A JOURNAL ON TAXONOMIC BOTANY,
PLANT -SOCIOLOGY AND ECOLOGY
REINWARDTIA
. Editors
MIEN A.
KIJS WAT A KARTAWINATA
N. WULIJARNI-SOETJIPTO
1 Published by '
HERBARIUM BOGORIENSE
LEMBAGA BIO-LOGI NASIONAL — LIPI
BO.SOR, INDONESIA
Eeinwardtia Vol. 9, Part 1, 1 —182 31 December 1974
10- ISSN 0(f34-365X
REINWARDTIA [VOL. 9
1/2-3/4 the leaf length, filiform, prominulous, reticulation regular, dense,
very minute; sometimes between-midrib and basals deep doniatia (slightly
hullate on the upper surface). Petiole slender, 6—10 mm long, laxly
sericeous, concave above. Inflorescences axillary, consisting of a very
short (1—2 mm) reduced main peduncle, bearing up to 5 flowers;
sometimes the flowers practically sessile. Pedicel filiform, 5—8 mm,
densely pilose. Tube broadly funnel shaped, 0.5 m, inside densely
sericeous. Tepals erect-patent, narrowly ovate, acute, stiff, 1.5—2 ram
long, densely sub-sericeous on both sides. Stamens 1.25 mm long, inserted
slightly below the tepals on the rim of the 0.5 mm broad tube; anthers
quadrangular or ovate-quadrangular, obtuse, slightly shorter than the
almost glabrous, thickish filaments; outer anthers with introrae upper
and lateral lower, large, slanting cells, almost in one plane, inner anthers
smaller, the lower large cells extrorse, the upper smaller ones lateral;
glands large, long-stalked. Staminodes sub-spathulate or oval on a thick
long filament. Ovary ellipsoid-ovoid, merging into a short, thick style
•with small stigma. Fruit ellipsoid to sub-globose, up to 6 x 10 mm,
apiculate. Cup fleshy, broadly funnel shaped, up to 6 mm long, the top
shallowly excavated, the persistent, sericeous tepals erect, 3 mm long,
2 mm wide at the base; pedicel 1 cm long, slender.
I have not seen the material, cited by Yang, but the description
fits N. delayayi.
CHINA. Yunnan, fl., Forrest lieea (BO, E ) ; Pcc-eha-ho, April, fl., Dda-ony s.n.
(BO, P ) ; Tenggueh, alt. 1E00 m, March, buds, fr., Farrezt 9 f « (BO, E) ; Yung-jen
Hsien, in ravine, alt. 1700 m, May, fl., Tsai 62891 (A, BO); Pa-ta-ouan, near Pee
tchouan, July, fl., Jean Py (Dueloux $292) (BO, P ) ; forest of Thou Ty, fl., Simeon
Ten (BO, P ) ; locality not indicated, fr., McLaren's Coll. U«c (BO, K).
RE I N W A R D T I A
Published by Herbarium Bogoriense — LBN,
Vol. 9, Part 1, pp. 97 —115 (1974)
MATERIALS FOR A REVISION OF LAURACEAE IV*
A. J. G. H. KOSTEBMANS
c/o Herbarium Bogoriense ,Bogor,Indonesia
ABSTRACT
In the genus Actinodaphtie Nees one new combination and one new
species ace presented; in LAndera Thunb. one new combination, 4 new
species and one new forma; in lAtsea Lam. one species has been reduced
to synonymy, one nomen novum is presented, 5 new species are described
and of 6 species new cecords are given; one species has been reinstated.
The obscure Mackilus sericea Bl. is a3sumed to be conspecific with
Persea bombyeina (King ex Hook, f.) Kosterm. In the genus Persea Mill.
one nomen novnm, two new combinations and one new species are created
and proposed. In the genus Phoebe Nees one new combination and two
ABSTRAK
DJIISJYI uiJiriTE ^ctijiodtiohns Neea satu korabinasi baru dan ssttu
jenis bum diusulkan; dalam Lindera Thunb. satu kambinasi baru, empat
jenis diakut kembalF dan daerah penyebaran enam jenis diperluaa oleh
terkumpulnya spesimen-speaimen baru. Machilus sericea Bl. dianggfap
aama dengan Perttea bombyema (King ex Hoof.f.) Kosterm.; da]am.
baru diciutakan Dalam matsa Pho&bG Nees satti kombinaai baru dasi du&
ienie baru diuaulkan.
ACTINODAPHNE Neea
Actinodaphne forrestii (Allen) Kosterm., comb. & stxtt. nov.
Actinadaphne reticuleta vac. forrestii Allen in Annals Missouri Bot. Gard.
25: 412. 1SSB. - Typus: Forrest 1S827 (AA, K|.
The species differs from A. retioulata, by the larger and thicker
leaves, which are not reticulate, but only show some "fielded" pattern,
the brown sublanuginose tomentum and the much shorter fruit pedice!-
Part I appear^ m Reinwardtia 7: 201-356. 19GE; part II in ibid. 451-635. 1969;
pact III in ibid. 8: 21-106. 1070.
REINWARDTIA [VOt. 9
Actinodaphne longipes Kosterm., spec. nov.
Aetijiodapkne reHculata var. glnbra Meissner in DC, Frodr. 15(1): 213. 1864;
Liou Ho, Laur. Chine et Indochine 158. 1932 and 1934; Allen in Ann. Missouri bot,
Gard, 25: 412. 1938; Kostermans, Bib]. Laur. 38. 1964. — Typus: J.S. Smith s.rc. (K).
Pedieellus fructiferus usque ad 4 cm longus; ramulis glabris.
Meissner's variety glabra was based on a specimen, collected by
Sir J.S. Smith. The provenance is obscure, Nepal with an interrogation
mark is written on the sheet at Kew. Hooker thought, that this specimen
was glabrous because of its older stage and thought, that the specimen
was collected in Khasia.
The type specimen shows shoots with a young stage of leaf develop-
ment and subsequently collected material, proves, that the plant is indeed
practically glabrous. Moreover, the leaves are not reticulate, but scrobi-
culate with a finer mesh thai) those of A. reticulata. Hooker was right,
that the specimen of J.S. Smith was from E. Bengal.
ASSAM. Paona, Naga Hills, 5800 ft. alt., Sept., fr., Bar 6%76 (K); Darjeelinsr,
Dilpa, alt. 6500 ft., Oct., fr., Otrmaston e.M. (K); E. Bengal, J.S. Smith s.n. (K).
LINDEEA Thunb.
Lindera aggregata (Sims) Kosterm., comb, nov,
Lauras aggregata John Sims in Curtis Bot. Mag. 51: tab. 21117 (1824);
Koatermans, Bibl. Laur.: 663 (1964).
Lindera etrycknifolia (S. & Z.) Villar in Blanco, Fl, Filip, ed. 3, Nov. App.: 182
(1880); Kostermans, I.e.: 773. — Dapknidium myrrha Sieb. & Zuec. (non Neesj
ex Meissn. in DC, Prodr. 15(1): 23 (1864); Kostermans, I.e.: 462. - Daphwidiu.'m
strychy.ifoliv.rn Sieb. & Zucc, Fl. Japon. Fern, nat. 2: 83 (1846); Kostermans, I.e.:
4SS. — Cinnamon™™ verlotii De Lukmanoff, Nomend, Iconogr. Canelliers ct Camphr.:
19, tab. 14, fig. 142 (1889); Kostermans, U,; S60.
In the description of Sims of this plant, it is mentioned, that it
was sent to England from Macao, China by Reeves in 1821 on a vessel,
named the Orwell, under Captain Lindsay (Sweet, Hort. Brit. 344. 1827
gives the date of introduction 1812, apparently a misprint).
No original material could be discovered so far, although a rather
recent collection of the Kew Gardens, is still labelled Laurus aggregata.
The plate is sufficiently detailed to make sure, that this is the shrub,
commonly called Lindera stryekmfolia. Loudon (Encyclop. 332, fig. 1855)
gives a crude copy of this plate.
The species is very close to Lindera sinensis (Bl.) Kosterm. of the
same provenance and also introduced by Reeves, the differences being
only in the size of the leaves and the glabrousness of L. sinensis.
1974] KOSTERMANS: Re of Lauraeeae IV
The type material of Cinnamomum verlotii de Lukmanoff is conserved
in Paris and labelled: cultivee au Musee sous le nom de Lawrus gbiueo-
pkylla, no. 90; it is a sterile specimen.
Lindera eberhardtii Lecomte {in Nouv. Arch. Mus., Ser. V, 5: 115.
1913) may belong here or it may be L. sinensis. The young shoots of
the material of L. eberhardtii are long-silky (not stated by Lecomte).
Lindera caesia, as accepted by Allen, is certainly L. sinensis.
In Paris a collection of Callery (no. 61) is from the type locality
(Macao).
Lindera delkata Kosterm., spec. nov.
Arbuscula? ramulis gracills glabris foliis alternantibus tenuiter
chartaceis glabris lanceolatis caudato-acuminatis basi acutis utrinque sub
lente prominule reticulatis opacis costis sat obscuris petiolis gracilis supra
eanaliculatis inflorescentiis pergracilis umbellulis paucis longe et gracile
pedunculatts glabris floribus dense sericeis.
TYPUS: Van Steenis 9880 (L).
Shrub ( ? ) ; branchlets slender, glabrous, smooth. End bud sericeous.
Leaves spirally arranged, thinly chartaceous, glabrous, lanceolate,
2 x 6—3.5 x IS cm, caudate-acuminate, base acute, both surfaces dull,
under the lens prominulously reticulate, upper surface with hardly
prominulous midrib, gradually impressed towards its base, lateral nerves
obscure, lower one with prominent midrib, the ca 10—15 pairs of lateral
nerves rather patent, obscure, hardly differentiated from the reticulation.
Petioles slender, 3—5 mm, channeled above. Inflorescences very slender,
up to 5 mm long, ending in a sericeous end bud, with one or two umbels,
or the inflorescence completely suppressed, umbel peduncles slender,
7—10 mm long. Florets densely sericeous, pedicels 2 mm; tepals narrow,
acute, 1.5 mm; stamens longer than the tepals, anthers ca 0.5 mm long.
Related to L. oxyphylla, but the leaves reticulate, caudate-acuminate,
shorter petioled; the inflorescences different.
SUMATRA. Atjeh Prov., Gajo Lands, S. of Blang Kadjeren, at the confluence
of rivers Kepi and river Auan, flat ridftc, alt. 1100 m, March, fl., Van Steenis 9SS9
(BO, L); perhaps also: Mt. Sago near Pajakumbuh, W. Sumatra, alt. 1500 m, ster.,
Meyer S905 (L).
Lindera moiUanoides Kosterm., spec. nov.
Arbor mediocris ramulis dense minute sublanuginoso velutinis foliis
alternant!bus chartaceis ovato-ellipticis acuminatis basi in petiolum
brevem contract!a supra glabra nitida minute dense reticuiata vel
sublaevia nerviis majoribus impressis subtus glauca perlaxe minute piiosa
REINWARDTIA [VOL. 9
nerviis majoribus prominentibus dense pilosis inflorescentiis aessilis
paueifloris innovation!bus perulatia sericeis fructus globosus pedicello
vix obeonico gracile piloso insidens.
TYPUB: Endert U57 (L).
Tree, 5—16 m high, 5—20 cm diam. Branchlets densely, minutely
rusty sublanuginose pilose. Leaves spirally arranged, in one plane,
chartaceous, ovate-elliptic, 3 x 6—5 x 14 cm, acuminate, base contracted
into the slender, pilose, 5—7 mm long petiole; both surfaces minutely
densely reticulate, upper one glossy, glabrous, the main nerves impressed,
lower one sparsely, minutely pilose on the veins, glaucous, midrib and
the 3—4 pairs of curved, ascendent lateral nerves densely pilose,
prominent, secondary nerves lax, parallel, subtiorizontal, prominulous,
the basal pair of lateral nerves may reach % of the leaf lamina.
Inflorescences axillary, sessile, few-flowered, surrounded by numerous
large, sericeous bracts. Fruit globose, up to 5 mm diam., red, on a slender,
slightly obconical, 5—7 mm long pedicel with the whithered flower parts
persistent.
MALAY PENINSULA. Pahatig, Cameron Highlands, valley of Bertam about Kuala
Habu, April, buds, Kep. SCSOS (L); ibid., G. Jasar, alt. 1600 m, tree 7 m, diam. 8 cm,
Oct., fl. yellowish green, Clie-ie Wee Lek 936 (L, SING); Pahang, Masu Hill, 1250 m
alt., tree 3 m, ster., Nur s.n. <K); Selangor, Sempang, April, fr., Ridley 1559T <K|.
SUMATRA. N. slope of Mt. Sago, Fajakumbuh, alt. 900 -1200 m, tree 15 m diam.
12 cm, bark yellow-brown with small tubercles, living bark 8 mm, dirty yellow
with brown spots, ster., Meyer 3536 (L); ibid., tree 3 m, Meyer SJ,10 (L); ibid,
alt. 900 m, Pebr., fr., Meyer U65 (L). BOKNEO. Sabah, Kinabalu, Ranau Distr.,
Sosopodon, alt. 1700 ro, tree 5 m, diam. 5 cm, bark brown, living bark yellow, wood
white, Febr., buds, San 4S819 (L); Kundasang, alt. 1700 m, treelet 1,5 m, Febr., buds,
San 29012 (K, L ) ; ibid., Liwago-Sosopodon Reserve, alt. 1E0O m, tree 13 m, diam.
20 cm, bark smooth, brown, living bark light brown, sapwood light yellowish, Febr.,
buds, San 1*7985 (K, L ) ; Gurulau Spur, alt. 2000 m, tree 16 m, diam. 10 cm, Nov.,
buds, /. & M.S. Clemens 50S8!) (K, L); Tenompok, 5000', ster., Clemens S840S (K) ;
Sosopodon-Kundasang, alt. 1800 m, tree 13 m, diam. 10 cm, bark purplish, inner bark
violet, wood white-yellowish, Nov., buds yellowish. Son S313S (K, L ) ; Ranau, above
Halting Bungalow Kinabalu near Kundussng, Sosopodon, alt. 1500 m, tree 15 m.
bole 3 m, diam 20 cm, bark 14 inch, soft., Apr., fl., Sow 33778 (K); Mesilau R.,
alt. 1700 m, tree 7 m, lower leaf surface glaucous, Jan., buds pink, RSNB i>17
(K, L); ibid., tree 13 m, Febr., buds, RSNB i283 (K, L); Sarawak, Bukit Mersing,
Anap, basalt ridge, submontane forest, alt. 1000 m, tree 8 m, diam. 5 cm, Sept., fr.,
S. 21970 (L); foot of Murut Mt., Dec, buds, Maulfm US (K); E. Kalimantan,
W. Kutei, Mt. Kemul, alt. 1B00 m, tree 5 m, Oct., fr. lightgreen, tinged with red,
Endert U57 (BO, K, L).
LlNDERA OBTUSILOJJA Bl.
Cf. Kostormans, Bibl. Laur, 762. 1964. - Typus: sine local., Kcishe s.n.,
ster. (10.
1974] KOSTERMANS: Re of Lauraceae IV
Forma villosa (Bl.) Kosterm., stat. nov. — Lindera obtiisiloba Bl. var.
villosa. BL, Mus. bot. Lugd. bat. 1(21}: 325. 1851 (basionym) ; cf. Koster-
mans, I.e. 763 - Typus: Siehold s.n. (L), "Sirumozi".
The type sheet at Leiden has 3 collections, all wrapped in Japanese
tissue paper. The label mentions K (Keishe) as collector and this label
mentions the three local names ("Wuhon bana, Dan kou bai and Kouzin
bana") cited by Blume. The sterile specimens belong all to the typical,
almost glabrous form.
The variety villosa of Blume is based on a Siebold apecimen at Leiden,
marked "Sirumozi". It is more a forma rather than a variety, young
plants are always densely sericeous.
Miquel mentioned in his Catalogue Mus. bot. Lugd. bat. I: 81.
1871 two more varieties: hirsuta and pubescens; these names were copied
from herbarium labels of var. pilosa El., they were published as nomina
nuda.
Lindera pilosa Kosterm., spec. nov.
Arbor ramulis apicem versus dense minutissime pilosis Mi is
alternantibus chartaceis elliptids acuminatis basi acutis supra glabris
nitidis nervus medianus impresses costis filiformis vix prominulis subtus
dense ferrugineo pilosis petiolis glabris supra anguste canaliculatis
umbellulis longe et gracile pedunculatis glabris.
TYPUS: Forbes 2iS (L).
Tree, 9 - 13 m, diam. 20 cm. Bark smooth, brown. Branchlets densely,
minutely dark rusty tomentellous. Leaves spirally arranged, chartaceous,
elliptic to subovate-elliptic, 5 x 10—7 x 15 cm, dinatinctly acuminate, base
acute; upper surface glabrous, glossy, midrib impressed (at least the
basal part), lateral nerves hardly prominulous or impressed, lower surface
densely rusty tomentellous, midrib prominent, the ca 6—7 pairs of lateral
nerves erect-patent, arcuate, prominent, slender, secondary nerves very
lax, subparallel. Petioles very slender, up to 2 cm long, narrowly channeled
above. Umbel bracts glabrous, peduncles slender, glabrous, 10—15 mm
long, glabrous, a few on very short branchlets in the leaf axils. Florets
densely, minutely pilose, tepals 1.5—2 mm long; pedicel slender, ca 3 mm
long.
The species is characterized by the dense rusty tomentum of the
lower leaf surface. In some specimens (For. Officer Sumatra's E. Coast
and Meyer 3300) it is much thinner. In the specimen Meyer 4048 the
umbels are minutely sericeous.
SUMATRA: Locality not indicated, fl., Forbes 2JfSl (L, 3 sheets); Mt. Sago near
Pajakumbuh, second, forest along Bukit Tjempago, alt. 900 -1000 m, tree 20 cm diam.,
bole 10 m, crown 5 m; outer bark smooth with saattered lenticels, 0.5 mm thick,
102 REINWARDTIA [VOL. 9
live b o r k r e d b r o w n , yellow w i t h r e d - b r o w n spots, common, Oct., fl., Meyer iOiS ( L ) ;
ibid., N. slope, a l t . 1000 m, s t e r . , t r e e d i a m . 20 cm, bole 4 c m , c r o w n 6 m, b a r k
b r o w n , smooth, Meyer SS09 ( L ) ; K a r a L a n d s , a l t . 1400 m, Oct., b u d s , f o r e s t Officer
Sumatra's E. Coast S ( B O , L ) ; S i b o l a n g i t , S.W. of B a n d a r b a r u , a l t . 9E0 m, Oct.,
b u d s , Lorzing 5S27 ( B O , L ) .
Lindera reticulosa Kosterm., spec. nov.
Arbor ramulis glabris gracilis foliis alternant!bus chartaceis glabris
lanceolatis distincte sensim attenuatis basi acutis utrinque prominule
reticulatia eostis supra obscuris subtus filiformis petiolis gracilis canali-
culatis inflorescentiis brevis pauci-umbellatis minutissime sericeia floribus
sericeis. . ,
TYPUS: F.R.I. 5016 (L).
Tree up to 27 m high, 30 cm diam.; bark smooth to cracked, lenti-
cellate; live bark light brown, cambium bright yellow, wood white.
Branches slender, glabrous. Leaves spirally arranged, chartaceous,
lanceolate to lanceolate-elliptic, 2 x 7—3 x 11 cm, long-acuminate with
sharp tip, base acute, both surfaces prominulously, rather obscurely
reticulate, upper one with filiform, prominulous midrib, lateral ribs
hardly visible, lower one paler, midrib prominent, the 7—9 pairs of
lateral nerves filiform, erect-patent, hardly prominulous. Petioles slender,
5—7 mm, channeled above. Inflorescence branches up to 8 mm long,
bearing a few umbels, minutely sericeous; umbel peduncles up to 1 cm
long, outer bud scales glabrous, inner ones sericeous. Florets densely
sericeous, tepals 1.75 ram, narrowly ovate, acute; stamens exerted.
Related to L. polyantka, but easily distinguished by the leaf reticu-
lation, which never occurs in L. polyantka. Also near L. dictyoneura,
which has stamens twice the length of the longer tepals.
MALAY PENINSULA, Pahang, Fraser's Hill, path to Pine Tree Hill, mossy ridge,
alt. 1300 m, tree 27 m, diam. 30 cm, bole fluted and knobly, bark smooth to cracked,
lenticellate, slash inner bark light brown, cambium bright yellow, wood white,
leaf glaucous beneath, March, male fls. pale yellow, F.R.I. 5016 (K, KEP, L | ;
trail to G. Berumbun, Cameron Highlands, Pahang, mountain Elope, tree 20 m,
short thick buttresses bark smooth slash inner bark brown granular outer layer
yellow towards cambium, wood pale yellow, Febr., female fls. pale yellow, F.R.I. HSU
(K, KEP, L), leaves broader, up to 4 X 9 cm; trail to G. Jasar, Cameron Highlands,
alt. 1400 m, tree 10 m, diam. 25 em, bark cracked, inner bark pale yellow, wood
white, March, buds green, fr. round, green with short, apical point (not seen),
leaves glaucous underneath, 2 X 6 - 3 X B em, F.R.I. 5060 (K, KEP, L).
LITSEA Lam.
Litsea alba Kosterm., spec. nov.
Arbor parva ramulis Elabris albo-brunneis sat crassis apicem versus
applanatis sulcatisque, foliis alternantibus chartaceis vel rigide chartaceis
1974] KOSTERMANS: Revision of Lauraceae IV 103
magnis glabris aubobovato-eliipticis breve obscure acuminatia basin versus
aensim attenuatis obtusis, supra subnitida laevia nervo mediano lato plana
costis obscuris, subtus pallidiora nervo mediano magno prominente
costis erecto-patentibus sat gracilis prominulis nerviis secundaris fili-
formis laxis, petiolis sat crassis albo-brunneis, umbellulis immaturis
axillaribua glomeratis minutissime adpresse piloais vel glabria breve
pedunculatis.
T Y P U S : BSIP 3851 ( L ) .
Small tree. Bark white, smooth, living bark soft, white; wood,
hard, white. Branchlets rather thick, towards their apices flattened and
sulcate, pale brownish white, glabrous, smooth. Leaves spirally arranged,
glabrous, chartaceous or rather stiffly chartaceous, subobovate-elliptic,
11 x 28—16 x 29 cm, shortly acuminate or acute, gradually tapered
towards the obtuse base; upper surface rather glossy, the broad midrib
flat, the lateral nerves obcure; lower surface paler, midrib strong,
prominulous, lateral nerves 12—14 pairs, erect-patent, rather straight,
curved near the margin, rather slender, prominulous, secondary nerves
filiform, prominulous, lax, somewhat parallel. Petiole pale brownish
white (as the branchlets), thick, 1—2 cm long. Immature umbels almost
sessile in the leaf axils, the (immature) peduncles short, minutely,
adpressed pilose, bud scales practically glabrous.
Characterized by the whitish bark of the branchlets and petioles.
The leaves dry a reddish black colour.
SOLOMON ISLANDS. Santa Ysabel, Allardyce Harbour, Sasakolo R., primary
forest on river bank in valley bottom, Jan., buds, B.S.I.P. $651 (K, L| ; S.E. Santa
Ysabel, Dadale Bay, ilat plain, primary forest, Aug., fr. light yellow (non vidi),
B.S.I.P. 7K61 (K, L).
LITSEA ALBAYANA Vidal
Cf. Kostermans, Bibl. Laur. 78G. 1364. — Typus: Vidal SCO <K), syntypes:
Vidal 1SS1, 7S17, 15148.
New synonym: Litsea, bulusaweiisiB Elmer, cf. KostcrmanE, I.e. 795. — Typus:
Elmer 1S792, Irosin, Mt. Buluaan, July, young fr. (CAL, L, US).
Other specimen: Elmer 15S75, same locality, April, fl. (L); ibid., alt. S00 m,
Jlay, f l , PNH S853S (L).
Litsea anamo Kosterm., spec. nov.
Arbor mediocris ramulis dense ferrugineo tomentellis foliis alternan-
tibus rigide chartaceis ellipticis obtusis vel obscure brevissime acuminatis
basi obtusis supra sat opaca sub lente minutissime dense reticulata nervo
Mediano impressa dense minute tomentella costis gracilis impress is
glabrescentis subtus dense minute rufo-tomentella nervo median a
prominente costis sat patentibus prominentibus venulis secundariis
REINWARDTIA
[VOL. fl
subparallelis connectis petiolis gracilis sat long is dense tomentellls supra
anguste canaliculatis inflorescentiis immaturis dense ferrugineo tomen-
tellis axillaris.
TYPUS: Hartley T.G.H. 115iS (L).
Tree 10—18 m tall, 10 and more cm diam.; bark grey brown to
brown, smooth or rather rough, under bark green, inner bark light brown,
or reddish brown, aromatic; wood straw. Branchlets densely rusty
tomentellous. Leaves spirally arranged, rather stiffly chartaceous, elliptic
(rarely subobovate-elliptic), 4.5 x 8—7 x 12 cm (2.5 x 6—3.5 x 10 cm in
the syntype), obscurely, very shortly apiculate or obtuse, base obtuse:
upper surface dull to slightly glossy, glabrous except on the nerves, under
the high power lens minutely reticulate, midrib impressed, tomentellous,
lateral nerves slender, impressed, glabrescent; lower surface densely
rusty tomentellous, midrib prominent, the 9—12 pairs of rather patent
lateral nerves prominent, connected by a rather lax reticulation. Petiole
rather slender, 10—15 mm long, densely tomentellous, narrowly sulcate
above. Umbels (immature) on very short, densely rusty-pilose axillary
branchlets, the buds densely tomentellous.
Belongs to the L. amicorum group.
TERBITOHY OF NEW GUINEA. E. Highl. Distr., Kainantu Subdistr., Kassam Pass,
alt. 1400 m, Jan., buds, local name Anamo (A, BRI, CANB, K, L ) ; Morofae Distr,
Kuali Greek, about 5 miles S. of Wail, April, buds, Hartley T.G.H. 11.H3 (K, L).
W. IRIAN. 15 km S.W. of Bernard Camp, Idcnburg R., alt. 1550 m, ster., Brass &
T/€?&tG@tih 11975 ( d i s t r i b u t e d as Elttsoc&Tnns't
LlTSEA AUREA K o s t e r m .
Cf. Kostcrmans ™ Reinwardtia H: 86, f. 40. 1ST0. — Typus: bb. SSSJ, (BO, L).
New records: JAVA. S. of Leuwiliang, alt. 400 m, Sept., buds. Backer 25244 (L);
BORNEO. Sarawak, Ulu Sg. Selada, Melinau, Kapit, 3d Div., tree 33 111, diam. 40 cm,
Iban name Anau, Aug., buds, S. 21,9,97 (BO, K, L, SING).
Litsea brookeana Kosterm., spec. nov.
Arbor ramulis dense minute ferrugineo tomentellis, foliis oppositis
rigide coriaceis ellipticis brevissime apiculatis basi obtusis, supra nitida
laevia neryo mediano impresso costia filiformis subimpressis, subtus
minute adpresse pilosis glabrescentibus nerviis exceptis nervo mediano
prominentibus costis sat patentibus gracilis marginem versus arcuatis
evanescent!bus, petiolis craesis brevis dense pilosis.
TYPUS: Miss W.M.A. Brooke S571 (L).
Tree; branches grey, smooth, flossy, thickened at the nodes;
branchlets stiff, densely, minutely rusty tomentellous. Leaves opposite,
1974] of Laurace
rigidly coriaceous, elliptic, 3 X 4.5—3 x 7—4 X 8 cm, shortly apiculate,
base obtuse; upper surface glossy, smooth, glabrous, midrib impressed
(pilose at the base), lateral nerves very thin, slightly impressed; lower
surface minutely adpressed, subsilky pilose, glabrescent, except on the
main nerves, midrib prominent, the 7—10 pairs of slender rather patent,
prominent lateral nerves arcuate near the margins. Petioles rather thick
5—7 mm long, densely pilose.
Although the specimen is sterile, it is easily recognizable by its thick
small opposite leaves, which place it in the L. ferruginea group.
BORNEO. Sarawak: Mt.
WM.A. Erooke 8571 (L).
lit, alt. 1700 m, 11 May 1954, Mi:
Cf. KoBtcrmi
jrbach 13S6.
LITSEA CAI.OPHYLLANTHA K. Schum.
8, Bibl. Laur. 706. 1964; E e rdtia 8: 1070. — Typus:
New records: NEW BRITAIN. Galilo village near Cape Hoskins, W. Nakanai,
tree 7 m, diam. 25 cm, Aug., young fr., NGF C501 (K, L, LAE). NEW
GUINEA. Morobe Distr., Bulolo, alt. 1400 m, lower montane forest with Caeianopsig,
Engelkardia, Gordonia, tree 30 m, bole 17 m, diam. 50 cm, June, fr., NGF 1YU1
(A, BISH, BO, BRI, CANB, K, L, LAE, PNH, SING, SYD, US), fr. slender,
subovste-cllipsoid, 8 X 12 mm, cup thin, 7 — 8 mm diam. merging into the swollen,
fleshy, obconical, 5 - 6 mm long pedicel; Bupu villaee, Warapit, alt. 1500 m,
tree 12 m, bole 3 jn, diam. 25 cm, stems hollow and inhabited by black ants,
veins yellow, July, fl., NGF 22982 (A, BO, BRI, CANB, K, L, SING, SYO);
above Bahaia, 15 m. SE of Garaina, alt. 1000 m, tree 17 m, diom. 30 cm, old second,
forest, bark light grey with warty lenticels, inner bark light straw, wood straw,
leaves pale glaucous underneath, Jan., after antheais. Hartley T.G.H. 13677 <L>;
E. Highlands Distr. near Sassaura, alt. JSOO m, tree 7 m, bark grey-brown, smooth,
inner bark straw, twigs hollow with ants, July, fr., Hartley T.G.H. 119HG (L);
S. Highlands, sawmill at lalibu, alt. 2200 m, tree 20 in, Sept., fr., NGF H70SI, (L),
fr. 8 x 14 mm, cup 3 - 5 mm, hardly different from fleshy pedicel. Irian Barat,
Japen Isl., Asau, Sebosiari, Serui Distr., alt. 210 m, tree 22 m, bole 15 m, diam.
40 cm, bark smooth, white, slash light brown, sapwood 8 cm, yellow, heart wood
brown, Sept., buds, BW 92i3 (L); steep coast near Djajapura (Hollandia), low alt.,
tree 7 m, diam. 10 cm, bark smooth, Aug., fr. black, Kostermans & Soegeng 295
(A, BO, G, K, L, P ) ; slope of Cyclope Mts near Djajapura, alt. 300 m, tree 6 m,
diam. 10 cm, bark thin, smooth, grey, wood yellow with cigarbox wood smell,
branchlets white, ster., Kostermjins & Soegeng 188 (BO, L).
LITSEA CAERII Kosterm.
Cf. Kostermans in Reinwardtia 8: 88, f. 41. 1970.
New records: TERRITORY OF NEW GUINEA, forests of the Somadjidji, alt. 450 m,
ster,, SchleshtBT 19S77 (L), leaves much larger than usual; Mt. Ganeae, alt. 1000 m,
ster.. Can- 15290 (L); Papua, Centr. Distr., track from Mt. Krone, E. of Woitape,
REINWARDTIA [VOL. 0
nit. 2300 m, in undergrowth of Castcmop*
2.6 cm, Jan., fr., NGF £1)352 (L).
forest, treekt bole 3 in, di;
LlTSEA CAULIFLORA S t a p f
Cf. Kostcrmans, Bibl. Lour. 797. 1964. - Typus: Haviland 1221.
Litsea cauliflora (Thw.) Trimen was published one year later,
as far as could be ascertained.
The fruit cups, contrary to those of L. cauiocarpa, which are also
eauliflorous, are small and merge into the obconical short pedicel.
New records: BORNEO. Sabah, Mt. Kinabalu area, fr., R.S.N.B. 6017 (K, L I ;
Tenompok, alt. 1700 m, Febr., fr., J. & M.S. Clemens 28507 (K, K ) ; Mesilau R.,
alt. 1700 m, tree 13 m, diam. 20 cm, April, fl., R.S.N.B. 4908 (K, L ] ; ibid., tree 7 m,
diam. IB cm, Jan., fl., R.S.N.B. 4057 (K, L); Ulu Langanani, alt. 1300 m, riverside
and hillside, common, Aug., fr., B.S.N.B. 1728 (K, I,).
LITSEA CAULOCARPA Merr.
Cf. Kostermans, Bibl. Laur. 797 1964. - Typus: Ramos 1894 (L, US), syntypes:
Ramoi 1591 (US); Villamil 309 (US).
The cups are large and thick, obseurely thickly ribbed, the fruit
diameter is less than the cup diameter.
New records: SUMATRA. W. Indragiri, Taluk region, Hutan Pulau Lawaa near
Taratik air Hitam, lowland Dipterocarp forest, sandy aoii, Jan., fr., Amiruddi-n 65 (L),
tree 5 m, diam. 12 cm, crown 3 m. BORNEO. Sabah, Sandakan, Kabili-Sepilok For.
Rea., low alt., tree 7 m, diom. 10 cm, Nov., fr. grey, BNBFD 1015), <L] ; Bagat Camp,
alt. 40 m, black soil, tree 15 m, bole 7 m, diam. 10 cm, April, fr. yellow, San S5/.B6
<L); Lungmanis, mile 9, alt. 50 m, tree 12 m, bole 10 m, diam. 13 cm, inner bark
white, aoft, Jan., fr. whitish green, Sera 27123 (L); mile 45, Labuk Rd., tree 10 m
near stream, inner bark yellowish, sapwood yellow, Jan., fr.. Sore iSU2 (L);
Keraton Camp, alt. 40 m, black soil near river bank, tree 17 m, bole 7 m, diam.
20 cm, bark brown, outer bark grey, inner brown, cambium yellow, Jan., young fr.,
San SSSS7 (L); mile 7.5 Brit. Borneo Timb. Co. Concession, Lungmanis, 25 miles
S.W. of Sandakan, alt. 20 ro, tree 15 m, March, fr., Sera A 29S1 (A, BO, K, KEP,
L, SING); Tawau, Balong For. Res., tree 13 m, bark grey brown, scaly. May,
buds, San S01S2 <L); ibid., tree 10 m, diam. 30 cm, bark scaly, alaah outer bark
thin, inner 13 mm, yellowish, May, buds, San SO1SS (L); Ulu SR. Serudong, tree
10 m, diam. 20 cm, Aug., young fr., Son 31190 (L); Quoin Hill, Cocoa Estate,
alt. SO m, tree 7 m, diam. 10 cm, outer bark thin, inner bark brown, sapwood white,
glossy, March, fr. white with peculiar smell, San 21525 (L); Ulu Sg. Kalumpang,
tree 7 m, diam. 30 cm, June, buda. Son S0A77 (L); Sarawak: S. Tubau above
Tubau village, steep spur, alt. 100 m, tree 13 m, diam. 20 cm, bark smooth,
grey-brown, mottled, April, fr. grey-green, S. 18S72 <L)i Kuching, Semengoh Arbor.,
leached yellow sandy clay, alt. 15 m, tree 13 m, diam. 15" cm, bark smooth, whitish,
May, fla. pale yellow, S. 1S959 (L); Sampadi For. Res., 25th mile Bau-Lundu Ed.,
1974] KOSTERMANS: Ravi: of Law IV 107
1st Div., alt. 250 m, k
buds greyish with gre
Central-East Kalimanta
cup light brownish, End
rangaa forest, yellow clay, tree 7 m, diam. 13 cm, June,
ish stalks, S. 26271 (BO, E, K, L, SING); Indonesian
, W. Kutei, alt. 150 m, tree 6 cm diam., Nov., fr. pink,
rt 475S (BO, L); ibid., tree 10 m, diam. 8 cm, Aug., fr. red,
cupule dirty brown, when young white and cup light green, Endert 2581 (BO, L).
LITSEA CHEWII Kosterm.
Cf. Kostermans, Bibl. Laur. 799. 1864; in K
Chew, Corner & Staintou 160 (BO, K).
Litaea. kelferi var. ovata Gamble, ef. Koster
-dtia 8: 81. 1970. - Typus:
8S0. - Typus: Wray 1600.
New records: BORNEO. Sabah, Mt. Kinabalu, Ranau Distr., E. Pinosok plateau,
alt. 1500 m, near stream, tree 7 m, bole 5 m, diam. 20 cm, inner bark brown,
sapwood yellow, July, buds, Sun SS07i (L); Meailau R., alt. 1700 m, tree 5 m, diam.
8 cm, lower leafsurface light glaucous, Febr., buds, RSNB 4203 (K, L); ibid.,
tree 5 m, Jan., buds yellowish green, RSNB 4105 (K, L ) ; ibid., tree 10 m, April,
fr.orange on one side, cup flat small, thick, ESA'B 7006 (K, L).
Litsea ehrysoneura Kosterm., spec. nov.
Arbor parva ramulis dense minute sublanuginoso-tomentellis, Joliis
alternantibus rigide chartaceis ellipticis acuminatis basi acutis supra
subnitida glabrescentia dense minute reticulata nervo mediano impressa
coatis filiformis subim press is, subtus sat laxe minute sublanuginoso
tomentella nervo mediano prominentibus costis erecto-patentibus promi-
nulis nerviis secundariia subparallelis conjunctus, petiolis sat longia
canaliculatis, infloreacentiia immaturis axillaris subpilosis.
TYPUS: BSIP 2Uk (L).
Small tree. Branchlets densely, minutely rusty sublanuginose-
tomentellous. Leaves spirally arranged, stiffly chartaceous, elliptic,
4 x 11—6 x 14.5 cm, inconspicuously acuminate, base acute; upper surface
rather glossy, densely sublanuginose-tornentellous, glabrescent (tardily on
the nerves), densely, minutely reticulate, midrib impressed, lateral nerves
filiform, slightly impressed, lower surface rather densely, minutely,
ferrugineoualy sublanuginose-tomentellous, minutely reticulate, midrib
prominent, densely pilose, lateral nerves 7—10 pairs, erect-patent, towards
the margin arcuate, prominulous, connected by subparallel slender
secondary veins. Petioles slender, 1.5— 2̂.5 cm long, channeled above,
densely pilose. Umbels (immature) on very short branchlets in the leaf
axils, the buds almost glabrous.
Solomon 1st. Santa Ysabel, near Maringe Lagoon, S. side of west ridge of
Mt. Sasari, forest on very steep hillaide, alt. 700 m, leaves grey below with golden
iins, petioles and twigs golden tomentose, Oct., buds, BSIP 2Ui (K, L).
REINWARDTIA [VOL. 9
LITSEA COREANA Lev.
Leveille in Fedde, Kepert. 10: 370. 1912; Allen apud Kehder in J. Arnold Arb.
17: 327. 1936; in Ann. Missouri bot. Gard. 25: 399. 1938; Kakai, Fl. sylv. Koreana
22: 38 and 55. 1839; Kostermans, Bibl. Laur. 807. 1964. - Typus: Taquet 13S5 ( E | ;
isotype: Taquet S171, iiOl (BO, E, K, LE).
Litsea zwccariim Koatermans in Eeinwardtia 8: i n . 1970.
Dapknidium lancifolmm Siebold & Zuccarini (non Thwaites), Flora Japon.
Fam. nat. 2: 83 (1846). 1S47; Kostermans, I.e. 461; in Reinwardtia, I.e.; lozoste
laneifolia Blume; cf. Kostermans, I.e. 461 (no. 12a) ; Actinodaphn* loncifolia IS. & Z.)
Meissner; cf. Kostermans, I.e. 857 (no. 226b).
I had the opportunity to study the original material of Leveille in
Edinburgh and discovered, that Leveille's description was based on
specimens, belonging to different genera: a real Litsea of which the
flowers were described and Persea thunbergii (specimen Taquet 1356)
of which the fruit were described.
Allen apparently saw only the latter specimen, and referred L. coreana
to Persea {Machilus) thunbergii. As, however, the main part of the
description covers Litsea coreana, this name should be used.
Litsea zuccarinii from Japan is identical with it, as was discovered
by Kakai.
Litsea krukovii Kosterm., nomen nov.
Actinodapkite pedicdlata Hayata (base) in Matsumlira & Hayata in Journ.
Coll. Sci. Tokyo 22: 351. 1906; Icon. 5: 172, fig. 60-C. 1915; ei. Kostermans, Bibl.
Laur. 35. 1964; Hui-Lin-Li, Woody Fl. Taiwan 198. 1963. - Fiwa pedicellata
(Hayata) Nakai in Journ. Japan. Bot. 14: 193. 1838; Kostermana, I.e. 521. — Litsea
pedicellata (Matsum. & Hay.) Hatusima in Journ. Geobot. 19 ( 1 - 2 ) : 25. 1971.
_ Typus: Myahe anno 1S99, Owatari anM 18SS, Makino 18S8, Hirar s.d.
Hatusima rightly transferred this species from Actinodaphne to
Litsea. However, Litsea pedicellata (M. & H.) Hatusima is antedated
by Litsea pedicellata Bartlett (1909), an american species.
It is a great pleasure to name this species after Dr. B.A. Krukoff,
the well known American botanist, who is supporting my work.
Litsea neohebridensis Kosterm., spec. nov.
Arbor mediocris ramulis glabris sat gracilis foliis alternantibus
lartaceis glabris ovatis vel late ellipticis brevissime late obscure
-juminatis vel ohtusis basi cuneatis, utrinque minutissime dense
reticulatis, supra sub-nitidis nervo mediano impressis costis filiformia
prominulis, subtus pallidiori^us nervo mediano prominentibus sat gracilia
ch;
3974] KOSTERMANS: Revision of Lav
costis erecto-patentibus utrinque 4—6 gracilis, petiolis gracilis perlongis
supra canaliculatis, infloreseentiis axillaris brevissirais sitbglabris
umbellulis paucis longe pedunculatis, fructus anguste ellpsoideis, cupulis
sessilis cupuliformis, basi obouicis margine integris.
TYPUS: A. Morrison s.n. fP).
Tree, 7 m high. Branchlets glabrous, rather slender. End bud very
minutely sericeous. Leaves spirally arranged, chartaceous, glabrous,
ovate to broadly elliptic, 3.5 x 6—6 x 11—6.5 x 10 cm, shortly, broadly,
rather obscurely acuminate or obtuse, base cuneate, both surfaces very
minutely, densely reticulate (more distinct underneath); upper surface
rather glossy, midrib slender, reddish, impressed, laterals filiform,
prominulous, lower surface paler, midrib rather slender, prominent,
reddish, laterals erect-patent, 4—6 pairs, prominulous, curved near the
margin, sometimes forked, secondary nerves obscure, very lax. Petioles
slender, 1.5—3 cm long, channeled above. Inflorescences axillary, ca 1—2
mm long, thickish (terminal bud minutely sericeous), slightly pilose,
bearing a few, peduncled umbels. Peduncles slender, up to 5 mm long
with a few sub-appressed hairs, umbel buds accutish with a few sub-
appressed hairs. Fruit narrowly ellipsoid, up to 7 x 15 mm. Cupula
cup-shaped, rather thin, sessile, 3—4 mm deep and 5 m diam. with
entire rim, merging gradually into an obconical, 2 mm long basal part.
Differs from L. aneityensis by its glabrous, thinner leaves with
longer petioles and the sessile cup.
NEW HEBRIDES. Tanna Issl., Lcnakcl, alt. 100
March, buds, Kajewski HO (P); Undine Bay, Efate,
Aug., fr., -4. Morrison s.n. (P).
rainforest, not common,
nit of Peak, 900 m alt.,
LITSEA VERBTEEGHII Allen
Cf. Kostermans, Bibl, Laur. 892. 1964; in Reinwardtia 7: 499. 1969; 8: 85. 1970.
- Typus: Brass & Versteegh 11162 (A, BO, K, L).
I formerly included this in L. alveolata Allen. Since more material
has become available, I must confess, that Allen was right in separating
the two. Superficially they are much alike, but the umbel stalks of
L. alveolata are much shorter, in L. versteeghii the umbels are smaller
on long peduncles; the fruits of L. alveolata have a cup continuous with
the obconic short pedicel, in L. versteeghii the cups are almost flat,
much smaller and abruptly narrowed into slender, longer pedicels.
Specimens examined: W. IRIAN. Hills N. of Api E., Kebar Valley, alt. 800 m,
clay soil, tree 26 m, bole 20 m, diam, 40 cm, buttresses up to 60 cm high, out 60 em,
and 4 cm thick, outer bark greenish brown with many shallow fissures, inner bark
yellow within, slash oehrous, rapidly turning brown, sap wood light yellow, no
iiO REINWARDTIA [VOL. 9 1974] K O S T E M M A N S : Re of Lau 111
heartwood, Oct., buds, BW $3$3 (L); Bale R., 18 km NE. of Lake Habbema,
alt. 2450 m, tree 5 — 6 m, Nov., fls. brown, Brass line (L). TERBITORY OF NEW
GUINEA. Sepik Distr., Hindenburg Range S. of Nerenavip village, alt. 2900 m,
secondary forest, tree 20 m, diam. 26 cm, wood yellow, Sept. ft., NGF 28517
(A, BRI, CANB, K, L); W. Highlands, Alipe, Kepaka, Upper Kangel Valley,
Subdistr. Hagen, alt. 2800 m, tree 17 m, diam. 20 cm, June, fr., Bowers T8i (L).
MACHILUS Nees (= PERSEA)
MACHILUS SERICEA (Nees) BI.
Blume, MUB. Bot. Lugd. Bat. 1 (21): 330. 1851; rf. Kostermans, Bibl. Laur. 926.
1061; Ocotea sericea Nees (non H.B.K., nee Goudot ex Meisn.) i« Wallich, PL Asiat.
rar. 2: 71. 1831, p.p. - Typua: Wallich Catal. 26OG -B, p.p., n. (L).
According to Hooker f. (Fl. Brit. India 5: 139. 1889), Laurus sericea
A, B, C might or might not belong to one species; it was described by
Nees as a Phoebe, but Blume separated later a specimen as Machilus
sericea, "observing that it differs from the Phoebe in the absence
(sphalm.! = presence) of rings, caused by the fall of budseales at the
base of the branches".
Blume's specimen, a small branchlet with leaves and inflorescence
is conserved in Leiden; it has a glaucous, sparsely sericeous lower leaf
surface, sericeous flowers and glabrous filaments. In my opinion it is
conspecific with Persea bombyeina (King ex Hooker f.) Kosterm.
The specific epithet "sericea" cannot be transferred to Persea
because of the American Persea sericea H.B.K. and consequently the
name Persea bombyeina {King ex Hooker f.) Kosterm. stands.
PERSEA Miller
Persea blumei Kosterm., nom. nov. - .
Bot. Lugd. Bat 1(21); 330. 1861;
iker f., FL Brit. India 5: 862. 1890;
!erb. Heland, e Ncpalia, Ocotea no. SO,
Machilv.i pubescent Blume (base), Mus
Meisner in DC, Prodi-, 15 <1): 40. 1864; Ho
Kostermans, Bibl. Laur. 924. 1064. - Typus: 1
fl. (L); isotypus (K).
The species was based oil a specimen sent from De Candolle to
Blume as Ocotea no. 20 (belonging to the set, sent by Wallich to
De Candolle in Geneve, Hornemann in Kjobnhavn and to D. Don of
Nepalese plants). The holotype in Leiden consists of a flowering
branchlet with two leaves, more complete material is in the De Candolle
herbarium and I found also a good specimen in Hooker's Herbarium
(Kew), overlooked by Hooker, who in his Flora of British India disposed
of it as a species being- unrecognizable from the description. It differs
from its nearest relative P. villosa by the leaves having a villose lower
surface without visible secondary nerves and the larger number of
laterals.
Persea declinata (Bl.) Kosterm., comb. nov.
Lzurua declinatQ. Blume (basionym), Catal. Gewasser. 'sLands Tlantent.
Buiteniorg 66. 1823; ef. Kostermans, Bibl. Laur. 611. 1964; Phoebe declinata (Bl.)
Nees, Syst. Laur. 114. 1838; cf. Kostermans, I.e. 1275; m Eeinwardtia 4: 203. 1957;
7: 353. 1968; Ocotea declinata Bl. (non Mez), Bjidr. FL Nederl. Indie l i e Stuk 573.
1S2S; cf. Kostermans, I.e. 1083.
I have hesitated to refer this to Persea, because of the non-reflexed,
slightly hardened tepals under the fruit. However, the leaf reticulation
is that of Persea and so are the globular fruit and since its closest
relation seems to be with P. excelsa, I believe that this species belongs
to Persea. In American species of Persea semi-erect persistent tepals
occur more often:
For synonymy see Kostermans in Reinwardtia 7: 353. 1968.
... Peraea endothrix Kosterm., spec. nov.
Arbor ramulis glabris foliis coriaceis glabris ellipticis utrinque
dense minute areolato-reticulatis breve acutis vel obtusis basi acutis,
supra sublaevia nervo mediano robusta prominentis costis gracilia,
petiolis bene euolutis, paniculis foliis subbrevioribus glabris paucifloris
breve et pauce ramosis, pedicellis longis tepalibus elongatis extus glabris
intus dense lanuginosis.
TYPUS; San 56521 (L).
Tree. Branchlets rather slender, glabrous, end bud small, glabrous.
Collar of prophyl scars lacking. Leaves spirally arranged, sub-aggregate,
coriaceous, glabrous, elliptic to subobovate-elliptic, 2.5 x 7 (2.5 x 9)—
C x 16 cm, shortly acuminate or obtuse, base acute, upper surface glossy,
smooth or obscurely pitted, midrib slender, strongly impressed; lower
surface densely areolate-reticulate, midrib strongly prominent, laterals
10—18 pairs, slender, erect-patent, arcuate near the margin, prominulous.
Petioles slender, 1—2 cm long. Panicles axillary, glabrous, 1.5—12 cm
long, few and shortly-branched near the apex, few-flowered. Pedicels
filiform, up to 7 mm. Tepals elongate, narrow, up to 4 mm long, glabrous
outside, densely woolly inside, tube infundibuliform, 1—1.5 cm; anthers
1—1.5 mm long, slender, pilose, anthers 0.5 mm, narrow, outer ones
with large lower cells and very small upper ones; inner with larger upper
REINWARDTIA
[VOL. 9 1374] KOSTERMANS: Revision of Lauraceae IV
cells, the lower more lateral; basal glands long-stalked; staminodes long-
stalked, ovate, acutish with thickened rims; ovary ellipsoid; style as
long as the filaments, stigma truncate, small.
No collars of narrow prophyl scars could be discovered on the
branches, which places this species near P. exceisa and P. declinata.
Characteristic is the woolly inside of the tepals (the inner ones are
leiss woolly).
BORNEO. Sabah, Eanau, alt. 1300 m, Tcnompok proposed toe
March, fl., San. SS5« (L).
eh Kinabalu,
Alseodapkne omeiensis Gamble in Sargent, PI. Wilson. 2: 70, 24 March 1914, cf.
Kostermans, I.e. 69; Notkapkoebe omeiemis (Gamble) Chun «i Contr. Biol. Lab. Sei.
Soc. China 1(5): 33. 1925; Kostcrmans, I.e. 1058.
Mackilns dunnianus and M. mairei have been already reduced to
synonyms of Alseodaphne omeiensis Gamble by Render and Liou Ho.
However, the former have been published 3 weeks earlier than A Iseodaphne
omeiensis.
Available material shows moreover the erect, indurate perianth
of the fruit, which places this in Phoebe.
Persea peduncula.ta (Bl.) Kosterm., comb. nov.
lozoste •pedu.nculata Blume (basionym), Mus. Bot. Lugd. Bat. 1 (23) : 364. JB51;
Kosternmns, Bibl. Lsur. 550. 1954. Actinodapkne pedunculate. (Bl.) Meisn. in DC,
Prodr. 15(1): 211. 1864; cf. Kostermans, Bibl. Laur. 3G. - Typus: Reeves s.n.
(e Herb. Lindley) (L).
Modulus velittina Champion ex Benth. in Hooker's Kew Journal & Gard. Miscell.
S: 198. 1853; Kostermans, Bibl. Laur. 930 (extensive references); in Reinwardtia
A- 1 £14 19fi2" a - 120 1970 f exclus f Tosea Liou Hot • PSTBSO vsivtiita I ChamD rat
Benth.) Kostermans in Eeinwardtia B: 194. 1962; Bibl., b.c. 1262. — Typus: Champion
181 (K).
Mackilus grij&ii Han*e in Ann. Sci. Nat., Sir. 4, 18: 226 (1862) 18G3; Koster-
mans, Bibl. I.e. 908; Persea grgsii (Hance) Kostermans in Eeinwardtia 6: 186.
1962; 6: 118. 1970. — Typus; Grqi iji Herb. Hance S7S8 (E, K).
Actinodaphne wagniflora Allen in Ann. Missouri Bot. Gard. 25: 403. 1938;
Kostermans, Bibl., l.c- 31; in Bull. Bot. Survey India 10: 287. 1968; in Eeinwardtia
8: 120. 1970.
This very common species, which flowers already when it is one
meter high, is based on a specimen, collected by Reeves in S. China and
in the position of Lindley, who sent a specimen to Blume.
Allen, who re-described it as an Actinodapkne, was apparently misled
by the aggregate leaves, although inflorescence and flower structure
excludes it definitely from Actinodapkne.
PHOEBE Nees
Phoebe dunniana (Levcille) Kosterm., comb. nov.
Mackilus dunnianus Ltveille in Fedde, Repert. 13: 174, 28 Febr. 1914; cf.
Kostermans, Bibl. Laur. 905. 1964. Machilus mmniana Yang in Journ. W. China
Border Res. Soe. 15, B: 75. 1945; Kostermans, I.e. 905 (sphalm. = dunniana).
— Typus: Maire s.n., coUines boisees a Long Ky, alt. 700 m, fleurs, Junt 1912 (E).
Machilue mairci Leveilte in Fedde, Repert. 13: 174. 1BI4, cf. Kostermans, I.e. 917.
- Typua: Maire s.n., fl., Lons Ky, May <E>.
Phoebe kjellhergii Kosterm., spec. nov.
Arbor ramulis apicem versus dense minutissime ferrugineo villosulis
foliia subaggregatis rigide chartaceis vel subcoriaceis obovato-ellipticis
vel suboblanceolatis breve vel brevissime acuminatis basin versus sensim
flttenuatis in petioliim sat crassum contractis vel acutis supra nitida
glabra nervo mediano impressa costis filiformis subimpressis subtus
opaca minutissime adpresse piloaa nervo mediano prominentis costis
erecto-patentis gracilis prominentis nerviis secundariis subparallelis
filiformis conjunctis infra ctescenti is foliia aequilongis vel longioribus vix
et breve ramosis glabrescentibus fructua ovoideus perianthus persistens
par vis minutisaime adpresse pilosis.
TYPUS: Kjellberg 2973 (BO).
Tree. Branches smooth, pinkish grey, the youngest ones towards
their tips densely, minutely, rusty villous. Leaves spirally arranged,
subaggregate (monopodial branching), rigidly chartaceous to subcoria-
ceous, obovate-elliptic to suboblanceolate, 4 x 11—6 x 17 cm, apex shortly
acuminate to apiculate, base gradually tapered, acutish or occasionally
somewhat rounded; upper surface smooth, glabrous, the midrib longitudi-
nally narrowly channeled, lateral nerves filiform, slightly impressed,
lower surface dull, finely, densely, very minutely appressed pilose, midrib
and the slender, erect-patent, ca 10 pairs of slightly arcuate lateral nerves
prominent, secondary nerves subparallel, filiform, prominulous, reticu-
lation fine, rather dense. Petioles rather stout, 10 mm long, puberulous,
Infruetescences axillary, subapical, up to 15 cm long with a long
peduncle and few, up to 2 cm long, subapical branches, glabrescent;
fruit ovoid, 11 x 13 mm, blue-black; persistent perianth rather flat
up to 6 mm diam., the tepals unequal, up to 3 mm long1, minutely, densely
appressed pubescent.
Characterized by the small persistent perianth, the obovate-elliptic,
thickish leaves with a very short, appressed pubescence.
The Kjellberg specimen was from a branch, picked up from the
ground.
114 REINWARDTIA
SULAWESI (Celebes). Diatr. Palopo, village Batang,
(BO); Todjambu, alt. TOO m, Dec, fr. black, peduncles
[VOL. 9
lit. 250 m, ster., 66. 20898
red, Kjellberg 2STS (BO).
Phoebe tenuifolia Kostcrm., spec. nov.
Phoebe ctercutwides (Elmer) Merrill, cf. Kostermans, Bibl. Laur. 1301. 1064, p.p.
(quoad Merritt 682i).
Lauraoeae, Spec. X, Koorders in Medsd. Plantent. Buitenzorg 19: 575. 1898.
Arbor ramulia griseis lenticellatia glabris apiee minutissime pilosis
foliis aggregates chartaceis diu glabreseentibus obovato-ellipticis brevis-
sime late acuminatis basin versus attenuatis acutis utrinque minutissime
dense reticulatis supra nitida nervo mediano impreasa costia obscuria
filiforrnis aubimpressis subtus pallid; ora subopaca nervo mediano
prominent! bus costis erecto-patentibua gracilis prominentibua nerviia
secundariis parallelis prominulis connectis petiolig brevis infructescentiis
foliis subbrevioribus apicem versus (sub lente) mmutissime dense pilosis
vix ramosis pedunculis perlongis robustis fruetus par vis ellipsoideo-
ovatis perianthus induratus minutiSRime pilosus amplectens.
TYPUS: de Boer 2s = bb. 2249 (BO).
Tree up to 20 m high and GO cm diam. Branchlets greyish white,
lenticellate, glabrous with very large, protruding- oval leaf scars, near
their apicea very minutely, densely pubescent. Leaves apirally arranged,
aggregate, ehartaceous, obovate-elliptic, 7 x 19—10 x 26 cm, obtuse with
a very short, broad acumen, base gradually tapered, acute, both surfaces
very minutely, densely reticulate, upper surface glabrous, glossy, midrib
impressed, lateral nerves obscure, filiform, slightly impressed, lower
surface more dull, in younger leaves minutely, rather densely appressed
pilose, soon glabrous, midrib prominent, lateral nerves ca 10—12 pairs,
erect-patent, slightly arcuate, prominent, connected by parallel, promrau-
lous, filiform secondary veina. Petiole 1—3 cm long. Fruiting panicles
up to 19 cm long with rather stout, long peduncle and a few up to 2 cm
long apical, widely spaced branches, towards their apices very minutely
appressed pubescent. Fruit ellipsoid ovoid, slender, yellowish-brown (in
aJcco), up to 6 x 10 mm, persistent perianth minutely, densely appressed
pilose, ca 5 aim long, tepals subequal.
A widespread species, characterized by its thin leaves with a very
fine pubescence on the lower leaf surface; the small persistent perianth
and the small fruit which dry yellowish brown.
It has a fairly constant vernacular name in Celebes: "Areepoongoo".
The specimen Merritt SSSi from Mindoro has somewhat broader
leaves with larger flowers, but the latter is due to the stage of development,
some flowers are diseased and have become gigantic.
1974] KOSTERMANS: Revision, of Lauraceae IV
PHILIPPINES. Mindoro, April, fl., Merritt SS2i (BO); Mindanao, Todaya
(Mt. Apo), Diatr. of Davao, May, fl., Elmer 10685 (BO). SULAWESI (Celebes), Minahasss,
Tondano, ster., Xoorders 17429 (BO, L ) ; ibid., near Eatatotok, ster., Koorders 17W1
(BO, L ) ; Eatahan, alt. 250-300 m, June, fr., Koorders 2J,O7O (BO, L) and 17/.SS
(BOli Kajuwatu, alt. 50 m, ster., Hoarders 17501, (BO, L| and 1751S (BO);
Lulumbulan near Paku-ure, April, f]., Hoarders 1T',S6 (BO, L ) ; near Modoinding
alt. 1060 m, Sept., buds, 66. 17120 (BO|; Palopo Distr., village Morante, alt. 200 m,
tree IS m, diam. 33 cm, ster., 66. gO8U (A, BO, L ) ; village Batang, alt. 250 m,
star., 66. 20898 (BO); Amurang, fr., sine call. (BO, K, L ) ; Is]. Buton, Distc.
Sampolawa, alt. 10 to, March, fr., de Boer 29 = bb. 2249 (BO, L ) ; Isl. Muna,
village Wapai, alt. 5 m, tree 20 m, diam. 60 cm, ster., bb. 21729 (A, BO, L).
i C O N T E N T S
P a g e
H A T T I N K , T. A. A revision of Malesian Caesalpinia, i n c l u d i n g
, Mezoneuroji ( L e g u m m o s a e - C a e s a l p i n i a c e a e ) 1
• J O N E S , H . G < Orchidaceae n a v a e vel m i n u s cogtlitae . . . . . . . 7 1
K E N G , H. Rediscovery of Cheilotheca malayana a n d t h e i d e n t i t y of .
Cheilotheca, Audresia and Mo.notropastmm (Ericaceae- .
M o n o t r o p o i d e a e ) 7 7
K O S T E R M A N S , A . J . G . H . A m o n o g r a p h o f t h e genus- N.eoanna-
•momum L i o u H o 8 5
M a t e r i a l s f o r a r e v i s i o n o f L a u r a c e a e I V . . . . . 9 7
r? A new Bornean species .of Mammea , . 117
Triadodapkne, a. new Jauraceous genua from Borneo . 119
— A monograph of Caryodaphnopsis A. Shaw . ., . . . 123
LARSEN, K. & LAKSEN, S. K. A new Amorphophallus from Thailand ' 139
NAYAK, M. P. A revision of Phtkiandra (Melastomataceae) . . 143
RAO, A. N. £ LEONG, F. L. Pollen morphology of certain tropical ,
• plants . . . . . . . - . • 153
SKVORTZOV, B. V. On some colourless flagellates from Java and Brasil 177
Distributor
BIBLIOTHECA BOGORIENSIS,
JALAN RAYA JUANDA 20,
EOGOK, INDONESIA
by ARCHIPEL
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