llir irt.r h;rlirnr di t'.rlconrologi:r c Strrtirnfìr pp. / l5-71 8 December 200,1 EQUISETALEAN STROBILUS IRAN PERS/COSTROBUS SHORT NOTE - NOTA BREVE VAEZ-JAVADT N. GEN.. A NE\l FROM THE TRIASSIC OF FATEMEH VAEZ-JAVAD] Receil..t .Seltenber /6, 2003; acel,tc.l Jub 22,20a1 (el' ao,r/. : Equiset.rlern srrobilus, Rhrcri:rn fìor.r, Shtrrsh.rk Fnr.. Alborz, NE Irîn. ,,ll,sn,lt. \ftlì preserred end :rbundenr pl;rnr ner.rirxsìls rerc lound in the tr.rs:l part oi thc Shem:hak Fornr;uion on Ozon Moun- t,rin, Alborz, NÈ Irrn. Perricosrroúris is e nerr eqtriseulcrn strobilus froùr rhe Rhrctiin (Upper Trlrss;c) srrxt. I'ctsico*robus ghd"-idelí is dcscribed .rs thc t1'pc spccies oi rhis genus- /'prriroslol,rs rp. is an inr- nrture conc ol this genur. Rarvrrto. l,teseìossìlì di pi.rnrc, rbbond.rnti e bcn conscn..rti, sono st:rti dnrcnuti nclh p.rrre brs.rle della F<,rnr.rzione Shemsh.rk dei mon!iOzon nclh carcn:r clell'Elburz. Nl: lran. Petsicostctbu òun nvo- ro srrobilo cli Fìluiserrlcs rinienuro rreglì strrri dicrì Reric.r (Triassico supcriore), h cui specie-ripo riene rlcscrirl.r con il noore di /'c,srcosoo- l,us ghaxìlcli.\iene in<'hre descrirro Pencosooúzs sp. conrc cono inr- nritufo d1 qucsto gencre. Introduction The area studied is locatcd on Ozon Mountain, 560 32' E and 37" 02' N, l5 Kn northerst of Jajarnr. Jxiarnl is iocated northwest of rhe Khorasan provincc, 175 Knr fronr thc citv of Boinurd (Fig. l).Thestudyareaisabout luuu m xDove sel tcve1. The Shemshak Formation is rvell-exposcd at Kuh- c-Ozon u,hich is rbout 800 Kni fron its tvpc locaÌiry in the Cennal Alborz-. The Shemshak Fornration has ir rhickness of 2080 m in Kuh-e-Ozon and consists mainly of sandstones, grey shiles intcrcalating with coaÌ seams rnd a fen linresrone beds. Afshar-Harb (1979) dcscribed thc Shemshak Formation in Kuh-e-Ozon, and on the ba- sis of Ammonite fossils he suggested a Middle Toarcian- Lorver Aalenian age for this unjt. Soheili (1982), prepar- ing a geological map for this area, menrioned that there are no lossils in the basal part of this unit up to 750 m. He suggested a Toarcian-Bathonian age for the upper part of this unit, based on Ammonire {ossils, Both Afshar-Harb and Soheiìi, while rccording presence of Ammonites in somc horizon of thc Shenr- shak Formation, did not report any planr megafbssils. Vrez-Javadi & Ghavidel-Syooki (2002) identified 1Z spe- cies of various orders. On the basis of D ictyoplryllum exile, Equisetites arcraceus, Podozamites latissimus and N ilssonia polynorpha the basal par t of the Shemshak Formation is dared as Rhactian. Stratigraphy The Shcrrshak Fornration in the Jajarm iÌrea con- sists mainlv of sandstone, shale, coal seams, and con- glonreratc horizons in thc lorver part and linìestones containing amnon;tes and bivalves in the uppcr part. The Shemshak Fornration disconformably overlies on the Elika Fornation and rhe upper contact of this for- rrrarion wirh thc Bash-Kalateh Forriation (Afshar-Harb 1979) is gradational. The lon'er part of the Shemshak For- marion begins with a conglomerate bed, l6 m thick, fol- lowed by medium-coarse grained sandstones alternating u'ith olive-grcy shales. The shales contain well-preserved and abundant plant megafossils sucl't as Equisetites drcn- Deprrrnenr of Earth Sciences, Shinz Unilersitr,, Shirrz, Isl,rnic Repubìic oi Iran 716 Fig.l - Location nrrp of stucliccl .rler. acet6 f Íom the Rh.rctian (Upper Tril,"ic.l (Vaez-Jar adi & Ghavidel Svooki 2002) and Persicostrobus gba,^ideli n. gen., n. sp.. The specimens described were collected ar .{0 m from the base of rhe alternations, i.e.56 nr from thc base of the unit. Systematic paleontology S,vsrenraric paleonrology based on Cleal (1993, pp. 779-791). Divis;on Pteridophyta Class Equiselopsida Takhtajan ex Nèmcjc, 1963 Order Equisetalss Trevisan, 1876 Persicostobus Vaez-Javadi n. gen. Tlrpe species: Pc'r.oi.roúar g/rar;.1r1i Vrcz-J.ìtidi n. sp. Derivative namg Persico from Persia means Iran, sfforrr' means cone. Diagnosis. Isolared, terminal strobilus, generally rectangular-cllipsoidal in shape, consìsting of whor ls of te- rragonalsporangiophores, arranged in colunns rnd rows in the nriddle parr; fine and linear leaves at the top of rhe strobilus and finc. short leaves at the basc, Age. Rhactirn. lcm Fig'2-A'Eqlìsetitesco/anrlarll(lÍlcrScb*'cirzrreri1997):R'Lquietitesbracteosas(a{rtKon'nol962)1C.fr7ais zoli l96l); D, fqu;refites arenacets (riter l'rentzen l9J3):E, E(lrisetitet merrJis,"i (îfrer Boureru l96t). Ne,i; cqtisetalun *robílus fon Trrassic of Irau 717 0 1cm Fig. I - *h, Pcrsico*robu: ghazile1i Vaez-Jrv:rdi n. sp.; c-d, Persìcastrobus sp Comparisons. Several species of cquiseralean stro- bili were introduced by Boureau (1964, p. a3l 135), Sch- werrzer et al. (1997,p. l3 5). Kelber & vln Konijnunburg- van Cittert (1998, p. 3, 4) but none oi these species has rectangular sporangiophores and upper or lower filiform leaves. In addition, the size of thjs genus is much big- ger than the previous kno*'n strobili (Fig. 2). Since the upper leaves have acute apices rnd an;rregular arrange- ment they can not be the stalks of sporangiophores. The sralks should have n'ider apiccs to attach the spo- rangrophores. Persicogtrobus ghavideli Vaez-Javadi n. sp. Holotype. Sample no. FJ-7; Text-Fig. 3a, b. Type locality. Shemshak Formation, 56" 32' E and 37" 02' N Ozon Mountain, East Alborz, NE Iran. Repository. Laboratory of Paleobotany, University of Shiraz, Iran. Sample Number FJ- 7, Derivation of name. In honor of the aurhor's su- pervisor Dr. Mohammad Ghavidel-Syooki. Description. An isolared terminal strobilus, big in size, 15.5 cm in lcngth, 4.5 cnr in width, with fìne and linear leaves at the top and short, filiform leaves, 4.5 cm in Ìength, at the base of rhe srrobilus . The body is the main parr of the cone, 5 cm ìn length and 3.5 cm in width. The body consists of several whorls includ- ing five tetragonal-rectanguÌar sporangiophores. Each of thesc sporangiophores varies in dimension (8-9 mm x l0-12 mm). The lower part of the cone is about 3 cm 718 E Vaez-Jat'adi in length. No three-dimensionallv specimens preserved were found. In loose material neither sporangia bornc on the pelrate sporangiophore nor basal scars of sporangia in an inner view of sporangiophorc heads have vet been observed, The strobilus atrached to a stem which con, tains 5-6 wide ridges. Comparisons. This new species diffcrs from previ- ous strobili of this family, and even of this order, by irs much bigger size and its tetragonal-rectangular shapc of the sporangiophore heads. For example, Equisetites are- naceus described by Kelber and van Konijnenburg-van Cirtert (1998) has smaller strobili (21-35 mm in length and 17-22 mm in maximunr width) and pentagonal-hexa- gonal sporangiophores. Persicostrobus sp. ftg, rc! o Description. Strobilus oval in shape,44 nrm in length and 35 nrnr in width, consistine of ninc whorls including five to six roundcd tetragonal sporangiophores in each ror.'- It is a ternrinal srrobilus. Discussion. This specimen secms to be an inrma- ture cone of Persicostrobus gbatlleÍ because of its rernri- nal position and the shape of its sporangiophores. Achn**ledgcneus.Thc.rurhor is mosr gratelul to Dr. Moham- nrrd Srdegh l;rkhr. Dr, Mohenrn;rd ch.rlidel-Slooki rnd Dr. Vrrrin Kirchner for rhcir uscful comnri.'nrs. Prol H. r'rrn Cirtcrt-Konjene'n- burg rerised an earlier version oi this paper REFERENCES Afshar-Harb A. 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Biostratigr 23:35-76 (in Russian). Schs'citzcr H. J., r'an Konijncnburg-r.an Cirrert J. H. A. & r'an der Burgh J. ( 1997) - The Rhacto - Jurassic flora of lran and Afghanisran. 10. Bryophyra, Lycophyra, Sphenophy- ta, Pterophvte - Euspor:rngiate rnd Protosporinqiàtae. Pa laeontographica, B, 2.13: I 03- I 92, Srurrgxrr. Sobcili M. (1982) - Explanatiorr of geological map of Kork- hud mounrlins. Geol. Sun'. Iran, Tehr.l-n, ll0 p. (ln Persian). VaezJavadi F 6e Ghavidel Svooki M. (2002) - PIant negafossil renrains from Shenrshak l:ormation ofJ,rjarm erea, NE Alborz, Iran. Palaeobotanist, 51 : 57-72, Lucknos'.