REINWARDTIA Vol. 22. No. 1. pp: 31‒35 DOI: 10.55981/reinwardtia.2023.4567 31 THE TYPIFICATION OF GIGANTOCHLOA TALUH WIDJAJA & ASTUTI (POACEAE, BAMBUSOIDEAE) Received March 30, 2023; accepted May 17, 2023 I PUTU GEDE P. DAMAYANTO Plant Biology Graduate Program, Department of Biology, Faculty of Mathematics and Natural Sciences, IPB Univer- sity. Jln. Raya Dramaga, Kampus IPB Dramaga, Bogor 16680, Indonesia. Herbarium Bogoriense, Research Center for Biosystematics and Evolution, National Research and Innovation Agency (BRIN). Jln. Raya Jakarta-Bogor Km 46, Cibinong, Bogor 16911, Indonesia. Email: iput004@brin.go.id https://orcid.org/0000-0001-8740-0696. HIMMAH RUSTIAMI Herbarium Bogoriense, Research Center for Biosystematics and Evolution, National Research and Innovation Agency (BRIN). Jln. Raya Jakarta-Bogor Km 46, Cibinong, Bogor 16911, Indonesia. Email: himmah.rustiami@brin.go.id https://orcid.org/0000-0002-2260-250X. MIFTAHUDIN Department of Biology, Faculty of Mathematics and Natural Sciences, IPB University. Jln. Raya Dramaga, Kampus IPB Dramaga, Bogor 16680, Indonesia. Email: miftahudin@apps.ipb.ac.id https://orcid.org/0000-0002-5641-1090. TATIK CHIKMAWATI Department of Biology, Faculty of Mathematics and Natural Sciences, IPB University. Jln. Raya Dramaga, Kampus IPB Dramaga, Bogor 16680, Indonesia. Email: tatikch@apps.ipb.ac.id https://orcid.org/0000-0001-9085-7590. ABSTRACT DAMAYANTO, I. P. G. P., RUSTIAMI, H., MIFTAHUDIN & CHIKMAWATI, T. 2023. The typification of Gigan- tochloa taluh Widjaja & Astuti (Poaceae, Bambusoideae). Reinwardtia 22(1): 31‒35. — The name of Gigantochloa taluh Widjaja & Astuti (Poaceae, Bambusoideae) was invalid because of failure to meet the requirements of the Inter- national Code of Nomenclature for Algae, Fungi, and Plants, that there was no herbarium location of type specimens was mentioned. A typification was provided here to address this issue. Key words: Bamboo, Gigantochloa taluh, holotype, typification. ABSTRAK DAMAYANTO, I. P. G. P., RUSTIAMI, H., MIFTAHUDIN & CHIKMAWATI, T. 2023. Tipifikasi Gigantochloa taluh Widjaja & Astuti (Poaceae, Bambusoideae). Reinwardtia 22(1): 31‒35. — Nama Gigantochloa taluh Widjaja & Astuti (Poaceae, Bambusoideae) dinyatakan tidak sah karena tidak memenuhi persyaratan Kode Tata Nama Inter- nasional untuk alga, jamur, dan tumbuh-tumbuhan, yaitu tidak menyebutkan lokasi herbarium dari spesimen tipe. Tipi- fikasi diberikan untuk mengatasi masalah ini. Kata kunci: Bambu, Gigantochloa taluh, holotipe, tipifikasi. INTRODUCTION Gigantochloa taluh Widjaja & Astuti (2004: 203) was first described in Reinwardtia volume 12(2) (Widjaja et al., 2004). Vorontsova et al. (2016) in their bamboo checklist assigned this taxon as hav- ing the status of “unplaced name”. After observing the specimen, scrutinizing the protologue, and reading relevant literature, we noticed that the name G. taluh was invalid due to it being contrary to Article 37.6 of the International Code of Botani- cal Nomenclature (ICBN) (Greuter et al., 2000), which was in force when the name was originally published. Article 37.6 of ICBN indicated that for the name of a new species or infraspecific taxon published on or after 1 January 1990 of which the type is a specimen or unpublished illustration, the single herbarium or collection or institution in which the type is conserved must be specified (Greuter et al., 2000). The name G. taluh is un- placed or cannot be put into synonymy because no type material is indicated. Therefore, it cannot be established to which species the name applies (POWO, 2023). This circumstance also aligns with the latest version of the International Code of No- menclature (ICN) for Algae, Fungi, and Plants, where G. taluh is contrary to Article 40.7 (Turland et al., 2018). Widjaja et al. (2004), did not mention http://dx.doi.org/10.14203/reinwardtia.v19i1.3850 mailto:himmah.rustiami@brin.go.id mailto:miftahudin@apps.ipb.ac.id mailto:tatikch@apps.ipb.ac.id 10.55981/reinwardtia.v22i1.4567 https://orcid.org/0000-0001-8740-0696 https://orcid.org/0000-0002-2260-250X https://orcid.org/0000-0002-5641-1090 https://orcid.org/0000-0001-9085-7590 https://dx.doi.org/10.55981/reinwardtia.v22i1.4567 https://orcid.org/0000-0001-8740-0696 https://orcid.org/0000-0002-2260-250X https://orcid.org/0000-0002-5641-1090 https://orcid.org/0000-0001-9085-7590 REINWARDTIA 32 [VOL.22 Fig. 1. Screen capture of Tropicos (2023) database portal and specimen of Inggit Puji A stuti EA W IP 456 (BO) with zoom-in on BO number and specimen label. DAMAYANTO et al.: Typification of Gigantochloa taluh Widjaja & Astuti 2023] 33 the herbarium where the type specimen of G. taluh was deposited. The original citation of the type specimen was less accurate as the collector’s name of the type specimen was not mentioned (see Article 9.2 ICN of Turland et al., 2018). Here, we provide the typification of G. taluh. Typification in plant taxonomy studies refers to the process of prescribing type (Rao, 2017). A nomenclature type (typus) is that element to which the name of a taxon is permanently attached (see Article 7.2 of Turland et al., 2018). Typification helps determine the correct application of taxa names and establishes nomenclatural types (see Turland et al., 2018). Typification can be pu- blished independently with the aim of providing valid and accepted taxon names as soon as possi- ble (see de Oliveira et al., 2009; Haevermans et al., 2013) or this is typically done during taxo- nomic revisions (see Vorontsova et al., 2013; Veldkamp, 2016), the preparation of synopsis (see Essi et al., 2017), monographs (see Snow et al., 2018), checklists (see Kellogg et al., 2020), pro- posals for conserving the taxon name (see Greuter and Rodríguez, 2015), the resurrection or rein- statement of submerged taxa (see Nobis et al., 2022), and the presentation of taxon name combi- nations (see Banfi et al., 2017). In Indonesian bamboos, several typifications have been carried out, such as Widjaja’s (1987) revision of the ge- nus Gigantochloa in Malesia, which also involved typification. Widjaja and Wong (2016) also per- formed typification when making a new combina- tion in the genus Chloothamnus in Malesia, and Widjaja (2020) conducted typification on the Fim- bribambusa soejatmiae Widjaja & Ervianti pu- blished by Ervianti et al. (2019), which has two different collection numbers of type specimens. Typification of G. taluh is required in this study to support the author’s ongoing bamboo taxonomic research in the Lesser Sunda Islands. MATERIALS AND METHODS Herbarium specimens of Gigantochloa taluh stored in Herbarium Bogoriense (BO) and Herba- rium Hortus Botanicus Baliensis (THBB) were gathered for observation. In addition, a photo- graph of a type specimen of G. taluh in the proto- logue was also consulted for comparison. TYPIFICATION GIGANTOCHLOA TALUH Widjaja & Astuti, Reinwardtia 12(2): 203. 2004. Type: Bali, Baturiti, Ekakarya Botanical Garden, Inggit Puji Astuti EAW IP 456 (holotype BO!, code: BO-1606188; isotype perhaps K [not seen]). Distribution. Endemic to Bangli, Bali and culti- vated in “Eka Karya” Bali Botanical Garden in Tabanan, Bali. Notes. Our r esear ch r evealed some inter esting findings regarding the type specimens of G. taluh. We found a discrepancy in the G. taluh herbarium holotype code on the Tropicos (2023) database portal, which should be BO-1606188, not BO- 1606183 (Fig. 1A, red arrow and 1B, BO number zoomed-in). In addition, the information on the label of the Inggit Puji Astuti’s specimen EA W IP 456 (Fig. 1B, label zoomed-in) mentions that a duplicate was sent to K. However, this is not listed in the Widjaja’s et al. protologue (2004). Unfortu- nately, we were unable to find scanned images of the Inggit Puji Astuti’s specimen EA W IP 456 in K through the GBIF (2023), JSTOR (2023), and KEW HERBARIUM CATALOGUE (2023) data- base portals. It should be noted that there may be duplicates of this specimen in K, yet this specimen in K may not be a current priority for scanning and online publication, or there was a typo in writing the label and it was never sent to K. In normal practice, when duplicates of specimens are distri- buted to other herbaria, the acronym(s) of recipient herbaria on the label is marked (checked or cir- cled). However, on the label of Inggit Puji Astuti’s specimen EA W IP 456 (Fig. 1B, label zoomed-in), the herbarium acronym is not marked, raising doubts about the shipment of duplicate of this specimen to K. Mrs. Inggit Puji Astuti (pers. comm., 2023) stated that the specimen EA W IP 456 was collected during the beginning of her study of bamboo. Unfortunately, she does not know if specimen EA W IP 456 was sent to K. This doubt is supported by information provided by Is- mail Apandi (pers. comm., 2023), the loan officer of BO who responsible for sending and receiving specimens, that he does not have any data regard- ing the shipment of Inggit Puji Astuti’s specimen EAW IP 456 to K or other herbaria. Therefore, it is possible that Inggit Puji Astuti’s specimen EA W IP 456 was never sent to K. Nevertheless, in this article, we are constrained by limited evidence and inconclusive interpretation and decide to write that the isotype of Inggit Puji Astuti’s specimen EA W IP 456 probably exists in K due to the information provided by the specimen label. In addition, the collection number of Inggit Puji Astuti’s specimen EA W IP 456 is also interesting because the additional code “EAW” is attached, referring to the abbreviation of the name of a re- tired senior bamboo researcher at BO, Prof. Dr. Elizabeth A. Widjaja. In the Widjaja et al. (2004), the previous label attached to the type specimen of G. taluh had IP as the sole collector. Thus, we pre- sume that the “EAW” was added later to a new label due to one reason or another and present to date. REINWARDTIA 34 [VOL.22 ACKNOWLEDGEMENTS We would like to express our sincere gratitude to the Director of Scientific Collection Manage- ment (Herbarium Bogoriense and Herbarium Hor- tus Botanicus Baliensis of “Eka Karya” Bali Bo- tanical Garden), National Research and Innovation Agency for granting us access to the specimens of Gigantochloa taluh. IPGPD want to thank Gusti Made Sudirga (“Eka Karya” Bali Botanical Gar- den) for his invaluable assistance, as well as, Inggit Puji Astuti (Bogor Botanical Garden) and Ismail Apandi (Herbarium Bogoriense) for the valuable information provided. We would like to express our sincere gratitude to the anonymous blind reviewers who provided their insightful comments and feedback. The author gratefully acknowledges Liam A. 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