01_goss A checklist of Navicula 1 A Checklist of Navicula (Class Bacillariophyceae) of the Philippines Milagrosa R. Martinez-Goss Professor, Institute of Biological Sciences and Curator, Museum of Natural History University of the Philippines Los Baños College, Laguna 4031, Philippines ABSTRACT A unified account of all known diatoms in the genus Navicula, class Bacillariophyceae, of the Philippines from 1886 to 1987 is presented. A total of 226 taxa are listed. Included herewith are the references that originally described or cited the taxon and the ecological/habitat records for each taxon. As much as possible, a list of exsciccata materials is given together with their repository places. Keywords: Navicula, Bacillariophyceae, Philippines, diatoms INTRODUCTION This compedium presents a unified account of all known diatoms under the genus Navicula from 1886 t o 1 9 8 7 o r w i t h i n a s p a n o f o v e r 1 0 0 y e a r s . Referencing was based mainly on available literature that deals in whole or in part on Philippine materials. In some cases Philippine exsciccata materials (diatom slides) that were found stored in the different diatom herbaria in the Western World were also noted. A wealth of these specimens were discovered at the Diatom Section of the British Museum of Natural History (BMNH) in London, England, which include the collections of Adam, Bess, Comber, etc. The collections of Mann from the USS Albatross were also observed at the Smithsonian Institution of the U.S. Museum of Natural History in Washington, D.C., U.S.A. I have not had a chance to see the holotypes of Hustedt and Skvortzow’s collections. But according to Dr. Charles W. Reimer of the Academy of Natural Sciences of Philadelphia (ANSP), there are some collections of Skvortzow that are in Hustedt’s collection, and the latter are stored in the herbarium of the Alfred Wegener Institute for Polar and Marine Research in Bremerhaven, Germany. The list includes 21 that are reported as new taxa (Table 1) and four species found only in Mexico (Campeche Bay) and in the marine waters of the Philippines and nowhere else according to M a n n ( 1 9 2 5 ) , i . e . , N . a p p ro x i m a t a v a r. substauroneiformis Grun., N. carinefera Grun., N. h e n n e d y i v a r. c a l i f o r n i c a ( G r e v. ) C l . a n d N . margarita Schmidt (?). There are 226 taxa recorded here that are recognized by Patrick and Reimer (1966), Round et al., (1990), and VanLandingham (1975). Unidentified specific epithets are recorded but not enumerated. The scope and use of the list is as in the previous paper (Martinez-Goss, 1995). Science Diliman (January-June 2001) 13:1, 1-32 2 Martinez-Goss Navicula biggemata N. caeca N. clavata f. corpulenta (Mann) Hust N. delecta Mann N. elegantoides Hust. N. funiculata Mann N. glabrissima Mann N. halophila f. tenuirostris Hust. N. hustedtii f. philippina Skv. N. imitans Mann N. insignita Hust. N. luzonensis Hust. N. mendica Mann N. mucicoloides Hust. N. philippina Skv. N. philippinarum Mann N. pulverulenta (pulvulenta) Mann N. pugio Mann N. retrostauros Mann* N. spiculifera Mann* N. transluscens Mann Marine Marine Marine Marine Laguna de Bay; Camarines Sur, Buhi Lake Marine Marine Rizal, Manila Quezon City, Balara Marine Laguna de Bay, Rizal, Lyon (ponds) Camarines Sur, Buhi Lake (rare) Marine Rizal, Lyon (ponds) Quezon City, Balara Marine Marine Marine Marine Marine Marine Mann, 1925 Mann, 1925 Hustedt, 1964 Mann, 1925 Hustedt, 1942a Mann, 1925 Mann, 1925 Hustedt, 1942a Skvortzow, 1937 Mann, 1925 Hustedt, 1942a Hustedt, 1942a Mann, 1925 Hustedt, 1942a Skvortzow, 1937 Mann, 1925 Mann, 1925 Mann, 1925 Mann, 1925 Mann, 1925 Marine Table 1. New taxa of Navicula reported from the Philippines *Not recognized by VanLandingham THE LIST NAVICULA Bory, p. 562, 1824; Patrick & Reimer, p. 439, 1966. abrupta (Greg.) Donkin Donkin, p. 13, 2/6, 1870; Mann, p. 94, 1925. Bottom marine samples taken near the islands by the United States Steamer (USS) Albatross, Mann (1.c.). abruptoides Hust. Hustedt, p. 515, f. 1557, 1964; Podzorski & Hakansson, p. 79, 31/8, 1987. LUZON: Palawan, Pto. Princesa (White Beach mangrove, root scrapings from Rhizophora), Podzorski & Hakansson (l.c.). americana Ehr. Ehrenberg, p. 129 (417), 1841 (1843); Hustedt, p. 280, f. 464, 1930; _______, p. 64, 1942a. LUZON: Laguna, Los Baños (ponds, Laguna de Bay); Camarines Sur, Buhi (river), Hustedt (1942a). MINDANAO: Lanao (Uyaan Lake), Hustedt (1942a). anglica Ralfs Ralfs in Pritchard, p. 900, 1861; Hustedt, p. 303, f. 530-531, 1930; _______, p. 73, 1942a. MINDANAO: Lanao (ponds), Hustedt (1942a). angusta Grun. Grunow, 10:528, 5/19, 1860. = Navicula cari var. angusta Grun. Grunow in Van Heurck, 7/17, 1885; Hustedt, p. 72, 1942a; Patrick & Reimer, p. 514-515, 49/5, 1966. LUZON: Laguna, Los Baños (ponds), Hustedt, 1942a. Taxon Source Reference A checklist of Navicula 3 VISAYAS: Leyte, Danao Lake, Hustedt, 1942a. approximata Grev. Greville, p. 28, 4/4, 1859; Mann, p. 94, 1925. = N. lyra var. approximata (Grev.) Cl. Cleve, p. 4 (1/1) 1878. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.); British Museum (BMNH), Adam’s slide #45.1/2; TS 60/64. _______ f. morei (moreii) (O’ Meara) Hust. Hustedt p. 416, f. 1491, 1964. = Navicula hennedyi var. niceaensis H. Perag. Peragallo, p. 47, 5/39 e.p., 1888. BMNH, Comber slide #31898. VISAYAS: Cebu. _______ var. substauroneiformis (Grun.) Cl. Grunow in Schmidt, et al., 2/20, 21, 1874; Mann, p. 94, 1925. Bottom marine samples, Mann (l.c.). _______ var. ? BMNH, Adam’s-Bess # TS497 3/2. arvensis Hust. Hustedt, p. 249, 20/19, 20, 1937b; _______, p. 60, 1942a; Patrick & Reimer, p. 483, 46/1-2, 1966. LUZON: Batangas, Taal (Crater Lake), Hustedt (1942a). bacillum Ehr. Ehrenberg, p. 130, 1838, (1839); _________, p. 418 (130), 4/5, f. 8, 1841 (1843); Hustedt, p. 64, 1942a. LUZON: Manila (Reservoir); Laguna, Los Baños; Camarines Sur, Buhi (lake), Hustedt (1942a). baileyana Grun. Grunow in Schmidt, et al, p. 88, 1/31, 1874; Castracane, p. 812, 1886. VISAYAS: Cebu, Castracane (l.c.) = Navicula polysticta Grev. Greville, p. 247,5/2, 1859. VISAYAS: Cebu, BMNH, Comber # 31892. barbitos Schmidt Schmidt, et al., 129/5, 1874; Mann, p. 95, 1925. Bottom marine samples taken around the islands by the USS Albatross, Mann (l.c.). biformis (Grun.) Mann Mann, p. 95, 20/6-7, 1925. Bottom marine samples taken around the islands by the USS Albatross, Mann (1.c.). = Mastoneis biformis (Grun.) Cl. Cleve, p. 194, 1894. LUZON: Manila, BMNH, Comber # 31904. VISAYAS: Cebu, BMNH, Comber # 31895. bigemmata Mann Mann, pp. 95-96, 21/1, 1925. Bottom marine samples taken around the islands by the USS Albatross. Type-Cat. No. 43639 deposited in the United States National Museum of Natural History (USNM), Mann (1.c.). 4 Martinez-Goss bipunctata O’ Meara O’ Meara, p. 286, 13/5, 1872; De Toni, pp. 78-79, 1891. MINDANAO: Brackishwaters coming from the volcanic mountain of the island in Cagayan, Sulu Archipelago (Chimmo), O’ Meara (l.c.); De Toni, (l.c.). borneoensis Hust. Hustedt, p. 735, f. 1712, 1966; Podzorski & Hakansson, p. 84, 33/7, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae encrusted on cove, marine), Podzorski & Hakansson (1.c.). brasiliensis Grun.* Grunow, p. 152, 14/10, 1863; Schmidt, Atlas, 6/19-21, 23/25, 31/33, 1874; Mann, p. 97, 1925. Bottom marine samples taken around the islands by the USS Albatross, Mann (1.c.). BMNH, Adam’s # F19.29, 40 2/18.883; TS 262 1/10-12, 6/7, TS 60/60. VISAYAS: Cebu, BMNH, Comber’s #31898, 31895, 31894. brasiliensis var. _______Cstr. Castracane, p. 812, 1886. BMNH, Adam’s-Bess #706, TS 60/66. VISAYAS: Cebu, Castracane (l. c.) brekkaensis Peters. Petersen, p. 389, f. 16, 1928; Hustedt, p. 58, 1942a. LUZON: Rizal, Pancipit River, Hustedt (l.c.). bruchii Grun. Grunow in Cleve, p. 13, 1928; Cleve, p. 36, 1895; De Toni, pp. 177-178, 1891. LUZON: Manila (marine), (Deby collection, BMNH). bryophila Peters. Petersen, p. 388, f. 13, 1928; Hustedt, p. 60, 1942a. MINDANAO: Lanao Lakes of Dagiangan and Uyaan, Hustedt (1.c.). caeca Mann Mann, p. 97, 21/3, 1925. Bottom marine samples taken near the islands by the USS Albatross. Type-Cat. No. 43641, USNM, Mann (1.c.). capitata Ehr.Ehrenberg, p. 185, 13/20, 1838; Van Heurck, p. 187, 25/719, 1896; Patrick & Reimer, pp. 536-537, 52/1-2, 1966; Podzorski & Hakansson, p. 81, 32/10, 1987. LUZON: Palawan, Taytay, Langen (Malapakan Cove, algae encrusted on the cove, marine), Podzorski & Hakansson (1.c.). cari Ehr. Ehrenberg, p. 83, 1836; _________, p. 174, 1838; Hustedt, p. 299, fig. 512, 1930; _______, p. 72, 1942a. LUZON: Rizal, Malabon (Dagatan swamp), Hustedt (1942a). MINDANAO: Uyaan Lake, Hustedt (1942a). carinifera Grun. Grunow in Schmidt, et al., 2/1, 1874; Mann, p. 98, 1925; Hustedt, p. 754, f. 1730, 1966. *Not recognized by VanLandingham, 1975. A checklist of Navicula 5 Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). cincta (Ehr.) Ralfs Van Heurck, p. 178, 3/105a, 1896; Schmidt, et al., 299/26-30, 1874; Hustedt, p. 298, f. 510, 1930; Pantastico, p. 194, 17/4, 1977; Patrick & Reimer, p. 516, 49/8, 1966. LUZON: Laguna, Los Baños, Mayondon (on shoreline of Laguna de Bay), Pantastico (1.c.). circumsecta (Grun.) Grun. Grunow in Schmidt, et al., 3/26-27, 1874; Cleve et Grunow, p. 42, 1880; Mann, p. 98, 1925; Hustedt, p. 409, f. 1487a-c, 1964. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). = Navicula polysticta var. circumsecta (Grun.) Grun. Grunow in Schmidt, et al., p. 89, 1/36, 42, 1874. BMNH, Adam’s # GC 2089. circumtexta Meister Meister in Schmidt, et al., 394/33-35, 1874; Podzorski & Hakansson, pp. 81-82, 32/4, 1987. LUZON: Palawan, Taytay, Langen (Malapakan Cove, scrapings from sand at low tide, marine), Podzorski & Hakansson (1.c.). citrus Krasske Krasske, p. 199, f. 14, 1923; Hustedt, p. 303, f. 529, 1930; _______, p. 66, 1942a. LUZON: Laguna de Bay, Hustedt (1942a). clavata Greg. Gregory, p. 46, 5/17, 1856. BMNH, Adam’s # F83 1/2; 4/3; 4. 120, 3/2.5. LUZON: Manila (on shell), BMNH, Comber # 30925, 31903. VISAYAS: Cebu, BMNH, Comber # 31900, 31898. = Navicula clavata var. caribaea Cl. Cleve, p. 61, 1895. VISAYAS: Cebu, BMNH, Comber # 31895. = Navicula caribaea Cl. Cleve in Schmidt, et al., 2/17, 70/48, 1874; Mann, pp. 99-100, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). = Navicula wrightii O’ Meara LUZON: Manila, BMNH, Comber # B. 31903. _______ f. corpulenta (Mann) Hust. Hustedt, p. 445, f. 1510-1511, 1964. = N. corpulenta Mann Mann, p. 99, 22/4, 1925. Bottom marine samples taken around the islands by the USS Albatross. Type-Cat. No. 43642, USNM, Mann (1.c.). _______ f. elliptica (Schmidt)Hust. Schmidt, et al., 3/13, 1874; Hustedt, p. 449, 1964. LUZON: Manila, BMNH, Comber # 31903. VISAYAS: Cebu, BMNH, Comber # 31898. _______ var. indica (Grev.) Cl. Cleve, p. 62, 1895; Schmidt, et al., 204/12, 1874. BMNH, Adam’s # GC 1865.6.1976; Bess #920; TS #497.2/3.4.522.2/4, F885. 6 Martinez-Goss = N. indica Grev. (non Ehr.) Greville, p. 95, 9/13, 1862; Cleve, p. 62, 1895; Mann, p. 104, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). BMNH, Adam’s # J 4201, M 68/4. LUZON: Manila (marine), BMNH, Rae’s collection. VISAYAS: Cebu, BMNH, Comber # 31898, Rae’s collection. _______ var. producta ? sensu Adam BMNH, Adam’s # F886. LUZON: Manila, BMNH, Comber # 31904. _______ var. ? BMNH, Adam’s # TS60/61.657. clipeiformis König König, p. 51, 59, 2/3, 3/6, 1959; Hustedt, p. 552, f. 1589, 1964; Podzorski & Hakansson, p. 76, 31/9, 1987. LUZON: Palawan, Taytay, Langen (Malapakan Cove, surface scrapings from sand at low tide, marine), Podzorski & Hakansson (1.c.). cluthensis Greg. Gregory, p. 478, 9/2, 1857; Hustedt, p. 651, f. 1653a-d, 1966. = N. erythraea Grun., p. 539, 5/17 (3/17), 1860; Mann, p. 101, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). confervacea (Ktz.) Grun. Grunow in Van Heurck, 14/36, 1880; Woltereck, p. 37, 1941; Hustedt, p. 63, 1942a; Prowse, p. 41, 12/n-o, 1962; Hustedt, p. 205, f. 1324a-d, 1962; Podzorski & Hakansson, pp. 77-78, 1987. LUZON: Benguet, Banaue (ponds), Woltereck, (1942); Rizal, Malabon, Dagatan (swamp), Marulay Water Reservoir, Lyon ponds; Laguna, Los Baños, Laguna de Bay; Batangas, Taal (Taal Lake, Yambo- Crater Lake, Crater Lake); Camarines Sur, Buhi (lake, river); Oriental Mindoro, Naujan (river), Calapan Lake, Hustedt (1942a); Palawan, Roxas (Caruray River, alt., c. 50 m, from rock surface), Taytay, Lake Manguao, (algae growing on floating grass roots, alt. c. 45 m), Podzorski & Hakansson, (1987). VISAYAS: Leyte, Danao Lake, Hustedt (1942a). MINDANAO: Lanao Lake, Hustedt (1942). = N. confervacea f. nipponica Skv. Skvortzow, p. 34, 2/3, 4/23, 1936; Hustedt, p. 61, f. 23, 1942a LUZON: Camarines Sur, Buhi Lake, Hustedt (l.c.). consors Sch. Schmidt, et al., 48/24-27, 1874; Mann, p. 98, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). contenta Grun. Grunow in Van Heurck p. 109, 1885; Van Heurck, p. 230, 5/239, 1896; Hustedt, p. 277, f. 458a, 1930; Skvortzow, p. 291, 1/40, 1937; Hustedt, p. 58, 1942a. A checklist of Navicula 7 LUZON: Batangas, Taal (Crater Lake), Hustedt (1942a). ______ var. biceps (Arn. Ms. Grun. in V.H.) Cleve Van Huerck, p. 230, 5/240, 1896. = N. contenta f. biceps (Arn. Grun in V.H.) Hust. Hustedt, p. 277, f. 458c, 1930; _______, p. 58, 1942a. LUZON: Pancipit River, Hustedt (1942a). _______ f. parallela (Peters.) Hust. Hustedt, p. 277, f. 458b, 1930; _______, p. 58, 1942a. LUZON: Camarines Sur, Buhi (lake), Hustedt (1942a). cryptocephala Ktz. Kützing, p. 95, 3/20, 26, 1844; Donkin, p. 37, 5/14, 1871; Van Heurck, p. 180, 3/122, 1896; Hustedt, p. 295, f. 496, 1930; Skvortzow, p. 291, 2/3, 1937; Hustedt, p. 67, 1942a; Podzorski & Hakansson, p. 87, 1987. LUZON: Rizal, Manila (Water Reservoir II), ponds in the Bureau of Science grounds; Laguna, Laguna de Bay; Camarines Sur, Buhi Lake; Oriental Mindoro, Naujan Lake, Hustedt (1942a); Rizal, Quezon City (Balara filters, Skvortzow, (l.c.); Palawan, Brooke’s Point (scrapings from a limestone rock in Tumarabong River, alt. c. 10 m), Podzorski & Hakansson (1987). cf. cryptolyra Brockmann Brockmann, p. 19, 3/22, 23, 1950; Hustedt, p. 534, f. 1570, 1964; Podzorski & Hakansson, p. 79, 32/5, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae encrusted on the cove, marine), Podzorski & Hakansson (1.c.). cuspidata (Ktz.) Ktz. Kützing, p. 94, 3/24, 37, 1844; W. Smith, p. 47, 16/131, 1853; Van Heurck, p. 214, 4/190, 1896; Mann, p. 99, 1925; Hustedt, p. 268, f. 433, 1930. Freshwater sample as detritus from the bottom marine samples taken by the USS Albatross, Mann (1.c.). _______ var. ambigua (Ehr.) Cl. Cleve, p. 110, 1894; Hustedt, p. 51, 1942a. LUZON: Rizal (ponds in the Bureau of Science grounds); Laguna, Los Baños, Laguna de Bay (lake and ponds); Camarines Sur, Buhi Lake, Hustedt (1942a); Palawan, Roxas, Pagdanan Range, Ibangley Brookside Hill (streamside pool, alt. c. 40 m), Podzorski & Hakansson (1.c.). MINDANAO: Cagayan de Oro, Singnan Lake; Lanao (lake), Hustedt (1.c.). cyclops Mann Mann, pp. 99-100, 21/5, 1925. Bottom marine samples taken near the islands by the USS Albatross; frequently observed. Type- Cat. No. 43643, USNM, Mann (1.c.). delecta Mann Mann, p. 100, 24/5-7, 1925. Bottom marine samples taken around the islands by the USS Albatross; frequently observed. Type-Cat. No. 43644, USNM, Mann (1.c.). BMNH, Adam’s # M58/31. dicephala Ehr. Ehrenberg, p. 45, 1837; _________, p. 185, 1838; Van Heurck, p. 188, 3/138, 1896; Hustedt, p. 302, f. 526, 1930; _______, p. 73, 1942a. 8 Martinez-Goss LUZON: Laguna, Los Baños (freshwater ponds), Hustedt (1942a). _______ var. undulata Östr. Östrup, p. 25, 3/33, 1918 (1920); Hustedt, p. 73, 1942a. LUZON: Camarines Sur, Buhi Lake, Hustedt (1942a). diffluens Schmidt = Navicula diffusa Schmidt Schmidt, et al., 2/15, 2/28, 1874; Mann, p. 101, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (l.c.). directa (W. Sm.) Ralfs Ralfs in Pritchard, p. 906, 1861. LUZON: Manila, BMNH, Comber # 31903, 31904. distans (W. Sm.) Ralfs Ralfs in Pritchard, p. 907, 1861; Schmidt, et al., 46/11-14, 1874. LUZON: Manila (on shell), BMNH, Comber # 30925, 30926. durandii (durrandii) Kitt. Kitton in Schmidt, et al., 129/1, 1874; Castracane, p. 812, 1886; Mann, p. 101, 21/6, 1925; Hustedt, p. 485, f. 1540, 1964. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.); BMNH, Adam’s # TS522.1/1. VanLandingham (1975) does not think the specimen is in accordance to Kitton. VISAYAS: Cebu, Castracane (l. c.) _______ var. rhomboidea (Cstr.) Schmidt Schmidt, et al., 129/2, 1874. = Navicula bullata Norman var. rhomboides Cstr. Castracane, p. 30, 30/8, 1886. VISAYAS: Cebu, Castracane (l.c.). _______ var ? BMNH, Adam’s # M 65/5. elegantoides Hust. Hustedt, p. 76, f. 142, 1942a; Woltereck, p. 52, 1941; Prowse, p. 42, 8/d, 1962. LUZON: Benguet, Banaue (ponds), Woltereck (l.c.); Rizal, Manila (Lyon ponds, ponds in Bureau of Science grounds); Laguna, Los Baños, Laguna de Bay; Buhi Lake, Hustedt (1.c.). exigua (Greg.) Grun. Grunow in Van Heurck, 8/32, 1880; Hustedt, p. 303, f. 538, 1930; _______, p. 73, 1942a. LUZON: Camarines Sur, Buhi Lake, Hustedt (1942a). MINDANAO: Uyaan Lake, Hustedt (1942a). forcipata Grev. Greville, p. 83, 6/10-11, 1859; Schmidt, et al., 70/17, 1874; Cleve, p. 65, 1895; Van Heurck, p. 203, 4/163, 1896; Peragallo & Peragallo, p. 130, 21/28, 1897-1908. BMNH, Adam’s # J 413. VISAYAS: Cebu, BMNH, Comber # 31898. _______ var. densestriata Schmidt Schmidt, et al., 70/14-16, 1874; Mann, p. 102, 1925. A checklist of Navicula 9 Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). _______ cf. forcipata var. elongata Perag & Perag. Peragallo & Peragallo, p. 131, 21/24, 1897-1908; Podzorski & Hakansson, p. 79, 33/1, 1987. LUZON: Palawan, Pto. Princesa, Pto. Princesa Bay (Table Head, on sandy bottom at 15 m depth), Podzorski & Hakansson (1.c.). _______ var. versicolor (Grun.) Grun. Grunow in Cleve et Möller, no. 208-210, 1879. VISAYAS: Cebu, BMNH, Comber # 31894, 31902. _______ var ? BMNH, Adam’s # G-211. funiculata Mann Mann, p. 102, 22/1-2, 1925. Bottom marine samples taken around the islands by the USS Albatross. Type-Cat. No. 43645, USNM, Mann (1.c.). BMNH, Adam’s # TS 60/2.76. genifera Schmidt Schmidt, et al., 2/6, 1874; Hustedt, p. 435, f. 1505, 1964. BMNH, Adam’s # W 935. glabrissima Mann Mann, p. 103, 22/3, 1925. MINDANAO: Jolo, Sulu, Sulu (bottom marine sample, rare). Type-Cat. No. 43646, USNM, Mann (1.c.). gracilis Ehr. Ehrenberg, p. 64, 69, 1830 (1832); _________, p. 155, 1854; Van Heurck, p. 179, 3/109, 1896; Hustedt, p. 299, f. 514, 1930. LUZON: Rizal, Manila (Tabaco River), sample taken by the ship Princess Louise, Ehrenberg (1854). granulata Bailey Bailey, p. 10, f. 16, 1854 (1853); Cleve, p. 48, 1895; Schmidt, et al., 244/7, 1874; Hustedt, p. 702, f. 1696, 1966; Podzorski & Hakansson, p. 84, 33/14, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (surface scrapings from the sand at low tide, by the cove), Podzorski & Hakansson (1.c.). cf. gregaria Donk. Donkin, p. 10, 1/10, 1861; Van Heurck, p. 181, 3/125, 1896; Hustedt, p. 269, f. 437, 1930; Podzorski & Hakansson, p. 78, 1987. =Navicula granulata cf. gregaria Donk. Donkin, p. 10, 1/10, 1861; Van Heurck, p. 181, 3/125, 1896; Hustedt, p. 269, f. 437, 1930; Podzorski & Hakansson, p. 78, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae encrusted on the cove, marine), Podzorski & Hakansson (1.c.). aff. grevillei (Ag.) Heiberg Heiberg, p. 83, 1863; Van Heurck, p. 232, 5/243, 1896; Prowse, p. 43, 12/6, 1962; Podzorski & Hakansson, p. 77, 30/9, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae encrusted on the cove, marine), Podzorski & Hakansson (1987). grimmii (grimmei) Krasske Krasske, p. 45, 1/14, 1925; Woltereck, p. 52, 1941; Hustedt, p. 274, f. 448, 1930; Hustedt, p. 55, 1942a. LUZON: Benguet, Banaue (ponds), Woltereck (l.c.); Rizal, Lyon (ponds); Camarines Sur, Buhi (river), Hustedt (1942a). MINDANAO: Lanao (lake), Hustedt (1942a). halophila (Grun.) Cl. Cleve, p. 109, 1894; Hustedt, p. 268, f. 436, 1930; _______, p. 51, 1942a; Woltereck, p. 52, 1941. LUZON: Mt. Province, Banaue (ponds) Woltereck (l.c.); Rizal, Lyon (ponds); Laguna, Los Baños (pond), Hustedt (1942a). MINDANAO: Lanao, Uyaan (lakes), Hustedt (1942a). _______ f. tenuirostris Hust. Hustedt, pp. 51-52, f. 76, 1942a. LUZON: Rizal, Manila (ponds in the Bureau of Science grounds, rare), Hustedt (1.c.). hamulifera Grun. Grunow in Cleve and Grunow, p. 44, 1880; Cleve, vol. 1, p. 154, 3/16-19, 1894; Peragallo & Peragallo, p. 65, 8/16, 1897-1908; Mann, p. 104, 1925; Hustedt, p. 312, f. 1430, 1962. Bottom sediments taken near the islands by the USS Albatross, Mann (1.c.). hassiaca Krasske Krasske, p. 47, 2/26, 1925; Hustedt, p. 279, f. 462, 1930; _______, p. 216, f. 1332, 1962; Podzorski & Hakansson, p. 81, 32/2, 1987. LUZON: Palawan, Taytay, Lake Manguao (Danao), Podzorski & Hakansson (l.c.). hennedyi (henedii) W. Sm. W. Smith, p. 93, 1856; Gregory, p. 40, 5/3, 1856; Van Heurck, p. 93, 9/14, 1880-1885; Cleve, p. 57, 1895; Mann, p. 104, 1925. Bottom sediments taken near the islands by the USS Albatross, Mann (1.c.); BMNH, Adam’s # E 530, M 58/14, F119.1/14.3/3. LUZON: Manila, BMNH, Comber #30925, 31903, 31904, 30926 (on shell). VISAYAS: Cebu, BMNH, Comber #31900. =Navicula hennedyi var. niceaensis (niceaeensis) H. Perag. H. Peragallo, p. 55, 5/39,1888. BMNH, Adam’s # M 58/22-3. ______ f. bacillifera (Pantoc.) Cl. Cleve, p. 58, 1895. = Navicula rugosa Janisch Janisch, 15/10, 11, 1888; Cleve, p. 59, 1895. BMNH, Adam’s # E 530. _______ var. californica (Grev.) Cl. Greville, p. 29, 4/5, 1859; Cleve, p. 58, 1895; Mann, p. 97, 1925. = Navicula californica Grev. Greville, p. 29, 4/5, 1859; Mann, p. 97, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (l.c.); BMNH, Adam’s # M58/20-1; 24.30. =Navicula californica var. campechiana Grun. Grunow in Schmidt, et al., 3/19, 1874; Mann, p. 97, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). 10 _______ var. furcata Perag. & Perag. Peragallo & Peragallo, p. 141, 24/16, 17, 1897-1908. BMNH, Adam’s # F 18 2/9.10. _______ f. granulata (Grun.) Hust. Grunow in Schmidt, et al., 3/3, 1874; Hustedt, p. 458, f. 1519, 1964. =Navicula hennedyi var. circumsecta Cl. Cleve p. 58, 1895 (non Grunow, 1874 in Schmidt, et al., 1874). It means Cleve thought the specimen is not the same as in Grunow in Schmidt, et al., but VanLandingham says they are the same. BMNH, Adam’s # TS 519/3. _______ var. neopolitana Cl. Cleve, p. 58, 1895; Podzorski & Hakansson, p. 80, 31/12, 1987. LUZON: Palawan, Taytay (beach on washed- ashore seaweed), Podzorski & Hakansson (1.c.). _______ f. ? BMNH, Adam’s # F 120.3/7. hennedyi Hust.* Podzorski & Hakansson, p. 79, 30/1, 1987. LUZON: Palawan, Pto. Princesa, Table Head, South of Pto. Princesa Bay (from surface at 15 m depth), Podzorski & Hakansson (1.c.). howeana Hagelstein Hagelstein, p. 385, 7/51, 1939; Podzorski & Hakansson, pp. 76-77, 30/6, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (surface scrapings from the sand at low tide in Malapakan Cove), Podzorski & Hakansson (1.c.). hustedtii Krasske Krasske, 3:198, f. 3, 1923; Patrick & Reimer, p. 527, 50/8, 1966. = Navicula hustedtii f. philippina Skv. Skvortzow, p. 290, 1/44, 1937a. LUZON: Rizal, Quezon City, Balara (water filter no. 8), Skvortzow (l.c.). imitans Mann Mann, p. 104, 22/5, 6, 1925. Bottom marine samples taken near the islands by the USS Albatross. Type-Cat. No. 43643, USNM, Mann (1.c.). indicatrix VanLan. VanLandingham, nom. nov. = N. alpha Chol. Cholnoky, p. 173, fig. 57, 1955 (non Cl. 1893). LUZON: Manila, BMNH, Comber #31903. inexacta Mann Mann, p. 105, 1925. = N. (Alloioneis) gruendleri (Cl. et Grun.) Cl. Mann, p. 104, 105, 22/7, 1925. Bottom marine samples taken near the islands by USS Albatross. No type designated, Mann (1.c.). inhalata Schmidt Schmidt, et al., 2/30, 1874; Cleve, p. 57, 1895; Mann, p. 107, 23/2, 1925; Hustedt, p. 390, f. 1475, 1964. Bottom marine samples taken near the islands by USS Albatross, Mann (1.c.); marine, Cleve (l.c.). insignita Hust. Hustedt, p. 73, f. 126, 139-141, 1942a; Woltereck, p. 52, 1941; Patrick & Reimer, p. 447, 40/3, 1966. 1 1 12 Martinez-Goss LUZON: Mt. Province, Bontoc, (Benguet, Banaue, ponds); Lyon (ponds), Woltereck (1.c.), Hustedt (1.c.); Rizal; Laguna, Laguna de Bay, Hustedt (1.c.). insociabilis Krasske Krasske, p. 114, 3/17, 1932; Schmidt, et al., 40/103-105, 1874; Hustedt, p. 63, 1942a; _______, p. 181, f. 1315a-h, 1962. LUZON: Rizal, Pancipit River, Hustedt (1942a). iota (jota) Cl. Cleve, p. 134, 5/22, 1894. VISAYAS: Cebu, BMNH, Comber # 31898. irrorata var. substauroneiformis (Grun.) Cl. Cleve, p. 56, 1895 =Navicula approximata var substauroneiformis (Grun.) Cl. Grunow in Schmidt, et al., 2/20, 21, 1874; Mann, p. 94, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (l.c.). jejuna Schmidt Schmidt, (1876) in Schmidt, et al., 46/76, 1874; Mann, p. 107, 1925; Skvortzow, p. 280, 4/7, 1932. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). laevissima Ktz. Kützing, p. 96, 21/14, 1844; Patrick & Reimer, p. 497, 47/13, 1966. BMNH, Adam’s # GC 1994. = Navicula bacilliformis Grun. Grunow in Cleve & Grunow, p. 44, 2/51, 1880; Hustedt, p. 55, 1942a. LUZON: Camarines Sur, Lake Buhi, Hustedt (l.c.). lagerheimii Cl. Cleve, p. 131, 1894; _____, p. 101, 7/1, 1894; Woltereck, 41:37-176,1941; Hustedt, in A. S. Atlas, T. 370, f. 19-21, 1874; Hustedt, p. 54, 1942a. LUZON: Mt. Province, Bontoc (Banaue, ponds), Woltereck (l.c.); Laguna de Bay, Los Baños (ponds), Pancipit River; Camarines Sur, Lake Buhi, river, Hustedt (1942a). MINDANAO: Lanao, Lanao Lake, Uyaan Lake, Hustedt (1942a). _______ f. intermedia Hust. Hustedt, in A. S. Atlas, T. 370, f. 22, 1874; LUZON: Oriental Mindoro, Calapan Lake, Hustedt (l.c.). lanceolata (Ag.) Ktz. Kützing, p. 94, 28/38, 30/48, 1844; Van Heurck, p. 186, 3/139, 1896; Hustedt, p. 305, f. 540, 1930; _______, p. 72, 1942a. MINDANAO: Lanao, Uyaan Lake, Hustedt (1942a). latissima Greg. Gregory, p. 40, 5/4, 1856. LUZON: Manila, BMNH, Comber #31904. longa (Greg.) Ralfs Ralfs in Pritchard, p. 906, 1861; Donkin, p. 55, 8/3a-b, 1872; Van Heurck, p. 185, 25/716, 1896; Mann, p. 108, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). A checklist of Navicula 13 lorenzii (Grun.) Hust. Hustedt, p. 29, f. 1188, 1961; Podzorski & Hakansson, p. 78, 3/1, 1a, 2, 2a, 1987. = Stichodesmis australis Grev. Greville, p. 535, 13/1-4, 1863. Bottom marine samples taken near the islands by the USS Albatross, Mann (l.c.). LUZON: Palawan, Pto. Princesa, Table Head (from sand surface at 15 m deep of Pto. Princesa Bay), Podzorski & Hakansson (1.c.). = Navicula scopulorum sensu Cl. Cleve, p. 151, e.p., 1894. VISAYAS: Cebu, BMNH, Comber # 31892. lunula Cl. Cleve, 5/5,1894; ______, p. 51, 1895. BMNH, Adam’s # G 211. luzonensis Hust. Hustedt, p. 59, f. 106, 1942a. LUZON: Camarines Sur, Buhi Lake, Hustedt (1.c.). lyra Ehr. Ehrenberg, p. 419, 1/1, f. 9a, 1841 (1843); Bailey, p. 431, 1853; Schmidt, et al., 2/16, 129/11-14, 1874; Cleve, p. 63, 1895; Mann, p. 108, 1925; Podzorski & Hakansson, p. 80, 31/10, 1987. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). BMNH, Adam’s-Bess #715, F 121, 2/7, M 58/ 15.16, TS 497.2/2. LUZON: Manila, BMNH, Comber #31904; Palawan, Taytay, Langen, Malapakan Is. (algae encrusted in Malapakan Cove, marine), Podzorski & Hakansson (1.c.). = Navicula gregoryana Grev. Greville, p. 10, 3/7, 1857. LUZON: Manila, (on sponge), BMNH, Comber #33424. VISAYAS: Cebu, BMNH, Comber # 31894. _______ var. cuneata Temp. & Perag. Tempere & Peragallo, p. 473, No. 988, 1914. VISAYAS: Cebu, BMNH, Comber # 31894, 31898. _______ var. elliptica Schmidt Schmidt in Schmidt, et al., 2/29, 2/34, 1874; Cleve, p. 64, 1895; Skvortzow, p. 272, 3/9, 1931; Mann, p. 108, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.); marine, Cleve (l.c.). BMNH, Adam’s #F 60.1/5. VISAYAS: Cebu, BMNH, Comber #31898. = Navicula lyra var. subelliptica Cl. Cleve, p. 64, 1895. Marine, Deby’s collection, Cleve (l.c.). _______ var. elliptica f. bullata (Norman) Cl. Cleve, p. 63, 1895; Schmidt, et al., 3/8-9, 1874. = N. bullata Norman Norman, p. 8, 2/7, 1861; Mann, p. 97, 1925. 14 Martinez-Goss Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). BMNH, Adam’s # F 40, 2/12.17. VISAYAS: Cebu, BMNH, Comber # 31894, 31897, 31898 (2), 31900. _______ var. granulata Perag. & Perag. Peragallo & Peragallo, 23/5, 1897; Schmidt, et al., 258/1,1874___. BMNH, Adam’s # F 45 1/10. _______ var. indica, sensu Adam BMNH, Adam’s # W 405. _______ var. insignis Schmidt Schmidt in Schmidt, et al., 2/27, 257/4, 1874; Mann, p. 108, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). = Navicula lyra var.? intermedia Perag. & Perag. Peragallo & Peragallo, p. 136 (141), 23/6 (11), 24/18, 1897. BMNH, Adam’s # F 20.2/8. _______ var. recta Grev. Greville, p. 28, 4/3, 1859; Mann, p. 108, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). BMNH, Adam’s # TS 497-2/5. = N. lyra var. recta f. typica Perag. Peragallo, 22/8, 1891. BMNH, Adam’s # TS 522 1/4.5. = N. lyra var. recta f. subelliptica Perag. & Perag. Peragallo & Peragallo, 22/7 (1897). BMNH, Adam’s # F 40 2/15.45.1/9.60 1/8. TS 497 2/10. _______ var. seductilis (Schmidt) O’Meara = Navicula seductilis Schmidt Schmidt in Schmidt, et al., 2/35, 1874; Mann, p. 119, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). _______ var. signata Schmidt Schmidt in Schmidt, et al., 2/4, 1874; Castracane, p. 33, 30/13, 1886. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). Frustules found in the vicinity of the Philippines, Castracane (l.c.). VISAYAS: Cebu, BMNH, Comber # 31893, 31895. _______ var. subtropica Schmidt Schmidt in Schmidt, et al., 2/24.25, 1874; Wolle, 16/14, 1890. BMNH, Adam’s # F 40 2/15.45.1/9.60 1/8. TS 497 2/10. _______ var. ? BMNH, Adam’s # F 20.2/7.45/12-14.119.3/8. G211; TS 262.1/6-9.6/3.522 1/2. maculata var. caribaea (Cl.) Cl. Cleve, p. 46, 1895. = Navicula caribaea Cl. Cleve in Schmidt, et al., 6/10-12, 1874; Mann, p. 97, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). maculosa Donk. Donkin, p. 25, 5/1, 1871; Schmidt, et al., 259/25, 1874. A checklist of Navicula 15 VISAYAS: Cebu, BMNH, Comber # 31899. = Navicula diffusa Schmidt Schmidt (1874) in Schmidt, et al., 2/28, 1874; Mann, p. 101, 1925. = Navicula madagascarensis Cl. Cleve, p. 23, 4/2, 1891; Mann, p. 108, 25/4, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). malombensis Otto Müller* Otto Müller, p. 84, 1/13, 1910. = Navicula mammalis Cstr. Castracane, p. 30, 20/2, 1886. Bottom marine samples taken near Philippine waters obtained by H.M.S. Challenger, Castracane (1.c.). mammalis Cstr. Castracane, p. 30, 20/2, 1886. Neighborhood of the Philippines, Castracane (l.c.). manokwariensis Choln. Cholnoky, p. 181, f. 71, 1963. = Navicula margarita Schmidt Schmidt (1892) in Schmidt, et al., 174/17, 1874; Mann, p. 108, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). margaritifera Truan y Luard et Witt Truan y Luard et Witt, p. 17, 4/10, 1888. = Navicula marginata Lewis Lewis, p. 64, 2/1, 1861; Mann, p. 108, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). mauriciana V.H. Van Heurck, p. 11, 2/182, 1909. = N. maxima Greg. Gregory, p. 41, 4/19, 1855; Mann, pp. 108-109, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). mendica Mann Mann, p. 109, 23/3, 1925. Bottom marine sample taken near the islands by the USS Albatross. Type-Cat. No. 43650, USNM, Mann (1.c.). menisculus Schum. Schumann, p. 56, 2/33, 1867; Skvortzow, p. 292, 2/8, 1937; Hustedt, p. 70, 1942a. LUZON: Rizal, Quezon City, Balara, Skvortzow (l.c.); Manila (ponds, Bureau of Science grounds); Batangas, Taal (lake); Camarines Sur, Buhi (lake); Hustedt (1.c.). minicans Hanna Hanna, p. 200, 13/4, 1932. = Navicula mimula Mann Mann, p. 110, 23/4, 1925. Bottom sediments taken near the islands by the USS Albatross, Mann (1.c.). minima Grun. Grunow in Van Heurck, p. 107, 14/15, 1880-1885. *Not recognized by VanLandingham, 1975. 16 Martinez-Goss = Navicula minima var. atomoides (Grun.) Cl. Cleve, p. 128, 1894; Hustedt, p. 56, 1942a. LUZON: Rizal, Manila (ponds, Bureau of Science grounds); Camarines Sur, Buhi Lake, Hustedt (l.c.). MINDANAO: Lanao (lake), Hustedt (1.c.). minuscula Grun. Grunow in Van Heurck, 14/3, 1880; Skvortzow, p. 291, 1/41, 1937; Hustedt, p. 59, 1942a. LUZON: Rizal, Quezon City (Balara Water Filters), Skvortzow (1.c.). MINDANAO: Lanao (lake, ponds), Hustedt (1.c.). mira Pant. & Greguss Greguss in Greguss, p. 212, 6/8, 1913; = Navicula mirabilis Leud.-Fortm. Leuduger Fortmorel, p. 31, 2/21, 1879; Mann, p. 110, 1925. Bottom marine sediments taken near the islands by the USS Albatross, rare, Mann (1.c.). modica Hust. Hustedt, p. 916, 41/21-23, 1945. = Navicula molesta Mann Mann, p. 110-111, 23/5, 1925. BMNH, Adam’s # F 119, 7/3. MINDANAO: Jolo, Sulu. Type-Cat. No. 43651, USNM, Mann (1.c.). monilifera f. constricta (Perag. & Perag.) Hust. Hustedt, p. 712, f. 1699B, 1966; Podzorski & Hakansson, pp. 84-85, 34/1, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (surface scrapings from the sand in the cove, at low tide); Podzorski & Hakansson (1.c.). mucicoloides Hust. Hustedt, p. 59, f. 98-102, 1942a. LUZON: Benguet, Bontoc; Rizal, Lyon (ponds), Hustedt (1.c.). muralis Grun. Grunow in Van Heurck, 14/27, 1880; Hustedt, p. 59, 1942a. LUZON: Camarines Sur, Buhi Lake, Hustedt (1.c.). mutica Ktz. Kützing, p. 93, 3/32, 1844; Woltereck, p. 52, 1941; Hustedt, p. 52, 1942a; Podzorski & Hakansson, p. 85, 33/8, 1987. LUZON: Benguet, Banaue (ponds), Woltereck (1.c.); Laguna, Los Baños (ponds, Laguna de Bay); Batangas, Taal Lake, Crater Lake; Camarines Sur, Buhi Lake, Hustedt (1.c.); Palawan, Taytay, Langen, Malapakan Is. (algae in the cove, marine); Taytay, Lake Manguao (Danao), (alt. c. 45 m, at the end of dry season, hence, lake at its shallowest); Roxas, Pagdangan Range, Ibangley Brookside Hill (alt. c. 40 m, on epiphyllous bryophyte beside a river), Podzorski & Hakansson (1.c.). VISAYAS: Leyte, Danao Lake, Hustedt (1.c.). MINDANAO: Lanao, Uyaan Lake, Hustedt (1.c.). _______ var. cohnii (Hilse) Grun. Grunow in Van Heurck, 10/17, 1880; Hustedt, p. 52-53, 1942a. LUZON: Rizal, Lyon; Laguna, Los Baños (ponds), Hustedt (1.c.). MINDANAO: Leyte, Uyaan Lake, Hustedt (1.c.). A checklist of Navicula 17 _______f. intermedia (Hust.) Hust. Hustedt, p. 586, f. 1593, 1966; Podzorski & Hakansson, p. 85, 33/9, 1987. LUZON: Palawan, Roxas, Pagdangan Range, (Ibangley Brookside Hill, from leaf surfaces), Podzorski & Hakansson (1.c.). = Navicula lagerheimii var. intermedia Hust. Hustedt in Schmidt, et al., 370/22, 1874; Hustedt, p. 54, 1942a. LUZON: Oriental Mindoro, Calapan Lake, Hustedt (1.c.). _______ var. nivalis (Ehr.) Hust. Hustedt, p. 290, 1911. = Navicula mutica var. tropica Hust. Hustedt, p. 233, 17/6, 1937b. LUZON: Laguna, Los Baños (ponds); Camarines Sur, Buhi Lake, Buhi River; Oriental Mindoro, Naujan (river), Hustedt (1.c.). nebulosa Greg. Gregory, p. 480, 9/8, 1857; Mann, p. 111, 1925. Bottom sediments taken near the islands by the USS Albatross, Mann (l.c.). VISAYAS: Cebu, BMNH, Comber #31900. nippon Skv. Skvortzow, p. 277, 5/17, 1936. = Navicula nitescens (Greg.) Ralfs* Ralfs in Pritchard, p. 898, 1861; Mann, p. 111, 1925. Bottom sediments taken near the islands by the USS Albatross, Mann (1.c.). northumbricaeformis M. Perag M. Pergallo in Heribaud, et al., p. 15, 1/19, 1920; = Navicula notabilis Grev. Greville, p. 18, 1/19, 1863; Mann, p. 111, 1925. Bottom sediments taken near the islands by the USS Albatross, Mann, (l.c.). nummularia Grev. Greville, p. 249, 5/6, 1859; De Toni, p. 96, 1891; Mann, p. 111, 1925. Bottom sediments taken near the islands by the USS Albatross, Mann (1.c.). = Navicula forcipata var. suborbicularis Grun. Grunow in Van Heurck, 10/5, 1880. Cleve, p. 66, 1895. MINDANAO:Sulu Sea(Deby Coll.). aff. ny Cl. Cleve, p. 75, 1/24, 1894; Podzorski & Hakansson, p. 75-76, 31/4, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (surface scrapings from sand at low tide in the cove, marine), Podzorski & Hakansson (l.c.). nympharum Hust. Hustedt, 1936, in Schmidt, et al., 401/85-87, 1874. = Navicula oamaruensis Grun.* Grunow, 1888, in Schmidt, et al., 129/9, 1874- __; Mann, p. 111, 1925. Bottom sediments taken near the islands by the USS Albatross, Mann (l.c.). obesa O. Müll. O. Müller, p. 84, 2/27, 1910. *Not recognized by VanLandingham, 1975. 18 Martinez-Goss = Navicula obesa (Grev.) Mann Mann, pp. 111-112, 23/6, 24/1, 1925. Bottom marine samples taken near the islands by the USS Albatross. No type designated, Mann (1.c.). occulta Krasske Krasske, p. 353, f. 12a-b, 1929. = Navicula ocellata Mann* Mann, p. 112, 24/2, 1925. Bottom marine samples taken near the islands by the USS Albatross. Type-Cat. No. 43652, USNM, Mann (1.c.). oscitans Schmidt Schmidt, 1875, in Schmidt, et al., 6/41, 1874; Mann, p. 112, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). ______ var. subundulata Cl. & Grove Grove, p. 67, 10/10, 1891. BMNH, Adam’s # TS 523/5.8. = Navicula suboscitans Mann Mann, p. 122, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). pacifica (Cstr.) Mann* Castracane p. 23, pl. 20, f. a, 1886; Mann, p. 113, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). palpebralis var. semiplena (Greg.) Cl. Cleve, p. 70, 1895. = Navicula angulosa var. beta Greg. Gregory, p. 42,5/8,1856; Mann, p. 94, 1925. Bottom sediments, Mann (l.c.). palustris var. obesa Zanon Zanon, p. 111, 2/10, 1949. =Navicula pandura Breb. Schmidt, et al., 11/1, 2, 4, 8, 9, 1874; Mann, p. 113, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). perrotettii (Grun.) Cl. Cleve, p. 110, 3/12, 1894; Hustedt, p. 51, 1942a. Diatom samples from the Philippines of slightly brackish waters derived from Dr. Rae’s collection, Cleve (1.c.). LUZON: Rizal, Manila (ponds, Bureau of Science grounds); Laguna, Los Baños (Laguna de Bay, ponds); Camarines Sur, Buhi (lake, river), Hustedt (l.c.). philippina Skv. Skvortzow, p. 291, 1/25, 45, 1937a. LUZON: Rizal, Quezon City, Balara (water filter no. 8), Skvortzow (l.c.). phillippinarum Mann Mann, pp. 114-115, 25/1, 1925. Bottom marine samples taken near the islands, including those from the Sulu group, taken by the USS Albatross. Type-Cat. No. 43655, USNM, Mann (1.c.). BMNH, Adam’s # F 83, 6/57; TS 523/9. platyventris Meist. Meister, p. 95, f. 33, 1934 (1935); Podzorski & Hakansson, p. 81, 32/3, 1987.*Not recognized by VanLandingham, 1975. A checklist of Navicula 19 LUZON: Palawan, Langen, Malapakan Is. (surface scrapings from the sand at low tide in the cove), Podzorski & Hakansson (1.c.). plicatula Grun. Cleve & Möller, slide #154-155; Grunow Ms. in Cleve, p. 155, 3/28, 1894; Mann, pp. 115-116, 24/8-9, 1925; Podzorski & Hakansson, p. 78, 30/8, 1987. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae from the cove, marine), Podzorski & Hakansson (1.c.). VISAYAS: Cebu, BMNH, Comber # 31894. plutonia O’ Meara* O’Meara, p. 286, 13/6, 1872; De Toni, p.78, 1891. MINDANAO: Sulu, Cagayan, (brackishwaters from the volcanic mountain), De Toni (l. c.). praetexta Ehr. Ehrenberg, p. 214, 1840 (1841); Castracane, p.812, 1886; Mann, p. 116, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). BMNH, Adam’s # GC 2094; F 119 3/9; TS 497.1/10; TS 523/11; TS 60/81. LUZON: Manila (on shell), BMNH, Comber # 30925. VISAYAS: Cebu, Castracane (l.c.). _______ var. abundans Schmidt Schmidt, et al., 129/8, 1874; Cleve, p. 55, 1895. VISAYAS: Cebu, BMNH, Comber # 31898, 31895, 31894. _______ var. ? BMNH, Adam’s # F 45.2/16. protracta (Grun.) Cl. Cleve, p. 140, 1894; Podzorski & Hakansson, p. 77, 33/13, 1987. LUZON: Palawan, Taytay (Lake Manguao, alt. c. 45 m., end of the dry season when it was at its shallowest, surrounded by semi-deciduous rainforest), Podzorski & Hakansson (1.c.). pseudobryophila Hust. Hustedt, p. 60, 1942a; _______, p. 114, 1942b. VISAYAS: Leyte (Danao Lake), Hustedt (1942a). pseudoclavata Mann Mann, p. 116, 25/2, 1925. Bottom marine samples taken near the islands by the USS Albatross. Type-Cat. No. 43656, USNM, Mann (1.c.). pseudoforcipata Hust. Hustedt, pp. 119-120 f. 11, 1942b; Podzorski & Hakansson, p. 80, 31/67, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (from the cove, marine); Pto. Princesa (south of Pto. Princesa Bay, from the rapidly deepening zone of the Sulu Sea, hand collected from corals at 10 m depth), Podzorski & Hakansson (1.c.). pseudony Hust. Hustedt, p. 23, 8/11, 1955; Podzorski & Hakansson, p. 80, 30/2, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae encrusted from the cove, marine), Podzorski & Hakansson (1.c.). *Not recognized by VanLandingham, 1975. 20 Martinez-Goss pugio Mann Mann, p. 117, 26/4, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). pulverulenta (pulvulenta) Mann Mann, pp. 117-118, 25/3, 1925. Bottom marine samples taken near the islands by the USS Albatross. Type-Cat. No. 43659, USNM, Mann (1.c.). pupula Ktz. Kützing, p. 93, 30/40, 1844; Woltereck, p. 52, 1941; Hustedt, p. 64, 1942a; Podzorski & Hakansson, p. 76, 30/3, 1987. LUZON: Benguet, Banaue (ponds), Woltereck, 1941; Rizal, Malabon, Dagatan (swamp), Manila (ponds in the Bureau of Science grounds, Pancipit River, Reservoir II); Laguna, Los Baños (ponds, Laguna de Bay), San Pablo (Bunot Lake, Bato Lake); Batangas, Taal (Yambo Crater Lake, Crater Lake); Camarines Sur, Buhi (lake, river); Payren Lake; Oriental Mindoro, Naujan Lake, Hustedt (1.c); Palawan, Taytay (Lake Manguao, alt. 45 m, near Public Works’ resthouse, at the end of the dry season, when lake was shallowest; small pool in stream bed flowing into NW side of the lake, 1.8 km from Public Work’s resthouse, alt. 130 m), Podzorski & Hakansson (1.c.). MINDANAO: Lanao (Uyaan Lake, Lanao Lake, river); Jolo, Sulu (Seit Lake), Hustedt (1.c.). _______ var. rectangularis (Greg.) Cl. & Grun. Cleve & Grunow, p. 45, 1880. = Navicula f. rectangularis (Greg.) Cl. & Grun. Hustedt, pp. 64-65, 1942a; Podzorski & Hakansson, p. 76, 30/4, 1987. LUZON: Palawan, Roxas, Pagdanan Range (Ibangley Brookside Hill, alt. 40 m, streamside pool), Podzorski & Hakansson (1.c.). VISAYAS: Leyte (Danao Lake), Hustedt (1.c.). MINDANAO: Lanao (Uyaan Lake), Hustedt (1.c.). _______ f. rostrata (Hust.) Hust. Hustedt, p. 291, 3/39, 1911; _______, p. 64, 1942a; _______, p. 282, 1957. LUZON: Laguna, Los Baños (Laguna de Bay, pond); Batangas, Taal (Taal Lake); Camarines Sur, Buhi (lake), Hustedt (1942a). MINDANAO: Lanao (pond), Hustedt (1942a). pygmaea Ktz. Kützing, p. 77, 1849; Woltereck, p. 52, 1941; Hustedt, pp. 66-67, 1942a; Podzorski & Hakansson, p. 80, 31/11, 1987. LUZON: Benguet, Banaue (ponds), Woltereck (1.c.); Rizal, Manila (Bureau of Science grounds, ponds), Pancipit River, Lyon (ponds); Laguna, Los Baños (ponds, Laguna de Bay); Camarines Sur, Buhi (river), Hustedt (1.c.); Palawan, Pto. Princesa (south of Pto. Princesa Bay, hand collected from sand surface at 15 m depth); Taytay, Langen, Malapakan Is. (surface scrapings from sand at low tide), Podzorski & Hakansson (1.c.). MINDANAO: Lanao (Lanao Lake), Jolo, Sulu (Seit Lake), Hustedt (1.c.). quarnerensoides Hust.* Hustedt, p. 49, f. 1203, 1961; (doubtful-fide: VanLandingham, p.2759,1975); Podzorski & Hakansson, 78, 1987. LUZON: Palawan, Pto. Princesa, Table Head (hand collected from sand surface at 15 m depth of Sulu Sea), Podzorski & Hakansson (1.c.). *Not recognized by VanLandingham, 1975. A checklist of Navicula 21 radiosa Ktz. Kützing, p. 91, 4/23, 1844; Woltereck, p. 52, 1941; Hustedt, pp. 69-70, 1942a; Podzorski & Hakansson, p. 82, 33/2, 3. LUZON: Benguet, Banaue (ponds); Rizal, Malabon (Dagatan swamp), Manila (Water Reservoir III, Ponds in Bureau of Science grounds, Lyon ponds), Hustedt (1.c.); Brooke’s Point (Tumarabong River over Oligocene-Miocene reef limestones, alt. 10 m, scrapings from the surface of a limestone rock), Podzorski & Hakansson (1.c.). _______ var. tenella (Breb. ex Ktz.) Cl. et Möll. (non Grun. fide VanLandingham, 1975); Cleve et Möller slide no. 224, 1881; Hustedt, pp. 69-70, 1942a; Podzorski & Hakansson, p. 82, 33/4, 1987. LUZON: Batangas, Taal (Volcano Lake), Hustedt (1.c.); Palawan, Pto. Princesa (Irawan River over mafic and mafic-plutonic rocks, alt. c. 150 m; scrapings from a bryophyte behind a waterfall), Podzorski & Hakansson, (1.c.). raena (Cstr.) De T. De Toni, p. 30, 1891; Mann, p. 118, 1925. Bottom marine samples taken near the islands by the USS Albatross. Type species from the Philippines (fide: Castracane below), Mann (1.c.). = Pinnularia raena Cstr. Castracane, p. 25-26, 15/3, 1886. VISAYAS: Cebu, from Dr. Rae’s collection, Castracane (1.c.). ramosissima (Ag.) Cl. Cleve, p. 26, 1895; Podzorski & Hakansson, p. 82, 34/3, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae encrusted from the cove, marine), Podzorski & Hakansson (1.c.). retracta Meist. Meister, p. 262, 4/6, 1937; = Navicula retrostauros Mann* Mann, p. 118, 25/2, 26/1, 2, 1925. BMNH, Adam’s #M 59.1/8; TS 60/59. Bottom marine samples taken near the islands by the USS Albatross. Type-Cat. No. 43660 USNM, Mann (l.c.). rhaphoneis (Ehr.) Grun. Grunow, p. 19, 91/17, 1867; Podzorski & Hakansson, pp. 82-83, 32/6, 7, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae scraped from the cove), Podzorski & Hakansson (1.c.). rhombica Greg. Gregory, p. 40, 4/16, 1855; Mann, p. 119, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). _______var. japonica (Brun.) Cl. Cleve, p. 152, 1894. BMNH, Adam’s #TS 523/2. rhynchocephala (rhynococephala) Ktz. Kützing, p. 152, 30/35, 1844; Hustedt, p. 67, 72, 1942a; delos Reyes, p. 59, pl. 4, f. 17, 1972. LUZON: Laguna, Los Baños (Laguna de Bay); Batangas, Taal (Crater Lake); Hustedt (1.c.). VISAYAS: (Tigbao water supply), delos Reyes (1.c.). riparia Hust. Hustedt, p. 52, f. 77-78, 1942a. *Not recognized by VanLandingham, 1975. 22 Martinez-Goss LUZON: Rizal, Malabon (Dagatan Swamp), Hustedt (1.c.). robertsiana (robertsoniana) Grev. Greville, p. 235, 3/9, 1865. VISAYAS: Cebu, BMNH, Comber #31895. _______ var. cuneata (Schmidt) Amosse’ Amosse’, p. 111, 1924. = N. prodiga Mann Mann, p. 252, 53/4, 1907; ______, p. 116, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1925). ruttneri Hust. Hustedt, 1936, in Schmidt, et al., 402/30-38, 49-52, 1874; _______, p. 55, 1942a. LUZON: Camarines Sur, Buhi (lake), Hustedt (1942a). _______ var. capitata Hust. Hustedt, 1936, in Schmidt, et al., 402/47, 48, 1874; _______, p. 55, 1942a. LUZON: Manila (Reservoir II), Oriental Mindoro, Naujan (river), Hustedt (l.c.). _______ var. rostrata Hust. Hustedt, 1936, in Schmidt, et al., 402/39-45, 1874; ______, p. 56, 1942a. LUZON: Rizal, Pancipit River; Oriental Mindoro, Naujan (Naujan Lake), Hustedt (1942a). MINDANAO: Lanao (pond), Hustedt (1942a). salinarum Grun. Grunow in Cleve & Möller, slide no. 107, 1878; Grunow in Cleve & Grunow, p. 33, 2/34, 1880; Pantastico, p. 195, 17/3, 1977. LUZON: Laguna, Los Baños, Mayondon, Pantastico (1.c.). schaarschmidtii Pantocsek Pantocsek, p. 28, 14/121, 1886. VISAYAS: Cebu, BMNH, Comber #31900. schoenfeldii Hust. Hustedt, p. 301, f. 520, 1930; _______, p. 70, 1942a. LUZON: Oriental Mindoro, Naujan (river), Hustedt (1942a). schroeteri Meist. Meister, p. 38, f. 100, 1932; Hustedt, p. 72, 1942a. LUZON: Laguna, Los Baños (Laguna de Bay); Batangas, Taal (Taal Lake); Oriental Mindoro, Naujan (river), Hustedt (1.c.). _______ var. escambica Patr. Patrick, p. 104, 7/3, 1959; Podzorski & Hakansson, p.83, 33/5, 1987. LUZON: Palawan, Roxas, Pagdanan Range, (Ibangley Brookside Hill streamside freshwater pool, alt. c. 40 m.), Podzorski & Hakansson (l.c.). scutelloides W. Sm. Smith, p.9, 1856; _____, in Gregory, p. 4, 1/15, 1856; Hustedt, p. 66, 1942a. LUZON: Batangas, Taal (Crater Lake); Camarines Sur, Buhi (lake), Hustedt. (l.c.) MINDANAO: Lanao (Lanao Lake, pond), Hustedt (1.c.). seminulum Grun.Grunow, p. 552, 4/3(2/3), 1860; Hustedt, p. 56, 1942a. LUZON: Camarines Sur, Buhi (lake); Batangas, Taal (Crater Lake). A checklist of Navicula 23 MINDANAO: Lanao (Lanao Lake, pond); Hustedt (l.c.). semistauros Mann* Mann, 119-120, 26/5, 1925. Bottom marine samples taken near the islands by the USS Albatross, Type-Cat. No. 43661, USNM, Mann (l.c.). sigma Brun Brun, p. 39, 16/5, 1891; Brun later homonym, Fide VanLandingham p. 2799, 1975; Podzorski & Hakansson, p. 75, 29/12, 1987. LUZON: Palawan, Pto. Princesa, Table Head (from sand surface at 15 m depth of Sulu Sea), Podzorski & Hakansson (1.c.). spectabilis Greg. Gregory, p. 481, 9/10, 1857; Cleve, p. 60, 1895; Mann, p. 121, 27/1-2, 1925; Podzorski & Hakansson, pp. 80-81, 32/1, 1987. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). Diatom samples (presumably from Dr. Rae’s collection), Cleve, 1894, p. 60, 1895. BMNH, Adam’s # J 4006; Bess 1709/3; G 211; F 40.2/16.45.1/11. LUZON: Palawan, Pto. Princesa, Table Head (from the sand surface at 15 m depth of Sulu Deep Sea), Podzorski & Hakansson (1.c.). VISAYAS: Cebu, BMNH, Comber #318931. _______ var. controversa (Schmidt) Cl. Schmidt, et al., 3/5, 1874; Cleve, p. 60, 1895. BMNH, Adam’s # TS 60/4. _______ var. emarginata Cl. Cleve p. 60, 1895. = Navicula excavata var. ________ Schmidt Schmidt in Schmidt, et al., 3/22-25, 1874. VISAYAS: Cebu, BMNH, Comber #31899. _______ var. excavata (Grev.) Cl. Cleve, p. 61, 1895; = N. excavata Grev. Greville, p. 130, 12/15, 1866; Mann, p. 102, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). BMNH, Adam’s # F 83.1/9.119.2/4; TS 522.2/2. = N. oswaldii Janisch Janisch, 1, 15/12, 1888; Mann, p. 113, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). _______ var. minuta Cl. Mereschkowsky, p. 320, 2/9-12, 1902. = Navicula hennedyi var. minuta Cl. Cleve, p. 7, 1/15, 1881. VISAYAS: Cebu, BMNH, Comber #31894. _______ var. ? BMNH, Adam’s # F 931; Bess 1710/3; TS 497. 2/7-9. spiculifera Mann* Mann, pp. 121-122, 26/8, 1925. *Not recognized by VanLandingham, 1975. 24 Martinez-Goss Bottom marine samples taken near the islands by the USS Albatross. Type-Cat. No. 4663, USNM, Mann (1.c.). spiralis O’ Meara* De Toni, p. 74, 1891. MINDANAO: Saline waters from the volcanic crater, Cagayan, Sulu Archipelago, De Toni (l.c.). studerri sensu Comber* LUZON: Manila (sponge), BMNH, Comber # 30924. subarvensis Hust. Hustedt, p. 60, f. 107-108, 1942a. LUZON: Camarines Sur, Buhi (Buhi Lake), Hustedt (1.c.). = N. lyra var. subcarinata Grun. Grunow, 1874, in Schmidt, et al., 2/5, 1874; Mann, p. 108, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). BMNH, Bess 62; F 40 2/19; TS 497.2/6, 522, 2/6, 3/8. LUZON: Manila, BMNH, Comber #31903. subdecipiens De T. De Toni, p. 147, 1891. = N. decipiens Cstr. Castracane, p. 32, 27/17, 1886. From Philippine marine diatom collection of Dr. Rae, Castracane (l.c.). subdecussis Hust. Hustedt, pp. 75-76, f. 136-138, 1942a. LUZON: Rizal, Manila (Reservoir II), Hustedt (1942a). subforcipata Hust. Hustedt, p. 533, f. 1569, 1964; Podzorski & Hakansson, p. 75, 31/51, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (surface scrapings from sand at low tide), Podzorski & Hakansson (1.c.). subrhynchocephala Hust. Hustedt, p. 156, 1/11, 1935; _______, p. 67, 1942a. LUZON: Rizal, Manila (Reservoir II, Pancipit River); Laguna, Los Baños (pond, Laguna de Bay, Payren Lake); Batangas, Taal (Crater Lake); Camarines Sur, Buhi (Buhi Lake, river), Hustedt (1942a). VISAYAS: Leyte (Danao Lake), Hustedt (1942a.). suluensis O’ Meara* De Toni, p. 80, 1891. MINDANAO: Brackishwaters in the crater of a volcanic island, Cagayan, Sulu, De Toni (l.c.). takoradiensis Hendey Hendey, p. 67, 1/8, 1958; Podzorski & Hakansson, p. 77, 30/7, 1987. LUZON: Palawan, Pto. Princesa, Table Head (from sand surface at 15 m depth of the Sulu Sea), Podzorski & Hakansson (1.c.). tenera Hust. Hustedt, p. 259, 18/11-12, 1937a; _______, p. 67, 1942a. LUZON: Rizal, Manila (Pancipit River); Batangas, Taal (Crater Lake); Hustedt (1942a). terminata Hust. Hustedt, p. 589, f. 1594, 1966; *Not recognized by VanLandingham, 1975. A checklist of Navicula 25 = N. mutica var. tropica Hust. Hustedt, 1936 in Schmidt, et al., 405/39-42, 1874; _______, pp. 53-54, 1942a. LUZON: Laguna, Los Baños (pond); Camarines Sur (Buhi Lake); Oriental Mindoro, Naujan (Naujan River), Hustedt (1942a). translucens Mann Mann, p. 123, 26/9, 1925. Bottom marine samples taken near the islands by the USS Albatross. Type-Cat. No. 43665, USNM, Mann (1.c.). tripunctata (O.F. Müll.) Bory* Bory, p. 128, 1822; Podzorski & Hakansson, p. 83, 33/6, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (scrapings from sand surface at low tide), Pto. Princesa, Honda Bay (scraping from coral reefs at the edge of an atoll), Podzorski & Hakansson (1.c.). uniseriata Östrup Östrup, p. 8, 1/9, 1913; Podzorski & Hakansson, pp. 77, 33/12, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae encrusted from cove), Podzorski & Hakansson (1.c.). variostriata Krasske Krasske, p. 197, f. 12, 1923; Hustedt, p. 55, 1942a. LUZON: Laguna (Laguna de Bay), Hustedt (1.c.). venusta Janisch Janisch, 15/17, 1888; Mann, p. 124, 1925. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). venustissima Kitt. Kitton, p. 101, 1889; Kitton in Leuduger-Fortmorel., p. 17, 2/3, 1892. BMNH, Adam’s # GC 2177. viminea Hust. Hustedt, 1934 in Schmidt, et al., 397/10-11, 1874; Podzorski & Hakansson, p. 83, 32/8-9, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (surface scrapings from sand at low tide), Podzorski & Hakansson (1.c.). viridula (Ktz.) Ehr. Ehrenberg, p. 53, 1836. = Navicula viridula Ktz. Kützing, p. 23, 13/12, 1833; Hustedt, p. 72, 1942a. LUZON: Laguna, Los Baños (pond); Camarines Sur, Buhi (Buhi Lake), Hustedt (1.c.). werestschangini (werestschangini) Skv. & Meyer Skvortzow & Meyer, p. 20, 1/64, 1928; Podzorski & Hakansson, p. 85, 34/5, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Is. (surface scrapings from sand at low tide), Podzorski & Hakansson (1.c.). yarrensis (jarrensis) Grun. Grunow 1876, in Schmidt, et al., 46/1-6, 1874; Mann, p. 126, 1925; Podzorski & Hakansson, p. 79, 31/3, 1987. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). BMNH, Adam’s # GC 2190; TS 262.1/ 4.5.18.6/1.2.89. *Not recognized by VanLandingham, 1975. 26 Martinez-Goss LUZON: Palawan, Taytay, Langen, Malapakan Is. (algae scraped from the cove), Podzorski & Hakansson (1.c.). zanzibarica Grev. var. zebuana Cstr. Greville, p. 129, 12/22, 1866; Castracane, p. 31, 28/8, 1886; A. Schmidt, 1888 in A. Schmidt, et al., 129/4, 1874. VISAYAS: Cebu, Castracane (1.c.). zostereti (zosteretii) Grun. Grunow, p. 528, 4/23, (2/23) 1860; Mann, p. 125, 1925; Podzorski & Hakansson, pp. 83-84, 33/10, 1987. Bottom marine samples taken near the islands by the USS Albatross, Mann (1.c.). LUZON: Palawan, Taytay, Langen, Malapakan Cove (scrapings from the cove), Pto. Princesa, White Beach (on mud surface in a mangrove), Honda Bay (coral scrapings from a bay), Podzorski & Hakansson (1.c.). sp. (miniscula) Podzorski & Hakansson, p. 81, 19/11, 1987. LUZON: Palawan, Taytay, Langen, Malapakan Cove (scraping from sand surface at low tide), Podzorski & Hakansson (1.c.). sp. 1 Esguerra, p. 174, 1951; Villadolid, p. 21, 1957. sp. 2 BMNH, Adam’s-Bess # M64-66; TS 80; Bess 22/ 6.714; TS 522.2/3.7.3/3. VISAYAS: Cebu, Castracane, p. 812, 1886. sp. 3 White chalk collected by the ship Princess Louise to Manila, under Capt. Wendt (1830-34). Identification by Dr. Meyer. Ehrenberg, Atlas, p. 177, pl. 1, 1854. ACKNOWLEDGMENTS This checklist was completed thru the financial assistance of the Jessup-McHenry Fund Grant of the Academy of Natural Sciences of Philadelphia (ANSP), U.S.A. and the British Council of England; the kind hospitality in making available their references and specimens of the ANSP and the British Museum of National History; the patience in encoding and revising the paper by C.M. Nuñez, M.V. Marasigan and W.F. Fabelllore. REFERENCES Amosse, A., 1924. Diatomees de la cote orientale d’ Afrique. 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