 cm, brown above and milky brown below when dry; leaf base cordate; leaf apex acute-very acute; petiole ca. 3 mm long, swollen and corky, drying pale brown; midrib furrowed on upper surface, and raised on lower surface, pale brown when dry; major lateral veins ca. 25 pairs, 1–1.5 cm apart, at an angle of 60°–70°, sometimes curved near the midrib and straight near intramarginal veins; minor lateral veins absent or present, oil dots between 2 major lateral veins a few or less than 20 per cm 2 ; intramarginal vein 1 or 2, faint, 1–3 mm from margin. Inflorescence not seen; flower with a pseudostipe 5 –7 mm long; hypanthial cup funnel-shaped; sepals triangular, 5–6 mm long, 5 mm wide; petals not seen; style 35 mm long. Fruits not seen. Notes. S. korthalsii is distinctive from the other Sumatran species in its leaf shape which is linear, 2012] 237 WIDODO: New nomenclature in Syzygium (Myrtaceae) from Indonesia and its vicinities reaching approximately 45 cm long, and only around 3.5 cm wide in average. Korthals (1846) described Jambosa lanceolata based on his collection from the forests of Mt. Malintang (―Melintang‖), W. Sumatra, Indonesia. Blume (1850) incorrectly concluded that this was a mixed collections, and proposed the names J. confusa Blume for material from Java: Blume s.n. (holo: L! sheet no 898.203-342), J. insignis Blume for that from Borneo: Korthals s.n. holo: L! (sheet no 898.203-347), and J. korthalsii Blume for the original collections from Mt. Malintang: Korthals s.n. (Holo L! sheet no 898.203-346). An additional specimen examined is Teijsmann 840 HB (BO!). He also suggested that Korthals‘s name was superflu- ous, as there was already a J. lanceolaria, based on Eugenia lanceolaria Roxb. (1832). Actually, Blume made it himself right here under J. korthalsii. These epithets are similar, but do not mean the same thing [lanceolarius = small (tip of a) spear; lanceolatus = lancet-shaped], and are not confusable under the ICBN. Contrary to Blume‘s opinion, Jambosa korthalsii therefore is a superfluous name and it is Jambosa lanceolata that stands as the correct name that now requires transferral to Syzygium. Currently in Syzygium the combination S. lanceolatum (Korth.) is taken by S. lanceolatum (Lam.) Wight & Arn. (1834). I therefore here propose a new name using Blume‘s illegitimate epithet as is allowed under Art. 58.1 of the ICBN (McNeill et al., 2006). 5. Syzygium suave (Ridl.) Widodo comb. nov. Basionym: Eugenia suavis Ridl., J. Fed. Malay States- Mus. 5 (1915)160. – Type: NE Malay Peninsula, Hills of Kol Samui, May 1913, H.C. Robinson s.n. (Holotype: K!). Big tree to 25 m tall, ca. 40 cm diameter. Bark grayish brown, peeling off in irregular strips 3–5 mm thick, 3–10 cm broad, 15–25 cm long. Sap- wood light brown, heartwood dark brown. Leaves oblong, 9–15 cm long, 2.5–4 cm wide; upper surface dark green, light green beneath; leaf drying grayish above and pale brown below; petiole slender, 10–15 mm long, canaliculate above, pale to dark brown when dry; base acuminate or obtuse, apex acuminate. Major lateral veins 9–14 pairs. Intramarginal vein 1–2 mm from margin. Flowers accented whitish. Peduncle branches and pedicel sharply 4-angled, pale brown when dry; hypanthial cup narrowly to broadly funnel-shaped; pseudostipe short or none. Calyxes yellowish white, especially the latter red tinged. Corolla white, stamen white, anthers yellow. 6. Syzygium sumatranum (Miq.) Widodo comb. nov. ― Fig. 4. Basionym: Jambosa sumatrana Miq., Fl. Ned. Ind. 1(1) (1855) 419. – Type: Sumatra, upper Angkola, Junghuhn s.n., Herb. Lugd. Bat No 898.203- 374 & 376 (Holotype: L!). A tree, size unknown. Twigs subterete, and slightly 4-angled-compressed below nodes, pale brown when dry. Petioles canaliculate, slender 4–6 mm long, yellowish brown when dry. Leaves opposite sometimes subopposite; elliptic or oblong sub abrupt acuminate, 7.5–13 cm long, 3.5–5.8 cm wide; drying dark brown above and reddish brown below; leaf base obtuse, without glandular dots; midrib sulcate above, paler beneath (brown when dry); major lateral veins 6–9 pairs; intramarginal vein 2–4 mm from margin. Inflorescence terminal, rarely axillary; peduncle terete, yellowish when dry. Flowers usually sessile; calyxes turbinate-clavate 2 – 4 mm long, subequal, when young. 7. Syzygium valetonianum (King) Widodo comb. nov. Basionym: Eugenia valetoniana King, J. Asiat. Soc. Ben- No Characters J. korthalsii J. confusa J. insignis 1 Twigs 4-angled to 4-winged, with smooth and whitish pale brown bark. terete and slighty compressed near the nodes. 4-angled. 2 Leaf form and size lanceolate-linear, 30 – 45 cm by 2.5 – 5 cm oblong-lanceolate, tapered gradually from below the middle to apex, 20 – 44 cm by 3 – 5 cm ovate-oblong or lanceolate, long acuminate, 4 – 10 cm by 1 – 2.75 cm 3 Leaf base cordate broadly obtuse or rounded rounded or subcordate 4 Locality Sumatra, Mount Malintang Java Borneo, Martapura Table 1. Morphological differences between J. korthalsii, J. confusa, and J. insignis REINWARDTIA 238 [VOL.13 gal, Pt. 2, Nat. Hist. 70 (1901)112. – Type: Malay Peninsula, Perak Larut 300 – 500 m alt. King s.n. (Holotype: K!). A tree, 20–25 m tall. Young branches thicker than a crow-quill, terete, the bark pale brown, flaky. Leaves elliptic–oblong, 7–10 cm long, 3–4 cm wide; both surfaces dark purple to blackish when dry; base cuneate-obtuse, apex acuminate or obtuse; petiole slender 10–15 mm long, canaliculate above, brown when dry; major lateral veins 8–12 pairs; intramarginal vein 1–2 mm from margin. Inflorescence terminal or axillary, paniculate, shorter than the leaves, with very short, terete, few flowered cymose branches, or in a small pedunculate cymes; peduncle terete, blackish when dry. Flowers sessile, calyxes ca. 7.5 mm long, hypanthial cup clavate-campanulate, constricted into a short pseudostipe; the wide mouth truncate, or with 4-deciduous, short, rounded lobes. Petals small, orbicular, deciduous. Fruits when unripe pyriform, when ripe globular, crowned by the wide calyx-limb, ca. 12 mm long. Notes. A species close to E. griffithii, and also close to E. clarkeana, and E. koordersiana, but with much smaller panicles and fruit at first pyriform. ACKNOWLEDGEMENTS I have been supported by a grant from the Directorate General of Higher Education, the Republic of Indonesia. I would like to thank Prof. Dr. Alex Hartana, Prof. Dr. Mien A. Rifai, and Dr. Tatik Chikmawati for advice. Herbarium Bogoriense (BO), National Herbarium of Netherlands (L) and Herbarium Kewense (K) are thanked for permission to study the collections deposited in these institutions. My special thanks to Prof. Peter van Welzen (L) for assistance and for critically reviewing some pa- pers. I would like to express my gratitude to Dr. Jan-Frits Veldkamp (L) for assistance and advice. To Lyn A. Cra- ven (CANB) and Prof. John Parnell (TCD) for advice and references. REFERENCES ALSTON, A. G. H. 1931. Myrtaceae. In: Supplement to the Handbook to the Flora of Ceylon 6: 109–121. Dulau, London. ASHTON, P. S. 1981. Myrtaceae. In M.D. Dassanayake, A revised handbook to the Flora of Ceylon 2: 403– 473. Smithsonian Institution & National Science Foundation, Washington (DC). BIFFIN, E. 2005. Sorting out the confusion: Phylogenetics of large genera and the lessons from Syzygium (Myrtaceae). Austral. Biol. Resources Study. Biologue 30. CSIRO Plant Industry, Canberra. Internet publication: http://www.environment.gov.au/ biodiversity/abrs/publications/other/biologue / 3 0 / index.html#syzygium BLUME, C. L. 1850. Jambosa korthalsii Blume Mus. Bot. 1: 107. DIELS, L. 1922. Die Myrtaceen von Papuasien. B o t a n i s c h e J a h r b ü c h e r f ü r S y s t e m a t i k , Pflanzengeschichte und Pflanzengeographie. 57: 356 –426. HENDERSON, M. R. 1949. The genus Eugenia (Myrtaceae) in Malaya. Gardens’ Bulletin Singapore 12: 1–293. KOCHUMMEN, K. M. 1978. Eugenia. In F.S.P. Ng, Tree flora of Malaya 3: 172–247. Longman, London. KORTHALS, P. W. 1846. Jambosa lanceolata Korth. Ned. Kruidk. Arch. 1:199. MCNEILL, J., BARRIE, F. R., BURDET, H. M., DE- MOULIN, V., HAWKSWORTH, D. L., MARHOLD, K., NICOLSON, D. H., PRADO, J., SILVA, P. C., SKOG, J. E., WIERSEMA, J. H. & TURLAND, N. J. 2006. International Code of Botanical Nomenclature (Vienna Code). (Regnum Vegetabile 146). Gantner, Ruggell, Liechtenstein, xviii + 568 p. URL: http:// ibot.sav.sk/icbn/main.html MERRILL, E. D. & PERRY, L. M. 1939. The Myrtaceous genus Syzygium Gaertner in Borneo. Mem. Amer. Acad. Arts 18 (3): 135–202. Reprinted in Mem. Gray Herb. Harvard Univ. 4. NIEDENZU, F. 1893. Myrtaceae. In A. Engler and K. Prantl, Die natürlichen Pflanzenfamilien 3, 7: 78–86. Engelmann, Leipzig. SCHMID, R. 1972. A resolution of the Eugenia- Syzygium controversy (Myrtaceae). American Journal of Botany 59 (4): 423–436. WIDODO, P. 2010. Enumeration of Sumatran Syzygium (Myrtaceae). Unpublished Dissertation. Institut Pertanian Bogor. Bogor. 2012] WIDODO: New nomenclature in Syzygium (Myrtaceae) from Indonesia and its vicinities 239 Fig. 1. Syzygium celebicum. Photo: P. Widodo. Fig. 2. Syzygium horsfieldii. Photo: P. Widodo. 240 REINWARDTIA [VOL.13 Fig. 3. Syzygium korthalsii Photo: P. Widodo. Fig. 4. Syzygium sumatranum. Photo: P. Widodo. 314 REINWARDTIA [VOL.13 ERRATUM REINWARDTIA Vol. 13, Part 2, 2010 1. Please change the existing word in p. 213, LINE 7 on ABSTRAK (written in Bahasa Indonesia version) with the following: Keberadaan dua jenis terakhir melampaui distribusi yang sebelumnya hanya diketahui di barat garis Wallace. 2. Please change the existing epithet name in p, 214, COLUMN 1, LINE 40 on Key to the species of Marantaceae in Sulawesi number 5.a. after Phrynium: longispicum INSTRUCTION TO AUTHORS Reinwardtia is a scientific journal on plant taxonomy, plant ecology, and ethnobotany. Manuscript intended for a publication should be written in English represent an article which has not been published in any other journal or proceedings. Every manuscript will be sent to two blind reviewers. Two printed copies (on A4 paper) of the manuscript of not more than 200 pages together with an electronic copy prepared on Word Processor computer program using Time New Romance letter type and saved in Rich Text File must be submitted. For the style of presentation, authors should follow the latest issue of Reinwardtia very closely. Title of the article should be followed by authors name and mailing address in one-paragraphed English abstract of not more than 250 words. Keywords should be given below each abstract. On a separated paper, author(s) should send the preferred running title of the article submitted. Taxonomic identification key should be prepared using the aligned couplet type. Strict adherence to the International Code of Botanical Nomenclature is observed, so that taxonomic and nomenclatural novelties should be clearly shown. Latin description for new taxon proposed should be provided and the herbaria where the type specimens area deposited should be presented in the long form that is name of taxon, authors name, year of publication, abbreviated journal or book title, volume, number and page. Map, line drawing illustration, or photograph preferably should be prepared in landscape presentation to occupy two columns. Illustration must be submitted as original art accompanying, but separated from the manuscript. On electronic copy, the illustration should be saved in jpg or gif format at least 350 pixels. Legends or illustration must be submitted separately at the end of the manuscript. Bibliography, list of literature cited or references follow the Harvard system. REINWARDTIA Vol. 13. No. 3. 2012 CONTENTS Page W.J.J.O. DE WILDE & BRIGITTA E.E. DUYFJES. Trichosanthes (Cucurbitaceae) in Malesia: additions and corrections, including a new species and a new variety 221 DEDEN GIRMANSYAH. Two new species of Begonia (Begoniaceae) from Bukit Tiga-puluh National Park, Riau, Sumatra 229 PUDJI WIDODO. New nomenclature in Syzygium (Myrtaceae) from Indonesia and its vicinities 235 ALEX SUMADIJAYA & JAN FRITS VELDKAMP. Non-Bambusoid Grasses (Gramineae) from Raja Ampat Archipelago, Papua Barat Province, Indonesia 241 ARY PRIHARDYANTO KEIM. New variety, records & discoveries of some species of Pandanus (Pandanaceae) in Sumatra and Kalimantan, Indonesia 255 HARRY WIRIADINATA. A new species of Begonia (Begoniaceae) from Sagea Lagoon, Weda Bay, Halmahera Island, North Moluccas, Indonesia 263 ARY PRIHARDYANTO KEIM. The Pandan flora of Foja-Mamberamo Game Reserve and Baliem Valley, Papua-Indonesia 271 JAN FRITS VELDKAMP. Koordersiochloa Merr. (Gramineae), the correct name for Streblochaete Hochst. exPilg. 299 SRI ENDARTI RAHAYU, KUSWATA KARTAWINATA, TATIEK CHIKMAWATI & ALEX HARTANA. Leaf anatomy of Pandanus species (Pandanaceae) from Java 305 Reinwardtia is a LIPI acredited Journal (258/AU 1/P2MBI/05/2010) Herbarium Bogoriense Botany Division Research Center for Biology- LIPI Cibinong, Indonesia dpn 444-651-2-PB blkng 451-665-3-PB.pdf dpn 451-665-2-PB_Page_3 blkng