Rivista Italiana di Paleontologia e Stratigrafia volume 102 numero 3 tavola 1 pagrne 413-476 Dicembre 1996 FLATSCHKOFELIA ANISICA GEN. ET SP. NOV. (FORAMINIFERIDA) FROM THE MIDDLE TRIASSIC (ANISIAN) OF NORTHERN DOLOMITES, ITALY ROBERTO RETTORI*. BABA SENOWBARI.DARYAN'S* & RAINER ZUHLKE*** Key-utords: Paleontologl Foraminifers, Triassic, Anisian, Alps, Nonhern Dolomites, Italy. Riassunto. Nel presente lavoro viene descritto un nuovo gener€ di foraminifero sessile a parete agglutinante: Flatscbkofelia (specie-tipo: Flatschkofelia anisica gen et sp. nov.). Questo foraminifero è stato rin- venuto nelle brecce di talus recifale di età Triassico medio (Anisico) de1la Formazione del Serla superiore (Dolomiti di Valdora, Italia). Flatscbhofelia, gen. nov., è riferito alla famiglia Placopsilinidae Rhum- b1er. Abstract. The new genus of sessile and agglutinated foramini- fer, Flaxschhofelia (type-spectes: Flatschhofelia anisica gen. et sp. nov.) from the Middle Triassic (Anisian) reef talus of the Olang Dolomites (lJpper Serla Formation, Dolomiti di Valdora, Italy) is introduced. Flaxcbkofelia, gen. nov., is referred to the family Placopsilinidae Rhumbler. lntroduction. After the end-Permian mass extinction, reefs and reefal carbonate buildups did not exist during the Late Permian to Early Triassic. First occurrence of reefs has been reported from the Early-Middle Anisian (Middle and Late Pelsonian) of the Dolomites, Italy and from Southern China (see Senowbari-Daryan er al., 1993). In the Northern part of the western Tethys additional Ani- sian reefs are known from the Carpathians and from the Karawanken Mountains. From the southern paft of the western Tethys no Anisian reefs are known. Most of Late Anisian reef carbonates in the Olang Dolomites (Southern Alps, Italy) are represented by al- lochthonous talus reef blocks of one or several cubicme- ter size. Autochthonous reef buildups are extremely rare. Both, the allochthonous reef blocks and autochtho- nous buildups occur within the Middle to Late Pelso- nian Recoaro Formation. The fossil content as well as the microfacies types of these reef carbonates are descri- bed by Bechstàdt 6c Brandner (1.970), Fois & Gaetani (198a) and by Senowbari-Daryan et al. (1993) in detail. Olangocoelia ozr Bechstadt & Brandner, a problematic fossil (sponge or alga?), is the most abundant organism in the Anisian reef carbonates, followed by different ty- pes of sphinctozoid and inozoid sponges, bryozoans, so- lenoporacean algae, corals, encrusting epibionts (poro- stromate algae, cyanophycean crusts, microproblematica, worms and foraminifers) (see for detaii Senowbari-Da- ryan et a1., 1,993). Foraminifers are relatively rare and mostly represented by Meandrospira dinarica Ko- chansky-Devidè & Pantic, Pilammina densa Pantic, En- doteba sp., Endotebanella sp., Endouiada sp., Endotria- della sp., Glomospira sp., Glomospirella sp. Duostomini- dae etc. Sessile foraminifers are also not abundant and represented by the questionable gena,s Bullopora? andby an agglutinated type, described here as Flatschhofelia ani- slca gen. et sp. nov. Systematic description Foraminiferida Eichwald, 1830 Suborder Text u I a ri i n a Delage & Hérouard, 1896 Superfamily L i t u o I a c ea de BlainviTle, 1827 Family Placopsilinidae Rhumbler, 1913 Subfamily Placopsilininae Rhumbler, 191.3 Genus Flatschkofelia gen. nov. Type-species: Flatschkofelia anisica gen. et sp. nov. Derivation of name. After the locality Flatschkofel, Olang Dolomites, Italy (Fig. 1), where the holotype was found. Diagnosis. Test simple, agglutinated and solid structured, attached on the early coiled stage, free and uncoiled on the final stage. Chambers are biserially ar- 'r Dipanimento di Scienze della Terra, Università degli Studi di Perugia, Piazza Università 4, 06100 Perugìa, Italy. 'r'r Institut fùr Palaontologie, Universitàt Erlangen-Nùrnberg, LoewenichstraBe 28, D-91054 Erlangen, Germany. ooo Geologisch-Palàontologisches Institut, Universitàt Heidelberg, Im NeuenheimerFeld234,D-69120 Heidelberg, Germany. /'"\:--' )--r-- )-^ ì-v;fir*.--'; / A N A B@en Mamolada o.-L--30* FurkeLPm Piz da Per6 Dreifrngerspitze LocalitY ,É Maurerkopf ^^Flatschkofel O12|lrn R. Rettori, B. Senoubari.Dartan €r R. Zùblke Fig. 1 - Location map of the Dolomites and Flatschkofel, where Flatscbkofelia anisica gen. et sp. nov. was found. ranged in the initial stage, bi- and uniserial in later stage. Aperture simple and terminal located in the first biserial stage, moving to the margin in the second uniserial stage. Composition of the genus. At the moment mono- typic, only represented by the type-species. Stratigraphic and geographic distribution. Middle Triassic (Anisian), Northern Dolomites, Italy. Remarks. Flatscbkofelia gen. nov. is herein inclu- ded in the family Placopsilinidae Rhumbler, 1913 deil- ned by I-oeblich & Tappan (1987, p. 80) as follows: "Test attached, early stage coiled, arcuate, or biserial, la- ter uncoiled; wall agglutinated, solid, aperture terminal, single or multiple. L. Jurassic to Flolocene" Tho f"-il-' i..l'161s5 two subfamilies: Placopsilini- nae Rhumblen I9I3 and Adhaerentiinae Loeblich Ec Tappan, L986. Flatscbkofelia gen. nov., is referred to Pla- copsilininae because having a coiled early stage even if all the genera belonging to the subfamily lacks a biserial Paft. The biserial arrangement of the chambers is a morphoiogical feature of Adhaerentiinae, but the unique genus (Adhaerentia Plummer, 1938, type-species Adhae- rentia midaayensis Plummer, 1938) included in the subfamily lacks an early coiled stage. The presence of a coiled early stage, a biserial se- cond part and a final uniserial part, which is common both in Placopsilininae and Adhaerentiinae, rogerher with aperturai features, could justify the introduction of a new subfamily for Flatschkofelia, gen. nov. Flowever, the material from the Northern Dolomites is not enough to introduce a new subfamily. At the momenr, Flatscbkofelia is referred to Pla- copsiiinidae also because the Lower Paleocene genus Adhaerentia shows, in the final srage, aperrure multiple with two to many separate openings and for strati- graphic reasons too. According to Loeblich & Tappan (1987) the strati- graphic range of Placopsilinidae spans from I-ower Juras- sic to Holocene, but we can affirm the range of the fa- mily starts from Triassic, nor only on rhe basis of the existence of the new genus Flatschkofelia, bur also becau- se of previous record of the genus Placopsilina in Trias- sic sediments (Trifonova 1967, 1992; Brónnimann & Za- nínettt. 1972\. Flatschkofelia anisica gen. et sp. nov. PL 1., Íig. 1-12 1993 Sessile agglutinated {oraminifer (gen. et sp. indet.) - Senowbari- Daryan, Ztihlke, Bechstid & Flùgel, p. 230, pl. 64, fig. 2,4,5. Derivation of name. Because rhe occurrence of the soecies in Ani- sirn limestones. Holotype. Longitudinal section illustrated in Pl. 1, fig. 1 (thin sec- tion: 89lV4,/18). Paratypes. All specirnens illustrated in Pl. 1, fig. 2-12 from different thin sections (see explanation of plate 1). Type locality. Flatschkofel, Olang Dolomites, Italy (see Fig. 1). Type level. Upper Serla Formation, Middle Triassic (Anisian, Pelso- nran). Material. Several specimens in different planes from thin sections: 89/Y7/49, 89/Vr/85, 89/V4/t8 (holotype), 89/Vr, 86/2, 89/y7/32, deposited at the Institute fùr Pa1àontologie, Universitv of Erlangen-Nùrnberg. Diagnosis. See diagnosis of the genus. Italy; Middle Triassic (Anisian, 8, Ia) 8e/Y4/1.8r 7) 89/Y1t e) PLATE 1 Fig. 1-12 - Flatscbhofelia anisica gen. et sp. nov. from Flatschkofel N, Upper Serla Formation, Olang Dolomites, Pelsonian). Fig. 1: Holotype, thin section 89/V4/1,8. Fìg. 2-72: Paratypes, thin sections: 2) 89/V7/32i 3, 4, 5, 6, 86/Vr/85r 1r, 12) 89/V1/8s. 415Pl. 1 Flatschkofelia anisica gen. et sp. no'u. .,,ts - O O.Smm R. Rettori, B. Senoubari-Daryan & R. Zilhlke Description. The test is attached, at least for the initial part, then, probably, may grow free of the at- tachment. It is irregulariy coiled in the reduced early stage, later uncoiled and rectilinear, slightly contorted above aii in the final part. The coiled early stage is fol- lowed by a short stage with 2-3 series of subangular chambers biserially arranged. The final part is uniserial and the chambers (5-7 ín the more developed specimens) are irregular in shape, increasing more rapidly in breadth than height. The aperture is simple, terminally located in the biserial stage, moving to the margin in the uniserial part, at the end of a short neck in any chamber offset toward the opposite margin in respect with the previous chamber. The chamber wall is solid, agglutinated and simple in structure. Dimensions of the test (in mm): Lensnt: u.)-1.+u Maximum diameter: 0.4 Height of the chmbers in the uniserial stage: 0.06-0.08 Breath of the chambers in the uniserial stage: 0.16-0.25 Thickness of the *all: 0.05-0.06 Stratigraphic and geographic distribution. Middle Triassic (Anisian, Pelsonian), Northern Dolomites, Italy. Association. Flaxcbkofelia anisica, gen. et sp. nov., was found within upper ramp and lagoonal facies of the prograding Upper Serla Formation. The underlaying Re- coaro Formation and the Upper Serla Formation contain allochthonous reef facies (reef talus biocks, reworked bio- clastic respectively). For details on Anisian reef orga- nisms of Olang Dolomites see Senowbary-Daryan et aI. (1993). The species is associated with rare foraminifers mainly represented by Endoteba sp., Endotebanella sp., Endotriada sp. and Endotriadella sp. in the reef facies; Meandrospira dinarica Kochansky-Devidè & Pantic and Pilammina densa Pantíc in the lagoonal facies. Acknouled.gments. The investigations were Forschungsgemeinschaft (Pro;eu Priority Program "Global and Sedimentation". supported by the Deutsche Be 641/17, Anis-RifQ in the Regionel Controls on Biogenic REFERENCES Bechstàdt T. & Brandner R. (1970) - Das Anis zwischen St. Vigil und dem Hóhensteintal (Pragser- und Olanger Do- lomiten, Sùdtirol). Festband Geol. Inst., 300-Jahr-Feier, Univ. Innsbruck, pp. 9-103. Brónnimann P. & Zaninetti L. (1,972) - Foraminifera from the basal Upper Muschelkalk at Hyères, western Basse-Pro- vence, Southern France. Rio. It. Paleont. Strat., v.78, n. l, pp. 43-44, Milano. Fois E. 8r Gaetani M. (1984) - The recovery of reef-building communities and the role of cnidarians in carbonate se- quences of the Middle Triassic (Anisian) in the Italian Dolomites. Paleontograpbica Amer., v. 54, pp. l9L2A0, Ithaca/New York. Loebiich A.R. E Tappan E. (1987) - Foraminiferal Genera and their classification. Van Nostrand Reinhold Comoanv Inc., Vol. oÎ 970 pp., New York. Senowbari-Daryan B., Zùhlke R., Bechstàd T. 6c Flùgei E. (1,993) - Anisian (Middle Triassic) Buildups of the Northern Dolomites (Italy): the recovery of reef corn- munities after the Permian,/Triassic Crisis. Facies. v. 28, pp. 181-256, Erlangen. Trifonova E. (1964 - Some new Triassic foraminifera in Bul- garla. Ann. Unio. Sofia, Geol., Geogr., Fac. 60 (1 Geol.) pp. 5-6, Sofia. Trifonova E. (1992) - Taxonomy of bulgarian Triassic Forami- nifera. I. Families Psammosphaeridae to Nodosinelli- dae. Geol. Balc., v. 22, n. 7, pp. 3-50, Sofia. Receioed June 20, 1996; accepted October 3, 1996