VOLUME 4 DECEMBER 1956 PART i

REINWARDTIA

BEING A CONTINUATION OF T H E

BULLETIN DU JARDIN BOTANIQUE DE BUITENZORG
(BULLETIN OF THE BOTANIC GARDENS, BUITENZORG)

EDITORS

A N W A R I DILMY

(Herbarium Bogoriense)

AND

C. G. G. J. VAN STEENIS
(Flora Malesiana)

Published by
HERBARIUM BOGORIENSE
KEBUN RAYA INDONESIA

Reinwardtia Vol.4, Part 1, pp. 1-118, BOGOR, December 1956



40 REINWARDTIA [Vol. 4

5. Dehaasia titanophylla (Airy Shaw) Kostermans, comb. nov.

Beilschmiedia titanophylla Airy Shaw (basonym) in Kew Bull. 1939: 534,
was described after the specimen: Native coll. 2438 from Borneo. A fragment of
this, conserved in the Singapore herbarium, could be examined. Although the
fruit are still unknown, the close relationship to D. longipedicellata (Ridley)
Kosterm. makes it advisable to relegate this species to Dehaasia.

R O S A C E A E

PARINARI Aubl.

Polyalthia? pulchrinervia Boerlage

This species was described by Boerlage (Catal. Hort. Bogor., fasc. 1: 20.
1899 and in Icones Bogor., fasc. 2: 106. 1899) after a sterile specimen, collected
from a tree, numbered IV G. 42 in the Bogor Botanical Gardens.

In the Bogor Herbarium there is one sheet labelled: Polyalthia ? pulchri-
nervia Boerlage (not in Boerlage's handwriting); no date of collecting is
indicated; the specimen was from the tree IV G. 42.

I consider this the type specimen, as it fits with Boerlage's description.
Duplicates of this have been sent to the Arnold Arboretum, the Kew, the Leiden
and the Singapore herbaria.

In 1915 additional flowering and fruiting material was collected from the
same numbered tree, which makes it clear that it represents Parinari corymbosa
Miq. Of the 1915 collections duplicates have been forwarded to the Arnold
Arboretum, the Kew and the Leiden herbaria.

S T E R C U L I A C E A E

STERCULIA

Sterculia perryae Kostermans, nom. nov.

Sterculia clemensiae Merrill & Perry in J. Arnold Arb. 30: 40. 1949 is,
because of its earlier homonym Sterculia clemensiae Ridley (in Kew Bull. 1933:
488), renamed here: Sterculia perryae Kostermans.

R E I N W A R D T I A

Published by Herbarium Bogoriense, Kebun Raya Indonesia
Volume 4, Part 1, pp. 41-68 (1956)

NOTES ON MALAYSIAN MALVACEAE—I *

J. VAN BORSSUM WAALKES **

SUMMARY

Twelve new species of Hibiscus sect. Azanza DC. from the Malaysian
region are described and illustrated. The genus Wilhelminia Hochr. is
reduced to a synonym of this section, which requires the new combination
Hisbiscus sciadiolepidns (Hochr.) Borss. for W. sciadiolepida Hochr., the
only species of the genus.

For some years past I have been engaged in studying the Malvaceae
of the Malaysian area. It is intended to publish the new facts and con-
clusions in a series of papers under the title indicated above, and eventual-
ly a concise treatment of the family will be included in the "Flora
Malesiana." The Malvaceae will be treated in a strict sense, i.e. the
Bombacaceae, which is considered a separate family, will be excluded.
The Bombacaceae are, it is true, very closely allied to the Malvaceae, but
nevertheless it had better be kept apart, both from a theoretical and from
a practical point of view. I hope to discuss the delimitation of the two
families in a later publication.

A critical revision of the family in the Malaysian area has not so far
been carried out, although numerous facts and conclusions have been re-
corded by various botanists, of whom the following may be mentioned here:
Masters, who gave a concise but accurate elaboration of the Indian species
in Hooker's "Flora of British India," Gagnepain on Malvaceae in French
Indo-China, Merrill especially on Philippine Malvaceae, F. von Miiller,
Schumann, Lauterbach, and Ulbrich on New Guinea species, E. G. Baker,
Garcke, Giirke, and last but not least Hochreutiner, who has an all-round
knowledge of the Malvaceae.

For the present paper I have studied specimens from Herbarium
Bogoriense (BO),1 and the Forestry Research Institute at Bogor (BZF),
as well as from the herbaria of Singapore (SING), Florence (FI), Lae
(LAE), Brisbane (BRI), Arnold Arboretum (A) and the Gray Herbarium

* This article was issued separately in June 1956.
** Botanist, Herbarium Bogoriense, Kebun Raya Indonesia.

1 The abbreviations are according to Lanjouw [in Regn. veg. 2 (Ed. 2) : 131-144.
1954]. . : :

— 41 —



42 R E I N W A R D T I A [VOL. 4

(GH). During European leave, collections in the Rijksherbarium at Leyden
(L), the Kew Herbarium (K), the British Museum (Natural History) (BM)
were consulted. I am very grateful to the Directors and Members of the
staff of these Institutes for the opportunity to study the collections
under their care.

FIG. 1. Map illustrating the localities of the species described; the numbers are those
given to the species in the text.

NEW SPECIES OF HIBISCUS SECTION AZANZA DC.

In the following pages I give a key of all Malaysian species of Hibiscus
section Azanza DC; the new species described are numbered. Hibiscus
papuamis Schum. & Lauterb., H. calodendron Ulbr. and H. sehlechteri
Lauterb. are omitted: these amount to nomina dubia since the type
specimens in Berlin were destroyed and apparently no duplicates exist in
other herbaria.

KEY TO THE MALAYSIAN SPECIES OF HIBISCUS SECT. AZANZA DC.

1. Ovary and capsule 10-celled as a result of 5 true and 5 false dissepiments.
2. Epicalyx segments 8—11, connate with each other, much shorter than the calyx,

forming a whorl of short, appressed to slightly patent dents. Pantropic species.
H. tiliaeeus L. emend. Hochr.

2. Epicalyx segments 4—9, mostly free, sometimes shortly connate with each other,
somewhat longer than the calyx or about as long as the calyx, rarely shorter
than the calyx, in the latter case 4—6 in number. Species from New Guinea and
surrounding islands.
3. Epicalyx segments 9 1. H. aruensis
3. Epicalyx segments 4—6.

4. Inside at base of corolla a cushion consisting- of long- woolly hairs; stigmas
capitellate.

1956] VAN BORSSUM WAALKES: Notes on Malaysian Malvaceae—/ 43

5. Indumentum consisting mainly of minute stellate hairs; flowers fairly
large; epicalyx deeply 6-partite, spreading more or less stellately.

2. H. pleijtei
5. Indumentum consisting mainly of scales; flowers small; epicalyx 5-fide,

cupulate to campanulate 3. H. pulvinulifer
4. Corolla without such a cushion. Stigmas as far as flowers present discoid.

6. Indumentum consisting mainly of stellate hairs.
7. Epicalyx segments widely ovate or orbicular, at base cordate.

8. Blade more or less orbicular, coarsely stellate hairy; epicalyx seg-
ments 4 4. H, leeuwenii

8. Blade ovate, with minute stellate hairs; epicalyx segments 5—6.
9. Green parts with scattered stellate hairs to glabrous; calyx

about as long as the epicalyx H. cfalbertisii F. Mull.
9. Green parts densely tomentose; calyx about twice as long as

the epicalyx 5. H. fluminis-idenburgii
7. Epicalyx segments lanceolate, at base narrowing gradually, not cord-

ate.
10. Blade with coarse stellate hairs, not tomentose; epicalyx segments 6.

6. H. ellipticifolius
10. Blade with minute stellate hairs, tomentose; epicalyx segments 5.

H. fluminis-aprilii Ulbr.
6. Indumentum consisting mainly of scales (fimbriate or not fimbriate).

11. Epicalyx more or less cupulate, 5-lobate to 5-fide.
12. Twigs and leaves with scales only; midrib of the leaves without

nectary H. sciadiolepidus (Hochr.) Borss.2

12. Twigs and leaves also with stellate hairs; midrib of the leaves
with an elliptical nectary 7. H. carrii

11. Epicalyx segments free or at base very shortly connate.
13. Blade ovate to elliptical; epicalyx segments 5, about 2 cm long.

8. H. archboldianus
13. Blade orbicular to very widely ovate; epicalyx segments 4 or 6.

" 14. Epicalyx segments 6, 1.5—2 cm long. 9. H. viomersleyanus
14. Epicalyx segments 4, 2—2.5 cm long. . . 10. H. lepidottts

1. Ovary and capsule 5-celled, without false dissepiments.
15. Calyx without scales.

16. Epicalyx segments 10—14; indumentum with very large stellate hairs.
H. macrophyllus Roxb. ex Hornem.

16. Epicalyx segments 7—9; indumentum with small stellate hairs.
17. Stipules more or less reniform; epicalyx deeply 7—9-partite, much shorter

than the calyx H. bomeensis Airy Shaw
17. Stipules ovate to lanceolate; epicalyx 7—fide, a little shorter than

the calyx 11. H. pseudotiliaceus
15. Calyx densely covered with scales.

18. Epicalyx segments 6—7, free, linear to lanceolate to ovate, almost as long as
the calyx. Seeds 2 per cell H. decaspermus Koord. & Valet.

2 Hibiscus sciadiolepidus (Hochreutiner) Borss., nov. comb.; basionym: Wilhelminia
seiadiolepida Hochreutiner in Nova Guinea 14: 162 t. 18. 1914. This species must be
classified under Hibiscus in my opinion. I hope to discuss this reduction of the genus
Wilhelminia in a later paper.



44 R E I N W A R D T I A [VOL. 4

18. Epicalyx consisting of a whorl of 7—8, at base connate, triangular dents,
much shorter than the calyx.
19. Blade orbicular, angular to slightly lobed. Seeds 6—7 per cell.

H. floccosus Mast.
19. Blade ovate, not lobed. Seeds 2 per cell 12. H. teijsmannii

1. Hibiscus aruensis Borss., nov. sp.—Fig. 2

Arbor altior. Ramuli teretes, tomentosi pilis stellatis minutis, pilis
quoque stellatis majoribus dispersis et pilis simplicibus longioribus paten-
tibus, denique glabrescentes. Stipulae plerumque appressae, late annexae,
ovatae vel triangulares, basi cordatae et auriculatae, apice obtusae vel
acutae vel subacuminatae, velutinae pilis stellatis minutis. Petiolus lamina
paulo brevior, teres, indumento sicut in ramulis. Lamina coriacea, late
elliptica, basi plerumque subcordata, saepe rotundata vel truncata, apice
acuta vel breviter cuspidata, margine integra, basi 5—9-nervata; costa va-
lidissima, in apicem percurrens, in pagina inferiore paulum supra basin
nectario lineari ornata; nervi laterales utrimque 3—5, sicut nervo basales
ceteri, erecto-patentes, recti sed prope marginem sursum curvati et in venis
ramosi; lamina in pagina superiore pilis stellatis minutis dispersis prae-
cipue in nervis, glabrescens, in pagina inferiore tomentosa pilis stellatis
minutis et pilis simplicibus dispersis. Flores solitarii in axillis superio-
ribus. Pedunculus brevis, teres, paulum supra basin articulatus, supra
articulum paulo crassior quam infra articulum, indumento sicut in ramulis.
Epicalicis segmenta 9, a calice et inter se separata, calice breviora, epi-
calicem cupulatum vel campanulatum componentia, lanceolata, basi sub-
auriculata, apice acuminata, in sicco prima specie linearia reduplicatione
marginis, extus intusque velutina pilis stellatis minutis. Calyx in statu
alabastri 5-alatus; calyx per anthesin subcampanulatus, 5-partitus, seg-
mentis lanceolatis vel longe triangularibus acutis vel breviter acuminatis,
5-nervis extus paulum prominentibus in apices segmentorum percurren-
tibus, extus velutinus pilis stellatis minutis, intus segmentis velutinus
pilis stellatis minutis, ceterum glaber. Corolla magna; petala longe obovata,
ad basin angustata, apice rotundata, extus praecipue ad basin pilis stella-
tis multibrachiatis densius vestita, intus ad marginem obtegentem pilis
stellatis tenuibus dispersis, et basi pilis stellatis longibrachiatis dense
ornata. Columna staminalis petalis circiter aequilonga, in parte superiore
stamina gerens, praecipue in parte inferiore pilis stellatis majoribus di-
spersis. Ovarium conicum, acutum, hirsutum, 10-loculare, multiovulatum;
stigmata discoidea. Pedunculus post anthesin paulum elongatus et incrassa-
tus; epicalyx non amplificatus; calyx paulum amplificatus. Capsula longe
ovoidea, acuminata, 10-locularis, hirsuta pilis simplicibus erectopatentibus
nitentibus, inter pilos hirsutos velutina pilis stellatis minutis. Semina
numerosa. reniformia, pilis stellatis lanatis ferrugineis dense vestita.

TYPE,—Buwalda 5270 (BO 116805).
Twigs 2—4 mm thick. Stipules 10—12 mm long and 7—10 mm wide.

Petiole 3—8 cm long and about 2 mm thick. Blade 6—17 cm long and
4.5—10.5 cm wide; nectary 1.5—2 cm long. Peduncle about 1.5—2.5 long
FIG. 2. Hibiscus aruensis Borss.: fruiting branch; a, corolla with staminal column. —

After Buwalda 5270 (BO 140251).

1956] VAN BORSSUM WAALKBS: Notes on Malaysian Malvaceae—/ 45



46 R E I N W A R D T I A [VOL. 4

and about 2 mm thick. Epicalyx segments about 2 cm long and 0.5 cm
wide. Calyx about 3 cm long and 1.5—2 cm wide; segments about 2 cm
long and 7—8 mm wide. Petals 7—8 cm long and 2—2.5 cm wide. Staminal
column 6—7 cm long; antheriferous part 1.5—2 cm long; filaments 5—7
mm long; anthers about 0.5 mm across. Ovary about 4 mm high and 3 mm
across. Style about 7 cm long; branches about 5 mm long; stigmas about
1 mm across. Capsule 3—4 cm long and 1.5 cm in diameter. Seeds about
2.5 mm long.

SPECIMENS EXAMINED.—MOLUCCAS. A r u I s . : Pulau Wokam, Selibatabata, pri-
mary forest, sloping ground, 40 m alt., tree 21 m, trunk 13 m high and 30 cm thick
breast high, flowers pink, local name Mala, June 18, 1938, Buwalda 5270 = Buwalda
For. Inst. 380 = 66. 25U1U (BO 116805 holotype, 116806, 140251, 57106, BZF, GH, L
950.298-391, K, SING; duplicates will be sent to A, BRI, PNH). — NEW GUINEA.
W e s t e r n p a r t : Idenburg R., Bernhard Camp, primary forest, on temporary
inundated ground, 50 m alt., tree 20 m, trunk 15 m thick breast high, flowers violet,
July 23, 1938, Meijer Drees 336 = bb. 25688 (BO 58593, BZF, L 950.251-514), 4 km
SW of Bernhard Camp, primary rain forest, 850 m alt., tree 22 m, trunk 45 cm in
diameter, flowers violet, March 1939, Brass & Versteegh 13126 (BO 58593, GH, L
950.51-763).

The type material was already recognised as belonging to a new
species by S. Hatusima during his stay at Bogor. He named it (in schedula)
Hibiscus aruensis. The species is closely related to H. d'albertisii F. Mull.
It differs especially in the greater number of epicalyx segments and the
nectaries on the midribs of the leaves. The specimens from New Guinea
differ from the type material in a somewhat thinner indumentum and in
somewhat smaller flowers and fruits.

2. Hibiscus pleijtei Borss., nov. sp.—Fig. 3

Arbor altior. Ramuli angulares, denique teretes, dense velutini pilis
stellatis minutissimis, glabrescentes, denique lenticulis paucis instructi.
Stipulae majores, appressae, late annexae, ovatae, apice obtusae vel acutae,
pilis stelltis minutis vestitae vel subglabrae. Petiolus lamina multo bre-
vior, teres, apice paulum incrassatus et ibi lenticulis transversis instructus,
pilis stellatis minutis singulis, vel glaber. Lamina coriacea, ovata, basi
rotundata et vulgo subcordata, apice obtuse cuspidata, margine integra,
basi 5-nervata; costa validissima, in apicem percurrens, basi in pagina
inferiore nectario lineari ornata; nervi laterales utrimque circa 6, sicut
nervi basales ceteri erecto-patentes, ante marginem sursum curvati et in
venis ramosi; lamina in utraque pagina subglabra sed nervi in pagina in-
feriore pilis stellatis minutis singulis. Flores solitarii in axillis superioribus
vel parvitate vel abortu bractearum in racemis terminalibus paucifloris.
Pedunculi sursum gradatim crassiores, sine articulo, velutini pilis stellatis
minutissimis, glabrescentes. Epicalyx a calice separatus, substellaris, 6-
partitus, segmentis ovatis vel lanceolatis acutis vel subacuminatis, extus
pilis stellatis minutis singulis, intus densius stellate pilosus. Calyx in statu

1956] VAN BORSSUM WAALKES: Notes on Malaysian Malvaceae—/ 47

FIG. 3. Hibiscus pleijtei Borss.: flowering branch; a, corolla with staminal column
(1 X) ; b, calyx and epicalyx from below (1 x) ; c, cross-section of ovary (5 x) ;

d, base of blade from below. — After Pleijte 510 (BO 116999).



48 R E I N W A R D T I A [VOL, 4

alabastri 5-alatus; calyx per anthesin campanulatus-stellaris, epicalice
longior, 5-partitus, segmentis ovatis acutis reduplicativis, obsolete nervatus,
extus squamis fimbriatis minutis densius vesititus, intus in segmentis velu-
tinus pilis stellatis minutis et basi annulo pilis brevibus sericeis ornatus.
Petala spathulata, apice rotundata, extus squamis fimbriatis minutis dense
vestita, intus basi pulvinulo pilorum lanatorum ornata, ad apicem pilis
stellatis minutis tenuibus dense vestita. Columna staminalis petalis bre-
vior, basi pulvinulis petalorum anguste circumdata, subglabra, in parte
superiore stamina gerens. Ovarium conicum, 5-angulare, acutum, 10-locu-
lare, minute squamatum, multiovulatum; stigmata capitellata. Capsula
ignota.

TYPE.—Pleijte 510 (BO 116999).
Twigs 2—5 mm thick. Stipules 10—12 mm long and 6—5 mm wide. Pe-

tiole 1—5 cm long and about 2 mm thick. Blade 7—20 cm long and 4.5—14
cm wide; nectary 5—10 mm above the base of the blade, 4—5 mm long.
Peduncle about 2 cm long and 1.5 mm thick. Epicalyx about 1.5 cm in
diameter, segments 8—9 mm long and about 4 mm wide. Calyx about 2 cm
in diameter, segments 8—10 mm long and about 5 mm wide. Petals 2.5—3
cm long and 0.5—1 cm wide. Staminal column 1.5—2 cm long, antheri-
ferous part 0.5—1 cm long. Ovary about 4 mm long and in diameter. Style
about 2 cm long; branches about 3 mm long.

SPECIMENS EXAMINED.—NEW G U I N E A . W e s t e r n p a r t : Kadamak (vicinity
of Sorong), open place in forest, 50 m alt., tree 25 m, t r u n k 30 cm in diameter, flowers
red, Aug. 11, 1948, Pleijte 510 (BO 116999 holotype, 117000; duplicates will be sent to
A, BRI, K, L, P, P N H , SING).

This species has the peculiar hair cushions on the petals and the
capitellate stigmas in common with the next species, but the epicalyx and
the indumentum point to relationship with H. d'albertisii F. Mull, and allies.

3. Hibiscus pulvinulifer Borss., nov. sp.—Fig. 4

Arbor altior. Ramuli angulares, denique teretes, paulum scabri, pilis
stellatis et squamis fimbriatis minutis dense vestiti, denique glabrescentes
et lenticulosi. Stipulae ignotae. Petiolus lamina multo brevior, teres, pilis
stellatis et squamis fimbriatis minutis dense vestitus. Lamina coriacea, late
elliptica, basi rotundata vel obtusa, apice obtusa vel acuta, margine integra,
basi 3-nervatis; costa validissima, in apicem percurrens, sine nectario;
nervi laterales utrimque 4—5, sicut nervi basales ceteri erecto-patentes, an-
te marginem sursum curvati et in venis ramosi; lamina squamis fimbriatis
minutis in pagina superiore paulum, in pagina inferiore dense vestita, et
ibi in angulis inter costam et nervos laterales stellate pilosa. Flores in
panicula laxa, parvi; paniculae axes teretes, indumento sicut in ramulis;
pedunculi in sicco sulcati, sine articulo, pilis stellatis et squamis fimbriatis
minutis dense vestiti; bracteae absentes. Epicalyx a calice separatus,
coriaceus, stellaris, 5-fidus, segmentis triangularibus acutis, extus squa-
matus, intus glaber. Calyx in statu alabastri 5-costatus; calyx per anthesin

1956] VAN BOKSSUM WAALKES: Notes on Malaysian Malvaceae—I 49

FIG. 4. Hibiscus pulvinulifer B o r s s . : f l o w e r i n g a n d f r u i t i n g b r a n c h : a, flower (2 x) ;
b , c a l y x w i t h o v a r y ( 2 x ) ; c , c a l y x a n d e p i c a l y x from below ( 2 X ) ; d , l o n g i t u d i n -
al section of c a p s u l e (2 x) ; e, cross-section of c a p s u l e (2 x ) . — A f t e r D o c t e r s

van L e e u w e n 9280 (BO 117013).



50 REINWARDTIA [VOL. 4

epicalice longior, coriaceus, cupulatus-campanulatus, 5-fidus, segmentis
patentibus, ovatis, acutis, obsolete nervatus, extus squamis majoribus
integris dense obtectus, intus velutinus pilis stellatis minutis. Petala
linearia vel lanceolata, apice acuta vel obtusa, extus squamibus magnis
fimbriatis dense vestita, intus in marginem et apicem versus pubescentia
pilis stellatis longibrachiatis, intus basi pulvinulo pilorum lanatorum
ornata. Columna staminalis petalis brevior, basi pulvinulis petalorum
anguste cireumdata, pro parte inferiore dense stellate pilosa, pro parte
superiore stamina gerens. Ovarium conicum, obsolete 5-angulare, squa-
matum, 10-loculare; stigmata capitellata. Pedunculus post anthesin paulum
elongatus; epicalyx et calyx paulum amplificati; calicis segmenta mar-
ginibus revoluta. Capsula calice circiter aequilonga, globosa vel obovoidea,
breviter rostrata, 5-costata, squamis integris magnis dense obtecta, 10-
locularis, loculis 1-seminiferis. Semina reniformia, pilis longis lanatis
ferrugineis dense ornata.

TYPE.—Docters van Leeuwen 9280 (BO 117012)
Twigs 2—3.5 mm thick. Petiole 1—-2.5 cm long and 1—1.5 mm thick.

Blade 5—10 cm long and 4.5—7.5 cm wide. Peduncle 0.5—1 cm long and
about 1 mm thick. Epicalyx 4—5 mm in diameter; segments 1.5—2 mm
long and about 1.5 mm wide. Calyx about 8 mm high and 10 mm wide;
segments 4—-5 mm long and about 3 mm wide. Corolla about 2.5 cm in
diameter; petals 1.5—2 cm long and 0.3—0.4 cm wide. Staminal tube
about 1 cm long. Ovary 2.5—3 mm high and 2.5—3 mm in diameter. Cap-
sule about 6 mm in diameter. Seeds 1.5—2 mm across.

SPECIMENS EXAMINED.—NEW G U I N E A . W e s t e r n p a r t : Van Rees Mts.,
Van Gelderen R., primary forest, 100 m alt., tree 25 m, flowers pink, May 1926,
Docters van Leeuwen 9280 (BO 117012 holotype, 117013, 117014; duplicates will be
sent to A, K, L, P N H , SING).

The species attractts attention by its small flowers with hair cushions
on the petals (hence the name). The form of the epicalyx and the scaly
indumentum indicates a relationship with H. sciadiolepidus (Hochr.) Borss.
which, however, has much larger flowers, discoid stigmas, and does not
possess hair cushions on the petals.

4. Hibiscus leeuwenii Borss., nov. sp.—Fig. 5

Arbor altior. Ramuli teretes, tomentosi pilis stellatis, scabriusculi,
glabrescentes, denique lenticulis minutis dispersis ornati. Stipulae erecto-
patentes vel patentes, late annexae, ovatae, apice breviter acuminatae, in
pagina superiore velutinae pilis stellatis minutis, in pagina inferiore to-
mentosae pilis stellatis. Petiolus lamina brevior, teres, densius tomentosus
pilis stellatis, paulum scaber. Lamina chartacea, orbicularis, basi cordata,
apice obtusa, acuta vel breviter acuminata, margine integra, basi 5—7-
nervata; costa validissima, in apicem percurrens, basi in pagina inferiore
nectario lineari ornata; nervi laterales utrimque 4—5, sicut nervi basales
erecto-patentes, sursum curvati et ante marginem in venis ramosi; lamina

1956] VAN BORSSUM WAALKES: Notes on Malaysian Malvaceae—I 51

d, seed (5 X ) . — A l t e r Docters van 9113 (BO 106686).



52
R E I N W A R D T I A [VOL. 4

in pagina superiore scabriuscula pilis stellatis minutis, in pagina inferiore
subtomentosa pilis stellatis. Flores solitarii in axillis superioribus. Pedun-
culus brevior, teres, sine articulo, in sicco sulcatus, tomentosus pilis stel-
latis. Epicalyx a calice separatus, subcampanulatus, 4-partitus, segmentis
inter se brevissime connatis ovatis basi cordatis subauriculatis apice acutis,
extus intusque scabriusculus pilis stellatis minutis, pilis quoque stellatis
magnis tenuibus dispersis. Calyx campanulatus, plerumque epicalice paulo
brevior, 5-fidus, segmentis ovatis obtusis vel acutis, 5 nervis extus pro-
minentibus in apices segmentorum percurrentibus atque 5 nervis extus
prominentibus ad sinum percurrentibus et paulum ante sinum furcatis,
extus scabriusculus pilis stellatis minutis, pilis quoque stellatis magnis
tenuibus dispersis, intus velutinus pilis stellatis minutis. Petala spathulata,
ad basin gradatim angustata, apice rotundata, extus praecipue ad basin
squamis fimbriatis minutis densius vestita, intus ad marginem obtegentem
pilis stellatis minutis, basi fimbriata pilis stellatis longis sericeis. Columna
staminalis petalis circiter aequilonga, basi pilis stellatis longis sericeis et
squamis fimbriatis, in parte superiore stamina gerens. Ovarium ovoideum,
obsolete 5-angulare, acutum, hirsutum, 10-loculare; stigmata discoidea.
Pedunculus post anthesin paulum elongatus et incrassatus; epicalyx post
anthesin non amplif icatus; calyx post anthesin amplificatus, epicalice
longior. Capsula longe ovoidea, acuminata, distincte 5-angularis, extus
hirsuta pilis simplicibus appressis nitentibus, pilis quoque stellatis minutis,
10-loeularis; valvis ligneis, intus minute breviter pilosis. Semina numerosa,
reniformia, valde curvata, indumento denso pilorum stellatorum lanato-
rum ferrugineorum.

TYPE.—Docters van Leeuwen 9113 (BO 106686).
Twigs 3—6 mm thick. Stipules about 12 mm long and 7—8 mm wide.

Petiole 4—10 cm long and about 1.5 mm thick. Blade 7—18 cm in diameter;
nectary 4—10 mm long. Peduncle 0.5—1.5 cm long and about 1.5 mm thick.
Epicalyx about 3 cm wide and high; segments 2.5—3.5 cm long and 1.5—2
cm wide. Calyx about 2.5 cm high and 2 cm wide; segments about 1.3 cm
long and 0.8 cm wide. Petals about 5 cm long and 1.5 cm wide. Staminal
column about 5 cm long; antheriferous part about 2 cm long; filaments
about 4 mm long; anthers about 1 mm long. Ovary about 6 mm long and
3—4 mm in diameter; style about 5 cm long; branches about 3 mm long;
stigmas about 1.5 mm across. Capsule about 3 cm long and 1 cm in dia-
meter. Seeds about 3 mm long.

SPECIMENS EXAMINED.—NEW G U I N E A . W e s t e r n p a r t : Mamberamo R.,
Albatros Bivouac, p r i m a r y forest, 30 m alt., tree 15 m, flowers red, May 1926, Docters
van Leeuwen 9113 (BO 106686 holotype, 106687; duplicates will be sent to A, K, L ) ,
Pioneers Bivouac, p r i m a r y forest, 30 m alt., tree 20 m, t r u n k 12 m high and 40—25 cm
thick, flowers lilac, Oct. 18, 1939, van Eechoud U = bb. 31108 (BO 57165, B Z P , L
950.252-443).

The species is apparently closely allied to H. d'albertisii F. Miill., but
differs by having four epicalyx segments, a coarser stellate indumentum,
and leaf blades which are more or less orbicular instead of ovate.

1956] VAN BORSSUM WAALKBS : Notes on Malaysian Malvaceae—/ 58

5. Hibiscus fluminis-idenburgii Borss., nov. sp.—Fig. 6

Arbor altior. Ramuli teretes, dense tomentosi pilis stellatis magnis et
pilis stellatis minutis. Stipulae patentes vel appressae, latius annexae,
ovatae, apice acutae, in pagina superiore velutinae pilis stellatis minutis,
in pagina inferiore tomentosae vel velutinae pilis stellatis magnis et pilis
stellatis minutis. Petiolus lamina brevior, teres, indumento sicut in ramu-
lis. Lamina coriacea, late ovata, basi cordata, apice acuta vel acuminata,
margine integra, basi 5—7-nervata; costa validissima, in apicem percur-
rens, sine nectario; nervi laterales utrimque 3—4, sicut nervi basales
ceteri erecto-patentes, sursum curvati et ante marginem in venis ramosi;
laminae pagina superior pilis stellatis dispersis sed in nervis tomentosa
pilis stellatis, pagina inferior dense tomentosa pilis stellatis magnis
minutisque. Flores solitarii in axillis superioribus. Pedunculus brevis,
non articulatus, teres, in sicco paulum sulcatus, dense tomentosus pilis
stellatis magnis minutisque. Epicalyx a calice separatus, ample cupulatus;
segmenta 5—6, inter se libera, erecto-patentes, ovata, basi subauriculata,
apice acuta vel paulum acuminata, margine plerumque reduplicativa, extus
tomentosus pilis stellatis magnis minutisque, intus velutinus pilis stellatis
minutis. Calyx in statu alabastri 5-alatus; per anthesin ample tubiformis,
paulum inflatus, epicalice circiter duplo longior, 5-fidus vel 5-partitus,
segmentis ovatis (ligulatis) apice obtusis, extus nervis 5 paulum promi-
nentibus in apicem segmentorum percurrentibus, extus densissime tomen-
tosus pilis stellatis magnis minutisque, intus velutinus pilis stellatis minu-
tis et ad basin pilis quoque stellatis magnis dispersis. Petala spathulata,
apice rotundata, extus praecipue ad basin squamis fimbriatis dense vestita,
intus ad marginem obtegentem pilis stellatis minutis, ceterum glabra.
Columna staminalis petalis paulo brevior, praecipue ad basin stellate
pilosa, in parte superiore stamina gerens. Ovarium ovoideum vel globosum,
acutum, obsolete 5-angulare, dense hirsutum; stigmata discoidea. Pedun-
culus post anthesin elongatus et incrassatus; epicalyx non amplif icatus;
calyx paulum amplificatus. Capsula longe ovoidea, acuta, extus hirsuta
pilis simplicibus erecto-patentibus longis nitentibus, pilis quoque stellatis
minutis, 10-locularis, valvis ligneis. Semina numerosa, reniformia, corona
densa pilorum stellatorum longorum lanatorum ferrugineorum ornata.

TYPE.—Brass 12978 (BO 116799).
Twigs 2—3 mm thick. Stipules 1.5—2 cm long and about 1 cm wide.

Petiole 3—5 cm long. Blade 8—13 cm long and 7—11 cm wide. Peduncle
1—1.5 cm long and about 2 mm thick. Epicalyx 1.5—2 cm high and about
3 cm wide; segments about 2 cm long and 1 cm wide. Calyx 3—-3.5 cm long
and about 3 cm wide; segments about 2 cm long and 1 cm wide. Petals
8—9 cm long and 2.5—3 cm wide. Staminal column 6—7 cm long; antheri-
ferous part about 2 cm long; filaments 3—4 mm long; anthers about
1.5 mm long. Ovary about 2.5 mm in diameter. Capsule about 4 cm long.
Seeds about 2.5 mm long.

SPECIMENS EXAMINED.—NEW G U I N E A . W e s t e r n p a r t : Idenburg R., 6 k m
SW of Bernhard Camp, rain forest, plentiful on the slopes of ridges and in ravines,
1200 m alt., tree up to 35 m, flowers deep pink, Feb. 1939, Brass 12978 (BO 116799



54
R E I N W A R D T I A

[VOL. 4

Fig.6. Hibiscus

1956] VAN BORSSUM WAALKES : Notes on Malaysian Malvaceae—7 55

holotype, BH, L 950.51-793), frequently in primary forest, on slope of ridge, 1100 m
alt., tree 37 m, trunk 48 cm in diameter, wood white, flowers dark red, Feb. 1939,
Brass & Versteegh 12525 (BO 116798, GH, L 950.51-794) = 66. 27343 (BZF), 15 km
SW of Bernhard Camp, occasionally in primary forest, on slopes of ridges, 1750 m
alt., tree 31 m, trunk 63 cm in diameter, sap-wood white, heart-wood dark brown,
Jan. 1939, Brass & Versteegh 11959 (BO 116797, GH, L 950.51-795) = 66. 26U8 (BZF).

This species is without doubt most nearly related to H. d'albertisii
F. Mull. It differs by the relatively short epicalyx segments and the ex-
tremely dense indumentum which determines the striking habitus. Brass
& Versteegh 11959 — bb. 26948 has an indumentum which is somewhat
less dense than the indumentum of the other numbers although still much
denser than in H. d'albertisii.

6. Hibiscus ellipticifolius Borss., nov. sp.—Fig. 7
Arbor humilis. Ramuli teretes, scabriusculi pilis stellatis dispersis,

glabrescentes, denique seriebus lenticulorum minutorum ornati. Stipulae
patentes, late annexae, lanceolatae vel ovatae, acutae vel paulum acumina-
tae. Petiolus lamina brevior, teres, apice paulum incrassatus, scabriusculus
pilis stellatis. Lamina chartacea, elliptica, raro ovata, basi rotundata vel
truncata vel paulum cordata, apice breviter cuspidata, margine integra,
basi 5—7-nervata; costa validissima, in apicem percurrens, in pagina
inferiore paulum supra basin nectario lineari ornata; nervi laterales
utrimque circiter 4, sicut nervi basales ceteri erecto-patentes, paulum sur-
sum curvati et ante marginem in venis ramosi; lamina in pagina superiore
pilis stellatis brevibus dispersis praecipue in nervis, glabrescens, in pagina
inferiore scabriuscula indumento densiore pilorum stellatorum praecipue
in nervis. Flores solitarii in axillis superioribus. Pedunculus brevis, prope
apicem articulatus vel non articulatus, teres, in sicco sulculatus, apice in-
crassatus, subtomentosus pilis stellatis minutis. Epicalicis segmenta 6,
libera, stellate patentia, lanceolata, calice breviora, basi angustata, apice
acuta vel acuminata, intus multo magis quam extus pilis stellatis minutis
dispersis. Calyx in statu alabastri ad apicem anguste 5-alatus; calyx per
anthesin campanulatus, 5-fidus, segmentis longe triangularibus, 5-nervis
extus prominentibus ad sinum percurrentibus et paulum ante sinum fur-
catis, extus scabriusculus pilis stellatis minutis multis et pilis stellatis
magnis paulis, intus velutinus pilis stellatis minutis. Petala longe obovata,
basi gradatim angustata, apice rotundata, extus praecipue ad basin pilis
stellatis densius vestita, margine fimbriata pilis stellatis longibrachiatis,
intus praecipue ad marginem obtegentem pilis stellatis minutis tenuibus.
Columna staminalis petalis brevior, in parte inferiore pubescens pilis stel-
latis longibrachiatis tenuibus, in parte superiore glabra et stamina gerens.
Ovarium extus hirsutum, 10-loculare; stigmata discoidea. Pedunculus post
anthesin paulum elongatus et incrassatus, glabrescens; epicalicis segmen-
ta reflexa, paulum elongata; calyx elongatus et amplificatus. Capsula
ovoidea, acuta, extus hirsuta pilis simplicibus longis erecto-patentibus
nitentibus, pilis quoque stellatis minutis, 10-locularis. Semina ignota.

TYPE.—N.G.F. 3689 (LAE).



56 R E I N W A R D T I A [ V O I J . 4 1956] VAN BORSSUM WAALKES; -.Notes on -Malaysian Malvaceae—/

FIG. 7. Hibiscus ellipticifolius Borss.: flowering branch; a,
column. — After N.G.F. 3689 (LAE).

corolla with staminal

Twigs 2—3 mm thick. Stipules 7—9 mm long and about 4 mm wide'.
Petiole 2.5—5.5 cm long and about 1 mm thick. Blade 9—13 cm long and
5 9 cm wide; nectary 5—10 mm long. Peduncle 10—15 mm long and
about 1.5 mm thick. Epicalyx segments 10—12 mm long and about 3 mm
wide. Calyx about 2 cm long and 0.8 cm wide; segments 8—9 mm long
and about 5 mm wide. Petals about 7 cm long and 2 cm wide. Staminal
column about 6 cm long; antheriferous part about 5.5 cm long. Capsule
about 2 cm long and 1.5 cm in diameter.

SPECIMENS EXAMINED.— N E W G U I N E A . E a s t e r n p a r t : Sepik Distr., near
edge of Karosomeri R., tree 7.5 m high, straight, flowers pink, wedges of white homo-
genous tissue penetrating the bark, Aug. 10, 1949, WomersUy N.G.F. S689 (LAE
holotype, BO 116807, BRI, GH, L 950.341-542, SING).

The species is without doubt closely allied to H. fluminis-wprilii Ulbr.
The name was choosen on account of its perfectly elliptical leave blades,
these being rare within the genus.

7. Hibiscus carrii Borss., nov. sp.—Fig. 8

Arbor humilis. Ramuli teretes, subtomentosi pilis stellatis, squamis
quoque fimbriatis minutis paucis, glabrescentes, denique lenticulis ornati.
Stipulae ignotae. Petiolus lamina brevier, basi et apice paulum incrassatus,
tomentosus pilis stellatis. Lamina coriacea, ovata, basi rotundata vel
truncata, apice acuta vel acuminata, margine integra, basi 3-nervata;
costa validissima, recta, in apicem percurrens, in pagina inferiore paulum
supra basin nectario elliptico ornata; nervi laterales utrimque 4—5, sicut
nervi basales ceteri, erecto-patentes, ante marginem sursum curvati et in
venis ramosi; nervi omnes in pagina inferiore prominentes; lamina in
pagina superiore squamis fimbriatis minutis paueis, in pagina inferiore
tomentosa pilis stellatis, squamis quoque fimbriatis minutis. Flores ignoti.
Capsulae in panicula paucibrachiata; bracteae et stipulae absentes; axes
teretes, subangulares, squamis minutis dense obtecti; pedunculus post an-
thesin teres, sensim crassior in apicem, sine articulo, squamis minutis den-
se obtectus. Epicalyx post anthesin a calice separatus, cupulatus, 5-fidus,
segmentis appressis late triangularibus obtusis vel acutis, extus squamis
minutis, intus glaber. Calyx post anthesin epicalice triplo longior, cam-
panulatus, capsulam anguste involvens, 5-lobatus, segmentis revolutis vel
reflexis longe triangularibus acutis, obsolete nervatus, extus dense minute
squamatus, intus in segmentis velutinus pilis stellatis minutissimis. Cap-
sula longe obovoidea, basi subacuta, apice conglobata sed centro cuspide
instructa, extus squamis majoribus dense obtecta, 10-locularis, valvis sub-
ligneis, intus levis; dissaepimenta vera per marginem intimam pilis ap-
pressis longis sericeis vestita. Semina numerosa, reniformia, corona densa
pilorum longorum lanatorum ferrugineorum ornata.

TYPE.—Carr 13219 (L 936.295-22).
Twigs 3—4 mm, thick. Petiole 4—8 cm long and 1.5—2 mm thick.

Blade 11—19 cm long and 3—12 cm wide; nectary 1—2 cm above the base,
about 1 mm long. Peduncle when in fruit 2—2.5 cm long and from base
to apex 2—4 mm thick. Epicalyx during fruiting 4—5 mm high and about
8 mm in diameter; segments about 2.5 mm long and 3.5 wide. Calyx
during fruiting 13—15 mm high and about 12 mm wide; segments about
7 mm long and 4—5 mm wide. Capsule about 2.5 cm long and 1 cm in
diameter. Seeds about 2 mm across.

S P E C I M E N S E X A M I N E D . — N E W G U I N E A . E a s t e r n p a r t : B o r i d i , f o r e s t ,

4200 ft. a l t . , t r e e of 40 ft., S e p t . 1 3 , 1935, Carr 13219 (L 936.295-22 h o l o t y p e ) .

It is a pity that Carr 13219 has no flowers, but the indumentum and
the epicalyx point to a very close relationship with H. sciadiolepidus
(Hochr.) Borss.



R E I N WARDTIA [VOL. 4

. . . U V

FlO. 8. H i b i a w s carrii Borss.: fruiting branch. — After Carr 13219 (L 986.295-22).

1956] VAN BoESSUM WAALKES: Notes on Malaysian Malvaceae—/

8. Hibiscus archboldianus Borss., nov. sp.—Fig. 9

Arbor altior. Ramuli angulares vel paulum applanati, denique teretes,
punctati squamis fimbriatis minutis dispersis, paulum scabri. Stipulae
erecto-patentes, late annexae, rigide coriaceae, ovatae, apice acutae, in
utrisque paginis squamis fimbriatis minutis dense vestitae. Petiolus lamina
brevior, teres, rigidus, praecipue apice squamis fimbriatis minutis dense
vestitus. Lamina rigide coriacea, ovata vel elliptica, basi rotundata vel
paulum cordata, apice cuspidata, margine integra, basi 5—7-nervata;
costa validissima, in apicem percurrens, sine nectario; nervi laterales
utrimque 3—4, sicut nervi basales ceteri erecto-patentes, paulum sursum
curvati et ante marginem in venis ramosi; lamina in pagina superiore in
nervis squamis fimbriatis minutis, in pagina inferiore paulum scabra,
praecipue in nervis squamis fimbriatis minutis. Flores solitarii in axillis
superioribus. Pedunculus brevis, teres, in sicco sulcatus, apice incrassatus,
sine artieulo, scaber indumento squamorum fimbriatorum minutorum.
Epicalicis segmenta 5, a calice et inter se separata, calice breviora, erecto-
patentia, coriacea, late ovata, basi abrupte angustata, apice acuminata,
extus intusque squamis fimbriatis minutis densius vestita. Calyx in statu
alabastri ad apicem 5-alatus; calyx per anthesin ample campanulatus,
5-fidus vel 5-partitus, segmentis ovatis acutis, 5 nervis extus prominenti-
bus in apicem segmentorum percurrentibus, extus scabrissimus, squamis
fimbriatis rigidis minutis dense vestitus, intus velutinus pilis stellatis
minutis. Petala magna, obovata vel spathulata, ad basin gradatim angus-
tata. apice rotundata, extus praecipue ad basin squamis fimbriatis rigidis
densius vestita, scabra, margine ad basin pilis stellatis lanatis, intus ad
basin squamis fimbriatis, basi pilis stellatis tenuibus, ceterum glabra.
Columna staminalis petalis circiter aequilonga, basi pilis stellatis lanatis,
praecipue ad basin squamis fimbriatis, in parte superiore stamina gerens.
Styli brachii brevi-pubescentes; stigmata discoidea. Fructus ignoti.

TYPE.—Brass 7092 (BO 116774).
Twigs 2—-4 mm thick. Stipules about 1.5 cm long and 0.8 cm wide.

Petiole 3—8 cm long and about 1.5 mm thick. Blade 8.5—16 cm long and
4.5—12 cm wide. Peduncle 1—1.5 cm long and about 2 mm thick. Epi-
calyx segments about 2 cm long and 1—1.5 cm wide. Calyx about 3 cm
high and 2.5 cm in diameter; segments 1.5—2 cm long and about 1 cm
wide. Petals about 10 cm long and 4 cm wide. Staminal column about
8.5 cm long; antheriferous part about 4 cm long; filaments 3—5 mm
long; anthers about 1 mm long. Style about 8.5 cm long; branches about
10 mm long; stigmas 2—2,5 mm across.

SPECIMENS EXAMINED.—NEW G U I N E A . E a s t e r n p a r t : Palmer R., 2 miles
below junction of Black R., common on low ridges along the river, 100 m alt., slender
canopy tree 25 m high, flowers pink, J u n e 1936, Brass 7092 (BO 116774 holotype,
BM, GH, L 950.88-561).

This species and the two next ones are closely related to each other
and together form a more or less natural group. As to the shape of the
epicalyx, the three species are related to H. d'albertisii F. Mull. They



60 R E I N W A R D T I A [VOL. 4

F I G . 9. Hibiscus archboldianus Borss.: flowering branch; a, petal; b, staminal tube
with stigmas. — After Brass 7092 (BO 116774).

19B6] VAN BORSSUM WAALKES: Notes on Malaysian Malvaceae—/ 61-

differ by their scaly indumentum. The name is given to commemorate R.
Archbold, the organizer of the famous expeditions which Brass joined as
botanist.

9. Hibiscus womersleyanus Borss., nov. sp.—Fig. 10

Arbor alta. Ramuli paulum angulares, denique teretes, squamis mi-
nutis densius vestiti, glabrescentes, denique lenticulis dispersis ornati,
paulum scabri. Stipulae erecto-patentes vel subreflexae, late annexae,
ovatae, apice obtusae, in pagina superiore squamis firnbriatis minutis et
pilis stellatis minutis vestitae, in pagina inferiore squamis densius ves-
titae. Petiolus lamina brevior, teres, in sicco paulum sulcatus, squamis
minutis densius vestitus. Lamina rigide coriacea, orbicularis vel latissime
ovata, basi paulum cordata, apice rotundata, obtusa vel breviter cuspidata,
margine integra, basi 7—5-nervata; costa validissima, recta, in apicem
percurrens, sine nectario; nervi laterales utrimque 3—5, sicut nervi ba-
sales ceteri erecto-patentes, paulum sursum curvati et ante marginem in
venis ramosi; nervi omnes in pagina inferiore valde prominentes; lamina
in pagina superiore pilis stellatis minutis dispersis, glabrescens, in pagina
inferiore squamis fimbriatis minutis dense vestita. Flores solitarii in
axillis superioribus. Pedunculus teres, apice incrassatus, in sicco paulum
sulcatus, sine articulo, squamis minutis densius vestitus. Epicalicis seg-
menta 6, coriacea, basi inter se breviter connata, a calice separata, calice
breviora, patentia vel reflexa, ovata, apice obtusa, extus squamis minutis
dispersis vestita, intus squamis fimbriatis minutis, glabrescentia. Calyx
in statu alabastri ad apicem 5-alatus; calyx per anthesin campanulatus,
paulum inflatus, coriaceus, 5-fidus vel 5-partitus, segmentis ovatis acutis,
obsolete nervatus, extus squamis minutis dense vestitus, intus velutinus
pilis stellatis minutis. Petala spathulata, apice rotundata, extus praecipue
ad basin scabra, squamis rigidis, intus ad marginem obtegentem pilis
stellatis tenuibus. Columna staminalis petalis brevior, praecipue ad basin
pilis stellatis majoribus dense vestita, in parte superiore stamina gerens.
U varium ovoideum-conicum, obtusum, obsolete 5-angulare, squamis fim-
briatis dense vestitum, 10-loculare; stigmata discoidea. Pedunculus post
anthesin elongatus et incrassatus; epicalyx et calyx post anthesin paulum
amplificati; calyx denique fissus et putescens. Capsula maxima, globosa,
acuta, extus squamis fimbriatis majoribus obtecta, 10-locularis; valvis
ligneis, intus levis, glaber. Semina numerosa, reniformia, indumento
denso lanato pilorum stellatorum longorum.

TYPE.—N.G.F. 3386 (LAE).
Twigs 2.5—5 mm thick. Stipules about 1.5 cm long and 0.5—0.8 cm

wide. Petiole 2—7.5 cm long and 1—2 mm thick. Blade 4—15 cm in dia-
meter. Peduncle 2—2.5 cm long and 2—3 mm thick. Epicalyx segments
1.5—2 cm long and 0.7—1 cm wide. Calyx about 3 cm high and 2.5 cm
wide; segments about 1.5 cm long and 0.8 cm wide. Petals about 7 cm long
and 2 cm wide. Staminal tube about 5 cm long; antheriferous part about
2.5 cm long; filaments about 2 mm long; anthers about 1 mm long. Ovary
about 5 mm high and 6 mm across; style about 5 cm long; branches about



62 R E I N W A R D T I A [VOL, 4

FIG. 10. Hibiscus womersleyanus Borss.: branch; a, calyx and epicalyx (2 X). — After
N.G.F. 4425 (BO 116804).

3 mm long; stigmas about 1.5 mm across. Capsule 4—4.5 cm in diameter.
Seeds 3—4 mm long:

S P E C I M E N S EXAMINED.—-NEW G U I N E A . E a s t e r n p a r t : A i y u r a , C e n t r a l
H i g h l a n d s , 1806 m alt., t r e e 24 m, t r u n k 15 m h i g h a n d 76 cm t h i c k b r e a s t h i g h ,

1956] VAN BOKSSUM WAALKES: Notes on Malaysian Malvaceae, 63

crown narrow, petals red, bark very fibrous and used by natives for making ropes,
wood white and soft, local names Papum (Aiyura) and Bidafu (Kamanu), Nov. 25,
1950, Womersley N.G.F, 3388 (LAE holotype, BO 116802, 116803, BRI, L 951.99-348,
SING), tree 30.5 ra, crown very sparse, flowers orange-red, bark fibrous and used
for string, local name Panpur (Aiyura), June 29, 1951, Womersley N.G.F. 4425 (BO
.116804, BRI, GH, L 952.64-357, LAE, SING), tall tree 40 m, trunk 35.5 m high and
61 cm thick breast high, inner bark brown with paler inwardly pointing teeth, bark
very fibrous, local name Tampo (Anona) and Pundenifa (Aiyura), Dec. 1, 1944,
s. coll. N.G.F. 1047 (LAE).

It may be that this species is simply a form of the previous one. More
material of this relationship should be collected.

10. Hibiscus lepidotus Borss., nov. sp.—Fig. 11

Arbor. Ramuli paulum applanati, denique teretes, punctati squamis
minutis densius vestiti, paulum scabri. Stipulae late ovatae, apice obtusae,
squamis minutis dense vestitae. Petiolus lamina brevior, teres, ad basin
in sicco sulcatus, squamis minutis densius vestitus. Lamina rigide coriacea,
orbicularis, basi paulum cordata, apice breviter acuminata, margine in-
tegra, basi 5-nervata; costa validissima, in apicem percurrens, sine nec-
tario; nervi laterales utrimque 3, sicut nervi basales ceteri erecto-patentes,
paulum sursum curvati et ante marginem in venis ramosi; lamina in pagina
superiore scabriuscula indumento denso pilorum stellatorum minutorum,
in pagina inferiore indumento densissimo squamorum fimbriatorum minu-
torum. Flores solitarii in axillis superioribus. Pedunculus teres, in sicco
sulcacus, sine articulo, squamis fimbriatis minutis dense vestitus, gla-
brescens. Epicalicis segmenta 4, a calice et inter se separata, calice paulum
breviora, coriacea, erecta, magna, late ovata vel suborbicularia, basi pau-
lum cordata, apice obtusa vel rotundata, extus squamis densius vestita, in-
tus squamis fimbriatis et pilis stellatis. Calyx in statu alabastri ad apicem
5-alatus; calyx per anthesin coriaceus, campanulatus, 5-lobatus vel 5-fidus,
segmentis ovatis acuminatis, obsolete nervatus, extus squamis minutis den-
se vestitus, intus velutinus pilis stellatis minutis. Petala spathulata, ad ba-
sin gradatim angustata, apice rotundata, extus praecipue ad basin squamis
fimbriatis majoribus dense vestita, intus ad marginem obtegentem in parte
inferiore pilis stellatis longibrachiatis, ceterum glabra. Columna stami-
nalis petalis circiter aequilonga, praecipue ad basin pilis stellatis longi-
brachiatis, quoque squamis fimbriatis dispersis vestita, in parte superiore
stamina gerens. Stigmata discoidea. Pedunculus post anthesin paulum
elongatus et incrassatus. Epicalyx et calyx paulum vel non amplificati.
Capsula ovoidea vel globosa, apice breviter acuminata, extus squamis
majoribus dense obtecta, 10-locularis. Semina numerosa (misera evoluta),
indumento pilorum lanatorum ferrugineorum circumdata.

TYPE.—Brass 4950 (BO 57181).
Twigs 3—5 mm thick. Stipules 1.5—2 cm long and about 1 cm wide.

Petiole 3—6 cm long and about 1.5 mm thick. Blade 7—11.5 cm in dia-
meter. Peduncle 1.5—2.5 cm long and 1.5 mm thick. Epicalyx segments
2—2.5 cm long and 1.2—2 cm wide. Calyx about 3.5 cm high and 2.5 cm.



R E I N W A R D T I A [VOL. 4

P I G . 11. Hibiscus lepidotus Borss.: flowering branches.
57181, B R I ) .

After Brass 4950 (BO

wide; segments about 1.5 cm long and 1 cm wide. Petals about 7 cm long
and 2 cm wide. Staminal column about 5.5 cm long; antheriferous part
about 2 cm long; filaments about 3 mm long, anthers about 1 mm long.
Style branches 5—6 mm long; stigmas about 2 mm across. Capsula 2—2.5
cm in diameter.

SPECIMENS EXAMINED.—NEW GUINEA, E a s t e r n p a r t : Mt. Tafa, sheltered
valley forests, alt. 2400 m, slender tree, open thinly foliaged crown, t r u n k about 20 cm
thick, bark thick and fibrous, wood soft and white, flowers red, April 1933, Brass
A950 (BO 57181 holotype, BM BRI, GH).

The remarks under the previous species also hold good for this
species.

1956] VAN BOESSUM WAALKES: Notes on Malaysian Malvaceae—/ 65

11. Hibiscus pseudotiliacus Borss., nov. sp.~—Fig. 12

Arbor altior. Ramuli paulum angulares, denique teretes, tomentosi
pilis stellatis majoribus et pilis stellatis minutis, glabrescentes, denique
lenticulis valde prominentibus ornati. Stipulae appressae vel paulum
erecto-patentes, late annexae, semi-amplexicaules, magnae, ovatae vel lan-
ceolatae, obtusae, in pagina superiore pubescentes pilis stellatis minutis
et pilis simplicibus brevibus, in pagina inferiore tomentosae pilis stellatis.
Petiolus lamina multo brevior, teres, apice paulum incrassatus, densius
tomentosus pilis stellatis majoribus et pilis stellatis minutis praecipue
apice. Lamina chartacea, orbicularis, basi penitus cordata, apice longe
cuspidata, margine integra vel leviter dentata, basi 5—7-nervata; costa
in apicem percurrens, sine nectario; nervi laterales utrimque 3—4, erec-
to-patentes, sursum curvatis et in venis ramosi; nervi basales ceteri quoque
versum radiantes, sursum curvati et ante marginem in venis ramosi;
lamina in pagina superiore pubescens pilis stellatis minutis praecipue in
nervis, glabrescens, in pagina inferiore laxe tomentosa pilis stellatis ma-
joribus praecipue in nervis. Flores in racemis compositis axillaribus pau-
cifloris. Axes primarii et secundarii teretes, indumento sicut in ramulis;
bracteae absentes; stipulae plerumque presentes, sicut stipulae foliorum.
Pedunculus brevis, teres, axe secundario tantum articulatus, densissime
tomentosus pilis stellatis magnis, pilis quoque stellatis minutis. Epicalyx
a calice separatus, magnus. calice paulum brevior, ample campanulatus vel
cupulatus, 7-fidus, segmentis ovatis acutis vel breviter acuminatis, extus
pilis stellatis majoribus et pilis stellatis minutis dense vestitus, paulum sca-
ber, intus ad marginem segmentorum velutinus pilis stellatis minutis.
Calyx campanulatus, 5-fidus, segmentis longe triangularibus acutis, 15-
nervatus, autem 5 costis validioribus extus prominentibus in apicem seg-
mentorum percurrentibus et 10 nervis tenuibus extus minus prominentibus
ante apicem segmentorum terminantibus, extus pilis majoribus et pilis
minutis, intus basi annulo pilorum simplicium appressorum instructus, ce-
terum glaber. Petala obovata, apice rotundata, extus inter nervos pilis stel-
latis longibrachiatis vestita praecipue ad basin, ad basin fimbriata, ceterum
glabra. Pedunculus post anthesin paulum elongatus; epicalyx et calyx
paulum amplificati. Capsula ovoidea, 5-angularis, apice breviter acumi-
nata, extus pilis simplicibus rigidis et pilis stellatis minutis ornati, 5-locu-
laris, valvis crassioribus, intus levis, glabra. Semina numerosa, reniformia,
angularia, pilis stellatis lanatis brevibus ferrugineis dense vestita.

TYPE.—bb. 33899 (BO 116796).
Twigs 2—6 mm thick. Stipules 2.5—3 cm long and about 1 cm wide.

Petiole 5.5—12 cm long and 1—2 mm thick. Blade 14—22 cm long and
12—20 cm wide. Peduncle 1—1.5 cm long and about 2 mm thick. Epicalyx
about 2 cm high and 3 cm wide; segments 1—1.5 cm long and 0.6—0.8 cm
wide. Calyx about 2.5 cm high and 2 cm wide; segments about 1.5 cm
long and 0.5—1 cm wide. Petals 6—7 cm long. Capsule about 2 cm long
and 1.7 cm in diameter. Seeds about 4 mm long.

SPECIMENS EXAMINED.—MOLUCCAS. M o r o t a i : Northern Tjao, p r i m a r y forest,
60 m alt., tree 22 m, trunk 15 m high and.S0:cm thick,breast high, flowers yellow, July 2,



R E I N W A R D T I A [VOL. 4

i v

Fig.12 Hibiscus

1956] VAN BORSSUM WAALKBS: Notes on Malaysian Malvaceae—-/ 67

1949, Tangkilisan (Exp. Kostermans) 229 = 66. 33899 (BO 116796 holotype, 116795,
BZP, L 952. 265-325, K, SING).

This species is at the most related to H. borneensis Airy Shaw, but
has nevertheless a more or less isolated position. At first sight the flowers
are reminiscent of H. d'albertisii F. Mull, and relatives, but the capsule is
5-celled. The species is named on account of the form of the leaves and
stipules which gives it the habitus of H. tiliaceus L.

12. Hibiscus teijsmannii Borss., nov. sp.—Fig. 13

Planta lignea (frutex an arbor?). Ramuli teretes, farinosi squamig
fimbriatis minutissimis, glabrescentes, lenticulis eircularibus dispersis.

FIG. 18. Hibiscus teysmamvii Borss.: fruiting branch. — After Teijsmann 12897HB
(BO 58063).



68 E E I N W A R D T I A [VOt. 4

Stipulae ignotae. Petiolus lamina brevior, teres, basi et apice paulum in-
crassatus, farinosus squamis fimbriatis minutissimis, glabrescens. Lamina
coriacea, ovata, basi rotundata vel paulum cordata, apice gradatim acumi-
nata, margine integra, basi 6—4-nervata; costa in apicem percurrens, basi
in pagina inf eriore nectario lineari ornata; nervi laterales utrimque 2—3,
sicut nervi basales ceteri erecto-patentes, paulum sursum curvati et ante
marginem in venis ramosi; lamina in pagina superiore praecipue basi,
squamis fimbriatis minutis dispersis, glabrescens, in pagina inferiore in
angulis inter nervos basales, et inter costam et nervos laterales fasciculis
pilorum simplicium brevium, glabrescens. Blores ignoti. Pedunculi soli-
tarii in axillis superioribus, post anthesin teretes, parte superiore arti-
culati, a basi ad articulum gradatim crassiores, supra articulum crassiores
quam infra articulum et sulcati, squamis fimbriatis minutissimis densius
vestrti. Epicalyx post anthesin cupulatus, a calice separatus, calice mult'o
brevior, 8-fidus, segmentis erecto-patentibus triangularibus acutis, extus
squamis fimbriatis minutis vestitus. Calyx post anthesin cupulatus-cam-
panulatus, capsulam anguste involvens, 5-lobatus, segmentis late triangu-
laribus (apices segmentorum in sicco destructi), 10-nervis extus paulum
prominentibus, squamis integris majoribus dense obtectus. Capsula obo-
voidea, apice acuta, extus dense sericeus pilis simplicibus, inter pilos sim-
plices pilis stellatis minutis, 5-locularis, valvis acutis, crassioribus, ligneis,
intus levibus, glabris et paulum nitentibus; loculus quisque plerumque in
parte superiore seminibus 2 adultis, in parte inferiore seminibus 2 abor-
tivis. Semina adulta magna, reniformia, corona densa pilorum appres-
sorum longorum mollium.

TYPE.—Teijsmann 12597HB (BO 58063).
Twigs 2.5—5 mm thick. Petiole 2—7.5 cm long and 1—2 mm thick.

Blade 7.5—15 long and 5.5—10 cm wide; nectary about 5 mm long. Pe-
duncle during fruiting 3—4.5 cm long; joint 10—13 mm from the apex;
peduncle below the joint about 2 mm thick, above the joint 4 mm thick.
Epicalyx during fruiting about 8 mm high and 14 mm wide; segments
about 0.5 cm long and 1 cm wide. Calyx about 2 cm wide; segments about
1 cm wide. Capsule about 2 cm long and 2.5 cm in diameter. Seeds about
5 mm long.

S P E C I M E N S E X A M I N E D . — C E L E B E S . S o u t h w e s t e r n p e n i n s u l a : S e h r o h

( P a n g k a d j e n e D i s t r . ) , Teijsmann 12597HB (BO 58063 h o l o t y p e , 5 8 0 6 1 , 58062, L

9 2 0 . 3 0 6 - 4 8 ) .

The species is without doubt very closely allied to H. floecosus Mast.
The most obvious differences are the ovate leaves and much smaller fruits.

R E I N W A R D T I A
Published by Herbarium Bogoriense, Kebun Raya Indonesia

Volume 4, Part 1, p.p. 69-74 (1956)

THE GENUS CULLENIA Wight * (Bombacaceae)

A. J. G. H. KOSTERMANS **

The monotypic genus Cullenia was established by Wight (Icones
PI. Ind. or. 5 (1) : pi. 1761—62 & text, 1851), who differentiated it from
Durio Adans. mainly by the lack of a corolla and the position and shape
of the anthers. The only species, originally described as Durio ceylanicus
by Gardner, was cited by Wight as Cullenia excelsa Wight. K. Schumann
corrected the specific epithet rather casually and atributed it (wrongly)
to Wight. Bentham (in Benth. & Hook., Gen. pi. 1: 212. 1867); Baillon
(Hist. pi. 4: 159. 1872), Masters (in Hook, f., Fl. Br. Ind. 1: 350. 1874)
and Beccari (Malesia 3: 219. 1889) accepted the genus.

Bakhuizen van den Brink (in Bull. Jard. bot. Buitenzorg III, 6: 228.
1924) incorporated the genus in Durio.

In my opinion Cullenia represents a "good" genus by its lack of
corolla. Alston, although accepting Bakhuizen's reduction, informed me
personally, that he, too, is inclined to consider Cullenia different from
Durio.

The pollen were described as being naked and pedicellate by Gardner;
this wrong statement was corrected by Wight; the anthers are pedicellate
and one-celled.

In this paper a new Cullenia species is described, which strengthens
the position of the genus; both species are restricted to the rain forest
region of Ceylon and the Southern Indian Peninsula.

Cullenia Wight
Trees; leaves alternate, lower surface covered with scales. Inflores-

cence pseudo-umbellate on old wood. Flowers covered by scales, in bud
covered by the epicalyx, which bursts at apex and drops before anthesis.
Calyx tubular, 5-toothed. Corolla 0. Staminal tube exserted; upper part of
filaments free, bearing along it the pedicellate, one-celled, glomerulate
anthers. Ovary 5-celled; ovules 2 or more in each cell, superposed; style
longer than staminal tube; stigma small, capitellate. Fruit globose, split-
ting into 3—4 valves, densely covered by long spines. Seeds covered by
a fleshy arill.

* Issued July 1956 as Communication no 51 of the Forest Research Institute,
Bogor, Indonesia.

** D. Sc, Botanist, Forest Service of Indonesia.

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